Carabidae

Carabidae image gallery

Region

Anthia mannerheimi afghana Anichtchenko, 2009 Holotype
Male adult
Afghanistan, Oruzgan, Sahrestan 2400m, 20.VII.1970 Kabakov leg.
male, 34 mm
deposited in ZIN
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Anthia mannerheimi afghana Anichtchenko, 2009 Paratype
Female adult
Afghanistan, Oruzgan, Sahrestan 2400m, 20.VII.1970 Kabakov leg.
female, 32 mm
deposited in ZIN
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Anthia (Anthia) artemis Gerstaecker, 1884
Male adult
Tanzanie, Nyanza, 11.1988
Copyright © Alexander Anichtchenko

Anthia (Anthia) cinctipennis cinctipennis Lequien, 1832
Male adult
length: 37-42 mm
Namibia, Etosha National Park
leg. U.Schmidt, 1994; det. W.Lorenz
Foto: U.Schmidt, 2006
Copyright © U.Schmidt

Anthia (Anthia) circumscripta circumscripta Klug, 1853
Female adult
Namitembo, Zomba, Malawi 11.1987
det. Vlasenko A. S. 2011
Copyright © Alexander Anichtchenko

Anthia (Anthia) mannerheimi mannerheimi Chaudoir, 1842
Male adult
length: 45 mm
Turkmenia, Finuza, 1992
Foto: U.Schmidt, 2006
Copyright © U.Schmidt

Anthia (Anthia) mannerheimi mannerheimi Chaudoir, 1842
Male adult

Copyright © Alexander Anichtchenko

Anthia (Anthia) circumscripta circumscripta Klug, 1853
Male adult
Tanzania, Mbonde area, Masasi, feb.2005
male, 52 mm
photo, det. & coll. Anichtchenko A. 2009
det. Vlasenko A. S. 2011
Copyright © Alexander Anichtchenko

Anthia (Anthia) circumscripta circumscripta Klug, 1853
Female adult
Tanzania, Mtandi (Masasi), XII.2006. Female, body lenght 48 mm det. Vlasenko A. S. 2011
Copyright © Denis Potanin

Anthia (Anthia) circumscripta circumscripta Klug, 1853
Male adult
Tanzania, Mtandi (Masasi), XII.2006. Male, body lenght 45 mm det. Vlasenko A. S. 2011
Copyright © Denis Potanin

Anthia (Anthia) sexguttata (Fabricius, 1775)
Female adult
India, Tiruvannamalai, Arcot, D. Tamil Nadu state, 10.9.1997 T.R.S. Nathan leg. (cAA) 46 mm; det. Anichtchenko A. 2009
Copyright © Alexander Anichtchenko

Anthia (Chilanthia) cavernosa Gerstaecker, 1866
Male adult
Sagala Hills, Kenya 12.1993
Copyright © Alexander Anichtchenko

Anthia (Chilanthia) cavernosa Gerstaecker, 1866
Male adult
length: 30-36 mm
Kenia, Tsavo National Park
leg. U.Schmidt, 1991; det. W.Lorenz
Foto: U.Schmidt, 2007
Copyright © U.Schmidt

Anthia pulcherrima H. W. Bates, 1888
Male adult
24/XII/2007-03/I/2008, Mikumi Village, Udzungwa Mts. Nat. Park, Tanzania
Copyright © Denis Potanin

Anthia (Termophilum) burchelli Hope, 1832
Male adult
Tanzania, Morogoro reg. long 46 mm
Copyright © Denis Potanin

Anthia (Termophilum) decemguttata (Linnaeus, 1764)
Male adult
S Africa,
38 mm
Copyright © Alexander Anichtchenko

Anthia (Termophilum) hexasticta Gerstaeker, 1866
Female adult
length: 40 mm Kenia, Tsavo National Parc, leg. U.Schmidt, 1991; det. W.Lorenz
Copyright © U.Schmidt

Anthia (Termophilum) omoplata parva Obst, 1901
Male adult
Tanzania, Mbonde area, Masasi, feb.2005
44 mm
photo, det. & coll. Anichtchenko A. 2009
det. Vlasenko A. S. 2011
Copyright © Alexander Anichtchenko

Anthia (Termophilum) omoplata omoplata Lequien, 1832
Male adult
length: 35 mm
Namibia, Windhoek
leg. U.Schmidt, 1994; det. W.Lorenz
Foto: U.Schmidt, 2006
Copyright © Udo Schmidt

Helluobrochus brevicollis (Dejean, 1831)
Male adult
Ecuador, Yasuni
Photo I.Sokolov
Copyright © Igor Sokolov

Helluomorpha eulinae Reichardt, 1974
Male adult
Paraguay, Dep. Pte. Hayes, Puerto Galileo,S25.04 W57.52 5-8.III.2008 leg. U.Drechsel. 19 mm; det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Macrocheilus saulcyi Chevrolat, 1854
Male adult
Libanon, Djezin, det. Ing. Jedlicka
Copyright © Ron Felix

Macrocheilus vanharteni Felix & Muilwijk, 2007 Holotype
Male adult
Seyun, 12-14.VIII.2002, light trap, leg. A.v. Harten &
A. Al Zubayri, Holotype, male 12,4 mm
Photo R. Felix
Copyright © Alexander Anichtchenko

Macrocheilus vanharteni Felix & Muilwijk, 2007 Paratype
Male adult
Al Kowd, VI.2002, light trap, leg. A.v.Harten & S. Al
Haruri, paratype, male, collection R. Felix
Photo R. Felix
Copyright © Alexander Anichtchenko

Anthia (Termophilum) sexmaculata sexmaculata (Fabricius, 1787)
Male adult
Sahrawi Arab Democratic Republic, Erqueyez Wadi 20.11.2003
male, 34 mm
photo, det. & coll. Anichtchenko A. 2009
Copyright © Alexander Anichtchenko

Anthia (Termophilum) venator Fabricius, 1792: 141
Male adult
Algeria, Oasis Beni Abbes,
male, 47 mm
photo, det. & coll. Anichtchenko A. 2009
Copyright © Alexander Anichtchenko

Anthia (Termophilum) fornasinii Bertoloni, 1845
Male adult
NE Zambia, 92 km NW Mpika, 27.XI.2006; ing. Snizek leg. Body lenght 42 mm.
Copyright © Denis Potanin

Cycloloba septemguttata (Fabricius, 1794)
Male adult
RSA, Kwazulu/Natal, Loteni Game Reserve, 1800m, 27.XI.2000
23 mm
det. P. Schьle
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Cypholoba alveolata (Breme, 1844)
Male adult

Cypholoba (Cypholoba) alveolata

Copyright © Alexander Anichtchenko

Cypholoba gracilis Dejean, 1831
Male adult

Copyright © Alexander Anichtchenko

Cypholoba tenuicollis errata Strohmeyer, 1928
Male adult

Copyright © Alexander Anichtchenko

Cypholoba caillaudi (Castelnau, 1835)
Male adult
Kafue, Zambie 11.1986
Copyright © Alexander Anichtchenko

Eccoptoptera cupricollis Chaudoir, 1878
Male adult
Ethiopia, Illubabor prov., 15 km W Abobo 10.9.88
Copyright © Alexander Anichtchenko

Eccoptoptera mutilloides lagenula (Gerstaecker, 1866)
Male adult
Uluguru Mts., Tanzania, July 2007 
Copyright © Denis Potanin

Gonogenia tabida (Fabricius, 1793)
Male adult

Copyright © Alexander Anichtchenko

Netrodera formicaria damarensis Kuntzen, 1919
Male adult
SW Africa, Kaokoveld, Anabib (Orupembe)
100 miles W Ohopobo 12-13.6.51
Copyright © Alexander Anichtchenko

Helluomorphoides praeustus bicolor (T.W.Harris, 1828)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Anthia (Termophilum) aemiliana Dohrn, 1881
Male adult
L=37 mm
NAMIBIA, Khomas reg., ~106 km ESE from Windhoek, Arnhem Cave & Lodge territory, Kalahari, on yellow-reddish sand, at night (20:00-0:00). H=1465 m. 13-18.XII.2010
det & photo Vlasenko A. S.
Copyright © Andrey Vlasenko

Anthia (Termophilum) aemiliana Dohrn, 1881
Female adult
L=41 mm
NAMIBIA, Khomas reg., ~106 km ESE from Windhoek, Arnhem Cave & Lodge territory, Kalahari, on yellow-reddish sand, at night (20:00-0:00). H=1465 m. 13-18.XII.2010
det & photo Vlasenko A. S.
Copyright © Andrey Vlasenko

Anthia (Termophilum) andersoni andersoni Chaudoir, 1861
Male adult
L=51 mm
NAMIBIA, Khomas reg., ~106 km ESE from Windhoek, Arnhem Cave & Lodge territory, Kalahari, on yellow-reddish sand, at night (20:00-0:00). H=1465 m. 13-18.XII.2010
det & photo Vlasenko A. S.
Copyright © Andrey Vlasenko

Anthia (Termophilum) burchelli burchelli Hope, 1832
Female adult
L=46 mm
NAMIBIA, Khomas reg., ~106 km ESE from Windhoek, Arnhem Cave & Lodge territory, Kalahari, on yellow-reddish sand, at night (20:00-0:00). H=1465 m. 13-18.XII.2010
det & photo Vlasenko A. S.
Copyright © Andrey Vlasenko

Anthia (Anthia) cinctipennis cinctipennis Lequien, 1832
Female adult
L=41 mm
NAMIBIA, Khomas reg., ~106 km ESE from Windhoek, Arnhem Cave & Lodge territory, Kalahari, on yellow-reddish sand, at night (20:00-0:00). H=1465 m. 13-18.XII.2010
det & photo Vlasenko A. S.
Copyright © Andrey Vlasenko

Anthia (Anthia) cinctipennis cinctipennis Lequien, 1832
Male adult
L=40 mm
NAMIBIA, Khomas reg., ~106 km ESE from Windhoek, Arnhem Cave & Lodge territory, Kalahari, on yellow-reddish sand, at night (20:00-0:00). H=1465 m. 13-18.XII.2010
det & photo Vlasenko A. S.
Copyright © Andrey Vlasenko

Anthia (Anthia) circumscripta circumscripta Klug, 1853
Female adult
L=46 mm
NAMIBIA, Hardap reg., 5,3 km W from Rehoboth, Lake Oanob Resort teritory, savanna, at night (20:00-23:00). H=1480 m. 8-10.XII.2010
det & photo Vlasenko A. S.
Copyright © Andrey Vlasenko

Anthia (Anthia) circumscripta circumscripta Klug, 1853
Male adult
L=45 mm
NAMIBIA, Khomas reg., 14 km N from Windhoek, Immanuel Wilderness Lodge, at night, under street lamp. H=1493 m. 28-30.XI.2010
det & photo Vlasenko A. S.
Copyright © Andrey Vlasenko

Anthia (Anthia) circumscripta circumscripta Klug, 1853
Male adult
L=41 mm
NAMIBIA, Khomas reg., 14 km N from Windhoek, Immanuel Wilderness Lodge, at night, under street lamp. H=1493 m. 28-30.XI.2010
det & photo Vlasenko A. S.
Copyright © Andrey Vlasenko

Anthia (Termophilum) limbata kolbei Obst, 1901
Female adult
L=33 mm
NAMIBIA, Khomas reg., ~106 km ESE from Windhoek, Arnhem Cave & Lodge territory, Kalahari, on yellow-reddish sand, at night (20:00-0:00). H=1465 m. 13-18.XII.2010
det & photo Vlasenko A. S.
Copyright © Andrey Vlasenko

Anthia (Anthia) pulcherrima H. W. Bates, 1888
Female adult
L=27 mm
TANZANIA, Iringa Region, Mtwivila env., Mkwawa village env. 18-24.XII.2010
det. & photo Vlasenko A. S.
Copyright © Andrey Vlasenko

Anthia (Anthia) pulcherrima H. W. Bates, 1888
Male adult
L=23 mm
TANZANIA, Iringa Region, Mtwivila env., Mkwawa village env. 18-24.XII.2010
det. & photo Vlasenko A. S.
Copyright © Andrey Vlasenko

Anthia (Termophilum) omoplata omoplata Lequien, 1832
Female adult
L=37 mm
NAMIBIA, Hardap reg., 5,3 km W from Rehoboth, Lake Oanob Resort teritory, savanna, at night (20:00-23:00). H=1480 m. 8-10.XII.2010
det & photo Vlasenko A. S.
Copyright © Andrey Vlasenko

Anthia (Termophilum) omoplata omoplata Lequien, 1832
Male adult
L=41 mm
NAMIBIA, Khomas reg., ~106 km ESE from Windhoek, Arnhem Cave & Lodge territory, Kalahari, on yellow-reddish sand, at night (20:00-0:00). H=1465 m. 13-18.XII.2010
det & photo Vlasenko A. S.
Copyright © Andrey Vlasenko

Anthia (Termophilum) omoplata omoplata Lequien, 1832
Male adult
L=40 mm
NAMIBIA, Khomas reg., ~106 km ESE from Windhoek, Arnhem Cave & Lodge territory, Kalahari, on yellow-reddish sand, at night (20:00-0:00). H=1465 m. 13-18.XII.2010
det & photo Vlasenko A. S.
Copyright © Andrey Vlasenko

Baeoglossa anthracina (Guerin-Meneville, 1847)
Female adult
L=32 mm
NAMIBIA, Khomas reg., ~106 km ESE from Windhoek, Arnhem Cave & Lodge territory, Kalahari, on yellow-reddish sand, at night (20:00-0:00). H=1465 m. 13-18.XII.2010
det & photo Vlasenko A. S.
Copyright © Andrey Vlasenko

Baeoglossa anthracina (Guerin-Meneville, 1847)
Male adult
L=25 mm
NAMIBIA, Khomas reg., 14 km N from Windhoek, Immanuel Wilderness Lodge, at night, under street lamp. H=1493 m. 28-30.XI.2010
det & photo Vlasenko A. S.
Copyright © Andrey Vlasenko

Cypholoba kavanaughi Basilewsky, 1983
Male adult
Angola, N of Caconda, 8.XI.2011, P. Schüle leg.
Photo: Peter Schüle
Copyright © Peter Schüle

Cypholoba sambesina (Peringuey, 1908)
Male adult
ZAMBIA, Kafue N.P., VI.2009, local collector” in PSC
(photo: P. Schüle)
Copyright © Peter Schüle

Cypholoba alstoni Peringuey, 1892
Male adult
Niekerk’s Hope in Griqualand West, Northern Cape Province, RSA
Copyright © ---

Cypholoba opulenta (Boheman, 1860)
Male adult
Us Pass, Khomas Hochland, Namibia
Copyright © ---

Cypholoba macilenta (Olivier, 1795)
Male adult
Groenkloof, Gauteng Province, RSA
Copyright © ---

Cypholoba notata (Perroud, 1846)
Male adult
Pongola River, KwaZulu/Natal Province, RSA
Copyright © ---

Cypholoba fritschi (Chaudoir In Oberthur, 1883)
Male adult
Twee Rivieren, Northern Cape Province, RSA
Copyright © ---

Cypholoba rutata (Peringuey, 1892)
Male adult
Pafuri, Kruger National Park, Limpopo Province, RSA
Copyright © ---

Cypholoba graphipteroides graphipteroides (Guerin-Meneville, 1845)
Male adult
Sabie River west of Paul Kruger Gate in the Kruger National Park
Copyright © ---

Cypholoba oberthueri seruana Strohmeyer, 1928
Male adult
Allemanskraal, Free State Province, RSA
Copyright © ---

Cypholoba leucospilota semilaevis (Chaudoir, 1861)
Male adult
E. Zululand, KwaZulu/Natal Province, RSA
Copyright © ---

Cypholoba tenuicollis aenigma (Dohrn, 1881)
Male adult
Zimbabwe
Copyright © ---

Cypholoba gracilis zuluana Basilewsky, 1948
Male adult
“E. Zululand,” KwaZulu/Natal Province, RSA
Copyright © ---

Cypholoba gracilis gracilis (Dejean, 1831)
Female adult
Vanwyksfontein Farm, Northern Cape Province, RSA
Copyright © ---

Cypholoba gracilis scrobiculata (Bertoloni, 1847)
Female adult
Inhambane, Mozambique
Copyright © ---

Cypholoba gracilis scrobiculata (Bertoloni, 1847)
Male adult
Zoutpansberg, Limpopo Province, RSA
Copyright © ---

Macrocheilus bicolor Andrewes, 1920 Holotype
Female adult
Kaoeqaoe, 740, Type, Macrochilus bicolor Type Andr., H. E. Andrewes det.; H. E. Andrewes Coll. B. M. 1945–97. (NHML)
photo Zhao & Tian, 2012
Copyright © ---

Macrocheilus impictus (Wiedemann, 1823)
Male adult
(MNHN)
photo Zhao & Tian, 2012
Copyright © ---

Macrocheilus immanis Andrewes, 1920 Holotype
Male adult
Toungoo; Type; Macrochilus immanis, Type, Andr., H. E. Andrewes det. (NHML)
photo Zhao & Tian, 2012

Copyright © ---

Macrocheilus niger Andrewes, 1920 Holotype
Female adult
type, 11, H. L. Andrewes, Nilgiri Hills, H. E. Andrewes Coll., B.M. 1945–97.; Macrochilus niger Type, Andr.,
H. E. Andrewes det.; Macrochilus niger Andrewes, 1920 (deposited in NHML)
photo Zhao & Tian, 2012
Copyright © ---

Macrocheilus asteriscus (White, 1844)
Male adult
photo Zhao & Tian, 2012
Copyright © ---

Macrocheilus vitalisi Aandrewes, 1920 Holotype
Female adult
China, Bowring 63·47, 986, 27/2/53 (NHML)
photo Zhao & Tian, 2012
Copyright © ---

Macrocheilus binotatus Andrewes, 1931
Female adult
Paggar Alam, Sumatra, J. Bouchaud (deposited in MNHN)
photo Zhao & Tian, 2012
Copyright © ---

Macrocheilus gigas Zhao & Tian, 2010 Holotype
Male adult
Guangdong: Zhanjiang, Jun.1983 (deposited in SCAU)
Photo Zhao & Tian, 2012
Copyright © ---

Macrocheilus parvimaculatus Zhao & Tian, 2010 Holotype
Male adult
Guangxi: Liuzhou: Luzhai, 26 May 1980, Shaozhou Ruan leg. (SCAU)
photo Zhao & Tian, 2012
Copyright © ---

Macrocheilus tripustulatus (Dejean, 1825) Holotype
Female adult
tripustulatus Dejean (non Fabr.), Java, Coll. Dejean, 3-pustulatus Wiedemann (Helluo). Fabr. (Brachinus), Western, Ex-Musaeo Chaudoir (MNHN)
photo Zhao & Tian, 2012
Copyright © ---

Macrocheilus chaudoiri Andrewes, 1919 Holotype
Female adult
Ex-Musaeo Chaudoir; Macrocheilus Chaudoiri Andr., H.S. Andrewes det.; 3-maculatus Chaud., Deccan, Coll. Jeakes (MNHN) photo Zhao & Tian, 2012
Copyright ©

Macrocheilus nigrotibialis Heller, 1900 Holotype
Male adult
Drs. Sarasin N. Celebes Panot-Maimang, “12623”, “Staatl. Museum fur Tierkunde. Dresden” (deposited in SNSD)
photo Zhao & Tian, 2012
Copyright © ---

Macrocheilus bensoni Hope, 1838
Male adult
photo Zhao & Tian, 2012
Copyright © ---

Macrocheilus deuvie Zhao & Tian, 2012 Holotype
Male adult
Philippines, Bohol. Ch. Semper (deposited in MNHN)
photo Zhao & Tian, 2012
Copyright © ---

Macrocheilus fuscipennis Zhao & Tian, 2010 Holotype
Male adult
Guangxi: Napo, 10 Oct.1970, by light trap
photo Zhao & Tian, 2012
Copyright © ---

Macrocheilus solidipalpis Zhao & Tian, 2010 Holotype
Male adult
Guangxi: Dibei, Oct.1980, Xiuzhen Mao leg., by light trap.
photo Zhao & Tian, 2012
Copyright © ---

Macrocheilus cheni Zhao & Tian, 2010 Holotype
Male adult
Guangxi: Tengxian, Oct.1980
Photo Zhao & Tian, 2012
Copyright © ---

Macrocheilus quadratus Zhao & Tian, 2010 Holotype
Male adult
Guangxi: Cangwu, 1980, in paddy field (SCAU)
photo Zhao & Tian, 2012
Copyright © ---

Macrocheilus sinuatilabris Zhao & Tian, 2010 Holotype
Male adult
Guangxi: Fenghuangcheng, Sep.1981, by light trap (SCAU)
photo Zhao & Tian, 2012
Copyright © ---

Anthia (Termophilum) nimrod (Fabricius, 1793)
Male adult
Sénegal, N’Gazobil, sous tas de bois 22/09/2099, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Macrocheilus biguttatus Gory, 1832
Female adult
Senegal, M’Bour Nianing, piège lumineux 24/09/2009, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Netrodera formicaria (Erichson, 1843)
Female adult
NAMIBIA, Khomas reg., 35.3 km SE from Windhoek, Auas Game Lodge territory H=1728 m 22°47.454'S 17°20.022' E 28.XI.2012 leg. E.Shankhiza
L=13.5 mm

Copyright © Kirill Makarov

Anthia (Termophilum) omoplata Lequien, 1832
Male adult
NAMIBIA, Oshana reg., 67,5 km NNW from Outjo, Toshari Lodge territory, H=1205 m S19°33.390' E15°52.679' 2-5.XII.2012 leg. E.Shankhiza
L=37.0 mm
Copyright © Kirill Makarov

Anthia (Termophilum) limbata Dejean, 1831
Male adult
NAMIBIA, Khomas reg., 35.3 km SE from Windhoek, Auas Game Lodge territory H=1728 m 22°47.454'S 17°20.022' E 28.XI.2012 leg. E.Shankhiza
L=30 mm
Copyright © Kirill Makarov

Anthia (Termophilum) aequilatera Klug, 1853
Male adult
C Tanzania, SW env. of Dodoma City XII.2011 leg. A.Shubin L=27 mm
Copyright © Kirill Makarov

Cypholoba tenuicollis (Chaudoir, 1878)
Male adult
C Tanzania, SW env. of Dodoma City XII.2011 leg. A.Shubin L=17 mm
Copyright © Kirill Makarov

Cypholoba caillaudi biscutellata (Fairmaire, 1887)
Female adult
C Tanzania, SW env. of Dodoma City XI.2011 leg. A.Shubin L=29 mm
Copyright © Kirill Makarov

Cypholoba caillaudi biscutellata (Fairmaire, 1887)
Male adult
C Tanzania, SW env. of Dodoma City XI.2011 leg. A.Shubin L=27 mm
Copyright © Kirill Makarov

Eccoptoptera cupricollis Chaudoir, 1878
Male adult
C Tanzania, SW env. of Dodoma City XII.2011 leg. A.Shubin L=18 mm
Copyright © Kirill Makarov

Anthia (Anthia) circumscripta circumscripta Klug, 1853
Male adult
C Tanzania, SW env. of Dodoma City I.2012 leg. A.Shubin L=42 mm
Copyright © Kirill Makarov

Omphra atrata (Klug, 1834)
Male adult
India, Goa, Cotigao Wildlife Senctuary, 25.03.2013, P. & L. Ustjuzhanin (ISEA)
R. Dudko det. et photo, 2013
Copyright © Roman Dudko

Cypholoba perspicillaris (Chaudoir, 1878)
Male adult
26 mm
Kenya, Tsavo National Park, Voi Lodge, 500-600 m, leg. U.Schmidt, 24.-26.XI.1991; det. J.Háva, 2011
Photo: U.Schmidt, 2013
Copyright © Udo Schmidt

Cypholoba cinereocincta (Fairmaire, 1885)
Male adult
32 mm
Kenya, Tsavo National Park, Voi Lodge, 500-600 m, leg. U.Schmidt, 24.-26.XI.1991; det. J.Háva, 2011
Photo: U.Schmidt, 2013
Copyright © Udo Schmidt

Foveocrotaphus burmanicus Anichtchenko, 2014 Holotype
Male adult
Myanmar N (Burma), 65 km NE Putao, 1250m, Zi Yar Dam vill., 18-21.05.1998, leg. S. Murzin & V. Siniaev (DUBC). 17 mm
Copyright © Alexander Anichtchenko

Gigadema obscurum Sloane, 1914
Male adult
Australia, Queensland, Brisbane. Image from: Mal, N., Drumont, A., Kerkhof, S. & P. Grootaert, 2012. Pictorial beetle collection of the Royal Belgian Institute of Natural Sciences. det. Anichtchenko A. 2020
Copyright ©

Anthia (Termophilum) venator (Fabricius, 1792)
Male adult
Lybia, Tajura, 16.x.1981-J. Roggeman Leg., 50mm, Coll. N. Mal
Copyright © Noël Mal

Anthia (Termophilum) nimrod nimrod (Fabricius, 1793)
Male adult
Senegal : Casamance, Sedhiou, 1891 - E. Laglaise, 41 mm, Coll. RBINS
Copyright © Noël Mal

Anthia (Termophilum) sulcata voltae Basilewsky, 1947
Male adult
Haute Volta : Ouagadagoudou, vii.1976, 35 mm, Coll. N. Mal
Copyright © Noël Mal

Anthia (Anthia) circumscripta circumscripta Klug, 1853
Male+Female adult
South Africa : Windhoeck, 52/43mm, Coll. N. Mal
Copyright © Noël Mal

Anthia (Anthia) cinctipennis cinctipennis Lequien, 1832
Male adult
South Rhodesia : Hilside, 11.xi.1922, Swimburne & Stevenson,45mm, Coll. RBINS
Copyright © Noël Mal

Anthia (Anthia) lunae lunae J.Thomson, 1859
Male adult
Congo : P.N. Upemba, ii/ee/3, 20.ix.1951, Mission H? de Saeger, 36mm , Coll. RBINS
Copyright © Noël Mal

Anthia (Termophilum) sexmaculata sexmaculata (Fabricius, 1787)
Male+Female adult
Morocco : Talfilalt-rissani iv.1981 - V. Oliviella Leg. (Size : 27-33 mm) Coll. N. Mal
Copyright © Noël Mal

Anthia (Termophilum) sulcata voltae Basilewsky, 1947
Male adult
Haute Volat : Ouagadagoudou vii.1976 (Size : 35 mm) Coll. N. Mal
Copyright © Noël Mal

Anthia (Anthia) thoracica thoracica (Thunberg, 1784)
Male adult
Rhodesia : Umali District, Mpudzi River, 10.i.1928,R. Stevenson, 42 mm, Coll. RBINS
Copyright © Noël Mal

Baeoglossa villosa (Thunberg, 1806)
Female adult
South Africa, Transvaal, ex Coll. Steinmetz, 30 mm, Coll. RBINS
Copyright © Noël Mal

Cypholoba notata (Perroud, 1846)
Male adult
South Africa : N. Krüger N.P., M. Shiloe x.1908 (Size : 19 mm), Coll. N. Mal
Copyright © Noël Mal

Atractonotus puncticollis Schüle & Heins, 2013 Holotype
Male adult
Zambia, North Western pr., 65 km N Kabompo, 14-15.XII.2007, 1110m, Ivo Martinu leg. (cPS).
Copyright © Peter Schüle

Helluodema brunneum Sloane, 1917 Syntypes
Male adult

Copyright © Museums Victoria

Gigadema minutum Laporte, 1867 Holotype
Male adult

Copyright © Museums Victoria

Helluapterus niger Sloane, 1914 Syntypes
Male adult

Copyright © Museums Victoria

Gigadema biordinatum Sloane, 1914 Syntypes
Male adult

Copyright © Museums Victoria

Gigadema mandibulare Blackburn, 1892 Syntypes
Male adult

Copyright © Museums Victoria

Pogonoglossus rufopiceus Baehr, 1993 Holotype
Male adult

Copyright © Museums Victoria

Erephognathus ranomafanae Kavanaugh and Rainio, 2016 Holotype
Female adult
MADAGASCAR: Province Fianarantsoa, Parc National Ranomafana, Vohiparara, at broken bridge, el 1110 m 8-15 November 2001”/ “21°13.57’S, 047°22.19’E collector: R. Harin’Hala California Acad of Sciences malaise trap in high altitude rainforest, MA-02-09A-02”; HOLOTYPE Erephognathus ranomafanae Kavanaugh & Rainio sp. n. 2015 (CAS). Image from Kavanaugh & Rainio, 2016
Copyright © - - -

Cypholoba tetrastigma tetrastigma (Chaudoir, 1848)
Male adult
S Ethiopia, Oromia reg., Borena Distr., Negele, 1343m, XI.2014. 20mm
Copyright © Alexander Anichtchenko

Colfax stewensi Andrewes, 1920
Male adult
10-11 mm Nekka Nalla, Ratnagiri District, Maharashtra, India, February 1973, M. Baburao leg. det. Danyang Zhao
Copyright © Shripad V. Manthen

Macrocheilus niger Andrewes, 1920
Male adult
16-16.50 mm Latitude 17.73625, Longitude 73.76983, Elevation (m) 671 Mhavashi, Satara District, Maharashtra, India, March 2017, leg. S.R Patil & Party; det. Shripad Manthen
Copyright © Shripad V. Manthen

Omphra complanata Reiche, 1843
Male adult
14 mm; GPS: Latitude 17.71172, Longitude 73.73987, Elevation (m) 773 Vengale, Satara District, Maharashtra, India, March 2017, leg. S.R Patil & Party; det. Shripad Manthen
Copyright © Shripad V. Manthen

Helluo insignis Sloane, 1890 Paratype
Male adult

Copyright © - - -

Neohelluo angulicollis Sloane, 1914
Male adult
Australia. 24 mm
Copyright © Mark Golding

Pleuracanthus sulcipennis Gray, 1832
Male adult
Bolivia, Dep. Santa Cruz, San Ignacio, 11.1996, leg. A. Mater (Collection R. Sciaky). 15 mm
Copyright © Alexander Anichtchenko

Dicranoglossus resplendens (Castelnau, 1867)
Male adult
Australia, Quennsl. Cloncurry, III.1993; 11mm (Collection R. Sciaky)
Copyright © Alexander Anichtchenko

Helluodes taprobanae Westwood, 1847
Female adult
Sri Lanka, Kandy, 1.1.1990, leg. Rautenstrauch. det. A. Anichtchenko, 2017. (Collection R. Sciaky) 17mm
Copyright © Alexander Anichtchenko

Meladroma angustata Basilewsky, 1970
Male adult
Rep. Centr. Africa, Maboke, M. Gombe, Doko, XI. 1985. (Collection and det by R. Sciaky as informicollis Liebke, 1928). 23mm ; det. Peter Schüle 2022
Copyright © Alexander Anichtchenko

Pogonoglossus danielsi Baehr, 2007 Holotype
Male adult
8 km S Mendooran, NSW 31’53’S 149’03’E 12 Feb 1992 mv lamp G. Daniels C. J. Burwell (UQIC Reg #90394) (QMT 123824). 8.4mm; Image from Baehr, 2007.
Copyright © - - -

Helluodes devagiriensis Sabu, Abhitha & Zhao, 2008 Holotype
Male adult
India,. Kerala, Arakulam (Thodupuzha) 17. V. 2004, Sabu T.K, rubber plantation litter (deposited in the Coleoptera collections of SJC). Image from Sabu, Abhitha & Zhao, 2008
Copyright © - - -

Pogonoglossus giganteus Baehr, 2005 Holotype
Male adult
Image from Sabu, Abhitha & Zhao, 2008
Copyright © - - -

Schuelea drumonti Baehr, 2004 Holotype
Female adult
Coll. I.R.Sc.N.B. Canopy mission P.N.G. Madang province, Baiteta, FOG AR 3-10 4.V. 1995, Leg. Olivier Missa (IRSNB). Image from Sabu, Abhitha & Zhao, 2008
Copyright © - - -

Physocrotaphus ceylonicus Parry, 1849 Lectotype
Male adult
(MNHN). Image from Sabu, Abhitha & Zhao, 2008
Copyright © - - -

Aenigma iris Newman, 1836
Female adult
Australia, Queensland, M. Garnet, 26.II.1992.
Copyright © Riccardo Sciaky

Helluodes taprobanae Westwood, 1847
Male adult
Sri Lanka, Kandy, 1.1.1990, leg. Rautenstrauch. det. A. Anichtchenko, 2017. (Collection R. Sciaky)
Copyright © Alexander Anichtchenko

Anguloderus mirabilis Anichtchenko & Sciaky, 2017 Holotype
Male adult
West Papua, Fak-Fak region, Fak-Fak, VI.2001 (RSC).
Copyright © Alexander Anichtchenko

Anguloderus mirabilis Anichtchenko & Sciaky, 2017 Holotype
Male adult
West Papua, Fak-Fak region, Fak-Fak, VI.2001 (RSC). 21 mm (without mandibles)
Copyright © Alexander Anichtchenko

Anthia (Anthia) kleinfeldi Puchner, 2013 Paratype
Female adult

Copyright © Alexander Dostal

Anthia (Anthia) kleinfeldi Puchner, 2013 Holotype
Male adult

Copyright © Alexander Dostal

Anthia (Anthia) calida Harold, 1878 Lectotype
Male adult

Copyright © Alexander Dostal

Anthia (Anthia) calida Harold, 1878
Female adult
Mayidi (Tervuren)
Copyright © Alexander Dostal

