Carabus (Acathaicus) alexandrae alexandrae Semenov, 1887 Holotype Holotype
Male adult
Kwantin and Li-tschara-pu, 22.VI.1885, leg. Potanin<br /> Deposited in Zoolog. Museum St. Petersburg.<br /> male, long. 22 mm<br /> photo B. Brezina
Copyright © Boleslav Brezina

Carabus (Acathaicus) alexandrae cratocephaloides Semenov, 1887
Female adult
female, 28mm
Copyright © Boleslav Brezina

Carabus (Acathaicus) interruptocostatus Semenov, 1887 Holotype Holotype
Female adult

Copyright © Boleslav Brezina

Carabus (Acoptolabrus) lopatini Morawitz, 1886
Male+Female adult
Russia,
Sakhalin isl., Tchekhov Mt, VI.1997
female, 40mm
photo & coll. Panin R.
Copyright © Ruslan Panin

Carabus (Acoptolabrus) schrencki schrencki (as hauryi Gehin, 1885 )
Male adult

Copyright © Boleslav Brezina

Carabus (Acoptolabrus) schrencki kimieae Kezuka & Imura, 1989
Male+Female adult
Mt. Wulongshan, Dandong City,
Liaoning Province, CHINE
25-30.VI.2008
Copyright © Safronov

Carabus (Coptolabrodes) haeckeli Brezina & Imura, 1997
Male adult
C China, Shaanxi,
Tsinling Mts., Taibaishan Mt., h=3000m, 33:55N/107:44 E, VI.2004, V.Siniaev & his team
female, 31 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Acoptolabrus) constricticollis constricticollis (as grallatorius Roeschke, 1921)
Male adult
NE-Korea:North Hamgyong Prov., Sechon-dong, VII.2004, Li leg.
L=28 mm
Copyright © Ruslan Panin

Carabus (Acoptolabrus) mirabilissimus rigoutianus Deuve & Li, 1998
Male adult
N-Korea, South Pyongan prov., Mt. Maeng-san, 6.VI.2010
L=25 mm
Copyright © Ruslan Panin

Carabus (Acoptolabrus) gehinii gehinii Fairmaire, 1886
Female adult
N-Japan, SW-Hokkaido, Chitose env.,VI.2011.
L=34 mm
Copyright © Ruslan Panin

Carabus (Acoptolabrus) schrencki minpongsanensis Deuve & Li, 2003
Male adult
North Korea, Chagang Prov., Manpo, Songhak-ri (villl.,) 1-16.VII.2011, L=26 mm
Copyright © Ruslan Panin

Carabus (Acoptolabrus) gehinii munakatai (Ishikawa, 1968)
Male adult
Japan, SW-Hokkaido, Yakumo town, 4.VI.2017, S.Shinichi leg., L=30 mm
Copyright © Ruslan Panin

Carabus (Acoptolabrus) constricticollis Kraatz, 1886
Male adult
China, Liaoning prov., Mt. Guanmen 20.6.2009 (cAA)
Copyright © Alexander Anichtchenko

Carabus (Carabus) gressittianus fuscomarginatus Kleinfeld, 1997 Holotype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Carabus) andreiianus Deuve, 2007
Female adult
C-China, N-Sichuan, Pingwu Co. h=1500 m, 27.VI.2011, L=24 mm
Copyright © Ruslan Panin

Carabus (Carabus) koidei koidei Imura, 1996
Male adult
C-China, S-Sichuan, Pass btw. Yanyaun / Muli, h=3244 m, 27.68638N; 101.22335E,,11-18.VI.2017, Ch. Reuter leg. L=22 mm
Copyright © Ruslan Panin

Carabus (s.str.) morphocaraboides zhaojueanus Imura, 1999
Female adult
China, S-Sichuan, SE Mianning, Pass btw Mianshanzhen & Yuexi 28°30’N/102°26’E, 22-23.VI.2017, Ch.Reuter leg., L=22 mm; det. B. Brezina, 2020
Copyright © Ruslan Panin

Carabus (s.str.) morphocaraboides zhaojueanus Imura, 1999
Male adult
China, S-Sichuan, road 56 km E of Xichang to Zhaojue, Mt.WenLin, Pass h=3200 m, 27°52’N/102°30E, 5-20.VI.2017, Ch.Reuter leg.; det B. Brezina, 2020
Copyright © Ruslan Panin

Carabus (Acoptopterus, ?Carabus) yunnanus yanyuanicus Cavazzuti, 1996
Endophallus
China, S-Sichuan, S Yanyuan, Chuandongzi Mt h=3300-3600 m
Copyright © Ruslan Panin

Carabus (Acoptopterus) yunnanus yanyuanicus Cavazzuti, 1996 (? or oblongior ssp.)
Male adult
China, S-Sichuan, S Yanyuan, Chuandongzi Mt h=3300-3600 m, 16-20.VI.2023, C.Reuter leg./det.Klienfeld, 2023, L=21.5 mm
Copyright © Ruslan Panin

Carabus (Acrocarabus) callisthenoides Semenov, 1888
Male adult
male, 22mm
Copyright © Boleslav Brezina

Carabus (Acrocarabus) guerini aldarkose Kabak, 1992
Male adult
male, 22mm
Copyright © Boleslav Brezina

Carabus (Acrocarabus) guerini guerini Fischer, 1842
Male adult
SE Kazakhstan, W Dzhungar Mts, Lepsinsk env., 1600 m., 18.06.2000, A.Klimenko
Copyright © Anton Menstschikov

Carabus (Alipaster) barovskii bayanbulak Deuve, 1993 Paratype Paratype
Female adult
female, 17mm
Copyright © Boleslav Brezina

Carabus (Alipaster) infantulus infantulus Morawitz, 1886
Male adult
male, 15mm
Copyright © Boleslav Brezina

Carabus (Cechenotribax) petri Semenov & Znojko, 1932
Female adult
female, 18mm
Copyright © Boleslav Brezina

Carabus (Eotribax) eous eous Morawitz, 1889
Male adult
Kirghyzstan, N Terskei Ala Too, Borskaun riv., h=2500 m., 7-14.07.2000
male
L=17 mm
Copyright © Maksim Smirnov

Carabus (Cechenotribax) petri Semenov & Znojko, 1932
Male adult
SE Kazakhstan, N-Tianshan:N-slopes Ketmen Mt.Rng. h=2250-2500m., near Ketmen vill.,1-5.VI.1990, Dolin V.G. leg.
L=13.5 mm
Copyright © Ruslan Panin

Carabus (Alipaster) pupulus pupulus Morawitz, 1889
Female adult
S Kyrgyzstan, Inner Tian Shan, Dzhany-Dzher Mt range, upper Mudyuryum riv. Ak-Baytal gorge h~3300 m, 15.VII.2011, S.Toporov leg. L=16 mm
Copyright © Ruslan Panin

Carabus (Apoplesius) deuvei hajdaji Hackel, 2007 Holotype Holotype
Male adult
Turkey- prov. Mus 60 km NW Mus 1630m
3856N 4108E Bulgan Gec. 23-24.06.03 E. & P. Hajdaj leg.
Copyright © Boleslav Brezina

Carabus (Apoplesius) omphreodes Reitter, 1898
Male adult

Copyright © Kirill Makarov

Carabus (Apoplesius) pseudodepressus Machard, 1988
Female adult
Turkey, Prov. Bitlis, S near Tatvan, 23.IV.2009 leg. M. & S. Nabozhenko
L=19.0 mm
Copyright © Kirill Makarov

Carabus (Apotomopterus) bousquetellus gengmaensis Kleinfeld, 2003 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) cantonensis metallaicus Kleinfeld, 1999 Holotype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) davidis neodavidis Kleinfeld & Schutze, 2001 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) casaleianus casaleianus Deuve, 1994
Female adult
China, SW Hunan Prov., Chengbu Co, Nan-Shan-Ding (26.34;110,08) 1-10.VIII,2006, I.Rapuzzi leg. L=33 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) davidis davidis Deyrolle & Fairmaire, 1878
Male adult
SE-China, Jiangxi prov., Yushan County, Mt Wangjing, VI,2012
L=38 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) patroclus vicarius Cavazzuti, 1996
Male adult
China, S Sichuan, / Yunnan border, W of Lugu lake, S of Yongning vill., h=2700m, 11-18.VI.2017, L=31 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) protenes ascendens Semenov, 1898
Male adult
China, Chongqing, N Jinfo Shan, Shanwangping h=1400m, 29.101944, 107.166389, 2-8.VI.2018, C.Reuter leg./det. F.Kleinfeld, L=26 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) protenes ascendens Semenov, 1898
Endophallus
China, Chongqing, N Jinfo Shan, Shanwangping h=1400m, 29.101944, 107.166389
Copyright © Ruslan Panin

Carabus (Apotomopterus) microtatos lingbaonis Imura, 2019
Endophallus
China, CW-Yunnan, Nánjiàn Yízú Zìzhìxiàn, Mt Lingbao h=2400 m, 24.768611, 100.500000
Copyright © Ruslan Panin

Carabus (Apotomopterus) mecynodes pinchuanensis Imura, 1999
Male adult
China, S-Sichuan, 40-50 km E Xichang (to Zhaojue) 27°51'N 102°30'E, h=2700-3000m, Pine forest, 5-20.VI.2017, C.Reuter leg., L= 29 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) monticola fontanae Born, 1906
Male+Female adult
N-Italy, Piemonte, Bielmonte, h1350 m, 14-15.VI.2010, Pascal Stefani leg.; female, 23 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) montivagus montivagus Palliardi, 1825
Male+Female adult
SW-Romania: Banat, Caras-Severin Reg., Sasca Montana – Stinapari, 21.V.-5.VI.2010, Krause Rene leg.<br /> female, 26 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) pseudomonticola Lapouge, 1908 ? nemoralis
Male adult
NE Spain, Catalonia, Sant Jaume de Frontanyà, h=1000 m, V-VI.2010, Pascal Stefani leg.
L=26 мм
Copyright © Ruslan Panin

Carabus (Archicarabus) steuartii Deyrolle, 1852
Female adult
NW-Spain, Pontevedra prov., Serra do Candán Mt range, 1.III.1992, S. Devesa leg./det. S. Sobrino
Copyright © Ruslan Panin

Carabus (Archicarabus) monticola roccae Born, 1923
Male adult
NW-Italy, Liguria: Savona env., Monte San Giorgio, 30.X.1977, Monauzzi leg. L=23 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) phoenix Lapouge, 1924
Male+Female adult
C-Lebanon, Byblos /Jbeil Distr., Kfardebian env., h=1200m, XI.2014, C. Reuter leg. female,23mm
Copyright © Ruslan Panin

Carabus (Archicarabus) wiedemanni congruus Motschulsky, 1865
Female adult
Turkey, Cankiri, Cerkes - Kursunlu, Atkaracalar 1250m 10.7.1996 Leg Staven and Skoupy (DUBC)
Copyright © Alexander Anichtchenko

Carabus (Archicarabus) wiedemanni wiedemanni (as mussardianus Breuning, 1965)
Female adult
NC-Turkey, Sinop province, Erfelek env., h=200m, 10-27.VII.2014, C. Reuter leg. L=24 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) wiedemanni (as mussardianus Breuning, 1965)
Male adult
NC-Turkey, Sinop province, Erfelek env., h=200m, 10-27.VII.2014, C. Reuter leg. L=23 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) monticola monticola (as maritimensis Born, 1923)
Female adult
SE France, Alpes-Maritimes, Col de Braus h=1000 m, 23.V-18.VI.2014, P. Cavazzuti & G. Proscia leg., L= 24 mm
Copyright © Ruslan Panin

Carabus (Aristocarabus) viridifossulatus viridifossulatus Fairmaire, 1887
Female adult
35mm
Copyright © Boleslav Brezina

Carabus (Aristocarabus) viridifossulatus seticollis Deuve & Mourzine, 2000
Male adult
28 mm
Copyright © Boleslav Brezina

Carabus (Aulonocarabus) gossarei imanensis Lafer, 1989
Male adult

Copyright © Kirill Makarov

Carabus (Aulonocarabus) gaschkewitschi czarensis Obydov, 1997
Female adult
Russia, CE-Siberia, N Chita reg., Kodar Mt. Range, Big Leprindo Lake env. h~2000m., 2-5.VII.2002 A. Prokhorov leg./det. A. Anichtchenko
L=19 mm
Copyright © Ruslan Panin

Carabus (Aulonocarabus) kurilensis ? daisetsuzanus Kono, 1936
Male adult
N-Japan, Hokkaido, Katō District,Kamishihoro, Horoka near Maruyama Bridge,19.VI.2013, T. Hirabayashi leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Aulonocarabus) gaschkewitschi gaschkewitschi Motschulsky, 1859
Male adult

Copyright © Kirill Makarov

Carabus (Aulonocarabus) kolymensis kodarensis Brinev, 1997
Male adult
Russia, SE Siberia, N Chita reg., Kodar Mt. Rng. h=1800m, Novaya Chara vill., 23.VI.2006. L=17 mm
Copyright © Ruslan Panin

Carabus (Aulonocarabus) truncaticollis fleischeri Reitter, 1898
Female adult
Russia, N Burajatia, Barguzinskiy Mts., the upper reaches of M.Chivyrkuy riv., h=2000 m, 26.VI-12.VII.2007, Vashchenko S leg. L=18.5 mm
Copyright © Ruslan Panin

Carabus (Aulonocarabus) truncaticollis fleischeri Reitter, 1898
Male adult
Russia, N Burajatia, Barguzinskiy Mts., the upper reaches of M.Chivyrkuy riv., h=2000 m, 26.VI-12.VII.2007, Vashchenko S leg. L=17 mm
Copyright © Ruslan Panin

Carabus (Axinocarabus) fedtschenkoi fedtschenkoi Solsky, 1874
Male adult
27 mm
Copyright © Boleslav Brezina

Carabus (Axinocarabus) fedtschenkoi kondarensis Kryzhanovskij, 1953
Female adult
NW-Tadzhikistan: Gissar Mt.Rng, Varzob river ravine, Gushari env., h~1400-1500m, 24.III.2010, L=29 mm
Copyright © Ruslan Panin

Carabus (Calocarabus) linxiaensis ponggartangensis Deuve, 1995 Holotype
Male adult
17 mm
Copyright © Boleslav Brezina

Carabus (Cryptocechenus) sifanicus Semenov, 1898 (as oxydocupreus)
Male adult
male, 17mm
Copyright © Boleslav Brezina

Carabus (Carabus) arvensis sylvaticus Dejean, 1826
Female adult
France, Centre reg., Eure-et-Loir, Forкt de Montecot, VI.2009, Pascal Stefani leg.
female, 20 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) arcensis sylvaticus Dejean, 1826 (as sugintensis Brunier, 1922)
Male adult
NW -France, Haut-Rhin Depart., Fulleren, XI.2004, Pascal Stefani leg.
Copyright © Ruslan Panin

Carabus (Carabus) arvensis arvensis Herbst, 1784
Male+Female adult
W Ukraine, NW Podolian, Lviv reg., Javoriv distr., near Stradch vill., 20-30.IV.2009, Panin R. leg.
Copyright © Ruslan Panin

Carabus (Carabus) arvensis arvensis Herbst, 1784
Male adult
W. Ukraine , NW. Podolian, Lviv reg., Nikolaev distr., near Krasiv vill., 9.II.08, Panin R. leg. L=17mm
Copyright © Ruslan Panin

