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Carabidae of the World
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Carabinae Latreille, 1802
→
Carabini Latreille, 1802
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Carabina sensu stricto
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Carabus Linne, 1758
→ Meganebrius Kraatz, 1895
Region
Afghanistan (AF)
Albania (AL)
Aleutian isl (ALU)
Algeria (AG)
Andorra (AN)
Angola (ANG)
Antarctica (Antarctica)
Antilles islands (ANT)
Antigua and Barbuda (Antigua and Barbuda)
Barbados (Barbados)
Cayman islands (Cayman islands)
Cuba (Cuba)
Dominica (Lesser Antilles) (Dominica (Lesser Antilles))
Dominican Republic (Hispaniola, Greater Antilles) (Dominican Republic (Greater Antilles))
Grenada (Grenada)
Guadeloupe (Guadeloupe)
Haiti (Haiti)
Jamaica (Jamaica)
Martinique (Martinique)
Puerto Rico (Puerto Rico)
Saint Kitts and Nevis (Saint Kitts and Nevis)
Saint Lucia (Saint Lucia)
St. Vincent (St. Vincent)
Trinidad and Tobago (Trinidad and Tobago)
Arab Emirates (AE)
Argentina (ARG)
Armenia (AR)
Australia (AST)
Christmas Island (Christmas Island)
Lord Howe Island (Lord Howe Island)
Macquarie Island (Macquarie Island)
Tasmania (Tasmania)
Austria (AU)
Azerbaijan (AB)
Azores (AZ)
BALTIC AMBER, Baltic See coast and Amber quarry Jantarny near Kaliningrad, Russia (BAM)
Bahamas (BHM)
Bahrain (BA)
Bangladesh (BGD)
Belgium (BE)
Belize (BLZ)
Bhutan (BT)
Bolivia (BOL)
Bosnia and Herzegovina (BH)
Botswana (BTW)
Brazil (BRZ)
Acre (AC)
Alagoas (AL)
Amazonas (AM)
Arnapá (AP)
Bahia (BA)
Ceará (CE)
District Federal (Brasilia) (DF)
Espirito Santo (ES)
Goiás (GO)
Maranhão (MA)
Mato Grosso (MT)
Mato Grosso do Sul (MS)
Minas Gerais (MG)
Paraná (PR)
Paraîba (PB)
Pará (PA)
Pernambouc (PE)
Piaui (PI)
Rio Grande do Norte (RN)
Rio Grande do Sul (RS)
Rio de Janeiro (RJ)
Rondônia (RO)
Roraima (RR)
Santa Catarina (SC)
Sergipe (SE)
São Paulo (SP)
Tocantins (TO)
Brunei (Brunei)
Bulgaria (BU )
Burundi (BUR)
Byelorussia (BY)
Cambodia (CBD)
Cameroon (CAM)
Canada (CAN)
Canary isls (CI)
Cape Verde (Cape Verde)
Central African Republic (CAR)
Chad (CHD)
Chile (CHL)
Juan Fernández Islands (Juan Fernández Islands)
China (CH)
Anhui (Anhui)
Beijing (Beijing)
Chongqing (Chongqing)
Fujian (Fujian)
Gansu (Gansu)
Guangdong (Guangdong)
Guangxi (Guangxi)
Guizhou (Guizhou)
Hainan (Hainan)
Hebei (Hebei)
Heilongjiang (Heilongjiang)
