Carabidae image gallery
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Acathaicus taxon page (17 photos)
Carabus (Acathaicus) alexandrae alexandrae Semenov, 1887 Holotype Holotype
Male adult
Kwantin and Li-tschara-pu, 22.VI.1885, leg. Potanin<br /> Deposited in Zoolog. Museum St. Petersburg.<br /> male, long. 22 mm<br /> photo B. Brezina
Copyright © Boleslav Brezina
Carabus (Acathaicus) alexandrae cratocephaloides Semenov, 1887
Female adult
female, 28mm
Copyright © Boleslav Brezina
Carabus (Acathaicus) interruptocostatus Semenov, 1887 Holotype Holotype
Female adult
Copyright © Boleslav Brezina -
Acoptolabrus taxon page (63 photos)
Carabus (Acoptolabrus) lopatini Morawitz, 1886
Male+Female adult
Russia,
Sakhalin isl., Tchekhov Mt, VI.1997
female, 40mm
photo & coll. Panin R.
Copyright © Ruslan Panin
Carabus (Acoptolabrus) schrencki kimieae Kezuka & Imura, 1989
Male+Female adult
Mt. Wulongshan, Dandong City,
Liaoning Province, CHINE
25-30.VI.2008
Copyright © Safronov
Carabus (Coptolabrodes) haeckeli Brezina & Imura, 1997
Male adult
C China, Shaanxi,
Tsinling Mts., Taibaishan Mt., h=3000m, 33:55N/107:44 E, VI.2004, V.Siniaev & his team
female, 31 mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Acoptolabrus) schrencki schrenck (as dongningensis Deuve & Li, 2005)
Male adult
NE China, Heilongjiang prov., Dongning, V.2009<br /> L=25 mm
Copyright © Ruslan Panin
Carabus (Acoptolabrus) mirabilissimus rigoutianus Deuve & Li, 1998
Male adult
N-Korea, South Pyongan prov., Mt. Maeng-san, 6.VI.2010
L=25 mm
Copyright © Ruslan Panin
Carabus (Acoptolabrus) gehinii gehinii Fairmaire, 1886
Female adult
N-Japan, SW-Hokkaido, Chitose env.,VI.2011.
L=34 mm
Copyright © Ruslan Panin
Carabus (Acoptolabrus) ? leechi huisaensis Deuve & Li, 1999
Male adult
NC-Korea, Ryanggang Prov., Puksubaek-san, 4-10.V.2010
28 mm
Copyright © Ruslan Panin
Carabus (Acoptolabrus) schrencki minpongsanensis Deuve & Li, 2003
Male adult
North Korea, Chagang Prov., Manpo, Songhak-ri (villl.,) 1-16.VII.2011, L=26 mm
Copyright © Ruslan Panin
Carabus (Acoptolabrus) gehinii munakatai (Ishikawa, 1968)
Male adult
Japan, SW-Hokkaido, Yakumo town, 4.VI.2017, S.Shinichi leg., L=30 mm
Copyright © Ruslan Panin
Carabus (Acoptolabrus) constricticollis Kraatz, 1886
Male adult
China, Liaoning prov., Mt. Guanmen 20.6.2009 (cAA)
Copyright © Alexander Anichtchenko -
Acoptopterus taxon page (12 photos)
Carabus (Carabus) gressittianus fuscomarginatus Kleinfeld, 1997 Holotype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Carabus) andreiianus Deuve, 2007
Female adult
C-China, N-Sichuan, Pingwu Co. h=1500 m, 27.VI.2011, L=24 mm
Copyright © Ruslan Panin
Carabus (Carabus) koidei koidei Imura, 1996
Male adult
C-China, S-Sichuan, Pass btw. Yanyaun / Muli, h=3244 m, 27.68638N; 101.22335E,,11-18.VI.2017, Ch. Reuter leg. L=22 mm
Copyright © Ruslan Panin
Carabus (s.str.) morphocaraboides zhaojueanus Imura, 1999
Female adult
China, S-Sichuan, SE Mianning, Pass btw Mianshanzhen & Yuexi 28°30’N/102°26’E, 22-23.VI.2017, Ch.Reuter leg., L=22 mm; det. B. Brezina, 2020
Copyright © Ruslan Panin
Carabus (s.str.) morphocaraboides zhaojueanus Imura, 1999
Male adult
China, S-Sichuan, road 56 km E of Xichang to Zhaojue, Mt.WenLin, Pass h=3200 m, 27°52’N/102°30E, 5-20.VI.2017, Ch.Reuter leg.; det B. Brezina, 2020
Copyright © Ruslan Panin
Carabus (Acoptopterus, ?Carabus) yunnanus yanyuanicus Cavazzuti, 1996
Endophallus
China, S-Sichuan, S Yanyuan, Chuandongzi Mt h=3300-3600 m
Copyright © Ruslan Panin
Carabus (Acoptopterus) yunnanus yanyuanicus Cavazzuti, 1996 (? or oblongior ssp.)
Male adult
China, S-Sichuan, S Yanyuan, Chuandongzi Mt h=3300-3600 m, 16-20.VI.2023, C.Reuter leg./det.Klienfeld, 2023, L=21.5 mm
Copyright © Ruslan Panin -
Acrocarabus taxon page (6 photos)
Carabus (Acrocarabus) callisthenoides Semenov, 1888
Male adult
male, 22mm
Copyright © Boleslav Brezina
Carabus (Acrocarabus) guerini aldarkose Kabak, 1992
Male adult
male, 22mm
Copyright © Boleslav Brezina
Carabus (Acrocarabus) guerini guerini Fischer, 1842
Male adult
SE Kazakhstan, W Dzhungar Mts, Lepsinsk env., 1600 m., 18.06.2000, A.Klimenko
Copyright © Anton Menstschikov -
Alipaster taxon page (18 photos)
Carabus (Alipaster) barovskii bayanbulak Deuve, 1993 Paratype Paratype
Female adult
female, 17mm
Copyright © Boleslav Brezina
Carabus (Alipaster) infantulus infantulus Morawitz, 1886
Male adult
male, 15mm
Copyright © Boleslav Brezina
Carabus (Cechenotribax) petri Semenov & Znojko, 1932
Female adult
female, 18mm
Copyright © Boleslav Brezina
Carabus (Eotribax) eous eous Morawitz, 1889
Male adult
Kirghyzstan, N Terskei Ala Too, Borskaun riv., h=2500 m., 7-14.07.2000
male
L=17 mm
Copyright © Maksim Smirnov
Carabus (Cechenotribax) petri Semenov & Znojko, 1932
Male adult
SE Kazakhstan, N-Tianshan:N-slopes Ketmen Mt.Rng. h=2250-2500m., near Ketmen vill.,1-5.VI.1990, Dolin V.G. leg.
L=13.5 mm
Copyright © Ruslan Panin
Carabus (Alipaster) pupulus pupulus Morawitz, 1889
Female adult
S Kyrgyzstan, Inner Tian Shan, Dzhany-Dzher Mt range, upper Mudyuryum riv. Ak-Baytal gorge h~3300 m, 15.VII.2011, S.Toporov leg. L=16 mm
Copyright © Ruslan Panin -
Apoplesius taxon page (5 photos)
Carabus (Apoplesius) deuvei hajdaji Hackel, 2007 Holotype Holotype
Male adult
Turkey- prov. Mus 60 km NW Mus 1630m
3856N 4108E Bulgan Gec. 23-24.06.03 E. & P. Hajdaj leg.
Copyright © Boleslav Brezina
Carabus (Apoplesius) pseudodepressus Machard, 1988
Female adult
Turkey, Prov. Bitlis, S near Tatvan, 23.IV.2009 leg. M. & S. Nabozhenko
L=19.0 mm
Copyright © Kirill Makarov -
Apotomopterus taxon page (197 photos)
Carabus (Apotomopterus) anxiensis Kleinfeld, 1998 Holotype & Paratype Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) cantonensis pervarius Kleinfeld, 1997 Holotype
Male adult
Frank Kleinfeld
Copyright © - - -
Carabus (Apotomopterus) cantonensis metallaicus Kleinfeld, 1999 Holotype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) davidis davidis Deyrolle & Fairmaire, 1878
Male adult
SE-China, Jiangxi prov., Yushan County, Mt Wangjing, VI,2012
L=38 mm
Copyright © Ruslan Panin
Carabus (Apotomopterus) protenes baoxingensis Deuve & Mourzine, 2003
Endophallus
China: С-Sichuan:Qionglai Shan Mts., Baoxing env. (TOPOTYPUS)
Copyright © Igor Solodovnikov
Carabus (Apotomopterus) delavayi parvipes Cavazzuti, 1996
Male adult
S-Sichuan: S of Yanyuan, Chuandongzi Massif h=3200 m, 27.351206, 101.507036, 8-19.VI.2017, C.Reuter leg., L=32 mm
Copyright © Ruslan Panin
Carabus (Apotomopterus) protenes ascendens Semenov, 1898
Endophallus
China, Chongqing, N Jinfo Shan, Shanwangping h=1400m, 29.101944, 107.166389
Copyright © Ruslan Panin
Carabus (Apotomopterus) patroclus apricalis Cavazzuti, 1997
Male adult
China, S Sichuan, W of Pingchuanzhen, near Galuo vill., h=2400 m, 7-18.VI.2017, C.Reuter leg./det. F.Kleinfeld, L=30 mm
Copyright © Ruslan Panin
Carabus (Apotomopterus) bousquetellus bousquetellus Deuve, 1998
Male adult
S-China, CW-Yunnan, W of Lincang, Daxue Shan h=2600m, 23.935556, 100.257778, 7-13.VI.2023, C.Reuter leg./det. F.Kleinfeld, L=24 mm
Copyright © Ruslan Panin
Carabus (Apotomopterus) tonkinensis Deuve, 1990
Male adult
Vietnam, Yen Bai, 5.2020 (cAA)
Copyright © Alexander Anichtchenko -
Archicarabus taxon page (40 photos)
Carabus (Archicarabus) monticola fontanae Born, 1906
Male+Female adult
N-Italy, Piemonte, Bielmonte, h1350 m, 14-15.VI.2010, Pascal Stefani leg.; female, 23 mm
Copyright © Ruslan Panin
Carabus (Archicarabus) montivagus montivagus Palliardi, 1825
Male+Female adult
SW-Romania: Banat, Caras-Severin Reg., Sasca Montana – Stinapari, 21.V.-5.VI.2010, Krause Rene leg.<br /> female, 26 mm
Copyright © Ruslan Panin
Carabus (Archicarabus) pseudomonticola Lapouge, 1908 ? nemoralis
Male adult
NE Spain, Catalonia, Sant Jaume de Frontanyà, h=1000 m, V-VI.2010, Pascal Stefani leg.
L=26 мм
Copyright © Ruslan Panin
Carabus (Archicarabus) gotschi gotschi Chaudoir, 1846
Male+Female adult
Transcaucasia, Armenia,
Ashotsk distr., Bandivan, 8-21.X.1991 P.V.Kazaryan leg.
female, 20mm
Copyright © Ruslan Panin
Carabus (Archicarabus) steuartii Deyrolle, 1852
Female adult
NW-Spain, Pontevedra prov., Serra do Candán Mt range, 1.III.1992, S. Devesa leg./det. S. Sobrino
Copyright © Ruslan Panin
Carabus (Archicarabus) monticola roccae Born, 1923
Male adult
NW-Italy, Liguria: Savona env., Monte San Giorgio, 30.X.1977, Monauzzi leg. L=23 mm
Copyright © Ruslan Panin
Carabus (Archicarabus) phoenix Lapouge, 1924
Male+Female adult
C-Lebanon, Byblos /Jbeil Distr., Kfardebian env., h=1200m, XI.2014, C. Reuter leg. female,23mm
Copyright © Ruslan Panin
Carabus (Archicarabus) wiedemanni (as mussardianus Breuning, 1965)
Male adult
NC-Turkey, Sinop province, Erfelek env., h=200m, 10-27.VII.2014, C. Reuter leg. L=23 mm
Copyright © Ruslan Panin
Carabus (Archicarabus) monticola monticola (as maritimensis Born, 1923)
Female adult
SE France, Alpes-Maritimes, Col de Braus h=1000 m, 23.V-18.VI.2014, P. Cavazzuti & G. Proscia leg., L= 24 mm
Copyright © Ruslan Panin
Carabus (Archicarabus) victor Fischer, 1836
Male adult
S Georgia, Samtskhe-Javakheti, Akhaltsikhe distr., near Andriatsminda, Anda vill h=1550 m, 14.VII.2021, E. Rutjan leg./ det. R.Panin l=21 mm
Copyright © Ruslan Panin -
Aristocarabus taxon page (13 photos)
Carabus (Aristocarabus) viridifossulatus viridifossulatus Fairmaire, 1887
Female adult
35mm
Copyright © Boleslav Brezina
Carabus (Aristocarabus) viridifossulatus seticollis Deuve & Mourzine, 2000
Male adult
28 mm
Copyright © Boleslav Brezina -
Aulonocarabus taxon page (70 photos)
Carabus (Aulonocarabus) gaschkewitschi czarensis Obydov, 1997
Female adult
Russia, CE-Siberia, N Chita reg., Kodar Mt. Range, Big Leprindo Lake env. h~2000m., 2-5.VII.2002 A. Prokhorov leg./det. A. Anichtchenko
L=19 mm
Copyright © Ruslan Panin
Carabus (Aulonocarabus) kurilensis ? daisetsuzanus Kono, 1936
Male adult
N-Japan, Hokkaido, Katō District,Kamishihoro, Horoka near Maruyama Bridge,19.VI.2013, T. Hirabayashi leg.
L=24 mm
Copyright © Ruslan Panin
Carabus (Aulonocarabus) gaschkewitschi gaschkewitschi Motschulsky, 1859
Male adult
Copyright © Kirill Makarov
Carabus (Aulonocarabus) kolymensis kodarensis Brinev, 1997
Male adult
Russia, SE Siberia, N Chita reg., Kodar Mt. Rng. h=1800m, Novaya Chara vill., 23.VI.2006. L=17 mm
Copyright © Ruslan Panin
Carabus (Aulonocarabus) truncaticollis fleischeri Reitter, 1898
Female adult
Russia, N Burajatia, Barguzinskiy Mts., the upper reaches of M.Chivyrkuy riv., h=2000 m, 26.VI-12.VII.2007, Vashchenko S leg. L=18.5 mm
Copyright © Ruslan Panin
Carabus (Aulonocarabus) truncaticollis fleischeri Reitter, 1898
Male adult
Russia, N Burajatia, Barguzinskiy Mts., the upper reaches of M.Chivyrkuy riv., h=2000 m, 26.VI-12.VII.2007, Vashchenko S leg. L=17 mm
Copyright © Ruslan Panin -
Axinocarabus taxon page (7 photos)
Carabus (Axinocarabus) fedtschenkoi fedtschenkoi Solsky, 1874
Male adult
27 mm
Copyright © Boleslav Brezina
Carabus (Axinocarabus) fedtschenkoi kondarensis Kryzhanovskij, 1953
Female adult
NW-Tadzhikistan: Gissar Mt.Rng, Varzob river ravine, Gushari env., h~1400-1500m, 24.III.2010, L=29 mm
Copyright © Ruslan Panin -
Calocarabus taxon page (61 photos)
Carabus (Calocarabus) linxiaensis ponggartangensis Deuve, 1995 Holotype
Male adult
17 mm
Copyright © Boleslav Brezina
Carabus (Cryptocechenus) sifanicus Semenov, 1898 (as oxydocupreus)
Male adult
male, 17mm
Copyright © Boleslav Brezina -
Carabus sensu stricto taxon page (112 photos)
Carabus (Carabus) arvensis arvensis Herbst, 1784
Male adult
W. Ukraine , NW. Podolian, Lviv reg., Nikolaev distr., near Krasiv vill., 9.II.08, Panin R. leg. L=17mm
Copyright © Ruslan Panin
Carabus (Carabus) granulatus leander Kraatz, 1878
Male+Female adult
Russia, C-Caucasus: Balkaria, Valley of riv. Cherek Balkarskyi, h=1750-1800m 10.VI-20.VII.1992, Zamotajlov A. leg.
female,22mm
Copyright © Ruslan Panin
Carabus (Carabus) arvensis deyrollei Gory, 1839
Female adult
NW Spain, Galicia Reg., Pontevedra Prov., A Lama, Xesta, h~650 m, 1.X.1994
L=17 mm
Copyright © Ruslan Panin
Carabus (Carabus) granulatus granulatus Linne, 1758
Male adult
W-Ukraine, Roztochia,
Lviv Reg., Javorov distr., near Ivano-Frankovo vill., Site Zalyvki, 17.IV-3.V.2010, Panin R. leg.
