Carabidae image gallery
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Acathaicus taxon page (17 photos)
Carabus (Acathaicus) alexandrae alexandrae Semenov, 1887 Holotype Holotype
Male adult
Kwantin and Li-tschara-pu, 22.VI.1885, leg. Potanin<br /> Deposited in Zoolog. Museum St. Petersburg.<br /> male, long. 22 mm<br /> photo B. Brezina
Copyright © Boleslav Brezina
Carabus (Acathaicus) alexandrae alexandrae Semenov, 1887
Male adult
male, 25mm<br /> photo B. Brezina
Copyright © Boleslav Brezina
Carabus (Acathaicus) alexandrae alexandrae Semenov, 1887
Male adult
female, 28mm<br /> photo B. Brezina
Copyright © Boleslav Brezina
Carabus (Acathaicus) alexandrae alexandrae Semenov, 1887
Male adult
female, 24mm<br /> photo B. Brezina
Copyright © Boleslav Brezina
Carabus (Acathaicus) alexandrae cratocephaloides Semenov, 1887 Holotype Holotype
Male adult
Copyright © Boleslav Brezina
Carabus (Acathaicus) alexandrae cratocephaloides Semenov, 1887
Male adult
male, 26mm
Copyright © Boleslav Brezina
Carabus (Acathaicus) alexandrae cratocephaloides Semenov, 1887
Male adult
male, 24mm
Copyright © Boleslav Brezina
Carabus (Acathaicus) alexandrae cratocephaloides Semenov, 1887
Female adult
female, 28mm
Copyright © Boleslav Brezina
Carabus (Acathaicus) idolon Semenov, 1898 Holotype Holotype
Male adult
male
Copyright © Boleslav Brezina
Carabus (Acathaicus) interruptocostatus interruptocostatus s.str.
Female adult
female, 27mm
Copyright © Boleslav Brezina
Carabus (Acathaicus) interruptocostatus interruptocostatus s.str.
Male adult
male, 29mm
Copyright © Boleslav Brezina
Carabus (Acathaicus) interruptocostatus Semenov, 1887 Holotype Holotype
Female adult
Copyright © Boleslav Brezina
Carabus (Acathaicus) alexandrae Semenov, 1887
Male adult
Carabus (Acathaicus) alexandrae alexandrae Semenov, 1897 Holotype
Kwantin and Li-tschara-pu, 22.VI.1885, leg. Potanin
Deposited in Zoolog. Museum St. Petersburg.
male, long. 22 mm
photo B. Brezina
Copyright © Boleslav Brezina
Carabus (Acathaicus) interruptocostatus fantingi Imura & Su, 1998 Holotype
Male adult
China, N.-Sichuan, above Jiuzhaigou, 2100-2200m, N bank of Riv. Bai He, Jiuzhaigou Xian" - 24 mm lang - in coll. Imura.
Copyright © Frank Kleinfeld
Carabus (Acathaicus) interruptocostatus nanpingicus Deuve, 1995 Holotype
Male adult
Chine, Sichuan, Nanping, VII.1995, S. J. L. Remond" - Länge 25 mm - in MNHN Paris.
Copyright © Frank Kleinfeld
Carabus (Acathaicus) interruptocostatus amplior Deuve, 1994 Holotype
Male adult
Chine, Gansu, Min Shan, 2000m, J. M. Bousquet, 1994" - Länge 29 mm - in MNHN Paris
Copyright © Frank Kleinfeld
Carabus (Acathaicus) interruptocostatus tewoicus Deuve, 1996 Holotype
Male adult
Chine, Gansu, Tewo, 2000m, J. L. Remond, SR. 24,5 mm, in MNHN Paris
Copyright © Frank Kleinfeld -
Acoptolabrus taxon page (63 photos)
Carabus (Acoptolabrus) constricticollis constricticollis Kraatz, 1886
Female adult
Copyright © Alexander Anichtchenko
Carabus (Acoptolabrus) gehinii gehinii (as aereicollis Hauser, 1921)
Male adult
Copyright © Alexander Anichtchenko
Carabus (Acoptolabrus) gehinii gehinii Fairmaire, 1886
Male+Female adult
Komari, Chitose, Hokkaido, Japan.
20-27.05.2007
Copyright © Safronov
Carabus (Acoptolabrus) gehinii gehinii Fairmaire, 1886
Male+Female adult
Bibi, Chitose, Hokkaido, Japan.
03.05.2000
Copyright © Safronov
Carabus (Acoptolabrus) gehinii gehinii (as manoianus (Imura, 1989)
Male adult
Copyright © Alexander Anichtchenko
Carabus (Acoptolabrus) gehinii munakatai (Ishikawa, 1968)
Male+Female adult
Mt.Shiribetsudake, Kishibetsu, Hokkaido, Japan.
05.06.2005
Copyright © Safronov
Carabus (Acoptolabrus) leechi nangminicola Deuve & Li, 2001
Female adult
N Coree, Cha Gangdo, Nangnimsan Mt., 6.2006. 31 mm. (coll. A. Montes, Madrid)
Copyright © Alexander Anichtchenko
Carabus (Acoptolabrus) lopatini Morawitz, 1886
Male+Female adult
Russia,
Sakhalin isl., Tchekhov Mt, VI.1997
female, 40mm
photo & coll. Panin R.
Copyright © Ruslan Panin
Carabus (Acoptolabrus) mirabilissimus rigoutianus Deuve & Li, 1998
Female adult
N Koree, Pyonganbukto, mt. Miaoxiang 6.2006
28 mm
coll. A.Montes
Copyright © Alexander Anichtchenko
Carabus (Acoptolabrus) schrencki schrencki (as hauryi Gehin, 1885 )
Male adult
Copyright © Boleslav Brezina
Carabus (Acoptolabrus) schrencki schrencki (as hauryi Gehin, 1885 )
Male adult
det. Brezina B.
Copyright © Boleslav Brezina
Carabus (Acoptolabrus) schrencki schrencki (as hauryi Gehin, 1885 )
Male adult
Copyright © Boleslav Brezina
Carabus (Acoptolabrus) schrencki schrencki Motschulsky, 1860
Female adult
Primorie, Lazovskii distr., Chandalaz mt., L.Voita leg 19.07.2008
photo A.Menstschicov
Copyright © Anton Menstschikov
Carabus (Acoptolabrus) schrencki kimieae Kezuka & Imura, 1989
Male+Female adult
Mt. Wulongshan, Dandong City,
Liaoning Province, CHINE
25-30.VI.2008
Copyright © Safronov
Carabus (Coptolabrodes) haeckeli Brezina & Imura, 1997
Male adult
C China, Shaanxi,
Tsinling Mts., Taibaishan Mt., h=3000m, 33:55N/107:44 E, VI.2004, V.Siniaev & his team
female, 31 mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Acoptolabrus) schrencki schrenck (as dongningensis Deuve & Li, 2005)
Male adult
NE China, Heilongjiang prov., Dongning, V.2009<br /> L=25 mm
Copyright © Ruslan Panin
Carabus (Acoptolabrus) constricticollis constricticollis (as grallatorius Roeschke, 1921)
Male adult
NE-Korea:North Hamgyong Prov., Sechon-dong, VII.2004, Li leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Acoptolabrus) mirabilissimus rigoutianus Deuve & Li, 1998
Male adult
N-Korea, South Pyongan prov., Mt. Maeng-san, 6.VI.2010
L=25 mm
Copyright © Ruslan Panin
Carabus (Acoptolabrus) gehinii gehinii Fairmaire, 1886
Female adult
N-Japan, SW-Hokkaido, Chitose env.,VI.2011.
L=34 mm
Copyright © Ruslan Panin
Carabus (Acoptolabrus) constricticollis pihyonensis Deuve & Li, 1998
Male adult
N-Korea, North Pyongan Prov., Tongnim County, 5-25.V.2010 <br /> 29 mm
Copyright © Ruslan Panin
Carabus (Acoptolabrus) ? leechi huisaensis Deuve & Li, 1999
Male adult
NC-Korea, Ryanggang Prov., Puksubaek-san, 4-10.V.2010
28 mm
Copyright © Ruslan Panin
Carabus (Acoptolabrus) mirabilissimus mirabilissimus Ishikawa & Deuve, 1982
Female adult
Mt.Taebaek-san, Kangwon-do,VI-2002, CE-Korea (L=26mm)
Copyright © Alain Mollard
Carabus (Acoptolabrus) mirabilissimus pseudokana Deuve & Li, 2002
Male adult
Mt.Wujia-san, Chagang-do, VII-2004, N-Korea (L=21.5mm)
Copyright © Alain Mollard
Carabus (Acoptolabrus) mirabilissimus kana Imura, 1989
Female adult
Mt.Togyu-san, Cholla-bukto, VII-2000, SC-Korea (L=23.5mm)
Copyright © Alain Mollard
Carabus (Acoptolabrus) mirabilissimus igniferescens Deuve, 1992
Male adult
Mt. Taebaek-san, Kangwon-do, VI-2002, CE-Korea (L=21.5mm)
Copyright © Alain Mollard
Carabus (Acoptolabrus) mirabilissimus mirabilissimus (as furumiensis)
Male adult
Mt.Sobaek-san, Ch’uich’ong bukto, VII-2001, CE-Korea (L=22mm)
Copyright © Alain Mollard
Carabus (Acoptolabrus) constricticollis igniferrimus Deuve & Li, 2003
Female adult
Mt.Nangnim-san, Chagang-do, VI-2002, N-Korea L=32mm
Copyright © Alain Mollard
Carabus (Acoptolabrus) constricticollis chonmasanensis Deuve, 1994
Female adult
Mt. Chonma-san, Pyongang-bukto VII-1996, N-Korea L=35mm
Copyright © Alain Mollard
Carabus (Acoptolabrus) constricticollis furumielloides Deuve, 1997
Male adult
Mt. Liuhe, Jilin Prov. VI-1997, Chine L=23mm
Copyright © Alain Mollard
Carabus (Acoptolabrus) constricticollis furumielloides Deuve, 1997
Male adult
Lanheyu, E-Benxi, Liaoning Prov. VII-1998, Chine L=33mm
Copyright © Alain Mollard
Carabus (Acoptolabrus) constricticollis samjiyonensis Deuve & Li, 2000
Male adult
Mt.Samjiyon-san, Yanggang-do, VI-2004, N-Korea L=22mm
Copyright © Alain Mollard
Carabus (Acoptolabrus) constricticollis myohyangensis Deuve & Li, 1998
Female adult
Mt.Myohyang-san, Pyongang-bukto VI-2002, N-Korea L=28mm
Copyright © Alain Mollard
Carabus (Acoptolabrus) gehinii gehinii (as konsenensis Ishikawa,1968)
Male adult
Shibecha (C-Hokkaido) Japan 01/05/1993
Copyright © Alain Mollard
Carabus (Acoptolabrus) gehinii gehinii (as radiatocostatus Ishikawa,1968)
Female adult
Erimo-Cho (SC-Hokkaido) Japan 15/06/1990.
Copyright © Alain Mollard
Carabus (Acoptolabrus) gehinii gehinii (as shimizui Imura, 1994)
Male adult
Tamaru-toge-Pass , (Shakotan),Japan 01/05/1995.
Copyright © Alain Mollard
Carabus (Acoptolabrus) gehinii gehinii (as sapporensis Uchida & Tamanuki,1927)
Female adult
Mt.Apodiake (SC-Kokkaido),Japan 12/06/1994
Copyright © Alain Mollard
Carabus (Acoptolabrus) gehinii munakatai (as nishijimai Imura, 1991)
Female adult
Mt. Obira-yama (Shimamaki) Japan, 12/07/1989
Copyright © Alain Mollard
Carabus (Acoptolabrus) gehinii munakatai (as okikulumi Nakajima,1993)
Male adult
Mt.Otobe-dake (Yamakoshi) Japan,07/06/1999
Copyright © Alain Mollard
Carabus (Acoptolabrus) gehinii munakatai (as ninetopa Nakajiima, 1993)
Male adult
Shimamaki-gun (SW-Hokkaido) Japan, 12/07/1999.
Copyright © Alain Mollard
Carabus (Acoptolabrus) gehinii munakatai (as furumii Mandl, 1982)
Male adult
Shiribeshi (Oshima) Japan, 23/07/1999.
Copyright © Alain Mollard
Carabus (Acoptolabrus) gehinii gehinii Fairmaire, 1886 Holotype
Male adult
Yokohama; http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec148
Copyright © - - -
Carabus (Acoptolabrus) lopatini Morawitz, 1886
Male adult
S Sakhalin, Yuzhno Sakhalinsk, Tschekhova Mt., 800 m.,, 28.VI.2017 ; Фотография с сайта www.zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov
Carabus (Acoptolabrus) schrencki minpongsanensis Deuve & Li, 2003
Male adult
North Korea, Chagang Prov., Manpo, Songhak-ri (villl.,) 1-16.VII.2011, L=26 mm
Copyright © Ruslan Panin
Carabus (Acoptolabrus) constricticollis constricticollis (as grallatorius)
Male adult
33 mm, Seishin Korea (= Chongjin, North Hamgyong province, D.R. of Korea) (CIR). Image from Rapuzzi, 2015
Copyright © - - -
Carabus (Acoptolabrus) constricticollis microcolasellus Rapuzzi, 2015 Holotype Paratype
Male+Female adult
D.R. of Korea, North Hamgyong Province, Paeksa, 1139 m, VII.2004, local collector leg. (CIR). Image from Rapuzzi, 2015
Copyright © - - -
Carabus (Acoptolabrus) costricticollis aff. limes Rapuzzi, 2009
Male adult
D.R. of Korea, South Hwanghae province, Unryul County, Mt. Kuwol, 954 m, local collector leg. (CIR). Image from Rapuzzi, 2015
Copyright © - - -
Carabus (Acoptolabrus) leechi drouini Rapuzzi, 2015 Holotype Paratype
Male+Female adult
D.R. of Korea, North Hamgyong Prov., Kichu, Mt. Muhak, 8/19.VII.2006, local collector leg. (CIR). Image from Rapuzzi, 2015
Copyright © - - -
Carabus (Acoptolabrus) leechi onjinsanicola Rapuzzi, 2009
Male adult
D. R. of Korea, South Hwanghae province, Unryul County, Mt. Kuwol, 954 m, local collector leg. (CIR). Image from Rapuzzi, 2015
Copyright © - - -
Carabus (Acoptolabrus) leechi viniciosalamii Rapuzzi, 2015 Holotype Paratype
Male+Female adult
D.R. of Korea, North Hamgyong Prov., Mt. Wanta, 24.VII.2004, local collector leg. (CIR). Image from Rapuzzi, 2015
Copyright © - - -
Carabus (Acoptolabrus) planicranion Rapuzzi, 2015 Holotype Paratype
Male+Female adult
D.R. of Korea, North Hamgyong Prov., Mt. Obong, (ca 42,40 NL; 129,80 EL), 1/15.VII.2012, local collector leg. (CIR). Image from Rapuzzi, 2015
Copyright © - - -
Carabus (Acoptolabrus) changeonleei Ishikawa & Kim, 1983
Female adult
R. of Korea, Jirisan Mt., I. Rapuzzi and L. Caldon leg. (CIR). Image from Rapuzzi, 2015
Copyright © - - -
Carabus (Acoptolabrus) mirabilissimus igniferescens Deuve, 1992
Male adult
R. of Korea, Gangwondo province, South from Gimhwa, 450 m (CIR). Image from Rapuzzi, 2015
Copyright © - - -
Carabus (Acoptolabrus) mirabilissimus mirabilissimus Ishikawa & Deuve, 1982
Male adult
R. of Korea, Gangwondo province, “Peace Dam”, NE Hwacheon 400 m (CIR). Image from Rapuzzi, 2015
Copyright © - - -
Carabus (Acoptolabrus) mirabilissimus furumiensis Deuve, 2002
Male adult
R. of Korea, Gangwondo province, Odaesan Mt., Jingogae, 800 m (CIR). Image from Rapuzzi, 2015
Copyright © - - -
Carabus (Acoptolabrus) leechi chiricola (Kwon & Lee, 1984) Holotype
Male adult
deposited in Kyungpook National University, Daegu, Republic of Korea, Scale bar = 5mm
Copyright © Jong-Bong Choi
Carabus (Acoptolabrus) leechi chiricola (Kwon & Lee, 1984) Paratype
Female adult
deposited in Kyungpook National University, Daegu, Republic of Korea, Scale bar = 5mm
Copyright © Jong-Bong Choi
Carabus (Acoptolabrus) leechi chiricola (Kwon & Lee, 1984) Paratype
Female adult
deposited in Kyungpook National University, Daegu, Republic of Korea, Scale bar = 5mm
Copyright © Jong-Bong Choi
Carabus (Acoptolabrus) gehinii munakatai (Ishikawa, 1968)
Male adult
Japan, SW-Hokkaido, Yakumo town, 4.VI.2017, S.Shinichi leg., L=30 mm
Copyright © Ruslan Panin
Carabus (Acoptolabrus) leechi yooni Deuve, 1990
Male+Female adult
Sorak-San (Kyongi) KOREE, VI-2002
Copyright © Alain Mollard
Carabus (Acoptolabrus) constricticollis fei Rapuzzi, Li & Zhang, 2012 Holotype Paratype
Male+Female adult
North East China, South West Liaoning Province, Liadong Peninsula, Haicheng City, Gusahanzhen, ErDaoHe, 700 m. 17/19-VI-2011 (Holotype female, in the collection of Key Laboratory of remote sensing monitoring of geographic environment, College of Heilongjiang Province, Harbin Normal University, Harbin, China)
Copyright © - - -
Carabus (Acoptolabrus) constricticollis Kraatz, 1886
Male adult
China, Liaoning prov., Mt. Guanmen 20.6.2009 (cAA)
Copyright © Alexander Anichtchenko
Carabus (Coptolabrus) osawai berezovskiianus Deuve & Mourzine, 2003
Male adult
Image courtesy A. Gorodinsky
Copyright © - - - -
Acoptopterus taxon page (12 photos)
Carabus (Carabus) dreuxioides wushanling Kleinfeld, 1998 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Carabus) emanuelei grabneri Kleinfeld, 2003 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Carabus) gressittianus daiyunshan Kleinfeld, 1998 Holotype
Male adult
Copyright © Frank Kleinfeld
Carabus (Carabus) gressittianus fuscomarginatus Kleinfeld, 1997 Holotype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Carabus) kucerai wuangxianus Kleinfeld, 1998 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Carabus) mianningensis Kleinfeld, 2000 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Carabus) nanosomus wufengensis Kleinfeld, 1998 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Carabus) oblongior yunlingicus Deuve, 1994
Male adult
China, NW Yunnan, Zhongdian env.
3400m, 5.7.2000
male, 22 mm
Copyright © Alexander Anichtchenko
Carabus (Carabus) pseudolatipennis wenxianensis Deuve, 1992
Male adult
China, S Gansu, Min Shan 2100 m,
SW Wudu 1.8.2000, V. Patrikeev leg.
Copyright © Alexander Anichtchenko
Carabus (Carabus) pseudolatipennis wenxianensis Deuve, 1992
Male adult
male, 20 mm
Copyright © Boleslav Brezina
Carabus (Carabus) xupuensis Kleinfeld, 1998 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Carabus) kucerai puchneri Kleinfeld, 2007
Male adult
China, Sichuan, road Pingwu-Nanping, Umg. Mupi Tibetan (Mupi 32.33N/104.33, 1250 m, 28.5.-4.6.2006),
27 mm, coll. Puchner
Copyright © Frank Kleinfeld
Carabus (Carabus) vigilax rautenstrauchi Rapuzzi, 2018 Holotype
Male adult
China, Sichuan, Leibo, Shanlinggang environs, VII.2018, Wang legit in collection Ivan Rapuzzi. Image from Rapuzzi, 2018
Copyright © - - -
Carabus (Carabus) andreiianus Deuve, 2007
Female adult
C-China, N-Sichuan, Pingwu Co. h=1500 m, 27.VI.2011, L=24 mm
Copyright © Ruslan Panin
Carabus (Carabus) koidei koidei Imura, 1996
Male adult
C-China, S-Sichuan, Pass btw. Yanyaun / Muli, h=3244 m, 27.68638N; 101.22335E,,11-18.VI.2017, Ch. Reuter leg. L=22 mm
Copyright © Ruslan Panin
Carabus (s.str.) morphocaraboides zhaojueanus Imura, 1999
Female adult
China, S-Sichuan, SE Mianning, Pass btw Mianshanzhen & Yuexi 28°30’N/102°26’E, 22-23.VI.2017, Ch.Reuter leg., L=22 mm; det. B. Brezina, 2020
Copyright © Ruslan Panin
Carabus (Carabus) vigilax vigilax Bates, 1890 Lectotype
Male adult
Yang-tsé sup. Wa-Shan 6000ft, 1889-06-01/1889-06-30 (MNHN)
Copyright © MNHN
Carabus (s.str.) morphocaraboides zhaojueanus Imura, 1999
Male adult
China, S-Sichuan, road 56 km E of Xichang to Zhaojue, Mt.WenLin, Pass h=3200 m, 27°52’N/102°30E, 5-20.VI.2017, Ch.Reuter leg.; det B. Brezina, 2020
Copyright © Ruslan Panin
Carabus (Carabus) pseudolatipennis nanpingensis Deuve, 1992 Holotype
Female adult
N-Sichuan: Nanping Xian, 2600m
Copyright © Thierry Deuve
Carabus (Carabus) latipennis latipennis Breuning, 1932 Holotype
Female adult
Copyright © Thierry Deuve
Carabus (Carabus) baogai subbaogai Deuve, 2013 Holotype
Female adult
Chine, Hunan, Nanshan, 1750-1850, 26°10'N/ 10°09'E, in coll. Jaroslav Tuma, Kostelec na Hane (République Tchèque).
Copyright © Thierry Deuve
Carabus (Carabus) donjuan Imura, 2011 Holotype Paratype
Male+Female adult
alpine zone of the Chuandongzi Massif, 27.21.04-34N / 101.30.25-33E, 3035-3290 m in alt., south of Yanyuan, Yanyuan Xian, Liangshan-yizu-zizhizhou, S Sichuan, SW China, 29.VI.2010 (ZIN). Image from Imura, 2011
Copyright © - - -
Carabus (Carabus) koidei yanyuan Cavazzuti & Rapuzzi, 2010
Male adult
Image from Imura, 2011
Copyright © - - -
Carabus (Archaeocarabus) mirandus Cavazzuti & Rapuzzi, 2005 Holotype
Male adult
Cina, Sichuan meridionale, Jinyang, 3000 m.
Copyright © - - -
Carabus (Carabus) gressittianus tiangongicus Deuve, 2019 Holotype
Female adult
Chine, Fujian méridional, Tiangong Shan, 1508 m, 25°15’N-117°03’E, in coll. Jaroslav Turna, Kostelec na Hane (République Tchèque)
Copyright © Thierry Deuve
Carabus (Acoptopterus) dongchuanicus fazhensis Deuve, Liu & Liang, 2024 Holotype
Male adult
Chine, province du Yunnan, district de Dongchuan, ville de Fazhe, Mont Jiaozi Shan, Yanzidong, 2910 m, 26,09758°N 102,89406°E, en bord de route (Hongliang Shi leg., 26.V.2010), in IOZ. Image from Deuve et al 2024
Copyright © - - -
Carabus (Acoptopterus) yunnanus dayaoicus Deuve, Liu & Liang, 2024 Holotype
Male adult
Chine, province du Yunnan, canton de Dayao, ville de Santai, col de Xiaobaicaoling, 3105 m, 26,0561°N 101,0544°E, en forêt mixte (Hongliang Shi & Ye Liu leg., 30.V.2012, dans des pièges), in IOZ. Image from Deuve et al, 2024
Copyright © - - -
Carabus (Acoptopterus) yunnanus yanyuanicus Cavazzuti, 1996 (? or oblongior ssp.)
Male adult
China, S-Sichuan, S Yanyuan, Chuandongzi Mt h=3300-3600 m, 16-20.VI.2023, C.Reuter leg./det.Klienfeld, 2023, L=21.5 mm
Copyright © Ruslan Panin
Carabus (Acoptopterus) rautenstrauchi alberti Rapuzzi, 2024 Holotype
Male adult
Sichuan, Gu Dui Xiang, Meigu to Shanlenggang; 2250 m asl; Jul. 2022
Copyright © - - -
Carabus (Acoptopterus) pseudolatipennis wenxianensis Deuve, 1992
Male adult
S Gansu, Minshan, 5-20km S Cagan, 2300-3200m, 7-20.7.2019, V. Patrikeev leg. (cAA)
Copyright © Alexander Anichtchenko
Carabus (Acoptopterus) baogai Hackel & Sehnal, 2009 Holotype Paratype
Male+Female adult
S China, NE Guangxi, Jiaojiang Xian: Baogai Shan, 25°32.598´N 110°57.502´E, 1571 m, VI-2009; in collection of M. Häckel. Image from Hackel & Sehnal, 2009
Copyright © - - - -
Acrocarabus taxon page (6 photos)
Carabus (Acrocarabus) callisthenoides Semenov, 1888
Male adult
male, 22mm
Copyright © Boleslav Brezina
Carabus (Acrocarabus) callisthenoides Semenov, 1888
Male adult
female, 23mm
Copyright © Boleslav Brezina
Carabus (Acrocarabus) guerini aldarkose Kabak, 1992
Male adult
male, 22mm
Copyright © Boleslav Brezina
Carabus (Acrocarabus) guerini guerini Fischer, 1842
Male adult
male, 21mm
Copyright © Boleslav Brezina
Carabus (Acrocarabus) guerini guerini Fischer, 1842
Female adult
female, 25mm
Copyright © Boleslav Brezina
Carabus (Acrocarabus) guerini guerini Fischer, 1842
Male adult
SE Kazakhstan, W Dzhungar Mts, Lepsinsk env., 1600 m., 18.06.2000, A.Klimenko
Copyright © Anton Menstschikov -
Alipaster taxon page (18 photos)
Carabus (Alipaster) barovskii bayanbulak Deuve, 1993 Paratype Paratype
Male adult
male, 15mm
Copyright © Boleslav Brezina
Carabus (Alipaster) barovskii bayanbulak Deuve, 1993 Paratype Paratype
Female adult
female, 17mm
Copyright © Boleslav Brezina
Carabus (Alipaster) infantulus infantulus Morawitz, 1886
Male adult
male, 15mm
Copyright © Boleslav Brezina
Carabus (Alipaster) pupulus pupulus Morawitz, 1889
Male adult
Inner Tien-Shan, W part of Terskei mnts., Ukek rng., 20-25 km E Kotshkorka, h=2800 m, 8.07.98 leg. Tretyakov<br /> male 13,6mm
Copyright © Maksim Smirnov
Carabus (Alipaster) pupulus pupulus Morawitz, 1889
Male adult
female, 16mm
Copyright © Boleslav Brezina
Carabus (Alipaster) pupulus pupulus Morawitz, 1889
Male adult
Kirgizstan, Terskei AlaToo Mt. rng, Kadzhi-Sai v., 2800 m.; A.Subankulov leg.; 30.05.2000
Copyright © Anton Menstschikov
Carabus (Eotribax) valikhanovi bagua Kabak, 2009 Holotype
Female adult
China, Xinjiang, N slope of Narat Mt. R., SSW of Tekes, Kokterek basin, Buryl Riv., 3145 m, 28.07.2009,
42°48'06»N / 81°41'20»E, I. Kabak leg. (ZIN)
Copyright © Ilya Kabak
Carabus (Alipaster) thianshanskii shatyensis Deuve et Tian, 2004 Paratype
Male adult
Copyright © Ilya Kabak
Carabus (Alipaster) thianshanskii sarkhobensis Kabak, 2009 Holotype
Female adult
CH, Xinjiang, N sl. of Khalyktau MR., N Kongurbulak R., H=3600 m, 42°35'50»N / 81°02'30»E, 31.07.2003, I. Kabak leg
(ZIN)
Copyright © Ilya Kabak
Carabus (Cechenotribax) petri alipastroides Kabak, 2009 Holotype
Male adult
E spur of Terskei Alatau Mt. R., 10 km S of Saryzhas Village, 2800-3000 m, 5.VII.1987, I. Kabak: 42°51’00”N / 79°37’00”E – 42°50’30”N / 79°39’00”E)
(ZIN)
Copyright © Ilya Kabak
Carabus (Cechenotribax) petri Semenov & Znojko, 1932
Female adult
female, 18mm
Copyright © Boleslav Brezina
Carabus (Cechenotribax) petri Semenov & Znojko, 1932
Male adult
male, 14mm
Copyright © Boleslav Brezina
Carabus (Eotribax) malkovskyi malkovskyi Kabak, 1990
Male adult
male, 19mm
Copyright © Boleslav Brezina
Carabus (Eotribax) malkovskyi malkovskyi (as erenhabergaensis Deuve, 1993) Paratype
Male adult
male, 16mm
Copyright © Boleslav Brezina
Carabus (Eotribax) malkovskyi malkovskyi (as erenhabergaensis Deuve, 1993)
Male adult
male, 17mm
Copyright © Boleslav Brezina
Carabus (Eotribax) malkovskyi malkovskyi (as erenhabergaensis Deuve, 1993)
Male adult
male, 17mm
Copyright © Boleslav Brezina
Carabus (Eotribax) malkovskyi malkovskyi (as erenhabergaensis Deuve, 1993)
Male adult
male, 18mm
Copyright © Boleslav Brezina
Carabus (Eotribax) hiekei Kabak & Kryzhanovskij, 1990
Female adult
female, 18mm
Copyright © Boleslav Brezina
Carabus (Eotribax) hiekei Kabak & Kryzhanovskij, 1990
Male adult
male, 17mm
Copyright © Boleslav Brezina
Carabus (Eotribax) eous ukokensis Belousov & Kabak, 1994 Paratype
Female adult
female, 18mm
Copyright © Boleslav Brezina
Carabus (Eotribax) eous ukokensis Belousov & Kabak, 1994 Paratype
Male adult
male, 17mm
Copyright © Boleslav Brezina
Carabus (Eotribax) eous eous Morawitz, 1889
Male adult
Kirgizstan, Terskey Ala-Too, Borskanu riv,
2600 m, 29.5.1999
Copyright © Alexander Anichtchenko
Carabus (Eotribax) eous eous Morawitz, 1889
Male adult
Kirghyzstan, N Terskei Ala Too, Borskaun riv., h=2500 m., 7-14.07.2000
male
L=17 mm
Copyright © Maksim Smirnov
Carabus (Cechenotribax) petri Semenov & Znojko, 1932
Male adult
SE Kazakhstan, N-Tianshan:N-slopes Ketmen Mt.Rng. h=2250-2500m., near Ketmen vill.,1-5.VI.1990, Dolin V.G. leg.
L=13.5 mm
Copyright © Ruslan Panin
Carabus (Alipaster) barovskii bayanbulak Deuve, 1993
Male adult
China, Xinjiang, S sl. of Khalyktau, road Kuga-Bayanbulak, Telemet Pass. S slope, h~3400-3500 9.VI.2001 leg.I.Kabak L=14.0 mm
Copyright © Kirill Makarov
Carabus (Alipaster) pupulus pupulus Morawitz, 1889
Female adult
S Kyrgyzstan, Inner Tian Shan, Dzhany-Dzher Mt range, upper Mudyuryum riv. Ak-Baytal gorge h~3300 m, 15.VII.2011, S.Toporov leg. L=16 mm
Copyright © Ruslan Panin
Carabus (Alipaster) foreli baiensis Deuve & Kabak, 2013 Holotype
Male adult
Chine Xinjiang, affluent droit de la rivière Terek, 3500 mètres (env. 58 km à l'ouest-nord-ouest de Baicheng), 42°0l'N/81°15'E, pelouses alpines et éboulis, in coll. Jaroslav Kalâb, à Jinacovice (République Tchèque)
Copyright © Thierry Deuve
Carabus (Alipaster) foreli baiensis Deuve & Kabak, 2013 Paratype
Male adult
Chine, Xinjiang, versant sud des Mts Khalyk-taou (Halik Shan), Bay Xian, au nord-ouest de Terekmeikon, 3200-3690 mètres, 42°02'22"N/81°26'17"E à 42°01'29"N/81°23'43"E (!. Kabak, 12 juillet 2006)
Copyright © Thierry Deuve
Carabus (Alipaster) semenoviellus Breuning, 1934 Holotype
Male adult
(NBC), «Tekkes-tal.», «semenoviellus t. Breuning c.», «collectie C. & O. Vogt Acq. 1960», «Typus», «Holotypus», «C. (Alipaster) semenoviellus Br. Gottwald det. 1984», «Alipaster semenoviellus Breuning, 1934 ZMAN type COLE. 0375.1».
Copyright © Ilya Kabak
Carabus (Alipaster) semenoviellus Breuning, 1934
Male adult
ESE of Tekes, Tagymbel, S of Karatogai Vill., 43°09ʹ19ʺN / 82°14ʹ45ʺE, 2200 m, 10.07.2014 (I.I. Kabak leg.)
Copyright © Ilya Kabak
Carabus (Alipaster) semenoviellus tokkuztaraensis Kabak, 2019 Holotype Paratype
Male adult
Holotype, left: China, Xinjiang, Narat Mt. R., Chashi Valley, Tuyukasha, 43°00ʹ29ʺN / 82°46ʹ02ʺE, 2275 m, 18.07.2014 (I.I. Kabak leg.); Paratypes: middle: China, Xinjiang, Narat Mt. R., Erbotu Valley, 43°06ʹ14ʺN / 83°00ʹ05ʺE, 1915 m, 21.07.2014 (I.I. Kabak leg.). Right: China, Xinjiang, Narat Mt. R., Aktuyuk River, S of Kurdelin, 43°04ʹ12ʺN / 82°51ʹ19ʺE, 2255 m, 31.07.2013 (I.I. Kabak leg.)
Copyright © Ilya Kabak -
Apoplesius taxon page (5 photos)
Carabus (Apoplesius) deuvei hajdaji Hackel, 2007 Holotype Holotype
Male adult
Turkey- prov. Mus 60 km NW Mus 1630m
3856N 4108E Bulgan Gec. 23-24.06.03 E. & P. Hajdaj leg.
Copyright © Boleslav Brezina
Carabus (Apoplesius) pseudodepressus Machard, 1988
Female adult
Turkey, Prov. Bitlis, S near Tatvan, 23.IV.2009 leg. M. & S. Nabozhenko
L=19.0 mm
Copyright © Kirill Makarov
Carabus (Apoplesius) omphreodes omphreodes Holotype
Male adult
Paraplesius omphreodes 1898 Mardin / coll. Rttr 5-IX-1900 / coll. Roeschke type / Holotypus / omphreodes Rttr. / coll. C. & O. Vogt Acq. 1969 - in coll. Zoolog. Inst. Univ. Amsterdam; 23,5 mm.
Copyright © Frank Kleinfeld
Carabus (Apoplesius) omphreodes rawandozi Kleinfeld & Reuter, 2018 Holotype Paratype
Male adult
Holotype: N. Irak, Umg. Rawandoz, 36°38’N/44°33’E, Akoian valley, 30/IV/12-V-2018, 1900-2000 m, (23,5mm). Paratypus ♀: id. but IV-2017 (25 mm)
Copyright © Frank Kleinfeld -
Apotomopterus taxon page (201 photos)
Carabus (Apotomopterus) anxiensis Kleinfeld, 1998 Holotype & Paratype Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) arrowi hector Hauser, 1924
Female adult
China, Guangxi pro., Maoershan Mountain, Liang H.B. coll. Photo by Liuye det. Anichtchenko A.
Copyright © Ye Liu
Carabus (Apotomopterus) bousquetellus gengmaensis Kleinfeld, 2003 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) cantonensis pervarius Kleinfeld, 1997 Holotype
Male adult
Frank Kleinfeld
Copyright © - - -
Carabus (Apotomopterus) casaleianus jigongshanicus Kleinfeld, 1998 Holotype & Paratype Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) clermontianus Breuning, 1933
Female adult
N Viet-Nam, 100 km NW Hanoi 5.1995 Rapaltovskii E. leg.
Copyright © Alexander Anichtchenko
Carabus (Apotomopterus) datianshanicus Kleinfeld, 1997 Holotype & Paratype Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) davidis davidis Deyrolle & Fairmaire, 1878
Female adult
Copyright © Alexander Anichtchenko
Carabus (Apotomopterus) cantonensis metallaicus Kleinfeld, 1999 Holotype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) davidis neodavidis Kleinfeld & Schutze, 2001 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) davidis viridithoracis Kleinfeld & Schutze, 2001 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) davidis yizhangensis Kleinfeld, 2002 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) delavayi medea Kleinfeld, 1999 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) delavayi tenuimanus Deuve & Imura, 1990
Male adult
Lijiang Yunnan Chine
Copyright © Daniel Prunier
Carabus (Apotomopterus) delavayi tenuimanus Deuve & Imura, 1990
Female adult
Lijiang, Yunnan, Chine
Copyright © Daniel Prunier
Carabus (Apotomopterus) leda Kleinfeld, 2000 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) korellianus Kleinfeld, 2002 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) infirmior homodynamus Deuve, 1994 (as jiucaishan) Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) hera Kleinfeld, 2000 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) guzhangensis Kleinfeld, 1998 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) guizhouensis yunwuensis Kleinfeld, 1996 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) guangxicus Deuve, 1989
Male adult
Guposhan, Hezhou, Guangxi prov. China
male,
photo Daniel Prunier
Copyright © Daniel Prunier
Carabus (Apotomopterus) grossefoveatus grossefoveatus Hauser, 1913 (as zhongnanshan) Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) grossefoveatus annikae Kleinfeld, 1996 Holotype
Male adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) delavayi yunlongensis Kleinfeld, 2000 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) longeantennatus angulihabitus Kleinfeld, 1998 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) longeantennatus changyangensis Kleinfeld, 1998 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) longeantennatus yongshan Kleinfeld, 1999 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) mecynodes helena Kleinfeld, 1999 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) penelope penelope Kleinfeld, 1997 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) rapuzzi rapuzzi Kleinfeld, 2000 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) roxane roxane Kleinfeld, 2002 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) yunanensis yunanensis Born, 1905 Paralectotypes
Female adult
Copyright © Alexander Anichtchenko
Carabus (Apotomopterus) sauteri fujianensis Kleinfeld, 1997 Holotype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) sauteri sanjiangensis Kleinfeld, 2001 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) sauteri tianpingensis Deuve & Yu, 1992 (as eleganticollis chalingensis) Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) sauteri tianpingensis Deuve & Yu, 1992 (as longhuiensis) Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) sauteri tianpingensis Deuve & Yu, 1992 (as peterfrankei) Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) schuetzei Kleinfeld, 1998 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) shun gigantothoracis Kleinfeld, 1997 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) shun paraschuetzei Kleinfeld, 2002 Holotype
Male+Female adult
Copyright © Kirschenhofer
Carabus (Apotomopterus) perangustus Kleinfeld & Schutze, 2003 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) songshanicus Kleinfeld, 1994 Holotype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) tonkinensis pandora Kleinfeld, 2000 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) tonkinensis tonkinensis Deuve, 1990
Female adult
N Viet Nam, Fan-si-pan mt., N seite Cha-pa (=Sapa) 1600 m, 20-30.4.95; 22.17N 103.44E Siniaev leg. (cAA)
Copyright © Alexander Anichtchenko
Carabus (Apotomopterus) torquatus lushanicus Kleinfeld, 1996 Holotype
Male adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) vitalisi bernardlassallei A.Muller, 2005 Paratype
Female adult
Copyright © Boleslav Brezina
Carabus (Apotomopterus) vogtae artemis Kleinfeld, 2000 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) vogtae menghaiensis Kleinfeld, 2000 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) vogtae mojiangensis Kleinfeld, 2000 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) vogtae paravogtae Kleinfeld, 2000 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) vogtae thetis Kleinfeld, 2000 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) yuanshanensis Kleinfeld, 1996 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) yunanensis kassandra Kleinfeld, 1999 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) yunanensis leiboensis Kleinfeld, 1999 Holotype Paratype
Male+Female adult
Copyright © Kirschenhofer
Carabus (Apotomopterus) angermaieri Kleinfeld, 2008 Holotype Paratype
Male+Female adult
China, Guangxi, Rong'an (=Chang'an: 25°13'109°23'), Durongshan, 1500m, V.2007, leg. Li et al. in coll. Kleinfeld
male 30mm, female 32 mm
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) lambrechti Kleinfeld, 2006 Holotype
Male adult
26 mm
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) pseudoguizhouensis Kleinfeld, 2006 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) shun emeifeng Kleinfeld, 2006 Holotype
Male adult
China, Fujian, Taining Co., SW Xinqiao: Emeifeng Shan (ca. 26.59N/117.05E), 1500 m, 34 mm, coll. Kleinfeld
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) delavayi planithoracis Kleinfeld, 2006 Holotype
Male adult
30 mm, <br /> China, Süd-Yunnan, Lüchün (23.00N/102.20E): Shanbashan, 2200 m, 9/2005, coll. Kleinfeld
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) casaleianus casaleianus Deuve, 1994
Female adult
China, SW Hunan Prov., Chengbu Co, Nan-Shan-Ding (26.34;110,08) 1-10.VIII,2006, I.Rapuzzi leg. L=33 mm
Copyright © Ruslan Panin
Carabus (Apotomopterus) dechambreianus dechambreianus Deuve & Li, 2000
Male+Female adult
Gailin, Guanxi prov., CHINA,
17-V-2011 loc. coll.
male - 36 mm, female - 38 mm.
Copyright © Alexandr Safronov
Carabus (Apotomopterus) shun floridus Cavazzuti & Ratti, 1998
Male+Female adult
CHINA, Guangdong prov., Mts. Nanling Reserve, 1-7.VII.2011 loc. coll. male - 38 mm, female - 40 mm.
Copyright © Alexandr Safronov
Carabus (Apotomopterus) kafka Kleinfeld & Reuter, 2010 Holotype
Male adult
China, Guangxi, Mao'er Shan (ca. 25°50N/110°22E), 1600-1900m, 29.V.-10.VI.2009, leg. Chr. Reuter (in coll. Reuter); 25,5 mm
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) eccoptopteroides Rapuzzi, 2012 Holotype Paratype
Male+Female adult
China, South Sichuan province, Pu-Ge County, Lianxiang, Kakaliangzi, 1-11.VI.2012 (deposited in private collection of Rapuzzi)<br /> Image Ivan Rapuzzi, 2012
Copyright © - - -
Carabus (Apotomopterus) francottei Rapuzzi, 2012 Holotype
Male adult
China, North Guangdong province, Mts. Nanling Shan, VI.2009 (deposited in collection of Rapuzzi). Image Ivan Rapuzzi, 2012
Copyright © - - -
Carabus (Apotomopterus) davidis davidis Deyrolle & Fairmaire, 1878
Male adult
SE-China, Jiangxi prov., Yushan County, Mt Wangjing, VI,2012
L=38 mm
Copyright © Ruslan Panin
Carabus (Apotomopterus) ngi ngi Deuve, 1994 Holotype
Female adult
in coll Lassalle
Copyright © Bernard Lassalle
Carabus (Apotomopterus) arrowi hector Hauser, 1924 Holotype
Female adult
Hunan, Liang to Tang; Ap. hector G.H. Typ; Zoolog. Museum Berlin. Museum für Naturkunde Berlin, Humboldt-Universität; 37 mm
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) yaophilus Deuve, 1990 (as dayaoshan) Holotype
Female adult
C-Guangxi: Dayao Shan: NE-part: Mengshan env.: 24:12N/110:31E 44 mm (in coll. Klienfeld)
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) elektra Kleinfeld, 2004 Holotype Paratype
Male+Female adult
29,5 mm
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) roxane miyi Kleinfeld, 2005 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) kouanping kouanping Maindron, 1906 Holotype
Female adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC158 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec158
Copyright © - - -
Carabus (Apotomopterus) arrowi shimenensis Deuve, 1997
Female adult
Qiangqiang, Sichuan, Chine. 40 mm
Copyright © Alain Mollard
Carabus (Apotomopterus) arrowi arrowi Hauser, 1913
Female adult
Simianshan, Guizhou, Chine. 31 mm
Copyright © Alain Mollard
Carabus (Apotomopterus) prattianus prattianus Bates, 1890
Female adult
Yichang, Hubei, Chine. 37 mm
Copyright © Alain Mollard
Carabus (Apotomopterus) prattianus prattianus Bates, 1890
Male adult
Wufeng, Hubei, Chine. 35 mm
Copyright © Alain Mollard
Carabus (Apotomopterus) breuningianus breuningianus Le Moult, 1930
Female adult
SC-China, Guangxi Reg., Pingle County, Mt. Shizishan h=1290 m,5-20,VII.2011, local coll. leg. L=43 mm
Copyright © Ruslan Panin
Carabus (Apotomopterus) penelope penelope Kleinfeld, 1997 Paratype
Male adult
Chine, C-Fujian: SE of Yong’an, Mt.Tianbaoyan, 25°58’ N / 117°22’ E, h = 1100 m, VII.1997, leg. Wong
Copyright © Igor Solodovnikov
Carabus (Apotomopterus) penelope penelope Kleinfeld, 1997 Paratype
Female adult
Chine, C-Fujian: SE of Yong’an, Mt.Tianbaoyan, 25°58’ N / 117°22’ E, h = 1100 m, VII.1997, leg. Wong
Copyright © Igor Solodovnikov
Carabus (Apotomopterus) vitalisi phoopanensis Imura, 2000
Male adult
NE-Laos: Xam Hua Prov.: Mt. Phoo-Pan (Phu Phan), N-slope, h = 2060 m (TOPOTYPUS)
Copyright © Igor Solodovnikov
Carabus (Apotomopterus) vitalisi phoopanensis Imura, 2000
Female adult
NE-Laos: Xam Hua Prov.: Mt. Phoo-Pan (Phu Phan), N-slope, h = 2060 m (TOPOTYPUS)
Copyright © Igor Solodovnikov
Carabus (Apotomopterus) arrowi hector Hauser, 1924 (as arrogantior)
Female adult
Yushan, Hunan, CHINA
Copyright © Alain Mollard
Carabus (Apotomopterus) arrowi trilobatus Cavazzuti, 1994
Male adult
Jinsha, Guizhou, CHINA
Copyright © Alain Mollard
Carabus (Apotomopterus) arrowi yubeicus Deuve & Tian, 1999
Female adult
Guanyang, Guangxi, CHINA
Copyright © Alain Mollard
Carabus (Apotomopterus) aeneocupreus Hauser, 1932
Female adult
Qiong Lai Shan, Sichuan, CHINA
Copyright © Alain Mollard
Carabus (Apotomopterus) prodigus prodigus Erichson, 1834
Male adult
Luofu Shan, Guangdong, CHINA
Copyright © Alain Mollard
Carabus (Apotomopterus) yuae arlequinus Deuve, 1994
Female adult
Yuanbao Shan, Guangxi, CHINA
Copyright © Alain Mollard
Carabus (Apotomopterus) tonkinensis tonkinensis Deuve, 1990
Male adult
North Vietnam, Tû Lé, (Yen Bai Prov.) 1200 m
Copyright © Alain Mollard
Carabus (Apotomopterus) tonkinensis jinpingensis Deuve & Tian, 2001
Female adult
Ailao Shan : Jinping Xian (Yunnan) China
Copyright © Alain Mollard
Carabus (Apotomopterus) mecynodes mecynodes Bates, 1890
Female adult
Wa-shan, Chengdu (Sichuan) China
Copyright © Alain Mollard
Carabus (Apotomopterus) mecynodes combustus Cavazzuti & Ratti, 1999
Female adult
Dabei Shan (Sichuan) China
Copyright © Alain Mollard
Carabus (Apotomopterus) mecynodes lilianae Cavazzuti, 1997
Male adult
Huidong Xian (Sichuan) China
Copyright © Alain Mollard
Carabus (Apotomopterus) arrowi yubeicus Deuve & Tian, 1999
Female adult
S-China, Guangxi, Jinxiu Co., Mt. Dayao,28.VII.2007, L=40 mm
Copyright © Ruslan Panin
Carabus (Apotomopterus) arrowi ghirettii Cavazzuti, 1995
Female adult
Fanjing Shan, Guizhou, 1900 m, Leg: C.Reuter. L= 42 mm
Copyright © Alain Mollard
Carabus (Apotomopterus) davidis davidis Deyrolle & Fairmaire, 1878 Holotype
Female adult
(MNHN-EC-EC137) coldb.mnhn.fr/catalognumber/mnhn/ec/ec137
Copyright © - - -
Carabus (Apotomopterus) delavayi delavayi Fairmaire, 1886 Lectotype
Female adult
(MNHN-EC-EC138) http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec138
Copyright ©
Carabus (Apotomopterus) vitalisi vitalisi Lapouge, 1916 Lectotype
Male adult
Xieng Khouang, Nan-Binh. deposited in MNHN (http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec514)
Copyright © - - -
Carabus (Apotomopterus) davidis helmutschuetzei Kleinfeld, 1995
Male adult
SE-China, SW-Jiangxi: S of Yongxin: Mt. Jinggangshan,V.2012, L=35 mm
Copyright © Ruslan Panin
Carabus (Apotomopterus) jingliae Deuve & Tian, 2019 Holotype
Male adult
Chine, Guangxi, Luzhai Xian, Xiangqiaoyan National Geopark, grotte She Dong, 194 mètres, 24°45’32’’N-109°39’21’’E, courant dans une salle de la partie totalement obscure de la grotte (Cheng Jingli leg., 29 janvier 2019), in coll. South China Agricultural University, Canton (Chine).
