Carabus (Neoplectes) apollo apollo Zolotarev, 1913
Male adult
Caucasus, Abchasia

Copyright © Dmitri Obydov

Carabus (Neoplectes) apollo phoebus Kurnakov, 1962
Male adult
Abchasia, Kodori, A. Zolotarew leg.
Photo by D.V. Obydov
from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Neoplectes) apollo tenebricosus Kurnakov, 1962
Male adult

Copyright © Safronov

Carabus (Neoplectes) apollo tenebricosus Kurnakov, 1962
Endophallus

Copyright © Alexandr Safronov

Carabus (Neoplectes) compressus rebellis Reitter, 1895
Male adult
Georgia, Kvemo-Svaneti, S slopes of Svanetskiy Mt.R., S slope of Latpari pass, h=2225m, 17.VI -13.VII.2017 leg. D.Fominykh L=24.0
Copyright © Kirill Makarov

Carabus (Neoplectes) compressus rebellis Reitter, 1895
Female adult
Georgia, Kvemo-Svaneti, S slopes of Svanetskiy Mt.R., S slope of Latpari pass, h=2225m, 17.VI -13.VII.2017 leg. D.Fominykh L=26.0 mm
Copyright © Kirill Makarov

Carabus (Neoplectes) zubovi Fominykh, Zamotajlov et Titarenko, 2018 Holotype
Male adult
S slopes of Svaneti Mt. Range near Lentekhi Town, S slopes of Goldash Mt., 42°49'N, 42°45'E, 2300–2800 m, 21.VI.2016, D. Fominykh, A. Zherebilo (ZISP).
Copyright © Kirill Makarov

Carabus (Tribax) lailensis stefaniianus Deuve, 2018 Holotype
Male adult
Géorgie, Basse-Svanétie, au nord de Lentekhi, versant sud de la chaîne de Svanétie, 1000 m., Th. Deuve et A. Matocq, juin 2016. (MNHN)
Copyright © Thierry Deuve

Carabus (Tribax) zubovi mananauriensis Deuve, 2022 Holotype
Male adult
Géorgie, Svanétie, 20 km à l’ouest de Lentekhi, 6 km à l’ouest de Mananaouri, 1350 m (A. Wřzecionko leg.), in coll. A. Wřzecionko, à Horni Sucha (République Tchèque).
Copyright © Thierry Deuve

Carabus (Neoplectes) compressus rebellis Reitter, 1895 Lectotype
Male adult
Image from Deuve, 2024
Copyright © Thierry Deuve

Carabus (Tribax) compressus pseudorebellis Deuve, 2024 Holotype
Male adult
Géorgie, Basse-Svanétie, 12 km au nord-est de Lentekhi, Mont Goldash, vallée de la Cholshoura, Durashi (« Dvashi »), env. 1150 mètres (W. Heinz, 1-4 juillet 2019), in coll. Muséum Stuttgart (Germany).
Copyright © Thierry Deuve

Carabus (Neoplectes) compressus zubovi Fominykh, Zamotajlov et Titarenko, 2018 Paratype
Male+Female adult
5-6: Paki Vill. environs; 7-8: Mannanauri Vill. environs. Scale bar 10.0 mm. Image from Fominykh, Zamotajlov et Titarenko, 2018
Copyright © - - -

Carabus (Neoplectes) compressus lailensis Belousov, 1992
Male adult
Carabus (Archiplectes) lailensis Belousov, 1992 (TOPOTYPUS) Georgia, Svaneti, S slopes of Svanetian Mts. rng., W slopes of Laila-lakheli Mt., upper of Skirili river, alpin zone, h = 3000 m, 27-30.VI.2014, leg. D. Fominykh, A. Zubov
Copyright © Igor Solodovnikov

Carabus (Neoplectes) compressus compressus Chaudoir, 1846 (? rebellis )
Male adult
Georgia, Caucasus, S of Sasashi vill., Lechkhumi Range N slope h=2430 m, alpine meadows, 242.759167, 42.973889, 20-21.VII.2025, I.Uspenskyi leg, L=26 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) daphnis askhicus Belousov, 1992 Paratype
Male adult
SC-Caucasus, W Georgia, valley of riv. Chkhorodukhu, h=1000-1200m, 14.V.-6.VII.1991, Zamotajlov A.S & Miroshnikov leg.
male, 29 mm
photo & collection Panin R.
Copyright © Ruslan Panin