Anthia (Anthia) convexipennis Putzeys, 1880
Female adult

Copyright © Alexander Dostal

Anthia (Anthia) convexipennis Putzeys, 1880
Male adult

Copyright © Alexander Dostal

Dailodontus clandestinus (Klug, 1834)
Male adult
Paraguay, Dep. Pte. Hayes, Gral. Bruguez, S24,44 W58,48 19-23.I.2006 leg. U. Drechsel. 12 mm. det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Helluomorpha eulinae Reichardt, 1974
Female adult
Paraguay, Dep Cordillera, Pirareta, 6-8.10.2012 U. Drechsel leg. 17 mm; det. Anichtchenko A. 2019
Copyright © Alexander Anichtchenko

Helluomorpha macroptera Chaudoir, 1850
Male adult
Paraguay, dep Guira, Ita Azul, 7-11.1.2011, U. Drechsel leg. det. A. Anichtchenko 2019
Copyright © Alexander Anichtchenko

Helluosoma cf. bouchardi Baehr, 2005
Male adult
Australia (coll. R. Sciaky)
Copyright © Riccardo Sciaky

Macrocheilus asteriscus (White, 1844)
Female adult
Laos
Copyright © Alexander Anichtchenko

Helluomorphoides squiresi (Chaudoir, 1872)
Male adult
Paraguay, Dep. Caazapa, Colonia Neufeld, 24.10-2.11.2008. 12 mm. det. Anichtchenko A. 2019
Copyright © Alexander Anichtchenko

Helluomorphoides rubricollis (Schaum, 1863)
Male adult
Paraguay, Dep. Caazapa, La Golondrina, S25,32 W55,29 14-19.III.2009 leg. U. Drechsel. det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Helluobrochus darlingtoni Reichardt, 1974
Male adult
Paraguay, dep. Caazapa, Colonia Neufeld, S26.28 W55.55, 24.10.2008, leg. U. Drechsel 15 mm; det. Anichtchenko A. 2019
Copyright © Alexander Anichtchenko

Macrocheilus chinnarensis Akhil, Divya & Sabu, 2019 Holotype
Male adult
Holotype, Male, India: Kerala: Chinnar, 420 m, 08.viii.2016; coll. S.V. Akhil . Image from Akhil, Divya & Sabu, 2019
Copyright © - - -

Macrocheilus bandipurensis Akhil, Divya & Sabu, 2019 Holotype
Male adult
Holotype, Male, India: Karnataka: Gundlupet, 06.v.2016; coll. S.V. Akhil. Image from Akhil et al, 2019
Copyright © - - -

Atractonotus mulsanti Perroud, 1846
Male adult

Copyright © Riccardo Sciaky

Anthia (Anthia) mannerheimi afghana Anichtchenko, 2009
Female adult
Afghanistan, Oruzgan, Sahrestan, h = 2500 m, 03-05.VII.1970, leg. O.N. Kabakov (TOPOTYPUS)
Copyright © Igor Solodovnikov

Anthia (Termophilum) duodecimguttata Bonelli, 1813
Male adult
CW IRAN, Khuzestan prov., Ahvaz County, Central District, Elhayi Rural District, 31 km N Ahvaz, near Borvayeh vill., Kerkhe riv. valley, Deshte-Mishen desert, h=31m, 31°35'40.80"N / 48°35'56.79"E, 9.V.2019, Yu.Skrylnik leg. L =35 mm
Copyright © Ruslan Panin

Anthia (Termophilum) duodecimguttata Bonelli, 1813
Female adult
CW IRAN, Khuzestan prov., Ahvaz County, Central District, Elhayi Rural District, 31 km N Ahvaz, near Borvayeh vill., Kerkhe riv. valley, Deshte-Mishen desert, h=31m, 31°35'40.80"N / 48°35'56.79"E, 9.V.2019, Yu.Skrylnik leg. L =36 mm
Copyright © Ruslan Panin

Anthia (Termophilum) sexmaculata (Fabricius, 1787)
Female adult
Morocco S Ouarzazate Prov., Tagounite env., sands 30°01'21"N 005°32'09"E 29-30.III.2011 leg. A.Gusakov L=33.0 mm
Copyright © Kirill Makarov

Cypholoba tenuicollis cf. salamensis (Kolbe, 1898)
Male adult
Tanzania, Morogoro area, Mvomero distr., Kibaoni v., 1.12.2001 G.Mpoyda leg. (DUBC)
Copyright © Alexander Anichtchenko

Cypholoba posticalis posticalis (Fairmaire, 1885)
Male adult
Somalia, Mogadiscio, Afgoi 22.4-5.5.1988
Copyright © Erich Kirschenhofer

Cypholoba tenuicollis kafuena (Peringuey, 1908)
Male adult
Zambia, Lusaka, Kafue city, Karue river, 22.11.1987
Copyright © Erich Kirschenhofer

Cypholoba rutata (Peringuey, 1892)
Male adult
Zambia, Lusaka, Kafue city, Karue river, 22.11.1987
Copyright © Erich Kirschenhofer

Cypholoba chanleri (Linell, 1896)
Male adult
Ostafrica, Waboniland
Copyright © Erich Kirschenhofer

Cypholoba trilunata (Gerstaecker, 1884)
Male adult
Brit. O. Africa, Athi
Copyright © Erich Kirschenhofer

Cypholoba semibrunnea Strohmeyer, 1928
Male adult
Reimer 1897, Mikindani, D.O. Africa
Copyright © Erich Kirschenhofer

Baeoglossa villosa (Thunberg, 1806)
Male adult

Copyright © Igor Solodovnikov

Anthia (Termophilum) legalli Lassalle, 2016 Holotype
Male adult
Dikomé Balué, env. de Kumba, XI.09, Fogoh John Muafor lg.; in coll. Institut de Recherche, Yaoundé.
Copyright © Bernard Lassalle

Anthia (Termophilum) sulcata adelpha J.Thomson, 1859
Male adult
Tchad : Moyen Chari, Ft de Manda et de Djoli Kera
Copyright © Bernard Lassalle

Anthia (Termophilum) galla lesnei Sternberg, 1906
Male adult
Cameroun : reg. de l’Adamaoua, Wak près de Ngaoundéré.
Copyright © Bernard Lassalle

Anthia (Anthia) maxillosa (Fabricius, 1793)
Male adult
S.Africa: RSA: Northern Cape, 80 km SW Springbok; 100 km W Urington town; Namaqua, Skilpad, 450 m, 51,5 mm
Copyright © Igor Solodovnikov

Anthia (Anthia) maxillosa (Fabricius, 1793)
Female adult
S.Africa: RSA: Northern Cape, 80 km SW Springbok; 100 km W Urington town; Namaqua, Skilpad, 450 m, 45 mm
Copyright © Igor Solodovnikov

Anthia (Termophilum) nimrod nimrod (Fabricius, 1793)
Female adult
Tchad, Abou Telfan près de Mongo
Copyright © Bernard Lassalle

Anthia (Anthia) lunae lunae J.Thomson, 1859
Male adult
Tchad, Ft de Manda. 39mm
Copyright © Bernard Lassalle

Anthia (Anthia) mirabilis Sternberg, 1906
Male adult
Gabon sept. 25 mm
Copyright © Bernard Lassalle

Eccoptoptera cupricollis viri Strohmeyer, 1928
Male adult
Tchad, Ft de Manda. 17mm
Copyright © Bernard Lassalle

Cypholoba tenuicollis crampelina Strohmeyer, 1928
Male adult
Cameroun sept., Rhumsiki. 18mm
Copyright © Bernard Lassalle

Cypholoba tenuicollis telfanensis Lassalle, 2016 Holotype
Female adult
Tchad, Guera, Abou Telfan près de Mongo, 20.VII.2014, Lassalle et Vanderbergh leg. (in coll. Lassalle).
Copyright © Bernard Lassalle

Cypholoba reflexicauda (Benard, 1925) Holotype
Male adult
République Centrafricaine : Hte-Sangha, environs de Carnot.
Copyright © Bernard Lassalle

Anthia (Termophilum) massilicata Guerin-Meneville, 1845
Male adult

Copyright © Igor Solodovnikov

Anthia (Termophilum) massilicata Guerin-Meneville, 1845
Female adult

Copyright © Igor Solodovnikov

Omphra hirta (Fabricius, 1801)
Male adult
S India, Kerala, Trivandrum Dt., Poonmudi mg., 3000 ft, 5.1989 (cAA)
Copyright © Alexander Anichtchenko

Meladroma umbraculata (Fabricius, 1801)
Male adult
Cote d'Ivoire, Lamto Toumodi, 23-08-1971
Copyright © Pascal Deschamps

Omphra erwini Akhil & Sabu, 2021 Holotype
Male adult
India: Gujarat: Ambaji, 19.ii.1992, coll. ZSI-DRC, det. Akhil S. V. Image from Akhil & Sabu, 2021. Aedeagus deformed.
Copyright © - - -

Omphra balli Akhil & Sabu, 2021 Holotype
Male adult
India: Telangana: Amrabad TR: View point Farhabad, 19.ix.2018, coll. S. S. Jadhav, det. Akhil S. V. Image from Akhil & Sabu, 2021
Copyright © - - -

Macrocheilus saulcyi Chevrolat, 1854
Female adult
Jordan, Petra (cAA). det. A. Anichtchenko, 2021
Copyright © Alexander Anichtchenko

Macrocheilus cambodianus Allegro & Giachino, 2021 Holotype
Male adult
Cambodia, Banteay Meanchey Province, near Sisophon, campus of the Mean Chey University, 13°14’45”n, 20°56’29”E, light trap, 22.x-23.xI.2019, P. Bun & V. Kong leg. (CAl).
Copyright © Gianni Allegro

Macrocheilus walterrossii Allegro & Giachino, 2021 Holotype
Male adult
Cambodia, Banteay Meanchey Province, near Sisophon, campus of the Mean Chey University, 13°14’45”n, 20°56’29”E, light trap, 22.x-23.xI.2019, P. Bun & V. Kong leg. (CGi).
Copyright © Gianni Allegro

Cypholoba cf intricata intricata (C.A.Dohrn, 1882)
Male adult
det. Peter Schüle 2021
Copyright © Igor Solodovnikov

Netrodera formicaria formicaria (Erichson, 1843)
Female adult

Copyright © Igor Solodovnikov

Cypholoba macilenta (Olivier, 1795)
Male adult

Copyright © Igor Solodovnikov

Cypholoba graphipteroides bilunata (Boheman, 1860)
Female adult

Copyright © Igor Solodovnikov

Cypholoba trilunata trilunata (Gerstaecker, 1884)
Male adult

Copyright © Igor Solodovnikov

Cypholoba trilunata trilunata (Gerstaecker, 1884)
Female adult

Copyright © Igor Solodovnikov

Cypholoba alveolata alveolata Breme, 1844
Male adult
S.Afr.: P.Natal, Kaffari
Copyright © Igor Solodovnikov

Gonogenia immerita immerita (Boheman, 1860)
Female adult

Copyright © Igor Solodovnikov

Anthia (Termophilum) sexmaculata marginata Latreille, 1823
Male adult
Mauritania
Copyright © Igor Solodovnikov

Anthia (Termophilum) sexmaculata marginata Latreille, 1823
Female adult
Anthia (Thermophilum) sexmaculata pro marginata Latreille, 1823 Sudan
Copyright © Igor Solodovnikov

Eccoptoptera mutilloides mutilliodes (Bertoloni, 1857)
Male adult
Eccoptoptera mutilloides mutilloides f.labrata Fairmaire, 1891 Republic of Mozambique
Copyright © Igor Solodovnikov

Anthia (Termophilum) sexmaculata sexmaculata (Fabricius, 1787)
Male adult

Copyright © Igor Solodovnikov

Anthia (Termophilum) sexmaculata sexmaculata (Fabricius, 1787)
Female adult

Copyright © Igor Solodovnikov

Anthia (Termophilum) ovampoa Peringuey, 1896
Female adult
C-Namibia, Otjozondjuba Region, 28 km E Otavi town, h = 1400 m
Copyright © Igor Solodovnikov

Anthia (Termophilum) sulcata voltae Basilewsky, 1947
Female adult
Afr.: Fort Archambault
Copyright © Igor Solodovnikov

Anthia (Termophilum) aequilatera Klug, 1853
Male adult
Tanzania: Dodoma, Mapinduzi
Copyright © Igor Solodovnikov

Anthia (Termophilum) discedens Sternberg, 1907
Male adult

Copyright © Igor Solodovnikov

Anthia (Termophilum) andersoni latelimbatum Kleinfeld, Puchner, 2012
Female adult
Zambia, Central Province, Chimoto vill. env., H=1150 m (TOPOTYPUS)
Copyright © Igor Solodovnikov

Anthia (Termophilum) andersoni latelimbatum Kleinfeld, Puchner, 2012
Male adult
Zambia, Central Province, Chimoto vill. env., H=1150 m (TOPOTYPUS)
Copyright © Igor Solodovnikov

Anthia (Termophilum) brevivittata Obst, 1901
Male adult
Zambie: Mfuenta, 1400m, Сentral Prov.
Copyright © Igor Solodovnikov

Anthia (Termophilum) namaqua Peringuey, 1896
Male adult

Copyright © Igor Solodovnikov

Anthia (Anthia) lefebvrei Guerin-Meneville, 1849
Male adult
South Somali: riv. Dzhuba (Jubba River)
Copyright © Igor Solodovnikov

Anthia (Anthia) sexguttata (Fabricius, 1775)
Male adult
India
Copyright © Igor Solodovnikov

Anthia (Anthia) mannerheimi mannerheimi Chaudoir, 1842
Male+Female adult
S.Iran: Sistan & Beloujistan prov., 20 km W Chobahor
Copyright © Igor Solodovnikov

Anthia (Anthia) mannerheimi mannerheimi Chaudoir, 1842
Female adult
Uzbekistan (border Turkmenistan)
Copyright © Igor Solodovnikov

Anthia (Anthia) mannerheimi mannerheimi Chaudoir, 1842
Male adult
Pakistan: Sindh, Shirak
Copyright © Igor Solodovnikov

Anthia (Anthia) mannerheimi mannerheimi Chaudoir, 1842
Male+Female adult
S-Turkmenistan
Copyright © Igor Solodovnikov

Helluonidius cyanipennis (Hope, 1842) Holotype
Male adult

Copyright © - - -

Ametroglossus ater (W.J.Macleay, 1887)
Male adult

Copyright © Creative Commons

Helluosoma atrum Castelnau, 1867
Male adult

Copyright © Creative Commons

Helluodema unicolor (Hope, 1842)
Male adult
Australia
Copyright © Creative Commons

Helluonidius aterrimus (W.J.Macleay, 1873)
Male adult

Copyright © Creative Commons

Triaenogenius (Triaenogenius) lugubrinus (Boheman, 1860)
Male adult
SW Africa, Kaokoveld, Sanitatas abr 85 miles WSW Ohopoho 14-16.6.1951 Swedish Sout Africa Expedition (cAA) 18 mm
Copyright © Alexander Anichtchenko

Epimicodema mastersii W.J.Macleay, 1871
Male adult

Copyright © - - -

Anthia (Termophilum) omoplata incolata Peringuey, 1892
Male adult
RSA, Kwazulu Natal, Pongola, h = 430 m
Copyright © Igor Solodovnikov

Anthia (Termophilum) omoplata incolata Peringuey, 1892
Female adult
RSA, Kwazulu Natal, Pongola, h = 430 m
Copyright © Igor Solodovnikov

Anthia (Termophilum) decemguttata (Linnaeus, 1764)
Male adult
RSA: W Cape, 7 rm E Calitzdorp 42,5 mm
Copyright © Igor Solodovnikov

Anthia (Termophilum) decemguttata (Linnaeus, 1764)
Female adult
RSA: W Cape, 7 rm E Calitzdorp 40 mm
Copyright © Igor Solodovnikov

Anthia (Termophilum) decemguttata (Linnaeus, 1764)
Male adult
26,55 mm
Copyright © Igor Solodovnikov

Anthia (Termophilum) decemguttata (Linnaeus, 1764)
Male adult
30,65 mm
Copyright © Igor Solodovnikov

Anthia (Termophilum) decemguttata (Linnaeus, 1764)
Male adult
36,5 mm
Copyright © Igor Solodovnikov

Anthia (Chilanthia) cavernosa Gerstaecker, 1866
Male adult
Kenya
Copyright © Igor Solodovnikov

Atractonotus mulsanti Perroud, 1846
Female adult
South Africa, Mkuze GR, Mantuma Camp 15.IX.2022 L=11.5
Copyright © Kirill Makarov

Baeoglossa villosa (Thunberg, 1806)
Female adult
South Africa, Karoo Padstal, 20 km NE Richmond 26.IX.2022 L=27.0
Copyright © Kirill Makarov

Cypholoba suturella (Chaudoir, 1866)
Male adult
Mikindani, D.O. Africa, Reimer 1897 (NMW)
Copyright © Erich Kirschenhofer

Cypholoba amatonga (Peringuey, 1892)
Male adult
Soutpansberg, Limpopo Province, RSA. Image from Mawdsley J.R. et al 2012
Copyright © - - -

Cypholoba puchneri Schüle, 2015 Holotype Paratype
Male+Female adult
ANGOLA, 27.XI.13, Cunene Prov. 20km NE Cahama, 1315m P. Schüle leg. (PSC).
Copyright © Peter Schüle

Cypholoba bottegoi G.Müller, 1938
Male adult
Somali, Daarbuduq. Image from Tomas Mazuch et al.
Copyright © - - -

Cypholoba caillaudi caillaudi (Castelnau, 1835)
Male adult
Somalia, Quljeet. Image from Tomas Mazuch et al.
Copyright © - - -

Cypholoba interrupta (Fairmaire, 1887)
Male adult
Somalia, Shanshacade. Image from Tomas Mazuch et al.
Copyright © - - -

Cypholoba cinereocincta cinereocincta (Fairmaire, 1885)
Male adult
Somalia, Shanshacade. Image from Tomas Mazuch et al.
Copyright © - - -

Cypholoba posticalis (Fairmaire, 1885)
Male adult
Somalia, Shanshacade. Image from Tomas Mazuch et al.
Copyright © - - -

Cypholoba obtusata (Fairmaire, 1887)
Female adult
Somalia, Daalo Forest, N of Erigavo. Image from Tomas Mazuch et al.
Copyright © - - -

Cypholoba tetrastigma tetrastigma (Chaudoir, 1848)
Male adult
Somalia, Boorama. Image from Tomas Mazuch et al.
Copyright © - - -

Anthia (Termophilum) revoili Lucas, 1881
Female adult
Somalia, 4.5 km NNW of Erigavo. Image from Tomas Mazuch et al.
Copyright © - - -

Anthia (Anthia) ferox J.Thomson, 1859
Male adult
Somalia, Boorama. Image from Tomas Mazuch et al.
Copyright © - - -

Anthia (Termophilum) revoili Lucas, 1881
Male adult
Somalia, Boorama. Image from Tomas Mazuch et al.
Copyright © - - -

Anthia (Anthia) ferox J.Thomson, 1859
Female adult
Somalia, 4.5 km NNW of Erigavo. Image from Tomas Mazuch et al.
Copyright © - - -

Anthia (Termophilum) fornasinii Bertoloni, 1845
Male adult
Anthia (Termophilum) fornasinii ssp.
Copyright © Igor Solodovnikov

Anthia (Termophilum) omoplata omoplata Lequien, 1832
Male adult
SAR, Mokala N.P., h = 1114 m, L = 36 mm
Copyright © Igor Solodovnikov

Anthia (Termophilum) omoplata omoplata Lequien, 1832
Male adult
Anthia (Termophilum) omoplata ? pro parva (SAR, Mokala N.P., h = 1114 m), 41 mm
Copyright © Igor Solodovnikov

Anthia (Termophilum) hexasticta Gerstaeker, 1866
Male adult
Kenya, Malandi, Gede, 4.2011
Copyright © Alexander Anichtchenko

Physoglossus devagiriensis Akhil & Sabu, 2020 Holotype
Male adult
India: Tamil Nadu: Dindigul dist: Kumbakarai falls; 369 m; coll. R. Babu; 21.iii.2016;
Copyright © Sabu K Thomas

Physoglossus devagiriensis Akhil & Sabu, 2020 Holotype
Male adult

Copyright © Sabu K Thomas

Cypholoba basilewskyi Schüle & Heinz, 2013 Paratype
Male+Female adult
Namibia, Kunene, 10-15 km SE Opuwo, 1200/1400 m 19.1.2012, HEINZ leg. Scale 5 mm
Copyright © Peter Schüle

Cypholoba heinzorum Schüle, 2013 Holotype
Male adult
ZAMBIA, North Western pr.\ 40 km SW Kabompo\ 15.Xll.2007\ lvo Martinu leg., 1110m (PSC)
Copyright © Peter Schüle

Macrocheilus biguttatus Gory, 1832
Female adult
Afrique de L`Ouest, Senegal Petit Cote, Mbour 20.IX.2009 IRD, Ultra Violet, Alain Coache
Copyright © Pascal Deschamps

Macrocheilus biguttatus Gory, 1832
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus allardi Basilewsky, 1957 Holotype
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus alluaudi Burgeon, 1937 Holotype
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus alluaudi Burgeon, 1937
Male adult
Republic of Congo, Réserve Naturelle des Gorilles de Lésio Louna 17–28.11.2017, Batéké Plateau H. Krahnstôver leg. (Collection P. Schüle, Herrenberg). Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus assmanni Serrano, 2023 Holotype
Female adult
Tanzania, 12-17.XII.1996\ Ruvuma, near Songea\ leg. Werner & Lizler. Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus angustatus Basilewsky, 1949 Holotype
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus basilewskyi A.Serrano, 2000 Holotype
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus bimaculatus (Dejean, 1831)
Male adult
Quebo (Coli), 28/06/1995, Col. A. Serrano\\ Miss. Ent. SPEN, à Guiné-Bissau, 25 Junho a 17 Julho 95. Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus biplagiatus (Boheman, 1848)
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus burgeoni Basilewsky, 1967 Holotype
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus clasispilus Basilewsky, 1967 Holotype
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus crampeli Alluaud, 1916 Paratype
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus cruciatus (Marc, 1840)
Male adult
Image from A. Serrano, 2023
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Macrocheilus diplospilus Basilewsky, 1967 Holotype
Male adult
Image from A. Serrano, 2023
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Macrocheilus dorsalis Klug, 1834
Male adult
Zambia NC., 82 km SSW Solwezi, 3.12.2004, Snižek, Tichẏ lgt. Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus dorsiger (Chaudoir, 1876)
Male adult
Burkina Faso: Nahouri, Forêt de Nazinga, Akwazena, 275 m, 11º 09′ 24″ N 001º 36′ 44″ O, 21.VII.2006, zone Soudanienne, savane boisée, piège lumineux, F & S. Génien, 2006-53. Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus elegantulus Burgeon, 1937 Holotype
Male adult
Image from A. Serrano, 2023
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Macrocheilus hybridus Peringuey, 1896
Female adult
Zimbabwe W Bulawayo, Shangani, Naletale ruins env. 20.12.2011, Snižek. Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus kudrnai Serrano, 2023 Holotype
Male adult
NE Benin\ E of Kandi\ 4 km W of Bensekou\ 24–25.06.2001\ A. Kudrna Jr. Lgt. Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus labrosus (Dejean, 1831)
Male adult
Central African Republic. Image from A. Serrano, 2023
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Macrocheilus longicollis Peringuey, 1904 Holotype
Male adult
Image from A. Serrano, 2023
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Macrocheilus longicollis Peringuey, 1904
Male adult
Zambia NE 2004, 30–60 km NW of Mpika 24.11. Snižek, Tichẏ lgt. Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus longicollis Peringuey, 1904
Male adult
Moçambique, 14.XII.2003 near Dombe, Werner & Lizler leg. Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus moraisi A.Serrano, 2000 Holotype
Male adult
Image from A. Serrano, 2023
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Macrocheilus ocellatus Basilewsky, 1953 Holotype
Male adult
Image from A. Serrano, 2023
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Macrocheilus overlaeti Burgeon, 1937 Holotype
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus persimilis Basilewsky, 1970 Holotype
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus persimilis Basilewsky, 1970
Male adult
Zambia, Northwestern Prov. 20 km SE Mwinilunga 7–8.01.2003 A. Kudrna Jr. Lgt. Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus philippeorum Serrano, 2023 Holotype
Male adult
Burkina Faso, Pama\ 24–28.VIII.2005,\ 11º 17′ 00′′ N, 0º 42′59E, 232m\ Phillippe Moretto leg. Image from A. Serrano, 2023
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Macrocheilus proximus Peringuey, 1896 Holotype
Male adult
Image from A. Serrano, 2023
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Macrocheilus proximus Peringuey, 1896
Female adult
Ethiopia, 25/27.IV.1998\ near Negele Borana\ Sidamo, leg. Werner. Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus quadrinotatus Burgeon, 1937 Holotype
Male adult
Image from A. Serrano, 2023
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Macrocheilus quadrinotatus Burgeon, 1937
Female adult
Kibundi (15 km N Nambua). Malanje, Angola, 19.XI.2017, 8º 54′ 33.31′′ S 16º 3′ 15.71′′ E, 1170 m. A. Serrano & R. Capela leg. Image from A. Serrano, 2023
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Macrocheilus scapularis Reiche, 1843
Male adult
Image from A. Serrano, 2023
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Macrocheilus schuelei Serrano, 2023 Holotype
Male adult
RCA, P.N. Ndoki, Camp 1, Chablis 1,\ 02º 28′ 40.5N 016º 13′ 02.6E, 24–25.II.\ 2012 forêt diversifiée, piège UV. Exp.\ Sangha 12, P. Moretto leg. -117-“ \\ “Collection P. Schüle, Herrenberg\ Shanga Projekt 2012” . Image from A. Serrano, 2023
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Macrocheilus spectandus Peringuey, 1904 Holotype
Male adult
Image from A. Serrano, 2023
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Macrocheilus spectandus Peringuey, 1904
Male adult
Tanzania, 9/13.XII.1997 near Songea, Ruvuma Prov. leg. Werner & Lizler. Image from A. Serrano, 2023
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Macrocheilus vinctus Basilewsky, 1960 Paratype
Male adult
Image from A. Serrano, 2023
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Macrocheilus taedatus Basilewsky, 1960 Paratype
Male adult
Image from A. Serrano, 2023
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Macrocheilus taedatus Basilewsky, 1960
Female adult
Tanzania, 10/16.XII.99, near Mitundo, Dodoma prov. Werner & Lizler leg. Image from A. Serrano, 2023
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Macrocheilus viduatus katanganus Basilewsky, 1960 Holotype
Male adult
Image from A. Serrano, 2023
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Macrocheilus viduatus katanganus Basilewsky, 1960
Female adult
Zambia. Image from A. Serrano, 2023
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Macrocheilus varians eugrammus Basilewsky, 1960 Holotype
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus varians Peringuey, 1904 Holotype
Male adult
Image from A. Serrano, 2023
Copyright © - - -
Anthiinae taxon page (335 photos)
Apotomus rufus (P. Rossi, 1790)
Male adult
S Spain,
8.IV.2010 Anichtchenko A. leg., det. & coll.
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Apotomus rufithorax Pecchioli, 1837: 445
Male adult
Iran, Esfahan, Kashan, near Maranjab 26.IV.2007 Anichtchenko A. leg. det. & coll.
Copyright © Alexander Anichtchenko

Apotomus testaceus Dejean, 1825
Male adult
Volgograd Reg., Pallas Distr., Elton Vill., on UV-light 12-15.VII.2006 K.Makarov leg. L=4.0 mm
Copyright © Kirill Makarov

Apotomus rufithorax Pecchioli, 1837
Male adult

Copyright © Kirill Makarov
Apotominae taxon page (4 photos)
Crepidogaster (Crepidogaster) nonstriata Chaudoir, 1876
Male adult
RSA, Limpopo Prov. 33km NNW Vaalwater
19.XI.2004 P. Schule leg.
Photo P.Schule
Copyright © Peter Schüle

Crepidogastrinus kochi Basilewsky, 1957
Male adult
S. Africa, Bushmanland, Pofadder,
30km E 23.8.1977 leg. Endroedy-Younga
Photo P.Schule
Copyright © Peter Schüle

Brachinus (Brachinus) plagiatus Reiche, 1868b: 39
Male adult
Azerbaijan, 10 km SE Agdzhabedi, 05.VII.2005, Pljushch I.G. leg. L=9mm. det. Anichtchenko 2010
Copyright © Ruslan Panin

Brachinus (Brachinus) alexandri F. Battoni, 1984c: 188
Male adult
S Ukraine, Krym Peninsula, Kizil-Koba, 12.III.2009, Shaporinskiy V. leg. L=9 mm photo Panin R.
Copyright © Ruslan Panin

Brachinus (Brachinus) crepitans Linne, 1758 (as nigripennis Letzner, 1851)
Male adult
W-UKRAINE, NW Podolian, Lviv Reg., Zolochiv Distr., near Osovitsa vill., in steppe. Panin R. leg. 1.V-16.VI.07 length = 8 mm. Photo Panin R.
Copyright © Ruslan Panin

Brachinus (Cnecostolus) exhalans P. Rossi, 1792: 84 (Carabus)
Male adult
Azerbaijan,
10 km SE Agdzhabedi, 05.VII.2005, Pljushch I.G. leg.
L=6mm
photo Panin R.
Copyright © Ruslan Panin

Brachinus (Cnecostolus) quadriguttatus Gebler, 1829: 29
Male adult
det. Anichtchenko A. photo M. Smirnov from www.zin.ru
Copyright © - - -

Brachinus (Cnecostolus) quadriguttatus Gebler, 1829: 29
Male adult
Volgograd Reg., Pallas Distr., N shore of Elton Lake, right bank of Khara Riv. 11.V.2006 A.Matalin leg.
L=8.6 mm
photo K. Makarov
Copyright © Kirill Makarov

Brachinus (Neobrachinus) medius Harris, 1828
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Brachinus (Neobrachinus) tenuicollis LeConte, 1844
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Brachinus (Brachynidius) macrocerus Chaudoir, 1876d: 67
Male adult
SE Primorskii reg., Lazo res., ~10 km SW Sokolchi vill.,
43°15` N, 134°7` E, 24.06.04 leg. M. & L. Smirnov
photo from www.zin.ru
Copyright © Alexander Anichtchenko

Mastax thermarum Steven, 1806: 166
Male adult
Astrakhan Reg., env. Baskunchak Lake, Bol. Bogdo Mt. 5.V.1995 leg. I.Melnik & E.Shankhiza
L=5.6 mm
Copyright © Kirill Makarov

Pheropsophus (Stenaptinus) hispanicus (Dejean, 1824)
Male adult
Spain, Motril
Copyright © Alexander Anichtchenko

Brachinus (Cnecostolus) bayardi Dejean, 1831
Female adult
E Turkmenistan, Charjev env., V.1996, V.Perepel leg.
12 mm
Copyright © Alexander Anichtchenko

Brachinus (Neobrachinus) cordicollis Dejean, 1826
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight

Pheropsophus (Stenaptinus) jessoensis A.Morawitz, 1862
Male adult
Collection ground Okushiri tou, Hokkaido Pref. 37781
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Brachinus (Brachynolomus) stenoderus Bates, 1873
Male adult
Collection ground Hobetsu cho, Hokkaido Pref. 33485
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Brachinus (Brachynolomus) scotomedes L. Redtenbacher, 1867
Male adult
Collection ground Tochigi Pref.Fujioka～machi 34784
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Brachinus (Brachynolomus) bantimurungensis Kirschenhofer, 2012 Holotype
Male adult
Indonesia, Südsulawesi, Bantimurung, 18./19.2.94 (collection David Wrase)
Copyright © ---

Brachinus (Cnecostolus) johorensis Kirschenhofer, 2012 Holotype
Female adult
Malaysia (Johor State) Kangkar Dohol, 30 km N Kota Tinggi 35 m (at light); 7.-11.IV.2007, P. Čechovsky.
(collection David Wrase)
Copyright © ---

Mastax histrio (Fabricius, 1801)
Male adult
N INDIA, Uttarakhand, 22-23.04.2012, 8 km NW Jaspur, lake shore, 29°21'10.54", 78°48' 30.99", Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Brachinus (Brachynidius) pokharaensis Kirschenhofer, 2012
Male adult
INDIA Uttarakhand, 8.04.2012 Rajaji Nat. Park, 4 km NE Chilla N30°0'13.88", E78°14'37.83" Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Mastax vegeta Andrewes, 1924
Male adult
India, Uttarakhand, 5 km NNE of Ramnagar, valley of Kosi River, 400-420 m, 29°26' N, 79°08,5' E, 21.04.2012, R. Dudko, I. Lyubechanskii
det. Anichtchenko A. 2012
Copyright © Roman Dudko

Mastax annulata Andrewes, 1924
Male adult
N INDIA, Uttarakhand, 10-12.04.2012, Uttarkashi distr.
N30°44.999' E 078°34.412' , 1446 m, 14 km E Uttarkashi Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Brachinus (Metabrachinus) servillei Marc, 1839
Male adult
Sénégal, Nianing, piège lumineux, 18/09/2009, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Brachinus (Dysbrachinus) dorsalis Dejean, 1831
Female adult
Sénégal, M’Bour, piège lumineux, 24/09/2009, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Aptinus (Aptinus) lugubris Schaum, 1862
Male adult
Grèce, Péloponèse, Taygète 9-07-2006, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Aptinus (Aptinus) merditanus Apfelbeck, 1918
Female adult
Grèce, Epire, Metzovon 9-06-1996, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Pheropsophus (Stenaptinus) africanus (Dejean, 1825)
Female adult
UAE, Al Markhanlya, Al Ain, 19.04.2000, C.Gillett leg.
Copyright © Alexander Anichtchenko