Copyright © Thierry Deuve
Carabus (Apotomopterus) bousquetellus luchunicus Deuve, 2019 Holotype
Male adult
Chine, Yunnan, Monts Ailao Shan, Lüchun (MNHN)
Copyright © Thierry Deuve
Carabus (Apotomopterus) caoyutangicus jiangsuensis Deuve, 2019 Holotype
Female adult
Chine, Jiangsu, Changzhou, Longhutang (Jatua), in coll. Jaroslav Turna, Kostelec na Hane (République Tchèque).
Copyright © Thierry Deuve
Carabus (Apotomopterus) shun pseudocantonensis Deuve & Tian, 2019 Holotype
Male adult
Chine, Guangdong, Guangning Xian, Shijian Zhen, Shanxin Cun, en forêt de bambous (Qin Changsheng, 25 mai 2016), in coll. South China Agricultural University (SCAU),
Copyright © Thierry Deuve
Carabus (Apotomopterus) cavazzutiellus neptunus Deuve, 2018 Holotype
Female adult
Chine, Sud-Est Sichuan, Huangjing Forest Park, 1480 mètres, 28°11’N-105°44’E (Jatua), in coll. J. Turna, à Kostelec na Hane (République Tchèque).
Copyright © Thierry Deuve
Carabus (Apotomopterus) longeantennatus acorepoides Deuve, 2018 Holotype
Female adult
Chine, Chongqing, Wulingshan Forest Park, 1690 m, 29°33’N-107°36’E (Jatua), in coll. J. Turna, à Kostelec na Hane (République Tchèque).
Copyright © Thierry Deuve
Carabus (Apotomopterus) yundongbeicus mopanensis Deuve, 2017 Holotype
Male adult
Chine, Yunnan, Mopan Shan, 2300 mètres, 23°57’N-101°58’E (Jatua leg.), in coll. J. Turna, à Kostelec na Hane (République Tchèque).
Copyright © Thierry Deuve
Carabus (Apotomopterus) protenes erlangbaensis Deuve, 2016 Holotype
Male adult
Chine, Guizhou nord occidental, Erlangba, 1200- 1350 m, 28°33’N-106°00’E (Jatua), in coll. Jaroslav Turna, à Kostelec na Hane (République Tchèque).
Copyright © Thierry Deuve
Carabus (Apotomopterus) rugulothorax Deuve, 2016 Holotype
Female adult
Chine, Guizhou nord occidental, Erlangba, 1200-1350 m, 28°33’N-106°00’E (Jatua), in coll. Jaroslav Turna, à Kostelec na Hane (République Tchèque).
Copyright © Thierry Deuve
Carabus (Apotomopterus) nigrovagans Deuve, 2016 Holotype
Male adult
Chine, Guangxi, Jinxiu Xian, Mont Dayao Shan, Laoshan. in coll. J.-L. & S. Remond, à Sainte-Adresse.
Copyright © Thierry Deuve
Carabus (Apotomopterus) nigrovagans Deuve, 2016 Paratype
Female adult
Chine, Guangxi, Jinxiu Xian, Mont Dayao Shan, Laoshan. in coll. J.-L. & S. Remond, à Sainte-Adresse.
Copyright © Thierry Deuve
Carabus (Apotomopterus) argenticornis Rapuzzi, 2020 Holotype
Male adult
China, South Guangxi province, Qinzhou city, Mt. Shiwandashan, 1200 m, V.2020. Image from Rapuzzi, 2020
Copyright © - - -
Carabus (Apotomopterus) longeantennatus transjinshaensis Rapuzzi, 2018 Holotype
Male adult
China, Sichuan, Leibo, Shanlinggang env., VII.2018, Wang legit, Ivan Rapuzzi collection. Image from Rapuzzi, 2018
Copyright © - - -
Carabus (Apotomopterus) benardi briseis Rapuzzi, 2018 Holotype
Female adult
China, Sichuan, Leibo, Shanlinggang environs, VII.2018, Wang legit, Ivan Rapuzzi collection. Image from Rapuzzi, 2018
Copyright ©
Carabus (Apotomopterus) benardi ebian Cavazutti & Rapuzzi, 2010
Male adult
specimen from type locality. Image from Rapuzzi, 2018
Copyright © - - -
Carabus (Apotomopterus) benardi alterego Rapuzzi, 2020 Holotype
Female adult
China, Southeast Sichuan province, leibo, 50 km South from Shanlinggangxiang, Vi.2019, Wang legit. Image from Rapuzzi, 2020
Copyright ©
Carabus (Apotomopterus) yunanensis planielongatus Rapuzzi, 2020 Holotype
Female adult
Southeast Sichuan province, leibo, 50 km South from Shanlinggangxiang, Vi.2019, Wang legit . Image from Rapuzi, 2020
Copyright © - - -
Carabus (Apotomopterus) cyanipennis atripennis Rapuzzi, 2020 Holotype
Male adult
China, Guangxi province, Yuecheng ling, 1800 m, 1/16.V.2008, V. Siniaev legit. Image from Rapuzzi, 2020
Copyright ©
Carabus (Apotomopterus) tonkinensis hagiangensis Rapuzzi, 2020 Holotype
Male adult
Vietnam, Ha Giang province, lung Cu, 1600m, iV.2019. Image from Rapuzzi, 2020
Copyright ©
Carabus (Apotomopterus) patroclus vicarius Cavazzuti, 1996
Male adult
China, S Sichuan, / Yunnan border, W of Lugu lake, S of Yongning vill., h=2700m, 11-18.VI.2017, L=31 mm
Copyright © Ruslan Panin
Carabus (Apotomopterus) arrowi ghirettii Cavazzuti, 1995
Female adult
S-China, NE-Guizhou prov., Fanjing Shan, 1800-2000 m, 5-11.VI.14, 27.54°N 108.42°E , L=40 mm
Copyright © Ruslan Panin
Carabus (Apotomopterus) ludivinae ludivinae Deuve, 1996
Male adult
S-China, SW-Yunnan: Lincang city, Mt. Daxue, 18.IV.2017, L=29 mm
Copyright © Ruslan Panin
Carabus (Apotomopterus) benardi benardi Breuning, 1931 Holotype
Male adult
http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec128
Copyright © MNHN
Carabus (Apotomopterus) prattianus prattianus Bates, 1890 Holotype
Female adult
Ichang (MNHN) http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec80
Copyright © MNHN
Carabus (Apotomopterus) protenes protenes Bates, 1889 Lectotype
Female adult
Ichang (MNHN) http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec81
Copyright © MNHN
Carabus (Apotomopterus) mecynodes mecynodes Bates, 1890
Female adult
NEOTYPE. Sichuan (MNHN) http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec82
Copyright © MNHN
Carabus (Apotomopterus) ichangensis Bates, 1889 Lectotype
Male adult
Ichang (MNHN) http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec83
Copyright © MNHN
Carabus (Apotomopterus) patroclus shiziensis Deuve, 2017 Holotype
Male adult
Chine, Yunnan, Wuding Xian, Shizi Shan, 2270 m, 25°32’N-102°23’E (Jatua), in coll. Jaroslav Turna, à Kostelec na Hane (République Tchèque)
Copyright © Thierry Deuve
Carabus (Apotomopterus) patroclus guzhouensis Deueve, 2017 Holotype
Male adult
Chine, Yunnan, Xinhua, parc forestier de Guzhou, 2010 m, 24°07’N-101°51’E (Jatua), in coll. Jaroslav Turna, à Kostelec na Hane (République Tchèque).
Copyright © Thierry Deuve
Carabus (Apotomopterus) protenes baoxingensis Deuve & Mourzine, 2003
Male adult
Copyright © Igor Solodovnikov
Carabus (Apotomopterus) protenes baoxingensis Deuve & Mourzine, 2003
Female adult
Copyright © Igor Solodovnikov
Carabus (Apotomopterus) yunanensis huizeanus Cavazzuti, 2009 Holotype
Male adult
Mt. SE di Huize, 2400-2800 m. N. Yunnan, Cina meridionale.
Copyright © - - -
Carabus (Apotomopterus) patroclus catagraphus Cavazzuti & Rapuzzi, 2005 Holotype
Male adult
China, Sichuan occidentale, Jiulong pref., Mai-Pu, 3000-3500 m. VII-2004, in coli. Rapuzzi
Copyright © - - -
Carabus (Apotomopterus) tibeticus Cavazzuti & Rapuzzi, 2005 Holotype
Female adult
Tibet orientale, Chamdo pref. (Yi Gong), Po-mi, 2300-2700 m. 20/27-VIII- 2004, in coli. Rapuzzi.
Copyright © - - -
Carabus (Apotomopterus) artemisioides Rapuzzi, 2023 Holotype Paratype
Male+Female adult
Vietnam, Lai Chau, Tam Duong, VII.2022. Image from Rapuzzi, 2023
Copyright © - - -
Carabus (Apotomopterus) tenuicostatus Rapuzzi, 2023 Holotype Paratype
Male+Female adult
Vietnam, Lai Chau, Tam Duong, IX.2022; Image from Rapuzzi, 2023
Copyright © - - -
Carabus (Apotomopterus) tonkinensis tonkinensis Deuve, 1990
Male adult
Vietnam, Lai Chau, Tam Duong, IX.2022. Image from Rapuzzi, 2023
Copyright © - - -
Carabus (Apotomopterus) guangxicus hongjianicus Deuve, 2019 Holotype
Male adult
Chine, Fujian méridional, Hongjian Shan, 1380 m, 25°04’N-117°11’E, in coll. Jaroslav Turna, Kostelec na Hane (République Tchèque).
Copyright © Thierry Deuve
Carabus (Apotomopterus) bousquetellus fengqingensis Deuve, Sota & Liang, 2023 (? bousquetellus) Holotype
Male adult
Chine, Yunnan Prov., Fengqing Distr., col de Guihuashu, 24,67361°N 99,84449°E, 2460 mètres, 11.IX.2004, H. Liang et T. Sota, in coll. Institut Zoologique, Pékin (Chine). Image from Deuve, Sota & Liang, 2023
Copyright © - - -
Carabus (Apotomopterus) anacoreta Rapuzzi, 2024 Holotype
Male adult
Ha Giang Province, Xin Man; Aug. 2023. Image from Rapuzzi, 2024; 31mm
Copyright © - - -
Carabus (Apotomopterus) vietnamicus Rapuzzi, 2023 Holotype Paratype
Male+Female adult
Vietnam, Xin Man, 2000/2100 m, VI.2023. Image from Rapuzzi, 2023; 32 mm
Copyright © - - -
Carabus (Apotomopterus) narcissus Rapuzzi, 2023 Holotype Paratype
Male+Female adult
Vietnam, Xin Man, 2000/2100 m, VI.2023. Image from Rapuzzi, 2023; 33 mm
Copyright © - - -
Carabus (Apotomopterus) tarasovinus toussainti Rapuzzi, 2024 Holotype
Female adult
Vietnam, Nghe An Province, Tuong Duong; Jun. 2023. Image from Rapuzzi, 2024
Copyright © - - -
Carabus (Apotomopterus) tonkinensis tropicus Rapuzzi, 2024 Holotype
Female adult
Vietnam, Da Nang Province, Ba Na hills, top of Mt. Chua; 1480 m asl; Aug. 2023. Image from Rapuzzi, 2024
Copyright © - - -
Carabus (Apotomopterus) delavayi parvipes Cavazzuti, 1996
Male adult
S-Sichuan: S of Yanyuan, Chuandongzi Massif h=3200 m, 27.351206, 101.507036, 8-19.VI.2017, C.Reuter leg., L=32 mm
Copyright © Ruslan Panin
Carabus (Apotomopterus) protenes ascendens Semenov, 1898
Male adult
China, Chongqing, N Jinfo Shan, Shanwangping h=1400m, 29.101944, 107.166389, 2-8.VI.2018, C.Reuter leg./det. F.Kleinfeld, L=26 mm
Copyright © Ruslan Panin
Carabus (Apotomopterus) laoshanicus Imura, 1995
Male adult
China, E Guangxi, Jinxiu Yao County, Tongmu vill., local collector, L=36 mm
Copyright © Ruslan Panin
Carabus (Apotomopterus) korellianus Kleinfeld, 2002
Male adult
China, SW-Yunnan, Dehong Distr., Ruili, Pijiazhai vill., h=1800 m, 24.048111, 97.738222, 9-12.VI.2023, C.Reuter leg., L=31 mm
Copyright © Ruslan Panin
Carabus (Apotomopterus) madefactus Cavazzuti, 1997
Male adult
China, CW-Yunnan, Nánjiàn Yízú Zìzhìxiàn, Mt Lingbao h=2400 m, 24.768611, 100.500000, 6-15.VI.2023, C.Reuter leg./det. F.Kleinfeld, L=27 mm
Copyright © Ruslan Panin
Carabus (Apotomopterus) mecynodes pinchuanensis Imura, 1999
Female adult
China, S-Sichuan, 40-50 km E Xichang (to Zhaojue) 27°51'N 102°30'E, h=2700-3000m, Pine forest, 5-20.VI.2017, C.Reuter leg., L=33 mm
Copyright © Ruslan Panin
Carabus (Apotomopterus) patroclus apricalis Cavazzuti, 1997
Male adult
China, S Sichuan, W of Pingchuanzhen, near Galuo vill., h=2400 m, 7-18.VI.2017, C.Reuter leg./det. F.Kleinfeld, L=30 mm
Copyright © Ruslan Panin
Carabus (Apotomopterus) microtatos lingbaonis Imura, 2019
Male adult
China, CW-Yunnan, Nánjiàn Yízú Zìzhìxiàn, Mt Lingbao h=2400 m, 24.768611, 100.500000, 6-15.VI.2023, C.Reuter leg., L=24 mm
Copyright © Ruslan Panin
Carabus (Apotomopterus) mecynodes pinchuanensis Imura, 1999
Male adult
China, S-Sichuan, 40-50 km E Xichang (to Zhaojue) 27°51'N 102°30'E, h=2700-3000m, Pine forest, 5-20.VI.2017, C.Reuter leg., L= 29 mm
Copyright © Ruslan Panin
Carabus (Apotomopterus) microtatos lingbaonis Imura, 2019
Female adult
China, CW-Yunnan, Nánjiàn Yízú Zìzhìxiàn, Mt Lingbao h=2400 m, 24.768611, 100.500000, 6-15.VI.2023, C.Reuter leg., L=25 mm
Copyright © Ruslan Panin
Carabus (Apotomopterus) patroclus patroclus Breuning, 1933
Male adult
China, S Sichuan, Pass 56 km SW Miannin, near Wawo h=2440 m, 21-24. VI.2017, C.Reuter leg./det. F.Kleinfeld, L=29 mm
Copyright © Ruslan Panin
Carabus (Apotomopterus) bousquetellus bousquetellus Deuve, 1998
Male adult
S-China, CW-Yunnan, W of Lincang, Daxue Shan h=2600m, 23.935556, 100.257778, 7-13.VI.2023, C.Reuter leg./det. F.Kleinfeld, L=24 mm
Copyright © Ruslan Panin
Carabus (Apotomopterus) delavayi patrikeevi Cavazzuti, 2001 Paratype
Male adult
Image courtesy A. Gorodinsky
Copyright © - - -
Carabus (Apotomopterus) dechambreianus yongzhouensis Deuve & Li, 2005
Female adult
Image courtesy A. Gorodinsky
Copyright © - - -
Carabus (Apotomopterus) grossefoveatus pingwuensis Deuve, 1997
Male adult
Image courtesy A. Gorodinsky
Copyright © - - -
Carabus (Apotomopterus) grossefoveatus achilles Hauser, 1924
Male adult
Image courtesy A. Gorodinsky
Copyright © - - -
Carabus (Apotomopterus) malaisei semelai Deuve, 1995
Female adult
Image courtesy A. Gorodinsky
Copyright © - - -
Carabus (Apotomopterus) tuxeni Mandl, 1979
Female adult
Image courtesy A. Gorodinsky
Copyright © - - -
Carabus (Apotomopterus) yaophilus Deuve, 1990
Female adult
Image courtesy A. Gorodinsky
Copyright © - - -
Carabus (Apotomopterus) cheni andreasmuelleri Lassalle, 2004 Holotype
Male adult
Vietnam sept., Ha Giang, Mt. Phu Tha Ca, 1850m, v.2004, Lassalle leg. (in coll Lassalle)
Copyright © Bernard Lassalle
Carabus (Apotomopterus) ngi guangwushanensis Lassalle, 2003 Paratype
Female adult
below the pass betw. Shanliangqu/Taoyuan (in coll Lassalle)
Copyright © Bernard Lassalle
Carabus (Apotomopterus) infirmior huaihuaensis Deuve, 2010 Holotype
Male adult
Hunan, Huaihua Xian, Liangzhi Shan, 1500m (in coll S. Dacatra)
Copyright © Thierry Deuve
Carabus (Apotomopterus) acorep Lassalle & Prunier, 1993 Holotype
Male adult
Xian: Mao'er Shan
Copyright © Bernard Lassalle
Carabus (Apotomopterus) toulgoeti Deuve, 1989
Male adult
NE Guangxi, Quanzhou Co., ZhenBaoDing. Image from Hackel & Sehnal, 2004
Copyright © - - -
Carabus (Apotomopterus) elliptipennis Deuve, 1995
Male adult
SE Guizhou, Leishan Co., Leigong Mt. Image from Hackel & Sehnal, 2004
Copyright © - - -
Carabus (Apotomopterus) breuningianus maolanensis Imura, Zhou & Su, 1999
Male+Female adult
SE Guizhou, Libo Co., Yaogu env. Image from Hackel & Sehnal, 2004
Copyright © - - -
Carabus (Apotomopterus) breuningianus breuningianus Le Moult, 1930
Male adult
C Guangxi, Guigang Pref., Zhenlongshan Mt. Image from Hackel & Sehnal, 2004
Copyright © - - -
Carabus (Apotomopterus) guibeicus Deuve & Tian, 1999
Female adult
SE Guizhou, Libo Co., Yaogu env. Image from Hackel & Sehnal, 2004
Copyright © - - -
Carabus (Apotomopterus) pangi Deuve & Tian, 1999
Male+Female adult
NE Guangxi, Jiaojiang: Baogai Mt. Image from Hackel & Sehnal, 2004
Copyright © - - -
Carabus (Apotomopterus) rapuzzi bobaiensis Kleinfeld, 2005 Holotype Paratype
Male+Female adult
S-Guangxi: Bobai, 22.13/109.55: Kueishanding, 1100m
Copyright © Frank Kleinfeld
Carabus (Apotomopterus) narcissus Rapuzzi, 2023
Male adult
Vietnam, Ha Giang, 5.2024
Copyright © Alexander Anichtchenko
Carabus (Apotomopterus) tonkinensis Deuve, 1990
Male adult
Vietnam, Yen Bai, 5.2020 (cAA)
Copyright © Alexander Anichtchenko
Carabus (Apotomopterus) tonkinensis Deuve, 1990
Female adult
Vietnam, Lai Chau, 7.2021 (cAA)
Copyright © Alexander Anichtchenko
Carabus (Apotomopterus) tonkinensis Deuve, 1990
Female adult
Vietnam, Yen Bai 5.2014 (cAA)
Copyright © Alexander Anichtchenko
Carabus (Apotomopterus) mecynodes mitis Cavazzuti, 2008 Holotype
Male adult
S-Sichuan: Mianning Xian: Pass SW of Mianning, 2650 m
Copyright © - - -
Carabus (Apotomopterus) vitalisi Lapouge, 1916
Female adult
Central Vietnam, Nghe An, 8.2025 (DUBC)
Copyright © Alexander Anichtchenko
Carabus (Apotomopterus) tamdaoensis Deuve, 2025 Holotype
Male adult
Vietnam septentrional, Province de Vinh Phuc, massif de Tam Dao, 1000 mètres, 21°27,577’N/105°38,489’E (M. Pejcha leg.), in coll. A. Müller, à Vienne (Autriche).
Copyright © Thierry Deuve
Carabus (Apotomopterus) longi maguanensis Deuve, 2025 Holotype
Male adult
Chine, Yunnan, Maguan Xian, Gulinqing, 1800 - 1850 mètres (Th. Deuve et Mingyi Tian leg., 9 août 2008), in coll. South China Agricultural University, Canton (Chine).
Copyright © Thierry Deuve -
Archicarabus taxon page (40 photos)
Carabus (Archicarabus) monticola fontanae Born, 1906
Male+Female adult
N-Italy, Piemonte, Bielmonte, h1350 m, 14-15.VI.2010, Pascal Stefani leg.; female, 23 mm
Copyright © Ruslan Panin
Carabus (Archicarabus) nemoralis lamadridae Born, 1895
Male adult
NW-Spain, León Province, Acebo, V.2007, Pascal Stefani leg.
Copyright © Ruslan Panin
Carabus (Archicarabus) montivagus montivagus Palliardi, 1825
Male+Female adult
SW-Romania: Banat, Caras-Severin Reg., Sasca Montana – Stinapari, 21.V.-5.VI.2010, Krause Rene leg.<br /> female, 26 mm
Copyright © Ruslan Panin
Carabus (Archicarabus) montivagus blandus Frivaldsky, 1865
Female adult
Slovacchia, Plastovce, 16.7.1995 Budka leg. 21 mm
Copyright © Alexander Anichtchenko
Carabus (Archicarabus) montivagus blandus Frivaldsky, 1865
Female adult
female, 23 mm<br /> S-Slovakia, Region de Nitra, Plastovce viil., 1.V.2009, Rudolf Kmeco leg.
Copyright © Ruslan Panin
Carabus (Archicarabus) pseudomonticola Lapouge, 1908 ? nemoralis
Male adult
NE Spain, Catalonia, Sant Jaume de Frontanyà, h=1000 m, V-VI.2010, Pascal Stefani leg.
L=26 мм
Copyright © Ruslan Panin
Carabus (Archicarabus) pseudomonticola Lapouge, 1908
Male adult
Spain, Montseny, Barcelona
19.VI.1986 M. Tomas leg.
male, 21 mm
Copyright © Alexander Anichtchenko
Carabus (Archicarabus) gotschi gotschi Chaudoir, 1846
Male+Female adult
Transcaucasia, Armenia,
Ashotsk distr., Bandivan, 8-21.X.1991 P.V.Kazaryan leg.
female, 20mm
Copyright © Ruslan Panin
Carabus (Archicarabus) nemoralis nemoralis Muller, 1764
Male+Female adult
W Ukraine, NW Podolian,
Lviv reg., Zolochiv distr., near Vilshanitsya vill., Lysa Gora, 16-30.V.2009 R.Panin leg.
female, 26 mm
Copyright © Ruslan Panin
Carabus (Archicarabus) nemoralis prasinotinctus Heyden, 1880
Male+Female adult
AUDE.Pic de Nore, h=1200m.IX.2005, A. Laforgue leg.
female,25mm/male, 24mm
Copyright © Ruslan Panin
Carabus (Archicarabus) nemoralis prasinotinctus Heyden, 1880
Male adult
N Spain, 3 km N Bera de Bidasoa 14.9.2002
Anichtchenko A. leg.
long. 25mm
Copyright © Alexander Anichtchenko
Carabus (Archicarabus) rossii Dejean, 1826
Male adult
Lazio (Roma), Bosco di
Nettuno 22.5.1986 C. Iacovone leg.
male, 23 mm
Copyright © Alexander Anichtchenko
Carabus (Archicarabus) steuartii Deyrolle, 1852
Male adult
Portugal, S. Martinho, 27.5.1987
male, 18 mm
Copyright © Alexander Anichtchenko
Carabus (Archicarabus) victor r Fischer, 1836
Male adult
Anatolia Or.<br /> Artvin Heinz leg.<br /> male, 18 mm
Copyright © Alexander Anichtchenko
Carabus (Archicarabus) victor wiedemanni Menetries, 1836
Male adult
Anatolia, Kocaeli, tra Kandira e Sile VII.2001
Solo Claudio leg.
long. 24mm
Copyright © Alexander Anichtchenko
Carabus (Archicarabus) wiedemanni wiedemanni Menetries, 1836
Male adult
SE-Bulgaria:Burgas env. Tsarevo (Michurin), 10-15.VII.1981, J.Slaba leg.<br /> L=22 mm
Copyright © Ruslan Panin
Carabus (Archicarabus) steuartii Deyrolle, 1852
Female adult
NW-Spain, Pontevedra prov., Serra do Candán Mt range, 1.III.1992, S. Devesa leg./det. S. Sobrino
Copyright © Ruslan Panin
Carabus (Archicarabus) monticola roccae Born, 1923
Male adult
NW-Italy, Liguria: Savona env., Monte San Giorgio, 30.X.1977, Monauzzi leg. L=23 mm
Copyright © Ruslan Panin
Carabus (Archicarabus) phoenix Lapouge, 1924
Male+Female adult
C-Lebanon, Byblos /Jbeil Distr., Kfardebian env., h=1200m, XI.2014, C. Reuter leg. female,23mm
Copyright © Ruslan Panin
Carabus (Archicarabus) victor Fischer, 1836
Female adult
NE-Turkey, Giresun, Sehitler Geçidi (Pass) h~2000 m, 23.VII.2001, S. Dacatra leg., L=19 mm
Copyright © Ruslan Panin
Carabus (Archicarabus) rossii Dejean, 1826
Female adult
NC Italy, Emilia reg., Bologna, San Martino, 29.V.2016, G..L.Agnoli leg. L=30 mm
Copyright © Ruslan Panin
Carabus (Archicarabus) gotschi caramanus Fairmaire, 1886 Holotype
Male adult
(MNHN-EC-EC141) http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec141
Copyright © - - -
Carabus (Archicarabus) alysidotus Illiger, 1798
Female adult
Italy, Reggio Emilia, Albinea env. h=150 m, VI.2018, A. Cartini leg., L=25 mm
Copyright © Ruslan Panin
Carabus (Archicarabus) wiedemanni pseudorugosus Machard, 1994
Male+Female adult
Col Buglan (Mus) TURQUIE VI-1981
Copyright © Alain Mollard
Carabus (Archicarabus) wiedemanni congruus Motschulsky, 1865
Female adult
Turkey, Cankiri, Cerkes - Kursunlu, Atkaracalar 1250m 10.7.1996 Leg Staven and Skoupy (DUBC)
Copyright © Alexander Anichtchenko
Carabus (Archicarabus) wiedemanni wiedemanni (as mussardianus Breuning, 1965)
Female adult
NC-Turkey, Sinop province, Erfelek env., h=200m, 10-27.VII.2014, C. Reuter leg. L=24 mm
Copyright © Ruslan Panin
Carabus (Archicarabus) wiedemanni (as mussardianus Breuning, 1965)
Male adult
NC-Turkey, Sinop province, Erfelek env., h=200m, 10-27.VII.2014, C. Reuter leg. L=23 mm
Copyright © Ruslan Panin
Carabus (Archicarabus) wiedemanni ilgazicus Breining, 1970
Female adult
Ilgaz (Cankiri) TURQUIE VI-1990
Copyright © Alain Mollard
Carabus (Archicarabus) wiedemanni ilgazicus Breining, 1970
Male adult
Ilgaz (Cankiri) TURQUIE VI-1990
Copyright © Alain Mollard
Carabus (Archicarabus) wiedemanni turcicus Motschulsky, 1865
Female adult
Camlibel(Tokat) TURQUIE VII-1991
Copyright © Alain Mollard
Carabus (Archicarabus) monticola monticola (as maritimensis Born, 1923)
Female adult
SE France, Alpes-Maritimes, Col de Braus h=1000 m, 23.V-18.VI.2014, P. Cavazzuti & G. Proscia leg., L= 24 mm
Copyright © Ruslan Panin
Carabus (Archicarabus) montivagus vellepiticus Hampe, 1850
Female adult
W Croatia, NP Northern Velebit, Stolak, 44.969492, 14.986169, 23.V-27.VI.2022, L=24 mm
Copyright © Ruslan Panin
Carabus (Archicarabus) montivagus vellepiticus Hampe, 1850
Male adult
W Croatia, NP Northern Velebit, Stolak, 44.969492, 14.986169, 23.V-27.VI.2022, L=23.5 mm
Copyright © Ruslan Panin
Carabus (Archicarabus) victor Fischer, 1836
Male adult
S Georgia, Samtskhe-Javakheti, Akhaltsikhe distr., near Andriatsminda, Anda vill h=1550 m, 14.VII.2021, E. Rutjan leg./ det. R.Panin l=21 mm
Copyright © Ruslan Panin -
Aristocarabus taxon page (13 photos)
Carabus (Aristocarabus) viridifossulatus viridifossulatus Fairmaire, 1887
Female adult
35mm
Copyright © Boleslav Brezina
Carabus (Aristocarabus) viridifossulatus viridifossulatus Fairmaire, 1887
Male adult
29 mm
Copyright © Boleslav Brezina
Carabus (Aristocarabus) viridifossulatus aleatus Cavazzuti, 2003
Male adult
26 mm
Copyright © Boleslav Brezina
Carabus (Aristocarabus) viridifossulatus gonggashanensis Deuve, 1992
Male adult
32 mm
Copyright © Boleslav Brezina
Carabus (Aristocarabus) viridifossulatus jiajiensis Deuve, 1998
Male adult
27 mm
Copyright © Boleslav Brezina
Carabus (Aristocarabus) viridifossulatus kvetoslavae Sehnal, Farkac & Hackel, 2006 Holotype
Female adult
36 mm
Copyright © Boleslav Brezina
Carabus (Aristocarabus) viridifossulatus kvetoslavae Sehnal, Farkac & Hackel, 2006 Paratype
Male adult
30 mm
Copyright © Boleslav Brezina
Carabus (Aristocarabus) viridifossulatus kvetoslavae Sehnal, Farkac & Hackel, 2006 Paratype
Female adult
35 mm
Copyright © Boleslav Brezina
Carabus (Aristocarabus) viridifossulatus romanovi Semenov, 1896
Male adult
27 mm
Copyright © Boleslav Brezina
Carabus (Aristocarabus) viridifossulatus seticollis Deuve & Mourzine, 2000
Male adult
28 mm
Copyright © Boleslav Brezina
Carabus (Aristocarabus) viridifossulatus viridifossulatus Fairmaire, 1887 Holotype
Male adult
Moupin
Copyright © - - -
Carabus (Aristocarabus) romanowi sandaguensis Imura, Zhou & Su, 2004 Holotype
Male adult
mixed forest, 2,875 m in altitude, Sandagu in northwestern Heishui Xian of northern Siclman, Southwest China, 24-26-VI I-200 1, XiaoDong Yu & Hong-Zhang ZHOU leg., in col l. Institute of Zoology, Chinese Academy of Sciences, Beijing. Image from Imura, Zhou & Su, 2004
Copyright © - - -
Carabus (Aristocarabus) viridifossulatus nobilissimus Cavazzuti & Rapuzzi, 2005 Paratype
Female adult
Cina, Sichuan occidentale, Jintang, Jia Ching Shan 3650 m
Copyright © - - - -
Aulonocarabus taxon page (70 photos)
Carabus (Aulonocarabus) canaliculatus jankowskiellus Deuve, 1991
Female adult
CHINA, Jilin prov., Hongshi, forest plant
Copyright © Anton Menstschikov
Carabus (Aulonocarabus) canaliculatus pseudocareniger Deuve, 1991
Male adult
S Primorie, Chernye mts., Poima riv., 30.07.- 6.08.1999, Anichtchenko A. leg.; 25 mm
Copyright © Alexander Anichtchenko
Carabus (Aulonocarabus) gossarei venustoides Deuve, 1989
Male adult
Primorskii reg., Chug. distr., Oblachnaya mt., Zabytyi, 11.7.1977 Kuznetsov V. leg. 19 mm
Copyright © Alexander Anichtchenko
Carabus (Aulonocarabus) kolymensis kodarensis Brinev, 1997
Male adult
NE Chita reg., Kodar mt. rng., up of Chara riv.,
15 km N Leprindo, 2200m 5-15.7.2001 Brinev A. leg.
Copyright © Alexander Anichtchenko
Carabus (Aulonocarabus) kolymensis viridicupreior Deuve & Mourzine, 1993
Male adult
SE Yakutia, Cherskii Mt.rng., Garmychan Mts., 20 km NNE Sasyr, <br /> Kalgar riv., 7.VIII.1998 Nogovitsyna leg.
Copyright © Alexander Anichtchenko
Carabus (Aulonocarabus) kolymensis viridicupreior Deuve & Mourzine, 1993
Female adult
10.VII.2001 NE RUSSIA, Magadan, near Karmaken vill
Copyright © Nil Belonogov
Carabus (Aulonocarabus) kurilensis kurilensis Lapouge, 1913
Male adult
Rusia, Sakhalin isl., 44 kordon
6.08.1991
male, 20mm
Copyright © Alexander Anichtchenko
Carabus (Aulonocarabus) kurilensis pseudodiamesus Ivanovs, 1994
Male adult
Russia, Sakhalin isl., Susunaiskii mg.
Chehov mt., 19.6.1990 Berlov E. leg.
male, 22 mm
Copyright © Alexander Anichtchenko
Carabus (Aulonocarabus) truncaticollis polaris Poppius, 1905
Male adult
Arctic Russia, NW Siberia, Yamal, 230 N from Labytnangi, Panta-Yakha Riv., 15-29.VI.2002; 18 mm.
Copyright © Ruslan Panin
Carabus (Aulonocarabus) truncaticollis polaris Poppius, 1905
Female adult
N.Russia, Polar Ural, Paypudynya River, Pompeliz Mt., h=500-700 m., 10-15.VII.2002 S. Andreev, A. Romanenko leg.
Copyright © Safronov
Carabus (Aulonocarabus) canaliculatus pseudocareniger Deuve, 1991
Male adult
N Korea, Ryanggang Prov., Mt. Tongjom Ridge h~1861m, (128.60E;41.20N) I. Rapuzzi leg.<br /> L=27 mm
Copyright © Ruslan Panin
Carabus (Aulonocarabus) mouthiezianus Deuve, 1991
Male adult
Yakutia, Verhoyanskii Mt.rng., âåðõîâüÿ Êåëå, ð. Íÿìíè, ëåâ. èñòîê. ð. Êîê÷èí 29.7.1994
Copyright © Alexander Anichtchenko
Carabus (Aulonocarabus) truncaticollis tungusensis Obydov, 2010 Holotype
Female adult
Evenk Autonomous Region, N. Tunguska Riv., Tura Vil. env., 12.VII.1979
Copyright © Dmitri Obydov
Carabus (Aulonocarabus) gaschkewitschi czarensis Obydov, 1997
Female adult
Russia, CE-Siberia, N Chita reg., Kodar Mt. Range, Big Leprindo Lake env. h~2000m., 2-5.VII.2002 A. Prokhorov leg./det. A. Anichtchenko
L=19 mm
Copyright © Ruslan Panin
Carabus (Aulonocarabus) canaliculatus cf hailinensis Deuve & Li, 2000
Male adult
NE China, Heilongjiang Prov., Mudanjiang City, Mt.Tai-Gu-Ding-zi h=800m, 15-25.VI.2010. L=29 mm
Copyright © Ruslan Panin
Carabus (Aulonocarabus) truncaticollis kodaricola Deuve, 1998
Female adult
N-Chita reg., Kodar Mt. Rng., h=1800m, Leprindo vill., 1.VII.2006; 17 mm
Copyright © Ruslan Panin
Carabus (Aulonocarabus) kurilensis ? daisetsuzanus Kono, 1936
Male adult
N-Japan, Hokkaido, Katō District,Kamishihoro, Horoka near Maruyama Bridge,19.VI.2013, T. Hirabayashi leg.
L=24 mm
Copyright © Ruslan Panin
Carabus (Aulonocarabus) gaschkewitschi gaschkewitschi Motschulsky, 1859
Male adult
Copyright © Kirill Makarov
Carabus (Aulonocarabus) kolymensis kodarensis Brinev, 1997
Male adult
Russia, SE Siberia, N Chita reg., Kodar Mt. Rng. h=1800m, Novaya Chara vill., 23.VI.2006. L=17 mm
Copyright © Ruslan Panin
Carabus (Aulonocarabus) gossarei huangshuni Kleinfeld & Reuter, 2009 Holotype
Male adult
China, Heilongjiang, Dongning (44°03'N/131°07'E), 22 mm
Copyright © Frank Kleinfeld
Carabus (Aulonocarabus) gossarei huangshuni Kleinfeld & Reuter, 2009 Paratype
Female adult
Copyright © Frank Kleinfeld
Carabus (Aulonocarabus) gossarei huangshuni Kleinfeld & Reuter, 2009 Paratype
Male adult
Copyright © Frank Kleinfeld
Carabus (Aulonocarabus) canaliculatus rufipennis Lapouge, 1910 Lectotype
Male adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC517
Copyright © - - -
Carabus (Aulonocarabus) canaliculatus penialis Lapouge, 1913 Lectotype
Male adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC516
Copyright © - - -
Carabus (Aulonocarabus) canaliculatus biraensis Deuve & Reuter, 2019 Holotype
Male adult
Russie, Extrême-Orient Russe, Oblast Autonome Juif (EAO), environs de Khabarovsk, au nord de Bira, Mont Studentcheskaïa, haut bassin de la rivière Rousskaïa, 1200 mètres, A. Brinev, 12-19.VII. 2014 (MNHN); 20 mm
Copyright © Thierry Deuve
Carabus (Aulonocarabus) gaschkewitschi brinevellus Deuve & Reuter, 2019 Holotype
Male adult
Russie, Sibérie, région de Khabarovsk, district de Verkhneboureinskij, Monts Ezop, 1900 mètres, 52°34’N-133°46’E (A. Brinev, 15-21 juillet 2005), in coll. C. Reuter, à Berlin (Allemagne).
Copyright © Thierry Deuve
Carabus (Aulonocarabus) gaschkewitschi ippolitovi Berlov, 2004 Paratype
Female adult
Иркутская область, Качугский район, заповедник "Байкало-Ленский", Байкальский хребет близ мыса Елохин, каменистая горная тундра на склоне западной экспозиции, 1500 м, 25-VI-2003, leg. О.Берлов. 20 мм. Image courtesy of O. Berlov, at www.zin.ru
Copyright © - - -
Carabus (Aulonocarabus) gaschkewitschi ippolitovi Berlov, 2004 Holotype
Male adult
Иркутская область, Качугский район, заповедник "Байкало-Ленский", Байкальский хребет близ мыса Елохин, каменистая горная тундра на склоне западной экспозиции, 1500 м, 25-VI-2003, leg. О.Берлов. 19 мм. Image courtesy of O. Berlov, at www.zin.ru
Copyright © - - -
Carabus (Aulonocarabus) gaschkewitschi s.str. Lectotype
Male adult
fl. Amur. Image courtesy of D.V. Obydov, at www.zin.ru
Copyright © - - -
Carabus (Aulonocarabus) gaschkewitschi gaschkewitschi Motschulsky, 1859
Male adult
Russia, Khabarovsk reg., Bureinskii Mt. rng., VII.64 det. Obydov. Image courtesy of D.V. Obydov, at www.zin.ru
Copyright © - - -
Carabus (Aulonocarabus) truncaticollis fleischeri Reitter, 1898
Female adult
Russia, N Burajatia, Barguzinskiy Mts., the upper reaches of M.Chivyrkuy riv., h=2000 m, 26.VI-12.VII.2007, Vashchenko S leg. L=18.5 mm
Copyright © Ruslan Panin
Carabus (Aulonocarabus) truncaticollis fleischeri Reitter, 1898
Male adult
Russia, N Burajatia, Barguzinskiy Mts., the upper reaches of M.Chivyrkuy riv., h=2000 m, 26.VI-12.VII.2007, Vashchenko S leg. L=17 mm
Copyright © Ruslan Panin
Carabus (Aulonocarabus) canaliculatus vojnitsi Mandl, 1980 Holotype
Male adult
Image from Xueping et al, 2013
Copyright © - - -
Carabus (Aulonocarabus) canaliculatus rufinus Beheim & Breuning, 1943
Male adult
Mt. Changbai Shan, Antu county, Jilin province, China. 13/15.VI.2012. Er-dao-bai-he, elevation 800 m. Image from Xueping et al, 2013
Copyright © - - -
Carabus (Aulonocarabus) canaliculatus vojnitsi Mandl, 1980
Male adult
Mt. Changbai Shan, Antu county, Jilin province, China. 13/15.VI.2012, Bai-shan-shang-fang, elevation 1850m. Ermans Birch forest. Image from Xueping et al, 2013
Copyright © - - -
Carabus (Aulonocarabus) gossarei mareschii Rapuzzi, 2010 Holotype
Male adult
Askol’d Island (CR). Image from Rapuzzi, 2012
Copyright © - - -
Carabus (Aulonocarabus) kurilensis pseudodiamesus Ivanovs, 1994
Male adult
Mt.Chehova (Sachalin Isl.) VIII-1992; 22 mm
Copyright © Alain Mollard
Carabus (Aulonocarabus) kurilensis rausuanus Ishikawa, 1966
Male adult
Horoka-Kamishihoro-cho,(Kato-Gun,Hokkaido) JAPON, VII-2016; 26 mm
Copyright © Alain Mollard
Carabus (Aulonocarabus) kurilensis rishirensis Nakane, 1957
Female adult
Mt. Rishiri, (Rishiri Isl. Hokkaido) JAPON, VIII-1998; 22 mm
Copyright © Alain Mollard
Carabus (Aulonocarabus) canaliculatus sichotensis Born, 1914
Male adult
Хабаровский край, Нанайский район, гол. Тардоки-Яни, 25-VI-1980, leg. В.Кузнецов. 20mm
Copyright © - - -
Carabus (Tomocarabus) filippovi Rapuzzi, 2023 Holotype
Male adult
Mongolia,Hentei highlands, NE Ulan Bator, VI.2009, Evgenii Filippov legit. (in coll. Rapuzzi). Image from Rapuzzi, 2023. New combination by Anichtchenko 9.9.2023
Copyright © - - -
Carabus (Aulonocarabus) truncaticollis s.str. Eschscholtz, 1833
Female adult
Copyright © Kirill Makarov
Carabus (Aulonocarabus) kurilensis pseudodiamesus Ivanovs, 1994
Male adult
Sakhalin, Makarovskiy Distr., env. Porech'e, 28-VII-1991, leg. E. Berlov; 23 mm.
Copyright © Oleg Berlov
Carabus (Aulonocarabus) kurilensis sichotensis Born, 1914
Male adult
Bureinski NP, up Bureya river, Karbohon lake, tundra 28.7.2006 (cAA)
Copyright © Alexander Anichtchenko
Carabus (Aulonocarabus) kurilensis sichotensis Born, 1914
Male adult
Khabarovsk reg., Verhnebureinskiidistr., Bureinskii NP., Lan riv. 31.5.2007 (cAA)
Copyright © Alexander Anichtchenko
Carabus (Aulonocarabus) canaliculatus careniger Chaudoir, 1863 (praedo)
Male adult
Primorskij Krai., Ussurijsky reg., Kaimanovka vill. 9-15.6.2008 (cAA)
Copyright © Alexander Anichtchenko
Carabus (Aulonocarabus) canaliculatus careniger Chaudoir, 1863 (praedo)
Male adult
Far East, Primorje, Spassk distr., Kalinovka vill. 170m, 44.4589N, 132.9834E, 20-31.7.2016, A. Kozlov leg. (cAA)
Copyright © Alexander Anichtchenko
Carabus (Aulonocarabus) canaliculatus canaliculatus Adams, 1812
Male adult
Buryatia, near Mondy vill., 1600m, 25.6.1979, E. Berlov leg. (cAA)
Copyright © Alexander Anichtchenko
Carabus (Aulonocarabus) canaliculatus jankowskiellus Deuve, 1991
Male adult
China, Liaoning prov., Kuandian Co., Banlaing, Ba-che-chuan 06.2007 (cAA)
Copyright © Alexander Anichtchenko
Carabus (Aulonocarabus) gossarei gossarei Haury, 1879 Holotype
Male adult
Amur or., Sibir or? 22 mm. (coll. Zool. Inst. Univ. Amsterdam)
Copyright © Alexander Anichtchenko -
Axinocarabus taxon page (7 photos)
Carabus (Axinocarabus) fedtschenkoi fedtschenkoi Solsky, 1874
Male adult
27 mm
Copyright © Boleslav Brezina
Carabus (Axinocarabus) fedtschenkoi fedtschenkoi Solsky, 1874
Female adult
30 mm
Copyright © Boleslav Brezina
Carabus (Axinocarabus) fedtschenkoi kondarensis Kryzhanovskij, 1953
Male adult
Tadzhikistan, Varzhobskoe usch., kishlak Navobob
4.1994 Avdeev I. leg.
Copyright © Alexander Anichtchenko
Carabus (Axinocarabus) fedtschenkoi kondarensis Kryzhanovskij, 1953
Female adult
31 mm
Copyright © Boleslav Brezina
Carabus (Axinocarabus) fedtschenkoi miles Semenov, 1886
Female adult
29 mm
Copyright © Boleslav Brezina
Carabus (Axinocarabus) fedtschenkoi kondarensis Kryzhanovskij, 1953
Female adult
NW-Tadzhikistan: Gissar Mt.Rng, Varzob river ravine, Gushari env., h~1400-1500m, 24.III.2010, L=29 mm
Copyright © Ruslan Panin -
Calocarabus taxon page (61 photos)
Carabus (Calocarabus) aristochroides aristochroides Deuve, 1992
Male adult
China Sichuan N-Occ. 25 km Nord Maniganggo, 4050 m, VI.2001 Cavazzuti leg.
Copyright © Pierfranco Cavazzuti
Carabus (Calocarabus) aristochroides qagcaensis Deuve, 1995
Male adult
17 mm
Copyright © Boleslav Brezina
Carabus (Calocarabus) aristochroides qagcaensis Deuve, 1995
Female adult
19 mm
Copyright © Boleslav Brezina
Carabus (Calocarabus) deuveianus Cavazzuti & Casale, 2006 Holotype
Male adult
Cina, Gansu mer., Qiling Shan, Passo a Nord di Tewo 4000m
Guan Gai Shan. Colle mer. 18.VII.2005 Cavazzuti leg.
Copyright © Pierfranco Cavazzuti
Carabus (Calocarabus) deuveianus Cavazzuti & Casale, 2006
Male adult
17 mm
Copyright © Boleslav Brezina
Carabus (Calocarabus) gratus nishidai Deuve & Imura, 1993 Paratype
Male adult
Copyright © Boleslav Brezina
Carabus (Calocarabus) gratus nishidai Deuve & Imura, 1993 Paratype
Female adult
19 mm
Copyright © Boleslav Brezina
Carabus (Calocarabus) guinanensis guinanensis Deuve, 1991
Male adult
16 mm
Copyright © Boleslav Brezina
Carabus (Calocarabus) guinanensis guinanensis Deuve, 1991
Female adult
17 mm
Copyright © Boleslav Brezina
Carabus (Calocarabus) guinanensis maqenicola Kalab, 2005 Paratype
Male adult
China, SE Qinghai mts. 15 km E Maqen, 34.27N/100.24E 4300m
J.Kalab leg., 22-24.7.2004 alpine meadows, screes
Copyright © Pierfranco Cavazzuti
Carabus (Calocarabus) juengerianus Kleinfeld, 1995 Paratype
Male adult
18 mm
Copyright © Boleslav Brezina
Carabus (Calocarabus) juengerianus Kleinfeld, 1995 (as yangguifei)
Male adult
17 mm
Copyright © Boleslav Brezina
Carabus (Calocarabus) juengerianus Kleinfeld, 1995 (as yangguifei)
Male adult
18 mm
Copyright © Boleslav Brezina
Carabus (Calocarabus) juengerianus Kleinfeld, 1995 (as yangguifei) Paratype
Female adult
20 mm
Copyright © Boleslav Brezina
Carabus (Calocarabus) linxiaensis darcangicus Deuve, 1995 Paratype
Female adult
20 mm
Copyright © Boleslav Brezina
Carabus (Calocarabus) linxiaensis darcangicus Deuve, 1995 Paratype
Male adult
18 mm
Copyright © Boleslav Brezina
Carabus (Calocarabus) linxiaensis linxiaensis Deuve, 1992
Male adult
17 mm
Copyright © Boleslav Brezina
Carabus (Calocarabus) linxiaensis linxiaensis Deuve, 1992
Female adult
18 mm
Copyright © Boleslav Brezina
Carabus (Calocarabus) linxiaensis ponggartangensis Deuve, 1995 Holotype
Male adult
17 mm
Copyright © Boleslav Brezina
Carabus (Calocarabus) linxiaensis pseudogratus Deuve, 1994 Paratype
Male adult
19 mm
Copyright © Boleslav Brezina
Carabus (Calocarabus) linxiaensis pseudogratus Deuve, 1994
Male adult
17 mm
Copyright © Boleslav Brezina
Carabus (Calocarabus) linxiaensis pseudogratus Deuve, 1994
Male adult
17 mm
Copyright © Boleslav Brezina
Carabus (Calocarabus) linxiaensis pseudogratus Deuve, 1994
Male adult
17 mm
Copyright © Boleslav Brezina
Carabus (Calocarabus) linxiaensis pseudogratus Deuve, 1994 Paratype
Male adult
China, NW Sichuan, 45 km S Zamtang 31.52N 100.59E,
4600 m, 15-19.7.2001 Rejsek leg.