Carabus (Neoplectes) daphnis askhicus Belousov, 1992 Paratype
Male+Female adult
W Georgia, valley of Chkhorodukhu riv., 1000-1200m
14.V-6.VII.1991
male 30mm, female 33mm
photo & collection Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) daphnis askhicus Belousov, 1992 Paratype
Male adult
Georgia, env. Martvilli, Askhi plateau
10-15.7.2001 A.Rubenian leg.
27 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) daphnis daphnis Kurnakov, 1962
Male adult
Georgia, Mingrelia, Lecheha vallis, Tehuri superior
Caucasus centr., 1200m, 27.6-7.7.1989 Retezar I. leg.
male, 28 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) daphnis daphnis Kurnakov, 1962
Endophallus
Georgia, Mingrelia, Lecheha vallis, Tehuri superior
Caucasus centr., 1200m, 27.6-7.7.1989 Retezar I. leg.
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) daphnis askhicus Belousov, 1992
Endophallus
Georgia, env. Martvilli, Askhi plateau
10-15.7.2001 A.Rubenian leg.
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) daphnis askhicus Belousov, 1992
Endophallus
W Georgia, valley of Chkhorodukhu riv., 1000-1200m
14.V-6.VII.1991
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) daphnis askhicus Belousov, 1992 Paratype
Endophallus

Copyright © Ruslan Panin

Carabus (Neoplectes) daphnis tsekuricus Belousov, 1992
Male adult
LEFT: Lakhashuri River valley; RIGHT: Tzikuri Mt. Scale bar 10.0 mm.
Copyright © Kirill Makarov

Carabus (Neoplectes) daphnis boulbenellus Deuve et Stéfani, 2017 Holotype
Male adult
Géorgie, Basse-Ratcha, à l’est de Nikortsminda, Skhartali, 1200-1300 m., Th. Deuve, P. Stéfani leg. (MNHN).
Copyright © Thierry Deuve

Carabus (Neoplectes) edithae exedithae Gottwald, 1985
Male adult
Karachaevo-Cherkessia, low cource of Daut river,
16-20.V.2009 Anichtchenko A. leg

Copyright © Alexander Anichtchenko

Carabus (Neoplectes) edithae exedithae Gottwald, 1985
Female adult
Karachaevo-Cherkessia, Zelenchukskii distr.,
near Bukovo vill., 8-13.VII.2007, S.Dementiev leg.

Copyright © Dmitri Obydov

Carabus (Neoplectes) edithae kabardensis Kurnakov, 1970
Female adult
Kabardino-Balkaria, Skalisty Mt Ridge, 1800 m,
20.VII.1996. Andreev leg.
Photo by D.V. Obydov

Copyright © Dmitri Obydov

Carabus (Archiplectes) edithae Reitter, 1893
Male adult

Copyright © Kirill Makarov

Carabus (Archiplectes) edithae Reitter, 1893
Habitat
Carabus (Archiplectes) edithae exedithae Gottwald, 1985
Karachaevo-Cherkessia, valley of Daut riv. 43.62' 42.09'
17-18.V.2009
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) edithae exedithae Gottwald, 1985
Male adult
NС-Caucasus, Karachaevo-Cherkessia, Abishir-Akhuba Mt.Range h=2250-2600 m, right bank Pkhiya river, 10-13.VI.2013, S. Pavluchuk leg.
L=25 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) edithae exedithae Gottwald, 1985
Endophallus
NС-Caucasus, Karachaevo-Cherkessia, right bank Pkhiya river, Abishir-Akhuba Mt.Range h=2250-2600 m,
Copyright © Ruslan Panin

Carabus (Neoplectes) edithae Reitter, 1893
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Carabus (Neoplectes) edithae miloslavskii Obydov, 2010 Holotype Paratype
Male+Female adult
Holotype: Caucasus centr., Archyz, zona subalp., 15.VI.1967, A. Bogachev leg; Paratype: KarachaiCherkessian Region, Zagedan Mt. Ridge, 2500 m., 6-7.VII.1994, Melnik leg.
Copyright © - - -

Carabus (Neoplectes) ibericus ibericus (as szekelyi Retezar, 2011)
Male+Female adult
Itrapula env. Image from Fominykh & Zamotajlov, 2016
Copyright © - - -

Carabus (Neoplectes) ibericus ibericus (as imereticus Retezar, 2011) Paratype
Endophallus
NC-Georgia, S-Caucasus: W slope Likhi Mt.Rng., 3 km W Rikoti Pass h~900 m,
Copyright © Ruslan Panin

Carabus (Neoplectes) titarenkoi kozlovantoni Fominykh, Zamotajlov & Titarenko, 2025 Paratype
Male adult
Georgia, Ratcha, right bank of river Gamrula, near Gomi vill., 1385 m, 3.V–12.VII.2018, pitfall traps, D. Fominykh, B. Dubinin, D. Safronov leg. Image from Fominykh, Zamotajlov & Titarenko, 2025
Copyright © - - -

Carabus (Neoplectes) titarenkoi kozlovantoni Fominykh, Zamotajlov & Titarenko, 2025 Paratype
Female adult
Georgia, Racha, Oni town env., near Ghari vill., btw rivers Rioni and Gharula., 870 m, 13.V–22.VI.2024, pitfall traps, D. Fominykh, A. Tatulyan, Ya. Talibov leg. Image from Fominykh, Zamotajlov & Titarenko, 2025
Copyright © - - -