L=20 mm
Copyright © Ruslan Panin
Carabus (Carabus) granulatus yezoensis Bates, 1883
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Carabus (Carabus) menetriesi menetriesi Hummel, 1827
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Carabus (Carabus) arvensis carpathus Born, 1902
Male adult
W Ukraine, NE Carpathians,Transcarpathian reg., Vihorlat, near Strychava vill., Mt Knyaginytsya h= 500 m, 18.V-22.VI. 2014, Yu.Kanarsky leg. L=18 mm
Copyright © Ruslan Panin
Carabus (Carabus) granulatus calabricus Spettoli & Vigna Taglianti, 2001
Female adult
S-Italy: Calabria, Nat.Park Aspromonte, near Gambarie h=1340 m, 38.158149, 15.823323, 3-6.IV.2025, M.Egger leg., L=22 mm
Copyright © Ruslan Panin
Carabus (Carabus) granulatus calabricus Spettoli & Vigna Taglianti, 2001
Male adult
S-Italy: Calabria, Nat.Park Aspromonte, near Gambarie h=1340 m, 38.158149, 15.823323, 3-6.IV.2025, M.Egger leg., L=19 mm
Copyright © Ruslan Panin -
Cathaicus taxon page (7 photos)
Carabus (Cathaicus) brandti brandti Faldermann, 1835 (as grossegranulatus) Syntypes Lectotype
Male adult
C. (Cathaicus) brandti grossegranulatus Breuning 1943<br /> Lectotypus/SyntypusLectotypus (25 mm) von "South Mandschou". Syntypus (25 mm) "South Mandschou, 30. V. 1937"
Copyright © Frank Kleinfeld -
Cathoplius taxon page (14 photos)
Carabus (Cathoplius) asperatus asperatus (Dejean, 1826)
Male adult
male, 24mm
Copyright © Boleslav Brezina -
Cechenochilus taxon page (45 photos)
Carabus (Procechenochilus) kokujewi Semenov, 1898
Female adult
RUSSIA, Karachay-Cherkessia, NW Caucasus Mts., Daut Mt. Range, Iskrovtsev - W.Ullu-Kel Mt. pass circus, h=2800-2600m
Copyright © Alexander Anichtchenko
Carabus (Cechenochilus) heydenianus Starck, 1889
Male+Female adult
NW-Caucasus,
Krasnodar reg., Mostovskoy distr., Skirda Mt Range, N slopes Yatyrgvart Mt. h=2300-2500m., 09-10.VII.2008, D. Fominykh leg.
female,19mm/male,18mm
Copyright © Ruslan Panin
Carabus (Cechenochilus) adangensis Gottwald, 1983
Female adult
Russia, NW-Caucasus:
Karacaj-Cherkessija, Abishira–Akhuba Mt. Range h=2000-2650m, Kyafar-Agur Riv. 14–17.VII.2009, Fominykh D. leg.
female,21mm
Copyright © Ruslan Panin
Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Paratype
Male adult
Russia, W Caucasus,
Karacaj-Cherkessija, Abishira-Akhuba Mnt Range, Area of pass Turii h~2650-2850m, 17.VII.2009, Fominykh D. leg.
male,17 mm
photo & coll. Panin R.
Copyright © Ruslan Panin
Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Paratype
Male adult
Russia, Karachai-Circassian Republic, Abishira-Akhuba Mount Range between passes Agur and Mylgvar, 2785–2830 m.
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) kokujewi Semenov, 1898
Male adult
Russia, Karachai-Cherkessia, vall. Daut Riv., near Glacier Ryndzhi-Murudzhu, h = 3050 m, 22.VIII.2011, leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) kokujewi Semenov, 1898
Male adult
Russia, Karachai-Cherkessia, vall. Daut Riv., near Glacier Ryndzhi-Murudzhu, h = 3050 m, 22.VIII.2011, leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) kokujewi Semenov, 1898
Female adult
Russia, Karachai-Cherkessia, vall. Daut Riv., near Glacier Ryndzhi-Murudzhu, h = 2550 m, 23.VIII.2011, leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) boeberi tschitscherini Semenov, 1896
Female adult
Russia, C Caucasus, Ingushetia, S slopes of Skalistyi Mt.R.
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) boeberi aequaliceps Reitter, 1896
Male adult
Russia, Dagestan, Andiyskiy Mt.R., near Andiyskiye vorota pass, 06.VI–18.VII.2021, leg. D. Fominykh
Copyright © Igor Solodovnikov -
Cephalornis taxon page (1 photos)
Carabus (Cephalornis) potanini (Semenov, 1887)
Male adult
China, S Gansu, 3400m, Minshan mts.,
Wenxian 8.92, J. M. Bousquet
Copyright © Boleslav Brezina -
Chaetocarabus taxon page (32 photos)
Carabus (Chaetocarabus) intricatus bayardi Solier, 1835
Male+Female adult
S-Italy,
Basilicata Reg., Potenza Prov., Latronico, F. Angelini leg. 14.VI.2002
female, 26mm
Copyright © Ruslan Panin
Carabus (Heterocarabus) marietti marietti (as ativsengueni Schweiger, 1962)
Male adult
male, 17mm
Copyright © Boleslav Brezina
Carabus (Chaetocarabus) intricatus krueperi Reitter, 1896
Male adult
CE-Greece: Pieria, Stomio (Larissa) Ossa Mt. 7-14.VII.2009, Krause Rene leg.
L=35 mm
Copyright © Ruslan Panin
Carabus (Chaetocarabus) arcadicus arcadicus Gistl, 1850
Female adult
CE-Greece, Kato Olympos, Karia env., 6-17.VII.2011, Krause Rene leg. L=37 mm
Copyright © Ruslan Panin
Carabus (Chaetocarabus) arcadicus merlini Schaum, 1861
Male adult
S-Greece:Peloponnesos: Taygetos Mt.Rng., 17-23.V.2005, V. Sima leg.
L=30 mm
Copyright © Ruslan Panin
Carabus (Heterocarabus) marietti marietti Cristoforis & Jan, 1837
Female adult
as ativsengueni Schweiger, 1962
NW Turkey, Bolu, Abant Mts.; 16mm
Copyright © Aleš Sedláček
Carabus (Chaetocarabus) arcadicus arcadicus Gistl, 1850
Male adult
Graecia, Pindus, Mt. Velouchi (Timfristos), 1200m, vi.1970-A. Sermet Leg., 30 mm
Copyright © Noël Mal
Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Male adult
W Ukraine, Lviv city, Sykhiv, Zubra forest, (Osvytskiy Grove), under bark, 49°47'28.10" N, 24° 2'20.09" E, 10.XI.2017, Panin R leg.
Copyright © Ruslan Panin
Carabus (Chaetocarabus) intricatus bayardi (as silaensis Haury, 1881)
Male adult
S-Italy, Calabria, Sila Grande Massif, Camigliatello h=1040 m, 19.XI.2009, G. Proscia leg. L=28 mm
Copyright © Ruslan Panin
Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Female adult
W Ukraine, Lviv city, Holosko, Bryukhovytsʹkyi forest 49.886833, 24.012083 under bark, 10.XI.2024, R. Panin leg., L=35 mm
Copyright © Ruslan Panin -
Chrysocarabus taxon page (83 photos)
Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860
Male+Female adult
SW -France,
Pyrenees-Atlantiques, Foret de Sare, XII.2009, Pascal Stefani leg.
female, 31 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) auronitens cupreonitens Chevrolat, 1861
Female adult
NW-FRANCE,
Basse-Normandie Reg., Orne, Forêt de Bellême, 14.XII.2008, Pascal Stefani leg.
female, 26mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) auronitens montanus Gehin, 1882
Male+Female adult
SC-FRANCE,
Haute-Garonne, Col de Mentй h~1300m., II.1980. Kretschmer leg.
female, 23mm.
photo/det. Panin R.
Copyright © Ruslan Panin
Carabus (Chrysocarabus) lineatus lateralis Chevrolat, 1840 (as quasilineatus Tarrier, 1975)
Male+Female adult
NW Spain,
E Asturias, Monte de Moal h~1000m., 5.VI.2008, Pascal Stefani leg.
female, 27mm.
Copyright © Ruslan Panin
Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896
Male adult
Anyos (ANDORRE), h=1450ms,
5-VI-2010, P. Stefani leg.
body lenght - 32 mm.
Copyright © Alexandr Safronov
Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896
Habitat
Andorra, Anyos, ii.2012, J. Nicolas Leg. & Photos
Copyright © Noël Mal
Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792
Male adult
C-Poland, Łódź sity, Łagiewniki Forest, in dead logs of Birch, 18-30.XII.2013, D. Nawak leg., L=25 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) solieri bonnetianus Colas, 1936 (loc.typ.)
Female adult
SE-France, Var, Tanneron, Saint-Cassien des Bois, 18.IV.2019, L=32 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) solieri bonnetianus Colas, 1936 (loc.typ.)
Male adult
SE-France, Var, Tanneron, Saint-Cassien des Bois, 18.IV.2019, L= 31 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) auronitens kraussi (as vindobonensis Kubik, 1901)
Female adult
Lower Austria, Baden, Klausen-Leopoldsdorf, 15.XII.2024, M. Egger leg., L=30 mm
Copyright © Ruslan Panin -
Coptolabrus taxon page (181 photos)
Carabus (Coptolabrus) formosus brockmanni (as nili Deuve, 2005 PT) Paratype
Male+Female adult
C China, Shaanxi,
Liupan Mts., Guan Shan Mt. h=800 m, , 34:50N/106:48 E, Team V.Siniaev leg.
female, 38 mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Coptolabrus) formosus elytraviridis Kleinfeld, 2007 Holotype Paratype
Male+Female adult
China, Sichuan, Aba pref., Nanping Co., Jiuzhaigou (ca. 33.17N/103.54E), 2000 m, 30.5.-6.6.1990, coll. Kleinfeld.
HT 32 mm, PT 36 mm
Copyright © Frank Kleinfeld
Carabus (Coptolabrus) formosus grumorum Semenov, 1898
Female adult
female, 30mm
Copyright © Boleslav Brezina
Carabus (Coptolabrus) elysii elysii Thomson, 1856 (as idai Kleinfeld, 2000) Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Coptolabrus) augustus tenuiorithorax Deuve & Li, 2006
Male+Female adult
SE-China, SW-Jiangxi, Mt Guan, V.2009, local coll. female, 47 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) augustus augustus Bates, 1888
Male adult
SE-China, Jiangxi prov., Yudu County, Ganzhou city, Yun Fengzang h=1000m, 5-20.V.2011
L=43 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) gemmifer pseudoformosus Deuve, 2006
Female adult
China, Tibet AR, Qamdo, Gonjo County, Kè Rì Xiāng vill.(Kerri Township), h=3100m, VI.2012, A. Mollard leg. L=38 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) nankotaizanus Kano, 1932
Male adult
NE-Taiwan, Yilan County, V.2017, L=39 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) smaragdinus furumiellus Deuve, 1994
Female adult
NE-China, E-Liaoning, Benxi, Mt Liuershan, VIII.1998. L=39 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) fruhstorferi Roeschke, 1900
Male adult
Japan, Tsushima Isl., 20.IX.2018, S. Shinichi leg., L=35 mm
Copyright © Ruslan Panin -
Cratocarabus taxon page (5 photos)
-
Cratocechenus taxon page (38 photos)
Carabus (Cratocechenus) jacobsoni Semenov, 1898
Male adult
S Kazakhstan, Almaty reg., Big Lake, h=2800 m, 25.VI.96
male
L=14 mm
Copyright © Maksim Smirnov
Carabus (Cratocechenus) akinini akinini Morawitz, 1886 (as oitalensis ) Paratype
Male adult
male, 20mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) akinini akinini Morawitz, 1886
Female adult
female, 21mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) ovtchinnikovi ovtchinnikovi Gottwald, 1987
Male adult
N-Kyrgyzstan, Kyungey Ala-Too Mt Rng, Cholpon-Ata env., h=2500m, 19.VI.2000,
L=19 mm
Copyright © Ruslan Panin -
Cratocephalus taxon page (17 photos)
Carabus (Cratocephalus) solskyi Ballion, 1878
Male adult
SE Kazakhstan, Ketmen Mt. rng., Tuyuk vill., 16.06.97 leg. S.Toropov
Copyright © Maksim Smirnov
Carabus (Cratocephalus) solskyi Ballion, 1878
Female adult
SE-KAZAKHSTAN, Ketmen Mt Rng, h=1800m, Tujuk vill.,15.V.2004
L=36 mm
Copyright © Ruslan Panin
Carabus (Cratocephalus) cicatricosus cicatricosus Fischer, 1842
Male adult
SE-Kazakhstan, Almaty reg., C Trans-Ili Alatau N-slopes, Kaskelen Valley, Izvestkovyi vill., h=1700 m, 6.V.2015, D.Kovalchuk leg., L=28.5 mm
Copyright © Ruslan Panin -
Cratophyrtus taxon page (15 photos)
Carabus (Cratophyrtus) turcosinensis medvedevi Kryzhanovskij, 1968
Female adult
S Kazakhstan, NW slope of Ugamskii mt.rng., 3 km W Sairamskii peak, 2200m
tributary of Karzhantau riv., 15.6.2004 Matalin A. leg.
20 mm
Copyright © Alexander Anichtchenko
Carabus (Cratophyrtus) kaufmanni segregatus Morawitz, 1886
Female adult
female, 19mm
Copyright © Boleslav Brezina -
Cryptocarabus taxon page (16 photos)
Carabus (Cryptocarabus) subparallelus Ballion, 1878
Female adult
Kazakhstan,
near Alma-Ata, V.2009, Pak O.leg.
female, 17mm
Copyright © Ruslan Panin
Carabus (Cryptocarabus) lindemanni Ballion, 1878
Female adult
SE Kazakhstan, Alma-Ata env., 16.04. 2000
L=16,0 mm
Copyright © Maksim Smirnov
Carabus (Cryptocarabus) ? turkestanus turkestanus Breuning, 1929
Male adult
Kyrgyzstan, W Kungei Ala-Too Mt Rng, Chon-Kemin riv., Kyzyl-Bairak vill., 25.IV.2010. L= 18 mm
Copyright © Ruslan Panin
Carabus (Cryptocarabus) tsharynensis Kabak 1994
Male adult
SE-Kazakhstan, Almaty Region, W-Zailiyskiy Alatau rng, Belshabdar valley h~1300 m, 25-27.VI.2002, A.Puchkov leg. L=17 mm
Copyright © Ruslan Panin
Carabus (Cryptocarabus) tsharynensis Kabak 1994
Female adult
E-Kazakhstan, Almaty Region, W-Zailiyskiy Alatau, Turgen distr., Tauturgen vill., h~1200 m, 8-13.VI.2002, A.Puchkov leg. L=19 mm
Copyright © Ruslan Panin
Carabus (Cryptocarabus) sacarum Kabak, 1998
Male adult
SE-Kazakhstan, Almaty reg., C-Zailiyskiy Alatau N-slopes, Kaskalen river Valley, Izvestkovy vill., h=1700 m, 6.V.2015, D.Kovalchuk leg./det. R.Panin L=18 mm
Copyright © Ruslan Panin
Carabus (Cryptocarabus) sacarum Kabak, 1998
Female adult
SE-Kazakhstan, Almaty reg., C-Zailiyskiy Alatau N-slopes, Kaskalen river Valley, Izvestkovy vill., h=1700 m, 6.V.2015, D.Kovalchuk leg./det. R.Panin L=20 mm
Copyright © Ruslan Panin -
Ctenocarabus taxon page (8 photos)
Carabus (Ctenocarabus) galicianus Gory, 1839
Male adult
Spain,
Sierra de la Cabrera Baja, 8 km SW Truchillas, 9.VII.2008, Anichtchenko A. leg.
