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Carabidae of the World
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Licininae Bonelli, 1810
→
Chlaeniini Brulle, 1834
→
Chlaeniina Brulle, 1834
→
Chlaenius Bonelli, 1810:
→ Chlaenioctenus Bates, 1892
Region
Afghanistan (AF)
Albania (AL)
Aleutian isl (ALU)
Algeria (AG)
Andorra (AN)
Angola (ANG)
Antarctica (Antarctica)
Antilles islands (ANT)
Antigua and Barbuda (Antigua and Barbuda)
Barbados (Barbados)
Cayman islands (Cayman islands)
Cuba (Cuba)
Dominica (Lesser Antilles) (Dominica (Lesser Antilles))
Dominican Republic (Hispaniola, Greater Antilles) (Dominican Republic (Greater Antilles))
Grenada (Grenada)
Guadeloupe (Guadeloupe)
Haiti (Haiti)
Jamaica (Jamaica)
Martinique (Martinique)
Puerto Rico (Puerto Rico)
Saint Kitts and Nevis (Saint Kitts and Nevis)
Saint Lucia (Saint Lucia)
St. Vincent (St. Vincent)
Trinidad and Tobago (Trinidad and Tobago)
Arab Emirates (AE)
Argentina (ARG)
Armenia (AR)
Australia (AST)
Christmas Island (Christmas Island)
Lord Howe Island (Lord Howe Island)
Macquarie Island (Macquarie Island)
Tasmania (Tasmania)
Austria (AU)
Azerbaijan (AB)
Azores (AZ)
BALTIC AMBER, Baltic See coast and Amber quarry Jantarny near Kaliningrad, Russia (BAM)
Bahamas (BHM)
Bahrain (BA)
Bangladesh (BGD)
Belgium (BE)
Belize (BLZ)
Bhutan (BT)
Bolivia (BOL)
Bosnia and Herzegovina (BH)
Botswana (BTW)
Brazil (BRZ)
Acre (AC)
Alagoas (AL)
Amazonas (AM)
Arnapá (AP)
Bahia (BA)
Ceará (CE)
District Federal (Brasilia) (DF)
Espirito Santo (ES)
Goiás (GO)
Maranhão (MA)
Mato Grosso (MT)
Mato Grosso do Sul (MS)
Minas Gerais (MG)
Paraná (PR)
Paraîba (PB)
Pará (PA)
Pernambouc (PE)
Piaui (PI)
Rio Grande do Norte (RN)
Rio Grande do Sul (RS)
Rio de Janeiro (RJ)
Rondônia (RO)
Roraima (RR)
Santa Catarina (SC)
Sergipe (SE)
São Paulo (SP)
Tocantins (TO)
Brunei (Brunei)
Bulgaria (BU )
Burundi (BUR)
Byelorussia (BY)
Cambodia (CBD)
Cameroon (CAM)
Canada (CAN)
Canary isls (CI)
Cape Verde (Cape Verde)
Central African Republic (CAR)
Chad (CHD)
Chile (CHL)
Juan Fernández Islands (Juan Fernández Islands)
China (CH)
Anhui (Anhui)
Beijing (Beijing)
Chongqing (Chongqing)
Fujian (Fujian)
Gansu (Gansu)
Guangdong (Guangdong)
Guangxi (Guangxi)
Guizhou (Guizhou)
Hainan (Hainan)
Hebei (Hebei)
Heilongjiang (Heilongjiang)
Henan (Henan)
Honkong (Hongkong)
Hubei (Hubei)