Carabus (Carabus) menetriesi menetriesi Hummel, 1827
Male adult
W-Ukraine, Roztochia, Lviv Reg., Yavorov distr., near Ivano-Frankovo vill., Site Zalyvki, Panin R. leg., 1-20.V.2005, male, 21 mm.
Copyright © Ruslan Panin

Carabus (Carabus) arvensis faldermanni Dejean, 1829
Male+Female adult
Russia, Far East, Primorskiy Kray, Ussuriysk Distr., Kaymanovka vill., 11.VII-1.VIII..2011, A. Korshunov leg.
female, 23 mm
Copyright © Ruslan Panin

Carabus (Carabus) arvensis hokkaidoensis Lapouge, 1924
Female adult

Copyright © Kirill Makarov

Carabus (Carabus) granulatus yezoensis Bates, 1883
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Carabus (Carabus) stscheglowi Mannerheim, 1827
Male adult
Russian, Chuvashia, Cheboksary, Zavolzhye distr., , near lake Astrakhanka, L=27 mm
Copyright © Ruslan Panin

Carabus (Carabus) sculpturatus (as gilanicus Mandl, 1967)
Female adult
N-Iran:Gilan Prov., Asalem Distr., Siahbil, deciduous forest (very wet) h= -10 m, 20-24.VI.2017, V. Čutka leg., L=22 mm
Copyright © Ruslan Panin

Carabus (Cathaicus) brandti brandti Faldermann, 1835 (as grossegranulatus) Syntypes Lectotype
Male adult
C. (Cathaicus) brandti grossegranulatus Breuning 1943<br /> Lectotypus/SyntypusLectotypus (25 mm) von "South Mandschou". Syntypus (25 mm) "South Mandschou, 30. V. 1937"
Copyright © Frank Kleinfeld

Carabus (Cathoplius) aliai Escalera, 1944
Male adult
male, 27mm
Copyright © Boleslav Brezina

Carabus (Cathoplius) asperatus asperatus (Dejean, 1826)
Male adult
male, 24mm
Copyright © Boleslav Brezina

Carabus (Cechenochilus) heydenianus Starck, 1889
Male+Female adult
NW-Caucasus,
Krasnodar reg., Mostovskoy distr., Skirda Mt Range, N slopes Yatyrgvart Mt. h=2300-2500m., 09-10.VII.2008, D. Fominykh leg.
female,19mm/male,18mm
Copyright © Ruslan Panin

Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Holotype
Male adult
Russia, Karachai-Circassian Republic, Abishira-Akhuba Mount Range between passes Agur and Mylgvar, 2785–2830 m.
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Paratype
Male adult
Russia, Karachai-Circassian Republic, Abishira-Akhuba Mount Range between passes Agur and Mylgvar, 2785–2830 m.
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) kokujewi Semenov, 1898
Male adult
Russia, Karachai-Cherkessia, vall. Daut Riv., near Glacier Ryndzhi-Murudzhu, h = 3050 m, 22.VIII.2011, leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) kokujewi Semenov, 1898
Male adult
Russia, Karachai-Cherkessia, vall. Daut Riv., near Glacier Ryndzhi-Murudzhu, h = 3050 m, 22.VIII.2011, leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) kokujewi Semenov, 1898
Female adult
Russia, Karachai-Cherkessia, vall. Daut Riv., near Glacier Ryndzhi-Murudzhu, h = 2550 m, 23.VIII.2011, leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) boeberi boeberi Adams, 1817
Female adult
Грузия, Мцхета-Мтианети, к северу от пос. Гудаури, ЮВ отроги г. Байдарская (нижний пик), по ущелью ручья, альпика, под камнями, точка 2, h = 2700 м, 42.495413° N, 44.487844° E, 10.08.2019, leg. И.А., А.И., С.В. Солодовниковы, О.И. Данилова 26 mm
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) boeberi tschitscherini Semenov, 1896
Male adult
Russia, C Caucasus, Ingushetia, S slopes of Skalistyi Mt.R.
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) heydenianus Starck, 1889
Male adult
Carabus (Procechenochilus) heydenianus euxinus Semenov, 1896 Abkhasia: Arabika plato (TOPOTYPUS)
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) boeberi aequaliceps Reitter, 1896
Female adult
Russia, Dagestan, Andiyskiy Mt.R., near Andiyskiye vorota pass, 06.VI–18.VII.2021, leg. D. Fominykh
Copyright © Igor Solodovnikov

Carabus (Cephalornis) potanini (Semenov, 1887)
Male adult
China, S Gansu, 3400m, Minshan mts.,
Wenxian 8.92, J. M. Bousquet
Copyright © Boleslav Brezina

Carabus (Chaetocarabus) intricatus bayardi Solier, 1835
Male+Female adult
S-Italy,
Basilicata Reg., Potenza Prov., Latronico, F. Angelini leg. 14.VI.2002
female, 26mm
Copyright © Ruslan Panin

Carabus (Heterocarabus) marietti marietti (as ativsengueni Schweiger, 1962)
Male adult
male, 17mm
Copyright © Boleslav Brezina

Carabus (Chaetocarabus) intricatus krueperi Reitter, 1896
Male adult
CE-Greece: Pieria, Stomio (Larissa) Ossa Mt. 7-14.VII.2009, Krause Rene leg.
L=35 mm
Copyright © Ruslan Panin

Carabus (Chaetocarabus) arcadicus arcadicus Gistl, 1850
Female adult
CE-Greece, Kato Olympos, Karia env., 6-17.VII.2011, Krause Rene leg. L=37 mm
Copyright © Ruslan Panin

Carabus (Heterocarabus) marietti marietti Cristoforis & Jan, 1837
Female adult
as ativsengueni Schweiger, 1962
NW Turkey, Bolu, Abant Mts.; 16mm

Copyright © Aleš Sedláček

Carabus (Chaetocarabus) arcadicus arcadicus Gistl, 1850
Male adult
Graecia, Pindus, Mt. Velouchi (Timfristos), 1200m, vi.1970-A. Sermet Leg., 30 mm
Copyright © Noël Mal

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Male adult
W Ukraine, Lviv city, Sykhiv, Zubra forest, (Osvytskiy Grove), under bark, 49°47'28.10" N, 24° 2'20.09" E, 10.XI.2017, Panin R leg.
Copyright © Ruslan Panin

Carabus (Chaetocarabus) intricatus bayardi (as silaensis Haury, 1881)
Female adult
S-Italy, Calabria, Sila Grande Massif, Camigliatello h=1040 m, 19.XI.2009, G. Proscia leg. L=31 mm
Copyright © Ruslan Panin

Carabus (Chaetocarabus) intricatus bayardi (as silaensis Haury, 1881)
Male adult
S-Italy, Calabria, Sila Grande Massif, Camigliatello h=1040 m, 19.XI.2009, G. Proscia leg. L=28 mm
Copyright © Ruslan Panin

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Female adult
W Ukraine, Lviv city, Holosko, Bryukhovytsʹkyi forest 49.886833, 24.012083 under bark, 10.XI.2024, R. Panin leg., L=35 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792
Male adult
Austria,
Weissenbach stt., III.1979, Kretschmer leg.
male, 24mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens costellatus Gehin, 1882
Male+Female adult
C-FRANCE,
Limousin Reg., Corrиze Dep., Aubazines, X.2005, Pascal Stefani leg.
female,26mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens escheri Palliardi, 1825 (ab. opacus Haury, 1878)
Male+Female adult
W Ukraine, NE Carpathians Mts.,<br /> Transcarpathian reg., Rakhov Distr., Chornogora range, h=2020m, alpin zone, 28.VI-22.VIII.2009, Panin R. leg.<br /> female, 27mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) lineatus lateralis Chevrolat, 1840 (as quasilineatus Tarrier, 1975)
Male+Female adult
NW Spain,
E Asturias, Monte de Moal h~1000m., 5.VI.2008, Pascal Stefani leg.
female, 27mm.
Copyright © Ruslan Panin

Carabus (Chrysocarabus) solieri solieri Dejean, 1826
Female adult
SE France, Alpes-Maritimes, Ascros, VI.2011
L=30 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896
Male adult
Anyos (ANDORRE), h=1450ms,
5-VI-2010, P. Stefani leg.
body lenght - 32 mm.
Copyright © Alexandr Safronov

Carabus (Chrysocarabus) lineatus lineatus (as pseudonumancianus Mollard, 2010)
Male adult
NC-Spain, Sierra de Cantabria , Sierra de Toloño Rng, Puerto de Herrera , h=1100m, 19.I,2007 25 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens escheri Palliardi, 1825 (as laevipennis Seidlitz, 1891)
Male adult
W Ukraine, NE Carpathians,Transcarpathian Region, Rakhiv Distr., Chornogora Mts Range, h=2000 m, alpin zone, V-28.VI..2014, Panin R. leg.
Copyright © Ruslan Panin

Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896
Habitat
Andorra, Anyos, ii.2012, J. Nicolas Leg. & Photos
Copyright © Noël Mal

Carabus (Chrysocarabus) solieri clairi Gehin, 1885 (as devillei Tarrier, 1972)
Male adult
SE-France, Alpes-Maritimes, Beuil env., h=1550m, 14.VII.1986
Copyright © Ruslan Panin

Carabus (Coptolabrus) pustulifer guerryi Born, 1903
Female adult
N Yunnan: Gaoligongshan, h =1300 m, 06.2003 Local people
Copyright © Igor Solodovnikov

Carabus (Coptolabrus) principalis Bates, 1889
Male+Female adult
Zi-Ding-Shan, h=1406 m, Guiyang County, Binzhou City, Hunan, CHINE
19-29.IV.2008
Copyright © Alexandr Safronov

Carabus (Coptolabrus) formosus sunpanensis Semenov, 1898 (as rostandi Deuve, 1991)
Male adult
China, N Sichuan province, Venchuan, h=1400 m., 30.VI.2007, A. Gorodinski leg.;
L=33 mm
Copyright © Anton Menstschikov

Carabus (Coptolabrus) elysii pulcher Kleinfeld, 1997 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Coptolabrus) augustus pustuliferus Kleinfeld, 1998 Holotype
Male adult

Copyright © Frank Kleinfeld

Carabus (Coptolabrus) formosus bousqueti Deuve, 1993
Male adult
C-China, S-Gansu prov., Wenxian County, Mt.Guanzigou,1-14.V.2010, local coll. leg.
L=32 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) formosus brockmanni Kleinfeld & Schutze, 1997
Female adult
C-China, S-Shaanxi: Shangzhou City, Shahezi,1-14.VI.,2013, local coll. leg.
L=39 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) formosus wanxianicus Imura, 1995
Male adult
China, NE-Sichuan, Dazhou, Wanyuan env., 6.VI.2005. L = 33mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) smaragdinus furumiellus Deuve, 1994
Female adult
NE-China, E-Liaoning, Benxi, Mt Liuershan, VIII.1998. L=39 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) pustulifer mirificus Kraatz, 1894
Male adult
SC-Chine, NW-Hunan, Longshan County, Mt. Maoping,10-30.VI.2008, L=44 mm
Copyright © Ruslan Panin

Carabus (Cratocarabus) puer Morawitz, 1886
Female adult
female, 16mm
Copyright © Boleslav Brezina

Carabus (Cratocechenus) jacobsoni Semenov, 1898
Male adult
S Kazakhstan, Almaty reg., Big Lake, h=2800 m, 25.VI.96
male
L=14 mm
Copyright © Maksim Smirnov

Carabus (Cratocechenus) akinini akinini Morawitz, 1886 (as oitalensis ) Paratype
Male adult
male, 20mm
Copyright © Boleslav Brezina

Carabus (Cratocechenus) akinini akinini Morawitz, 1886
Female adult
female, 21mm
Copyright © Boleslav Brezina

Carabus (Cratocechenus) ovtchinnikovi ovtchinnikovi Gottwald, 1987
Male adult
N-Kyrgyzstan, Kyungey Ala-Too Mt Rng, Cholpon-Ata env., h=2500m, 19.VI.2000,
L=19 mm
Copyright © Ruslan Panin

Carabus (Cratocephalus) solskyi Ballion, 1878
Male adult
SE Kazakhstan, Ketmen Mt. rng., Tuyuk vill., 16.06.97 leg. S.Toropov
Copyright © Maksim Smirnov

Carabus (Cratocephalus) solskyi Ballion, 1878
Female adult
SE-KAZAKHSTAN, Ketmen Mt Rng, h=1800m, Tujuk vill.,15.V.2004
L=36 mm
Copyright © Ruslan Panin

Carabus (Cratocephalus) cicatricosus cicatricosus Fischer, 1842
Male adult
SE-Kazakhstan, Almaty reg., C Trans-Ili Alatau N-slopes, Kaskelen Valley, Izvestkovyi vill., h=1700 m, 6.V.2015, D.Kovalchuk leg., L=28.5 mm
Copyright © Ruslan Panin

Carabus (Cratophyrtus) turcosinensis medvedevi Kryzhanovskij, 1968
Female adult
S Kazakhstan, NW slope of Ugamskii mt.rng., 3 km W Sairamskii peak, 2200m
tributary of Karzhantau riv., 15.6.2004 Matalin A. leg.
20 mm
Copyright © Alexander Anichtchenko

Carabus (Cratophyrtus) kaufmanni segregatus Morawitz, 1886
Female adult
female, 19mm
Copyright © Boleslav Brezina

Carabus (Cryptocarabus) subparallelus Ballion, 1878
Female adult
Kazakhstan,
near Alma-Ata, V.2009, Pak O.leg.
female, 17mm
Copyright © Ruslan Panin

Carabus (Cryptocarabus) lindemanni Ballion, 1878
Female adult
SE Kazakhstan, Alma-Ata env., 16.04. 2000
L=16,0 mm
Copyright © Maksim Smirnov

Carabus (Cryptocarabus) ? turkestanus turkestanus Breuning, 1929
Male adult
Kyrgyzstan, W Kungei Ala-Too Mt Rng, Chon-Kemin riv., Kyzyl-Bairak vill., 25.IV.2010. L= 18 mm
Copyright © Ruslan Panin

Carabus (Cryptocarabus) tsharynensis Kabak 1994
Male adult
SE-Kazakhstan, Almaty Region, W-Zailiyskiy Alatau rng, Belshabdar valley h~1300 m, 25-27.VI.2002, A.Puchkov leg. L=17 mm
Copyright © Ruslan Panin

Carabus (Cryptocarabus) tsharynensis Kabak 1994
Female adult
E-Kazakhstan, Almaty Region, W-Zailiyskiy Alatau, Turgen distr., Tauturgen vill., h~1200 m, 8-13.VI.2002, A.Puchkov leg. L=19 mm
Copyright © Ruslan Panin

Carabus (Cryptocarabus) sacarum Kabak, 1998
Male adult
SE-Kazakhstan, Almaty reg., C-Zailiyskiy Alatau N-slopes, Kaskalen river Valley, Izvestkovy vill., h=1700 m, 6.V.2015, D.Kovalchuk leg./det. R.Panin L=18 mm
Copyright © Ruslan Panin

Carabus (Cryptocarabus) sacarum Kabak, 1998
Female adult
SE-Kazakhstan, Almaty reg., C-Zailiyskiy Alatau N-slopes, Kaskalen river Valley, Izvestkovy vill., h=1700 m, 6.V.2015, D.Kovalchuk leg./det. R.Panin L=20 mm
Copyright © Ruslan Panin

Carabus (Ctenocarabus) galicianus Gory, 1839
Male adult
Spain,
Sierra de la Cabrera Baja, 8 km SW Truchillas, 9.VII.2008, Anichtchenko A. leg.
L= 20 mm
Copyright © Alexander Anichtchenko