L= 20 mm
Copyright © Alexander Anichtchenko
Carabus (Ctenocarabus) galicianus Gory, 1839
Female adult
NW-Spain, Pontevedra prov., Campo Lameiro env., 15.VIII.1994, S.Sobrino leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Ctenocarabus) melancholicus costatus Germar, 1824
Female adult
C-Spain, Madrid region, Rozas de Puerto Real env.,21.XII.2008, T.Gazurek leg.
L=27 mm
Copyright © Ruslan Panin
Carabus (Ctenocarabus) melancholicus melancholicus Fabricius, 1798
Male adult
NW Morocco, Kénitra, Souk El Arbaa env., II.1985, V.Olivella leg. L=25 mm
Copyright © Ruslan Panin
Carabus (Ctenocarabus) melancholicus submeridionalis Breuning, 1975
Male adult
S- Spain, Cádiz, Tarifa, El Cabrito, 28.XII.2019, F. Garcia leg. L=26 mm
Copyright © Ruslan Panin
Carabus (Ctenocarabus) melancholicus costatus Germar, 1824
Male adult
NW Portugal, Viana do Castelo distr., Sierra de Agra, 41°47'52.3"N 8°41'27.6"W, h=786 m, 3.VII.2016, D. Prunier leg.
Copyright © Ruslan Panin -
Cupreocarabus taxon page (45 photos)
Carabus (Cupreocarabus) kangdingi kangdingi Korell, Kleinfeld & Gorgner, 1992 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Cupreocarabus) huangi huangi Deuve, 1992
Male adult
C-China, E Tibet, N-Sichuan, Hongyuan,valley 10 km SE Sanggarpar, h=4200m, 32.300053, 102.584031, alpine meadow / scree, 19.VII.1995, B.Brezina leg.<br /> L=21 mm
Copyright © Ruslan Panin
Carabus (Cupreocarabus) lixianensis lixianensis Deuve, 1990
Male adult
China, E Tibet, N-Sichuan, (Barkam),valley SW of Barkam, h=3200m, 31*53’ N; 102*12’E, mixed/coniferous forest, 22.VII.1995, K.& B.Brezina
L=23 mm
Copyright © Ruslan Panin
Carabus (Cupreocarabus) alpherakii alpherakii Semenov, 1898
Female adult
China, W Sichuan, Litang env., h~5000 m,29.VII -3.VIIII.1995, J. Schneider leg. L=23 mm
Copyright © Ruslan Panin
Carabus (Cupreocarabus) ? siguniangensis jiachingensis Lassalle, 2003
Male adult
Chine, Sichuan, Lushan, Mt. Jiajin Shan, 10.VII.2016 local collector leg., L=21 mm
Copyright © Ruslan Panin
Carabus (Cupreocarabus) balangicus balangicus Cavazzuti, 2002
Male adult
Chine, C-Sichuan, Xiaojin Co, Wolong valley, Balang Shan Pass h=3448 m, 30.884617, 102.973117, 9.VI.-1.VII.2011, G. Proscia leg. L=19 mm
Copyright © Ruslan Panin
Carabus (Cupreocarabus) balangicus balangicus Cavazzuti, 2002
Male adult
Chine, C-Sichuan, Xiaojin Co, Wolong valley, Balang Shan Pass h=3448 m, 30.884617, 102.973117, 9.VI.-1.VII.2011, G. Proscia leg. L=20 mm
Copyright © Ruslan Panin
Carabus (Cupreocarabus) huangi huangi Deuve, 1992
Male adult
China, E Tibet, N-Sichuan, 53 km NE of Barkam, Pass Sandaoban env., h=4293 m, 32.232950, 102.598533, 11-29.VI.2011, G. Proscia leg., L=21 mm
Copyright © Ruslan Panin
Carabus (Cupreocarabus) huangi huangi Deuve, 1992
Female adult
China, E Tibet, N-Sichuan, 53 km NE of Barkam, Pass Sandaoban env., h=4293 m, 32.232950, 102.598533, 11-29.VI.2011, G. Proscia leg., L=21.5 mm
Copyright © Ruslan Panin -
Cychrostomus taxon page (8 photos)
Carabus (Cychrostomus) pseudoprosodes Semenov & Znojko, 1932
Male adult
male, 19mm
Copyright © Boleslav Brezina
Carabus (Cychrostomus) anchocephalus lanzhouensis Deuve, 1991 Paratype
Male adult
male, 21mm
Copyright © Boleslav Brezina
Carabus (Cychrostomus) anchocephalus anchocephalus Reitter, 1896
Male adult
male, 22mm
Copyright © Boleslav Brezina -
Cyclocarabus taxon page (14 photos)
Carabus (Cyclocarabus) aulacocnemus galinae Obydov, 2003 Paratype
Male+Female adult
W-Tianshan:Karatau Mts.,N-part:Boialdyr River valley,
N of Ken-su, 600-750m, 22-V-2000 М. Danilevsky leg.
male - 20 мм, female - 19 мм.
Copyright © Alexandr Safronov
Carabus (Cyclocarabus) namanganensis namanganensis Heyden, 1886
Male+Female adult
Kazakhstan, S. Karatau Mts. R., Mashat vill. env.
23-IV-1993 М. Danilevsky leg.
male - 21 мм, female - 22 мм.
Copyright © Alexandr Safronov
Carabus (Cyclocarabus) zarudnyi Semenov & Znojko, 1932
Female adult
NE-Uzbekistan, W-Tian-Shan, NW slopes Pskem Mt. Rng, 8-14 km E Pskem, 2100-2500 m, 16.VI.1998, A. Puchkov leg., L=21 mm
Copyright © Ruslan Panin
Carabus (Cyclocarabus) karaterekensis hemicallisthenes Kabak & Kalab, 1997
Male adult
NW- Kyrgyzstan, W-Tian-Shan, Chandalash Mt.Rng, Chakmak-Suu vill., 3.VI.2000, L=17 mm
Copyright © Ruslan Panin
Carabus (Cyclocarabus) brosciformis Semenov, 1896
Male adult
NE-Uzbekistan, W-Tian-Shan, Chatkal Mt Range, Tashkent reg., Yangiabad env., h=2100m, 30.IV.2002, L=23.5 mm
Copyright © Ruslan Panin -
Damaster taxon page (15 photos)
Carabus (Damaster) blaptoides oxuroides Schaum, 1862
Female adult
Japan, C-Honshu, Ofuse-cho, Kamitakai, Nagano Pref., 12.VI.2006, Ohshima leg.
L=34 mm
Copyright © Ruslan Panin
Carabus (Damaster) blaptoides capito Lewis, 1881
Male adult
Japan: Niigato pref., Sado Isl., Sado, Yamato
Copyright © Igor Solodovnikov
Carabus (Damaster) blaptoides oxuroides Schaum, 1862
Female adult
Japan, C Honshu, Nagano Prefecture, Nagano city, 3.IV.2012, S. Shinichi leg. L=42 mm. det. Anichtchenko A. 2021
Copyright © Ruslan Panin
Carabus (Damaster) blaptoides rugipennis Motschulsky, 1861
Male adult
Kunashir isl.
Copyright © Kirill Makarov
Carabus (Damaster) blaptoides blaptoides (Kollar, 1836)
Female adult
Japan, SW Honshu, Shimane Pref., Unnan city, 16.I2016, S.Shinichi leg., L=48 mm
Copyright © Ruslan Panin -
Deroplectes taxon page (32 photos)
Carabus (Deroplectes) staudingeri Ganglbauer, 1886
Female adult
N Tajikistan, SW Pamir-Alay, Fann Mts h=1800m, Padrut vill., 2-3.V.1997
L=28 mm
Copyright © Ruslan Panin
Carabus (Deroplectes) dokhtouroffi dokhtouroffi Ganglbauer, 1886
Male adult
NC Tadzhikistan, Pamir, N-slope Peter I.Mt Rng, h=2100m,Ganishou vill., 10-12.VI.2006, Ivanova E. leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Deroplectes) klapperichianus coiffaitianus Deuve, 1990
Male adult
Tadzhikistan: Badakhshan: Ishkashim Mt.R.: Kuh-i-Lal, VI.1979 leg. I.K. Lopatin (TOPOTYPUS)
Copyright © Igor Solodovnikov
Carabus (Deroplectes) klapperichianus coiffaitianus Deuve, 1990
Female adult
Tadzhikistan: Badakhshan: Ishkashim Mt.R.: Kuh-i-Lal, VI.1979 leg. I.K. Lopatin (TOPOTYPUS)
Copyright © Igor Solodovnikov
Carabus (Deroplectes) klapperichianus salangicus Kleinfeld & Reuter, 2010 Paratype
Female adult
N-Afghanistan, Hindukush Mt.Rng, Salang Pass, Nordseite, 25°27'.494N, 68°58.399'E, ca. reg. subalp., h = 2200-2400 m, V.2010, leg. Christoph Reuter
Copyright © Igor Solodovnikov -
Diocarabus taxon page (45 photos)
Carabus (Diocarabus) slovtzovi slovtzovi Mannerheim, 1849
Male adult
S Siberia, Burjatia reg., E Sayan Mts., Khugaituma mt., h=1700 m, 1-5. VII.2001
L=14,3 mm
Copyright © Maksim Smirnov
Carabus (Diocarabus) linnaei linnaei Panzer, 1812
Male adult
W-Ukraine, NW podolian, <br /> Lviv Reg., Zhydachiv distr., near Vybranivka, 17.II.2007,Panin R. leg.<br /> female, 21 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) massagetus massagetus Motschulsky, 1846 (alpin form)
Male adult
Russsia, Altay, Ukok upland, Kaldzhin-Kul Lake h=2400-2550 m, 49 *19*N/87*26*E, 16-27.VII.2008, R.Dudko leg. L=17.5 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) linnaei linnaei (as scopolii Dejean, 1826)
Male adult
W Ukraine, Lviv reg., NW Podolian Upland, forest near Vybranivka vill., 12.X.2007, Panin R. leg. L=19 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI.-5,VIII.2017, R. Panin R. & Yu.Kanarsky leg.
Copyright © Ruslan Panin
Carabus (Diocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=16.5 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=17 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg.
Copyright © Ruslan Panin
Carabus (Diocarabus) fraterculus neochinensis Deuve & Li, 1998
Female adult
NE-China: Liaoning prov., Fengcheng City, Shicheng, 20-28.VI.2011, Li Jingke leg., L=22 mm
Copyright © Ruslan Panin -
Eccoptolabrus taxon page (22 photos)
Carabus (Lasiocoptolabrus) sunwukong sunwukong Imura, 1993 Paratype
Male adult
Copyright © Boleslav Brezina
Carabus (Lasiocoptolabrus) sunwukong baotianmanus (Imura, 2005) Paratype
Male adult
Copyright © Boleslav Brezina
Carabus (Eccoptolabrus) exiguus nivium Breuning, 1933
Female adult
female, 20mm
Copyright © Boleslav Brezina
Carabus (Eccoptolabrus) exiguus exiguus Semenov, 1898 Holotype
Male adult
male, 18 mm
Copyright © Frank Kleinfeld -
Eucarabus taxon page (111 photos)
Carabus (Eucarabus) obsoletus carpathicus Palliardi, 1825
Male+Female adult
female, 25 mm
SW-Romania, Banat, Resita - Valiug, 3-8.VI.2007, Krause R. leg.
Copyright © Ruslan Panin
Carabus (Eucarabus) obsoletus obsoletus Sturm, 1815
Male adult
NW Ukrainian Carpathians, Lviv Reg., Staryi Sambor Distr., near Dobromyl , h=500 m., 15-24.VI.2005, Panin R. leg., male, 28mm
Copyright © Ruslan Panin
Carabus (Eucarabus) manfredeggeri Müller 2012 Paratype
Male+Female adult
N-Korea, Chagang Prov., Manpo City, Songhak-ri, 2-16.VII.2011<br /> female, 21 mm<br /> male, 19 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825
Female adult
SW Romania, Banat - Eibenthal; 33 mm
Copyright © Aleš Sedláček
Carabus (Eucarabus) ulrichi arrogans Schaum, 1858
Male adult
NE Serbia, Majdanpek; 23 mm
Copyright © Aleš Sedláček
Carabus (Eucarabus) obsoletus obsoletus (as csikii Mallasz, 1901)
Female adult
NE Carpathians, Marmarosh Mts., Transcarphatian Reg., near Lug vill., h=350 m, 26.IV-8.VI.2014, R.Panin & Yu. Kanarsky leg. L=32 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) parreyssi parreyssi Palliardi, 1825
Male adult
W-Croatia, NP Northern Velebit, Zavižan,15.V-13.VI.1996, Colombetta leg., L=21 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) billbergi kuandianicus Rapuzzi, 2007
Female adult
NE China, Liaoning, Kuandian County, Dachuantou env., 20-29.VI.2015,
Copyright © Ruslan Panin
Carabus (Eucarabus) billbergi kuandianicus Rapuzzi, 2007
Male adult
NE China, Liaoning, Kuandian County, Dachuantou env., 20-29.VI.2015, L=23 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) catenulatus fontanellae Reitter, 1896
Male adult
W-Croatia, Dalmatia, Isl. Rab, 16-30.V.2019, G. Proscia leg.
Copyright © Ruslan Panin -
Euleptocarabus taxon page (5 photos)
Carabus (Euleptocarabus) porrecticollis porrecticollis Bates, 1883
Male adult
Japan: Niigata pref., Chayagahara Jyouetsu City,
18.XI.2006 A.Toba leg.
Copyright © Alexander Anichtchenko
Carabus (Euleptocarabus) porrecticollis Bates, 1883
Female adult
Atsushiokanomachikano, Kitakata City, Fukushima Pref., Japan, 29.III.2014, T. Nakamura leg.
Copyright © Alexander Anichtchenko
Carabus (Euleptocarabus) porrecticollis porrecticollis Bates, 1883
Male adult
Japan, C Honshu, Nagano Pre., Toyooka town, 28.XII.2019, S. Shinichi leg. L=30 mm
Copyright © Ruslan Panin -
Eupachys taxon page (3 photos)
Carabus (Eupachys) glyptopterus Fischer, 1827
Male+Female adult
Russia, SE-Siberia,
Buryat Republic, Ivolginsky Distr., 04.VII.2010, Filippov A.E. leg.
female, 32 mm
photo Panin R.
Copyright © Ruslan Panin -
Eurycarabus taxon page (29 photos)
Carabus (Eurycarabus) famini lucasi Gaubil, 1849
Male adult
N-Tunisia, Bizerte reg., Menzel Bourguiba city,h=5 m, XII.1981, S. Remond leg.