Copyright © Alexander Anichtchenko
Carabus (Calocarabus) linxiaensis subacidus Cavazzuti, 2002 Holotype
Male adult
China, Sichuan N-Occ., Sertar-Garze, 50 km a Nord del bivio Luhuo-Garze,
4400.VI.2001 Cavazzuti leg.
Copyright © Pierfranco Cavazzuti
Carabus (Calocarabus) turnaianus Deuve, 1995
Male adult
China, Sichuan N-Occ., 20 km Sud di Qagca, Lala Shan 42-4500m
VI.2001 Cavazzuti leg.
Copyright © Pierfranco Cavazzuti
Carabus (Calocarabus) trichothorax trichothorax Brezina & Imura, 1997
Male adult
22 mm
Copyright © Boleslav Brezina
Carabus (Calocarabus) trichothorax trichothorax Brezina & Imura, 1997
Female adult
22 mm
Copyright © Boleslav Brezina
Carabus (Calocarabus) pingpong Brezina & Hackel, 2004 Holotype Paratype
Male adult
CH, S Gansu, SSW Minxian (12) W Qilian Shan
11 km W of Kunda 34:08N/103:45E/ 4000-4300m
alpine zone 9-10.VII.2002 Belousov & Kabak
Copyright © Boleslav Brezina
Carabus (Calocarabus) pingpong Brezina & Hackel, 2004
Male adult
CH, S Gansu, SSW Minxian (12) W Qilian Shan
11 km W of Kunda 34:08N/103:45E/ 4000-4300m
alpine zone 9-10.VII.2002 Belousov & Kabak
Copyright © Boleslav Brezina
Carabus (Calocarabus) pingpong Brezina & Hackel, 2004 Paratype
Female adult
20 mm
Copyright © Boleslav Brezina
Carabus (Calocarabus) pingpong Brezina & Hackel, 2004 Paratype
Male adult
18 mm
Copyright © Boleslav Brezina
Carabus (Calocarabus) przewalskii somoli Deuve, 2007 Holotype
Male adult
16 mm
Copyright © Boleslav Brezina
Carabus (Calocarabus) przewalskii przewalskii Morawitz, 1886 (as elnarae)
Male adult
CINA, Qinghai Nord-Or. Passo fre Daliang e Obo
3600 m, 11.VII.2005 Cavazzuti leg.
Copyright © Pierfranco Cavazzuti
Carabus (Calocarabus) przewalskii przewalskii Morawitz, 1886
Male adult
17 mm
Copyright © Boleslav Brezina
Carabus (Calocarabus) przewalskii przewalskii Morawitz, 1886
Female adult
Copyright © Boleslav Brezina
Carabus (Calocarabus) przewalskii przewalskii Morawitz, 1886
Female adult
19 mm
Copyright © Boleslav Brezina
Carabus (Calocarabus) przewalskii sewertzowi Semenov, 1898 Holotype
Male adult
17 mm
Copyright © Boleslav Brezina
Carabus (Calocarabus) przewalskii sewertzowi Semenov, 1898
Male adult
19 mm
Copyright © Boleslav Brezina
Carabus (Calocarabus) przewalskii sewertzowi Semenov, 1898
Female adult
20 mm
Copyright © Boleslav Brezina
Carabus (Calocarabus) przewalskii paulusianus Deuve, 2007
Male adult
19 mm
Copyright © Boleslav Brezina
Carabus (Cryptocechenus) sifanicus Semenov, 1898 (as oxydocupreus)
Male adult
male, 17mm
Copyright © Boleslav Brezina
Carabus (Cryptocechenus) kalabi Deuve, 1990 (as cuprifuscatus) Paratype
Female adult
Copyright © Boleslav Brezina
Carabus (Calocarabus) worahniki Kleinfeld & Puchner, 2011 Holotype
Male adult
CHINA, Prov. Sichuan, zwischen Garzê und Baiyü, Berg 25km südlich von Garzê, Nordseite, 3780m, /31°32.611N/099°58.596´E, 30.V-14.VI.2010 - leg. et in coll. Puchner
21 mm
Foto Kleinfeld XI.2010
Copyright © Frank Kleinfeld
Carabus (Calocarabus) miraculix Kleinfeld & Puchner, 2011 Holotype
Male adult
CHINA, Prov. Sichuan, zwischen Garzê und Baiyü, Berg 25 km südlich von Garzê, Nordseite, 4100m, leg. Puchner 30.V-14.VI.2010 - N 31°28.298´ E 099°59.930´
24 mm
Foto Kleinfeld XI.2010
Copyright © Frank Kleinfeld
Carabus (Calocarabus) aurocostulus Deuve & Kaláb, 2018 Holotype
Male adult
Chine, Ouest-Sichuan, montagnes à 15 km à l’ouest de Keluodong, 4600-4700 mètres, 31º58’N/98º32’E (pelouses alpines, éboulis), in coll. J. Kaláb, à Jinačovice (République Tchèque).
Copyright © Thierry Deuve
Carabus (Calocarabus) punctatocostulus Deuve & Kaláb, 2018 Holotype
Female adult
Chine, Ouest-Sichuan, montagnes à 15 km à l’ouest de Keluodong, 4600-4700 mètres, 31º58’N/98º32’E (pelouses alpines, éboulis), in coll. J. Kaláb, à Jinačovice (République Tchèque).
Copyright © Thierry Deuve
Carabus (Calocarabus) aristochroides luoxuensis Deauve & Kalab, 2019 Holotype
Male adult
Chine, Sichuan occidental, 4800-4900 mètres, col à 23 km au nord-nord-est de Luoxu, 32º40’N/98º03-04’E (prairies alpines et éboulis). in coll. J. Kaláb, à Jinačovice (République Tchèque)
Copyright © Thierry Deuve
Carabus (Calocarabus) przewalskii menyuanensis Deuve, Liu & Liang, 2024 Holotype
Male adult
Chine, province du Qinghai, canton de Menyuan, ville de Zhugu, Sigou, 2602 m, 37,1353°N 102,3887°E (Yizhou Liu & Wenqi Yin leg., 26.VII.2020, dans des pièges), in IOZ. Image from Deuve 2024
Copyright © - - -
-
Carabus sensu stricto taxon page (112 photos)
Carabus (Carabus) arvensis sylvaticus Dejean, 1826
Female adult
France, Centre reg., Eure-et-Loir, Forкt de Montecot, VI.2009, Pascal Stefani leg.
female, 20 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) arcensis sylvaticus Dejean, 1826 (as sugintensis Brunier, 1922)
Male adult
NW -France, Haut-Rhin Depart., Fulleren, XI.2004, Pascal Stefani leg.
Copyright © Ruslan Panin
Carabus (Carabus) arvensis sylvaticus Dejean, 1826 (as liebmanni Lengerken, 1911 )
Female adult
France 67, Weifersweller 350m, Vosges massif
25.8.91
long. 21mm
Copyright © Alexander Anichtchenko
Carabus (Carabus) arvensis sylvaticus Dejean, 1826
Male adult
France 61, Longny-au-Perche. 30.V.1991
male, 19 mm
Copyright © Alexander Anichtchenko
Carabus (Carabus) arvensis arvensis Herbst, 1784
Male+Female adult
W Ukraine, NW Podolian, Lviv reg., Javoriv distr., near Stradch vill., 20-30.IV.2009, Panin R. leg.
Copyright © Ruslan Panin
Carabus (Carabus) arvensis arvensis Herbst, 1784
Male adult
W Ukraine, Male Polissya, Lviv reg., near Kamenka-Bugzkaya, under the bark of oak tree, in the forest, 29.III.08, Panin R. leg. male,17mm
Copyright © Ruslan Panin
Carabus (Carabus) arvensis arvensis Herbst, 1784
Male adult
W. Ukraine , NW. Podolian, Lviv reg., Nikolaev distr., near Krasiv vill., 9.II.08, Panin R. leg. L=17mm
Copyright © Ruslan Panin
Carabus (Carabus) arvensis conciliator Fischer, 1822 (as jurgitae)
Male adult
Russland, Altai, Aktasch
leg. Nikolaevsky, 1991; det. U.Schmidt, 1995
Foto: U.Schmidt, 2008
Copyright © U.Schmidt
Carabus (Carabus) arvensis baschkiricus Breuning, 1936
Female adult
Chebarkul, Chelyabinsk area<br /> 2.08.2008
Copyright © Evgeniy Chibilov
Carabus (Carabus) arvensis carpathus Born, 1902
Male+Female adult
Ukrainian Carpathians,
Transcarpathian reg., Rakhov Distr., Chornohora range, h=1850 -2000m., 29.VI-28.VII.2007 Panin R. leg.
18mm.
Copyright © Ruslan Panin
Carabus (Carabus) arvensis deyrollei Gory, 1839
Male adult
Spain, Asturias, 1-8.7.1999
long. 17mm
Copyright © Alexander Anichtchenko
Carabus (Carabus) arvensis faldermanni Dejean, 1829
Female adult
Rusia, S Primorskii reg., Poima riv., 30.VII-6.VIII.1999
Anichtchenko A. leg.
long. 22mm
Copyright © Alexander Anichtchenko
Carabus (Carabus) arvensis faldermanni Dejean, 1829
Female adult
Rusia, Evreiskaja AO., near Obluchie 20.6.93
Chichvarhin A. leg.
long. 22mm
Copyright © Alexander Anichtchenko
Carabus (Carabus) granulatus leander Kraatz, 1878
Male+Female adult
Russia, C-Caucasus: Balkaria, Valley of riv. Cherek Balkarskyi, h=1750-1800m 10.VI-20.VII.1992, Zamotajlov A. leg.
female,22mm
Copyright © Ruslan Panin
Carabus (Carabus) granulatus calabricus Spettoli & Vigna Taglianti, 2001
Male adult
Italia, Pollino, San Sever. Luc. Bosco Magnano,
(PZ) 13.X.2002 800m, F.Angelini leg.
long.18mm
Copyright © Alexander Anichtchenko
Carabus (Carabus) granulatus duarius Fischer, 1844
Male+Female adult
SE Siberia, Russia
Irkutsk reg., near Pivovarikha vill., 7.V.07, D.Fominyh leg.
female, 22mm
Copyright © Ruslan Panin
Carabus (Carabus) granulatus granulatus Linne, 1758
Male adult
W Ukraine,
Lviv Reg., Javoriv distr., near Ivano-Frankovo vill., 30.IV-18.V.2009, Panin R. leg.
male, 21mm
Copyright © Ruslan Panin
Carabus (Carabus) granulatus granulatus Linne, 1758
Male adult
23 mm
Germania, Oberfranken, Bad Steben
leg.det. U.Schmidt, 2006
Foto: U.Schmidt, 2008
Copyright © U.Schmidt
Carabus (Carabus) granulatus granulatus Linne, 1758
Female adult
Polonia, Verh. Zalesie, V.1994 Strelnikov D. leg.
long. 21mm
Copyright © Alexander Anichtchenko
Carabus (Carabus) granulatus granulatus Linne, 1758
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Carabus (Carabus) granulatus interstitialis Duftschmid, 1812
Male adult
Austria, Murek, X.1971 leg. Orselli
long. 22mm
Copyright © Alexander Anichtchenko
Carabus (Carabus) granulatus yezoensis Bates, 1883
Male adult
S Sakhalin isl., Novoaleksandrovka, Susun riv., 29.V.90<br /> long. 19mm
Copyright © Alexander Anichtchenko
Carabus (Carabus) menetriesi menetriesi Hummel, 1827
Male adult
W-Ukraine, Roztochia, Lviv Reg., Yavorov distr., near Ivano-Frankovo vill., Site Zalyvki, Panin R. leg., 1-20.V.2005, male, 21 mm.
Copyright © Ruslan Panin
Carabus (Carabus) stscheglowi Mannerheim, 1827
Female adult
E Ukraine,
Dniepropetrovsk reg.,Novomoskovskiy distr. near Andriivka vill.16.VII.06 Kanarsky Yu.leg.
female 23mm
Copyright © Ruslan Panin
Carabus (Carabus) stscheglowi Mannerheim, 1827
Male+Female adult
Central Russia , Voronezh region, near Grafskoe, 12.VIII.1984 long=20mm
Copyright © Ruslan Panin
Carabus (Carabus) sculpturatus Menetries, 1832
Male+Female adult
Azerbaijan,
Talysh Mts. Hirkan National Park, Moscow Forest, 8-9.VII.1988, P.V.Kazaryan leg.
female, 22 mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Carabus) granulatus interstitialis Duftschmid, 1812
Male adult
Italy, Emilia reg., Modena, Castelnuovo Rangone, 31.XII.1991, Sola C. leg.
L=19 mm
Copyright © Ruslan Panin
Carabus (Carabus) arvensis deyrollei Gory, 1839
Female adult
NW Spain, Galicia Reg., Pontevedra Prov., A Lama, Xesta, h~650 m, 1.X.1994
L=17 mm
Copyright © Ruslan Panin
Carabus (Carabus) arvensis conciliator Fischer, 1822 (as jurgitae Paratype) Paratype
Male adult
Russia, Altai, Aktash, VI.1992, V.Shott leg.
Copyright © Dmitri Obydov
Carabus (Carabus) arvensis conciliator Fischer, 1822
Male adult
Russia, Barguzin Nat. res., Davshe vill env., hot springs, 14.VII.1990, T. Ananina leg.
Copyright © Dmitri Obydov
Carabus (Carabus) arvensis hokkaidoensis Lapouge, 1924
Male adult
Russia, South Kuriles ils., Kunashir isl., S slope of Golovinka volcan, 28.VII.1990, S. Pushkarev leg.
Copyright © Dmitri Obydov
Carabus (Carabus) arvensis klitini Obydov, 2007 Holotype
Male adult
Russia, South Kuriles, Urup isl., Schukina bay, spring 700 м SW Vasina, 24-31.VII.2006, A. Klitin leg.
Copyright © Dmitri Obydov
Carabus (Carabus) arvensis klitini Obydov, 2007 Paratype
Female adult
Russia, South Kuriles, Urup isl., Schukina bay, spring near Vasina, 22-24.VII.2006, A. Klitin leg.
Copyright © Dmitri Obydov
Carabus (Carabus) arvensis faldermanni Dejean, 1829
Male+Female adult
Russia, Far East, Primorskiy Kray, Ussuriysk Distr., Kaymanovka vill., 11.VII-1.VIII..2011, A. Korshunov leg.
female, 23 mm
Copyright © Ruslan Panin
Carabus (Carabus) vinctus Weber, 1801
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -
Carabus (Carabus) arvensis carpathus (as pseudonoricus Kenyery, 1983)
Male adult
Slovenia: Julske Alpy, Podvelka, h=450m, 24.X.2010, M.Janalik leg.
L=19 mm
Copyright © Ruslan Panin
Carabus (Carabus) granulatus yezoensis Bates, 1883
Male adult
Russia, S-Sakhalin isl., Makarov distr., Mt. Chekhova h=400m, 12.VI.1998, M. Nesterov leg.<br /> L=23 mm
Copyright © Ruslan Panin
Carabus (Carabus) stscheglowi Mannerheim, 1827
Male adult
S Ukraine, Crimea,<br /> Chatyr-Dag jayla, h=1000m, 11-23.VIII.2000, Solodovnikov I. leg.<br /> L=19 mm
Copyright © Ruslan Panin
Carabus (Carabus) arvensis hokkaidoensis Lapouge, 1924
Male adult
N Japan: Hokkaido, Hamamasu env.,1-5.VI.2002, Eiji Yamauchi leg.
L=20 mm
Copyright © Ruslan Panin
Carabus (Carabus) granulatus granulatus Linne, 1758
Male adult
W-Ukraine, Roztochia,
Lviv Reg., Javorov distr., near Ivano-Frankovo vill., Site Zalyvki, 17.IV-3.V.2010, Panin R. leg.
L=20 mm
Copyright © Ruslan Panin
Carabus (Carabus) menetriesi pacholei Sokolar, 1911
Male+Female adult
Czech Republic, SW Bohemia, Šumava-Volary, 01.V.2007, Slavoj Budka leg.
Copyright © Ruslan Panin
Carabus (Carabus) arvensis venetianus Bernau, 1914
Male adult
N Italy, Veneto, Dolomiten Mts.; 13mm
Copyright © Aleš Sedláček
Carabus (Carabus) arvensis venetianus Bernau, 1914
Female adult
N Italy, Veneto, Dolomiten Mts.; 15mm
Copyright © Aleš Sedláček
Carabus (Carabus) arvensis arvensis Herbst, 1784 (as noricus Sokolar, 1910)
Female adult
CZ, C Moravia, Dřevohostice - Líšná, 13mm
Copyright © Aleš Sedláček
Carabus (Carabus) arvensis carpathus Born, 1902
Male adult
as <i>eremita </i>Fischer, 1823<br /> NW Slovakia, Malá Fatra Mts., 12mm
Copyright © Aleš Sedláček
Carabus (Carabus) arvensis deyrollei Gory, 1839
Female adult
NW Spain, Lugo env./Sarria/; 15mm
Copyright © Aleš Sedláček
Carabus (Carabus) granulatus interstitialis Duftschmid, 1812
Female adult
S Austria, Kärnten, Vellach; 17mm
Copyright © Aleš Sedláček
Carabus (Carabus) arvensis deyrollei Gory, 1839
Male adult
NW-Spain,
Cantabria, Udías env., Monte Corona h=100m,14.VI.2002, Javier de Diego leg.
L=16 mm
Copyright © Ruslan Panin
Carabus (Carabus) granulatus granulatus (as hibernicus Lindroth, 1956)
Female adult
W-Ireland, County Mayo, Letterneevoge (R312),16-30.VI.2013, J.Stypula leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Carabus) granulatus yezoensis Bates, 1883
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Carabus (Carabus) menetriesi menetriesi Hummel, 1827
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Carabus (Carabus) stscheglowi Mannerheim, 1827
Male adult
Russian, Chuvashia, Cheboksary, Zavolzhye distr., , near lake Astrakhanka, L=27 mm
Copyright © Ruslan Panin
Carabus (Carabus) arvensis carpathus Born, 1902
Male adult
W Ukraine, NE Carpathians,Transcarpathian reg., Vihorlat, near Strychava vill., Mt Knyaginytsya h= 500 m, 18.V-22.VI. 2014, Yu.Kanarsky leg. L=18 mm
Copyright © Ruslan Panin
Carabus (Carabus) arvensis arvensis Herbst, 1784
Female adult
NW Ukraine, Polesia, Rovno reg., Rokytne distr., near Bil’sk vill., 4-6.VII.2015, Panin R. leg. L=20 mm
Copyright © Ruslan Panin
Carabus (Carabus) sculpturatus (as gilanicus Mandl, 1967)
Male adult
N-Iran:Gilan Prov., Asalem Distr., Siahbil, deciduous forest (very wet) h= -10 m, 20-24.VI.2017, V. Čutka leg., L=19 mm
Copyright © Ruslan Panin
Carabus (Carabus) sculpturatus (as gilanicus Mandl, 1967)
Female adult
N-Iran:Gilan Prov., Asalem Distr., Siahbil, deciduous forest (very wet) h= -10 m, 20-24.VI.2017, V. Čutka leg., L=22 mm
Copyright © Ruslan Panin
Carabus (Carabus) vinctus Weber, 1801
Male adult
LOUISIANA, East Baton Rouge Parish, 1.5mi W Denham Springs, 30.482 ̊ N 90.981 ̊ W, bottomland hardwood forest, hand collected, 4 Sep 2017
Copyright © Igor Sokolov
Carabus (Carabus) arvensis sachalinensis Lapouge, 1906
Male adult
Kuriles isl., Kunashir
Copyright © Kirill Makarov
Carabus (Carabus) granulatus granulatus Linne, 1758
Male adult
ugly specimens
Copyright © Igor Solodovnikov
Carabus (Carabus) granulatus granulatus Linne, 1758
Male adult
Carabus (s.str.) granulatus granulatus ab. wimmeli Schulz, 1900 S-Belarus, Brest reg., env. Tomashovka, pine forest, 26.VI-13.VII.2022, Kuznetzov V.A.
Copyright © Igor Solodovnikov
Carabus (Carabus) granulatus granulatus Linne, 1758
Female adult
S-Belarus, Brest reg., near vill. Tomashovka
Copyright © Igor Solodovnikov
Carabus (Carabus) vanvolxemi Putzeys, 1875
Male adult
Japan, Tochigi pref., Nikko city, Dorobu 17.8.2000
Copyright © Alexander Anichtchenko
Carabus (Carabus) goryi Dejean, 1831
Male adult
WV, Pacahontas Co, 6mi N of Durbin, 25.6.2010 (cAA)
Copyright © Alexander Anichtchenko
Carabus (Carabus) arcensis arcensis Herbst, 1784
Female adult
Lower Austria, Baden, Klausen-Leopoldsdorf, 15.XII.2024, M. Egger leg. L=22 mm
Copyright © Ruslan Panin
Carabus (Carabus) granulatus calabricus Spettoli & Vigna Taglianti, 2001
Female adult
S-Italy: Calabria, Nat.Park Aspromonte, near Gambarie h=1340 m, 38.158149, 15.823323, 3-6.IV.2025, M.Egger leg., L=22 mm
Copyright © Ruslan Panin
Carabus (Carabus) granulatus calabricus Spettoli & Vigna Taglianti, 2001
Male adult
S-Italy: Calabria, Nat.Park Aspromonte, near Gambarie h=1340 m, 38.158149, 15.823323, 3-6.IV.2025, M.Egger leg., L=19 mm
Copyright © Ruslan Panin
Carabus (Carabus) menetriesi menetriesi Hummel, 1827
Male adult
W Ukraine, Lviv city, Landscape Reserve “Peatbog Bilogorshcha” NW part, 17.IV-14.V.2018, Panin R. leg., L=21.5 mm
Copyright © Ruslan Panin -
Cathaicus taxon page (7 photos)
Carabus (Cathaicus) brandti aishanensis Kleinfeld, 1999 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Cathaicus) brandti brandti Faldermann, 1835
Male adult
China, Beijing, Zhongguanchun, in the flower bed, 2007, Liu ye leg.;<br /> Photo & det. by liuye
Copyright © Ye Liu
Carabus (Cathaicus) brandti brandti Faldermann, 1835 (as grossegranulatus) Syntypes Lectotype
Male adult
C. (Cathaicus) brandti grossegranulatus Breuning 1943<br /> Lectotypus/SyntypusLectotypus (25 mm) von "South Mandschou". Syntypus (25 mm) "South Mandschou, 30. V. 1937"
Copyright © Frank Kleinfeld
Carabus (Cathaicus) brandti brandti Faldermann, 1835 Syntypes Lectotype
Male adult
C. (Cathaicus) brandti grossegranulatus Breuning 1943
Lectotypus/SyntypusLectotypus (25 mm) von "South Mandschou". Syntypus (25 mm) "South Mandschou, 30. V. 1937"
Copyright © Frank Kleinfeld
Carabus (Cathaicus) brandti reginae Deuve, 1997 Holotype
Male adult
Ex Ningxia, Suyuko, Helan Shan 2800 m, 2.VII.1996.
MNHN Paris. 30 mm
Copyright © Frank Kleinfeld -
Cathoplius taxon page (14 photos)
Carabus (Cathoplius) asperatus asperatus (Dejean, 1826)
Male adult
male, 24mm
Copyright © Boleslav Brezina
Carabus (Cathoplius) asperatus asperatus (Dejean, 1826)
Female adult
female, 26mm
Copyright © Boleslav Brezina
Carabus (Cathoplius) asperatus ifniensis (Zarco, 1941)
Male adult
male, 30mm
Copyright © Boleslav Brezina
Carabus (Cathoplius) asperatus ifniensis (Zarco, 1941)
Male adult
male, 34mm
Copyright © Boleslav Brezina
Carabus (Cathoplius) stenocephalus Lucas, 1866 (as escalerae)
Male adult
male, 29mm
Copyright © Boleslav Brezina
Carabus (Cathoplius) stenocephalus Lucas, 1866 (as escalerae)
Female adult
female, 33mm
Copyright © Boleslav Brezina
Carabus (Cathoplius) asperatus susicus Antoine, 1941
Male adult
male, 27mm
Copyright © Boleslav Brezina
Carabus (Cathoplius) asperatus susicus Antoine, 1941
Female adult
female, 29mm
Copyright © Boleslav Brezina
Carabus (Cathoplius) asperatus asperatus (Dejean, 1826)
Male adult
Morocco, 4 km S Saft 32°10'28"N 009°15'27"E 20-21.III.2011 leg. S.Gusakov L=24.0 mm
Copyright © Kirill Makarov
Carabus (Cathoplius) stenocephalus ifniensis (Zarco, 1941)
Female adult
beach of Sidi Ifni city, 10 m a.s.l., 29°23'52"N; 10°10'40"W, Sidi Ifni province, central Morocco. Image from Ghittino, Busato & Casale, 2015.
Copyright © - - -
Carabus (Cathoplius) aliai Escalera, 1944
Male adult
Tan Tan. Image from Ghittino, Busato & Casale, 2015.
Copyright © - - - -
Cechenochilus taxon page (46 photos)
Carabus (Procechenochilus) kokujewi Semenov, 1898
Female adult
RUSSIA, Karachay-Cherkessia, NW Caucasus Mts., Daut Mt. Range, Iskrovtsev - W.Ullu-Kel Mt. pass circus, h=2800-2600m
Copyright © Alexander Anichtchenko
Carabus (Procechenochilus) ? kokujewi Semenov, 1898
Female adult
Russia, NW Caucasus, Karachay-Cherkessia, Karachaevsk distr., near Uchkula vill., valley Uchkulanichi riv. h=2800-2900m, 25-28.VII.2010, Fominykh D. leg.
female, 23 mm
Copyright © Ruslan Panin
Carabus (Cechenochilus) heydenianus Starck, 1889
Male adult
Caucasian nat. park., S slope Pseashkho mt., 2500 m
21.7.1975 Ovtsharenko V. leg.
Copyright © Alexander Anichtchenko
Carabus (Cechenochilus) heydenianus Starck, 1889
Male+Female adult
NW-Caucasus,
Krasnodar reg., Mostovskoy distr., Skirda Mt Range, N slopes Yatyrgvart Mt. h=2300-2500m., 09-10.VII.2008, D. Fominykh leg.
female,19mm/male,18mm
Copyright © Ruslan Panin
Carabus (Cechenochilus) adangensis Gottwald, 1983
Female adult
female, 18mm
Copyright © Boleslav Brezina
Carabus (Cechenochilus) adangensis Gottwald, 1983
Female adult
Russia, NW-Caucasus:
Karacaj-Cherkessija, Abishira–Akhuba Mt. Range h=2000-2650m, Kyafar-Agur Riv. 14–17.VII.2009, Fominykh D. leg.
female,21mm
Copyright © Ruslan Panin
Carabus (Cechenochilus) lazorum Belousov & Zamotajlov, 1999
Male adult
male, 20mm
Copyright © Boleslav Brezina
Carabus (Cechenochilus) boeberi longiceps Chaudoir, 1846
Male adult
C-Caucasus, N Ossetia, near Kozskii, h=2800,
A.G. Koval leg. 10.08.2001; 12 mm
Copyright © Anton Menstschikov
Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Holotype
Male adult
Russia, Karachai-Circassian Republic, Abishira-Akhuba Mount Range between passes Agur and Mylgvar, 2785–2830 m.
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009
Male adult
male, 17mm
Copyright © Boleslav Brezina
Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Paratype
Male adult
Russia, W Caucasus,
Karacaj-Cherkessija, Abishira-Akhuba Mnt Range, Area of pass Turii h~2650-2850m, 17.VII.2009, Fominykh D. leg.
male,17 mm
photo & coll. Panin R.
Copyright © Ruslan Panin
Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Paratype
Female adult
Russia, Karachai-Circassian Republic, Abishira-Akhuba Mount Range between passes Agur and Mylgvar, 2785–2830 m. PARATYPUS
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009
Female adult
Russia, Karachai-Circassian Republic, Abishira-Akhuba Mount Range between passes Agur and Mylgvar, 2785–2830 m. PARATYPUS
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Paratype
Female adult
Russia, Karachai-Circassian Republic, Abishira-Akhuba Mount Range between passes Agur and Mylgvar, 2785–2830 m.
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Paratype
Male adult
Russia, Karachai-Circassian Republic, Abishira-Akhuba Mount Range between passes Agur and Mylgvar, 2785–2830 m.
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Paratype
Female adult
female, 21mm
Copyright © Boleslav Brezina
Carabus (Cechenochilus) boeberi buschi Semenov & Znojko, 1931
Female adult
female, 19mm
Copyright © Boleslav Brezina
Carabus (Cechenochilus) boeberi buschi Semenov & Znojko, 1931
Female adult
female, 23mm
Copyright © Boleslav Brezina
Carabus (Cechenochilus) boeberi boeberi Adams, 1817
Male adult
male, 18mm
Copyright © Boleslav Brezina
Carabus (Cechenochilus) boeberi buschi Semenov & Znojko, 1931
Male adult
Male, 19 mm.
C Caucasus, Kabardino-Balkaria, upper reach. riv. Adylsu, h=2550, 20-24.07.2008, Leg. E.Derzhinsky
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) boeberi aequaliceps Reitter, 1896
Male adult
Checheno-Ingusheti, 20 kilometres eastward Tebulosmta, 2600-3000 m, 9.VI.1986, Komarov leg.
Specimen from the collection of М. Shestopalova (Moscow).
Copyright © Dmitri Obydov
Carabus (Cechenochilus) adangensis Gottwald, 1983 (as gusevi) Paratype
Male adult
Paratypus: Carabus (Cechenochilus) gusevi Zamotajlov et Koval, 1989; Abkhazia, Dzyshra, 26.VII.1987, I. Belousov leg.
Specimen from the collection of М. Shestopalov (Moscow).
Copyright © Dmitri Obydov
Carabus (Cechenochilus) kokujewi Semenov, 1898
Male adult
Russia, Karachai-Cherkessia, vall. Daut Riv., near Glacier Ryndzhi-Murudzhu, h = 3050 m, 22.VIII.2011, leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) kokujewi Semenov, 1898
Female adult
Russia, Karachai-Cherkessia, vall. Daut Riv., near Glacier Ryndzhi-Murudzhu, h = 3100 m, 22.VIII.2011, leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) kokujewi Semenov, 1898
Male adult
Russia, Karachai-Cherkessia, vall. Daut Riv., near Glacier Ryndzhi-Murudzhu, h = 3050 m, 22.VIII.2011, leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) kokujewi Semenov, 1898
Female adult
Russia, Karachai-Cherkessia, vall. Daut Riv., near Glacier Ryndzhi-Murudzhu, h = 2550 m, 23.VIII.2011, leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) adangensis Gottwald, 1983
Female adult
Russia, Karachai-Cherkessia, Abischira-Akhuba Mt.R., env. vill. Romantic, alpin zone, h = 2470 m, 10.07-22.08.2019, leg. S.M. Pavlyuchuk, A.P. Tatulyan
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) boeberi boeberi Adams, 1817
Female adult
Грузия, Мцхета-Мтианети, к северу от пос. Гудаури, ЮВ отроги г. Байдарская (нижний пик), по ущелью ручья, альпика, под камнями, точка 2, h = 2700 м, 42.495413° N, 44.487844° E, 10.08.2019, leg. И.А., А.И., С.В. Солодовниковы, О.И. Данилова 26 mm
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) boeberi tschitscherini Semenov, 1896
Male adult
Russia, C Caucasus, Ingushetia, S slopes of Skalistyi Mt.R.
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) boeberi tschitscherini Semenov, 1896
Female adult
Russia, C Caucasus, Ingushetia, S slopes of Skalistyi Mt.R.
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) heydenianus Starck, 1889
Male adult
Carabus (Procechenochilus) heydenianus euxinus Semenov, 1896 Abkhasia: Arabika plato (TOPOTYPUS)
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) heydenianus Starck, 1889
Female adult
Carabus (Procechenochilus) heydenianus euxinus Semenov, 1896 Abkhasia: Arabika plato (TOPOTYPUS)
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) boeberi aequaliceps Reitter, 1896
Male adult
Russia, Dagestan, Andiyskiy Mt.R., near Andiyskiye vorota pass, 06.VI–18.VII.2021, leg. D. Fominykh
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) boeberi aequaliceps Reitter, 1896
Female adult
Russia, Dagestan, Andiyskiy Mt.R., near Andiyskiye vorota pass, 06.VI–18.VII.2021, leg. D. Fominykh
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) boeberi buschi Semenov & Znojko, 1931
Male adult
Russia: Kabardino-Balkaria: u.r. Adylsu Riv.
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) boeberi boeberi Adams, 1817
Female adult
Georgia, Mtskheta-Mtianeti, near Jvari Pass, left bank of the Baidara River, near mineral springs, scree, under rocks, h = 2265 m, 42.531180° N, 44.471374° E, 26.07.2025, leg. I.A., S.V. Solodovnikov, 25 mm
Copyright © Igor Solodovnikov
Carabus (Cechenochilus) heydenianus dzhentui Zamotajlov, 1997
Female adult
Copyright © Kirill Makarov -
Cephalornis taxon page (1 photos)
Carabus (Cephalornis) potanini (Semenov, 1887)
Male adult
China, S Gansu, 3400m, Minshan mts.,
Wenxian 8.92, J. M. Bousquet
Copyright © Boleslav Brezina -
Chaetocarabus taxon page (32 photos)
Carabus (Chaetocarabus) arcadicus arcadicus Gistl, 1850
Female adult
female, 35mm
Copyright © Boleslav Brezina
Carabus (Chaetocarabus) arcadicus arcadicus Gistl, 1850
Male adult
male, 29mm
Copyright © Boleslav Brezina
Carabus (Chaetocarabus) arcadicus merlini Schaum, 1861
Male adult
male, 30mm
Copyright © Boleslav Brezina
Carabus (Chaetocarabus) arcadicus merlini Schaum, 1861
Female adult
female, 36mm
Copyright © Boleslav Brezina
Carabus (Chaetocarabus) intricatus bayardi Solier, 1835
Male+Female adult
S-Italy,
Basilicata Reg., Potenza Prov., Latronico, F. Angelini leg. 14.VI.2002
female, 26mm
Copyright © Ruslan Panin
Carabus (Chaetocarabus) intricatus bayardi Solier, 1835
Male adult
male, 25mm
Copyright © Boleslav Brezina
Carabus (Chaetocarabus) intricatus bayardi Solier, 1835
Female adult
female, 27mm
Copyright © Boleslav Brezina
Carabus (Chaetocarabus) intricatus intricatus Linne, 1761 (as bohemicus)
Female adult
female, 33mm
Copyright © Boleslav Brezina
Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Male adult
male, 26mm
Copyright © Boleslav Brezina
Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Male adult
W-Ukraine,
Lviv Reg., Zhydachiv distr., near Vybranivka Vill. Panin R.leg. 12.X.2007
31mm.
Copyright © Ruslan Panin
Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Female adult
36 mm
Fundort: Germania, Oberfranken, Kulmbach
leg.det. U.Schmidt, 1977
Foto: U.Schmidt, 2008
Copyright © U.Schmidt
Carabus (Chaetocarabus) intricatus krueperi Reitter, 1896
Male adult
male, 35mm
Copyright © Boleslav Brezina
Carabus (Chaetocarabus) intricatus krueperi Reitter, 1896
Female adult
female, 38mm
Copyright © Boleslav Brezina
Carabus (Chaetocarabus) intricatus lefebvrei Dejean, 1826
Male adult
male, 28mm
Copyright © Boleslav Brezina
Carabus (Chaetocarabus) intricatus lefebvrei Dejean, 1826
Female adult
female, 30mm
Copyright © Boleslav Brezina
Carabus (Heterocarabus) marietti sapancaensis Blumenthal & Breuning, 1967
Female adult
female, 22mm
Copyright © Boleslav Brezina
Carabus (Heterocarabus) marietti sapancaensis Blumenthal & Breuning, 1967
Male adult
male, 22mm
Copyright © Boleslav Brezina
Carabus (Heterocarabus) marietti ornatus Heinz, 1976 Paratype
Female adult
female, 20mm
Copyright © Boleslav Brezina
Carabus (Heterocarabus) marietti ormanensis Blumenthal & Breuning, 1967
Male adult
male, 21mm
Copyright © Boleslav Brezina
Carabus (Heterocarabus) marietti muchei Breuning, 1961
Male adult
male, 19mm
Copyright © Boleslav Brezina
Carabus (Heterocarabus) marietti inexspectatus Basquin & Darge, 1986
Male adult
male, 20mm
Copyright © Boleslav Brezina
Carabus (Heterocarabus) marietti inexspectatus Basquin & Darge, 1986
Female adult
female, 21mm
Copyright © Boleslav Brezina
Carabus (Heterocarabus) marietti marietti Cristoforis & Jan, 1837 (as charetianus Auvray, 1992)
Male adult
male, 21mm
Copyright © Boleslav Brezina
Carabus (Heterocarabus) marietti marietti Cristoforis & Jan, 1837 (as charetianus Auvray, 1992)
Female adult
female, 23mm
Copyright © Boleslav Brezina
Carabus (Heterocarabus) marietti bischoffi Chaudoir, 1848
Female adult
female, 20mm
Copyright © Boleslav Brezina
Carabus (Heterocarabus) marietti bischoffi Chaudoir, 1848
Male adult
male, 18mm
Copyright © Boleslav Brezina
Carabus (Heterocarabus) marietti bischoffi Chaudoir, 1848
Male adult
male, 19mm
Copyright © Boleslav Brezina
Carabus (Heterocarabus) marietti marietti (as ativsengueni Schweiger, 1962)
Male adult
male, 17mm
Copyright © Boleslav Brezina
Carabus (Chaetocarabus) intricatus krueperi Reitter, 1896
Male adult
CE-Greece: Pieria, Stomio (Larissa) Ossa Mt. 7-14.VII.2009, Krause Rene leg.
L=35 mm
Copyright © Ruslan Panin
Carabus (Chaetocarabus) arcadicus arcadicus Gistl, 1850
Female adult
CE-Greece, Kato Olympos, Karia env., 6-17.VII.2011, Krause Rene leg. L=37 mm
Copyright © Ruslan Panin
Carabus (Chaetocarabus) arcadicus merlini Schaum, 1861
Male adult
S-Greece:Peloponnesos: Taygetos Mt.Rng., 17-23.V.2005, V. Sima leg.
L=30 mm
Copyright © Ruslan Panin
Carabus (Heterocarabus) marietti marietti Cristoforis & Jan, 1837
Female adult
as ativsengueni Schweiger, 1962
NW Turkey, Bolu, Abant Mts.; 16mm
Copyright © Aleš Sedláček
Carabus (Heterocarabus) marietti marietti Cristoforis & Jan, 1837
Male adult
as ativsengueni Schweiger, 1962
NW Turkey, Bolu, Abant Mts.; 14mm
Copyright © Aleš Sedláček
Carabus (Chaetocarabus) arcadicus arcadicus Gistl, 1850
Male adult
Graecia, Pindus, Mt. Velouchi (Timfristos), 1200m, vi.1970-A. Sermet Leg., 30 mm
Copyright © Noël Mal
Carabus (Chaetocarabus) intricatus krueperi Reitter, 1896
Male+Female adult
Graecia : Mt. Ossa, 1000m, 31.x.1975, Leg P. Ratti, Coll. N. Mal
Copyright © Noël Mal
Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Male adult
W Ukraine, Lviv city, Sykhiv, Zubra forest, (Osvytskiy Grove), under bark, 49°47'28.10" N, 24° 2'20.09" E, 10.XI.2017, Panin R leg.
Copyright © Ruslan Panin
Carabus (Chaetocarabus) intricatus bayardi (as silaensis Haury, 1881)
Female adult
S-Italy, Calabria, Sila Grande Massif, Camigliatello h=1040 m, 19.XI.2009, G. Proscia leg. L=31 mm
Copyright © Ruslan Panin
Carabus (Chaetocarabus) intricatus bayardi (as silaensis Haury, 1881)
Male adult
S-Italy, Calabria, Sila Grande Massif, Camigliatello h=1040 m, 19.XI.2009, G. Proscia leg. L=28 mm
Copyright © Ruslan Panin
Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Female adult
W Ukraine, Lviv city, Holosko, Bryukhovytsʹkyi forest 49.886833, 24.012083 under bark, 10.XI.2024, R. Panin leg., L=35 mm
Copyright © Ruslan Panin -
Chrysocarabus taxon page (83 photos)
Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860
Male+Female adult
SW -France,
Pyrenees-Atlantiques, Foret de Sare, XII.2009, Pascal Stefani leg.
female, 31 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860
Female adult
Spain,
Navarra, Sierra de Aralar, Altxueta Mt., 5.V.2006
Anichtchenko A. leg.
L= 17mm
Copyright © Alexander Anichtchenko
Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860
Male adult
Spain,
Pais Vasco, Bera de Bidasoa, Larrun 450m,
9.9.2001 Anichtchenko A. leg.
male, 29mm
Copyright © Alexander Anichtchenko
Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as delacrei Tarrier, 1975)
Male+Female adult
NE Spain, Catalonia, Sant Jaume de Frontanyа, h=1000 m, V-VI.2010, Pascal Stefani leg.
female, 38 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as delacrei Tarrier, 1975)
Male adult
Spain,
Catalunya, San Jaume de Frontana,
May 2001
Copyright © Ruslan Panin
Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826
Female adult
Auriac alt.
500m. (France : Aude) 25.V.200
Copyright © Ruslan Panin
Carabus (Chrysocarabus) olympiae Sella, 1855
Male+Female adult
N-Italy,
Piemonte, Bielmonte, 14-15.VI.1981, Pascal Stefani leg.
female, 34 mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Chrysocarabus) olympiae Sella, 1855
Male adult
Italy, Piemonte, Sessera, 2500m 23.III.1987
Copyright © Alexander Anichtchenko
Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792
Female adult
NE-FRANCE,
Lorraine Reg., Moselle, 15.II.2010, Pascal Stefani leg.
female, 26 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792
Male adult
Austria,
Weissenbach stt., III.1979, Kretschmer leg.
male, 24mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792 (as aureus)
Male adult
France 28, Saint-Eliph, V.1991
female, 25mm
Copyright © Alexander Anichtchenko
Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792
Female adult
France 28, Saint-Eliph, V.1991
female, 24 mm
Copyright © Alexander Anichtchenko
Carabus (Chrysocarabus) auronitens costellatus Gehin, 1882 (as subfestivus Oberthur, 1884)
Female adult
W- FRANCE,
Bretagne Reg., Finistrere, Foret de Coatloc'h, XI.2006
female, 26 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) auronitens costellatus Gehin, 1882
Male+Female adult
C-FRANCE,
Limousin Reg., Corrиze Dep., Aubazines, X.2005, Pascal Stefani leg.
female,26mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) auronitens costellatus Gehin, 1882
Male adult
France 42, Mount de Madeleine, 1300m, les Biefs
18.VIII.1991
male, 19mm
Copyright © Alexander Anichtchenko
Carabus (Chrysocarabus) auronitens cupreonitens Chevrolat, 1861
Female adult
NW-FRANCE,
Basse-Normandie Reg., Orne, Forêt de Bellême, 14.XII.2008, Pascal Stefani leg.
female, 26mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) auronitens cupreonitens Chevrolat, 1861
Male+Female adult
NW-FRANCE,
Basse-Normandie Reg,, Orne, Foret d' Ecouves, V.2009, Pascal Stefani leg.
male, 25 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) auronitens escheri (as laevipennis Seidlitz, 1891)
Male adult
W Ukraine, NE Carpathians Mts.
Transcarpathian, Rakhov Distr., Chornogora range, Petros Mt.h=2010m, 9.VI.2009, R.Panin leg
male, 25 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) auronitens escheri Palliardi, 1825 (ab. opacus Haury, 1878)
Male+Female adult
W Ukraine, NE Carpathians Mts.,<br /> Transcarpathian reg., Rakhov Distr., Chornogora range, h=2020m, alpin zone, 28.VI-22.VIII.2009, Panin R. leg.<br /> female, 27mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) auronitens escheri Palliardi, 1825
Male adult
W.Ukraine,
Carpatians, Grebeniv env., Zelemyanka Mts.range h=1000 m., 9-20.VII.2007, R.Panin leg.
Copyright © Safronov
Carabus (Chrysocarabus) auronitens farinesi Dejean, 1826 (as barthei Lapouge, 1912)
Male+Female adult
SC -France,
Midi-Pyrйnйes reg., Ariиge, Foret de Belesta, XI.2008, J.Nicolas leg.
L=31 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) auronitens festivus Dejean, 1826
Male adult
male, 22mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) auronitens montanus Gehin, 1882
Male+Female adult
SC-FRANCE,
Haute-Garonne, Col de Mentй h~1300m., II.1980. Kretschmer leg.
female, 23mm.
photo/det. Panin R.
Copyright © Ruslan Panin
Carabus (Chrysocarabus) lineatus lateralis Chevrolat, 1840
Male adult
Spain, Villoslada de Cameros, Logroño, 19.6.87
F. Rodriguez leg.
male, 24mm
Copyright © Alexander Anichtchenko
Carabus (Chrysocarabus) lineatus lineatus Dejean, 1826
Female adult
N-Spain,
Navarra, Sierra de Urbasa, Puerto de Opacua, III.2003
female, 27mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) lineatus salmantinus Bolivar, 1922
Male adult
Spain, Candelario, Salamanca, 19.XI.1988
leg. E. Carabajal
long. 24mm
Copyright © Alexander Anichtchenko
Carabus (Chrysocarabus) solieri clairi Gehin, 1885
Male adult
France, Caussols, VII.1984
male, 26 mm
Copyright © Alexander Anichtchenko
Carabus (Chrysocarabus) solieri liguranus Breuning, 1933
Female adult
NW Italy,
Altare, Savona,15.VI.1995, Cartini A.leg.
female, 33mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) solieri solieri Dejean, 1826
Male adult
SE-France: Alpes-Maritimes
Copyright © Safronov
Carabus (Chrysocarabus) splendens Olivier, 1790
Male+Female adult
SE France,
Midi-Pyrйnйes reg., Ariиge, Vaychis, 22.X.2002, A.Laforgue leg.
female, 30mm.
Copyright © Ruslan Panin
Carabus (Chrysocarabus) splendens Olivier, 1790
Male adult
Spain, Pais Vasco, Bera de Bidasoa, 15.9.2001
Anichtchenko A. leg.
male, 30mm
Copyright © Alexander Anichtchenko
Carabus (Chrysocarabus) hispanus Fabricius, 1787
Male adult
FRANCE,
Tarn, Foret de la Gresigne, h=400m.VII.2005 Laforgue leg.
male-30mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) hispanus Fabricius, 1787
Male adult
Auriac alt. 500m.
(France : Aude) 25.V.2000
Copyright © Pascal Stefani
Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896 (as juberriensis Mollard, 1999)
Male+Female adult
ANDORRA,
E-Pyrenees, Juberri, h=1350-1400m, 22.VI-18.VII.2007
female, 33mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) lineatus lateralis Chevrolat, 1840 (as quasilineatus Tarrier, 1975)
Male+Female adult
NW Spain,
E Asturias, Monte de Moal h~1000m., 5.VI.2008, Pascal Stefani leg.
female, 27mm.
Copyright © Ruslan Panin
Carabus (Chrysocarabus) solieri solieri Dejean, 1826
Female adult
SE France, Alpes-Maritimes, Ascros, VI.2011
L=30 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860 (as ursuius Lapouge, 1911)
Male+Female adult
FRANCE (64), Ursuya Mt.
XII-2009, P. Stefani leg.
male - 30 mm, female - 32 mm.
Copyright © Alexandr Safronov
Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896
Male adult
Anyos (ANDORRE), h=1450ms,
5-VI-2010, P. Stefani leg.
body lenght - 32 mm.
Copyright © Alexandr Safronov
Carabus (Chrysocarabus) lineatus salmantinus Bolivar, 1922
Male adult
CW Spain, Cáceres, San Martín de Trevejo,24.VI.2011, A. Diez leg.
L=24 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) lineatus lineatus (as pseudonumancianus Mollard, 2010)
Male adult
NC-Spain, Sierra de Cantabria , Sierra de Toloño Rng, Puerto de Herrera , h=1100m, 19.I,2007 25 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) auronitens escheri Palliardi, 1825 (as laevipennis Seidlitz, 1891)
Male adult
W Ukraine, NE Carpathians,Transcarpathian Region, Rakhiv Distr., Chornogora Mts Range, h=2000 m, alpin zone, V-28.VI..2014, Panin R. leg.
Copyright © Ruslan Panin
Carabus (Chrysocarabus) auronitens festivus Dejean, 1826 (ab. holochrysum)
Female adult
SC-FRANCE, Tarn, Le Linas h=550 m, XI.2008, Pascal Stefani leg. L=24 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896
Male adult
Andorra, Leg. Gerts, vi.1966, 32 mm
Copyright © Noël Mal
Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as aragonensis Lapouge,1913)
Male adult
Male adult ,Sierra de Montseny (Barcelone) Spain.
