Carabus (Acathaicus) alexandrae alexandrae Semenov, 1887 Holotype Holotype
Male adult
Kwantin and Li-tschara-pu, 22.VI.1885, leg. Potanin<br /> Deposited in Zoolog. Museum St. Petersburg.<br /> male, long. 22 mm<br /> photo B. Brezina
Copyright © Boleslav Brezina

Carabus (Acathaicus) alexandrae cratocephaloides Semenov, 1887
Female adult
female, 28mm
Copyright © Boleslav Brezina

Carabus (Acathaicus) interruptocostatus Semenov, 1887 Holotype Holotype
Female adult

Copyright © Boleslav Brezina

Carabus (Acoptolabrus) lopatini Morawitz, 1886
Male+Female adult
Russia,
Sakhalin isl., Tchekhov Mt, VI.1997
female, 40mm
photo & coll. Panin R.
Copyright © Ruslan Panin

Carabus (Acoptolabrus) schrencki schrencki (as hauryi Gehin, 1885 )
Male adult

Copyright © Boleslav Brezina

Carabus (Acoptolabrus) schrencki kimieae Kezuka & Imura, 1989
Male+Female adult
Mt. Wulongshan, Dandong City,
Liaoning Province, CHINE
25-30.VI.2008
Copyright © Safronov

Carabus (Coptolabrodes) haeckeli Brezina & Imura, 1997
Male adult
C China, Shaanxi,
Tsinling Mts., Taibaishan Mt., h=3000m, 33:55N/107:44 E, VI.2004, V.Siniaev & his team
female, 31 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Acoptolabrus) schrencki schrenck (as dongningensis Deuve & Li, 2005)
Male adult
NE China, Heilongjiang prov., Dongning, V.2009<br /> L=25 mm
Copyright © Ruslan Panin

Carabus (Acoptolabrus) constricticollis constricticollis (as grallatorius Roeschke, 1921)
Male adult
NE-Korea:North Hamgyong Prov., Sechon-dong, VII.2004, Li leg.
L=28 mm
Copyright © Ruslan Panin

Carabus (Acoptolabrus) gehinii gehinii Fairmaire, 1886
Female adult
N-Japan, SW-Hokkaido, Chitose env.,VI.2011.
L=34 mm
Copyright © Ruslan Panin

Carabus (Acoptolabrus) constricticollis pihyonensis Deuve & Li, 1998
Male adult
N-Korea, North Pyongan Prov., Tongnim County, 5-25.V.2010 <br /> 29 mm
Copyright © Ruslan Panin

Carabus (Acoptolabrus) ? leechi huisaensis Deuve & Li, 1999
Male adult
NC-Korea, Ryanggang Prov., Puksubaek-san, 4-10.V.2010
28 mm

Copyright © Ruslan Panin

Carabus (Acoptolabrus) constricticollis Kraatz, 1886
Male adult
China, Liaoning prov., Mt. Guanmen 20.6.2009 (cAA)
Copyright © Alexander Anichtchenko

Carabus (Carabus) gressittianus fuscomarginatus Kleinfeld, 1997 Holotype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Carabus) andreiianus Deuve, 2007
Female adult
C-China, N-Sichuan, Pingwu Co. h=1500 m, 27.VI.2011, L=24 mm
Copyright © Ruslan Panin

Carabus (Carabus) koidei koidei Imura, 1996
Male adult
C-China, S-Sichuan, Pass btw. Yanyaun / Muli, h=3244 m, 27.68638N; 101.22335E,,11-18.VI.2017, Ch. Reuter leg. L=22 mm
Copyright © Ruslan Panin

Carabus (s.str.) morphocaraboides zhaojueanus Imura, 1999
Female adult
China, S-Sichuan, SE Mianning, Pass btw Mianshanzhen & Yuexi 28°30’N/102°26’E, 22-23.VI.2017, Ch.Reuter leg., L=22 mm; det. B. Brezina, 2020
Copyright © Ruslan Panin

Carabus (s.str.) morphocaraboides zhaojueanus Imura, 1999
Male adult
China, S-Sichuan, road 56 km E of Xichang to Zhaojue, Mt.WenLin, Pass h=3200 m, 27°52’N/102°30E, 5-20.VI.2017, Ch.Reuter leg.; det B. Brezina, 2020
Copyright © Ruslan Panin

Carabus (Acoptopterus, ?Carabus) yunnanus yanyuanicus Cavazzuti, 1996
Endophallus
China, S-Sichuan, S Yanyuan, Chuandongzi Mt h=3300-3600 m
Copyright © Ruslan Panin

Carabus (Acoptopterus) yunnanus yanyuanicus Cavazzuti, 1996 (? or oblongior ssp.)
Male adult
China, S-Sichuan, S Yanyuan, Chuandongzi Mt h=3300-3600 m, 16-20.VI.2023, C.Reuter leg./det.Klienfeld, 2023, L=21.5 mm
Copyright © Ruslan Panin

Carabus (Acrocarabus) callisthenoides Semenov, 1888
Male adult
male, 22mm
Copyright © Boleslav Brezina

Carabus (Acrocarabus) guerini aldarkose Kabak, 1992
Male adult
male, 22mm
Copyright © Boleslav Brezina

Carabus (Acrocarabus) guerini guerini Fischer, 1842
Male adult
SE Kazakhstan, W Dzhungar Mts, Lepsinsk env., 1600 m., 18.06.2000, A.Klimenko
Copyright © Anton Menstschikov

Carabus (Alipaster) barovskii bayanbulak Deuve, 1993 Paratype Paratype
Female adult
female, 17mm
Copyright © Boleslav Brezina

Carabus (Alipaster) infantulus infantulus Morawitz, 1886
Male adult
male, 15mm
Copyright © Boleslav Brezina

Carabus (Eotribax) eous eous Morawitz, 1889
Male adult
Kirghyzstan, N Terskei Ala Too, Borskaun riv., h=2500 m., 7-14.07.2000
male
L=17 mm
Copyright © Maksim Smirnov

Carabus (Alipaster) pupulus pupulus Morawitz, 1889
Female adult
S Kyrgyzstan, Inner Tian Shan, Dzhany-Dzher Mt range, upper Mudyuryum riv. Ak-Baytal gorge h~3300 m, 15.VII.2011, S.Toporov leg. L=16 mm
Copyright © Ruslan Panin

Carabus (Apoplesius) deuvei hajdaji Hackel, 2007 Holotype Holotype
Male adult
Turkey- prov. Mus 60 km NW Mus 1630m
3856N 4108E Bulgan Gec. 23-24.06.03 E. & P. Hajdaj leg.
Copyright © Boleslav Brezina

Carabus (Apoplesius) omphreodes Reitter, 1898
Male adult

Copyright © Kirill Makarov

Carabus (Apoplesius) pseudodepressus Machard, 1988
Female adult
Turkey, Prov. Bitlis, S near Tatvan, 23.IV.2009 leg. M. & S. Nabozhenko
L=19.0 mm
Copyright © Kirill Makarov

Carabus (Apotomopterus) bousquetellus gengmaensis Kleinfeld, 2003 Holotype & Paratype Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) cantonensis pervarius Kleinfeld, 1997 Holotype
Male adult
Frank Kleinfeld
Copyright © - - -

Carabus (Apotomopterus) delavayi s.str. (as medea Kleinfeld, 1999) Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) casaleianus casaleianus Deuve, 1994
Female adult
China, SW Hunan Prov., Chengbu Co, Nan-Shan-Ding (26.34;110,08)1-10.VIII,2006, I.Rapuzzi leg.
L=33 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) protenes baoxingensis Deuve & Mourzine, 2003
Female adult

Copyright © Igor Solodovnikov

Carabus (Apotomopterus) protenes ascendens Semenov, 1898
Male adult
China, Chongqing, N Jinfo Shan, Shanwangping h=1400m, 29.101944, 107.166389, 2-8.VI.2018, C.Reuter leg./det. F.Kleinfeld, L=26 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) madefactus Cavazzuti, 1997
Male adult
China, CW-Yunnan, Nánjiàn Yízú Zìzhìxiàn, Mt Lingbao h=2400 m, 24.768611, 100.500000, 6-15.VI.2023, C.Reuter leg./det. F.Kleinfeld, L=27 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) patroclus apricalis Cavazzuti, 1997
Male adult
China, S Sichuan, W of Pingchuanzhen, near Galuo vill., h=2400 m, 7-18.VI.2017, C.Reuter leg./det. F.Kleinfeld, L=30 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) patroclus patroclus Breuning, 1933
Male adult
China, S Sichuan, Pass 56 km SW Miannin, near Wawo h=2440 m, 21-24. VI.2017, C.Reuter leg./det. F.Kleinfeld, L=29 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) bousquetellus bousquetellus Deuve, 1998
Male adult
S-China, CW-Yunnan, W of Lincang, Daxue Shan h=2600m, 23.935556, 100.257778, 7-13.VI.2023, C.Reuter leg./det. F.Kleinfeld, L=24 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) monticola fontanae Born, 1906
Male+Female adult
N-Italy,
Piemonte, Bielmonte, h1350 m, 14-15.VI.2010, Pascal Stefani leg.
female, 23 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Archicarabus) montivagus montivagus Palliardi, 1825
Male+Female adult
SW-Romania: Banat, Caras-Severin Reg., Sasca Montana – Stinapari, 21.V.-5.VI.2010, Krause Rene leg.
female, 26 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) phoenix Lapouge, 1924
Male+Female adult
C-Lebanon, Byblos /Jbeil Distr., Kfardebian env., h=1200m, XI.2014, C. Reuter leg. female,23mm
Copyright © Ruslan Panin

Carabus (Archicarabus) victor Fischer, 1836
Female adult
NE-Turkey, Giresun, Sehitler Geçidi (Pass) h~2000 m, 23.VII.2001, S. Dacatra leg., L=19 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) alysidotus Illiger, 1798
Female adult
Italy, Reggio Emilia, Albinea env. h=150 m, VI.2018, A. Cartini leg., L=25 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) wiedemanni congruus Motschulsky, 1865
Female adult
Turkey, Cankiri, Cerkes - Kursunlu, Atkaracalar 1250m 10.7.1996 Leg Staven and Skoupy (DUBC)
Copyright © Alexander Anichtchenko

Carabus (Archicarabus) wiedemanni wiedemanni (as mussardianus Breuning, 1965)
Female adult
NC-Turkey, Sinop province, Erfelek env., h=200m, 10-27.VII.2014, C. Reuter leg. L=24 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) wiedemanni (as mussardianus Breuning, 1965)
Male adult
NC-Turkey, Sinop province, Erfelek env., h=200m, 10-27.VII.2014, C. Reuter leg. L=23 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) montivagus vellepiticus Hampe, 1850
Female adult
W Croatia, NP Northern Velebit, Stolak, 44.969492, 14.986169, 23.V-27.VI.2022, L=24 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) montivagus vellepiticus Hampe, 1850
Male adult
W Croatia, NP Northern Velebit, Stolak, 44.969492, 14.986169, 23.V-27.VI.2022, L=23.5 mm
Copyright © Ruslan Panin

Carabus (Aristocarabus) viridifossulatus viridifossulatus Fairmaire, 1887
Female adult
35mm
Copyright © Boleslav Brezina

Carabus (Aristocarabus) viridifossulatus seticollis Deuve & Mourzine, 2000
Male adult
28 mm
Copyright © Boleslav Brezina

Carabus (Aulonocarabus) truncaticollis polaris Poppius, 1905
Male adult
Arctic Russia, NW Siberia,
Yamal, 230 N from Labytnangi, Panta-Yakha Riv., 15-29.VI.2002
male,18mm.
Copyright © Ruslan Panin

Carabus (Aulonocarabus) canaliculatus pseudocareniger Deuve, 1991
Male adult
N Korea, Ryanggang Prov., Mt. Tongjom Ridge h~1861m, (128.60E;41.20N) I. Rapuzzi leg.
L=27 mm
Copyright © Ruslan Panin

Carabus (Aulonocarabus) gaschkewitschi czarensis Obydov, 1997
Female adult
Russia, CE-Siberia, N Chita reg., Kodar Mt. Range, Big Leprindo Lake env. h~2000m., 2-5.VII.2002 A. Prokhorov leg./det. A. Anichtchenko
L=19 mm
Copyright © Ruslan Panin

Carabus (Aulonocarabus) canaliculatus hailinensis Deuve & Li, 2000
Male adult
NE China, Heilongjiang Prov., Mudanjiang City, Mt.Tai-Gu-Ding-zi h=800m, 15-25.VI.2010
L=29 mm
Copyright © Ruslan Panin

Carabus (Aulonocarabus) truncaticollis kodaricola Deuve, 1998
Female adult
SE-Siberia:N-Chita reg., Kodar Mt.Rng.,h=1800m, Leprindo vill., 1.VII.2006,
L=17 mm
Copyright © Ruslan Panin

Carabus (Aulonocarabus) kurilensis ? daisetsuzanus Kono, 1936
Male adult
N-Japan, Hokkaido, Katō District,Kamishihoro, Horoka near Maruyama Bridge,19.VI.2013, T. Hirabayashi leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Aulonocarabus) gaschkewitschi gaschkewitschi Motschulsky, 1859
Male adult

Copyright © Kirill Makarov

Carabus (Aulonocarabus) kolymensis kodarensis Brinev, 1997
Male adult
Russia, SE Siberia, N Chita reg., Kodar Mt. Rng. h=1800m, Novaya Chara vill., 23.VI.2006. L=17 mm
Copyright © Ruslan Panin

Carabus (Aulonocarabus) truncaticollis fleischeri Reitter, 1898
Female adult
Russia, N Burajatia, Barguzinskiy Mts., the upper reaches of M.Chivyrkuy riv., h=2000 m, 26.VI-12.VII.2007, Vashchenko S leg. L=18.5 mm
Copyright © Ruslan Panin

Carabus (Aulonocarabus) truncaticollis fleischeri Reitter, 1898
Male adult
Russia, N Burajatia, Barguzinskiy Mts., the upper reaches of M.Chivyrkuy riv., h=2000 m, 26.VI-12.VII.2007, Vashchenko S leg. L=17 mm
Copyright © Ruslan Panin

Carabus (Axinocarabus) fedtschenkoi fedtschenkoi Solsky, 1874
Male adult
27 mm
Copyright © Boleslav Brezina

Carabus (Axinocarabus) fedtschenkoi kondarensis Kryzhanovskij, 1953
Female adult
NW-Tadzhikistan: Gissar Mt.Rng, Varzob river ravine, Gushari env., h~1400-1500m, 24.III.2010, L=29 mm
Copyright © Ruslan Panin

Carabus (Calocarabus) linxiaensis ponggartangensis Deuve, 1995 Holotype
Male adult
17 mm
Copyright © Boleslav Brezina

Carabus (Cryptocechenus) sifanicus Semenov, 1898 (as oxydocupreus)
Male adult
male, 17mm
Copyright © Boleslav Brezina

Carabus (Carabus) arvensis arvensis Herbst, 1784
Male+Female adult
W Ukraine, NW Podolian, Lviv reg., Javoriv distr., near Stradch vill., 20-30.IV.2009, Panin R. leg.
Copyright © Ruslan Panin

Carabus (Carabus) granulatus granulatus Linne, 1758
Female adult
long. 22mm
Copyright © Ruslan Panin