L=25 mm
Copyright © Ruslan Panin
Carabus (Eurycarabus) famini fezzanus Lapouge, 1915
Male adult
Morocco, Middle Atlas Mt Rng, Azrou city h=1500m, IV.1999, S. Remond leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Eurycarabus) famini piraticus (as mamorensis (Kocher, 1964)
Female adult
NW-Morocco, betw.Meknes/Kenitra, Foret de Mamora, II.2008
L= 27 mm
Copyright © Ruslan Panin
Carabus (Eurycarabus) famini atlantis Alluaud, 1923
Female adult
C Morocco, Haut Atlas, SW of Oukaïmeden h=2900m, 31.173644, -7.844442, 29.IV-11.V.2024, C.Reuter leg., L=21 mm
Copyright © Ruslan Panin -
Fulgenticarabus taxon page (2 photos)
Carabus (Fulgenticarabus) flutschi koreus Deuve, 2006
Male adult
N Korea, Samjiyon, Ryang Gang Province
6-12.6.2006 leg. Wong
Copyright © Boleslav Brezina -
Gnathocarabus taxon page (2 photos)
Carabus (Gnathocarabus) kusnetzovi Semenov, 1903
Female adult
female, 21mm
Copyright © Boleslav Brezina -
Goniocarabus taxon page (14 photos)
Carabus (Goniocarabus) sogdianus Semenov, 1898
Male adult
N Tajikistan, SW Pamir-Alay, Zarafshan Range, Fann Mts, Marguzor Lake circ., Padrut vill., 16.V.1997
L=21 mm
Copyright © Ruslan Panin
Carabus (Goniocarabus) grombczewskii grombczewskii Semenov, 1891
Male adult
NC Tadzhikistan, Pamir, N-slope Peter I.Mt Rng, h=2100m, Ganishou vill.,12.VII.2006, Pak O. leg.
L=19 mm
Copyright © Ruslan Panin
Carabus (Goniocarabus) perelloi Casale, 1979
Male adult
N-Afghanistan, Hindukush, Provinz Baghlan, N of Kabul, N Salang Pass, subalp., h = 2200 m, 35°27'.494N / 68°58.399'E, Mai 2011, leg. Christoph Reuter (TOPOTYPUS)
Copyright © Igor Solodovnikov -
Hemicarabus taxon page (31 photos)
Carabus (Hemicarabus) tuberculosus Dejean, 1829
Male adult
Far East, Lazovsky Dist., shore of Japan Sea, 3 km NE Preobrazhenie Vill., Ezhovaya (Tasovaya) Bay 13.07.06 M. & L. Smirnov's leg.
male
L=20,0 mm
Copyright © Maksim Smirnov
Carabus (Hemicarabus) nitens Linne, 1758
Male+Female adult
Belarus,
Vitebsk reg., Ushachskiy distr., near Bolshie Dolcy vill. 9.V.1990, I.A.Solodovnikov leg.
female, 18mm.
Copyright © Ruslan Panin
Carabus (Hemicarabus) serratus Say, 1825
Male adult
E-USA,Virginia, Leesburg env., Ball's Bluff Battlefield Regional Park, 27.VIII-4.IX.2015,V. Čutka leg. L=17 mm
Copyright © Ruslan Panin
Carabus (Hemicarabus) serratus Say, 1825
Female adult
E-USA, Virginia, Leesburg env., Ball's Bluff Battlefield Regional Park, 27.VIII-4.IX.2015,V. Čutka leg. L=18 mm
Copyright © Ruslan Panin -
Homoeocarabus taxon page (6 photos)
Carabus (Homoeocarabus) maeander maeander s.str.
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Carabus (Homoeocarabus) maeander paludis Gehin, 1885
Female adult
Japan, Hokkaido; Tomakomai city, Benten, 17.V.2015, Takemoto leg., L=21 mm
Copyright © Ruslan Panin -
Hygrocarabus taxon page (5 photos)
Carabus (Hygrocarabus) nodulosus Creutzer, 1799
Male+Female adult
Austria, Styria Region (Steiermark), Leibnitz, 31.X.1999, I.Rapuzzi leg.<br /> female, 29 mm
Copyright © Ruslan Panin
Carabus (Hygrocarabus) variolosus Fabricius, 1787
Male adult
W-Ukraine, NW Podolian,<br /> Lviv reg., Zhydachiv distr.,near Vybranivka vill., 15.V-8.VI.2008, Panin R. leg.<br /> male, 27mm
Copyright © Ruslan Panin
Carabus (Hygrocarabus) variolosus Fabricius, 1787
Male+Female adult
W-Ukraine,<br /> Lviv Reg., Zhydachiv distr., near Vybranivka vill., Panin R. leg.<br /> female, 30mm
Copyright © Ruslan Panin -
Imaibius taxon page (51 photos)
Carabus (Imaibius) baronii battalensis Heinz & Staven, 1998 Paratype
Male adult
Copyright © Alexander Anichtchenko
Carabus (Imaibius) dardiellus perobscurus Deuve, 1980
Male adult
N India.Kashmir, Loran, W part Pir Panchal range,h=2400-2500m, 33°49'"N, 74°20'E
L=25 mm
Copyright © Ruslan Panin
Carabus (Imaibius) dardiellus ? subpunctulus Roeschke, 1907
Male adult
N India, Kashmir, Pahalgam, Aru Valley h=2400m, VII.2007
L=25 mm
Copyright © Ruslan Panin
Carabus (Imaibius) wittmerorum Heinertz, 1978
Male adult
N. Pakistan: Frontier prov., Miandam, Swat vall., h=1800-2600 m, leg.S.Remond (TOPOTYPUS)
Copyright © Igor Solodovnikov
Carabus (Imaibius) barysomus hazarensis Deuve, 1982 (TOPOTYP)
Male adult
N-Pakistan: Mansehra prov., Hazara, Kaghan valley: Shocgran env., h = 2750 m L= 42 mm
Copyright © Igor Solodovnikov
Carabus (Imaibius) boysi boysi Tatum, 1851
Male adult
NW-India, Himachal Pradesh, Manikaran env., Pulga vill., 4.VII.2008. L=26 mm
Copyright © Ruslan Panin
Carabus (Imaibius) stoliczkanus brauni Heinz, 1993
Female adult
NE Pakistan, Azad Kashmir, Bagh Distr., Sudhan-Gali h~2300 m,15.VIII.1994, S Dacatra leg. L=37 mm
Copyright © Ruslan Panin
Carabus (Imaibius) caschmirensis fortetuberculatus Heinz, 1983 Paratype
Male adult
Copyright © Alexander Anichtchenko
Carabus (Imaibius) nouristani nouristani (Ledoux, 1977)
Female adult
NE-Afghanistan: Nuristan: Darah-I-Nur valley, 2900m 34°45'N 70°40'E, VIII.2012, C.Reuter leg., L=26 mm
Copyright © Ruslan Panin
Carabus (Imaibius) nouristani nouristani (Ledoux, 1977)
Male adult
NE-Afghanistan: Nuristan: Darah-I-Nur valley, 2900m 34°45'N 70°40'E, VIII.2012, C.Reuter leg., L=26 mm
Copyright © Ruslan Panin -
Iniopachus taxon page (7 photos)
Carabus (Iniopachus) pyrenaeus jeanneli Deuve, 1994
Male adult
1-2: Pirineos, Canfranc, Ibon de Iserias 2200 m 16-17.7.05<br /> 3: Pirineos, Monte Perdido, Collata Anisolo 2400m 25.8.05
Copyright © Alexander Anichtchenko
Carabus (Iniopachus) auriculatus ubinensis (Puissegur, 1955)
Female adult
Spain, Montes Cantabricos, Peña Ubiña, 2400m
5-6.VII.2009 Anichtchenko A. leg.
female, 19 mm
Copyright © Alexander Anichtchenko
Carabus (Iniopachus) pyrenaeus pyrenaeus (as cephalicus Csiki, 1927)
Male adult
SW France, Pyrenees-Atlantiques, Mt.Cirque d'Esturou h=1800m, 29.VIII.2014, Pascal Stefani leg. L=20 mm
Copyright © Ruslan Panin -
Ischnocarabus taxon page (4 photos)
Carabus (Ischnocarabus) tenuitarsis tenuitarsis Kraatz, 1877
Male adult
male, 24mm
Copyright © Boleslav Brezina -
Isiocarabus taxon page (24 photos)
Carabus (Isiocarabus) fiduciarius fiduciarius (as fiduciarius tim) Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Isiocarabus) hunanicola liangshanicus (as dreuxi liangshanicus) HT PT Kleinfeld, 1998 Holotype Paratype
Male+Female adult
Carabus (Isiocarabus) hunanicola liangshanicus (as dreuxi liangshanicus) HT PT
Copyright © Frank Kleinfeld
Carabus (Isiocarabus) miaorum Lassalle & Prunier, 1993
Male adult
S-China, N-Guangxi, Mao'er Shan, h~1600 m, V.2009, L= 26 mm
Copyright © Ruslan Panin
Carabus (Isiocarabus) fiduciarius fiduciarius Thomson, 1856
Female adult
CE-China, Hubei, Xiaogan City, Shuangfeng Mountain, 26.VI.2016, L=30 mm
Copyright © Ruslan Panin
Carabus (Isiocarabus) miaorum Lassalle & Prunier, 1993
Male adult
China, NW-Guangxi, Mao'er Shan, h~1600 m 25°52'N 110°29'E, 28.V-9.VI.2009, C.Reuter leg.
Copyright © Ruslan Panin -
Lamprostus taxon page (31 photos)
Carabus (Lamprostus) calleyi pseudoprasinus Lapouge, 1912
Male+Female adult
Transcaucasia, Armenia,
Ashotsk distr. Tsoghamarg vill, .21-22.V.1997, P.V.Kazaryan leg
female, 24 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) torosus torosus Frivaldsky, 1835
Male adult
NE-Bulgaria, Dobrich Region, Kavarna env.
L=31 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) saulcyi saulcyi Piochard, 1875 (as chambouqensis)
Female adult
N-Lebanon, Fnaпdek, Djebel Qammoua, h=1100-1200m, 29.X.2012, C. Reuter leg.
L=27 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) saulcyi saulcyi Piochard, 1875
Male adult
C Lebanon, Jbeil Distr., Ehmej vill., h=1100 m.V.2012, C. Reuter leg.
L=24 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) hemprichi elonensis Schweiger, 1970
Male adult
N-Israel, Midlle Galilea, Nazareth env., Hoshaya viil., 3-12..III.2015, B.Vas’ko leg./det. Panin R.
Copyright © Ruslan Panin
Carabus (Lamprostus) syrus cheikensis Deuve, 1992
Male adult
N-Israel, Midlle Galilea, Nazareth env., Hoshaya viil., 13.XII.2012, Vas’ko, leg./det. Panin R. L=30мм
Copyright © Ruslan Panin
Carabus (Lamprostus) robustus triumphans Cavazzuti, 1987
Male adult
NE-Turkey:Trabzon, Tonya env., VIII.2004. L=41 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) calleyi pseudoprasinus Lapouge, 1912
Male adult
S Georgia, Tsalka (Paravani) 41°29′18.7″N, 43°51′12.3″E, 8-9.V.2016, Pavel Turek leg. / det. I. Solodovnikov, L=23 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) syrus moukarzelae Deuve, 2002
Female adult
S-Lebanon: Marjaayoun, 18.XI.-11.XII.2017, C. Reuter leg., L=35 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) syrus cheikensis Deuve, 1992
Female adult
SE Lebanon, Rashaya Distr., Tannoura, oak forest, h=700 m, 11-19.IV.2016, C. Reuter leg., L= 32 mm
Copyright © Ruslan Panin -
Leptocarabus taxon page (46 photos)
Carabus (Leptocarabus) yokoae marcilhaci Deuve, 1992
Male adult
CHINA, Shanxi, Tschingling Mts., Taibeichan Nat. reserv.;
Copyright © Anton Menstschikov
Carabus (Leptocarabus) harmandi fujisan Ishikawa, 1972
Female adult
Japan, Yamanashi pref., Fuji-Rinds 1600m, Narusawa vill. 3.V.2005 Kurihara K. leg. 25 mm
Copyright © Alexander Anichtchenko
Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931
Male adult
NE-China:Liaoning prov., Fengcheng City, Miaoyang, 12-17.VI.2011<br /> L=26 mm
Copyright © Ruslan Panin
Carabus (Leptocarabus) procerulus Chaudoir, 1862
Male adult
Japan:S Honshu, Matsubara-ko, VIII.1997, S. Remond leg.
L=30 mm
Copyright © Ruslan Panin
Carabus (Leptocarabus) procerulus Chaudoir, 1862
Male adult
Japan, S Honshu, Mie Pref., Hakusan, VII.2001, S. Remond leg.<br /> L=33 mm
Copyright © Ruslan Panin
Carabus (Leptocarabus) arboreus pararboreus (Ishikawa, 1992)
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Carabus (Leptocarabus) arboreus fujisanus Bates, 1883
Male adult
Japan, CE-Honshu, Yamanashi pref., Mt. Fujisan h~1600m, 23.IX.1982, Y. Imura leg. 23 mm
Copyright © Ruslan Panin
Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931
Male adult
NE-China:E-Liaoning: NP Kuandian Tianqiaogou , VI.2010, L=25 mm, det. Panin R.
Copyright © Ruslan Panin
Carabus (Leptocarabus) arboreus fujisanus Bates, 1883
Female adult
Japan, Honshu, Mt. Fuji h~1900m, Shizuoka pref., Fujinomiya city, 23.VI.2007, M. Matsunaga leg., L=25 mm
Copyright © Ruslan Panin -
Leptoplesius taxon page (6 photos)
Carabus (Leptoplesius) merzbacheri kirgisiensis Mandl, 1967
Female adult
22mm
Copyright © Boleslav Brezina -
Limnocarabus taxon page (15 photos)
Carabus (Limnocarabus) clathratus clathratus Linne, 1761
Female adult
W-Ukraine, Maloe Polesie, Lviv Region, near Kamenka-Bugzkaya, 28.III.2008, Panin R. leg. L=31 mm
Copyright © Ruslan Panin
Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
Female adult
SW Slovakia, Bratislava env., IV.2004, Krause Rene leg.
L=32 mm
Copyright © Ruslan Panin
Carabus (Limnocarabus) clathratus clathratus (as jansoni Kraatz, 1890)
Male adult
W-Ireland, County Mayo, Bellacorick bog, 9.VII.2013, J.Stypula leg. L=27 mm
Copyright © Ruslan Panin
Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
Male adult
W Ukraine, Transcarpathian Reg., 4 km NE Chop, forest Peresh, Panin R. leg. L=30 mm
Copyright © Ruslan Panin -
Lipaster taxon page (18 photos)
Carabus (Lipaster) stjernvalli stjernvalli Mannerheim, 1830
Male+Female adult
Armenia,
Ashotsk, 22.VI.1990, P.V.Kazaryan leg.
female 34mm
Copyright © Ruslan Panin
Carabus (Lipaster) stjernvalli osellai Breuning & Ruspoli, 1970
Male+Female adult
NE-Turkey: Artvin:Hopa, Borçka env., h=750 m, 9.VI-7.VII.2011, M.Janalik leg.
female, 42 mm
Copyright © Ruslan Panin
Carabus (Lipaster) stjernvalli osellai Breuning & Ruspoli, 1970
Male adult
NE-Turkey: Artvin, Hopa / Çifteköprü, VII.1977, L=36 mm
Copyright © Ruslan Panin
Carabus (Lipaster) stjernvalli stjernvalli Mannerheim, 1830
Male adult
SC Georgia, Tsalka distr. 7 km NE of Paravani Lake, 41.540750, 43.879806, h=2100 m, 12-16.VII.2025, I.Uspenskyi leg., L=25 mm
Copyright © Ruslan Panin -
Macrothorax taxon page (26 photos)
Carabus (Macrothorax) morbillosus alternans Palliardi, 1825
Male adult
S Italy, Sicily, Province Palermo, Piana degli Albanesi h~700m, XI.2010, I.Rapuzzi leg.
L=32 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) aumonti maroccanus Bedel, 1895
Male adult
N Morocco (Spain:Melilla), hipodromo, X.1969, leg. Pardo Alcaide
L=38 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) rugosus boeticus Deyrolle, 1852
Male adult
SC Spain, Jaen reg., Puente de la Sierra, 10.VII.2011, M.Lopez leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) rugosus rugosus Fabricius, 1792
Male adult
N Morocco, El Biutz –Taghramt,20.I.2005, Manfred Egger leg.
