Carabus (Coptolabrus) augustus ignigena Hauser, 1914
Male adult
Chine, CE Hunan, Changde 07.2004 h=700m Local people<br /> Det.I.A.Solodovnikov 2006
Copyright © Igor Solodovnikov

Carabus (Coptolabrus) augustus cristianofonti Deuve & Font, 1998
Female adult
China, Guizhou Prov., Sinan, 14.V.2009. L= 41mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) augustus chalingxian Kleinfeld, 1998 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Coptolabrus) augustus augustus Bates, 1888
Male adult
SE-China, Jiangxi prov., Yudu County, Ganzhou city, Yun Fengzang h=1000m, 5-20.V.2011
L=43 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) augustus augustus Bates, 1888 Lectotype
Male adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC105 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec105
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Carabus (Coptolabrus) augustus ignigena Hauser, 1914 (as lautus)
Male+Female adult
Mt.Yingpan Shan (Qiubei Xian, Yunnan) CHINE, VII-2008
Copyright © Alain Mollard

Carabus (Coptolabrus) augustus tongrenensis Deuve & Li, 2006
Male adult
Tongren Xian (Jiangkou, Guizhou) CHINE
Copyright © Alain Mollard

Carabus (Coptolabrus) augustus haiyangicus Deuve, 2012
Male+Female adult
Xing’an (Guilin, Guangxi) CHINE
Copyright © Alain Mollard

Carabus (Coptolabrus) augustus mugepoensis Deuve & Li in Deuve, 2009
Male+Female adult
Mt.Gupo Shan (Zhongshan, Guangxi) CHINE
Copyright © Alain Mollard

Carabus (Coptolabrus) augustus divus Roeschke, 1908
Male+Female adult
Pingxiang (Zhongshan, Jiangxi) CHINE
Copyright © Alain Mollard

Carabus (Coptolabrus) elysii elysii Thomson, 1856
Male+Female adult
S. Shennongjia, Fangxian county, Hubei, CHINE
1-11.06.2007
Copyright © Alexandr Safronov

Carabus (Coptolabrus) elysii rothschildi Born, 1899 Holotype
Female adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC123 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec123
Copyright © - - -

Carabus (Coptolabrus) elysii rothschildi Born, 1899
Label
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC123 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec123
Copyright © - - -

Carabus (Coptolabrus) elysii wangguofeni Rapuzzi, 2016 Holotype
Male adult
China, Anhui province, Taihu, Wangling vill., South slope of Mt. Dabieshan, 400 m, 10/30.IV.2015, (30º31'18'' N; 116º16'39'' E), Xihuangshun legit; preserved in Ivan Rapuzzi collection. Image from Rapuzzi, 2016
Copyright ©

Carabus (Coptolabrus) elysii xiaoganicus Deuve et Remond, 2016
Male adult
China, Hubei, Xiaogan, Mt Shuangfeng Shan, VII.2016 L=37 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) elysii xiaoganicus Deuve et Remond, 2016
Endophallus
China, Hubei, Xiaogan, Mt Shuangfeng Shan,
Copyright © Ruslan Panin

Carabus (Coptolabrus) elysii xiaoganicus Deuve et Remond, 2016
Male+Female adult
Shuangfeng Shan (Xiaogan, Hubei) CHINE VI-2016
Copyright © Alain Mollard

Carabus (Coptolabrus) elysii xiaoganicus Deuve et Remond, 2016
Male+Female adult
Shuangfeng Shan (Xiaogan, Hubei) CHINE VI-2016
Copyright © Alain Mollard

Carabus (Coptolabrus) elysii amucronatus Deuve, 2010
Male adult
Zhongtiao Shan (Shanxi) CHINE VII-2002
Copyright © Alain Mollard

Carabus (Coptolabrus) elysii viridicyaneus Lassalle, 2016 Paratype
Male adult

Copyright © Bernard Lassalle

Carabus (Coptolabrus) formosus brockmanni (as nili Deuve, 2005 PT) Paratype
Male+Female adult
C China, Shaanxi,
Liupan Mts., Guan Shan Mt. h=800 m, , 34:50N/106:48 E, Team V.Siniaev leg.
female, 38 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Coptolabrus) formosus sunpanensis Semenov, 1898
Male adult

Copyright © Alexander Anichtchenko

Carabus (Coptolabrus) formosus grumorum Semenov, 1898
Female adult
female, 30mm
Copyright © Boleslav Brezina