Carabus (Neoplectes) szekelyi tatulyani Fominykh, Zamotajlov & Titarenko, 2025 Paratype
Male adult
Kontanto env. Image from Fominykh, Zamotajlov & Titarenko, 2025
Copyright © - - -

Carabus (Neoplectes) szekelyi tatulyani Fominykh, Zamotajlov & Titarenko, 2025 Paratype
Female adult
Kontanto env. Image from Fominykh, Zamotajlov & Titarenko, 2025
Copyright © - - -

Carabus (Neoplectes) szekelyi tatulyani Fominykh, Zamotajlov & Titarenko, 2025 Paratype
Male adult
Tli env. Image from Fominykh, Zamotajlov & Titarenko, 2025
Copyright © - - -

Carabus (Neoplectes) szekelyi tatulyani Fominykh, Zamotajlov & Titarenko, 2025 Paratype
Female adult
Tli env. Image from Fominykh, Zamotajlov & Titarenko, 2025
Copyright © - - -

Carabus (Neoplectes) ibericus pseudoszekelyi Fominykh, Zamotajlov & Titarenko, 2025 Paratype
Female adult
Shkmeri env. Image from Fominykh, Zamotajlov & Titarenko, 2025
Copyright © - - -

Carabus (Neoplectes) ibericus Fischer, 1823
Endophallus
Image from Fominykh, Zamotajlov & Titarenko, 2025 with modifications
Copyright © - - -

Carabus (Archiplectes) jason Semenov, 1898
Male adult
Caucasus, W Georgia, Svanetia, upper reaches of riv. Shoudidi, 1700-2100m, 20-21.V.1986
leg. A. Zamotajlov
Copyright © Sunvas

Carabus (Neoplectes) jason Semenov, 1898
Male adult
Abkhasia, up river Shaudidi, 21.V - 2.VII.1986, Zamotailov A. leg.
Photo by D.V. Obydov

Copyright © Dmitri Obydov

Carabus (Neoplectes) jason Semenov, 1898
Male adult
Abhazia, Tchkhaltinskiy (Abkhazskiy) Mt.R., 20-26.VII.2022 leg. D. Fominykh
Copyright © Igor Solodovnikov

Carabus (Neoplectes) jason Semenov, 1898
Endophallus

Copyright © Igor Solodovnikov

Carabus (Neoplectes) khipstaensis Solodovnikov, 2015 Paratype
Male adult
Абхазия, Гудаутский р-н, Ю скл. Бзыбского хр., водораздельный холм в истоках р. Хипста, 43°17'52.03" N / 40°40'8.72" E, h = 1435 m, Линия № 64-4, буковый спелый лес с папоротниками и ежевикой, 09-13.08.2014, leg. И.А. Солодовников, Е.В. Татун
Copyright © Igor Solodovnikov

Carabus (Neoplectes) khipstaensis Solodovnikov, 2015 Paratype
Male adult
Абхазия, Гудаутский р-н, Ю скл. Бзыбского хр., водораздельный холм в истоках р. Хипста, 43°17'52.03" N / 40°40'8.72" E, h = 1435 m, Линия № 64-4, буковый спелый лес с папоротниками и ежевикой, 09-13.08.2014, leg. И.А. Солодовников, Е.В. Татун
Copyright © Igor Solodovnikov

Carabus (Neoplectes) khipstaensis Solodovnikov, 2015 Paratype
Female adult
Абхазия, Гудаутский р-н, Ю скл. Бзыбского хр., водораздельный холм в истоках р. Хипста, 43°17'52.03" N / 40°40'8.72" E, h = 1435 m, Линия № 64-4, буковый спелый лес с папоротниками и ежевикой, 09-13.08.2014, leg. И.А. Солодовников, Е.В. Татун
Copyright © Igor Solodovnikov

Carabus (Neoplectes) khipstaensis Solodovnikov, 2015 Paratype
Female adult
Абхазия, Гудаутский р-н, Ю скл. Бзыбского хр., водораздельный холм в истоках р. Хипста, 43°17'52.03" N / 40°40'8.72" E, h = 1435 m, Линия № 64-4, буковый спелый лес с папоротниками и ежевикой, 09-13.08.2014, leg. И.А. Солодовников, Е.В. Татун
Copyright © Igor Solodovnikov

Carabus (Neoplectes) polychrous khipstaensis Solodovnikov, 2015
Endophallus

Copyright © Igor Solodovnikov

Carabus (Neoplectes) koltzei faunulus Kurnakov, 1970
Female adult
Abchasia, Akarmara, 6500 f, A. Zolotarew leg.
Photo by D.V. Obydov

Copyright © Dmitri Obydov

Carabus (Neoplectes) koltzei faunus Kurnakov, 1970
Male adult
Abkhazia, Kadorsky Mt Ridge, 1000 m, Chlou, 22.V.1993, A. Evstigneev leg.
Photo by D.V. Obydov