Carabus (Ctenocarabus) galicianus Gory, 1839
Female adult
NW-Spain, Pontevedra prov., Campo Lameiro env., 15.VIII.1994, S.Sobrino leg.
L=28 mm
Copyright © Ruslan Panin

Carabus (Ctenocarabus) melancholicus costatus Germar, 1824
Female adult
C-Spain, Madrid region, Rozas de Puerto Real env.,21.XII.2008, T.Gazurek leg.
L=27 mm
Copyright © Ruslan Panin

Carabus (Ctenocarabus) melancholicus melancholicus Fabricius, 1798
Male adult
NW Morocco, Kénitra, Souk El Arbaa env., II.1985, V.Olivella leg. L=25 mm
Copyright © Ruslan Panin

Carabus (Ctenocarabus) melancholicus submeridionalis Breuning, 1975
Male adult
S- Spain, Cádiz, Tarifa, El Cabrito, 28.XII.2019, F. Garcia leg. L=26 mm
Copyright © Ruslan Panin

Carabus (Ctenocarabus) melancholicus costatus Germar, 1824
Male adult
NW Portugal, Viana do Castelo distr., Sierra de Agra, 41°47'52.3"N 8°41'27.6"W, h=786 m, 3.VII.2016, D. Prunier leg.
Copyright © Ruslan Panin

Carabus (Cupreocarabus) kangdingi kangdingi Korell, Kleinfeld & Gorgner, 1992 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Cupreocarabus) huangi huangi Deuve, 1992
Male adult
C-China, E Tibet, N-Sichuan, Hongyuan,valley 10 km SE Sanggarpar, h=4200m, 32.300053, 102.584031, alpine meadow / scree, 19.VII.1995, B.Brezina leg.<br /> L=21 mm
Copyright © Ruslan Panin

Carabus (Cupreocarabus) lixianensis lixianensis Deuve, 1990
Male adult
China, E Tibet, N-Sichuan, (Barkam),valley SW of Barkam, h=3200m, 31*53’ N; 102*12’E, mixed/coniferous forest, 22.VII.1995, K.& B.Brezina
L=23 mm
Copyright © Ruslan Panin

Carabus (Cupreocarabus) alpherakii alpherakii Semenov, 1898
Female adult
China, W Sichuan, Litang env., h~5000 m,29.VII -3.VIIII.1995, J. Schneider leg. L=23 mm
Copyright © Ruslan Panin

Carabus (Cupreocarabus) mollardianus Deuve, 2016
Female adult
Chine, Sichuan, Lushan, Mt. Jiajin Shan, 10.VII.2016 local collector leg./ det. R.Panin, L=22 mm
Copyright © Ruslan Panin

Carabus (Cupreocarabus) ? siguniangensis jiachingensis Lassalle, 2003
Male adult
Chine, Sichuan, Lushan, Mt. Jiajin Shan, 10.VII.2016 local collector leg., L=21 mm
Copyright © Ruslan Panin

Carabus (Cupreocarabus) balangicus balangicus Cavazzuti, 2002
Male adult
Chine, C-Sichuan, Xiaojin Co, Wolong valley, Balang Shan Pass h=3448 m, 30.884617, 102.973117, 9.VI.-1.VII.2011, G. Proscia leg. L=19 mm
Copyright © Ruslan Panin

Carabus (Cupreocarabus) balangicus balangicus Cavazzuti, 2002
Male adult
Chine, C-Sichuan, Xiaojin Co, Wolong valley, Balang Shan Pass h=3448 m, 30.884617, 102.973117, 9.VI.-1.VII.2011, G. Proscia leg. L=20 mm
Copyright © Ruslan Panin

Carabus (Cupreocarabus) huangi huangi Deuve, 1992
Male adult
China, E Tibet, N-Sichuan, 53 km NE of Barkam, Pass Sandaoban env., h=4293 m, 32.232950, 102.598533, 11-29.VI.2011, G. Proscia leg., L=21 mm
Copyright © Ruslan Panin

Carabus (Cupreocarabus) huangi huangi Deuve, 1992
Female adult
China, E Tibet, N-Sichuan, 53 km NE of Barkam, Pass Sandaoban env., h=4293 m, 32.232950, 102.598533, 11-29.VI.2011, G. Proscia leg., L=21.5 mm
Copyright © Ruslan Panin

Carabus (Cychrostomus) pseudoprosodes Semenov & Znojko, 1932
Male adult
male, 19mm
Copyright © Boleslav Brezina

Carabus (Cychrostomus) anchocephalus lanzhouensis Deuve, 1991 Paratype
Male adult
male, 21mm
Copyright © Boleslav Brezina

Carabus (Cychrostomus) anchocephalus anchocephalus Reitter, 1896
Male adult
male, 22mm
Copyright © Boleslav Brezina

Carabus (Cyclocarabus) aulacocnemus galinae Obydov, 2003 Paratype
Male+Female adult
W-Tianshan:Karatau Mts.,N-part:Boialdyr River valley,
N of Ken-su, 600-750m, 22-V-2000 М. Danilevsky leg.
male - 20 мм, female - 19 мм.
Copyright © Alexandr Safronov

Carabus (Cyclocarabus) namanganensis namanganensis Heyden, 1886
Male+Female adult
Kazakhstan, S. Karatau Mts. R., Mashat vill. env.
23-IV-1993 М. Danilevsky leg.
male - 21 мм, female - 22 мм.
Copyright © Alexandr Safronov

Carabus (Cyclocarabus) zarudnyi Semenov & Znojko, 1932
Female adult
NE-Uzbekistan, W-Tian-Shan, NW slopes Pskem Mt. Rng, 8-14 km E Pskem, 2100-2500 m, 16.VI.1998, A. Puchkov leg., L=21 mm
Copyright © Ruslan Panin

Carabus (Cyclocarabus) karaterekensis hemicallisthenes Kabak & Kalab, 1997
Male adult
NW- Kyrgyzstan, W-Tian-Shan, Chandalash Mt.Rng, Chakmak-Suu vill., 3.VI.2000, L=17 mm
Copyright © Ruslan Panin

Carabus (Cyclocarabus) brosciformis Semenov, 1896
Male adult
NE-Uzbekistan, W-Tian-Shan, Chatkal Mt Range, Tashkent reg., Yangiabad env., h=2100m, 30.IV.2002, L=23.5 mm
Copyright © Ruslan Panin

Carabus (Damaster) blaptoides oxuroides Schaum, 1862
Female adult
Japan, C-Honshu, Ofuse-cho, Kamitakai, Nagano Pref., 12.VI.2006, Ohshima leg.
L=34 mm
Copyright © Ruslan Panin

Carabus (Damaster) blaptoides capito Lewis, 1881
Male adult
Japan: Niigato pref., Sado Isl., Sado, Yamato
Copyright © Igor Solodovnikov

Carabus (Damaster) blaptoides oxuroides Schaum, 1862
Female adult
Japan, C Honshu, Nagano Prefecture, Nagano city, 3.IV.2012, S. Shinichi leg. L=42 mm. det. Anichtchenko A. 2021
Copyright © Ruslan Panin

Carabus (Damaster) blaptoides rugipennis Motschulsky, 1861
Male adult
Kunashir isl.
Copyright © Kirill Makarov

Carabus (Damaster) blaptoides blaptoides (Kollar, 1836)
Female adult
Japan, SW Honshu, Shimane Pref., Unnan city, 16.I2016, S.Shinichi leg., L=48 mm
Copyright © Ruslan Panin

Carabus (Deroplectes) staudingeri Ganglbauer, 1886
Female adult
N Tajikistan, SW Pamir-Alay, Fann Mts h=1800m, Padrut vill., 2-3.V.1997
L=28 mm
Copyright © Ruslan Panin

Carabus (Deroplectes) arcanus arcanus Semenov, 1898
Male adult

Copyright © Boleslav Brezina

Carabus (Deroplectes) dokhtouroffi dokhtouroffi Ganglbauer, 1886
Male adult
NC Tadzhikistan, Pamir, N-slope Peter I.Mt Rng, h=2100m,Ganishou vill., 10-12.VI.2006, Ivanova E. leg.
L=28 mm
Copyright © Ruslan Panin

Carabus (Deroplectes) klapperichianus coiffaitianus Deuve, 1990
Male adult
Tadzhikistan: Badakhshan: Ishkashim Mt.R.: Kuh-i-Lal, VI.1979 leg. I.K. Lopatin (TOPOTYPUS)
Copyright © Igor Solodovnikov

Carabus (Deroplectes) klapperichianus coiffaitianus Deuve, 1990
Female adult
Tadzhikistan: Badakhshan: Ishkashim Mt.R.: Kuh-i-Lal, VI.1979 leg. I.K. Lopatin (TOPOTYPUS)
Copyright © Igor Solodovnikov

Carabus (Deroplectes) klapperichianus salangicus Kleinfeld & Reuter, 2010 Paratype
Female adult
N-Afghanistan, Hindukush Mt.Rng, Salang Pass, Nordseite, 25°27'.494N, 68°58.399'E, ca. reg. subalp., h = 2200-2400 m, V.2010, leg. Christoph Reuter
Copyright © Igor Solodovnikov

Carabus (Diocarabus) linnaei macairei Dejean, 1826
Female adult
SW-Romania, Banat, Resita - Valiug, 29.V- 9.VI.2005, Krause R. , Baros, Pittner leg.<br /> L=19 mm
Copyright © Ruslan Panin

Carabus (Diocarabus) linnaei linnaei Panzer, 1812
Male adult
W-Ukraine, NW podolian, <br /> Lviv Reg., Zhydachiv distr., near Vybranivka, 17.II.2007,Panin R. leg.<br /> female, 21 mm
Copyright © Ruslan Panin

Carabus (Diocarabus) fraterculus fraterculus Reitter, 1895
Male adult
N-Korea, N Pyongan Prov.,Sakju, Mt Ponsan, 20-29.VI.2009, L. Bieber leg.<br /> L=19 mm
Copyright © Ruslan Panin

Carabus (Diocarabus) linnaei linnaei (as scopolii Dejean, 1826)
Male adult
W Ukraine, Lviv reg., NW Podolian Upland, forest near Vybranivka vill., 12.X.2007, Panin R. leg. L=19 mm
Copyright © Ruslan Panin

Carabus (Diocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI.-5,VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=17 mm
Copyright © Ruslan Panin

Carabus (Diocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Female adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=18 mm
Copyright © Ruslan Panin

Carabus (ODiocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=16.5 mm
Copyright © Ruslan Panin

Carabus (Diocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Gutyn-Tomnatyk h~2015 m alpin zone, 8.VI-28.VII.2011, R. Panin R. leg. L=14 mm
Copyright © Ruslan Panin

Carabus (Diocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg.
Copyright © Ruslan Panin

Carabus (Diocarabus) linnaei folgariacus (as tauernensis Mandl, 1985)
Female adult
Austria, Steiermark, Rottenmanner Tauern, h=1000 m, Strechengraben, 13.VII.2012, L=20 mm
Copyright © Ruslan Panin

Carabus (Lasiocoptolabrus) sunwukong sunwukong Imura, 1993 Paratype
Male adult

Copyright © Boleslav Brezina

Carabus (Lasiocoptolabrus) sunwukong baotianmanus (Imura, 2005) Paratype
Male adult

Copyright © Boleslav Brezina

Carabus (Eccoptolabrus) exiguus nivium Breuning, 1933
Female adult
female, 20mm
Copyright © Boleslav Brezina

Carabus (Eccoptolabrus) exiguus exiguus Semenov, 1898 Holotype
Male adult
male, 18 mm
Copyright © Frank Kleinfeld

Carabus (Eucarabus) sternbergi Roeschke, 1898
Female adult
Korea, Gyeongi Do,
Bokwang Sa 30.2.1976
female, 28 mm
Copyright © Alexander Anichtchenko

Carabus (Eucarabus) manifestus pyonganicola Deuve & Li, 1998
Male+Female adult
N-Korea: Pyonganam-Do, YangTok County,(39;127), 12-19,VI.2006
Copyright © Ruslan Panin

Carabus (Eucarabus) obsoletus obsoletus Sturm, 1815
Male adult
NW Ukrainian Carpathians, Lviv Reg., Staryi Sambor Distr., near Dobromyl , h=500 m., 15-24.VI.2005, Panin R. leg., male, 28mm
Copyright © Ruslan Panin

Carabus (Eucarabus) obsoletus nagyagensis Seidlitz, 1888
Female adult
NW-Romania, Mts Ses (Plopis),Cluj Reg., Bucea env, h=580 m, 8.VI.2011, Sl.Budka leg.
L=26 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825
Male+Female adult
Male - SW Romania, Banat - Bocsa/Coltan,
21.V-05.VI.2010, R. Krause leg., L=29 mm.
Female - SW Romania, Banat - Saska Montana, Stinapari,
25-VI-2010, R. Krause leg., L=31 mm.
Copyright © Alexandr Safronov

Carabus (Eucarabus) obsoletus nagyagensis (as paranagyagensis Lie, 1990)
Male adult
W Romania, Arad reg., Zarand Mts (Muntii Zarandului), Nadas-Barzavah h=142 m, J. Kynkor leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) manfredeggeri Müller 2012 Paratype
Male+Female adult
N-Korea, Chagang Prov., Manpo City, Songhak-ri, 2-16.VII.2011<br /> female, 21 mm<br /> male, 19 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) ulrichi arrogans Schaum, 1858
Male adult
NE Serbia, Majdanpek; 23 mm
Copyright © Aleš Sedláček

Carabus (Eucarabus) obsoletus nagyagensis (as fossulifer Fleischer, 1893)
Male adult
NW-Romania, Bihor, Mts Seș (Plopis), 20 km N Aleșd, Pădurea Neagră env., VI.2014, J.Kynkor leg. L=24 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) parreyssi parreyssi Palliardi, 1825
Male adult
W-Croatia, NP Northern Velebit, Zavižan,15.V-13.VI.1996, Colombetta leg., L=21 mm
Copyright © Ruslan Panin

Carabus (Euleptocarabus) porrecticollis porrecticollis Bates, 1883
Male adult
Japan: Niigata pref., Chayagahara Jyouetsu City,
18.XI.2006 A.Toba leg.
Copyright © Alexander Anichtchenko

Carabus (Euleptocarabus) porrecticollis Bates, 1883
Female adult
Atsushiokanomachikano, Kitakata City, Fukushima Pref., Japan, 29.III.2014, T. Nakamura leg.
Copyright © Alexander Anichtchenko

Carabus (Euleptocarabus) porrecticollis porrecticollis Bates, 1883
Male adult
Japan, C Honshu, Nagano Pre., Toyooka town, 28.XII.2019, S. Shinichi leg. L=30 mm
Copyright © Ruslan Panin

Carabus (Eupachys) glyptopterus Fischer, 1827
Male+Female adult
Russia, SE-Siberia,
Buryat Republic, Ivolginsky Distr., 04.VII.2010, Filippov A.E. leg.
female, 32 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Eurycarabus) famini lucasi Gaubil, 1849
Male adult
N-Tunisia, Bizerte reg., Menzel Bourguiba city,h=5 m, XII.1981, S. Remond leg.
L=25 mm
Copyright © Ruslan Panin

Carabus (Eurycarabus) famini fezzanus Lapouge, 1915
Male adult
Morocco, Middle Atlas Mt Rng, Azrou city h=1500m, IV.1999, S. Remond leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Eurycarabus) famini piraticus (as mamorensis (Kocher, 1964)
Female adult
NW-Morocco, betw.Meknes/Kenitra, Foret de Mamora, II.2008
L= 27 mm
Copyright © Ruslan Panin

