Carabidae image gallery
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Acathaicus taxon page (17 photos)
Carabus (Acathaicus) alexandrae alexandrae Semenov, 1887 Holotype Holotype
Male adult
Kwantin and Li-tschara-pu, 22.VI.1885, leg. Potanin<br /> Deposited in Zoolog. Museum St. Petersburg.<br /> male, long. 22 mm<br /> photo B. Brezina
Copyright © Boleslav Brezina
Carabus (Acathaicus) alexandrae cratocephaloides Semenov, 1887
Female adult
female, 28mm
Copyright © Boleslav Brezina
Carabus (Acathaicus) interruptocostatus Semenov, 1887 Holotype Holotype
Female adult
Copyright © Boleslav Brezina -
Acoptolabrus taxon page (63 photos)
Carabus (Acoptolabrus) lopatini Morawitz, 1886
Male+Female adult
Russia,
Sakhalin isl., Tchekhov Mt, VI.1997
female, 40mm
photo & coll. Panin R.
Copyright © Ruslan Panin
Carabus (Acoptolabrus) schrencki schrencki (as hauryi Gehin, 1885 )
Male adult
Copyright © Boleslav Brezina
Carabus (Acoptolabrus) schrencki kimieae Kezuka & Imura, 1989
Male+Female adult
Mt. Wulongshan, Dandong City,
Liaoning Province, CHINE
25-30.VI.2008
Copyright © Safronov
Carabus (Coptolabrodes) haeckeli Brezina & Imura, 1997
Male adult
C China, Shaanxi,
Tsinling Mts., Taibaishan Mt., h=3000m, 33:55N/107:44 E, VI.2004, V.Siniaev & his team
female, 31 mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Acoptolabrus) schrencki schrenck (as dongningensis Deuve & Li, 2005)
Male adult
NE China, Heilongjiang prov., Dongning, V.2009<br /> L=25 mm
Copyright © Ruslan Panin
Carabus (Acoptolabrus) constricticollis constricticollis (as grallatorius Roeschke, 1921)
Male adult
NE-Korea:North Hamgyong Prov., Sechon-dong, VII.2004, Li leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Acoptolabrus) gehinii gehinii Fairmaire, 1886
Female adult
N-Japan, SW-Hokkaido, Chitose env.,VI.2011.
L=34 mm
Copyright © Ruslan Panin
Carabus (Acoptolabrus) constricticollis pihyonensis Deuve & Li, 1998
Male adult
N-Korea, North Pyongan Prov., Tongnim County, 5-25.V.2010 <br /> 29 mm
Copyright © Ruslan Panin
Carabus (Acoptolabrus) ? leechi huisaensis Deuve & Li, 1999
Male adult
NC-Korea, Ryanggang Prov., Puksubaek-san, 4-10.V.2010
28 mm
Copyright © Ruslan Panin
Carabus (Acoptolabrus) gehinii munakatai (Ishikawa, 1968)
Male adult
Japan, SW-Hokkaido, Yakumo town, 4.VI.2017, S.Shinichi leg., L=30 mm
Copyright © Ruslan Panin -
Acoptopterus taxon page (12 photos)
Carabus (Carabus) gressittianus fuscomarginatus Kleinfeld, 1997 Holotype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Carabus) andreiianus Deuve, 2007
Female adult
C-China, N-Sichuan, Pingwu Co. h=1500 m, 27.VI.2011, L=24 mm
Copyright © Ruslan Panin
Carabus (Carabus) koidei koidei Imura, 1996
Male adult
C-China, S-Sichuan, Pass btw. Yanyaun / Muli, h=3244 m, 27.68638N; 101.22335E,,11-18.VI.2017, Ch. Reuter leg. L=22 mm
Copyright © Ruslan Panin
Carabus (s.str.) morphocaraboides zhaojueanus Imura, 1999
Female adult
China, S-Sichuan, SE Mianning, Pass btw Mianshanzhen & Yuexi 28°30’N/102°26’E, 22-23.VI.2017, Ch.Reuter leg., L=22 mm; det. B. Brezina, 2020
Copyright © Ruslan Panin
Carabus (s.str.) morphocaraboides zhaojueanus Imura, 1999
Male adult
China, S-Sichuan, road 56 km E of Xichang to Zhaojue, Mt.WenLin, Pass h=3200 m, 27°52’N/102°30E, 5-20.VI.2017, Ch.Reuter leg.; det B. Brezina, 2020
Copyright © Ruslan Panin
Carabus (Acoptopterus, ?Carabus) yunnanus yanyuanicus Cavazzuti, 1996
Endophallus
China, S-Sichuan, S Yanyuan, Chuandongzi Mt h=3300-3600 m
Copyright © Ruslan Panin
Carabus (Acoptopterus) yunnanus yanyuanicus Cavazzuti, 1996 (? or oblongior ssp.)
Male adult
China, S-Sichuan, S Yanyuan, Chuandongzi Mt h=3300-3600 m, 16-20.VI.2023, C.Reuter leg./det.Klienfeld, 2023, L=21.5 mm
Copyright © Ruslan Panin -
Acrocarabus taxon page (6 photos)
Carabus (Acrocarabus) callisthenoides Semenov, 1888
Male adult
male, 22mm
Copyright © Boleslav Brezina
Carabus (Acrocarabus) guerini aldarkose Kabak, 1992
Male adult
male, 22mm
Copyright © Boleslav Brezina
Carabus (Acrocarabus) guerini guerini Fischer, 1842
Male adult
SE Kazakhstan, W Dzhungar Mts, Lepsinsk env., 1600 m., 18.06.2000, A.Klimenko
Copyright © Anton Menstschikov -
Alipaster taxon page (18 photos)
Carabus (Alipaster) barovskii bayanbulak Deuve, 1993 Paratype Paratype
Female adult
female, 17mm
Copyright © Boleslav Brezina
Carabus (Alipaster) infantulus infantulus Morawitz, 1886
Male adult
male, 15mm
Copyright © Boleslav Brezina
Carabus (Cechenotribax) petri Semenov & Znojko, 1932
Female adult
female, 18mm
Copyright © Boleslav Brezina
Carabus (Eotribax) eous eous Morawitz, 1889
Male adult
Kirghyzstan, N Terskei Ala Too, Borskaun riv., h=2500 m., 7-14.07.2000
male
L=17 mm
Copyright © Maksim Smirnov
Carabus (Cechenotribax) petri Semenov & Znojko, 1932
Male adult
SE Kazakhstan, N-Tianshan:N-slopes Ketmen Mt.Rng. h=2250-2500m., near Ketmen vill.,1-5.VI.1990, Dolin V.G. leg.
L=13.5 mm
Copyright © Ruslan Panin
Carabus (Alipaster) pupulus pupulus Morawitz, 1889
Female adult
S Kyrgyzstan, Inner Tian Shan, Dzhany-Dzher Mt range, upper Mudyuryum riv. Ak-Baytal gorge h~3300 m, 15.VII.2011, S.Toporov leg. L=16 mm
Copyright © Ruslan Panin -
Apoplesius taxon page (5 photos)
Carabus (Apoplesius) deuvei hajdaji Hackel, 2007 Holotype Holotype
Male adult
Turkey- prov. Mus 60 km NW Mus 1630m
3856N 4108E Bulgan Gec. 23-24.06.03 E. & P. Hajdaj leg.
Copyright © Boleslav Brezina
Carabus (Apoplesius) pseudodepressus Machard, 1988
Female adult
Turkey, Prov. Bitlis, S near Tatvan, 23.IV.2009 leg. M. & S. Nabozhenko
L=19.0 mm
Copyright © Kirill Makarov -
Apotomopterus taxon page (205 photos)
Carabus (Apotomopterus) cantonensis pervarius Kleinfeld, 1997 Holotype
Male adult
Frank Kleinfeld
Copyright © - - -
Carabus (Apotomopterus) cantonensis metallaicus Kleinfeld, 1999 Holotype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) delavayi medea Kleinfeld, 1999 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) davidis davidis Deyrolle & Fairmaire, 1878
Male adult
SE-China, Jiangxi prov., Yushan County, Mt Wangjing, VI,2012
L=38 mm
Copyright © Ruslan Panin
Carabus (Apotomopterus) davidis helmutschuetzei Kleinfeld, 1995
Male adult
SE-China, SW-Jiangxi: S of Yongxin: Mt. Jinggangshan,V.2012, L=35 mm
Copyright © Ruslan Panin
Carabus (Apotomopterus) patroclus vicarius Cavazzuti, 1996
Male adult
China, S Sichuan, / Yunnan border, W of Lugu lake, S of Yongning vill., h=2700m, 11-18.VI.2017, L=31 mm
Copyright © Ruslan Panin
Carabus (Apotomopterus) protenes baoxingensis Deuve & Mourzine, 2003
Endophallus
China: С-Sichuan:Qionglai Shan Mts., Baoxing env. (TOPOTYPUS)
Copyright © Igor Solodovnikov
Carabus (Apotomopterus) protenes ascendens Semenov, 1898
Endophallus
China, Chongqing, N Jinfo Shan, Shanwangping h=1400m, 29.101944, 107.166389
Copyright © Ruslan Panin
Carabus (Apotomopterus) mecynodes pinchuanensis Imura, 1999
Female adult
China, S-Sichuan, 40-50 km E Xichang (to Zhaojue) 27°51'N 102°30'E, h=2700-3000m, Pine forest, 5-20.VI.2017, C.Reuter leg., L=33 mm
Copyright © Ruslan Panin
Carabus (Apotomopterus) mecynodes pinchuanensis Imura, 1999
Male adult
China, S-Sichuan, 40-50 km E Xichang (to Zhaojue) 27°51'N 102°30'E, h=2700-3000m, Pine forest, 5-20.VI.2017, C.Reuter leg., L= 29 mm
Copyright © Ruslan Panin -
Archicarabus taxon page (40 photos)
Carabus (Archicarabus) monticola fontanae Born, 1906
Male+Female adult
N-Italy, Piemonte, Bielmonte, h1350 m, 14-15.VI.2010, Pascal Stefani leg.; female, 23 mm
Copyright © Ruslan Panin
Carabus (Archicarabus) gotschi gotschi Chaudoir, 1846
Male+Female adult
Transcaucasia, Armenia,
Ashotsk distr., Bandivan, 8-21.X.1991 P.V.Kazaryan leg.
female, 20mm
Copyright © Ruslan Panin
Carabus (Archicarabus) wiedemanni wiedemanni Menetries, 1836
Male adult
SE-Bulgaria:Burgas env. Tsarevo (Michurin), 10-15.VII.1981, J.Slaba leg.<br /> L=22 mm
Copyright © Ruslan Panin
Carabus (Archicarabus) phoenix Lapouge, 1924
Male+Female adult
C-Lebanon, Byblos /Jbeil Distr., Kfardebian env., h=1200m, XI.2014, C. Reuter leg. female,23mm
Copyright © Ruslan Panin
Carabus (Archicarabus) victor Fischer, 1836
Female adult
NE-Turkey, Giresun, Sehitler Geçidi (Pass) h~2000 m, 23.VII.2001, S. Dacatra leg., L=19 mm
Copyright © Ruslan Panin
Carabus (Archicarabus) rossii Dejean, 1826
Female adult
NC Italy, Emilia reg., Bologna, San Martino, 29.V.2016, G..L.Agnoli leg. L=30 mm
Copyright © Ruslan Panin
Carabus (Archicarabus) alysidotus Illiger, 1798
Female adult
Italy, Reggio Emilia, Albinea env. h=150 m, VI.2018, A. Cartini leg., L=25 mm
Copyright © Ruslan Panin
Carabus (Archicarabus) wiedemanni congruus Motschulsky, 1865
Female adult
Turkey, Cankiri, Cerkes - Kursunlu, Atkaracalar 1250m 10.7.1996 Leg Staven and Skoupy (DUBC)
Copyright © Alexander Anichtchenko
Carabus (Archicarabus) monticola monticola (as maritimensis Born, 1923)
Female adult
SE France, Alpes-Maritimes, Col de Braus h=1000 m, 23.V-18.VI.2014, P. Cavazzuti & G. Proscia leg., L= 24 mm
Copyright © Ruslan Panin
Carabus (Archicarabus) victor Fischer, 1836
Male adult
S Georgia, Samtskhe-Javakheti, Akhaltsikhe distr., near Andriatsminda, Anda vill h=1550 m, 14.VII.2021, E. Rutjan leg./ det. R.Panin l=21 mm
Copyright © Ruslan Panin -
Aristocarabus taxon page (13 photos)
Carabus (Aristocarabus) viridifossulatus viridifossulatus Fairmaire, 1887
Female adult
35mm
Copyright © Boleslav Brezina
Carabus (Aristocarabus) viridifossulatus seticollis Deuve & Mourzine, 2000
Male adult
28 mm
Copyright © Boleslav Brezina -
Aulonocarabus taxon page (70 photos)
Carabus (Aulonocarabus) gaschkewitschi czarensis Obydov, 1997
Female adult
Russia, CE-Siberia, N Chita reg., Kodar Mt. Range, Big Leprindo Lake env. h~2000m., 2-5.VII.2002 A. Prokhorov leg./det. A. Anichtchenko
L=19 mm
Copyright © Ruslan Panin
Carabus (Aulonocarabus) canaliculatus hailinensis Deuve & Li, 2000
Male adult
NE China, Heilongjiang Prov., Mudanjiang City, Mt.Tai-Gu-Ding-zi h=800m, 15-25.VI.2010. L=29 mm
Copyright © Ruslan Panin
Carabus (Aulonocarabus) kurilensis ? daisetsuzanus Kono, 1936
Male adult
N-Japan, Hokkaido, Katō District,Kamishihoro, Horoka near Maruyama Bridge,19.VI.2013, T. Hirabayashi leg.
L=24 mm
Copyright © Ruslan Panin
Carabus (Aulonocarabus) gaschkewitschi gaschkewitschi Motschulsky, 1859
Male adult
Copyright © Kirill Makarov
Carabus (Aulonocarabus) kolymensis kodarensis Brinev, 1997
Male adult
Russia, SE Siberia, N Chita reg., Kodar Mt. Rng. h=1800m, Novaya Chara vill., 23.VI.2006. L=17 mm
Copyright © Ruslan Panin
Carabus (Aulonocarabus) truncaticollis fleischeri Reitter, 1898
Female adult
Russia, N Burajatia, Barguzinskiy Mts., the upper reaches of M.Chivyrkuy riv., h=2000 m, 26.VI-12.VII.2007, Vashchenko S leg. L=18.5 mm
Copyright © Ruslan Panin
Carabus (Aulonocarabus) truncaticollis fleischeri Reitter, 1898
Male adult
Russia, N Burajatia, Barguzinskiy Mts., the upper reaches of M.Chivyrkuy riv., h=2000 m, 26.VI-12.VII.2007, Vashchenko S leg. L=17 mm
Copyright © Ruslan Panin -
Axinocarabus taxon page (7 photos)
Carabus (Axinocarabus) fedtschenkoi fedtschenkoi Solsky, 1874
Male adult
27 mm
Copyright © Boleslav Brezina
Carabus (Axinocarabus) fedtschenkoi kondarensis Kryzhanovskij, 1953
Female adult
NW-Tadzhikistan: Gissar Mt.Rng, Varzob river ravine, Gushari env., h~1400-1500m, 24.III.2010, L=29 mm
Copyright © Ruslan Panin -
Calocarabus taxon page (61 photos)
Carabus (Calocarabus) linxiaensis ponggartangensis Deuve, 1995 Holotype
Male adult
17 mm
Copyright © Boleslav Brezina
Carabus (Cryptocechenus) sifanicus Semenov, 1898 (as oxydocupreus)
Male adult
male, 17mm
Copyright © Boleslav Brezina -
Carabus sensu stricto taxon page (112 photos)
Carabus (Carabus) arvensis arvensis Herbst, 1784
Male+Female adult
W Ukraine, NW Podolian, Lviv reg., Javoriv distr., near Stradch vill., 20-30.IV.2009, Panin R. leg.