L=29 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) morbillosus constantinus Kraatz, 1899
Male adult
NW Tunisia; Tebarka, E Sejenane,26.I.2004, Manfred Egger leg.
L=33 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) rugosus celtibericus Germar, 1824
Female adult
SW-Portugal, Setúbal Distr., Cercal do Alentejo/ Tanganheira vill. 8.XII.2006, T.Gazurek leg.
L=31 m
Copyright © Ruslan Panin
Carabus (Macrothorax) morbillosus constantinus (as galloprovincialis Lapouge, 1910)
Female adult
SE-France, Var, near Le Muy, VIII.1980, Michon leg.
L=31 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) morbillosus lampedusae Born, 1925
Male adult
S Italy, Isl. Lampedusa, Baia dei Conigli, VI.2003, L=29 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) rugosus laufferi Breuning, 1927
Female adult
C-Spain: Avila, El Hornillo env., 27.V.2017, J. Ferrero leg. 30 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) rugosus celtibericus Germar, 1824
Male adult
SW-Portugal, Serra do Cercal h=217 m, 37°47'15.6"N 8°42'02.0"W, road N390, 19.III.2011, D. Prunier leg., L=27 mm
Copyright © Ruslan Panin -
Meganebrius taxon page (35 photos)
Carabus (Meganebrius) kadoudali (Morvan, 1981)
Male+Female adult
C-Nepal, SW Dhaulagiri, W Jaljala h=3150 m, 28°30'05"N, 83°11'48"E, 23.IX.2012, J. Schmidt leg.
female, 20 mm
Copyright © Ruslan Panin
Carabus (Meganebrius) pseudoharmandi Mandl, 1965
Female adult
E-Nepal, Kosi zone, Khandbari,(1.5km SW of Chichila vill., 27*45’843’’N/ 87*22’473’E, h=1940m), damaged primary forest, 13 .V.2010, B.Brezina & M.Häckel leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Meganebrius) arunensis Heinertz, 1980
Male adult
E-Nepal, Kosi zone, Khandbari, 1.5km SW of Chichila vill., 27*47’175’’N/ 87*23’622’E, h=1880m, damaged primary forest, 11.V.2010, B.Brezina & M.Häckel leg.
L=27 mm
Copyright © Ruslan Panin
Carabus (Meganebrius) quinlani smetanai Häckel, 2011 Paratype
Male+Female adult
C-Nepal:Dhaulagiri Massif:Baglung Distr., (1.5km W of Okhale vill., 28.32’152’’N/ 83.48’673’E, h=2600m) damaged primary forest, 19.V.2010, B.Brezina & M.Häckel leg.
male, 21mm
female, 23 mm
Copyright © Ruslan Panin
Carabus (Meganebrius) quinlani quinlani Mandl, 1965
Male adult
C-Nepal, Gandaki, Manang Distr., Timang env., h~2500-2600m, old open forest, grassland, 3-5.V.2007, Fouqué R. leg. L=19 mm
Copyright © Ruslan Panin
Carabus (Meganebrius) tuberculipennis ganjalaensis Deuve, 2000
Male adult
Nepal, Helambu, GanjaLa Pass S-slope, Dhukpu Kharka h=4100m, 28°03'50"N, 85°34'21"E, 12.VIII.2010, S.Tamang leg., L=21 mm
Copyright © Ruslan Panin -
Megodontoides taxon page (11 photos)
-
Megodontus taxon page (162 photos)
Carabus (Megodontus) violaceus andrzejuscii (as mehelyi Ganglbauer, 1896 loc. typ.)
Male+Female adult
N-Romania, Rodna Mts., Pasul Setref - Sacel 10km E, h=810 m, 8-19.VI.2009, Krause Rene leg. female, 31mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus fiorii Born, 1901
Male+Female adult
NW-Italy,
Reggio Emilia, Regnano, 5.IX.1996 leg. Cartini A.
female, 30 mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823
Male+Female adult
W. Ukraine, W. Podolian, Lviv reg., Nikolaev distr., near Krasiv vill., under the bark of oak tree, 8.II.2009, Panin R. leg. female, 29mm
Copyright © Ruslan Panin
Carabus (Megodontus) exaratus Quensel, 1806
Male+Female adult
NW Caucasus, Russia,
Krasnodar reg., Krasnodar City, 27.IX-07.X.07, D.D.Fominykh leg.
female,29mm/male, 27mm
Copyright © Ruslan Panin
Carabus (Megodontus) croaticus croaticus Dejean, 1826
Male adult
SW Slovenia, Ilirska Bistrica, Sviščaki, Snežnik,17-28.VI.2009, Manfred Egger leg.
L=32 mm
Copyright © Ruslan Panin
Carabus (Megodontus) stroganowi transfuga Semenov, 1898
Male adult
SE-Azerbaijan: Talysh, Kalvaz, 21-30.VI., Dolin V.G. leg.
L=32 mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus aurichalceus Kraatz, 1879
Male adult
NC-Spain, Cantabria, Santander, Cueto env.XII.2013 L=28 mm
Copyright © Ruslan Panin
Carabus (Megodontus) persianus javahardehensis Deuve, 2000
Male adult
N-Iran, Mazandaran Prov., Ramsar Co., Javahar Deh env., 16-25.VI.2017, Václav Čutka leg. L=28 mm
Copyright © Ruslan Panin
Carabus (Megodontus) obovatus Fischer, 1827
Endophallus
C.obovatus taizhensis Obydov, 2006 Khakassia: Kuznetskyi Alatau, env. vill. Priiskovyi, Bobrovaya Mt., h = 1200-1300 m
Copyright © Igor Solodovnikov -
Mesocarabus taxon page (150 photos)
Carabus (Mesocarabus) problematicus planiusculus Haury in Gehin, 1885
Male adult
NE Spain, Catalonia, Sant Jaume de Frontanyà, h=1000 m, V-VI.2010, Pascal Stefani leg.
L=24 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) problematicus harcyniae (as procedens Csiki, 1927)
Male adult
England, North Yorkshire, Fylingdales Moor, Grouse Hill, VII.2006 L=24 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) problematicus islandicus Lindroth, 1968
Female adult
SE-Iceland:Höfn env.:Hornafjördur, 24-28.VIII.2008, C. Reuter leg.
L=24 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) cantabricus cantabricus Chevrolat, 1840 (loc. typ.)
Male adult
NW-Spain, Pontevedra prov., A Cañiza viil., Alto de Fontefría h=800m, 31.XII.2007, T.Gazurek leg./det.Panin R.<br /> L=27 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) latus cf. molossoides Lassalle, 1984
Male adult
C-Spain, Toledo, Mts de Toledo, Sierra del Castañar Rng, Puerto del Milagro env, h=920m, 7.II.2009, T.Gazurek leg./det.R. Panin.
L=24 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826
Male adult
CW-Spain: Salamanca, Brincones env., h=750m, X.2003<br /> L=25 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) breuningi Csiki, 1927
Female adult
NW-Spain, Prov. Zamora, San Martín de Castañeda, 5.VII.1984, M.Toribio leg. L=28 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) breuningi Csiki, 1927 (loc.typ.)
Male adult
NW-Spain, Orense, Orensano Central Massif, Monte de Cabeza de Manzaneda, 25.VII.1988, F. Rodrigues leg. L=25 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) lusitanicus helluo (as alicantinus Ganglbauer, 1886)
Male adult
SE-Spain, Murcia reg., Jumilla, Sierra del Carche, h =1300 m, 30.XI.2019, F. Garcia leg.
Copyright © Ruslan Panin
Carabus (Mesocarabus) lusitanicus trabuccarius Fairmaire, 1857
Female adult
NE Spain, Catalonia, Tarragona, Sierra de Montsant h=800 m, Albarca env.,7.XI.2018, T. Rafales leg., L=27 mm
Copyright © Ruslan Panin -
Microplectes taxon page (14 photos)
Carabus (Microplectes) riedeli Menetries, 1832
Male adult
N-Ossetia,
Khazniaush Pass, h=2500-2700m, 9.VI-31.VII.1994, Zamotajlov A.leg.
male, 17mm
Copyright © Ruslan Panin
Carabus (Microplectes) convallium convallium Starck, 1889
Male adult
male, 19mm
Copyright © Boleslav Brezina
Carabus (Microplectes) convallium convallium Starck, 1889
Male+Female adult
female:NW-Caucasus, Russia,
Krasnodar reg., Apsheron Distr., Lagonaki Mt., near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominyh leg.
male:Karachaevo-Cherkessia,the right source of the river Beskess, h=2100m, 10.VIII.1997, I.A. Solodovnikov leg.
female, 21mm/male, 19mm
Copyright © Ruslan Panin
Carabus (Microplectes) convallium argonautarum Semenov, 1898
Male adult
male, 18mm
Copyright © Boleslav Brezina -
Mimocarabus taxon page (11 photos)
Carabus (Mimocarabus) maurus maurus Adams, 1817
Male+Female adult
Armenia, Ashotsk distr., Tsoghamarg vill., 24-27.V.1999 P.V.Kazaryan leg. female, 20mm/male,18mm
Copyright © Ruslan Panin
Carabus (Mimocarabus) pumilio Kuster, 1846
Male adult
S-Armenia, Vayots Dzor Region, E Ughedzor vill., h=2630 m, 9.VI.2014, L=16 mm
Copyright © Ruslan Panin
Carabus (Mimocarabus) maurus maurus (Adams, 1817)
Male adult
E Georgia, Kakheti, near Patardzeuli vill., h=750-850 m, 10.IV.2015, E. Rutjan leg., L=22 mm
Copyright © Ruslan Panin -
Morphocarabus taxon page (432 photos)
Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
Male adult
Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
S.Bohemia,
Hluboka nad Vltavou, 24.VI-18.VII.1990, S. Budka leg.
male, 27mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) spasskianus shoriensis Obydov, 1999
Male adult
Russia, Khakassia,
Abakanskii Mt Range, Askiz river vall., 7 km E Kazanovka vill., 15.VI.2006, A.& N. Menstschikovy leg.
L=24 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi hampei Kuster, 1846
Male adult
C Romania, Târgu Mureş, Ernei, 46°35′471″N 24°37’351″E, h=300 m,20.VIII.2004, Sl.Budka leg.
L=30 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878 (as viridipennis Kraatz, 1886, nec Lewis,18
Male+Female adult
Russia, Far East,Primorsky reg., S. Sikhote-Alin Mts. R., Kavalerovo vill. env.,
5-25.VII.1992, С. Kunitzyn leg.
male - 25 mm, female - 25 mm.
Copyright © Alexandr Safronov
Carabus (Morphocarabus) rothi hampei as telekii Csiki, 1937 (? hampei )
Female adult
CW Romania, Arad distr., Zarandulu Mt Range, near Ilteu vill., h=276 m, 46°66′487″N 22°20’455″E, 25.VI.2005, Sl.Budka leg.
L=30 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi (? incompsus as spectabilis Csiki, 1906)
Female adult
NC Romania, Bistrita Nasaud reg., Călimani Mts, Colibita vill., mixed forest, meadows h=800m, 15-21.V.2011, M.Pasek leg.<br /> L=34 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens ciscarpathicus (as polonicus Lomnicki, 1886)
Female adult
W Ukraine, Lviv Region, Zhydahciv distr., Borodchytsi vill.,the right bank of Dniester river, 11-16.VII.2012, Panin. R leg. L=27 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Female adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky forest, near Pyatnychany vill., 28.V-9.VI.2013, Panin R. leg. L=30 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) henningi oviformis Beheim & Breuning, 1943
Male adult
Russia, Polar Ural, Yamalo-Nenets reg.,Harp station envir., 15.VII.1995, G. Chekura leg. L=19 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) scheidleri bergeri Peeters & I.Rapuzzi, 2025
Endophallus
W-Austria,Tyrol, Vergötschen (Kaunertal), Feichten, h= 1300 m
Copyright © Ruslan Panin -
Morvanicarabus taxon page (1 photos)
Carabus (Morvanicarabus) scheibei scheibei Eidam, 1937
Female adult
NE-Afghanistan, Hindu Kush, Nuristan Prov., near Parun, 35°19'N 70°52'E h=2900 m, Pine forest, VI.2022, local collector/det. Kleinfeld, L= 24 mm
Copyright © Ruslan Panin -
Neoplectes taxon page (286 photos)
Carabus (Neoplectes) daphnis askhicus Belousov, 1992 Paratype
Male adult
SC-Caucasus, W Georgia, valley of riv. Chkhorodukhu, h=1000-1200m, 14.V.-6.VII.1991, Zamotajlov A.S & Miroshnikov leg.
male, 29 mm
photo & collection Panin R.
Copyright © Ruslan Panin
Carabus (Neoplectes) kratkyi Ganglbauer, 1890
Male+Female adult
W Caucas Mts., 2500m, Mt. Yatyrgvarta,
25.VI-12.VIII.1992 Shchurov V. leg.
male 20mm, female 21mm
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890
Female adult
NW-Caucasus, Russia,
Karacaj-Cherkessija, Abishira-Akhuba range h=2650m, upper Kyafar-Agur Riv., 4-9.VIII.2009, Solodovnikov I.A. leg.
female, 23 mm
Copyright © Ruslan Panin
Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988 (as shchessiensis) Paratype
Male+Female adult
NW Caucasus,
Krasnodar reg., Tuapsinskyi distr. Shessi Mt.
30.V.-25.VI.1993 Mt.Shessi, h=1800m., Imura & A.S.Zamotajlov, 1993 leg.
female, 29mm
Copyright © Ruslan Panin
Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
Male adult
NW-Caucasus, Russia,
Krasnodar reg., Apsheron Distr., Lagonaki Mt. ,near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominykh leg.
male, 28mm
Copyright © Ruslan Panin
Carabus (Neoplectes) satyrus duripshensis Kurnakov, 1972
Male adult
W.Caucasus, Abkhasia, Gudauta distr.,
Duripsh village env., Khipsta valley, h=500-700 m.
07-12.VI.2009 A.Safronov leg.
male - 30 mm.
Copyright © Alexandr Safronov
Carabus (Neoplectes) polychrous polychrous Rost, 1892
Female adult
W.Caucasus, Abkhasia, Gudauta distr., Duripsh village env., Khipsta valley, h=500-700 m. 8-11.06.2009 A. Safronov leg. lenght 40-43 mm.
Copyright ©
Carabus (Neoplectes) reitteri gagrinus Starck, 1894
Female adult
ABKHASIA, Gagra distr., Mamdsyshkha Mt., near Gagra, Fagus forest + karst, h = 1300-1330 m, biotope N 23, 43º18’186" N / 040º18’44" E, 12.05-05.07.2014, Leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
Female adult
Russia, Krasnodar Territory, West-Caucasus, Apsheronsky district, Guam ridge, env. Mezmay village, by the road, beech, oak, blackberry, h = 1200-1250 m, 03.IV-14.VI.2020, leg. S.M. Pavlyuchuk, A.P. Tatulyan
Copyright © Igor Solodovnikov
Carabus (Neoplectes) ibericus lafertei Chaudoir, 1846
Female adult
SC-Georgia, Samtskhe–Javakheti, Meskheti Mt Range, Borjomi, Likani env., h=897 m, 41°50.125'N 43°19.905'E, VI.2018, D. Prunier leg., L=37 mm
Copyright © Ruslan Panin -
Neoplesius taxon page (34 photos)
Carabus (Eocechenus) kaznakovi nangqenicus Deuve, 1995 Paratype
Male adult
Copyright © Boleslav Brezina -
Nesaeocarabus taxon page (9 photos)
Carabus (Nesaeocarabus) abbreviatus Brulle, 1835
Male adult
Canarias Isle, NC Tenerife; 22mm
Copyright © Aleš Sedláček
Carabus (Nesaeocarabus) faustus cabrerai Enderlein, 1929
Male adult
Spain, Canarias Isl.,Teneriffe: NW-part:Teno Massif, Erjos env., h=1000m, 18-21.III.2013, V.Slovak leg.