Copyright © Dmitri Obydov

Carabus (Neoplectes) koltzei kobachidzei Kurnakov, 1970
Female adult
Caucasus, E Abkhasia, SW slopes of Akiba mt. rng., 1200m, 7.V.-2.VII.1991<br /> Zamotajlov & Miroshnikov leg.
Copyright © Sunvas

Carabus (Neoplectes) koltzei kobachidzei Kurnakov, 1970
Female adult
Caucasus, E Abkhasia, SW slopes of Akiba mt. rng., 1200m, 7.V.-2.VII.1991
Zamotajlov & Miroshnikov leg.
Copyright © Sunvas

Carabus (Neoplectes) koltzei kobachidzei Kurnakov, 1970
Male+Female adult
Caucasus, E Abkhasia, SW slope of Akiba mt.rng., 1200m, 7.V.-2.VII.1991 Zamotajlov & Miroshnikov leg.
male 37mm, female 38mm
photo & collection Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Archiplectes) koltzei Rost, 1889
Male adult
Abkhasia, Habyu 3.7.2002
Ivliev P. leg.
32 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) koltzei koltzei Rost, 1889
Male adult
Abkhasia, Tsebeldy, 23.V.1986, A.Zamotailov leg.
Photo by D.V. Obydov

Copyright © Dmitri Obydov

Carabus (Neoplectes) koltzei lennoni Gottwald, 1985
Male adult
Abkhasia, Amtkel lake, 22.V - 4.VII.1986, A.Zamotailov leg.
Photo by D.V. Obydov
from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Neoplectes) komarowi komarowi Reitter, 1882
Female adult
Centr. Caucasus, Svanetia, coll. A. Zolotarew
Photo by D.V. Obydov
from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Neoplectes) komarowi komarowi Reitter, 1882
Female adult
NW Georgia, Upper Svaneti, Mestia distr., Lakhamula env. h~1250 m, 19.VI.2014, V.Sima leg. L=33 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) komarowi mestiaicus Deuve, 2016 Holotype
Female adult
Géorgie, Haute Svanétie, 10 km au nord de Mestia, vers les lacs de Koruldi, env. 2500 mètres, in coll. A. Wřzecionko, à Horní Sucha (République Tchèque).
Copyright © Thierry Deuve

Carabus (Neoplectes) komarowi komarowi Reitter, 1882 Lectotype Paralectotypes
Male+Female adult
LT - female «Kaukasus Swanetien Leder Reitter » , «Komarowi m.», «Roeschke C.», « MA.INS.COLE.194.1», «Lectotype», «Carabus komarowi Reitter, Lectotype, Th. Deuve des. 2024», ex coll. Vogt, in coll. Naturalis Biodiversity Center, Leyde (Pays-Bas). Image from Deuve, 2024
Copyright © Thierry Deuve

Carabus (Neoplectes) komarowi komarowi Reitter, 1882 Lectotype Paralectotypes
Female adult
LT, left, female: «Kaukas Leder», «Komarowi v. mutabilis m.», «Roeschke C. », « Collectie C. & O. Vogt, Acq. 1960», «Plectes Komarowi mutabilis Reitter, 1896, ZMAN type COLE.0264.1», «Lectotype», «Carabus komarowi mutabilis Reitter, Lectotype, Th. Deuve des. 2024 » , ex coll. Vogt, in coll. Naturalis Biodiversity Center, Leyde (Pays-Bas). Image from Deuve, 2024
Copyright © Thierry Deuve

Carabus (Tribax) perditus Deuve, 2024 Holotype
Male adult
MNHN
Copyright © Thierry Deuve

Carabus (Tibax) komarowi antoni Deuve, 2024 Holotype
Male adult
«Kaukasus Swanetien Leder Reitter», ex coll. M. Maindron, ex coll. G. Babault, in coll. Muséum National d’Histoire Naturelle, Paris (France).
Copyright © Thierry Deuve

Carabus (Neoplectes) kratkyi felicitanus Reitter, 1893 (as porcellus)
Male adult
NW Caucasus, Resp. Adygea, Mt.Tybga, 28.VI.1997
male,20mm
Copyright © Ruslan Panin

Carabus (Neoplectes) kratkyi felicitanus Reitter, 1893 (as porcellus)
Male adult
Prov. Kuban, mt. Tybga, 1300 m, VI.1913, Shaposhnikov leg.

Copyright © Dmitri Obydov

Carabus (Neoplectes) kratkyi felicitanus Reitter, 1893
Female adult
Rusia, Caucas, Dzhuga mt., 2200 m, 24.VI. - 22.VIII.1986, Zamotajlov A. leg.