Henan (Henan)
Honkong (Hongkong)
Hubei (Hubei)
Hunan (Hunan)
Inner Mongolia (Inner Mongolia)
Jiangsu (Jiangsu)
Jiangxi (Jiangxi)
Jilin (Jilin)
Liaoning (Liaoning)
Macau (Macau)
Ningxia (Ningxia)
Qinghai (Qinghai)
Shaanxi (Shaanxi)
Shandong (Shandong)
Shanghai (Shanghai)
Shanxi (Shanxi)
Sichuan (Sichuan)
Tianjin (Tsiensin) (Tianjin)
Xinjiang (Xinjiang)
Xizang (Tibet) (Xizang (Tibet))
Yunnan (Yunnan)
Zhejiang (Zhejiang)
Colombia (COL)
Comoros isl. (COM)
Costa Rica (CRC)
Croatia (CR)
Crozet Islands (CRZ)
Cyprus (CY)
Czech (CZ)
DOMINICAN AMBER, Dominican Republic, Lower Miocene. (DAM)
Democratic Republic of the Congo (CNG)
Denmark (DE)
Diego Garcia isls (DGC)
Djibouti (DJB)
Ecuador (ECD)
Egypt (EG)
Sinai (Sinai)
El Salvador (SVD)
Equatorial Guinea (EQG)
Bioco (Fernando Pó) island (Bioco (Fernando Pó) island)
Eritrea (ERT)
Estonia (EN)
Ethiopia (ETP)
Falkland il. (FLK)
Faroe Islands (FA)
Finland (FI)
France (FR)
Corse (Corse)
French Guiana (GNF)
Gabon (GAB)
Galapagos Islands (GAL)
Georgia (GG)
Abkhazia (Abkhazia)
Ajaria (Ajaria)
South Ossetia (South Ossetia)
Germany (GE)
Great Britain (GB)
Greece (GR)
Aegean Islands (Aegean Islands)
Crete (Crete)
Dodecanese Islands (Dodecanese Islands)
Euboea (Evia) Island (Euboea (Evia) Island)
Ionian Islands (Ionian Islands)
Peloponnese (Peloponnese)
Saronic Islands (Saronic Islands)
Sporades Islands (Sporades Islands)
The Cyclades (The Cyclades)
Guatemala (GTM)
Guyana (GYN)
Honduras (HDR)
Hungary (HU)
Iceland (IC)
India (IND)
Andaman and Nicobar isl. (Andaman and Nicobar isl.)
Andhra Pradesh (Andhra Pradesh)
Arunachal Pradesh (Arunachal Pradesh)
Assam (Assam)
Bihar (Bihar)
Chhattisgarh (Chhattisgarh)
Delhi (Delhi)
Goa (Goa)
Gujarat (Gujarat)
Haryana (Haryana)
Himachal Pradesh (Himachal Pradesh)
Jammu and Kashmir (Jammu and Kashmir)
Jharkhand (Jharkhand)
Karnataka (Karnataka)
Kerala (Kerala)
Ladakh (Ladakh)
Madhya Pradesh (Madhya Pradesh)
Maharashtra (Maharashtra)
Manipur (Manipur)
Meghalaya (Meghalaya)
Mizoram (Mizoram)
Nagaland (Nagaland)
Odisha (Odisha)
Puducherry (Puducherry)
Punjab (Punjab)
Rajasthan (Rajasthan)
Sikkim (Sikkim)
Tamil Nadu (Tamil Nadu)
Telangana (Telangana)
Tripura (Tripura)
Uttar Pradesh (Uttar Pradesh)
Uttarakhand (Uttarakhand)
West Bengal (West Bengal)
Indonesia (IDS)
Borneo (Kalimantan) (Borneo)
Java (Java)
Lesser Sunda Islands (Lesser Sunda Ils.)
Maluku Islands (Maluku Isl.)