Carabus (Carabus) menetriesi menetriesi Hummel, 1827
Male adult
W-Ukraine, Roztochia, Lviv Reg., Yavorov distr., near Ivano-Frankovo vill., Site Zalyvki, Panin R. leg., 1-20.V.2005, male, 21 mm.
Copyright © Ruslan Panin

Carabus (Carabus) sculpturatus Menetries, 1832
Male+Female adult
Azerbaijan,
Talysh Mts. Hirkan National Park, Moscow Forest, 8-9.VII.1988, P.V.Kazaryan leg.
female, 22 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Carabus) granulatus interstitialis Duftschmid, 1812
Male adult
Italy, Emilia reg., Modena, Castelnuovo Rangone, 31.XII.1991, Sola C. leg.
L=19 mm
Copyright © Ruslan Panin

Carabus (Carabus) arvensis carpathus (as pseudonoricus Kenyery, 1983)
Male adult
Slovenia: Julske Alpy, Podvelka, h=450m, 24.X.2010, M.Janalik leg.
L=19 mm
Copyright © Ruslan Panin

Carabus (Carabus) granulatus granulatus Linne, 1758
Male adult
W-Ukraine, Roztochia,
Lviv Reg., Javorov distr., near Ivano-Frankovo vill., Site Zalyvki, 17.IV-3.V.2010, Panin R. leg.
L=20 mm

Copyright © Ruslan Panin

Carabus (Carabus) arvensis hokkaidoensis Lapouge, 1924
Female adult

Copyright © Kirill Makarov

Carabus (Carabus) granulatus yezoensis Bates, 1883
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Carabus (Carabus) menetriesi menetriesi Hummel, 1827
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Carabus (Cathaicus) brandti brandti Faldermann, 1835 (as grossegranulatus) Syntypes Lectotype
Male adult
C. (Cathaicus) brandti grossegranulatus Breuning 1943<br /> Lectotypus/SyntypusLectotypus (25 mm) von "South Mandschou". Syntypus (25 mm) "South Mandschou, 30. V. 1937"
Copyright © Frank Kleinfeld

Carabus (Cathoplius) aliai Escalera, 1944
Male adult
male, 27mm
Copyright © Boleslav Brezina

Carabus (Cathoplius) asperatus asperatus (Dejean, 1826)
Male adult
male, 24mm
Copyright © Boleslav Brezina

Carabus (Procechenochilus) kokujewi Semenov, 1898
Female adult
RUSSIA, Karachay-Cherkessia, NW Caucasus Mts., Daut Mt. Range, Iskrovtsev - W.Ullu-Kel Mt. pass circus, h=2800-2600m
Copyright © Alexander Anichtchenko

Carabus (Procechenochilus) ? kokujewi Semenov, 1898
Female adult
Russia, NW Caucasus, Karachay-Cherkessia, Karachaevsk distr., near Uchkula vill., valley Uchkulanichi riv. h=2800-2900m, 25-28.VII.2010, Fominykh D. leg.
female, 23 mm
Copyright © Ruslan Panin

Carabus (Cechenochilus) adangensis Gottwald, 1983
Female adult
Russia, NW-Caucasus:
Karacaj-Cherkessija, Abishira–Akhuba Mt. Range h=2000-2650m, Kyafar-Agur Riv. 14–17.VII.2009, Fominykh D. leg.
female,21mm
Copyright © Ruslan Panin

Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Holotype
Male adult
Russia, Karachai-Circassian Republic, Abishira-Akhuba Mount Range between passes Agur and Mylgvar, 2785–2830 m.
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Paratype
Male adult
Russia, W Caucasus,
Karacaj-Cherkessija, Abishira-Akhuba Mnt Range, Area of pass Turii h~2650-2850m, 17.VII.2009, Fominykh D. leg.
male,17 mm
photo & coll. Panin R.
Copyright © Ruslan Panin

Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Paratype
Male adult
Russia, Karachai-Circassian Republic, Abishira-Akhuba Mount Range between passes Agur and Mylgvar, 2785–2830 m.
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) kokujewi Semenov, 1898
Female adult
Russia, Karachai-Cherkessia, vall. Daut Riv., near Glacier Ryndzhi-Murudzhu, h = 3100 m, 22.VIII.2011, leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) kokujewi Semenov, 1898
Male adult
Russia, Karachai-Cherkessia, vall. Daut Riv., near Glacier Ryndzhi-Murudzhu, h = 3050 m, 22.VIII.2011, leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) boeberi boeberi Adams, 1817
Female adult
Грузия, Мцхета-Мтианети, к северу от пос. Гудаури, ЮВ отроги г. Байдарская (нижний пик), по ущелью ручья, альпика, под камнями, точка 2, h = 2700 м, 42.495413° N, 44.487844° E, 10.08.2019, leg. И.А., А.И., С.В. Солодовниковы, О.И. Данилова 26 mm
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) boeberi aequaliceps Reitter, 1896
Male adult
Russia, Dagestan, Andiyskiy Mt.R., near Andiyskiye vorota pass, 06.VI–18.VII.2021, leg. D. Fominykh
Copyright © Igor Solodovnikov

Carabus (Cechenotribax) petri Semenov & Znojko, 1932
Female adult
female, 18mm
Copyright © Boleslav Brezina

Carabus (Cechenotribax) petri Semenov & Znojko, 1932
Male adult
SE Kazakhstan, N-Tianshan:N-slopes Ketmen Mt.Rng. h=2250-2500m., near Ketmen vill.,1-5.VI.1990, Dolin V.G. leg.
L=13.5 mm
Copyright © Ruslan Panin

Carabus (Cephalornis) potanini (Semenov, 1887)
Male adult
China, S Gansu, 3400m, Minshan mts.,
Wenxian 8.92, J. M. Bousquet
Copyright © Boleslav Brezina

Carabus (Chaetocarabus) intricatus bayardi Solier, 1835
Male+Female adult
S-Italy,
Basilicata Reg., Potenza Prov., Latronico, F. Angelini leg. 14.VI.2002
female, 26mm
Copyright © Ruslan Panin

Carabus (Chaetocarabus) intricatus krueperi Reitter, 1896
Male adult
CE-Greece: Pieria, Stomio (Larissa) Ossa Mt. 7-14.VII.2009, Krause Rene leg.
L=35 mm
Copyright © Ruslan Panin

Carabus (Chaetocarabus) arcadicus arcadicus Gistl, 1850
Female adult
CE-Greece, Kato Olympos, Karia env., 6-17.VII.2011, Krause Rene leg. L=37 mm
Copyright © Ruslan Panin

Carabus (Chaetocarabus) arcadicus merlini Schaum, 1861
Male adult
S-Greece:Peloponnesos: Taygetos Mt.Rng., 17-23.V.2005, V. Sima leg.
L=30 mm
Copyright © Ruslan Panin

Carabus (Chaetocarabus) arcadicus arcadicus Gistl, 1850
Male adult
Graecia, Pindus, Mt. Velouchi (Timfristos), 1200m, vi.1970-A. Sermet Leg., 30 mm
Copyright © Noël Mal

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Male adult
W Ukraine, Lviv city, Sykhiv, Zubra forest, (Osvytskiy Grove), under bark, 49°47'28.10" N, 24° 2'20.09" E, 10.XI.2017, Panin R leg.
Copyright © Ruslan Panin

Carabus (Chaetocarabus) intricatus bayardi (as silaensis Haury, 1881)
Female adult
S-Italy, Calabria, Sila Grande Massif, Camigliatello h=1040 m, 19.XI.2009, G. Proscia leg. L=31 mm
Copyright © Ruslan Panin

Carabus (Chaetocarabus) intricatus bayardi (as silaensis Haury, 1881)
Male adult
S-Italy, Calabria, Sila Grande Massif, Camigliatello h=1040 m, 19.XI.2009, G. Proscia leg. L=28 mm
Copyright © Ruslan Panin

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Female adult
W Ukraine, Lviv city, Holosko, Bryukhovytsʹkyi forest 49.886833, 24.012083 under bark, 10.XI.2024, R. Panin leg., L=35 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860
Male+Female adult
SW -France,
Pyrenees-Atlantiques, Foret de Sare, XII.2009, Pascal Stefani leg.
female, 31 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as delacrei Tarrier, 1975)
Male+Female adult
NE Spain, Catalonia, Sant Jaume de Frontanyа, h=1000 m, V-VI.2010, Pascal Stefani leg.
female, 38 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens costellatus Gehin, 1882 (as subfestivus Oberthur, 1884)
Female adult
W- FRANCE,
Bretagne Reg., Finistrere, Foret de Coatloc'h, XI.2006
female, 26 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens cupreonitens Chevrolat, 1861
Female adult
NW-FRANCE,
Basse-Normandie Reg., Orne, Forêt de Bellême, 14.XII.2008, Pascal Stefani leg.
female, 26mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens cupreonitens Chevrolat, 1861
Male+Female adult
NW-FRANCE,
Basse-Normandie Reg,, Orne, Foret d' Ecouves, V.2009, Pascal Stefani leg.
male, 25 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens montanus Gehin, 1882
Male+Female adult
SC-FRANCE,
Haute-Garonne, Col de Mentй h~1300m., II.1980. Kretschmer leg.
female, 23mm.
photo/det. Panin R.
Copyright © Ruslan Panin

Carabus (Chrysocarabus) lineatus lateralis Chevrolat, 1840 (as quasilineatus Tarrier, 1975)
Male+Female adult
NW Spain,
E Asturias, Monte de Moal h~1000m., 5.VI.2008, Pascal Stefani leg.
female, 27mm.
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens festivus Dejean, 1826 (ab. holochrysum)
Female adult
SC-FRANCE, Tarn, Le Linas h=550 m, XI.2008, Pascal Stefani leg. L=24 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) solieri bonnetianus Colas, 1936 (loc.typ.)
Female adult
SE-France, Var, Tanneron, Saint-Cassien des Bois, 18.IV.2019, L=32 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) solieri bonnetianus Colas, 1936 (loc.typ.)
Male adult
SE-France, Var, Tanneron, Saint-Cassien des Bois, 18.IV.2019, L= 31 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) smaragdinus longipennis Chaudoir, 1863
Male+Female adult

Copyright © Alexandr Safronov

Carabus (Coptolabrus) pustulifer pustulifer Lucas, 1869
Female adult
Chine, C.Sichuan, Ya'an, 07.2005 Local people
Copyright © Igor Solodovnikov

Carabus (Coptolabrus) pustulifer (as cyanelongatus Deuve, 2006)
Female adult
Zhagyab, Quamdu City, Tibet<br /> 8-15.VI.2008
Copyright ©

Carabus (Coptolabrus) lafossei tiantai Kleinfeld, 1997 Holotype
Female adult

Copyright © Frank Kleinfeld

Carabus (Coptolabrus) elysii magnificens Kleinfeld, 1999 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Coptolabrus) smaragdinus liaodongensis (Li, 1992)
Male adult
NE-China, E-Liaoning: Anshan: Mt.Qian Shan, 9-14.VI.2011. L=39 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) gemmifer pseudoformosus Deuve, 2006
Female adult
China, Tibet AR, Qamdo, Gonjo County, Kè Rì Xiāng vill.(Kerri Township), h=3100m, VI.2012, A. Mollard leg. L=38 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) jankowskii fusanus (as obtusipennis)
Female adult
SC-Korea: Chiri-san Massif, Mt. Jirisan, 28.VI.2011, L=35 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) elysii xiaoganicus Deuve et Remond, 2016
Male adult
China, Hubei, Xiaogan, Mt Shuangfeng Shan, VII.2016 L=37 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) ignimitella angulicollis Hauser, 1911
Male adult
China Hunan, Guidong pr., 20.5.2009 (cAA) 44 mm
Copyright © Alexander Anichtchenko

Carabus (Cratocarabus) puer Morawitz, 1886
Female adult
female, 16mm
Copyright © Boleslav Brezina

Carabus (Cratocechenus) jacobsoni Semenov, 1898
Male adult
S Kazakhstan, Almaty reg., Big Lake, h=2800 m, 25.VI.96
male
L=14 mm
Copyright © Maksim Smirnov

Carabus (Cratocechenus) akinini akinini Morawitz, 1886 (as oitalensis ) Paratype
Male adult
male, 20mm
Copyright © Boleslav Brezina

Carabus (Cratocechenus) akinini akinini Morawitz, 1886
Female adult
female, 21mm
Copyright © Boleslav Brezina

Carabus (Cratocechenus) ovtchinnikovi ovtchinnikovi Gottwald, 1987
Male adult
N-Kyrgyzstan, Kyungey Ala-Too Mt Rng, Cholpon-Ata env., h=2500m, 19.VI.2000,
L=19 mm
Copyright © Ruslan Panin

Carabus (Cratocephalus) solskyi Ballion, 1878
Male adult
SE Kazakhstan, Ketmen Mt. rng., Tuyuk vill., 16.06.97 leg. S.Toropov
Copyright © Maksim Smirnov

Carabus (Cratocephalus) solskyi Ballion, 1878
Female adult
SE-KAZAKHSTAN, Ketmen Mt Rng, h=1800m, Tujuk vill.,15.V.2004
L=36 mm
Copyright © Ruslan Panin

Carabus (Cratocephalus) cicatricosus cicatricosus Fischer, 1842
Male adult
SE-Kazakhstan, Almaty reg., C Trans-Ili Alatau N-slopes, Kaskelen Valley, Izvestkovyi vill., h=1700 m, 6.V.2015, D.Kovalchuk leg., L=28.5 mm
Copyright © Ruslan Panin

Carabus (Cratophyrtus) turcosinensis medvedevi Kryzhanovskij, 1968
Female adult
S Kazakhstan, NW slope of Ugamskii mt.rng., 3 km W Sairamskii peak, 2200m
tributary of Karzhantau riv., 15.6.2004 Matalin A. leg.
20 mm
Copyright © Alexander Anichtchenko

Carabus (Cratophyrtus) kaufmanni segregatus Morawitz, 1886
Female adult
female, 19mm
Copyright © Boleslav Brezina

Carabus (Cryptocarabus) subparallelus Ballion, 1878
Female adult
Kazakhstan,
near Alma-Ata, V.2009, Pak O.leg.
female, 17mm
Copyright © Ruslan Panin

Carabus (Cryptocarabus) lindemanni Ballion, 1878
Female adult
SE Kazakhstan, Alma-Ata env., 16.04. 2000
L=16,0 mm
Copyright © Maksim Smirnov

Carabus (Cryptocarabus) ? turkestanus turkestanus Breuning, 1929
Male adult
Kyrgyzstan, W Kungei Ala-Too Mt Rng, Chon-Kemin riv., Kyzyl-Bairak vill., 25.IV.2010. L= 18 mm
Copyright © Ruslan Panin

Carabus (Cryptocarabus) tsharynensis Kabak 1994
Male adult
SE-Kazakhstan, Almaty Region, W-Zailiyskiy Alatau rng, Belshabdar valley h~1300 m, 25-27.VI.2002, A.Puchkov leg. L=17 mm
Copyright © Ruslan Panin

Carabus (Cryptocarabus) tsharynensis Kabak 1994
Female adult
E-Kazakhstan, Almaty Region, W-Zailiyskiy Alatau, Turgen distr., Tauturgen vill., h~1200 m, 8-13.VI.2002, A.Puchkov leg. L=19 mm
Copyright © Ruslan Panin