Copyright © Ruslan Panin
Carabus (Carabus) granulatus leander Kraatz, 1878
Male+Female adult
Russia, C-Caucasus: Balkaria, Valley of riv. Cherek Balkarskyi, h=1750-1800m 10.VI-20.VII.1992, Zamotajlov A. leg.
female,22mm
Copyright © Ruslan Panin
Carabus (Carabus) granulatus granulatus Linne, 1758
Male adult
W Ukraine,
Lviv Reg., Javoriv distr., near Ivano-Frankovo vill., 30.IV-18.V.2009, Panin R. leg.
male, 21mm
Copyright © Ruslan Panin
Carabus (Carabus) arvensis deyrollei Gory, 1839
Male adult
NW-Spain,
Cantabria, Udías env., Monte Corona h=100m,14.VI.2002, Javier de Diego leg.
L=16 mm
Copyright © Ruslan Panin
Carabus (Carabus) sculpturatus (as gilanicus Mandl, 1967)
Female adult
N-Iran:Gilan Prov., Asalem Distr., Siahbil, deciduous forest (very wet) h= -10 m, 20-24.VI.2017, V. Čutka leg., L=22 mm
Copyright © Ruslan Panin
Carabus (Carabus) goryi Dejean, 1831
Male adult
WV, Pacahontas Co, 6mi N of Durbin, 25.6.2010 (cAA)
Copyright © Alexander Anichtchenko
Carabus (Carabus) granulatus calabricus Spettoli & Vigna Taglianti, 2001
Male adult
S-Italy: Calabria, Nat.Park Aspromonte, near Gambarie h=1340 m, 38.158149, 15.823323, 3-6.IV.2025, M.Egger leg., L=19 mm
Copyright © Ruslan Panin
Carabus (Carabus) menetriesi menetriesi Hummel, 1827
Male adult
W Ukraine, Lviv city, Landscape Reserve “Peatbog Bilogorshcha” NW part, 17.IV-14.V.2018, Panin R. leg., L=21.5 mm
Copyright © Ruslan Panin -
Cathaicus taxon page (7 photos)
Carabus (Cathaicus) brandti brandti Faldermann, 1835 (as grossegranulatus) Syntypes Lectotype
Male adult
C. (Cathaicus) brandti grossegranulatus Breuning 1943<br /> Lectotypus/SyntypusLectotypus (25 mm) von "South Mandschou". Syntypus (25 mm) "South Mandschou, 30. V. 1937"
Copyright © Frank Kleinfeld -
Cathoplius taxon page (14 photos)
Carabus (Cathoplius) asperatus asperatus (Dejean, 1826)
Male adult
male, 24mm
Copyright © Boleslav Brezina -
Cechenochilus taxon page (46 photos)
Carabus (Procechenochilus) ? kokujewi Semenov, 1898
Female adult
Russia, NW Caucasus, Karachay-Cherkessia, Karachaevsk distr., near Uchkula vill., valley Uchkulanichi riv. h=2800-2900m, 25-28.VII.2010, Fominykh D. leg.
female, 23 mm
Copyright © Ruslan Panin
Carabus (Cechenochilus) heydenianus Starck, 1889
Male+Female adult
NW-Caucasus,
Krasnodar reg., Mostovskoy distr., Skirda Mt Range, N slopes Yatyrgvart Mt. h=2300-2500m., 09-10.VII.2008, D. Fominykh leg.
female,19mm/male,18mm
Copyright © Ruslan Panin
Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Holotype
Male adult
Russia, Karachai-Circassian Republic, Abishira-Akhuba Mount Range between passes Agur and Mylgvar, 2785–2830 m.
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) kokujewi Semenov, 1898
Male adult
Russia, Karachai-Cherkessia, vall. Daut Riv., near Glacier Ryndzhi-Murudzhu, h = 3050 m, 22.VIII.2011, leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) kokujewi Semenov, 1898
Female adult
Russia, Karachai-Cherkessia, vall. Daut Riv., near Glacier Ryndzhi-Murudzhu, h = 3100 m, 22.VIII.2011, leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) kokujewi Semenov, 1898
Female adult
Russia, Karachai-Cherkessia, vall. Daut Riv., near Glacier Ryndzhi-Murudzhu, h = 2550 m, 23.VIII.2011, leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) boeberi tschitscherini Semenov, 1896
Male adult
Russia, C Caucasus, Ingushetia, S slopes of Skalistyi Mt.R.
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) heydenianus Starck, 1889
Male adult
Carabus (Procechenochilus) heydenianus euxinus Semenov, 1896 Abkhasia: Arabika plato (TOPOTYPUS)
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) boeberi aequaliceps Reitter, 1896
Female adult
Russia, Dagestan, Andiyskiy Mt.R., near Andiyskiye vorota pass, 06.VI–18.VII.2021, leg. D. Fominykh
Copyright © Igor Solodovnikov
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Cephalornis taxon page (1 photos)
Carabus (Cephalornis) potanini (Semenov, 1887)
Male adult
China, S Gansu, 3400m, Minshan mts.,
Wenxian 8.92, J. M. Bousquet
Copyright © Boleslav Brezina -
Chaetocarabus taxon page (32 photos)
Carabus (Chaetocarabus) intricatus bayardi Solier, 1835
Male+Female adult
S-Italy,
Basilicata Reg., Potenza Prov., Latronico, F. Angelini leg. 14.VI.2002
female, 26mm
Copyright © Ruslan Panin
Carabus (Heterocarabus) marietti marietti (as ativsengueni Schweiger, 1962)
Male adult
male, 17mm
Copyright © Boleslav Brezina
Carabus (Chaetocarabus) intricatus krueperi Reitter, 1896
Male adult
CE-Greece: Pieria, Stomio (Larissa) Ossa Mt. 7-14.VII.2009, Krause Rene leg.
L=35 mm
Copyright © Ruslan Panin
Carabus (Chaetocarabus) arcadicus arcadicus Gistl, 1850
Female adult
CE-Greece, Kato Olympos, Karia env., 6-17.VII.2011, Krause Rene leg. L=37 mm
Copyright © Ruslan Panin
Carabus (Chaetocarabus) arcadicus merlini Schaum, 1861
Male adult
S-Greece:Peloponnesos: Taygetos Mt.Rng., 17-23.V.2005, V. Sima leg.
L=30 mm
Copyright © Ruslan Panin
Carabus (Heterocarabus) marietti marietti Cristoforis & Jan, 1837
Female adult
as ativsengueni Schweiger, 1962
NW Turkey, Bolu, Abant Mts.; 16mm
Copyright © Aleš Sedláček
Carabus (Chaetocarabus) arcadicus arcadicus Gistl, 1850
Male adult
Graecia, Pindus, Mt. Velouchi (Timfristos), 1200m, vi.1970-A. Sermet Leg., 30 mm
Copyright © Noël Mal
Carabus (Chaetocarabus) intricatus bayardi (as silaensis Haury, 1881)
Female adult
S-Italy, Calabria, Sila Grande Massif, Camigliatello h=1040 m, 19.XI.2009, G. Proscia leg. L=31 mm
Copyright © Ruslan Panin
Carabus (Chaetocarabus) intricatus bayardi (as silaensis Haury, 1881)
Male adult
S-Italy, Calabria, Sila Grande Massif, Camigliatello h=1040 m, 19.XI.2009, G. Proscia leg. L=28 mm
Copyright © Ruslan Panin
Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Female adult
W Ukraine, Lviv city, Holosko, Bryukhovytsʹkyi forest 49.886833, 24.012083 under bark, 10.XI.2024, R. Panin leg., L=35 mm
Copyright © Ruslan Panin -
Chrysocarabus taxon page (83 photos)
Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as delacrei Tarrier, 1975)
Male+Female adult
NE Spain, Catalonia, Sant Jaume de Frontanyа, h=1000 m, V-VI.2010, Pascal Stefani leg.
female, 38 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) olympiae Sella, 1855
Male+Female adult
N-Italy,
Piemonte, Bielmonte, 14-15.VI.1981, Pascal Stefani leg.
female, 34 mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Chrysocarabus) auronitens costellatus Gehin, 1882 (as subfestivus Oberthur, 1884)
Female adult
W- FRANCE,
Bretagne Reg., Finistrere, Foret de Coatloc'h, XI.2006
female, 26 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) auronitens escheri Palliardi, 1825 (ab. opacus Haury, 1878)
Male+Female adult
W Ukraine, NE Carpathians Mts.,<br /> Transcarpathian reg., Rakhov Distr., Chornogora range, h=2020m, alpin zone, 28.VI-22.VIII.2009, Panin R. leg.<br /> female, 27mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) splendens Olivier, 1790
Male+Female adult
SE France,
Midi-Pyrйnйes reg., Ariиge, Vaychis, 22.X.2002, A.Laforgue leg.
female, 30mm.
Copyright © Ruslan Panin
Carabus (Chrysocarabus) hispanus Fabricius, 1787
Male adult
FRANCE,
Tarn, Foret de la Gresigne, h=400m.VII.2005 Laforgue leg.
male-30mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) lineatus lateralis Chevrolat, 1840 (as quasilineatus Tarrier, 1975)
Male+Female adult
NW Spain,
E Asturias, Monte de Moal h~1000m., 5.VI.2008, Pascal Stefani leg.
female, 27mm.
Copyright © Ruslan Panin
Carabus (Chrysocarabus) lineatus lineatus (as pseudonumancianus Mollard, 2010)
Male adult
NC-Spain, Sierra de Cantabria , Sierra de Toloño Rng, Puerto de Herrera , h=1100m, 19.I,2007 25 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896
Habitat
Andorra, Anyos, ii.2012, J. Nicolas Leg. & Photos
Copyright © Noël Mal
Carabus (Chrysocarabus) auronitens kraussi Lapouge, 1898
Female adult
C-Austria, Styria, Ennstal Alps, Totes Gebirge Mt Rng, Mt Loser h~1100 m, 16-20.VII.2014, R.Janak leg. L=27 mm
Copyright © Ruslan Panin -
Coptolabrus taxon page (181 photos)
Carabus (Coptolabrus) pustulifer guerryi Born, 1903
Female adult
N Yunnan: Gaoligongshan, h =1300 m, 06.2003 Local people
Copyright © Igor Solodovnikov
Carabus (Coptolabrus) pustulifer (as cyanelongatus Deuve, 2006)
Female adult
Zhagyab, Quamdu City, Tibet<br /> 8-15.VI.2008
Copyright ©
Carabus (Coptolabrus) jankowskii benxiensis Imura, 1991
Male adult
NE-China,
Liaoning Prov., Donggou, Mt. Guanmen, 15.VI.2009
male, 36mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) smaragdinus liaodongensis (Li, 1992)
Male adult
NE-China, E-Liaoning: Anshan: Mt.Qian Shan, 9-14.VI.2011. L=39 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) smaragdinus dolichognathus Deuve, 1995
Male adult
NW-Korea, North Pyongan Province, Mt. Donggorim, 1-20.VI.2013, local coll. leg.<br /> L=37 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) augustus augustus Bates, 1888
Male adult
SE-China, Jiangxi prov., Yudu County, Ganzhou city, Yun Fengzang h=1000m, 5-20.V.2011
L=43 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) augustus ignigena Hauser, 1914
Female adult
China : Guizhoh, Sinan vi.2006 (Size : 35 mm) Coll. N. Mal
Copyright © Noël Mal
Carabus (Coptolabrus) jankowskii fusanus (as obtusipennis)
Female adult
SC-Korea: Chiri-san Massif, Mt. Jirisan, 28.VI.2011, L=35 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) gemmifer gemmifer (as loczyi Csiki, 1927)
Female adult
S-China, W-Yunnan: Dali city, Mt Jizushan, VI.2018, L=34 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) pustulifer mirificus Kraatz, 1894
Male adult
SC-Chine, NW-Hunan, Longshan County, Mt. Maoping,10-30.VI.2008, L=44 mm
Copyright © Ruslan Panin -
Cratocarabus taxon page (5 photos)
-
Cratocechenus taxon page (38 photos)
Carabus (Cratocechenus) jacobsoni Semenov, 1898
Male adult
S Kazakhstan, Almaty reg., Big Lake, h=2800 m, 25.VI.96
male
L=14 mm
Copyright © Maksim Smirnov
Carabus (Cratocechenus) akinini akinini Morawitz, 1886 (as oitalensis ) Paratype
Male adult
male, 20mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) akinini akinini Morawitz, 1886
Female adult
female, 21mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) ovtchinnikovi ovtchinnikovi Gottwald, 1987
Male adult
N-Kyrgyzstan, Kyungey Ala-Too Mt Rng, Cholpon-Ata env., h=2500m, 19.VI.2000,
L=19 mm
Copyright © Ruslan Panin -
Cratocephalus taxon page (17 photos)
Carabus (Cratocephalus) solskyi Ballion, 1878
Male adult
SE Kazakhstan, Ketmen Mt. rng., Tuyuk vill., 16.06.97 leg. S.Toropov
Copyright © Maksim Smirnov
Carabus (Cratocephalus) solskyi Ballion, 1878
Female adult
SE-KAZAKHSTAN, Ketmen Mt Rng, h=1800m, Tujuk vill.,15.V.2004
L=36 mm
Copyright © Ruslan Panin
Carabus (Cratocephalus) cicatricosus cicatricosus Fischer, 1842
Male adult
SE-Kazakhstan, Almaty reg., C Trans-Ili Alatau N-slopes, Kaskelen Valley, Izvestkovyi vill., h=1700 m, 6.V.2015, D.Kovalchuk leg., L=28.5 mm
Copyright © Ruslan Panin -
Cratophyrtus taxon page (15 photos)
Carabus (Cratophyrtus) turcosinensis medvedevi Kryzhanovskij, 1968
Female adult
S Kazakhstan, NW slope of Ugamskii mt.rng., 3 km W Sairamskii peak, 2200m
tributary of Karzhantau riv., 15.6.2004 Matalin A. leg.
20 mm
Copyright © Alexander Anichtchenko
Carabus (Cratophyrtus) kaufmanni segregatus Morawitz, 1886
Female adult
female, 19mm
Copyright © Boleslav Brezina -
Cryptocarabus taxon page (16 photos)
Carabus (Cryptocarabus) subparallelus Ballion, 1878
Female adult
Kazakhstan,
near Alma-Ata, V.2009, Pak O.leg.
female, 17mm
Copyright © Ruslan Panin
Carabus (Cryptocarabus) lindemanni Ballion, 1878
Female adult
SE Kazakhstan, Alma-Ata env., 16.04. 2000
L=16,0 mm
Copyright © Maksim Smirnov
Carabus (Cryptocarabus) ? turkestanus turkestanus Breuning, 1929
Male adult
Kyrgyzstan, W Kungei Ala-Too Mt Rng, Chon-Kemin riv., Kyzyl-Bairak vill., 25.IV.2010. L= 18 mm
Copyright © Ruslan Panin
Carabus (Cryptocarabus) tsharynensis Kabak 1994
Male adult
SE-Kazakhstan, Almaty Region, W-Zailiyskiy Alatau rng, Belshabdar valley h~1300 m, 25-27.VI.2002, A.Puchkov leg. L=17 mm
Copyright © Ruslan Panin
Carabus (Cryptocarabus) tsharynensis Kabak 1994
Female adult
E-Kazakhstan, Almaty Region, W-Zailiyskiy Alatau, Turgen distr., Tauturgen vill., h~1200 m, 8-13.VI.2002, A.Puchkov leg. L=19 mm
Copyright © Ruslan Panin
Carabus (Cryptocarabus) sacarum Kabak, 1998
Male adult
SE-Kazakhstan, Almaty reg., C-Zailiyskiy Alatau N-slopes, Kaskalen river Valley, Izvestkovy vill., h=1700 m, 6.V.2015, D.Kovalchuk leg./det. R.Panin L=18 mm
Copyright © Ruslan Panin
Carabus (Cryptocarabus) sacarum Kabak, 1998
Female adult
SE-Kazakhstan, Almaty reg., C-Zailiyskiy Alatau N-slopes, Kaskalen river Valley, Izvestkovy vill., h=1700 m, 6.V.2015, D.Kovalchuk leg./det. R.Panin L=20 mm
Copyright © Ruslan Panin -
Ctenocarabus taxon page (8 photos)
Carabus (Ctenocarabus) galicianus Gory, 1839
Male adult
Spain,
Sierra de la Cabrera Baja, 8 km SW Truchillas, 9.VII.2008, Anichtchenko A. leg.