L=23 mm
Copyright © Ruslan Panin -
Ohomopterus taxon page (38 photos)
Carabus (Ohomopterus) albrechti albrechti Morawitz, 1862
Female adult
Japan, S-Hokkaido:Oshima Peninsula,Pass btw Otoshibe - Assabu, h~250-300 m, 10-20.VII.2010, 42:01’53’Т; 140:22’09’E, I.Rapuzzi & L.Caldon leg.
Copyright © Ruslan Panin
Carabus (Ohomopterus) japonicus japonicus Motschulsky, 1857
Male adult
S Japan, Kyushu, Kumamoto Pref., Izumi viil., h=600 m,VII.2001, S. Remond leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Ohomopterus) yamato kitai (Imura, 2004)
Male adult
Japan, S Honshu, Mie Pref., Hakusan, VII.2001, S. Remond leg.<br /> L=21 mm
Copyright © Ruslan Panin
Carabus (Ohomopterus) dehaanii dehaanii Chaudoir, 1848
Female adult
Japan, S Honshu, Hyogo Pref., Nishiwaki city, 20.VI.2010, Shinichi S. leg., L=37 mm
Copyright © Ruslan Panin
Carabus (Ohomopterus) dehaanii dehaanii Chaudoir, 1848
Male adult
Japan, S Honshu, Mie Pref., Inabe city, 21.II.2021, Shinichi S. leg., L=33 mm
Copyright © Ruslan Panin
Carabus (Ohomopterus) japonicus japonicus Motschulsky, 1857
Male adult
MT. Narabara-san, Ryuokakami Tamagawa town, Imabari city, Ehime pref. Japan 20.6.2015 Y. Amano (cAA). det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko
Carabus (Ohomopterus) japonicus japonicus Motschulsky, 1857
Female adult
Fukuoka pref., Omuta city, Hagio, 16.1.2012, S. Nishimoto leg. (cAA)
Copyright © Alexander Anichtchenko
Carabus (Ohomopterus) albrechti Morawitz, 1862
Male adult
Horoman, Samani town, Samani Co., Hokkaido, 16.6.2015 T. Nakamura leg. (cAA). det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko
Carabus (Ohomopterus) insulicola nishikawai (Ishikawa, 1966)
Male adult
Yamada, Isumi city, Chiba pref. Japan 20.5.2007 K. Osozawa leg. (cAA). 28mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko
Carabus (Ohomopterus) insulicola insulicola Chaudoir, 1869
Male adult
Japan, Kiyose, Tokyo 20.1.1961 (cAA). 29 mm
Copyright © Alexander Anichtchenko -
Ophiocarabus taxon page (34 photos)
Carabus (Ophiocarabus) regeli obscurior Semenov, 1888
Male+Female adult
Kyrgyzstan, NE Tian-Shan, E. Terskey Alatau Mts. R.,
Kok-kia River, h=2800 м.
7-8.VI.1997 М. Danilevsky leg.
Body lenght 15-18 mm.
Copyright © Alexandr Safronov
Carabus (Ophiocarabus) ballionis Kraatz, 1879
Male adult
NW-China, Xinjiang prov., 40 km N Narat, h=1500, 12-12.VI.2004, Benes & Kaiser leg.
L=22 mm
Copyright © Ruslan Panin
Carabus (Ophiocarabus) successor koeksuensis Kabak, 2013 Paratype
Male adult
China, Xinjiang, Koeksu Bas., Tshon-Aidak, SSE Tekes, 42* 32' 47'' N / 82* 05' 56'' E -- 42* 32' 51'' N / 82* 05' 36'' E, 3330-3550 m, 11.07.2012, Kabak I.I. leg.
Copyright © Ilya Kabak
Carabus (Ophiocarabus) successor successor Reitter, 1896
Male adult
SE-Kazakhstan, Narynkol distr., 4 km SW Bayankol, h~2400-2500 m, 2-14.VI.2007, Shumov leg. L=17.5 mm
Copyright © Ruslan Panin
Carabus (Ophiocarabus) successor successor Reitter, 1896
Female adult
SE-Kazakhstan, Narynkol distr., 4 km SW Bayankol, h~2400-2500 m, 2-14.VI.2007, Shumov leg. L=17 mm
Copyright © Ruslan Panin -
Oreocarabus taxon page (61 photos)
Carabus (Pachystus) hortensis neumeyeri Schaum, 1856
Female adult
Italy, Basilicata, Serra Calvello (PZ)
1400 m, 7.2003 F. Angelini leg.
female, 25 mm
Copyright © Alexander Anichtchenko
Carabus (Oreocarabus) carinthiacus Sturm, 1815
Male adult
NE-Italy, Bolzano, Brenner, Zirog h=2000 m, 26.VII.1992, Manfred Egger leg.
L=20 mm
Copyright © Ruslan Panin
Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847
Male adult
WC Spain, Cáceres, Hervás, h=1000m, V.2007, Pascal Stefani leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Oreocarabus) latro minshanensis Deuve, 1987
Male adult
China, N-Sichuan, Jiuzhaigou env., 13.VI.2001,E. Kučera leg. L=23 mm
Copyright © Ruslan Panin
Carabus (Oreocarabus) luetgensi pseudosteuarti Lapouge, 1924
Female adult
Portugal, Guarda, Sierra de Estrela h=800 m, 26.VI-4.VIII.2015, L=26 mm
Copyright © Ruslan Panin
Carabus (Oreocarabus) luetgensi pseudosteuarti Lapouge, 1924
Male adult
Portugal, Guarda, Sierra de Estrela h=800 m, 26.VI-4.VIII.2015, L=24 mm
Copyright © Ruslan Panin
Carabus (Pachystus) hortensis preslii Dejean, 1830
Male adult
S-Greece:, N Peloponnes, Erymanthos Reg., Kalentzi, 26.VI.2007, Manfred Egger leg. L=26 mm
Copyright © Ruslan Panin
Carabus (Pachystus) hortensis hortensis Linne, 1758
Female adult
Natural hybrid Carabus hortensis X Carabus granulatus. Belarus, Minsk reg., 8,7 km of Borisov, near vill. Lyubatovstchina, pine forest (Pinetum pleuroziosum), h = 213 m, 54.31934° N, 28.511486° E, 20-27.09.2018, leg. Klychenok Snezhana
Copyright © Igor Solodovnikov
Carabus (Hypsocarabus) mikhaili mikhaili Deuve & Mourzine, 1997
Female adult
L=23 mm
Copyright © Ruslan Panin
Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847
Female adult
Spain, Sierra de Guadarrama, Puerto de la Quesera, 1720m, 41.214640, -3.430896, Anichtchenko A. leg. 10.X.2025 (cAA).
Copyright © Alexander Anichtchenko -
Orinocarabus taxon page (68 photos)
Carabus (Orinocarabus) alpestris hoppei Germar, 1824
Male adult
W Austria, Tirol, Innsbruck, Hafelekar Mts h=2200m, Manfred Egger leg.
L=19 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) fairmairei fairmairei Thomson, 1875
Male adult
N Italy, Alpi Graie; 20 mm
Copyright © Aleš Sedláček
Carabus (Orinocarabus) fairmairei heteromorphus Daniel, 1896
Female adult
NW-Italy, Aosta region, Champorcher env., Mt.Dondenaz, VIII.2008, V.Monzini leg.
L=19 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) alpestris dolomitanus Mandl, 1956
Male adult
NE-Italy, South Tyrol, Dolomites-Alps, 500 m left of Würzjoch, Gömaalm - at the foot Mt.Peitlerkofel h=2100m, 28.VII-17.VIII.2013, Manfred Egger leg.
l=18 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) pedemontanus raynaudianus Deuve & Simard, 1977
Male adult
SE France, Alpes-Maritimes, Isola env., Colle Santa-Anna h=2309 m, 44*13',127" N, 07*06',242"E, 31-7.VIII..2012. L=21 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) concolor lepontinus Born, 1908
Male adult
NW-Italy, Piemonte (VB), Lepontine Alps, Mt. Zeda , 6.VII.1998, L Diotti leg., L=20 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) concolor concolor Fabricius, 1792
Female adult
NW-Italy, Breuil-Cervinia h~2500 m, 2.IX.1971, Georgeon leg. L=21 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) sylvestris transsylvanicus Dejean, 1826 (aberrant)
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg.
Copyright © Ruslan Panin
Carabus (Orinocarabus) sylvestris haberfelneri Ganglbauer, 1892
Female adult
Austria, Styria, Dietmannsdorf, Herrenwaldrücken, h=1550-1600m, 29.VII-15.VIII.2024, M.Fuchs leg., L=25 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) sylvestris sylvestris (as micklitzi Sokolar, 1910)
Endophallus
W Austria, Styria, Mt Hochwechsel W slope, 47.531667, 15.906000 h=1640 m,
Copyright © Ruslan Panin -
Oxycarabus taxon page (8 photos)
Carabus (Oxycarabus) saphyrinus pseudosaphyrinus Schweiger, 1964 (as rabaronianus)
Male adult
Anatolia, Kastamonu, Yaraligoz verso Catalzeitin,
VI.2001 Sola Claudio leg.
male, 28 mm
Copyright © Alexander Anichtchenko
Carabus (Oxycarabus) saphyrinus pirithous Reitter, 1896
Male adult
NW-Turkey,Istanbul (Asian part),Polonezköy env., 2.VI.2009. S. Dacatra leg. L=27 mm
Copyright © Ruslan Panin -
Pachycarabus taxon page (27 photos)
Carabus (Pachycarabus) roseri soganliensis Heinz & Korge, 1964
Male+Female adult
Turkey, Rize prov., Ikizdere distr., Rize Daglari,
47 km to SE from lyidere, Sivrikaya Koyu, h=2025 m.
30.VI-5.VII.2011, A. Vlasenko leg.
male - 23 mm, female - 24 mm.
Copyright © Alexandr Safronov
Carabus (Pachycarabus) staehlini Adams, 1817
Male adult
S Russia, E-Caucasus, Dagestan, Nukatl Mt Rng h=2600m, 25.VII.1997, V. Savizky leg. L=21 mm
Copyright © Ruslan Panin
Carabus (Pachycarabus) imitator katherinae Reitter, 1898
Endophallus
Abkhasia, Sukhumi distr., Anchkho Mt. Pass E- part h=1870-1920 m. det. I. Solodovnikov, 2025
Copyright © Ruslan Panin
Carabus (Pachycarabus) imitator imitator Reitter, 1883
Endophallus
Caucasus Major, NW Georgia, Utviri Pass env. h=2630 m, 43.083111, 42.255944
Copyright © Ruslan Panin
Carabus (Pachycarabus) roseri roseri Faldermann, 1835
Female adult
Georgia, Lesser Caucasus, Imereti prov., N slope Meskheti Mt Rng, Zekari Pass, h~1950 m, 24.VII.2021, E.Rujan leg., L=22 mm
Copyright © Ruslan Panin
Carabus (Pachycarabus) swaneticus swaneticus Reitter, 1883
Endophallus
Georgia, Caucasus, Svaneti Range E part, Latpari Pass h=2700-2800 m,
Copyright © Ruslan Panin
Carabus (Pachycarabus) swaneticus swaneticus Reitter, 1883
Female adult
Georgia, Caucasus, Svaneti Range E part, Latpari Pass h=2700-2800 m, 23.VI.2024, I.Uspenskyi leg., L=19 mm
Copyright © Ruslan Panin
Carabus (Pachycarabus) swaneticus swaneticus Reitter, 1883
Male adult
Georgia , Caucasus, E part Egrisi Mt Range, Mt. Tsekuri N slope h=2370 m, 18-20.VI..2025, I.Uspenskyi leg. L=19 mm
Copyright © Ruslan Panin
Carabus (Pachycarabus) staehlini rionicus Belousov, 1992
Male adult
Georgia, Caucasus, S of Sasashi, Lechkhumi Range N sope h=2430m, alpine meadows 242.759167, 42.973889, 20-21.VII.2025, I.Uspenskyi leg./det. R. Panin, L=19.5 mm
Copyright © Ruslan Panin -
Pachystus taxon page (45 photos)
Carabus (Pachystus) tamsi tamsi Menetries, 1832
Male+Female adult
Transcaucasia, Armenia, <br /> Ashotsk, 18.VI.2001, P.V.Kazaryan leg.<br /> female, 26mm//male, 24mm
Copyright © Ruslan Panin
Carabus (Pachystus) hungaricus hungaricus Fabricius, 1792
Female adult
C Hungary, Pest reg., Örkény, 9-10.V.1997, S.Budka leg.
Copyright © Ruslan Panin
Carabus (Pachystus) hungaricus cribellatus Adams, 1812
Male adult
E Siberia, Russia, Krasnoyarsk Reg., near Krasnoyarsk, forest-steppe,1.VII-20.VIII,2011,M.Ivanov leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Pachystus) hungaricus scythus Motschulsky, 1847
Female adult
S Ukraine, Zaporizhzhya reg., Khortytsa island, 16-20.VII.2011, M. Mulenko leg., det. Panin R.
L=28 mm
Copyright © Ruslan Panin
Carabus (Pachystus) cribratus staneki Sterba, 1931
Female adult
NC Turkey, Küre Mts., 60km N Kastamonu; 27mm
Copyright © Aleš Sedláček
Carabus (Pachystus) cavernosus cavernosus Frivaldsky, 1837
Female adult
W Bulgaria, Cerovo, 31 mm
Copyright © Aleš Sedláček
Carabus (Pachystus) cavernosus variolatus Costa, 1839
Male adult
C-Italy:Abruzzo reg., Gran Sasso d'Italia Massif, h~1600 m, VIII.1997, Seichert leg. L=26 mm
Copyright © Ruslan Panin
Carabus (Pachystus) cavernosus rozneri Retezár & Gy. Szél, 2013 Paratype
Male adult
Romania, Apuseni : Trascău Mts. L=27 mm
Copyright © Ruslan Panin
Carabus (Pachystus) cavernosus variolatus Costa, 1839
Female adult
C-Italy: Abruzzo reg., Gran Sasso massif, Campo Imperatore, h=1900 m, VIII.2021, G.Proscia leg., L=27 mm
Copyright © Ruslan Panin
Carabus (Pachystus) cribratus cribratus Quensel, 1806
Male adult
NW Armenia, Amasia, (Shirak prov.), 20-24.VIII.1998, Kazaryan P.V. leg.
Copyright © Ruslan Panin -
Pagocarabus taxon page (49 photos)
Carabus (Pagocarabus) crassesculptus diruptus Morawitz, 1886
Male adult
China, Gansu, Umg. Xiahe 3000-3200m
24-26.6.1993 Heinz leg.
20 mm
Copyright © Alexander Anichtchenko
Carabus (Pagocarabus) crassesculptus eleganticolor Deuve, 1997
Female adult
S Gansu, Min Shan, 5-20 km S Cagan, 2300-3200m 7-20.7.2019 V. Patrikeev leg. (cAA). det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko -
Pantophyrtus taxon page (20 photos)
Carabus (Pantophyrtus) turcomanorum turcomanorum Thieme, 1881
Male+Female adult
Kyrgyzstan, Kyrgyzskyi Mts. R., Uzun Bulak Gorge, h=1800 ms.
01-09.VI.2002 loc. coll.
male - 28 mm, female - 33 mm.