Copyright © Dmitri Obydov

Carabus (Neoplectes) kratkyi Ganglbauer, 1890
Male+Female adult
W Caucas, 1900m, Khatsavitaya Mt., 4.VI-13.VII.1994
A.Miroshnikov leg.
male 25mm, female 26mm
photo & collection Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) kratkyi Ganglbauer, 1890
Male adult
Russia, NW-Caucasus: <br /> Krasnodar reg., Mostovskoy distr., Khatsavitaja Mt. h=1900m, 4.VI.-13.VII.1994, A. Miroshnikov leg.<br /> male, 26mm<br /> photo Panin R./det.Solodovnikov I.A.
Copyright © Ruslan Panin

Carabus (Neoplectes) kratkyi Ganglbauer, 1890
Male+Female adult
W Caucas Mts., 2500m, Mt. Yatyrgvarta,
25.VI-12.VIII.1992 Shchurov V. leg.
male 20mm, female 21mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890
Female adult
NW-Caucasus, Russia,
Karacaj-Cherkessija, Abishira-Akhuba range h=2650m, upper Kyafar-Agur Riv., 4-9.VIII.2009, Solodovnikov I.A. leg.
female, 23 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890
Male adult
Caucasus, Kuban, Meyer-Darcis-Patria

Copyright © Dmitri Obydov

Carabus (Neoplectes) kratkyi solodovnikovi Zamotajlov, 1992
Male adult
W. Caucasus, M. Bambak Mt., VI.1931, Vvedensky leg.
from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890
Endophallus
Endophallus
Copyright © Ruslan Panin

Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
Female adult
female, 49mm
Copyright © Boleslav Brezina

Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
Male adult
male, 43mm
Copyright © Boleslav Brezina

Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
Female adult

Copyright © Kirill Makarov

Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
Female adult
W Georgia, Martvili Distr., 5 km N T'aleri, 11-18.VI.2014, V. Sima leg., L=50 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) martviliensis mzekalae Fominykh, Zamotajlov et Titarenko, 2019 Paratype
Male+Female adult
Imereti, left bank of river Semi near Melouri, 475 m, 19.V–12.VII.2019, D. Fominykh, A. Zubov, B. Dubinin & D. Safronov leg.
Copyright © Kirill Makarov

Carabus (Neoplectes) martviliensis mzekalae Fominykh, Zamotajlov et Titarenko, 2019 Paratype
Male+Female adult
Imereti, left bank of river Tskhenistskali near Dzedzileti, 620 m, 16.V–13.VII.2019, D. Fominykh, A. Zubov, B. Dubinin & D. Safronov leg.
Copyright © Kirill Makarov

Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
Male+Female adult
Imereti, S slopes of Askhi plateau near Zeda-Gordi, NE slope of Mt. Gulkuna, 980 m, 20.V–14.VII.2019, D. Fominykh & A. Zubov leg.
Copyright © Kirill Makarov

Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
Male+Female adult
Samegrelo, Doberazeni vic., left bank of river Tekhuri, 500 m, 16.IV–31.V.2017, D. Fominykh leg.
Copyright © Kirill Makarov

Carabus (Neoplectes) martviliensis Retezar & Djavelidze, 1992
Endophallus
Zeda Gordi env., 980 m and Doberazeni env., 600 m.
Copyright © Kirill Makarov

Carabus (Neoplectes) mellyi boulbenorum Deuve, 2015 Holotype
Male adult
Géorgie, au nord de Lentekhi, 1000 m. R. et A. Boulben (MNHN)
Copyright © Thierry Deuve

Carabus (Archiplectes) plasoni dalensis Rost, 1893
Male adult
Abkhasia, near Yu. Ptysh vill.
14.7.1986 Zamotailov A. leg.
31 mm
Copyright © Alexander Anichtchenko

Carabus (Archiplectes) plasoni dalensis Rost, 1893
Male adult
Caucasus merid., Dal-Suchumi, N.S. Brianskyj leg.
from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Archiplectes) plasoni plasoni Ganglbauer, 1886
Female adult
Gruzia, Centr. Caucasus, Svanetia, coll. A. Zolotarew
det. D.V. Obydov
from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Neoplectes) polychrous Rost, 1892
Male adult
Abkhasia, Gudaut distr., near Huap vill., 600 m, 9.VI.1990, M. Danilevskyi leg. det. D.V. Obydov
Copyright © Dmitri Obydov

Carabus (Neoplectes) polychrous polychrous Rost, 1892
Female adult
W.Caucasus, Abkhasia, Gudauta distr., Duripsh village env., Khipsta valley, h=500-700 m. 8-11.06.2009 A. Safronov leg. lenght 40-43 mm.
Copyright ©

Carabus (Neoplectes) rousianus rousianus Gottwald, 1985
Male+Female adult
Абхазия, Гудаутский р-н, 5-8 км ЮВ от пос. Лдзаа (Ludzava) долина р. Риапши, левый борт; балки, поросшие орехом, каштаном, рододендроном понтийским и кленом, h = 58-70 m, 43º10’43” N / 40º25’37” E, 16-22.07.2011, leg. И.А. Солодовников, Н.Ю. Пичугин, С.M. Павлючук
Copyright © Igor Solodovnikov