L= 20 mm
Copyright © Alexander Anichtchenko
Carabus (Ctenocarabus) galicianus Gory, 1839
Female adult
NW-Spain, Pontevedra prov., Campo Lameiro env., 15.VIII.1994, S.Sobrino leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Ctenocarabus) melancholicus costatus Germar, 1824
Female adult
C-Spain, Madrid region, Rozas de Puerto Real env.,21.XII.2008, T.Gazurek leg.
L=27 mm
Copyright © Ruslan Panin
Carabus (Ctenocarabus) melancholicus melancholicus Fabricius, 1798
Male adult
NW Morocco, Kénitra, Souk El Arbaa env., II.1985, V.Olivella leg. L=25 mm
Copyright © Ruslan Panin
Carabus (Ctenocarabus) melancholicus submeridionalis Breuning, 1975
Male adult
S- Spain, Cádiz, Tarifa, El Cabrito, 28.XII.2019, F. Garcia leg. L=26 mm
Copyright © Ruslan Panin
Carabus (Ctenocarabus) melancholicus costatus Germar, 1824
Male adult
NW Portugal, Viana do Castelo distr., Sierra de Agra, 41°47'52.3"N 8°41'27.6"W, h=786 m, 3.VII.2016, D. Prunier leg.
Copyright © Ruslan Panin -
Cupreocarabus taxon page (46 photos)
Carabus (Cupreocarabus) kangdingi kangdingi Korell, Kleinfeld & Gorgner, 1992 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Cupreocarabus) huangi huangi Deuve, 1992
Male adult
C-China, E Tibet, N-Sichuan, Hongyuan,valley 10 km SE Sanggarpar, h=4200m, 32.300053, 102.584031, alpine meadow / scree, 19.VII.1995, B.Brezina leg.<br /> L=21 mm
Copyright © Ruslan Panin
Carabus (Cupreocarabus) lixianensis lixianensis Deuve, 1990
Male adult
China, E Tibet, N-Sichuan, (Barkam),valley SW of Barkam, h=3200m, 31*53’ N; 102*12’E, mixed/coniferous forest, 22.VII.1995, K.& B.Brezina
L=23 mm
Copyright © Ruslan Panin
Carabus (Cupreocarabus) alpherakii alpherakii Semenov, 1898
Female adult
China, W Sichuan, Litang env., h~5000 m,29.VII -3.VIIII.1995, J. Schneider leg. L=23 mm
Copyright © Ruslan Panin
Carabus (Cupreocarabus) mollardianus Deuve, 2016
Female adult
Chine, Sichuan, Lushan, Mt. Jiajin Shan, 10.VII.2016 local collector leg./ det. R.Panin, L=22 mm
Copyright © Ruslan Panin
Carabus (Cupreocarabus) balangicus balangicus Cavazzuti, 2002
Male adult
Chine, C-Sichuan, Xiaojin Co, Wolong valley, Balang Shan Pass h=3448 m, 30.884617, 102.973117, 9.VI.-1.VII.2011, G. Proscia leg. L=19 mm
Copyright © Ruslan Panin
Carabus (Cupreocarabus) balangicus balangicus Cavazzuti, 2002
Male adult
Chine, C-Sichuan, Xiaojin Co, Wolong valley, Balang Shan Pass h=3448 m, 30.884617, 102.973117, 9.VI.-1.VII.2011, G. Proscia leg. L=20 mm
Copyright © Ruslan Panin
Carabus (Cupreocarabus) huangi huangi Deuve, 1992
Female adult
China, E Tibet, N-Sichuan, 53 km NE of Barkam, Pass Sandaoban env., h=4293 m, 32.232950, 102.598533, 11-29.VI.2011, G. Proscia leg., L=21.5 mm
Copyright © Ruslan Panin
Carabus (Cupreocarabus) lixianensis lixianensis Deuve, 1990
Male adult
China, E Tibet, N-Sichuan, (Barkam), valley SW of Barkam, h=3200m, 31*53’ N; 102*12’E ,mixed/coniferous forest, 22.VII.1995, K.& B.Brezina leg., L=23 mm
Copyright © Ruslan Panin -
Cychrostomus taxon page (8 photos)
Carabus (Cychrostomus) pseudoprosodes Semenov & Znojko, 1932
Male adult
male, 19mm
Copyright © Boleslav Brezina
Carabus (Cychrostomus) anchocephalus lanzhouensis Deuve, 1991 Paratype
Male adult
male, 21mm
Copyright © Boleslav Brezina
Carabus (Cychrostomus) anchocephalus anchocephalus Reitter, 1896
Male adult
male, 22mm
Copyright © Boleslav Brezina -
Cyclocarabus taxon page (14 photos)
Carabus (Cyclocarabus) aulacocnemus galinae Obydov, 2003 Paratype
Male+Female adult
W-Tianshan:Karatau Mts.,N-part:Boialdyr River valley,
N of Ken-su, 600-750m, 22-V-2000 М. Danilevsky leg.
male - 20 мм, female - 19 мм.
Copyright © Alexandr Safronov
Carabus (Cyclocarabus) namanganensis namanganensis Heyden, 1886
Male+Female adult
Kazakhstan, S. Karatau Mts. R., Mashat vill. env.
23-IV-1993 М. Danilevsky leg.
male - 21 мм, female - 22 мм.
Copyright © Alexandr Safronov
Carabus (Cyclocarabus) zarudnyi Semenov & Znojko, 1932
Female adult
NE-Uzbekistan, W-Tian-Shan, NW slopes Pskem Mt. Rng, 8-14 km E Pskem, 2100-2500 m, 16.VI.1998, A. Puchkov leg., L=21 mm
Copyright © Ruslan Panin
Carabus (Cyclocarabus) karaterekensis hemicallisthenes Kabak & Kalab, 1997
Male adult
NW- Kyrgyzstan, W-Tian-Shan, Chandalash Mt.Rng, Chakmak-Suu vill., 3.VI.2000, L=17 mm
Copyright © Ruslan Panin
Carabus (Cyclocarabus) brosciformis Semenov, 1896
Male adult
NE-Uzbekistan, W-Tian-Shan, Chatkal Mt Range, Tashkent reg., Yangiabad env., h=2100m, 30.IV.2002, L=23.5 mm
Copyright © Ruslan Panin -
Damaster taxon page (15 photos)
Carabus (Damaster) blaptoides oxuroides Schaum, 1862
Female adult
Japan, C-Honshu, Ofuse-cho, Kamitakai, Nagano Pref., 12.VI.2006, Ohshima leg.
L=34 mm
Copyright © Ruslan Panin
Carabus (Damaster) blaptoides capito Lewis, 1881
Male adult
Japan: Niigato pref., Sado Isl., Sado, Yamato
Copyright © Igor Solodovnikov
Carabus (Damaster) blaptoides oxuroides Schaum, 1862
Female adult
Japan, C Honshu, Nagano Prefecture, Nagano city, 3.IV.2012, S. Shinichi leg. L=42 mm. det. Anichtchenko A. 2021
Copyright © Ruslan Panin
Carabus (Damaster) blaptoides rugipennis Motschulsky, 1861
Male adult
Kunashir isl.
Copyright © Kirill Makarov
Carabus (Damaster) blaptoides blaptoides (Kollar, 1836)
Female adult
Japan, SW Honshu, Shimane Pref., Unnan city, 16.I2016, S.Shinichi leg., L=48 mm
Copyright © Ruslan Panin -
Deroplectes taxon page (32 photos)
Carabus (Deroplectes) staudingeri Ganglbauer, 1886
Female adult
N Tajikistan, SW Pamir-Alay, Fann Mts h=1800m, Padrut vill., 2-3.V.1997
L=28 mm
Copyright © Ruslan Panin
Carabus (Deroplectes) dokhtouroffi dokhtouroffi Ganglbauer, 1886
Male adult
NC Tadzhikistan, Pamir, N-slope Peter I.Mt Rng, h=2100m,Ganishou vill., 10-12.VI.2006, Ivanova E. leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Deroplectes) klapperichianus coiffaitianus Deuve, 1990
Male adult
Tadzhikistan: Badakhshan: Ishkashim Mt.R.: Kuh-i-Lal, VI.1979 leg. I.K. Lopatin (TOPOTYPUS)
Copyright © Igor Solodovnikov
Carabus (Deroplectes) klapperichianus coiffaitianus Deuve, 1990
Female adult
Tadzhikistan: Badakhshan: Ishkashim Mt.R.: Kuh-i-Lal, VI.1979 leg. I.K. Lopatin (TOPOTYPUS)
Copyright © Igor Solodovnikov
Carabus (Deroplectes) klapperichianus salangicus Kleinfeld & Reuter, 2010 Paratype
Female adult
N-Afghanistan, Hindukush Mt.Rng, Salang Pass, Nordseite, 25°27'.494N, 68°58.399'E, ca. reg. subalp., h = 2200-2400 m, V.2010, leg. Christoph Reuter
Copyright © Igor Solodovnikov -
Diocarabus taxon page (46 photos)
Carabus (Diocarabus) loschnikovi loschnikovi Fischer, 1823
Male adult
Komi Resp., Pecero-Ilych Nat. Res., Zajachij Kamen h~786 62°10'06" N 59°12'53" E 8.VIII.2004 M. Chernjakhovski leg.
L=15.0 mm
Copyright © Evgeniy Zinoviev
Carabus (Diocarabus) slovtzovi slovtzovi Mannerheim, 1849
Male adult
S Siberia, Burjatia reg., E Sayan Mts., Khugaituma mt., h=1700 m, 1-5. VII.2001
L=14,3 mm
Copyright © Maksim Smirnov
Carabus (Diocarabus) beybienkoi Kryzhanovskij, 1973
Male adult
S Sakhalin, Susunaj Mt.R. 10.08 - 13.09.2003 D. Kochetkov leg.
L=14,5 mm
Copyright © Maksim Smirnov
Carabus (Diocarabus) linnaei linnaei (as scopolii Dejean, 1826)
Female adult
W Ukraine, NW-Podolian, near Lviv city, Vynnykivskyi forest, 20.IV.-13.V.2011, R. Panin leg.<br /> L=22 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) linnaei linnaei (as scopolii Dejean, 1826)
Male adult
W Ukraine, Lviv reg., NW Podolian Upland, forest near Vybranivka vill., 12.X.2007, Panin R. leg. L=19 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L =17 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=19 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Gutyn-Tomnatyk h~2015 m alpin zone, 8.VI-28.VII.2011, R. Panin R. leg. L=14 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Female adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Gutyn-Tomnatyk h~2015 m, alpin zone, 8.VI-28.VII.2011, R. Panin R. leg. L=18 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) fraterculus neochinensis Deuve & Li, 1998
Male adult
NE-China: Liaoning prov., Fengcheng City, Shicheng, 20-28.VI.2011, Li Jingke leg., L=20 mm
Copyright © Ruslan Panin -
Eccoptolabrus taxon page (22 photos)
Carabus (Lasiocoptolabrus) sunwukong sunwukong Imura, 1993 Paratype
Male adult
Copyright © Boleslav Brezina
Carabus (Lasiocoptolabrus) sunwukong baotianmanus (Imura, 2005) Paratype
Male adult
Copyright © Boleslav Brezina
Carabus (Eccoptolabrus) exiguus nivium Breuning, 1933
Female adult
female, 20mm
Copyright © Boleslav Brezina
Carabus (Eccoptolabrus) exiguus exiguus Semenov, 1898 Holotype
Male adult
male, 18 mm
Copyright © Frank Kleinfeld -
Eucarabus taxon page (111 photos)
Carabus (Eucarabus) obsoletus obsoletus Sturm, 1815
Male adult
NW Ukrainian Carpathians, Lviv Reg., Staryi Sambor Distr., near Dobromyl , h=500 m., 15-24.VI.2005, Panin R. leg., male, 28mm
Copyright © Ruslan Panin
Carabus (Eucarabus) catenulatus catenulatus Scopoli, 1763
Female adult
NW Croatia, Bosanci, 1-12,VI.2004, Baros Slovak leg.<br /> L=33 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825
Male+Female adult
Male - SW Romania, Banat - Bocsa/Coltan,
21.V-05.VI.2010, R. Krause leg., L=29 mm.
Female - SW Romania, Banat - Saska Montana, Stinapari,
25-VI-2010, R. Krause leg., L=31 mm.
Copyright © Alexandr Safronov
Carabus (Eucarabus) ulrichi rhilensis (as stussineri Géhin, 1885)
Male adult
W Croatia, Ivanščica Mts , Ivanec, 24.VI-7.VII.2006, Krause Rene leg.
L=29 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) ulrichi ulrichi (as intercessor Sokolar, 1912)
Male adult
N Hungary, Pilis Mts., Piliszentlelek, 20-27.VII.2011, Krause Rene leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825
Male adult
SW Romania, Banat - Eibenthal; 29 mm
Copyright © Aleš Sedláček
Carabus (Eucarabus) obsoletus nagyagensis (as fossulifer Fleischer, 1893)
Male adult
NW-Romania, Bihor, Mts Seș (Plopis), 20 km N Aleșd, Pădurea Neagră env., VI.2014, J.Kynkor leg. L=24 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) obsoletus obsoletus (as csikii Mallasz, 1901)
Female adult
NE Carpathians, Marmarosh Mts., Transcarphatian Reg., near Lug vill., h=350 m, 26.IV-8.VI.2014, R.Panin & Yu. Kanarsky leg. L=30 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) parreyssi parreyssi Palliardi, 1825
Male adult
W-Croatia, NP Northern Velebit, Zavižan,15.V-13.VI.1996, Colombetta leg., L=21 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) billbergi kuandianicus Rapuzzi, 2007
Female adult
NE China, Liaoning, Kuandian County, Dachuantou env., 20-29.VI.2015,
Copyright © Ruslan Panin -
Euleptocarabus taxon page (5 photos)
Carabus (Euleptocarabus) porrecticollis porrecticollis Bates, 1883
Male adult
Japan: Niigata pref., Chayagahara Jyouetsu City,
18.XI.2006 A.Toba leg.
Copyright © Alexander Anichtchenko
Carabus (Euleptocarabus) porrecticollis Bates, 1883
Female adult
Atsushiokanomachikano, Kitakata City, Fukushima Pref., Japan, 29.III.2014, T. Nakamura leg.
Copyright © Alexander Anichtchenko
Carabus (Euleptocarabus) porrecticollis porrecticollis Bates, 1883
Male adult
Japan, C Honshu, Nagano Pre., Toyooka town, 28.XII.2019, S. Shinichi leg. L=30 mm
Copyright © Ruslan Panin -
Eupachys taxon page (3 photos)
Carabus (Eupachys) glyptopterus Fischer, 1827
Male+Female adult
Russia, SE-Siberia,
Buryat Republic, Ivolginsky Distr., 04.VII.2010, Filippov A.E. leg.
female, 32 mm
photo Panin R.