Papua (Papua)
Riau Islands (Riau Islands)
Sulawesi (Celebes) (Sulawesi)
Sumatra (Sumatra)
Timor (Timor)
Iran (IN)
Iraq (IQ)
Ireland (IR)
Israel (IS)
Italy (IT)
Sardegna (Sardegna)
Sicily (Sicily)
Japan (JA)
Hokkaido (Hokkaido)
Honshu (Honshu)
Kyushu (Kyushu)
Ryukyu Islands (Ryukyu Islands)
Shikoku (Shikoku)
Jordan (JO)
Kashmir (KA)
Kazakhstan (KZ)
Kenya (KEN)
Kerguelen Islands (Kerguelen Islands)
Kuwait (KU)
Kyrgyzstan (KI)
Laos (LAO)
Latvia (LA)
Lebanon (LE)
Lesotho (LSO)
Libya (LB)
Liechtenstein (LS)
Lithuania (LT)
Luxenbourg (LU)
Macedonia (MC)
Madagascar (MAD)
Madeira Arch. (MR)
Malawi (MLW)
Malaysia (MLS)
Sabah (Borneo) (Sabah (Borneo))
Sarawak (Borneo) (Sarawak (Borneo))
Maldives (MLD)
Malta (MA)
Mauritania (MAU)
Mauritius (MRC)
Melanesia (Melanesia)
Bismarck Archipelago (Bismarck Archipelago)
Fiji (Fiji)
New Caledonia (New Caledonia)
Papua New Guinea (Papua New Guinea)
Solomon Islands (Solomon Islands)
Vanuatu ils (Vanuatu ils)
Mexico (MEX)
Aguascalientes (Aguascalientes)
Baja California (Baja California)
Baja California Sur (Baja California Sur)
Campeche (Campeche)
Chiapas (Chiapas)
Chihuahua (Chihuahua)
Coahuila (Coahuila)
Colima (Colima)
Durango (Durango)
Federal District (Federal District)
Guanajuato (Guanajuato)
Guerrero (Guerrero)
Hidalgo (Hidalgo)
Jalisco (Jalisco)
Mexico State (Mexico State)
Michoacan (Michoacan)
Morelos (Morelos)
Nayarit (Nayarit)
Nuevo Leon (Nuevo Leon)
Oaxaca (Oaxaca)
Puebla (Puebla)
Queretaro (Queretaro)
Quintana Roo (Quintana Roo)
San Luis Potosi (San Luis Potosi)
Sinaloa (Sinaloa)
Sonora (Sonora)
Tabasco (Tabasco)
Tamaulipas (Tamaulipas)
Tlaxcala (Tlaxcala)
Veracruz (Veracruz)
Yucatan (Yucatan)
Zacatecas (Zacatecas)
Micronesia (MIC)
Caroline islands (Caroline islands)
Guam (Guam)
Kiribati (Kiribati)
Marshall ils (Marshall ils)
North Marians isl. (North Marians isl.)
Palau archipelago (Palau archipelago)
Moldavia (MD)
Mongolia (MG)
Montenegro (ME)
Mordor (RU)
Altai Republic and Altaisky Province (Altai Republic and Altaisky Province)
Amurskaya Area (Amurskaya Area)
Bashkiria, E of Chelyabinsk regs. (S Ural mts. Bashkiria, E of Chelyabinsk regs.)
Buryat Republic (Buryat Republic)
Chukot autonomous region (Chukotka)
East Sayan Mountains (East Sayan Mountains)
Ingushetia, Chechnya, Daghestan (E Caucas) (Ingushetia, Chechnya, Daghestan (E Caucas))
Irkutsk Area (Irkutsk Area)
Kabardino-Balkaria, Ossetia (C Caucas) (Kabardino-Balkaria, Ossetia (C Caucas))
Kaliningrad, Briansk, Smolensk, Tula, Orlov, Kaluga, Moskow, Riaz (C of Russian plains)
Kamchatskaya Area (Kamchatka)
Karachaevo-Cherkessia (W Caucas) (Karachaevo-Cherkessia)
Karelia, Leningrad reg., Pskov reg., Kostroma reg., Perm reg., Komi, Pechora plains. (N of Russian plains)
Khabarovsky Province (Khabarovsky Province)
Khamar-Daban Mountains (Khamar-Daban Mountains (E Siberia))
Komi, Yamalo-Nenetsk, Hanty-Mansi (N Ural mts.)