Hunan (Hunan)
Inner Mongolia (Inner Mongolia)
Jiangsu (Jiangsu)
Jiangxi (Jiangxi)
Jilin (Jilin)
Liaoning (Liaoning)
Macau (Macau)
Ningxia (Ningxia)
Qinghai (Qinghai)
Shaanxi (Shaanxi)
Shandong (Shandong)
Shanghai (Shanghai)
Shanxi (Shanxi)
Sichuan (Sichuan)
Tianjin (Tsiensin) (Tianjin)
Xinjiang (Xinjiang)
Xizang (Tibet) (Xizang (Tibet))
Yunnan (Yunnan)
Zhejiang (Zhejiang)
Colombia (COL)
Comoros isl. (COM)
Costa Rica (CRC)
Croatia (CR)
Crozet Islands (CRZ)
Cyprus (CY)
Czech (CZ)
DOMINICAN AMBER, Dominican Republic, Lower Miocene. (DAM)
Democratic Republic of the Congo (CNG)
Denmark (DE)
Diego Garcia isls (DGC)
Djibouti (DJB)
Ecuador (ECD)
Egypt (EG)
Sinai (Sinai)
El Salvador (SVD)
Equatorial Guinea (EQG)
Bioco (Fernando Pó) island (Bioco (Fernando Pó) island)
Eritrea (ERT)
Estonia (EN)
Ethiopia (ETP)
Falkland il. (FLK)
Faroe Islands (FA)
Finland (FI)
France (FR)
Corse (Corse)
French Guiana (GNF)
Gabon (GAB)
Galapagos Islands (GAL)
Georgia (GG)
Abkhazia (Abkhazia)
Ajaria (Ajaria)
South Ossetia (South Ossetia)
Germany (GE)
Great Britain (GB)
Greece (GR)
Aegean Islands (Aegean Islands)
Crete (Crete)
Dodecanese Islands (Dodecanese Islands)
Euboea (Evia) Island (Euboea (Evia) Island)
Ionian Islands (Ionian Islands)
Peloponnese (Peloponnese)
Saronic Islands (Saronic Islands)
Sporades Islands (Sporades Islands)
The Cyclades (The Cyclades)
Guatemala (GTM)
Guyana (GYN)
Honduras (HDR)
Hungary (HU)
Iceland (IC)
India (IND)
Andaman and Nicobar isl. (Andaman and Nicobar isl.)
Andhra Pradesh (Andhra Pradesh)
Arunachal Pradesh (Arunachal Pradesh)
Assam (Assam)
Bihar (Bihar)
Chhattisgarh (Chhattisgarh)
Delhi (Delhi)
Goa (Goa)
Gujarat (Gujarat)
Haryana (Haryana)
Himachal Pradesh (Himachal Pradesh)
Jammu and Kashmir (Jammu and Kashmir)
Jharkhand (Jharkhand)
Karnataka (Karnataka)
Kerala (Kerala)
Ladakh (Ladakh)
Madhya Pradesh (Madhya Pradesh)
Maharashtra (Maharashtra)
Manipur (Manipur)
Meghalaya (Meghalaya)
Mizoram (Mizoram)
Nagaland (Nagaland)
Odisha (Odisha)
Puducherry (Puducherry)
Punjab (Punjab)
Rajasthan (Rajasthan)
Sikkim (Sikkim)
Tamil Nadu (Tamil Nadu)
Telangana (Telangana)
Tripura (Tripura)
Uttar Pradesh (Uttar Pradesh)
Uttarakhand (Uttarakhand)
West Bengal (West Bengal)
Indonesia (IDS)
Borneo (Kalimantan) (Borneo)
Java (Java)
Lesser Sunda Islands (Lesser Sunda Ils.)
Maluku Islands (Maluku Isl.)