Copyright © Ruslan Panin -
Eurycarabus taxon page (29 photos)
Carabus (Eurycarabus) famini lucasi Gaubil, 1849
Male adult
N-Tunisia, Bizerte reg., Menzel Bourguiba city,h=5 m, XII.1981, S. Remond leg.
L=25 mm
Copyright © Ruslan Panin
Carabus (Eurycarabus) famini fezzanus Lapouge, 1915
Male adult
Morocco, Middle Atlas Mt Rng, Azrou city h=1500m, IV.1999, S. Remond leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Eurycarabus) famini piraticus (as mamorensis (Kocher, 1964)
Female adult
NW-Morocco, betw.Meknes/Kenitra, Foret de Mamora, II.2008
L= 27 mm
Copyright © Ruslan Panin
Carabus (Eurycarabus) famini atlantis Alluaud, 1923
Female adult
C Morocco, Haut Atlas, SW of Oukaïmeden h=2900m, 31.173644, -7.844442, 29.IV-11.V.2024, C.Reuter leg., L=21 mm
Copyright © Ruslan Panin -
Fulgenticarabus taxon page (2 photos)
Carabus (Fulgenticarabus) flutschi koreus Deuve, 2006
Male adult
N Korea, Samjiyon, Ryang Gang Province
6-12.6.2006 leg. Wong
Copyright © Boleslav Brezina -
Gnathocarabus taxon page (2 photos)
Carabus (Gnathocarabus) kusnetzovi Semenov, 1903
Female adult
female, 21mm
Copyright © Boleslav Brezina -
Goniocarabus taxon page (14 photos)
Carabus (Goniocarabus) sogdianus Semenov, 1898
Male adult
N Tajikistan, SW Pamir-Alay, Zarafshan Range, Fann Mts, Marguzor Lake circ., Padrut vill., 16.V.1997
L=21 mm
Copyright © Ruslan Panin
Carabus (Goniocarabus) grombczewskii grombczewskii Semenov, 1891
Male adult
NC Tadzhikistan, Pamir, N-slope Peter I.Mt Rng, h=2100m, Ganishou vill.,12.VII.2006, Pak O. leg.
L=19 mm
Copyright © Ruslan Panin
Carabus (Goniocarabus) perelloi Casale, 1979
Male adult
N-Afghanistan, Hindukush, Provinz Baghlan, N of Kabul, N Salang Pass, subalp., h = 2200 m, 35°27'.494N / 68°58.399'E, Mai 2011, leg. Christoph Reuter (TOPOTYPUS)
Copyright © Igor Solodovnikov -
Hemicarabus taxon page (31 photos)
Carabus (Hemicarabus) tuberculosus Dejean, 1829
Male adult
Far East, Lazovsky Dist., shore of Japan Sea, 3 km NE Preobrazhenie Vill., Ezhovaya (Tasovaya) Bay 13.07.06 M. & L. Smirnov's leg.
male
L=20,0 mm
Copyright © Maksim Smirnov
Carabus (Hemicarabus) nitens Linne, 1758
Male+Female adult
Belarus,
Vitebsk reg., Ushachskiy distr., near Bolshie Dolcy vill. 9.V.1990, I.A.Solodovnikov leg.
female, 18mm.
Copyright © Ruslan Panin
Carabus (Hemicarabus) serratus Say, 1825
Male adult
E-USA,Virginia, Leesburg env., Ball's Bluff Battlefield Regional Park, 27.VIII-4.IX.2015,V. Čutka leg. L=17 mm
Copyright © Ruslan Panin
Carabus (Hemicarabus) serratus Say, 1825
Female adult
E-USA, Virginia, Leesburg env., Ball's Bluff Battlefield Regional Park, 27.VIII-4.IX.2015,V. Čutka leg. L=18 mm
Copyright © Ruslan Panin -
Homoeocarabus taxon page (6 photos)
Carabus (Homoeocarabus) maeander maeander s.str.
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Carabus (Homoeocarabus) maeander paludis Gehin, 1885
Female adult
Japan, Hokkaido; Tomakomai city, Benten, 17.V.2015, Takemoto leg., L=21 mm
Copyright © Ruslan Panin -
Hygrocarabus taxon page (5 photos)
Carabus (Hygrocarabus) nodulosus Creutzer, 1799
Male+Female adult
Austria, Styria Region (Steiermark), Leibnitz, 31.X.1999, I.Rapuzzi leg.<br /> female, 29 mm
Copyright © Ruslan Panin
Carabus (Hygrocarabus) variolosus Fabricius, 1787
Male adult
W-Ukraine, NW Podolian,<br /> Lviv reg., Zhydachiv distr.,near Vybranivka vill., 15.V-8.VI.2008, Panin R. leg.<br /> male, 27mm
Copyright © Ruslan Panin
Carabus (Hygrocarabus) variolosus Fabricius, 1787
Male+Female adult
W-Ukraine,<br /> Lviv Reg., Zhydachiv distr., near Vybranivka vill., Panin R. leg.<br /> female, 30mm
Copyright © Ruslan Panin -
Imaibius taxon page (55 photos)
Carabus (Imaibius) baronii battalensis Heinz & Staven, 1998 Paratype
Male adult
Copyright © Alexander Anichtchenko
Carabus (Imaibius) dardiellus perobscurus Deuve, 1980
Male adult
N India.Kashmir, Loran, W part Pir Panchal range,h=2400-2500m, 33°49'"N, 74°20'E
L=25 mm
Copyright © Ruslan Panin
Carabus (Imaibius) dardiellus ? subpunctulus Roeschke, 1907
Male adult
N India, Kashmir, Pahalgam, Aru Valley h=2400m, VII.2007
L=25 mm
Copyright © Ruslan Panin
Carabus (Imaibius) wittmerorum Heinertz, 1978
Male adult
N. Pakistan: Frontier prov., Miandam, Swat vall., h=1800-2600 m, leg.S.Remond (TOPOTYPUS)
Copyright © Igor Solodovnikov
Carabus (Imaibius) barysomus hazarensis Deuve, 1982 (TOPOTYP)
Male adult
N-Pakistan: Mansehra prov., Hazara, Kaghan valley: Shocgran env., h = 2750 m L= 42 mm
Copyright © Igor Solodovnikov
Carabus (Imaibius) boysi boysi Tatum, 1851
Male adult
NW-India, Himachal Pradesh, Manikaran env., Pulga vill., 4.VII.2008. L=26 mm
Copyright © Ruslan Panin
Carabus (Imaibius) stoliczkanus brauni Heinz, 1993
Female adult
NE Pakistan, Azad Kashmir, Bagh Distr., Sudhan-Gali h~2300 m,15.VIII.1994, S Dacatra leg. L=37 mm
Copyright © Ruslan Panin
Carabus (Imaibius) caschmirensis fortetuberculatus Heinz, 1983 Paratype
Male adult
Copyright © Alexander Anichtchenko
Carabus (Imaibius) nouristani nouristani (Ledoux, 1977)
Female adult
NE-Afghanistan: Nuristan: Darah-I-Nur valley, 2900m 34°45'N 70°40'E, VIII.2012, C.Reuter leg., L=26 mm
Copyright © Ruslan Panin
Carabus (Imaibius) caschmirensis fortetuberculatus Heinz, 1983
Male adult
N Pakistan, Kaghan Valley, Shogran forest h=2150 m, 34.644722, 73.460556, 7-14,VI.2025, Ch.Reuter leg., L=34 mm
Copyright © Ruslan Panin -
Iniopachus taxon page (7 photos)
Carabus (Iniopachus) pyrenaeus jeanneli Deuve, 1994
Male adult
1-2: Pirineos, Canfranc, Ibon de Iserias 2200 m 16-17.7.05<br /> 3: Pirineos, Monte Perdido, Collata Anisolo 2400m 25.8.05
Copyright © Alexander Anichtchenko
Carabus (Iniopachus) auriculatus ubinensis (Puissegur, 1955)
Female adult
Spain, Montes Cantabricos, Peña Ubiña, 2400m
5-6.VII.2009 Anichtchenko A. leg.
female, 19 mm
Copyright © Alexander Anichtchenko
Carabus (Iniopachus) pyrenaeus pyrenaeus (as cephalicus Csiki, 1927)
Male adult
SW France, Pyrenees-Atlantiques, Mt.Cirque d'Esturou h=1800m, 29.VIII.2014, Pascal Stefani leg. L=20 mm
Copyright © Ruslan Panin -
Ischnocarabus taxon page (4 photos)
Carabus (Ischnocarabus) tenuitarsis tenuitarsis Kraatz, 1877
Male adult
male, 24mm
Copyright © Boleslav Brezina -
Isiocarabus taxon page (25 photos)
Carabus (Isiocarabus) fiduciarius fiduciarius (as fiduciarius tim) Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Isiocarabus) hunanicola liangshanicus (as dreuxi liangshanicus) HT PT Kleinfeld, 1998 Holotype Paratype
Male+Female adult
Carabus (Isiocarabus) hunanicola liangshanicus (as dreuxi liangshanicus) HT PT
Copyright © Frank Kleinfeld
Carabus (Isiocarabus) miaorum Lassalle & Prunier, 1993
Male adult
S-China, N-Guangxi, Mao'er Shan, h~1600 m, V.2009, L= 26 mm
Copyright © Ruslan Panin
Carabus (Isiocarabus) fiduciarius fiduciarius Thomson, 1856
Female adult
CE-China, Hubei, Xiaogan City, Shuangfeng Mountain, 26.VI.2016, L=30 mm
Copyright © Ruslan Panin
Carabus (Isiocarabus) miaorum Lassalle & Prunier, 1993
Male adult
China, NW-Guangxi, Mao'er Shan, h~1600 m 25°52'N 110°29'E, 28.V-9.VI.2009, C.Reuter leg.
Copyright © Ruslan Panin
Carabus (Isiocarabus) kiukiangensis orphniopterus Hauser, 1920
Female adult
SE-China, NW-Fujian, Emei Feng Shan, 27°01'N 117°05'E, h=1500 m, (reg. silv.), 17.V.2025, Ch.Reuter leg., L=30 mm
Copyright © Ruslan Panin -
Lamprostus taxon page (31 photos)
Carabus (Lamprostus) torosus torosus Frivaldsky, 1835
Male adult
NE-Bulgaria, Dobrich Region, Kavarna env.
L=31 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) saulcyi saulcyi Piochard, 1875
Male adult
C Lebanon, Jbeil Distr., Ehmej vill., h=1100 m.V.2012, C. Reuter leg.
L=24 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) torosus sinopensis Breuning & Ruspoli, 1970
Male adult
NC-Turkey, Sinop province, Erfelek env., h=200m, 10-27.VII.2014, C. Reuter leg. L=31 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) hemprichi elonensis Schweiger, 1970
Male adult
N-Israel, Midlle Galilea, Nazareth env., Hoshaya viil., 3-12..III.2015, B.Vas’ko leg./det. Panin R.
Copyright © Ruslan Panin
Carabus (Lamprostus) syrus cheikensis Deuve, 1992
Male adult
N-Israel, Midlle Galilea, Nazareth env., Hoshaya viil., 13.XII.2012, Vas’ko, leg./det. Panin R. L=30мм
Copyright © Ruslan Panin
Carabus (Lamprostus) torosus giresuni Breuning, 1964 (as paululus Heinz & Korge, 1964)
Male adult
NE-Turkey, Giresun, Sehitler Geçidi (Pass) h~2000 m,24.VII.2001, S. Dacatra leg., L=26 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) torosus nordmanni Chaudoir, 1848 (as enricoi )
Male adult
NE-Turkey:Ordu prov.,3 km N Gölköy h~1100 m, VII.2006, L=37 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) calleyi pseudoprasinus Lapouge, 1912
Male adult
S Georgia, Tsalka (Paravani) 41°29′18.7″N, 43°51′12.3″E, 8-9.V.2016, Pavel Turek leg. / det. I. Solodovnikov, L=23 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) syrus moukarzelae Deuve, 2002
Female adult
S-Lebanon: Marjaayoun, 18.XI.-11.XII.2017, C. Reuter leg., L=35 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) syrus cheikensis Deuve, 1992
Female adult
SE Lebanon, Rashaya Distr., Tannoura, oak forest, h=700 m, 11-19.IV.2016, C. Reuter leg., L= 32 mm
Copyright © Ruslan Panin -
Leptocarabus taxon page (46 photos)
Carabus (Leptocarabus) yokoae marcilhaci Deuve, 1992
Male adult
CHINA, Shanxi, Tschingling Mts., Taibeichan Nat. reserv.;
Copyright © Anton Menstschikov
Carabus (Leptocarabus) harmandi fujisan Ishikawa, 1972
Female adult
Japan, Yamanashi pref., Fuji-Rinds 1600m, Narusawa vill. 3.V.2005 Kurihara K. leg. 25 mm
Copyright © Alexander Anichtchenko
Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931
Male adult
NE-China:Liaoning prov., Fengcheng City, Miaoyang, 12-17.VI.2011<br /> L=26 mm
Copyright © Ruslan Panin
Carabus (Leptocarabus) procerulus Chaudoir, 1862
Male adult
Japan:S Honshu, Matsubara-ko, VIII.1997, S. Remond leg.
L=30 mm
Copyright © Ruslan Panin
Carabus (Leptocarabus) procerulus Chaudoir, 1862
Male adult
Japan, S Honshu, Mie Pref., Hakusan, VII.2001, S. Remond leg.<br /> L=33 mm
Copyright © Ruslan Panin
Carabus (Leptocarabus) arboreus pararboreus (Ishikawa, 1992)
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Carabus (Leptocarabus) arboreus fujisanus Bates, 1883
Male adult
Japan, CE-Honshu, Yamanashi pref., Mt. Fujisan h~1600m, 23.IX.1982, Y. Imura leg. 23 mm
Copyright © Ruslan Panin
Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931
Male adult
NE-China:E-Liaoning: NP Kuandian Tianqiaogou , VI.2010, L=25 mm, det. Panin R.
Copyright © Ruslan Panin
Carabus (Leptocarabus) arboreus fujisanus Bates, 1883
Female adult
Japan, Honshu, Mt. Fuji h~1900m, Shizuoka pref., Fujinomiya city, 23.VI.2007, M. Matsunaga leg., L=25 mm
Copyright © Ruslan Panin -
Leptoplesius taxon page (6 photos)
Carabus (Leptoplesius) merzbacheri kirgisiensis Mandl, 1967
Female adult
22mm
Copyright © Boleslav Brezina -
Limnocarabus taxon page (15 photos)
Carabus (Limnocarabus) clathratus clathratus Linne, 1761
Female adult
W-Ukraine, Maloe Polesie, Lviv Region, near Kamenka-Bugzkaya, 28.III.2008, Panin R. leg. L=31 mm
Copyright © Ruslan Panin
Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
Female adult
SW Slovakia, Bratislava env., IV.2004, Krause Rene leg.
L=32 mm
Copyright © Ruslan Panin
Carabus (Limnocarabus) clathratus clathratus (as jansoni Kraatz, 1890)
Male adult
W-Ireland, County Mayo, Bellacorick bog, 9.VII.2013, J.Stypula leg. L=27 mm
Copyright © Ruslan Panin
Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
Male adult
W Ukraine, Transcarpathian Reg., 4 km NE Chop, forest Peresh, Panin R. leg. L=30 mm
Copyright © Ruslan Panin -
Lipaster taxon page (18 photos)
Carabus (Lipaster) stjernvalli stjernvalli Mannerheim, 1830
Male+Female adult
Armenia,
Ashotsk, 22.VI.1990, P.V.Kazaryan leg.
female 34mm
Copyright © Ruslan Panin
Carabus (Lipaster) stjernvalli osellai Breuning & Ruspoli, 1970
Male+Female adult
NE-Turkey: Artvin:Hopa, Borçka env., h=750 m, 9.VI-7.VII.2011, M.Janalik leg.
female, 42 mm
Copyright © Ruslan Panin
Carabus (Lipaster) stjernvalli osellai Breuning & Ruspoli, 1970
Male adult
NE-Turkey: Artvin, Hopa / Çifteköprü, VII.1977, L=36 mm
Copyright © Ruslan Panin
Carabus (Lipaster) stjernvalli stjernvalli Mannerheim, 1830
Male adult
SC Georgia, Tsalka distr. 7 km NE of Paravani Lake, 41.540750, 43.879806, h=2100 m, 12-16.VII.2025, I.Uspenskyi leg., L=25 mm
Copyright © Ruslan Panin -
Macrothorax taxon page (26 photos)
Carabus (Macrothorax) rugosus rugosus Fabricius, 1792
Male adult
N Morocco, El Biutz –Taghramt,20.I.2005, Manfred Egger leg.