Carabus (Coptolabrus) formosus grumorum Semenov, 1898
Male adult
male, 24mm
Copyright © Boleslav Brezina

Carabus (Coptolabrus) formosus bousqueti Deuve, 1993
Male adult
C-China, S-Gansu prov., Wenxian County, Mt.Guanzigou,1-14.V.2010, local coll. leg.
L=32 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) formosus brockmanni Kleinfeld & Schutze, 1997
Female adult
C-China, S-Shaanxi: Shangzhou City, Shahezi,1-14.VI.,2013, local coll. leg.
L=39 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) formosus formosus Semenov, 1887
Male+Female adult
Wu-Chang-xi, (Gansu) CHINE VI-1998
Copyright © Alain Mollard

Carabus (Coptolabrus) formosus subformosus Semenov, 1887
Male+Female adult
Wenxian, (Gansu) CHINE VIII-1992
Copyright © Alain Mollard

Carabus (Coptolabrus) formosus akane Imura & Yamaya, 1994 Paratype
Female adult
Tongchuan (Xi’an, Shaanxi) CHINE VI-1993
Copyright © Alain Mollard

Carabus (Coptolabrus) formosus purpureodiscoides Kleinfeld, 1999
Male+Female adult
Dalong Shan, Yunxian (Shiyan, Hubei) CHINE X-2010
Copyright © Alain Mollard

Carabus (Coptolabrus) fruhstorferi Roeschke, 1900
Male adult
Collection ground Izuhara cho, Nagasaki Pref. 35764
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Carabus (Coptolabrus) fruhstorferi Roeschke, 1900
Male adult
Japan, Tsushima Isl., 20.IX.2018, S. Shinichi leg., L=35 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) gemmifer pseudoformosus Deuve, 2006
Female adult
China, Tibet AR, Qamdo, Gonjo County, Kè Rì Xiāng vill.(Kerri Township), h=3100m, VI.2012, A. Mollard leg. L=38 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) gemmifer mesites Fairmaire, 1897 Holotype
Female adult
"Siao-Lou-Lou-Chan, Chasseurs thibétains, 1898". Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC149 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec149
Copyright © MNHN

Carabus (Coptolabrus) gemmifer gemmifer Fairmaire, 1887 Holotype
Male adult
Yunnan (MNHN) http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec150
Copyright © MNHN

Carabus (Coptolabrus) gemmifer loczyi Csiki, 1927 Holotype
Male adult
Yunnan; reçu de Bang Haes le 23/IV/1912 comme yunnanus (MNHN) http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec124
Copyright © MNHN

Carabus (Coptolabrus) gemmifer mesites Fairmaire, 1897
Male adult
China, Shimian 1200m Shimian Yaan Sichuan VII.2017 (DUBC). det. Anichtchenko A. 2018
Copyright © Alexander Anichtchenko

Carabus (Coptolabrus) gemmifer gemmifer (as loczyi Csiki, 1927)
Female adult
S-China, W-Yunnan: Dali city, Mt Jizushan, VI.2018, L=34 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) gemmifer mesites Fairmaire, 1897
Female adult
China, Sichuan, Xichang City, Muli Co., Ke-Er-xiang 2-12.7.2009 (cAA) 32 mm
Copyright © Alexander Anichtchenko

Carabus (Coptolabrus) gemmifer rectipustulatus Rapuzzi, 2024 Holotype
Male adult
Sichuan, Yanyuan to Wodixiang; 2700 m asl; 16 Aug. 2022
Copyright © - - -

Carabus (Coptolabrus) ignimitella Bates, 1888
Female adult
China, Yunnan, Tali (error?) Image from: Mal, N., Drumont, A., Kerkhof, S. & P. Grootaert, 2012. Pictorial beetle collection of the Royal Belgian Institute of Natural Sciences.
Copyright © - - -

Carabus (Coptolabrus) ignimitella antaeus Hauser, 1914 (as guidongicus Cavazzuti & Ratti, 1998)
Female adult
China : Jianxi, Jing Gangshan City 10-18.vi.2007 - J. Li Leg. (Size : 42 mm) Coll. N. Mal
Copyright © Noël Mal

Carabus (Coptolabrus) ignimitella guangxicola Deuve, 1998
Male adult
S-China, Guangxi prov., Jingxiu, Mt. Dayao Shan, 1-17.V.2006, L=47 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) ignimitella ertli Born, 1910 Holotype
Male adult
Foothow (MNHN-EC-EC136)
Copyright © - - -