Pheropsophus (Stenaptinus) catoirei (Dejean, 1825)
Male adult
N India, Uttarakhand, left side of Kosi riv., 5 km N Ramnagar, N29*432 E79*140, 7-11.06.2011 Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Pheropsophus (Stenaptinus) hispanicus (Dejean, 1824)
Female adult
Spain, Andalucia, San Pedro de Alcantara, 7.04.2010, Anichtchenko A. leg.
Copyright © Alexander Anichtchenko

Pheropsophus (Stenaptinus) occipitalis s.str. (as nebulosus Chaudoir, 1876)
Male adult
Indonesia as nebulosus det. Pier Mauro Giachino, 2013
Copyright © Alexander Anichtchenko

Pheropsophus (Stenaptinus) indicus Akhil & Sabu, 2019 Paratype
Male adult
India, Karnataka, 2013, leg. Anichtchenko A. (as occipitalis (W.S.Macleay, 1825) det. Fedorenko, 2013)
Copyright © Alexander Anichtchenko

Pheropsophus (Stenaptinus) occipitalis (W.S.Macleay, 1825)
Male adult
Indonesia, Palau Banyak, Tuangku isl., Haloban, R. Cibulskis leg.
det. Fedorenko, 2013
Copyright © Alexander Anichtchenko

Aptinus (Aptinidius) displosor (L. Dufour, 1811)
Male adult
S-Spain,
Cádiz prov.,Alcalá de los Gazules,1.II.2003,T. Gazurek leg.
L=18 mm

Copyright © Ruslan Panin

Brachinus (Brachynolomus) sclopeta (Fabricius, 1792)
Male adult
S Bosnia, Mostar; 6mm
Copyright © Aleš Sedláček

Brachinus (Brachynolomus) nigricornis Gebler, 1829
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pheropsophus (Stenaptinus) javanus (Dejean, 1825)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Brachinus (Aploa) nobilis Dejean, 1831
Male adult
SE Iran, Sistan-va-Baluchestan, Tange-Surhkeh v., 1000m, 19.IV.2013, O. Legezin leg.
11 mm
det. Anichtchenko A. 2013
Copyright © Alexander Anichtchenko

Brachinus (Aploa) nobilis Dejean, 1831
Male adult
12,5 mm
Namibia, Etosha National Park, Okaukuejo, 1400 m, leg. U.Schmidt, 18.-20.III.1994; det. M.Baehr, 2012
Photo: U.Schmidt, 2013
Copyright © Udo Schmidt

Pheropsophus (Stenaptinus) cinctus (Gory, 1833)
Male adult
17,7 mm
Senegal, M`Bour, environment Club Aldiana, leg. U.Schmidt, 8.-20.X.1992; det. M.Baehr, 2012
Photo: U.Schmidt, 2014
Copyright © Udo Schmidt

Pheropsophus (Pheropsophus) debauvii (Guerin-Meneville, 1838)
Male adult
Peru: Loreto, Cmp. Terry 14 May 90, 75*28'W 05*07' S, T.L. Erwin Coll.; Igapo forest floor
21 mm
det. T.L. Erwin, 2001
Copyright © Alexander Anichtchenko

Pheropsophus (Pheropsophus) biplagiatus Chaudoir, 1876
Male adult
Mexico, Oaxaca, 22.4 mi. n. Puerto Escondido, 2100, VII.20-21.1966, George E. Ball, D.R. Whitehead collectors
det. T.L. Erwin, 2001
15 mm
Copyright © Alexander Anichtchenko

Aptinus (Aptinus) acutangulus Chaudoir, 1876
Female adult
SW-Montenegro, Dinaric Alps, Herceg Novi, Vrbanje, Mt Orjen h=1800m, 13-19.VII.2012, Manfred Egger leg.
L=13 mm

Copyright © Ruslan Panin

Styphlomerus (Styphlomerus) placidus (Peringuey, 1896)
Male adult
Namibia, Etosha National Park, Okaukuejo, 1400 m, leg. U.Schmidt, 18.-20.III.1994;
det. M.Baehr, 2012
Photo: U.Schmidt, 2014
6,0 mm
Copyright © Udo Schmidt

Brachinus (INCERTAE) dalatensis Fedorenko, 2013 Paratype
Male adult

Copyright © Fedorenko

Brachinus (INCERTAE) belyaevae Fedorenko, 2013 Holotype
Female adult
South Vietnam, ~95 km SEE of Ho Chi Minh, Xuyen Moc Nature Reserve, 15.V.1991, leg. N. Belyaeva.
Copyright © Fedorenko

Brachinus (INCERTAE) ovalipennis Fedorenko, 2013 Paratype
Male adult

Copyright © Fedorenko

Pheropsophus (Stenaptinus) similis Fedorenko, 2013 Holotype
Male adult
Central Vietnam, Ha Tinh Prov., Vu Quang Nature Reserve, h = 400–700 m, 27.VIII–4.X.1997, leg. M. Kalyakin
Copyright © Fedorenko

Pheropsophus (Stenaptinus) ngoclinhensis Fedorenko, 2013 Paratype
Male adult
Central Vietnam, Kon Tum Province, N[orthern] part., Ngoc Linh Nature Reserve, Ngoc Linh env., pitfall traps, V.2005, leg. A. Anichkin
Copyright © Fedorenko

Pheropsophus (Stenaptinus) bidoupensis Fedorenko, 2013 Paratype
Female adult
S[outh] Vietnam, Lam Dong Prov., Bi Doup – Nui Ba [Nature] Reserve, env. Long Lanh, 12°10´44´´N 108°40´44´´E, h = 1400-1600 m, 20–21.IV.2008, leg. D. Fedorenko
Copyright © Fedorenko

Pheropsophus (Stenaptinus) obliteratus Fedorenko, 2013 Holotype
Male adult
S[outh] Vietnam, Dak Lak province, Chu Yang Sin Natn. Park, 12°23´48´´N 108°20´59´´E, upper flow of Krong Kmar river, h = 1000 m, 30.III–14.IV.2012, leg. D. Fedorenko
Copyright © Fedorenko

Brachinus (Metabrachinus) armiger Dejean, 1831
Male adult
10,0 mm
Namibia, Etosha National Park, Okaukuejo, 1400 m, leg. U.Schmidt, 18.-20.III.1994; det. M.Baehr, 2012
Photo: U.Schmidt, 2014
Copyright © Udo Schmidt

Pheropsophus (Pheropsophus) platycephalus Reichardt, 1967
Male adult
Natal. Brazil, W. M. Mann. 1954
det. T.L. Erwin, 2001
14 mm
Copyright © Alexander Anichtchenko

Pheropsophus (Pheropsophus) rivieri (Demay, 1838)
Male adult
Brasil, Para, Mosqueiro, 7.V. 1977, A. M. Barbanti
det. T.L. Erwin, 2001
17 mm
Copyright © Alexander Anichtchenko

Styphlomerus (Styphlomerus) equestris (Dejean, 1831)
Male adult
10,0 mm Senegal, M`Bour, 14 km south, environment Club Aldiana, leg. U.Schmidt, 8.-20.X.1992; det. M.Baehr, 2012 Photo: U.Schmidt, 2014
Copyright © Udo Schmidt

Brachinus (Brachinus) crepitans (Linnaeus, 1758)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Brachinus (Brachinoaptinus) boeticus Rambur, 1837
Male adult
Sierra Nevada. det. Anichtchenko A.
Copyright © Kirill Makarov

Brachinus (INCERTAE) longipalpis Wiedemann, 1821
Male adult
India, Uttarakhand, Rajaji Nat. Park, 4 km NE Chilla, N30°0'13.88'', E78°14'37.83'', 8.04.2012, leg. Anichtchenko A.; 7 mm. det. D. Fedorenko, 2015
Copyright © Alexander Anichtchenko

Brachinus (INCERTAE) fuscipennis Dejean, 1825
Male adult
6,8 mm; Namibia, Etosha National Park, Okaukuejo, 1400 m, leg. U.Schmidt, 18.-20.III.1994; det. M.Baehr, 2015; Photo: U.Schmidt, 2016
Copyright © Udo Schmidt

Crepidogaster (Crepidogaster) pauliani Basilewsky, 1953
Male adult
MADAGASCAR, Tulear Prov., nr. Tsimanampetsotsa NP, P. Schüle det.
Copyright © Peter Schüle

Pheropsophus (Aptinomorphus) bipartitus Fairmaire, 1868
Male adult

Copyright © Erich Kirschenhofer

Pheropsophus (Pheropsophus) ecuadorensis Hubenthal, 1911
Male adult
Ecuador, Napo, Pto. Misahualli, 30.10.1996
Copyright © Erich Kirschenhofer

Pheropsophus (Stenaptinus) jurinei Dejean, 1825 Holotype
Male adult
Senegal
Copyright © Erich Kirschenhofer

Pheropsophus (Stenaptinus) longipennis Chaudoir, 1843 Holotype
Male adult
Senegal
Copyright © Erich Kirschenhofer

Pheropsophus (Stenaptinus) planti Chaudoir, 1876 Holotype
Male adult

Copyright © Erich Kirschenhofer

Brachinus (Metabrachinus) connectus Dejean, 1831 Holotype
Male adult
Senegal
Copyright © Erich Kirschenhofer

Pheropsophus (Stenaptinus) dux Chaudoir, 1876
Male adult
Gambia (Mus Budapest)
Copyright © Erich Kirschenhofer

Brachinus (Brachynolomus) stenoderus Bates, 1873
Male adult

Copyright © Kirill Makarov

Brachinus (Cnecostolus) cruciatus (Quensel In Schoenherr, 1806)
Male adult
S Казахстан, 8.6 км NE с. Балтаколь, берег р. Сырдарья, тугай, 43°9.40' N / 67°50.99' E, 7-12.VI.2016, И.Забалуев leg. Фотография с сайта www.zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Mastax nepalensis Morvan, 1977
Male adult
N India, Uttarakhand, left side of Kosi riv., 5 km N Ramnagar, N29*432 E79*140, 7-11.06.2011 Anichtchenko A. leg.; det. Anichtchenko A., 2016
Copyright © Alexander Anichtchenko

Mastax florida Andrewes, 1924
Male adult
India, Goa, Talpona riv., 24-27.IV.2013, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Mastax laeviceps Bates, 1891
Male adult
N India, Uttarakhand, left side of Kosi riv., 5 km N Ramnagar, N29*432 E79*140, 7-11.06.2011 Anichtchenko A. leg.; det. Anichtchenko A., 2016
Copyright © Alexander Anichtchenko

Pheropsophus (Stenaptinus) akhaensis Kirschenhofer, 2010 Holotype
Male adult

Copyright © Erich Kirschenhofer

Pheropsophus (Stenaptinus) yunnanensis Kirschenhofer, 2010 Holotype
Female adult
China, Yunnan, Nujiang Lisu Aut. Pref., creek valley 3 km SE Gongshan, 1450-1500m, 27*43'02"N, 98*41'27"E, litter, moss, sifted, 5.VI.2007, leg. A. Putz
Copyright © Erich Kirschenhofer

Pheropsophus (Stenaptinus) tamdaoensis Kirschenhofer, 2010 Holotype
Female adult
N Vietnam, Tam Dao Nat Park
Copyright © Erich Kirschenhofer

Brachinus (Metabrachinus) sexnotatus gerardi Burgeon, 1937 Holotype
Female adult
Tanganika: Moero, Kiambi VI-VII.1930, Dr. P. Gerard. Image from Hrdlicka, 2008
Copyright © - - -

Pheropsophus (Stenaptinus) guanxiensis Kirschenhofer, 2010 Holotype
Male adult

Copyright © Erich Kirschenhofer

Pheropsophus (Stenaptinus) andrewesi Jedlicka, 1964 Holotype
Male adult
India, Assam
Copyright © Erich Kirschenhofer

Pheropsophus (Stenaptinus) aptinoides Chaudoir, 1876 Holotype
Male adult
(coll Chaudoir, Mus Paris)
Copyright © Erich Kirschenhofer

Pheropsophus (Stenaptinus) bimaculatus (Linnaeus, 1771)
Male adult
(coll Chaudoir, Mus Paris)
Copyright © Erich Kirschenhofer

Pheropsophus (Stenaptinus) picicollis Chaudoir, 1876
Male adult
Burma, Rangoon distr, Hlegu - Goygon, III.1997. det. Kirschenhofer
Copyright © Erich Kirschenhofer

Pheropsophus (Stenaptinus) marginatus goudotii (Dejean, 1831)
Male adult
Madagascar (Mus. of Berlin)
Copyright © Erich Kirschenhofer

Pheropsophus (Stenaptinus) marginatus marginatus (Dejean, 1825)
Male adult
? Type ? (Mus of Paris)
Copyright © Erich Kirschenhofer

Brachinus (Brachinus) elegans Chaudoir, 1842
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Brachinus (Cnecostolus) hamatus Fischer von Waldheim, 1828
Male adult

Copyright © Kirill Makarov

Brachinus (Cnecostolus) hamatus Fischer von Waldheim, 1828
Male adult

Copyright © Kirill Makarov

Brachinus (INCERTAE) chinensis Chaudoir, 1850
Male adult
Thailand
Copyright © Erich Kirschenhofer

Brachinus (Brachynidius) lombokensis Kirschenhofer, 2010 Holotype
Male adult
Lombok, Pasu, 1.12.2003, leg. Y. Yokoi (ZSM).
Copyright © Erich Kirschenhofer

Brachinus (INCERTAE) caligatus Bates, 1890 Holotype
Male adult
Mus Genova
Copyright © Erich Kirschenhofer

Brachinus (INCERTAE) exquisitus Bates, 1892 Holotype
Male adult

Copyright © Erich Kirschenhofer

Brachinus (INCERTAE) jakli Hrdlicka, 2009 Holotype
Female adult
Indonesia - Sumba Is. / Luku - Melolo N.P. / 7.-9.i.2001 / St. Jákl lgt. [p, white label]. Image from HRDLIČKA, 2009
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Brachinus (INCERTAE) praestans Andrewes, 1931
Male adult
? Paratype. Image from HRDLIČKA, 2009
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Brachinus (INCERTAE) votrubai Hrdlicka, 2009 Holotype
Male adult
Sumba East / 20 km S of Waingapu / Wairinding, 300m / 30.i.-2.ii.2001 Votruba lgt.
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Brachinus (Brachynidius) pokharaensis Kirschenhofer, 2012 Holotype
Male adult
Nepal, W. Pokhara Baglung 5/96, Kali Gandaki
Copyright © Erich Kirschenhofer

Brachinus (Brachynidius) pokharaensis Kirschenhofer, 2012 Paratype
Male adult

Copyright © Erich Kirschenhofer

Brachinus (Brachynidius) cambodgensis Kirschenhofer, 2012 Holotype
Female adult
Cambodia SW, 20 km SE Koh Kong, Tatai river, 11°34 N, 103°07E, 17.X – 15.XI.2011, 50–300m, Z. LINEK lgt. (D. Wrase collection)
Copyright © Erich Kirschenhofer

Brachinus (Brachynidius) circumtinctus Bates, 1892
Female adult
Burma, Rangoon Distr., Hlegu Goygon, lgt. Klicha M. (cDW). det. E. Kirschenhofer
Copyright © Erich Kirschenhofer

Brachinus (Brachynidius) limbellus Chaudoir, 1876
Male adult
India (Assam State, Dibrugarth distr.) Dibrugarth (at light), 2.–3.VI.2006, Guillaume DE ROUGEMONT (cDW). det. E. Kirschenhofer
Copyright © Erich Kirschenhofer

Brachinus (Brachynidius) modestus Schmidt-Goebel, 1846
Male adult
Myanmar (Yangon) Pegu, 60 km NNE Yangon 17°19' N/96°28' E (lux) 22.XI.2003, M. HORNBURG (cDW). det. E. Kischenhofer
Copyright © Erich Kirschenhofer

Brachinus (Brachynidius) suturellus Chaudoir, 1876
Male adult
N Laos, 10 km N Luang Prabang Mekong riv. (240 km N Vientiane), XII. 1992
Copyright © Erich Kirschenhofer

Brachinus (INCERTAE) cychroides Hrdlicka, 2009 Holotype
Female adult
Vietnam, Tam Dao, 25.v.1991, M. Dudycha lgt. Deposited in collection National Museum of Prague. Image from HRDLIČKA, 2009
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Brachinus (INCERTAE) bolovenensis Hrdlicka, 2009 Holotype
Male adult
LAOS SOUTH / BOLOVEN PLATEAU / BAN ITOU / 23-27.v.2007 / B. MAKOVSKÝ Lgt. Holotype is deposited in author´s collection. Image from HRDLIČKA, 2009
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Brachinus (INCERTAE) pelengensis Hrdlicka, 2009 Holotype
Male adult
INDONESIA - / PELENG ISL., vii.2007 / lgt. local collectors. Holotype is deposited in author´s collection. Image from HRDLIČKA, 2009
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Brachinus (INCERTAE) fukienensis Hrdlicka, 2009 Holotype
Male adult
China, SW Fujian, 28.iv.-24.vii. GUANG ZHAI SHAN, 25°42´N 116°47´E, 450-480 m, 2006. Holotype is deposited in collection National Museum of Prague. Image from HRDLIČKA, 2009
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Brachinus (INCERTAE) hubeiensis Hrdlicka, 2009 Holotype
Male adult
China, W Hubei, 26.v.-10.vii., BADONG N env., 31.1N 110.4E, stream valley , 2003. Holotype is deposited in author´s collection. Image from HRDLIČKA, 2009
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Brachinus (INCERTAE) turnai Hrdlicka, 2009 Holotype
Male adult
China, Fujian c., 22.vi.-13.vii., Ziyungdongshan, NW slopes, 25°46´N 117°20´E, 900-1100 m 2007. Holotype is deposited in collection National Museum of Prague. Image from HRDLIČKA, 2009
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Brachinus (Brachinoaptinus) pateri oscuratus Mateu, 1958
Male adult
Spain, Andalusia, Sierra Magina, 1800m, 25-28.3.2010, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Brachinus (Brachinoaptinus) testaceus Rambur, 1837
Male adult
Spain, Andalisia, near Algeciras, 6.4.2003, A. Anichtchenko leg & det.
Copyright © Alexander Anichtchenko

Brachinus (Brachinoaptinus) kryzhanovskii Belousov & Kabak, 1992
Male adult
Kirgizstan, Uzun-Akmat Mt.rng., 4.6.1995, S. Ovtchinnikov leg. det. A. Anichtchenko
Copyright © Alexander Anichtchenko

Brachinus (Brachinoaptinus) pecoudi pecoudi Puel, 1925
Male adult
Spain, Peña Ubiña Mt., Montes Cantabricos, 2000m, 28.6.2009, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Brachinus (Brachinoaptinus) andalusiacus Rambur, 1837
Male adult
Spain, Andalusia, Sierra de las Nieves, 1750-1800m, 6-8.4.2010, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Brachinus (Brachinoaptinus) bellicosus L. Dufour, 1820
Male adult
Spain, Cacerez, Jaraiz de la Vera, 30.4.2006. det. Anichtchenko A.
Copyright © Alexander Anichtchenko

Brachinus (Neobrachinus) geniculatus Dejean, 1831
Male adult
Nicaragua, Managua Distr., Laguna de Xiloa, 30.11.1994, E. Van den Berghe leg.; det. R. Davidson
Copyright © Alexander Anichtchenko

Brachinus (Brachinus) atripensis Ballion, 1871
Female adult
Uzbekistan, Zeravshan mt.rng., Urgut v., 2200m, 24.6.2003, O. Legezin leg.
Copyright © Alexander Anichtchenko

Brachinus (Brachynolomus) immaculicornis Dejean, 1826
Male adult
Italy, Calabria, Asprom, San Luca (RC), 200m, 28.4.2002, leg. F. Angelini.
Copyright © Alexander Anichtchenko

Brachinus (Brachynolomus) tianshanicus Mikhailov, 1976
Male adult
Tashkent reg., 15 km E. Gazalkent, Karankulsai riv., 1000m, 7.4.2002; 6 mm
Copyright © Alexander Anichtchenko

Brachinus (Brachynolomus) brevicollis Motschulsky, 1844
Male adult
Uzbekistan, Tashkent distr., 15 km E Gazalkent, Karankolsai riv., 1000m, 7.4.2002. Klimenko leg. det. A. Anichtchenko
Copyright © Alexander Anichtchenko

Brachinus (Cnecostolus) bipustulatus Quensel In Schoenherr, 1806
Male adult
Greece, Etolia Akarnania, Neohori, 1.5.1999. F. Angelini leg.
Copyright © Alexander Anichtchenko

Brachinus (Brachinoaptinus) italicus Dejean, 1831
Male adult
Italy, Calabria, Aspromonte, San Luca, 200m, 24.1.2003, leg. F. Angelini
Copyright © Alexander Anichtchenko

Brachinus (Brachynolomus) nigricornis Gebler, 1829
Male adult
Italy, Basilicata, Policoro (MT), 14.9.1999, leg. F. Angelini
Copyright © Alexander Anichtchenko

Brachinus (Brachynolomus) nigricornis Gebler, 1829
Male adult
Italy, Puglia, S. Cataldo, WWF Le Cesine, 1.7.1995, leg. F. Angelini
Copyright © Alexander Anichtchenko

Brachinus (Brachynolomus) sichemita Reiche & Saulcy, 1855
Male adult
Greece, Ahaia, Kalavryta, 28-29.4.1999, leg. F. Angelini
Copyright © Alexander Anichtchenko

Brachinus (Brachinus) efflans Dejean, 1830
Male adult
Spain, Andalusia, San Pedro de Alcantara, 7.4.2010, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Brachinus (Brachinus) ejaculans Fischer von Waldheim, 1828
Male adult
Bulgaria, Chernomorets, 24.11.2006, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Brachinus (Brachinus) alexandri F. Battoni, 1984
Male adult
SE Azerbajan, Talysh, Mistan v., 7-10.10.1993. det. A. Anichtchenko
Copyright © Alexander Anichtchenko

Brachinus (Brachinus) psophia Audinet-Serville, 1821
Male adult
Georgia, Ajara, Meskhetskii mt. rng., Taginauri Mt., 27.6.2000. det. A. Anichtchenko
Copyright © Alexander Anichtchenko

Brachinus (Metabrachinus) connectus seydeli Burgeon, 1937 Holotype
Female adult
MUSÉE DU CONGO Katanga : Kisunka -2-IV-1925 Ch. Seydel, Kisunka 2 Avril 1925, Ch.Seydel. Image from Hrdlicka, 2008
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Brachinus (Metabrachinus) connectus ovalis Liebke, 1934 Paratype
Male adult
MUSÉE DU CONGO, Kamerun: Joko, Don. Moser (MRAC). Image from Hrdlicka, 2008
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Brachinus (Metabrachinus) stappersi Liebke, 1934 Holotype
Female adult
MUSÉE DU CONGO, Rives de la Ruzizi, 24.VII.1912, Dr. Stappers (MRAC). Image from Hrdlicka, 2008
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Styphlomerus (Styphlomerus) collarti Basilewsky, 1948 Paratype
Male adult
Congo-belge: Nizi, Blukwa-2-II-1929, A. Collart (MRAC). Image from Hrdlicka, 2008
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Aptinus (Aptinus) pyranaeus Dejean, 1824
Male adult
Spain, Pireneos, ref. de Estos, 1800m, 22.8. 2005, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Brachinus (Brachynolomus) scotomedes L. Redtenbacher, 1867
Female adult
Japan, Higashihagio Town, Omuta City, Fukuoka Pref., 19.6.2011, S. Nishimoto leg.
Copyright © Alexander Anichtchenko

Aptinus (Aptinus) alpinus Dejean, 1829
Female adult
Italy, SV, Colle del Melogno, 1030m, 25.5.2009
Copyright © Alexander Anichtchenko

Brachinus (Brachynolomus) variventris L.Schaufuss, 1862
Male adult
Spain, Teruel, Bronchales, 11.5.1996
Copyright © Alexander Anichtchenko

Brachinus (Neobrachinus) fumans (Fabricius, 1781)
Male adult
Penssylvania, Luzerne Vo., Nescopeck State Park, 17.5.2004. det. R. Davidson. 13 mm
Copyright © Alexander Anichtchenko

Brachinus (Neobrachinus) adustipennis Erwin, 1969
Male adult
Tenesse, Lake Co., Mississippi r., 361420N, 893500W, 30.8.2002, Robert D. leg. det. Davidson
Copyright © Alexander Anichtchenko

Brachinus (Brachinus) plagiatus Reiche, 1868
Male adult

Copyright © Alexander Anichtchenko

Brachinus (Neobrachinus) janthinipennis (Dejean, 1831)
Male adult

Copyright © Alexander Anichtchenko

Brachinus (Metabrachinus) quadrisignatus (Jeannel, 1949) Paratype
Male adult

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Brachinus (INCERTAE) papua Darlington, 1968 Holotype
Male adult

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Brachinus (Cnecostolus) bagdatensis Pic, 1902 Holotype
Male adult
Image from Muilwijk & Seiedy, 2017
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Lebia zaboli Jedlièka 1968 Holotype
Male adult
Image from Muilwijk & Seiedy, 2017
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Brachinus (INCERTAE) strictus Morvan 1971 Paratype
Female adult
Iran, Vll de Chalus/ 2700 m VII.1968/ nord ouest P. Morvan
Copyright © Jan Muilwijk

Brachinus (Metabrachinus) fasciatocollis (Fairmaire, 1901)
Male adult
Madagascar
Copyright © Vittorino Monzini

Mastax brittoni Quentin, 1952 Holotype
Male adult
BMNH
Copyright © - - -

Pheropsophus (Stenaptinus) galloi Giachino, 2005
Male adult
Indonesia, Papua prov. det. Anichtchenko A. 2019
Copyright © Alexander Anichtchenko

Pheropsophus (Stenaptinus) cf riedeli Giachino, 2005
Male adult

Copyright © Alexander Anichtchenko

Styphlomerus (Styphlomerus) bimaculatus Hrdlička, 2017 Holotype
Female adult
N 27°08´22´´ E 76°20´38´´ / INDIA BOR. OCC. / RAJASTHAN state, Alwar di. / Naranimata env., 460 m / 3.-12. 8. 2002, lgt. P. Šrámek [p, white label] // HOLOTYPE / Styphlomerus / bimaculatus sp. n. / det. Jan Hrdlička, 2017 [p, red label]“. Holotype is deposited in coll. National Museum of Prague. Image from Jan Hrdlička, 2017
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Pheropsophus (Stenaptinus) azoulayi Lassalle & Schnell, 2018 Holotype
Male adult
Philippines, Visayas Orientales,Leyte, Baybay, Lagolago, forêt en surplomb de Visayas State University, 100 m, 27.04.17, Lassalle et Schnell leg., in coll Lassalle.
Copyright © Bernard Lassalle

Brachinus (INCERTAE) bendanilloi Lassalle & Schnell, 2018 Holotype
Female adult
Philippines, Visayas central, île de Cebu, Ft d’Alcoy, 31.08.17, Lassalle et Schnell leg., in coll. San Carlos University à Cebu Talamban (Philippines).
Copyright © Bernard Lassalle

Brachinus (INCERTAE) leytensis Lassalle & Schnell, 2018 Holotype
Male adult
Philippines, Visayas orientales, Leyte, Baybay, Lagolago, forêt en surplomb de Visayas State University, 100 m, 27.04.17, Lassalle et Schnell leg., in coll Schnell.
Copyright © Bernard Lassalle

Brachinus (INCERTAE) grootaerti Tian & Deuve, 2013 Holotype
Male adult
Coll. I. R. Sc. N. B, the Philippines, C. Palawan, 5 km E. of Napsan, cleared primary forest, 09.42.256 N/118.31.034 E, 1–2. Dec. 2007, J. H. Lourens leg.” and “Coll. IRSNB, Achat J. H. Lourens, I. G.:30.852. (in RISNB, Brusssels)
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Brachinus (INCERTAE) barclayi Hrdlička, 2019 Holotype
Male adult
Rangoon / Burma / 28.5.1922 / E. A. Dabreu [hw, white label] // Brit. Mus. [p] / 1958- [p] 385 [hw], [white label] // Brachynus / sp. nov. [hw, white label] // HOLOTYPE / Brachinus / barclayi sp. n. / det. Jan Hrdlička, 2018 [p, red label]. Image from Jan HRDLIČKA, 2019
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Brachinus (Cnecostolus) geiseri Hrdlička, 2019 Holotype
Female adult
INDIA: / Poona. At light. / P.V. Joshi. / B.M. 1961 - 164 // 17 [hw, white label] // Brachynus sp. / (nov. in B. M.) / E.B.Britton.det.1961 // HOLOTYPE / Brachinus / geiseri sp. n. / det. Jan Hrdlička, 2018. 9.8 mm Image from Jan HRDLIČKA, 2019
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Brachinus (INCERTAE) garnerae Hrdlička, 2019 Holotype
Male adult
Nyew-tew Id. / nr Shei-pu / Walker Coll. / 93.-18. // IO 194 // Brachynus sp. // HOLOTYPE / Brachinus / garnerae sp. n. / det. Jan Hrdlička, 2018. 7.23 mm; Image from Jan HRDLIČKA, 2019
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Pheropsophus (Stenaptinus) lei Hrdlička, 2019 Holotype
Male adult
VIETNAM Central, / Quang Nam Province, / Tay Giang, Mt. Axan, / 1.300 m, IV.2017, / Thanh Luong Le lgt.; 20.9 mm; Image from Jan HRDLIČKA, 2019
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Pheropsophus (Stenaptinus) snizeki Hrdlička, 2019 Paratype
Male adult
NW Thailand, Chieng Dao, / Ban San Pakia, 5.- 10.v. / 2004, 1200m, Sv. Bílý leg. (NMPC); 16 mm; Image from Jan HRDLIČKA, 2019
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Pheropsophus (Stenaptinus) worthingtoni Hrdlička, 2019 Holotype
Male adult
S.E. THAILAND / Khao Soi Dao N.P. / 7.XI.2018 / Leonard Worthington lgt. // HOLOTYPE / Pheropsophus / worthingtoni sp. n. / det. Jan Hrdlička, 2018 . Holotype is deposited in coll. Jan Hrdlička, Babice u Říčan (CZE). Image from Jan HRDLIČKA, 2019
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Pheropsophus (Stenaptinus) lumawigi Hrdlička, 2019
Male adult
PHILIPPINES-MINDANAO / Davao del Sur, / Kapatagan, VIII.2016, / lgt. Ismael Lumawig. 16.3 mm; Image from Jan HRDLIČKA, 2019
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Pheropsophus (Stenaptinus) jandai Hrdlička, 2019 Holotype
Female adult
PNG, MADANG Prov. / Halopa / xii.2001 / M. Janda leg. 17.6 mm; Image from Jan HRDLIČKA, 2019
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Pheropsophus (Stenaptinus) fastigiatus (Linnaeus, 1764)
Female adult
Congo. det. Anichtchenko A. 2019
Copyright © Alexander Anichtchenko

Pheropsophus (Stenaptinus) capensis Chaudoir, 1876
Female adult
S Arf.: Waterberg Geelhoutbosch farm, 24.21S -27.38E, 7.10.1995. compared with typus f.typ.
Copyright © Erich Kirschenhofer

Pheropsophus (Pheropsophus) ecuadorensis Hubenthal, 1911
Male adult
Ecuador, Orellana prov., rio Napo, 8.2014. (DUBC) det. Anichtchenko A. 2019
Copyright © Alexander Anichtchenko

Pheropsophus (Stenaptinus) jessoensis A.Morawitz, 1862
Male adult
Japan, Fukuoka pref., Omuta, Higashihagio, 17.5.2014. det. Anichtchenko A., 2019
Copyright © Alexander Anichtchenko

Pheropsophus (Stenaptinus) lumawigi Hrdlička, 2019
Male adult
Mindanao, Davao del Sur, Apo Mt., 3.2014. det. Anichtchenko A. 2019
Copyright © Alexander Anichtchenko

Pheropsophus (Stenaptinus) nigerrimus Jedlicka, 1935
Male adult
Mindanao, Bukidnon, Panamokan, 12.2014. det. Anichtchenko A., 2019
Copyright © Alexander Anichtchenko

Pheropsophus (Stenaptinus) capitatus Jedlicka, 1935
Male adult
Philippines, Luzon, Nueva Vizcaya, Belance, 3.2014. det. Anichtchenko A. 2019
Copyright © Alexander Anichtchenko

Pheropsophus (Stenaptinus) windjanae Baehr, 2012 Paratype
Male adult
Australia, Windjana Gorge National Park, Lennard River, 17°24.72’S, 124°56.53’E, 103 m, 21–22 November 2007, M. Baehr
Copyright © Martin Baehr

Pheropsophus (Stenaptinus) abbreviatus Arrow, 1901
Male adult
Urw. Mawambi
Copyright © Erich Kirschenhofer

Pheropsophus (Stenaptinus) amnicola Darlington, 1968 Holotype
Male adult
vic. Nadzub, Brit. N. G. July 1944 Darlington; A46; Outline Todd1961; meas. male; Drawn 1966 Mary Catron No. 335 (MCZ)
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Pheropsophus (Stenaptinus) catulus Darlington, 1968 Holotype
Male adult
Dobodura Papua N. G. Mar. -July, 1944, Darlington; Darlington at B.M. 1947-48 Notes p.; meas. male; Drawn 1966 Mary Catron No. 81 (MCZ).
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Pheropsophus (Stenaptinus) bisulcatus Chaudoir, 1843
Female adult
Niger (NMW)
Copyright © Erich Kirschenhofer

Pheropsophus (Stenaptinus) lei Hrdlička, 2019
Male adult
Vietnam Tay Giang, Quang Nam 1300m, 4.2017. 21 mm; det. D. Fedorenko, 2019
Copyright © Alexander Anichtchenko