Copyright © Alexandr Safronov
Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993
Male adult
Uzbekistan, Chatkal Mt. rng., Irtash, 41°10'25.7"N 70°16'20.4"E, 5-7.8.2021, 2400m, A. Anichtchenko leg. (cAA). 27mm
Copyright © Alexander Anichtchenko
Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993
Female adult
Uzbekistan, Chatkal Mt. rng., Irtash, 41°10'25.7"N 70°16'20.4"E, 5-7.8.2021, 2400m, A. Anichtchenko leg. (cAA). 30 mm
Copyright © Alexander Anichtchenko -
Piocarabus taxon page (5 photos)
Carabus (Piocarabus) titanus Breuning, 1933
Male adult
C-China Sichuan prov., Nanping, h=1500m, VI.2000, S. Remond leg.<br /> L=34 mm
Copyright © Ruslan Panin
Carabus (Piocarabus) nanwutai Kleinfeld, Korell & Wrase, 1996
Male adult
China, W-Shaanxi, Taibai Shan Mts, btw. Houzhenzi/Ba Xian Tai, nr.Nan Tian Men h=3200m, 33°55'17"N/107°46'53"E, 25-30.VII.2011, Li Wuyang leg.<br /> L=19 mm
Copyright © Ruslan Panin
Carabus (Piocarabus) sui Imura & Zhou, 1998
Female adult
CE-China, Henan, Mt Wangwu, V.2007, local collector. L= 33 mm
Copyright © Ruslan Panin
Carabus (Piocarabus) vladimirskyi Dejean & Boisduval, 1830
Male adult
Buryatia, Kyahta distr. [Khyagh], 5 km SE Chikoi vill., 16.8.1998, Anichtchenko A. leg.
Copyright © Alexander Anichtchenko
Carabus (Piocarabus) titanus Breuning, 1933
Female adult
China Sichuan prov., Nanping, h=1500m, VI.2000, S. Remond leg.
L=38 mm
Copyright © Ruslan Panin -
Platycarabus taxon page (37 photos)
Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Female adult
W Ukraine, NE Carpathians, Transcarpathian Reg., Rakhov Distr., Chornogora Mts Range h=2030m, alpin zone, 13-24.VIII.2010, Panin R. leg., female, 22 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) irregularis montandoni Buysson, 1882
Male adult
W-Ukraine, NW Podolian,
Lviv Reg., Zhydachiv distr., near Vybranivka, under the bark of oak tree, in the forest,12.X.2007 Panin R. leg
L= 22 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Male adult
W.Ukraine, Transcarpathian Reg., Rakhov Distr., Chornogora Mt range h=2020m alpine zone, Panin R. leg. 28.VI-9.VIII.2008, male 19 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii fabricii Panzer, 1812
Male adult
NE-Italy,
Trentino-Alto Adige Reg,. Bolzano-Bozen Prov.,Val Aurina – Rottal 2050-2150m, 28.VI.1997, I.Rapuzzi leg.
male, 19mm
Copyright © Ruslan Panin
Carabus (Platycarabus) irregularis ramanus Sokolar, 1909 (as hayeki Kobmann, 1925)
Male adult
N Slovenia, Maribor – Dravograd, Pomorje Mts.,13.XI.2005, I.Rapuzzi leg.
Copyright © Ruslan Panin
Carabus (Platycarabus) depressus lucens Schaum, 1856
Female adult
SE-France: Alpi Marittime, Monte Viso, 14.VII.2010
L=23 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii fabricii (as koralpicus Sokolar, 1910)
Female adult
S Austria, Carinthia reg.(Kärnten), Wolfsberg, Koralpe Mt Range,18.VI.2005, Manfred Egger leg.
L=22 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii ucrainicus Lazorko, 1951
Male adult
Ukrainian Carpathians, Ivano-Frankivsk reg., Yaremcha distr., Gorgany Mts., h=1590 m, 22.VII-3.VIII.08, Panin R. leg., male, 23mm
Copyright © Ruslan Panin
Carabus (Platycarabus) creutzeri kircheri Germar, 1824
Male adult
S Austria, Karnische Alpen; male, 24mm
Copyright © Aleš Sedláček
Carabus (Platycarabus) irregularis ramanus Sokolar, 1909
Female adult
NW Croatia, Ivanščica Mts , Ivanec, 4-18.VI.2011, Krause Rene leg., L=33 mm
Copyright © Ruslan Panin -
Procerus taxon page (54 photos)
Carabus (Procerus) gigas gigas Creutzer, 1799
Male+Female adult
N Croatia, Ivanščica Mts , Ivanec, 4-18.VI.2011, Krause Rene leg.
female, 54 mm
Copyright © Ruslan Panin
Carabus (Procerus) scabrosus caucasicus Adams, 1817
Male adult
N Caucasus,
Stavropol Krai, Pyatigorsk, Mt.Mashuk, VI.2007 Tikhonov Yu. leg.
44mm
Copyright © Ruslan Panin
Carabus (Procerus) gigas duponcheli Dejean, 1831 (as kusdasi Mandl, 1966)
Male adult
S-Greece: Peloponnes, Mt.Chelmos, 20.V.2007.
L=46 mm
Copyright © Ruslan Panin -
Procrustes taxon page (48 photos)
Carabus (Procrustes) chevrolati chevrolati Cristoforis & Jan, 1837
Male+Female adult
female, 31mm: Ashotsk,Vardaghbyur vill., 30.V.1997
male, 30 mm: Amasia, Chaldash vill.,8-11.VIII.1997
P.V. Kazaryan leg.
Copyright © Ruslan Panin
Carabus (Procrustes) coriaceus hopffgarteni (as emgei Ganglbauer, 1888)
Male adult
CE-Greece: Larissa reg. Stomio, Ossa Mt. 6-17.VII.2011, Krause Rene leg.
L=32 mm
Copyright © Ruslan Panin
Carabus (Procrustes) chevrolati thirki Chaudoir, 1857
Male+Female adult
Turkey, Rize prov., Ikizdere distr., Rize Daglari,
47 km to SE from lyidere, Sivrikaya Koyu, h=2025 m.
30.VI-5.VII.2011, A. Vlasenko leg.
male - 28 mm, female - 31 mm.
Copyright © Alexandr Safronov
Carabus (Procrustes) banoni Dejean, 1829
Male adult
Greece,W Crete Island, Mili gorge ,12.V.2009, Manfred Egger leg.
L=26 mm
Copyright © Ruslan Panin
Carabus (Procrustes) talyschensis talyschensis Menetries, 1832
Female adult
Azerbaijan, Talysh Mts., Zuvand, Kalvaz vill., h=1650 m, 15.VII.2003, Dolin V.G. leg.
L=32 mm
Copyright © Ruslan Panin
Carabus (Procrustes) piochardi piochardi Gehin, 1883
Male adult
C-Lebanon, Faraya vill., Mt. Djebel Sannin h=1800m, X.2012, C. Reuter leg. <br /> L=31 mm
Copyright © Ruslan Panin
Carabus (Procrustes) paiafa acuticollis Motschulsky, 1858
Male adult
SC-Turkey: Alania (Irmasan), rte 695 au km 25 Akseki, h= 1225 m, 02.X.2003
Copyright © Igor Solodovnikov
Carabus (Procrustes) piochardi praestigiator Morawitz, 1886
Male adult
C-Lebanon, Byblos /Jbeil Distr., Kfardebian env., h=1200m, XI.2014, C. Reuter leg. L=31 mm
Copyright © Ruslan Panin
Carabus (Procrustes) coriaceus ressli Mandl, 1961
Female adult
SC-Turkey: Hatay Prov, Dörtyol, Nur Mts, Topaktaş h=900-1200 m, 12.V.2005, D. Prunier leg. L=35 mm
Copyright © Ruslan Panin
Carabus (Procrustes) coriaceus excavatus Charpentier, 1825
Female adult
W Croatia, Velebit Mts, Jablanac Stirovaca h=998 m, 44.71124, 14.95673,3.VI-7.VII.2017, G. Proscia leg., L= 38 mm
Copyright © Ruslan Panin -
Pseudocoptolabrus taxon page (42 photos)
Carabus (Pseudocoptolabrus) burmanensis Breuning, 1932
Male adult
male, 23mm
Copyright © Boleslav Brezina
Carabus (Pseudocoptolabrus) businskyi Deuve, 1990
Male adult
CH, NW-Yunnan (Lijiang) Yulongxue Shan, E-slope
27:10:50N/100:12:26E/ 4400m, alpine meadows 12.VII.2000
K. & B. Brezina, M.Brychcin, M. Hackel
Copyright © Boleslav Brezina -
Pseudocranion taxon page (14 photos)
Carabus (Pseudocranion) sackeni sackeni Semenov, 1898
Male adult
male, 23mm
Copyright © Boleslav Brezina
Carabus (Pseudocranion) sackeni remondianus Deuve, 1996 Paratype
Female adult
Museum of Institute of Systematic Biology (Daugavpils, Latvia)
Copyright © Alexander Anichtchenko
Carabus (Pseudocranion) lazikouensis lazikouensis (as kundaensis Deuve, 2003) Paratype
Female adult
C-China, S-Gansu: SSW of Minxian: Qilian Shan: W Kunda, 34:10N/103:47E, h=2800-3000m, forest, 7.VII.2002, Belousov & Kabak leg.
L=24 mm
Copyright © Ruslan Panin
Carabus (Pseudocranion) taibaishanicus taibaishanicus Deuve, 1989
Female adult
China, W-Shaanxi, Taibai Shan Mts, btw. Houzhenzi/Ba Xian Tai, nr. Nan Tian Men h=3200m, 33°55'17"N/ 107°46'53"E, 25-30.VII.2011, Li Wuyang leg. L=29 mm
Copyright © Ruslan Panin -
Relictocarabus taxon page (1 photos)
Carabus (Relictocarabus) meurguesianus Ledoux, 1990
Male adult
Image from molbiol.ru
Copyright © - - - -
Rhigocarabus taxon page (163 photos)
Carabus (Rhigocarabus) sehnali Brezina & Hackel, 2006 Holotype Paratype
Male adult
male, 20mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) sehnali Brezina & Hackel, 2006
Female adult
female, 23mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) gentleman Brezina & Hackel, 2004 Holotype
Male adult
male, 19mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) buddaicus gansuicus Deuve, 1991
Male adult
China, NC-Sichuan (ZOIGE), LANGMUSI (=DOGCANGLHAMO), 34:03:48N/102:38:08E, h=4100m, alpine meadows, screes, 16-18.VII.1999, B.Brezina leg.
L=20 mm
Copyright © Ruslan Panin
Carabus (Rhigocarabus) buddaicus linxiaicus Deuve, 1995
Male adult
China, S-Gansu (LABRANG):, valley.E of Ponggartang, 35:12-13N/102:54-55E, h=3000-3700m, alpine meadows, coniferous forests,10.-14.VII.1999, B.Brezina leg. L=20 mm
Copyright © Ruslan Panin
Carabus (Rhigocarabus) cateniger cateniger (as sommitalis Cavazzuti, 2006)
Female adult
China, C-Gansu, Mts Qilian h=3378 m, road G227, Yangxiongzi env. 38.050917, 100.887267, 18-23.VI.2011, G.Proscia leg., L=19 mm
Copyright © Ruslan Panin
Carabus (Rhigocarabus) cateniger cateniger (as sommitalis Cavazzuti, 2006)
Male adult
China, C-Gansu, Mts Qilian h=3378 m, road G227, Yangxiongzi env. 38.050917, 100.887267, 18-23.VI.2011, G.Proscia leg., L=17 mm
Copyright © Ruslan Panin
Carabus (Rhigocarabus) roborowskii roborowskii Semenov, 1887
Male adult
China, E-Qinghai, Mts NW of Tongren city, 35.562778, 101.843667, h=3870 m, 15-26.VI.2011, G. Proscia leg./det.R.Panin, L=20 mm
Copyright © Ruslan Panin
Carabus (Rhigocarabus) roborowskii roborowskii Semenov, 1887
Female adult
China, E-Qinghai, Mts NW of Tongren city, 35.562778, 101.843667, h=3870 m, 15-26.VI.2011, G. Pros cia leg./det.R.Panin, L=20 mm
Copyright © Ruslan Panin
Carabus (Rhigocarabus) rhododendron Deuve & Imura, 1991
Male adult
China, N Yunnan: Baimaxue Shan, S of Deqen, h=4300m, 28°20'N 99°03'E, 16-20.VI.2016, C.Reuter leg., L=18 mm
Copyright © Ruslan Panin -
Scambocarabus taxon page (6 photos)
Carabus (Scambocarabus) kruberi laobeiensis Deuve & Li, 2000
Male adult
NE-China: Liaoning prov., Fengcheng City, Miaoyang, 12-17.VI.2011
L=27 mm
Copyright © Ruslan Panin -
Semnocarabus taxon page (29 photos)
Carabus (Semnocarabus) carbonicolor carbonicolor Morawitz, 1886
Male+Female adult
Kyrgyzstan, NC Tianshan, Kyrgyz Mt. R., Tuyuk riv., h=2000 м.
25-VI-1997 М. Danilevsky leg.
male - 20 mm, female - 21 mm.
Copyright © Alexandr Safronov
Carabus (Semnocarabus) erosus kuldshaensis Ballion, 1878
Male adult
S-Kazakhstan:Ili River valley,:near Bakanas,3-5.V.2002
L=21 mm
Copyright © Ruslan Panin
Carabus (Semnocarabus) carbonicolor callosus Semenov, 1896
Male+Female adult
KAZAKHSTAN,
Bolshoe Almaatinskoe Lake,30.VI.1992/ det.Panin R.
female 19mm
Copyright © Ruslan Panin
Carabus (Semnocarabus) carbonicolor korolkowi Semenov, 1896
Male adult
NC-Kyrgyzstan, Kyrgyz Mt.Rng , 10 km SW of Bishkek, Chon-Aryk env, h=1100m, 24.V.1998, /det. Panin R.
22 mm
Copyright © Ruslan Panin
Carabus (Semnocarabus) erosus corax Semenov, 1896
Endophallus
SE-Kazakhstan, Almaty reg., C Trans-Ili Alatau N-slopes, Kaskalen river Valley, Izvestkovaya vill., h=1700 m, 6.V.2015, D.Kovalchuk leg./det. R.Panin,
Copyright © Ruslan Panin
Carabus (Semnocarabus) erosus corax Semenov, 1896
Male adult
SE-Kazakhstan, Almaty reg., C Trans-Ili Alatau N-slopes, Kaskalen river Valley, Izvestkovaya vill., h=1700 m, 6.V.2015, D.Kovalchuk leg./det. R.Panin,
Copyright © Ruslan Panin
Carabus (Semnocarabus) erosus corax Semenov, 1896
Female adult
SE-Kazakhstan, Almaty reg., C Trans-Ili Alatau N-slopes, Kaskalen river Valley, Izvestkovaya vill., h=1700 m, 6.V.2015, D.Kovalchuk leg./det. R.Panin, L=24 mm
Copyright © Ruslan Panin
Carabus (Semnocarabus) carbonicolor carbonicolor Morawitz, 1886
Female adult
SE-Kazakhstan, Almaty reg., Khan Tengri Massif, Narynkol env.h=1800 m, 30-31.-V.2007, Shumov leg., L=21 mm
Copyright © Ruslan Panin
Carabus (Semnocarabus) erosus erosus Motschulsky, 1865
Endophallus
SE-Kazakhstan, 70 km E of Almaty, E Trans-Ili Alatau N-slopes, valley Assy (Asy) riv. Assy plateau h=1700 m, 12.V.2015
Copyright © Ruslan Panin
Carabus (Semnocarabus) rustemi Kabak, 2009
Male adult
SE-Kazakhstan, Almaty reg., near Bodokhudzir 43.916667, 79.716667 h=500 m, 12.VIII.2019, det. R.Panin, L=22 mm
Copyright © Ruslan Panin -
Shunichiocarabus taxon page (1 photos)
Carabus (Shunichiocarabus) uenoianus uenoianus Imura, 1995
Male adult
Copyright © - - - -
Sigenacarabus taxon page (6 photos)
Carabus (Pagocarabus) caelestinus caelestinus (Imura, 2007) Paratype
Male adult
Copyright © Boleslav Brezina
Carabus (Pagocarabus) sichuanicola sichuanicola (as drolmae Lassalle, 2003)
Female adult
China, Sichuan, NW Chengdu, Mt. Jiajin Shan h~4000 m,VI. 2016, local collector, L= 24 mm
Copyright © Ruslan Panin
Carabus (Sigenacarabus) trachynodes moxiensis Deuve, 1992
Male adult
China, W-Sichuan, W of Moxi, Daxue Range, Mt.Gongga Shan h=3000-3100m, 29°34′29.6″N /101°59′44.0″E, 10-12.VI.2018, Ch. Reuter leg. L=21 mm
Copyright © Ruslan Panin
Carabus (Sigenacarabus) trachynodes moxiensis Deuve, 1992
Male adult
China, W-Sichuan, Daxue Range, Mt.Gongga Shan h=3000-3100m, Hailuogou Glacier Forest Park, 29.574889, 101.995556, 10-12.VI.2018, Ch. Reuter leg., L=21 mm
Copyright © Ruslan Panin -
Sphodristocarabus taxon page (73 photos)
Carabus (Sphodristocarabus) varians varians Fischer, 1827
Male+Female adult
female: Transcaucasia, Armenia, Amasia distr., Karakhachskiy mountain pass h=2270m, 3.VIII.1985 P.V.Kazaryan leg., 22mm<br /> male: Armenia, Amasia distr., Arpalach vill. VI.1999, P.V. Kazaryan leg.