Carabus (Neoplectes) polychrous Rost, 1892
Female adult
Абхазия, Гудаутский р-н, окр. п. Тванаарху, дол. р. Хипсты, ущелье небольшого ручейка, поросшее буками, ежевикой и папоротником, Линия 25-3, 43°14’ N / 040°39’ E, h = 500 m,08.05-05.07.2013, leg. И.А. Солодовников, С.В. Солодовникова, С.M. Павлючук, Н.Ю. Пичугин, В.М. Коцур
Copyright © Igor Solodovnikov

Carabus (Neoplectes) polychrous Rost, 1892
Male+Female adult
Абхазия, Гудаутский р-н, окр. п. Тванаарху, дол. р. Хипсты, ущелье небольшого ручейка, поросшее буками, ежевикой и папоротником, Линия 25-3, 43°14’ N / 040°39’ E, h = 500 m,08.05-05.07.2013, leg. И.А. Солодовников, С.В. Солодовникова, С.M. Павлючук, Н.Ю. Пичугин, В.М. Коцур
Copyright © Igor Solodovnikov

Carabus (Neoplectes) rousianus svetlanae Solodovnikov, 2015 Paratype
Male+Female adult
Абхазия, Гудаутский р-н, 12 км дороги на оз. Рица, лев. борт р. Бзыбь, буковый лес + каштан на щебнисто-глинистой почве, уроч. Авджуляр, Линия № 20, h = 210-230 m, 43º19’07” N / 040º25’06” E, 09.V-03.VII.2013, leg. И.А. Солодовников, С.В. Солодовникова, С.М. Павлючук, Н.Ю. Пичугин, В.М. Коцур
Copyright © Igor Solodovnikov

Carabus (Neoplectes) rousianus svetlanae Solodovnikov, 2015 Paratype
Male adult
Абхазия, Гудаутский р-н, 12 км дороги на оз. Рица, лев. борт р. Бзыбь, буковый лес + каштан на щебнисто-глинистой почве, уроч. Авджуляр, Линия № 20, h = 210-230 m, 43º19’07” N / 040º25’06” E, 09.V-03.VII.2013, leg. И.А. Солодовников, С.В. Солодовникова, С.М. Павлючук, Н.Ю. Пичугин, В.М. Коцур
Copyright © Igor Solodovnikov

Carabus (Neoplectes) polychrous svetlanae Solodovnikov, 2015
Endophallus

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Carabus (Neoplectes) polychrous rousianus Gottwald, 1985
Endophallus

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Carabus (Neoplectes) polychrous polychrous
Endophallus

Copyright © Igor Solodovnikov

Carabus (Archiplectes) prometheus zolotarevi Zamotajlov, 1988 Paratype
Male adult
Paratypus, male - Fl. Min. Laba, VII.1918, Shaposhnikov leg.
from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990 (as hephaestus)
Male adult
Paratypus Carabus (Archiplectes) hephaestus
Maikop, Ust. Urusht, 26.IV.1910

Copyright © Dmitri Obydov

Carabus (Archiplectes) prometheus pschadensis Zamotajlov, 1990
Female adult
NW Caucas, 5 km N from Pshada vill., 25.04.1989, Smorodinov leg.
specimen from collection of M.Shestopalov (Moscow).
det. D.V. Obydov
from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Neoplectes) prometheus prometheus Reitter, 1887
Male adult
NW-Caucasus,Krasnodar reg., Apsheron Distr., Lagonaki Mt., near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominykh leg.
L= 42mm.
Copyright © Ruslan Panin

Carabus (Archiplectes) prometheus prometheus Reitter, 1887 (as nubicola)
Male+Female adult
W Caucasus, Adygea,
h~1700m. upper res. Riv. Kurdzhips, 11-21.VIII.2007, I.A.Solodovnikov leg./det.
L=32mm.
Copyright © Ruslan Panin

Carabus (Neoplectes) prometheus prometheus Reitter, 1887
Male adult
Krasnodar distr., Mezmai
20.6.1989
38 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) prometheus prometheus Reitter, 1887
Endophallus
aedeagus
Copyright © Ruslan Panin

Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990
Endophallus

Copyright © Ruslan Panin

Carabus (Neoplectes) zolotarevi Zamotajlov, 1988
Endophallus
Carabus (Archiplectes) zolotarevi zolotarevi Zamotajlov, 1988 Russia: Krasnodar reg., SW slopes of Malye Balkany Mt.R., env. Allous pass, h = 1960 m, 08-18.VII.2023 leg. D. Fominykh
Copyright © Igor Solodovnikov

Carabus (Neoplectes) zolotarevi dzhugensis Zamotajlov, 1988
Male adult
Russia, Adygea, E slopes of Dzuga Mt., left bank of Urushten Riv., near Meteostation “Dzhuga”, h = 1970 m, 21.VII-04.IX.2016, leg. Yu. Chumachenko
Copyright © Igor Solodovnikov