Carabus (Cryptocarabus) sacarum Kabak, 1998
Male adult
SE-Kazakhstan, Almaty reg., C-Zailiyskiy Alatau N-slopes, Kaskalen river Valley, Izvestkovy vill., h=1700 m, 6.V.2015, D.Kovalchuk leg./det. R.Panin L=18 mm
Copyright © Ruslan Panin

Carabus (Cryptocarabus) sacarum Kabak, 1998
Female adult
SE-Kazakhstan, Almaty reg., C-Zailiyskiy Alatau N-slopes, Kaskalen river Valley, Izvestkovy vill., h=1700 m, 6.V.2015, D.Kovalchuk leg./det. R.Panin L=20 mm
Copyright © Ruslan Panin

Carabus (Ctenocarabus) galicianus Gory, 1839
Male adult
Spain,
Sierra de la Cabrera Baja, 8 km SW Truchillas, 9.VII.2008, Anichtchenko A. leg.
L= 20 mm
Copyright © Alexander Anichtchenko

Carabus (Ctenocarabus) galicianus Gory, 1839
Female adult
NW-Spain, Pontevedra prov., Campo Lameiro env., 15.VIII.1994, S.Sobrino leg.
L=28 mm
Copyright © Ruslan Panin

Carabus (Ctenocarabus) melancholicus costatus Germar, 1824
Female adult
C-Spain, Madrid region, Rozas de Puerto Real env.,21.XII.2008, T.Gazurek leg.
L=27 mm
Copyright © Ruslan Panin

Carabus (Ctenocarabus) melancholicus melancholicus Fabricius, 1798
Male adult
NW Morocco, Kénitra, Souk El Arbaa env., II.1985, V.Olivella leg. L=25 mm
Copyright © Ruslan Panin

Carabus (Ctenocarabus) melancholicus submeridionalis Breuning, 1975
Male adult
S- Spain, Cádiz, Tarifa, El Cabrito, 28.XII.2019, F. Garcia leg. L=26 mm
Copyright © Ruslan Panin

Carabus (Ctenocarabus) melancholicus costatus Germar, 1824
Male adult
NW Portugal, Viana do Castelo distr., Sierra de Agra, 41°47'52.3"N 8°41'27.6"W, h=786 m, 3.VII.2016, D. Prunier leg.
Copyright © Ruslan Panin

Carabus (Cupreocarabus) kangdingi kangdingi Korell, Kleinfeld & Gorgner, 1992 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Cupreocarabus) brezinai Deuve, 1994
Male adult

Copyright © Boleslav Brezina

Carabus (Cupreocarabus) huangi huangi Deuve, 1992
Male adult
C-China, E Tibet, N-Sichuan, Hongyuan,valley 10 km SE Sanggarpar, h=4200m, 32.300053, 102.584031, alpine meadow / scree, 19.VII.1995, B.Brezina leg.<br /> L=21 mm
Copyright © Ruslan Panin

Carabus (Cupreocarabus) lixianensis lixianensis Deuve, 1990
Male adult
China, E Tibet, N-Sichuan, (Barkam),valley SW of Barkam, h=3200m, 31*53’ N; 102*12’E, mixed/coniferous forest, 22.VII.1995, K.& B.Brezina
L=23 mm
Copyright © Ruslan Panin

Carabus (Cupreocarabus) mollardianus Deuve, 2016
Female adult
Chine, Sichuan, Lushan, Mt. Jiajin Shan, 10.VII.2016 local collector leg./ det. R.Panin, L=22 mm
Copyright © Ruslan Panin

Carabus (Cupreocarabus) ? siguniangensis jiachingensis Lassalle, 2003
Male adult
Chine, Sichuan, Lushan, Mt. Jiajin Shan, 10.VII.2016 local collector leg., L=21 mm
Copyright © Ruslan Panin

Carabus (Cupreocarabus) balangicus balangicus Cavazzuti, 2002
Male adult
Chine, C-Sichuan, Xiaojin Co, Wolong valley, Balang Shan Pass h=3448 m, 30.884617, 102.973117, 9.VI.-1.VII.2011, G. Proscia leg. L=19 mm
Copyright © Ruslan Panin

Carabus (Cupreocarabus) balangicus balangicus Cavazzuti, 2002
Male adult
Chine, C-Sichuan, Xiaojin Co, Wolong valley, Balang Shan Pass h=3448 m, 30.884617, 102.973117, 9.VI.-1.VII.2011, G. Proscia leg. L=20 mm
Copyright © Ruslan Panin

Carabus (Cupreocarabus) huangi huangi Deuve, 1992
Male adult
China, E Tibet, N-Sichuan, 53 km NE of Barkam, Pass Sandaoban env., h=4293 m, 32.232950, 102.598533, 11-29.VI.2011, G. Proscia leg., L=21 mm
Copyright © Ruslan Panin

Carabus (Cupreocarabus) huangi huangi Deuve, 1992
Female adult
China, E Tibet, N-Sichuan, 53 km NE of Barkam, Pass Sandaoban env., h=4293 m, 32.232950, 102.598533, 11-29.VI.2011, G. Proscia leg., L=21.5 mm
Copyright © Ruslan Panin

Carabus (Cychrostomus) pseudoprosodes Semenov & Znojko, 1932
Male adult
male, 19mm
Copyright © Boleslav Brezina

Carabus (Cychrostomus) anchocephalus lanzhouensis Deuve, 1991 Paratype
Male adult
male, 21mm
Copyright © Boleslav Brezina

Carabus (Cychrostomus) anchocephalus anchocephalus Reitter, 1896
Male adult
male, 22mm
Copyright © Boleslav Brezina

Carabus (Cyclocarabus) aulacocnemus galinae Obydov, 2003 Paratype
Male+Female adult
W-Tianshan:Karatau Mts.,N-part:Boialdyr River valley,
N of Ken-su, 600-750m, 22-V-2000 М. Danilevsky leg.
male - 20 мм, female - 19 мм.
Copyright © Alexandr Safronov

Carabus (Cyclocarabus) namanganensis namanganensis Heyden, 1886
Male+Female adult
Kazakhstan, S. Karatau Mts. R., Mashat vill. env.
23-IV-1993 М. Danilevsky leg.
male - 21 мм, female - 22 мм.
Copyright © Alexandr Safronov

Carabus (Cyclocarabus) zarudnyi Semenov & Znojko, 1932
Female adult
NE-Uzbekistan, W-Tian-Shan, NW slopes Pskem Mt. Rng, 8-14 km E Pskem, 2100-2500 m, 16.VI.1998, A. Puchkov leg., L=21 mm
Copyright © Ruslan Panin

Carabus (Cyclocarabus) karaterekensis hemicallisthenes Kabak & Kalab, 1997
Male adult
NW- Kyrgyzstan, W-Tian-Shan, Chandalash Mt.Rng, Chakmak-Suu vill., 3.VI.2000, L=17 mm
Copyright © Ruslan Panin

Carabus (Cyclocarabus) brosciformis Semenov, 1896
Male adult
NE-Uzbekistan, W-Tian-Shan, Chatkal Mt Range, Tashkent reg., Yangiabad env., h=2100m, 30.IV.2002, L=23.5 mm
Copyright © Ruslan Panin

Carabus (Damaster) blaptoides oxuroides Schaum, 1862
Female adult
Japan, C-Honshu, Ofuse-cho, Kamitakai, Nagano Pref., 12.VI.2006, Ohshima leg.
L=34 mm
Copyright © Ruslan Panin

Carabus (Damaster) blaptoides capito Lewis, 1881
Male adult
Japan: Niigato pref., Sado Isl., Sado, Yamato
Copyright © Igor Solodovnikov

Carabus (Damaster) blaptoides oxuroides Schaum, 1862
Female adult
Japan, C Honshu, Nagano Prefecture, Nagano city, 3.IV.2012, S. Shinichi leg. L=42 mm. det. Anichtchenko A. 2021
Copyright © Ruslan Panin

Carabus (Damaster) blaptoides rugipennis Motschulsky, 1861
Male adult
Kunashir isl.
Copyright © Kirill Makarov

Carabus (Damaster) blaptoides blaptoides (Kollar, 1836)
Female adult
Japan, SW Honshu, Shimane Pref., Unnan city, 16.I2016, S.Shinichi leg., L=48 mm
Copyright © Ruslan Panin

Carabus (Deroplectes) staudingeri Ganglbauer, 1886
Female adult
N Tajikistan, SW Pamir-Alay, Fann Mts h=1800m, Padrut vill., 2-3.V.1997
L=28 mm
Copyright © Ruslan Panin

Carabus (Deroplectes) arcanus arcanus Semenov, 1898
Male adult

Copyright © Boleslav Brezina

Carabus (Deroplectes) dokhtouroffi dokhtouroffi Ganglbauer, 1886
Male adult
NC Tadzhikistan, Pamir, N-slope Peter I.Mt Rng, h=2100m,Ganishou vill., 10-12.VI.2006, Ivanova E. leg.
L=28 mm
Copyright © Ruslan Panin

Carabus (Deroplectes) klapperichianus coiffaitianus Deuve, 1990
Male adult
Tadzhikistan: Badakhshan: Ishkashim Mt.R.: Kuh-i-Lal, VI.1979 leg. I.K. Lopatin (TOPOTYPUS)
Copyright © Igor Solodovnikov

Carabus (Deroplectes) klapperichianus coiffaitianus Deuve, 1990
Female adult
Tadzhikistan: Badakhshan: Ishkashim Mt.R.: Kuh-i-Lal, VI.1979 leg. I.K. Lopatin (TOPOTYPUS)
Copyright © Igor Solodovnikov

Carabus (Deroplectes) klapperichianus salangicus Kleinfeld & Reuter, 2010 Paratype
Female adult
N-Afghanistan, Hindukush Mt.Rng, Salang Pass, Nordseite, 25°27'.494N, 68°58.399'E, ca. reg. subalp., h = 2200-2400 m, V.2010, leg. Christoph Reuter
Copyright © Igor Solodovnikov

Carabus (Diocarabus) loschnikovi loschnikovi Fischer, 1823
Male adult
Komi Resp., Pecero-Ilych Nat. Res., Zajachij Kamen h~786 62°10'06" N 59°12'53" E 8.VIII.2004 M. Chernjakhovski leg.
L=15.0 mm
Copyright © Evgeniy Zinoviev

Carabus (Diocarabus) lucepunctus ignipunctatus Rapuzzi, 2009 PT Paratype
Male+Female adult
China, SW Hunan Prov., Chengbu Co, Nan-Shan-Ding (26.34;110,08)1-10.VIII,2006, I.Rapuzzi leg.
femele, 18 mm
Copyright © Ruslan Panin

Carabus (Diocarabus) loschnikovi tsaganensis Obydov, 2001
Male adult
Russia, W Tuva,
Kyzyl-Tayga Mt Range h=1500 m, Ak-Suk riv vall., 21.VI.2006, A.& N. Menstschikovy leg.
L=16 mm
Copyright © Ruslan Panin

Carabus (Diocarabus) beybienkoi Kryzhanovskij, 1973
Male adult
S Sakhalin, Susunaj Mt.R. 10.08 - 13.09.2003 D. Kochetkov leg.
L=14,5 mm
Copyright © Maksim Smirnov

Carabus (Diocarabus) fraterculus fraterculus Reitter, 1895
Male adult
N-Korea, N Pyongan Prov.,Sakju, Mt Ponsan, 20-29.VI.2009, L. Bieber leg.
L=19 mm
Copyright © Ruslan Panin

Carabus (Diocarabus) aurocinctus Motschulsky, 1846
Female adult
det. K. Makarov
Copyright © Alexander Anichtchenko

Carabus (Diocarabus) opaculus opaculus Putzeys, 1875
Female adult

Copyright © Kirill Makarov

Carabus (Diocarabus) massagetus massagetus Motschulsky, 1846 (alpin form)
Male adult
Russsia, Altay, Ukok upland, Kaldzhin-Kul Lake h=2400-2550 m, 49 *19*N/87*26*E, 16-27.VII.2008, R.Dudko leg. L=17.5 mm
Copyright © Ruslan Panin

Carabus (Diocarabus) fraterculus neochinensis Deuve & Li, 1998
Female adult
NE-China: Liaoning prov., Fengcheng City, Shicheng, 20-28.VI.2011, Li Jingke leg., L=22 mm
Copyright © Ruslan Panin

Carabus (Diocarabus) fraterculus neochinensis Deuve & Li, 1998
Male adult
NE-China: Liaoning prov., Fengcheng City, Shicheng, 20-28.VI.2011, Li Jingke leg., L=20 mm
Copyright © Ruslan Panin

Carabus (Lasiocoptolabrus) sunwukong sunwukong Imura, 1993 Paratype
Male adult

Copyright © Boleslav Brezina

Carabus (Lasiocoptolabrus) sunwukong baotianmanus (Imura, 2005) Paratype
Male adult

Copyright © Boleslav Brezina

Carabus (Eccoptolabrus) exiguus nivium Breuning, 1933
Female adult
female, 20mm
Copyright © Boleslav Brezina

Carabus (Eccoptolabrus) exiguus exiguus Semenov, 1898 Holotype
Male adult
male, 18 mm
Copyright © Frank Kleinfeld

Carabus (Eucarabus) obsoletus carpathicus Palliardi, 1825
Male+Female adult
female, 25 mm
SW-Romania, Banat, Resita - Valiug, 3-8.VI.2007, Krause R. leg.
Copyright © Ruslan Panin

Carabus (Eucarabus) manifestus pyonganicola Deuve & Li, 1998
Male+Female adult
N-Korea: Pyonganam-Do, YangTok County,(39;127), 12-19,VI.2006
Copyright © Ruslan Panin

Carabus (Eucarabus) obsoletus nagyagensis Seidlitz, 1888
Female adult
NW-Romania, Mts Ses (Plopis),Cluj Reg., Bucea env, h=580 m, 8.VI.2011, Sl.Budka leg.
L=26 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) ulrichi rhilensis (as stussineri Géhin, 1885)
Male adult
W Croatia, Ivanščica Mts , Ivanec, 24.VI-7.VII.2006, Krause Rene leg.
L=29 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) ulrichi arrogans Schaum, 1858
Male adult
NE-Serbia:Majdanpek, 9-11,V.2010, L.Brhel leg.
L=28 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) ulrichi arrogans Schaum, 1858
Male adult
NE Serbia, Majdanpek; 23 mm
Copyright © Aleš Sedláček

Carabus (Eucarabus) ulrichi arrogans Schaum, 1858
Female adult
NE Serbia, Majdanpek; 29 mm
Copyright © Aleš Sedláček

Carabus (Eucarabus) cartereti omisanensis Lassalle, 1999
Male adult
South Korea, Gangwon Province, Kyebang-san, h=1100m, VI.1998 L=24 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) obsoletus obsoletus (as csikii Mallasz, 1901)
Female adult
NE Carpathians, Marmarosh Mts., Transcarphatian Reg., near Lug vill., h=350 m, 26.IV-8.VI.2014, R.Panin & Yu. Kanarsky leg. L=30 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) catenulatus fontanellae Reitter, 1896
Male adult
W-Croatia, Dalmatia, Isl. Rab, 16-30.V.2019, G. Proscia leg.
Copyright © Ruslan Panin