Krasnodar reg. (W Caucas)
Krasnoyarsky Province (Krasnoyarsk)
Kuriles Islands (Kuriles Islands)
Magadanskaya Area (Magadan)
Omsk, S of Tyumen, W Novosibirsk regs. (S of W Siberia)
Perm, Sverdlovsk regs. (C Ural mts)
Primorsky District (Primorie) (Maritime Prov)
Sakhalin Island (Sakhalin Island)
Tuva Republic (Tuva)
Voronezh, Rostov, Saratov, Kalmykia, Volgograd, Astrahan regs. (S of Russian plains)
West Sayan Mountains (West Sayan)
Yakut-Saha Republic (Yakut-Saha Republic)
Zabaykalsky Krai (ex Chita reg.) (Zabaykalsky Krai)
plains of Krasnodar reg., S Rostov reg., Stavropol reg. (Ciscaucas reg. (plains of Krasnodar reg., S Rostov)
taiga from Ural mts to Yenisei riv. (Middle of W Siberia)
tundra of Tyumen, Yamalo-Nenetsk regs. (N of W Siberia)
Morocco (MO)
Mozambique (MZB)
Myanmar (Burma) (BRM)
Namibia (NAM)
Nepal (NP)
Netherlands (NL)
New Zealand (NZL)
Antipodes Islands (Antipodes Islands)
Auckland Islands (Auckland Islands)
Campbell Island (Campbell Island)
Chatham Islands (Chatham Islands)
Nicaragua (NCG)
North Korea (NC)
Norway (NR)
Svalbard (Spitzbergen) (SB)
Oman (OM)
Pakistan (PA)
Panama (PNM)
Paraguay (PRG)
Peru (PER)
Philippines (PLP)
Basilan (Basilan)
Cebu (Cebu)
Guimaras (Guimaras)
Leyte (Leyte)
Luzon (Luzon)
Marinduque (Marinduque)
Masbate (Masbate)
Mindanao (Mindanao)
Mindoro (Mindoro)
Negros (Negros)
Palawan (Palawan)
Panay (Panay)
Polillo Islands (Polillo Islands)
Romblon (Romblon)
Samar (Samar)
Sulu Archipelago (Sulu Archipelago)
Poland (PL)
Polynesia (POL)
Cook Islands (Cook Islands)
Hawaiian Islands (HAW)
Marquesas Islands (Marquesas Islands)
Niue (Niue)
Norfolk ils (Norfolk ils)
Pitcairn ils (Pitcairn ils)
Rapa Nui (Easter Island) (Rapa Nui)
Samoa (Samoa)
Society Islands (Society Islands)
Tahiti (Tahiti)
Tokelau (TKL)
Tonga (Tonga)
Tuvalu ils (Tuvalu ils)
Portugal (PT)
Qatar (QA)
Republic of South Africa (RSA)
Republic of the Congo (CG)
Reunion isl. (Reunion isl.)
Rodriguez isls. (ROD)
Romania (RO)
Rwanda (RWA)
Santa Helena isl (SH)
Sao Tome e Principe (STM)
Saudi Arabia (SA)
Serbia (SRB)
Seychelles (SEY)
Singapore (SGP)
Slovakia (SK)
Slovenia (SL)
Somalia (SOM)
South Georgia and the South Sandwich Islands (SGSSI)
South Korea (SC)
Spain (SP)
Balearic Islands (Balearic Islands)
Sri Lanka (SRL)
St. Paul isls (STP)
Sudan (SUD)
Surinam (SUR)
Swaziland (SWZ)
Sweden (SV)
Switzerland (SZ)
Syria (SY)
Tadzhikistan (TD)
Taiwan (Formosa) (Taiwan)
Tanzania (TAN)
Zanzibar isl. (Zanzibar isl.)
Thailand (TAI)
Tunisia (TU)
Turkey (TR)
Turkmenistan (TM)
Uganda (UGD)
Ukraine (UK)
Crimea (Crimea)
United States (USA)
Alabama (Alabama)
Alaska (Alaska)
Arizona (Arizona)
Arkansas (Arkansas)
California (California)
Colorado (Colorado)
Connecticut (Connecticut)
Delaware (Delaware)
Florida (Florida)
Georgia (Georgia)
Idaho (Idaho)
Illinois (Illinois)
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Iowa (Iowa)
Kansas (Kansas)
Kentucky (Kentucky)
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Maine (Maine)
Maryland (Maryland)
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Michigan (Michigan)
Minnesota (Minnesota)
Mississippi (Mississippi)
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Pennsylvania (Pennsylvania)
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South Dakota (South Dakota )
Tennessee (Tennessee)
Texas (Texas)
Utah (Utah)
Vermont (Vermont)
Virginia (Virginia)
Washington (Washington)
West Virginia (West Virginia)
Wisconsin (Wisconsin)
Wyoming (Wyoming)
Uruguay (URG)
Uzbekistan (UZ)
Venezuela (VNZ)
Vietnam (VTN)
West Africa (WAF)
Benin (Benin)
Burkina Faso (Burkina Faso)
Côte d'Ivoire (Côte d'Ivoire)
Gambia (Gambia)
Ghana (Ghana)
Guinea (Guinea)
Guinea Bissau (Guinea Bissau)
Liberia (Liberia)
Mali (Mali)
Niger (Niger)
Nigeria (Nigeria)
Senegal (Senegal)
Sierra Leone (Sierra Leone)
Togo (Togo)
Yemen (YE)
Socotra (Socotra)
Zambia (ZMB)
Zimbabwe (ZIM)
Main |
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Adult
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Endophallus
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Labels
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Biotope
Carabus (Meganebrius) arunensis Heinertz, 1980
Female adult
Copyright ©
Boleslav Brezina
Carabus (Meganebrius) arunensis Heinertz, 1980
Habitat
Nepal
Copyright ©
Boleslav Brezina
Carabus (Meganebrius) arunensis Heinertz, 1980
Male adult
E-Nepal, Kosi zone, Khandbari, 1.5km SW of Chichila vill., 27*47’175’’N/ 87*23’622’E, h=1880m, damaged primary forest, 11.V.2010, B.Brezina & M.Häckel leg.