Copyright © Alain Mollard
Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as frontanyaensis Mollard,1999)
Female adult
N of Boreda, (Lerida) Spain.
Copyright © Alain Mollard
Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as brevicollis Lapouge,1910)
Female adult
Corbières (Aude) France.
Copyright © Alain Mollard
Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as brevicollis var. viridis)
Female adult
Rennes les bains (Aude) France.
Copyright © Alain Mollard
Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as delacrei Tarrier,1975)
Female adult
Nostra Signora de Queralt, Berga (Barcelona) Spain.
Copyright © Alain Mollard
Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as occideneissimus Meyer,1995)
Male adult
Collado de Boixols, (Lérida) Spain.
Copyright © Alain Mollard
Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as urgelensis Mollard,1999)
Female adult
Sierra del Cadi,Toloriu, (Lérida) Spain.
Copyright © Alain Mollard
Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as jeannei)
Female adult
Sierra de la Mussara (Tarragona) Spain.
Copyright © Alain Mollard
Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as merangensis Mollard,1999)
Male adult
Méranges, near Puigcerda, (Catalunya) Spain
Copyright © Alain Mollard
Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as monserraticus Csiki,1917)
Female adult
Sierra de Montserrat, (Barcelone) Spain
Copyright © Alain Mollard
Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as palensis Mollard,1999)
Male adult
Col de N’hug, (N-Catalunya) Spain.
Copyright © Alain Mollard
Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as quillansis Mollard,1999)
Male adult
Granès (Aude) France.
Copyright © Alain Mollard
Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (f. typ.)
Female adult
Prats de Mollo (Pyr.Or) France
Copyright © Alain Mollard
Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as soredensis Mollard,1999)
Male adult
Ft.de Sorède, Mts. Albères (Pyr.Or.) France
Copyright © Alain Mollard
Carabus (Chrysocarabus) rutilans opulentus Oberthür, 1898
Female adult
East-Sierra de Aigues-Tortes, Tahull,Bohi (Lerida) Spain
Copyright © Alain Mollard
Carabus (Chrysocarabus) rutilans opulentus Oberthür, 1898 (as alinsis, Mollard 1999)
Male adult
Alins
Copyright © Alain Mollard
Carabus (Chrysocarabus) rutilans opulentus Oberthür, 1898 (as turbonensis, Mollard 1999)
Male adult
Massif du Turbon, (Huesca) Spain
Copyright © Alain Mollard
Carabus (Chrysocarabus) rutilans opulentus Oberthür, 1898 (as cotiellensis, Mollard 1999)
Female adult
East-Mt. Cotiella (Huesca) Spain
Copyright © Alain Mollard
Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896 (as casteillensis, Mollard 1999)
Female adult
Cap del Prat d’Orient,Castells (Lerida) Spain
Copyright © Alain Mollard
Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896 (as benesensis, Mollard 1999)
Female adult
S-Sierra de Aigues-Tortes, Malpas-Benes
Copyright © Alain Mollard
Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896 (as pervesensis, Mollard 1999)
Female adult
Col de Pervers, Sierra de St. Gervas
Copyright © Alain Mollard
Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896 (as basetaensis, Mollard 1999)
Female adult
Guils, Sierra de la Baseta (Lerida) Spain
Copyright © Alain Mollard
Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896 (as capdellensis, Mollard 1999)
Female adult
La Torre de Capdella (Lerida) Spain
Copyright © Alain Mollard
Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896 (as juberriensis, Mollard 1999)
Male adult
Cesters, 1600m, Andorra
Copyright © Alain Mollard
Carabus (Chrysocarabus) auronitens kraussi Lapouge, 1898
Female adult
C-Austria, Styria, Ennstal Alps, Totes Gebirge Mt Rng, Mt Loser h~1100 m, 16-20.VII.2014, R.Janak leg. L=27 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) olympiae Sella, 1855
Male adult
Carabus (Chrysocarabus) olympiae n. fucatus Lapouge, 1902; un Italy: Piemont (TO), Bielmonte, h = 800 m.
Copyright © Igor Solodovnikov
Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792
Male adult
C-Poland, Łódź sity, Łagiewniki Forest, in dead logs of Birch, 18-30.XII.2013, D. Nawak leg., L=25 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) solieri bonnetianus Colas, 1936 (loc.typ.)
Female adult
SE-France, Var, Tanneron, Saint-Cassien des Bois, 18.IV.2019, L=32 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) solieri bonnetianus Colas, 1936 (loc.typ.)
Male adult
SE-France, Var, Tanneron, Saint-Cassien des Bois, 18.IV.2019, L= 31 mm
Copyright © Ruslan Panin
Carabus (Chrysocarabus) solieri clairi Gehin, 1885 (as devillei Tarrier, 1972)
Male adult
SE-France, Alpes-Maritimes, Beuil env., h=1550m, 14.VII.1986
Copyright © Ruslan Panin
Carabus (Chrysocarabus) hispanus (f. castaneus Lapouge, 1902)
Male adult
collection Galibert (Museum de Toulouse). Image from Laforgue, 2016
Copyright © - - -
Carabus (Chrysocarabus) bugareti Jeannel, 1943 Holotype
Female adult
Cammazes (MNHN). Image from Laforgue, 2016
Copyright © - - -
Carabus (Chrysocarabus) auronitens kraussi (as vindobonensis Kubik, 1901)
Female adult
Lower Austria, Baden, Klausen-Leopoldsdorf, 15.XII.2024, M. Egger leg., L=30 mm
Copyright © Ruslan Panin -
Coptolabrus taxon page (181 photos)
Carabus (Coptolabrus) formosus nili Deuve, 2005 Paratype
Male+Female adult
C China, Shaanxi, Liupan Mts., Guan Shan Mt. h=800 m, , 34:50N/106:48 E, Team V.Siniaev leg. female, 38 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) formosus elytraviridis Kleinfeld, 2007 Holotype Paratype
Male+Female adult
China, Sichuan, Aba pref., Nanping Co., Jiuzhaigou (ca. 33.17N/103.54E), 2000 m, 30.5.-6.6.1990, coll. Kleinfeld.
HT 32 mm, PT 36 mm
Copyright © Frank Kleinfeld
Carabus (Coptolabrus) smaragdinus yanggangensis Deuve & Li, 2000
Male+Female adult
Mt. Tongnimpeak, h=1520 m, Ryang Gong Province, N.Korea, 1-18.VI.2008
Copyright ©
Carabus (Coptolabrus) smaragdinus mandschuricus Semenov, 1898
Male adult
Russia, Far East, South Primoriye, Kaymanovka village, 10-20.07.2000; P.Gorbunov leg.
Copyright © Anton Menstschikov
Carabus (Coptolabrus) smaragdinus mandschuricus Semenov, 1898
Male adult
Primorje, Lasovskii distr., Chandalaz mt., L.Voita leg. 19.VII.2008
Copyright © Anton Menstschikov
Carabus (Coptolabrus) smaragdinus longipennis Chaudoir, 1863
Male+Female adult
Copyright © Alexandr Safronov
Carabus (Coptolabrus) smaragdinus longipennis Chaudoir, 1863
Female adult
Huolongling, Beitaizi, Huadian City, Jilin Prov., CHINE
1-8.VI.2007
Copyright © Alexandr Safronov
Carabus (Coptolabrus) smaragdinus longipennis (as mucdenensis Hauser, 1921)
Male adult
Mt. Wulongshan, Dandong City,
Liaoning Province, CHINE
25-30.VI.2008
Copyright © Alexandr Safronov
Carabus (Coptolabrus) smaragdinus honanensis Hauser, 1921
Male+Female adult
Mt. Wangwu, Henan Prov., CHINA
03.06.2006
Copyright © Alexandr Safronov
Carabus (Coptolabrus) smaragdinus hoenggandoensis (Kwon & Lee, 1984)
Female adult
Chujado Isl., S. Korea, 21.07.2006; 40 mm.
Copyright ©
Carabus (Coptolabrus) smaragdinus furumiellus Deuve, 1994
Male+Female adult
Hua-Bo-zi-Shan, Huanren County,Liaoning Province, CHINE, 20-30.VI.2008
Copyright ©
Carabus (Coptolabrus) pustulifer shangzhouicus Deuve & Li, 2003
Male+Female adult
Shahezi, Shangzhou City, Shaanxi Province, CHINE 11-22.VI.2008
Copyright ©
Carabus (Coptolabrus) pustulifer pustulifer Lucas, 1869
Female adult
China, Sichuan, Leshan, June,
800m,
female, 40 mm
Copyright © Alexander Anichtchenko
Carabus (Coptolabrus) pustulifer pustulifer Lucas, 1869
Female adult
Chine, C.Sichuan, Ya'an, 07.2005 Local people
Copyright © Igor Solodovnikov
Carabus (Coptolabrus) pustulifer pustulifer Lucas, 1869
Male adult
Chine, C.Sichuan, Ya'an, 07.2005 Local people
Copyright © Igor Solodovnikov
Carabus (Coptolabrus) pustulifer mirificus Kraatz, 1894
Male+Female adult
Mt. Julong, Baokang, Hubei Prov., CHINE
25.VII.2007
Copyright © Alexandr Safronov
Carabus (Coptolabrus) pustulifer mirificus Kraatz, 1894
Male adult
Chine, SW-Hubei, Enshi, VI.2005 Local people
Copyright © Igor Solodovnikov
Carabus (Coptolabrus) pustulifer mirificus Kraatz, 1894
Male adult
Chine, SW-Hubei, Enshi, 06.2005, Local people
Copyright © Igor Solodovnikov
Carabus (Coptolabrus) pustulifer guerryi Born, 1903
Female adult
N Yunnan: Gaoligongshan, h =1300 m, 06.2003 Local people
Copyright © Igor Solodovnikov
Carabus (Coptolabrus) pustulifer (as cyanelongatus Deuve, 2006)
Female adult
Zhagyab, Quamdu City, Tibet<br /> 8-15.VI.2008
Copyright ©
Carabus (Coptolabrus) principalis Bates, 1889
Male+Female adult
Zi-Ding-Shan, h=1406 m, Guiyang County, Binzhou City, Hunan, CHINE
19-29.IV.2008
Copyright © Alexandr Safronov
Carabus (Coptolabrus) principalis Bates, 1889
Male adult
C.Chine, W Hubei, Changyang, VIII.2005,
Det. I.A.Solodovnikov 2006
Copyright © Igor Solodovnikov
Carabus (Coptolabrus) principalis Bates, 1889
Male adult
C.Chine, W Hubei, Changyang, 08.2005,
Det. I.A.Solodovnikov 2006
Copyright © Igor Solodovnikov
Carabus (Coptolabrus) principalis Bates, 1889
Female adult
C.Chine, W Hubei, Changyang, 08.2005,
Det. I.A.Solodovnikov 2006
Copyright © Igor Solodovnikov
Carabus (Coptolabrus) principalis Bates, 1889
Female adult
C Chine, W Hubei, Changyang, 08.2005,
Det. I.A.Solodovnikov 2006
Copyright © Igor Solodovnikov
Carabus (Coptolabrus) nankotaizanus Kano, 1932
Male adult
Taiwan,
Tatumi, h=200 m, 20.02.1971, D.Muting leg.
Det. I.A.Solodovniov 2006
Copyright © Igor Solodovnikov
Carabus (Coptolabrus) nankotaizanus Kano, 1932
Male adult
Taiwan,
Tatumi, h=200 m, 20.II.1971, D.Muting leg.
Det. I.A.Solodovniov 2006
Copyright © Igor Solodovnikov
Carabus (Coptolabrus) lafossei tungchengensis Li, 1993
Male+Female adult
Tiantou, Yuexi County,
Anhui Province, CHINE
2-8.VI.2008
male 45mm, female 50 mm
Copyright © Alexandr Safronov
Carabus (Coptolabrus) lafossei tungchengensis Li, 1993 (as dabieshanus Imura, 1996)
Male adult
Chine, Hubei, Dabieshan: Yuexi,h=700m., VI.2004, Local people
Det. I.A.Solodovniov 2006
Copyright © Igor Solodovnikov
Carabus (Coptolabrus) lafossei tungchengensis Li, 1993 (as dabieshanus Imura, 1996)
Male adult
Chine, Hubei, Dabieshan: Yuexi,h=700m., VI.2004, Local people
Det. I.A.Solodovniov 2006
Copyright © Igor Solodovnikov
Carabus (Coptolabrus) lafossei tiantai Kleinfeld, 1997 Holotype
Female adult
Copyright © Frank Kleinfeld
Carabus (Coptolabrus) lafossei saturatus Hauser, 1913
Female adult
N Fujian, Jung Ling Shan, 18-24.7.96
female 53 mm
Copyright © Alexander Anichtchenko
Carabus (Coptolabrus) lafossei pseudocoelestis Kleinfeld, 1999 Holotype
Male adult
Copyright © Frank Kleinfeld
Carabus (Coptolabrus) lafossei montigradus Hauser, 1920
Female adult
Chine, Anhui, Huangshan, 07.2005 h=700m Local people
Det. I.A.Solodovnikov 2006
Copyright © Igor Solodovnikov
Carabus (Coptolabrus) lafossei montigradus Hauser, 1920
Female adult
Chine, Anhui, Huangshan, 07.2005 h=700m Local people
Det. I.A.Solodovnikov 2006
Copyright © Igor Solodovnikov
Carabus (Coptolabrus) lafossei montigradus Hauser, 1920
Male adult
Chine, Anhui, Huangshan, 07.2005 h=700m Local people
Det. I.A.Solodovnikov 2006
Copyright © Igor Solodovnikov
Carabus (Coptolabrus) lafossei montigradus Hauser, 1920
Male adult
Chine, Anhui, Huangshan, 07.2005 h=700m Local people
Det. I.A.Solodovnikov 2006
Copyright © Igor Solodovnikov
Carabus (Coptolabrus) lafossei lafossei Feisthamel, 1845 (as donckieri Born,1898)
Male adult
Chine, Zhejiang: Zhousan Archipelago
Copyright © Igor Solodovnikov
Carabus (Coptolabrus) lafossei lafossei Feisthamel, 1845 (as donckieri Born,1898)
Male adult
Chine, Zhejiang: Zhousan Archipelago
Copyright © Igor Solodovnikov
Carabus (Coptolabrus) lafossei buchi Hauser, 1913
Male+Female adult
Yushanwu, Anji County,
Zhejiang Province, CHINE
16-29.IV.2008
Copyright © Alexandr Safronov
Carabus (Coptolabrus) lafossei lafossei Feisthamel, 1845
Male adult
China, Zhejiang pro., Liang H.B. coll;
Copyright © Ye Liu
Carabus (Coptolabrus) jankowskii jankowskii Oberthur, 1883
Male adult
N.Korea, Chagang Do, Mt. Langlin, 30.07.2005
Copyright © Anton Menstschikov
Carabus (Coptolabrus) jankowskii benxiensis Imura, 1991
Female adult
CHINA, Jilin prov., Mt. Longgang., 15.07.2005
Copyright © Anton Menstschikov
Carabus (Coptolabrus) jankowskii benxiensis Imura, 1991
Male+Female adult
Miaoyang, Shicheng, Fengcheng City,
Liaoning Province, CHINE
25-30.VI.2008
Copyright © Alexandr Safronov
Carabus (Coptolabrus) jankowskii benxiensis Imura, 1991
Male adult
NE-China,
Liaoning Prov., Donggou, Mt. Guanmen, 15.VI.2009
male, 36mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) ishizukai Deuve & Ohshima, 1989
Male adult
Chine, E.Sichuan, Wushan 07,2003 h=800m, Local people
Det.I.A.Solodovnikov 2006
Copyright © Igor Solodovnikov
Carabus (Coptolabrus) ishizukai Deuve & Ohshima, 1989
Male adult
Chine, E.Sichuan, Wushan 07,2003 h=800m, Local people
Det.I.A.Solodovnikov 2006
Copyright © Igor Solodovnikov
Carabus (Coptolabrus) augustus ignigena Hauser, 1914
Female adult
Chine, CE Hunan, Changde 07.2004 h=700m Local people
Det.I.A.Solodovnikov 2006
Copyright © Igor Solodovnikov
Carabus (Coptolabrus) augustus ignigena Hauser, 1914
Male adult
Chine, CE Hunan, Changde 07.2004 h=700m Local people<br /> Det.I.A.Solodovnikov 2006
Copyright © Igor Solodovnikov
Carabus (Coptolabrus) formosus sunpanensis Semenov, 1898
Male adult
Copyright © Alexander Anichtchenko
Carabus (Coptolabrus) formosus grumorum Semenov, 1898
Female adult
female, 30mm
Copyright © Boleslav Brezina
Carabus (Coptolabrus) formosus grumorum Semenov, 1898
Male adult
male, 24mm
Copyright © Boleslav Brezina
Carabus (Coptolabrus) formosus brockmanni Kleinfeld & Schutze, 1997 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Coptolabrus) formosus bousqueti Deuve, 1993
Male adult
Wenxian, N.Sichuan, CHINA
21.07.2006
Copyright © Alexandr Safronov
Carabus (Coptolabrus) formosus rostandi Deuve, 1991
Male adult
China, N Sichuan province, Venchuan, h=1400 m., 30.VI.2007, A. Gorodinski leg.; L=33 mm
Copyright © Anton Menstschikov
Carabus (Coptolabrus) elysii pulcher Kleinfeld, 1997 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Coptolabrus) elysii magnificens Kleinfeld, 1999 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Coptolabrus) elysii elysii Thomson, 1856 (as idai Kleinfeld, 2000) Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Coptolabrus) elysii elysii Thomson, 1856
Male+Female adult
S. Shennongjia, Fangxian county, Hubei, CHINE
1-11.06.2007
Copyright © Alexandr Safronov
Carabus (Coptolabrus) elysii elysii Thomson, 1856
Female adult
China, Sichuan province, Wanyuan; 15.06.2009; Local people leg.;
L=32 mm.
Copyright © Anton Menstschikov
Carabus (Coptolabrus) augustus pustuliferus Kleinfeld, 1998 Holotype
Male adult
Copyright © Frank Kleinfeld
Carabus (Coptolabrus) augustus cristianofonti Deuve & Font, 1998
Female adult
China, Guizhou Prov., Sinan, 14.V.2009. L= 41mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) augustus chalingxian Kleinfeld, 1998 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Coptolabrus) augustus augustus Bates, 1888
Male+Female adult
Male: Mt. Wugong, Pingxiang, Jiangxi, CHINA. 06.06.2007
Female: Mt. Jinngang, Jiangxi, CHINA. 26.06.2007
Copyright © Alexandr Safronov
Carabus (Coptolabrus) augustus adriaenssensi (as pseudoignigena Kleinfeld, 1996) Holotype
Female adult
Copyright © Frank Kleinfeld
Carabus (Coptolabrus) augustus yangminshanicus Font, 1997
Male adult
Guangxi prov., Gongcheng county, N. Lianhua, CHINA<br /> 9-17.06.2007, L (mail) = 40 mm, L (female) = 45 mm.
Copyright ©
Carabus (Coptolabrus) augustus yangminshanicus Font, 1997
Female adult
Mt. Yangmingshan, Yongzhou City, Hunan, CHINA<br /> 20-28.06.2007
Copyright ©
Carabus (Coptolabrus) fruhstorferi Roeschke, 1900
Male adult
Collection ground Izuhara cho, Nagasaki Pref. 35764
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Carabus (Coptolabrus) pustulifer shangzhouicus Deuve & Li, 2003
Male adult
C-Chine, SE-Shaanxi Province, Shahezi, Shangzhou City, 6-18.VII.2010<br /> L=43 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) smaragdinus dachangensis Lu & Müller, 2009
Female adult
NE-China, Liaoning Prov., Changhai County, Dachangshan Dao Isl.,12.VI.2011
L=40 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) smaragdinus liaodongensis (Li, 1992)
Male adult
NE-China, E-Liaoning: Anshan: Mt.Qian Shan, 9-14.VI.2011. L=39 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) smaragdinus sungkii J.K. Park & J. Park, 2013 Holotype Paratype
Male+Female adult
Holotype male: Is.Gureopdo (37°19′71.42″N, 125°98′72.62″E), Deokjeok archipelago, Gyeonggi-do, 19.viii.2010, J.K.Park; Paratype female: Is. Gureopdo, Deokjeok archipelago, Gyeonggi-do, J.K.Park. Image from original describtion J.K. Park & J. Park, 2013
Copyright ©
Carabus (Coptolabrus) ? lafossei coelestis Stewart, 1845
Female adult
CE-China, Shangai, 3.V.2007 L=34 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) augustus tenuiorithorax Deuve & Li, 2006
Male+Female adult
SE-China, SW-Jiangxi, Mt Guan, V.2009, local coll. female, 47 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) formosus bousqueti Deuve, 1993
Male adult
C-China, S-Gansu prov., Wenxian County, Mt.Guanzigou,1-14.V.2010, local coll. leg.
L=32 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) lafossei saturatus Hauser, 1913
Male adult
SE-China, Jiangxi prov.,Yushan County, Mt Wangjing, VI.2012
40 мм
Copyright © Ruslan Panin
Carabus (Coptolabrus) pustulifer (as cyanelongatus Deuve, 2006)
Male adult
C-Chine, Tibet, Qamdo, Gonjo County, Ke Ri Xiang env., VI.2012, Alain Mollard leg.<br /> L=43 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) formosus brockmanni Kleinfeld & Schutze, 1997
Female adult
C-China, S-Shaanxi: Shangzhou City, Shahezi,1-14.VI.,2013, local coll. leg. L=39 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) smaragdinus dolichognathus Deuve, 1995
Male adult
NW-Korea, North Pyongan Province, Mt. Donggorim, 1-20.VI.2013, local coll. leg.<br /> L=37 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) augustus augustus Bates, 1888
Male adult
SE-China, Jiangxi prov., Yudu County, Ganzhou city, Yun Fengzang h=1000m, 5-20.V.2011
L=43 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) augustus ignigena (as subpustulatus Breuuning 1961) Holotype
Male adult
Hunan, China. ignigena subpustulatus (In coll Lassalle).
Copyright © Bernard Lassalle
Carabus (Coptolabrus) augustus adriaenssensi Lassalle & Prunier, 1993 Holotype
Male adult
Chine, Guangxi, Mts Miao Er Shan, 1800m, 5.92 (in coll B. Lassalle) ? pustulifer adriaenssensi Lassalle & Prunier, 1993
Copyright © Bernard Lassalle
Carabus (Coptolabrus) gemmifer pseudoformosus Deuve, 2006
Female adult
China, Tibet AR, Qamdo, Gonjo County, Kè Rì Xiāng vill.(Kerri Township), h=3100m, VI.2012, A. Mollard leg. L=38 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) ignimitella Bates, 1888
Female adult
China, Yunnan, Tali (error?) Image from: Mal, N., Drumont, A., Kerkhof, S. & P. Grootaert, 2012. Pictorial beetle collection of the Royal Belgian Institute of Natural Sciences.
Copyright © - - -
Carabus (Coptolabrus) smaragdinus monilifer Tatum, 1847 (as tyrannus)
Female adult
Korea, Jeju, Quelpaert Island (= Jeju-Do). Image from: Mal, N., Drumont, A., Kerkhof, S. & P. Grootaert, 2012. Pictorial beetle collection of the Royal Belgian Institute of Natural Sciences.
Copyright © - - -
Carabus (Coptolabrus) gemmifer mesites Fairmaire, 1897 Holotype
Female adult
"Siao-Lou-Lou-Chan, Chasseurs thibétains, 1898". Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC149 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec149
Copyright © MNHN
Carabus (Coptolabrus) elysii rothschildi Born, 1899 Holotype
Female adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC123 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec123
Copyright © - - -
Carabus (Coptolabrus) augustus augustus Bates, 1888 Lectotype
Male adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC105 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec105
Copyright © - - -
Carabus (Coptolabrus) pustulifer pustulifer Lucas, 1869 Holotype
Female adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC163 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec163
Copyright © - - -
Carabus (Coptolabrus) augustus ignigena Hauser, 1914
Female adult
China : Guizhoh, Sinan vi.2006 (Size : 35 mm) Coll. N. Mal
Copyright © Noël Mal
Carabus (Coptolabrus) ignimitella antaeus Hauser, 1914 (as guidongicus Cavazzuti & Ratti, 1998)
Female adult
China : Jianxi, Jing Gangshan City 10-18.vi.2007 - J. Li Leg. (Size : 42 mm) Coll. N. Mal
Copyright © Noël Mal
Carabus (Coptolabrus) pustulifer shangzhouicus Deuve & Li, 2003
Female adult
China : Shaanxi, Shazhou City, Shahezi 10.vii.2007 - J. Li Leg. (Size : 42 mm), Coll. N. Mal
Copyright © Noël Mal
Carabus (Coptolabrus) ignimitella guangxicola Deuve, 1998
Male adult
S-China, Guangxi prov., Jingxiu, Mt. Dayao Shan, 1-17.V.2006, L=47 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) augustus tenuitarsatus Rapuzzi, 2016 Holotype
Male adult
China, Chongqing province, Pengshui county, Mt. Heimending. Image from Rapuzzi, 2016
Copyright ©
Carabus (Coptolabrus) smaragdinus losevi Rapuzzi, 2016 Holotype
Male adult
Far East of Russia, South Primorye, Khasanskiy district, Furugelm Island, 11/13.VII.2013, O. Losev legit. Image from Rapuzzi, 2016
Copyright © - - -
Carabus (Coptolabrus) elysii wangguofeni Rapuzzi, 2016 Holotype
Male adult
China, Anhui province, Taihu, Wangling vill., South slope of Mt. Dabieshan, 400 m, 10/30.IV.2015, (30º31'18'' N; 116º16'39'' E), Xihuangshun legit; preserved in Ivan Rapuzzi collection. Image from Rapuzzi, 2016
Copyright ©
Carabus (Coptolabrus) ignimitella ertli Born, 1910 Holotype
Male adult
Foothow (MNHN-EC-EC136)
Copyright © - - -
Carabus (Coptolabrus) gemmifer gemmifer Fairmaire, 1887 Holotype
Male adult
Yunnan (MNHN) http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec150
Copyright © MNHN
Carabus (Coptolabrus) gemmifer loczyi Csiki, 1927 Holotype
Male adult
Yunnan; reçu de Bang Haes le 23/IV/1912 comme yunnanus (MNHN) http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec124
Copyright © MNHN
Carabus (Coptolabrus) ignimitella antaeus Hauser, 1914 (as guidongicus)
Female adult
China, E-Hunan, Zhuzhou, Yanling County, Wanyang Shan Mts, VI.2003 , L= 47 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) jankowskii fusanus Born, 1907
Female adult
South Korea, S Gyeongsang Prov., Busan, Mt Geumwonsan, 17.VI.2017, L=37 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) formosus wanxianicus Imura, 1995
Male adult
China, NE-Sichuan, Dazhou, Wanyuan env., 6.VI.2005. L = 33mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) nankotaizanus Kano, 1932
Male adult
NE-Taiwan, Yilan County, V.2017, L=39 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) jankowskii fusanus (as obtusipennis)
Female adult
SC-Korea: Chiri-san Massif, Mt. Jirisan, 28.VI.2011, L=35 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) gemmifer mesites Fairmaire, 1897
Male adult
China, Shimian 1200m Shimian Yaan Sichuan VII.2017 (DUBC). det. Anichtchenko A. 2018
Copyright © Alexander Anichtchenko
Carabus (Coptolabrus) elysii xiaoganicus Deuve et Remond, 2016
Male adult
China, Hubei, Xiaogan, Mt Shuangfeng Shan, VII.2016 L=37 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) gemmifer gemmifer (as loczyi Csiki, 1927)
Female adult
S-China, W-Yunnan: Dali city, Mt Jizushan, VI.2018, L=34 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) lafossei buchi Hauser, 1913
Male adult
China Zhejiang Yuyao - Leg. Buch 1925
Copyright © Vittorino Monzini
Carabus (Coptolabrus) smaragdinus furumiellus Deuve, 1994
Female adult
NE-China, E-Liaoning, Benxi, Mt Liuershan, VIII.1998. L=39 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) pustulifer lengchongensis Deuve, 2016 Holotype
Male adult
Chine, Guizhou nord-occidental, Lengchong, 1900 mètres, 27°06’N-105°15’E (Jatua), in coll. Jaroslav Turna, à Kostelec na Hane (République Tchèque).
Copyright © Thierry Deuve
Carabus (Coptolabrus) elysii xiaoganicus Deuve et Remond, 2016 Holotype
Male adult
Chine, Hubei, Xiaogan, Mt Shuangfeng Shan, in coll. J.-L. & S. Remond, à Sainte-Adresse.
Copyright ©
Carabus (Coptolabrus) smaragdinus shantungensis Born, 1910 Holotype
Male adult
Shandong, Qingdao reçu de Bang Haes le 23/IV/1912 . http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec122
Copyright © MNHN
Carabus (Coptolabrus) smaragdinus branickii Taczanowski, 1887
Male adult
N Korea, S. Hamgyong Prov., Mt.Chail Peak, 1-16.VII.2010, l=38 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) fruhstorferi Roeschke, 1900
Male adult
Japan, Tsushima Isl., 20.IX.2018, S. Shinichi leg., L=35 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) smaragdinus euviridis (Ishikawa & Kim, 1983)
Male adult
38 mm, R. of Korea, Jirisan Mt., I. Rapuzzi and L. Caldon leg. (CIR). Image from Rapuzzi, 2015
Copyright © - - -
Carabus (Coptolabrus) smaragdinus euaureus Rapuzzi, 2015 Holotype Paratype
Male+Female adult
R. of Korea, Jirisan Nat. Park, Yeongrieongchi, 1200 m, 7/16.VII.2012, I. Rapuzzi and L. Caldon leg. (CIR). Image from Rapuzzi, 2015
Copyright © - - -
Carabus (Coptolabrus) smaragdinus buangun Rapuzzi, 2015 Holotype Paratype
Male+Female adult
R. of Korea, Jeollabuk-do Province, Buan-gun, Byeonsanbando, 100/200m, 13/20.VII.2012, I. Rapuzzi and L. Caldon leg. (CIR). Image from Rapuzzi, 2015
Copyright © - - -
Carabus (Coptolabrus) jankowskii byeoksanensis Rapuzzi, 2015 Holotype Paratype
Male+Female adult
R. of Korea, Jeollabuk-do Province, Buan-gun, Byeonsanbando, 100/200 m, 13/20.VII.2012, I. Rapuzzi and L. Caldon leg. (CIR). Image from Rapuzzi, 2015
Copyright © - - -
Carabus (Coptolabrus) smaragdinus robinzoni Rapuzzi, 2010 Holotype
Male adult
Reyneke island (CR). Image from Rapuzzi, 2012
Copyright © - - -
Carabus (Coptolabrus) smaragdinus hoenggandoensis (Kwon & Lee, 1984) Holotype
Male adult
deposited in Kyungpook National University, Daegu, Republic of Korea, Scale bar = 5mm
Copyright © Jong-Bong Choi
Carabus (Coptolabrus) smaragdinus hoenggandoensis (Kwon & Lee, 1984) Paratype
Female adult
deposited in Kyungpook National University, Daegu, Republic of Korea, Scale bar = 5mm
Copyright © Jong-Bong Choi
Carabus (Coptolabrus) smaragdinus hongdoensis (Kwon & Lee, 1984) Holotype
Male adult
deposited in Kyungpook National University, Daegu, Republic of Korea, Scale bar = 5mm
Copyright © Jong-Bong Choi
Damaster (Coptolabrus) jankowskii irwolsanensis Kim, 2009 Holotype
Male adult
Mt. Irwol, Yeongyang-gun, Gyeongsangbuk-do, 26, vii, 2007
Copyright © - - -
Damaster (Coptolabrus) jankowskii unmunsanensis Kim, 2009 Paratype
Female adult
Mt. Unmun, Cheongdo-gun, Gyeongsangbuk-do, 12, viii, 2004
Copyright © - - -
Damaster (Coptolabrus) jankowskii ganghwadoensis Kim, 2009 Paratype
Female adult
Mt. Goryeo, Ganghwa-gun, Gyeonggi-do, 16, viii, 2007
Copyright © - - -
Carabus (Coptolabrus) nankotaizanus Kano, 1932 (as shimonoyai Kurosawa & Kudo, 1980)
Male adult
Mt.Lala-Shan, (Fuxing,Taoyuan) TAIWAN (FORMOSE) II-1997
Copyright © Alain Mollard
Carabus (Coptolabrus) augustus ignigena Hauser, 1914 (as lautus)
Male+Female adult
Mt.Yingpan Shan (Qiubei Xian, Yunnan) CHINE, VII-2008
Copyright © Alain Mollard
Carabus (Coptolabrus) ignimitella angulicollis Hauser, 1911
Male+Female adult
Mt.Datian Ding (Baishi,Guangdong) CHINE, VII-2008 (F 52mm, M 46mm)
Copyright © Alain Mollard
Carabus (Coptolabrus) ignimitella angulicollis Hauser, 1911 (as mellianus Kolbe, 1914)
Male+Female adult
Tianjing Shan (Luyuan, Lecheng), Guangdong) CHINE, VII-2008 (m 48mm, fem 50mm)
Copyright © Alain Mollard
Carabus (Coptolabrus) kubani Deuve, 1990 Paratype
Male adult
Daju, (Yulongxue Shan, Yunnan) CHINE, VII-1990
Copyright © Alain Mollard
Carabus (Coptolabrus) ignimitella stillatus Cavazzuti & Ratti, 1998
Male+Female adult
Mt.Nanshan (Nanping, Fujian) CHINE VI-2001
Copyright © Alain Mollard
Carabus (Coptolabrus) ignimitella ignimitella Bates, 1888 (as nyukmingi Deuve, 1991)
Male+Female adult
Wuyi Shan (Fujian) CHINE VI-2001
Copyright © Alain Mollard
Carabus (Coptolabrus) ignimitella ignimitella Bates, 1888
Male+Female adult
Luozhou (Fuzhou, Fujian) CHINE VI-2001
Copyright © Alain Mollard
Carabus (Coptolabrus) pustulifer aeriviolaceus Cavazzuti & Rapuzzi, 2010
Male+Female adult
Meigu (Xichang, Sichuan) CHINE, VII-2008
Copyright © Alain Mollard
Carabus (Coptolabrus) pustulifer elisabettianus Font, 1997
Male+Female adult
Huanjan Shan (Wuchuan, Guizhou) CHINE, VII-2008
Copyright © Alain Mollard
Carabus (Coptolabrus) pustulifer inusitatus Cavazzuti, 1997
Male+Female adult
Fanjing Shan (Jiangkou, Guizhou) CHINE, VI-2007
Copyright © Alain Mollard
Carabus (Coptolabrus) pustulifer katsumii Deuve & Ohshima, 1992 Paratype
Female adult
Wenxian (Gansu) CHINE, VII-2008
Copyright © Alain Mollard
Carabus (Coptolabrus) pustulifer mirandior Deuve, 1993
Male+Female adult
Dayao (Yunnan) CHINE, VII-2008
Copyright © Alain Mollard
Carabus (Coptolabrus) pustulifer mirificoides Deuve & Li, 2002
Male+Female adult
Yangtian (Gansu) CHINE, VII-2008
Copyright © Alain Mollard
Carabus (Coptolabrus) pustulifer qiannaicus Deuve & Tian, 2000
Male+Female adult
Longzjia (Maoping, Guizhou) CHINE, V-2008
Copyright © Alain Mollard
Carabus (Coptolabrus) pustulifer szechwanensis Hauser, 1931
Male+Female adult
Jinfo Shan (Chongqing) CHINE, V-2008
Copyright © Alain Mollard
Carabus (Coptolabrus) pustulifer wakoi Imura, 1995
Female adult
Mts. Dabashan (Chengkou, Sichuan) CHINE, V-2008
Copyright © Alain Mollard
Carabus (Coptolabrus) pustulifer xianfengensis Deuve & Li, 2009
Male+Female adult
Mt.Maoping (Zhaoshi, Hunan/Hubei) CHINE, V-2008
Copyright © Alain Mollard
Carabus (Coptolabrus) augustus chenzhouensis Deuve & Sehnal, 2010
Male+Female adult
Yizhang Xian (Hunan) CHINE
Copyright © Alain Mollard
Carabus (Coptolabrus) augustus tongrenensis Deuve & Li, 2006
Male adult
Tongren Xian (Jiangkou, Guizhou) CHINE
Copyright © Alain Mollard
Carabus (Coptolabrus) augustus haiyangicus Deuve, 2012
Male+Female adult
Xing’an (Guilin, Guangxi) CHINE
Copyright © Alain Mollard
Carabus (Coptolabrus) pustulifer adriaenssensi (as guizhouicus Deuve, 1994)
Male+Female adult
Leigong Shan (Kaili, Guizhou) CHINE
Copyright © Alain Mollard
Carabus (Coptolabrus) augustus mugepoensis Deuve & Li in Deuve, 2009
Male+Female adult
Mt.Gupo Shan (Zhongshan, Guangxi) CHINE
Copyright © Alain Mollard
Carabus (Coptolabrus) augustus divus Roeschke, 1908
Male+Female adult
Pingxiang (Zhongshan, Jiangxi) CHINE
Copyright © Alain Mollard
Carabus (Coptolabrus) formosus formosus Semenov, 1887
Male+Female adult
Wu-Chang-xi, (Gansu) CHINE VI-1998
Copyright © Alain Mollard
Carabus (Coptolabrus) formosus subformosus Semenov, 1887
Male+Female adult
Wenxian, (Gansu) CHINE VIII-1992
Copyright © Alain Mollard
Carabus (Coptolabrus) formosus brockmannioides Deuve & Murzin, 1997 Paratype
Male+Female adult
Shihoshian, (Lixian, Gansu) CHINE VI-2003
Copyright © Alain Mollard
Carabus (Coptolabrus) formosus akane Imura & Yamaya, 1994 Paratype
Female adult
Tongchuan (Xi’an, Shaanxi) CHINE VI-1993
Copyright © Alain Mollard
Carabus (Coptolabrus) formosus purpureodiscoides Kleinfeld, 1999
Male+Female adult
Dalong Shan, Yunxian (Shiyan, Hubei) CHINE X-2010
Copyright © Alain Mollard
Carabus (Coptolabrus) elysii xiaoganicus Deuve et Remond, 2016
Male+Female adult
Shuangfeng Shan (Xiaogan, Hubei) CHINE VI-2016
Copyright © Alain Mollard
Carabus (Coptolabrus) elysii xiaoganicus Deuve et Remond, 2016
Male+Female adult
Shuangfeng Shan (Xiaogan, Hubei) CHINE VI-2016
Copyright © Alain Mollard
Carabus (Coptolabrus) elysii chikongschanensis Hauser, 1914
Male+Female adult
Zhuguang, (Hunan) CHINE V-1991
Copyright © Alain Mollard
Carabus (Coptolabrus) elysii amucronatus Deuve, 2010
Male adult
Zhongtiao Shan (Shanxi) CHINE VII-2002
Copyright © Alain Mollard
Carabus (Coptolabrus) elysii viridicyaneus Lassalle, 2016 Paratype
Male adult
Copyright © Bernard Lassalle
Carabus (Coptolabrus) smaragdinus smaragdinus Fischer, 1823
Male+Female adult
Mt. Nerchinski (Tchita, Sibérie) RUSSIE VII-1995
Copyright © Alain Mollard
Carabus (Coptolabrus) smaragdinus lengxuemeiae Deuve & Li, 2000
Male+Female adult
Lujiaogou Shan (Kuandian, Liaoning) CHINE VI-1997
Copyright © Alain Mollard
Carabus (Coptolabrus) jankowskii chindoensis Kwon & Park, 1989
Male+Female adult
Chongue san (Chindo Isle, ) KOREE VII-1992
Copyright © Alain Mollard
Carabus (Coptolabrus) jankowskii quelpartianus Breuning, 1932
Male+Female adult
Hallasam, (Cheju-do Isle, ) KOREE VII-1990
Copyright © Alain Mollard
Carabus (Coptolabrus) lafossei tungchengensis Li, 1993
Male+Female adult
Mt.Huang-Bai-Shan (Shangcheng, Henan) VI-2012
Copyright © Alain Mollard
Carabus (Coptolabrus) elysii amucronatus Deuve, 2010 Holotype
Male adult
Chine, Shanxi, Yongji. 29mm
Copyright © Thierry Deuve
Carabus (Coptolabrus) elysii elysii Thomson, 1856 (as idai)
Female adult
China, Qinling, Zhouzhi co., Shaanxi pr., 8-14.6.2009 (cAA).
Copyright © Alexander Anichtchenko
Carabus (Coptolabrus) lafossei tungchengensis Li, 1993
Male adult
China, Hubei, Shangzhi co., Shimenyan 8-11.6.2009 (cAA) 42 mm
Copyright © Alexander Anichtchenko
Carabus (Coptolabrus) gemmifer mesites Fairmaire, 1897
Female adult
China, Sichuan, Xichang City, Muli Co., Ke-Er-xiang 2-12.7.2009 (cAA) 32 mm
Copyright © Alexander Anichtchenko
Carabus (Coptolabrus) ignimitella angulicollis Hauser, 1911
Male adult
China Hunan, Guidong pr., 20.5.2009 (cAA) 44 mm
Copyright © Alexander Anichtchenko
Carabus (Coptolabrus) gemmifer rectipustulatus Rapuzzi, 2024 Holotype
Male adult
Sichuan, Yanyuan to Wodixiang; 2700 m asl; 16 Aug. 2022
Copyright © - - -
Carabus (Coptolabrus) elysii pulcher Kleinfeld, 1997
Male adult
EC-China, S-Henan, Jigong Shan, (?Hanjinggou), V. 2015, L=40 mm
Copyright © Ruslan Panin
Carabus (Coptolabrus) pustulifer mirificus Kraatz, 1894
Male adult
SC-Chine, NW-Hunan, Longshan County, Mt. Maoping,10-30.VI.2008, L=44 mm
Copyright © Ruslan Panin -
Cratocarabus taxon page (5 photos)
Carabus (Cratocarabus) kryzhanovskii molossulus Deuve, 1993 Paratype
Male adult
mele, 16mm
Copyright © Boleslav Brezina
Carabus (Cratocarabus) kryzhanovskii molossulusas (as laevimolossulus Deuve, 1993) Paratype
Male adult
female, 17mm
Copyright © Boleslav Brezina
Carabus (Cratocarabus) kryzhanovskii kryzhanovskii Bogachev, 1965
Male adult
male, 15mm
Copyright © Boleslav Brezina
Carabus (Cratocarabus) kryzhanovskii kryzhanovskii Bogachev, 1965
Female adult
Copyright © Ilya Kabak -
Cratocechenus taxon page (38 photos)
Carabus (Cratocechenus) wrzecionkoi HT (HYBRID ?Eotribax valikhanovi/Cratocechenus akinini puellus ) Holotype
Male adult
Copyright © Boleslav Brezina
Carabus (Cratocechenus) tshistjakovae tshistjakovae Kabak, 2001 Paratype
Male adult
male, 24mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) redikortzewi Semenov & Breuning, 1931
Female adult
female, 17mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) redikortzewi Semenov & Breuning, 1931
Male adult
male, 15mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) ovtchinnikovi ovtchinnikovi Gottwald, 1987
Male adult
Copyright © Alexander Anichtchenko
Carabus (Cratocechenus) ovtchinnikovi ovtchinnikovi Gottwald, 1987
Female adult
female, 18mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) ovtchinnikovi letellierianus Deuve, 1990
Female adult
female, 21mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) ovtchinnikovi letellierianus Deuve, 1990
Male adult
male, 18mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) ovtchinnikovi kafkai Deuve, 1994
Female adult
female, 21mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) jacobsoni Semenov, 1898
Female adult
female, 16mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) jacobsoni x akinini elisabethae HYBRID
Female adult
female, 18mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) jacobsoni Semenov, 1898
Male adult
S Kazakhstan, Almaty reg., Big Lake, h=2800 m, 25.VI.96
male
L=14 mm
Copyright © Maksim Smirnov
Carabus (Cratocechenus) akinini puellus Lapouge, 1924
Male adult
male, 19mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) akinini puellus Lapouge, 1924
Female adult
female, 21mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) akinini akinini Morawitz, 1886 (as oitalensis ) Paratype
Male adult
male, 20mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) akinini musartianus Gottwald, 1987
Female adult
female, 22mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) akinini musartianus Gottwald, 1987
Male adult
male, 21mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) akinini loudai Gottwald, 1987
Female adult
female, 23mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) akinini loudai Gottwald, 1987
Male adult
male, 20mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) akinini loudai Gottwald, 1987
Male adult
male, 21mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) akinini ketmenensis Dolin, 1991
Male adult
male, 22mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) akinini ketmenensis Dolin, 1991
Male adult
male, 21mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) akinini loudai (as issykatae Chromý, 1993) Paratype
Male adult
male, 21mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) akinini loudai (as issykatae Chromý, 1993) Paratype
Male adult
female, 23mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) akinini elisabethae Semenov, 1908
Female adult
female, 23mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) akinini elisabethae Semenov, 1908
Female adult
female, 22mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) akinini elisabethae Semenov, 1908
Male adult
male, 20mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) akinini elisabethae Semenov, 1908
Male adult
male, 21mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) akinini buffi Deuve & Kalab, 1992
Male adult
male, 18mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) akinini akinini Morawitz, 1886
Female adult
female, 21mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) akinini akinini Morawitz, 1886
Male adult
male, 18mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) akinini akinini Morawitz, 1886
Male adult
male, 20mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) akinini akinini Morawitz, 1886
Male adult
male, 22mm
Copyright © Boleslav Brezina
Carabus (Cratocechenus) akinini akinini Morawitz, 1886 (as yiningensis ) Holotype
Female adult
China, Kuldzha (Yining) VI.1936
Copyright © Dmitri Obydov
Carabus (Cratocechenus) ovtchinnikovi ovtchinnikovi Gottwald, 1987
Male adult
N-Kyrgyzstan, Kyungey Ala-Too Mt Rng, Cholpon-Ata env., h=2500m, 19.VI.2000,
L=19 mm
Copyright © Ruslan Panin
Carabus (Cratocechenus) tshistjakovae gracilidens Kabak, 2004
Female adult
China, Xinjiang, Narat mt.rng.
Copyright © Ilya Kabak
Carabus (Cratocechenus) tshistjakovae gracilidens Kabak, 2004
Female adult
China, Xinjiang, Narat Mt.R.,, h~2275 Tshashi vall.,Tuyukasha Riv. 43°00'29"N 82°46'02"E 18.VII.2014 leg. I.Kabak L=22.0
Copyright © Kirill Makarov -
Cratocephalus taxon page (17 photos)
Carabus (Cratocephalus) solskyi Ballion, 1878
Male adult
SE Kazakhstan, Ketmen Mt. rng., Tuyuk vill., 16.06.97 leg. S.Toropov
Copyright © Maksim Smirnov
Carabus (Cratocephalus) corrugis Dohrn, 1882
Male adult
Copyright © Alexandr Safronov
Carabus (Cratocephalus) cicatricosus cicatricosus Fischer, 1842
Male adult
SE Kazakhstan, Ketmen mt.rng., Tuyuk vill., 1800m, 15.5.2000
Copyright © Alexander Anichtchenko
Carabus (Cratocephalus) cicatricosus cicatricosus Fischer, 1842
Male adult
female, 33mm
Copyright © Boleslav Brezina
Carabus (Cratocephalus) cicatricosus cicatricosus Fischer, 1842
Male adult
male, 28mm
Copyright © Boleslav Brezina
Carabus (Cratocephalus) cicatricosus cicatricosus Fischer, 1842
Male adult
KAZAKHSTAN, Bolshoe Almatinskoe Lake, 30.VI.1992
L=26mm
Copyright © Ruslan Panin
Carabus (Cratocephalus) balassogloi Dohrn, 1882
Male adult
KIRGIZSTAN, N Issyk-Kul Lake, Ananievo env., 1800 m; 20.04.2000
Copyright © Anton Menstschikov
Carabus (Cratocephalus) solskyi Ballion, 1878
Female adult
SE-KAZAKHSTAN, Ketmen Mt Rng, h=1800m, Tujuk vill.,15.V.2004
L=36 mm
Copyright © Ruslan Panin
Carabus (Cratocephalus) solskyi chengyii Rapuzzi & Yang, 2021 Holotype Paratype
Male+Female adult
China, Xinjiang province, Yining County, Kashi Township Reservoir, VII.2020 (deposited in collection Ivan Rapuzzi, Prepotto, Italy). Image from Rapuzzi & Yang, 2021
Copyright © - - -
Carabus (Cratocephalus) cicatricosus cicatricosus Fischer, 1842
Male adult
SE-Kazakhstan, Almaty reg., C Trans-Ili Alatau N-slopes, Kaskelen Valley, Izvestkovyi vill., h=1700 m, 6.V.2015, D.Kovalchuk leg., L=28.5 mm
Copyright © Ruslan Panin -
Cratophyrtus taxon page (15 photos)
Carabus (Cratophyrtus) turcosinensis medvedevi Kryzhanovskij, 1968
Female adult
S Kazakhstan, NW slope of Ugamskii mt.rng., 3 km W Sairamskii peak, 2200m
tributary of Karzhantau riv., 15.6.2004 Matalin A. leg.