L=29 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) morbillosus constantinus Kraatz, 1899
Male adult
NW Tunisia; Tebarka, E Sejenane,26.I.2004, Manfred Egger leg.
L=33 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) morbillosus macilentus Lapouge, 1899
Female adult
SE Spain, Murcia reg., P.N. de Calblanque, 1.V.2003, T.Gazurek leg.
L=34 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) rugosus celtibericus Germar, 1824
Female adult
SW-Portugal, Setúbal Distr., Cercal do Alentejo/ Tanganheira vill. 8.XII.2006, T.Gazurek leg.
L=31 m
Copyright © Ruslan Panin
Carabus (Macrothorax) morbillosus constantinus (as galloprovincialis Lapouge, 1910)
Female adult
SE-France, Var, near Le Muy, VIII.1980, Michon leg.
L=31 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) planatus Chaudoir, 1843
Male adult
S Italy, N Sicily, Nebrodi; 33 mm
Copyright © Aleš Sedláček
Carabus (Macrothorax) morbillosus morbillosus Fabricius, 1792 (as cheminorum Deuve, 1988)
Female adult
SE-France, Var, Fréjus env., 26.II.1997, <br /> L=31 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) rugosus laufferi Breuning, 1927
Female adult
C-Spain: Avila, El Hornillo env., 27.V.2017, J. Ferrero leg. 30 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) rugosus levantinus Lauffer, 1905
Male adult
SE-Spain, Murcia, Jumillla, Sierra del Carche, 6.VII.2019, F. Garcia leg., L=29 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) planatus Chaudoir, 1843
Female adult
S Italy, N Sicily, Nebrodi Mts, Portella di Fémmina Morta h=1500 m, 2.IX.1989, V.Aliquo leg. L=33 mm
Copyright © Ruslan Panin -
Meganebrius taxon page (35 photos)
Carabus (Meganebrius) kadoudali (Morvan, 1981)
Male+Female adult
C-Nepal, SW Dhaulagiri, W Jaljala h=3150 m, 28°30'05"N, 83°11'48"E, 23.IX.2012, J. Schmidt leg.
female, 20 mm
Copyright © Ruslan Panin
Carabus (Meganebrius) pseudoharmandi Mandl, 1965
Female adult
E-Nepal, Kosi zone, Khandbari,(1.5km SW of Chichila vill., 27*45’843’’N/ 87*22’473’E, h=1940m), damaged primary forest, 13 .V.2010, B.Brezina & M.Häckel leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Meganebrius) arunensis Heinertz, 1980
Male adult
E-Nepal, Kosi zone, Khandbari, 1.5km SW of Chichila vill., 27*47’175’’N/ 87*23’622’E, h=1880m, damaged primary forest, 11.V.2010, B.Brezina & M.Häckel leg.
L=27 mm
Copyright © Ruslan Panin
Carabus (Meganebrius) quinlani smetanai Häckel, 2011 Paratype
Male+Female adult
C-Nepal:Dhaulagiri Massif:Baglung Distr., (1.5km W of Okhale vill., 28.32’152’’N/ 83.48’673’E, h=2600m) damaged primary forest, 19.V.2010, B.Brezina & M.Häckel leg.
male, 21mm
female, 23 mm
Copyright © Ruslan Panin
Carabus (Meganebrius) quinlani quinlani Mandl, 1965
Male adult
C-Nepal, Gandaki, Manang Distr., Timang env., h~2500-2600m, old open forest, grassland, 3-5.V.2007, Fouqué R. leg. L=19 mm
Copyright © Ruslan Panin
Carabus (Meganebrius) tuberculipennis ganjalaensis Deuve, 2000
Male adult
Nepal, Helambu, GanjaLa Pass S-slope, Dhukpu Kharka h=4100m, 28°03'50"N, 85°34'21"E, 12.VIII.2010, S.Tamang leg., L=21 mm
Copyright © Ruslan Panin -
Megodontoides taxon page (11 photos)
-
Megodontus taxon page (162 photos)
Carabus (Megodontus) violaceus andrzejuscii (as wolffii Dejean, 1826)
Male+Female adult
SW-Romania: Banat, Moldova Noua, Ravenska viil., 21.V.-5.VI.2010, Krause Rene leg. female, 32 mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus fiorii Born, 1901
Male+Female adult
NW-Italy,
Reggio Emilia, Regnano, 5.IX.1996 leg. Cartini A.
female, 30 mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus violaceus Linne, 1758
Male adult
S.Bohemia, Hluboka nad Vltavou, h=380 m, 6.VIII.2004, S. Budka leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus germarii Sturm, 1815
Female adult
Croatia, Ivanščica Mts , Ivanec, 6-13.VI.2003, Krause Rene leg.
L=36 mm
Copyright © Ruslan Panin
Carabus (Megodontus) vietinghoffi bowringi (as lazoensis Obydov, 1999)
Male+Female adult
Russia, Far East, Primorsky reg., S. Sichote-Alin Mts.R., Kavalerovo vill. env.,
5-25.VII.1992, S. Kunitzyn leg.
male - 28 mm, female - 30 mm.
Copyright © Alexandr Safronov
Carabus (Megodontus) caelatus caelatus (as malissorum Apfelbeck, 1919)
Male adult
Montenegro, National park Durmitor; near Zabljak vill., Mt. Sedlo h=2200 m, 6-22.VIII.2010, Franko Sandel leg.
29 мм
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus aurichalceus Kraatz, 1879
Male adult
NC-Spain, Cantabria, Santander, Cueto env.XII.2013 L=28 mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus violaceus trans violaceus purpurascens Fabricius,1787
Male+Female adult
E-Netherlands, Gelderland, Winterswijk Woold env.,15.II.2015, H.de Haan leg.
Copyright © Ruslan Panin
Carabus (Megodontus) persianus javahardehensis Deuve, 2000
Female adult
N-Iran, Mazandaran Prov., Ramsar Co., Javahar Deh env., 16-25.VI.2017, Václav Čutka leg. L=30 mm
Copyright © Ruslan Panin -
Mesocarabus taxon page (150 photos)
Carabus (Mesocarabus) problematicus harcyniae Sturm, 1815 (as obenbergerianus Breuning, 1927)
Male adult
Czech, Bohemia, Plzeň Reg., Domažlice Distr., Babylon, h=500 m, 30.VIII.2008, S.I. Budka leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) dufouri dufouri Dejean, 1829
Male adult
SC Spain, Jaen reg., Monte Javalcuz h=1436 m, 6.X.2010, M.A. Lopez leg.
L=27 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) cantabricus cantabricus Chevrolat, 1840 (loc. typ.)
Male adult
NW-Spain, Pontevedra prov., A Cañiza viil., Alto de Fontefría h=800m, 31.XII.2007, T.Gazurek leg./det.Panin R.<br /> L=27 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) latus cf. molossoides Lassalle, 1984
Male adult
C-Spain, Toledo, Mts de Toledo, Sierra del Castañar Rng, Puerto del Milagro env, h=920m, 7.II.2009, T.Gazurek leg./det.R. Panin.
L=24 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826
Male adult
CW-Spain: Salamanca, Brincones env., h=750m, X.2003<br /> L=25 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) problematicus harcyniae (as belgicus Lapouge, 1913)
Male adult
Luxembourg, Bollendorf-Pont env., 3.III.1999 /det.R Panin
L=24 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) lusitanicus gougeleti (as valderepisensis Branes, 1988)
Male adult
Spain, Jaen prov., Despeñaperros, Arroyo de los Arcos, 38.381847, -3.515651, 1-2.05.2014, Anichtchenko A. leg. & det.; 18 mm
Copyright © Alexander Anichtchenko
Carabus (Mesocarabus) dufouri Dejean, 1829
Female adult
S Spain, Granada, Sierra Arana, Píñar env., 14.X.2018, F. Garcia leg. L=27 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) lusitanicus lusitanicus (as vieirae Oliveira, 1875)
Male adult
CW-Portugal: Leiria distr., near Marinha Grande, 39°45'57.8"N 8°58'10.3"W, h-79 m, 19.IX.2016, D. Prunir leg., L=24 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) riffensis Fairmaire, 1875
Endophallus
N-Morocco, Riff Mts h=1650 m, 26 km E of Bab Berred
Copyright © Ruslan Panin -
Microplectes taxon page (14 photos)
Carabus (Microplectes) riedeli Menetries, 1832
Male adult
N-Ossetia,
Khazniaush Pass, h=2500-2700m, 9.VI-31.VII.1994, Zamotajlov A.leg.
male, 17mm
Copyright © Ruslan Panin
Carabus (Microplectes) convallium convallium Starck, 1889
Male adult
male, 19mm
Copyright © Boleslav Brezina
Carabus (Microplectes) convallium convallium Starck, 1889
Male+Female adult
female:NW-Caucasus, Russia,
Krasnodar reg., Apsheron Distr., Lagonaki Mt., near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominyh leg.
male:Karachaevo-Cherkessia,the right source of the river Beskess, h=2100m, 10.VIII.1997, I.A. Solodovnikov leg.
female, 21mm/male, 19mm
Copyright © Ruslan Panin
Carabus (Microplectes) convallium argonautarum Semenov, 1898
Male adult
male, 18mm
Copyright © Boleslav Brezina -
Mimocarabus taxon page (11 photos)
Carabus (Mimocarabus) maurus maurus Adams, 1817
Male+Female adult
Armenia, Ashotsk distr., Tsoghamarg vill., 24-27.V.1999 P.V.Kazaryan leg. female, 20mm/male,18mm
Copyright © Ruslan Panin
Carabus (Mimocarabus) pumilio Kuster, 1846
Male adult
S-Armenia, Vayots Dzor Region, E Ughedzor vill., h=2630 m, 9.VI.2014, L=16 mm
Copyright © Ruslan Panin
Carabus (Mimocarabus) maurus maurus (Adams, 1817)
Male adult
E Georgia, Kakheti, near Patardzeuli vill., h=750-850 m, 10.IV.2015, E. Rutjan leg., L=22 mm
Copyright © Ruslan Panin -
Morphocarabus taxon page (437 photos)
Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
Male adult
W Sayan Mts., Kazhyr-Suk riv., 80 km W Oleniya<br /> Rechka, W part of Oiskii mt. rng., 30.6.-7.7.1999<br /> 2100 m, Brinev A. leg.<br /> male, 19 mm
Copyright © Alexander Anichtchenko
Carabus (Rhigoidocarabus) zhubajie laojunensis Kleinfeld, 1999 Holotype
Male adult
Copyright © Frank Kleinfeld
Carabus (Morphocarabus) rothi hampei Kuster, 1846 (as dacicus Csiki, 1906)
Female adult
NW Romania , Bistrita Nasaud, Beclean env., 17.VI.2009, S.Budka leg.
female, 31 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906 loc.typ.
Male+Female adult
W-UKRAINE,<br /> Transcarpathian region, Uzhgorod distr., h=250 m, 8.VII-25.VIII.2011<br /> Panin R. leg./det.<br /> female, 33 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as vertesensis Retezár, 1974)
Male adult
NW Hungary: Fejer: Vértes Mt Rng, Tatabanya, 28.VII-2.VIII.1996
L=29 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Female adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 20.V-25.VI.2008, Panin R. leg. L= 31 mm.
Copyright © Ruslan Panin
Carabus (Morphocarabus) henningi henningi Fischer, 1817
Male adult
Russia, S Siberia, NW Buryatia, E Sayans Mts., Tunka Range, near Mondy vill., h=2400 m, alpine tundra, 1-15.VI.2001, Mishustin R. leg./det. R.Panin
Copyright © Ruslan Panin
Carabus (Morphocarabus) spasskianus putus Motschulsky, 1846 (loc.typ.)
Male adult
Russia, W Altay, 18 km N of Zmeinogorsk, 23.VI.1998, A.& N. Menstschikovy leg. L=23 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Male adult
SE Poland, N Carpathians, Subcarpathia Prov., National Park Magura, VII.2016, M.Ranocha leg./ det. R. Panin, L=30 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) scheidleri bergeri Peeters & I.Rapuzzi, 2025
Male adult
W-Austria, Tyrol, Ötztal Alps, Kaunertal, Feichten, h=1281 m, 47.035200, 10.748700, 1.VIII.2025, M.Egger leg., L=26 mm
Copyright © Ruslan Panin -
Morvanicarabus taxon page (2 photos)
Carabus (Morvanicarabus) scheibei scheibei Eidam, 1937
Female adult
NE-Afghanistan, Hindu Kush, Nuristan Prov., near Parun, 35°19'N 70°52'E h=2900 m, Pine forest, VI.2022, local collector/det. Kleinfeld, L= 24 mm
Copyright © Ruslan Panin
Carabus (Morvanicarabus) scheibei swatensis Heinertz, 1979
Male adult
NW-Pakistan, Kumrat Valley Dir h=2400 m,1-3.VI.2025,Ch.Reuter leg./det., l=18,5 mm
Copyright © Ruslan Panin -
Neoplectes taxon page (286 photos)
Carabus (Neoplectes) daphnis askhicus Belousov, 1992 Paratype
Male adult
SC-Caucasus, W Georgia, valley of riv. Chkhorodukhu, h=1000-1200m, 14.V.-6.VII.1991, Zamotajlov A.S & Miroshnikov leg.
male, 29 mm
photo & collection Panin R.
Copyright © Ruslan Panin
Carabus (Neoplectes) edithae exedithae Gottwald, 1985
Male adult
Karachaevo-Cherkessia, low cource of Daut river, 16-20.V.2009 Anichtchenko A. leg
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) juenthneri avadcharensis Kurnakov, 1970 (loc.typ.)
Male+Female adult
Abkhasia, near Avadhara, Site Pyv h~1850m, 43.490730 40.686433, 1.VI-12.VII.2009, Zamotajlov A.S., Bondarenko A.S. leg.
L=27mm
Copyright © Ruslan Panin
Carabus (Neoplectes) kratkyi felicitanus Reitter, 1893 (as porcellus)
Male adult
NW Caucasus, Resp. Adygea, Mt.Tybga, 28.VI.1997
male,20mm
Copyright © Ruslan Panin
Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
Male adult
NW-Caucasus, Russia,
Krasnodar reg., Apsheron Distr., Lagonaki Mt. ,near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominykh leg.
male, 28mm
Copyright © Ruslan Panin
Carabus (Neoplectes) reitteri reitteri Retowski, 1885
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Carabus (Neoplectes) reitteri gagrinus Starck, 1894
Female adult
ABKHASIA, Gagra distr., Mamdsyshkha Mt., near Gagra, Fagus forest + karst, h = 1300-1330 m, biotope N 23, 43º18’186" N / 040º18’44" E, 12.05-05.07.2014, Leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Neoplectes) starckianus theseus Briansky, 1910
Female adult
Russia, Krasnodar Territory, Tuapse district, env. vill. Psebe, left bank of the Psebe River, hornbeam + oak + walnut, h = 146 m, 03.05-01.07.2014, leg. I.A. Solodovnikov, S.V. Solodovnikova, S.M. Pavlyuchuk
Copyright © Igor Solodovnikov
Carabus (Neoplectes) starckianus adelaidae Starck, 1889
Female adult
Russia, Krasnodar area, Adler, Ader distr., right bank vall. Riv. Msymta, near vill. Chvesipse, h = 800 m, 06.VII.1999, B. Saratovskikh leg.