L=27 mm
Copyright ©
Ruslan Panin
arunensis
taxon page
(3 photos)
Carabus (Meganebrius) deliae colasianus (Morvan & Mandl, 1974)
Holotype
Male adult
Copyright ©
Igor Solodovnikov
Carabus (Meganebrius) deliae colasianus (Morvan & Mandl, 1974)
Male adult
Copyright © - - -
Carabus (Meganebrius) deliae deliae (Morvan, 1972)
Holotype
Male adult
Copyright © - - -
Carabus (Meganebrius) deliae deliae (Morvan, 1972)
Holotype
Label
Copyright © - - -
deliae
taxon page
(4 photos)
Carabus (Meganebrius) everesti everesti Andrewes, 1929
Endophallus
Kemba Kharka
Copyright ©
Joachim Schmidt
everesti
taxon page
(1 photos)
Carabus (Meganebrius) franzi Mandl, 1974
Female adult
Nepal, Lantang, near Dunche, Songta vill,
3500 m, 15.5.2000
Copyright ©
Alexander Anichtchenko
franzi
taxon page
(1 photos)
Carabus (Meganebrius) hongliangi Deuve, Liu & Liang, 2024
Holotype
Male adult
Chine, Tibet, Préfecture de Xigazê, canton de Gyirong, ville de Gyirong, village de Cha, col de Latuo La, 3995 m, 28,3908°N 85,4074°E, pelouses alpines (Hongliang Shi leg., 10.VII.2019, sous des pierres), in IOZ. Image from Deuve et al 2024
Copyright ©
- - -
hongliangi
taxon page
(1 photos)
Carabus (Meganebrius) indicus indicus Fairmaire, 1889
Endophallus
Tonglu
Copyright ©
Joachim Schmidt
indicus
taxon page
(1 photos)
Carabus (Meganebrius) kadoudali (Morvan, 1981)
Male+Female adult
C-Nepal, SW Dhaulagiri, W Jaljala h=3150 m, 28°30'05"N, 83°11'48"E, 23.IX.2012, J. Schmidt leg.
female, 20 mm
Copyright ©
Ruslan Panin
Carabus (Meganebrius) kadoudali (Morvan, 1981)
Paratype
Female adult
Vallée de Uttar Ganga, Dhorpatan, col de Jhala-Jélé , 3300m, Morvan P. 1977-04-01/1977-04-30 (MNHN) http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec184
Copyright ©
MNHN
Carabus (Meganebrius) davatchii (Morvan, 1981)
Paratype
Male adult
Vallée de Uttar Ganga, Dhorpatan, col de Jhala-Jélé (MNHN) http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec186
Copyright ©
MNHN
kadoudali
taxon page
(3 photos)
Carabus (Meganebrius) koganae koganae Colas, 1961
Holotype
Male adult
Copyright ©
Igor Solodovnikov
koganae
taxon page
(1 photos)
Carabus (Meganebrius) lebretae lebretae Colas, 1961
Holotype
Male adult
Copyright © - - -
lebretae
taxon page
(1 photos)
Carabus (Meganebrius) pseudoharmandi Mandl, 1965
Female adult
E-Nepal, Kosi zone, Khandbari,(1.5km SW of Chichila vill., 27*45’843’’N/ 87*22’473’E, h=1940m), damaged primary forest, 13 .V.2010, B.Brezina & M.Häckel leg.