20 mm
Copyright © Alexander Anichtchenko
Carabus (Cratophyrtus) turcosinensis turcosinensis (as korelli Kleinfeld, 1988) Holotype
Male adult
Copyright © Frank Kleinfeld
Carabus (Cratophyrtus) kaufmanni segregatus Morawitz, 1886
Female adult
female, 19mm
Copyright © Boleslav Brezina
Carabus (Cratophyrtus) kaufmanni segregatus Morawitz, 1886
Male adult
male, 18mm
Copyright © Boleslav Brezina
Carabus (Cratophyrtus) kaufmanni lebedevianus Breuning, 1933
Male adult
male, 19mm
Copyright © Boleslav Brezina
Carabus (Cratophyrtus) kaufmanni lebedevianus Breuning, 1933
Male adult
male, 20mm
Copyright © Boleslav Brezina
Carabus (Cratophyrtus) kaufmanni kaufmanni Solsky, 1874
Female adult
female, 19mm
Copyright © Boleslav Brezina
Carabus (Cratophyrtus) kaufmanni kaufmanni Solsky, 1874
Male adult
male, 19mm
Copyright © Boleslav Brezina
Carabus (Cratophyrtus) katajevi katajevi Gottwald, 1989
Male adult
male, 25mm
Copyright © Boleslav Brezina
Carabus (Cratophyrtus) nikolajevi Kabak, 1998 Holotype
Male adult
W part of Kirgiz Mt. R., valley of Makpal River, 21.07.1996, G.V. Nikolajev leg. [Southern Kazakhstan, ca. 42°53'N 72°01'E] (ZIN).
Copyright © Ilya Kabak
Carabus (Cratophyrtus) nikolajevi Kabak, 1998
Male adult
S Kazakhstan, Kyrgyz Ala-Too Mt. R., right source of Shungir Riv., 27 km SSW of Lugovoy Vill., 2073 m, 42°43'05"N / 72°36'31"E, 7.06.2025, I.I. Kabak, A.M. Shapovalov leg. Image from Kabak & Shapovalov, 2025
Copyright © Ilya Kabak -
Cryptocarabus taxon page (16 photos)
Carabus (Cryptocarabus) subparallelus Ballion, 1878
Female adult
Kazakhstan,
near Alma-Ata, V.2009, Pak O.leg.
female, 17mm
Copyright © Ruslan Panin
Carabus (Cryptocarabus) sacarum Kabak, 1998
Male adult
SE Kazakhstan, Zailiiskii Ala too, Kaskelen riv.,
1400m 24.V.2000
Copyright © Alexander Anichtchenko
Carabus (Cryptocarabus) mullerellus kasantsevi Kabak, 1993
Male adult
SE Kazakhstan, Alma-Ata reg., Narynkol vill.
12.V.2000 Klimenko A. leg.
Copyright © Alexander Anichtchenko
Carabus (Cryptocarabus) lindemanni Ballion, 1878
Male adult
SE Kazakhstan, Alma-Ata env., Kok-Tube 16.IV.2000
L=15mm
Copyright © Alexander Anichtchenko
Carabus (Cryptocarabus) lindemanni Ballion, 1878
Female adult
SE Kazakhstan, Alma-Ata env., 16.04. 2000
L=16,0 mm
Copyright © Maksim Smirnov
Carabus (Cryptocarabus) kadyrbekovi Kabak 1994
Male adult
SE Kazakhstan, Zailijskii Ala Too, Karakastek riv., 27.5.2000
Copyright © Alexander Anichtchenko
Carabus (Cryptocarabus) ? turkestanus turkestanus Breuning, 1929
Male adult
Kyrgyzstan, W Kungei Ala-Too Mt Rng, Chon-Kemin riv., Kyzyl-Bairak vill., 25.IV.2010. L= 18 mm
Copyright © Ruslan Panin
Carabus (Cryptocarabus) kasantsevi Kabak, 1993
Male adult
SE Kasakhstan, E part of Terskey Alatau, Bayankol riv., tugai.
Copyright © Ilya Kabak
Carabus (Cryptocarabus) sororius Morawitz, 1886
Male adult
China, Xinjiang, S slope of Boro-Horo Mt.R., env. of Ulastai vill.,h~1690 43°49'50"N 83°08'45"E 8.V.2017 leg. I.Kabak L=18.5 mm
Copyright © Kirill Makarov
Carabus (Cryptocarabus) sororius Morawitz, 1886
Female adult
China, Xinjiang, S slope of Boro-Horo Mt.R., env. of Ulastai vill.,h~1690 43°49'50"N 83°08'45"E 8.V.2017 leg. I.Kabak L=20.0 mm
Copyright © Kirill Makarov
Carabus (Cryptocarabus) tsharynensis Kabak 1994
Male adult
SE-Kazakhstan, Almaty Region, W-Zailiyskiy Alatau rng, Belshabdar valley h~1300 m, 25-27.VI.2002, A.Puchkov leg. L=17 mm
Copyright © Ruslan Panin
Carabus (Cryptocarabus) tsharynensis Kabak 1994
Female adult
E-Kazakhstan, Almaty Region, W-Zailiyskiy Alatau, Turgen distr., Tauturgen vill., h~1200 m, 8-13.VI.2002, A.Puchkov leg. L=19 mm
Copyright © Ruslan Panin
Carabus (Cryptocarabus) sacarum Kabak, 1998
Male adult
SE-Kazakhstan, Almaty reg., C-Zailiyskiy Alatau N-slopes, Kaskalen river Valley, Izvestkovy vill., h=1700 m, 6.V.2015, D.Kovalchuk leg./det. R.Panin L=18 mm
Copyright © Ruslan Panin
Carabus (Cryptocarabus) sacarum Kabak, 1998
Female adult
SE-Kazakhstan, Almaty reg., C-Zailiyskiy Alatau N-slopes, Kaskalen river Valley, Izvestkovy vill., h=1700 m, 6.V.2015, D.Kovalchuk leg./det. R.Panin L=20 mm
Copyright © Ruslan Panin -
Ctenocarabus taxon page (8 photos)
Carabus (Ctenocarabus) galicianus Gory, 1839
Male adult
Spain,
Sierra de la Cabrera Baja, 8 km SW Truchillas, 9.VII.2008, Anichtchenko A. leg.
L= 20 mm
Copyright © Alexander Anichtchenko
Carabus (Ctenocarabus) galicianus Gory, 1839
Female adult
NW-Spain, Pontevedra prov., Campo Lameiro env., 15.VIII.1994, S.Sobrino leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Ctenocarabus) melancholicus costatus Germar, 1824
Female adult
C-Spain, Madrid region, Rozas de Puerto Real env.,21.XII.2008, T.Gazurek leg.
L=27 mm
Copyright © Ruslan Panin
Carabus (Ctenocarabus) melancholicus melancholicus Fabricius, 1798
Male adult
NW Morocco, Kénitra, Souk El Arbaa env., II.1985, V.Olivella leg. L=25 mm
Copyright © Ruslan Panin
Carabus (Ctenocarabus) melancholicus submeridionalis Breuning, 1975
Male adult
S- Spain, Cádiz, Tarifa, El Cabrito, 28.XII.2019, F. Garcia leg. L=26 mm
Copyright © Ruslan Panin
Carabus (Ctenocarabus) melancholicus costatus Germar, 1824
Male adult
NW Portugal, Viana do Castelo distr., Sierra de Agra, 41°47'52.3"N 8°41'27.6"W, h=786 m, 3.VII.2016, D. Prunier leg.
Copyright © Ruslan Panin -
Cupreocarabus taxon page (45 photos)
Carabus (Pseudocranion) fumigatus fumigatus Semenov, 1898 Lectotype
Male adult
male, 22mm
Copyright © Boleslav Brezina
Carabus (Pseudocranion) fumigatus fumigatus Semenov, 1898 Syntypes
Female adult
female, 22mm
Copyright © Boleslav Brezina
Carabus (Pagocarabus) sinotibeticola xiaodongi (Imura, Zhou & Su, 2004)
Male adult
Copyright © Boleslav Brezina
Carabus (Pagocarabus) sinotibeticola sinotibeticola Mandl, 1975
Male adult
Copyright © Boleslav Brezina
Carabus (Pagocarabus) sinotibeticola sinotibeticola Mandl, 1975
Female adult
Copyright © Boleslav Brezina
Carabus (Pagocarabus) sinotibeticola pantholops Brezina, Hackel & Sehnal, 2007 Holotype
Male adult
SW-Sichuan: W of Zhier: btw. 28°22.293'N/101°32.701'E & 28°21.113'N/101°29.384'E, 3500 m, 23mm
Copyright © Boleslav Brezina
Carabus (Pagocarabus) sinotibeticola pantholops Brezina, Hackel & Sehnal, 2007 Paratype
Female adult
24mm
Copyright © Boleslav Brezina
Carabus (Pagocarabus) alpherakii takin Hackel & Sehnal, 2007 Holotype Paratype
Male adult
male, 19mm
Copyright © Boleslav Brezina
Carabus (Pagocarabus) alpherakii takin Hackel & Sehnal, 2007
Male adult
male, 17mm
Copyright © Boleslav Brezina
Carabus (Pagocarabus) alpherakii takin Hackel & Sehnal, 2007 Paratype
Female adult
female, 22mm
Copyright © Boleslav Brezina
Carabus (Pagocarabus) alpherakii takin Hackel & Sehnal, 2007 Paratype
Female adult
female, 23mm
Copyright © Boleslav Brezina
Carabus (Pagocarabus) alpherakii takin Hackel & Sehnal, 2007 Paratype
Male adult
male, 18 mm
Copyright © Boleslav Brezina
Carabus (Pagocarabus) alpherakii pignataroi Cavazzuti, 1996 Paratype
Female adult
female, 20mm
Copyright © Boleslav Brezina
Carabus (Pagocarabus) alpherakii pignataroi Cavazzuti, 1996
Male adult
male, 20mm
Copyright © Boleslav Brezina
Carabus (Pagocarabus) alpherakii pignataroi Cavazzuti, 1996 Paratype
Female adult
Copyright © Boleslav Brezina
Carabus (Cupreocarabus) alpherakii alpherakii Semenov, 1898
Male adult
male, 23mm
Copyright © Boleslav Brezina
Carabus (Cupreocarabus) alpherakii alpherakii Semenov, 1898
Female adult
female, 24mm
Copyright © Boleslav Brezina
Carabus (Cupreocarabus) alpherakii alpherakii Semenov, 1898
Male adult
Chine, W Sichuan, Litang env., <br /> 3-6.7.99 Dr. V. Banes leg.
Copyright © Alexander Anichtchenko
Carabus (Pagocarabus) alpherakii budha Breuning, 1950
Female adult
female, 20mm
Copyright © Boleslav Brezina
Carabus (Pagocarabus) alpherakii budha Breuning, 1950
Female adult
female, 24mm
Copyright © Boleslav Brezina
Carabus (Pagocarabus) alpherakii budha Breuning, 1950
Male adult
male, 20 mm
Copyright © Boleslav Brezina
Carabus (Pagocarabus) alpherakii budha Breuning, 1950
Male adult
male, 19mm
Copyright © Boleslav Brezina
Carabus (Cupreocarabus) lixianensis jinchuanicus Deuve & Tian, 2007
Male adult
male, 22 mm
Copyright © Boleslav Brezina
Carabus (Cupreocarabus) lixianensis jinchuanicus Deuve & Tian, 2007
Female adult
female, 24 mm
Copyright © Boleslav Brezina
Carabus (Cupreocarabus) lixianensis gamdaensis Deuve & Tian, 2002 Paratype
Male adult
male, 22mm
Copyright © Boleslav Brezina
Carabus (Cupreocarabus) laevithorax taiyanghensis Deuve & Tian, 2006
Male adult
male, 21 mm
Copyright © Boleslav Brezina
Carabus (Cupreocarabus) laevithorax taiyanghensis Deuve & Tian, 2006
Female adult
female, 23 mm
Copyright © Boleslav Brezina
Carabus (Cupreocarabus) kangdingi sculptior Deuve, 1992
Female adult
female, 25mm
Copyright © Boleslav Brezina
Carabus (Cupreocarabus) kangdingi sculptior Deuve, 1992
Male adult
male, 22mm
Copyright © Boleslav Brezina
Carabus (Cupreocarabus) kangdingi sculptior Deuve, 1992
Male adult
male, 23mm
Copyright © Boleslav Brezina
Carabus (Cupreocarabus) kangdingi kangdingi Korell, Kleinfeld & Gorgner, 1992 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Cupreocarabus) drahoslavae Brezina B. & Hackel M., 2006 Paratype
Female adult
Copyright © Boleslav Brezina
Carabus (Cupreocarabus) drahoslavae Brezina B. & Hackel M., 2006 Holotype
Male adult
Copyright © Boleslav Brezina
Carabus (Cupreocarabus) huangi huangi Deuve, 1992
Male adult
C-China, E Tibet, N-Sichuan, Hongyuan,valley 10 km SE Sanggarpar, h=4200m, 32.300053, 102.584031, alpine meadow / scree, 19.VII.1995, B.Brezina leg.<br /> L=21 mm
Copyright © Ruslan Panin
Carabus (Cupreocarabus) lixianensis lixianensis Deuve, 1990
Male adult
China, E Tibet, N-Sichuan, (Barkam),valley SW of Barkam, h=3200m, 31*53’ N; 102*12’E, mixed/coniferous forest, 22.VII.1995, K.& B.Brezina
L=23 mm
Copyright © Ruslan Panin
Carabus (Cupreocarabus) alpherakii alpherakii Semenov, 1898
Female adult
China, W Sichuan, Litang env., h~5000 m,29.VII -3.VIIII.1995, J. Schneider leg. L=23 mm
Copyright © Ruslan Panin
Carabus (Cupreocarabus) tianmingyii Deuve, 2015 Holotype
Male adult
Chine, Sichuan, Barkam Xian, Kangshan Xiang, Ya’erzhu, 3500-4000 mètres (Th. Deuve, Tian Mingyi et Song Zhao, 14-18 juillet 2007), in coll. South China Agricultural University, Canton.
Copyright © Thierry Deuve
C. (Cupreocarabus) kuceraianus subemili Deuve & Kaláb, 2017 Holotype
Male adult
Chine, Sichuan centro-occidental, montagnes à 15 km à l’est-nord-est de Jintang, 4100 mètres, 30º25’N/102º25’E (pelouses alpines, éboulis), in coll. J. Kaláb, à Jinačovice (République Tchèque).
Copyright © Thierry Deuve
Carabus (Cupreocarabus) mollardianus Deuve, 2016 Holotype
Female adult
Chine, Sichuan, Baoxing Xian, Monts Jiajin Shan, Lushan
Copyright © Thierry Deuve
Carabus (Cupreocarabus) balangicus qiaoqiensis Deuve & Kaláb, 2015 Holotype
Male adult
Chine, Sichuan central, montagnes à 13 km au nord-est de Qiaoqi, 3750-4200 mètres, 30°47’N/102°50-51’E, en pelouse alpine et éboulis, in coll. Jaroslav Kaláb, à Jinačovice (République Tchèque).
Copyright © Thierry Deuve
Carabus (Cupreocarabus) kuceraianus emili Deuve et Kaláb, 2015 Holotype
Male adult
Chine, Sichuan central, env. 11 km au nord-ouest / nord-nord-ouest de Jintang, abords orientaux du col de Muzhuliangzi, 4000 mètres, 30°27’N/102°13’E, en pelouse alpine et éboulis, in coll. Jaroslav Kaláb, à Jinačovice (République Tchèque).
Copyright © Thierry Deuve
Carabus (Cupreocarabus) mollardianus Deuve, 2016
Female adult
Chine, Sichuan, Lushan, Mt. Jiajin Shan, 10.VII.2016 local collector leg./ det. R.Panin, L=22 mm
Copyright © Ruslan Panin
Carabus (Cupreocarabus) ? siguniangensis jiachingensis Lassalle, 2003
Male adult
Chine, Sichuan, Lushan, Mt. Jiajin Shan, 10.VII.2016 local collector leg., L=21 mm
Copyright © Ruslan Panin
Carabus (Cupreocarabus) alpherakii exilior Deuve, 2017 Holotype
Male adult
Chine, Sichuan, 40 km S Hekou (Yajiang), mts à l’ouest de Tanggang, 3800 m, 29°41’31’’N-101°01’08’’E, in coll. Ross Sehnal, à Unhošt (République Tchèque).
Copyright © Thierry Deuve
Carabus (Cupreocarabus) alpherakii younglimae Deuve, 2017 Holotype
Male adult
Chine, Sichuan, vall. 8 km NE Shade, 4575 m, 29°42’33’’N-101°19’10’’E, in coll. M. Janata, à Prague (République Tchèque).
Copyright © Thierry Deuve
Carabus (Cupreocarabus) sinotibeticola feicuipennis (Imura, Zhou & Su, 2004) Holotype
Male adult
shrubs of Rhododendron spp., 4, 1 35 m in altitude, in southwestern Jiulong Xian of western Sichuan, Southwest China, 1 0-13-VI I-200 1, Xiao-Dong Yu & Hong-Zhang ZHOU leg., in coll. Institute of Zoology, Chinese Academy of Sciences, Beijing. Image from (Imura, Zhou & Su, 2004)
Copyright © - - -
Carabus (Cupreocarabus) sculptior geshizhanus (Imura, Zhou & Su, 2004) Holotype
Male adult
mixed forest, 2,610 m in altitude, Geshizha in central Danba Xian of west-central Sichuan, Southwest China, 20-22-V II-200 1, Xiao-Dong Yu & Hong-Zhang ZHOU leg. Image from (Imura, Zhou & Su, 2004)
Copyright © - - -
Carabus (Cupreocarabus) minimocupreus zhegushanus Cavazzuti & Rapuzzi, 2005 Holotype
Female adult
Cina, Sichuan centrale, Lixian, Colle di Zhegushan 4100 m.
Copyright © - - -
Carabus (Cupreocarabus) melii Cavazzuti & Ratti, 2005 Holotype
Male adult
Cina, Sichuan centrale, Valle a nord-ovest di Heishui, 2700 m
Copyright © - - -
Carabus (Cupreocarabus) balangicus balangicus Cavazzuti, 2002
Male adult
Chine, C-Sichuan, Xiaojin Co, Wolong valley, Balang Shan Pass h=3448 m, 30.884617, 102.973117, 9.VI.-1.VII.2011, G. Proscia leg. L=19 mm
Copyright © Ruslan Panin
Carabus (Cupreocarabus) balangicus balangicus Cavazzuti, 2002
Male adult
Chine, C-Sichuan, Xiaojin Co, Wolong valley, Balang Shan Pass h=3448 m, 30.884617, 102.973117, 9.VI.-1.VII.2011, G. Proscia leg. L=20 mm
Copyright © Ruslan Panin
Carabus (Cupreocarabus) huangi huangi Deuve, 1992
Male adult
China, E Tibet, N-Sichuan, 53 km NE of Barkam, Pass Sandaoban env., h=4293 m, 32.232950, 102.598533, 11-29.VI.2011, G. Proscia leg., L=21 mm
Copyright © Ruslan Panin
Carabus (Cupreocarabus) huangi huangi Deuve, 1992
Female adult
China, E Tibet, N-Sichuan, 53 km NE of Barkam, Pass Sandaoban env., h=4293 m, 32.232950, 102.598533, 11-29.VI.2011, G. Proscia leg., L=21.5 mm
Copyright © Ruslan Panin
Carabus (Cupreocarabus) laevithorax arduus Cavazzuti, 2008 Holotype
Male adult
C-Sichuan: Xiaojin Xian: Buma Shan: pass N of Fubian, 31.33159N 102.24679 (102.26405) 4550m
Copyright © - - - -
Cychrostomus taxon page (8 photos)
Carabus (Cychrostomus) pseudoprosodes Semenov & Znojko, 1932
Male adult
male, 19mm
Copyright © Boleslav Brezina
Carabus (Cychrostomus) anchocephalus anchocephalus (as facetus Semenov, 1903)
Female adult
L=24mm
Copyright © Boleslav Brezina
Carabus (Cychrostomus) anchocephalus lanzhouensis Deuve, 1991
Male adult
male, 24mm
Copyright © Boleslav Brezina
Carabus (Cychrostomus) anchocephalus lanzhouensis Deuve, 1991 Paratype
Male adult
male, 21mm
Copyright © Boleslav Brezina
Carabus (Cychrostomus) anchocephalus anchocephalus Reitter, 1896
Male adult
male, 22mm
Copyright © Boleslav Brezina
Carabus (Cychrostomus) anchocephalus anchocephalus Reitter, 1896
Male adult
China, Shanxi pro., Jiaochen
Copyright © Ye Liu
Carabus (Cychrostomus) anchocephalus farinosus Rapuzzi, 2023 Holotype
Male adult
China, Inner Mongolia Province, 30 Km NW Hohhot, 3/7.VI.2010, M. Janata legit. Image from Rapuzzi, 2023
Copyright © - - -
Carabus (Cychrostomus) anchocephalus rectigranulatus Rapuzzi, 2024 Holotype
Male adult
Shanxi Province, Lu Liang, Kang Cheng Zhen; Nov. 2022. Image from Rapuzzi, 2024
Copyright © - - - -
Cyclocarabus taxon page (14 photos)
Carabus (Cyclocarabus) belousovi Kabak, 1992
Female adult
Uzbekistan, Karzhantau Mt., Aktash vill.
1500 m, 12.4.1999
female, 23 mm
Copyright © Alexander Anichtchenko
Carabus (Cyclocarabus) pseudolamprostus Kalab, 1996
Male adult
Kirgizstan, S slope of Pskem mt. rng.,
Chandalash riv., 2400m, 1.6.2000
male, 24 mm
Copyright © Alexander Anichtchenko
Carabus (Cyclocarabus) namanganensis namanganensis Heyden, 1886
Female adult
Uzbekistan, Karzhantau Mt., Aktash vill.
1500 m, 12.4.1999
L= 23 mm
Copyright © Alexander Anichtchenko
Carabus (Cyclocarabus) namanganensis namanganensis Heyden, 1886
Male adult
Uzbekistan, Karzhantau Mt., Aktash vill.
1500 m, 12.4.1999
L= 22 mm
Copyright © Alexander Anichtchenko
Carabus (Cyclocarabus) karaterekensis hemicallisthenes Kabak & Kalab, 1997
Male adult
Kirgizstan, Chandalash, Chakmak-Suu riv.,
2800 m, 1-2.6.2000
male, 17 mm
Copyright © Alexander Anichtchenko
Carabus (Cyclocarabus) aulacocnemus uzbek Deuve & Kalab, 1992
Female adult
Uzbekistan, 2800m, Chatkalskii mt. rng.,
Jangi-Abad vill., bass. Angren riv., 6.6.2000 Tihonov D. leg.
L= 24 mm
Copyright © Alexander Anichtchenko
Carabus (Cyclocarabus) aulacocnemus nurbeki Kalab, 1997
Male adult
Kirgizstan, S slope Pskem, Chandalash riv.,
2200 m, 31.5.2000
male, 21 mm
Copyright © Alexander Anichtchenko
Carabus (Cyclocarabus) aulacocnemus aulacocnemus ( as minusculus Semenov, 1903)
Male adult
W Kirgizstan, Talas mt. rng, Manas mt., 1800 m
22.4.2000
male, 18 mm
Copyright © Alexander Anichtchenko
Carabus (Cyclocarabus) aulacocnemus galinae Obydov, 2003 Paratype
Male+Female adult
W-Tianshan:Karatau Mts.,N-part:Boialdyr River valley,
N of Ken-su, 600-750m, 22-V-2000 М. Danilevsky leg.
male - 20 мм, female - 19 мм.
Copyright © Alexandr Safronov
Carabus (Cyclocarabus) namanganensis namanganensis Heyden, 1886
Male+Female adult
Kazakhstan, S. Karatau Mts. R., Mashat vill. env.
23-IV-1993 М. Danilevsky leg.
male - 21 мм, female - 22 мм.
Copyright © Alexandr Safronov
Carabus (Cyclocarabus) zarudnyi Semenov & Znojko, 1932
Female adult
NE-Uzbekistan, W-Tian-Shan, NW slopes Pskem Mt. Rng, 8-14 km E Pskem, 2100-2500 m, 16.VI.1998, A. Puchkov leg., L=21 mm
Copyright © Ruslan Panin
Carabus (Cyclocarabus) karaterekensis hemicallisthenes Kabak & Kalab, 1997
Male adult
NW- Kyrgyzstan, W-Tian-Shan, Chandalash Mt.Rng, Chakmak-Suu vill., 3.VI.2000, L=17 mm
Copyright © Ruslan Panin
Carabus (Cyclocarabus) brosciformis Semenov, 1896
Male adult
NE-Uzbekistan, W-Tian-Shan, Chatkal Mt Range, Tashkent reg., Yangiabad env., h=2100m, 30.IV.2002, L=23.5 mm
Copyright © Ruslan Panin -
Damaster taxon page (15 photos)
Carabus (Damaster) blaptoides oxuroides Schaum, 1862
Female adult
Japan, C-Honshu, Ofuse-cho, Kamitakai, Nagano Pref., 12.VI.2006, Ohshima leg.
L=34 mm
Copyright © Ruslan Panin
Carabus (Damaster) blaptoides babaianus Ishikawa, 1983
Male adult
Japan: C Honshu: Niigata pref., Mysato: Sakae Abiko-City; Iwafune distr., Awashima Isl. (LABEL ERROR)
Copyright © Igor Solodovnikov
Carabus (Damaster) blaptoides babaianus Ishikawa, 1983
Female adult
Japan: C Honshu: Niigata pref., Mysato: Sakae Abiko-City; Iwafune distr., Awashima Isl. (LABEL ERROR)
Copyright © Igor Solodovnikov
Carabus (Damaster) blaptoides capito Lewis, 1881
Male adult
Japan: Niigato pref., Sado Isl., Sado, Yamato
Copyright © Igor Solodovnikov
Carabus (Damaster) blaptoides capito Lewis, 1881
Male adult
Japan: Niigato pref., Sado Isl., Sado, Yamato
Copyright © Igor Solodovnikov
Carabus (Damaster) blaptoides oxuroides Schaum, 1862
Female adult
Japan, C Honshu, Nagano Prefecture, Nagano city, 3.IV.2012, S. Shinichi leg. L=42 mm. det. Anichtchenko A. 2021
Copyright © Ruslan Panin
Carabus (Damaster) blaptoides rugipennis Motschulsky, 1861
Male adult
Kunashir isl.
Copyright © Kirill Makarov
Carabus (Damaster) blaptoides babaianus Ishikawa, 1983
Male adult
Hagyu., Iide Town, Nishiokitama Co., Yamagata pref., Japan, 17 3.2016.T. Nakamura leg. (DUBC). 35mm
Copyright © Alexander Anichtchenko
Carabus (Damaster) blaptoides oxuroides Schaum, 1862
Female adult
Japan, Kiyose, Tokyo, 20.1.1961 (DUBC). 40 mm
Copyright © Alexander Anichtchenko
Carabus (Damaster) blaptoides blaptoides (Kollar, 1836)
Female adult
Japan, SW Honshu, Shimane Pref., Unnan city, 16.I2016, S.Shinichi leg., L=48 mm
Copyright © Ruslan Panin
Carabus (Damaster) blaptoides blaptoides (Kollar, 1836)
Female adult
Shiobara (Tochigi), JAPON, IV-1990
Copyright © Alain Mollard
Carabus (Damaster) blaptoides blaptoides (Kollar, 1836) (as lewisi Rye, 1872)
Female adult
Kitakata (Myakasi, Kyushu) JAPON, VI-1989
Copyright © Alain Mollard
Carabus (Damaster) blaptoides fortunei Adams, 1861
Male+Female adult
Awa-Shima Isle (Niigata) JAPON III-1990
Copyright © Alain Mollard -
Deroplectes taxon page (32 photos)
Carabus (Deroplectes) staudingeri Ganglbauer, 1886
Female adult
N Tajikistan, SW Pamir-Alay, Fann Mts h=1800m, Padrut vill., 2-3.V.1997
L=28 mm
Copyright © Ruslan Panin
Carabus (Deroplectes) staudingeri Ganglbauer, 1886
Male adult
Uzbekhistan, W Gissar mt.rng., 60 km Sakhrisaby 17.5.1994, Klimenko leg.
male, 25 mm
Copyright © Alexander Anichtchenko
Carabus (Deroplectes) staudingeri Ganglbauer, 1886
Female adult
female, 30mm
Copyright © Boleslav Brezina
Carabus (Deroplectes) sphinx sphinx (as pseudocoiffaitianus Deuve, 1990)
Female adult
L=30 mm
Copyright © Boleslav Brezina
Carabus (Deroplectes) sphinx sphinx (as pseudocoiffaitianus Deuve, 1990)
Male adult
L=25 mm
Copyright © Boleslav Brezina
Carabus (Deroplectes) sphinx dardazicus Deuve, 1990
Female adult
female, 28mm
Copyright © Boleslav Brezina
Carabus (Deroplectes) sphinx dardazicus Deuve, 1990
Male adult
male, 27mm
Copyright © Boleslav Brezina
Carabus (Deroplectes) dokhtouroffi pseudomidas Mandl, 1967
Male adult
male, 28mm
Copyright © Boleslav Brezina
Carabus (Deroplectes) dokhtouroffi pseudomidas Mandl, 1967
Male adult
male, 28mm
Copyright © Boleslav Brezina
Carabus (Deroplectes) dokhtouroffi dokhtouroffi Ganglbauer, 1886
Female adult
female, 32mm
Copyright © Boleslav Brezina
Carabus (Deroplectes) dokhtouroffi dokhtouroffi Ganglbauer, 1886
Male adult
male, 28mm
Copyright © Boleslav Brezina
Carabus (Deroplectes) dokhtouroffi ? pseudomidas Mandl, 1967
Male adult
Vahshskii Mt. rng., Saryhosor, Mullohomi 28.4.1990 S. L. Zonshtein leg.
Copyright © Alexander Anichtchenko
Carabus (Deroplectes) arcanus arcanus Semenov, 1898
Female adult
female, 25mm
Copyright © Boleslav Brezina
Carabus (Deroplectes) dokhtouroffi dokhtouroffi Ganglbauer, 1886
Male adult
NC Tadzhikistan, Pamir, N-slope Peter I.Mt Rng, h=2100m,Ganishou vill., 10-12.VI.2006, Ivanova E. leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Deroplectes) klapperichianus klapperichianus Mandl, 1955 Holotype
Male adult
image from carabus-online.de
Copyright © Frank Kleinfeld
Carabus (Deroplectes) klapperichianus salangicus Kleinfeld & Reuter, 2010 Holotype
Male adult
Afghanistan, Salang Pass, Nordseite, 25°27'N/68°58'E, ca. 2300m, 8.-18.IV.2010 (reg. subalp.) (in coll. Reuter)
23 mm
Copyright © Frank Kleinfeld
Carabus (Deroplectes) klapperichianus salangicus Kleinfeld & Reuter, 2010 Paratype
Female adult
Afghanistan, Salang Pass, Nordseite, 25°27'N/68°58'E, ca. 2300m, 8.-18.IV.2010 (reg. subalp.) (in coll. Reuter)
27 mm
Copyright © Frank Kleinfeld
Carabus (Deroplectes) arcanus hissarianus Semenov, 1900
Male adult
Tadzhikistan, near Dushanbe, env. Vill. Varzob, h = 1120 m, 38º46’27” N / 68º49’03” Е, 19.IV.1992, leg. V.Kukuruza
Copyright © Igor Solodovnikov
Carabus (Deroplectes) klapperichianus coiffaitianus Deuve, 1990
Male adult
Tadzhikistan: Badakhshan: Ishkashim Mt.R.: Kuh-i-Lal, VI.1979 leg. I.K. Lopatin (TOPOTYPUS)
Copyright © Igor Solodovnikov
Carabus (Deroplectes) klapperichianus coiffaitianus Deuve, 1990
Female adult
Tadzhikistan: Badakhshan: Ishkashim Mt.R.: Kuh-i-Lal, VI.1979 leg. I.K. Lopatin (TOPOTYPUS)
Copyright © Igor Solodovnikov
Carabus (Deroplectes) klapperichianus salangicus Kleinfeld & Reuter, 2010 Paratype
Female adult
N-Afghanistan, Hindukush Mt.Rng, Salang Pass, Nordseite, 25°27'.494N, 68°58.399'E, ca. reg. subalp., h = 2200-2400 m, V.2010, leg. Christoph Reuter
Copyright © Igor Solodovnikov
-
Diocarabus taxon page (45 photos)
Carabus (Diocarabus) loschnikovi loschnikovi Fischer, 1823
Male adult
Komi Resp., Pecero-Ilych Nat. Res., Zajachij Kamen h~786 62°10'06" N 59°12'53" E 8.VIII.2004 M. Chernjakhovski leg.
L=15.0 mm
Copyright © Evgeniy Zinoviev
Carabus (Carpatophilus) linnaei linnaei Panzer, 1812
Male adult
Carabus (Carpatophilus) linnaei linnaei Panzer, 1812
Copyright © Safronov
Carabus (Diocarabus) opaculus opaculus Putzeys, 1875
Male adult
Rusia, Kunashir isl.,<br /> 8.1991 Shestopalov M. leg.<br /> male, 15 mm
Copyright © Alexander Anichtchenko
Carabus (Diocarabus) opaculus opaculus Putzeys, 1875
Male adult
Japan, Hokkaido, <br /> Hokuryu-cho 260 m., Museushi (West), 43:48’37’N;141:40’13’E, 6-26.VII.2010, I.Rapuzzi & L.Caldon leg.
Copyright © Ruslan Panin
Carabus (Diocarabus) lucepunctus ignipunctatus Rapuzzi, 2009 Paratype
Male+Female adult
femele, 18 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) loschnikovi tsaganensis Obydov, 2001
Male adult
Russia, W Tuva,
Kyzyl-Tayga Mt Range h=1500 m, Ak-Suk riv vall., 21.VI.2006, A.& N. Menstschikovy leg.
L=16 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) caustomarginatus Imura & Mizusawa, 1994 (as paegamensis) Holotype Paratype
Male+Female adult
1.-20.6.2006, Nordkorea, Chonchon County, Mt.Paegam, 1825m, (40.80/126.20), leg. einheimischer Sammler; coll. A. Mьller (Wien)
Copyright © A.Muller
Carabus (Diocarabus) fraterculus odaesaniculus Deuve, 1991
Female adult
S Korea, South Odea Mt., 21.06.2003
Copyright © Anton Menstschikov
Carabus (Diocarabus) linnaei macairei Dejean, 1826
Female adult
SW-Romania, Banat, Resita - Valiug, 29.V- 9.VI.2005, Krause R. , Baros, Pittner leg.<br /> L=19 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) slovtzovi rasae Obydov, 2001 Holotype
Male adult
Russia, SE Tuva, pass Balyktyk-Hem, 2300 м, 50°17'N 96°23'E, 27.VI-5.VII.1996, D. Obydov leg.
Copyright © Dmitri Obydov
Carabus (Diocarabus) slovtzovi slovtzovi Mannerheim, 1849
Female adult
Russia, East Sayans,Tunkunskye Goltsy Mts., near Hoito-Gol vill., D.E.Lomakin leg
L=12,5 mm.
Copyright © Anton Menstschikov
Carabus (Diocarabus) slovtzovi slovtzovi Mannerheim, 1849
Female adult
SW Burjatia, 20 km. S. Kyren vill., Zungol r., 21.06.2007 D.E. Lomakin leg.
Copyright © Anton Menstschikov
Carabus (Diocarabus) slovtzovi slovtzovi Mannerheim, 1849
Male adult
S Siberia, Burjatia reg., E Sayan Mts., Khugaituma mt., h=1700 m, 1-5. VII.2001
L=14,3 mm
Copyright © Maksim Smirnov
Carabus (Diocarabus) massagetus massagetus Motschulsky, 1846
Male adult
Khakassia, Bograd Distr., env. Borozda Vill., forest VII.2006, M.Shestopalov leg.
L=18.0 mm
Copyright © Kirill Makarov
Carabus (Diocarabus) loschnikovi loschnikovi Fischer, 1823
Male adult
RUSSIA, Tuva, Tanu-Ola Mts., near Shurmak village, h=1500m , 23-30.VI.1997
Copyright © - - -
Carabus (Diocarabus) fraterculus fraterculus Reitter, 1895
Male adult
Kaolingzu st., Kirin pr., 18.VII.1990<br /> L=16 mm
Copyright © Alexander Anichtchenko
Carabus (Diocarabus) dorogostaiskii Shilenkov, 1983
Female adult
Chita reg., Sokhondo, Kyra vill., 18.VII.1990 Shilenkov V. leg.
L= 15 mm
Copyright © Alexander Anichtchenko
Carabus (Diocarabus) chamissonis Fischer, 1822
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Carabus (Diocarabus) caustomarginatus Imura & Mizusawa, 1994
Male adult
Copyright © Alexander Anichtchenko
Carabus (Diocarabus) beybienkoi Kryzhanovskij, 1973
Male adult
S Sakhalin, Susunaj Mt.R. 10.08 - 13.09.2003 D. Kochetkov leg.
L=14,5 mm
Copyright © Maksim Smirnov
Carabus (Diocarabus) linnaei linnaei Panzer, 1812
Male adult
W-Ukraine, NW podolian, <br /> Lviv Reg., Zhydachiv distr., near Vybranivka, 17.II.2007,Panin R. leg.<br /> female, 21 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) fraterculus fraterculus Reitter, 1895
Male adult
N-Korea, N Pyongan Prov.,Sakju, Mt Ponsan, 20-29.VI.2009, L. Bieber leg.<br /> L=19 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) linnaei linnaei (as scopolii Dejean, 1826)
Female adult
W Ukraine, NW-Podolian, near Lviv city, Vynnykivskyi forest, 20.IV.-13.V.2011, R. Panin leg.<br /> L=22 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) aurocinctus Motschulsky, 1846
Female adult
det. K. Makarov
Copyright © Alexander Anichtchenko
Carabus (Diocarabus) massagetus massagetus Motschulsky, 1846 (alpin form)
Male adult
Russsia, Altay, Ukok upland, Kaldzhin-Kul Lake h=2400-2550 m, 49 *19*N/87*26*E, 16-27.VII.2008, R.Dudko leg. L=17.5 mm
Copyright © Ruslan Panin
Carabus groenlandicus Dejean, 1831 (=chamissonis) Lectotype
Male adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC64 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec64
Copyright © - - -
Carabus (Diocarabus) massagetus pusillus Lapouge, 1915 Lectotype
Female adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC526
Copyright © - - -
Carabus (Diocarabus) linnaei folgariacus Born, 1913
Male adult
NE-Italy, Trentino-Alto Adige (Südtirol), N of vill. San Michele vill., 13-17.VI.2016, R. Janak leg. L=20 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) linnaei linnaei Panzer, 1812
Male adult
W.UKRAINE, Lviv Reg., NE Carpathians, Skolivski Beskidy Mts. Zelemyanka Mt. Rng h=600m, 15.VII-12VIII.2014, Panin R. leg. L=19 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) linnaei linnaei Panzer, 1812
Female adult
W Ukraine, Transcarpathian Reg., Rakhiv distr., Marmarosh Mts., h~400 m, 19-23.VII.2010, Reserve "Kuziy", Panin R. leg. L=23 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) linnaei linnaei Panzer, 1812
Male adult
W-UKRAINE, Ivano-Frankivsk region, S Gorgany Mts, Synyk Mt. h =1630 м., 15.V-18.VI.2010, Panin R. leg. L=19 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) linnaei linnaei (as scopolii Dejean, 1826)
Male adult
W Ukraine, Lviv reg., NW Podolian Upland, forest near Vybranivka vill., 12.X.2007, Panin R. leg. L=19 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI.-5,VIII.2017, R. Panin R. & Yu.Kanarsky leg.
Copyright © Ruslan Panin
Carabus (Diocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI.-5,VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=17 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Female adult
Male adult W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI.-5,VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=18 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI.-5,VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=17.5 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Holotype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=17.5 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L =17 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=16.5 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=17 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Female adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=17 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Female adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=18 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Female adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=19 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=19 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Female adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=20 mm
Copyright © Ruslan Panin
Carabus (ODiocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=16.5 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=19 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Gutyn-Tomnatyk h~2015 m alpin zone, 8.VI-28.VII.2011, R. Panin R. leg. L=14 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Female adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Gutyn-Tomnatyk h~2015 m, alpin zone, 8.VI-28.VII.2011, R. Panin R. leg. L=18 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg.
Copyright © Ruslan Panin
Carabus (Diocarabus) pusillus Lapouge, 1915 Paralectotypes
Female adult
Image courtesy of MNHN
Copyright © MNHN
Carabus (Diocarabus) fraterculus neochinensis Deuve & Li, 1998
Female adult
NE-China: Liaoning prov., Fengcheng City, Shicheng, 20-28.VI.2011, Li Jingke leg., L=22 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) fraterculus neochinensis Deuve & Li, 1998
Male adult
NE-China: Liaoning prov., Fengcheng City, Shicheng, 20-28.VI.2011, Li Jingke leg., L=20 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) slovtzovi sainakensis Obydov, 2010 Holotype
Male adult
Tuva, Akademika Obrucheva Mt Ridge, 2100 m, N52°00’ E95º34’, Sainak Pass, 2.VII.1998, D. Obydov leg.
Copyright © - - -
Carabus (Diocarabus) loschnikovi tsaganensis Obydov, 2001 Paratype
Male adult
Россия, Ю-З Тува, хр. Цаган-Шибэту, 2230 м, окр. Мугур-Аксу, 50°20'N 90°40'E, 22.VII.2000, Д. Обыдов leg.
Copyright © Dmitri Obydov
Carabus (Diocarabus) slovtzovi mugurensis Gottwald, 1993
Male adult
Ю-З Тува, окр. Мугур-Аксу, около оз. Хиндиктиг-Холь, 2500 м, 50°21'N 89°56'E, 24-25.VII.2000, Д. Обыдов leg.
Copyright © Dmitri Obydov
Carabus (Diocarabus) opaculus sladkovi Obydov, 2008 Holotype Paratype
Male+Female adult
Россия, Курильские острова, о. Итуруп, Баранский вулкан, 16.VII.1976, А. Сладков leg.
Copyright © Dmitri Obydov
Carabus (Diocarabus) beybienkoi medioinsularis Deuve, 1991
Male adult
Россия, Сахалин, Тымовск, 14.VII.1993, Г. Васильев leg.
Copyright © Dmitri Obydov
Carabus (Diocarabus) lucepunctus Cavazzuti & Rapuzzi, 2005 Holotype
Male adult
Korea sett., Yanggangdo, Mt. Kuandu-Hyesan. 20/28-VII-2003 (in coll. Rapuzzi)
Copyright © - - -
Carabus (Diocarabus) dorogostaiskii Shilenkov, 1983
Female adult
Chita reg., Malhanskii Mt. rng., Cheremhovskii pass, 1640m, 23.08.1998, 50.539176, 108.743628, Anichtchenko A. leg.
Copyright © Alexander Anichtchenko
Carabus (Diocarabus) beybienkoi Kryzhanovskij, 1973
Female adult
Sakhalin isl., 44 kordon, 6.8.1991, Generalov A. leg. (cAA).
Copyright © Alexander Anichtchenko
Carabus (Diocarabus) aurocinctus Motschulsky, 1846
Male adult
Magadan reg. Palatka vill., 26.7.1992 Kuznetsov D. leg. (cAA)
Copyright © Alexander Anichtchenko
Carabus (Diocarabus) aurocinctus Motschulsky, 1846
Female adult
Buryatia, lake Baikal shore, 52.987835, 108.304803, Goriachinsk vill. 12.8.1997б Anichtchenko A. leg (cAA)
Copyright © Alexander Anichtchenko
Carabus (Diocarabus) linnaei folgariacus (as tauernensis Mandl, 1985)
Female adult
Austria, Steiermark, Rottenmanner Tauern, h=1000 m, Strechengraben, 13.VII.2012, L=20 mm
Copyright © Ruslan Panin
Carabus (Diocarabus) massagetus sajanicola Deuve, 2010 Holotype
Male adult
Siberie, Sayanogorsk, mount Sayan, 25.5.1991 (MNHN)
Copyright © Thierry Deuve -
Eccoptolabrus taxon page (22 photos)
Carabus (Lasiocoptolabrus) sunwukong sunwukong Imura, 1993 Paratype
Male adult
Copyright © Boleslav Brezina
Carabus (Lasiocoptolabrus) sunwukong baotianmanus (Imura, 2005) Paratype
Male adult
Copyright © Boleslav Brezina
Carabus (Eccoptolabrus) exiguus wudumontanus Imura, 1998
Male adult
male, 19mm
Copyright © Boleslav Brezina
Carabus (Eccoptolabrus) exiguus tagong Kleinfeld & Puchner, 2007 Holotype Paratype
Female adult
female, 20 mm
Copyright © Frank Kleinfeld
Carabus (Eccoptolabrus) exiguus tagong Kleinfeld & Puchner, 2007 Paratype
Female adult
female, 20 mm
Copyright © Frank Kleinfeld
Carabus (Eccoptolabrus) exiguus tagong Kleinfeld & Puchner, 2007 Holotype
Male adult
male, 19 mm
Copyright © Frank Kleinfeld
Carabus (Eccoptolabrus) exiguus ochotonarum Brezina, 1996 Holotype Paratype
Female adult
Copyright © Frank Kleinfeld
Carabus (Eccoptolabrus) exiguus ochotonarum Brezina, 1996
Female adult
CH. S-Gansu (Labrang) valley E of Ponggartang
35:14N/102:55E. ca. 2700m conoferous forest
9.VII.1995 K. & B. Brezina
Copyright © Frank Kleinfeld
Carabus (Eccoptolabrus) exiguus ochotonarum Brezina, 1996 Paratype
Male adult
Copyright © Frank Kleinfeld
Carabus (Eccoptolabrus) exiguus nivium Breuning, 1933
Male adult
male, 18mm
Copyright © Boleslav Brezina
Carabus (Eccoptolabrus) exiguus nivium Breuning, 1933
Female adult
female, 20mm
Copyright © Boleslav Brezina
Carabus (Eccoptolabrus) exiguus nivium Breuning, 1933
Male adult
female, 18mm
Copyright © Boleslav Brezina
Carabus (Eccoptolabrus) exiguus nivium Breuning, 1933 Holotype Paratype
Female adult
female, 19 mm
Copyright © Frank Kleinfeld
Carabus (Eccoptolabrus) exiguus nivium Breuning, 1933
Male adult
male, 18 mm
Copyright © Frank Kleinfeld
Carabus (Eccoptolabrus) exiguus lanzhouicus Deuve, 1989
Male adult
male, 17mm
Copyright © Boleslav Brezina
Carabus (Eccoptolabrus) exiguus lanzhouicus Deuve, 1989
Male adult
male, 19mm
Copyright © Boleslav Brezina
Carabus (Eccoptolabrus) exiguus fanianus Imura, 1993 Paratype
Male adult
male, 18mm
Copyright © Boleslav Brezina
Carabus (Eccoptolabrus) exiguus fanianus Imura, 1993
Female adult
female, 19mm
Copyright © Boleslav Brezina
Carabus (Eccoptolabrus) exiguus exiguus Semenov, 1898
Female adult
female, 19mm
Copyright © Boleslav Brezina
Carabus (Eccoptolabrus) exiguus exiguus Semenov, 1898 Holotype
Male adult
male, 18 mm
Copyright © Frank Kleinfeld
Carabus (Eccoptolabrus) exiguus evae Brezina et Hackel, 2007 Holotype
Male adult
China, S-Qinghai, 5km S Baima, 30.VI.-1.VII.2006 V. Banes
long of imago 19 mm
Copyright © Boleslav Brezina
Carabus (Eccoptolabrus) exiguus cordimeatus Cavazzuti, 1997 Holotype
Male adult
male, 19mm
Copyright © Boleslav Brezina
Carabus (Eccoptolabrus) exiguus cithara Cavazzuti, 2002 Holotype
Female adult
female, 22mm
Copyright © Boleslav Brezina
Carabus (Eccoptolabrus) exiguus alucinatus Cavazzuti, 2006 Holotype
Female adult
female, 21mm
Copyright © Boleslav Brezina -
Eucarabus taxon page (111 photos)
Carabus (Eucarabus) obsoletus carpathicus Palliardi, 1825
Male+Female adult
female, 25 mm
SW-Romania, Banat, Resita - Valiug, 3-8.VI.2007, Krause R. leg.
Copyright © Ruslan Panin
Carabus (Eucarabus) sternbergi Roeschke, 1898
Female adult
Korea, Gyeongi Do,
Bokwang Sa 30.2.1976
female, 28 mm
Copyright © Alexander Anichtchenko
Carabus (Eucarabus) pawlowskianus Deuve, 1989
Female adult
N Korea, Pong-San Mt.
12.7.1997
female, 25 mm
Copyright © Alexander Anichtchenko
Carabus (Eucarabus) nitididorsus Ishikawa & Kim, 1983
Male adult
S Korea, Deagu, 25.8.2003
Copyright © Alexander Anichtchenko
Carabus (Eucarabus) nitididorsus Ishikawa & Kim, 1983
Female adult
Korea, Peak Chonwongbong, 1600-1900m
Mt. Chiri-San, Kyongsangnam-Do, 18.7.87 Y. Imura leg.