Carabus (Eurycarabus) famini atlantis Alluaud, 1923
Female adult
C Morocco, Haut Atlas, SW of Oukaïmeden h=2900m, 31.173644, -7.844442, 29.IV-11.V.2024, C.Reuter leg., L=21 mm
Copyright © Ruslan Panin

Carabus (Fulgenticarabus) flutschi koreus Deuve, 2006
Male adult
N Korea, Samjiyon, Ryang Gang Province
6-12.6.2006 leg. Wong
Copyright © Boleslav Brezina

Carabus (Gnathocarabus) kusnetzovi Semenov, 1903
Female adult
female, 21mm
Copyright © Boleslav Brezina

Carabus (Goniocarabus) sogdianus Semenov, 1898
Male adult
male, 21mm
Copyright © Boleslav Brezina

Carabus (Goniocarabus) sogdianus Semenov, 1898
Male adult
N Tajikistan, SW Pamir-Alay, Zarafshan Range, Fann Mts, Marguzor Lake circ., Padrut vill., 16.V.1997
L=21 mm
Copyright © Ruslan Panin

Carabus (Goniocarabus) grombczewskii grombczewskii Semenov, 1891
Male adult
NC Tadzhikistan, Pamir, N-slope Peter I.Mt Rng, h=2100m, Ganishou vill.,12.VII.2006, Pak O. leg.
L=19 mm
Copyright © Ruslan Panin

Carabus (Goniocarabus) perelloi Casale, 1979
Male adult
N-Afghanistan, Hindukush, Provinz Baghlan, N of Kabul, N Salang Pass, subalp., h = 2200 m, 35°27'.494N / 68°58.399'E, Mai 2011, leg. Christoph Reuter (TOPOTYPUS)
Copyright © Igor Solodovnikov

Carabus (Hemicarabus) tuberculosus Dejean, 1829
Male adult
Far East, Lazovsky Dist., shore of Japan Sea, 3 km NE Preobrazhenie Vill., Ezhovaya (Tasovaya) Bay 13.07.06 M. & L. Smirnov's leg.
male
L=20,0 mm
Copyright © Maksim Smirnov

Carabus (Hemicarabus) nitens Linne, 1758
Male+Female adult
Belarus,
Vitebsk reg., Ushachskiy distr., near Bolshie Dolcy vill. 9.V.1990, I.A.Solodovnikov leg.
female, 18mm.
Copyright © Ruslan Panin

Carabus (Hemicarabus) serratus Say, 1825
Male adult
E-USA,Virginia, Leesburg env., Ball's Bluff Battlefield Regional Park, 27.VIII-4.IX.2015,V. Čutka leg. L=17 mm
Copyright © Ruslan Panin

Carabus (Hemicarabus) serratus Say, 1825
Female adult
E-USA, Virginia, Leesburg env., Ball's Bluff Battlefield Regional Park, 27.VIII-4.IX.2015,V. Čutka leg. L=18 mm
Copyright © Ruslan Panin

Carabus (Homoeocarabus) maeander maeander s.str.
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Carabus (Homoeocarabus) maeander paludis Gehin, 1885
Female adult
Japan, Hokkaido; Tomakomai city, Benten, 17.V.2015, Takemoto leg., L=21 mm
Copyright © Ruslan Panin

Carabus (Hygrocarabus) nodulosus Creutzer, 1799
Male+Female adult
Austria, Styria Region (Steiermark), Leibnitz, 31.X.1999, I.Rapuzzi leg.<br /> female, 29 mm
Copyright © Ruslan Panin

Carabus (Hygrocarabus) variolosus Fabricius, 1787
Male adult
W-Ukraine, NW Podolian,<br /> Lviv reg., Zhydachiv distr.,near Vybranivka vill., 15.V-8.VI.2008, Panin R. leg.<br /> male, 27mm
Copyright © Ruslan Panin

Carabus (Hygrocarabus) variolosus Fabricius, 1787
Male+Female adult
W-Ukraine,<br /> Lviv Reg., Zhydachiv distr., near Vybranivka vill., Panin R. leg.<br /> female, 30mm
Copyright © Ruslan Panin

Carabus (Imaibius) baronii battalensis Heinz & Staven, 1998 Paratype
Male adult

Copyright © Alexander Anichtchenko

Carabus (Imaibius) dardiellus perobscurus Deuve, 1980
Male adult
N India.Kashmir, Loran, W part Pir Panchal range,h=2400-2500m, 33°49'"N, 74°20'E
L=25 mm
Copyright © Ruslan Panin

Carabus (Imaibius) dardiellus ? subpunctulus Roeschke, 1907
Male adult
N India, Kashmir, Pahalgam, Aru Valley h=2400m, VII.2007
L=25 mm
Copyright © Ruslan Panin

Carabus (Imaibius) wittmerorum Heinertz, 1978
Male adult
N. Pakistan: Frontier prov., Miandam, Swat vall., h=1800-2600 m, leg.S.Remond (TOPOTYPUS)
Copyright © Igor Solodovnikov

Carabus (Imaibius) barysomus hazarensis Deuve, 1982 (TOPOTYP)
Male adult
N-Pakistan: Mansehra prov., Hazara, Kaghan valley: Shocgran env., h = 2750 m L= 42 mm
Copyright © Igor Solodovnikov

Carabus (Imaibius) boysi boysi Tatum, 1851
Male adult
NW-India, Himachal Pradesh, Manikaran env., Pulga vill., 4.VII.2008. L=26 mm
Copyright © Ruslan Panin

Carabus (Imaibius) stoliczkanus brauni Heinz, 1993
Female adult
NE Pakistan, Azad Kashmir, Bagh Distr., Sudhan-Gali h~2300 m,15.VIII.1994, S Dacatra leg. L=37 mm
Copyright © Ruslan Panin

Carabus (Imaibius) caschmirensis fortetuberculatus Heinz, 1983 Paratype
Male adult

Copyright © Alexander Anichtchenko

Carabus (Imaibius) nouristani nouristani (Ledoux, 1977)
Female adult
NE-Afghanistan: Nuristan: Darah-I-Nur valley, 2900m 34°45'N 70°40'E, VIII.2012, C.Reuter leg., L=26 mm
Copyright © Ruslan Panin

Carabus (Imaibius) nouristani nouristani (Ledoux, 1977)
Male adult
NE-Afghanistan: Nuristan: Darah-I-Nur valley, 2900m 34°45'N 70°40'E, VIII.2012, C.Reuter leg., L=26 mm
Copyright © Ruslan Panin

Carabus (Iniopachus) pyrenaeus jeanneli Deuve, 1994
Male adult
1-2: Pirineos, Canfranc, Ibon de Iserias 2200 m 16-17.7.05<br /> 3: Pirineos, Monte Perdido, Collata Anisolo 2400m 25.8.05
Copyright © Alexander Anichtchenko

Carabus (Iniopachus) auriculatus ubinensis (Puissegur, 1955)
Female adult
Spain, Montes Cantabricos, Peña Ubiña, 2400m
5-6.VII.2009 Anichtchenko A. leg.
female, 19 mm
Copyright © Alexander Anichtchenko

Carabus (Iniopachus) pyrenaeus pyrenaeus (as cephalicus Csiki, 1927)
Male adult
SW France, Pyrenees-Atlantiques, Mt.Cirque d'Esturou h=1800m, 29.VIII.2014, Pascal Stefani leg. L=20 mm
Copyright © Ruslan Panin

Carabus (Ischnocarabus) tenuitarsis tenuitarsis Kraatz, 1877
Male adult
male, 24mm
Copyright © Boleslav Brezina

Carabus (Isiocarabus) fiduciarius fiduciarius (as fiduciarius tim) Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Isiocarabus) hunanicola liangshanicus (as dreuxi liangshanicus) HT PT Kleinfeld, 1998 Holotype Paratype
Male+Female adult
Carabus (Isiocarabus) hunanicola liangshanicus (as dreuxi liangshanicus) HT PT
Copyright © Frank Kleinfeld

Carabus (Isiocarabus) miaorum Lassalle & Prunier, 1993
Male adult
S-China, N-Guangxi, Mao'er Shan, h~1600 m, V.2009, L= 26 mm
Copyright © Ruslan Panin

Carabus (Isiocarabus) fiduciarius fiduciarius Thomson, 1856
Female adult
CE-China, Hubei, Xiaogan City, Shuangfeng Mountain, 26.VI.2016, L=30 mm
Copyright © Ruslan Panin

Carabus (Isiocarabus) miaorum Lassalle & Prunier, 1993
Male adult
China, NW-Guangxi, Mao'er Shan, h~1600 m 25°52'N 110°29'E, 28.V-9.VI.2009, C.Reuter leg.
Copyright © Ruslan Panin

Carabus (Lamprostus) calleyi pseudoprasinus Lapouge, 1912
Male+Female adult
Transcaucasia, Armenia,
Ashotsk distr. Tsoghamarg vill, .21-22.V.1997, P.V.Kazaryan leg
female, 24 mm
Copyright © Ruslan Panin

Carabus (Lamprostus) robustus ducalis Schaufuss, 1882
Male adult
NE-Turkey:Trabzon, Zigana Pass h=2000m,14-24.VI.2010, V. Slovák leg. <br /> L=32 mm
Copyright © Ruslan Panin

Carabus (Lamprostus) saulcyi saulcyi Piochard, 1875 (as chambouqensis)
Female adult
N-Lebanon, Fnaпdek, Djebel Qammoua, h=1100-1200m, 29.X.2012, C. Reuter leg.
L=27 mm
Copyright © Ruslan Panin

Carabus (Lamprostus) saulcyi saulcyi Piochard, 1875
Male adult
C Lebanon, Jbeil Distr., Ehmej vill., h=1100 m.V.2012, C. Reuter leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Lamprostus) syrus pseudopinguis Heinz & Staven, 2000
Male adult
NW-Jordan, Ajlun env., oak forests on dry limestone, h=897m, 32°21′33.1″N/ 035°43’47.4″E, 4.V.2010, Schnitter/Schellhorn leg.<br /> L=26 mm
Copyright © Ruslan Panin

Carabus (Lamprostus) torosus sinopensis Breuning & Ruspoli, 1970
Male adult
NC-Turkey, Sinop province, Erfelek env., h=200m, 10-27.VII.2014, C. Reuter leg. L=31 mm
Copyright © Ruslan Panin

Carabus (Lamprostus) syrus cheikensis Deuve, 1992
Male adult
N-Israel, Midlle Galilea, Nazareth env., Hoshaya viil., 13.XII.2012, Vas’ko, leg./det. Panin R. L=30мм
Copyright © Ruslan Panin

Carabus (Lamprostus) torosus nordmanni Chaudoir, 1848 (as enricoi )
Male adult
NE-Turkey:Ordu prov.,3 km N Gölköy h~1100 m, VII.2006, L=37 mm
Copyright © Ruslan Panin

Carabus (Lamprostus) syrus moukarzelae Deuve, 2002
Female adult
S-Lebanon: Marjaayoun, 18.XI.-11.XII.2017, C. Reuter leg., L=35 mm
Copyright © Ruslan Panin

Carabus (Lamprostus) syrus cheikensis Deuve, 1992
Female adult
SE Lebanon, Rashaya Distr., Tannoura, oak forest, h=700 m, 11-19.IV.2016, C. Reuter leg., L= 32 mm
Copyright © Ruslan Panin

Carabus (Leptocarabus) yokoae marcilhaci Deuve, 1992
Male adult
CHINA, Shanxi, Tschingling Mts., Taibeichan Nat. reserv.;
Copyright © Anton Menstschikov

Carabus (Leptocarabus) harmandi fujisan Ishikawa, 1972
Female adult
Japan, Yamanashi pref., Fuji-Rinds 1600m, Narusawa vill. 3.V.2005 Kurihara K. leg. 25 mm
Copyright © Alexander Anichtchenko

Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931
Male adult
NE-China:Liaoning prov., Fengcheng City, Miaoyang, 12-17.VI.2011<br /> L=26 mm
Copyright © Ruslan Panin

Carabus (Leptocarabus) procerulus Chaudoir, 1862
Male adult
Japan:S Honshu, Matsubara-ko, VIII.1997, S. Remond leg.
L=30 mm
Copyright © Ruslan Panin

Carabus (Leptocarabus) procerulus Chaudoir, 1862
Male adult
Japan, S Honshu, Mie Pref., Hakusan, VII.2001, S. Remond leg.<br /> L=33 mm
Copyright © Ruslan Panin

Carabus (Leptocarabus) arboreus pararboreus (Ishikawa, 1992)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Carabus (Leptocarabus) arboreus fujisanus Bates, 1883
Male adult
Japan, CE-Honshu, Yamanashi pref., Mt. Fujisan h~1600m, 23.IX.1982, Y. Imura leg. 23 mm
Copyright © Ruslan Panin

Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931
Male adult
NE-China:E-Liaoning: NP Kuandian Tianqiaogou , VI.2010, L=25 mm, det. Panin R.
Copyright © Ruslan Panin

Carabus (Leptocarabus) arboreus fujisanus Bates, 1883
Female adult
Japan, Honshu, Mt. Fuji h~1900m, Shizuoka pref., Fujinomiya city, 23.VI.2007, M. Matsunaga leg., L=25 mm
Copyright © Ruslan Panin

Carabus (Leptoplesius) merzbacheri kirgisiensis Mandl, 1967
Female adult
22mm
Copyright © Boleslav Brezina

Carabus (Limnocarabus) clathratus clathratus Linne, 1761
Female adult
W-Ukraine, Maloe Polesie, Lviv Region, near Kamenka-Bugzkaya, 28.III.2008, Panin R. leg. L=31 mm
Copyright © Ruslan Panin

Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
Female adult
SW Slovakia, Bratislava env., IV.2004, Krause Rene leg.
L=32 mm
Copyright © Ruslan Panin

Carabus (Limnocarabus) clathratus clathratus (as jansoni Kraatz, 1890)
Male adult
W-Ireland, County Mayo, Bellacorick bog, 9.VII.2013, J.Stypula leg. L=27 mm
Copyright © Ruslan Panin

Carabus (Limnocarabus) clathratus clathratus Linne, 1761
Female adult

Copyright © Kirill Makarov

Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
Male adult
W Ukraine, Transcarpathian Reg., 4 km NE Chop, forest Peresh, Panin R. leg. L=30 mm
Copyright © Ruslan Panin

Carabus (Lipaster) stjernvalli stjernvalli Mannerheim, 1830
Male+Female adult
Armenia,
Ashotsk, 22.VI.1990, P.V.Kazaryan leg.
female 34mm
Copyright © Ruslan Panin

Carabus (Lipaster) stjernvalli osellai Breuning & Ruspoli, 1970
Male+Female adult
NE-Turkey: Artvin:Hopa, Borçka env., h=750 m, 9.VI-7.VII.2011, M.Janalik leg.
female, 42 mm
Copyright © Ruslan Panin

Carabus (Lipaster) stjernvalli osellai Breuning & Ruspoli, 1970
Male adult
NE-Turkey: Artvin, Hopa / Çifteköprü, VII.1977, L=36 mm
Copyright © Ruslan Panin