Copyright © Igor Solodovnikov
Carabus (Neoplectes) starckianus adelaidae Starck, 1889
Female adult
Russia, Krasnodar area, Adler, Ader distr., right bank vall. Riv. Msymta, near vill. Chvesipse (Fagus+Quercus+Castanea forest) [Sochi national park], h = 720 m, 03.V-08.VII.2016, I.A. Solodovnikov, S.V.Solodovnikova, V.M.Kotsur leg.
Copyright © Igor Solodovnikov -
Neoplesius taxon page (34 photos)
Carabus (Eocechenus) kaznakovi nangqenicus Deuve, 1995 Paratype
Male adult
Copyright © Boleslav Brezina -
Nesaeocarabus taxon page (9 photos)
Carabus (Nesaeocarabus) abbreviatus Brulle, 1835
Male adult
Canarias Isle, NC Tenerife; 22mm
Copyright © Aleš Sedláček
Carabus (Nesaeocarabus) faustus cabrerai Enderlein, 1929
Male adult
Spain, Canarias Isl.,Teneriffe: NW-part:Teno Massif, Erjos env., h=1000m, 18-21.III.2013, V.Slovak leg.
L=23 mm
Copyright © Ruslan Panin -
Ohomopterus taxon page (38 photos)
Carabus (Ohomopterus) japonicus japonicus Motschulsky, 1857
Male adult
S Japan, Kyushu, Kumamoto Pref., Izumi viil., h=600 m,VII.2001, S. Remond leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Ohomopterus) yamato kitai (Imura, 2004)
Male adult
Japan, S Honshu, Mie Pref., Hakusan, VII.2001, S. Remond leg.<br /> L=21 mm
Copyright © Ruslan Panin
Carabus (Ohomopterus) dehaanii dehaanii Chaudoir, 1848
Female adult
Japan, S Honshu, Hyogo Pref., Nishiwaki city, 20.VI.2010, Shinichi S. leg., L=37 mm
Copyright © Ruslan Panin
Carabus (Ohomopterus) dehaanii dehaanii Chaudoir, 1848
Male adult
Japan, S Honshu, Mie Pref., Inabe city, 21.II.2021, Shinichi S. leg., L=33 mm
Copyright © Ruslan Panin
Carabus (Ohomopterus) japonicus japonicus Motschulsky, 1857
Male adult
MT. Narabara-san, Ryuokakami Tamagawa town, Imabari city, Ehime pref. Japan 20.6.2015 Y. Amano (cAA). det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko
Carabus (Ohomopterus) japonicus japonicus Motschulsky, 1857
Female adult
Fukuoka pref., Omuta city, Hagio, 16.1.2012, S. Nishimoto leg. (cAA)
Copyright © Alexander Anichtchenko
Carabus (Ohomopterus) lewisianus awakazusanus Ishikawa, 1981
Male adult
Mt. Iyogatake 280m, Tomiyama town, Awa Co., Chiba pref, 4.2.1996 K. Watanabe leg. (cAA). det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko
Carabus (Ohomopterus) albrechti Morawitz, 1862
Male adult
Horoman, Samani town, Samani Co., Hokkaido, 16.6.2015 T. Nakamura leg. (cAA). det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko
Carabus (Ohomopterus) insulicola nishikawai (Ishikawa, 1966)
Male adult
Yamada, Isumi city, Chiba pref. Japan 20.5.2007 K. Osozawa leg. (cAA). 28mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko
Carabus (Ohomopterus) insulicola insulicola Chaudoir, 1869
Male adult
Japan, Kiyose, Tokyo 20.1.1961 (cAA). 29 mm
Copyright © Alexander Anichtchenko -
Ophiocarabus taxon page (34 photos)
Carabus (Ophiocarabus) regeli obscurior Semenov, 1888
Male+Female adult
Kyrgyzstan, NE Tian-Shan, E. Terskey Alatau Mts. R.,
Kok-kia River, h=2800 м.
7-8.VI.1997 М. Danilevsky leg.
Body lenght 15-18 mm.
Copyright © Alexandr Safronov
Carabus (Ophiocarabus) ballionis Kraatz, 1879
Male adult
NW-China, Xinjiang prov., 40 km N Narat, h=1500, 12-12.VI.2004, Benes & Kaiser leg.
L=22 mm
Copyright © Ruslan Panin
Carabus (Ophiocarabus) successor koeksuensis Kabak, 2013 Paratype
Male adult
China, Xinjiang, Koeksu Bas., Tshon-Aidak, SSE Tekes, 42* 32' 47'' N / 82* 05' 56'' E -- 42* 32' 51'' N / 82* 05' 36'' E, 3330-3550 m, 11.07.2012, Kabak I.I. leg.
Copyright © Ilya Kabak
Carabus (Ophiocarabus) successor successor Reitter, 1896
Male adult
SE-Kazakhstan, Narynkol distr., 4 km SW Bayankol, h~2400-2500 m, 2-14.VI.2007, Shumov leg. L=17.5 mm
Copyright © Ruslan Panin
Carabus (Ophiocarabus) successor successor Reitter, 1896
Female adult
SE-Kazakhstan, Narynkol distr., 4 km SW Bayankol, h~2400-2500 m, 2-14.VI.2007, Shumov leg. L=17 mm
Copyright © Ruslan Panin -
Oreocarabus taxon page (61 photos)
Carabus (Pachystus) hortensis neumeyeri Schaum, 1856
Female adult
Italy, Basilicata, Serra Calvello (PZ)
1400 m, 7.2003 F. Angelini leg.
female, 25 mm
Copyright © Alexander Anichtchenko
Carabus (Oreocarabus) carinthiacus Sturm, 1815
Male adult
NE-Italy, Bolzano, Brenner, Zirog h=2000 m, 26.VII.1992, Manfred Egger leg.
L=20 mm
Copyright © Ruslan Panin
Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847
Male adult
WC Spain, Cáceres, Hervás, h=1000m, V.2007, Pascal Stefani leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Oreocarabus) carinthiacus Sturm, 1815
Female adult
NW Slovenia, Trnovski Gozd; 24 mm
Copyright © Aleš Sedláček
Carabus (Oreocarabus) ghiliani La Ferte-Senectere, 1847
Female adult
C-Spain: Sierra de Guadarrama, Puerto de Cotos (Madrid), 20.IX.2018, F.Rodriguez leg., L=22 mm
Copyright © Ruslan Panin
Carabus (Oreocarabus) luetgensi pseudosteuarti Lapouge, 1924
Female adult
Portugal, Guarda, Sierra de Estrela h=800 m, 26.VI-4.VIII.2015, L=26 mm
Copyright © Ruslan Panin
Carabus (Oreocarabus) luetgensi pseudosteuarti Lapouge, 1924
Male adult
Portugal, Guarda, Sierra de Estrela h=800 m, 26.VI-4.VIII.2015, L=24 mm
Copyright © Ruslan Panin
Carabus (Pachystus) hortensis hortensis Linne, 1758
Female adult
Natural hybrid Carabus hortensis X Carabus granulatus. Belarus, Minsk reg., 8,7 km of Borisov, near vill. Lyubatovstchina, pine forest (Pinetum pleuroziosum), h = 213 m, 54.31934° N, 28.511486° E, 20-27.09.2018, leg. Klychenok Snezhana
Copyright © Igor Solodovnikov
Carabus (Hypsocarabus) mikhaili mikhaili Deuve & Mourzine, 1997
Female adult
L=23 mm
Copyright © Ruslan Panin
Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847
Female adult
Spain, Sierra de Guadarrama, Puerto de la Quesera, 1720m, 41.214640, -3.430896, Anichtchenko A. leg. 10.X.2025 (cAA).
Copyright © Alexander Anichtchenko -
Orinocarabus taxon page (68 photos)
Carabus (Orinocarabus) castanopterus A. & J.B. Villa, 1833
Female adult
NC-Italy - S-Switzerland, Val Colla – Gazzirola, 22.VI.1999, C. Sola leg.
L=21mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) sylvestris sylvestris Panzer, 1793 (as slabai Niedl, 1968)
Female adult
Czech, WS Bohemia,
Plzen Reg., Domazlice Distr., Babylon, h=500m, 30.VIII.2008, S.I. Budka leg.
L=24mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) fairmairei heteromorphus Daniel, 1896
Female adult
NW-Italy, Aosta region, Champorcher env., Mt.Dondenaz, VIII.2008, V.Monzini leg.
L=19 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) fairmairei fenestrellanus Beuthin, 1899
Male adult
NW-Italy, Piemonte prov., Cottian Alps, Mt Albergian h=2450 m.<br /> L=20 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) bertolinii Kraatz, 1878
Male adult
NE-Italy, Dolomites-Alps, Trentino prov., Rolle Pass env., Malga Fosse h=2000m, 8-28.VI.2013, Manfred Egger leg.
L=19 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) alpestris dolomitanus Mandl, 1956
Male adult
NE-Italy, South Tyrol, Dolomites-Alps, 500 m left of Würzjoch, Gömaalm - at the foot Mt.Peitlerkofel h=2100m, 28.VII-17.VIII.2013, Manfred Egger leg.
l=18 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) concolor lepontinus Born, 1908
Male adult
NW-Italy, Piemonte (VB), Lepontine Alps, Mt. Zeda , 6.VII.1998, L Diotti leg., L=20 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) concolor concolor Fabricius, 1792
Female adult
NW-Italy, Breuil-Cervinia h~2500 m, 2.IX.1971, Georgeon leg. L=21 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) sylvestris transsylvanicus Dejean, 1826
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=21 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) bertolinii Kraatz, 1878
Male adult
NE-Italy, Trentino prov. Dolomites Alps, Plateau Pale di San Martino h=2500 m, 17-19.VIII.2019, M. Egger leg.
Copyright © Ruslan Panin -
Oxycarabus taxon page (8 photos)
Carabus (Oxycarabus) saphyrinus pseudosaphyrinus Schweiger, 1964 (as rabaronianus)
Male adult
Anatolia, Kastamonu, Yaraligoz verso Catalzeitin,
VI.2001 Sola Claudio leg.
male, 28 mm
Copyright © Alexander Anichtchenko
Carabus (Oxycarabus) saphyrinus pirithous Reitter, 1896
Male adult
NW-Turkey,Istanbul (Asian part),Polonezköy env., 2.VI.2009. S. Dacatra leg. L=27 mm
Copyright © Ruslan Panin -
Pachycarabus taxon page (27 photos)
Carabus (Pachycarabus) koenigi Ganglbauer, 1886
Male adult
NW Caucasus, S Russia, Adygea Rep., Lagonaki plateau, Site Murzikal h=1800-2000m, 24.V-18.IX.2010, D.Fominykh leg.
L= 22 mm
Copyright © Ruslan Panin
Carabus (Pachycarabus) roseri soganliensis Heinz & Korge, 1964
Male+Female adult
Turkey, Rize prov., Ikizdere distr., Rize Daglari,
47 km to SE from lyidere, Sivrikaya Koyu, h=2025 m.
30.VI-5.VII.2011, A. Vlasenko leg.
male - 23 mm, female - 24 mm.
Copyright © Alexandr Safronov
Carabus (Pachycarabus) imitator imitator Reitter, 1883
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Carabus (Pachycarabus) swaneticus kvirensis Belousov, 1992
Male adult
SC-Caucasus: NW-Georgia, Mingrelia, Egrissky Mt.Rng., h~1800-2000 m, 23-26.VII.1990, I.Retezár leg. L=20 mm
Copyright © Ruslan Panin
Carabus (Pachycarabus) roseri roseri Faldermann, 1835
Female adult
Georgia, Lesser Caucasus, Imereti prov., N slope Meskheti Mt Rng, Zekari Pass, h~1950 m, 24.VII.2021, E.Rujan leg., L=22 mm
Copyright © Ruslan Panin
Carabus (Pachycarabus) swaneticus swaneticus Reitter, 1883
Endophallus
Georgia, Caucasus, Svaneti Range E part, Latpari Pass h=2700-2800 m,
Copyright © Ruslan Panin
Carabus (Pachycarabus) swaneticus swaneticus Reitter, 1883
Male adult
Georgia , Caucasus, E part Egrisi Mt Range, Mt. Tsekuri N slope h=2370 m, 18-20.VI..2025, I.Uspenskyi leg. L=19 mm
Copyright © Ruslan Panin
Carabus (Pachycarabus) staehlini rionicus Belousov, 1992
Female adult
Georgia, Caucasus, S of Sasashi, Lechkhumi Range N sope h=2430m, alpine meadows 242.759167, 42.973889, 20-21.VII.2025, I.Uspenskyi leg./det. R. Panin, L=20.5 mm
Copyright © Ruslan Panin
Carabus (Pachycarabus) koenigi Ganglbauer, 1886
Male adult
NW Caucasus, Republic Adygea, Mt.Tybga h=2100 m, 28.VI.2006, Tret’yakov Yu. leg., L=21 mm
Copyright © Ruslan Panin -
Pachystus taxon page (45 photos)
Carabus (Pachystus) hungaricus mingens Quensel, 1806
Male adult
male, 31 mm
SW-Russia: Stavropol Krai, near Stavropol city , Russkiy Les, 11.VI-28.IX.2010, Pavluchuk S. leg.
Copyright © Ruslan Panin
Carabus (Pachystus) hungaricus cribellatus Adams, 1812
Male adult
E Siberia, Russia, Krasnoyarsk Reg., near Krasnoyarsk, forest-steppe,1.VII-20.VIII,2011,M.Ivanov leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Pachystus) hungaricus scythus Motschulsky, 1847
Female adult
S Ukraine, Zaporizhzhya reg., Khortytsa island, 16-20.VII.2011, M. Mulenko leg., det. Panin R.
L=28 mm
Copyright © Ruslan Panin
Carabus (Pachystus) hungaricus hungaricus Fabricius, 1792
Male adult
Czech, S Moravia, Pouzdrany, 8,VIII.2005, V. Slovak lgt.
L=27 mm
Copyright © Ruslan Panin
Carabus (Pachystus) cribratus staneki Sterba, 1931
Male adult
NC Turkey, Küre Mts., 60km N Kastamonu; 25mm
Copyright © Aleš Sedláček
Carabus (Pachystus) cribratus staneki Sterba, 1931
Female adult
NC Turkey, Küre Mts., 60km N Kastamonu; 27mm
Copyright © Aleš Sedláček
Carabus (Pachystus) cavernosus variolatus Costa, 1839
Male adult
C-Italy:Abruzzo reg., Gran Sasso d'Italia Massif, h~1600 m, VIII.1997, Seichert leg. L=26 mm
Copyright © Ruslan Panin
Carabus (Pachystus) glabratus gibbosus Heyden, 1866
Male adult
N Croatia, Ivanščica Mts, Ivanec, 4-18.VI.2011, Krause Rene leg., L=32 mm
Copyright © Ruslan Panin
Carabus (Pachystus) cribratus cribratus Quensel, 1806
Male adult
NW Armenia, Amasia, (Shirak prov.), 20-24.VIII.1998, Kazaryan P.V. leg.
Copyright © Ruslan Panin -
Pagocarabus taxon page (49 photos)
Carabus (Pagocarabus) crassesculptus diruptus Morawitz, 1886
Male adult
China, Gansu, Umg. Xiahe 3000-3200m
24-26.6.1993 Heinz leg.
20 mm
Copyright © Alexander Anichtchenko
Carabus (Pagocarabus) crassesculptus eleganticolor Deuve, 1997
Female adult
S Gansu, Min Shan, 5-20 km S Cagan, 2300-3200m 7-20.7.2019 V. Patrikeev leg. (cAA). det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko -
Pantophyrtus taxon page (20 photos)
Carabus (Pantophyrtus) turcomanorum turcomanorum Thieme, 1881
Male+Female adult
Kyrgyzstan, Kyrgyzskyi Mts. R., Uzun Bulak Gorge, h=1800 ms.