Carabus (Neoplectes) protensus olegi Belousov, 1992 Paratype
Male adult
Caucasus, W Georgia, Svanetia, right bank of riv. Magana, 800m, 12.V.-4.VII.1991, Zamotajlov & Miroshnikov leg.
male, 31mm
photo and collection Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) protensus protensoides Novotny & Vorisek, 1988
Male adult
Georgia, Martvilli env., Akshi plateau 10-15.7.2001
Rubenian leg.
32 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) protensus protensoides Novotny & Vorisek, 1988
Endophallus
W Georgia, Martvili distr., W slope of Askhi Plateau h=1200-1800 m,
Copyright © Ruslan Panin

Carabus (Neoplectes) protensus protensoides Novotny & Vorisek, 1988
Male adult
W Georgia, Martvili distr., W slope of Askhi Plateau h=1200-1800 m, 18-19.VI.2005, Shumov leg., L=33 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) protensus protensus Schaum, 1864
Female adult
Georgia, Caucasus, Imereti reg., Dzedzileti env, h=500-600m, 18-24.VI.2024, I.Uspenskyi leg., L=36.5 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) protensus protensus Schaum, 1864
Male adult
Georgia, Caucasus, Imereti reg., Dzedzileti env, h=500-600m, 18-24.VI.2024, I.Uspenskyi leg., L=32 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) juenthneri atchibachi Zamotajlov, 1991 Paratype
Female adult
W Caucas, Abkhasia, Valley of river Bzyb near Mt. Achibakhu. h=1500m.,
24.IV.-6.VI.1991 A.S. Zamotajlov leg.
coll. Anichtchenko
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) juenthneri atchibachi Zamotajlov, 1991 Paratype
Male adult
W Caucas, Abkhasia, Valley of river Bzyb near Mt. Achibakhu. h=1500m.,
24.IV.-6.VI.1991 A.S. Zamotajlov leg.
coll. Anichtchenko
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) juenthneri avadcharensis Kurnakov, 1970
Male adult
Abkhasia, 7,4km NE from Ritza lake, 1,5 km N Kutykuh vill.,
1635 m, 19-25.VII.2008, A. Vlasenko leg.
Copyright © Dmitri Obydov

Carabus (Neoplectes) reitteri reitteri Retowski, 1885 (as achunensis)
Male adult
W. Caucasus, Sochi Distr., Chernomoretz Vill. env.,
15.VII.1992, P. Gorbachev leg.
det. D.V. Obydov

Copyright © Dmitri Obydov

Carabus (Neoplectes) besleticus dsychvensis Gottwald, 1985
Male+Female adult
Abkhazia, Bzybian Mt. Range, SW slopes of Mt. Dzykhva, 2000-2300 m, alpine zone, N 43º13’, E 041º08’, 29.VI-03.VII.2013, leg. D. Fominykh
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus dsychvensis Gottwald, 1985
Male+Female adult
Abkhazia, Bzybian Mt. Range, SW slopes of Mt. Dzykhva, 2000-2300 m, alpine zone, N 43º13’, E 041º08’, 29.VI-03.VII.2013, leg. D. Fominykh
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus dsychvensis Gottwald, 1985
Endophallus

Copyright © Igor Solodovnikov

Carabus (Neoplectes) reitteri gagrinus Starck, 1894
Male adult
ABKHASIA, Gagra distr., Mamdsyshkha Mt., near Gagra, Fagus forest + karst, h = 1300-1330 m, biotope N 23, 43º18’186" N / 040º18’44" E, 12.05-05.07.2014, Leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Neoplectes) reitteri gagrinus Starck, 1894
Female adult
ABKHASIA, Gagra distr., Mamdsyshkha Mt., near Gagra, Fagus forest + karst, h = 1300-1330 m, biotope N 23, 43º18’186" N / 040º18’44" E, 12.05-05.07.2014, Leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Neoplectes) reitteri reitteri Retowski, 1885
Male adult

Copyright © Kirill Makarov

Carabus (Neoplectes) satyrus duripshensis Kurnakov, 1972
Male adult
W.Caucasus, Abkhasia, Gudauta distr.,
Duripsh village env., Khipsta valley, h=500-700 m.
07-12.VI.2009 A.Safronov leg.
male - 30 mm.
Copyright © Alexandr Safronov

Carabus (Neoplectes) satyrus satyrus Kurnakov, 1962
Male+Female adult
Abkhazia, Gulripsh Distr., NW slopes of Gurzul Mt. Range near Merkheul Village, 170-230 m, left tributary of Machara River, hornbeam, alder, rhododendron (Rhododendron ponticum) forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014
Endophallus