Carabus (Lipaster) stjernvalli stjernvalli Mannerheim, 1830
Male adult
SC Georgia, Tsalka distr. 7 km NE of Paravani Lake, 41.540750, 43.879806, h=2100 m, 12-16.VII.2025, I.Uspenskyi leg., L=25 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) rugosus rugosus Fabricius, 1792
Male adult
N Morocco, El Biutz –Taghramt,20.I.2005, Manfred Egger leg.
L=29 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) morbillosus constantinus Kraatz, 1899
Male adult
NW Tunisia; Tebarka, E Sejenane,26.I.2004, Manfred Egger leg.
L=33 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) morbillosus macilentus Lapouge, 1899
Female adult
SE Spain, Murcia reg., P.N. de Calblanque, 1.V.2003, T.Gazurek leg.
L=34 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) rugosus celtibericus Germar, 1824
Female adult
SW-Portugal, Setúbal Distr., Cercal do Alentejo/ Tanganheira vill. 8.XII.2006, T.Gazurek leg.
L=31 m
Copyright © Ruslan Panin

Carabus (Macrothorax) morbillosus constantinus (as galloprovincialis Lapouge, 1910)
Female adult
SE-France, Var, near Le Muy, VIII.1980, Michon leg.
L=31 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) planatus Chaudoir, 1843
Male adult
S Italy, N Sicily, Nebrodi; 33 mm
Copyright © Aleš Sedláček

Carabus (Macrothorax) morbillosus lampedusae Born, 1925
Male adult
S Italy, Isl. Lampedusa, Baia dei Conigli, VI.2003, L=29 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) rugosus levantinus Lauffer, 1905
Male adult
SE-Spain, Murcia, Jumillla, Sierra del Carche, 6.VII.2019, F. Garcia leg., L=29 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) planatus Chaudoir, 1843
Female adult
S Italy, N Sicily, Nebrodi Mts, Portella di Fémmina Morta h=1500 m, 2.IX.1989, V.Aliquo leg. L=33 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) rugosus celtibericus Germar, 1824
Male adult
SW-Portugal, Serra do Cercal h=217 m, 37°47'15.6"N 8°42'02.0"W, road N390, 19.III.2011, D. Prunier leg., L=27 mm
Copyright © Ruslan Panin

Carabus (Meganebrius) arunensis Heinertz, 1980
Female adult

Copyright © Boleslav Brezina

Carabus (Meganebrius) kadoudali (Morvan, 1981)
Male+Female adult
C-Nepal, SW Dhaulagiri, W Jaljala h=3150 m, 28°30'05"N, 83°11'48"E, 23.IX.2012, J. Schmidt leg.
female, 20 mm
Copyright © Ruslan Panin

Carabus (Meganebrius) pseudoharmandi Mandl, 1965
Female adult
E-Nepal, Kosi zone, Khandbari,(1.5km SW of Chichila vill., 27*45’843’’N/ 87*22’473’E, h=1940m), damaged primary forest, 13 .V.2010, B.Brezina & M.Häckel leg.
L=23 mm

Copyright © Ruslan Panin

Carabus (Meganebrius) arunensis Heinertz, 1980
Male adult
E-Nepal, Kosi zone, Khandbari, 1.5km SW of Chichila vill., 27*47’175’’N/ 87*23’622’E, h=1880m, damaged primary forest, 11.V.2010, B.Brezina & M.Häckel leg.
L=27 mm

Copyright © Ruslan Panin

Carabus (Meganebrius) quinlani smetanai Häckel, 2011 Paratype
Male+Female adult
C-Nepal:Dhaulagiri Massif:Baglung Distr., (1.5km W of Okhale vill., 28.32’152’’N/ 83.48’673’E, h=2600m) damaged primary forest, 19.V.2010, B.Brezina & M.Häckel leg.
male, 21mm
female, 23 mm
Copyright © Ruslan Panin

Carabus (Meganebrius) quinlani quinlani Mandl, 1965
Male adult
C-Nepal, Gandaki, Manang Distr., Timang env., h~2500-2600m, old open forest, grassland, 3-5.V.2007, Fouqué R. leg. L=19 mm
Copyright © Ruslan Panin

Carabus (Meganebrius) tuberculipennis ganjalaensis Deuve, 2000
Male adult
Nepal, Helambu, GanjaLa Pass S-slope, Dhukpu Kharka h=4100m, 28°03'50"N, 85°34'21"E, 12.VIII.2010, S.Tamang leg., L=21 mm
Copyright © Ruslan Panin

Carabus (Pagocarabus) erwini erwini s.str.
Male adult

Copyright © Boleslav Brezina

Carabus (Megodontus) violaceus andrzejuscii (as wolffii Dejean, 1826)
Male+Female adult
SW-Romania: Banat, Moldova Noua, Ravenska viil., 21.V.-5.VI.2010, Krause Rene leg. female, 32 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus andrzejuscii (as mehelyi Ganglbauer, 1896 loc. typ.)
Male+Female adult
N-Romania, Rodna Mts., Pasul Setref - Sacel 10km E, h=810 m, 8-19.VI.2009, Krause Rene leg. female, 31mm
Copyright © Ruslan Panin

Carabus (Megodontus) exaratus Quensel, 1806
Male adult
Russia, C.Caucasus, N.Ossetia,
valley of River Khazniaush,
h=1400 m, 10.VI-31.VII.1993
A. Zamotajlov leg.
Copyright © Safronov

Carabus (Megodontus) violaceus violaceus Linne, 1758
Male adult
S.Bohemia, Hluboka nad Vltavou, h=380 m, 6.VIII.2004, S. Budka leg.
L=28 mm
Copyright © Ruslan Panin

Carabus (Megodontus) vietinghoffi bowringi (as lazoensis Obydov, 1999)
Male+Female adult
Russia, Far East, Primorsky reg., S. Sichote-Alin Mts.R., Kavalerovo vill. env.,
5-25.VII.1992, S. Kunitzyn leg.
male - 28 mm, female - 30 mm.
Copyright © Alexandr Safronov

Carabus (Megodontus) vietinghoffi bowringi Chaudoir, 1863
Male+Female adult
Russia, Far East, Primorsky reg., Khasansky distr.,
Zanadvorovka vill., Gusevsky pit.
M - 19-21.VI.2007, body lenght 29 mm.
F - 15.VII.2008, body lenght 30 mm.
S. Veriga leg.
Copyright © Alexandr Safronov

Carabus (Megodontus) kolbei hidakamontanus Ishikawa, 1966
Male adult
N-Japan, C-Hokkaido, Hidaka Mt.Rng, Mt.Apoidak near Samani town, 20.VIII.1982,T. Akaishi leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823
Male adult
W-Ukraine, Podolian Tovtry, Ternopol reg., Pidvolochysk distr., between Chagari-Zbarazhki & Shevchenkove vill., steppe sites, 6.V-7.VI.2012, Panin R. leg. L=25 mm
Copyright © Ruslan Panin

Carabus (Megodontus) persianus javahardehensis Deuve, 2000
Male adult
N-Iran, Mazandaran Prov., Ramsar Co., Javahar Deh env., 16-25.VI.2017, Václav Čutka leg. L=28 mm
Copyright © Ruslan Panin

Carabus (Megodontus) kolbei hidakamontanus Ishikawa, 1966
Female adult
E-Hokkaido: Samani distr., Horomankyo env.
Copyright © Igor Solodovnikov

Carabus (Mesocarabus) problematicus planiusculus Haury in Gehin, 1885
Male adult
NE Spain, Catalonia, Sant Jaume de Frontanyà, h=1000 m, V-VI.2010, Pascal Stefani leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) latus latus Dejean, 1826
Female adult
CW Spain, Cáceres, San Martín de Trevejo, 24.VI.2011, A. Diez leg./det.Panin R.
L=27 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) cantabricus cantabricus Chevrolat, 1840 (loc. typ.)
Male adult
NW-Spain, Pontevedra prov., A Cañiza viil., Alto de Fontefría h=800m, 31.XII.2007, T.Gazurek leg./det.Panin R.<br /> L=27 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) macrocephalus Dejean, 1826
Male adult
NW-Spain,Cantabrian Mts, Picos de Europa Mt Rng, Puerto de Panderruedas h=1460 m, 25.IX.2004,T.Gazurek leg.
L=28 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus harcyniae (as procedens Csiki, 1927)
Male adult
W-Ireland, County Mayo, Clare Isl., 17-28.VI.2002, S. McCormack leg.<br /> L=25 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) (? problematicus trans lusitanicus) planiusculus (as xaxarsi Born, 1917)
Male adult
NE Spain, Catalonia, Girona Prov., San Hilario Sacalm env., 20.II.1988, A. Mollard leg.<br /> L=25 m
Copyright © Ruslan Panin

Carabus (Mesocarabus) breuningi Csiki, 1927
Female adult
NW-Spain, Prov. Zamora, San Martín de Castañeda, 5.VII.1984, M.Toribio leg. L=28 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus lusitanicus (as vieirae Oliveira, 1875)
Male adult
CW-Portugal: Leiria distr., near Marinha Grande, 39°45'57.8"N 8°58'10.3"W, h-79 m, 19.IX.2016, D. Prunir leg., L=24 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) riffensis Fairmaire, 1875
Endophallus
N-Morocco, Riff Mts h=1650 m, 26 km E of Bab Berred
Copyright © Ruslan Panin

Carabus (Mesocarabus) riffensis Fairmaire, 1875
Male adult
N-Morocco, Riff Mts h=1650 m, 26 km E of Bab Berred, 34.961333, -4.669417, Cedar forest, C. Reuter leg. L=27 mm
Copyright © Ruslan Panin

Carabus (Microplectes) riedeli Menetries, 1832
Female adult
female, 19mm
Copyright © Boleslav Brezina

Carabus (Microplectes) riedeli Menetries, 1832
Male adult
N-Ossetia,
Khazniaush Pass, h=2500-2700m, 9.VI-31.VII.1994, Zamotajlov A.leg.
male, 17mm
Copyright © Ruslan Panin

Carabus (Microplectes) convallium convallium Starck, 1889
Male adult
male, 19mm
Copyright © Boleslav Brezina

Carabus (Microplectes) convallium convallium Starck, 1889
Male+Female adult
female:NW-Caucasus, Russia,
Krasnodar reg., Apsheron Distr., Lagonaki Mt., near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominyh leg.
male:Karachaevo-Cherkessia,the right source of the river Beskess, h=2100m, 10.VIII.1997, I.A. Solodovnikov leg.
female, 21mm/male, 19mm
Copyright © Ruslan Panin

Carabus (Microplectes) convallium argonautarum Semenov, 1898
Male adult
male, 18mm
Copyright © Boleslav Brezina

Carabus (Mimocarabus) maurus maurus Adams, 1817
Male+Female adult
Armenia, Ashotsk distr., Tsoghamarg vill., 24-27.V.1999 P.V.Kazaryan leg. female, 20mm/male,18mm
Copyright © Ruslan Panin

Carabus (Mimocarabus) pumilio Kuster, 1846
Male adult
S-Armenia, Vayots Dzor Region, E Ughedzor vill., h=2630 m, 9.VI.2014, L=16 mm
Copyright © Ruslan Panin

Carabus (Mimocarabus) maurus maurus (Adams, 1817)
Male adult
E Georgia, Kakheti, near Patardzeuli vill., h=750-850 m, 10.IV.2015, E. Rutjan leg., L=22 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei (as ormayi Reitter, 1896) loc.typ.
Male adult
W. UKRAINE,
Transcarpathian reg., near Beregovo, 27.IV-22.V..2011,, Panin R. & Chyzhevsky O. leg.
L=28 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens trans scheidleri
Male adult
NC Croatia, Ivanščica Mts , Ivanec,4-18.VI.2011, Krause Rene leg.<br /> L=31 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878 (as viridipennis Kraatz, 1886, nec Lewis,18
Male+Female adult
Russia, Far East,Primorsky reg., S. Sikhote-Alin Mts. R., Kavalerovo vill. env.,
5-25.VII.1992, С. Kunitzyn leg.
male - 25 mm, female - 25 mm.
Copyright © Alexandr Safronov

Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as pseudojucundus Retezár, 1974)
Male adult
NW-Hungary::Komarom distr., Tardos, Gerecse Mts., 20.-27.V.2011, Krause Rene leg.
L=26 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi rothi (as equistriatus Kraatz, 1878)
Female adult
C Romania, Sibiu reg., left bank Olt River, Avrig env., 12.VI.2013, V. Slovak leg.
L=30 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
Female adult
W Ukraine, Transcarpathian lowland, near Chop, 11.VI-19.VII.2019, Panin R leg. L=29 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Female adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 28.V-9.VI.2013, Panin R. leg. L= 28 mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) scheidleri Panzer, 1799
Endophallus
E-Austria, Wienerwald, Hochstraß am Lengbachl, h=575 m,
Copyright © Ruslan Panin

Carabus (Morphocarabus) scheidleri Panzer, 1799
Male adult
E-Austria, Wienerwald, Hochstraß am Lengbachl, h=575 m, 25.VI-2.VII.2024, M. Fuchs leg. L= 30 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) henningi oviformis Beheim & Breuning, 1943
Male adult
Russia, Polar Ural, Yamalo-Nenets reg.,Harp station envir., 15.VII.1995, G. Chekura leg. L=19 mm
Copyright © Ruslan Panin

Carabus (Morvanicarabus) scheibei scheibei Eidam, 1937
Female adult
NE-Afghanistan, Hindu Kush, Nuristan Prov., near Parun, 35°19'N 70°52'E h=2900 m, Pine forest, VI.2022, local collector/det. Kleinfeld, L= 24 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) daphnis daphnis Kurnakov, 1962
Male adult
Georgia, Mingrelia, Lecheha vallis, Tehuri superior
Caucasus centr., 1200m, 27.6-7.7.1989 Retezar I. leg.
male, 28 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) reitteri laevisternis Starck, 1890
Male adult
Abkhasia,W.slope of Gagrinsky Mt., Krepostnaya River source, h=1325m., A.S.Zamotajlov leg. 27.VII.2000
male, 29 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) edithae exedithae Gottwald, 1985
Male adult
NС-Caucasus, Karachaevo-Cherkessia, Abishir-Akhuba Mt.Range h=2250-2600 m, right bank Pkhiya river, 10-13.VI.2013, S. Pavluchuk leg.
L=25 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) satyrus satyrus Kurnakov, 1962
Male adult
Abkhazia, Gulripsh Distr., NW slopes of Gurzul Mt. Range near Merkheul Village, 170-230 m, left tributary of Machara River, hornbeam, alder, rhododendron (Rhododendron ponticum) forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus besleticus Kurnakov, 1972
Male+Female adult
Abkhazia, Sukhum Distr., vicinities of Village Kaman, N slope of Mt. Birtskha, 160 m, beech, linden, box, elder forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Neoplectes) komarowi komarowi Reitter, 1882
Female adult
NW Georgia, Upper Svaneti, Mestia distr., Lakhamula env. h~1250 m, 19.VI.2014, V.Sima leg. L=33 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) starckianus adelaidae Starck, 1889
Male adult
Russia, Krasnodar area, Adler, Ader distr., right bank vall. Riv. Msymta, near vill. Chvesipse (Fagus+Quercus+Castanea forest) [Sochi national park], h = 720 m, 03.V-08.VII.2016, I.A. Solodovnikov, S.V.Solodovnikova, V.M.Kotsur leg.
Copyright © Igor Solodovnikov

Carabus (Neoplectes) protensus protensoides Novotny & Vorisek, 1988
Endophallus
W Georgia, Martvili distr., W slope of Askhi Plateau h=1200-1800 m,
Copyright © Ruslan Panin