Carabus (Euleptocarabus) porrecticollis porrecticollis Bates, 1883
Male adult
Japan: Niigata pref., Chayagahara Jyouetsu City,
18.XI.2006 A.Toba leg.
Copyright © Alexander Anichtchenko

Carabus (Euleptocarabus) porrecticollis Bates, 1883
Female adult
Atsushiokanomachikano, Kitakata City, Fukushima Pref., Japan, 29.III.2014, T. Nakamura leg.
Copyright © Alexander Anichtchenko

Carabus (Euleptocarabus) porrecticollis porrecticollis Bates, 1883
Male adult
Japan, C Honshu, Nagano Pre., Toyooka town, 28.XII.2019, S. Shinichi leg. L=30 mm
Copyright © Ruslan Panin

Carabus (Eupachys) glyptopterus Fischer, 1827
Male+Female adult
Russia, SE-Siberia,
Buryat Republic, Ivolginsky Distr., 04.VII.2010, Filippov A.E. leg.
female, 32 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Eurycarabus) famini lucasi Gaubil, 1849
Male adult
N-Tunisia, Bizerte reg., Menzel Bourguiba city,h=5 m, XII.1981, S. Remond leg.
L=25 mm
Copyright © Ruslan Panin

Carabus (Eurycarabus) famini fezzanus Lapouge, 1915
Male adult
Morocco, Middle Atlas Mt Rng, Azrou city h=1500m, IV.1999, S. Remond leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Eurycarabus) famini piraticus (as mamorensis (Kocher, 1964)
Female adult
NW-Morocco, betw.Meknes/Kenitra, Foret de Mamora, II.2008
L= 27 mm
Copyright © Ruslan Panin

Carabus (Eurycarabus) famini atlantis Alluaud, 1923
Female adult
C Morocco, Haut Atlas, SW of Oukaïmeden h=2900m, 31.173644, -7.844442, 29.IV-11.V.2024, C.Reuter leg., L=21 mm
Copyright © Ruslan Panin

Carabus (Fulgenticarabus) flutschi koreus Deuve, 2006
Male adult
N Korea, Samjiyon, Ryang Gang Province
6-12.6.2006 leg. Wong
Copyright © Boleslav Brezina

Carabus (Gnathocarabus) kusnetzovi Semenov, 1903
Female adult
female, 21mm
Copyright © Boleslav Brezina

Carabus (Goniocarabus) sogdianus Semenov, 1898
Male adult
male, 21mm
Copyright © Boleslav Brezina

Carabus (Goniocarabus) sogdianus Semenov, 1898
Male adult
N Tajikistan, SW Pamir-Alay, Zarafshan Range, Fann Mts, Marguzor Lake circ., Padrut vill., 16.V.1997
L=21 mm
Copyright © Ruslan Panin

Carabus (Goniocarabus) grombczewskii grombczewskii Semenov, 1891
Male adult
NC Tadzhikistan, Pamir, N-slope Peter I.Mt Rng, h=2100m, Ganishou vill.,12.VII.2006, Pak O. leg.
L=19 mm
Copyright © Ruslan Panin

Carabus (Goniocarabus) perelloi Casale, 1979
Male adult
N-Afghanistan, Hindukush, Provinz Baghlan, N of Kabul, N Salang Pass, subalp., h = 2200 m, 35°27'.494N / 68°58.399'E, Mai 2011, leg. Christoph Reuter (TOPOTYPUS)
Copyright © Igor Solodovnikov

Carabus (Hemicarabus) tuberculosus Dejean, 1829
Male adult
Far East, Lazovsky Dist., shore of Japan Sea, 3 km NE Preobrazhenie Vill., Ezhovaya (Tasovaya) Bay 13.07.06 M. & L. Smirnov's leg.
male
L=20,0 mm
Copyright © Maksim Smirnov

Carabus (Hemicarabus) nitens Linne, 1758
Male+Female adult
Belarus,
Vitebsk reg., Ushachskiy distr., near Bolshie Dolcy vill. 9.V.1990, I.A.Solodovnikov leg.
female, 18mm.
Copyright © Ruslan Panin

Carabus (Hemicarabus) serratus Say, 1825
Male adult
E-USA,Virginia, Leesburg env., Ball's Bluff Battlefield Regional Park, 27.VIII-4.IX.2015,V. Čutka leg. L=17 mm
Copyright © Ruslan Panin

Carabus (Hemicarabus) serratus Say, 1825
Female adult
E-USA, Virginia, Leesburg env., Ball's Bluff Battlefield Regional Park, 27.VIII-4.IX.2015,V. Čutka leg. L=18 mm
Copyright © Ruslan Panin

Carabus (Heterocarabus) marietti marietti (as ativsengueni Schweiger, 1962)
Male adult
male, 17mm
Copyright © Boleslav Brezina

Carabus (Heterocarabus) marietti marietti Cristoforis & Jan, 1837
Female adult
as ativsengueni Schweiger, 1962
NW Turkey, Bolu, Abant Mts.; 16mm

Copyright © Aleš Sedláček

Carabus (Homoeocarabus) maeander maeander s.str.
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Carabus (Homoeocarabus) maeander paludis Gehin, 1885
Female adult
Japan, Hokkaido; Tomakomai city, Benten, 17.V.2015, Takemoto leg., L=21 mm
Copyright © Ruslan Panin

Carabus (Hygrocarabus) nodulosus Creutzer, 1799
Male+Female adult
Austria, Styria Region (Steiermark), Leibnitz, 31.X.1999, I.Rapuzzi leg.<br /> female, 29 mm
Copyright © Ruslan Panin

Carabus (Hygrocarabus) variolosus Fabricius, 1787
Male adult
W-Ukraine, NW Podolian,<br /> Lviv reg., Zhydachiv distr.,near Vybranivka vill., 15.V-8.VI.2008, Panin R. leg.<br /> male, 27mm
Copyright © Ruslan Panin

Carabus (Hygrocarabus) variolosus Fabricius, 1787
Male+Female adult
W-Ukraine,<br /> Lviv Reg., Zhydachiv distr., near Vybranivka vill., Panin R. leg.<br /> female, 30mm
Copyright © Ruslan Panin

Carabus (Imaibius) baronii battalensis Heinz & Staven, 1998 Paratype
Male adult

Copyright © Alexander Anichtchenko

Carabus (Imaibius) dardiellus perobscurus Deuve, 1980
Male adult
N India.Kashmir, Loran, W part Pir Panchal range,h=2400-2500m, 33°49'"N, 74°20'E
L=25 mm
Copyright © Ruslan Panin

Carabus (Imaibius) dardiellus ? subpunctulus Roeschke, 1907
Male adult
N India, Kashmir, Pahalgam, Aru Valley h=2400m, VII.2007
L=25 mm
Copyright © Ruslan Panin

Carabus (Imaibius) wittmerorum Heinertz, 1978
Male adult
N. Pakistan: Frontier prov., Miandam, Swat vall., h=1800-2600 m, leg.S.Remond (TOPOTYPUS)
Copyright © Igor Solodovnikov

Carabus (Imaibius) barysomus hazarensis Deuve, 1982 (TOPOTYP)
Male adult
N-Pakistan: Mansehra prov., Hazara, Kaghan valley: Shocgran env., h = 2750 m L= 42 mm
Copyright © Igor Solodovnikov

Carabus (Imaibius) boysi boysi Tatum, 1851
Male adult
NW-India, Himachal Pradesh, Manikaran env., Pulga vill., 4.VII.2008. L=26 mm
Copyright © Ruslan Panin

Carabus (Imaibius) stoliczkanus brauni Heinz, 1993
Female adult
NE Pakistan, Azad Kashmir, Bagh Distr., Sudhan-Gali h~2300 m,15.VIII.1994, S Dacatra leg. L=37 mm
Copyright © Ruslan Panin

Carabus (Imaibius) caschmirensis fortetuberculatus Heinz, 1983 Paratype
Male adult

Copyright © Alexander Anichtchenko

Carabus (Imaibius) nouristani nouristani (Ledoux, 1977)
Female adult
NE-Afghanistan: Nuristan: Darah-I-Nur valley, 2900m 34°45'N 70°40'E, VIII.2012, C.Reuter leg., L=26 mm
Copyright © Ruslan Panin

Carabus (Imaibius) nouristani nouristani (Ledoux, 1977)
Male adult
NE-Afghanistan: Nuristan: Darah-I-Nur valley, 2900m 34°45'N 70°40'E, VIII.2012, C.Reuter leg., L=26 mm
Copyright © Ruslan Panin

Carabus (Iniopachus) pyrenaeus jeanneli Deuve, 1994
Male adult
1-2: Pirineos, Canfranc, Ibon de Iserias 2200 m 16-17.7.05<br /> 3: Pirineos, Monte Perdido, Collata Anisolo 2400m 25.8.05
Copyright © Alexander Anichtchenko

Carabus (Iniopachus) auriculatus ubinensis (Puissegur, 1955)
Female adult
Spain, Montes Cantabricos, Peña Ubiña, 2400m
5-6.VII.2009 Anichtchenko A. leg.
female, 19 mm
Copyright © Alexander Anichtchenko

Carabus (Iniopachus) pyrenaeus pyrenaeus (as cephalicus Csiki, 1927)
Male adult
SW France, Pyrenees-Atlantiques, Mt.Cirque d'Esturou h=1800m, 29.VIII.2014, Pascal Stefani leg. L=20 mm
Copyright © Ruslan Panin

Carabus (Ischnocarabus) tenuitarsis tenuitarsis Kraatz, 1877
Male adult
male, 24mm
Copyright © Boleslav Brezina

Carabus (Isiocarabus) fiduciarius fiduciarius (as fiduciarius tim) Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Isiocarabus) hunanicola liangshanicus (as dreuxi liangshanicus) HT PT Kleinfeld, 1998 Holotype Paratype
Male+Female adult
Carabus (Isiocarabus) hunanicola liangshanicus (as dreuxi liangshanicus) HT PT
Copyright © Frank Kleinfeld

Carabus (Isiocarabus) miaorum Lassalle & Prunier, 1993
Male adult
S-China, N-Guangxi, Mao'er Shan, h~1600 m, V.2009, L= 26 mm
Copyright © Ruslan Panin

Carabus (Isiocarabus) fiduciarius fiduciarius Thomson, 1856
Female adult
CE-China, Hubei, Xiaogan City, Shuangfeng Mountain, 26.VI.2016, L=30 mm
Copyright © Ruslan Panin

Carabus (Isiocarabus) miaorum Lassalle & Prunier, 1993
Male adult
China, NW-Guangxi, Mao'er Shan, h~1600 m 25°52'N 110°29'E, 28.V-9.VI.2009, C.Reuter leg.
Copyright © Ruslan Panin

Carabus (Lamprostus) calleyi pseudoprasinus Lapouge, 1912
Male+Female adult
Transcaucasia, Armenia,
Ashotsk distr. Tsoghamarg vill, .21-22.V.1997, P.V.Kazaryan leg
female, 24 mm
Copyright © Ruslan Panin

Carabus (Lamprostus) robustus ducalis Schaufuss, 1882
Male adult
NE-Turkey:Trabzon, Zigana Pass h=2000m,14-24.VI.2010, V. Slovák leg.
L=32 mm
Copyright © Ruslan Panin

Carabus (Lamprostus) torosus torosus Frivaldsky, 1835
Male adult
NE-Bulgaria, Dobrich Region, Kavarna env.
L=31 mm
Copyright © Ruslan Panin

Carabus (Lamprostus) syrus pseudopinguis Heinz & Staven, 2000
Male adult
NW-Jordan, Ajlun env., oak forests on dry limestone, h=897m, 32°21′33.1″N/ 035°43’47.4″E, 4.V.2010, Schnitter/Schellhorn leg.
L=26 mm
Copyright © Ruslan Panin

Carabus (Lamprostus) syrus cheikensis Deuve, 1992
Male adult
N-Israel, Midlle Galilea, Nazareth env., Hoshaya viil., 13.XII.2012, Vas’ko, leg./det. Panin R. L=30мм
Copyright © Ruslan Panin

Carabus (Lamprostus) torosus giresuni Breuning, 1964 (as paululus Heinz & Korge, 1964)
Male adult
NE-Turkey, Giresun, Sehitler Geçidi (Pass) h~2000 m,24.VII.2001, S. Dacatra leg., L=26 mm
Copyright © Ruslan Panin

Carabus (Lamprostus) torosus nordmanni Chaudoir, 1848 (as enricoi )
Male adult
NE-Turkey:Ordu prov.,3 km N Gölköy h~1100 m, VII.2006, L=37 mm
Copyright © Ruslan Panin

Carabus (Lamprostus) calleyi pseudoprasinus Lapouge, 1912
Male adult
S Georgia, Tsalka (Paravani) 41°29′18.7″N, 43°51′12.3″E, 8-9.V.2016, Pavel Turek leg. / det. I. Solodovnikov, L=23 mm
Copyright © Ruslan Panin

Carabus (Lamprostus) syrus moukarzelae Deuve, 2002
Female adult
S-Lebanon: Marjaayoun, 18.XI.-11.XII.2017, C. Reuter leg., L=35 mm
Copyright © Ruslan Panin

Carabus (Lamprostus) syrus cheikensis Deuve, 1992
Female adult
SE Lebanon, Rashaya Distr., Tannoura, oak forest, h=700 m, 11-19.IV.2016, C. Reuter leg., L= 32 mm
Copyright © Ruslan Panin

Carabus (Leptocarabus) yokoae marcilhaci Deuve, 1992
Male adult
CHINA, Shanxi, Tschingling Mts., Taibeichan Nat. reserv.;
Copyright © Anton Menstschikov

Carabus (Leptocarabus) harmandi fujisan Ishikawa, 1972
Female adult
Japan, Yamanashi pref., Fuji-Rinds 1600m, Narusawa vill. 3.V.2005 Kurihara K. leg. 25 mm
Copyright © Alexander Anichtchenko

Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931
Male adult
NE-China:Liaoning prov., Fengcheng City, Miaoyang, 12-17.VI.2011
L=26 mm
Copyright © Ruslan Panin

Carabus (Leptocarabus) procerulus Chaudoir, 1862
Male adult
Japan:S Honshu, Matsubara-ko, VIII.1997, S. Remond leg.
L=30 mm
Copyright © Ruslan Panin

Carabus (Leptocarabus) procerulus Chaudoir, 1862
Male adult
Japan, S Honshu, Mie Pref., Hakusan, VII.2001, S. Remond leg.<br /> L=33 mm
Copyright © Ruslan Panin

Carabus (Leptocarabus) arboreus pararboreus (Ishikawa, 1992)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Carabus (Leptocarabus) arboreus fujisanus Bates, 1883
Male adult
Japan, CE-Honshu, Yamanashi pref., Mt. Fujisan h~1600m, 23.IX.1982, Y. Imura leg. L=23 mm
Copyright © Ruslan Panin

Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931
Male adult
NE-China:E-Liaoning: NP Kuandian Tianqiaogou , VI.2010, L=25 mm, det. Panin R.
Copyright © Ruslan Panin

Carabus (Leptocarabus) arboreus fujisanus Bates, 1883
Female adult
Japan, Honshu, Mt. Fuji h~1900m, Shizuoka pref., Fujinomiya city, 23.VI.2007, M. Matsunaga leg., L=25 mm
Copyright © Ruslan Panin