Copyright © Alexander Anichtchenko
Carabus (Eucarabus) manifestus guanmenshanus Imura, 1991
Female adult
China, Liaoning, Mt. Lao Gu Go, Dandong, 29.5.97
Copyright © Alexander Anichtchenko
Carabus (Eucarabus) billbergi billbergi Mannerheim, 1827
Male adult
Russia, E Siberia, Buryatia, lake Baikal, near Il'inka vill. Anichtchenko A. leg.
Copyright © Alexander Anichtchenko
Carabus (Eucarabus) ? manifestus Kraatz, 1881
Male adult
Tian tou, Yueshi Counti, Anhui, China, 1-10.08.2004
Copyright © Safronov
Carabus (Eucarabus) ulrichi s.str. (as volhynicus Lutshnik, 1927)
Male+Female adult
W Ukraine, NW Podolian,
Ternopil reg., Pidvolochisk distr., forest Bogdanivskiy, near Romanove Selo, 7-24.V.2008, Panin R. & Solodovnikov I. leg.
Copyright © Ruslan Panin
Carabus (Eucarabus) ulrichi ulrichi Germar, 1824 (as podolicus Semenov, 1898 loc.typ)
Female adult
C-UKRAINE,<br /> Vinnitsa Reg., near Khmilnyk, forest, 10-20.VIII.2002, Demidov G.N. leg., det. Panin R<br /> female, 30mm
Copyright © Ruslan Panin
Carabus (Eucarabus) ulrichi ulrichi Germar, 1824
Male adult
NW Ukrainian Carpathians
Lviv Reg., Staryi Sambir Distr., near Dobromyl , h=500m 9.VI-25.VII.2007, Panin R. Leg.
Copyright © Safronov
Carabus (Eucarabus) billbergi billbergi Mannerheim, 1827
Male adult
Rusia, Primorskii reg., Kiparisovo vill., 5.8.1993
Chichvarhin A. leg.
male, 20 mm
Copyright © Alexander Anichtchenko
Carabus (Eucarabus) billbergi billbergi Mannerheim, 1827
Male adult
Rusia, Primorskii reg., Anuchinskii distr.,
near LZU-3, 7.8.1979 Bakurov leg.
male, 21 mm
Copyright © Alexander Anichtchenko
Carabus (Eucarabus) manifestus pyonganicola Deuve & Li, 1998
Male+Female adult
N-Korea: Pyonganam-Do, YangTok County,(39;127), 12-19,VI.2006
Copyright © Ruslan Panin
Carabus (Eucarabus) manifestus guanmenshanus Imura, 1991
Female adult
NE China, Liaoning, Anshan City, Mt.Qian Shan, 4-16.VII.2005, I.Rapuzzi leg.
L=26 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) obsoletus obsoletus Sturm, 1815
Male adult
NW Ukrainian Carpathians, Lviv Reg., Staryi Sambor Distr., near Dobromyl , h=500 m., 15-24.VI.2005, Panin R. leg., male, 28mm
Copyright © Ruslan Panin
Carabus (Eucarabus) obsoletus obsoletus Sturm, 1815
Male adult
NW Ukrainian Carpathians,
Lviv reg., Staryi Sambor Distr., near Dobromyl , h=500m 6-28.VI.2009, Panin R. leg.
male, 27 mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Eucarabus) catenulatus catenulatus Scopoli, 1763
Male adult
Lombardy - North Italy
Copyright © Monzini
Carabus (Eucarabus) catenulatus catenulatus Scopoli, 1763
Female adult
NW Croatia, Bosanci, 1-12,VI.2004, Baros Slovak leg.<br /> L=33 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) catenulatus catenulatus Scopoli, 1763
Male adult
Italia, Riva San Vitale (Chleti), VI.1996 Rubizzani leg., 29 mm
Copyright © Alexander Anichtchenko
Carabus (Eucarabus) cumanus Fischer, 1823
Male+Female adult
NW Caucasus, Russia,
Krasnodar reg.,Tuapse distr., Mt.Semashkho 04.V.2008, D.Fominyh leg.
female, 24mm
Copyright © Ruslan Panin
Carabus (Eucarabus) italicus italicus Dejean, 1826
Male adult
Italia, Emilia Romagna, Serramazzoni (Modena)<br /> 700m, 9.VI.1990 M. Po leg.<br /> male, 22 mm
Copyright © Alexander Anichtchenko
Carabus (Eucarabus) italicus italicus Dejean, 1826
Male adult
NE-ITALY, Veneto, Hiane, 14.V.1986 Sandec F. leg. L= 23mm
Copyright © Ruslan Panin
Carabus (Eucarabus) manifestus manifestus Kraatz, 1881
Male adult
China, Beijing, Xiaolongmeng, 2005, Liu Ye coll.;
L=20mm
Copyright © Ye Liu
Carabus (Eucarabus) ulrichi arrogans Schaum, 1858 (as gornjakensis Pavicevic & Toševski, 19
Male+Female adult
NE-Serbia, Serbian Carpathians,
Branicevskom reg., Šetonje, Gornjak, 27.V-10.VI.2006, R.Krause leg.
Copyright © Ruslan Panin
Carabus (Eucarabus) ulrichi arrogans Schaum, 1858 (as ceremosnjensis Pavicevic & Toševski,
Male+Female adult
NE-Serbia,
Serbian Carpathians, Homolje Mts., Krepoljin 20-31.V.2006, Sima leg.
female, 29mm
Copyright © Ruslan Panin
Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825 (as superboides Lie, 1989)
Female adult
SW-Romania, Banat,
Oravita, 5-7.VI.2007, Krause Rene leg.
female,30mm
Copyright © Ruslan Panin
Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825
Male adult
NE Serbien, Negotim .
V.1981
male, 25 mm
Copyright © Alexander Anichtchenko
Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825 (as superbus Kraatz, 1878)
Male+Female adult
SW-Romania,
Banat, Moldova Noua, Ravenska viil., 29.V-12.VI.2006, Krause Rene leg.
female,31mm
Copyright © Ruslan Panin
Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825
Female adult
female, 32mm
Copyright © Boleslav Brezina
Carabus (Eucarabus) ulrichi rhilensis Kraatz, 1876 (as sokolari Born, 1904)
Male+Female adult
Czech Rep., W Bohemia,
Plzen reg., Tachov Distr., Plana u Marianskych Lazni, 17-29.V.2008, S.I.Budka leg.
female, 26 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) obsoletus nagyagensis Seidlitz, 1888
Female adult
NW-Romania, Mts Ses (Plopis),Cluj Reg., Bucea env, h=580 m, 8.VI.2011, Sl.Budka leg.
L=26 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825
Male+Female adult
Male - SW Romania, Banat - Bocsa/Coltan,
21.V-05.VI.2010, R. Krause leg., L=29 mm.
Female - SW Romania, Banat - Saska Montana, Stinapari,
25-VI-2010, R. Krause leg., L=31 mm.
Copyright © Alexandr Safronov
Carabus (Eucarabus) obsoletus nagyagensis (as paranagyagensis Lie, 1990)
Male adult
W Romania, Arad reg., Zarand Mts (Muntii Zarandului), Nadas-Barzavah h=142 m, J. Kynkor leg.
L=24 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) ulrichi rhilensis (as stussineri Géhin, 1885)
Male adult
W Croatia, Ivanščica Mts , Ivanec, 24.VI-7.VII.2006, Krause Rene leg.
L=29 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) ulrichi ulrichi (as intercessor Sokolar, 1912)
Male adult
N Hungary, Pilis Mts., Piliszentlelek, 20-27.VII.2011, Krause Rene leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) ulrichi arrogans Schaum, 1858
Male adult
NE-Serbia:Majdanpek, 9-11,V.2010, L.Brhel leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) obsoletus obsoletus Sturm, 1815 (as prunneri Mallasz, 1901)
Male adult
NW Romania, Apuseni Mts, Trascău Mt Rng, Lungesti vill., 25 km W Turda,.VI.2012
L=26 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) manfredeggeri Müller 2012 Paratype
Male+Female adult
N-Korea, Chagang Prov., Manpo City, Songhak-ri, 2-16.VII.2011<br /> female, 21 mm<br /> male, 19 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) sternbergi gureopensis J.K. Park & J. Park, 2013 Holotype Paratype
Male+Female adult
Is. Gureopdo (37°19′71.42″N, 125°98′72.62″E), Deokjeok archipelago, Gyeonggi-do, Korea, 19.viii.2010, J.K.Park
Image from original describtion J.K. Park & J. Park, 2013
Copyright © - - -
Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825
Male adult
SW Romania, Banat - Eibenthal; 29 mm
Copyright © Aleš Sedláček
Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825
Female adult
SW Romania, Banat - Eibenthal; 33 mm
Copyright © Aleš Sedláček
Carabus (Eucarabus) ulrichi arrogans Schaum, 1858
Male adult
NE Serbia, Majdanpek; 23 mm
Copyright © Aleš Sedláček
Carabus (Eucarabus) ulrichi arrogans Schaum, 1858
Female adult
NE Serbia, Majdanpek; 29 mm
Copyright © Aleš Sedláček
Carabus (Eucarabus) catenulatus fontanellae Reitter, 1896
Male adult
W-Croatia, Cres island,VIII.1994, G. Zannna leg. L=33 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) cartereti omisanensis Lassalle, 1999
Male adult
South Korea, Gangwon Province, Kyebang-san, h=1100m, VI.1998 L=24 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) obsoletus nagyagensis (as fossulifer Fleischer, 1893)
Male adult
NW-Romania, Bihor, Mts Seș (Plopis), 20 km N Aleșd, Pădurea Neagră env., VI.2014, J.Kynkor leg. L=24 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) obsoletus obsoletus (as csikii Mallasz, 1901)
Female adult
NE Carpathians, Marmarosh Mts., Transcarphatian Reg., near Lug vill., h=350 m, 26.IV-8.VI.2014, R.Panin & Yu. Kanarsky leg. L=30 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) obsoletus obsoletus (as csikii Mallasz, 1901)
Female adult
NE Carpathians, Marmarosh Mts., Transcarphatian Reg., near Lug vill., h=350 m, 26.IV-8.VI.2014, R.Panin & Yu. Kanarsky leg. L=32 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) obsoletus obsoletus var sacheri Thomson, 1875 Syntypes
Male adult
3958:2 (MZLU)
Copyright © Lund University
Carabus (Eucarabus) italicus rostagnoi Luigioni, 1904
Female adult
C-Italy, Lazio, Fondi env.,10.II.1979, L = 25 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) parreyssi parreyssi Palliardi, 1825
Male adult
W-Croatia, NP Northern Velebit, Zavižan,15.V-13.VI.1996, Colombetta leg., L=21 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) angustus dopyeong Rapuzzi, 2015
Male adult
scale bar = 5mm
Copyright © Jong-Bong Choi
Carabus (Eucarabus) sternbergi sternbergi Roeschke, 1898
Male adult
Scale bar = 5mm
Copyright © Jong-Bong Choi
Carabus (Eucarabus) cartereti cartereti Deuve, 1982
Male adult
Scale bar = 5mm
Copyright © Jong-Bong Choi
Carabus (Eucarabus) nitididorsus nitididorsus Ishikawa & Kim, 1983
Male adult
Scale bar = 5mm
Copyright © Jong-Bong Choi
Carabus (Eucarabus) nitididorsus nitididorsus Ishikawa & Kim, 1983
Male adult
Scale bar = 5mm
Copyright © Jong-Bong Choi
Carabus (Eucarabus) nitididorsus nitididorsus Ishikawa & Kim, 1983
Male adult
Scale bar = 5mm
Copyright © Jong-Bong Choi
Carabus (Eucarabus) nitididorsus paegunsanus Imura, 1997
Male adult
Scale bar = 5mm
Copyright © Jong-Bong Choi
Carabus (Eucarabus) nitididorsus baegyangsanus (Choi & Park, 2019) Holotype
Male adult
Scale bar = 5mm
Copyright © Jong-Bong Choi
Carabus (Eucarabus) nitididorsus baegyangsanus (Choi & Park, 2019) Paratype
Male adult
Scale bar = 5mm
Copyright © Jong-Bong Choi
Carabus (Eucarabus) namhaedoensis Kwon & Lee, 1984
Male adult
Scale bar = 5mm
Copyright © Jong-Bong Choi
Carabus (Eucarabus) namhaedoensis Kwon & Lee, 1984
Male adult
Scale bar = 5mm
Copyright © Jong-Bong Choi
Carabus (Eucarabus) ulrichii (as podolicus Semenov, 1898 loc.typ. )
Male adult
CW-UKRAINE, Vinnytsia Distr., Zarvantsi vill., 1-25.V.2020, Kokhan S. leg./det. R.Panin,
Copyright © Ruslan Panin
Carabus (Eucarabus) manifestus guanmenshanus Imura, 1991
Male adult
China, Liaoning, Tieling County, Sandagou env., h=850 m, 13-18.VI.2011
Copyright © Ruslan Panin
Carabus (Eucarabus) billbergi kuandianicus Rapuzzi, 2007
Female adult
NE China, Liaoning, Kuandian County, Dachuantou env., 20-29.VI.2015,
Copyright © Ruslan Panin
Carabus (Eucarabus) xiuyanensis Deuve & Li, 1998
Male adult
NE-China: SE-Liaoning, Xiuyan County, Dayingzi, Mt. Laopingding, 16.VI.2015, L=24.5 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) xiuyanensis Deuve & Li, 1998
Female adult
NE-China: SE-Liaoning, Xiuyan County, Dayingzi, Mt. Laopingding, 16.VI.2015,
Copyright © Ruslan Panin
Carabus (Eucarabus) billbergi kuandianicus Rapuzzi, 2007
Male adult
NE China, Liaoning, Kuandian County, Dachuantou env., 20-29.VI.2015, L=23 mm
Copyright © Ruslan Panin
Carabus (Carabus) szeli obong Rapuzzi, 2015 Holotype Paratype
Male+Female adult
Democratic Republic of Korea, North Hamgyong Province, Mt. Obong, (ca 42.40 NL; 129.80 EL), 1/15.VII.2012. Image from Rapuzzi, 2015
Copyright ©
Carabus (Carabus) angustus dopyeong Rapuzzi, 2015 Holotype Paratype
Male+Female adult
R. of Korea, Geonggi-do/Gangwondo provincial border, Pass North from Dopyeong, 600 m, 3/13.VII.2012, I. Rapuzzi and L. Caldon leg. Image from Rapuzzi, 2015
Copyright © - - -
Carabus (Carabus) sternbergi gimhwa Rapuzzi, 2015 Holotype Paratype
Male+Female adult
R. of Korea, Gyeonggi-do/Gangwon-do provincial border, Pass North from Dopyeong, 600 m, 3/13.VII.2012, I. Rapuzzi and L. Caldon leg. Image from Rapuzzi, 2015
Copyright © - - -
Carabus (Carabus) sternbergi goheungicus Rapuzzi, 2015 Holotype
Female adult
Republic of Korea, Goheung peninsula, Paryeongsan, 200m, 10/17.VII.2012, I. Rapuzzi and L. Caldon legit. Image from Rapuzzi, 2015
Copyright © - - -
Carabus (Carabus) sternbergi jindoensis Rapuzzi, 2015 Holotype Paratype
Male+Female adult
R. of Korea, Jindo Island, 8.VIII.2010. Image from Rapuzzi, 2015
Copyright © - - -
Carabus (Carabus) sternbergi deogyusan Rapuzzi, 2015 Holotype Paratype
Male+Female adult
Republic of Korea, Cholla Bukdo, Deogyusan, 10 km North from Gucheondong, 550 m, 7/15.VII.2007, I. Rapuzzi and L. Caldon leg. Image from Rapuzzi, 2015
Copyright © - - -
Carabus (Eucarabus) cartereti peacedam Rapuzzi, 2015 Holotype Paratype
Male+Female adult
R. of Korea, Gangwondo, “Peace Dam”, NE Hwacheon 400 m, 3/13.VII.2012, I. Rapuzzi and L. Caldon leg. Image from Rapuzzi, 2015
Copyright © - - -
Carabus (Eucarabus) parreyssi parreyssi Palliardi, 1825
Female adult
SW-Croatia, Mts Velebit, Mali Halan, h=1390 m, 23.V-20.VI.2022, L=24 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) parreyssi parreyssi Palliardi, 1825
Male adult
SW-Croatia, Mts Velebit, Mali Halan, h=1390 m, 23.V-20.VI.2022, L=22.5 mm
Copyright © Ruslan Panin
Carabus (Eucarabus) szeli szeli Holotype
Male adult
Image from: Korean type specimens of insects deposited in Hungarian Natural History Museum.
Copyright © - - -
Carabus (Eucarabus) catenulatus fontanellae Reitter, 1896
Male adult
W-Croatia, Dalmatia, Isl. Rab, 16-30.V.2019, G. Proscia leg.
Copyright © Ruslan Panin
Carabus (Eucarabus) manifestus pyonganicola Deuve & Li, 1998
Male adult
Mt. Changbai Shan, Antu county, Jilin province, China. 13/15.VI.2012. Er-dao-bai-he, 800 m. 22.4 mm. Image from Xueping et al 2013
Copyright © - - -
Carabus (Eucarabus) szeli changbaicus Rapuzzi & Li, 2013 Holotype
Male adult
Huang-song-pu-lin-chang, 1300 m., Mt. Changbai Shan, Antu county, Jilin province, China. 13/15.VI.2012 (Coniferous forest); preserved in Ivan Rapuzzi collection, Prepotto (UD), Italy
Copyright © - - -
Carabus (Eucarabus) manifestus guanmenshanus Imura, 1991
Male adult
China, Liaoning Pr., Kuandian Co., Banlaing, Ba-Che-chuan (cAA). det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko
Carabus (Eucarabus) ulrichi rhilensis (as stussineri Géhin, 1885)
Female adult
S Austria, Styria, Leutschach, Schmirnberg, 46.634400, 15.502400, 1-25.VI.2021, M.Egger leg., L=30 mm
Copyright © Ruslan Panin -
Euleptocarabus taxon page (5 photos)
Carabus (Euleptocarabus) porrecticollis porrecticollis Bates, 1883
Male adult
Japan: Niigata pref., Chayagahara Jyouetsu City,
18.XI.2006 A.Toba leg.
Copyright © Alexander Anichtchenko
Carabus (Euleptocarabus) porrecticollis Bates, 1883
Female adult
Atsushiokanomachikano, Kitakata City, Fukushima Pref., Japan, 29.III.2014, T. Nakamura leg.
Copyright © Alexander Anichtchenko
Carabus (Euleptocarabus) porrecticollis porrecticollis Bates, 1883
Male adult
Japan, C Honshu, Nagano Pre., Toyooka town, 28.XII.2019, S. Shinichi leg. L=30 mm
Copyright © Ruslan Panin
Carabus (Euleptocarabus) porrecticollis kansaiensis Nakane, 1961
Female adult
Takatsuki (Osaka) JAPON, XI-1985
Copyright © Alain Mollard -
Eupachys taxon page (3 photos)
Carabus (Eupachys) glyptopterus Fischer, 1827
Male+Female adult
Russia, SE-Siberia,
Buryat Republic, Ivolginsky Distr., 04.VII.2010, Filippov A.E. leg.
female, 32 mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Eupachys) glyptopterus Fischer, 1827
Female adult
Mongolia, 15 km N Bayanteeg, 2000 m, 17.7.2006 Anichtchenko A. leg.
Copyright © Alexander Anichtchenko
Carabus (Eupachys) acutithorax Deuve, 1989
Male adult
China, Shanxi, Yongzhi [Yongji]
Copyright © Thierry Deuve -
Eurycarabus taxon page (29 photos)
Carabus (Eurycarabus) genei Gene, 1839
Male adult
Corse de Sud, env. de
Rondinara, VI.1998 Tingaud M. leg.
male, 23 mm
Copyright © Alexander Anichtchenko
Carabus (Eurycarabus) famini fezzanus Lapouge, 1915
Female adult
Marokko, Moy Atlas, Azrou, Ibel Hebri, 1950 m
leg. Staven/Gremmels, 2000; det. K.Staven, 2000
length: 22 mm
Foto: U.Schmidt, 2009
Copyright © U.Schmidt
Carabus (Eurycarabus) famini piraticus Fairmaire, 1880
Male adult
Morocco, Safi 10 km
Khattazakane env. 29.XII.2001, h 100m
male, 22 mm
Copyright © Alexander Anichtchenko
Carabus (Eurycarabus) famini fezzanus Lapouge, 1915
Male adult
Marokko, Moyen Atlas, s. Azrou
Jbel Hebri, zedernwald, 1950m, 31.III.2000 Staven/Gremmels leg.
male, 22 mm
Copyright © Alexander Anichtchenko
Carabus (Eurycarabus) famini favieri Fairmaire, 1859
Female adult
N Africa, 2 km W Ceuta Allabacal, 17-18.II.2004, Anichtchenko A. leg.
female, 22 mm
Copyright © Alexander Anichtchenko
Carabus (Eurycarabus) faminii romanoi Rapuzzi & Sparacio, 2012 Holotype Paratype
Male+Female adult
Holotypus male, Godrano (PA), 25.XI.1978, legit I.Sparacio.<br /> photo from Ivan Rapuzzi & Ignazio Sparacio, 2012
Copyright © - - -
Carabus (Eurycarabus) famini famini Dejean, 1826 Holotype
Male adult
(MNHN)
photo from Ivan Rapuzzi & Ignazio Sparacio, 2012
Copyright © ---
Carabus (Eurycarabus) famini lucasi Gaubil, 1849
Male adult
N-Tunisia, Bizerte reg., Menzel Bourguiba city,h=5 m, XII.1981, S. Remond leg.
L=25 mm
Copyright © Ruslan Panin
Carabus (Eurycarabus) famini fezzanus Lapouge, 1915
Male adult
Morocco, Middle Atlas Mt Rng, Azrou city h=1500m, IV.1999, S. Remond leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Eurycarabus) famini piraticus (as mamorensis (Kocher, 1964)
Female adult
NW-Morocco, betw.Meknes/Kenitra, Foret de Mamora, II.2008
L= 27 mm
Copyright © Ruslan Panin
Carabus (Eurycarabus) famini itoensis Antoine, 1939 Paratype
Female adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC1049 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec1049
Copyright © - - -
Carabus (Eurycarabus) famini favieri Fairmaire, 1859 Holotype
Female adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC143 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec143
Copyright © - - -
Carabus (Eurycarabus) famini grasselyi Antoine, 1941 Holotype
Female adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC1044 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec1044
Copyright © - - -
Carabus (Eurycarabus) famini mairei Antoine, 1925 Holotype
Female adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC1047 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec1047
Copyright © - - -
Carabus (Eurycarabus) famini itoensis Antoine, 1939 Holotype
Male adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC1048 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec1048
Copyright © - - -
Carabus (Eurycarabus) famini atlantis Alluaud, 1923 Holotype
Male adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC1058 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec1058
Copyright © - - -
Carabus (Eurycarabus) famini algiricus Géhin, 1885 Holotype
Male adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC433 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec433
Copyright © - - -
Carabus (Eurycarabus) famini piraticus Fairmaire, 1880 Lectotype
Male adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC144 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec144
Copyright © - - -
Carabus (Eurycarabus) famini atlantis Alluaud, 1923
Female adult
C Morocco, Haut Atlas, SW of Oukaïmeden h=2900m, 31.173644, -7.844442, 29.IV-11.V.2024, C.Reuter leg., L=21 mm
Copyright © Ruslan Panin -
Fulgenticarabus taxon page (2 photos)
Carabus (Fulgenticarabus) flutschi koreus Deuve, 2006
Male adult
N Korea, Samjiyon, Ryang Gang Province
6-12.6.2006 leg. Wong
Copyright © Boleslav Brezina
Carabus (Fulgenticarabus) flutschi koreus Deuve, 2006
Female adult
female, 22mm
Copyright © Boleslav Brezina -
Gnathocarabus taxon page (2 photos)
Carabus (Gnathocarabus) kusnetzovi Semenov, 1903
Female adult
female, 21mm
Copyright © Boleslav Brezina
-
Goniocarabus taxon page (14 photos)
Carabus (Goniocarabus) tadzhikistanus Kryzhanovskij, 1968
Female adult
female, 28mm
Copyright © Boleslav Brezina
Carabus (Goniocarabus) tadzhikistanus Kryzhanovskij, 1968
Male adult
male, 25mm
Copyright © Boleslav Brezina
Carabus (Goniocarabus) gussakovskii Kryzhanovskij, 1971
Female adult
female, 23mm
Copyright © Boleslav Brezina
Carabus (Goniocarabus) gussakovskii Kryzhanovskij, 1971
Male adult
male, 22mm
Copyright © Boleslav Brezina
Carabus (Goniocarabus) gussakovskii Kryzhanovskij, 1971
Male adult
male, 21mm
Copyright © Boleslav Brezina
Carabus (Goniocarabus) grombczewskii Semenov, 1891
Female adult
female, 22mm
Copyright © Boleslav Brezina
Carabus (Goniocarabus) caerulans Moravitz, 1886
Female adult
female, 24mm
Copyright © Boleslav Brezina
Carabus (Goniocarabus) sogdianus Semenov, 1898
Male adult
N Tajikistan, SW Pamir-Alay, Zarafshan Range, Fann Mts, Marguzor Lake circ., Padrut vill., 16.V.1997
L=21 mm
Copyright © Ruslan Panin
Carabus (Goniocarabus) grombczewskii grombczewskii Semenov, 1891
Male adult
NC Tadzhikistan, Pamir, N-slope Peter I.Mt Rng, h=2100m, Ganishou vill.,12.VII.2006, Pak O. leg.
L=19 mm
Copyright © Ruslan Panin
Carabus (Goniocarabus) perelloi Casale, 1979
Male adult
N-Afghanistan, Hindukush, Provinz Baghlan, N of Kabul, N Salang Pass, subalp., h = 2200 m, 35°27'.494N / 68°58.399'E, Mai 2011, leg. Christoph Reuter (TOPOTYPUS)
Copyright © Igor Solodovnikov -
Hemicarabus taxon page (31 photos)
Carabus (Hemicarabus) tuberculosus Dejean, 1829
Male adult
China, Shanxi pro., Jiaochen
Copyright © Ye Liu
Carabus (Hemicarabus) tuberculosus Dejean, 1829
Male adult
Far East, Lazovsky Dist., shore of Japan Sea, 3 km NE Preobrazhenie Vill., Ezhovaya (Tasovaya) Bay 13.07.06 M. & L. Smirnov's leg.
male
L=20,0 mm
Copyright © Maksim Smirnov
Carabus (Hemicarabus) serratus Say, 1825 (as tatumi Motschulsky, 1865 )
Female adult
female, 18mm
Copyright © Boleslav Brezina
Carabus (Hemicarabus) serratus Say, 1825 (as tatumi Motschulsky, 1865 )
Female adult
female, 19mm
Copyright © Boleslav Brezina
Carabus (Hemicarabus) serratus Say, 1825 (as tatumi Motschulsky, 1865)
Male adult
male, 17mm
Copyright © Boleslav Brezina
Carabus (Hemicarabus) nitens Linne, 1758
Male+Female adult
Belarus,
Vitebsk reg., Ushachskiy distr., near Bolshie Dolcy vill. 9.V.1990, I.A.Solodovnikov leg.
female, 18mm.
Copyright © Ruslan Panin
Carabus (Hemicarabus) nitens Linne, 1758
Male adult
Russia, Ural, riv. Rezh, vill. Kamenka; 10.VII.2009, N. Menstschicova leg.
male, 14 mm.,
Copyright © Anton Menstschikov
Carabus (Hemicarabus) macleayi macleayi Dejean, 1826
Male adult
male, 15mm
Copyright © Boleslav Brezina
Carabus (Hemicarabus) macleayi macleayi Dejean, 1826
Male adult
Russia, Far East,<br /> Lazovskij Nat. Res., Proselochnaja Riv. near nouth, Tumannaja Mt. 42°59.888' N 134°06.935' E 18.VIII.2007, K.Makarov & A.Zaitsev leg.<br /> L=16 mm
Copyright © Kirill Makarov
Carabus (Hemicarabus) nitens Linne, 1758
Male adult
16,6 mm, England, Dorset, Studland
leg.det. J.W.Yerbury, 18.-29.VIII.2006
Coll. Oxford University Museum of Natural History
Photo: U.Schmidt, 2011
Copyright © Udo Schmidt
Carabus (Hemicarabus) tuberculosus Dejean, 1829
Male adult
Иркутск, Новоленино, 20 июня 1985, leg. Э.Берлов
Copyright © Oleg Berlov
Carabus (Hemicarabus) serratus Say, 1825
Male adult
E-USA,Virginia, Leesburg env., Ball's Bluff Battlefield Regional Park, 27.VIII-4.IX.2015,V. Čutka leg. L=17 mm
Copyright © Ruslan Panin
Carabus (Hemicarabus) serratus Say, 1825
Female adult
E-USA, Virginia, Leesburg env., Ball's Bluff Battlefield Regional Park, 27.VIII-4.IX.2015,V. Čutka leg. L=18 mm
Copyright © Ruslan Panin
Carabus (Hemicarabus) macleayi amanoi (Imura, 2004)
Male adult
alpine zone of Mt. Rishiri-zan, on Is. Rishiri-to, off the western coast of the northern tip of Hokkaido, Northeast Japan. Image from Imura, 2005
Copyright © - - -
Carabus (Hemicarabus) macleayi amanoi (Imura, 2004)
Male adult
alpine zone of Mt. Rishiri-zan, on Is. Rishiri-to, off the western coast of the northern tip of Hokkaido, Northeast Japan. Image from Imura, 2005
Copyright © - - -
Carabus (Hemicarabus) nitens Linne, 1758
Male adult
Belarus, Vitebsk reg., Miory distr. Swamp «Mokh», 09.05.2011, leg. G.G. Sushko
Copyright © Igor Solodovnikov
Carabus (Hemicarabus) tuberculosus Dejean, 1829
Male adult
Carabus (Hemicarabus) tuberculosus n. strophium F.-W., 1842 SE Kazakhstan: NW Tarbogatai Mt.R.
Copyright © Igor Solodovnikov
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Homoeocarabus taxon page (6 photos)
Carabus (Homoeocarabus) maeander paludis Gehin, 1885
Female adult
female, 21mm
Copyright © Boleslav Brezina
Carabus (Homoeocarabus) maeander maeander s.str.
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Carabus (Homoeocarabus) maeander paludis Gehin, 1885
Female adult
Japan, Hokkaido; Tomakomai city, Benten, 17.V.2015, Takemoto leg., L=21 mm
Copyright © Ruslan Panin
Carabus (Homoeocarabus) maeander evenkiensis Obydov, 2010 Holotype Paratype
Male+Female adult
North Siberia, Evenk Autonomous Region, N Tunguska River, 60 km up to Tura Village, Tabachnyi Stream, 19.VI.1972
Copyright © - - - -
Hygrocarabus taxon page (5 photos)
Carabus (Hygrocarabus) nodulosus Creutzer, 1799
Male+Female adult
Austria, Styria Region (Steiermark), Leibnitz, 31.X.1999, I.Rapuzzi leg.<br /> female, 29 mm
Copyright © Ruslan Panin
Carabus (Hygrocarabus) variolosus Fabricius, 1787
Male adult
W-Ukraine, NW Podolian,<br /> Lviv reg., Zhydachiv distr.,near Vybranivka vill., 15.V-8.VI.2008, Panin R. leg.<br /> male, 27mm
Copyright © Ruslan Panin
Carabus (Hygrocarabus) variolosus Fabricius, 1787
Male+Female adult
W-Ukraine,<br /> Lviv Reg., Zhydachiv distr., near Vybranivka vill., Panin R. leg.<br /> female, 30mm
Copyright © Ruslan Panin -
Imaibius taxon page (51 photos)
Carabus (Imaibius) reuteri Kleinfeld, 2008 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Imaibius) pachtoun kabakovianus Gottwald, 1992 Holotype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Imaibius) nouristani nouristani (Ledoux, 1977)
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Imaibius) nouristani bandaensis Kleinfeld & Reuter, 2007 Holotype
Male adult
Afghanistan, prov. Konar, NE Jalalabad, Schamschehr - Banda-Wald (ca. 34.44N/70.50E), 2800 m
23 mm
Copyright © Frank Kleinfeld
Carabus (Imaibius) dardiellus polymorphus Heinz, 1993 Paratype
Female adult
Museum of Institute of Systematic Biology (Daugavpils, Latvia)
Copyright © Alexander Anichtchenko
Carabus (Imaibius) caschmirensis caschmirensis Redtenbacher, 1844
Female adult
length: 32 mm<br /> Pakistan, Mansehra<br /> leg. Heinz & Staven, 2001; det. K.Staven, 2001<br /> Foto: U.Schmidt, 2008
Copyright © U.Schmidt
Carabus (Imaibius) barysomus hazarensis Deuve, 1982
Male adult
length: 35 mm<br /> Pakistan, Mansehra<br /> leg.det. K.Staven, 2001<br /> Topotypus<br /> Foto: U.Schmidt, 2008
Copyright © U.Schmidt
Carabus (Imaibius) baronii sharanicus Deuve & Letellier, 1980 Paratype
Male adult
Copyright © Alexander Anichtchenko
Carabus (Imaibius) baronii sharanicus Deuve & Letellier, 1980
Male adult
Copyright © Alexander Anichtchenko
Carabus (Imaibius) baronii baronii Heinertz, 1977
Male adult
length: 23 mm
Pakistan, Mansehra
leg.det. K.Staven, 1997
Topotypus
Foto: U.Schmidt, 2008
Copyright © U.Schmidt
Carabus (Imaibius) baronii battalensis Heinz & Staven, 1998 Paratype
Male adult
Copyright © Alexander Anichtchenko
Carabus (Imaibius) dardiellus perobscurus Deuve, 1980
Male adult
N India.Kashmir, Loran, W part Pir Panchal range,h=2400-2500m, 33°49'"N, 74°20'E
L=25 mm
Copyright © Ruslan Panin
Carabus (Imaibius) dardiellus ? subpunctulus Roeschke, 1907
Male adult
N India, Kashmir, Pahalgam, Aru Valley h=2400m, VII.2007
L=25 mm
Copyright © Ruslan Panin
Carabus (Imaibius) wittmerorum Heinertz, 1978
Male adult
N. Pakistan: Frontier prov., Miandam, Swat vall., h=1800-2600 m, leg.S.Remond (TOPOTYPUS)
Copyright © Igor Solodovnikov
Carabus (Imaibius) barysomus hazarensis Deuve, 1982 (TOPOTYP)
Male adult
N-Pakistan: Mansehra prov., Hazara, Kaghan valley: Shocgran env., h = 2750 m L= 42 mm
Copyright © Igor Solodovnikov
Carabus (Imaibius) wittmerorum Heinertz, 1978
Male adult
N. Pakistan: Frontier prov., Miandam, Swat vall., h=1800-2600 m, leg. S.Remond (TOPOTYPUS)
Copyright © Igor Solodovnikov
Carabus (Imaibius) boysi boysi Tatum, 1851
Male adult
NW-India, Himachal Pradesh, Manikaran env., Pulga vill., 4.VII.2008. L=26 mm
Copyright © Ruslan Panin
Carabus (Imaibius) dardiellus var.granulisparsus Bates, 1889 Lectotype
Female adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC100 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec100
Copyright © - - -
Carabus (Imaibius) stoliczkanus brauni Heinz, 1993
Female adult
NE Pakistan, Azad Kashmir, Bagh Distr., Sudhan-Gali h~2300 m,15.VIII.1994, S Dacatra leg. L=37 mm
Copyright © Ruslan Panin
Carabus (Imaibius) leepai sabatinellii Deuve & Kaláb, 2017 Holotype
Male adult
Pakistan, Azad Kashmir, District de Muzaffarabad, Pir Chanasi (G. Sabatinelli, 15 août 2012), in coll. J. Kaláb, à Jinačovice (République Tchèque).
Copyright © Thierry Deuve
Carabus (Imaibius) rostianus Semenov, 1906
Female adult
Natural History Museum (2014). Dataset: Collection specimens. Resource: Specimens. Natural History Museum Data Portal (data.nhm.ac.uk). © The Trustees of the Natural History Museum, London
Copyright © - - -
Carabus (Imaibius) gridellii naranensis Heinertz, 1978
Female adult
Natural History Museum (2014). Dataset: Collection specimens. Resource: Specimens. Natural History Museum Data Portal (data.nhm.ac.uk). © The Trustees of the Natural History Museum, London
Copyright © - - -
Carabus (Imaibius) epipleuralis Semenov, 1906
Male adult
Natural History Museum (2014). Dataset: Collection specimens. Resource: Specimens. Natural History Museum Data Portal (data.nhm.ac.uk). © The Trustees of the Natural History Museum, London
Copyright © - - -
Carabus (Imaibius) gridellii babussarensis Deuve & Kalab, 2000
Female adult
Pakistan, Hazara, Babursar pass., 4100m, 10.Viii.1982, W.G. Tremewan. Natural History Museum (2014). Dataset: Collection specimens. Resource: Specimens. Natural History Museum Data Portal (data.nhm.ac.uk). © The Trustees of the Natural History Museum, London
Copyright © - - -
Carabus (Imaibius) wittmerorum vojtechi Schneider & Rapuzzi, 2021 Holotype Paratype
Male+Female adult
Pakistan, Province Khyber, District Swabi, Bir Gali env., 1900 m, 03.VI.2020, Ikram Ullah legit.
Copyright © - - -
Carabus (Imaibius) barysomus heroicus Semenov, 1906
Female adult
Sudhangali (Muzaffarba) CASHMIRE, VIII-2008
Copyright © Alain Mollard
Carabus (Imaibius) barysomus huegeli Roeschke, 1907
Female adult
Daksum (Hindoustan) CASHMIRE, VIII-1980
Copyright © Alain Mollard
Carabus (Imaibius) cavifrons Mandl, 1974
Female adult
Bumra Uhel (Kuz-Heol) NEPAL, VI-1980
Copyright © Alain Mollard
Carabus (Imaibius) dardiellus polymorphus Heinz, 1993 Holotype
Male adult
Reshian, 2100-2500 m, W. Heinz leg. Image from Deuve & Schmidt, 2023
Copyright © Thierry Deuve
Carabus (Imaibius) dardiellus walterheinzi Deuve & Schmidt, 2023 Holotype
Male adult
Pakistan, Azad Jammu & Kashmir, District de Muzaffarabad, environs de Reshian, 2100-2500 mètres (W. Heinz leg.), in coll. Muséum de Stuttgart, Allemagne
Copyright © Thierry Deuve
Carabus (Imaibius) caschmirensis fortetuberculatus Heinz, 1983 Paratype
Male adult
Copyright © Alexander Anichtchenko
Carabus (Imaibius) boysi boysi Tatum, 1851
Female adult
N India, Uttarakhand, Ghangaria, N30.42.245, E079.35.634, 3000m, 11-13.6.2011, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko
Carabus (Imaibius) caschmirensis fortetuberculatus Heinz, 1983 Paratype
Female adult
Copyright © Alexander Anichtchenko
Carabus (Imaibius) nouristani nouristani (Ledoux, 1977)
Female adult
NE-Afghanistan: Nuristan: Darah-I-Nur valley, 2900m 34°45'N 70°40'E, VIII.2012, C.Reuter leg., L=26 mm
Copyright © Ruslan Panin
Carabus (Imaibius) nouristani nouristani (Ledoux, 1977)
Male adult
NE-Afghanistan: Nuristan: Darah-I-Nur valley, 2900m 34°45'N 70°40'E, VIII.2012, C.Reuter leg., L=26 mm
Copyright © Ruslan Panin -
Iniopachus taxon page (7 photos)
Carabus (Iniopachus) pyrenaeus jeanneli Deuve, 1994
Male adult
1-2: Pirineos, Canfranc, Ibon de Iserias 2200 m 16-17.7.05<br /> 3: Pirineos, Monte Perdido, Collata Anisolo 2400m 25.8.05
Copyright © Alexander Anichtchenko
Carabus (Iniopachus) auriculatus ubinensis (Puissegur, 1955)
Female adult
Spain, Montes Cantabricos, Peña Ubiña, 2400m
5-6.VII.2009 Anichtchenko A. leg.
female, 19 mm
Copyright © Alexander Anichtchenko
Carabus (Iniopachus) auriculatus auriculatus Putzeys, 1872
Female adult
Spain, Picos de Europa, Macizo Central, Las Colladinas,
ref. Hermoso 2100m, 8.8.2003
Copyright © Alexander Anichtchenko
Carabus (Iniopachus) pyrenaeus pyrenaeus (as cephalicus Csiki, 1927)
Male adult
SW France, Pyrenees-Atlantiques, Mt.Cirque d'Esturou h=1800m, 29.VIII.2014, Pascal Stefani leg. L=20 mm
Copyright © Ruslan Panin
Carabus (Iniopachus) pyrenaeus cerdanus Lapouge, 1924
Male+Female adult
Cirque des Peyssons, ANDORRE, VI-1985
Copyright © Alain Mollard
Carabus (Iniopachus) pyrenaeus costulus Gehin, 1885
Male+Female adult
Mt. Canigou, 2400m (Pyrénées Orientales, Pyrénées) FRANCE, VI-1985
Copyright © Alain Mollard
Carabus (Iniopachus) pyrenaeus vallierensis Colas, 1965
Male+Female adult
Mt.Vallier (Ariege,Pyrénées) FRANCE, VI-1998
Copyright © Alain Mollard -
Ischnocarabus taxon page (4 photos)
Carabus (Ischnocarabus) tenuitarsis tenuitarsis Kraatz, 1877
Male adult
male, 24mm
Copyright © Boleslav Brezina
Carabus (Ischnocarabus) tenuitarsis tenuitarsis Kraatz, 1877
Female adult
female, 26mm
Copyright © Boleslav Brezina
Carabus (Ischnocarabus) cychropalpus Peyron, 1858
Female adult
female, 28mm
Copyright © Boleslav Brezina -
Isiocarabus taxon page (24 photos)
Carabus (Isiocarabus) strandiellus dayongensis Kleinfeld, 1996 HT Holotype
Male adult
Carabus (Isiocarabus) strandiellus dayongensis HT
Copyright © Frank Kleinfeld
Carabus (Isiocarabus) dargei dargei Deuve, 1987 (as kulingensis Holotype) Holotype
Male adult
Carabus (Isiocarabus) dargei dargei (as kulingensis Holotype)
Copyright © Frank Kleinfeld
Carabus (Isiocarabus) dargei yulingensis (as dreuxi yulingensis) Kleinfeld, 2002 HT PT Holotype Paratype
Male+Female adult
Carabus (Isiocarabus) dargei yulingensis (as dreuxi yulingensis) HT PT
Copyright © Frank Kleinfeld
Carabus (Isiocarabus) fiduciarius fiduciarius (as fiduciarius tim) Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Isiocarabus) hunanicola jinxiuensis Imura, 1995 (as dreuxi xinyingensis) HT PT Holotype Paratype
Male+Female adult
Carabus (Isiocarabus) hunanicola jinxiuensis (as dreuxi xinyingensis) HT PT
Copyright © Frank Kleinfeld
Carabus (Isiocarabus) hunanicola liangshanicus (as dreuxi liangshanicus) HT PT Kleinfeld, 1998 Holotype Paratype
Male+Female adult
Carabus (Isiocarabus) hunanicola liangshanicus (as dreuxi liangshanicus) HT PT
Copyright © Frank Kleinfeld
Carabus (Isiocarabus) kiukiangensis orphniopterus Hauser, 1920
Male adult
Carabus (Isiocarabus) kuikiangensis orphniopterus Hauser, 1920
China, Fujian, Yun Ling Shan,
Fokien, 4.6.1998
Copyright © Alexander Anichtchenko
Carabus (Isiocarabus) kiukiangensis alzonai Deuve, 1990 Holotype
Male adult
Copyright ©
Carabus (Isiocarabus) kiukiangensis alzonai Deuve, 1990 (aedeagus) Holotype
Male adult
Copyright ©
Carabus (Isiocarabus) kiukiangensis alzonai Deuve, 1990 (Label) Holotype
Male adult
Copyright ©
Carabus (Isiocarabus) dargei dargei Deuve, 1987 (as anhuinus xingziensis Kleinfeld 1996) Holotype
Male adult
China, Jiangxi, Lu Shan mts (ca. 29°26'N/115°57'E). 24 mm
Copyright © Frank Kleinfeld
Carabus (Isiocarabus) hienfoungi hienfoungi Thomson, 1857 Lectotype
Male adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC164 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec164
Copyright © - - -
Carabus (Isiocarabus) miaorum Lassalle & Prunier, 1993
Male adult
S-China, N-Guangxi, Mao'er Shan, h~1600 m, V.2009, L= 26 mm
Copyright © Ruslan Panin
Carabus (Isiocarabus) fiduciarius fiduciarius Thomson, 1856
Female adult
CE-China, Hubei, Xiaogan City, Shuangfeng Mountain, 26.VI.2016, L=30 mm
Copyright © Ruslan Panin
Carabus (Isiocarabus) kiukiangensis kiukiangensis Bates, 1888 Lectotype
Male adult
Jiujiang (MNHN) http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec88
Copyright © MNHN
Carabus (Isiocarabus) fiduciarius fiduciarius Thomson, 1856
Male adult
27.5 mm. China, Sichuan, Wanyuan, Mt. Hua-e-shan. Image from Rapuzzi, 2015
Copyright © - - -
Carabus (Isiocarabus) fiduciarius kirinicus Csiki, 1927
Male+Female adult
Male: 34 mm, R. of Korea, Kyonggi-Do, Nam-yang-Ju-shi. Female: 35 mm, R. of Korea, Kyonggi-Do, East from Seoul, Yangpyeong (CIR). Image from Rapuzzi, 2015
Copyright © - - -
Carabus (Isiocarabus) fiduciarius saishutoicus Csiki, 1927
Male+Female adult
Male: 26.5 mm, R. of Korea, Cheju Island, Mt. Hallasan. Female: 31 mm, R. of Korea, Cheju Island, Mt. Hangla (CIR). Image from Rapuzzi, 2015
Copyright © - - -
Carabus (Isiocarabus) kiukiangensis Bates, 1888
Male adult
Gutianshan National Park (Gutianshan NP), formerly Gutianshan National Nature Reserve, is located in the western part of Zhejiang Province in southeast China (29°14'N, 118°07'E). Image from Zumstein et al, 2021
Copyright © - - -
Carabus (Isiocarabus) miaorum Lassalle & Prunier, 1993
Male adult
China, NW-Guangxi, Mao'er Shan, h~1600 m 25°52'N 110°29'E, 28.V-9.VI.2009, C.Reuter leg.
Copyright © Ruslan Panin
Carabus (Isiocarabus) miaorum baodingensis Häckel & Sehnal, 2009 Holotype Paratype
Male+Female adult
S China, NE Guangxi, Quanzhou Co. ZhenBaoDing, 26°8.113´N, 110°49.36´E, 2100 m, VI-2009 (in collection of M. Häckel)
Copyright © - - - -
Lamprostus taxon page (31 photos)
Carabus (Lamprostus) calleyi pseudoprasinus Lapouge, 1912
Male+Female adult
Transcaucasia, Armenia,
Ashotsk distr. Tsoghamarg vill, .21-22.V.1997, P.V.Kazaryan leg
female, 24 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) calleyi nigrinus Motschulsky, 1865
Male adult
Azerbaijan,
Talysh Mts., Palikesh, Zuvand, 08.VI.2001, Dolin V.G. leg.
male, 24mm
Copyright © Ruslan Panin
Carabus (Lamprostus) torosus spinolae Cristoforis & Jan, 1837
Male adult
length: 29 mm
Türkei, Cankiri, Cerkes, Kurtcimeni
leg.det. K.Staven, 1989
Foto: U.Schmidt, 2009
Copyright © U.Schmidt
Carabus (Lamprostus) robustus ducalis Schaufuss, 1882
Male adult
NE-Turkey:Trabzon, Zigana Pass h=2000m,14-24.VI.2010, V. Slovák leg. <br /> L=32 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) torosus torosus Frivaldsky, 1835
Male adult
NE-Bulgaria, Dobrich Region, Kavarna env.
L=31 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) saulcyi saulcyi Piochard, 1875 (as chambouqensis)
Female adult
N-Lebanon, Fnaпdek, Djebel Qammoua, h=1100-1200m, 29.X.2012, C. Reuter leg.
L=27 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) saulcyi saulcyi Piochard, 1875
Male adult
C Lebanon, Jbeil Distr., Ehmej vill., h=1100 m.V.2012, C. Reuter leg.
L=24 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) syrus pseudopinguis Heinz & Staven, 2000
Male adult
NW-Jordan, Ajlun env., oak forests on dry limestone, h=897m, 32°21′33.1″N/ 035°43’47.4″E, 4.V.2010, Schnitter/Schellhorn leg.<br /> L=26 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) torosus sinopensis Breuning & Ruspoli, 1970
Male adult
NC-Turkey, Sinop province, Erfelek env., h=200m, 10-27.VII.2014, C. Reuter leg. L=31 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) hemprichi elonensis Schweiger, 1970
Male adult
N-Israel, Midlle Galilea, Nazareth env., Hoshaya viil., 3-12..III.2015, B.Vas’ko leg./det. Panin R.