Pheropsophus (Stenaptinus) sonae Hrdlička, 2015 Holotype
Male adult
NIGERIA, 13.-14.V.2011, / 25 km E of ABUJA, / 400 m, 09°05´32,67´´N, / 07°45´14,82´´E, / V. Kremitovský leg. [p, white label] // HOLOTYPE / Pheropsophus sonae / sp. n. / det. Jan Hrdlička, 2014. Image from Jan HRDLIČKA, 2015
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Pheropsophus (Stenaptinus) jakli Hrdlička, 2015 Holotype
Male adult
INDONESIA, TANIMBAR / Isls., YAMDENA Isl., / 21 km NE SAUMLAKI, / MAM´S vill., 27.11.- / 11.xii.2005, St. Jakl lgt. [p, white label] // HOLOTYPE / Pheropsophus jakli / sp. n. / det. Jan Hrdlička, 2014. Image from Jan HRDLIČKA, 2015
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Pheropsophus (Stenaptinus) kabelkai Hrdlička, 2015 Holotype
Male adult
INDONESIA / (Tanimbar Isls.), / YAMDENA ISL., / Lorulun vill. env. // (20 km NE of Saumlaki), / 150 m alt., / 25.XI. – 24.XII.2006 / lgt. St. Jakl [p, white label] // HOLOTYPE / Pheropsophus kabelkai / sp. n. / det. Jan Hrdlička, 2014. Image from Hrdlička, 2015
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Pheropsophus (Stenaptinus) agnatus (Chaudoir, 1876)
Male adult
Thailand, Khao Soi Dao N.P., Chanthaburi, leg. L.Worthington, 14.II.1918; det. Hrdlicka, 2019 as worthingtoni; Photo: U.Schmidt, 2019. 17.3 mm
Copyright © Udo Schmidt

Brachinus (Brachynolomus) sanchi Azadbakhsh & Kirschenhofer, 2018 Paratype
Male adult

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Brachinus (Brachinus) mansorii Azadbakhsh & Kirschenhofer, 2018 Holotype
Male adult
Iran, Hormozgan Province, Ahmadi env., 4.IV.2017, leg. S. Azadbakhsh (ZMUT)
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Brachynillus varendorffi Reitter, 1904 Holotype
Male adult

Copyright © Erich Kirschenhofer

Styphlomerus (Styphlomerus) burgeoni Liebke, 1934
Male adult
RCA, 45 km N Sibut, 400m, 11.5.2009. det. E. Kirschenhofer
Copyright © Alexander Anichtchenko

Styphlomerus (Styphlomerus) quadrimaculatus (Dejean, 1831)
Female adult
Ethiopia, Oromia, 8 km S Dolomena, 1200m, 11-14.4.2016. det. E. Kirschenhofer
Copyright © Erich Kirschenhofer

Brachinus (Metabrachinus) congicus (Basilewsky, 1958) Paratype
Male adult
Coll. Mus. Congo, L. Leopold II: Bolobo, 6/XII-1952, P. Basilewsky
Copyright © Erich Kirschenhofer

Brachinus (INCERTAE) apicalis adansoni Alluaud, 1926 Paratype
Male adult
Afrique Or Anglaise, Pobi de Seringheti, (De Bura a Taveta), Ch. Alluaud I-IV.1904
Copyright © Erich Kirschenhofer

Styphlomerus (Styphlomerus) devagiriensis Akhil & Thomas, 2019 Holotype
Male adult
Image from Akhil & Thomas, 2019
Copyright © - - -

Styphlomerus (Styphlomerus) striatus Akhil & Thomas, 2019 Holotype
Male adult
Image from Akhil & Thomas, 2019
Copyright © - - -

Styphlomerus (Styphlomerinus) fusciceps (Schmidt-Gobel, 1846)
Male adult
Image from Akhil & Thomas, 2019
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Styphlomerus (Styphlomerinus) fusciceps (Schmidt-Gobel, 1846) Holotype
Male adult
(NMPC) . Image from Akhil & Thomas, 2019
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Styphlomerus (Styphlomerus) aulicus (Dejean, 1831)
Male adult
RCA, 60 km SE Bonar, 860m, N05*42' E16*00', 1.6.2009. det. E. Kirschenhofer
Copyright © Erich Kirschenhofer

Styphlomerus (Styphlomerus) exilis (Laferte-Senectere, 1850)
Male adult
Tanzania. 5 mm; det. E. Kirschenhofer
Copyright © Alexander Anichtchenko

Styphlomerus (Styphlomerus) kisantuus Liebke, 1934 Holotype
Male adult
Kisantu, P. Goossens (MRAC)
Copyright © Erich Kirschenhofer

Styphlomerus (Styphlomerus) titschacki lamottei Basilewsky, 1951 Paratype
Male adult
Keoulenta. Kimba (Guinee), M. Lamotte 11.VI.48 (MRAC)
Copyright © Erich Kirschenhofer

Styphlomerus (Styphlomerus) 4-maculatus var katanganus Liebke, 1934 Holotype
Male adult
Bukama, 23.IV.1911, Dr. Bequaert (MRAC)
Copyright © Erich Kirschenhofer

Aptinus (Aptinus) bombarda (Illiger, 1800)
Male adult
Autriche, Carinthie Sankt Jakob im Rosental, 20-06-2007
Copyright © Pascal Deschamps

Brachinus (Brachynolomus) devagiriensis Akhil, Divya & Sabu, 2020
Female adult
India, Karnataka, Kerwa Di, N14.52.49 E 74.15.52, lake, 18-19.4.2013 (cAA); 5.5 mm; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Brachinus (Brachynolomus) devagiriensis Akhil, Divya & Sabu, 2020
Male adult
India, Karnataka, Kerwa Di, N14.52.49 E 74.15.52, lake, 18-19.4.2013 4.5 mm (cAA). det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Brachinus (Cnecostolus) tetrastigma Fairmaire, 1897
Male adult
Madagascar, Mariaramo, Majunga 17.11.1983 (cAA) 4.5 mm; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Brachinus (INCERTAE) mindanaoensis Tian & Deuve, 2013
Male adult
Mindanao, Agusan del Sur, Esperanza, 4.2014 (DUBC) 16 mm; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Brachinus (INCERTAE) solidipalpis Tian & Deuve, 2007
Female adult
Philippines, C Luzon, Zambales, Subic, 11.2015 (DUBC) det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Brachinus (Metabrachinus) tetraspilotus Chaudoir, 1876
Male adult
Ethiopia, Illubabor prov., 25 SE Abobo (cAA) 11 mm; det. Anichtchenko A., 2020
Copyright © Alexander Anichtchenko

Brachinus (INCERTAE) nigrovirescens Basilewsky, 1951
Male adult
Ethiopia, Illubabor prov., 18km SE Abobo, 25.8-5.9.1988 (DUBC) 7.5 mm; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Brachinus (INCERTAE) solidipalpis Tian & Deuve, 2007
Male adult
Philippines C Luzon, Zambales, Subic, 11.2015 (DUBC) det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Brachinus (Brachynolomus) costatulus (Quensel In Schoenherr, 1806)
Male adult
NW Kazakhstan, Aktjubinsk reg., Emba vill., 16.5.2000 (DUBC) 11 mm; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Brachinus (Brachynolomus) explodens Duftschmid, 1812
Female adult
Bulgaria Chernomorets, 24.11.2006, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Brachinus (Brachynolomus) flores Kirschenhofer, 2011 Holotype
Female adult
West Flores, Rana Mese, 20. - 30. VI. 1927, Rensch S.G. (MHB).
Copyright © Erich Kirschenhofer

Brachinus (Brachynolomus) hunanensis Kirschenhofer, 2011 Holotype
Female adult
China, Wukang, Proz. Hunan (MHB). 16 mm
Copyright © Erich Kirschenhofer

Brachinus (Brachynolomus) batuensis Kirschenhofer, 2011 Holotype
Male adult
Batoe Doelang, Batoe Lantch Gebirge, 1300 m, Rensch, 14.5.27 (MHB), = Indonesien. Provinz Nord Sumatra, Batu Inseln. 14.3mm
Copyright © Erich Kirschenhofer

Brachinus (Brachynolomus) cf. sanchi Azadbakhsh & Kirschenhofer, 2018
Female adult
Iran, Kerman, Deh Bakri, Gebal Barez Mts., 29.08N 57.91E, 2300m, 2-3.6.2010 (cAA) det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Brachinus (Brachynolomus) inops Andrewes, 1932 Holotype
Male adult
India, W. Almora Du. Almora U.P. (BMNH) Natural History Museum (2014). Dataset: Collection specimens. Resource: Specimens. Natural History Museum Data Portal (data.nhm.ac.uk). https://doi.org/10.5519/0002965
Copyright © - - -

Brachinus (Brachynolomus) peltastes Andrewes, 1931 Holotype
Female adult
“Dehra Dun: Dhobalwala, ‘2,304 ft.’, H. E. Andrewes Coll. B.M.1945-97, ‘bought from Staudinger & Bang-Haas, 1930, det. H. E. Andrewes" (BMNH) Natural History Museum (2014). Dataset: Collection specimens. Resource: Specimens. Natural History Museum Data Portal (data.nhm.ac.uk). https://doi.org/10.5519/0002965
Copyright © - - -

Brachinus (Metabrachinus) sexmaculatus Dejean, 1825
Male adult
Myanmar, Shan state, Taunggyi env., VI.2008 (coll Wrase); E. Kirchenhofer det as B. hexagrammus.; det. Anichtchenko A. 2020
Copyright © Erich Kirschenhofer

Brachinus (Brachynolomus) stevensi Andrewes, 1924 Holotype
Female adult
“Gielle Khola, Tista Valley, Darjiling. 28.xi-3.xii.19, 600 ft., coll. H. Stevens, ‘H. E. Andrewes Coll. B.M. 1945-97’, det. H. E. Andrewes” (BMNH). Natural History Museum (2014). Dataset: Collection specimens. Resource: Specimens. Natural History Museum Data Portal (data.nhm.ac.uk). https://doi.org/10.5519/0002965
Copyright © - - -

Brachinus (Brachynolomus) thieleni Kirschenhofer, 1987
Male adult
29 km NW Lan Sak. det. E. Kirschenhofer
Copyright © Erich Kirschenhofer

Brachinus (Brachynolomus) puncticollis Schmidt-Goebel, 1846
Male adult
25 km NW Lan Sak
Copyright © Erich Kirschenhofer

Brachinus (INCERTAE) daliensis Kirschenhofer, 2011 Holotype
Male adult
China (Yunnan) Dali Bai Aut. Pref., Babuab Jiang river bank, 1800 m, 24*51'26.6''N, 100*29'22.1''E (under gravel/stones, flood debris) 16.IX.2009 D.W.WRASE
Copyright © Erich Kirschenhofer

Brachinus (Metabrachinus) hexagrammus Chaudoir, 1876
Male adult
Natural History Museum (2014). Dataset: Collection specimens. Resource: Specimens. Natural History Museum Data Portal (data.nhm.ac.uk). https://doi.org/10.5519/0002965
Copyright © - - -

Brachinus (Brachynolomus) pallidipes Reitter, 1919
Male adult
N India, Uttarakhand, Rajaji Nat. Park., 4km NE Chilla, 8.4.2012 (cAA) 11 mm
Copyright © Alexander Anichtchenko

Styphlomerus (Styphlomerus) paralleloides Lorenz, 1998
Male adult

Copyright © Erich Kirschenhofer

Pheropsophus (Stenaptinus) glabricollis Fedorenko, 2020 Paratype
Male adult
1 - male, N-Vietnam, 40 km W of Cao Bang, Phia Oac Mt., 22º36´30´´N/ 105º52´20´´E, h~1600–1650 m, deciduous forest, 3–11.X.2018, leg. D. Fedorenko. 2-3 - Vietnam, Tam Dao, 8–13.IV.1986, Gorokhov
Copyright © Dmitri Fedorenko

Pheropsophus (Stenaptinus) tamdaoensis Kirschenhofer, 2010
Male adult
Vietnam. det. Fedorenko D., 2020
Copyright © Dmitri Fedorenko

Pheropsophus (Stenaptinus) guanxiensis Kirschenhofer, 2010
Male adult
Vietnam. det. Fedorenko, 2020
Copyright © Dmitri Fedorenko

Pheropsophus (Stenaptinus) obliteratus Fedorenko, 2013
Male adult
Vietnam. det. D. Fedorenko 2020
Copyright © Dmitri Fedorenko

Pheropsophus (Stenaptinus) montanus Fedorenko, 2020 Paratype
Male adult
Vietnam, Gia Lai Province, ~40 km ENE of Pleiku, 14º13´21´´N108º19´55´´E, Kon Ka Kinh Nat[io]n[al]. Park, h=1490 m, 21–30.V.2017, D. Fedorenko leg.
Copyright © Dmitri Fedorenko

Pheropsophus (Stenaptinus) dissolutus (Andrewes, 1923)
Male adult
1 — from N Vietnam; 2 — China, Southern Sichuan, env. Xichang, h=1600 m, 21.V.2018, leg. A. Gorodinski. det. D. Fedorenko, 2020
Copyright © Dmitri Fedorenko

Styphlomerus dissimilis Basilewsky, 1943 Holotype
Male adult

Copyright © Erich Kirschenhofer

Neobrachinus sp.
Male adult
NE-ARGENTINA, Jujuy Prov., San Salvador de Jujuy, Tumbaya, h = 2130 m, S 23°52', W 65°27', 27.II.2015, leg. G.L.Agnoli + G.Garsaniti
Copyright © Igor Solodovnikov

Pheropsophus (Stenaptinus) capensis Chaudoir, 1876
Female adult
RSA Kwazulu-Natal. det. Anichtchenko A. 2021
Copyright © Igor Solodovnikov

Pheropsophus (Stenaptinus) consularis flaviceps Fedorenko, 2021 Paratype
Female adult

Copyright © Dmitri Fedorenko

Pheropsophus (Stenaptinus) dongnaiensis Fedorenko, 2021 Paratype
Male adult
S[outhern] Vietnam, N[orthern part of] Dongnai Province, Nam Cat Tien Nat[ional]. Park, Exped[ition of]. Russ[ian].-Vietnamese Tropical Centre, at light HQL450, 5.XII.2004, leg. D.Fedorenko’.
Copyright © Dmitri Fedorenko

Pheropsophus (Stenaptinus) sumatrensis Fedorenko, 2021 Holotype
Male adult
(ZMMU) labeled ‘Sumatra, Barat Province, Maninjau vill., 00°17´09´´N/ 100°13´54´´E, h=498 m, 18–19.XII.2014, leg. I. Melnik’.
Copyright © Dmitri Fedorenko

Pheropsophus (Stenaptinus) stenoderus (Chaudoir, 1850)
Male adult
1- India, 2 - N Thailand; 3 - Song Thanh NP.
Copyright © Dmitri Fedorenko

Pheropsophus (Stenaptinus) javanus (Dejean, 1825)
Male adult
1 - Loc Bao; 2-3 - Thailand
Copyright © Dmitri Fedorenko

Pheropsophus (Stenaptinus) fimbriatus (Chaudoir, 1876)
Male adult
1 - Sumatra; 2-3 - Cambodia; 4-5 - Vietnam, Cat Tien NP.
Copyright © Dmitri Fedorenko

Styphlomerus (Styphlomerus) gebieni Liebke, 1927
Male+Female adult
Cote d'Ivoire, Lamto Toumodi, 21-08-1971
Copyright © Pascal Deschamps

Brachinus (Neobrachinus) bilineatus Castelnau, 1835
Male adult
Paraguay, Dep. Caazapa, Colonia Neufeld, S26.28 W55.55, 24.10.2008, U. Drechsel leg. (cAA) 8 mm; det. Anichtchenko A.
Copyright © Alexander Anichtchenko

Brachinus (Metabrachinus) tetragrammus Chaudoir, 1876 Syntypes
Male adult
Ind.or.bor., coll. Dr. Bacon, det. Chaudoir (MNHN)
Copyright © - - -

Brachinus (Metabrachinus) eucosmus Andrewes, 1937 Holotype
Female adult
Type, Dehra Dun, U.P., coll. J. C. M. Gardner, 20.III.1933, Ex. Descented termite mound, 2328, det. H. E. Andrewes, Brit.Mus.1937.548. (BMNH)
Copyright © - - -

Brachinus (INCERTAE) illotus Chaudoir, 1876 Syntypes
Male adult
Deccan, coll. Stevens, det. Chaudoir (MNHN)
Copyright © - - -

Brachinus (Metabrachinus) reyi Andrewes, 1924 Holotype
Male adult
Madras: Surada, coll. R. P. J. Rey, ex. coll. Babault (BMNH)
Copyright © - - -

Brachinus (INCERTAE) andamanensis Tian et Deuve, 2016 Holotype
Male adult
Andaman (MNHN)
Copyright © Mingyi Tian

Brachinus (INCERTAE) drumonti Tian & Deuve, 2016 Paratype
Male adult
scale bar: 5.0 mm;
Copyright © Mingyi Tian

Brachinus (INCERTAE) guangdongensis Tian & Deuve, 2016 Holotype
Male adult
China: Guangdong: Longmen Xian, Nankun Shan, alt. 1000 m”, “VIII-XI, 2000, T. Deuve & M. Y. Tian leg. (SCAU)
Copyright © Mingyi Tian

Brachinus (Brachinoaptinus) angustatus (Dejean, 1831)
Male adult
Spain, Cadiz, Vejer de la Frontera, El Palmar21.10.2008 (cAA). det. A. Anichtchenko 7.5mm
Copyright © Alexander Anichtchenko

Aptinus (Aptinus) creticus (Pic, 1903)
Female adult
Turkey, Akseki 29.3.98 (DUBC)
Copyright © Alexander Anichtchenko

Aptinus (Aptinus) creticus (Pic, 1903)
Male adult
Turkey, Mugla prov., Oludeniz distr., Babadag Mt., 1000m, 15.5.2012 (DUBC)
Copyright © Alexander Anichtchenko

Aptinus (Aptinus) cordicollis Chaudoir, 1843
Male adult
Turkey Abant 1.6.1991 (DUBC)
Copyright © Alexander Anichtchenko

Pheropsophus (Stenaptinus) lisaae Écharoux & Roux, 2021 Holotype Paratype
Male+Female adult
Image from Écharoux & Roux, 2021
Copyright © - - -

Pheropsophus (Stenaptinus) kersteni Gerstaecker, 1866
Male adult
Image from Écharoux & Roux, 2021
Copyright © - - -

Pheropsophus (Stenaptinus) kersteni quadrimaculatus Hubenthal, 1914 Syntypes
Male adult
Image from Écharoux & Roux, 2021
Copyright © - - -

Pheropsophus (Stenaptinus) amadori Lassalle & Roux, 2021 Holotype
Male adult
Holotype, male. Paratype - Coxostyles.
Copyright © Bernard Lassalle

Pheropsophus (Stenaptinus) bicolor Lassalle & Roux, 2021 Holotype Paratype
Male+Female adult

Copyright © Bernard Lassalle

Pheropsophus (Stenaptinus) delahayei Lassalle & Roux, 2021 Holotype
Male adult

Copyright © Bernard Lassalle

Pheropsophus (Stenaptinus) drumonti Lassalle & Roux, 2021 Holotype
Male adult

Copyright © Bernard Lassalle

Pheropsophus (Stenaptinus) fallax Lassalle & Roux, 2021 Holotype
Male adult

Copyright © Bernard Lassalle

Pheropsophus (Stenaptinus) fallax Lassalle & Roux, 2021 Paratype
Female adult

Copyright © Bernard Lassalle

Pheropsophus (Stenaptinus) insolitus Lassalle & Roux, 2021 Holotype
Male adult
Holotype male + Coxostyles of PT
Copyright © Bernard Lassalle

Pheropsophus (Stenaptinus) kivuensis Lassalle & Roux, 2021 Holotype
Male adult

Copyright © Bernard Lassalle

Pheropsophus (Stenaptinus) legalli Lassalle & Roux, 2021 Holotype Paratype
Male+Female adult

Copyright © Bernard Lassalle

Pheropsophus (Stenaptinus) mirandus Lassalle & Roux, 2021 Holotype
Male adult
Coxostyles of PT
Copyright © Bernard Lassalle

Pheropsophus (Stenaptinus) pseudomashunus Lassalle & Roux, 2021 Holotype
Male adult

Copyright © Bernard Lassalle

Pheropsophus (Stenaptinus) simplex Lassalle & Roux, 2021 Holotype
Male adult

Copyright © Bernard Lassalle

Pheropsophus (Stenaptinus) simplex Lassalle & Roux, 2021 Paratype
Female adult

Copyright © Bernard Lassalle

Pheropsophus (Stenaptinus) simulator Lassalle & Roux, 2021 Holotype
Male adult
Coxostyles PT
Copyright © Bernard Lassalle

Pheropsophus (Stenaptinus) unicolor Lassalle & Roux, 2021 Holotype
Male adult

Copyright © Bernard Lassalle

Pheropsophus (Stenaptinus) fimbriatus (Chaudoir, 1876) Lectotype
Male adult
Image courtesy Azadeh Taghavian-Azari (MNHN)
Copyright © - - -

Brachinus (Neobrachinus) elongatulus Chaudoir, 1876
Male adult
Size variation within the species. Scale bar: 5 mm. Image from Ikagawa & Moore, 2022
Copyright © - - -

Brachinus (Neobrachinus) cibolensis Erwin, 1970
Male adult
Image from Ikagawa & Moore, 2022
Copyright © - - -

Brachinus (Neobrachinus) azureipennis Chaudoir, 1876
Male adult
Image from Ikagawa & Moore, 2022. Scale bar: 1 cm.
Copyright © - - -

Brachinus (Neobrachinus) gebhardis Erwin, 1965
Male adult
Image from Ikagawa & Moore, 2022. Scale bar: 1 cm.
Copyright © - - -

Brachinus (Neobrachinus) elongatulus Chaudoir, 1876
Male adult
Image from Ikagawa & Moore, 2022. Scale bar: 1 cm.
Copyright © - - -

Brachinus (Neobrachinus) mexicanus Dejean, 1831
Male adult
Image from Ikagawa & Moore, 2022. Scale bar: 1 cm.
Copyright © - - -

Brachinus (Neobrachinus) costipennis Motschulsky, 1859
Male adult
Image from Ikagawa & Moore, 2022. Scale bar: 1 cm.
Copyright © - - -

Brachinus (Neobrachinus) hirsutus Bates, 1884
Male adult
Image from Ikagawa & Moore, 2022. Scale bar: 1 cm.
Copyright © - - -

Brachinus (Neobrachinus) lateralis Dejean, 1831
Male adult
Image from Ikagawa & Moore, 2022. Scale bar: 1 cm.
Copyright © - - -

Brachinus (Neobrachinus) favicollis Erwin, 1965
Male adult
Image from Ikagawa & Moore, 2022. Scale bar: 1 cm.
Copyright © - - -

Brachinus (Brachynolomus) hajdajorum Wrase & Hrdlička, 2023 Holotype
Male adult
NW Syria / Slinfah (1500m) E Latakia / 26.-29.V.1998 / lgt. E. & P. Hajdaj (cDW). Image from Wrase & Hrdlička, 2023
Copyright © - - -
Brachininae taxon page (330 photos)
Barypus calchaquensis Roig-Junent, 1992
Male adult
Argentina, Salta Piedra del Molino, 3600 m 2.86 Ratel L. leg. (cAA). det. A. Cicchino, 2023
Copyright © Alexander Anichtchenko

Barypus (Cardiophthalmus) clivinoides (Curtis, 1839)
Male adult

Copyright © Alexander Anichtchenko

Broscodera (Broscodera) insignis Mannerheim, 1852
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Broscus cephalotes Linne, 1758: 414 (Carabus)
Male adult
length: 17-25 mm
Helvetia, Graubünden, Ilanz
leg.det. U.Schmidt, 2003
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Broscus nobilis Dejean, 1828: 432
Male adult
Greece, Isl. Rhodos, 10 km NNW Lindos 10-20m, 18.4.1994 A Korell leg.
17 mm
photo, det. & coll. Anichtchenko A. 2009
Copyright © Alexander Anichtchenko

Broscus uhagoni Bolivar y Pieltain, 1912: 374
Male adult
Spain, Toledo, Quero 30.3.2008
16 mm
photo & det. Anichtchenko A. 2009
Copyright © Alexander Anichtchenko

Chaetobroscus (Chaetobroscus) kezukai Dostal, 1984
Male adult
N India, Himachal Pradesh, W Himalayae, Pirpandzhal Mt.R., Ratang Pass, h~3600 15.VII.2003 A.Gorodinski leg.
L=12.5 mm
Copyright © Kirill Makarov

Craspedonotus margellanicus Kraatz, 1884c: 229
Male adult
length: 19 mm Pamir, South Tadzhikistan, Gonishou vill. 5.06.1989
Copyright © Evgeniy Zinoviev

Miscodera arctica Paykull, 1798: 85
Male adult
SW Tuva, West Tannu-Ola Mt.R., valley of Chas-Adyr Riv. near mouth of Bol.Bug Riv., h~1620 13.VI.2007 R.Dudko leg.
L=7.0 mm
Copyright © Kirill Makarov

Miscodera arctica Paykull, 1798: 85
Male adult
length: 6,3-8 mm
England, Dorback Bum
leg.det. M.G.Telfer, 2004
Foto: U.Schmidt, 2009
Copyright © U.Schmidt

Zacotus matthewsii LeConte, 1869
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Creobius eudouxii (Guerin-Meneville, 1839)
Male adult
Chile, Osomo, Chan-chan 27.11.2004 G.Aceituno leg.
19 mm
photo & coll. Anichtchenko A. 2009
Copyright © Alexander Anichtchenko

Craspedonotus tibialis Schaum, 1863
Male adult
China, Guangxi pro., Yuanbaoshan, 2009.10, Liu ye & Shi hongliang coll.;
photo by liuye
Copyright © Ye Liu

Broscus cephalotes (Linne, 1758)
Female adult
L=21,1 mm
FINLAND, Uusimaa, Rayskala env., 21-31.VII.2008
leg. & det. Ovsiannikov I.
photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Chylnus ater (Putzeys, 1868)
Male adult
Grove, S.J. (2010). Tasmanian Forest Insect Collection web-site
Copyright © Simon Grove

Axonya championi Andrewes, 1923
Male adult
N India, Uttarakhand, left side of Kosi riv., 5 km N Ramnagar, N29*432 E79*140, 7-11.06.2011, Anichtchenko A. leg. & det.
10 mm
Copyright © Alexander Anichtchenko

Craspedonotus himalayanus Semenov, 1910: 217
Male adult
N INDIA, Uttarakhand, 13-15.04.2012, Uttarkashi distr. N30°57'41.57", E78°41'54.75" left tributary of Bhagirathi riv., 2300 m,
Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Eobroscus (Eobroscus) lutshniki Roubal, 1928d: 90
Male adult
Far East of Russia, S Sakhalin isl., Aniva dist., Taranai vill., 30.07.2006, O. Legesin leg.
Anichtchenko A. det.
Copyright © Alexander Anichtchenko

Broscosoma valainisi Barsevskis, 2010 Holotype
Female adult
China, Shanxi, Tschingling Mts., Taibeichan Nat. res., 33*53'N 107*49'E, 2000 m, 10.08.2005, V. Patrikeev leg.
Copyright © Alexander Anichtchenko

Eobroscus (Eobroscus) lutshniki Roubal, 1928d: 90
Female adult
S Sakhalin, Krilion, env. Shebunini 46°24'59"N 141°51'53"E 9-18.VI.2003 leg. I.Melnik
L=14.1 mm
Copyright © Kirill Makarov

Broscosoma baldense Rosenhauer, 1846: 4
Male adult
Italy, Veneto, Verona, Mt. Baldo, 2088 m, 45*41'40'' N, 10*49'32 E, 26.VIII.2011, E. Ruzzier leg. (Collection Enrico Ruzzier) det. A. Anichtchenko, 2014 ; 9.5 mm
Copyright © Alexander Anichtchenko

Broscus semistriatus (Dejean, 1828)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Mecodema manaia Seldon & Leschnen, 2011
Male adult
Photo taken by S.E. Thorpe 26 mm
Copyright © - - -

Broscus laevigatus Dejean, 1828: 431 (as illustris)
Male adult
ISRAËL région d'Haifa forêt de Kiryat Ala 06-12-2013
Copyright © Pascal Deschamps

Broscus glaber Brulle, 1836
Male adult
Gran Canaria, 1200m, Artenara, El Pinar, 12.1995. det. A. Anichtchenko, 2016
Copyright © Alexander Anichtchenko

Broscus punctatus Dejean, 1828: 431
Female adult
India, Uttarakhand, 17-18.04.2012, Uttarkashi distr., N30*45'15.72" E78*34'32.80, 1374 m, 14 km E Uttarkashi, Anichtchenko A. leg & det.
Copyright © Alexander Anichtchenko

Broscus przewalskii Semenov, 1889d: 393
Male adult
China, Quinghai prov, Lajishan Mts., Sinin vill., 3000m, 22.7.2007, Patrikeev leg. det. A. Anichtchenko 2016
Copyright © Alexander Anichtchenko

Broscus declivis Semenov, 1889d: 392
Male adult
SE Kazakhstan, Ketmen Mt., Tujuk vill., 1800m, 8.V.2000. det. A. Anichtchenko, 2016
Copyright © Alexander Anichtchenko

Broscus crassimargo Wollaston, 1865: 6
Male adult
La Gomera, El Cedro, 23-30.3.2009. (DUBC). det. A. Anichtchenko, 2016
Copyright © Alexander Anichtchenko

Broscus laevigatus Dejean, 1828: 431
Male adult
Israel, Rosh-Pinna, 5.X.2001, A. Zamesov leg. (DUBC). det. A. Anichtchenko, 2016
Copyright © Alexander Anichtchenko

Broscus karelini Zoubkoff, 1837: 65
Male adult
Iran, Golestan, Til Abad, route to Sharood, 6.6.2001. det. A. Anichtchenko, 2016
Copyright © Alexander Anichtchenko

Broscodes karumicus Bolivar & Pieltain, 1914
Male adult
Iran, Fars prov., Tange Tizab, 2015
Copyright © Alexander Anichtchenko

Adotela noctis (Sloane, 1893) Syntypes
Male adult

Copyright © Museums Victoria

Adotela bicolor (Castelnau, 1867) Holotype
Male adult

Copyright © Museums Victoria

Chylnus substriatum (Moore, 1960) Holotype
Male adult

Copyright © Museums Victoria

Mecodema alternans Castelnau, 1867 Holotype
Male adult

Copyright © Museums Victoria

Eurylychnus kershawi Sloane, 1915 Syntypes
Male adult

Copyright © Museums Victoria

Brithysternus nodosum Sloane, 1910 Syntypes
Female adult

Copyright © Museums Victoria

Brithysternus macleayi Sloane, 1910 Holotype
Male adult

Copyright © Museums Victoria

Cerotalis versicolor Castelnau, 1867 Holotype
Male adult

Copyright © Museums Victoria

Adotela frenchi Sloane, 1890 Syntypes
Male adult

Copyright © Museums Victoria

Promecoderus viridiaeneus Sloane, 1915 Holotype
Male adult

Copyright © Museums Victoria

Promecoderus cordicollis Sloane, 1908 Syntypes
Male adult

Copyright © Museums Victoria

Acallistus cuprescens (Sloane, 1920) Syntypes
Male adult
Promecoderus cuprescens Sloane, 1920
Copyright © Museums Victoria

Mecodema kokoromatua Seldon, Leschen & Liebherr, 2012 Paratype
Female adult
Image from Seldon, Leschen & Liebherr, 2012
Copyright © - - -

Mecodema chaiup Seldon, 2015 Holotype
Male adult
NEW ZEALAND, HB, Mohi Bush Scenic Reserve,2 January 2008. D.S. Seldon & C.P. Martin, S39◦5124.5" E176◦5356.2. Image from Seldon, 2015
Copyright © - - -

Brithysternus nodosum Sloane, 1910
Male adult
Australia. 29 mm
Copyright © Mark Golding

Bembidiomorphum convexum Champion, 1918
Male adult
3.5 mm (Collection R. Sciaky)
Copyright © Alexander Anichtchenko

Broscodera (Sinobrosculus) dreuxi Deuve, 1990: 187
Male adult
China, C Sichuan, Jintang, Jiajin Shan 3400 m., 6.VII.2001.
Copyright © Riccardo Sciaky

Kashmirobroscus mirabilis Schmidt, Wrase & Sciaky, 2013 Holotype
Male adult
N Pakistan, NW Frontier prov., Khagan Valley, 2500-3500 m, Naran SE env., 26.27.VII.1998, Leg. Kalab (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Broscosoma farkaci Sciaky & Facchini, 2005
Female adult
China, Tibet, Motuo, Hanmi, 08.2013. det. Anichtchenko A. 2018
Copyright © Alexander Anichtchenko

Broscosoma relictum Weissmandl, 1936
Male adult
Italy, Lombardia, Pizzo Presolana (BG) 2000m, 23.6.2005 F. Angelini
Copyright © Alexander Anichtchenko

Broscosoma kalabianum viridescens Deuve, 2018 Holotype
Male adult
Chine, Sichuan, montagnes à 12 km au SE-SSE de Wodi, 30°40’N-102°14’E, 4400 mètres (pelouses alpines et éboulis), in coll. A. Dostal, Vienne
Copyright © Thierry Deuve

Cascellius gravesii Curtis, 1839
Male adult
Argentina, Tierra del Fuego, Glaciar Martial, 20.II.1996, Leg. Sciaky (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Nothobroscus chilensis Roig-Junient and Ball, 1995
Male adult
Chile, Cherquenco, II.1995 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Percolestus blackburni Sloane, 1892
Male adult
Australia, NSW, Kosciusko Mt., 1900 m, 16.I.1991 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Broscosoma gaoligongense Deuve & Wrase, 2015 Holotype
Male adult
Chine, Yunnan, monts Gaoligong Shan, col à 21 km au nord-ouest de Liuku, 3150 mètres, 25°58’22’’N - 98°41’00’’E, D. W. Wrase, 9.VI.2006 (in coll. D. W. Wrase, Germany). Image from Deuve & Wrase, 2015
Copyright © - - -