Carabus (Coptolabrus) ignimitella antaeus Hauser, 1914 (as guidongicus)
Female adult
China, E-Hunan, Zhuzhou, Yanling County, Wanyang Shan Mts, VI.2003 , L= 47 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) ignimitella angulicollis Hauser, 1911
Male+Female adult
Mt.Datian Ding (Baishi,Guangdong) CHINE, VII-2008 (F 52mm, M 46mm)
Copyright © Alain Mollard

Carabus (Coptolabrus) ignimitella angulicollis Hauser, 1911 (as mellianus Kolbe, 1914)
Male+Female adult
Tianjing Shan (Luyuan, Lecheng), Guangdong) CHINE, VII-2008 (m 48mm, fem 50mm)
Copyright © Alain Mollard

Carabus (Coptolabrus) ignimitella stillatus Cavazzuti & Ratti, 1998
Male+Female adult
Mt.Nanshan (Nanping, Fujian) CHINE VI-2001
Copyright © Alain Mollard

Carabus (Coptolabrus) ignimitella ignimitella Bates, 1888 (as nyukmingi Deuve, 1991)
Male+Female adult
Wuyi Shan (Fujian) CHINE VI-2001
Copyright © Alain Mollard

Carabus (Coptolabrus) ignimitella ignimitella Bates, 1888
Male+Female adult
Luozhou (Fuzhou, Fujian) CHINE VI-2001
Copyright © Alain Mollard

Carabus (Coptolabrus) ishizukai Deuve & Ohshima, 1989
Male adult
Chine, E.Sichuan, Wushan 07,2003 h=800m, Local people
Det.I.A.Solodovnikov 2006
Copyright © Igor Solodovnikov

Carabus (Coptolabrus) ishizukai Deuve & Ohshima, 1989
Male adult
Chine, E.Sichuan, Wushan 07,2003 h=800m, Local people
Det.I.A.Solodovnikov 2006
Copyright © Igor Solodovnikov

Carabus (Coptolabrus) jankowskii jankowskii Oberthur, 1883
Male adult
N.Korea, Chagang Do, Mt. Langlin, 30.07.2005
Copyright © Anton Menstschikov

Carabus (Coptolabrus) jankowskii benxiensis Imura, 1991
Male+Female adult
Miaoyang, Shicheng, Fengcheng City,
Liaoning Province, CHINE
25-30.VI.2008
Copyright © Alexandr Safronov

Carabus (Coptolabrus) jankowskii fusanus Born, 1907
Female adult
South Korea, S Gyeongsang Prov., Busan, Mt Geumwonsan, 17.VI.2017, L=37 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) jankowskii fusanus (as obtusipennis)
Female adult
SC-Korea: Chiri-san Massif, Mt. Jirisan, 28.VI.2011, L=35 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) jankowskii byeoksanensis Rapuzzi, 2015 Holotype Paratype
Male+Female adult
R. of Korea, Jeollabuk-do Province, Buan-gun, Byeonsanbando, 100/200 m, 13/20.VII.2012, I. Rapuzzi and L. Caldon leg. (CIR). Image from Rapuzzi, 2015
Copyright © - - -

Damaster (Coptolabrus) jankowskii irwolsanensis Kim, 2009 Holotype
Male adult
Mt. Irwol, Yeongyang-gun, Gyeongsangbuk-do, 26, vii, 2007
Copyright © - - -

Damaster (Coptolabrus) jankowskii unmunsanensis Kim, 2009 Paratype
Female adult
Mt. Unmun, Cheongdo-gun, Gyeongsangbuk-do, 12, viii, 2004
Copyright © - - -

Damaster (Coptolabrus) jankowskii ganghwadoensis Kim, 2009 Paratype
Female adult
Mt. Goryeo, Ganghwa-gun, Gyeonggi-do, 16, viii, 2007
Copyright © - - -

Carabus (Coptolabrus) jankowskii chindoensis Kwon & Park, 1989
Male+Female adult
Chongue san (Chindo Isle, ) KOREE VII-1992
Copyright © Alain Mollard

Carabus (Coptolabrus) jankowskii quelpartianus Breuning, 1932
Male+Female adult
Hallasam, (Cheju-do Isle, ) KOREE VII-1990
Copyright © Alain Mollard