Carabus (Leptoplesius) merzbacheri kirgisiensis Mandl, 1967
Female adult
22mm
Copyright © Boleslav Brezina

Carabus (Limnocarabus) clathratus clathratus Linne, 1761
Female adult
W-Ukraine, Maloe Polesie, Lviv Region, near Kamenka-Bugzkaya, 28.III.2008, Panin R. leg. L=31 mm
Copyright © Ruslan Panin

Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
Female adult
SW Slovakia, Bratislava env., IV.2004, Krause Rene leg.
L=32 mm
Copyright © Ruslan Panin

Carabus (Limnocarabus) clathratus clathratus (as jansoni Kraatz, 1890)
Male adult
W-Ireland, County Mayo, Bellacorick bog, 9.VII.2013, J.Stypula leg. L=27 mm
Copyright © Ruslan Panin

Carabus (Limnocarabus) clathratus clathratus Linne, 1761
Female adult

Copyright © Kirill Makarov

Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
Male adult
W Ukraine, Transcarpathian Reg., 4 km NE Chop, forest Peresh, Panin R. leg. L=30 mm
Copyright © Ruslan Panin

Carabus (Lipaster) stjernvalli stjernvalli Mannerheim, 1830
Male+Female adult
Armenia,
Ashotsk, 22.VI.1990, P.V.Kazaryan leg.
female 34mm
Copyright © Ruslan Panin

Carabus (Lipaster) stjernvalli osellai Breuning & Ruspoli, 1970
Male+Female adult
NE-Turkey: Artvin:Hopa, Borçka env., h=750 m, 9.VI-7.VII.2011, M.Janalik leg.
female, 42 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) morbillosus alternans Palliardi, 1825
Male adult
S Italy, Sicily, Province Palermo, Piana degli Albanesi h~700m, XI.2010, I.Rapuzzi leg.
L=32 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) aumonti maroccanus Bedel, 1895
Male adult
N Morocco (Spain:Melilla), hipodromo, X.1969, leg. Pardo Alcaide
L=38 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) rugosus boeticus Deyrolle, 1852
Male adult
SC Spain, Jaen reg., Puente de la Sierra, 10.VII.2011, M.Lopez leg.
L=28 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) morbillosus constantinus Kraatz, 1899
Male adult
NW Tunisia; Tebarka, E Sejenane,26.I.2004, Manfred Egger leg.
L=33 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) morbillosus macilentus Lapouge, 1899
Female adult
SE Spain, Murcia reg., P.N. de Calblanque, 1.V.2003, T.Gazurek leg.
L=34 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) planatus Chaudoir, 1843
Male adult
S Italy, N Sicily, Nebrodi; 33 mm
Copyright © Aleš Sedláček

Carabus (Macrothorax) morbillosus morbillosus Fabricius, 1792 (as cheminorum Deuve, 1988)
Female adult
SE-France, Var, Fréjus env., 26.II.1997,
L=31 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) morbillosus lampedusae Born, 1925
Male adult
S Italy, Isl. Lampedusa, Baia dei Conigli, VI.2003, L=29 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) rugosus levantinus Lauffer, 1905
Male adult
SE-Spain, Murcia, Jumillla, Sierra del Carche, 6.VII.2019, F. Garcia leg., L=29 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) planatus Chaudoir, 1843
Female adult
S Italy, N Sicily, Nebrodi Mts, Portella di Fémmina Morta h=1500 m, 2.IX.1989, V.Aliquo leg. L=33 mm
Copyright © Ruslan Panin

Carabus (Meganebrius) arunensis Heinertz, 1980
Female adult

Copyright © Boleslav Brezina

Carabus (Meganebrius) kadoudali (Morvan, 1981)
Male+Female adult
C-Nepal, SW Dhaulagiri, W Jaljala h=3150 m, 28°30'05"N, 83°11'48"E, 23.IX.2012, J. Schmidt leg.
female, 20 mm
Copyright © Ruslan Panin

Carabus (Meganebrius) pseudoharmandi Mandl, 1965
Female adult
E-Nepal, Kosi zone, Khandbari,(1.5km SW of Chichila vill., 27*45’843’’N/ 87*22’473’E, h=1940m), damaged primary forest, 13 .V.2010, B.Brezina & M.Häckel leg.
L=23 mm

Copyright © Ruslan Panin

Carabus (Meganebrius) arunensis Heinertz, 1980
Male adult
E-Nepal, Kosi zone, Khandbari, 1.5km SW of Chichila vill., 27*47’175’’N/ 87*23’622’E, h=1880m, damaged primary forest, 11.V.2010, B.Brezina & M.Häckel leg.
L=27 mm

Copyright © Ruslan Panin

Carabus (Meganebrius) quinlani smetanai Häckel, 2011 Paratype
Male+Female adult
C-Nepal:Dhaulagiri Massif:Baglung Distr., (1.5km W of Okhale vill., 28.32’152’’N/ 83.48’673’E, h=2600m) damaged primary forest, 19.V.2010, B.Brezina & M.Häckel leg.
male, 21mm
female, 23 mm
Copyright © Ruslan Panin

Carabus (Meganebrius) quinlani quinlani Mandl, 1965
Male adult
C-Nepal, Gandaki, Manang Distr., Timang env., h~2500-2600m, old open forest, grassland, 3-5.V.2007, Fouqué R. leg. L=19 mm
Copyright © Ruslan Panin

Carabus (Meganebrius) tuberculipennis ganjalaensis Deuve, 2000
Male adult
Nepal, Helambu, GanjaLa Pass S-slope, Dhukpu Kharka h=4100m, 28°03'50"N, 85°34'21"E, 12.VIII.2010, S.Tamang leg., L=21 mm
Copyright © Ruslan Panin

Carabus (Pagocarabus) erwini erwini s.str.
Male adult

Copyright © Boleslav Brezina

Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823
Male adult
W-UKRAINE
NW Podolian, Lviv Reg., Zolochiv Distr., near Osovitsa, steppe sites, Panin R. leg. 1.V-6.VI.07
male, 26mm
Copyright © Safronov

Carabus (Megodontus) violaceus violaceus Linne, 1758
Male adult
S.Bohemia, Hluboka nad Vltavou, h=380 m, 6.VIII.2004, S. Budka leg.
L=28 mm
Copyright © Ruslan Panin

Carabus (Megodontus) caelatus schreiberi Kraatz, 1887
Male adult
SW Slovenia, Obrov, Poljane, 14-27.VIII.2010, Manfed Egger leg.
L=33 mm
Copyright © Ruslan Panin

Carabus (Megodontus) croaticus zepcensis (as bosiljevici Drovenik & Pavićević, 1985)
Male adult
CW-Serbia, Zlatar Mts, Nova Varoš, Zlatar Lake env., 1-12.VII.2007, P.Sima leg. L=30 mm
Copyright © Ruslan Panin

Carabus (Megodontus) caelatus dalmatinus (as procerus Reitter, 1885)
Male adult
S Bosnia, Potoci env. h=500 m, VII.2006, Bartoloni leg.
L=38 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus purpurascens Fabricius, 1787 (as osismius Lapouge, 1922)
Male adult
W-FRANCE, Brittany , Finistère, Botvarec Forest, 19.XI.2006, Pascal Stefani leg.
L=29 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus violaceus trans violaceus purpurascens Fabricius,1787
Male+Female adult
E-Netherlands, Gelderland, Winterswijk Woold env.,15.II.2015, H.de Haan leg.
Copyright © Ruslan Panin

Carabus (Megodontus) persianus javahardehensis Deuve, 2000
Female adult
N-Iran, Mazandaran Prov., Ramsar Co., Javahar Deh env., 16-25.VI.2017, Václav Čutka leg. L=30 mm
Copyright © Ruslan Panin

Carabus (Megodontus) kolbei taikinus Ishikawa, 1993
Female adult
Japan, SC-Hokkaido:Tokachi, Hiroo Distr., Taiki-cho env., 3-7.IX.2019, S.Shinichi leg.
Copyright © Ruslan Panin

Carabus (Megodontus) leachi leachi Fischer, 1823
Male adult
NE Kazakhstan, W Altay, near Shemonaikha city., flood plain, 18-28.VII.2018, S.Vashchenko leg., L=21.5 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus lusitanicus Fabricius, 1801(as schaumi Gaubil, 1849)
Male+Female adult
Portugal,
Viseu distr., Gosende, h~1010m, 15.X.2009, D.Prunier leg.
female,26 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus harcyniae (as procedens Csiki, 1927)
Male adult
England, North Yorkshire, Fylingdales Moor, Grouse Hill, VII.2006 L=24 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus brevis (as sorianus Jeanne, 1972)
Male adult
NC-Spain, Soria, Puerto de Oncala, 2.VI.2006
L=24 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus islandicus Lindroth, 1968
Female adult
SE-Iceland:Höfn env.:Hornafjördur, 24-28.VIII.2008, C. Reuter leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) cantabricus cantabricus Chevrolat, 1840 (loc. typ.)
Male adult
NW-Spain, Pontevedra prov., A Cañiza viil., Alto de Fontefría h=800m, 31.XII.2007, T.Gazurek leg./det.Panin R.<br /> L=27 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus harcyniae (as procedens Csiki, 1927)
Female adult
N-Ireland, Fermanagh Distr., Lough Navar Forest, 17.VII.2013, D.Novak & J.Stypula leg.<br /> L=27mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus helluo Dejean, 1826
Female adult
SE-Spain, Prov.Alicante, Port de la Carrasqueta, 26.XII.1987, M.Toribio leg. L=25 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus gougeleti (as valderepisensis Branes, 1988)
Male adult
Spain, Jaen prov., Despeñaperros, Arroyo de los Arcos, 38.381847, -3.515651, 1-2.05.2014, Anichtchenko A. leg. & det.; 18 mm
Copyright © Alexander Anichtchenko

Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
Male adult
NC-Spain, Basque Co, Álava, Sierra de Gorbea, VI.2014, L=29 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus trabuccarius Fairmaire, 1857
Male adult
NE Spain, Catalonia, Tarragona, Sierra de Montsant h=800 m, Albarca env.,7.XI.2018, T. Rafales leg., L=25 mm
Copyright © Ruslan Panin

Carabus (Microplectes) riedeli Menetries, 1832
Female adult
female, 19mm
Copyright © Boleslav Brezina

Carabus (Microplectes) riedeli Menetries, 1832
Male adult
N-Ossetia,
Khazniaush Pass, h=2500-2700m, 9.VI-31.VII.1994, Zamotajlov A.leg.
male, 17mm
Copyright © Ruslan Panin

Carabus (Microplectes) convallium convallium Starck, 1889
Male adult
male, 19mm
Copyright © Boleslav Brezina

Carabus (Microplectes) convallium convallium Starck, 1889
Male+Female adult
female:NW-Caucasus, Russia,
Krasnodar reg., Apsheron Distr., Lagonaki Mt., near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominyh leg.
male:Karachaevo-Cherkessia,the right source of the river Beskess, h=2100m, 10.VIII.1997, I.A. Solodovnikov leg.
female, 21mm/male, 19mm
Copyright © Ruslan Panin

Carabus (Microplectes) convallium argonautarum Semenov, 1898
Male adult
male, 18mm
Copyright © Boleslav Brezina

Carabus (Mimocarabus) maurus maurus Adams, 1817
Male+Female adult
Armenia,
Ashotsk distr., Tsoghamarg vill., 24-27.V.1999 P.V.Kazaryan leg.
female, 20mm/male,18mm
Copyright © Ruslan Panin

Carabus (Mimocarabus) pumilio Kuster, 1846
Male adult
S-Armenia, Vayots Dzor Region, E Ughedzor vill., h=2630 m, 9.VI.2014, L=16 mm
Copyright © Ruslan Panin

Carabus (Mimocarabus) maurus maurus (Adams, 1817)
Male adult
E Georgia, Kakheti, near Patardzeuli vill., h=750-850 m, 10.IV.2015, E. Rutjan leg., L=22 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi incompsus (as mendax Csiki,1906)
Male+Female adult
W.Ukraine,
Transcarpathian Reg., Rakhov Distr., Chornogora Mt Range, h=1900m. alpine zone, Panin R. leg/det. 28.VI-9.VIII.2008
female, 32 mm
photo endophallus Solodovnikov I.A
Copyright © Ruslan Panin

Carabus (Morphocarabus) odoratus bargusinicus Shilenkov, 1996
Male+Female adult

    Carabus (Morphocarabus) odoratus bargusinicus Shilenkov, 1996
    Russia, N Burajatia,
    Barguzinskiy Mts., the upper reaches of M.Chiverkuy riv., h=2000m, Vashchenko S. leg.
    female, 21mm/male,19mm
    photo Panin R.

Copyright © Ruslan Panin

Carabus (Morphocarabus) michailovi Kabak, 1992
Male adult
NE-Kazakhstan,
S Altai Mts.,Sarym-Sakty Range, Sarym-Sakty riv., h=2400m. 21.VI.1999., A.Klimenko leg.
male,18mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
Male adult
Russia, S Siberia,
S Khakassia, N Tuva reg., Sailyg-Khem-Taiga mts. range h=2400m, 10.VII.2004
male, 18mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
Male adult
W Sayan Mts., Kazhyr-Suk riv., 80 km W Oleniya
Rechka, W part of Oiskii mt. rng., 30.6.-7.7.1999
2100 m, Brinev A. leg.
male, 19 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) rothi (? incompsus as spectabilis Csiki, 1906)
Female adult
NC Romania, Bistrita Nasaud reg., Călimani Mts, Colibita vill., mixed forest, meadows h=800m, 15-21.V.2011, M.Pasek leg.<br /> L=34 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi distinguendus Csiki, 1906
Male adult
NC-Hungary: E- part of Budapest, Cinkota,28.VIII-17.IX.1980, I.Retezár leg. L=27 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens (as lomnitzkii Reitter, 1896)
Male adult
W Ukraina, Podolian Upland, steppe reserve Kasova Gora near Burshtyn vill., 15.VI -28.VII.2020, R. Panin & D. Khrapov leg., L=24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Male adult
SE Poland, N Carpathians, Subcarpathia Prov., Nadleśnictwo Baligród, (209a) 22.VII.2021, M.Kosibowicz leg./det. R.Panin, L=31 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Male adult
SE Poland, N Carpathians, Subcarpathia Prov., National Park Magura, VII.2016, M.Ranocha leg./ det. R. Panin, L=30 mm
Copyright © Ruslan Panin

Carabus (Morvanicarabus) scheibei scheibei Eidam, 1937
Female adult
NE-Afghanistan, Hindu Kush, Nuristan Prov., near Parun, 35°19'N 70°52'E h=2900 m, Pine forest, VI.2022, local collector/det. Kleinfeld, L= 24 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) kratkyi felicitanus Reitter, 1893 (as porcellus)
Male adult
NW Caucasus, Resp. Adygea, Mt.Tybga, 28.VI.1997
male,20mm
Copyright © Ruslan Panin

Carabus (Neoplectes) kratkyi Ganglbauer, 1890
Male+Female adult
W Caucas Mts., 2500m, Mt. Yatyrgvarta,
25.VI-12.VIII.1992 Shchurov V. leg.
male 20mm, female 21mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) reitteri laevisternis Starck, 1890
Male adult
Abkhasia,W.slope of Gagrinsky Mt., Krepostnaya River source, h=1325m., A.S.Zamotajlov leg. 27.VII.2000
male, 29 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) reitteri laevisternis Starck, 1890
Male adult
W-Caucasus:Sochi env., Aibga Mt. h=2000m., 06.VII.2000
male, 29mm
Copyright © Ruslan Panin