Broscosoma sehnali Deuve, 2006 Paratype
Male adult
China, W Sichuan, 5.VI.2006, West of Zier (Zi'er) 4241m, N28*20.886 E101*28.361, R. Sehnal & M. Tryzna leg. (DUBC)
Copyright © Alexander Anichtchenko

Orthoglymma wangapeka Liebherr, Marris, Emberson, Syrett & Roig-Juñent, 2011 Holotype
Male adult
Image courtesy from Liebherr, Marris, Emberson, Syrett & Roig-Juñent, 2011
Copyright © - - -

Broscosoma businskae Dvorak, 1998
Male adult
China, Tibet, Linzhi area, Mt. Serkyla 4000m 8.9.2019 (DUBC) det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Oregus septentrionalis Pawson, 2003 Holotype
Male adult
Image from Lincoln University Entomology Research Collection
Copyright © - - -

Broscosoma qiului Jiang, Liu & Wang, 2021 Holotype
Male adult
China: Chongqing City, Beibei District, Jinyun Village, Jinyun Mountain, elevation 790 m, 29°51’12.21”N, 106°23’01.04”E, 06.VI.2020, Qiu Lu leg. (SCAU).
Copyright © - - -

Broscosoma ribbei rougeriei Deuve & Tian, 2002
Male+Female adult
China, Yunnan Province, Derung-Nu Autonomous County Gongshan, near Gong-Du Tunnel, elevation 3500 m, 2017.07.28, Chen Chao-Ming leg. (LPYC).
Copyright © - - -

Broscosoma guoliangi Jiang, Liu & Wang, 2021 Holotype Paratype
Male+Female adult
China: Fujian Province, Sanming City, Luobading, elevation 1400-1500 m, 2020.05.18, Guo Liang leg. ( SCAU).
Copyright © - - -

Broscosoma zhengyuandongi Jiang, Feng & Wang, 2020 Holotype
Male adult
China: Fujian Province, Longyan City, Xinluo District, Jiangshan Township, Huanglianymeng Mountain, elevation 900 m, 12.II.2018, Yuan-Dong Zheng leg. (SCAU)
Copyright © - - -

Eobroscus bhutanensis Morvan, 1982
Female adult
NEPAL: Lamjung distr. , Manaslu Mountains , Dudh Pokhari Lekh , between Simia Kharka and Malamche Kharka, 3300–3500 m, 12–13 September 1995, J. Schmidt leg. (cSCHM)
Copyright © Joachim Schmidt

Kashmirobroscus ortrudae Schmidt, Wrase & Sciaky, 2013 Paratype
Male adult
Pakistan (Azad Jammu & Kashmir) Noori-Top 3600–4000 m (s. Jalkhad) 15 / 16.VII.2003 Heinz leg.
Copyright © Joachim Schmidt

Chaetobroscus bhutanensis Morvan, 1980
Male adult
Bhutan, 3 October 2017, at Camp 4 Wangchelakh 4170 m, 27°22'32.2''N, 90°19'56.5''E
Copyright © Joachim Schmidt

Broscosoma ribbei Putzeys, 1877
Male adult
CASENT1021037; 2.8 km above Shibali, Lishadi Township,Fugong County, Yunnan, China. scale line = 1.0 mm
Copyright © David Kavanaugh

Broscodera (Sinobrosculus) chukuai Kavanaugh & Liang, 2021 Paratype
Male adult
CASENT1026168; NNE of Chukuai Lake, Bingzhongluo, Gongshan County, Yunnan, China, scale line = 1.0 mm. Image from Kavanaugh & Liang, 2021
Copyright © David Kavanaugh

Broscodera (Sinobrosculus) gaoligongensis Kavanaugh & Liang, 2021 Paratype
Male adult
CASENT1020104; South Fork of Yamu He, LumadengTownship, Fugong County, Yunnan, China, scale line = 1.0 mm
Copyright © David Kavanaugh

Broscosoma bicoloratum Kavanaugh & Liang, 2021 Paratype
Male adult
CASENT1026704; slope NW of Heipu Yakou, DulongjiangTownhip, Gongshan County, Yunnan, China, scale line = 1.0 mm
Copyright © David Kavanaugh

Broscosoma danzhuense Kavanaugh & Liang, 2021 Holotype
Female adult
IOZ, labeled: CASENT 1004637”/ “CHINA, YunnanProvince, Gaoligong Shan, Nujiang Prefecture, Gongshan County, Danzhu He drainage, 13.5-13.8airkm SSW of Gongshan [Cikai], 2720-2840 m,”/ “N27.63267°/E98.60861° to”/“N27.63331°/E98.60356°, 30 June – 5 July 2000, Stop#00-17E, D.H. Kavanaugh, C.E. Groswold[sic], Liang H.-B., D. Ubick, & Dong D.-Z. Collectors
Copyright © David Kavanaugh

Broscosoma furvum Kavanaugh & Liang, 2021 Paratype
Male adult
CASENT1020001; first cirque S of Shibali Yakou, Fugong County, Yunnan, China), scale line = 1.0 mm
Copyright © David Kavanaugh

Broscosoma gaoligongense Deuve & Wrase, 2015
Male adult
CASENT1027285; just W of Fenxue Yakou, Pianma Township, County, Yunnan, China, scale line = 1.0 mm
Copyright © David Kavanaugh

Broscosoma gongshanense Kavanaugh & Liang, 2021 Paratype
Male adult
CASENT1008139; Dongshaofang area, Gongshan County, Yunnan, China, scale line = 1.0 mm
Copyright © David Kavanaugh

Broscosoma holomarginatum Kavanaugh & Liang, 2021 Paratype
Male adult
CASENT1015162; 0.5 km N of Kongdang, DulongjiangTownship, Gongshan County, Yunnan, China, scale line = 1.0 mm
Copyright © David Kavanaugh

Broscosoma parvum Kavanaugh & Liang, 2021 Paratype
Male adult
CASENT1020001; first cirque S of Shibali Yakou, LumadengTownship, Fugong County, Yunnan, China, scale line = 1.0 mm
Copyright © David Kavanaugh

Broscosoma purpureum Kavanaugh & Liang, 2021 Paratype
Male adult
CASENT1015338; 0.5 km N of Kongdang, DulongjiangTownship, Gongshan County, Yunnan, China, scale line = 1.0 mm
Copyright © David Kavanaugh

Broscosoma resbecqi Kavanaugh & Liang, 2021 Holotype
Male adult
IOZ, labeled: “ CASENT 1016251”/ “CHINA, Yunnan,Gongshan County, Dulongjiang Township, Siran Wang, 0.2 km above confluence with DulongJiang, N28.01347°/ E098.32117°,”/ “1720 m, 1 November 2004, Stop # DHK-2004-066, D.H.Kavanaugh & D.-Z. Dong collectors”/ “HOLOTYPE Broscosomaresbecqi Kavanaugh & Liangsp. nov. designated 2021” [red label].
Copyright © David Kavanaugh

Broscosoma viridicollare Kavanaugh & Liang, 2021 Paratype
Male adult
CASENT1022787; 8.5 km above Shibali, Lishadi Twonship, Fugong County, Yunnan, China, scale line = 1.0 mm
Copyright © David Kavanaugh

Anheterus gracilis (Germain, 1848)
Male adult

Copyright © Creative Commons

Mecodema hector Britton, 1949 Holotype
Male adult
New Zealand, Mt Hector, Otago 1910-12-26
Copyright © - - -

Mecodema bullatum Lewis, 1902 Holotype
Male adult
F Sandager. New Zealand, Puysegur [Point]. Museum of New Zealand Te Papa Tongarewa
Copyright © - - -

Mecodema obesum Townsend, 1965
Male adult

Copyright © - - -

Bountya insularis Townsend, 1971 Holotype
Male adult
held at the Auckland War Memorial Museum
Copyright © - - -

Rawlinsius papillatus Davidson and Ball, 1998 Paratype
Male adult
Mexico, Guerrero, 26 km NW El Paraiso, 1986-08-08 (Image from IREKANI, Instituto de Biología)
Copyright © - - -

Barypus (Barypus) speciosus Dejean, 1831
Male adult
Pueblo9 Liebig, provincia de Entre Rios, II-1989
Copyright © Armando Cicchino

Barypus (Barypus) pulchellus Burmeister, 1868
Female adult
Mar del Plata, Provincia de Buenos Aires, XII-2001
Copyright © Armando Cicchino

Barypus (Barypus) rivalis (Germar, 1824)
Male adult
Cinco Cerros, Partido der Balcarce, Bs. As. II-2001
Copyright © Armando Cicchino

Barypus (Barypus) comechingonensis Roig-Junent, 1992
Female adult
Tulumba, Provincia de Cordoba II-1996
Copyright © Armando Cicchino

Barypus (Cardiophthalmus) mendozensis Roig-Junent and Cicchino, 1989
Female adult
La Consulta, Provincia de Mendoza I-2010
Copyright © Armando Cicchino

Barypus (Cardiophthalmus) gentilii Roig-Junent, 1992
Female adult
Collon Cura, Provincia de Neuquen XI-1988
Copyright © Armando Cicchino

Barypus (Cardiophthalmus) clivinoides (Curtis, 1839)
Female adult
Lapataia, Provincia de Tierra del Fuego, I-1974
Copyright © Armando Cicchino
Broscinae taxon page (108 photos)
Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
Female adult

Copyright © Evgeniy Chibilov

Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
Male adult
Russia,
S Ural Mts, Mt.Iremel h=800m, 4.VI.2000 Filimonov V. leg.
male,21mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
Male adult
Carabus (Morphocarabus) karpinskii Rusia, S Ural mts., Jremel Mt., 1300 m 4.6.2000male, 20 mm
19.08.2008
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) karpinskii Kryzhanovskij & Matveev, 1993
Female adult

Copyright © Evgeniy Chibilov

Carabus (Morphocarabus) karpinskii Kryzhanovskij & Matveev, 1993
Female adult

Copyright © Evgeniy Chibilov

Carabus (Diocarabus) loschnikovi loschnikovi Fischer, 1823
Male adult
Komi Resp., Pecero-Ilych Nat. Res., Zajachij Kamen h~786 62°10'06" N 59°12'53" E 8.VIII.2004 M. Chernjakhovski leg.
L=15.0 mm
Copyright © Evgeniy Zinoviev

Aplothorax burchelli Waterhouse, 1841
Male adult
St. Helena island
Copyright © Boleslav Brezina

Calosoma (Calosoma) inquisitor cupreum Dejean, 1826: 196
Female adult
Armenia, Zangezur range, Meghri suburbs., Vahravar vill., Rivulet vall., h=1400 m Yuriy Shevnin leg.; 23.05.2007; 21 mm.
Copyright © Anton Menstschikov

Calosoma (Calosoma) inquisitor cupreum Dejean, 1826: 196
Male adult
Iran, Ardabil, 30 km NE Ardabil,
5 km SE Namin, 18-19.5.2007 Anichtchenko A. leg.
male, 20 mm
Copyright © Alexander Anichtchenko

Calosoma (Calosoma) schayeri schayeri Erichson, 1842
Male adult
Australia N.T. Alice Springs, Airport
19.5.2000 M. Langer leg.
male, 21 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisphaena) reticulatum reticulatum s.str.
Female adult
Germania, or. DDR, Bezirk Frankfurt, 24.V.1989
female, 25 mm
photo & det. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Callistenia) moniliatus Le Conte, 1851: 200
Male adult
SE Canada, Nueva Escocia
19.5.1970
male, 17 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) anthrax Semenov, 1900 Holotype
Male adult
collection of ZIN
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) breviusculus Mannerheim, 1830
Male adult
Armenia, near Ashtarik 15.6.1997 male, 22 mm photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) elegans s.str. (as declivis Dohrn, 1884b)
Male adult
SE Kazakhstan Alma-Ata reg., 30 km N Sary-Ozek 28.4.1998 Klimenko leg. male, 22 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) elegans s.str. (as declivis Dohrn, 1884b)
Female adult
SE. Kazakhstan, Sary-Olek vill., 26.IV.1997
Copyright © Anton Menstschikov

Calosoma (Callisthenes) elegans amethystinus Semenov & Redikorzev, 1928
Male adult
SE Kazakhstan, E Zailiyski mt. rng. Chilik riv.
Malybai vill. 28.4.1999 Klimenko A. leg.
male, 25 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) elegans amethystinus Semenov & Redikorzev, 1928
Male adult
26 mm.
Zailiyski Alatay.
06.1997 Chibilov E. leg.
Copyright © Sunvas

Calosoma (Callisthenes) elegans amethystinus Semenov & Redikorzev, 1928
Female adult
SE Kazakhstan, E Zailiyski mt. rng. Chilik riv.
Malybai vill. 28.4.1999 Klimenko A. leg.
female, 26 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) elegans elegans s.str.
Male adult
SE Kazakhstan, Tchu-Iliyskiy Mts., Kordai pass, h=1500; A. Klimenko leg.; 20.04.1997;
24 mm.
photo A. Menstschicov
Copyright © Anton Menstschikov

Calosoma (Callisthenes) elegans karagaicus Lapouge, 1924
Male adult
Inner Tian-Shan, 20 km N jf Naryn
27.5.1997 Klimenko A. leg.
male, 27 mm
photo, det. & coll. Anichtchenko A. 2009
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) elegans karagaicus Lapouge, 1924
Male adult
KIRGHIZSTAN, Inner Tian-Shan Mts., near Naryn tn., Naryn riv. Valley, 18-20.V.1999, Leg. Irtlatch A.
Copyright © Anton Menstschikov

Calosoma (Callisthenes) elegans manderstjernae Ballion, 1870
Female adult
Kirgizia, S Kirgizskii mt. rng., 2200m
Kichi-Kimin riv., Ak-Tuz vill. 05.1998 Klimenko leg.
female, 32 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) elegans semenovi Motschulsky, 1859
Female adult
SE Kazakhstan, Zailijsky Mt. rng.,
Talgar env., 1500 m, 21.5.2000
female, 27 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) eversmanni Chaudoir, 1850
Male adult
Syria, Antilibanon
1.3.1991
male, 22 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) fischeri Fischer, 1842
Male adult
Mongolia, Mandalgob, 20 km
N Amardalai vill. 11-12.6.2003 Anichtchenko A. leg
male, 19 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) grumi Semenov, 1900 Holotype
Female adult
collection ZIN
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) breuningi Mandl, 1954
Female adult
E Kazakhstan, S Alakol lake, Koktuma vill., 1-10.5.1999 Nekroshevich S. leg. female, 22 mm; det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) breuningi Mandl, 1954
Male adult
E.Kazakhstan, Alakol lake, Koktuma vill., 07.V.1999
Copyright © Anton Menstschikov

Calosoma (Callisthenes) breuningi iliense Kabak, 2022
Male adult
SE Kazakhstan, near Sary Ozek vill., A. Klimenko leg; 22.04.1997; 22 mm.; det. Anichtchenko A. 2022
Copyright © Anton Menstschikov

Calosoma (Callisthenes) kuschakewitschi kuschakewitschi s.str.
Male adult
Karzhantau mt. rng., Susungen riv.,
2300 m, 7.6.1998 Tretjakov leg.
male, 23 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) kuschakewitschi plasoni Born, 1917
Male adult
Kirgizia, 40 km S Bishkek
10.5.1990
male, 26 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) kuschakewitschi plasoni Born, 1917
Female adult
Kirgizia, Chon-Aryk. 23.V.1998 V. Dolin leg.
Copyright © Denis Potanin

Calosoma (Callisthenes) marginatus Gebler, 1829
Female adult
NE Kazakhstan, N Zaisan lake, 20.6.2000
Narymskii mt.rng. Klimenko A. leg.
female, 18 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) mirificus Casale, 1979
Female adult
Afganistan, S slope of Paropamiz Mt. rng., near Sabzan pass, 2400m, 22.VI.1967 M.Stoljarov leg.
Collection of ZIN
det. D. Obydov 1999
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) panderi Fischer, 1822
Male adult
NW Kazakhstan, Aktiubinsk reg., Baiganinsk distr.,
near Zharly vill. 10-12.05.1997 leg. M.& L. Smirnov
male, 20 mm
Photo from www.zin.ru
Copyright © Maksim Smirnov

Calosoma (Callisthenes) pavlovskii Kryzhanovskij, 1955
Male adult
Kirgizstan, Talas Mt. rng., Besh-Tash
riv., 1000m, 15.5.2003
male, 20 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) pseudocarabus Semenov et Redikorzev, 1928
Male adult
NE Kazakhstan, Saur mt.rng. 2000m
Sargar mt., 14.6.2001 Klimenko leg.
male, 16 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) pseudocarabus Semenov et Redikorzev, 1928
Female adult
E Kazakstan, NE Saur, 2 km. W m. Tas, h=2500; 10.06.2001, Ivanov A.V. leg.,
15 mm., female
photo, det. & coll. Menstschikov
Copyright © Anton Menstschikov

Calosoma (Callisthenes) pseudocarabus Semenov et Redikorzev, 1928
Male adult
E Kazakstan, NE Saur, 2 km. W m. Tas, h=2500; 10.06.2001, Ivanov A.V. leg., 14 mm., male
photo, det. & coll. Menstschikov
Copyright © Anton Menstschikov

Calosoma (Callisthenes) regelianus A. Morawitz, 1886
Female adult
Tadzikistan, Darvaz Ezgan 2.5.1971
female, 25 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) usgentensis usgentensis Solskyi, 1874
Male adult
Kirghizstan, near Jalal-Abad town, A. Klimenko leg., 05.04.1997; 30 mm.
photo, det. & coll. A. Menstschicov
Copyright © Anton Menstschikov

Calosoma (Callisthenes) usgentensis usgentensis Solskyi, 1874
Female adult
S Kirgizia, Dzhalal-Abad reg.,
Suzak vill., 1000m, 25.5.1999
male, 27 mm
photo, det. & coll. A. Anichtchenko
Copyright © Alexander Anichtchenko

Calosoma (Callitropa) externum Say, 1823: 149
Female adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Calosoma (Calosoma) scrutator Fabricius, 1775
Female adult
USA, Illinois, Peoria Co., Orange Prairie, 31.5.1990 32 mm coll. A.Montes photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Calosoma) scrutator Fabricius, 1775
Male adult
3 m SE Tishomingo , Tishomingo Co., Mississippi, USA; 24.VI.2005 Eric B. Lago leg.
Copyright © Denis Potanin

Calosoma (Calosoma) wilcoxi Le Conte, 1848
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Calosoma (Calosoma) wilcoxi Le Conte, 1848
Male adult
2 m. W Rawlings, Allegany Co., Maryland, USA; 26 May 1983 J.D. Glaser leg.
Copyright © Denis Potanin

Calosoma (Calosoma) cyanescens Motschulsky, 1859
Male adult
Primorskii reg., Kedrovaja Pad Nat.res., 11.VII.1994 Chichvarhin A. leg
male, 19 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Calosoma) frigidum Kirby, 1837: 19
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Calosoma (Calosoma) frigidum Kirby, 1837: 19
Male adult
2 m W Rawlings, Allegany Co., Maryland, USA; 26.V.1983 J.D. Glaser leg.
Copyright © Denis Potanin

Calosoma (Calosoma) maximowiczi A. Morawitz, 1863: 20
Female adult
China, Mt. Wangfoshan (1940 m.), Baokang Countri, Hubei; 20-28.06.2008, 30 mm.
Copyright © Anton Menstschikov

Calosoma (Calosoma) oceanicum klynstrai Breuning, 1927
Female adult
INDONESIA, Tanimbar Island, III.2009
photo & coll. N. Belonogov
Copyright © Nil Belonogov

Calosoma (Calosoma) oceanicum klynstrai Breuning, 1927
Male adult
INDONESIA, Tanimbar Island, III.2009
photo & coll. N. Belonogov
Copyright © Nil Belonogov

Calosoma (Calosoma) sycophanta Linne, 1758
Male adult
Spain, Sierra Nevada, Pico de Veleta
2800m
Copyright © Alexander Anichtchenko

Calosoma (Carabosoma) prominens LeConte, 1853
Male adult
USA Arizona, Cochise Co., Interstate 8 at Texas, Canyon Rest Area, 1500m, 32º00.3N 110º08.4W, 24.7.1996 Kavanaugh D. leg.; 30 mm
Copyright © Alexander Anichtchenko

Calosoma (Carabosoma) peregrinator Guerin-Meneville, 1844
Male adult
Fort Davis, Jeff Davis Co., Texas, USA; July 1998
Copyright © Denis Potanin

Calosoma (Caminara) denticolle Gebler, 1833: 274
Female adult
Burjatia, Selenga riv., Deben vill. 12.7.2003 Anichtchenko A. leg. female, 25 mm
Copyright © Alexander Anichtchenko

Calosoma (Caminara) imbricatum deserticola Semenov, 1897d: 242
Male adult
W Kazakstan, Ustjurtskij res., Kordon Kenderli; 01.05.2009, Gorbunov P.J. leg.,
25 mm., male
photo, det. & coll. Menstschikov
Copyright © Anton Menstschikov

Calosoma (Caminara) imbricatum deserticola Semenov, 1897d: 242
Male adult
W Kazakstan, Ustjurtskij res., Kordon Kenderli; 01.05.2009, Gorbunov P.J. leg., 24 mm., female photo, det. & coll. Menstschikov
Copyright © Anton Menstschikov

Calosoma (Campalita) chlorostictum Dejean, 1831a: 558
Male adult
RSA, Kwazulu/Natal, Badplaas, 1000m, 1.XII.2000 male, 24 mm det. P. Schüle photo &. coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Carabophanus) arrowi Breuning, 1928
Male adult
Ethiopia, Oromia reg., Arsi highlands, Wahoro env., Tiliku Lensho river valley10.-12.II.2010, J.Štastny lgt
photo Jaroslav Stastny
Copyright © Jaroslav Štastny

Calosoma (Castrida) alternans granulatum Perty, 1830
Male adult
Paraguay, Dep.Caaguazu, Tayao S25.28 W56.07 12-16.III.2003 leg. U.Drechsel
male, 28 mm
photo, det. & coll. Anichtchenko A. 2009
Copyright © Alexander Anichtchenko

Calosoma (Castrida) alternans granulatum Perty, 1830
Female adult
Argentina, Entre Rios,
Liebig, nov.1997
long. 32 mm
photo, det. & coll. Anichtchenko A. 2009
Copyright © Alexander Anichtchenko

Calosoma (Castrida) argentinensis Csiki, 1927
Male adult
Argentina, Prov. Santa Fe Coronda, 22.12.1965 leg. Mahunka, 23 mm
Copyright © Alexander Anichtchenko

Calosoma (Castrida) argentinensis Csiki, 1927
Male adult
ARGENTINA: Salta; November 1995
Copyright © Denis Potanin

Calosoma (Castrida) vagans Dejean,
Male adult
Chile, Santiago de Chile, Quilicura
19.9.1991 G. Aceituno leg.
male, 21 mm
Copyright © Alexander Anichtchenko

Calosoma (Charmosta) investigator Illiger, 1798
Male adult
Lake Baikal, Buriatia, Iliynka vill., 1.8.1997 Anichtchenko A. leg.; 20 mm
Copyright © Alexander Anichtchenko

Calosoma (Charmosta) investigator Illiger, 1798
Male adult
S. Ural, Bashkiria, near Mesjagutovo vill., 09.07.2001 A. Menstschicov leg.
Copyright © Anton Menstschikov

Calosoma (Charmosta) lugens Chaudoir, 1869f 372
Male adult
Wen-dong, North Fengcheng City, Liaoning, China; 1-5.VII.2006
Copyright © Denis Potanin

Calosoma (Chrysostigma) calidum Fabricius, 1775
Male adult
Canada Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Calosoma (Chrysostigma) semilaeve LeConte, 1851: 199
Male adult
Orange Co., California, USA; May 1973
Copyright ©

Calosoma (Ctenosta) bastardi Alluaud, 1925
Female adult

Copyright © Alexander Anichtchenko

Calosoma (Ctenosta) orientale Hope, 1838
Male adult
China, Yunnan pro.
Copyright © Ye Liu

Calosoma (Ctenosta) planicolle Chaudoir, 1869
Male adult
length: 30 mm. Kenia, Tsavo National Parc, leg. U.Schmidt, 1991; det. W.Lorenz, 2000. Foto: U.Schmidt, 2006
Copyright © U.Schmidt

Calosoma (Ctenosta) planicolle Chaudoir, 1869
Male adult
SOUTH AFRICA: Northern Transvaal
Copyright © Denis Potanin

Calosoma (Ctenosta) senegalense Dejean, 1831
Female adult
length: 22-27 mm
Kenia Tsavo National Parc
leg. U.Schmidt, 1991; det. W.Lorenz, 2000
Foto: U.Schmidt, 2006
Copyright © U.Schmidt

Calosoma (Microcallisthenes) relictum Apfelbeck, 1918c
Male adult
Macedonia, Shar mt., Peak Tserapashina
1850-2530m 9.7.1995
male, 13 mm
Copyright © Alexander Anichtchenko

Calosoma (Microcallisthenes) relictum Apfelbeck, 1918c
Male adult
Macedonia, Shar mt., Peak Titov Vrah
2747 m, 14.7.1995 V.Sakalian leg.
male, 13 mm
Copyright © Alexander Anichtchenko

Calosoma (Calosoma) inquisitor inquisitor Linne, 1758: 414
Male adult
length: 13-24 mm
England, Dendless Wood
leg.det. M.G.Telfer, 2003
Foto: U.Schmidt, 2009
Copyright © U.Schmidt

Calosoma (Calosoma) himalayanum Gestro, 1875b: 851
Female adult
China, Tibet
Copyright © Ye Liu

Calosoma (Carabops) abyssinicum Gestro, 1880
Male adult
Ethiopie, Shemene 08.2004
19 mm
coll. A.Montes
Copyright © Alexander Anichtchenko

Calosoma (Carabops) abyssinicum Gestro, 1880
Male adult
Ethiopia, Bale prov., Str. ZW. Sasheme u. Dodola, Mt. Mascall 3100-3300 m
21.VII.2005 Puchner leg.
male, 18 mm
det. Puchner F. 2005
photo & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Ceroglossus buqueti deuvei Jiroux, 1996
Male adult
Chile, VIII Reg., 1000-1250 m, P.N. Nahuelbuta, (Piedra del Aguila) 7-12.XII.1999 Heinz leg.
male, 22 mm
photo, det. & coll. Anichtchenko A. 2009
Copyright © Alexander Anichtchenko

Ceroglossus buqueti andestus Kraatz-Koschlau, 1887
Male adult
CHILE, Liquine; 05/12.02.2005; C.Saavedra leg.; Determination confirmed by Anichtchenko, 2019
Copyright © Anton Menstschikov

Ceroglossus buqueti lepidus Kraatz-Koschlau, 1891 (as cherquencoensis)
Male adult
Chile, IX reg., Cherquenco Vn. Llaima, 800-1000m, 29-30.I.1999 Heinz leg. 21 mm; det. & coll. Anichtchenko A. 2019
Copyright © Alexander Anichtchenko

Ceroglossus buqueti sybarita Gerstaecker, 1858
Female adult
Chile, Chiloe, 160m, 8 km N Butalcura 22-24.XII.1998 det. Heinz
female, 24 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Ceroglossus chilensis cyanicollis Kraatz, 1887
Male adult
Chile (VIII reg), Umg. Lag. de la Laja, 950 m, 19-20.12.1998 Heinz leg. 26 mm; det. Anichtchenko A., 2019
Copyright © Alexander Anichtchenko

Ceroglossus chilensis fallaciosus Kraatz, 1880
Male adult
CHILE, Talca prov., Constitucion tn. env., II.2008. det. Anichtchenko A., 2019
Copyright © Nil Belonogov

Ceroglossus chilensis ficheti Jiroux, 1996
Male adult
Chile (VIII reg), P.N. Nahuelbuta, (Piedra del Aguila) 700 m, 7-12.12.1999, Heinz leg.; 28 mm. Determination confirmed by Anichtchenko, 2019
Copyright © Alexander Anichtchenko

Ceroglossus chilensis gloriosus Gerstaecker, 1858
Male adult
Chile (IX reg.), Malalcahuello, 800 m, 9-10.12.1999 Heinz leg. 25 mm; Determination confirmed by Anichtchenko, 2019
Copyright © Alexander Anichtchenko

Ceroglossus chilensis latemarginatus Kraatz-Koschlau, 1889
Male adult
Chile (VIII. Reg.) Pata de Gallina, 650m, b.Contulmo 7-12.12.1999 Heinz leg. 27 mm; Determination confirmed by Anichtchenko, 2019
Copyright © Alexander Anichtchenko

Ceroglossus chilensis mochae Reed, 1874
Female adult
CHILE, Arauco prov., Isla Mocha. - II.2008. photo Belonogov N.; Determination confirmed by Anichtchenko, 2019
Copyright © Nil Belonogov

Ceroglossus chilensis resplendens Jaffrezic & Rataj, 2006
Male adult
CHILE, Huife, IX Region. Determination confirmed by Anichtchenko, 2019
Copyright © Anton Menstschikov

Ceroglossus chilensis seladonicus Kraatz-Koschlau, 1887
Male adult
CHILE, Malleco prov., Malalcahuello Nat. Res. - I.2008
photo Belonogov N.
Copyright © Nil Belonogov

Ceroglossus chilensis seladonicus Kraatz-Koschlau, 1887
Female adult
Chile, Malleco prov., Madalcahuello
enero 2003 G. Aceituno leg.
28 mm
Copyright © Alexander Anichtchenko

Ceroglossus chilensis solieri Roeschke, 1900
Female adult
Chile, Chiloe, 160m, 8 km N Butalcura 22-24.XII.1998 det. Heinz
female, 28 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Ceroglossus darwini darwini s.str.
Male adult
Chile, Chiloe, ca. 25 km NW Queilen, 350 m, (n. Castro), 26-27.XII.1999 Leg. Heinz
male, 22 mm
photo, det. & coll. Anichtchenko A. 2009
Copyright © Alexander Anichtchenko

Ceroglossus resplendens Jaffrezic & Rataj, 2006
Female adult
CHILE. X Region, Termas de Carranco; 01/05 February. 2007; C.Saavedra leg.; det. Anichtchenko A. 2019
Copyright © Anton Menstschikov

Ceroglossus speciosus Gerstaecker, 1858
Female adult
Chile, Chiloe, S. Pedro de Piruquiña 350 m, (n. Castro), 24-25.XII.1999 Heinz leg.
female, 23 mm
photo, det. & coll. Anichtchenko A. 2009
Copyright © Alexander Anichtchenko

Calosoma (Teratexis) fabulosum Semenov & Znojko, 1933 Holotype
Male adult
collection of ZIN
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko

Calosoma (Teratexis) fabulosum Semenov & Znojko, 1933 Paratype
Male adult
collection of ZIN
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko

Cychrus sehnali Haeckel, 2007 Holotype
Male adult

Copyright © Boleslav Brezina

Cychropsis (Sinocychropsis) tryznai Haeckel & Sehnal, 2007 Holotype
Male adult
China - W Sichuan, 5.6.2006 West of Zhier (Zier) 4118 M
alpine forest N 28.20.507. E 101.28.014 leg. R Sehnal & M. Tryzna
Holotype
Copyright © Boleslav Brezina

Cychrus angustatus Hoppe & Hornschuch, 1825
Male adult
Italia, Lombardia (BG), Arera mt., 2100m Zambla, 19.6.2003 A. Degiovanni leg. det. V. Monzini, 2014 male, 20 mm
Copyright © Alexander Anichtchenko

Cychrus belousovi O.Berlov, 1997 Holotype
Male adult

Copyright © Oleg Berlov

Cychrus inessae O.Berlov, 1997 Holotype
Male adult

Copyright © Oleg Berlov

Cychrus tuberculatus Harris, 1839
Female adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Cychropsis (Shuocychropsis) paramontana Sehnal & Hackel, 2006 Holotype Holotype
Male adult

Copyright © Boleslav Brezina

Cychrus volkovi O.Berlov, 1997 Holotype Holotype
Male adult

Copyright © Oleg Berlov

Scaphinotus (Scaphinotus) viduus Dejean, 1826
Male adult
Canada Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Scaphinotus (Steniridia) guyoti LeConte, 1866
Male adult
USA Georgia, Brasstown, Towns Co.,
Bald Mt., 16.7.2000 Sehnal leg.
male, 23 mm
Copyright © Alexander Anichtchenko

Scaphinotus (Nomaretus) bilobus Say, 1825
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Sphaeroderus canadensis Chaudoir, 1861
Male adult
Canada Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Sphaeroderus stenostomus lecontei Dejean, 1826
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Sphaeroderus nitidicollis (as brevoorti LeConte, 1848)
Female adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Pamborus alternans Latreille, 1817
Female adult
Australia, NSW. Kerewong S.F.
21.12.1978
30 mm
Copyright © Alexander Anichtchenko

Pamborus brisbanensis Castelnau, 1867
Male adult
Museum of Zurich
photo Anichtchenko
Copyright © Alexander Anichtchenko

Pamborus guerini Gory, 1830
Male adult
Museum of Zurich
photo Anichtchenko
Copyright © Alexander Anichtchenko

Pamborus opacus Gehin, 1885
Male adult
Museum of Zurich
photo Anichtchenko
Copyright © Alexander Anichtchenko