Papua (Papua)
Riau Islands (Riau Islands)
Sulawesi (Celebes) (Sulawesi)
Sumatra (Sumatra)
Timor (Timor)
Iran (IN)
Iraq (IQ)
Ireland (IR)
Israel (IS)
Italy (IT)
Sardegna (Sardegna)
Sicily (Sicily)
Japan (JA)
Hokkaido (Hokkaido)
Honshu (Honshu)
Kyushu (Kyushu)
Ryukyu Islands (Ryukyu Islands)
Shikoku (Shikoku)
Jordan (JO)
Kashmir (KA)
Kazakhstan (KZ)
Kenya (KEN)
Kerguelen Islands (Kerguelen Islands)
Kuwait (KU)
Kyrgyzstan (KI)
Laos (LAO)
Latvia (LA)
Lebanon (LE)
Lesotho (LSO)
Libya (LB)
Liechtenstein (LS)
Lithuania (LT)
Luxenbourg (LU)
Macedonia (MC)
Madagascar (MAD)
Madeira Arch. (MR)
Malawi (MLW)
Malaysia (MLS)
Sabah (Borneo) (Sabah (Borneo))
Sarawak (Borneo) (Sarawak (Borneo))
Maldives (MLD)
Malta (MA)
Mauritania (MAU)
Mauritius (MRC)
Melanesia (Melanesia)
Bismarck Archipelago (Bismarck Archipelago)
Fiji (Fiji)
New Caledonia (New Caledonia)
Papua New Guinea (Papua New Guinea)
Solomon Islands (Solomon Islands)
Vanuatu ils (Vanuatu ils)
Mexico (MEX)
Aguascalientes (Aguascalientes)
Baja California (Baja California)
Baja California Sur (Baja California Sur)
Campeche (Campeche)
Chiapas (Chiapas)
Chihuahua (Chihuahua)
Coahuila (Coahuila)
Colima (Colima)
Durango (Durango)
Federal District (Federal District)
Guanajuato (Guanajuato)
Guerrero (Guerrero)
Hidalgo (Hidalgo)
Jalisco (Jalisco)
Mexico State (Mexico State)
Michoacan (Michoacan)
Morelos (Morelos)
Nayarit (Nayarit)
Nuevo Leon (Nuevo Leon)
Oaxaca (Oaxaca)
Puebla (Puebla)
Queretaro (Queretaro)
Quintana Roo (Quintana Roo)
San Luis Potosi (San Luis Potosi)
Sinaloa (Sinaloa)
Sonora (Sonora)
Tabasco (Tabasco)
Tamaulipas (Tamaulipas)
Tlaxcala (Tlaxcala)
Veracruz (Veracruz)
Yucatan (Yucatan)
Zacatecas (Zacatecas)
Micronesia (MIC)
Caroline islands (Caroline islands)
Guam (Guam)
Kiribati (Kiribati)
Marshall ils (Marshall ils)
North Marians isl. (North Marians isl.)
Palau archipelago (Palau archipelago)
Moldavia (MD)
Mongolia (MG)
Montenegro (ME)
Mordor (RU)
Altai Republic and Altaisky Province (Altai Republic and Altaisky Province)
Amurskaya Area (Amurskaya Area)
Bashkiria, E of Chelyabinsk regs. (S Ural mts. Bashkiria, E of Chelyabinsk regs.)
Buryat Republic (Buryat Republic)
Chukot autonomous region (Chukotka)
East Sayan Mountains (East Sayan Mountains)
Ingushetia, Chechnya, Daghestan (E Caucas) (Ingushetia, Chechnya, Daghestan (E Caucas))
Irkutsk Area (Irkutsk Area)
Kabardino-Balkaria, Ossetia (C Caucas) (Kabardino-Balkaria, Ossetia (C Caucas))
Kaliningrad, Briansk, Smolensk, Tula, Orlov, Kaluga, Moskow, Riaz (C of Russian plains)
Kamchatskaya Area (Kamchatka)
Karachaevo-Cherkessia (W Caucas) (Karachaevo-Cherkessia)
Karelia, Leningrad reg., Pskov reg., Kostroma reg., Perm reg., Komi, Pechora plains. (N of Russian plains)
Khabarovsky Province (Khabarovsky Province)
Khamar-Daban Mountains (Khamar-Daban Mountains (E Siberia))
Komi, Yamalo-Nenetsk, Hanty-Mansi (N Ural mts.)