Copyright © Igor Solodovnikov

Carabus (Neoplectes) satyrus s.str.
Endophallus

Copyright © Igor Solodovnikov

Carabus (Neoplectes) pseudopshuensis Zamotajlov, 1991
Male adult
1 male - Abkhazia, Sukhum Distr., vicinities of Village Sanchara, River Baul, beech forest site, 20.VII.1991, leg. M.N. Maksimenkov 2 male - Abkhazia, Sukhum Distr., SE of Village Pskhu, hamlet Bitaga, Mt. Bzybskaya, 550-650 m, beech forest site, 12.V-30.VI.2010, leg. I. Solodovnikov, D. Fominykh
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972
Male+Female adult
Abkhazia, Gudauta Distr, vicinities of Mtsara (Chiryuta), Zashirbara Mt. Range, source of River Mtsara, beech, blackberry, fern site, 575-690 m, 11.V-05.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus napraensis Belousov & Zamotajlov, 1993 Paratype
Female adult
Abkhazia, Gudauta Distr., Bzybian Mt. Range, vicinities of Mt. Chibzharga, 1900-2000 m, 20.VI-26.VII.1992, leg. I. Belousov, A. Zamotajlov, A. Miroshnikov
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus anastasiae Solodovnikov, 2014 Holotype Paratype
Female adult
Abkhasia, Gudauta distr., S slope Bzybian Mt.R., upper reach. Riv. Khipsta, 43°18' N / 40°41' E, h = 1907-1915 m, subalpine meadows with karst blocks, 07-12.08.2014, leg. I.A. Solodovnikov, E.V. Tatun
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus anastasiae Solodovnikov, 2014 Paratype
Female adult
Abkhasia, Gudaura distr., S slope Bzybian Mt.R., upper reach. Riv. Khipsta, 43°18' N / 40°41' E, h = 1907-1915 m, subalpine meadows with karst blocks, 07-12.08.2014, leg. I.A. Solodovnikov, E.V. Tatun
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus anastasiae Solodovnikov, 2014 Paratype
Male adult
Abkhasia, Gudauta distr., S slope Bzybian Mt.R., upper reach. Riv. Khipsta, 43°18' N / 40°41' E, h = 1984-1990 m, subalpine meadows with big karst blocks, 07-12.08.2014, leg. I.A. Solodovnikov, E.V. Tatun
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starcki Heyden, 1885
Male+Female adult
NW-Caucasus,
Krasnodar reg., Mt. Aishkho, Malaya Laba River, h=2200m., 14.VII.2008, D.Fominyh leg.
female-21mm
Copyright © Ruslan Panin

Carabus (Neoplectes) starcki Heyden, 1885
Male adult
Caucasian Nat. Park, S slope
Pseashkho mt. 2500m 21.7.1975, Ovtsharenko leg.
20 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) starcki Heyden, 1885
Male adult
NW Caucasus, Adygeya, Pshekho-Su Mt. env., h=1500 m., 21.07.2006, D.Safronov leg.
Copyright © Safronov

Carabus (Neoplectes) starcki Heyden, 1885 (as parvulus)
Male adult
Prov. Kuban, mt. Aischa, VI.1906, Shaposhnikov leg.
det. D.V. Obydov

Copyright © Dmitri Obydov

Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988 (as shchessiensis) Paratype
Male+Female adult
NW Caucasus,
Krasnodar reg., Tuapsinskyi distr. Shessi Mt.
30.V.-25.VI.1993 Mt.Shessi, h=1800m., Imura & A.S.Zamotajlov, 1993 leg.
female, 29mm
Copyright © Ruslan Panin

Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988
Male adult
West. Caucasus, Shakhe Valley, 1.VI.1989, A. Shamaev leg.
det. D.V. Obydov

Copyright © Dmitri Obydov

Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
Male+Female adult
NW-Caucasus, Russia,
Krasnodar reg.,the right bank of Mzymta river, Engelmanovs glades, h=1200m., A.S.Zamotajlov leg.
female-30mm\male-28mm
Copyright © Ruslan Panin

Carabus (Neoplectes) starckianus starckianus Ganglbauer, 1886
Female adult
W Caucas, Sochi distr., near Lazorevskoe vill., 9.06.1993, P.Gorbachev leg.
det. D.V. Obydov


Copyright © Dmitri Obydov

Carabus (Neoplectes) starckianus theseus Briansky, 1910
Female adult
Tuapse distr., Anastasievka vill., Semiglavaya mt.
bass. Pshekaho riv. 20.7.1995
female, 30 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988 (as shchessiensis) Paratype
Endophallus

Copyright © Ruslan Panin

Carabus (Neoplectes) starckianus olgae Semenov, 1898
Label

Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
Female adult
Russia, Krasnodar Territory, West-Caucasus, Apsheronsky district, Guam ridge, env. Mezmay village, by the road, beech, oak, blackberry, h = 1200-1250 m, 03.IV-14.VI.2020, leg. S.M. Pavlyuchuk, A.P. Tatulyan
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
Male adult
W Caucasus, Aibga Mt.R., 02.VIII.1994, A.S. Zamotailov leg.
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus adelaidae Starck, 1889
Female adult
Russia, Krasnodar area, Adler, Ader distr., right bank vall. Riv. Msymta, near vill. Chvesipse (Fagus+Quercus+Castanea forest) [Sochi national park], h = 720 m, 03.V-08.VII.2016, I.A. Solodovnikov, S.V.Solodovnikova, V.M.Kotsur leg.
Copyright © Igor Solodovnikov