Carabus (Neoplectes) satyrus napraensis Belousov & Zamotajlov, 1993 Paratype
Male adult
Abkhasia, Bzybian Mt. Range near Mt. Chibzharga., h=1900-2000m., A.S. Zamotajlov leg.
male, 28mm
photo & coll.Panin R.
Copyright © Ruslan Panin

Carabus (Neoplectes) satyrus duripshensis Kurnakov, 1972
Male+Female adult
W.Caucasus, Abkhasia, Gudauta distr.,
Duripsh village env., Khipsta valley, h=500-700 m.
07-12.VI.2009 A.Safronov leg.
Body lenght 30-33 mm.
Copyright © Alexandr Safronov

Carabus (Neoplectes) reitteri gagrinus Starck, 1894 (as gegicus Gottwald, 1985)
Male+Female adult
W.Caucasus, Abkhasia, Gagra distr., Gega riv. valley,
13-15.07.2005 D. Kasatkin. leg.
male - 30 mm, female - 34 mm.
Copyright © Alexandr Safronov

Carabus (Neoplectes) reitteri reitteri Retowski, 1885
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014 Paratype
Male adult
Abkhazia, N Slopes of Bzybian Mt. Range, right bank of River Reshevie (= Reshava) (left tributary of River Bzyb), 700-1000 m, 13.V-02.VII.2010, leg. D. Fominykh, I. Solodovnikov
Copyright © Igor Solodovnikov

Carabus (Neoplectes) protensus protensoides Novotny & Vorisek, 1988
Male adult
W Georgia, Martvili distr., W slope of Askhi Plateau h=1200-1800 m, 18-19.VI.2005, Shumov leg., L=33 mm
Copyright © Ruslan Panin

Carabus (Eocechenus) kaznakovi nangqenicus Deuve, 1995 Paratype
Male adult

Copyright © Boleslav Brezina

Carabus (Nesaeocarabus) abbreviatus Brulle, 1835
Male adult
Canarias Isle, NC Tenerife; 22mm
Copyright © Aleš Sedláček

Carabus (Nesaeocarabus) faustus cabrerai Enderlein, 1929
Male adult
Spain, Canarias Isl.,Teneriffe: NW-part:Teno Massif, Erjos env., h=1000m, 18-21.III.2013, V.Slovak leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Ohomopterus) insulicola insulicola Chaudoir, 1869
Male adult
Japan, Yamanashi pref., Kodachi, Kawaguchiko-cho, Minamitusru distr.
2.IX.2000 T. Kurihara leg.
male, 29 mm
Copyright © Alexander Anichtchenko

Carabus (Ohomopterus) insulicola insulicola Chaudoir, 1869
Female adult
15.VII.1998 JAPAN, Ichihara, env. Chiba
Copyright © Nil Belonogov

Carabus (Ohomopterus) lewisianus lewisianus Breuning, 1932
Male adult
Japan, Yamanashi pref., Koganesawa-rindo, Otsuki city,
13.6.1990 K. Kurihara leg.
male, 20 mm
Copyright © Alexander Anichtchenko

Carabus (Ohomopterus) albrechti albrechti Morawitz, 1862
Female adult
Japan, S-Hokkaido:Oshima Peninsula,Pass btw Otoshibe - Assabu, h~250-300 m, 10-20.VII.2010, 42:01’53’Т; 140:22’09’E, I.Rapuzzi & L.Caldon leg.
Copyright © Ruslan Panin

Carabus (Ohomopterus) japonicus japonicus Motschulsky, 1857
Male adult
S Japan, Kyushu, Kumamoto Pref., Izumi viil., h=600 m,VII.2001, S. Remond leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Ohomopterus) yamato kitai (Imura, 2004)
Male adult
Japan, S Honshu, Mie Pref., Hakusan, VII.2001, S. Remond leg.
L=21 mm
Copyright © Ruslan Panin

Carabus (Ohomopterus) dehaanii dehaanii Chaudoir, 1848
Female adult
Japan, S Honshu, Hyogo Pref., Nishiwaki city, 20.VI.2010, Shinichi S. leg., L=37 mm
Copyright © Ruslan Panin

Carabus (Ohomopterus) dehaanii dehaanii Chaudoir, 1848
Male adult
Japan, S Honshu, Mie Pref., Inabe city, 21.II.2021, Shinichi S. leg., L=33 mm
Copyright © Ruslan Panin

Carabus (Ophiocarabus) regeli obscurior Semenov, 1888
Male+Female adult
Kyrgyzstan, NE Tian-Shan, E. Terskey Alatau Mts. R.,
Kok-kia River, h=2800 м.
7-8.VI.1997 М. Danilevsky leg.
Body lenght 15-18 mm.
Copyright © Alexandr Safronov

Carabus (Ophiocarabus) ballionis Kraatz, 1879
Male adult
NW-China, Xinjiang prov., 40 km N Narat, h=1500, 12-12.VI.2004, Benes & Kaiser leg.
L=22 mm
Copyright © Ruslan Panin

Carabus (Ophiocarabus) successor koeksuensis Kabak, 2013 Paratype
Male adult
China, Xinjiang, Koeksu Bas., Tshon-Aidak, SSE Tekes, 42* 32' 47'' N / 82* 05' 56'' E -- 42* 32' 51'' N / 82* 05' 36'' E, 3330-3550 m, 11.07.2012, Kabak I.I. leg.
Copyright © Ilya Kabak

Carabus (Ophiocarabus) successor successor Reitter, 1896
Male adult
SE-Kazakhstan, Narynkol distr., 4 km SW Bayankol, h~2400-2500 m, 2-14.VI.2007, Shumov leg. L=17.5 mm
Copyright © Ruslan Panin

Carabus (Ophiocarabus) successor successor Reitter, 1896
Female adult
SE-Kazakhstan, Narynkol distr., 4 km SW Bayankol, h~2400-2500 m, 2-14.VI.2007, Shumov leg. L=17 mm
Copyright © Ruslan Panin

Carabus (Pachystus) hortensis neumeyeri Schaum, 1856
Female adult
Italy, Basilicata, Serra Calvello (PZ)
1400 m, 7.2003 F. Angelini leg.
female, 25 mm
Copyright © Alexander Anichtchenko

Carabus (Oreocarabus) luetgensi getschmanni Lapouge, 1924
Male adult
NW-Spain, León Province, Acebo, h=1000m, V.2007, Pascal Stefani leg.
L=22 mm
Copyright © Ruslan Panin

Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847
Male adult
WC Spain, Cáceres, Hervás, h=1000m, V.2007, Pascal Stefani leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (?Oreocarabus) qinlingensis Imura, 1993
Male adult
China, W-Shaanxi, Taibai Shan Mts, btw. Houzhenzi/Ba Xian Tai, nr.Nan Tian Men h=3200m, 33°55'17"N/107°46'53"E, 25-30.VII.2011, Li Wuyang leg.
L=19 mm
Copyright © Ruslan Panin

Carabus (Oreocarabus) latro minshanensis Deuve, 1987
Male adult
China, N-Sichuan, Jiuzhaigou env., 13.VI.2001,E. Kučera leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Oreocarabus) carinthiacus Sturm, 1815
Female adult
NW Slovenia, Trnovski Gozd; 24 mm
Copyright © Aleš Sedláček

Carabus (Oreocarabus) luetgensi pseudosteuarti Lapouge, 1924
Female adult
Portugal, Guarda, Sierra de Estrela h=800 m, 26.VI-4.VIII.2015, L=26 mm
Copyright © Ruslan Panin

Carabus (Oreocarabus) luetgensi pseudosteuarti Lapouge, 1924
Male adult
Portugal, Guarda, Sierra de Estrela h=800 m, 26.VI-4.VIII.2015, L=24 mm
Copyright © Ruslan Panin

Carabus (Pachystus) hortensis preslii Dejean, 1830
Male adult
S-Greece:, N Peloponnes, Erymanthos Reg., Kalentzi, 26.VI.2007, Manfred Egger leg. L=26 mm
Copyright © Ruslan Panin

Carabus (Pachystus) hortensis hortensis Linne, 1758
Female adult
Natural hybrid Carabus hortensis X Carabus granulatus. Belarus, Minsk reg., 8,7 km of Borisov, near vill. Lyubatovstchina, pine forest (Pinetum pleuroziosum), h = 213 m, 54.31934° N, 28.511486° E, 20-27.09.2018, leg. Klychenok Snezhana
Copyright © Igor Solodovnikov

Carabus (Orinocarabus) linnaei macairei Dejean, 1826
Female adult
SW-Romania, Banat, Resita - Valiug, 29.V- 9.VI.2005, Krause R. , Baros, Pittner leg.
L=19 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) fairmairei fairmairei Thomson, 1875
Male adult
N Italy, Alpi Graie; 20 mm
Copyright © Aleš Sedláček

Carabus (Orinocarabus) alpestris dolomitanus Mandl, 1956
Male adult
NE-Italy, South Tyrol, Dolomites-Alps, 500 m left of Würzjoch, Gömaalm - at the foot Mt.Peitlerkofel h=2100m, 28.VII-17.VIII.2013, Manfred Egger leg.
l=18 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei folgariacus Born, 1913
Male adult
NE-Italy, Trentino-Alto Adige (Südtirol), N of vill. San Michele vill., 13-17.VI.2016, R. Janak leg. L=20 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei linnaei Panzer, 1812
Female adult
W Ukraine, Transcarpathian Reg., Rakhiv distr., Marmarosh Mts., h~400 m, 19-23.VII.2010, Reserve "Kuziy", Panin R. leg. L=23 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI.-5,VIII.2017, R. Panin R. & Yu.Kanarsky leg.
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L =17 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Female adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=17 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=19 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Gutyn-Tomnatyk h~2015 m alpin zone, 8.VI-28.VII.2011, R. Panin R. leg. L=14 mm
Copyright © Ruslan Panin

Carabus (Oxycarabus) saphyrinus pseudosaphyrinus Schweiger, 1964 (as rabaronianus)
Male adult
Anatolia, Kastamonu, Yaraligoz verso Catalzeitin,
VI.2001 Sola Claudio leg.
male, 28 mm
Copyright © Alexander Anichtchenko

Carabus (Oxycarabus) saphyrinus pirithous Reitter, 1896
Male adult
NW-Turkey,Istanbul (Asian part),Polonezköy env., 2.VI.2009. S. Dacatra leg. L=27 mm
Copyright © Ruslan Panin

Carabus (Pachycarabus) koenigi Ganglbauer, 1886
Male adult
NW Caucasus, S Russia, Adygea Rep., Lagonaki plateau, Site Murzikal h=1800-2000m, 24.V-18.IX.2010, D.Fominykh leg.
L= 22 mm
Copyright © Ruslan Panin

Carabus (Pachycarabus) roseri soganliensis Heinz & Korge, 1964
Male+Female adult
Turkey, Rize prov., Ikizdere distr., Rize Daglari,
47 km to SE from lyidere, Sivrikaya Koyu, h=2025 m.
30.VI-5.VII.2011, A. Vlasenko leg.
male - 23 mm, female - 24 mm.
Copyright © Alexandr Safronov

Carabus (Pachycarabus) imitator imitator Reitter, 1883
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Carabus (Pachycarabus) staehlini Adams, 1817
Male adult
S Russia, E-Caucasus, Dagestan, Nukatl Mt Rng h=2600m, 25.VII.1997, V. Savizky leg. L=21 mm
Copyright © Ruslan Panin

Carabus (Pachycarabus) swaneticus kvirensis Belousov, 1992
Male adult
SC-Caucasus: NW-Georgia, Mingrelia, Egrissky Mt.Rng., h~1800-2000 m, 23-26.VII.1990, I.Retezár leg. L=20 mm
Copyright © Ruslan Panin

Carabus (Pachycarabus) koenigi Ganglbauer, 1886
Male adult

Copyright © Kirill Makarov

Carabus (Pachycarabus) imitator katherinae Reitter, 1898
Endophallus
Abkhasia, Sukhumi distr., Anchkho Mt. Pass E- part h=1870-1920 m. det. I. Solodovnikov, 2025
Copyright © Ruslan Panin

Carabus (Pachycarabus) imitator katherinae Reitter, 1898
Male adult
Abkhasia, Sukhumi distr., Anchkho Mt. Pass E- part h=1870-1920 m, 15-20.VII.2011, Pavluchuk S. leg., L= 21 mm; det. I. Solodovnikov, 2025
Copyright © Ruslan Panin

Carabus (Pachycarabus) imitator imitator Reitter, 1883
Male adult
Caucasus Major, NW Georgia, Utviri Pass env. h=2630 m, 43.083111, 42.255944
Copyright © Ruslan Panin

Carabus (Pachycarabus) roseri roseri Faldermann, 1835
Female adult
Georgia, Lesser Caucasus, Imereti prov., N slope Meskheti Mt Rng, Zekari Pass, h~1950 m, 24.VII.2021, E.Rujan leg., L=22 mm
Copyright © Ruslan Panin

Carabus (Pachystus) tamsi tamsi Menetries, 1832
Male+Female adult
Transcaucasia, Armenia,
Ashotsk, 18.VI.2001, P.V.Kazaryan leg.
female, 26mm//male, 24mm
Copyright © Ruslan Panin

Carabus (Pachystus) hungaricus hungaricus Fabricius, 1792
Female adult
C Hungary, Pest reg., Örkény, 9-10.V.1997, S.Budka leg.
Copyright © Ruslan Panin

Carabus (Pachystus) hungaricus scythus Motschulsky, 1847
Female adult
S Ukraine, Zaporizhzhya reg., Khortytsa island, 16-20.VII.2011, M. Mulenko leg., det. Panin R.
L=28 mm
Copyright © Ruslan Panin

Carabus (Pachystus) hungaricus hungaricus Fabricius, 1792
Male adult
Czech, S Moravia, Pouzdrany, 8,VIII.2005, V. Slovak lgt.
L=27 mm
Copyright © Ruslan Panin

Carabus (Pachystus) cribratus staneki Sterba, 1931
Male adult
NC Turkey, Küre Mts., 60km N Kastamonu; 25mm
Copyright © Aleš Sedláček

Carabus (Pachystus) cribratus staneki Sterba, 1931
Female adult
NC Turkey, Küre Mts., 60km N Kastamonu; 27mm
Copyright © Aleš Sedláček

Carabus (Pachystus) cavernosus rozneri Retezár & Gy. Szél, 2013 Paratype
Male adult
Romania, Apuseni : Trascău Mts. L=27 mm
Copyright © Ruslan Panin

Carabus (Pachystus) glabratus glabratus Paykull, 1790
Male adult

Copyright © Kirill Makarov

Carabus (Pachystus) cavernosus variolatus Costa, 1839
Female adult
C-Italy: Abruzzo reg., Gran Sasso massif, Campo Imperatore, h=1900 m, VIII.2021, G.Proscia leg., L=27 mm
Copyright © Ruslan Panin