L=23 mm
Copyright ©
Ruslan Panin
pseudoharmandi
taxon page
(1 photos)
Carabus (Meganebrius) quinlani smetanai Häckel, 2011
Paratype
Male+Female adult
C-Nepal:Dhaulagiri Massif:Baglung Distr., (1.5km W of Okhale vill., 28.32’152’’N/ 83.48’673’E, h=2600m) damaged primary forest, 19.V.2010, B.Brezina & M.Häckel leg.
male, 21mm
female, 23 mm
Copyright ©
Ruslan Panin
Carabus (Meganebrius) quinlani smetanai Häckel, 2011
Paratype
Endophallus
C-Nepal:Dhaulagiri Massif:Baglung Distr.,(1.5km W of Okhale vill., 28.32’152’’N/ 83.48’673’E, h=2600m)
Copyright ©
Ruslan Panin
Carabus (Meganebrius) quinlani quinlani Mandl, 1965
Holotype
Male adult
Copyright © - - -
Carabus (Meganebrius) quinlani quinlani Mandl, 1965
Male adult
C-Nepal, Gandaki, Manang Distr., Timang env., h~2500-2600m, old open forest, grassland, 3-5.V.2007, Fouqué R. leg. L=19 mm
Copyright ©
Ruslan Panin
quinlani
taxon page
(4 photos)
Carabus (Meganebrius) salpansis Deuve, 1984
Endophallus
Kemba Kharka
Copyright ©
Joachim Schmidt
salpansis
taxon page
(1 photos)
Carabus (Meganebrius) santostamangi thanglaensis Deuve & Schmidt, 2017
Holotype
Male adult
Est-Népal, limite des districts de Sankhuwasabha et de Taplejung, Jaljale Himal, Thangla Phedi, 4 150 m, 27°39’09’’ N – 87°35’56’’ E (S. Tamang leg., 27-vi-2011), in coll. J. Schmidt (Admannshagen, Allemagne).
Copyright ©
Joachim Schmidt
Carabus (Meganebrius) santostamangi santostamangi
Holotype
Male adult
Est-Népal, district de Taplejung, versant oriental de la Palung Khola, 3 720 m, 27° 35’ 43’’ N – 87° 41’ 50’’ E (S. Tamang leg., 7-vii-2011), in coll. J. Schmidt (Admannshagen, Allemagne).
Copyright ©
Joachim Schmidt
Carabus (Meganebrius) santostamangi santostamangi
Endophallus
Copyright ©
Joachim Schmidt
santostamangi
taxon page
(3 photos)
Carabus (Meganebrius) tamang gurkhaorum Lassalle, 1985
Female adult
CW Nepal, Manaslu massif 2300 m,
Bara Pokhari Lekh, 5.2001 M.Hackel leg.
Copyright ©
Alexander Anichtchenko
tamang
taxon page
(1 photos)
Carabus (Meganebrius) thudamensis Deuve & Schmidt, 2017
Holotype
Male adult
Est-Népal, District de Sankhuwasabha, Thudam, 3500 – 3600 m, 27°45’23’’N – 87°32’56’’E (S. Tamang leg., 1-vii-2011), in coll. J. Schmidt (Admannshagen, Allemagne).
Copyright ©
Joachim Schmidt
Carabus (Meganebrius) thudamensis Deuve & Schmidt, 2017
Holotype
Endophallus
Copyright ©
Joachim Schmidt
thudamensis
taxon page
(2 photos)
Carabus (Meganebrius) tuberculipennis ganjalaensis Deuve, 2000
Male adult
Nepal, Helambu, GanjaLa Pass S-slope, Dhukpu Kharka h=4100m, 28°03'50"N, 85°34'21"E, 12.VIII.2010, S.Tamang leg., L=21 mm
Copyright ©
Ruslan Panin
tuberculipennis
taxon page
(1 photos)
Carabus (Meganebrius) wallichi wittmeri Mandl, 1974
Female adult
Nepal, Lantang, near Dunche, Songta vill.
3500 m 15.5.2000
Copyright ©
Alexander Anichtchenko
wallichi
taxon page
(1 photos)
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