Carabus (Neoplectes) protensus protensoides Novotny & Vorisek, 1988
Male adult
W Georgia, Martvili distr., W slope of Askhi Plateau h=1200-1800 m, 18-19.VI.2005, Shumov leg., L=33 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) ibericus montreuilianus Deuve, 2014
Endophallus
SC Georgia, Lesser Caucasus, Samsari Range, Mt Egoisar h=2100-2200m, 41.651528, 43.770083
Copyright © Ruslan Panin

Carabus (Eocechenus) kaznakovi nangqenicus Deuve, 1995 Paratype
Male adult

Copyright © Boleslav Brezina

Carabus (Nesaeocarabus) abbreviatus Brulle, 1835
Male adult
Canarias Isle, NC Tenerife; 22mm
Copyright © Aleš Sedláček

Carabus (Nesaeocarabus) faustus cabrerai Enderlein, 1929
Male adult
Spain, Canarias Isl.,Teneriffe: NW-part:Teno Massif, Erjos env., h=1000m, 18-21.III.2013, V.Slovak leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Ohomopterus) albrechti albrechti Morawitz, 1862
Female adult
Japan, S-Hokkaido:Oshima Peninsula,Pass btw Otoshibe - Assabu, h~250-300 m, 10-20.VII.2010, 42:01’53’Т; 140:22’09’E, I.Rapuzzi & L.Caldon leg.
Copyright © Ruslan Panin

Carabus (Ohomopterus) yamato kitai (Imura, 2004)
Male adult
Japan, S Honshu, Mie Pref., Hakusan, VII.2001, S. Remond leg.<br /> L=21 mm
Copyright © Ruslan Panin

Carabus (Ohomopterus) dehaanii dehaanii Chaudoir, 1848
Female adult
Japan, S Honshu, Hyogo Pref., Nishiwaki city, 20.VI.2010, Shinichi S. leg., L=37 mm
Copyright © Ruslan Panin

Carabus (Ohomopterus) dehaanii dehaanii Chaudoir, 1848
Male adult
Japan, S Honshu, Mie Pref., Inabe city, 21.II.2021, Shinichi S. leg., L=33 mm
Copyright © Ruslan Panin

Carabus (Ohomopterus) japonicus japonicus Motschulsky, 1857
Male adult
MT. Narabara-san, Ryuokakami Tamagawa town, Imabari city, Ehime pref. Japan 20.6.2015 Y. Amano (cAA). det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Carabus (Ohomopterus) japonicus japonicus Motschulsky, 1857
Female adult
Fukuoka pref., Omuta city, Hagio, 16.1.2012, S. Nishimoto leg. (cAA)
Copyright © Alexander Anichtchenko

Carabus (Ohomopterus) lewisianus awakazusanus Ishikawa, 1981
Male adult
Mt. Iyogatake 280m, Tomiyama town, Awa Co., Chiba pref, 4.2.1996 K. Watanabe leg. (cAA). det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Carabus (Ohomopterus) albrechti Morawitz, 1862
Male adult
Horoman, Samani town, Samani Co., Hokkaido, 16.6.2015 T. Nakamura leg. (cAA). det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Carabus (Ohomopterus) insulicola nishikawai (Ishikawa, 1966)
Male adult
Yamada, Isumi city, Chiba pref. Japan 20.5.2007 K. Osozawa leg. (cAA). 28mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Carabus (Ohomopterus) insulicola insulicola Chaudoir, 1869
Male adult
Japan, Kiyose, Tokyo 20.1.1961 (cAA). 29 mm
Copyright © Alexander Anichtchenko

Carabus (Ophiocarabus) regeli obscurior Semenov, 1888
Male+Female adult
Kyrgyzstan, NE Tian-Shan, E. Terskey Alatau Mts. R.,
Kok-kia River, h=2800 м.
7-8.VI.1997 М. Danilevsky leg.
Body lenght 15-18 mm.
Copyright © Alexandr Safronov

Carabus (Ophiocarabus) ballionis Kraatz, 1879
Male adult
NW-China, Xinjiang prov., 40 km N Narat, h=1500, 12-12.VI.2004, Benes & Kaiser leg.
L=22 mm
Copyright © Ruslan Panin

Carabus (Ophiocarabus) successor koeksuensis Kabak, 2013 Paratype
Male adult
China, Xinjiang, Koeksu Bas., Tshon-Aidak, SSE Tekes, 42* 32' 47'' N / 82* 05' 56'' E -- 42* 32' 51'' N / 82* 05' 36'' E, 3330-3550 m, 11.07.2012, Kabak I.I. leg.
Copyright © Ilya Kabak

Carabus (Ophiocarabus) successor successor Reitter, 1896
Male adult
SE-Kazakhstan, Narynkol distr., 4 km SW Bayankol, h~2400-2500 m, 2-14.VI.2007, Shumov leg. L=17.5 mm
Copyright © Ruslan Panin

Carabus (Ophiocarabus) successor successor Reitter, 1896
Female adult
SE-Kazakhstan, Narynkol distr., 4 km SW Bayankol, h~2400-2500 m, 2-14.VI.2007, Shumov leg. L=17 mm
Copyright © Ruslan Panin

Carabus (Pachystus) hortensis neumeyeri Schaum, 1856
Female adult
Italy, Basilicata, Serra Calvello (PZ)
1400 m, 7.2003 F. Angelini leg.
female, 25 mm
Copyright © Alexander Anichtchenko

Carabus (Oreocarabus) carinthiacus Sturm, 1815
Male adult
NE-Italy, Bolzano, Brenner, Zirog h=2000 m, 26.VII.1992, Manfred Egger leg.
L=20 mm
Copyright © Ruslan Panin

Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847
Male adult
WC Spain, Cáceres, Hervás, h=1000m, V.2007, Pascal Stefani leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Oreocarabus) latro minshanensis Deuve, 1987
Male adult
China, N-Sichuan, Jiuzhaigou env., 13.VI.2001,E. Kučera leg. L=23 mm
Copyright © Ruslan Panin

Carabus (Oreocarabus) luetgensi pseudosteuarti Lapouge, 1924
Female adult
Portugal, Guarda, Sierra de Estrela h=800 m, 26.VI-4.VIII.2015, L=26 mm
Copyright © Ruslan Panin

Carabus (Oreocarabus) luetgensi pseudosteuarti Lapouge, 1924
Male adult
Portugal, Guarda, Sierra de Estrela h=800 m, 26.VI-4.VIII.2015, L=24 mm
Copyright © Ruslan Panin

Carabus (Pachystus) hortensis preslii Dejean, 1830
Male adult
S-Greece:, N Peloponnes, Erymanthos Reg., Kalentzi, 26.VI.2007, Manfred Egger leg. L=26 mm
Copyright © Ruslan Panin

Carabus (Pachystus) hortensis hortensis Linne, 1758
Female adult
Natural hybrid Carabus hortensis X Carabus granulatus. Belarus, Minsk reg., 8,7 km of Borisov, near vill. Lyubatovstchina, pine forest (Pinetum pleuroziosum), h = 213 m, 54.31934° N, 28.511486° E, 20-27.09.2018, leg. Klychenok Snezhana
Copyright © Igor Solodovnikov

Carabus (Hypsocarabus) mikhaili mikhaili Deuve & Mourzine, 1997
Female adult
L=23 mm
Copyright © Ruslan Panin

Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847
Female adult
Spain, Sierra de Guadarrama, Puerto de la Quesera, 1720m, 41.214640, -3.430896, Anichtchenko A. leg. 10.X.2025 (cAA).
Copyright © Alexander Anichtchenko

Carabus (Orinocarabus) sylvestris sylvestris Panzer, 1793 (as slabai Niedl, 1968)
Female adult
Czech, WS Bohemia,
Plzen Reg., Domazlice Distr., Babylon, h=500m, 30.VIII.2008, S.I. Budka leg.
L=24mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) alpestris hoppei Germar, 1824
Male adult
W Austria, Tirol, Innsbruck, Hafelekar Mts h=2200m, Manfred Egger leg.
L=19 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) fairmairei fairmairei Thomson, 1875
Male adult
N Italy, Alpi Graie; 20 mm
Copyright © Aleš Sedláček

Carabus (Orinocarabus) bertolinii Kraatz, 1878
Male adult
NE-Italy, Dolomites-Alps, Trentino prov., Rolle Pass env., Malga Fosse h=2000m, 8-28.VI.2013, Manfred Egger leg.
L=19 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) alpestris dolomitanus Mandl, 1956
Male adult
NE-Italy, South Tyrol, Dolomites-Alps, 500 m left of Würzjoch, Gömaalm - at the foot Mt.Peitlerkofel h=2100m, 28.VII-17.VIII.2013, Manfred Egger leg.
l=18 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) pedemontanus raynaudianus Deuve & Simard, 1977
Male adult
SE France, Alpes-Maritimes, Isola env., Colle Santa-Anna h=2309 m, 44*13',127" N, 07*06',242"E, 31-7.VIII..2012. L=21 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) concolor lepontinus Born, 1908
Male adult
NW-Italy, Piemonte (VB), Lepontine Alps, Mt. Zeda , 6.VII.1998, L Diotti leg., L=20 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) concolor concolor Fabricius, 1792
Female adult
NW-Italy, Breuil-Cervinia h~2500 m, 2.IX.1971, Georgeon leg. L=21 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) latreilleanus Csiki, 1927
Female adult
NW-Italy, Piemonte, Biella, Biellese Alps Rng, Mt.Camino, 25.VI.1997, V.Monzini leg. L=18 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) bertolinii Kraatz, 1878
Male adult
NE-Italy, Trentino prov. Dolomites Alps, Plateau Pale di San Martino h=2500 m, 17-19.VIII.2019, M. Egger leg.
Copyright © Ruslan Panin

Carabus (Oxycarabus) saphyrinus pseudosaphyrinus Schweiger, 1964 (as rabaronianus)
Male adult
Anatolia, Kastamonu, Yaraligoz verso Catalzeitin,
VI.2001 Sola Claudio leg.
male, 28 mm
Copyright © Alexander Anichtchenko

Carabus (Oxycarabus) saphyrinus pirithous Reitter, 1896
Male adult
NW-Turkey,Istanbul (Asian part),Polonezköy env., 2.VI.2009. S. Dacatra leg. L=27 mm
Copyright © Ruslan Panin

Carabus (Pachycarabus) koenigi Ganglbauer, 1886
Male adult
NW Caucasus, S Russia, Adygea Rep., Lagonaki plateau, Site Murzikal h=1800-2000m, 24.V-18.IX.2010, D.Fominykh leg.
L= 22 mm
Copyright © Ruslan Panin

Carabus (Pachycarabus) roseri soganliensis Heinz & Korge, 1964
Male+Female adult
Turkey, Rize prov., Ikizdere distr., Rize Daglari,
47 km to SE from lyidere, Sivrikaya Koyu, h=2025 m.
30.VI-5.VII.2011, A. Vlasenko leg.
male - 23 mm, female - 24 mm.
Copyright © Alexandr Safronov

Carabus (Pachycarabus) roseri soganliensis Heinz & Korge, 1964
Female adult
NE-Turkey: Rize Prov.Ovit Pass (40.37;40.46) h-2700m,15.VII.2008, I.Rapuzzi & Caldon leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Pachycarabus) staehlini Adams, 1817
Male adult
S Russia, E-Caucasus, Dagestan, Nukatl Mt Rng h=2600m, 25.VII.1997, V. Savizky leg. L=21 mm
Copyright © Ruslan Panin

Carabus (Pachycarabus) koenigi Ganglbauer, 1886
Male adult

Copyright © Kirill Makarov

Carabus (Pachycarabus) roseri roseri Faldermann, 1835
Female adult
Georgia, Lesser Caucasus, Imereti prov., N slope Meskheti Mt Rng, Zekari Pass, h~1950 m, 24.VII.2021, E.Rujan leg., L=22 mm
Copyright © Ruslan Panin

Carabus (Pachycarabus) swaneticus swaneticus Reitter, 1883
Male adult
Georgia, Caucasus, Svaneti Range E part, Latpari Pass h=2700-2800 m, 23.VI.2024, I.Uspenskyi leg./ det. R. Panin, L=18 mm
Copyright © Ruslan Panin

Carabus (Pachycarabus) swaneticus swaneticus Reitter, 1883
Female adult
Georgia, Caucasus, Svaneti Range E part, Latpari Pass h=2700-2800 m, 23.VI.2024, I.Uspenskyi leg., L=19 mm
Copyright © Ruslan Panin

Carabus (Pachycarabus) swaneticus swaneticus Reitter, 1883
Male adult
Georgia , Caucasus, E part Egrisi Mt Range, Mt. Tsekuri N slope h=2370 m, 18-20.VI..2025, I.Uspenskyi leg. L=19 mm
Copyright © Ruslan Panin

Carabus (Pachycarabus) staehlini rionicus Belousov, 1992
Male adult
Georgia, Caucasus, S of Sasashi, Lechkhumi Range N sope h=2430m, alpine meadows 242.759167, 42.973889, 20-21.VII.2025, I.Uspenskyi leg./det. R,Panin, L=20 mm
Copyright © Ruslan Panin

Carabus (Pachystus) tamsi tamsi Menetries, 1832
Male+Female adult
Transcaucasia, Armenia, <br /> Ashotsk, 18.VI.2001, P.V.Kazaryan leg.<br /> female, 26mm//male, 24mm
Copyright © Ruslan Panin

Carabus (Pachystus) hungaricus hungaricus Fabricius, 1792
Female adult
C Hungary, Pest reg., Örkény, 9-10.V.1997, S.Budka leg.
Copyright © Ruslan Panin

Carabus (Pachystus) hungaricus scythus Motschulsky, 1847
Female adult
S Ukraine, Zaporizhzhya reg., Khortytsa island, 16-20.VII.2011, M. Mulenko leg., det. Panin R.
L=28 mm
Copyright © Ruslan Panin

Carabus (Pachystus) hungaricus hungaricus Fabricius, 1792
Male adult
Czech, S Moravia, Pouzdrany, 8,VIII.2005, V. Slovak lgt.
L=27 mm
Copyright © Ruslan Panin

Carabus (Pachystus) cribratus staneki Sterba, 1931
Male adult
NC Turkey, Küre Mts., 60km N Kastamonu; 25mm
Copyright © Aleš Sedláček

Carabus (Pachystus) cribratus staneki Sterba, 1931
Female adult
NC Turkey, Küre Mts., 60km N Kastamonu; 27mm
Copyright © Aleš Sedláček

Carabus (Pachystus) cavernosus variolatus Costa, 1839
Male adult
C-Italy:Abruzzo reg., Gran Sasso d'Italia Massif, h~1600 m, VIII.1997, Seichert leg. L=26 mm
Copyright © Ruslan Panin

Carabus (Pachystus) cavernosus rozneri Retezár & Gy. Szél, 2013 Paratype
Male adult
Romania, Apuseni : Trascău Mts. L=27 mm
Copyright © Ruslan Panin

Carabus (Pachystus) glabratus gibbosus Heyden, 1866
Male adult
N Croatia, Ivanščica Mts, Ivanec, 4-18.VI.2011, Krause Rene leg., L=32 mm
Copyright © Ruslan Panin

Carabus (Pachystus) cavernosus variolatus Costa, 1839
Female adult
C-Italy: Abruzzo reg., Gran Sasso massif, Campo Imperatore, h=1900 m, VIII.2021, G.Proscia leg., L=27 mm
Copyright © Ruslan Panin

Carabus (Pagocarabus) crassesculptus diruptus Morawitz, 1886
Male adult
China, Gansu, Umg. Xiahe 3000-3200m
24-26.6.1993 Heinz leg.
20 mm
Copyright © Alexander Anichtchenko