Carabus (Pachystus) cribratus cribratus Quensel, 1806
Male adult
NW Armenia, Amasia, (Shirak prov.), 20-24.VIII.1998, Kazaryan P.V. leg.
Copyright © Ruslan Panin

Carabus (Pagocarabus) crassesculptus diruptus Morawitz, 1886
Male adult
China, Gansu, Umg. Xiahe 3000-3200m
24-26.6.1993 Heinz leg.
20 mm
Copyright © Alexander Anichtchenko

Carabus (Pagocarabus) crassethoracis Kleinfeld, 1999 Holotype
Male adult

Copyright © Frank Kleinfeld

Carabus (Pagocarabus) crassesculptus eleganticolor Deuve, 1997
Female adult
S Gansu, Min Shan, 5-20 km S Cagan, 2300-3200m 7-20.7.2019 V. Patrikeev leg. (cAA). det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Carabus (Pantophyrtus) turcomanorum turcomanorum Thieme, 1881
Male+Female adult
Kyrgyzstan, Kyrgyzskyi Mts. R., Uzun Bulak Gorge, h=1800 ms.
01-09.VI.2002 loc. coll.
male - 28 mm, female - 33 mm.
Copyright © Alexandr Safronov

Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993
Male adult
Uzbekistan, Chatkal Mt. rng., Irtash, 41°10'25.7"N 70°16'20.4"E, 5-7.8.2021, 2400m, A. Anichtchenko leg. (cAA). 27mm
Copyright © Alexander Anichtchenko

Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993
Female adult
Uzbekistan, Chatkal Mt. rng., Irtash, 41°10'25.7"N 70°16'20.4"E, 5-7.8.2021, 2400m, A. Anichtchenko leg. (cAA). 30 mm
Copyright © Alexander Anichtchenko

Carabus (Piocarabus) titanus Breuning, 1933
Male adult
C-China Sichuan prov., Nanping, h=1500m, VI.2000, S. Remond leg.<br /> L=34 mm
Copyright © Ruslan Panin

Carabus (Piocarabus) sui Imura & Zhou, 1998
Female adult
CE-China, Henan, Mt Wangwu, V.2007, local collector. L= 33 mm
Copyright © Ruslan Panin

Carabus (Piocarabus) vladimirskyi Dejean & Boisduval, 1830
Male adult
Buryatia, Kyahta distr. [Khyagh], 5 km SE Chikoi vill., 16.8.1998, Anichtchenko A. leg.
Copyright © Alexander Anichtchenko

Carabus (Piocarabus) titanus Breuning, 1933
Female adult
China Sichuan prov., Nanping, h=1500m, VI.2000, S. Remond leg.
L=38 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Male adult
W.Ukraine, Transcarpathian Reg., Rakhov Distr., Chornogora range., h=2020m. Panin R. leg. 28.VI-9.VIII.2008, male, 19 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Female adult
Ukrainian Carpathians, Transcarpathian reg., Rakhov Distr., Chornogora range, h=2010m, 29.VI-28.VII.2007, Panin R. leg., female, 23 mm., photo/det.Panin R
Copyright © Ruslan Panin

Carabus (Platycarabus) irregularis ramanus Sokolar, 1909 (as hayeki Kobmann, 1925)
Male adult
N Slovenia, Maribor – Dravograd, Pomorje Mts.,13.XI.2005, I.Rapuzzi leg.
Copyright © Ruslan Panin

Carabus (Platycarabus) creutzeri creutzeri Fabricius, 1801 (as trnovensis Bernau, 1914)
Male adult
W Slovenia,
Lokve, Trnovski Gozd h=1100m, 17.IV.2006, leg. I.Rapuzzi
male, 26mm
Copyright © Ruslan Panin

Carabus (Platycarabus) creutzeri creutzeri Fabricius, 1801(as posarinii Rapuzzi, 1999 PT) Paratype
Male+Female adult
Slovenia:Koper:Kras Mt.R.:Brkini Mts.:Suhorije env., VIII.1998, I.Rapuzzi leg.
photo & koll. Panin R.
female, 32 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) fabricii fabricii (as fassatii Smetana, 1955)
Male adult
C-Slovakia:Nízke Tatry Mts., 4.VI.2011, Krause Rene leg.
L=19 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) fabricii fabricii (as fassatii Smetana, 1955)
Male adult
C-Slovakia:Nizke Tatry Mts., h=1750 m,.9.VI.2007, V.Slovak leg.
L=20 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Female adult
N- Romania, E-Carpathians, Rodna Mts, h=2000m, 7-8.VI.2012, Cartuz Z.leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) creutzeri kircheri Germar, 1824
Male adult
S Austria, Karnische Alpen; male, 24mm
Copyright © Aleš Sedláček

Carabus (Platycarabus) creutzeri baldensis Schaum, 1857
Male adult
NC-Italy: Trento, Sega di Ala, IX.1999, A. Vicini leg. , L=24 mm
Copyright © Ruslan Panin

Carabus (Procerus) scabrosus tauricus Bonelli, 1811
Female adult

Copyright © Safronov

Carabus (Procerus) gigas gigas Creutzer, 1799
Male+Female adult
N Croatia, Ivanščica Mts , Ivanec, 4-18.VI.2011, Krause Rene leg.
female, 54 mm
Copyright © Ruslan Panin

Carabus (Procerus) scabrosus caucasicus Adams, 1817
Male adult
N Caucasus,
Stavropol Krai, Pyatigorsk, Mt.Mashuk, VI.2007 Tikhonov Yu. leg.
44mm
Copyright © Ruslan Panin

Carabus (Procerus) gigas duponcheli Dejean, 1831 (as kusdasi Mandl, 1966)
Male adult
S-Greece: Peloponnes, Mt.Chelmos, 20.V.2007.
L=46 mm
Copyright © Ruslan Panin

Carabus (Procrustes) chevrolati chevrolati Cristoforis & Jan, 1837
Male+Female adult
female, 31mm: Ashotsk,Vardaghbyur vill., 30.V.1997
male, 30 mm: Amasia, Chaldash vill.,8-11.VIII.1997
P.V. Kazaryan leg.
Copyright © Ruslan Panin

Carabus (Procrustes) chevrolati thirki Chaudoir, 1857
Male+Female adult
Turkey, Rize prov., Ikizdere distr., Rize Daglari,
47 km to SE from lyidere, Sivrikaya Koyu, h=2025 m.
30.VI-5.VII.2011, A. Vlasenko leg.
male - 28 mm, female - 31 mm.
Copyright © Alexandr Safronov

Carabus (Procrustes) anatolicus anatolicus Chaudoir, 1857
Male adult
Cyprus, Troodos, 17.2.1V.2010,. Manfred Egger leg.
L=27 mm
Copyright © Ruslan Panin

Carabus (Procrustes) banoni Dejean, 1829
Male adult
Greece,W Crete Island, Mili gorge ,12.V.2009, Manfred Egger leg.
L=26 mm
Copyright © Ruslan Panin

Carabus (Procrustes) talyschensis talyschensis Menetries, 1832
Female adult
Azerbaijan, Talysh Mts., Zuvand, Kalvaz vill., h=1650 m, 15.VII.2003, Dolin V.G. leg.
L=32 mm
Copyright © Ruslan Panin

Carabus (Procrustes) piochardi praestigiator Morawitz, 1886
Male adult
C-Lebanon, Byblos /Jbeil Distr., Kfardebian env., h=1200m, XI.2014, C. Reuter leg. L=31 mm
Copyright © Ruslan Panin

Carabus (Procrustes) coriaceus ressli Mandl, 1961
Male adult
SC-Turkey: Hatay Prov, Dörtyol, Nur Mts, Topaktaş h=900-1200 m, 12.V.2005, D. Prunier leg.
Copyright © Ruslan Panin

Carabus (Procrustes) chevrolati internatus Heinz & Korge, 1964
Female adult
Turkey, Bolu, Gerede, Esentepe 1350m, 1.7.1994, leg. K. Staven
Copyright © Alexander Anichtchenko

Carabus (Procrustes) coriaceus excavatus Charpentier, 1825
Male adult
W Croatia, Velebit Mts, Jablanac Stirovaca h=998 m, 44.71124, 14.95673,3.VI-7.VII.2017, G. Proscia leg., L= 37 mm
Copyright © Ruslan Panin

Carabus (Procrustes) coriaceus excavatus Charpentier, 1825
Female adult
W Croatia, Velebit Mts, Jablanac Stirovaca h=998 m, 44.71124, 14.95673,3.VI-7.VII.2017, G. Proscia leg., L= 38 mm
Copyright © Ruslan Panin

Carabus (Pseudocoptolabrus) burmanensis Breuning, 1932
Male adult
male, 23mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) businskyi Deuve, 1990
Male adult
CH, NW-Yunnan (Lijiang) Yulongxue Shan, E-slope
27:10:50N/100:12:26E/ 4400m, alpine meadows 12.VII.2000
K. & B. Brezina, M.Brychcin, M. Hackel
Copyright © Boleslav Brezina

Carabus (Pseudocranion) sackeni sackeni Semenov, 1898
Male adult
male, 23mm
Copyright © Boleslav Brezina

Carabus (Pseudocranion) sackeni remondianus Deuve, 1996 Paratype
Female adult
Museum of Institute of Systematic Biology (Daugavpils, Latvia)
Copyright © Alexander Anichtchenko

Carabus (Pseudocranion) lazikouensis lazikouensis (as kundaensis Deuve, 2003) Paratype
Female adult
C-China, S-Gansu: SSW of Minxian: Qilian Shan: W Kunda, 34:10N/103:47E, h=2800-3000m, forest, 7.VII.2002, Belousov & Kabak leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Pseudocranion) taibaishanicus taibaishanicus Deuve, 1989
Female adult
China, W-Shaanxi, Taibai Shan Mts, btw.Houzhenzi/Ba Xian Tai, nr. Nan Tian Men h=3200m,
33°55'17"N/ 107°46'53"E, 25-30.VII.2011, Li Wuyang leg.
L=29 mm
Copyright © Ruslan Panin

Carabus (Pseudotribax) ferghanicus Breuning, 1933
Male adult
male, 29mm
Copyright © Boleslav Brezina

Carabus (Relictocarabus) meurguesianus Ledoux, 1990
Male adult
Image from molbiol.ru
Copyright © - - -

Carabus (Rhigocarabus) sehnali Brezina & Hackel, 2006
Female adult
female, 23mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) gentleman Brezina & Hackel, 2004 Holotype
Male adult
male, 19mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) indigestus confucius Breuning, 1933
Female adult
C-China, E Tibet,W-Sichuan prov., Kangding, Zheduo Pass h=4300m 30.VI.2005 , V.Benes leg,
L=17 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) buddaicus linxiaicus Deuve, 1995
Male adult
China, S-Gansu (LABRANG):, valley.E of Ponggartang, 35:12-13N/102:54-55E, h=3000-3700m, alpine meadows, coniferous forests,10.-14.VII.1999, B.Brezina leg.
L=20 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) morawitzianus morawitzianus Semenov, 1887
Male adult
C China, Qinghai, Zadoi County, Toramarkog h~4600 m, VI.2001, P. Cavazzuti leg. L=17 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) turnai turnai Deuve, 1994
Male adult
China, Tibet, W-Sichuan, pass btw Sêrtar / Luhuo h=4200 m, 28.VI.2011, V.Patrikeev leg.L=19 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) turnai turnai Deuve, 1994
Female adult
China, Tibet, W-Sichuan, pass btw Sêrtar / Luhuo h=4200 m, 28.VI.2011, V.Patrikeev leg. L=21 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) cateniger cateniger (as sommitalis Cavazzuti, 2006)
Female adult
China, C-Gansu, Mts Qilian h=3378 m, road G227, Yangxiongzi env. 38.050917, 100.887267, 18-23.VI.2011, G.Proscia leg., L=19 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) cateniger dabanicola Deuve, 1998
Male adult
China, Qinghai, Xining city, 55km NE of Datong, Daban Shan h=3936 m, 37.348450, 101.397600, 17-23.VI.2011, G.Proscia leg., L= 16 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) roborowskii roborowskii Semenov, 1887
Male adult
China, E-Qinghai, Mts NW of Tongren city, 35.562778, 101.843667, h=3870 m, 15-26.VI.2011, G. Proscia leg./det.R.Panin, L=20 mm
Copyright © Ruslan Panin

Carabus (Scambocarabus) kruberi laobeiensis Deuve & Li, 2000
Male adult
NE-China: Liaoning prov., Fengcheng City, Miaoyang, 12-17.VI.2011
L=27 mm
Copyright © Ruslan Panin

Carabus (Semnocarabus) regulus regulus Dohrn, 1882
Male adult
photo M. Smirnov from www.zin.ru
Copyright © Maksim Smirnov

Carabus (Semnocarabus) carbonicolor carbonicolor Morawitz, 1886
Male+Female adult
Kyrgyzstan, NC Tianshan, Kyrgyz Mt. R., Tuyuk riv., h=2000 м.
25-VI-1997 М. Danilevsky leg.
male - 20 mm, female - 21 mm.
Copyright © Alexandr Safronov

Carabus (Semnocarabus) carbonicolor korolkowi Semenov, 1896
Male adult
NC-Kyrgyzstan, Kyrgyz Mt.Rng , 10 km SW of Bishkek, Chon-Aryk env, h=1100m, 24.V.1998, /det. Panin R.
22 mm

Copyright © Ruslan Panin

Carabus (Semnocarabus) erosus corax Semenov, 1896
Endophallus
SE-Kazakhstan, Almaty reg., C Trans-Ili Alatau N-slopes, Kaskalen river Valley, Izvestkovaya vill., h=1700 m, 6.V.2015, D.Kovalchuk leg./det. R.Panin,
Copyright © Ruslan Panin

Carabus (Semnocarabus) erosus corax Semenov, 1896
Male adult
SE-Kazakhstan, Almaty reg., C Trans-Ili Alatau N-slopes, Kaskalen river Valley, Izvestkovaya vill., h=1700 m, 6.V.2015, D.Kovalchuk leg./det. R.Panin,
Copyright © Ruslan Panin

Carabus (Semnocarabus) carbonicolor korolkowi Semenov, 1896
Female adult
SE-Kazakhstan, Almaty, E Zailisky Alatau, Bokaydin-tau, Belshabdar valley h=1300 m, 5-27.VI.2002, A.Puchkov leg.L=21 mm
Copyright © Ruslan Panin

Carabus (Semnocarabus) carbonicolor carbonicolor Morawitz, 1886
Female adult
SE-Kazakhstan, Almaty reg., Khan Tengri Massif, Narynkol env.h=1800 m, 30-31.-V.2007, Shumov leg., L=21 mm
Copyright © Ruslan Panin

Carabus (Semnocarabus) erosus erosus Motschulsky, 1865
Endophallus
SE-Kazakhstan, 70 km E of Almaty, E Trans-Ili Alatau N-slopes, valley Assy (Asy) riv. Assy plateau h=1700 m, 12.V.2015
Copyright © Ruslan Panin

Carabus (Semnocarabus) erosus prope erosus Motschulsky, 1865
Male adult
SE-Kazakhstan, 70 km E of Almaty, E Trans-Ili Alatau N-slopes, valley Assy (Asy) riv. Assy plateau h=1700 m, 12.V.2015, L=21 mm,/ det. R. Panin
Copyright © Ruslan Panin