Carabus (Coptolabrus) kubani Deuve, 1990 Paratype
Male adult
Daju, (Yulongxue Shan, Yunnan) CHINE, VII-1990
Copyright © Alain Mollard

Carabus (Coptolabrus) lafossei tungchengensis Li, 1993 (as dabieshanus Imura, 1996)
Male adult
Chine, Hubei, Dabieshan: Yuexi,h=700m., VI.2004, Local people
Det. I.A.Solodovniov 2006
Copyright © Igor Solodovnikov

Carabus (Coptolabrus) lafossei tiantai Kleinfeld, 1997 Holotype
Female adult

Copyright © Frank Kleinfeld

Carabus (Coptolabrus) lafossei saturatus Hauser, 1913
Female adult
N Fujian, Jung Ling Shan, 18-24.7.96
female 53 mm
Copyright © Alexander Anichtchenko

Carabus (Coptolabrus) lafossei pseudocoelestis Kleinfeld, 1999 Holotype
Male adult

Copyright © Frank Kleinfeld

Carabus (Coptolabrus) lafossei montigradus Hauser, 1920
Male adult
Chine, Anhui, Huangshan, 07.2005 h=700m Local people
Det. I.A.Solodovnikov 2006
Copyright © Igor Solodovnikov

Carabus (Coptolabrus) ? lafossei coelestis Stewart, 1845
Female adult
CE-China, Shangai, 3.V.2007 L=34 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) lafossei saturatus Hauser, 1913
Male adult
SE-China, Jiangxi prov.,Yushan County, Mt Wangjing, VI.2012
40 мм
Copyright © Ruslan Panin

Carabus (Coptolabrus) lafossei buchi Hauser, 1913
Male adult
China Zhejiang Yuyao - Leg. Buch 1925
Copyright © Vittorino Monzini

Carabus (Coptolabrus) lafossei tungchengensis Li, 1993
Male+Female adult
Mt.Huang-Bai-Shan (Shangcheng, Henan) VI-2012
Copyright © Alain Mollard

Carabus (Coptolabrus) lafossei tungchengensis Li, 1993
Endophallus
China, Hubei, Shangzhi co., Shimenyan 8-11.6.2009 (cAA)
Copyright © Alexander Anichtchenko

Carabus (Coptolabrus) nankotaizanus Kano, 1932
Male adult
Taiwan,
Tatumi, h=200 m, 20.02.1971, D.Muting leg.
Det. I.A.Solodovniov 2006
Copyright © Igor Solodovnikov

Carabus (Coptolabrus) nankotaizanus Kano, 1932
Male adult
Taiwan,
Tatumi, h=200 m, 20.II.1971, D.Muting leg.
Det. I.A.Solodovniov 2006
Copyright © Igor Solodovnikov

Carabus (Coptolabrus) nankotaizanus Kano, 1932
Male adult
NE-Taiwan, Yilan County, V.2017, L=39 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) nankotaizanus Kano, 1932 (as shimonoyai Kurosawa & Kudo, 1980)
Male adult
Mt.Lala-Shan, (Fuxing,Taoyuan) TAIWAN (FORMOSE) II-1997
Copyright © Alain Mollard

Carabus (Coptolabrus) osawai berezovskiianus Deuve & Mourzine, 2003
Male adult
Image courtesy A. Gorodinsky
Copyright © - - -

Carabus (Coptolabrus) principalis Bates, 1889
Male+Female adult
Zi-Ding-Shan, h=1406 m, Guiyang County, Binzhou City, Hunan, CHINE
19-29.IV.2008
Copyright © Alexandr Safronov

Carabus (Coptolabrus) principalis Bates, 1889
Male adult
C.Chine, W Hubei, Changyang, VIII.2005,
Det. I.A.Solodovnikov 2006
Copyright © Igor Solodovnikov

Carabus (Coptolabrus) principalis Bates, 1889
Male adult
C.Chine, W Hubei, Changyang, 08.2005,
Det. I.A.Solodovnikov 2006
Copyright © Igor Solodovnikov

Carabus (Coptolabrus) principalis Bates, 1889
Female adult
C.Chine, W Hubei, Changyang, 08.2005,
Det. I.A.Solodovnikov 2006
Copyright © Igor Solodovnikov

Carabus (Coptolabrus) principalis Bates, 1889
Female adult
C Chine, W Hubei, Changyang, 08.2005,
Det. I.A.Solodovnikov 2006
Copyright © Igor Solodovnikov