Pamborus pradieri darlingtoni Banninger, 1940 Paratype
Male adult
Museum of Zurich photo Anichtchenko
Copyright © Alexander Anichtchenko

Pamborus pradieri darlingtoni Banninger, 1940
Female adult
Australia, N.S.W. New England rng.,
Moonan Flats, 18.12.2000 T.Pollak leg.
21 mm
Copyright © Alexander Anichtchenko

Pamborus pradieri pradieri Chaudoir, 1869
Male adult
Museum of Zurich
photo Anichtchenko
Copyright © Alexander Anichtchenko

Pamborus punctatus Darlington, 1961 Paratype Paratype
Male adult
Museum of Zurich
photo Anichtchenko
Copyright © Alexander Anichtchenko

Pamborus tropicus Darlington, 1961 Paratype Paratype
Male adult
Museum of Zurich
photo Anichtchenko
Copyright © Alexander Anichtchenko

Carabus (Acathaicus) alexandrae alexandrae Semenov, 1887 Holotype Holotype
Male adult
Kwantin and Li-tschara-pu, 22.VI.1885, leg. Potanin Deposited in Zoolog. Museum St. Petersburg. male, long. 22 mm photo B. Brezina
Copyright © Boleslav Brezina

Carabus (Acathaicus) alexandrae alexandrae Semenov, 1887
Male adult
male, 25mm photo B. Brezina
Copyright © Boleslav Brezina

Carabus (Acathaicus) alexandrae alexandrae Semenov, 1887
Male adult
female, 28mm photo B. Brezina
Copyright © Boleslav Brezina

Carabus (Acathaicus) alexandrae alexandrae Semenov, 1887
Male adult
female, 24mm photo B. Brezina
Copyright © Boleslav Brezina

Carabus (Acathaicus) alexandrae cratocephaloides Semenov, 1887 Holotype Holotype
Male adult

Copyright © Boleslav Brezina

Carabus (Acathaicus) alexandrae cratocephaloides Semenov, 1887
Male adult
male, 26mm
Copyright © Boleslav Brezina

Carabus (Acathaicus) alexandrae cratocephaloides Semenov, 1887
Male adult
male, 24mm
Copyright © Boleslav Brezina

Carabus (Acathaicus) alexandrae cratocephaloides Semenov, 1887
Female adult
female, 28mm
Copyright © Boleslav Brezina

Carabus (Acathaicus) idolon Semenov, 1898 Holotype Holotype
Male adult
male
Copyright © Boleslav Brezina

Carabus (Acathaicus) interruptocostatus interruptocostatus s.str.
Female adult
female, 27mm
Copyright © Boleslav Brezina

Carabus (Acathaicus) interruptocostatus interruptocostatus s.str.
Male adult
male, 29mm
Copyright © Boleslav Brezina

Carabus (Acathaicus) interruptocostatus Semenov, 1887 Holotype Holotype
Female adult

Copyright © Boleslav Brezina

Carabus (Acoptolabrus) constricticollis constricticollis Kraatz, 1886
Female adult

Copyright © Alexander Anichtchenko

Carabus (Acoptolabrus) gehinii gehinii (as aereicollis Hauser, 1921)
Male adult

Copyright © Alexander Anichtchenko

Carabus (Acoptolabrus) gehinii gehinii Fairmaire, 1886
Male+Female adult
Komari, Chitose, Hokkaido, Japan.
20-27.05.2007
Copyright © Safronov

Carabus (Acoptolabrus) gehinii gehinii Fairmaire, 1886
Male+Female adult
Bibi, Chitose, Hokkaido, Japan.
03.05.2000
Copyright © Safronov

Carabus (Acoptolabrus) gehinii gehinii (as manoianus (Imura, 1989)
Male adult

Copyright © Alexander Anichtchenko

Carabus (Acoptolabrus) gehinii munakatai (Ishikawa, 1968)
Male+Female adult
Mt.Shiribetsudake, Kishibetsu, Hokkaido, Japan.
05.06.2005
Copyright © Safronov

Carabus (Acoptolabrus) leechi leechi Bates, 1888
Male adult

Copyright © Boleslav Brezina

Carabus (Acoptolabrus) leechi nangminicola Deuve & Li, 2001
Female adult
N Coree, Cha Gangdo, Nangnimsan Mt., 6.2006. 31 mm. (coll. A. Montes, Madrid)
Copyright © Alexander Anichtchenko

Carabus (Acoptolabrus) lopatini Morawitz, 1886
Male+Female adult
Russia,
Sakhalin isl., Tchekhov Mt, VI.1997
female, 40mm
photo & coll. Panin R.
Copyright © Ruslan Panin

Carabus (Acoptolabrus) mirabilissimus rigoutianus Deuve & Li, 1998
Female adult
N Koree, Pyonganbukto, mt. Miaoxiang 6.2006
28 mm
coll. A.Montes
Copyright © Alexander Anichtchenko

Carabus (Acoptolabrus) schrencki schrencki (as hauryi Gehin, 1885 )
Male adult

Copyright © Boleslav Brezina

Carabus (Acoptolabrus) schrencki schrencki (as hauryi Gehin, 1885 )
Male adult
det. Brezina B.
Copyright © Boleslav Brezina

Carabus (Acoptolabrus) schrencki schrencki (as hauryi Gehin, 1885 )
Male adult

Copyright © Boleslav Brezina

Carabus (Acoptolabrus) schrencki schrencki Motschulsky, 1860
Female adult
Primorie, Lazovskii distr., Chandalaz mt., L.Voita leg 19.07.2008
photo A.Menstschicov
Copyright © Anton Menstschikov

Carabus (Acoptolabrus) schrencki kimieae Kezuka & Imura, 1989
Male+Female adult
Mt. Wulongshan, Dandong City,
Liaoning Province, CHINE
25-30.VI.2008
Copyright © Safronov

Carabus (Acrocarabus) callisthenoides Semenov, 1888
Male adult
male, 22mm
Copyright © Boleslav Brezina

Carabus (Acrocarabus) callisthenoides Semenov, 1888
Male adult
female, 23mm
Copyright © Boleslav Brezina

Carabus (Acrocarabus) guerini aldarkose Kabak, 1992
Male adult
male, 22mm
Copyright © Boleslav Brezina

Carabus (Acrocarabus) guerini guerini Fischer, 1842
Male adult
male, 21mm
Copyright © Boleslav Brezina

Carabus (Acrocarabus) guerini guerini Fischer, 1842
Female adult
female, 25mm
Copyright © Boleslav Brezina

Carabus (Acrocarabus) guerini guerini Fischer, 1842
Male adult
SE Kazakhstan, W Dzhungar Mts, Lepsinsk env., 1600 m., 18.06.2000, A.Klimenko
Copyright © Anton Menstschikov

Carabus (Alipaster) barovskii bayanbulak Deuve, 1993 Paratype Paratype
Male adult
male, 15mm
Copyright © Boleslav Brezina

Carabus (Alipaster) barovskii bayanbulak Deuve, 1993 Paratype Paratype
Female adult
female, 17mm
Copyright © Boleslav Brezina

Carabus (Alipaster) infantulus infantulus Morawitz, 1886
Male adult
male, 15mm
Copyright © Boleslav Brezina

Carabus (Alipaster) pupulus pupulus Morawitz, 1889
Male adult
Inner Tien-Shan, W part of Terskei mnts., Ukek rng., 20-25 km E Kotshkorka, h=2800 m, 8.07.98 leg. Tretyakov male 13,6mm
Copyright © Maksim Smirnov

Carabus (Alipaster) pupulus pupulus Morawitz, 1889
Male adult
male, 13mm
Copyright © Boleslav Brezina

Carabus (Alipaster) pupulus pupulus Morawitz, 1889
Male adult
female, 16mm
Copyright © Boleslav Brezina

Carabus (Alipaster) pupulus pupulus Morawitz, 1889
Male adult
Kirgizstan, Terskei AlaToo Mt. rng, Kadzhi-Sai v., 2800 m.; A.Subankulov leg.; 30.05.2000
Copyright © Anton Menstschikov

Carabus (Apoplesius) deuvei hajdaji Hackel, 2007 Holotype Holotype
Male adult
Turkey- prov. Mus 60 km NW Mus 1630m
3856N 4108E Bulgan Gec. 23-24.06.03 E. & P. Hajdaj leg.
Copyright © Boleslav Brezina

Carabus (Apotomopterus) anxiensis Kleinfeld, 1998 Holotype & Paratype Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) arrowi hector Hauser, 1924
Female adult
China, Guangxi pro., Maoershan Mountain, Liang H.B. coll. Photo by Liuye det. Anichtchenko A.
Copyright © Ye Liu

Carabus (Apotomopterus) bousquetellus gengmaensis Kleinfeld, 2003 Holotype & Paratype Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) cantonensis pervarius Kleinfeld, 1997 Holotype Holotype
Male adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) cantonensis pervarius Kleinfeld, 1997
Male adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) casaleianus jigongshanicus Kleinfeld, 1998 Holotype & Paratype Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) clermontianus Breuning, 1933
Female adult
N Viet-Nam, 100 km NW Hanoi 5.1995 Rapaltovskii E. leg.
Copyright © Alexander Anichtchenko

Carabus (Apotomopterus) datianshanicus Kleinfeld, 1997 Holotype & Paratype Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) davidis davidis Deyrolle & Fairmaire, 1878
Female adult

Copyright © Alexander Anichtchenko

Carabus (Apotomopterus) cantonensis metallaicus Kleinfeld, 1999 Holotype Holotype
Female adult
as pervarius metallaicus
Copyright © Frank Kleinfeld

Carabus (Apotomopterus) davidis neodavidis Kleinfeld & Schutze, 2001 Holotype & Paratype Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) davidis viridithoracis Kleinfeld & Schutze, 2001 Holotype & Paratype Holotype Paratype
Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) davidis yizhangensis Kleinfeld, 2002 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) delavayi s.str. (as medea Kleinfeld, 1999) Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) delavayi tenuimanus Deuve & Imura, 1990
Male adult
Lijiang Yunnan Chine
photo Daniel Prunier
Copyright © Daniel Prunier

Carabus (Apotomopterus) delavayi tenuimanus Deuve & Imura, 1990
Female adult
Lijiang, Yunnan, Chine
female
photo Daniel Prunier
Copyright © Daniel Prunier

Calosoma (Campalita) chlorostictum Dejean, 1831a: 558
Male adult
La Mandraka, Centre Est, Madagascar; December 1995
Copyright © Denis Potanin

Carabus (Morphocarabus) illigeri illigeri (as bjelasnicensis Apfelbeck, 1902)
Male+Female adult
Serbia, Debelo Brdo, 5.VI.2007, S. Sapaly leg. L=32 mm photo Panin R.
Copyright © Ruslan Panin

Carabus (Archicarabus) monticola fontanae Born, 1906
Male+Female adult
N-Italy,
Piemonte, Bielmonte, h1350 m, 14-15.VI.2010, Pascal Stefani leg.
female, 23 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Morphocarabus) kollari Palliardi, 1825
Female adult
female, 24 mm
Copyright © Boleslav Brezina

Carabus (Morphocarabus) kollari Palliardi, 1825
Male adult
male, 27 mm
Copyright © Boleslav Brezina

Carabus (Trachycarabus) estreicheri Fischer, 1822
Male+Female adult
W UKRAINE, NW Podolian,
Lviv Region, Brody distr., near Yasenov vill., steppe sites, 2-28.V.2010, Panin R. leg.
female, 23mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) estreicheri Fischer, 1822
Male adult

Copyright © Safronov

Carabus (Trachycarabus) estreicheri Fischer, 1822
Female adult
W-UKRAINE, NW Podolian,
Lviv Region, Brody distr., near Yasenov vill., steppe sites, 12.V-2.VI.2007 Panin R. leg.
female, 22mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei Kuster, 1846
Male+Female adult
W.Ukraine.
Transcarpathian region,Vinogradiv distr., near Malaya Kopanya vill., Chorna Gora Mt., h=180m. 7-18.VII.2008. Panin R. leg.
female 33 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei Kuster, 1846
Male adult
W.Ukraine.
Transcarpathian region,Vinogradov distr., near Malaya Kopanya vill., Chorna Gora Mt., h=180m. 7-18.VII.08. Panin R. leg.
male, 28 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei Kuster, 1846 (as ormayi Reitter, 1896) loc.typ.
Female adult
W.UKRAINE,
Transcarpathian reg., near Beregovo,15.V-16.VI.2009 Kanarsky Yu. & Geryak Yu. leg.
female, 30 mm
photo/det. Panin R.
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei Kuster, 1846
Female adult
Carabus (Morphocarabus) rothi hampei
W Ukraine, Zakarpatian reg., Vinogradov
1.7.1989 Avdeev I. leg.
female, 31 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) rothi hampei Kuster, 1846
Female adult
arabus (Morphocarabus) rothi hampei
W Ukraine, Zakarpatian reg., Vinogradov
1.7.1989 Avdeev I. leg.
female, 31 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) spasskianus schestopalovi Plutenko, 1995
Male adult
NE-Kazakhstan,
S-Altay, Sarym-Sakty Mt. h~2500m., Sarym-Sakty riv. 24.VI.1999
male, 23mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus schestopalovi Plutenko, 1995
Male adult
SE Kazakhstan,
S Altay Mts., Sarymsakty mt. rng., Sarymsakty riv.
22.VI.2000 2400m, Nekroshevich S. leg.
male, 20 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) kollari Palliardi, 1825
Male+Female adult
Romania, SW Carpathians,
Banat, Anina Mts. (Muntii Aninei) 8 km NE Oravita, 7.VI.2007, S.I. Budka leg.
female, 34 mm
photo Panin R
Copyright © Ruslan Panin

Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860
Male+Female adult
SW -France,
Pyrenees-Atlantiques, Foret de Sare, XII.2009, Pascal Stefani leg.
female, 31 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860
Female adult
Spain,
Navarra, Sierra de Aralar, Altxueta Mt., 5.V.2006
Anichtchenko A. leg.
L= 17mm
Copyright © Alexander Anichtchenko

Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860
Male adult
Spain,
Pais Vasco, Bera de Bidasoa, Larrun 450m,
9.9.2001 Anichtchenko A. leg.
male, 29mm
Copyright © Alexander Anichtchenko

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as delacrei Tarrier, 1975)
Male+Female adult
NE Spain, Catalonia, Sant Jaume de Frontanyа, h=1000 m, V-VI.2010, Pascal Stefani leg.
female, 38 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as delacrei Tarrier, 1975)
Male adult
Spain,
Catalunya, San Jaume de Frontana,
May 2001
Copyright © Ruslan Panin

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826
Female adult
Auriac alt.
500m. (France : Aude) 25.V.200
Copyright © Ruslan Panin

Carabus (Carabus) arvensis sylvaticus Dejean, 1826
Female adult
France, Centre reg., Eure-et-Loir, Forкt de Montecot, VI.2009, Pascal Stefani leg.
female, 20 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) arcensis sylvaticus Dejean, 1826 (as sugintensis Brunier, 1922)
Male adult
NW -France, Haut-Rhin Depart., Fulleren, XI.2004, Pascal Stefani leg.
Copyright © Ruslan Panin

Carabus (Carabus) arvensis sylvaticus Dejean, 1826 (as liebmanni Lengerken, 1911 )
Female adult
France 67, Weifersweller 350m, Vosges massif
25.8.91
long. 21mm
Copyright © Alexander Anichtchenko

Carabus (Carabus) arvensis sylvaticus Dejean, 1826
Male adult
France 61, Longny-au-Perche. 30.V.1991
male, 19 mm
Copyright © Alexander Anichtchenko

Carabus (Chrysocarabus) olympiae Sella, 1855
Male+Female adult
N-Italy,
Piemonte, Bielmonte, 14-15.VI.1981, Pascal Stefani leg.
female, 34 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Chrysocarabus) olympiae Sella, 1855
Male adult
Italy, Piemonte, Sessera, 2500m 23.III.1987
Copyright © Alexander Anichtchenko

Carabus (Coptolabrodes) haeckeli Brezina & Imura, 1997
Male adult
C China, Shaanxi,
Tsinling Mts., Taibaishan Mt., h=3000m, 33:55N/107:44 E, VI.2004, V.Siniaev & his team
female, 31 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus tscherkessicus Korge, 1964
Male adult
S Russia, NW-Caucasus,
Karachay-Cherkessia,Teberda, Mukhinsky (Mukhu) Mt. pass, h=2750-2800 m , 8.VIII.2010, I.A. Solodovnikov leg.
male, 23 mm

Copyright © Ruslan Panin

Carabus (Megodontus) imperialis Fischer, 1823
Female adult
Carabus (Megodontus) imperialis
photo M. Smirnov
from www.zin.ru
Copyright © Maksim Smirnov

Carabus (Eupachys) glyptopterus Fischer, 1827
Male+Female adult
Russia, SE-Siberia,
Buryat Republic, Ivolginsky Distr., 04.VII.2010, Filippov A.E. leg.
female, 32 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Eupachys) glyptopterus Fischer, 1827
Female adult
Mongolia, 15 km N Bayanteeg, 2000 m, 17.7.2006 Anichtchenko A. leg.
Copyright © Alexander Anichtchenko

Carabus (Megodontus) schoenherri sajanus Breuning, 1927
Male+Female adult
Carabus (Megodontus) schoenherri sajanus Breuning, 1927 Russia, SW Siberia, Krasnoyarsk reg., S of Abakan, Sayanogorsk distr., Cheryomushki vill. env. 07-VII-1994 M. Danilevsky leg. male 27mm, female 31 mm.
Copyright © Safronov

Carabus (Megodontus) schoenherri sajanus Breuning, 1927
Male adult
Carabus (Megodontus) schoenherri sajanus Breuning, 1927
W Sayan Mts, Borus mt. rng., near Cheremushki vill.,
29-31.7.2000 Brinev A. leg.
male, 30mm
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Coptolabrus) formosus brockmanni (as nili Deuve, 2005 PT) Paratype
Male+Female adult
C China, Shaanxi,
Liupan Mts., Guan Shan Mt. h=800 m, , 34:50N/106:48 E, Team V.Siniaev leg.
female, 38 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus fulgens Charpentier, 1825
Male adult
NC-Spain,
Navarra Reg., Sierra de Andнa, IX.2004, Pascal Stefani leg.
male, 30 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus fulgens Charpentier, 1825 (as gerundensis Born, 1916)
Male adult
Spain, Gerona, Garriguella, XI.1990 J. Soler leg.
male, 31mm
Copyright © Alexander Anichtchenko

Carabus (Procerus) gigas gigas Creutzer, 1799
Male adult
Croatia
Copyright © Stanislav Krejcik

Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792
Female adult
NE-FRANCE,
Lorraine Reg., Moselle, 15.II.2010, Pascal Stefani leg.
female, 26 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792
Male adult
Austria,
Weissenbach stt., III.1979, Kretschmer leg.
male, 24mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792 (as aureus)
Male adult
France 28, Saint-Eliph, V.1991
female, 25mm
Copyright © Alexander Anichtchenko

Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792
Female adult
France 28, Saint-Eliph, V.1991
female, 24 mm

Copyright © Alexander Anichtchenko

Carabus (Archicarabus) nemoralis lamadridae Born, 1895 (as layrei Tarrier, 1975)
Male adult
NW-Spain, León Province, Acebo, V.2007, Pascal Stefani leg.
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii zawadzkii Kraatz, 1854
Male adult
Ukrainian Carpathians
Lviv Reg., Skole Distr., near Grebeniv vill., h=1000m 9 - 20.VII.07, Panin R. leg.
male, 31 mm
Copyright © Safronov

Carabus (Acathaicus) alexandrae Semenov, 1887
Male adult
Carabus (Acathaicus) alexandrae alexandrae Semenov, 1897 Holotype
Kwantin and Li-tschara-pu, 22.VI.1885, leg. Potanin
Deposited in Zoolog. Museum St. Petersburg.
male, long. 22 mm
photo B. Brezina
Copyright © Boleslav Brezina

Carabus (Carpatophilus) linnaei linnaei Panzer, 1812
Male adult
Carabus (Carpatophilus) linnaei linnaei Panzer, 1812
Copyright © Safronov

Carabus (Isiocarabus) strandiellus dayongensis Kleinfeld, 1996 HT Holotype
Male adult
Carabus (Isiocarabus) strandiellus dayongensis HT
Copyright © Frank Kleinfeld

Carabus (Isiocarabus) dargei dargei Deuve, 1987 (as kulingensis Holotype) Holotype
Male adult
Carabus (Isiocarabus) dargei dargei (as kulingensis Holotype)
Copyright © Frank Kleinfeld

Carabus (Isiocarabus) dargei yulingensis (as dreuxi yulingensis) Kleinfeld, 2002 HT PT Holotype Paratype
Male+Female adult
Carabus (Isiocarabus) dargei yulingensis (as dreuxi yulingensis) HT PT
Copyright © Frank Kleinfeld

Carabus (Isiocarabus) fiduciarius fiduciarius (as fiduciarius tim) Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Isiocarabus) hunanicola jinxiuensis Imura, 1995 (as dreuxi xinyingensis) HT PT Holotype Paratype
Male+Female adult
Carabus (Isiocarabus) hunanicola jinxiuensis (as dreuxi xinyingensis) HT PT
Copyright © Frank Kleinfeld

Carabus (Isiocarabus) hunanicola liangshanicus (as dreuxi liangshanicus) HT PT Kleinfeld, 1998 Holotype Paratype
Male+Female adult
Carabus (Isiocarabus) hunanicola liangshanicus (as dreuxi liangshanicus) HT PT
Copyright © Frank Kleinfeld

Carabus (Isiocarabus) kiukiangensis orphniopterus Hauser, 1920
Male adult
Carabus (Isiocarabus) kuikiangensis orphniopterus Hauser, 1920
China, Fujian, Yun Ling Shan,
Fokien, 4.6.1998
Copyright © Alexander Anichtchenko

Carabus (Ulocarabus) theanus Reitter, 1895
Male adult
Vakhskiy Mt Range,Saryhosor, Mullonomi,
29.4.1990, S.Zonshteyn leg.
Copyright © Alexander Anichtchenko

Carabus (Tribax) titan titan Zolotarev, 1913
Male+Female adult
Russia,NW Caucasus,
Krasnodar reg.,Chernogorie Mnt. Range, Agulova Balka h=950m V-VII.1999, A.S.Zamotajlov leg.
female,32mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Tribax) titan titan Zolotarev, 1913
Male adult
Carabus (Tribax) titan titan Zolotarev, 1913
Tuapse distr., Anastasievka vill., Semiglavaya mt.,
Pshekaho riv., 20.7.1995
male, 30 mm
Copyright © Alexander Anichtchenko

Carabus (Tribax) titan abadzechus Zamotajlov, 1988
Male adult
Carabus (Tribax) titan abadzechus Zamotajlov, 1988
W Caucasus, Adygea, Asish-Tau Mt.R., vall riv. Bzykha, Fagus forest, h=1450m, 14-22.08.2008, I.A.Solodovnikov, O.Beljavskyi
male, 25 mm

Copyright © Igor Solodovnikov

Carabus (Tribax) steveni steveni Menetries, 1832
Male adult
Carabus (Tribax) steveni steveni Menetries, 1832
Karachaevo-Cherkessia,
Hudes riv., near Tsarskie vorota,
V.2009 Anichtchenko A. leg.
male, 25mm
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Tribax) steveni steveni Menetries, 1832
Male adult
Carabus (Tribax) steveni steveni Menetries, 1832
C Caucasus, Kabardino-Balkaria,
Elbrusskyi distr., riv. Malka, upper reaches riv. Sultan-Bashi, h=2400-2600 m, 26, VII, 2008. Leg. I.A.Solodovnikov.
male, 20,5 mm
Copyright © Igor Solodovnikov

Carabus (Tribax) steveni steveni Menetries, 1832
Female adult
Carabus (Tribax) steveni steveni Menetries, 1832
C Caucasus, Kabardino-Balkaria,
Elbrusskyi distr., riv. Malka, upper reaches riv. Sultan-Bashi, h=2400-2600 m, 26, VII, 2008. Leg. I.A.Solodovnikov.
female 24,5 mm
Copyright © Igor Solodovnikov

Carabus (Tribax) steveni schamyli Hampe, 1852
Female adult
S Russia, N Ossetia, Alagir distr., near Nizhniy Tsey vill.,VII.1981
female, 25mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Tribax) steveni schamyli Hampe, 1852
Male adult
C Caucasus, N Ossetia,
N slope of Pastbistshnyi Mt.R., Tagardon env., h=1100m, 7-9.08.1997, leg. A.G.Koval
male 25,5 mm
Copyright © Igor Solodovnikov

Carabus (Tribax) steveni schamyli Hampe, 1852
Male adult
C Caucasus, N Ossetia,
N slope of Pastbistshnyi Mt.R., Tagardon env., h=1100m, 7-9.08.1997, leg. A.G.Koval
female 27 mm
Copyright © Igor Solodovnikov

Carabus (Tribax) steveni schamyli Hampe, 1852
Male adult
N Osetia, B. Balkary, 1700 m, 2.9.85
Abdurahmanov leg.
male, 24 mm
Copyright © Alexander Anichtchenko

Carabus (Tribax) steveni satanas Semenov, 1896
Male adult
Carabus (Tribax) steveni satanas Semenov, 1896
S Russia, C-Caucasus:
Balkaria, Valley of riv. Cherek Balkarskyi, h=1750-1800m 10.VI-20.VII.1992 Leg. Zamotajlov A.
male, 23 mm
photo Panin R., det. Solodovnikov I.A
Copyright © Ruslan Panin

Carabus (Tribax) steveni satanas Semenov, 1896
Male+Female adult
Carabus (Tribax) steveni satanas Semenov, 1896
Russia, C.Caucasus, Kabardino-Balkaria,
valley of River Cherek Balkarsky, h=1800 m
10.VI-20.VII.1992
A. Zamotajlov leg.
Copyright © Safronov

Carabus (Tribax) puschkini puschkini Adams, 1817
Male adult
Armenia, Shirak Province, Ashotsk, 18.VI.2001, P.V.Kazaryan leg. male 25mm
Copyright © Ruslan Panin

Carabus (Tribax) puschkini ishikawaianus Breuning & Ruspoli, 1970
Male adult

Copyright © Alexander Anichtchenko

Carabus (Tribax) kasbekianus veselyi Gottwald, 1980
Female adult
Carabus (Tribax) kasbekianus veselyi Gottwald, 1980
Caucasus, Georgia, Egrissky Mt.R., riverheads Tekhuri river, h=2400, A.G.Koval leg.; 04.07.1989; 25 mm.; photo A. Menstschicov
Copyright © Anton Menstschikov

Carabus (Tribax) kasbekianus onerosus Belousov & Zamotajlov, 1993 Paratype
Male+Female adult
Carabus (Tribax) kasbekianus onerosus Belousov & Zamotajlov, 1993 Paratypes
Caucasus, W Georgia, Svanetia, right bank o Magana riv.,
800m, 12.V.-4.VII.1991 Zamotajlov & Miroshnikov leg.
male 27mm, female 33mm
photo & collection Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Tribax) edmundi Semenov, 1896
Male adult
Carabus (Tribax) edmundi Semenov, 1896
Copyright © Boleslav Brezina

Carabus (Tribax) circassicus teberdensis Zolotarev, 1913
Male+Female adult
W-Caucasus: Karacaj-Cherkessija,
Abishira–Akhuba Mnt. Range h=2000-2650m, Kyafar-Agur Riv., 14–17.VII.2009, Fominykh D. leg.
female, 25mm
Copyright © Ruslan Panin

Carabus (Tribax) circassicus kubaniensis Semenov, 1896
Male adult
N Caucasus, Krasnodar reg., 30 km S Maykop
august 1993
male, 26 mm
Copyright © Alexander Anichtchenko

Carabus (Tribax) circassicus kubaniensis Semenov, 1896
Male+Female adult
Russia, NW Caucasus,
Krasnodar reg., Chernogorie mts., 24.V-29.VII.1999. A.S. Zamotajlov leg.
female 29mm/male 25mm
Copyright © Ruslan Panin

Carabus (Tribax) circassicus circassicus Ganglbauer, 1886
Male+Female adult
Abkhasia,
W.slope of Gagrinsky Mt., Krepostnaya River source, h=930-980m., A.S.Zamotajlov leg. 27.VII.2000
female, 29 mm/male, 26mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Tribax) circassicus abagonensis Starck, 1894
Male+Female adult
S Russia, NW Caucasus,
Adygea Rep., Mt.Abago, 26.VI.1999
L=24mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Tribax) biebersteini ulgeni Gottwald, 1982
Male adult
S Russia, Karacaj-Cherkessija,
near Kart-Djurt vill., upper Kylan riv. h=1300m, 26-27.V.2009, Fominykh D. leg.
male, 28mm
Copyright © Ruslan Panin

Carabus (Tribax) biebersteini otcharensis Kurnakov, 1970
Male adult
Caucasus, W Georgia, Svanetia, upper reaches of riv. Shoudidi, 1700-2100m, 20-21.V.1986 leg. A. Zamotajlov
Copyright © Evgeniy Chibilov

Carabus (Tribax) biebersteini ingusch Zolotarev, 1913
Male+Female adult
Carabus (Tribax) biebersteini ingusch Zolotarev, 1913
Russia, C.Caucasus, N.Ossetia,
valley of River Khazniaush
10.VI-31.VII.1993
A. Zamotajlov leg.
Copyright © Safronov

Carabus (Tribax) biebersteini constantinowi Starck, 1894
Male adult
Carabus (Tribax) biebersteini constantinowi Starck, 1894
NW-Caucasus, S Russia,
Karacaj-Cherkessija, Nizhniy Arkhyz, 21-30.V.2001, Tikhonov V. leg.
male,30mm
Copyright © Ruslan Panin

Carabus (Tribax) apschuanus apschuanus Rost, 1893
Male+Female adult
Carabus (Tribax) apschuanus apschuanus Rost, 1893
Russia, Caucasus, Krasnodar reg.,
h=1200 m, 29.VII.2000
A. Zamotajlov leg.
Copyright © Safronov

Carabus (Tribax) agnatus agnatus Ganglbauer, 1889
Male adult
NW-Caucasus,
Krasnodar reg., Mostovskoj distr., Skirda Mts. N. slope Yatyrgvart Mt., h=2300-2500m. 09-10.VII.2008, D.Fominyh leg.
male, 23mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus sibiricus Fischer, 1822
Male adult

Copyright © Kirill Makarov

Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886
Male+Female adult
W Ukraine, Ternopol reg., Pidvolochisk distr., Supranivka vill. 3.VII.1908. Leg. Stobetskii. S. Photo R. Panin
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886
Male adult
W Ukraine,
Ternopil reg., Pidvolohcisk distr., Supranivka vill.
male, 26mm
The end of XIX the beginning of XX century
photo Panin R.
coll. SMNH Ukraine, Lviv
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus obliteratus Fischer, 1827
Male adult
Altai mts., Kalbinskii mt.rng.
Ognevka vill. VII.1994, Klimenko A. leg.
male, 28 mm
Copyright © Alexander Anichtchenko

Carabus (Trachycarabus) sibiricus obliteratus Fischer, 1827
Male adult
NE Kazakhstan,
S Altai Mts., Ulbinsky range h~500m.,Turgusun vill., 20.VI.2001
male 25mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus kolosovi Zinoviev, 1997 loc.typ.
Female adult
S. Ural, Bashkiria, near Mesjagutovo vill., 09.07.2001 A. Menstschicov leg.
photo A. Menstschicov
Copyright © Anton Menstschikov

Carabus (Trachycarabus) sibiricus kolosovi Zinoviev, 1997
Female adult
Russia, North-East Bashkortostan. Ai river, near Sikiyaz-Tamak vill.
12.07.1997, Zinoviev E.leg
length: 21,8 mm
Copyright © Evgeniy Zinoviev

Carabus (Trachycarabus) sibiricus haeres (as fossulatus Dejean, 1826)
Female adult
Carabus (Trachycarabus) sibiricus fossulatus Dejean, 1826 NE Ukraine, Kharkov reg.,1974 female, 27mm.
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus haeres (as fossulatus Dejean, 1826)
Male+Female adult
E.UKRAINE, Kharkov reg., Zmiev distr.,in steppe,10-20.V I.2007, D.Kovalchuk leg. female, 26mm/male, 24mm. photo/det. Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus errans Fischer, 1823
Female adult
Carabus (Trachycarabus) sibiricus errans Fischer, 1823
S.Russia, Rostov-Don reg., Avilo-Uspenskaya env.
V.2000
Copyright © Safronov

Carabus (Trachycarabus) sibiricus errans Fischer, 1823
Male adult
SW-Russia,
Rostov reg. env. stanitsa Razdorskaya, A. Ponomarev leg. 9-15.V.2001
male 25mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus errans Fischer, 1823
Male+Female adult
female:SE Ukraine,
Donetsk, 25.VII.2009, A.I.Gubin leg. L=30mm.
male:E Ukraine, Lugansk reg., near Slavyanoserbk, 5-15.VIII.2009, S.Konovalov leg. L=26mm.
photo/det.Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus bosphoranus Fischer, 1823
Female adult
Carabus (Trachycarabus) sibiricus bosphoranus Fischer, 1823
S Ukraine,
Krym Peninsula, Nikitskaya Jaila, S.Mosyanin leg. 10.VIII.2000
female 23mm.
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus bosphoranus Fischer, 1823
Male adult
Carabus (Trachycarabus) sibiricus bosphoranus Fisch. 1823 S Ukraine, Krym Peninsula, Dolgorukovskaya jaila, 25.VIII.1991 I.A.Solodovnikov leg. male, 23mm 
Copyright © Ruslan Panin

Carabus (Trachycarabus) scabriusculus scabriusculus (as erythropus Fischer, 1822)
Male+Female adult
W.Ukraine,
NW Podolian, Lviv Reg., Zolochiv distr., near Luky vill., steppe sites, 7.V-8.VI. 2010
leg. Panin R.
female, 22mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) scabriusculus scabriusculus (as erythropus Fischer, 1822)
Male adult
W-UKRAINE
NW Podolian, Lviv Reg., Zolochiv Distr., near Luka vill., in steppe. Panin R. leg. 1.V - 16.VI.07
Copyright © Safronov

Carabus (Trachycarabus) scabriusculus inapertus Motschulsky, 1850 (loc.typ)
Male+Female adult
NE UKAINE,
Kharkov reg., Zmiev distr., in steppe, 15.VI-14.VII.2009, Kovalchuk D. leg.
female, 20mm
photo/det. Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus planus Gehin, 1885
Male+Female adult
S Russia, NW Caucasus, Karacaj-Cherkessija, near Kart-Djurt vill., upper Kylan riv. h=1300m, 26-27.V.2009 , Fominykh D. leg. L=24mm photo/det. Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus planus Gehin, 1885
Male adult
NW Caucasus, Rostov reg., Avilo-Uspenskaya vill. 22.5.2000 Arzanov Yu. leg. male, 25 mm
Copyright © Alexander Anichtchenko

Carabus (Trachycarabus) planus planus Gehin, 1885
Male adult
S Russia, Krasnodar reg., Anapa distr., B. Utrish vill., env., 11-20.VIII.2008, Khatchikov E. leg. male, 26mm photo Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus campicola Reitter, 1896
Male adult
Krym Peninsula, near Sudak. Leg.Vashchenko S. 1-5.V.2004 27 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus campicola Reitter, 1896
Male+Female adult
S Ukraine, Krym Peninsula, near Perevalnoe vill., in steppe, 15.IV.2009, Shaporinskiy V. leg. female, 28 mm photo/det.Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) mandibularis mandibularis Fischer, 1827
Male adult
NE Kazakhstan, S Altai Mt., Narymskii mt. rng.
Bolshenarymskoe vill. 26.7.2001, 1800m Klimenko leg.
male, 21 mm
Copyright © Alexander Anichtchenko

Carabus (Trachycarabus) mandibularis buchtarmensis Kryzhanovskij, 1953
Male adult
Kazakhstan, Kalbinsky mt. rng., 600m ,
Ognevka vill., 23.6.98 Klimenko A. leg.
male, 23 mm
Copyright © Alexander Anichtchenko

Carabus (Trachycarabus) mandibularis buchtarmensis Kryzhanovskij, 1953
Male adult
NE-Kazakhstan,
S-Altays, Kalbinky Mt. h=500m., near Ognevka vill., 20.VI.1998.A. Klimenko leg.
male, 23mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
Male adult
Burjatia, Dzhida distr., near Beloozersk vill. 4-9.VII.1997 Anichtchenko A. leg.
male, 23 mm
Copyright © Alexander Anichtchenko

Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
Male adult
Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
China, Beijing, Mentougou, 2005;
Photo & det. by liuye

Copyright © Ye Liu

Carabus (Trachycarabus) besseri Fischer, 1822
Male adult

Copyright © Safronov

Carabus (Trachycarabus) besseri Fischer, 1822
Male+Female adult
W Ukraine.NW Podolian,
Lviv reg., Zolochiv distr., Lysa Gora near Stinka vill., steppe sites, 25.IV-9.V.2009 Panin R. leg
female, 30mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) scabripennis scabripennis Chaudoir, 1850
Male adult
Carabus (Tomocarabus) scabripennis scabripennis Chaudoir, 1850
C. Armenia,
Kotayk reg., Yeghvard, Arailer Mt., 16.VI.1999, P.V.Kazaryan leg.
male, 20mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Tomocarabus) scabripennis marthaiellus Rapuzzi & Kleinfeld, 2000 Holotype Paratype
Male+Female adult
Carabus (Tomocarabus) scabripennis marthaiellus Rapuzzi & Kleinfeld, 2000
Copyright © Frank Kleinfeld

Carabus (Tomocarabus) marginalis Fabricius, 1794
Male adult
male, 26 mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) marginalis Fabricius, 1794
Male+Female adult
E Ukraine,
Kharkov reg.,near Vasishchevo vill.10-20.VI.2006 D.Kovalchuk leg.
female 24mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) decolor Fischer, 1823
Female adult
Carabus (Tomocarabus) decolor Fischer, 1823
Copyright © Alexander Anichtchenko

Carabus (Tomocarabus) decolor Fischer, 1823
Male adult
S Russia, NW-Caucasus,
Karacaj-Cherkessija,Skalisty Mnt. Range, Site Banditskoe h=1400m, 2-3.V.2009,
Fominykh D., Bondarenko A.S. leg.
male,17mm
Copyright © Ruslan Panin

Calosoma (Calosoma) cyanescens Motschulsky, 1859
Male adult
China,
Jilin prov, P'an-shih
42° 56' 32" N
126° 03' 22" E
20.06.2009
Local people leg.