Carabus (Pagocarabus) crassethoracis Kleinfeld, 1999 Holotype
Male adult

Copyright © Frank Kleinfeld

Carabus (Pagocarabus) crassesculptus eleganticolor Deuve, 1997
Female adult
S Gansu, Min Shan, 5-20 km S Cagan, 2300-3200m 7-20.7.2019 V. Patrikeev leg. (cAA). det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Carabus (Pseudotribax) ferghanicus Breuning, 1933
Male adult
male, 29mm
Copyright © Boleslav Brezina

Carabus (Pantophyrtus) turcomanorum turcomanorum Thieme, 1881
Male+Female adult
Kyrgyzstan, Kyrgyzskyi Mts. R., Uzun Bulak Gorge, h=1800 ms.
01-09.VI.2002 loc. coll.
male - 28 mm, female - 33 mm.
Copyright © Alexandr Safronov

Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993
Male adult
Uzbekistan, Chatkal Mt. rng., Irtash, 41°10'25.7"N 70°16'20.4"E, 5-7.8.2021, 2400m, A. Anichtchenko leg. (cAA). 27mm
Copyright © Alexander Anichtchenko

Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993
Female adult
Uzbekistan, Chatkal Mt. rng., Irtash, 41°10'25.7"N 70°16'20.4"E, 5-7.8.2021, 2400m, A. Anichtchenko leg. (cAA). 30 mm
Copyright © Alexander Anichtchenko

Carabus (Piocarabus) titanus Breuning, 1933
Male adult
C-China Sichuan prov., Nanping, h=1500m, VI.2000, S. Remond leg.<br /> L=34 mm
Copyright © Ruslan Panin

Carabus (Piocarabus) nanwutai Kleinfeld, Korell & Wrase, 1996
Male adult
China, W-Shaanxi, Taibai Shan Mts, btw. Houzhenzi/Ba Xian Tai, nr.Nan Tian Men h=3200m, 33°55'17"N/107°46'53"E, 25-30.VII.2011, Li Wuyang leg.<br /> L=19 mm
Copyright © Ruslan Panin

Carabus (Piocarabus) sui Imura & Zhou, 1998
Female adult
CE-China, Henan, Mt Wangwu, V.2007, local collector. L= 33 mm
Copyright © Ruslan Panin

Carabus (Piocarabus) vladimirskyi Dejean & Boisduval, 1830
Male adult
Buryatia, Kyahta distr. [Khyagh], 5 km SE Chikoi vill., 16.8.1998, Anichtchenko A. leg.
Copyright © Alexander Anichtchenko

Carabus (Piocarabus) titanus Breuning, 1933
Female adult
China Sichuan prov., Nanping, h=1500m, VI.2000, S. Remond leg.
L=38 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Female adult
W Ukraine, NE Carpathians, Transcarpathian Reg., Rakhov Distr., Chornogora Mts Range h=2030m, alpin zone, 13-24.VIII.2010, Panin R. leg., female, 22 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Female adult
Ukrainian Carpathians, Transcarpathian reg., Rakhov Distr., Chornogora range, h=2010m, 29.VI-28.VII.2007, Panin R. leg., female, 23 mm., photo/det.Panin R
Copyright © Ruslan Panin

Carabus (Platycarabus) irregularis montandoni Buysson, 1882
Male adult
W-Ukraine, NW Podolian,
Lviv Reg., Zhydachiv distr., near Vybranivka, under the bark of oak tree, in the forest,12.X.2007 Panin R. leg
L= 22 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) irregularis ramanus Sokolar, 1909 (as bucephalus Kraatz, 1879)
Male+Female adult
W Slovenia, Goriška region Trnovo, h~1000 m, IX.2002, I.Rapuzzi leg.
female, 29 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) depressus lucens Schaum, 1856
Female adult
SE-France: Alpi Marittime, Monte Viso, 14.VII.2010
L=23 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) fabricii fabricii (as feichtauensis Müller & Sach, 1999)
Male adult
NC Austria, E-Alps, Windischgarsten, Mt Hoher Nock, VII.2004, Manfred Egger leg.
L=18 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) fabricii ucrainicus Lazorko, 1951
Male adult
Ukrainian Carpathians, Ivano-Frankivsk reg., Yaremcha distr., Gorgany Mts., h=1590 m, 22.VII-3.VIII.08, Panin R. leg., male, 23mm
Copyright © Ruslan Panin

Carabus (Platycarabus) fabricii ucrainicus Lazorko, 1951
Male adult
Ukrainian Carpathians, Ivano-Frankivsk reg., Gorgany Mts., h=1590 m, 17.VI-26.VII.2011, Panin R. leg.<br /> male, 23mm
Copyright © Ruslan Panin

Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Female adult
N- Romania, E-Carpathians, Rodna Mts, h=2000m, 7-8.VI.2012, Cartuz Z.leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Female adult
W Ukraine, NE Carpathians, Transcarpathian Region, Chornogora Mt Range, h=2000 m, alpin zone, 28.VII-11.IX.2014, Panin R. leg. L=22 mm
Copyright © Ruslan Panin

Carabus (Procerus) scabrosus tauricus Bonelli, 1811
Female adult

Copyright © Safronov

Carabus (Procerus) gigas gigas Creutzer, 1799
Male+Female adult
N Croatia, Ivanščica Mts , Ivanec, 4-18.VI.2011, Krause Rene leg.
female, 54 mm
Copyright © Ruslan Panin

Carabus (Procerus) scabrosus caucasicus Adams, 1817
Male adult
N Caucasus,
Stavropol Krai, Pyatigorsk, Mt.Mashuk, VI.2007 Tikhonov Yu. leg.
44mm
Copyright © Ruslan Panin

Carabus (Procerus) gigas duponcheli Dejean, 1831 (as kusdasi Mandl, 1966)
Male adult
S-Greece: Peloponnes, Mt.Chelmos, 20.V.2007.
L=46 mm
Copyright © Ruslan Panin

Carabus (Procrustes) chevrolati chevrolati Cristoforis & Jan, 1837
Male+Female adult
female, 31mm: Ashotsk,Vardaghbyur vill., 30.V.1997
male, 30 mm: Amasia, Chaldash vill.,8-11.VIII.1997
P.V. Kazaryan leg.
Copyright © Ruslan Panin

Carabus (Procrustes) chevrolati thirki Chaudoir, 1857
Male+Female adult
Turkey, Rize prov., Ikizdere distr., Rize Daglari,
47 km to SE from lyidere, Sivrikaya Koyu, h=2025 m.
30.VI-5.VII.2011, A. Vlasenko leg.
male - 28 mm, female - 31 mm.
Copyright © Alexandr Safronov

Carabus (Procrustes) anatolicus anatolicus Chaudoir, 1857
Male adult
Cyprus, Troodos, 17.2.1V.2010,. Manfred Egger leg.
L=27 mm
Copyright © Ruslan Panin

Carabus (Procrustes) banoni Dejean, 1829
Male adult
Greece,W Crete Island, Mili gorge ,12.V.2009, Manfred Egger leg.
L=26 mm
Copyright © Ruslan Panin

Carabus (Procrustes) piochardi piochardi Gehin, 1883
Male adult
C-Lebanon, Faraya vill., Mt. Djebel Sannin h=1800m, X.2012, C. Reuter leg. <br /> L=31 mm
Copyright © Ruslan Panin

Carabus (Procrustes) coriaceus rugifer Kraatz, 1877
Female adult
W.UKRAINE, NE Carpathians, Lviv Reg., Skole distr., Zelemyanka Mt.Range h=700m, 26.V-1.VII.2010, Panin R. leg. L=43 mm
Copyright © Ruslan Panin

Carabus (Procrustes) coriaceus coriaceus Linne, 1758
Female adult
Belgique : Prov. Luxembourg, Harnoncourt 31.vii.1975 - M. Grotz Leg. (Size : 36 mm)
Copyright © Noël Mal

Carabus (Procrustes) coriaceus ressli Mandl, 1961
Male adult
SC-Turkey: Hatay Prov, Dörtyol, Nur Mts, Topaktaş h=900-1200 m, 12.V.2005, D. Prunier leg.
Copyright © Ruslan Panin

Carabus (Procrustes) coriaceus ressli Mandl, 1961
Female adult
SC-Turkey: Hatay Prov, Dörtyol, Nur Mts, Topaktaş h=900-1200 m, 12.V.2005, D. Prunier leg. L=35 mm
Copyright © Ruslan Panin

Carabus (Procrustes) chevrolati internatus Heinz & Korge, 1964
Female adult
Turkey, Bolu, Gerede, Esentepe 1350m, 1.7.1994, leg. K. Staven
Copyright © Alexander Anichtchenko

Carabus (Pseudocoptolabrus) burmanensis Breuning, 1932
Male adult
male, 23mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) businskyi Deuve, 1990
Male adult
CH, NW-Yunnan (Lijiang) Yulongxue Shan, E-slope
27:10:50N/100:12:26E/ 4400m, alpine meadows 12.VII.2000
K. & B. Brezina, M.Brychcin, M. Hackel
Copyright © Boleslav Brezina

Carabus (Pseudocranion) sackeni sackeni Semenov, 1898
Male adult
male, 23mm
Copyright © Boleslav Brezina

Carabus (Pseudocranion) sackeni remondianus Deuve, 1996 Paratype
Female adult
Museum of Institute of Systematic Biology (Daugavpils, Latvia)
Copyright © Alexander Anichtchenko

Carabus (Pseudocranion) lazikouensis lazikouensis (as kundaensis Deuve, 2003) Paratype
Female adult
C-China, S-Gansu: SSW of Minxian: Qilian Shan: W Kunda, 34:10N/103:47E, h=2800-3000m, forest, 7.VII.2002, Belousov & Kabak leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Pseudocranion) taibaishanicus taibaishanicus Deuve, 1989
Female adult
China, W-Shaanxi, Taibai Shan Mts, btw. Houzhenzi/Ba Xian Tai, nr. Nan Tian Men h=3200m, 33°55'17"N/ 107°46'53"E, 25-30.VII.2011, Li Wuyang leg. L=29 mm
Copyright © Ruslan Panin

Carabus (Relictocarabus) meurguesianus Ledoux, 1990
Male adult
Image from molbiol.ru
Copyright © - - -

Carabus (Rhigocarabus) buddaicus gansuicus Deuve, 1991
Male adult
China, NC-Sichuan (ZOIGE), LANGMUSI (=DOGCANGLHAMO), 34:03:48N/102:38:08E, h=4100m, alpine meadows, screes, 16-18.VII.1999, B.Brezina leg.
L=20 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) indigestus confucius Breuning, 1933
Female adult
C-China, E Tibet,W-Sichuan prov., Kangding, Zheduo Pass h=4300m 30.VI.2005 , V.Benes leg,
L=17 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) ladygini degensis Deuve, 1989
Male adult
C-China, NW- Sichuan prov., Garzê Tibetan AP, Pass Garzê – Sêrtar, 19.VII.2007, V.Benes leg.
L=17 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) buddaicus linxiaicus Deuve, 1995
Male adult
China, S-Gansu (LABRANG):, valley.E of Ponggartang, 35:12-13N/102:54-55E, h=3000-3700m, alpine meadows, coniferous forests,10.-14.VII.1999, B.Brezina leg. L=20 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) morawitzianus morawitzianus Semenov, 1887
Male adult
C China, Qinghai, Zadoi County, Toramarkog h~4600 m, VI.2001, P. Cavazzuti leg. L=17 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) turnai turnai Deuve, 1994
Male adult
China, Tibet, W-Sichuan, pass btw Sêrtar / Luhuo h=4200 m, 28.VI.2011, V.Patrikeev leg.L=19 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) turnai turnai Deuve, 1994
Female adult
China, Tibet, W-Sichuan, pass btw Sêrtar / Luhuo h=4200 m, 28.VI.2011, V.Patrikeev leg. L=21 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) roborowskii roborowskii Semenov, 1887
Male adult
China, E-Qinghai, Mts NW of Tongren city, 35.562778, 101.843667, h=3870 m, 15-26.VI.2011, G. Proscia leg./det.R.Panin, L=20 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) roborowskii roborowskii Semenov, 1887
Female adult
China, E-Qinghai, Mts NW of Tongren city, 35.562778, 101.843667, h=3870 m, 15-26.VI.2011, G. Pros cia leg./det.R.Panin, L=20 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) rhododendron Deuve & Imura, 1991
Male adult
China, N Yunnan: Baimaxue Shan, S of Deqen, h=4300m, 28°20'N 99°03'E, 16-20.VI.2016, C.Reuter leg., L=18 mm
Copyright © Ruslan Panin

Carabus (Scambocarabus) kruberi laobeiensis Deuve & Li, 2000
Male adult
NE-China: Liaoning prov., Fengcheng City, Miaoyang, 12-17.VI.2011
L=27 mm
Copyright © Ruslan Panin

Carabus (Semnocarabus) regulus regulus Dohrn, 1882
Male adult
photo M. Smirnov from www.zin.ru
Copyright © Maksim Smirnov

Carabus (Semnocarabus) carbonicolor carbonicolor Morawitz, 1886
Male+Female adult
Kyrgyzstan, NC Tianshan, Kyrgyz Mt. R., Tuyuk riv., h=2000 м.
25-VI-1997 М. Danilevsky leg.
male - 20 mm, female - 21 mm.
Copyright © Alexandr Safronov

Carabus (Semnocarabus) carbonicolor callosus Semenov, 1896
Male+Female adult
KAZAKHSTAN,
Bolshoe Almaatinskoe Lake,30.VI.1992/ det.Panin R.
female 19mm
Copyright © Ruslan Panin

Carabus (Semnocarabus) carbonicolor korolkowi Semenov, 1896
Male adult
NC-Kyrgyzstan, Kyrgyz Mt.Rng , 10 km SW of Bishkek, Chon-Aryk env, h=1100m, 24.V.1998, /det. Panin R.
22 mm

Copyright © Ruslan Panin

Carabus (Semnocarabus) erosus corax Semenov, 1896
Male adult
SE-Kazakhstan, Almaty reg., C Trans-Ili Alatau N-slopes, Kaskalen river Valley, Izvestkovaya vill., h=1700 m, 6.V.2015, D.Kovalchuk leg./det. R.Panin,
Copyright © Ruslan Panin

Carabus (Semnocarabus) erosus corax Semenov, 1896
Female adult
SE-Kazakhstan, Almaty reg., C Trans-Ili Alatau N-slopes, Kaskalen river Valley, Izvestkovaya vill., h=1700 m, 6.V.2015, D.Kovalchuk leg./det. R.Panin, L=24 mm
Copyright © Ruslan Panin

Carabus (Semnocarabus) carbonicolor korolkowi Semenov, 1896
Female adult
SE-Kazakhstan, Almaty, E Zailisky Alatau, Bokaydin-tau, Belshabdar valley h=1300 m, 5-27.VI.2002, A.Puchkov leg.L=21 mm
Copyright © Ruslan Panin

Carabus (Semnocarabus) carbonicolor carbonicolor Morawitz, 1886
Female adult
SE-Kazakhstan, Almaty reg., Khan Tengri Massif, Narynkol env.h=1800 m, 30-31.-V.2007, Shumov leg., L=21 mm
Copyright © Ruslan Panin

Carabus (Semnocarabus) erosus erosus Motschulsky, 1865
Endophallus
SE-Kazakhstan, 70 km E of Almaty, E Trans-Ili Alatau N-slopes, valley Assy (Asy) riv. Assy plateau h=1700 m, 12.V.2015
Copyright © Ruslan Panin