Carabus (Coptolabrus) pustulifer pustulifer Lucas, 1869
Female adult
Chine, C.Sichuan, Ya'an, 07.2005 Local people
Copyright © Igor Solodovnikov

Carabus (Coptolabrus) pustulifer pustulifer Lucas, 1869
Male adult
Chine, C.Sichuan, Ya'an, 07.2005 Local people

Copyright © Igor Solodovnikov

Carabus (Coptolabrus) pustulifer mirificus Kraatz, 1894
Male adult
Chine, SW-Hubei, Enshi, 06.2005, Local people
Copyright © Igor Solodovnikov

Carabus (Coptolabrus) augustus adriaenssensi (as pseudoignigena Kleinfeld, 1996) Holotype
Female adult

Copyright © Frank Kleinfeld

Carabus (Coptolabrus) pustulifer shangzhouicus Deuve & Li, 2003
Male adult
C-Chine, SE-Shaanxi Province, Shahezi, Shangzhou City, 6-18.VII.2010<br /> L=43 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) pustulifer (as cyanelongatus Deuve, 2006)
Male adult
C-Chine, Tibet, Qamdo, Gonjo County, Ke Ri Xiang env., VI.2012, Alain Mollard leg.<br /> L=43 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) augustus adriaenssensi Lassalle & Prunier, 1993 Holotype
Male adult
Chine, Guangxi, Mts Miao Er Shan, 1800m, 5.92 (in coll B. Lassalle) ? pustulifer adriaenssensi Lassalle & Prunier, 1993
Copyright © Bernard Lassalle

Carabus (Coptolabrus) pustulifer pustulifer Lucas, 1869
Label
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC163 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec163
Copyright © - - -

Carabus (Coptolabrus) pustulifer elisabettianus Font, 1997
Male+Female adult
Huanjan Shan (Wuchuan, Guizhou) CHINE, VII-2008
Copyright © Alain Mollard

Carabus (Coptolabrus) pustulifer mirandior Deuve, 1993
Male+Female adult
Dayao (Yunnan) CHINE, VII-2008
Copyright © Alain Mollard

Carabus (Coptolabrus) smaragdinus mandschuricus Semenov, 1898
Male adult
Primorje, Lasovskii distr., Chandalaz mt., L.Voita leg. 19.VII.2008
Copyright © Anton Menstschikov

Carabus (Coptolabrus) smaragdinus longipennis Chaudoir, 1863
Male+Female adult

Copyright © Alexandr Safronov

Carabus (Coptolabrus) smaragdinus furumiellus Deuve, 1994
Male+Female adult
Hua-Bo-zi-Shan, Huanren County,Liaoning Province, CHINE, 20-30.VI.2008
Copyright ©

Carabus (Coptolabrus) smaragdinus dachangensis Lu & Müller, 2009
Female adult
NE-China, Liaoning Prov., Changhai County, Dachangshan Dao Isl.,12.VI.2011
L=40 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) smaragdinus liaodongensis (Li, 1992)
Male adult
NE-China, E-Liaoning: Anshan: Mt.Qian Shan, 9-14.VI.2011. L=39 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) smaragdinus sungkii J.K. Park & J. Park, 2013 Holotype Paratype
Male+Female adult
Holotype male: Is.Gureopdo (37°19′71.42″N, 125°98′72.62″E), Deokjeok archipelago, Gyeonggi-do, 19.viii.2010, J.K.Park; Paratype female: Is. Gureopdo, Deokjeok archipelago, Gyeonggi-do, J.K.Park. Image from original describtion J.K. Park & J. Park, 2013
Copyright ©

Carabus (Coptolabrus) smaragdinus dolichognathus Deuve, 1995
Male adult
NW-Korea, North Pyongan Province, Mt. Donggorim, 1-20.VI.2013, local coll. leg.<br /> L=37 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) smaragdinus shantungensis Born, 1910 Holotype
Male adult
Shandong, Qingdao reçu de Bang Haes le 23/IV/1912 . http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec122
Copyright © MNHN

Carabus (Coptolabrus) smaragdinus robinzoni Rapuzzi, 2010 Holotype
Male adult
Reyneke island (CR). Image from Rapuzzi, 2012
Copyright © - - -

Carabus (Coptolabrus) smaragdinus hongdoensis (Kwon & Lee, 1984) Holotype
Male adult
deposited in Kyungpook National University, Daegu, Republic of Korea, Scale bar = 5mm
Copyright © Jong-Bong Choi