Copyright © Anton Menstschikov

Calosoma (Caminara) reitteri Roeschke, 1897
Male adult
W Kazakstan, Ustjurtskij res., Kordon Kenderli; 01.05.2009, Gorbunov P.J. leg.,
26 mm.; photo A. Menstschicov
Copyright © Anton Menstschikov

Calosoma (Caminara) reitteri Roeschke, 1897
Male adult
Turkmenia, Badkhyz,
Akar-Cheshme 9.4.1985
23 mm
Copyright © Alexander Anichtchenko

Calosoma (Caminara) olivieri Dejean, 1831
Male adult
З Казахстан, среднее течение р. Эмба, окр. п. Миялы, П.Ю.Горбунов leg. 02.07.2009; photo A. Menstschicov
Copyright © Anton Menstschikov

Carabus (Morphocarabus) wulffiusi wulffiusi Morawitz, 1862
Male+Female adult
Russia, Far East,
Primorskiy Kray, near Kamenushka vill., 2-3.VIII.1992, K.E.Dovgaylo leg.
female, 24mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) wulffiusi wulffiusi Morawitz, 1862
Female adult

Copyright © Safronov

Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Male adult
Serbia, Majdanpek, Ceremosja 15.6.2001 Z.Zlatic leg. male, 32 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Male+Female adult
NE-Serbia, Krs Mts., Majdanpek env.,h=300m.20-31.V.2006, Pr.Sima leg. female, 32mm/male, 30mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) venustus venustus Morawitz, 1862
Male adult
SE Primorskii reg., Lazovskii Nat.Res., 12 km S Staraya Kamenka vill., near Amerika kor. 43°15N, 134°1E, 19.VII.04 leg. M.& L. Smirnovy photo from www.zin.ru
Copyright © Maksim Smirnov

Carabus (Morphocarabus) tarbagataicus pietrorattii Deuve, 1991(as pseudogebleri Deuve, 1994)
Male adult
Carabus (Morphocarabus) tarbagataicus pietrorattii Deuve, 1991(as pseudogebleri Deuve, 1994)
S Tarbagatai mt.rng., Alexeevka vill.,
600m, 11.5.2001 Klimenko A. leg.
male, 30 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) tarbagataicus pietrorattii Deuve, 1991
Female adult

Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997 HT Holotype
Male adult
Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997 HT
Rusia, SE Tuva, Sangilen mt.rng., Kargy riv., 50°31'N 97°01'E,
1800 m, 29.07.1996, D. Obydov leg.

Copyright © Dmitri Obydov

Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997
Female adult
Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997
Tuva, Akademika Obrucheva mt.rng., 15 km N Cherbi vill.
ur. Chailag 7.VII.1999 A. & N. Menschikovy
female, 22 mm
det. Menschikov A.
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997
Male adult
Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997
Tuva, Akademika Obrucheva mt.rng., 15 km N Cherbi vill.
ur. Chailag 7.VII.1999 A. & N. Menschikovy
male, 21 mm
det. Menschikov A.
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) spasskianus tungus Obydov, 1999 HT Holotype
Male adult
Carabus (Morphocarabus) spasskianus tungus Obydov, 1999 HT
Rusia, N Krasnoyarsk reg., Ilimpeya riv and Dovogpokan riv.
VII.1974, Lunin leg.
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) spasskianus spasskianus Breuning, 1932
Male adult
Siberia, Lake Baikal, Listvianka vill. 25.8.90
Anichtchenko A. leg.
male, 23 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) spasskianus spasskianus Breuning, 1932
Male+Female adult
SE Siberia, Russia
Irkutsk reg., near Pivovarikha vill., 7.V.07, D. Fominyh leg.
male, 24mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus shoriensis Obydov, 1999 HT Holotype
Male adult
Gornaya Shoria, 8.VIII.1978, S. Nikireev leg.

Copyright © Dmitri Obydov

Carabus (Morphocarabus) spasskianus saragaschensis Obydov, 1999
Male adult
W Sayan Mts., Batenevskii Mt.rng., 12 km E Pervomaiskoe vill.,
Enisey riv., mountain steppe, 400m, 5.VII.2000 A. & N. Menschikovy leg.
female, 26mm

Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) spasskianus saragaschensis Obydov, 1999 Holotype
Male adult
Rusia, Khakassia, Saragash, 400 m, 6.VII.1994, D. Obydov leg.

Copyright © Dmitri Obydov

Carabus (Morphocarabus) spasskianus putus Motschulsky, 1846
Male adult
Carabus (Morphocarabus) spasskianus putus Motschulsky, 1846
SW Siberia, Altay Mts., Shebalino distr., Kumalyr env.
02.07.2003, A.Safronov leg.
Copyright © Safronov

Carabus (Morphocarabus) spasskianus microputus O.Berlov & Shilenkov, 1996
Female adult
Tuva, 25 km E Ak-Dovurak 23.VI.2006
A. & N. Menschikovy leg.
female, 24 mm
det. Menschikov A.

Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) spasskianus microputus O.Berlov & Shilenkov, 1996
Male adult
Tuva, 25 km E Ak-Dovurak 23.VI.2006
A. & N. Menschikovy leg
female, 24 mm
det. Menschikov A.

Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) spasskianus microputus O.Berlov & Shilenkov, 1996
Male adult
Rusia, SW Tuva, W Tannu-Ola mt.rng., bass. of Barlyk riv.,
50°29'N 90°44'E, 30.VII.2000, D. Obydov leg.
Copyright © Dmitri Obydov

Carabus (Morphocarabus) spasskianus korostelevi Obydov, 2004loc.typ.
Male adult
E Kazahstan, Ivanovskii Mts., Prohodnoy Belok, 03.06.2000, A. Ivanov leg.

Copyright © Anton Menstschikov

Carabus (Morphocarabus) spasskianus geblerianus Obydov, 1999 Holotype
Male adult
Rusia, Barnaul, 27.5.1965
Copyright © Dmitri Obydov

Carabus (Morphocarabus) spasskianus cracens Obydov, 1999
Male adult
Carabus (Morphocarabus) spasskianus cracens Obydov, 1999
NE Kazakhstan, S Altay Mts., Kalbinskii range,
Ognevka vill., 600m, 18.6.1999 A. Klimenko leg.
male, 22 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) spasskianus cracens Obydov, 1999 Paratype
Male adult
Museum of Institute of Systematic Biology (Daugavpils, Latvia)
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) spasskianus cracens Obydov, 1999
Female adult
Carabus (Morphocarabus) spasskianus cracens Obydov, 1999
NE-Kazakhstan:
S-Altay, Kalbinsky Mt., Ognevka vill., h~600m 18.VI.1999 A. Klimenko leg.
female, 24mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus cracens Obydov, 1999 Holotype
Male adult
Carabus (Morphocarabus) spasskianus cracens HT
E Kazakhstan, Kolbinskii Mt.rng., 600m, near Ognevka vill.,
13-18.07.1999, D. Obydov leg
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Cyclocarabus) belousovi Kabak, 1992
Female adult
Uzbekistan, Karzhantau Mt., Aktash vill.
1500 m, 12.4.1999
female, 23 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
Female adult
Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
Moravia mer.,Vranov, 26-29.06.1979, Dr. Resl leg.
photo A. Menstschicov
Copyright © Anton Menstschikov

Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
Male adult
Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
SE Moravia mer.,Vranov, 26-29.06.1979, Dr. Resl leg.
photo A. Menstschicov
Copyright © Anton Menstschikov

Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
Male adult
Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
S.Bohemia,
Hluboka nad Vltavou, 24.VI-18.VII.1990, S. Budka leg.
male, 27mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799 (as scheidleri floriani)
Male adult
Carabus (Morphocarabus) scheidleri scheidleri (as scheidleri floriani)
A.: Steiermark, Rein Umg. VIII.1983 Kenyery
female, 32 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) excellens excellens Fabricius, 1798
Male adult
NW Podolian
Copyright © Safronov

Carabus (Morphocarabus) excellens excellens Fabricius, 1798
Male adult
Moldova, 2003
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens (as lomnitzkii Reitter, 1896 )
Male adult
W-UKRAINE, NW Podolian, Lviv Reg., Zolochiv Distr., W Ukraine, near Gologirki vill., steppe sites, 19.V.-7.VI.2007, Panin R. leg. 25 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens Fabricius, 1798 (ab.erythromerus Dejean, 1826)
Male adult
Moldova, Anenii Noi distr., near Garbovets vill. 20.VI.08 Bodnar' I.I.leg. male, 26mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens (as lomnitzkii Reitter, 1896)
Male adult
W Ukraine, NW Podolian, Ternopil reg., Pidvilochisk distr., near Rosokhuvatets & Kamyanki vill., steppe sites, 24.V-15.VI.2008, Panin R. leg.
male, 24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) helleri Ganglbauer, 1893
Male adult
Moravia Mer., Bile Karpaty 28.5.1993 Budka leg. male, 28 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) helleri Ganglbauer, 1893
Male+Female adult
SE Moravia, Strani, V.2002, M.Snizek leg. male bronze colour-26mm male blue colour-28mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi incompsus (as mendax Csiki,1906)
Male+Female adult
W.Ukraine,
Transcarpathian Reg., Rakhov Distr., Chornogora Mt Range, h=1900m. alpine zone, Panin R. leg/det. 28.VI-9.VIII.2008
female, 32 mm
photo endophallus Solodovnikov I.A
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi comptus Dejean, 1831
Male adult
Carabus (Morphocarabus) rothi comptus Dejean, 1831
Carpathians
male, 23mm
coll.SMNH Ukraine ,Lvi
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi alutensis Savulescu, 1972
Male adult
C Romania,
Olt River: Rimnica Vilcea VII.1971, Savulescu N. leg.

Copyright © Igor Solodovnikov

Carabus (Eucarabus) obsoletus carpathicus Palliardi, 1825
Male+Female adult
female, 25 mm
SW-Romania, Banat, Resita - Valiug, 3-8.VI.2007, Krause R. leg.
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus tibiscinus Csiki, 1905
Male+Female adult
female,28 mm
SE Hungary, Csongrбd, Makу env., 1-8.VIII.2004, Krause R., Slovak, Pittner leg
Copyright © Ruslan Panin

Carabus (Tribax) apschuanus pseudoplatessa Gottwald, 1982
Male+Female adult
NW-Caucasus, Karachay-Cherkessia, Teberda: Dzhamagat gorge, the left bank of the riv. Dzhamagat, 10.VII-28.VIII.2010 S.Pavluchuk leg.
female, 27 mm
Copyright © Ruslan Panin

Carabus (Pachystus) hungaricus mingens Quensel, 1806
Male adult
male, 31 mm
SW-Russia: Stavropol Krai, near Stavropol city , Russkiy Les, 11.VI-28.IX.2010, Pavluchuk S. leg.
Copyright © Ruslan Panin

Carabus (Archicarabus) montivagus montivagus Palliardi, 1825
Male+Female adult
SW-Romania: Banat, Caras-Severin Reg., Sasca Montana – Stinapari, 21.V.-5.VI.2010, Krause Rene leg.
female, 26 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) montivagus blandus Frivaldsky, 1865
Female adult
Carabus (Archicarabus) montivagus blandus
Slovacchia, Plastovce,
16.7.1995 Budka leg.
female, 21 mm
Copyright © Alexander Anichtchenko

Carabus (Archicarabus) montivagus blandus Frivaldsky, 1865
Female adult
female, 23 mm
S-Slovakia, Region de Nitra, Plastovce viil., 1.V.2009, Rudolf Kmeco leg.
Copyright © Ruslan Panin

Carabus (Procechenochilus) kokujewi Semenov, 1898
Female adult
RUSSIA, Karachay-Cherkessia, NW Caucasus Mts., Daut Mt. Range, Iskrovtsev - W.Ullu-Kel Mt. pass circus, h=2800-2600m
Copyright © Alexander Anichtchenko

Carabus (Procechenochilus) ? kokujewi Semenov, 1898
Female adult
Russia, NW Caucasus, Karachay-Cherkessia, Karachaevsk distr., near Uchkula vill., valley Uchkulanichi riv. h=2800-2900m, 25-28.VII.2010, Fominykh D. leg.
female, 23 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) starckianus ganglbaueri Reitter, 1888
Male adult
Russia, NW Caucasus, Krasnodar reg., Adygeia distr., Guzeripl env. 20.5.2003
male, 26 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) starckianus ganglbaueri Reitter, 1888
Female adult
NW Caucasus, S Russia, Adygea Rep., Confluence of the river Belaya & Kisha h~700m, 24.V-7.VIII.2010, D.Fominyh leg.
female,32 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus andrzejuscii (as wolffii Dejean, 1826)
Male+Female adult
SW-Romania: Banat, Moldova Noua, Ravenska viil., 21.V.-5.VI.2010, Krause Rene leg. female, 32 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus andrzejuscii (as mehelyi Ganglbauer, 1896 loc. typ.)
Male+Female adult
N-Romania, Rodna Mts., Pasul Setref - Sacel 10km E, h=810 m, 8-19.VI.2009, Krause Rene leg. female, 31mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus andrzejuscii Dejean, 1826 (as pygmaeus Petri, 1912)
Male adult
W Ukraine, NE Carpathians, Transcarpathian Reg., Rakhov Distr., Chornogora Mt Range h=2030m, alpine zone, 13-24.VI.2010 Panin R. leg./det. male, 25 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) regalis regalis Fischer, 1822
Male adult
Carabus (Morphocarabus) regalis regalis Fischer, 1822
S. Ural, Bashkiria, near Mesjagutovo vill., 09.07.2001 A. Menstschicov leg.
photo A. Menstschicov
Copyright © Anton Menstschikov

Carabus (Morphocarabus) regalis jenissoni Dejean, 1831
Male adult
Carabus (Morphocarabus) regalis jenissoni Dejean, 1831
Rusia, Tuva, Todzh, Azas lake, 990 m,
52°24'N 96°33'E, 8.VII.1998, D. Obydov leg.

Copyright © Dmitri Obydov

Carabus (Morphocarabus) regalis jenissoni Dejean, 1831
Female adult
Carabus (Morphocarabus) regalis jenissoni Dejean, 1831
SW Siberia, Altays Mts, Choya distr., Sukhoy Karasuk env.
23.07.2004, A.Safronov leg.
Copyright © Safronov

Carabus (Morphocarabus) regalis eltoni Obydov, 2006 Holotype
Male adult
Carabus (Morphocarabus) regalis eltoni Obydov, 2006 HT
Rusia, Tver distr., near Ilyinskoe vill.,
29-30.VII.2006, D. Obydov leg.
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) odoratus septentrionalis Breuning 1932
Female adult
N. Russia, arctic Urals, Paypudynya Riv., Pompeliz Mt.h=500-700 m.,10-15.07.2002
S. Andreev, A. Romanenko leg.
Copyright © Safronov

Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844: 100
Male adult
Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844 Buriatia, W Hamar-Daban Mts., up riv. M.Zangisan, Baishint Ula Mt. 2300m 23.5.1999 Anichtchenko A. leg. male, 18 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844: 100 Lectotype
Male adult
Carabus (Morphocarabus) odoratus odoratus LT Lectotype Hamar-Gaban Mts. photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844 (as melleus Lapouge, 1909)
Male+Female adult
Russia, S Siberia, East Sayans, Mondy circ., h~2300m., 5-8.VI.2001, R. Mishustin leg. 20mm.
Copyright © Ruslan Panin

Carabus (Semnocarabus) rustemi Kabak, 2009 Paratype
Male adult
Paratype
Copyright © Ilya Kabak

Carabus (Eotribax) valikhanovi bagua Kabak, 2009 Holotype
Female adult
China, Xinjiang, N slope of Narat Mt. R., SSW of Tekes, Kokterek basin, Buryl Riv., 3145 m, 28.07.2009,
42°48'06»N / 81°41'20»E, I. Kabak leg. (ZIN)
Copyright © Ilya Kabak

Carabus (Alipaster) thianshanskii shatyensis Deuve et Tian, 2004 Paratype
Male adult

Copyright © Ilya Kabak

Carabus (Alipaster) thianshanskii sarkhobensis Kabak, 2009 Holotype
Female adult
CH, Xinjiang, N sl. of Khalyktau MR., N Kongurbulak R., H=3600 m, 42°35'50»N / 81°02'30»E, 31.07.2003, I. Kabak leg
(ZIN)
Copyright © Ilya Kabak

Carabus (Cechenotribax) petri alipastroides Kabak, 2009 Holotype
Male adult
E spur of Terskei Alatau Mt. R., 10 km S of Saryzhas Village, 2800-3000 m, 5.VII.1987, I. Kabak: 42°51’00”N / 79°37’00”E – 42°50’30”N / 79°39’00”E)
(ZIN)
Copyright © Ilya Kabak

Carabus (Tomocarabus) convexus s.str. (as pyrenaicus Born, 1907)
Male adult
Pirineos, Canfranc, 1300m, Barranco de Izas
15.6.2005 Anichtchenko A. leg.
male, 16 mm
Copyright © Alexander Anichtchenko

Carabus (Tomocarabus) convexus s.str. (as paganetti Born, 1905)
Male adult
Lazio, Roma, Bosco di Nettuno
22.5.1986 C. Iacovone leg.
male, 20 mm
Copyright © Alexander Anichtchenko

Carabus (Tomocarabus) convexus convexus Fabricius, 1775
Male adult
W.Ukraine,
Transcarpathian region,Vinogradov distr., near Malaya Kopanya vill., Chorna Gora Mt., h=180m. 7-18.VII.08. Panin R. leg.
male 18 mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) convexus s.str. (ab. niger)
Male adult
W Ukraine, NW Podolia, Lviv Region, Zolochiv distr., near Gologirky vill., in steppe 1-19.V.2007 Panin R. leg.
16mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) convexus erzeliki Schweiger, 1962
Male adult
Turkey, Sivas 1560 m. 19.7.1991
male, 19 mm
Copyright © Alexander Anichtchenko

Carabus (Tomocarabus) bessarabicus bessarabicus Fischer, 1823
Male+Female adult
UKRAINE,
Kherson Reg., Golaya Pristan distr. near Rybalche vill. 6.IV.2000
long (female) 20mm
Copyright © Ruslan Panin

Carabus (Tachypus) cristoforii Spence, 1823 (as nicolasi Reitter, 1888)
Male adult
S-France, Hautes-Pyrenees, Pic de Nere h=2200m,15.VII.1999,
male, 14mm
Copyright © Ruslan Panin

Carabus (Oreocarabus) sui Imura & Zhou, 1998
Female adult
China, Beijing, Yunmengshan,2005, Liu ye coll.;
35mm
Copyright © Ye Liu

Carabus (Teratocarabus) azrael azrael Semenov & Znojko, 1932
Female adult

Copyright © Alexander Anichtchenko

Carabus (Teratocarabus) azrael mizunumaianus Imura, 1991
Male adult
China, Liaoning, Benxi, Gongchangung
20.VI.2004 Hong
Copyright © Boleslav Brezina

Carabus (Tanaocarabus) sylvosus Say, 1825
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Carabus (Tanaocarabus) sylvosus Say, 1825
Female adult
female, 26mm
Copyright © Boleslav Brezina

Carabus (Tanaocarabus) sylvosus Say, 1825
Male adult
male, 25mm
Copyright © Boleslav Brezina

Carabus (Tanaocarabus) finitimus Haldeman, 1852
Female adult

Copyright © Boleslav Brezina

Carabus (Tanaocarabus) hendrichsi Bolivar y Pieltain, Rotger & Coronado, 1967
Male adult
male, 23mm
Copyright © Boleslav Brezina

Carabus (Tanaocarabus) forreri Bates, 1882
Male adult
Mexico, Durango, Sierra Madre Occ.,
Buenos Aires w., 2700 m, La Ciudad, 14.6.90 Heinz leg.
Copyright © Alexander Anichtchenko

Carabus (Tanaocarabus) forreri forreri Bates, 1882
Female adult
female, 23mm
Copyright © Boleslav Brezina

Carabus (Tanaocarabus) forreri forreri Bates, 1882
Male adult
male, 22mm
Copyright © Boleslav Brezina

Carabus (Tachypus) cancellatus fusus Palliardi, 1825
Male+Female adult
France, Riveres Charente, leg. Laforgue VI.1996
female, 24mm
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus s.str. (as sulinensis Born, 1902)
Male+Female adult
S UKRAINE,
Krym Peninsula, Staryi Krym, 17.V.2002
L= 26 mm.
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus s.str. (as conspersus Lapouge, 1902)
Male+Female adult
E Byelorussia,
Mogilev Reg. Bykhov distr., near Voronino vill., 20.V-3.VI.2007, Komyagunova N. leg.
female, 26mm
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus s.str. (as tuberculatus Dejean, 1826)
Male adult
W Ukraine,NW Podolian,
Ternopil reg., Pidvolochisk distr., forest Bogdanivskiy, near Romanove Selo, 7-24.V.08, R.Yu.Panin & I.A.Solodovnikov leg.
male, 23mm
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus s.str. (as scythicus Schaum, 1857)
Male adult
Ukrainian Carpathians,
Transcarpathian reg., Rakhov Distr., Chornogora Mnt.Range h=1900m. Panin R. leg. 28.VI-9.VII.08
male, 24mm.
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus s.str. (as scythicus Schaum, 1857)
Female adult
Ukrainian Carpathians,
Transcarpathian reg., Rakhov Distr., Chornohora range h=1850m, 29.VI-28.VII.2007 Panin R. leg.
female, 27 mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) odoratus martjanovianus Obydov 1999 HT Holotype
Male adult
Carabus (Morphocarabus) odoratus martjanovianus Obydov 1999
Rusia, Sayan Mts., near Sayanogorsk, 7.VII.1993
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) odoratus magadanicus Obydov 1999 PT Paratype
Male adult
Carabus (Morphocarabus) odoratus magadanicus Obydov 1999 PT
Rusia, Far East, E of Magadan, W Nyuklia vill.
VII.1997, Yu. Marusik leg.
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) odoratus krugeri Obydov 1999 PT Paratype
Female adult
Carabus (Morphocarabus) odoratus krugeri Obydov 1999 PT
Rusia, Tuva, Akademika Obrucheva Mt.rng., 2100, 51°56'N 095°32'E,
Koptu pass, tundra 2.VII.1998, D. Obydov leg.
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) odoratus kamtchatensis Breuning 1942
Male adult
Carabus (Morphocarabus) odoratus kamtchatensis Breuning 1942
Rusia, Kamchatka peninsula, VII.21
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) odoratus dohrni Gebler, 1847
Male adult
Carabus (Morphocarabus) odoratus dohrni Gebler, 1847
Rusia, Kuznetskii Alatau, Taizhe-Su riv., 53°55'N 89°10'E,
1250-1650 m, 25.VI-30.VII.2002, V. Murzin and V. Sinyaev leg.
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) odoratus dabanensis Shilenkov 1996 loc.typ.
Male+Female adult
Carabus (Morphocarabus) odoratus dabanensis Shilenkov 1996 loc.typ.
Buryatia, Hamar-Daban mt. rng., Snezhnaya riv.,
4.6.2007 Anichtchenko A. leg.
male 17 mm, female 19 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) odoratus czernyscheviellus Obydov, 2002 PT Paratype
Female adult
Carabus (Morphocarabus) odoratus czernyscheviellus PT
Rusia, SE Transbaikalia, Chita reg., Nerchinsko-Kugenskii Mt.rng.,
near Vershino-Darasunskii vill., Bogatui Mt. 1221 m.,
15.VII.2002, D. Obydov leg.
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) odoratus bargusinicus Shilenkov, 1996
Male+Female adult

 Carabus (Morphocarabus) odoratus bargusinicus Shilenkov, 1996
 Russia, N Burajatia,
 Barguzinskiy Mts., the upper reaches of M.Chiverkuy riv., h=2000m, Vashchenko S. leg.
 female, 21mm/male,19mm
 photo Panin R.

Copyright © Ruslan Panin

Carabus (Morphocarabus) odoratus antropovi Shilenkov 1996
Male+Female adult
Russia, S Tuva, Khorumnug -Tayga mts. Range, h=1600-2300m, Buren riv. valley, 4-26.VI.2000 Vashchenko S & A. leg female, 20mm/male,18mm photo Panin R.
Copyright © Ruslan Panin

Carabus (Morphocarabus) odoratus ?antropovi Shilenkov 1996
Male adult
S Tuva, Horumnut-Taiga Mt.rng., Bureya riv., 2000 m, tundra, Vaschenko S. leg. 18 mm
Copyright © Alexander Anichtchenko

Carabus (Tachypus) auratus auratus Linne, 1761
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Carabus (Tachypus) auratus s.str. (as brunieri Barthe, 1921)
Male adult
SC-FRANCE, Aude, Rennes les Bains, 6.VII.1989, Sunard A. leg.
male, 26mm
Copyright © Ruslan Panin

Carabus (Tachypus) auratus auratus Linne, 1761
Female adult
30 mm
Germania, Oberfranken, Pegnitz
leg.det. U.Schmidt, 1972
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Carabus (Tachypus) auratus lasserrei Doue, 1855
Male adult
SC-FRANCE, Midi-Pyrуnуes reg. Tarn, Saint Paul cap de Joux, 24.V.2003, A. Laforgue leg.
male, 24mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) monilis Fabricius, 1792
Male adult
France 39, Foncina le Haut 300m, 23.6.1979; 29 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) monilis Fabricius, 1792 (as taunicus)
Male adult
W Germany, Vogelsberg Mt., 700m, 13.5.1985
male, 24 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) monilis Fabricius, 1792 (as interpositus Géhin, 1880)
Male adult
NW France,
Basse-Normandie, Orne, Forкt d'Йcouves, V.2009, Pascal Stefani leg.
male, 29mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) monilis (as interpositus Gehin, 1880)
Male+Female adult
NW France,
Haute-Normandie reg., Bois Les Loges, Etretat, Seine-Maritime,10.V.1988
female, 28mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) monilis (as amoenus Baudet-Lafarge, 1836)
Female adult
C France,
Puy-de-Dome, Col de la Croix Saint Robert h=1300m, VII.2008, Pascal Stefani leg.
female, 24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) monilis (as helveticus Heer, 1837 Holotype) Holotype
Male adult
Carabus (Morphocarabus) monilis (as helveticus Heer, 1837 Holotype)
?hybrid of monilis/cancellatus
Museum of Zurich
photo Anichtchenko
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) monilis (as dupeuxi Deuve, 1974)
Male+Female adult

France,
Charente, Foret de Braconne VII.1998, Laforgue leg.
famale, 27mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) michailovi Kabak, 1992
Male adult
NE Kazakhstan, S Altay Mts., Sarymsakty mt. rng.,
Sarymsakty riv., 2400 m, 22.6.2000 Nekroshevich S. leg.
male, 18 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) michailovi Kabak, 1992
Male adult
NE Kazakhstan, S Altay Mts., Sarymsakty mt. rng.,
Sarymsakty riv., 2400 m, 22.6.2000 Nekroshevich S. leg.
male, 17 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) michailovi Kabak, 1992
Male adult
NE-Kazakhstan,
S Altai Mts.,Sarym-Sakty Range, Sarym-Sakty riv., h=2400m. 21.VI.1999., A.Klimenko leg.
male,18mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) mestscherjakovi telezkianus Shilenkov, 1996
Male adult
Carabus (Morphocarabus) mestscherjakovi telezkianus Shilenkov, 1996
Rusia, Altai, S shore of Teletskoe lake, 5.VII.1968
Copyright © Dmitri Obydov

Carabus (Morphocarabus) mestscherjakovi telezkianus Shilenkov, 1996
Male adult
Carabus (Morphocarabus) mestscherjakovi telezkianus Shilenkov, 1996
SW Khakasia, Abakanskii Mt.rng., Choochek Mt., 20 km SSE Mrassu vill.,
1800m, 7-19.7.1999 Lomakin D. leg.
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
Male adult
W Sayan mts., Kopen mt., Dzhabash riv., 75 km
S from Arbaty vill., 2235m, 26-30.6.2003 Brinev A. leg.
long. 17 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
Male adult
Russia, S Siberia,
S Khakassia, N Tuva reg., Sailyg-Khem-Taiga mts. range h=2400m, 10.VII.2004
male, 18mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
Male+Female adult
Russia, S Siberia,
S Khakassia, N Tuva reg., Sailyg-Khem-Taiga mts. range h=2000-2600m,10-30.VI.2005, S.Vashchenko & S.Alexeenko leg.
male,17.5mm/female,19mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Morphocarabus) mestscherjakovi pseudoodoratus Brinev, 2002 Paratype Paratype
Male adult
Krasnojarsk reg., Aradan distr., 5 km SW Aradan vill., Us riv., forest, 52є32`N 93є27`E 9-12.VII.1998 A.Brinev leg male, 18 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) mestscherjakovi pseudoodoratus Brinev, 2002 Paratype Paratype
Female adult
Krasnojarsk reg., Aradan distr., 5 km SW Aradan vill., vall. of Us riv., A.Brinev leg. female, 20 mm photo & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
Male adult
Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
W Sayan, Olenya rechka
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
Male adult
Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
W Sayan, Kulumys Mt.R., env. Oiskoje Lake, mountain tundra, 12-14. 07.1999 A.Semin leg.
male
L=17,0 mm
Copyright © Maksim Smirnov

Carabus (Morphocarabus) kozhantschikovi martjanovi Lutshnik, 1924
Female adult
Khakassia, 1800 m, near Cheremushki vill., 12.6.-20.7.2000
female, 21 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
Male adult
W Sayan Mts., Kazhyr-Suk riv., 80 km W Oleniya
Rechka, W part of Oiskii mt. rng., 30.6.-7.7.1999
2100 m, Brinev A. leg.
male, 19 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
Male adult
W Sayan, low fl. Kazyr-Suk riv., 80km W meteostation Olenia Rechka, W part of Oiskii mt.rng., 30.VI-7.VII.1999 2100m, Brinev A. leg.
male, 20mm
det. Brinev A.
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 18789 (as viridipennis Kraatz, 1886, nec Lewis,
Male adult
E Rusia, Khabarovsk reg.,
near Birobidzhan VI.1991
male, 26 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
Male adult
Rusia, S Primorie, Vladivostok distr., Razdolnaya riv., near Kluchevoi vill., 10-14.VII.1988, S. Muhanov leg.; Obydov det. as tristiculus Kraatz, 1879
Copyright © Dmitri Obydov

Carabus (Morphocarabus) hummeli cf pusongensis Imura, 1993
Male adult
China, Beijing, Wulingshan mountain, 1985
Copyright © Ye Liu

Carabus (Morphocarabus) hummeli ssp.
Male adult
Carabus (Morphocarabus) hummeli ssp.
China, East Inner Mongolia, Chifen, 2006, Shi H.L. leg.
Photo & det. by liuye
Copyright © Ye Liu

Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
Male adult
China, Jinling pro.;
Photo by liuye
det. Anichtchenko A.
Copyright © Ye Liu

Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
Male adult
Russia, Far East,
Primorskiy Kray, near Kamenushka vill., 2-3.VIII.1992, K.E.Dovgaylo leg.
male, 27mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) hummeli pusongensis Imura, 1993
Male adult
S Primorie, Chernye gory, Poima riv.,
30.7-5.8.1999 Anichtchenko A. leg.
male, 26 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) hummeli nevelskii Shilenkov, 1996
Female adult
Rusia, S Sakhalin isl., VII.1993, G. Vasiliev leg.