01-09.VI.2002 loc. coll.
male - 28 mm, female - 33 mm.
Copyright © Alexandr Safronov
Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993
Male adult
Uzbekistan, Chatkal Mt. rng., Irtash, 41°10'25.7"N 70°16'20.4"E, 5-7.8.2021, 2400m, A. Anichtchenko leg. (cAA). 27mm
Copyright © Alexander Anichtchenko
Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993
Female adult
Uzbekistan, Chatkal Mt. rng., Irtash, 41°10'25.7"N 70°16'20.4"E, 5-7.8.2021, 2400m, A. Anichtchenko leg. (cAA). 30 mm
Copyright © Alexander Anichtchenko -
Piocarabus taxon page (5 photos)
Carabus (Piocarabus) titanus Breuning, 1933
Male adult
C-China Sichuan prov., Nanping, h=1500m, VI.2000, S. Remond leg.<br /> L=34 mm
Copyright © Ruslan Panin
Carabus (Piocarabus) nanwutai Kleinfeld, Korell & Wrase, 1996
Male adult
China, W-Shaanxi, Taibai Shan Mts, btw. Houzhenzi/Ba Xian Tai, nr.Nan Tian Men h=3200m, 33°55'17"N/107°46'53"E, 25-30.VII.2011, Li Wuyang leg.<br /> L=19 mm
Copyright © Ruslan Panin
Carabus (Piocarabus) sui Imura & Zhou, 1998
Female adult
CE-China, Henan, Mt Wangwu, V.2007, local collector. L= 33 mm
Copyright © Ruslan Panin
Carabus (Piocarabus) vladimirskyi Dejean & Boisduval, 1830
Male adult
Buryatia, Kyahta distr. [Khyagh], 5 km SE Chikoi vill., 16.8.1998, Anichtchenko A. leg.
Copyright © Alexander Anichtchenko
Carabus (Piocarabus) titanus Breuning, 1933
Female adult
China Sichuan prov., Nanping, h=1500m, VI.2000, S. Remond leg.
L=38 mm
Copyright © Ruslan Panin -
Platycarabus taxon page (37 photos)
Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Male adult
W.Ukraine, Transcarpathian Reg., Rakhov Distr., Chornogora range., h=2020m. Panin R. leg. 28.VI-9.VIII.2008, male, 19 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) irregularis montandoni Buysson, 1882
Male adult
W-Ukraine, NW Podolian,
Lviv Reg., Zhydachiv distr., near Vybranivka, under the bark of oak tree, in the forest,12.X.2007 Panin R. leg
L= 22 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Male adult
W.Ukraine, Transcarpathian Reg., Rakhov Distr., Chornogora Mt range h=2020m alpine zone, Panin R. leg. 28.VI-9.VIII.2008, male 19 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) creutzeri kircheri Germar, 1824
Male+Female adult
NE-Italy, Veneto Reg., Belluno, Forcella Lavardet h=2000m, VII.2002, I.Rapuzzi leg.
female, 24 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii fabricii (as fassatii Smetana, 1955)
Male adult
C-Slovakia:Nizke Tatry Mts., h=1750 m,.9.VI.2007, V.Slovak leg.
L=20 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii fabricii (as poloniensis Géhin, 1876)
Female adult
N-Slovakia, Orava Beskids Mts, Mt Babia Góra h=1720 m, 5.VII.20.VIII.2010, Rudolf Kmeco leg.
L=22 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Female adult
N- Romania, E-Carpathians, Rodna Mts, h=2000m, 7-8.VI.2012, Cartuz Z.leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Female adult
W Ukraine, NE Carpathians, Transcarpathian Region, Chornogora Mt Range, h=2000 m, alpin zone, 28.VII-11.IX.2014, Panin R. leg. L=22 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) irregularis irregularis (as jurassicus Born, 1911)
Male+Female adult
France, Ain (01), Jura Mts, Grand Colombier, Forêt d' Arvières h~1250 m, 25.X.2014, Ya. Vasseur leg., L=23 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) irregularis ramanus Sokolar, 1909
Female adult
NW Croatia, Ivanščica Mts , Ivanec, 4-18.VI.2011, Krause Rene leg., L=33 mm
Copyright © Ruslan Panin -
Procerus taxon page (54 photos)
Carabus (Procerus) gigas gigas Creutzer, 1799
Male+Female adult
N Croatia, Ivanščica Mts , Ivanec, 4-18.VI.2011, Krause Rene leg.
female, 54 mm
Copyright © Ruslan Panin
Carabus (Procerus) scabrosus caucasicus Adams, 1817
Male adult
N Caucasus,
Stavropol Krai, Pyatigorsk, Mt.Mashuk, VI.2007 Tikhonov Yu. leg.
44mm
Copyright © Ruslan Panin
Carabus (Procerus) gigas duponcheli Dejean, 1831 (as kusdasi Mandl, 1966)
Male adult
S-Greece: Peloponnes, Mt.Chelmos, 20.V.2007.
L=46 mm
Copyright © Ruslan Panin -
Procrustes taxon page (51 photos)
Carabus (Procrustes) chevrolati thirki Chaudoir, 1857
Male adult
NE Turkey, Trabzon, Zigana Pass, h~2000m, V. Slovak leg.
L=27 mm
Copyright © Ruslan Panin
Carabus (Procrustes) coriaceus hopffgarteni (as emgei Ganglbauer, 1888)
Male adult
CE-Greece: Larissa reg. Stomio, Ossa Mt. 6-17.VII.2011, Krause Rene leg.
L=32 mm
Copyright © Ruslan Panin
Carabus (Procrustes) anatolicus anatolicus Chaudoir, 1857
Male adult
Cyprus, Troodos, 17.2.1V.2010,. Manfred Egger leg.
L=27 mm
Copyright © Ruslan Panin
Carabus (Procrustes) coriaceus mopsucrenae Peyron, 1858
Male adult
SC-Turkey, Adana Prov., Doganbeyli, 8.V.2001, Snizhek M. leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Procrustes) talyschensis talyschensis Menetries, 1832
Female adult
Azerbaijan, Talysh Mts., Zuvand, Kalvaz vill., h=1650 m, 15.VII.2003, Dolin V.G. leg.
L=32 mm
Copyright © Ruslan Panin
Carabus (Procrustes) piochardi praestigiator Morawitz, 1886
Male adult
C-Lebanon, Byblos /Jbeil Distr., Kfardebian env., h=1200m, XI.2014, C. Reuter leg. L=31 mm
Copyright © Ruslan Panin
Carabus (Procrustes) coriaceus ressli Mandl, 1961
Male adult
SC-Turkey: Hatay Prov, Dörtyol, Nur Mts, Topaktaş h=900-1200 m, 12.V.2005, D. Prunier leg.
Copyright © Ruslan Panin
Carabus (Procrustes) coriaceus ressli Mandl, 1961
Female adult
SC-Turkey: Hatay Prov, Dörtyol, Nur Mts, Topaktaş h=900-1200 m, 12.V.2005, D. Prunier leg. L=35 mm
Copyright © Ruslan Panin
Carabus (Procrustes) chevrolati internatus Heinz & Korge, 1964
Female adult
Turkey, Bolu, Gerede, Esentepe 1350m, 1.7.1994, leg. K. Staven
Copyright © Alexander Anichtchenko -
Pseudocoptolabrus taxon page (42 photos)
Carabus (Pseudocoptolabrus) burmanensis Breuning, 1932
Male adult
male, 23mm
Copyright © Boleslav Brezina
Carabus (Pseudocoptolabrus) businskyi Deuve, 1990
Male adult
CH, NW-Yunnan (Lijiang) Yulongxue Shan, E-slope
27:10:50N/100:12:26E/ 4400m, alpine meadows 12.VII.2000
K. & B. Brezina, M.Brychcin, M. Hackel
Copyright © Boleslav Brezina -
Pseudocranion taxon page (14 photos)
Carabus (Pseudocranion) sackeni sackeni Semenov, 1898
Male adult
male, 23mm
Copyright © Boleslav Brezina
Carabus (Pseudocranion) sackeni remondianus Deuve, 1996 Paratype
Female adult
Museum of Institute of Systematic Biology (Daugavpils, Latvia)
Copyright © Alexander Anichtchenko
Carabus (Pseudocranion) lazikouensis lazikouensis (as kundaensis Deuve, 2003) Paratype
Female adult
C-China, S-Gansu: SSW of Minxian: Qilian Shan: W Kunda, 34:10N/103:47E, h=2800-3000m, forest, 7.VII.2002, Belousov & Kabak leg.
L=24 mm
Copyright © Ruslan Panin
Carabus (Pseudocranion) taibaishanicus taibaishanicus Deuve, 1989
Female adult
China, W-Shaanxi, Taibai Shan Mts, btw. Houzhenzi/Ba Xian Tai, nr. Nan Tian Men h=3200m, 33°55'17"N/ 107°46'53"E, 25-30.VII.2011, Li Wuyang leg. L=29 mm
Copyright © Ruslan Panin -
Relictocarabus taxon page (1 photos)
Carabus (Relictocarabus) meurguesianus Ledoux, 1990
Male adult
Image from molbiol.ru
Copyright © - - - -
Rhigocarabus taxon page (164 photos)
Carabus (Rhigocarabus) gentleman Brezina & Hackel, 2004 Holotype
Male adult
male, 19mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) buddaicus gansuicus Deuve, 1991
Male adult
China, NC-Sichuan (ZOIGE), LANGMUSI (=DOGCANGLHAMO), 34:03:48N/102:38:08E, h=4100m, alpine meadows, screes, 16-18.VII.1999, B.Brezina leg.
L=20 mm
Copyright © Ruslan Panin
Carabus (Rhigocarabus) indigestus confucius Breuning, 1933
Female adult
C-China, E Tibet,W-Sichuan prov., Kangding, Zheduo Pass h=4300m 30.VI.2005 , V.Benes leg,
L=17 mm
Copyright © Ruslan Panin
Carabus (Rhigocarabus) ladygini degensis Deuve, 1989
Male adult
C-China, NW- Sichuan prov., Garzê Tibetan AP, Pass Garzê – Sêrtar, 19.VII.2007, V.Benes leg.
L=17 mm
Copyright © Ruslan Panin
Carabus (Rhigocarabus) buddaicus linxiaicus Deuve, 1995
Male adult
China, S-Gansu (LABRANG):, valley.E of Ponggartang, 35:12-13N/102:54-55E, h=3000-3700m, alpine meadows, coniferous forests,10.-14.VII.1999, B.Brezina leg. L=20 mm
Copyright © Ruslan Panin
Carabus (Rhigocarabus) morawitzianus morawitzianus Semenov, 1887
Male adult
C China, Qinghai, Zadoi County, Toramarkog h~4600 m, VI.2001, P. Cavazzuti leg. L=17 mm
Copyright © Ruslan Panin
Carabus (Rhigocarabus) turnai turnai Deuve, 1994
Male adult
China, Tibet, W-Sichuan, pass btw Sêrtar / Luhuo h=4200 m, 28.VI.2011, V.Patrikeev leg.L=19 mm
Copyright © Ruslan Panin
Carabus (Rhigocarabus) turnai turnai Deuve, 1994
Female adult
China, Tibet, W-Sichuan, pass btw Sêrtar / Luhuo h=4200 m, 28.VI.2011, V.Patrikeev leg. L=21 mm
Copyright © Ruslan Panin
Carabus (Rhigocarabus) morawitzianus morawitzianus Semenov, 1887
Male adult
Qinghai, Anemaqen mts pass, 20 km N of Maduo town, 4500 m, 6.7.2012, V. Patrikeev leg.; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko
Carabus (Rhigocarabus) rhododendron Deuve & Imura, 1991
Male adult
China, N Yunnan: Baimaxue Shan, S of Deqen, h=4300m, 28°20'N 99°03'E, 16-20.VI.2016, C.Reuter leg., L=18 mm
Copyright © Ruslan Panin -
Scambocarabus taxon page (6 photos)
Carabus (Scambocarabus) kruberi laobeiensis Deuve & Li, 2000
Male adult
NE-China: Liaoning prov., Fengcheng City, Miaoyang, 12-17.VI.2011
L=27 mm
Copyright © Ruslan Panin -
Semnocarabus taxon page (29 photos)
Carabus (Semnocarabus) regulus regulus Dohrn, 1882
Male adult
photo M. Smirnov from www.zin.ru
Copyright © Maksim Smirnov
Carabus (Semnocarabus) carbonicolor carbonicolor Morawitz, 1886
Male+Female adult
Kyrgyzstan, NC Tianshan, Kyrgyz Mt. R., Tuyuk riv., h=2000 м.
25-VI-1997 М. Danilevsky leg.
male - 20 mm, female - 21 mm.
Copyright © Alexandr Safronov
Carabus (Semnocarabus) erosus kuldshaensis Ballion, 1878
Male adult
S-Kazakhstan:Ili River valley,:near Bakanas,3-5.V.2002
L=21 mm
Copyright © Ruslan Panin
Carabus (Semnocarabus) carbonicolor callosus Semenov, 1896
Male+Female adult
KAZAKHSTAN,
Bolshoe Almaatinskoe Lake,30.VI.1992/ det.Panin R.
female 19mm
Copyright © Ruslan Panin
Carabus (Semnocarabus) carbonicolor korolkowi Semenov, 1896
Male adult
NC-Kyrgyzstan, Kyrgyz Mt.Rng , 10 km SW of Bishkek, Chon-Aryk env, h=1100m, 24.V.1998, /det. Panin R.
22 mm
Copyright © Ruslan Panin
Carabus (Semnocarabus) erosus corax Semenov, 1896
Endophallus
SE-Kazakhstan, Almaty reg., C Trans-Ili Alatau N-slopes, Kaskalen river Valley, Izvestkovaya vill., h=1700 m, 6.V.2015, D.Kovalchuk leg./det. R.Panin,
Copyright © Ruslan Panin
Carabus (Semnocarabus) carbonicolor carbonicolor Morawitz, 1886
Female adult
SE-Kazakhstan, Almaty reg., Khan Tengri Massif, Narynkol env.h=1800 m, 30-31.-V.2007, Shumov leg., L=21 mm
Copyright © Ruslan Panin
Carabus (Semnocarabus) erosus erosus Motschulsky, 1865
Endophallus
SE-Kazakhstan, 70 km E of Almaty, E Trans-Ili Alatau N-slopes, valley Assy (Asy) riv. Assy plateau h=1700 m, 12.V.2015
Copyright © Ruslan Panin
Carabus (Semnocarabus) erosus prope erosus Motschulsky, 1865
Male adult
SE-Kazakhstan, 70 km E of Almaty, E Trans-Ili Alatau N-slopes, valley Assy (Asy) riv. Assy plateau h=1700 m, 12.V.2015, L=21 mm,/ det. R. Panin
Copyright © Ruslan Panin
Carabus (Semnocarabus) rustemi Kabak, 2009
Male adult
SE-Kazakhstan, Almaty reg., near Bodokhudzir 43.916667, 79.716667 h=500 m, 12.VIII.2019, det. R.Panin, L=22 mm
Copyright © Ruslan Panin -
Shunichiocarabus taxon page (1 photos)
Carabus (Shunichiocarabus) uenoianus uenoianus Imura, 1995
Male adult
Copyright © - - - -
Sigenacarabus taxon page (6 photos)
Carabus (Pagocarabus) caelestinus caelestinus (Imura, 2007) Paratype
Male adult
Copyright © Boleslav Brezina
Carabus (Pagocarabus) sichuanicola sichuanicola (as drolmae Lassalle, 2003)
Female adult
China, Sichuan, NW Chengdu, Mt. Jiajin Shan h~4000 m,VI. 2016, local collector, L= 24 mm
Copyright © Ruslan Panin
Carabus (Sigenacarabus) trachynodes moxiensis Deuve, 1992
Male adult
China, W-Sichuan, W of Moxi, Daxue Range, Mt.Gongga Shan h=3000-3100m, 29°34′29.6″N /101°59′44.0″E, 10-12.VI.2018, Ch. Reuter leg. L=21 mm
Copyright © Ruslan Panin
Carabus (Sigenacarabus) trachynodes moxiensis Deuve, 1992
Male adult
China, W-Sichuan, Daxue Range, Mt.Gongga Shan h=3000-3100m, Hailuogou Glacier Forest Park, 29.574889, 101.995556, 10-12.VI.2018, Ch. Reuter leg., L=21 mm
Copyright © Ruslan Panin -
Sphodristocarabus taxon page (73 photos)
Carabus (Sphodristocarabus) varians varians Fischer, 1827
Male+Female adult
female: Transcaucasia, Armenia, Amasia distr., Karakhachskiy mountain pass h=2270m, 3.VIII.1985 P.V.Kazaryan leg., 22mm<br /> male: Armenia, Amasia distr., Arpalach vill. VI.1999, P.V. Kazaryan leg.