Krasnodar reg. (W Caucas)
Krasnoyarsky Province (Krasnoyarsk)
Kuriles Islands (Kuriles Islands)
Magadanskaya Area (Magadan)
Omsk, S of Tyumen, W Novosibirsk regs. (S of W Siberia)
Perm, Sverdlovsk regs. (C Ural mts)
Primorsky District (Primorie) (Maritime Prov)
Sakhalin Island (Sakhalin Island)
Tuva Republic (Tuva)
Voronezh, Rostov, Saratov, Kalmykia, Volgograd, Astrahan regs. (S of Russian plains)
West Sayan Mountains (West Sayan)
Yakut-Saha Republic (Yakut-Saha Republic)
Zabaykalsky Krai (ex Chita reg.) (Zabaykalsky Krai)
plains of Krasnodar reg., S Rostov reg., Stavropol reg. (Ciscaucas reg. (plains of Krasnodar reg., S Rostov)
taiga from Ural mts to Yenisei riv. (Middle of W Siberia)
tundra of Tyumen, Yamalo-Nenetsk regs. (N of W Siberia)
Morocco (MO)
Mozambique (MZB)
Myanmar (Burma) (BRM)
Namibia (NAM)
Nepal (NP)
Netherlands (NL)
New Zealand (NZL)
Antipodes Islands (Antipodes Islands)
Auckland Islands (Auckland Islands)
Campbell Island (Campbell Island)
Chatham Islands (Chatham Islands)
Nicaragua (NCG)
North Korea (NC)
Norway (NR)
Svalbard (Spitzbergen) (SB)
Oman (OM)
Pakistan (PA)
Panama (PNM)
Paraguay (PRG)
Peru (PER)
Philippines (PLP)
Basilan (Basilan)
Cebu (Cebu)
Guimaras (Guimaras)
Leyte (Leyte)
Luzon (Luzon)
Marinduque (Marinduque)
Masbate (Masbate)
Mindanao (Mindanao)
Mindoro (Mindoro)
Negros (Negros)
Palawan (Palawan)
Panay (Panay)
Polillo Islands (Polillo Islands)
Romblon (Romblon)
Samar (Samar)
Sulu Archipelago (Sulu Archipelago)
Poland (PL)
Polynesia (POL)
Cook Islands (Cook Islands)
Hawaiian Islands (HAW)
Marquesas Islands (Marquesas Islands)
Niue (Niue)
Norfolk ils (Norfolk ils)
Pitcairn ils (Pitcairn ils)
Rapa Nui (Easter Island) (Rapa Nui)
Samoa (Samoa)
Society Islands (Society Islands)
Tahiti (Tahiti)
Tokelau (TKL)
Tonga (Tonga)
Tuvalu ils (Tuvalu ils)
Portugal (PT)
Qatar (QA)
Republic of South Africa (RSA)
Republic of the Congo (CG)
Reunion isl. (Reunion isl.)
Rodriguez isls. (ROD)
Romania (RO)
Rwanda (RWA)
Santa Helena isl (SH)
Sao Tome e Principe (STM)
Saudi Arabia (SA)
Serbia (SRB)
Seychelles (SEY)
Singapore (SGP)
Slovakia (SK)
Slovenia (SL)
Somalia (SOM)
South Georgia and the South Sandwich Islands (SGSSI)
South Korea (SC)
Spain (SP)
Balearic Islands (Balearic Islands)
Sri Lanka (SRL)
St. Paul isls (STP)
Sudan (SUD)
Surinam (SUR)
Swaziland (SWZ)
Sweden (SV)
Switzerland (SZ)
Syria (SY)
Tadzhikistan (TD)
Taiwan (Formosa) (Taiwan)
Tanzania (TAN)
Zanzibar isl. (Zanzibar isl.)