Copyright © Dmitri Obydov

Carabus (Morphocarabus) hummeli nevelskii Shilenkov, 1996
Male+Female adult
Russia, Sakhalin isl.,
Makarov distr., Lesnaja riv., VII.1992, M. Nesterov leg.
female,23mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) hummeli hummeli Fischer, 1823
Male adult
Carabus (Morphocarabus) hummeli hummeli Fischer, 1823
Rusia, Chita reg., Darasun, VII.1991
I. Avdeev leg.
male, 23 mm
Copyright © Alexander Anichtchenko

Carabus (Sphodristocarabus) varians varians Fischer, 1827
Male+Female adult
female: Transcaucasia, Armenia, Amasia distr., Karakhachskiy mountain pass h=2270m, 3.VIII.1985 P.V.Kazaryan leg., 22mm male: Armenia, Amasia distr., Arpalach vill. VI.1999, P.V.Kazaryan leg.
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) varians rugatus Breuning, 1934
Female adult

Copyright © Ruslan Panin

Carabus (Sphodristocarabus) blumenthaliensis basilewskianus Breuning & Ruspoli, 1970
Male adult
SW Georgia, Adzharia 10 km Baku 2000m
18.6.98
25 mm
Copyright © Alexander Anichtchenko

Carabus (Sphodristocarabus) blumenthaliensis basilewskianus Breuning & Ruspoli, 1970
Male+Female adult
NE-Turkey,
Artvin Prov., Hopa vers Borcka, Cankutaran Gecidi h=800m, 17.VI.2007 leg. D. Prunier
female, 24mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) varians janthinus s.str.
Male+Female adult
NW-Caucasus, Russia, Krasnodar reg.,Tuapse distr., Mt.Semashkho 04.V.2008, D.Fominyh leg. female, 24mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) varians armeniacus Mannerheim, 1830
Female adult
NE Turkey, Artvin Prov., Borçka, Macahel Gecidi h=1450m, 18.VI.2007, D.Prunier leg. female, 26mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) tokatensis almushensis Mandl, 1975
Male adult
Turkey, Sivas, 20 km N Zara, N Arapca, 1750m; 11.VII.1993
Copyright © Alexander Anichtchenko

Carabus (Sphodristocarabus) scovitzi czipkai Heinz, 1980
Male adult
Turkey, Artvin, Kafkasor, 1600 m
4.7.1996 Staven and Skoupy leg.
male, 22 mm
Copyright © Alexander Anichtchenko

Carabus (Sphodristocarabus) macrogonus simonianus Breuning & Ruspoli, 1970
Female adult
Turkey, Ordu, Golkoy, 1220 m, 19.7.91
Copyright © Alexander Anichtchenko

Carabus (Sphodristocarabus) macrogonus resplendens Breuning, 1934
Female adult
Turkey; Cayirkent Samsun 12.VII.1996
Copyright © Alexander Anichtchenko

Carabus (Sphodristocarabus) georgia Cavazzuti, 1984
Male adult

Copyright © Boleslav Brezina

Carabus (Sphodristocarabus) bohemani bohemani Menetries, 1832
Female adult
S Azerbaydzan, Lenkoran distr., Vazha riv.,
2 km NW Dashdatuk vill. 38º40*09N 48º42*06E
17.4.2004 Makarov K. leg.
female, 22 mm
Copyright © Alexander Anichtchenko

Carabus (Sphodristocarabus) bohemani bohemani Menetries, 1832
Male adult
Azerbaijan,
Talysh Mts., Polikesh, h~1800m.,2-6.VI.2001 Dolin V.G. leg.
male, 20mm.
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) hollbergi Mannerheim, 1827
Male adult
N Azerbaijan,
4km W Ismayilli, h=700m, 9.VII.2008, Pljushch I.G. leg.
male,22mm
Copyright © Ruslan Panin

Carabus (Semnocarabus) transiliensis Semenov, 1896
Male adult
Kirgizstan, S slope Kungei Ala Too
Oi-Tal riv. 3000m 14.7.1999
13 mm
Copyright © Alexander Anichtchenko

Carabus (Semnocarabus) transiliensis Semenov, 1896
Male+Female adult
E KAZAKHSTAN,
Bolshoe Almaatinskoe Lake,30.VI.1992
15mm
Copyright © Ruslan Panin

Carabus (Semnocarabus) transiliensis Semenov, 1896
Male adult
SE-KAZAKHSTAN,
Ketmen Mts.,N-slopes h=2250-2500m.1-5.VI.1990
16mm
Copyright © Ruslan Panin

Carabus (Semnocarabus) regulus regulus Dohrn, 1882
Male adult
photo M. Smirnov from www.zin.ru
Copyright © Maksim Smirnov

Carabus (Semnocarabus) regulus regulus Dohrn, 1882
Male adult
E Kirgizia, Kaindy-Ketta mts. rng. Sary-Dzhas riv.
Maidadyr near Enelchek vill. 6-8.7.1999 Klimenko leg.
14 mm
Copyright © Alexander Anichtchenko

Carabus (Semnocarabus) erosus erosus Motschulsky, 1865
Male adult
Kirgizia, S Kungei Ala-Too, Kichi-Yuryukty
2800m 1.7.1999 Subankulov leg.
male, 21 mm
Copyright © Alexander Anichtchenko

Carabus (Semnocarabus) erosus corax Semenov, 1896
Male adult
SE Kazakhstan, Ili riv., Bakanas vill.,
1-5.5.1999 Klimenko A. leg.
Copyright © Alexander Anichtchenko

Carabus (Semnocarabus) carbonicolor carbonicolor Morawitz, 1886
Male adult
E Kirgizia, Kaindy-Ketta mts. rng., Sary-Dzas riv.,
Maidadyr near Enelchek vill., 6-8.7.99 Klimenko leg.
Copyright © Alexander Anichtchenko

Carabus (Semnocarabus) carbonicolor callosus Semenov, 1896
Male adult
Kazakhstan, Bol. Alma-Atinka riv. 1600m 1.6.1999 Klimenko A. leg.
male, 19 mm
Copyright © Alexander Anichtchenko

Carabus (Scambocarabus) sculptipennis Chaudoir, 1887
Male adult
China, Beijing, Xiaolongmeng, 2005, Liu ye coll.;
22mm; photo by Liuye
Copyright © Ye Liu

Carabus (Scambocarabus) kruberi kruberi Fischer, 1822
Male adult
Rusia, Chita reg., 23 km SW Borzia, Borzinskoe lake,
1.VI.2002 Anichtchenko A. leg.
Copyright © Alexander Anichtchenko

Carabus (Scambocarabus) kruberi Fischer, 1822
Male adult
China, Beijing, labagoumeng, 2004, Li fei coll.;
21mm; photo by Liuye
Copyright © Ye Liu

Carabus (Morphocarabus) hummeli biamensis Obydov, 2007 Paratypus Paratype
Male adult
Russia, North Primorie, B. Svetlovodnaya riv. (Biame riv.), 40 km from source, 30.07.1976, A. Klitin leg.
Copyright © Dmitri Obydov

Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
Female adult
E.RUSSIA, Primorsky ter., Chuguevsky distr., 30 km ESE from Chuguevka, 9-14.VIII.2007
Copyright © Nil Belonogov

Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
Male adult
E.RUSSIA, Primorsky ter., Chuguevsky distr., 30 km ESE from Chuguevka, 9-14.VIII.2007
Copyright © Nil Belonogov

Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 18789 (as viridipennis Kraatz, 1886, nec Lewis,
Female adult
Primorskii reg., near Kovalerovo
VII.1990 Evstigneev A. leg.
female, 25 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) hummeli ? middendorfi Menetries, 1851
Male adult
Carabus (Morphocarabus) hummeli ? middendorfi Menetries, 1851
Rusia, Khabarovsk reg., Ohotsk sea, Port Ayan, 08.24
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) henningi pelevini Obydov 2006
Female adult
NE Kazakhstan, S Altai mt.rng., Sarymskiy mt. rng., Sarymsakty riv., 2400m, 22.VI.2000 Nekroshevich S. leg. female 17mm det. Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) henningi pelevini Obydov 2006 HT
Male adult
Carabus (Morphocarabus) henningi pelevini Obydov 2006 HT
Kazakhstan, S Altai, Sarym-Sakty Mt.rng., 3000-3400 m,
18-20.VI.1998, A. Klimenko leg.
Copyright © Dmitri Obydov

Carabus (Morphocarabus) henningi peczoricus Obydov 2000 HT Holotype
Male adult
Carabus (Morphocarabus) henningi peczoricus Obydov 2000 HT
Rusia, Komi republic, Kosiyu River mouth, (Pechora basin) 21.VII.1980
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) henningi oviformis Beheim & Breuning, 1943
Male adult
Carabus (Morphocarabus) henningi oviformis Beheim & Breuning, 1943
Rusia, Krasnoyarsk reg., Taimyr AO, near Norilsk,
07.2004, M. Shestopalov leg.
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) henningi melyachi Shilenkov & Menstshikov, 1996
Male adult
Carabus (Morphocarabus) henningi melyachi Shilenkov & Menstshikov, 1996
C Altai mts, Seminskii mt.rng., Katun riv., 3km S Ust-Sema vill.
4.VII.2002 A. & N. Menschikovy
male, 17mm
det. Menschikov A.
photo Anichtchenko
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) henningi henningi Fischer, 1817
Male adult
S. Ural, Bashkiria, near Mesjagutovo vill., 09.07.2001 A. Menstschicov leg.
Copyright © Anton Menstschikov

Carabus (Morphocarabus) henningi henningi Fischer, 1817
Male adult
SE Siberia, Russia, Irkutsk reg., near Pivovarikha vill., 7.V.07, D. Fominykh leg. female, 21mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) henningi hamarensis Shilenkov, 1996
Female adult
S Siberia, Hamar-Daban mts., Snezhnaya riv., 1-6.VI.2007, Anichtchenko A. leg. & det., 20 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) henningi hamarensis Shilenkov, 1996
Female adult
Carabus (Morphocarabus) henningi hamarensis Shilenkov, 1996 loc.typ.
Copyright © Alexander Anichtchenko

Carabus (Rhigocarabus) xiei xiei Deuve, 1992
Male adult
male, 18mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) xiei xiei Deuve, 1992
Female adult
female, 19mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) xiei xiei Deuve, 1992
Male adult
male, 20mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) xiei xiei Deuve, 1992
Male adult
male, 17mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) xiei sanggarparicus Deuve, 1996 Holotype Paratype
Male adult
Holotype
male, 19mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) xiei sanggarparicus Deuve, 1996
Female adult
Paratype
female, 20mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) xiei barkamensis Deuve, 1994 Holotype Paratype
Male adult
Holotype, male, 19mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) xiei barkamensis Deuve, 1994
Male adult
Paratype, female, 19mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) xiangchengicus qingmaicus Deuve, 2007
Male adult
male, 15mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) xiangchengicus qingmaicus Deuve, 2007
Female adult
female, 18mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) xiangchengicus qingmaicus Deuve, 2007
Female adult
female, 16mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) xiangchengicus sangpiensis Deuve, 2007 Paratype
Male adult
male, 15 mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) thais thais Heinz, 1997
Male adult
male, 16mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) thais thais Heinz, 1997
Male adult
male, 19mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) thais thais Heinz, 1997
Male adult
male, 20mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) subindigestus subindigestus Deuve, 1994
Female adult
female, 15mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) subindigestus subindigestus Deuve, 1994
Male adult
male, 16mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) subindigestus subindigestus Deuve, 1994
Male adult
male, 15mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) subindigestus subindigestus Deuve, 1994
Male adult
male, 16mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) sehnali Brezina & Hackel, 2006 Holotype Paratype
Male adult
male, 20mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) sehnali Brezina & Hackel, 2006
Female adult
female, 23mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) roborowskii roborowskii Semenov, 1887
Male adult
China, E Tibet, Chegarla 4800m, 20 km E
of Rioche, 26.6.97 A. Wrzecionko leg.
Copyright © Alexander Anichtchenko

Carabus (Rhigocarabus) rhododendron Deuve & Imura, 1991
Male adult
male, 18mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) rhododendron Deuve & Imura, 1991
Female adult
female, 19mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) rhododendron Deuve & Imura, 1991
Male adult
male, 17mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) quindecim Cavazzuti, 2002 Holotype
Male adult
Holotype, male, 15mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) pseudopusio Deuve, 1996 Paratype
Male adult
male, 15mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) pseudopusio Deuve, 1996
Female adult
female, 16mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) poeta Semenov, 1898 Holotype Paratype Lectotype
Male adult
male, 17mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) poeta Semenov, 1898
Male adult
male, 18mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) poeta (as korsakowi Semenov, 1898) Holotype
Female adult
Holotype, female 18 mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) poeta (as korsakowi Semenov, 1898) Syntypes
Female adult
female, 16mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) poeta (as korsakowi Semenov, 1898) Lectotype
Male adult
male, 17mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) pepek Imura & Kalab, 2006 Paratype
Male adult
male, 17mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) pepek Imura & Kalab, 2006 Paratype
Female adult
female, 18mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) morawitzianus breviculus Cavazzuti & Ratti, 1998 Paratype
Male adult
male, 15 mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) maleki Deuve, 1991 Paratype
Male adult
male, 14mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) maleki Deuve, 1991
Female adult
female, 15mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) maleki Deuve, 1991 Paratype
Male adult
male, 15mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) itzingeri rugulosior Deuve, 1996 Paratype Paratype
Male adult
male, 18mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) itzingeri itzingeri Breuning, 1934
Male adult
male, 19mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) itzingeri itzingeri Breuning, 1934
Female adult
female, 20mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) itzingeri kozaburoi Imura, 1999 Paratype
Male adult
male, 16mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) itzingeri kozaburoi Imura, 1999 Paratype
Female adult
female, 18mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) handelmazzettii handelmazzettii Mandl, 1955
Male adult
male, 18mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) handelmazzettii handelmazzettii Mandl, 1955
Female adult
female, 20mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) handelmazzettii handelmazzettii Mandl, 1955
Female adult
female, 21mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) handelmazzettii habaicus Deuve, 2005 Paratype
Male adult
male, 16mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) pusio pusio s.str.
Male adult
male, 16mm
(as hylonomus)
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) pusio pusio s.str.
Female adult
female, 17mm
(as hylonomus)
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) gracilicollis gracilicollis Semenov, 1887
Male adult
male, 15mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) gracilicollis gracilicollis Semenov, 1887
Female adult
female, 16mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) gigoloides priapeius Cavazzuti, 2006
Male adult
male, 20mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) gigoloides jone Brezina & Hackel, 2004 Holotype Paratype
Male adult
male, 19mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) gigoloides jone Brezina & Hackel, 2004
Female adult
female, 20mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) gigolo (as qilingianus Cavazzuti, 2006)
Male adult
18mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) gigolo Heinz & Brezina, 1996 Paratype
Male adult
20mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) gigolo Heinz & Brezina, 1996 Paratype
Male adult
20 mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) gigolo Heinz & Brezina, 1996 Paratype
Female adult
21 mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) gentleman Brezina & Hackel, 2004 Holotype
Male adult
male, 19mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) gentleman Brezina & Hackel, 2004 Paratype
Female adult
female, 20mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) dacatraianus Deuve, 1995
Male adult
China, E Tibet, Angenla 5000-5100m
45 km W of Bachen 29.6-2.7.1997 Wrzecionko leg.
16 mm
Copyright © Alexander Anichtchenko

Carabus (Rhigocarabus) choui Deuve, 1989
Male adult
male, 16mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) choui Deuve, 1989
Female adult
female, 17mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) cateniger cateniger s.str. (as sommitalis)
Male adult
male, 16 mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) cateniger cateniger Morawitz, 1886
Female adult
female, 19mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) cateniger cateniger Morawitz, 1886
Male adult
male, 18mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) cateniger saltuensis Cavazzuti, 2006 Paratype
Female adult
female, 19mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) cateniger dabanicola Deuve, 1998
Male adult
male, 16mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) cateniger dabanicola Deuve, 1998
Female adult
female, 18mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) buddaicus bocsiki Deuve, 2007 Holotype
Male adult
male, 20 mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) buddaicus angenlaicus Deuve, 2007 Paratype
Male adult
male, 18 mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) broukpytlik Brezina & Hackel, 2004 Holotype Paratype
Male adult
male, 19mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) broukpytlik Brezina & Hackel, 2004
Female adult
female, 20mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) broukpytlik Brezina & Hackel, 2004
Male adult
male, 16mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) morawitzianus breviculus Cavazzuti & Ratti, 1998 Paratype
Male adult
China, N Tibet, Tangula Shan Mts., Tangula
pass 5100-5300m, 8-11.7.1997 A. Wrzecionko leg.
Copyright © Alexander Anichtchenko

Carabus (Rhigocarabus) batangicus fickleri Korell & Richter, 1996 Paratype
Male adult
male, 16 mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) poeta Semenov, 1898
Male adult
male, 17mm
Copyright © Boleslav Brezina

Carabus (Oreocarabus) reitterianus Breuning, 1934
Male adult
China, Gansu, ca 25 km E Xiahe 2700-2900m, Wagatan Timber
30.7.-2.8.1993 Heinz leg.
male, 23 mm
Copyright © Alexander Anichtchenko

Carabus (Oreocarabus) nanwutai Kleinfeld, Korell & Wrase, 1996 Holotype Paratype
Male adult
China, Shaanxi, Qing Ling Shan, track Hou Zen Zi vill.
to Taiban Shan, 2500m, 27-29.6.1998 mixed forest
male, long. 19 mm
Copyright © Alexander Anichtchenko

Carabus (Oreocarabus) nanwutai Kleinfeld, Korell & Wrase, 1996
Male+Female adult

Copyright © Frank Kleinfeld

Carabus kitawakianus kitawakianus Imura, 1993
Male adult
China, Shaanxi, Tschinling mts.,
Taibeichan Nat. res. 2000m 5-15.8.2005
male, 27 mm
Copyright © Alexander Anichtchenko

Carabus (Pseudotribax) ferghanicus Breuning, 1933
Male adult
male, 29mm
Copyright © Boleslav Brezina

Carabus (Pseudotribax) validus Kraatz, 1884
Male adult
male, 31mm
Copyright © Boleslav Brezina

Carabus (Pseudotribax) validus Kraatz, 1884
Female adult
female, 32mm
Copyright © Boleslav Brezina

Carabus (Pseudocranion) sackeni sackeni Semenov, 1898
Male adult
male, 23mm
Copyright © Boleslav Brezina

Carabus (Pseudocranion) sackeni remondianus Deuve, 1996 Paratype
Female adult
Museum of Institute of Systematic Biology (Daugavpils, Latvia)
Copyright © Alexander Anichtchenko

Carabus (Pseudocranion) labrangicus macilentior Deuve, 1996
Female adult
female, 21mm
Copyright © Boleslav Brezina

Carabus (Pseudocranion) gansuensis jedompansis Deuve, 1989
Male adult
China, Sichuan, Langmusi 3200m 7.1999
Copyright © Alexander Anichtchenko

Carabus (Pseudocranion) fumigatus fumigatus Semenov, 1898 Lectotype
Male adult
male, 22mm
Copyright © Boleslav Brezina

Carabus (Pseudocranion) fumigatus fumigatus Semenov, 1898 Syntypes
Female adult
female, 22mm
Copyright © Boleslav Brezina

Carabus (Morphocarabus) gebleri ultimus Obydov, 1999
Female adult
Carabus (Morphocarabus) gebleri ultimus Obydov, 1999 PT
E Kazakhstan, Altai, near Zyrjanovska, 6.VII.1995, D. Obydov leg.
female
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) gebleri ultimus Obydov, 1999 Holotype
Male adult
Carabus (Morphocarabus) gebleri ultimus Obydov, 1999 HT
E Kazakhstan, Altai, near Zyrjanovska, 6.VII.1995, D. Obydov leg.
male
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) gebleri gebleri Fischer, 1817
Female adult
Carabus (Morphocarabus) gebleri gebleri Fischer, 1817
Russia, Altai, Zmeinogorsk
female
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) gebleri gebleri Fischer, 1817
Male adult
Carabus (Morphocarabus) gebleri gebleri Fischer, 1817
Rusia, Altai, Zmeinogorsk
male
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) chaudoiri zaikai Obydov, 1999 PT Paratype
Female adult
Carabus (Morphocarabus) chaudoiri zaikai Obydov, 1999 PT
Rusia, Tuva, Akademik Obruchev Mt. rng., 2100 m;, 52°00'N 95°34'E,
Sainak pass, tundra, 2.VII.1998, D. Obydov leg.
female,
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) chaudoiri juliae Obydov, 2002
Male adult
male, 18mm
Copyright © Boleslav Brezina

Carabus (Morphocarabus) chaudoiri czadanicus Obydov, 1997 HT Holotype
Male adult
Carabus (Morphocarabus) chaudoiri czadanicus Obydov, 1997 HT
Rusia, Tuva, near Hadyn vill., 28-30.VI.1994, R. Shmarzh leg.
male,
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) chaudoiri beladici Obydov, 2001
Male adult
S-Tuva
East Tannu-Ola,Biche-Sailig r.h=2100m, 15.VII.1996, D.E.Lomakin leg.
Copyright © Anton Menstschikov

Carabus (Morphocarabus) chaudoiri beladici Obydov, 2001 PT Paratype
Female adult
Carabus (Morphocarabus) chaudoiri beladici Obydov, 2001 PT
Rusia, Tuva, E Tannu-Ola Mt.rng., near Kara-Hol lake, 29-31.V.2000, M. Beladzhik leg
female,
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) aeruginosus zyrjanovskianus Shilenkov & O.Berlov, 1996
Male+Female adult
NE KAZAKHSTAN,
S-Altays, Ulbinsky Mt. h=900m., Bobrovka vill., 24.VI.01, A. Klimenko leg.
female,25 mm
Copyright © Ruslan Panin

Carabus (Pseudocoptolabrus) taliensis taliensis (Fairmaire, 1886)
Male adult
China, Yunnan, Lijiang env., 2700 m, 10.6.1998
Copyright © Alexander Anichtchenko

Carabus (Pseudocoptolabrus) taliensis s.str. (as yulongxuensis)
Male adult
male, 26mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) taliensis s.str. (as yulongxuensis)
Male adult
male, 24mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) taliensis s.str. (as yulongxuensis)
Female adult
female, 29mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) taliensis yanmenensis Deuve, 1996
Male adult
male, 27mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) taliensis yanmenensis Deuve, 1996
Female adult
female, 28mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) taliensis xueshanicola Deuve, 1992
Female adult
female, 30mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) taliensis xueshanicola Deuve, 1992
Male adult
male, 26mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) taliensis xueshanicola Deuve, 1992
Male adult
male, 27mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) taliensis xueshanicola Deuve, 1992
Male adult
male, 28mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) taliensis s.str. (as peterheisei) Holotype
Male adult
Holotype
Copyright © Frank Kleinfeld

Carabus (Pseudocoptolabrus) taliensis myanmarensis Imura, 1999
Female adult
female, 36 mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) taliensis mulianus Deuve, 1995
Male adult
male, 27mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) taliensis liqiuensis Imura, 1999
Male adult
male, 32mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) taliensis s.str. (as lijiangensis) Paratype
Female adult
female, 33mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) taliensis s.str. (as lijiangensis)
Male adult
male, 31mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) taliensis jingpingensis Deuve, 2005 Paratype
Male adult
male, 26mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) taliensis jingpingensis Deuve, 2005 Paratype
Female adult
female, 30mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) nosei taronensis Imura, 2001 Paratype
Female adult
female, 28mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) burmanensis Breuning, 1932
Male adult
male, 23mm
Copyright © Boleslav Brezina

Carabus (Morphocarabus) aeruginosus verzhutzkii O.Berlov & Shilenkov, 1996
Male adult
Carabus (Morphocarabus) aeruginosus verzhutzkii O.Berlov & Shilenkov, 1996
C Altays, Katun r., 5 km. N Katanda vill., 21.06.2002, A. Menstschicov leg.
Copyright © Anton Menstschikov

Carabus (Pseudocoptolabrus) armiger armiger Imura, 1997
Female adult
female, 30mm
Copyright © Boleslav Brezina

Carabus (Morphocarabus) aeruginosus verzhutzkii O.Berlov & Shilenkov, 1996
Male adult
C Altai Mts., Seminskii Mt.rng., Katun riv., 3 km S Ust'-Sema riv.,
4.VII.2002 A. & N. Menschikovy leg.
det. Mentschikov A.
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) aeruginosus salechardensis Obydov, 1999 Paratype
Male adult
Rusia, Selehard, tundra, 30.VIII.1953, I. Telyshev leg

Copyright © Dmitri Obydov

Carabus (Morphocarabus) aeruginosus hermani Mannerheim, 1827 loc.typ.
Male adult
C Rusia, S Ural Mts., Zlatoust, 22-28.VI.1952, S. Nikulina leg.
male,

Copyright © Dmitri Obydov

Carabus (Procrustes) paiafa acuticollis Motschulsky, 1858
Male adult

Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) aeruginosus eschscholtzi Mannerheim, 1827
Male adult
Russia, W Altai, Zmeinogorsk distr., near Cherepanovskii vill., 2-16.6.1990, D. Obydov leg.
Copyright © Dmitri Obydov

Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
Male adult
SE Siberia, E Sayan, Nizhneudinsk, h=500m, 21-26.VI.2006,
male, 26mm
Copyright © Ruslan Panin

Carabus (Mimocarabus) maurus osculatii Osculati, 1844
Male adult
Carabus (Mimocarabus) maurus osculatii Osculati, 1844
Copyright © Alexander Anichtchenko

Carabus (Mimocarabus) maurus maurus Adams, 1817
Male+Female adult
Armenia,
Ashotsk distr., Tsoghamarg vill., 24-27.V.1999 P.V.Kazaryan leg.
female, 20mm/male,18mm
Copyright © Ruslan Panin

Carabus (Procrustes) impressus Klug, 1832 (as carmelita)
Male adult

Copyright © Safronov

Carabus (Procrustes) iimpressus Klug, 1832 (as drusum) Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Procrustes) impressus Klug, 1832 (as desertum) Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Procrustes) coriaceus rugifer Kraatz, 1877
Male adult

Copyright © Safronov

Carabus (Procrustes) coriaceus kindermanni Waltl, 1838
Male+Female adult
NW TURKEY,
Stambul Avdzhilar, (Eropean part), h~20 m, 10-29.X.2007 M.V.Cheriavskaya leg./det. I.A.Solodovnikov
female,31mm
Copyright © Ruslan Panin

Carabus (Procrustes) coriaceus hopffgarteni Kraatz, 1878
Male adult

Copyright © Safronov

Carabus (Procrustes) coriaceus cerisyi Dejean, 1826 (as foudrasi)
Male adult

Copyright © Boleslav Brezina

Carabus (Procrustes) coriaceus coriaceus Linne, 1758
Female adult
The neighborhood Nicolo-Berezovskoe republic of
Bashkortostan 5.07
Copyright © Sunvas

Carabus (Procrustes) coriaceus coriaceus Linne, 1758
Male adult
length: 32.42 mm
Germania, Oberfranken, Weismain
leg.det. U.Schmidt, 1983
Foto: U.Schmidt, 2005
Copyright © U.Schmidt

Carabus (Procrustes) coriaceus cerisyi Dejean, 1826
Male adult
male, 30mm
Copyright © Boleslav Brezina

Carabus (Procrustes) coriaceus cerisyi Dejean, 1826
Female adult
female, 31mm
Copyright © Boleslav Brezina

Carabus (Procrustes) clypeatus clypeatus Adams, 1817
Female adult
C-Caucasus, N Ossetia, vall. Mamyhdon riv., h=2100
Koval A.G. leg; 08.08.2001
Copyright © Anton Menstschikov

Carabus (Procrustes) chevrolati korbi Breuning, 1928
Male adult
length: 28 mm
Türkei, Tülü/Sathan, Erzincan
leg. Staven & Willmer; det. K.Staven, 2003
Foto: U.Schmidt, 2009
Copyright © U.Schmidt

Carabus (Procrustes) anatolicus lycicus Breuning, 1934
Male adult

Copyright © Safronov

Carabus (Procrustes) chevrolati chevrolati Cristoforis & Jan, 1837
Male+Female adult
female, 31mm: Ashotsk,Vardaghbyur vill., 30.V.1997
male, 30 mm: Amasia, Chaldash vill.,8-11.VIII.1997
P.V. Kazaryan leg.
Copyright © Ruslan Panin

Carabus (Procerus) scabrosus tauricus Bonelli, 1811
Female adult

Copyright © Safronov

Carabus (Cechenochilus) heydenianus Starck, 1889
Male adult
Caucasian nat. park., S slope Pseashkho mt., 2500 m
21.7.1975 Ovtsharenko V. leg.
Copyright © Alexander Anichtchenko

Carabus (Cechenochilus) heydenianus Starck, 1889
Male+Female adult
NW-Caucasus,
Krasnodar reg., Mostovskoy distr., Skirda Mt Range, N slopes Yatyrgvart Mt. h=2300-2500m., 09-10.VII.2008, D. Fominykh leg.
female,19mm/male,18mm
Copyright © Ruslan Panin

Carabus (Cechenochilus) adangensis Gottwald, 1983
Male adult
male, 19mm
Copyright © Boleslav Brezina

Carabus (Cechenochilus) adangensis Gottwald, 1983
Female adult
female, 18mm
Copyright © Boleslav Brezina

Carabus (Cechenochilus) adangensis Gottwald, 1983
Female adult
Russia, NW-Caucasus:
Karacaj-Cherkessija, Abishira–Akhuba Mt. Range h=2000-2650m, Kyafar-Agur Riv. 14–17.VII.2009, Fominykh D. leg.
female,21mm
Copyright © Ruslan Panin

Carabus (Cechenochilus) adangensis (as gusevi)
Male adult
male, 19mm
Copyright © Boleslav Brezina

Carabus (Cechenochilus) adangensis (as gusevi)
Male adult
male, 19mm
Copyright © Boleslav Brezina

Carabus (Cechenochilus) lazorum Belousov & Zamotajlov, 1999
Male adult
male, 20mm
Copyright © Boleslav Brezina

Carabus (Cechenochilus) boeberi longiceps Chaudoir, 1846
Male adult
C-Caucasus, N Ossetia, near Kozskii, h=2800,
A.G. Koval leg. 10.08.2001; 12 mm
Copyright © Anton Menstschikov

Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Holotype
Male adult
Russia, Karachai-Circassian Republic, Abishira-Akhuba Mount Range between passes Agur and Mylgvar, 2785–2830 m.
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009
Male adult
male, 17mm
Copyright © Boleslav Brezina

Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Paratype
Male adult
Russia, W Caucasus,
Karacaj-Cherkessija, Abishira-Akhuba Mnt Range, Area of pass Turii h~2650-2850m, 17.VII.2009, Fominykh D. leg.
male,17 mm
photo & coll. Panin R.
Copyright © Ruslan Panin

Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Paratype
Female adult
Russia, Karachai-Circassian Republic, Abishira-Akhuba Mount Range between passes Agur and Mylgvar, 2785–2830 m. PARATYPUS
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009
Female adult
Russia, Karachai-Circassian Republic, Abishira-Akhuba Mount Range between passes Agur and Mylgvar, 2785–2830 m. PARATYPUS
Copyright © Igor Solodovnikov