Copyright © Ruslan Panin
Carabus (Lamprostus) syrus cheikensis Deuve, 1992
Male adult
N-Israel, Midlle Galilea, Nazareth env., Hoshaya viil., 13.XII.2012, Vas’ko, leg./det. Panin R. L=30мм
Copyright © Ruslan Panin
Carabus (Lamprostus) erenleriensis erenleriensis Schweiger, 1966, loc.typ.
Male adult
NW-Turkey: Bolu Prov., Abant Lake, 25.VIII.1990 L=35 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) torosus giresuni Breuning, 1964 (as paululus Heinz & Korge, 1964)
Male adult
NE-Turkey, Giresun, Sehitler Geçidi (Pass) h~2000 m,24.VII.2001, S. Dacatra leg., L=26 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) robustus triumphans Cavazzuti, 1987
Male adult
NE-Turkey:Trabzon, Tonya env., VIII.2004. L=41 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) torosus nordmanni Chaudoir, 1848 (as enricoi )
Male adult
NE-Turkey:Ordu prov.,3 km N Gölköy h~1100 m, VII.2006, L=37 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) calleyi pseudoprasinus Lapouge, 1912
Male adult
S Georgia, Tsalka (Paravani) 41°29′18.7″N, 43°51′12.3″E,
Copyright © Ruslan Panin
Carabus (Lamprostus) calleyi pseudoprasinus Lapouge, 1912
Male adult
S Georgia, Tsalka (Paravani) 41°29′18.7″N, 43°51′12.3″E, 8-9.V.2016, Pavel Turek leg. / det. I. Solodovnikov, L=23 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) hemprichi hemprichi Dejean, 1826
Male adult
Lebanon. det. Mingyi Tian
Copyright © Mingyi Tian
Carabus (Lamprostus) saulcyi saulcyi Piochard, 1875 (as chambouqensis)
Male adult
Lebanon
Copyright © Mingyi Tian
Carabus (Lamprostus) punctatus Castelnau, 1835
Female adult
Anatolia mer., Umg. Yayladagi (Antakya), 600-900m, 13.4.1981, Heinz leg. (cAA)
Copyright © Alexander Anichtchenko
Carabus (Lamprostus) syrus moukarzelae Deuve, 2002
Female adult
S-Lebanon: Marjaayoun, 18.XI.-11.XII.2017, C. Reuter leg., L=35 mm
Copyright © Ruslan Panin
Carabus (Lamprostus) hemprichi sidonius Lapouge, 1907
Male adult
Israel, Menashe Forest, b. Megiddo, 200-300mm 7.04.1987 Heinz leg (cAA). 27 mm
Copyright © Alexander Anichtchenko
Carabus (Lamprostus) syrus cheikensis Deuve, 1992
Female adult
SE Lebanon, Rashaya Distr., Tannoura, oak forest, h=700 m, 11-19.IV.2016, C. Reuter leg., L= 32 mm
Copyright © Ruslan Panin -
Leptocarabus taxon page (46 photos)
Carabus (Leptocarabus) yokoae marcilhaci Deuve, 1992
Male adult
CHINA, Shanxi, Tschingling Mts., Taibeichan Nat. reserv.;
Copyright © Anton Menstschikov
Carabus (Leptocarabus) yokoae marcilhaci Deuve, 1992
Male adult
China, Shanxi, Taschingling Mts., Taibeishan Nat. res.
2000 m., 5-15.08.2005
Copyright © Alexander Anichtchenko
Carabus (Leptocarabus) semiopacus Reitter, 1895
Female adult
N Korea, Nangnim, 14.VII.1981
female, long. 24 mm
Copyright © Alexander Anichtchenko
Carabus (Leptocarabus) procerulus Chaudoir, 1862
Male adult
Japan, Yamanashi pref., Otawa-rindo,
Narusawa vill., 9.IX.2002 T. Kurihara leg
male, 31 mm
Copyright © Alexander Anichtchenko
Carabus (Leptocarabus) harmandi fujisan Ishikawa, 1972
Female adult
Japan, Yamanashi pref., Fuji-Rinds 1600m, Narusawa vill. 3.V.2005 Kurihara K. leg. 25 mm
Copyright © Alexander Anichtchenko
Carabus (Leptocarabus) arboreus exarboreus (Ishikawa, 1992)
Male+Female adult
E.Russia, Far East, Sakhalin Isl., forest of M.Tym river, 11-IX-2000 . Egorov leg. male 27mm, female 30mm.
Copyright © Safronov
Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931
Male adult
NE-China:Liaoning prov., Fengcheng City, Miaoyang, 12-17.VI.2011<br /> L=26 mm
Copyright © Ruslan Panin
Carabus (Leptocarabus) hiurai Kamiyoshi & Mizoguchi, 1960
Male adult
Collection ground Hojyo city, Ehime Pref. 34358
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Carabus (Leptocarabus) kumagai kumagai Kimura & Komiya, 1974
Male adult
Collection ground Otsu city, Shiga Pref. 35330
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Carabus (Leptocarabus) procerulus Chaudoir, 1862
Male adult
Japan:S Honshu, Matsubara-ko, VIII.1997, S. Remond leg.
L=30 mm
Copyright © Ruslan Panin
Carabus (Leptocarabus) procerulus Chaudoir, 1862
Male adult
Japan, S Honshu, Mie Pref., Hakusan, VII.2001, S. Remond leg.<br /> L=33 mm
Copyright © Ruslan Panin
Carabus (Leptocarabus) arboreus arboreus Lewis, 1882
Male adult
N-Japan, SW-Hokkaido, Chitose sity, 25.V-8.VI.2013, T. Hirabayashi leg.<br /> 32 mm
Copyright © Ruslan Panin
Carabus (Leptocarabus) arboreus pararboreus (Ishikawa, 1992)
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Carabus (Leptocarabus) arboreus pararboreus (Ishikawa, 1992)
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Carabus (Leptocarabus) arboreus fujisanus Bates, 1883
Male adult
Japan, CE-Honshu, Yamanashi pref., Mt. Fujisan h~1600m, 23.IX.1982, Y. Imura leg. 23 mm
Copyright © Ruslan Panin
Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931
Male adult
NE-China:E-Liaoning: NP Kuandian Tianqiaogou , VI.2010, L=25 mm, det. Panin R.
Copyright © Ruslan Panin
Carabus (Leptocarabus) arboreus fujisanus Bates, 1883
Female adult
Japan, Honshu, Mt. Fuji h~1900m, Shizuoka pref., Fujinomiya city, 23.VI.2007, M. Matsunaga leg., L=25 mm
Copyright © Ruslan Panin
Carabus (Leptocarabus) harmandi Lapouge, 1909 Lectotype
Male adult
Honshu, Tochigi, Mts near Nikko. Muséum national d’Histoire naturelle, Paris (France) Collection: Insects - Coleoptera (EC) Specimen MNHN-EC-EC549
Copyright © MNHN
Carabus (Leptocarabus) vogtianus cf horvatovichi Deuve, 1992
Male adult
R. of Korea, Geonggido/Gangwondo provincial border, Pass North from Dopyeong, 600 m, 3/13.VII.2012, I. Rapuzzi and L. Caldon leg. Image from Rapuzzi, 2015
Copyright © - - -
Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931 Neotype
Male adult
Chongjin Muséum national d’Histoire naturelle, Paris (France). Collection: Insects - Coleoptera (EC). Specimen MNHN-EC-EC518
Copyright © MNHN
Carabus (Leptocarabus) kyushuensis cerberus Ishikawa, 1962
Female adult
Fuchu (Mitsugi,Hiroshima) JAPON, VII-2001. 34 mm
Copyright © Alain Mollard
Carabus (Leptocarabus) kyushuensis nakatomii Ishikawa, 1966
Male adult
Tombara (Honshu,Shimane) JAPON, VII-2001. 36 mm
Copyright © Alain Mollard
Carabus (Leptocarabus) kyushuensis kyushuensis Nakane, 1960
Female adult
Izumi, (Honshu, Kumamoto) JAPON, VII-2001. 31 mm
Copyright © Alain Mollard
Carabus (Leptocarabus) arboreus akitanus (Ishikawa, 1992)
Male adult
Muriyoshi (Akita), JAPON, VIII-2001; 26 mm
Copyright © Alain Mollard
Carabus (Leptocarabus) arboreus babai Ishikawa, 1968
Male adult
Hiychiyama (Niigata, Honshu), JAPON, XI-1984; 20 mm
Copyright © Alain Mollard
Carabus (Leptocarabus) arboreus exilis Bates, 1883
Male adult
Sado Isle ( Honshu), JAPON, X-1994; 27 mm
Copyright © Alain Mollard
Carabus (Leptocarabus) arboreus gracillimus Bates, 1883
Female adult
Mt.Tate-yama, 2000m (Toyama, Honshu), JAPON, X-1994; 20 mm
Copyright © Alain Mollard
Carabus (Leptocarabus) arboreus karibanus (Ishikawa, 1992)
Female adult
Kuromatsunai (Hokkaido), JAPON, X-1994; 30 mm
Copyright © Alain Mollard
Carabus (Leptocarabus) arboreus kitakamisanus (Ishikawa, 1992)
Male adult
Niisato (Iwate, Honshu), JAPON, VIII-2001; 30 mm
Copyright © Alain Mollard
Carabus (Leptocarabus) arboreus nepta (Ishikawa, 1984)
Male adult
North, Hakkoda-san (Aomori, Honshu), JAPON, VI-1986 ; 25 mm
Copyright © Alain Mollard
Carabus (Leptocarabus) arboreus parexilis Nakane, 1961
Male adult
Towada-Cho (Honshu), JAPON, X-1990; 30 mm
Copyright © Alain Mollard
Carabus (Leptocarabus) arboreus ogurai Ishikawa, 1969
Female adult
Tabayama (Yamanashi, Honshu), JAPON, X-1990; 24 mm
Copyright © Alain Mollard
Carabus (Leptocarabus) arboreus pronepta (Ishikawa, 1992)
Male adult
Hakkode-san (Aomori, Honshu), JAPON, XI-1990; 24 mm
Copyright © Alain Mollard
Carabus (Leptocarabus) arboreus shinanensis Born, 1922
Male adult
Mt. Yatsugatake (Nagano, Honshu), JAPON, VII-1985; 23 mm
Copyright © Alain Mollard
Carabus (Leptocarabus) arboreus tenuiformis Bates, 1883
Female adult
Mt. Osaki-Yama,1900m (Tochigi, Honshu), JAPON, VI-1988; 26 mm
Copyright © Alain Mollard
Carabus (Leptocarabus) yokoae xunyangbanus Imura, 1998
Male adult
Qinling Shan, (Xi’an, Shanxi) CHINE VI-1992; 27 mm
Copyright © Alain Mollard
Carabus (Leptocarabus) yokoae nanjiangensis Imura & Su, 2000
Female adult
Gangwu Shan (Taoyuan, Sichuan) CHINE VI-1992; 27 mm
Copyright © Alain Mollard
Carabus (Leptocarabus) seishinensis seunglaki (Kwon & Lee, 1984)
Male adult
Taebaeck-san (Kangwon-sud) KOREE, VII-1998; 21 mm
Copyright © Alain Mollard
Carabus (Leptocarabus) vogtianus horvatovichi Deuve, 1992 Holotype
Male adult
Image from: Korean type specimens of insects deposited in Hungarian Natural History Museum.
Copyright © - - -
Carabus (Leptocarabus) semiopacus sstr (left); semiopacus gwangyangensis (right)
Male adult
Image from T. Kwon et al (2023)
Copyright © - - -
Carabus (Leptocarabus) semiopacus Reitter, 1895
Male adult
1A and 3B: A. (A.) semiopacus semiopacus; 2A and 4B: A. (A.) semiopacus gwangyangensis Kwon & Park, 2023 (A: right lateral view; B: left and right basal lobes), (Scale bars: 1 mm). Image from T. Kwon et al, 2023
Copyright © - - -
Carabus (Leptocarabus) koreanus Reitter, 1895
Male adult
Habitus of A. koreanus ssp. and male aedeagus tips: A. koreanus koreanus; B: A. koreanus minor; C: A. koreanus mineti; D: A. koreanus rectokoreanus (Scale bars: 1 cm). Image from T. Kwon et al 2023
Copyright © - - - -
Leptoplesius taxon page (6 photos)
Carabus (Leptoplesius) merzbacheri kirgisiensis Mandl, 1967
Female adult
22mm
Copyright © Boleslav Brezina
Carabus (Leptoplesius) merzbacheri kirgisiensis Mandl, 1967
Female adult
female, 27mm
Copyright © Boleslav Brezina
Carabus (Leptoplesius) merzbacheri kirgisiensis Mandl, 1967
Female adult
female, 24mm
Copyright © Boleslav Brezina
Carabus (Leptoplesius) shokalskii kuqaensis Deuve, 1994 Paratype
Female adult
female, 24mm
Copyright © Boleslav Brezina
Carabus (Leptoplesius) schotti Obydov, 1996 Holotype
Female adult
Киргизия, хр. Терскей, дол. р. Тургей
Copyright © Dmitri Obydov
Carabus (Leptoplesius) itshkibashi Kabak, 2004
Female adult
China, Xinjiang, S slope of Khalyktau
Copyright © Ilya Kabak -
Limnocarabus taxon page (15 photos)
Carabus (Limnocarabus) clathratus clathratus Linne, 1761
Female adult
W-Ukraine, Maloe Polesie, Lviv Region, near Kamenka-Bugzkaya, 28.III.2008, Panin R. leg. L=31 mm
Copyright © Ruslan Panin
Carabus (Limnocarabus) clathratus clathratus Linne, 1761
Male adult
length: 20-36 mm, Germania, Mecklenburg, Wismar, leg. F.Wolf, 1998; det. A.Skale, Photo: U.Schmidt, 2009
Copyright © U.Schmidt
Carabus (Limnocarabus) clathratus clathratus (as clathratus foveolatoseriatus)
Female adult
The Chelyabinsk area, Kizilskyi township, Cheka mountain (steppe). 14-15/07/2008
Copyright © Sunvas
Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
Male adult
Transcaucasia, Armenia,
Amasia distr., Arpalach vill., VI-VII.1999, P.V.Kazaryan leg.
male, 25mm
Copyright © Ruslan Panin
Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
Female adult
SW Slovakia, Bratislava env., IV.2004, Krause Rene leg.
L=32 mm
Copyright © Ruslan Panin
Carabus (Limnocarabus) clathratus maacki (as maacki aquatilis)
Male adult
Collection ground Jusanko, Aomori Pref. 35305
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Carabus (Limnocarabus) clathratus clathratus (as jansoni Kraatz, 1890)
Male adult
W-Ireland, County Mayo, Bellacorick bog, 9.VII.2013, J.Stypula leg. L=27 mm
Copyright © Ruslan Panin
Carabus (Limnocarabus) clathratus auraniensis Muller, 1902 (as antonellii Luigioni, 1921)
Male adult
Gaggiano Milan, Italy
Copyright © Vittorino Monzini
Carabus (Limnocarabus) clathratus maacki Morawitz, 1862
Female adult
Приморский край, Романовка, VI.1986
Copyright © Alexander Anichtchenko
Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
Male adult
W Ukraine, Transcarpathian Reg., 4 km NE Chop, forest Peresh, Panin R. leg. L=30 mm
Copyright © Ruslan Panin
Carabus (Limnocarabus) clathratus arelatensis Lapouge, 1903
Female adult
Aigues-Tortes, Camargue, France VI-1984
Copyright © Alain Mollard
Carabus (Limnocarabus) clathratus stygius Ganglbauer, 1890
Male adult
Lenkoran (Narimanabad) AZERBAIDJAN III-1970
Copyright © Alain Mollard -
Lipaster taxon page (18 photos)
Carabus (Lipaster) stjernvalli stjernvalli Mannerheim, 1830
Male+Female adult
Armenia,
Ashotsk, 22.VI.1990, P.V.Kazaryan leg.
female 34mm
Copyright © Ruslan Panin
Carabus (Lipaster) stjernvalli (as tabackuriensis Novotny & Vorisek, 1988)
Male adult
Copyright © Alexander Anichtchenko
Carabus (Lipaster) stjernvalli osellai Breuning & Ruspoli, 1970
Male+Female adult
NE-Turkey: Artvin:Hopa, Borçka env., h=750 m, 9.VI-7.VII.2011, M.Janalik leg.
female, 42 mm
Copyright © Ruslan Panin
Carabus (Lipaster) stjernvalli paravanicus Fominykh et al 2020 Paratype
Male+Female adult
Georgia, Samtskhe-Dzhavakheti, W bank of Paravani Lake near Aspara vill., E slopes of Kyerogly Mt., 2100-2200 m, N 41°28’24.99” / E 43°46’19.65”, 04.-05.06.2015 (D. Fominykh, A. Zubov & A. Safronov). Image from Fominykh et al 2020
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Carabus (Lipaster) stjernvalli bogatshevi Zamotajlov, 1990 Holotype
Male adult
Transcaucasus, mt. Tshareli (=Chareli) near Borzhomi, 2100 m, 30 VII 1929, A. Bogačev leg.” (ZISP). Image from Fominykh et al 2020
Copyright © - - -
Carabus (Lipaster) stjernvalli tabackuriensis Novotný & Vořišek, 1988
Male+Female adult
Georgia, Kvemo-Kartli, Tsalka distr., 4.5 km SW of Kavta (=Chapaevka) vill., NE slopes of Buladag Mt., 1700 m, N 41°32’ / E44°01’, pitfall traps, 20.05.-04.06.2015 (A. Zubov & R. Khryapin). Image from Fominykh et al 2020
Copyright © - - -
Carabus (Lipaster) stjernvalli tabackuriensis Novotný & Vořišek, 1988
Male+Female adult
Georgia, Kvemo-Kartli, Trialeti Mt. Range near Tukhmatash Pass, E slopes of Dalidag Mt., 2070 m, N 41°32’ / E43°52’, pitfall traps, 06.06.–07.2017 (D. Fominykh). Image from Fominykh et al 2020
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Carabus (Lipaster) stjernvalli kartalinicus Fominykh et al 2020 Paratype
Male+Female adult
Georgia, Kvemo-Kartli, Trialeti Mt. Range near Manglisi vill., NW part of Kartli Mt. Range, N slopes of Shikhan Mt., 2070 m, N 41°45’ / E 44°22’, pitfall traps, 07.06.-17.07.2017 (D. Fominykh). Image from Fominykh et al 2020
Copyright © - - -
Carabus (Lipaster) stjernvalli kartalinicus Fominykh et al 2020 Paratype
Male adult
Georgia, Shida-Kartli, Trialeti Mt. Range near Dre vill., NW part of Razmeti Mt. Range, 1920 m, N 41°50’ / E 44°05’, 08.06.2017 (D. Fominykh). Image from Fominykh et al 2020
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Carabus (Lipaster) stjernvalli stjernvalli Mannerheim, 1830
Male+Female adult
1-2: Georgia, Adjaria, Arsiani (=Arsiyan) Mt. Range near Goderdzi Pass, Chanchakhi Mt., 2300 m, N 41°36’35.00” / E42°31’34.15”, 10.06.2015 (D. Fominykh & A. Zubov); 3-4: Georgia, Adjaria (=Adjara, Adzharia), Baki (Bako) vill. env., 08.-26.06.1998 (V. Filippov). Image from Fominykh et al 2020
Copyright © - - -
Carabus (Lipaster) stjernvalli stjernvalli Mannerheim, 1830
Male+Female adult
Georgia, Guria, N slopes of Adzhar-Imereti Mt. Range near Gomi vill. between Bzhuzhi and Kalova rivers, 1440 m, N 41°50’16,60” / E 42°08’29,20”, pitfall traps, 10.04.-07.06.2016 (D. Fominykh). Image from Fominykh et al 2020
Copyright © - - -
Carabus (Lipaster) stjernvalli stjernvalli Mannerheim, 1830
Male+Female adult
Georgia, Samtskhe-Dzhavakheti, Adigeni Distr., Khormani Mt. Range, E slopes of Tail’ Mt., 2290 m, N 41°36’57.00” / E 42°40’32.40”, 22.-24.05.2015 (A. Zubov & R. Khryapin) Image from Fominykh et al 2020
Copyright © - - -
Carabus (Lipaster) stjernvalli gvalijai Retezár, 1992
Male+Female adult
Georgia, Racha, Racha Mt. Range, N slopes of Khikhata Mt. Range near Mravaldzali vill., 2100 m, N 42°29’13,64” / E 43°21’12,19”, 08.-09.06.2016 (A. Zubov). Image from Fominykh et al 2020
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Carabus (Lipaster) stjernvalli gvalijai Retezár, 1992
Male+Female adult
Georgia, Kvemo-Svaneti, S slopes of Svaneti Mt. Range near Lentekhi vill., S slopes of Goldash Mt., 2615 m, N 42°50’00,10” / E 42°45’16,90”, pitfall traps, 18.06.-13.07.2017 (D. Fominykh). Image from Fominykh et al 2020
Copyright © - - -
Carabus (Lipaster) stjernvalli osellai Breuning & Ruspoli, 1970
Male adult
NE-Turkey: Artvin, Hopa / Çifteköprü, VII.1977, L=36 mm
Copyright © Ruslan Panin
Carabus (Lipaster) stjernvalli stjernvalli Mannerheim, 1830
Male adult
SC Georgia, Tsalka distr. 7 km NE of Paravani Lake, 41.540750, 43.879806, h=2100 m, 12-16.VII.2025, I.Uspenskyi leg., L=25 mm
Copyright © Ruslan Panin -
Macrothorax taxon page (26 photos)
Carabus (Macrothorax) morbillosus alternans Palliardi, 1825
Male adult
S Italy, Sicily, Province Palermo, Piana degli Albanesi h~700m, XI.2010, I.Rapuzzi leg.
L=32 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) planatus Chaudoir, 1843
Male adult
Sicilia, Madonie, Piano Battaglia (PA) 1500m
4.6.1992 Aliquo leg.
male, 31 mm
Copyright © Alexander Anichtchenko
Carabus (Macrothorax) morbillosus macilentus Lapouge, 1899
Male adult
Spain, Alicante, Santa Pola, 20.2.1988 M. Godoy leg.
male, 29 mm
Copyright © Alexander Anichtchenko
Carabus (Macrothorax) rugosus boeticus Deyrolle, 1852
Female adult
Spain, Cadiz, Novo Sancti Petri, 10.6.2003
female, 32 mm
Copyright © Alexander Anichtchenko
Carabus (Macrothorax) rugosus rugosus Fabricius, 1792
Male adult
N Africa, 2 km W Ceuta, Allabacal,
17-18.2.2004 Anichtchenko A. leg.
male, 26 mm
Copyright © Alexander Anichtchenko
Carabus (Macrothorax) rugosus laufferi Breuning, 1927
Female adult
Spain, Gredos Mts., 2 km E from La Mira Mt.
4-7.7.2007 Anichtchenko A. leg.
female, 29 mm
Copyright © Alexander Anichtchenko
Carabus (Macrothorax) rugosus celtibericus Germar, 1824
Male adult
Portugal, Vila Fresca de Azeitas,
Santana, 13.12.2001
male, 30 mm
Copyright © Alexander Anichtchenko
Carabus (Macrothorax) aumonti maroccanus Bedel, 1895
Male adult
N Morocco (Spain:Melilla), hipodromo, X.1969, leg. Pardo Alcaide
L=38 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) rugosus boeticus Deyrolle, 1852
Male adult
SC Spain, Jaen reg., Puente de la Sierra, 10.VII.2011, M.Lopez leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) rugosus rugosus Fabricius, 1792
Male adult
N Morocco, El Biutz –Taghramt,20.I.2005, Manfred Egger leg.
L=29 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) morbillosus constantinus Kraatz, 1899
Male adult
NW Tunisia; Tebarka, E Sejenane,26.I.2004, Manfred Egger leg.
L=33 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) morbillosus macilentus Lapouge, 1899
Female adult
SE Spain, Murcia reg., P.N. de Calblanque, 1.V.2003, T.Gazurek leg.
L=34 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) rugosus celtibericus Germar, 1824
Female adult
SW-Portugal, Setúbal Distr., Cercal do Alentejo/ Tanganheira vill. 8.XII.2006, T.Gazurek leg.
L=31 m
Copyright © Ruslan Panin
Carabus (Macrothorax) morbillosus constantinus (as galloprovincialis Lapouge, 1910)
Female adult
SE-France, Var, near Le Muy, VIII.1980, Michon leg.
L=31 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) planatus Chaudoir, 1843
Male adult
S Italy, N Sicily, Nebrodi; 33 mm
Copyright © Aleš Sedláček
Carabus (Macrothorax) morbillosus morbillosus Fabricius, 1792 (as cheminorum Deuve, 1988)
Female adult
SE-France, Var, Fréjus env., 26.II.1997, <br /> L=31 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) morbillosus lampedusae Born, 1925
Female adult
AG, Sicily, Island of Lampedusa, leg V. Monzini
Copyright © Vittorino Monzini
Carabus (Macrothorax) morbillosus lampedusae Born, 1925
Male adult
S Italy, Isl. Lampedusa, Baia dei Conigli, VI.2003, L=29 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) rugosus laufferi Breuning, 1927
Female adult
C-Spain: Avila, El Hornillo env., 27.V.2017, J. Ferrero leg. 30 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) rugosus levantinus Lauffer, 1905
Male adult
SE-Spain, Murcia, Jumillla, Sierra del Carche, 6.VII.2019, F. Garcia leg., L=29 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) planatus Chaudoir, 1843
Female adult
S Italy, N Sicily, Nebrodi Mts, Portella di Fémmina Morta h=1500 m, 2.IX.1989, V.Aliquo leg. L=33 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) rugosus celtibericus Germar, 1824
Male adult
SW-Portugal, Serra do Cercal h=217 m, 37°47'15.6"N 8°42'02.0"W, road N390, 19.III.2011, D. Prunier leg., L=27 mm
Copyright © Ruslan Panin
Carabus (Macrothorax) aumonti Lucas, 1849 (as sculptus Lapouge, 1925)
Male adult
Lalla Maghrnia , ALGERIE, V-1927; 42 mm
Copyright © Alain Mollard -
Meganebrius taxon page (35 photos)
Carabus (Meganebrius) wallichi wittmeri Mandl, 1974
Female adult
Nepal, Lantang, near Dunche, Songta vill.
3500 m 15.5.2000
Copyright © Alexander Anichtchenko
Carabus (Meganebrius) franzi Mandl, 1974
Female adult
Nepal, Lantang, near Dunche, Songta vill,
3500 m, 15.5.2000
Copyright © Alexander Anichtchenko
Carabus (Meganebrius) tamang gurkhaorum Lassalle, 1985
Female adult
CW Nepal, Manaslu massif 2300 m,
Bara Pokhari Lekh, 5.2001 M.Hackel leg.
Copyright © Alexander Anichtchenko
Carabus (Meganebrius) kadoudali (Morvan, 1981)
Male+Female adult
C-Nepal, SW Dhaulagiri, W Jaljala h=3150 m, 28°30'05"N, 83°11'48"E, 23.IX.2012, J. Schmidt leg.
female, 20 mm
Copyright © Ruslan Panin
Carabus (Meganebrius) pseudoharmandi Mandl, 1965
Female adult
E-Nepal, Kosi zone, Khandbari,(1.5km SW of Chichila vill., 27*45’843’’N/ 87*22’473’E, h=1940m), damaged primary forest, 13 .V.2010, B.Brezina & M.Häckel leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Meganebrius) arunensis Heinertz, 1980
Male adult
E-Nepal, Kosi zone, Khandbari, 1.5km SW of Chichila vill., 27*47’175’’N/ 87*23’622’E, h=1880m, damaged primary forest, 11.V.2010, B.Brezina & M.Häckel leg.
L=27 mm
Copyright © Ruslan Panin
Carabus (Meganebrius) quinlani smetanai Häckel, 2011 Paratype
Male+Female adult
C-Nepal:Dhaulagiri Massif:Baglung Distr., (1.5km W of Okhale vill., 28.32’152’’N/ 83.48’673’E, h=2600m) damaged primary forest, 19.V.2010, B.Brezina & M.Häckel leg.
male, 21mm
female, 23 mm
Copyright © Ruslan Panin
Carabus (Meganebrius) deliae colasianus (Morvan & Mandl, 1974) Holotype
Male adult
Copyright © Igor Solodovnikov
Carabus (Meganebrius) deliae colasianus (Morvan & Mandl, 1974)
Male adult
Copyright © - - -
Carabus (Meganebrius) deliae deliae (Morvan, 1972) Holotype
Male adult
Copyright © - - -
Carabus (Meganebrius) quinlani quinlani Mandl, 1965 Holotype
Male adult
Copyright © - - -
Carabus (Meganebrius) lebretae lebretae Colas, 1961 Holotype
Male adult
Copyright © - - -
Carabus (Meganebrius) quinlani quinlani Mandl, 1965
Male adult
C-Nepal, Gandaki, Manang Distr., Timang env., h~2500-2600m, old open forest, grassland, 3-5.V.2007, Fouqué R. leg. L=19 mm
Copyright © Ruslan Panin
Carabus (Meganebrius) tuberculipennis ganjalaensis Deuve, 2000
Male adult
Nepal, Helambu, GanjaLa Pass S-slope, Dhukpu Kharka h=4100m, 28°03'50"N, 85°34'21"E, 12.VIII.2010, S.Tamang leg., L=21 mm
Copyright © Ruslan Panin
Carabus (Meganebrius) kadoudali (Morvan, 1981) Paratype
Female adult
Vallée de Uttar Ganga, Dhorpatan, col de Jhala-Jélé , 3300m, Morvan P. 1977-04-01/1977-04-30 (MNHN) http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec184
Copyright © MNHN
Carabus (Meganebrius) davatchii (Morvan, 1981) Paratype
Male adult
Vallée de Uttar Ganga, Dhorpatan, col de Jhala-Jélé (MNHN) http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec186
Copyright © MNHN
Carabus (Meganebrius) thudamensis Deuve & Schmidt, 2017 Holotype
Male adult
Est-Népal, District de Sankhuwasabha, Thudam, 3500 – 3600 m, 27°45’23’’N – 87°32’56’’E (S. Tamang leg., 1-vii-2011), in coll. J. Schmidt (Admannshagen, Allemagne).
Copyright © Joachim Schmidt
Carabus (Meganebrius) santostamangi thanglaensis Deuve & Schmidt, 2017 Holotype
Male adult
Est-Népal, limite des districts de Sankhuwasabha et de Taplejung, Jaljale Himal, Thangla Phedi, 4 150 m, 27°39’09’’ N – 87°35’56’’ E (S. Tamang leg., 27-vi-2011), in coll. J. Schmidt (Admannshagen, Allemagne).
Copyright © Joachim Schmidt
Carabus (Meganebrius) santostamangi santostamangi Holotype
Male adult
Est-Népal, district de Taplejung, versant oriental de la Palung Khola, 3 720 m, 27° 35’ 43’’ N – 87° 41’ 50’’ E (S. Tamang leg., 7-vii-2011), in coll. J. Schmidt (Admannshagen, Allemagne).
Copyright © Joachim Schmidt
Carabus (Meganebrius) hongliangi Deuve, Liu & Liang, 2024 Holotype
Male adult
Chine, Tibet, Préfecture de Xigazê, canton de Gyirong, ville de Gyirong, village de Cha, col de Latuo La, 3995 m, 28,3908°N 85,4074°E, pelouses alpines (Hongliang Shi leg., 10.VII.2019, sous des pierres), in IOZ. Image from Deuve et al 2024
Copyright © - - -
Carabus (Meganebrius) heterocostatus s.str. Deuve & Schmidt, 2023 Holotype
Male adult
Népal oriental, versant méridional du Makalu Himal, Siga Pani, 2800 m, 27°17,531’N-87°02,053’E, 7.VI.2008, leg. S. Tamang (SCHM).
Copyright © Joachim Schmidt
Carabus (Meganebrius) heterocostatus pseudarunensis Deuve & Schmidt, 2023 Holotype
Male adult
Népal oriental, Kimalung, 2500 m, 4.VI.1989, leg. P. Morvan (MRV).
Copyright © Joachim Schmidt
Carabus (Meganebrius) tuberculipennis bhairavkundensis Deuve & Schmidt, 2023 Holotype
Male adult
Népal, Bhairav Kunda Lekh, Chormu Kharka, 4000-4100 m, 27°58’34’’N/85°52’18’’E, 20-21.V.2011, leg. J. Schmidt (SCHM).
Copyright © Joachim Schmidt
Carabus (Meganebrius) tuberculipennis sprecherae Deuve & Schmidt, 2023 Holotype
Male adult
Népal, Massif d’Helambu, environs de Paanch Pokhari, 4000-4200 m, 28°02’33’’N/85°42’48’’E, 7.V.2010, leg. J. Schmidt (SCHM).
Copyright © Joachim Schmidt -
Megodontoides taxon page (11 photos)
Carabus (Pagocarabus) qiangding Kleinfeld & Puchner, 2007 Holotype Paratype
Male+Female adult
China. Sichuan, Str. zwischen Yajiang und Kangding, Tagong Grasslands (30°02'59,3''N/101°34'14,3''E)
28 mm
Copyright © Frank Kleinfeld
Carabus (Pagocarabus) thibetanus Breuning, 1950 Holotype
Male adult
Tong-ho River, Szetschuan, VII.1947, 4.000m / C. Plesius thibetanus mihi, Typ / HOLOTYPE Coll. Frey, NHM Basel ? 27 mm
Copyright © Frank Kleinfeld
Carabus (Pagocarabus) promachus danbaensis Kleinfeld, 1999
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Pagocarabus) promachus promachus s.str. (as konkerianus)
Female adult
Copyright © Boleslav Brezina
Carabus (Pagocarabus) poschingerianus Korell & Kleinfeld, 1989 Holotype
Male adult
Copyright © Frank Kleinfeld
Carabus (Megodontoides) promachus promachus Holotype
Male adult
Kangding (MNHN) http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec96
Copyright © MNHN
Carabus (Megodontoides) promachus wujiapeng (Imura, Zhou & Su, 2004) Holotype
Male adult
mixed forest, 2,320 m in altitude, Wujiapeng in southwestrn Danba Xian of west-central Siclman, Southwest China, 1 9-2 1 -VII.2001, Xiao-Dong Yu & Hong-Zhang ZHOU leg. Image from (Imura, Zhou & Su, 2004)
Copyright © - - - -
Megodontus taxon page (162 photos)
Carabus (Megodontus) imperialis Fischer, 1823
Female adult
Carabus (Megodontus) imperialis
photo M. Smirnov
from www.zin.ru
Copyright © Maksim Smirnov
Carabus (Megodontus) schoenherri sajanus Breuning, 1927
Male+Female adult
<b>Carabus (Megodontus) schoenherri sajanus Breuning, 1927</b><br /> Russia, SW Siberia, Krasnoyarsk reg., S of Abakan,<br /> Sayanogorsk distr., Cheryomushki vill. env.<br /> 07-VII-1994<br /> M. Danilevsky leg.<br /> male 27mm, female 31 mm.
Copyright © Safronov
Carabus (Megodontus) schoenherri sajanus Breuning, 1927
Male adult
Carabus (Megodontus) schoenherri sajanus Breuning, 1927
W Sayan Mts, Borus mt. rng., near Cheremushki vill.,
29-31.7.2000 Brinev A. leg.
male, 30mm
photo Anichtchenko A.
Copyright © Alexander Anichtchenko
Carabus (Megodontus) violaceus fulgens Charpentier, 1825
Male adult
NC-Spain,
Navarra Reg., Sierra de Andнa, IX.2004, Pascal Stefani leg.
male, 30 mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus fulgens Charpentier, 1825 (as gerundensis Born, 1916)
Male adult
Spain, Gerona, Garriguella, XI.1990 J. Soler leg.
male, 31mm
Copyright © Alexander Anichtchenko
Carabus (Megodontus) violaceus andrzejuscii (as wolffii Dejean, 1826)
Male+Female adult
SW-Romania: Banat, Moldova Noua, Ravenska viil., 21.V.-5.VI.2010, Krause Rene leg. female, 32 mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus andrzejuscii (as mehelyi Ganglbauer, 1896 loc. typ.)
Male+Female adult
N-Romania, Rodna Mts., Pasul Setref - Sacel 10km E, h=810 m, 8-19.VI.2009, Krause Rene leg. female, 31mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus andrzejuscii Dejean, 1826 (as pygmaeus Petri, 1912)
Male adult
W Ukraine, NE Carpathians,<br /> Transcarpathian Reg., Rakhov Distr., Chornogora Mt Range h=2030m, alpine zone, 13-24.VI.2010 Panin R. leg./det.<br /> male, 25 mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus rilvensis Kolbe, 1887
Female adult
Bulgaria, Stara Zagora distr. Stara Planina Mts., Shipka Mt., 16.08.2005, A. Safronov leg.
Copyright © Safronov
Carabus (Megodontus) violaceus picenus A.& J.B.Villa, 1838
Male adult
Italia, Marche Pesaro
leg.det. K.Staven, 1998
Foto: U.Schmidt, 2008
length: 26-30 mm
Copyright © U.Schmidt
Carabus (Megodontus) violaceus mixtus Gehin, 1876
Female adult
France, Eloise 74, pont de Gresin 14.07.1991
female, 29mm
Copyright © Alexander Anichtchenko
Carabus (Megodontus) violaceus fiorii Born, 1901
Male+Female adult
NW-Italy,
Reggio Emilia, Regnano, 5.IX.1996 leg. Cartini A.
female, 30 mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus baeterrensis Lapouge, 1901
Male+Female adult
S France,
Midi-Pyrénées Region, Ariège, Col du Chioula h=1500m, V-VI.2005, Pascal Stefani leg.
photo Panin R.
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus aurolimbatus Dejean, 1829
Female adult
Kazakhstan,
North Kazakhstan Province (Kokchetavski region)1.VII.1986
female, 24mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus aurolimbatus Dejean, 1829
Male adult
C.UKRAINE,
Poltava reg., near Novi Sanzhary, bank of Vorskla River, 1-2.VII.07, D.Kovalchuk leg.
male 25mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus aurichalceus Kraatz, 1879
Male adult
Spain, Pais Vasco, Aranzazu, 6.I.2005
Anichtchenko A. leg.
male, 27 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko
Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823
Male adult
W Ukraine, NW Podolian, Ternopol reg., Kremenets distr., near Chornyi Lis vill., steppe sites, 4-22.V.2009, Panin R. leg. female, 29 mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823
Male adult
W-UKRAINE
NW Podolian, Lviv Reg., Zolochiv Distr., near Osovitsa, steppe sites, Panin R. leg. 1.V-6.VI.07
male, 26mm
Copyright © Safronov
Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823
Male+Female adult
W. Ukraine, W. Podolian, Lviv reg., Nikolaev distr., near Krasiv vill., under the bark of oak tree, 8.II.2009, Panin R. leg. female, 29mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823 (as carbonatus Schaufuss, 1882)
Male adult
W.UKRAINE, Ukrainian Carpathians, Lviv Reg., Skolivski Beskidy Mts., near Skole, h=700m, 20.IX.2009, Panin R. leg. male, 33mm
Copyright © Ruslan Panin
Carabus (Megodontus) vietinghoffi vietinghoffi Adams, 1812
Male adult
Yakutia, near Kular vill., 27.7.2000, 20 mm det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko
Carabus (Megodontus) vietinghoffi vietinghoffi Adams, 1812
Female adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Carabus (Megodontus) vietinghoffi fulgidus Fischer, 1827
Male adult
Russia, Evreiskaya AO., near Obluchie 14.6.1993
Chichvarhin A. leg.
male, 20 mm
photo, det. & coll. Anichtchenko A
Copyright © Alexander Anichtchenko
Carabus (Megodontus) tuvensis Shilenkov, 1996
Male adult
Photo A. Menstschicov
Copyright © Anton Menstschikov
Carabus (Megodontus) tuvensis Shilenkov, 1996
Female adult
Photo A. Mentschikov
Copyright © Anton Menstschikov
Carabus (Megodontus) stroganowi tiedemanni Breuning, 1934
Male adult
IRAN, Gilan, Masuleh, 1550 m., 17.06.2002., O.Montrieull leg.
photo A. Menstschicov
Copyright © Anton Menstschikov
Carabus (Megodontus) stroganowi stroganowi Zoubkoff, 1837
Male adult
Iran,
Mazanderan, Naharkoran, b. Gorgan
500m, 4-11.VII.1975 Heinz leg.
male, 30 mm
photo Anichtchenko A.
Copyright © Alexander Anichtchenko
Carabus (Megodontus) schoenherri schoenherri Fischer, 1822
Male adult
enza Reg., Kuznetsk Distr., env. Tikhmenevo Vill., upper course of Sura River, pine forest VI.2005 I.Lebezhinskaja leg.
L=28.0 mm
Copyright © Kirill Makarov
Carabus (Megodontus) schoenherri schoenherri Fischer, 1822 (as sangilenus Obydov, 1997)
Male adult
Tyva, Akademika Obrucheva mt.rng., 15km N Cherbi vill.,
Chailag ur., 7.7.1999 A. & N. Menshikovy leg.
male, 26mm
photo Anichtchenko A.
Copyright © Alexander Anichtchenko
Carabus (Megodontus) persianus persianus Roeschke, 1896
Male adult
Iran, Gilan, pas. N.Dailiman, 1100m, 12-19.VII.1999 Heinz leg.
male, 32 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko
Carabus (Megodontus) obovatus Fischer, 1827
Female adult
Kemerovo reg., Kuznetskii Mt. rng., Berikul, 1.6.1982
female, 16mm
Copyright © Alexander Anichtchenko
Carabus (Megodontus) obovatus Fischer, 1827
Male adult
Kemerovo reg., Kuznetskii Mt. rng., Berikul, 1.6.1982
male, 17mm
Copyright © Alexander Anichtchenko
Carabus (Megodontus) obovatus Fischer, 1827
Female adult
Kuznetskii Mt., Gluhari vill, A. Menstschicov leg., 07.06.2006
female
Copyright © Anton Menstschikov
Carabus (Megodontus) leachi klimenkoi Obydov, 1999 Holotype
Male adult
Museum of Institute of Systematic Biology (Daugavpils, Latvia)
Copyright © Alexander Anichtchenko
Carabus (Megodontus) leachi panzeri Dejean, 1829 (as klimenkoi Obydov, 1999)
Male adult
Kazakhstan, S Altay mts., Ognevka vill.,<br /> Kalbinskii mt. rng., 16.6.1999 Klimenko A. leg.
Copyright © Alexander Anichtchenko
Carabus (Megodontus) gyllenhali Fischer, 1827
Male adult
Krym Peninsula
male, 29 mm
Copyright © Ruslan Panin
Carabus (Megodontus) gyllenhali Fischer, 1827
Male adult
Krym Peninsula, near Sevastopol, 05.05.2000, A.V. Konovalov leg.
Copyright © Anton Menstschikov
Carabus (Megodontus) exaratus Quensel, 1806
Male adult
Russia, C.Caucasus, N.Ossetia,
valley of River Khazniaush,
h=1400 m, 10.VI-31.VII.1993
A. Zamotajlov leg.
Copyright © Safronov
Carabus (Megodontus) exaratus Quensel, 1806
Male+Female adult
NW Caucasus, Russia,
Krasnodar reg., Krasnodar City, 27.IX-07.X.07, D.D.Fominykh leg.
female,29mm/male, 27mm
Copyright © Ruslan Panin
Carabus (Megodontus) kantaikensis Gehin, 1885
Female adult
Russia, Lake Baikal, vall. of Goloustnaya riv., Nizhnii Kochergat vill.,
27.VII.1994 Anichtchenko A. leg.
female, 18 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko
Carabus (Megodontus) kantaikensis Gehin, 1885
Male adult
E Sayan Mts., Sentsa riv. (Oka riv. confluence), near Shasnur vill.
7-16.7.1998 Anichtchenko A. leg.
male, 18 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko
Carabus (Megodontus) kantaikensis Gehin, 1885
Male adult
Krasnoyarsk Area, Sayanogorsk Distr, env. Cheremushki Vill., right bank of Enisej Riv., h= 500 - 800 m, 23.07.1999 A. Semin leg.<br /> male<br /> L=17,0 mm
Copyright © Maksim Smirnov
Carabus (Megodontus) viridicans Obydov, 1993
Male adult
W Sayan, Cheremushki vill., Borus Mt. rng., h= 500m, 29-30.7.2000, Brinev A. leg.; 16 mm
Copyright © Alexander Anichtchenko
Carabus (Megodontus) viridicans Obydov, 1993
Female adult
Photo A. Menstschicov
Copyright © Anton Menstschikov
Carabus (Megodontus) danilevskii Obydov, 1993
Male adult
Krasnoyarsk Area, Sayanogorsk Distr, env. Cheremushki Vill., right bank of Enisej Riv., h= 500 - 800 m, 23.07.1999 A. Semin leg.
L=17,0 mm
Copyright © Maksim Smirnov
Carabus (Megodontus) croaticus croaticus Dejean, 1826
Male adult
YU- Slovenija, Krim 1000 m, 30.10.1975 Drovenik leg.
Copyright © Alexander Anichtchenko
Carabus (Megodontus) caelatus caelatus Fabricius, 1801
Male adult
Bosnia, Trebevic, Vidikovac-Brus, 1200m
21.11.1985 Mihljevic B. leg.
Copyright © Alexander Anichtchenko
Carabus (Megodontus) bonvouloiri bonvouloiri Chaudoir, 1863
Female adult
Anatolia Orientale, Macka-Zigana VIII.1981
Copyright © Alexander Anichtchenko
Carabus (Megodontus) avinovi Semenov & Znojko, 1932
Male adult
Russia, Sakhalin isl., Susunaiskyi rng.,
Chehov Mt., 400m, 19.6.1990 Berlov E. leg.
female, 22 mm
photo Anichtchenko A.
Copyright © Alexander Anichtchenko
Carabus (Megodontus) violaceus violaceus Linne, 1758
Male adult
S.Bohemia, Hluboka nad Vltavou, h=380 m, 6.VIII.2004, S. Budka leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus germarii Sturm, 1815
Female adult
Croatia, Ivanščica Mts , Ivanec, 6-13.VI.2003, Krause Rene leg.
L=36 mm
Copyright © Ruslan Panin
Carabus (Megodontus) vietinghoffi bowringi Chaudoir, 1863
Male adult
Russia, Far East, Primorskiy Kray, Ussuriysk Distr., Kaymanovka vill , 26.VII-3.VIII.2011, A. Korshunov leg.
L=29
Copyright © Ruslan Panin
Carabus (Megodontus) obovatus Fischer, 1827
Male adult
SW Siberia, Russia, Kemerevo reg., Krapivino distr., Azhendarovo, forest, 26-31.VII.2011, D. Efimov leg.
L=18 mm
Copyright © Ruslan Panin
Carabus (Megodontus) vietinghoffi bowringi (as lazoensis Obydov, 1999)
Male+Female adult
Russia, Far East, Primorsky reg., S. Sichote-Alin Mts.R., Kavalerovo vill. env.,
5-25.VII.1992, S. Kunitzyn leg.
male - 28 mm, female - 30 mm.
Copyright © Alexandr Safronov
Carabus (Megodontus) vietinghoffi bowringi Chaudoir, 1863
Male+Female adult
Russia, Far East, Primorsky reg., Khasansky distr.,
Zanadvorovka vill., Gusevsky pit.
M - 19-21.VI.2007, body lenght 29 mm.
F - 15.VII.2008, body lenght 30 mm.
S. Veriga leg.
Copyright © Alexandr Safronov
Carabus (Megodontus) vietinghoffi bowringi Chaudoir, 1863
Female adult
Tian Tou, Yueshi County, Anhui Prov., CHINA
1-10.VIII.2004 local coll.
female - 10 mm.
Copyright © Alexandr Safronov
Carabus (Megodontus) danilevskii Obydov, 1993
Male+Female adult
Russia, SW Siberia, Krasnoyarsk reg., W. Sayan, near Cheryomushki.
26-29.VII.1996 loc. coll.
male - 18mm, female - 20 mm.
Copyright © Alexandr Safronov
Carabus (Megodontus) kantaikensis Gehin, 1885
Female adult
Russia, SW Siberia, W. Sayan, Krasnoyarsk reg.,
S of Abakan, near Cheryomushki, Dzhoysky Pass.
07-VII-1994 M. Danilevsky leg.
body lenght 18 mm.
Copyright © Alexandr Safronov
Carabus (Megodontus) obovatus Fischer, 1827
Male adult
SW Siberia, Kemerovo reg., Krapivino distr.,
8 km SSW Saltymakovo vill., h=150 m.
20-30.VI.2010 D. Efimov, A. Korshunov leg.
body lenght 18 mm.
Copyright © Alexandr Safronov
Carabus (Megodontus) imperialis Fischer, 1823
Male adult
NE Kazakhstan, Zyrjanovsk distr., Putincevo vill. env.
25-VI-2005 M. Danilevsky leg.
male 24 mm, female 26 mm.