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) varians armeniacus Mannerheim, 1830
Female adult
NE Turkey,<br /> Artvin Prov., Borçka, Macahel Gecidi h=1450m, 18.VI.2007, D.Prunier leg.<br /> female, 26mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) macrogonus pseudokolatensis Heinz & Korge, 1967
Male adult
NE-Turkey::Kackar Dagi Mt, Ayder S of Ardesen h=1600m, 17-21.VI.2010, M.Janalik leg.
L=20 mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) varians armeniacus Mannerheim, 1830
Male adult
NE Turkey, Artvin Prov., Şavşat, Çamlıca,18.VI.1993, M. Janata leg.<br /> L=22 mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) separatus persa Lapouge, 1924
Male adult
N-Iran: Mazandaran Province, Elborz Mt.Rng, Vandarbon env., 10.VII.2002 L=23 mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) separatus separatus Gehin, 1885
Male adult
N-Iran:Gilan Prov., Asalem - Khalkal road, 25-30.V.2018, Václav Čutka leg. L= 21 mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) separatus separatus Gehin, 1885
Female adult
N-Iran: Gilan Prov., Asalem - Khalkal road, 25-30.V.2018, Václav Čutka leg. L= 23 mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) separatus haerii Heinz, 1973
Female adult
N-Iran:Gilan Prov., Rudbar, Harzevil env.,23-24.V.2018, Václav Čutka leg. L=24 mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) separatus haerii Heinz, 1973
Male adult
N-Iran: Gilan Prov., E of Rudbar, Barehsar env., 19-23.VI.2017, Václav Čutka leg. L=22 mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) varians armeniacus Mannerheim, 1830
Male adult
C Georgia, Imereti, Caucasus, Likhi Mt Range, Rikoti Pass h=900-1000 m, 14-16.V.2017, E. Rutjan E. leg., L=22 mm
Copyright © Ruslan Panin -
Stephanocarabus taxon page
-
Tachypus taxon page (50 photos)
Carabus (Tachypus) cancellatus s.str. (as sulinensis Born, 1902)
Male+Female adult
S UKRAINE, <br /> Krym Peninsula, Staryi Krym, 17.V.2002<br /> L= 26 mm.
Copyright © Ruslan Panin
Carabus (Tachypus) auratus brunieri Barthe, 1921
Male adult
SC-FRANCE, Aude, Rennes les Bains, 6.VII.1989, Sunard A. leg., 26mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
Male adult
Italy-Switzerland, Riviera – Preonzo, 18.II.1998, C.Sola leg.
L=26 mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus graniger Palliardi, 1825
Male adult
SW-Romania, Banatului Montan, Resita - Valiug, 29.V-9.VI.2005. Krause R. leg.
L=25 mm
Copyright © Ruslan Panin
Carabus (Tachypus) cristoforii Spence, 1823
Female adult
S France, Pyrénés Mts., Tarascon; 15mm
Copyright © Aleš Sedláček
Carabus (Tachypus) cancellatus durus Reitter, 1896
Male+Female adult
W Ukraine, Transcarpathian reg., Vinogradov distr., near Malaya Kopanya vill.,7-18.VII.2008, Panin R. leg. <br /> female, 32mm.
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus intermedius Dejean, 1826
Female adult
NW -Serbia,Fruška Gora,Vrdnik env. h=380m, 9-10.VII.2013, M. Janalik leg. L=29 mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus durus Reitter, 1896
Female adult
W Ukraine, Transcarpathian, Velykyi Bereznyi distr., Stuzhytsya (Stužica), 2-29.VI.2011, Yu. Kanarsky leg. L=33.5 mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus graniger (as biharicus Reitter, 1896)
Female adult
CW-Romania, Arad distr., Săvârşin - Ilteu, 24-30.V.2017, V. Šima leg., l=30 mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus cancellatus Illiger, 1798
Male adult
W-Ukraine, Podolian Upland (Lviv Opillia) W of Vybranivka vill., under the bark, 8.II.2008, R.Panin leg., L=22 mm
Copyright © Ruslan Panin -
Tanaocarabus taxon page (14 photos)
Carabus (Tanaocarabus) hendrichsi Bolivar y Pieltain, Rotger & Coronado, 1967
Male adult
male, 23mm
Copyright © Boleslav Brezina
Carabus (Oreocarabus) taedatus agassi Le Conte, 1850
Male adult
SC-USA, New Mexico, Santa Fe,VIII.1999, Sehnal R. leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Oreocarabus) taedatus bicanaliceps Casey, 1920
Male adult
NW-USA, Washington, Thurston County, Mima Mound Prairie, 15.IX.2005, A. Laforgue leg. L=23 мм
Copyright © Ruslan Panin -
Teratocarabus taxon page (2 photos)
Carabus (Teratocarabus) azrael mizunumaianus Imura, 1991
Male adult
China, Liaoning, Benxi, Gongchangung
20.VI.2004 Hong
Copyright © Boleslav Brezina -
Tomocarabus taxon page (30 photos)
Carabus (Tomocarabus) scabripennis scabripennis Chaudoir, 1850
Male adult
Carabus (Tomocarabus) scabripennis scabripennis Chaudoir, 1850
C. Armenia,
Kotayk reg., Yeghvard, Arailer Mt., 16.VI.1999, P.V.Kazaryan leg.
male, 20mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Tomocarabus) marginalis Fabricius, 1794
Male+Female adult
E Ukraine,
Kharkov reg.,near Vasishchevo vill.10-20.VI.2006 D.Kovalchuk leg.
female 24mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) convexus s.str. (as pyrenaicus Born, 1907)
Male adult
Pirineos, Canfranc, 1300m, Barranco de Izas
15.6.2005 Anichtchenko A. leg.
male, 16 mm
Copyright © Alexander Anichtchenko
Carabus (Tomocarabus) bessarabicus bessarabicus Fischer, 1823
Male+Female adult
UKRAINE,
Kherson Reg., Golaya Pristan distr. near Rybalche vill. 6.IV.2000
long (female) 20mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) convexus gracilior Gehin, 1885
Male adult
SE-Bulgaria, Burgas Prov., Primorsko env., 18-30.VI.2000, Krause Rene leg.
L=18 mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) bessarabicus ? concretus Fischer, 1823
Male adult
E Ukraine, Lugannsk Reg., Sverdlovsk distr., Steppe Reserve “Provalsky", 3.V-3.VI.2011, leg.Glotov S./det.Panin R.<br /> female, 22 mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) convexus dilatatus (as perplexus Schaum, 1857)
Male adult
CE-Greece: Larissa reg., Stomio, Ossa Mt., 6-17.VII.2011, Krause Rene leg.
L=19 mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) convexus dilatatus Dejean, 1826
Male adult
NE Serbia, Majdanpek; 13mm
Copyright © Aleš Sedláček
Carabus (Tomocarabus) convexus erzeliki Schweiger, 1962
Female adult
NC-Turkey, Sinop province, Erfelek env., h=200m, 10-27.VII.2014, C. Reuter leg. L=20 mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) marginalis Fabricius, 1794
Endophallus
N Ukraine, Kiev reg., Vyshgorod disrt., near Khotyanivka vill.,
Copyright © Ruslan Panin -
Trachycarabus taxon page (9 photos)
Carabus (Trachycarabus) sibiricus tscherkessicus Korge, 1964
Male adult
S Russia, NW-Caucasus,<br /> Karachay-Cherkessia,Teberda, Mukhinsky (Mukhu) Mt. pass, h=2750-2800 m , 8.VIII.2010, I.A. Solodovnikov leg.<br /> male, 23 mm<br />
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886
Male adult
W Ukraine,
Ternopil reg., Pidvolohcisk distr., Supranivka vill.
male, 26mm
The end of XIX the beginning of XX century
photo Panin R.
coll. SMNH Ukraine, Lviv
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus errans Fischer, 1823
Male+Female adult
female:SE Ukraine,
Donetsk, 25.VII.2009, A.I.Gubin leg. L=30mm.
male:E Ukraine, Lugansk reg., near Slavyanoserbk, 5-15.VIII.2009, S.Konovalov leg. L=26mm.
photo/det.Panin R.
Copyright © Ruslan Panin
Carabus (Trachycarabus) scabriusculus scabriusculus (as erythropus Fischer, 1822)
Male+Female adult
W.Ukraine,<br /> NW Podolian Upland , Lviv Reg., Zolochiv distr., near Luky vill., steppe sites, 7.V-8.VI. 2010<br /> leg. Panin R.<br /> female, 22mm
Copyright © Ruslan Panin
Carabus (Trachycarabus) planus campicola Reitter, 1896
Male adult
Krym Peninsula, near Sudak. Leg.Vashchenko S. 1-5.V.2004<br /> 27 mm
Copyright © Ruslan Panin
Carabus (Trachycarabus) besseri Fischer, 1822
Male+Female adult
W Ukraine.NW Podolian,
Lviv reg., Zolochiv distr., Lysa Gora near Stinka vill., steppe sites, 25.IV-9.V.2009 Panin R. leg
female, 30mm
Copyright © Ruslan Panin
Carabus (Trachycarabus) estreicheri Fischer, 1822 (as jaegeri Ménétriés, 1827)
Female adult
E UKAINE, Lugansk city, Site Luganskoe, 20-30.VII.2010, V.A.Landik leg.
L=21 mm
det.Panin.R.
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus haeres (trans s. errans)
Female adult
C Ukraine, S of Poltava city, edge forest-lane, 2-26.IX.2010, O. Pozharov leg. /det. R.Panin. L=30 mm
Copyright © Ruslan Panin
Carabus (Trachycarabus) estreicheri Fischer, 1822 (ab.rufofemoratus Lomnicki, 1892)
Female adult
W Ukraina, Podolian Upland, Lviv reg., Brody distr., Mt. Makitra, steppe site, 19.V-5.VI.2020, Panin R. leg. L=23 mm
Copyright © Ruslan Panin
Carabus (Trachycarabus) scabriusculus inapertus Motschulsky, 1850
Male adult
NC Ukraine, Kiev city, Lysa Gora, 1-15.VI.2012, Gerasimov R.leg./det. R.Panin, L=19 mm
Copyright © Ruslan Panin -
Tribax taxon page (145 photos)
Carabus (Tribax) titan titan Zolotarev, 1913
Male+Female adult
Russia,NW Caucasus,
Krasnodar reg.,Chernogorie Mnt. Range, Agulova Balka h=950m V-VII.1999, A.S.Zamotajlov leg.
female,32mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Tribax) steveni steveni Menetries, 1832
Male adult
Carabus (Tribax) steveni steveni Menetries, 1832
Karachaevo-Cherkessia,
Hudes riv., near Tsarskie vorota,
V.2009 Anichtchenko A. leg.
male, 25mm
photo Anichtchenko A.
Copyright © Alexander Anichtchenko
Carabus (Tribax) biebersteini otcharensis Kurnakov, 1970
Male adult
W. Caucasus, Abkhazia, Gudauta distr.,
Duripsh vill. env., Khipsta River, h=700 m,
11-06-2009 A. Safronov leg.
male - 40 mm
Copyright © Alexandr Safronov
Carabus (Tribax) puschkini platypterus Ganglbauer, 1886
Male adult
SW Georgia, Lesser Caucasus, Adzhar-Imereti (Meskheti) Mt Range, 10 km S Sairme h~1850 m, 27.VI-2.VII.2013, A.Puchkov & Martynov leg. L=27 mm
Copyright © Ruslan Panin
Carabus (Tribax) puschkini kolenatii (as suramensis Kraatz, 1887)
Male adult
S-Caucasus: NC-Georgia, W slope Likhi Mt.Range, 3 km W Rikoti Pass h~900m 11-14.VI -2006, A.Puchkov leg. L=27 mm
Copyright © Ruslan Panin
Carabus (Tribax) nothus fausti Dohrn, 1873
Male adult
Carabus (Microtribax) fausti fausti Dohrn, 1873 Russia: Daghestan, Kyabyak Mt.R.
Copyright © Igor Solodovnikov
Carabus (Tribax) lederi lederi Reitter, 1882
Female adult
Georgia, Kvemo-Svaneti, Svanetian Mt.R., near Latpari pass., alpin zone, 2700-3000m (TOPOTYPUS)
Copyright © Igor Solodovnikov
Carabus (Tribax) kasakorum kasakorum Semenov, 1896
Female adult
Russia: N.Osetia: Pastbyshchniy Mt.R., S slopes of Khoshkharanrag Mt.R., right bank of Amusadon riv., h=1500 m
Copyright © Igor Solodovnikov
Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Female adult
Russia, Chechnya, Caucasus, Skalystyi Mt. Range upper river Gekhi, Mt. Ekkyrkort, N 42°52', E 45°26', h=2230 m, 26.V–21.VII.2018, D.Fominykh leg.
Copyright © Igor Solodovnikov
Carabus (Tribax) kasbekianus mingrelicus Reitter, 1889
Endophallus
Georgia, Caucasus, 3 km W Nakerala Pass 1450-1500 m, 42.386528, 42.977944
Copyright © Ruslan Panin -
Ulocarabus taxon page (14 photos)
Carabus (Ulocarabus) stschurovskii Solsky, 1874
Male adult
Tadzhikistan, Pamir-Alay, S slope Gissar Mt.Range, Takob, h-1700m, 10.V.1996, Dolin V.G. leg.
det. Panin R.Yu.
L=22 mm
Copyright © Ruslan Panin
Carabus (Ulocarabus) stschurovskii Solsky, 1874
Female adult
Tadzhikistan, Pamir, N-slope Peter I.Mt Rng,h=2100m,Ganishou vill., Dolin V.G. leg.
det.Panin R.Yu.
L=24 mm
Copyright © Ruslan Panin
Carabus (Ulocarabus) theanus delerei Mandl, 1955
Male adult
N-Afghanistan, Hindukush Mt.Rng, NE Salang Pass h=2300 m, 35°27'494"N, 68°58'399"E, IV.2011, C. Reuter leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Ulocarabus) theanus theanus Reitter, 1895
Male adult
Tadzhikistan, S slope Gissar Mt.Rng, left bank of the river Sorbo, Haidar gorge,1.VI.1984, Dolin V.G. leg./det.Panin R.Yu.
L=21 mm
Copyright © Ruslan Panin
Carabus (Ulocarabus)? theanus theanus Reitter, 1895
Male adult
NC Tadzhikistan, Pamir, N-slope Peter I.Mt Rng, h=2100m, Ganishou vill., 10-12.VI.2006, Ivanova E. leg.
L=23 mm
Copyright © Ruslan Panin