Thailand (TAI)
Tunisia (TU)
Turkey (TR)
Turkmenistan (TM)
Uganda (UGD)
Ukraine (UK)
Crimea (Crimea)
United States (USA)
Alabama (Alabama)
Alaska (Alaska)
Arizona (Arizona)
Arkansas (Arkansas)
California (California)
Colorado (Colorado)
Connecticut (Connecticut)
Delaware (Delaware)
Florida (Florida)
Georgia (Georgia)
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Illinois (Illinois)
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Iowa (Iowa)
Kansas (Kansas)
Kentucky (Kentucky)
Louisiana (Louisiana)
Maine (Maine)
Maryland (Maryland)
Massachusetts (Massachusetts)
Michigan (Michigan)
Minnesota (Minnesota)
Mississippi (Mississippi)
Missouri (Missouri)
Montana (Montana)
Nebraska (Nebraska)
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North Dakota (North Dakota)
Ohio (Ohio)
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Pennsylvania (Pennsylvania)
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South Dakota (South Dakota )
Tennessee (Tennessee)
Texas (Texas)
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Virginia (Virginia)
Washington (Washington)
West Virginia (West Virginia)
Wisconsin (Wisconsin)
Wyoming (Wyoming)
Uruguay (URG)
Uzbekistan (UZ)
Venezuela (VNZ)
Vietnam (VTN)
West Africa (WAF)
Benin (Benin)
Burkina Faso (Burkina Faso)
Côte d'Ivoire (Côte d'Ivoire)
Gambia (Gambia)
Ghana (Ghana)
Guinea (Guinea)
Guinea Bissau (Guinea Bissau)
Liberia (Liberia)
Mali (Mali)
Niger (Niger)
Nigeria (Nigeria)
Senegal (Senegal)
Sierra Leone (Sierra Leone)
Togo (Togo)
Yemen (YE)
Socotra (Socotra)
Zambia (ZMB)
Zimbabwe (ZIM)
Chlaenius (Chlaenioctenus) aodai Liu & Liang, 2013
Holotype
Male adult
(NHMB), N. Vietnam (Tonkin), Tamdao, 12−24.5.1989, Pacholatko leg. / Freiwlliger, Museumsverein, Basel, 1989. Image from Liu & Liang, 2013
Copyright © - - -
aodai
taxon page
(1 photos)
Chlaenius (Chlaenioctenus) bilyi Kirschenhofer, 2009
Holotype
Female adult
Copyright ©
Erich Kirschenhofer
bilyi
taxon page
(1 photos)
Chlaenius (Chlaenioctenus) bodhidharma Liu & Liang, 2013
Holotype Paratype
Male+Female adult
Holotype male (NHMB), “Darjeeling D., Ch. J. Rai’’ / “Alghera, 2900m, 25.IV.1982. Image from Liu & Liang, 2013
Copyright © - - -
bodhidharma
taxon page
(1 photos)
Chlaenius (Chlaenioctenus) cheni Liu & Liang, 2013
Holotype Paratype
Male+Female adult
Holotype male (SCAU), China, Guangdong Prov., Chebaling Nature Reserve,1989.3.21, Chen Shoujian leg. Image from Liu & Liang, 2013
Copyright © - - -
cheni
taxon page
(1 photos)
Chlaenius (Chlaenioctenus) chiangdaoensis Kirschenhofer, 2020
Holotype
Male adult
Thailand N., 2018 \ Chiang Dao, 8.5.\ Wiang Haeng env.\ M. Snížek lgt. (in coll. Martin Häckel, Prag). © Harald Schillhammer
Copyright © - - -
chiangdaoensis
taxon page
(1 photos)
Chlaenius (Chlaenioctenus) eneides Bates, 1892
Lectotype Paralectotypes
Male+Female adult
Lectotype male (MNHN), “Assam plains”/ “Ex-Musaeo, H.W. Bates, 1892”/ “Museum Paris, coll. R. Oberthür, 1952”/ “Chlaenius eneides Bates”. Paralectotype female (MNHN), “Assam plains’’ / “Ex-Musaeo, H.W. Bates, 1892’’ / “Museum Paris, coll. R. Oberthür, 1952”. Image from Liu & Liang, 2013
Copyright © - - -
eneides
taxon page
(1 photos)
Chlaenius (Chlaenioctenus) freyi Jedlicka, 1960
Female adult
China, North Guangdong pro., Nanling Natural Reserve, 2006.5.1-7, night, Xu Chunguo leg;
16.7mm; Compare with the Holotype
Photo by Liuye
Copyright ©
Ye Liu
Chlaenius (Chlaenioctenus) freyi Jedlicka, 1960
Female adult
Guangdong Prov., Nanling Natural Reserve, 2006.5.1–7, night,
Chunguo Xu leg.