Carabus (Semnocarabus) rustemi Kabak, 2009
Male adult
SE-Kazakhstan, Almaty reg., near Bodokhudzir 43.916667, 79.716667 h=500 m, 12.VIII.2019, det. R.Panin, L=22 mm
Copyright © Ruslan Panin

Carabus (Shunichiocarabus) uenoianus uenoianus Imura, 1995
Male adult

Copyright © - - -

Carabus (Pagocarabus) draco Brezina, 1999
Male adult

Copyright © Boleslav Brezina

Carabus (Pagocarabus) caelestinus caelestinus (Imura, 2007) Paratype
Male adult

Copyright © Boleslav Brezina

Carabus (Pagocarabus) sichuanicola sichuanicola (as drolmae Lassalle, 2003)
Female adult
China, Sichuan, NW Chengdu, Mt. Jiajin Shan h~4000 m,VI. 2016, local collector, L= 24 mm
Copyright © Ruslan Panin

Carabus (Sigenacarabus) trachynodes moxiensis Deuve, 1992
Male adult
China, W-Sichuan, W of Moxi, Daxue Range, Mt.Gongga Shan h=3000-3100m, 29°34′29.6″N /101°59′44.0″E, 10-12.VI.2018, Ch. Reuter leg. L=21 mm
Copyright © Ruslan Panin

Carabus (Sigenacarabus) trachynodes moxiensis Deuve, 1992
Male adult
China, W-Sichuan, Daxue Range, Mt.Gongga Shan h=3000-3100m, Hailuogou Glacier Forest Park, 29.574889, 101.995556, 10-12.VI.2018, Ch. Reuter leg., L=21 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) varians janthinus s.str.
Male+Female adult
NW-Caucasus, Russia, Krasnodar reg.,Tuapse distr., Mt.Semashkho 04.V.2008, D.Fominyh leg.<br /> female, 24mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) adamsi Adams, 1817 (as chiragricus Fischer, 1828)
Male+Female adult
Russia, N. Caucasus, Stavropol reg., Zheleznovodsk, Zheleznaya Mt., h=600 m. 5-11.VI.2011, Е. Shankhiza leg. male - 24 mm, female - 25 mm.
Copyright ©

Carabus (Sphodristocarabus) varians varians Fischer, 1827
Male+Female adult
N. Armenia, Idzhevan distr., Acharkut vill. env., h=800 m.<br /> 26-28.VII.2008 S. Andreev leg.<br /> male - 22 mm, female - 23 mm.
Copyright ©

Carabus (Sphodristocarabus) macrogonus macrogonus Chaudoir, 1847
Male adult
NE-Turkey:Trabzon env., Sumela, 17-21.VI.2010, M.Janalik leg.
L=20 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) macrogonus pseudokolatensis Heinz & Korge, 1967
Male adult
NE-Turkey::Kackar Dagi Mt, Ayder S of Ardesen h=1600m, 17-21.VI.2010, M.Janalik leg.
L=20 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) separatus persa Lapouge, 1924
Male adult
N-Iran: Mazandaran Province, Elborz Mt.Rng, Vandarbon env., 10.VII.2002 L=23 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) separatus bararii Deuve, 2000
Male adult
N-Iran: Mazendaran: Ramsar env., deciduous forest, h~305 m, 16-24.VI.2017, Václav Čutka leg. L=22 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) separatus separatus Gehin, 1885
Male adult
N-Iran:Gilan Prov., Asalem - Khalkal road, 25-30.V.2018, Václav Čutka leg. L= 21 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) separatus haerii Heinz, 1973
Female adult
N-Iran:Gilan Prov., Rudbar, Harzevil env.,23-24.V.2018, Václav Čutka leg. L=24 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) varians armeniacus Mannerheim, 1830
Male adult
C Georgia, Imereti, Caucasus, Likhi Mt Range, Rikoti Pass h=900-1000 m, 14-16.V.2017, E. Rutjan E. leg., L=22 mm
Copyright © Ruslan Panin

Carabus (Tachypus) auratus brunieri Barthe, 1921
Male adult
SC-FRANCE, Aude, Rennes les Bains, 6.VII.1989, Sunard A. leg., 26mm
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
Male adult
Italy-Switzerland, Riviera – Preonzo, 18.II.1998, C.Sola leg.
L=26 mm
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus graniger Palliardi, 1825
Male adult
SW-Romania, Banatului Montan, Resita - Valiug, 29.V-9.VI.2005. Krause R. leg.
L=25 mm
Copyright © Ruslan Panin

Carabus (Tachypus) cristoforii Spence, 1823
Female adult
S France, Pyrénés Mts., Tarascon; 15mm
Copyright © Aleš Sedláček

Carabus (Tachypus) cancellatus durus Reitter, 1896
Male+Female adult
W Ukraine, Transcarpathian reg., Vinogradov distr., near Malaya Kopanya vill.,7-18.VII.2008, Panin R. leg. <br /> female, 32mm.
Copyright © Ruslan Panin

Carabus (Tachypus) vagans Olivier, 1795
Female adult
SE-France, Var: Le Muy env., 30.VII.2008., A. Paulian leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus intermedius Dejean, 1826
Female adult
NW -Serbia,Fruška Gora,Vrdnik env. h=380m, 9-10.VII.2013, M. Janalik leg. L=29 mm
Copyright © Ruslan Panin

Carabus (Tachypus) auratus honnoratii Dejean, 1826
Male+Female adult
France : Vaucluse, Mt. Ventoux-Mt. Serein, 1400-1800m, vii.1969-N. Mal Leg.-Coll.RBINS, N. Mal - including f.i. ventouxensis and nicolasi
Copyright © Noël Mal

Carabus (Tachypus) cancellatus strictus Kolbe, 1913
Female adult
W Ukraine, NE Carpathians, Maramaros Mts (Hutsul Alps), Gorge of Kuziy (Kizi) stream, h = 400m, 20-23.V.2021, Panin R. leg., L=32 mm
Copyright © Ruslan Panin

Carabus (Tanaocarabus) hendrichsi Bolivar y Pieltain, Rotger & Coronado, 1967
Male adult
male, 23mm
Copyright © Boleslav Brezina

Carabus (Oreocarabus) taedatus agassi Le Conte, 1850
Male adult
SC-USA, New Mexico, Santa Fe,VIII.1999, Sehnal R. leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Oreocarabus) taedatus bicanaliceps Casey, 1920
Male adult
NW-USA, Washington, Thurston County, Mima Mound Prairie, 15.IX.2005, A. Laforgue leg. L=23 мм
Copyright © Ruslan Panin

Carabus (Teratocarabus) azrael mizunumaianus Imura, 1991
Male adult
China, Liaoning, Benxi, Gongchangung
20.VI.2004 Hong
Copyright © Boleslav Brezina

Carabus (Tomocarabus) scabripennis scabripennis Chaudoir, 1850
Male adult
Carabus (Tomocarabus) scabripennis scabripennis Chaudoir, 1850
C. Armenia,
Kotayk reg., Yeghvard, Arailer Mt., 16.VI.1999, P.V.Kazaryan leg.
male, 20mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Tomocarabus) marginalis Fabricius, 1794
Male+Female adult
E Ukraine,
Kharkov reg.,near Vasishchevo vill.10-20.VI.2006 D.Kovalchuk leg.
female 24mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) decolor Fischer, 1823
Male adult
S Russia, NW-Caucasus,
Karacaj-Cherkessija,Skalisty Mnt. Range, Site Banditskoe h=1400m, 2-3.V.2009,
Fominykh D., Bondarenko A.S. leg.
male,17mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) convexus s.str. (as pyrenaicus Born, 1907)
Male adult
Pirineos, Canfranc, 1300m, Barranco de Izas
15.6.2005 Anichtchenko A. leg.
male, 16 mm
Copyright © Alexander Anichtchenko

Carabus (Tomocarabus) convexus gracilior Gehin, 1885
Male adult
SE-Bulgaria, Burgas Prov., Primorsko env., 18-30.VI.2000, Krause Rene leg.
L=18 mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) convexus moreanus Reitter, 1896
Female adult
Greece:, N Peloponnes, Xylokastro, Zemeno,h=1000m, 6.VI.1997, Manfred Egger leg. L=21 mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) rumelicus syriensis Breuning, 1943
Male adult
N Lebanon, Fnaпdek, Djebel Qammoua, h=1100-1200m, C. Reuter leg.
L=17.5 mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) convexus dilatatus Dejean, 1826
Male adult
NE Serbia, Majdanpek; 13mm
Copyright © Aleš Sedláček

Carabus (Tomocarabus) convexus gracilior Gehin, 1885
Female adult
SW Romania, Banat; 15mm
Copyright © Aleš Sedláček

Carabus (Tomocarabus) convexus erzeliki Schweiger, 1962
Female adult
NC-Turkey, Sinop province, Erfelek env., h=200m, 10-27.VII.2014, C. Reuter leg. L=20 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886
Male+Female adult
W Ukraine,<br /> Ternopol reg., Pidvolochisk distr., Supranivka vill.<br /> 3.VII.1908. Leg. Stobetskii. S.<br /> Photo R. Panin
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus bosphoranus Fischer, 1823
Male adult
Carabus (Trachycarabus) sibiricus bosphoranus Fisch. 1823<br /> S Ukraine, Krym Peninsula,<br /> Dolgorukovskaya jaila, 25.VIII.1991 I.A.Solodovnikov leg.<br /> male, 23mm<br />
Copyright © Ruslan Panin

Carabus (Trachycarabus) scabriusculus scabriusculus (as erythropus Fischer, 1822)
Male+Female adult
W.Ukraine,<br /> NW Podolian Upland , Lviv Reg., Zolochiv distr., near Luky vill., steppe sites, 7.V-8.VI. 2010<br /> leg. Panin R.<br /> female, 22mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) besseri Fischer, 1822
Male+Female adult
W Ukraine.NW Podolian,
Lviv reg., Zolochiv distr., Lysa Gora near Stinka vill., steppe sites, 25.IV-9.V.2009 Panin R. leg
female, 30mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) estreicheri Fischer, 1822
Female adult
Volgograd Reg., Serafimovka Distr., Ust' -Khoperskaya Station 5.V.2009 leg.O.Sveshnikov
18.5 mm
Copyright © Kirill Makarov

Carabus (Trachycarabus) sibiricus sibiricus Fischer, 1822
Male adult
E Siberia, Russia, Krasnoyarsk Reg., near Krasnoyarsk, on dry steppe slopes, 7.VIII,2011, M.Ivanov leg.
L=21 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) latreillei ? semicoriaceus Kraatz, 1881
Male adult
NE-China: Liaodong Peninsula, Liaoning prov., Gaizhou distr., Mt. Buyun Shan,VI.2007, Hao leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
Male adult
N Mongolia, Khövsgöl reg., Lake Khovsgol, h=1640m, 51*:12’N; 100*:45’E, steppificated sites,
20-25.VII.2010. A.Dergalev leg.
L=22 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) estreicheri Fischer, 1822 (ab.rufofemoratus Lomnicki, 1892)
Female adult
W Ukraina, Podolian Upland, Lviv reg., Brody distr., Mt. Makitra, steppe site, 19.V-5.VI.2020, Panin R. leg. L=23 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus campicola Dejean, 1831
Female adult
S Ukraine, Crimea, S slope Mt Demerdzhi h=650 m, sibljak, VII.2004, R. Panin leg., L=29 mm
Copyright © Ruslan Panin

Carabus (Tribax) puschkini puschkini Adams, 1817
Male adult
Armenia,<br /> Shirak Province, Ashotsk, 18.VI.2001, P.V.Kazaryan leg.<br /> male 25mm
Copyright © Ruslan Panin

Carabus (Tribax) puschkini ishikawaianus Breuning & Ruspoli, 1970
Male adult
NE-Turkey: Artvin:Hopa, Borçka env., h=450 m, 29.VI-7.VII.2011, M.Janalik leg.
L=33 mm
Copyright © Ruslan Panin

Carabus (Tribax) agnatus pseudagnatus Novotný & Voříšek, 1988
Female adult
NW-Caucasus, Karachay-Cherkessia, Bol'shaya Marka River h=2500-2600m, 12.VIII.2010, I.A. Solodovnikov leg./det.
Copyright © Ruslan Panin

Carabus (Tribax) puschkini platypterus Ganglbauer, 1886
Male adult
SW Georgia, Lesser Caucasus, Adzhar-Imereti (Meskheti) Mt Range, 10 km S Sairme h~1850 m, 27.VI-2.VII.2013, A.Puchkov & Martynov leg. L=27 mm
Copyright © Ruslan Panin

Carabus (Tribax) nothus sharoiensis Fominykh, Zamotajlov, Avtaeva et Titarenko, 2019 Paratype
Male adult

Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum kasakorum Semenov, 1896
Female adult
Russia: N.Osetia: Pastbyshchniy Mt.R., S slopes of Khoshkharanrag Mt.R., right bank of Amusadon riv., h=1500 m
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
Female adult
Russia: N.Osetia, Bokovoy Mt.R., near Arkhonskiy pass., alpin zone, h = 2700 m, 23.V-21VII.2018, D. Fominykh leg.
Copyright © Igor Solodovnikov

Carabus (Tribax) puschkini kolenatii Chaudoir, 1846
Male adult
C-Georgia, Trialeti Mt Range, 2 km SE of Borjomi h=916 m, 41°48'35.3"N 43°26'11.8"E, 17.VI.2013, D. Prunier leg./det. Panin R., L=28 mm
Copyright © Ruslan Panin

Carabus (Tribax) kasbekianus veselyi Gottwald, 1980
Male adult
Georgia, Caucasus, S of Sasashi vill., Lechkhumi Range N slope h=2430 m, alpine meadows, 242.759167, 42.973889, 20-21.VII.2025, I.Uspenskyi leg./det. R.Panin, L=24 mm
Copyright © Ruslan Panin

Carabus (Tribax) kasbekianus veselyi Gottwald, 1980
Endophallus
Georgia, Caucasus, S of Sasashi vill., Lechkhumi Range N slope h=2430 m, alpine meadows, 242.759167, 42.973889,
Copyright © Ruslan Panin

Carabus (Ulocarabus) stschurovskii Solsky, 1874
Male adult
Tadzhikistan, Pamir-Alay, S slope Gissar Mt.Range, Takob, h-1700m, 10.V.1996, Dolin V.G. leg.
det. Panin R.Yu.
L=22 mm
Copyright © Ruslan Panin

Carabus (Ulocarabus) stschurovskii Solsky, 1874
Female adult
Tadzhikistan, Pamir, N-slope Peter I.Mt Rng,h=2100m,Ganishou vill., Dolin V.G. leg.
det.Panin R.Yu.
L=24 mm
Copyright © Ruslan Panin

Carabus (Ulocarabus) theanus delerei Mandl, 1955
Male adult
N-Afghanistan, Hindukush Mt.Rng, NE Salang Pass h=2300 m, 35°27'494"N, 68°58'399"E, IV.2011, C. Reuter leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Ulocarabus) theanus theanus Reitter, 1895
Male adult
Tadzhikistan, S slope Gissar Mt.Rng, left bank of the river Sorbo, Haidar gorge,1.VI.1984, Dolin V.G. leg./det.Panin R.Yu.
L=21 mm
Copyright © Ruslan Panin

Carabus (Ulocarabus)? theanus theanus Reitter, 1895
Male adult
NC Tadzhikistan, Pamir, N-slope Peter I.Mt Rng, h=2100m, Ganishou vill., 10-12.VI.2006, Ivanova E. leg.
L=23 mm
Copyright © Ruslan Panin