Carabus (Semnocarabus) rustemi Kabak, 2009
Male adult
SE-Kazakhstan, Almaty reg., near Bodokhudzir 43.916667, 79.716667 h=500 m, 12.VIII.2019, det. R.Panin, L=22 mm
Copyright © Ruslan Panin

Carabus (Shunichiocarabus) uenoianus uenoianus Imura, 1995
Male adult

Copyright © - - -

Carabus (Pagocarabus) draco Brezina, 1999
Male adult

Copyright © Boleslav Brezina

Carabus (Pagocarabus) caelestinus caelestinus (Imura, 2007) Paratype
Male adult

Copyright © Boleslav Brezina

Carabus (Pagocarabus) sichuanicola sichuanicola (as drolmae Lassalle, 2003)
Female adult
China, Sichuan, NW Chengdu, Mt. Jiajin Shan h~4000 m,VI. 2016, local collector, L= 24 mm
Copyright © Ruslan Panin

Carabus (Sigenacarabus) trachynodes moxiensis Deuve, 1992
Male adult
China, W-Sichuan, W of Moxi, Daxue Range, Mt.Gongga Shan h=3000-3100m, 29°34′29.6″N /101°59′44.0″E, 10-12.VI.2018, Ch. Reuter leg. L=21 mm
Copyright © Ruslan Panin

Carabus (Sigenacarabus) trachynodes moxiensis Deuve, 1992
Male adult
China, W-Sichuan, Daxue Range, Mt.Gongga Shan h=3000-3100m, Hailuogou Glacier Forest Park, 29.574889, 101.995556, 10-12.VI.2018, Ch. Reuter leg., L=21 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) varians varians Fischer, 1827
Male+Female adult
female: Transcaucasia, Armenia, Amasia distr., Karakhachskiy mountain pass h=2270m, 3.VIII.1985 P.V.Kazaryan leg., 22mm<br /> male: Armenia, Amasia distr., Arpalach vill. VI.1999, P.V. Kazaryan leg.
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) varians armeniacus Mannerheim, 1830
Female adult
NE Turkey,<br /> Artvin Prov., Borçka, Macahel Gecidi h=1450m, 18.VI.2007, D.Prunier leg.<br /> female, 26mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) adamsi Adams, 1817 (as chiragricus Fischer, 1828)
Male+Female adult
Russia, N. Caucasus, Stavropol reg., Zheleznovodsk, Zheleznaya Mt., h=600 m. 5-11.VI.2011, Е. Shankhiza leg. male - 24 mm, female - 25 mm.
Copyright ©

Carabus (Sphodristocarabus) macrogonus macrogonus Chaudoir, 1847
Male adult
NE-Turkey:Trabzon env., Sumela, 17-21.VI.2010, M.Janalik leg.
L=20 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) macrogonus pseudokolatensis Heinz & Korge, 1967
Male adult
NE-Turkey::Kackar Dagi Mt, Ayder S of Ardesen h=1600m, 17-21.VI.2010, M.Janalik leg.
L=20 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) separatus persa Lapouge, 1924
Male adult
N-Iran: Mazandaran Province, Elborz Mt.Rng, Vandarbon env., 10.VII.2002 L=23 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) blumenthaliensis aequicostalis Staven & Heinz, 1995
Male adult
NE-Turkey:NE of Artvin:Dalis Dağ ,h=1600-1800m, 8-18.VII.2014, C. Reuter leg. L= 25 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) separatus separatus Gehin, 1885
Male adult
N-Iran:Gilan Prov., Asalem - Khalkal road, 25-30.V.2018, Václav Čutka leg. L= 21 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) separatus separatus Gehin, 1885
Female adult
N-Iran: Gilan Prov., Asalem - Khalkal road, 25-30.V.2018, Václav Čutka leg. L= 23 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) separatus separatus Gehin, 1885
Male adult
N-Iran:Gilan Prov., Asalem - Khalkal road, 25-30.V.2018, Václav Čutka leg. L= 22 mm
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus tibiscinus Csiki, 1905
Male+Female adult
female,28 mm
SE Hungary, Csongrбd, Makу env., 1-8.VIII.2004, Krause R., Slovak, Pittner leg
Copyright © Ruslan Panin

Carabus (Tachypus) auratus s.str. (as brunieri Barthe, 1921)
Male adult
SC-FRANCE, Aude, Rennes les Bains, 6.VII.1989, Sunard A. leg.
male, 26mm
Copyright © Ruslan Panin

Carabus (Tachypus) auratus lasserrei Doue, 1855
Male adult
SC-FRANCE, Midi-Pyrуnуes reg. Tarn, Saint Paul cap de Joux, 24.V.2003, A. Laforgue leg.
male, 24mm
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
Male adult
Italy-Switzerland, Riviera – Preonzo, 18.II.1998, C.Sola leg.
L=26 mm
Copyright © Ruslan Panin

Carabus (Tachypus) cristoforii Spence, 1823
Female adult
S France, Pyrénés Mts., Tarascon; 15mm
Copyright © Aleš Sedláček

Carabus (Tachypus) cancellatus durus Reitter, 1896 (as carpathicus Kolbe, 1913)
Male+Female adult
Ukrainian Carpathians,<br /> Ivano-Frankivsk reg., Gorgany Mts., 20km SW Nadvorna, 22.VI-3.VIII.20, Panin R. leg.<br /> female, 26mm
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus durus Reitter, 1896
Male+Female adult
W Ukraine,<br /> Transcarpathian reg., Vinogradov distr., near Malaya Kopanya vill.,7-18.VII.2008, Panin R. leg. <br /> female, 32mm.
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus intermedius Dejean, 1826
Female adult
NW -Serbia,Fruška Gora,Vrdnik env. h=380m, 9-10.VII.2013, M. Janalik leg. L=29 mm
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus durus (as strictus Kolbe, 1913)
Male adult
W Ukraine, NE Carpathians, Maramaros Mts (Hutsul Alps) Gorge of Kuziy (Kizi) stream, h = 400m, 20-23.V.2021, Panin R. leg., L=30 mm
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus durus (as strictus Kolbe, 1913)
Female adult
W Ukraine, NE Carpathians, Maramaros Mts (Hutsul Alps), Gorge of Kuziy (Kizi) stream, h = 400m, 20-23.V.2021, Panin R. leg., L=32 mm
Copyright © Ruslan Panin

Carabus (Tanaocarabus) hendrichsi Bolivar y Pieltain, Rotger & Coronado, 1967
Male adult
male, 23mm
Copyright © Boleslav Brezina

Carabus (Teratocarabus) azrael mizunumaianus Imura, 1991
Male adult
China, Liaoning, Benxi, Gongchangung
20.VI.2004 Hong
Copyright © Boleslav Brezina

Carabus (Tomocarabus) scabripennis scabripennis Chaudoir, 1850
Male adult
Carabus (Tomocarabus) scabripennis scabripennis Chaudoir, 1850
C. Armenia,
Kotayk reg., Yeghvard, Arailer Mt., 16.VI.1999, P.V.Kazaryan leg.
male, 20mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Tomocarabus) marginalis Fabricius, 1794
Male+Female adult
E Ukraine,
Kharkov reg.,near Vasishchevo vill.10-20.VI.2006 D.Kovalchuk leg.
female 24mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) convexus s.str. (as pyrenaicus Born, 1907)
Male adult
Pirineos, Canfranc, 1300m, Barranco de Izas
15.6.2005 Anichtchenko A. leg.
male, 16 mm
Copyright © Alexander Anichtchenko

Carabus (Tomocarabus) convexus moreanus Reitter, 1896
Female adult
Greece:, N Peloponnes, Xylokastro, Zemeno,h=1000m, 6.VI.1997, Manfred Egger leg.
L=21 mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) rumelicus syriensis Breuning, 1943
Male adult
N Lebanon, Fnaпdek, Djebel Qammoua, h=1100-1200m, C. Reuter leg.
L=17.5 mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) convexus moreanus Reitter, 1896
Male adult
S Greece, Taygetos Mts.; 13mm
Copyright © Aleš Sedláček

Carabus (Tomocarabus) convexus dilatatus Dejean, 1826
Male adult
NE Serbia, Majdanpek; 13mm
Copyright © Aleš Sedláček

Carabus (Tomocarabus) convexus gracilior Gehin, 1885
Female adult
SW Romania, Banat; 15mm
Copyright © Aleš Sedláček

Carabus (Oreocarabus) taedatus bicanaliceps Casey, 1920
Male adult
NW-USA, Washington, Thurston County, Mima Mound Prairie, 15.IX.2005, A. Laforgue leg. L=23 мм
Copyright © Ruslan Panin

Carabus (Tomocarabus) convexus erzeliki Schweiger, 1962
Female adult
NC-Turkey, Sinop province, Erfelek env., h=200m, 10-27.VII.2014, C. Reuter leg. L=20 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus sibiricus Fischer, 1822
Male adult

Copyright © Kirill Makarov

Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886
Male+Female adult
W Ukraine,<br /> Ternopol reg., Pidvolochisk distr., Supranivka vill.<br /> 3.VII.1908. Leg. Stobetskii. S.<br /> Photo R. Panin
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus haeres (as fossulatus Dejean, 1826)
Male+Female adult
E.UKRAINE,<br /> Kharkov reg., Zmiev distr.,in steppe,10-20.V I.2007, D.Kovalchuk leg.<br /> female, 26mm/male, 24mm.<br /> photo/det. Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus errans Fischer, 1823
Male+Female adult
female:SE Ukraine,
Donetsk, 25.VII.2009, A.I.Gubin leg. L=30mm.
male:E Ukraine, Lugansk reg., near Slavyanoserbk, 5-15.VIII.2009, S.Konovalov leg. L=26mm.
photo/det.Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus planus Gehin, 1885
Endophallus
<b>endophalus</b> Karacaj-Cherkessija, near Kart-Djurt vill., upper Kylan riv. h=1300m
Copyright © Ruslan Panin

Carabus (Trachycarabus) besseri Fischer, 1822
Male+Female adult
W Ukraine.NW Podolian,
Lviv reg., Zolochiv distr., Lysa Gora near Stinka vill., steppe sites, 25.IV-9.V.2009 Panin R. leg
female, 30mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus haeres (trans s. errans)
Female adult
C Ukraine, S of Poltava city, edge forest-lane, 2-26.IX.2010, O. Pozharov leg. /det. R.Panin. L=30 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus retowskianus Mandl, 1955
Male adult
S Ukraine, Zaporizhia reg., Melitopol distr., near Troitskoe vill., 1.VI.2010, I.V. Kovalev leg., L=27 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) estreicheri Fischer, 1822 (ab.rufofemoratus Lomnicki, 1892)
Female adult
W Ukraina, Podolian Upland, Lviv reg., Brody distr., Mt. Makitra, steppe site, 19.V-5.VI.2020, Panin R. leg. L=23 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus campicola Dejean, 1831
Female adult
S Ukraine, Crimea, S slope Mt Demerdzhi h=650 m, sibljak, VII.2004, R. Panin leg., L=29 mm
Copyright © Ruslan Panin

Carabus (Tribax) circassicus circassicus (as tshchaltensis Novotný & Voříšek
Male adult
W-Caucasus: Abkhasia, Gudauta distr., E slope Mt Anchkho h=2200-2350 m, 14-20.VII.2011, S.Pavluchuk, I. Solodovnikov leg./det. I. A. Solodovnikov.
25 mm

Copyright © Ruslan Panin

Carabus (Tribax) agnatus pseudagnatus Novotný & Voříšek, 1988
Male adult
W-Caucasus: Abkhasia, Gudauta distr., Mt Anchkho h=2200-2350 m, 14-20.VII.2011, S.Pavluchuk, I. Solodovnikov leg./det. I. A. Solodovnikov.
L=25 mm
Copyright © Ruslan Panin

Carabus (Tribax) agnatus agnatus Ganglbauer, 1889
Male adult

Copyright © Kirill Makarov

Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
Male adult
Russia, C-Caucasus: N Ossetia, Chekhatsirtit Mt range, Mt Tepli h=3000 m, 21.VII.2009. /det. Solodovnikov I.A, L=20.5 mm
Copyright © Ruslan Panin

Carabus (Tribax) steveni schamyli Hampe, 1852
Male adult
C Caucasus, North Ossetia, the road on the village of Coban (Koban), the left river bank Gizeldon, the deciduous wood, h = 895-905 m, N 42°55'32.91", E 44°31'53.53", 20.05-03.07.2017, leg. A.P. Tatulyan
Copyright © Igor Solodovnikov

Carabus (Tribax) steveni schamyli Hampe, 1852
Male adult
C Caucasus, North Ossetia, the road on the village of Coban (Koban), the left river bank Gizeldon, the deciduous wood, h = 895-905 m, N 42°55'32.91", E 44°31'53.53", 20.05-03.07.2017, leg. A.P. Tatulyan
Copyright © Igor Solodovnikov

Carabus (Tribax) nothus fausti Dohrn, 1873
Female adult
Carabus (Microtribax) fausti fausti Dohrn, 1873 Russia: Daghestan, Kyabyak Mt.R.
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Endophallus
Russia, Caucasus, Ingushetia, South slopes of Skalistyi Mt Range, Stolovaya Mt., near Beyni, alpin zone, h = 2800 m, N 42°51’35.30",E 44°42’13.70", h = 2800m, 21.V–19.VII.2018, D. Fominykh leg. (TOPOTYPUS)
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Male adult
Russia, Caucasus, Ingushetia, Skalistyi Mt. Range, S slopes of Tsoreylam Mt. Range, 5 km W of Mt. Khakhalgi, N 42°50’07.60", E 45°05’02.20", h=2500 m, 25.V–21.VII.2017, D. Fominykh leg.
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
Male adult
Russia: N.Osetia, Bokovoy Mt.R., near Arkhonskiy pass., alpin zone, h = 2700 m, 23.V-21VII.2018, D. Fominykh leg.
Copyright © Igor Solodovnikov

Carabus (Ulocarabus) stschurovskii Solsky, 1874
Male adult
Tadzhikistan, Pamir-Alay, S slope Gissar Mt.Range, Takob, h-1700m, 10.V.1996, Dolin V.G. leg.
det. Panin R.Yu.
L=22 mm
Copyright © Ruslan Panin

Carabus (Ulocarabus) stschurovskii Solsky, 1874
Female adult
Tadzhikistan, Pamir, N-slope Peter I.Mt Rng,h=2100m,Ganishou vill., Dolin V.G. leg.
det.Panin R.Yu.
L=24 mm
Copyright © Ruslan Panin

Carabus (Ulocarabus) theanus delerei Mandl, 1955
Male adult
N-Afghanistan, Hindukush Mt.Rng, NE Salang Pass h=2300 m, 35°27'494"N, 68°58'399"E, IV.2011, C. Reuter leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Ulocarabus) theanus theanus Reitter, 1895
Male adult
Tadzhikistan, S slope Gissar Mt.Rng, left bank of the river Sorbo, Haidar gorge,1.VI.1984, Dolin V.G. leg./det.Panin R.Yu.
L=21 mm
Copyright © Ruslan Panin

Carabus (Ulocarabus)? theanus theanus Reitter, 1895
Male adult
NC Tadzhikistan, Pamir, N-slope Peter I.Mt Rng, h=2100m, Ganishou vill., 10-12.VI.2006, Ivanova E. leg.
L=23 mm
Copyright © Ruslan Panin