Copyright ©
Ye Liu
Chlaenius (Chlaenioctenus) freyi Jedlicka, 1960
Holotype
Male adult
NHMB<br /> Photo by Yuxia Yang
Copyright ©
- - -
freyi
taxon page
(3 photos)
Chlaenius (Ocybatus) langsonensis Kirschenhofer, 2008
Holotype
Male adult
Tonkin, Montes Mauson, April, Mai 2-3000' H. Fruhstorfer (MNHUB) (Provinz Langson).
Copyright ©
Erich Kirschenhofer
langsonensis
taxon page
(1 photos)
Chlaenius (Chlaenioctenus) li Liu & Liang, 2013
Holotype Paratype
Male+Female adult
Holotype male (IZCAS), “China, Hainan Prov., Diaoluoshan Nature Reserve, on tree trunk, N18.72510, E109.86861’’ / “921m, 2012.4.18 N, Liu Y. & Shi H.L. leg., Institute of Zoology. Image from Liu & Liang, 2013
Copyright © - - -
li
taxon page
(1 photos)
Chlaenius (Chlaenioctenus) pectinipes Bates, 1892
Holotype
Male adult
Carin Ghecu, 1300–1400m, L. Fea, V. 1888, L. Fea”/ “Typus”/“pectinipes Bates”/“Chlaenius (Chlaenioctenus) pectinipes Bates”/ “Chlaenius (Chlaenioctenus) pectinipes Bates, Typus!”/ “Mus. Civ. Genova”/ “Holotypus male, Chlaenius (Chlaenioctenus) pectinipes Bates, 1892” [red label]. (MCSN).
Copyright ©
Ye Liu
pectinipes
taxon page
(1 photos)
Chlaenius (Chlaenioctenus) piligenys Liu & Liang, 2010
Paratype
Female adult
Copyright ©
Ye Liu
Chlaenius (Chlaenioctenus) piligenys Liu & Liang, 2010
Holotype
Male adult
China, Taiwan Prov., Pingtung, Dahanshan, 2008.IV.6, night, Wensin Lin
leg. (NMNS)
Copyright ©
Ye Liu
piligenys
taxon page
(2 photos)
Chlaenius (Chlaenioctenus) rotundithorax Liu & Kavanaugh, 2010
Holotype
Male adult
“Yunnan Prov., Tengchong, Jietou, Datang, Dahelingganjiao, 25.73947°N,98.69630°E’’/ “2010m, 2006.5.15 day, Kavanaugh D., Brett R. California Academy & IOZ, Chinese Acad.Sci.”/ “HOLOTYPE, Chlaenius (Chlaenioctenus) rotundithorax n. sp., designated by Y. Liu & D. Kavanaugh 2009”
Copyright ©
Ye Liu
rotundithorax
taxon page
(1 photos)
Chlaenius (Chlaenioctenus) schillhammeri Kirschenhofer, 2003
Holotype
Female adult
Copyright ©
Schillhammer
schillhammeri
taxon page
(1 photos)
Chlaenius (Chlaenioctenus) sehnali Kirschenhofer, 2008
Holotype
Female adult
S China, NE Guangxi, Xing'an co: Mao'er Mountain, Natur. Reserv. 493 m, V. 2007, N:25°51'11.4'', E: 110° 29' 15'' lgt. M. Häckel & R. Sehnal (CollSEHN).
Copyright ©
Erich Kirschenhofer
sehnali
taxon page
(1 photos)
Chlaenius (Chlaenioctenus) tingaudi Lassalle, 2015
Holotype
Male adult
Copyright ©
Bernard Lassalle
Chlaenius (Chlaenioctenus) cf tingaudi Lassalle, 2015
Male adult
NE Laos, Mt. Phu Phan, 2060m, VI.2018. (DUBC). det. Anichtchenko A. 2019
Copyright ©
Alexander Anichtchenko
tingaudi
taxon page
(2 photos)
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