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) varians janthinus s.str.
Male+Female adult
NW-Caucasus, Russia, Krasnodar reg.,Tuapse distr., Mt.Semashkho 04.V.2008, D.Fominyh leg.<br /> female, 24mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) macrogonus pseudokolatensis Heinz & Korge, 1967
Male adult
NE-Turkey::Kackar Dagi Mt, Ayder S of Ardesen h=1600m, 17-21.VI.2010, M.Janalik leg.
L=20 mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) separatus persa Lapouge, 1924
Male adult
N-Iran: Mazandaran Province, Elborz Mt.Rng, Vandarbon env., 10.VII.2002 L=23 mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) blumenthaliensis aequicostalis Staven & Heinz, 1995
Male adult
NE-Turkey:NE of Artvin:Dalis Dağ ,h=1600-1800m, 8-18.VII.2014, C. Reuter leg. L= 25 mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) separatus bararii Deuve, 2000
Male adult
N-Iran: Mazendaran: Ramsar env., deciduous forest, h~305 m, 16-24.VI.2017, Václav Čutka leg. L=22 mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) separatus separatus Gehin, 1885
Female adult
N-Iran: Gilan Prov., Asalem - Khalkal road, 25-30.V.2018, Václav Čutka leg. L= 23 mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) separatus separatus Gehin, 1885
Male adult
N-Iran:Gilan Prov., Asalem - Khalkal road, 25-30.V.2018, Václav Čutka leg. L= 22 mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) varians janthinus Ganglbauer, 1887
Female adult
Carabus (Sphrodistocarabus) janthinus dvorschaki Mandl, 1975 Abkhazia, Gudautsky district, S- Bzybsky Ridge, a watershed hill at the head of the river Khipsta, 43°17'56.63" N, 40°40'16.74" E, h= 1490 m, mature beech-maple forest with butterbur, moist soil, 05.07-17.08.2017, leg. I.A. Solodovnikov, S.V. Solodovnikova, E.V. Tatun
Copyright © Igor Solodovnikov
Carabus (Sphodristocarabus) varians armeniacus Mannerheim, 1830
Male adult
C Georgia, Imereti, Caucasus, Likhi Mt Range, Rikoti Pass h=900-1000 m, 14-16.V.2017, E. Rutjan E. leg., L=22 mm
Copyright © Ruslan Panin -
Stephanocarabus taxon page
-
Tachypus taxon page (40 photos)
Carabus (Tachypus) auratus brunieri Barthe, 1921
Male adult
SC-FRANCE, Aude, Rennes les Bains, 6.VII.1989, Sunard A. leg., 26mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
Male adult
Italy-Switzerland, Riviera – Preonzo, 18.II.1998, C.Sola leg.
L=26 mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus graniger Palliardi, 1825
Male adult
SW-Romania, Banatului Montan, Resita - Valiug, 29.V-9.VI.2005. Krause R. leg.
L=25 mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus durus ( as carpathicus Kolbe, 1913)
Male+Female adult
Ukrainian Carpathians, Ivano-Frankivsk reg., Gorgany Mts., 20km SW Nadvorna, 22.VI-3.VIII.20, Panin R. leg.<br /> female, 26mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus durus Reitter, 1896
Male+Female adult
W Ukraine, Transcarpathian reg., Vinogradov distr., near Malaya Kopanya vill.,7-18.VII.2008, Panin R. leg. <br /> female, 32mm.
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus intermedius Dejean, 1826
Female adult
NW -Serbia,Fruška Gora,Vrdnik env. h=380m, 9-10.VII.2013, M. Janalik leg. L=29 mm
Copyright © Ruslan Panin
Carabus (Tachypus) auratus honnoratii Dejean, 1826
Male+Female adult
France : Vaucluse, Mt. Ventoux-Mt. Serein, 1400-1800m, vii.1969-N. Mal Leg.-Coll.RBINS, N. Mal - including f.i. ventouxensis and nicolasi
Copyright © Noël Mal
Carabus (Tachypus) cancellatus durus (as strictus Kolbe, 1913)
Male adult
W Ukraine, NE Carpathians, Maramaros Mts (Hutsul Alps) Gorge of Kuziy (Kizi) stream, h = 400m, 20-23.V.2021, Panin R. leg., L=30 mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus durus (as strictus Kolbe, 1913)
Female adult
W Ukraine, NE Carpathians, Maramaros Mts (Hutsul Alps), Gorge of Kuziy (Kizi) stream, h = 400m, 20-23.V.2021, Panin R. leg., L=32 mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus durus Reitter, 1896
Female adult
W Ukraine, Transcarpathian, Velykyi Bereznyi distr., Stuzhytsya (Stužica), 2-29.VI.2011, Yu. Kanarsky leg. L=33.5 mm
Copyright © Ruslan Panin -
Tanaocarabus taxon page (14 photos)
Carabus (Tanaocarabus) hendrichsi Bolivar y Pieltain, Rotger & Coronado, 1967
Male adult
male, 23mm
Copyright © Boleslav Brezina
Carabus (Oreocarabus) taedatus agassi Le Conte, 1850
Male adult
SC-USA, New Mexico, Santa Fe,VIII.1999, Sehnal R. leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Oreocarabus) taedatus bicanaliceps Casey, 1920
Male adult
NW-USA, Washington, Thurston County, Mima Mound Prairie, 15.IX.2005, A. Laforgue leg. L=23 мм
Copyright © Ruslan Panin -
Teratocarabus taxon page (2 photos)
Carabus (Teratocarabus) azrael mizunumaianus Imura, 1991
Male adult
China, Liaoning, Benxi, Gongchangung
20.VI.2004 Hong
Copyright © Boleslav Brezina -
Tomocarabus taxon page (28 photos)
Carabus (Tomocarabus) scabripennis scabripennis Chaudoir, 1850
Male adult
Carabus (Tomocarabus) scabripennis scabripennis Chaudoir, 1850
C. Armenia,
Kotayk reg., Yeghvard, Arailer Mt., 16.VI.1999, P.V.Kazaryan leg.
male, 20mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Tomocarabus) marginalis Fabricius, 1794
Male+Female adult
E Ukraine,
Kharkov reg.,near Vasishchevo vill.10-20.VI.2006 D.Kovalchuk leg.
female 24mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) decolor Fischer, 1823
Male adult
S Russia, NW-Caucasus,
Karacaj-Cherkessija,Skalisty Mnt. Range, Site Banditskoe h=1400m, 2-3.V.2009,
Fominykh D., Bondarenko A.S. leg.
male,17mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) bessarabicus bessarabicus Fischer, 1823
Male+Female adult
UKRAINE,
Kherson Reg., Golaya Pristan distr. near Rybalche vill. 6.IV.2000
long (female) 20mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) convexus gracilior Gehin, 1885
Male adult
SE-Bulgaria, Burgas Prov., Primorsko env., 18-30.VI.2000, Krause Rene leg.
L=18 mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) bessarabicus ? concretus Fischer, 1823
Male adult
E Ukraine, Lugannsk Reg., Sverdlovsk distr., Steppe Reserve “Provalsky", 3.V-3.VI.2011, leg.Glotov S./det.Panin R.<br /> female, 22 mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) convexus moreanus Reitter, 1896
Female adult
Greece:, N Peloponnes, Xylokastro, Zemeno,h=1000m, 6.VI.1997, Manfred Egger leg. L=21 mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) rumelicus syriensis Breuning, 1943
Male adult
N Lebanon, Fnaпdek, Djebel Qammoua, h=1100-1200m, C. Reuter leg.
L=17.5 mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) convexus dilatatus Dejean, 1826
Male adult
NE Serbia, Majdanpek; 13mm
Copyright © Aleš Sedláček
Carabus (Tomocarabus) convexus erzeliki Schweiger, 1962
Female adult
NC-Turkey, Sinop province, Erfelek env., h=200m, 10-27.VII.2014, C. Reuter leg. L=20 mm
Copyright © Ruslan Panin -
Trachycarabus taxon page (3 photos)
Carabus (Trachycarabus) estreicheri Fischer, 1822
Male+Female adult
W UKRAINE, NW Podolian,
Lviv Region, Brody distr., near Yasenov vill., steppe sites, 2-28.V.2010, Panin R. leg.
female, 23mm
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus tscherkessicus Korge, 1964
Male adult
S Russia, NW-Caucasus,<br /> Karachay-Cherkessia,Teberda, Mukhinsky (Mukhu) Mt. pass, h=2750-2800 m , 8.VIII.2010, I.A. Solodovnikov leg.<br /> male, 23 mm<br />
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus haeres (as fossulatus Dejean, 1826)
Male+Female adult
E.UKRAINE,<br /> Kharkov reg., Zmiev distr.,in steppe,10-20.V I.2007, D.Kovalchuk leg.<br /> female, 26mm/male, 24mm.<br /> photo/det. Panin R.
Copyright © Ruslan Panin
Carabus (Trachycarabus) scabriusculus scabriusculus (as erythropus Fischer, 1822)
Male+Female adult
W.Ukraine,<br /> NW Podolian Upland , Lviv Reg., Zolochiv distr., near Luky vill., steppe sites, 7.V-8.VI. 2010<br /> leg. Panin R.<br /> female, 22mm
Copyright © Ruslan Panin
Carabus (Trachycarabus) planus planus Gehin, 1885
Male adult
S Russia,<br /> Krasnodar reg., Anapa distr., B. Utrish vill., env., 11-20.VIII.2008, Khatchikov E. leg.<br /> male, 26mm<br /> photo Panin R.
Copyright © Ruslan Panin
Carabus (Trachycarabus) planus planus Gehin, 1885
Endophallus
<b>endophalus</b> Karacaj-Cherkessija, near Kart-Djurt vill., upper Kylan riv. h=1300m
Copyright © Ruslan Panin
Carabus (Trachycarabus) besseri Fischer, 1822
Male+Female adult
W Ukraine.NW Podolian,
Lviv reg., Zolochiv distr., Lysa Gora near Stinka vill., steppe sites, 25.IV-9.V.2009 Panin R. leg
female, 30mm
Copyright © Ruslan Panin
Carabus (Trachycarabus) estreicheri Fischer, 1822
Female adult
Volgograd Reg., Serafimovka Distr., Ust' -Khoperskaya Station 5.V.2009 leg.O.Sveshnikov
18.5 mm
Copyright © Kirill Makarov
Carabus (Trachycarabus) estreicheri Fischer, 1822 (ab.rufofemoratus Lomnicki, 1892)
Female adult
W Ukraina, Podolian Upland, Lviv reg., Brody distr., Mt. Makitra, steppe site, 19.V-5.VI.2020, Panin R. leg. L=23 mm
Copyright © Ruslan Panin
Carabus (Trachycarabus) planus campicola Dejean, 1831
Female adult
S Ukraine, Crimea, S slope Mt Demerdzhi h=650 m, sibljak, VII.2004, R. Panin leg., L=29 mm
Copyright © Ruslan Panin -
Tribax taxon page (145 photos)
Carabus (Tribax) titan titan Zolotarev, 1913
Male+Female adult
Russia,NW Caucasus,
Krasnodar reg.,Chernogorie Mnt. Range, Agulova Balka h=950m V-VII.1999, A.S.Zamotajlov leg.
female,32mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Tribax) circassicus abagonensis Starck, 1894
Male+Female adult
S Russia, NW Caucasus,
Adygea Rep., Mt.Abago, 26.VI.1999
L=24mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Tribax) steveni schamyli Hampe, 1852
Female adult
C Caucasus, North Ossetia, the road on the village of Coban (Koban), the left river bank Gizeldon, the deciduous wood, h = 895-905 m, N 42°55'32.91", E 44°31'53.53", 20.05-03.07.2017, leg. A.P. Tatulyan
Copyright © Igor Solodovnikov
Carabus (Tribax) nothus sharoiensis Fominykh, Zamotajlov, Avtaeva et Titarenko, 2019 Paratype
Male adult
Copyright © Igor Solodovnikov
Carabus (Tribax) nothus fausti Dohrn, 1873
Female adult
Carabus (Microtribax) fausti fausti Dohrn, 1873 Russia: Daghestan, Kyabyak Mt.R.
Copyright © Igor Solodovnikov
Carabus (Tribax) puschkini kolenatii Chaudoir, 1846
Male adult
C-Georgia, Trialeti Mt Range, 2 km SE of Borjomi h=916 m, 41°48'35.3"N 43°26'11.8"E, 17.VI.2013, D. Prunier leg./det. Panin R., L=28 mm
Copyright © Ruslan Panin
Carabus (Tribax) kasbekianus veselyi Gottwald, 1980
Female adult
Georgia, Caucasus, Svaneti Range E part, Latpari Pass h=2700-2800 m, 23.VI.2024, I.Uspenskyi leg., L= 24 mm
Copyright © Ruslan Panin
Carabus (Tribax) kasbekianus veselyi Gottwald, 1980
Male adult
Georgia, Caucasus, Svaneti Range E part, Latpari Pass h=2700-2800 m, 23.VI.2024, I.Uspenskyi leg., L= 22 mm
Copyright © Ruslan Panin
Carabus (Tribax) kasbekianus veselyi Gottwald, 1980
Male adult
Georgia, Caucasus, S of Sasashi vill., Lechkhumi Range N slope h=2430 m, alpine meadows, 242.759167, 42.973889, 20-21.VII.2025, I.Uspenskyi leg./det. R.Panin, L=24 mm
Copyright © Ruslan Panin -
Ulocarabus taxon page (14 photos)
Carabus (Ulocarabus) stschurovskii Solsky, 1874
Male adult
Tadzhikistan, Pamir-Alay, S slope Gissar Mt.Range, Takob, h-1700m, 10.V.1996, Dolin V.G. leg.
det. Panin R.Yu.
L=22 mm
Copyright © Ruslan Panin
Carabus (Ulocarabus) stschurovskii Solsky, 1874
Female adult
Tadzhikistan, Pamir, N-slope Peter I.Mt Rng,h=2100m,Ganishou vill., Dolin V.G. leg.
det.Panin R.Yu.
L=24 mm
Copyright © Ruslan Panin
Carabus (Ulocarabus) theanus delerei Mandl, 1955
Male adult
N-Afghanistan, Hindukush Mt.Rng, NE Salang Pass h=2300 m, 35°27'494"N, 68°58'399"E, IV.2011, C. Reuter leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Ulocarabus) theanus theanus Reitter, 1895
Male adult
Tadzhikistan, S slope Gissar Mt.Rng, left bank of the river Sorbo, Haidar gorge,1.VI.1984, Dolin V.G. leg./det.Panin R.Yu.
L=21 mm
Copyright © Ruslan Panin
Carabus (Ulocarabus)? theanus theanus Reitter, 1895
Male adult
NC Tadzhikistan, Pamir, N-slope Peter I.Mt Rng, h=2100m, Ganishou vill., 10-12.VI.2006, Ivanova E. leg.
L=23 mm
Copyright © Ruslan Panin