Copyright © Safronov
Carabus (Megodontus) violaceus violaceus Linne, 1758 (as intermarinus Blumenthal & Staven, 1985)
Male+Female adult
Poland, Gdansk distr., Gdynya env., Reservat «Cisowa»,
5-9.08.1998 I. Solodovnikov leg.
female - 26 mm.
Copyright © Alexandr Safronov
Carabus (Megodontus) bonvouloiri bonvouloiri (as kaschkarensis Blumenthal & Breuning, 1967)
Male adult
NE-Turke,Rize vill., S Ayder h=1740 m,28.VI-6.VII.2011, M.Janalik leg.
L=32 mm
Copyright © Ruslan Panin
Carabus (Megodontus) croaticus croaticus Dejean, 1826
Male adult
SW Slovenia, Ilirska Bistrica, Sviščaki, Snežnik,17-28.VI.2009, Manfred Egger leg.
L=32 mm
Copyright © Ruslan Panin
Carabus (Megodontus) caelatus schreiberi Kraatz, 1887
Male adult
SW Slovenia, Obrov, Poljane, 14-27.VIII.2010, Manfed Egger leg.
L=33 mm
Copyright © Ruslan Panin
Carabus (Megodontus) planicollis verae Csiki, 1905
Male+Female adult
♂ C-Romania:S Carpathians, Fagaraş Mts, near Avrig Lake, h=2100 m, 5.VI.2005
♀ C-Romania:S Carpathians, Fagaraş Mts, Saratii-Valley h=1700 m, 6.VI.2005
female, 28 мм
Copyright © Ruslan Panin
Carabus (Megodontus) stroganowi transfuga Semenov, 1898
Male adult
SE-Azerbaijan: Talysh, Kalvaz, 21-30.VI., Dolin V.G. leg.
L=32 mm
Copyright © Ruslan Panin
Carabus (Megodontus) kolbei kolbei Roeschke, 1887
Male adult
N-Japan, C-Hokkaido, Hidaka distr., Niikappu env., 27.VIII.1989, Sasaki leg,
L=27 mm
Copyright © Ruslan Panin
Carabus (Megodontus) kolbei hidakamontanus Ishikawa, 1966
Male adult
N-Japan, C-Hokkaido, Hidaka Mt.Rng, Mt.Apoidak near Samani town, 20.VIII.1982,T. Akaishi leg.<br /> L=24 mm
Copyright © Ruslan Panin
Carabus (Megodontus) croaticus zepcensis (as bosiljevici Drovenik & Pavićević, 1985)
Male adult
CW-Serbia, Zlatar Mts, Nova Varoš, Zlatar Lake env., 1-12.VII.2007, P.Sima leg. L=30 mm
Copyright © Ruslan Panin
Carabus (Megodontus) caelatus dalmatinus (as procerus Reitter, 1885)
Male adult
S Bosnia, Potoci env. h=500 m, VII.2006, Bartoloni leg.
L=38 mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus dryas Gistl, 1857
Male adult
NE-Greece, Mt. Olympus, Litochoro env., 20-30.V.2005, Sima leg.
L=27 mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus vlasuljensis Apfelbeck, 1894
Male adult
WC-Serbia, Borak - Gornje Babine, 1-10.VII.2007, M. Sima leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus azurescens Dejean, 1826 (loc.typ.)
Male adult
W-Croatia, Velebit Mts., betw.Karlobag & Brusane vill., VI.2002, Manfred Egger leg.
L=30 mm
Copyright © Ruslan Panin
Carabus (Megodontus) caelatus caelatus (as malissorum Apfelbeck, 1919)
Male adult
Montenegro, National park Durmitor; near Zabljak vill., Mt. Sedlo h=2200 m, 6-22.VIII.2010, Franko Sandel leg.
29 мм
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus germarii Sturm, 1815
Male adult
N Italy, Trent, Ampola, Mt. Tremalzo h=1700m, 3-13.VI.2010, Manfred Egger leg.
29 mm
Copyright © Ruslan Panin
Carabus (Megodontus) bonvouloiri graciliformis Breuning, 1964
Male adult
NC Turkey, Gölköy env., Harcbeli pass, 35mm
Copyright © Aleš Sedláček
Carabus (Megodontus) bonvouloiri graciliformis Breuning, 1964
Female adult
NC Turkey, Gölköy env., Harcbeli pass, 37mm
Copyright © Aleš Sedláček
Carabus (Megodontus) kolbei chishimanus Nakane, 1961
Female adult
SC Kurils islands, 28mm
Copyright © Aleš Sedláček
Carabus (Megodontus) violaceus purpurascens Fabricius, 1787 (as osismius Lapouge, 1922)
Male adult
W-FRANCE, Brittany , Finistère, Botvarec Forest, 19.XI.2006, Pascal Stefani leg.
L=29 mm
Copyright © Ruslan Panin
Carabus (Megodontus) caelatus dalmatinus Duftschmid, 1812
Female adult
Bosnia, Mostar; 40 mm
Copyright © Aleš Sedláček
Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823
Male adult
W-Ukraine, Podolian Tovtry, Ternopol reg., Pidvolochysk distr., between Chagari-Zbarazhki & Shevchenkove vill., steppe sites, 6.V-7.VI.2012, Panin R. leg. L=25 mm
Copyright © Ruslan Panin
Carabus (Megodontus) bonvouloiri casalei Cavazzuti, 1986
Male adult
NE-Turkey: Artvin: Kabaca, env. Murgul, h = 1000 m, VII.1991, leg. St. Il. Remond
Copyright © Igor Solodovnikov
Carabus (Megodontus) bonvouloiri casalei Cavazzuti, 1986
Male adult
NE-Turkey: Artvin: Kafkäsor, h = 1700 m, VII.2000, leg. St. Il. Remond 32,5 mm
Copyright © Igor Solodovnikov
Carabus (Megodontus) bonvouloiri graciliformis Breuning, 1964
Female adult
NE-Turkey: Ordu: Sud Gölköy env., Harçbeli pass, h= 1500 m, VI.1999, leg. St. Il. Remond 40,5 mm
Copyright © Igor Solodovnikov
Carabus (Megodontus) bonvouloiri graciliformis Breuning, 1964
Male adult
NE-Turkey: Ordu: Sud Gölköy env., Harçbeli pass, h= 1400 m, VI.1999, leg. St. Il. Remond
Copyright © Igor Solodovnikov
Carabus (Megodontus) bonvouloiri bonvouloiri Chaudoir, 1863
Male adult
NE-Turkey: Rize: Ovitdagi Geç, h = 1700 m, VII.2000, leg. St. Il. Remond (as bonvouloiri kaschkarensis Blumenthal & Breuning, 1967; un)
Copyright © Igor Solodovnikov
Carabus (Megodontus) stroganowi galugahensis Lassalle 2001 Holotype
Male adult
Iran, Mazanderan, Mts 50km sud de Galugah, 5.2000 (in coll Lassalle)
Copyright © Bernard Lassalle
Carabus (Megodontus) violaceus aurichalceus Kraatz, 1879
Male adult
NC-Spain, Cantabria, Santander, Cueto env.XII.2013 L=28 mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus mixtus (as neomixtus Tarrier, 1975)
Male adult
L=29mm SE-France, Drôme, Foret de Saou, VII.2014, Pascal Stefani leg.
Copyright © Ruslan Panin
Carabus (Megodontus) kolbei hidakamontanus Ishikawa, 1966
Female adult
JAPAN, Hokkaido, Samani distr., Samani town, Horomankyo, 16.08.2013, Tsukasa, Nakamura leg. 25 мм.
Copyright © Sergey Kakunin
Carabus (Megodontus) leachi panzeri Dejean, 1829 Holotype
Male adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC48 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec48
Copyright © - - -
Carabus (Megodontus) violaceus violaceus trans violaceus purpurascens Fabricius,1787
Male+Female adult
E-Netherlands, Gelderland, Winterswijk Woold env.,15.II.2015, H.de Haan leg.
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus exasperatus Duftschmid, 1812
Male adult
Hungary, Budapest, Mt Buda, Harsbokor-hill,16-19.V.1992, I.Retezár leg., L=27 mm
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus aurolimbatus Dejean, 1829 Holotype
Male adult
Loc: "Moscow". http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec51 Muséum national d’Histoire naturelle, Paris (France) Collection: Insects - Coleoptera (EC) Specimen MNHN-EC-EC51
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Carabus (Megodontus) kolbei taikinus Ishikawa, 1993
Male adult
N-Japan, SC-Hokkaido:Tokachi, Hiroo Distr., Taiki-cho env., 28.VIII-2.IX.2017, Teruo Hirabayashi leg. L=24 mm
Copyright © Ruslan Panin
Carabus (Megodontus) persianus espeyliensis Deuve, 2001
Male adult
N-Iran, Gilan Prov., Deylaman Distr., Balarud Mts, Espili env., 18-24.VI.2017, Václav Čutka leg., L= 32 mm
Copyright © Ruslan Panin
Carabus (Megodontus) stroganowi morgani Lapouge, 1907
Male adult
Iran, Mazandaran prov., Nowshahr env, Khirood forest, 24.5.2013, 1200 m.
Copyright © Alexander Anichtchenko
Carabus (Megodontus) stroganowi tiedemanni Breuning, 1934
Male adult
Copyright © Alexander Anichtchenko
Carabus (Megodontus) persianus javahardehensis Deuve, 2000
Male adult
N-Iran, Mazandaran Prov., Ramsar Co., Javahar Deh env., 16-25.VI.2017, Václav Čutka leg. L=28 mm
Copyright © Ruslan Panin
Carabus (Megodontus) persianus javahardehensis Deuve, 2000
Female adult
N-Iran, Mazandaran Prov., Ramsar Co., Javahar Deh env., 16-25.VI.2017, Václav Čutka leg. L=30 mm
Copyright © Ruslan Panin
Carabus (Megodontus) avinovi Semenov & Znojko, 1932
Female adult
S Sakhalin, Dolinsky distr., Firsovo vill. env., 3.VII.2017 . Фотография с сайта www.zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov
Carabus (Megodontus) vietinghoffi rugicolor Rapuzzi, 2010 Holotype
Male adult
Reyneke island (CR). Image from Rapuzzi, 2012
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Carabus (Megodontus) obovatus Fischer, 1827
Male adult
C.obovatus taizhensis Obydov, 2006 Khakassia: Kuznetskyi Alatau, env. vill. Priiskovyi, Bobrovaya Mt., h = 1200-1300 m
Copyright © Igor Solodovnikov
Carabus (Megodontus) kolbei taikinus Ishikawa, 1993
Female adult
Japan, SC-Hokkaido:Tokachi, Hiroo Distr., Taiki-cho env., 3-7.IX.2019, S.Shinichi leg.
Copyright © Ruslan Panin
Carabus (Megodontus) kolbei kosugei Nakane, 1955
Male adult
Japan, SW-Hokkaido, Otaru city, 2-16.VIII.2020, S.Shinichi leg.
Copyright © Ruslan Panin
Carabus (Megodontus) kolbei hidakamontanus Ishikawa, 1966
Male adult
SE-Hokkaido: Samani distr.
Copyright © Igor Solodovnikov
Carabus (Megodontus) kolbei hidakamontanus Ishikawa, 1966
Male adult
E-Hokkaido: Samani distr., Horomankyo env.
Copyright © Igor Solodovnikov
Carabus (Megodontus) kolbei hidakamontanus Ishikawa, 1966
Female adult
E-Hokkaido: Samani distr., Horomankyo env.
Copyright © Igor Solodovnikov
Carabus (Megodontus) kolbei urupiensis Obydov, 2007 Holotype
Female adult
South Kuril Islands, Urup Island, north-western coast of Tokotan Lake, 21-29.VII. 2006, A. K. Klitin leg.
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Carabus (Megodontus) kolbei ushishirensis Obydov & Saldaitis, 1996 Holotype Paratype
Male+Female adult
Holotype: Ushishir Islands, Yankicha, N and SE part of Kraternaya, 47°30'N, 152°48'E, 20.VIII.1995, Yu. Marusik leg.". Paratype: Ketoy Island
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Carabus (Megodontus) leachi semiensis Obydov, 2010 Holotype
Male adult
E. Kazakhstan, Semei (Semipalatinsk) env., 12-26.VII.1992, V. Sinyaev leg.
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Carabus (Megodontus) croaticus antoniocaldoni Rapuzzi, 2014 Holotype Paratype
Male+Female adult
Serbia, Kopaonik mountain, 1500m, IX.2013, I. Rapuzzi and L. Caldon leg; (Ivan Rapuzzi collection).
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Carabus (Megodontus) tuvensis Shilenkov, 1996 Holotype Paratype
Male+Female adult
Holotypus (m) - Россия, Тува, 80 км С пос. Тээли, 25.06.1972, Б.А. Коротяев leg. Paratypus (f) - Россия, Тува, Саяны, р. Ак-Суг, 26.06.1972, Б.А. Коротяев leg.
Copyright © Dmitri Obydov
Carabus (Megodontus) obovatus pseudoleachi Obydov, 1999 Holotype
Male adult
Россия, Алтай, окр. пос. Яйлю, 06.1988, Водянов leg.
Copyright © Dmitri Obydov
Carabus (Megodontus) obovatus taizhensis Obydov, 2006 Paratype
Male adult
Россия, Кузне-кий Алатау, долина р. Тайже-Су, около Хакасской границы, 1250-1650 м, 53°55'N, 89°10'E, 25.VI-30.VII.2002, В. Мурзин и В. Синяев leg.
Copyright © Dmitri Obydov
Carabus (Megodontus) leachi sarymsaktensis Obydov, 1999 Paratype
Male adult
Казахстан, Южный Алтай, 2500 - 3400 м, хр. Сарым-Сакты, 21.VI.1998, A. Клименко leg.
Copyright © Dmitri Obydov
Carabus (Megodontus) danilevskii Obydov, 1993 Holotype Paratype
Male+Female adult
Россия, Зап. Саян, левый берег р. Енисей, хр. Джойский, 700-800 м, около пос. Черемушки, 20-28.VII.1992, Д. Обыдов leg.
Copyright © Dmitri Obydov
Carabus (Megodontus) vietinghoffi fulgidus Fischer, 1827 (as bureianus Shilenkov, 1996)
Male adult
Far East, Khabarovsk reg., Verhnebureinskiy distr., Bureinskiy N.R., Lan riv. 10.6.2007 (DUBC) 20 mm
Copyright © Alexander Anichtchenko
Carabus (Megodontus) planicollis verae Csiki, 1905
Female adult
C-Romania: Carpathians, E of Olt River: Fagaraş Mts, Balea Cascada, h = 1150 m
Copyright © Igor Solodovnikov
Carabus (Megodontus) leachi leachi Fischer, 1823
Female adult
NE Kazakhstan, W Altay, near Shemonaikha city., flood plain, 18-28.VII.2018, S.Vashchenko leg., L=22.5 mm
Copyright © Ruslan Panin
Carabus (Megodontus) leachi leachi Fischer, 1823
Male adult
NE Kazakhstan, W Altay, near Shemonaikha city., flood plain, 18-28.VII.2018, S.Vashchenko leg., L=21.5 mm
Copyright © Ruslan Panin
Carabus (Megodontus) croaticus croaticus Dejean, 1826
Female adult
W-Croatia, Velebit Mts., h=950 m, Oltari - Zavizan, 30.VI. 2018, L=32 mm
Copyright © Ruslan Panin
Carabus (Megodontus) croaticus croaticus Dejean, 1826
Male adult
W-Croatia, Velebit Mts., h=950 m, Oltari - Zavizan, 30.VI.2018
Copyright © Ruslan Panin
Carabus (Megodontus) vietinghoffi fulgidiformis Obydov, 2008 Holotype Paratype
Male+Female adult
North China, Gaolinzy env., 2-24.07.1996, A. Doroshenko leg.
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Carabus (Megodontus) caelatus dalmatinus Duftschmid, 1812
Female adult
Croatia, Brač island, 2025, D. Strelnikoff leg. (cAA)
Copyright © Alexander Anichtchenko
Carabus (Megodontus) caelatus dalmatinus Duftschmid, 1812
Male adult
Croatia, Brač island, 2025, D. Strelnikoff leg. (cAA)
Copyright © Alexander Anichtchenko
Carabus (Megodontus) caelatus dalmatinus Duftschmid, 1812
Male adult
Croatia, Brač island, 2025, D. Strelnikoff leg. (cAA)
Copyright © Alexander Anichtchenko -
Mesocarabus taxon page (150 photos)
Carabus (Mesocarabus) macrocephalus Dejean, 1826
Male adult
Spain, Picos de Europa, Macizo Central,
Llagu Bajero 1890m 6.8.2003
Copyright © Alexander Anichtchenko
Carabus (Mesocarabus) lusitanicus lusitanicus Fabricius, 1801
Male adult
Portugal, Vale de Estrela, Guarda, VI.1976
Copyright © Alexander Anichtchenko
Carabus (Mesocarabus) lusitanicus lusitanicus Fabricius, 1801(as schaumi Gaubil, 1849)
Male+Female adult
Portugal,
Viseu distr., Gosende, h~1010m, 15.X.2009, D.Prunier leg.
female,26 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) lusitanicus brevis Dejean, 1826
Female adult
Spain, Sierra Guadarrama, Montes Carpetinos, Pto. de Somosierra 1600-1700m, 3.VI.2006
Copyright © Alexander Anichtchenko
Carabus (Mesocarabus) breuningi Csiki, 1927
Male adult
Spain, San Martin de Castaneda, Zamora<br /> 11 oct 1984,h~1500 m
Copyright © Alexander Anichtchenko
Carabus (Mesocarabus) lusitanicus gougeleti Reiche, 1863 (as alcaracinus Kraatz, 1886)
Male adult
Spain, Riopa (Albacete)
Copyright © Alexander Anichtchenko
Carabus (Mesocarabus) dufouri dufouri Dejean, 1829 (as papillatus Lapouge, 1902)
Male adult
Spain, Algeciras, Molino de las Cuevas
5-6.4.2003 Anichtchenko A. leg.
Copyright © Alexander Anichtchenko
Carabus (Mesocarabus) baguenai Breuning, 1926
Male adult
Spain, Almeria, Sierra de Baza 10.III.1995<br /> male<br /> photo Anichtchenko A.<br />
Copyright © Alexander Anichtchenko
Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
Male adult
SW-France,
Foret d'Iraty Pyrenees-Atlantiques, d'Occabe, 15.VIII.1988
male,30 mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876 Loc.typ.
Female adult
Spain, Navarra, Sierra de Aralar,
Altxueta 1200-1300m 6.1.2001
Copyright © Alexander Anichtchenko
Carabus (Mesocarabus) problematicus harcyniae Sturm, 1815 (as obenbergerianus Breuning, 1927)
Male adult
Czech, Bohemia, Plzeň Reg., Domažlice Distr., Babylon, h=500 m, 30.VIII.2008, S.I. Budka leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) problematicus inflatus Kraatz, 1878
Male adult
SE–France, Var departament, Foret royale -Saint-Paul-en-Foret vill.,18.VI.2002, Thelot J.P. leg.
L=26 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) problematicus planiusculus Haury in Gehin, 1885
Male adult
NE Spain, Catalonia, Sant Jaume de Frontanyà, h=1000 m, V-VI.2010, Pascal Stefani leg.
L=24 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) latus latus Dejean, 1826
Female adult
CW Spain, Cáceres, San Martín de Trevejo, 24.VI.2011, A. Diez leg./det.Panin R.
L=27 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) dufouri dufouri Dejean, 1829
Male adult
SC Spain, Jaen reg., Monte Javalcuz h=1436 m, 6.X.2010, M.A. Lopez leg.
L=27 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) problematicus harcyniae (as procedens Csiki, 1927)
Male adult
England, North Yorkshire, Fylingdales Moor, Grouse Hill, VII.2006 L=24 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) lusitanicus brevis (as sorianus Jeanne, 1972)
Male adult
NC-Spain, Soria, Puerto de Oncala, 2.VI.2006
L=24 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) lusitanicus gougeleti Reiche, 186
Male adult
CW Spain, Province Huelva, Arroyomolinos de León,30.X.2011, T. Alcantara leg.; L=23 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) problematicus islandicus Lindroth, 1968
Female adult
SE-Iceland:Höfn env.:Hornafjördur, 24-28.VIII.2008, C. Reuter leg.
L=24 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) cantabricus cantabricus Chevrolat, 1840 (loc. typ.)
Male adult
NW-Spain, Pontevedra prov., A Cañiza viil., Alto de Fontefría h=800m, 31.XII.2007, T.Gazurek leg./det.Panin R.<br /> L=27 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) baguenai Breuning, 1926
Male adult
SE Spain, Almería Province, Carboneras env., 25.XI.2006, T.Gazurek leg./det. R.Panin <br /> L=26 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) macrocephalus Dejean, 1826
Male adult
NW-Spain,Cantabrian Mts, Picos de Europa Mt Rng, Puerto de Panderruedas h=1460 m, 25.IX.2004,T.Gazurek leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) latus cf. molossoides Lassalle, 1984
Male adult
C-Spain, Toledo, Mts de Toledo, Sierra del Castañar Rng, Puerto del Milagro env, h=920m, 7.II.2009, T.Gazurek leg./det.R. Panin.
L=24 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) problematicus harcyniae (as procedens Csiki, 1927)
Male adult
W-Ireland, County Mayo, Clare Isl., 17-28.VI.2002, S. McCormack leg.<br /> L=25 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) problematicus harcyniae (as procedens Csiki, 1927)
Female adult
N-Ireland, Fermanagh Distr., Lough Navar Forest, 17.VII.2013, D.Novak & J.Stypula leg.<br /> L=27mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) lusitanicus helluo (as albarracinus Ganglbauer, 1886)
Male adult
CE-Spain, Provincia Teruel, Sierra de Javalón Rng, El Cañigral env., h=1460m, X.2010, A. Mollard leg.<br /> L=23 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826
Male adult
CW-Spain: Salamanca, Brincones env., h=750m, X.2003<br /> L=25 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) problematicus harcyniae (as belgicus Lapouge, 1913)
Male adult
Luxembourg, Bollendorf-Pont env., 3.III.1999 /det.R Panin
L=24 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) (? problematicus trans lusitanicus) planiusculus (as xaxarsi Born, 1917)
Male adult
NE Spain, Catalonia, Girona Prov., San Hilario Sacalm env., 20.II.1988, A. Mollard leg.<br /> L=25 m
Copyright © Ruslan Panin
Carabus (Mesocarabus) lusitanicus helluo Dejean, 1826
Female adult
SE-Spain, Prov.Alicante, Port de la Carrasqueta, 26.XII.1987, M.Toribio leg. L=25 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) latus antiquus Dejean, 1826 Holotype
Female adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC36 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec36
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Carabus (Mesocarabus) lusitanicus brevis Dejean, 1826 Lectotype
Female adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC32 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec32
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Carabus (Mesocarabus) lusitanicus egesippei La Ferté-Sénectere, 1847 Holotype
Female adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC154 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec154
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Carabus (Mesocarabus) lusitanicus mertolaensis Breuning, 1975 (= brevis) Holotype
Female adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC130 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec130
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Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826 Holotype
Male adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC34 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec34
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Carabus (Mesocarabus) lusitanicus latus Dejean, 1826 Lectotype
Male adult
"Navalmoral" [Navalmoral de la Mata]. Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC35 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec35
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Carabus (Mesocarabus) lusitanicus trypalsi Breuning, 1975 Paratype
Female adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC132 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec132
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Carabus (Mesocarabus) lusitanicus trypalsi Breuning, 1975 Holotype
Male adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC133 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec133
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Carabus (Mesocarabus) lusitanicus helluo Dejean, 1826 Holotype
Female adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC31 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec31
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Carabus (Mesocarabus) dufouri papillatus Lapouge, 1902 Lectotype
Male adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC576 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec576
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Carabus (Mesocarabus) lusitanicus gougeleti (as valderepisensis Branes, 1988)
Male adult
Spain, Jaen prov., Despeñaperros, Arroyo de los Arcos, 38.381847, -3.515651, 1-2.05.2014, Anichtchenko A. leg. & det.; 18 mm
Copyright © Alexander Anichtchenko
Carabus (Mesocarabus) riffensis Fairmaire, 1875 Lectotype
Female adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC142 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec142
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Carabus (Mesocarabus) riffensis Fairmaire, 1875
Male adult
Morocco, Rif, Foresta di Ketama, 25.06.1998, 1600m, leg. D. Sechi (immature)
Copyright © Alexander Anichtchenko
Carabus (Mesocarabus) cantabricus moroderi (as rodriguezianus Meyer & Mollard in Deuve, 2003)
Female adult
Poza de la Sal, Burgos, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Mesocarabus) problematicus inflatus (as solidus)
Female adult
Ripoll, Barcelone, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Mesocarabus) problematicus planiusculus (as xaxarsi)
Male adult
Sant Sadurni, Gerone, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Mesocarabus) problematicus planiusculus Haury in Gehin, 1885
Male adult
La Seo de Urgel, Gerone, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Mesocarabus) problematicus planiusculus (as bepmalei Lapouge, 1904)
Female adult
Pic de Bassibes, Huesca, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Mesocarabus) problematicus planiusculus (as navarrensis Breuning, 1933)
Male adult
Mt.Urculo, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) problematicus planiusculus (as trapeti Bleuse, 1885)
Female adult
Panticosa, Huesca, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) problematicus planiusculus (as lagardei Mollard, 1996) Paratype
Female adult
Sierra de Andia, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Mesocarabus) problematicus planiusculus (as andorranus Lapouge, 1902)
Female adult
Col d'envalira, Pas de la Case, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Mesocarabus) lusitanicus lusitanicus (as vieirae Oliveira, 1875)
Male adult
Guia, Leiria, Portugal. det. A. Anichtchenko; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) lusitanicus lusitanicus (as schaumi Gaubil, 1849)
Male adult
S.de Gata, Extramadure, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Mesocarabus) lusitanicus castilianus (as fuentei Breuning, 1926)
Male adult
Retortillo, Salamanque, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) lusitanicus castilianus (as bolivari Breuning, 1926)
Male adult
Ledrada, Salamanque, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826
Female adult
Villasdardo, Salamanque, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) lusitanicus castilianus (as sayagoensis Mollard 2006) Paratype
Male adult
Moral de Sayago, Zamora, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) lusitanicus brevis (as olmedoensis Mollard, 2007) Paratype
Female adult
Olmedo, Valladolid, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826
Male adult
S.de Candelario, Salamanca ("Avila" error). Spain. det. A. Anichtchenko, 2016; (in Mollard, 2013 as lusitanicus brevis) Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) lusitanicus antiquus Dejean, 1826
Female adult
Portugal, Ribatejo, Coruche, 13.x.1974, A. Zuzarte leg., 25 mm, Coll. N. Mal
Copyright © Noël Mal
Carabus (Mesocarabus) lusitanicus latus (as molossoides Lassalle, 1984)
Male adult
Cabezabellosa, Extramadure, det. A. Mollard; Copyright © Magellanes
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Carabus (Mesocarabus) lusitanicus gougeleti (as nevadensis Brañes, 1989) Paratype
Male adult
Guadix, Granada, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Mesocarabus) lusitanicus gougeleti Reiche, 1863
Female adult
S.Norte de Sevilla, Huelva, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) lusitanicus antiquus Dejean, 1826
Male adult
Aliseda, Extramadure, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) lusitanicus helluo Dejean, 1826
Female adult
Puerto de Tortosa, Tarragone, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) lusitanicus trabuccarius Fairmaire, 1857
Female adult
Gaia, Barcelona, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) lusitanicus trabuccarius (as galardonensis Meyer & Mollard, 1996) Paratype
Female adult
Puerto de Monrepos, Huesca, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) cantabricus moroderi (as magdalenai Meyer & Mollard, 1996) Paratype
Female adult
Pto. de Magdalena, Leon, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) cantabricus moroderi Breuning, 1926
Male adult
Foncebadon, Leon, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) cantabricus moroderi (as almanzaensis Meyer & Mollard in Deuve, 2003) Paratype
Female adult
Almaza, Leon, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) breuningi Csiki, 1927
Male adult
Villanueva, Orense, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) cantabricus cantabricus (as asturicus Born, 1925)
Female adult
Pravia, Asturies, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) cantabricus cantabricus (as liebei Eidam, 1925)
Male adult
Fonsagrada, Galice, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) cantabricus cantabricus Chevrolat, 1840 (loc. typ.)
Female adult
La Caniza, Galice, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) cantabricus moroderi (as joaquini Breuning & Toulgoet, 1974)
Male adult
Pena Ubina, Leon, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) cantabricus amayensis Lassalle, 1984
Male adult
Pto.de Amaya, Burgos, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) macrocephaloides (Jeanne, 1972)
Female adult
Sierra de Andia, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) macrocephaloides (Jeanne, 1972)
Female adult
Sierra de Andia, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) macrocephaloides (Jeanne, 1972)
Female adult
Sierra de Andia, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) macrocephaloides (Jeanne, 1972)
Female adult
Sierra de Andia, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) riffensis Fairmaire, 1875
Female adult
Morocco : Rif, Ketama vi.1980 - V. Oliviella Leg. (Size : 33 mm)
Copyright © Noël Mal
Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
Female adult
Sierra de Aralar, Navarra, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) cantabricus brabeus (as barcelecoanus)
Male adult
Col de Velate, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) cantabricus brabeus (as barcelecoanus)
Female adult
Col de Velate, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
Male adult
Col de Velate, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) lusitanicus brevis (as segovianus Mollard, 2011) Paratype
Male adult
S.de Gorda, Soria, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) lusitanicus gougeleti (as alcaracinus)
Female adult
Alcaraz, Albacete, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) lusitanicus helluo (as alicantinus)
Female adult
S.de Bernia, Alicante, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) lusitanicus trabuccarius (as leyrensis)
Female adult
Sierra de Leyre, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) lusitanicus trabuccarius (as trypalsi)
Male adult
Plan du Trypals, Huesca, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) lusitanicus trabuccarius (as solanensis)
Male adult
S.de Bolave, Huesca, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) lusitanicus trabuccarius (as troncedoensis) Paratype
Female adult
Troncedo, Huesca, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) lusitanicus trabuccarius (as benabarrensis) Paratype
Male adult
Benabarre, Huesca, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) lusitanicus brevis (as pironensis) Paratype
Female adult
Pennasrubias, Segovia, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) lusitanicus brevis (as lermaensis) Paratype
Female adult
Espinosa de Cervera, Burgos, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) lusitanicus brevis (as logronicus)
Male adult
Pto.de Piqueras, La Rioja, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Mesocarabus) lusitanicus trabuccarius (as catalonicus)
Male adult
Ft.de Roquerolles, Prades, Tarragone, SPAIN
Copyright © Alain Mollard
Carabus (Mesocarabus) lusitanicus brevis (as segovianus)
Male adult
Sierra de Pardos, South Calatayud, Saragosse, SPAIN
Copyright © Alain Mollard
Carabus (Mesocarabus) lusitanicus helluo (as albarracinus)
Female adult
Altos de Cabrera, Pineda de Ciguela, Cuenca, SPAIN
Copyright © Alain Mollard
Carabus (Mesocarabus) riffensis Fairmaire, 1875 (as dominicae)
Male adult
Ft.de Ketama , Rif, MAROC
Copyright © Alain Mollard
Carabus (Mesocarabus) lusitanicus cf. antiquus-gougeleti
Male adult
Bicada, South Beja, PORTUGAL
Copyright © Alain Mollard
Carabus (Mesocarabus) problematicus problematicus Herbst, 1786
Female adult
W-Hungary, NE-Alps, Mts. Kőszeg, Velem env., Borha-forrás, 26.VII.1981, Gy. Somorijai leg. L=26 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) leptopus Thomson, 1875 Holotype
Female adult
3957:2 (MZLU)
Copyright © Lund University
Carabus (Mesocarabus) breuningi Csiki, 1927
Female adult
NW-Spain, Prov. Zamora, San Martín de Castañeda, 5.VII.1984, M.Toribio leg. L=28 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) dufouri Dejean, 1829
Female adult
S Spain, Granada, Sierra Arana, Píñar env., 14.X.2018, F. Garcia leg. L=27 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) breuningi Csiki, 1927 (loc.typ.)
Male adult
NW-Spain, Orense, Orensano Central Massif, Monte de Cabeza de Manzaneda, 25.VII.1988, F. Rodrigues leg. L=25 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
Male adult
NC-Spain, Basque Co, Álava, Sierra de Gorbea, VI.2014, L=29 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) lusitanicus helluo (as alicantinus Ganglbauer, 1886)
Male adult
SE-Spain, Murcia reg., Jumilla, Sierra del Carche, h =1300 m, 30.XI.2019, F. Garcia leg.
Copyright © Ruslan Panin
Carabus (Mesocarabus) lusitanicus antiquus Dejean, 1826
Female adult
SW Portugal, Beja Distr., 3,7 km NW Brunheiras, road CM1072, 37°46'31.9"N 8°47'02.2"W, 15.III.2011, D. Prunier leg., L=26 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) lusitanicus trabuccarius Fairmaire, 1857
Male adult
NE Spain, Catalonia, Tarragona, Sierra de Montsant h=800 m, Albarca env.,7.XI.2018, T. Rafales leg., L=25 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) lusitanicus trabuccarius Fairmaire, 1857
Female adult
NE Spain, Catalonia, Tarragona, Sierra de Montsant h=800 m, Albarca env.,7.XI.2018, T. Rafales leg., L=27 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) lusitanicus lusitanicus (as vieirae Oliveira, 1875)
Female adult
CW-Portugal: Leiria distr., near Marinha Grande, 39°45'57.8"N 8°58'10.3"W, h-79 m, 19.IX.2016, D. Prunir leg., L=26 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) lusitanicus lusitanicus (as vieirae Oliveira, 1875)
Male adult
CW-Portugal: Leiria distr., near Marinha Grande, 39°45'57.8"N 8°58'10.3"W, h-79 m, 19.IX.2016, D. Prunir leg., L=24 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) problematicus kolskianus Obydov, 2008 Holotype Paratype
Male+Female adult
Holotypus, male - Россия, Кольский полуостров, Мурман (Мурманск), Александровск (г. Полярный), Л. Зенкевич leg. Paratypus, female - Россия, Кольский полуостров, Мурман, окр. Александровска, 1914, Л. Зенкевич leg.
Copyright © Dmitri Obydov
Carabus (Mesocarabus) riffensis Fairmaire, 1875
Female adult
N-Morocco, Riff Mts h=1650 m, 26 km E of Bab Berred, 34.961333, -4.669417, Cedar forest, C. Reuter leg. L=33 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) riffensis Fairmaire, 1875
Male adult
N-Morocco, Riff Mts h=1650 m, 26 km E of Bab Berred, 34.961333, -4.669417, Cedar forest, C. Reuter leg. L=27 mm
Copyright © Ruslan Panin
Carabus (Mesocarabus) lusitanicus brevis Dejean, 1826
Female adult
Spain, Sierra de Guadarrama, Puerto de la Quesera, 1720m, 41.214640, -3.430896, Anichtchenko A. leg. 10.X.2025 (cAA).
Copyright © Alexander Anichtchenko -
Microplectes taxon page (14 photos)
Carabus (Microplectes) riedeli Menetries, 1832
Male adult
N-Ossetia,
Khazniaush Pass, h=2500-2700m, 9.VI-31.VII.1994, Zamotajlov A.leg.
male, 17mm
Copyright © Ruslan Panin
Carabus (Microplectes) convallium convallium Starck, 1889
Male adult
male, 19mm
Copyright © Boleslav Brezina
Carabus (Microplectes) convallium convallium Starck, 1889
Male adult
male, 19mm
Copyright © Boleslav Brezina
Carabus (Microplectes) convallium convallium Starck, 1889
Male+Female adult
female:NW-Caucasus, Russia,
Krasnodar reg., Apsheron Distr., Lagonaki Mt., near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominyh leg.
male:Karachaevo-Cherkessia,the right source of the river Beskess, h=2100m, 10.VIII.1997, I.A. Solodovnikov leg.
female, 21mm/male, 19mm
Copyright © Ruslan Panin
Carabus (Microplectes) convallium argonautarum Semenov, 1898
Male adult
male, 18mm
Copyright © Boleslav Brezina
Carabus (Microplectes) convallium argonautarum Semenov, 1898
Male adult
male, 19mm
Copyright © Boleslav Brezina
Carabus (Microplectes) convallium argonautarum Semenov, 1898
Female adult
female, 20mm
Copyright © Boleslav Brezina
Carabus (Microplectes) convallium argonautarum Semenov, 1898
Male+Female adult
Abkhasia,
W.slope of Gagrinsky Mt., Krepostnaya River source, h=930-980m., A.S.Zamotajlov leg. 27.VII.2000
female, 19mm
photo Panin R.
Copyright © Ruslan Panin -
Mimocarabus taxon page (11 photos)
Carabus (Mimocarabus) maurus osculatii Osculati, 1844
Male adult
Iran
Copyright © Alexander Anichtchenko
Carabus (Mimocarabus) maurus maurus Adams, 1817
Male+Female adult
Armenia, Ashotsk distr., Tsoghamarg vill., 24-27.V.1999 P.V.Kazaryan leg. female, 20mm/male,18mm
Copyright © Ruslan Panin
Carabus (Mimocarabus) pumilio Kuster, 1846
Male adult
S-Armenia, Vayots Dzor Region, E Ughedzor vill., h=2630 m, 9.VI.2014, L=16 mm
Copyright © Ruslan Panin
Carabus (Mimocarabus) maurus smyrnensis Deuve, 2009 Holotype
Male adult
Turquie, Anatolie occidentale, Izmir (« Smyrne, Asie Mineure »), in Muséum national d’Histoire naturelle, Paris.
Copyright © Thierry Deuve
Carabus (Mimocarabus) maurus paphius Redtenbacher, 1843 Lectotype
Male+Female adult
Lectotype female (left) from collection Kotschy (Muséum de Vienne); right - « Chypre » Muséum de Paris
Copyright © Thierry Deuve
Carabus (Mimocarabus) maurus maurus (Adams, 1817)
Male adult
E Georgia, Kakheti, near Patardzeuli vill., h=750-850 m, 10.IV.2015, E. Rutjan leg., L=22 mm
Copyright © Ruslan Panin
Carabus (Mimocarabus) maurus osculatii Osculati, 1844
Female adult
Iran, Chaharmahal and Bakhtiari prov., 5 km W Samsami, C. Zagros mts., 3500m, 5-6.6.2008, Anichtchenko A. leg. (cAA)
Copyright © Alexander Anichtchenko -
Morphocarabus taxon page (436 photos)
Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
Female adult
Copyright © Evgeniy Chibilov
Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
Male adult
Russia,
S Ural Mts, Mt.Iremel h=800m, 4.VI.2000 Filimonov V. leg.
male,21mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
Male adult
Carabus (Morphocarabus) karpinskii Rusia, S Ural mts., Jremel Mt., 1300 m 4.6.2000male, 20 mm
19.08.2008
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) karpinskii Kryzhanovskij & Matveev, 1993
Female adult
Copyright © Evgeniy Chibilov
Carabus (Morphocarabus) karpinskii Kryzhanovskij & Matveev, 1993
Female adult
Copyright © Evgeniy Chibilov
Carabus (Morphocarabus) illigeri illigeri (as bjelasnicensis Apfelbeck, 1902)
Male+Female adult
Serbia, Debelo Brdo, 5.VI.2007, S. Sapaly leg. L=32 mm photo Panin R.
Copyright © Ruslan Panin
Carabus (Morphocarabus) kollari Palliardi, 1825
Female adult
female, 24 mm
Copyright © Boleslav Brezina
Carabus (Morphocarabus) rothi hampei Kuster, 1846
Male+Female adult
W.Ukraine.
Transcarpathian region,Vinogradiv distr., near Malaya Kopanya vill., Chorna Gora Mt., h=180m. 7-18.VII.2008. Panin R. leg.
female 33 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi hampei Kuster, 1846
Male adult
W.Ukraine.
Transcarpathian region,Vinogradov distr., near Malaya Kopanya vill., Chorna Gora Mt., h=180m. 7-18.VII.08. Panin R. leg.
male, 28 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi hampei Kuster, 1846 (as ormayi Reitter, 1896) loc.typ.
Female adult
W.UKRAINE,
Transcarpathian reg., near Beregovo,15.V-16.VI.2009 Kanarsky Yu. & Geryak Yu. leg.
female, 30 mm
photo/det. Panin R.
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi hampei Kuster, 1846
Female adult
Carabus (Morphocarabus) rothi hampei
W Ukraine, Zakarpatian reg., Vinogradov
1.7.1989 Avdeev I. leg.
female, 31 mm
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) rothi hampei Kuster, 1846
Female adult
arabus (Morphocarabus) rothi hampei
W Ukraine, Zakarpatian reg., Vinogradov
1.7.1989 Avdeev I. leg.
female, 31 mm
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) spasskianus schestopalovi Plutenko, 1995
Male adult
NE-Kazakhstan,
S-Altay, Sarym-Sakty Mt. h~2500m., Sarym-Sakty riv. 24.VI.1999
male, 23mm.
Copyright © Ruslan Panin
Carabus (Morphocarabus) spasskianus schestopalovi Plutenko, 1995
Male adult
SE Kazakhstan,
S Altay Mts., Sarymsakty mt. rng., Sarymsakty riv.
22.VI.2000 2400m, Nekroshevich S. leg.
male, 20 mm
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) kollari Palliardi, 1825
Male+Female adult
Romania, SW Carpathians,
Banat, Anina Mts. (Muntii Aninei) 8 km NE Oravita, 7.VI.2007, S.I. Budka leg.
female, 34 mm
photo Panin R
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii zawadzkii Kraatz, 1854
Male adult
Ukrainian Carpathians
Lviv Reg., Skole Distr., near Grebeniv vill., h=1000m 9 - 20.VII.07, Panin R. leg.
male, 31 mm
Copyright © Safronov
Carabus (Morphocarabus) wulffiusi wulffiusi Morawitz, 1862
Male+Female adult
Russia, Far East,
Primorskiy Kray, near Kamenushka vill., 2-3.VIII.1992, K.E.Dovgaylo leg.
female, 24mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Male adult
Serbia, Majdanpek, Ceremosja 15.6.2001 Z.Zlatic leg. male, 32 mm
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Male+Female adult
NE-Serbia, Krs Mts., Majdanpek env.,h=300m.20-31.V.2006, Pr.Sima leg. female, 32mm/male, 30mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) venustus venustus Morawitz, 1862
Male adult
SE Primorskii reg., Lazovskii Nat.Res., 12 km S Staraya Kamenka vill., near Amerika kor. 43°15N, 134°1E, 19.VII.04 leg. M.& L. Smirnovy photo from www.zin.ru
Copyright © Maksim Smirnov
Carabus (Morphocarabus) tarbagataicus pietrorattii Deuve, 1991(as pseudogebleri Deuve, 1994)
Male adult
Carabus (Morphocarabus) tarbagataicus pietrorattii Deuve, 1991(as pseudogebleri Deuve, 1994)
S Tarbagatai mt.rng., Alexeevka vill.,
600m, 11.5.2001 Klimenko A. leg.
male, 30 mm
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997 Holotype
Male adult
Rusia, SE Tuva, Sangilen mt.rng., Kargy riv., 50°31'N 97°01'E, 1800 m, 29.07.1996, D. Obydov leg.
Copyright © Dmitri Obydov
Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997
Female adult
Tuva, Akademika Obrucheva mt.rng., 15 km N Cherbi vill. ur. Chailag 7.VII.1999 A. & N. Menschikovy. 22 mm. det. Menschikov A.
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997
Male adult
Tuva, Akademika Obrucheva mt.rng., 15 km N Cherbi vill. ur. Chailag 7.VII.1999 A. & N. Menschikovy (cAA). 21 mm. det. Menschikov A.
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) spasskianus tungus Obydov, 1999 Holotype
Male adult
Rusia, N Krasnoyarsk reg., Ilimpeya riv and Dovogpokan riv. VII.1974, Lunin leg. photo from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Morphocarabus) spasskianus spasskianus Breuning, 1932
Male adult
Siberia, Lake Baikal, Listvianka vill. 25.8.90
Anichtchenko A. leg.
male, 23 mm
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) spasskianus spasskianus Breuning, 1932
Male+Female adult
SE Siberia, Russia
Irkutsk reg., near Pivovarikha vill., 7.V.07, D. Fominyh leg.
male, 24mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) spasskianus shoriensis Obydov, 1999 Holotype
Male adult
Gornaya Shoria, 8.VIII.1978, S. Nikireev leg.
Copyright © Dmitri Obydov
Carabus (Morphocarabus) spasskianus saragaschensis Obydov, 1999
Male adult
W Sayan Mts., Batenevskii Mt.rng., 12 km E Pervomaiskoe vill., Enisey riv., mountain steppe, 400m, 5.VII.2000 A. & N. Menschikovy leg. (cAA). 26 mm
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) spasskianus saragaschensis Obydov, 1999 Holotype
Male adult
Rusia, Khakassia, Saragash, 400 m, 6.VII.1994, D. Obydov leg.
Copyright © Dmitri Obydov
Carabus (Morphocarabus) spasskianus putus Motschulsky, 1846
Male adult
Carabus (Morphocarabus) spasskianus putus Motschulsky, 1846
SW Siberia, Altay Mts., Shebalino distr., Kumalyr env.
02.07.2003, A.Safronov leg.
Copyright © Safronov
Carabus (Morphocarabus) spasskianus microputus O.Berlov & Shilenkov, 1996
Female adult
Tuva, 25 km E Ak-Dovurak 23.VI.2006, A. & N. Menschikovy leg. 24 mm det. Menschikov A.
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) spasskianus microputus O.Berlov & Shilenkov, 1996
Male adult
Tuva, 25 km E Ak-Dovurak 23.VI.2006, A. & N. Menschikovy leg. 24 mm. det. Menschikov A.
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) spasskianus microputus O.Berlov & Shilenkov, 1996
Male adult
Rusia, SW Tuva, W Tannu-Ola mt.rng., bass. of Barlyk riv., 50°29'N 90°44'E, 30.VII.2000, D. Obydov leg.
Copyright © Dmitri Obydov
Carabus (Morphocarabus) spasskianus korostelevi Obydov, 2004 (loc.typ.)
Male adult
E Kazahstan, Ivanovskii Mts., Prohodnoy Belok, 03.06.2000, A. Ivanov leg.
Copyright © Anton Menstschikov
Carabus (Morphocarabus) spasskianus geblerianus Obydov, 1999 Holotype
Male adult
Rusia, Barnaul, 27.5.1965
Copyright © Dmitri Obydov
Carabus (Morphocarabus) spasskianus cracens Obydov, 1999
Male adult
NE Kazakhstan, S Altay Mts., Kalbinskii range, Ognevka vill., 600m, 18.6.1999 A. Klimenko leg. (cAA). 22 mm
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) spasskianus cracens Obydov, 1999 Paratype
Female adult
DUBC
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) spasskianus cracens Obydov, 1999
Female adult
NE-Kazakhstan: S-Altay, Kalbinsky Mt., Ognevka vill., h~600m 18.VI.1999 A. Klimenko leg. 24 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) spasskianus cracens Obydov, 1999 Holotype
Male adult
E Kazakhstan, Kolbinskii Mt.rng., 600m, near Ognevka vill., 13-18.07.1999, D. Obydov leg<br /> photo from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
Female adult
Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
Moravia mer.,Vranov, 26-29.06.1979, Dr. Resl leg.
photo A. Menstschicov
Copyright © Anton Menstschikov
Carabus (Morphocarabus) scheidleri Panzer, 1799
Male adult
SE Moravia mer.,Vranov, 26-29.06.1979, Dr. Resl leg.<br /> photo A. Menstschicov
Copyright © Anton Menstschikov
Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
Male adult
Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
S.Bohemia,
Hluboka nad Vltavou, 24.VI-18.VII.1990, S. Budka leg.
male, 27mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799 (as scheidleri floriani)
Male adult
A.: Steiermark, Rein Umg. VIII.1983 Kenyery<br /> female, 32 mm
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) excellens excellens Fabricius, 1798
Male adult
NW Podolian
Copyright © Safronov
Carabus (Morphocarabus) excellens excellens Fabricius, 1798
Male adult
Moldova, 2003
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens (as lomnitzkii Reitter, 1896 )
Male adult
W-UKRAINE, NW Podolian, Lviv Reg., Zolochiv Distr., W Ukraine, Lysa Gora steppe massif, 19.V.-7.VI.2007, Panin R. leg. 25 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens excellens Fabricius, 1798 (ab.erythromerus Dejean, 1826)
Male adult
Moldova, Anenii Noi distr., near Garbovets vill. 20.VI.08 Bodnar' I.I.leg. male, 26mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens (as lomnitzkii Reitter, 1896)
Male adult
W Ukraine, NW Podolian, Ternopil reg., Pidvilochisk distr., near Rosokhuvatets & Kamyanki vill., steppe sites, 24.V-15.VI.2008, Panin R. leg.
male, 24 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) helleri Ganglbauer, 1893
Male adult
Moravia Mer., Bile Karpaty 28.5.1993 Budka leg. male, 28 mm
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) helleri Ganglbauer, 1893
Male+Female adult
SE Moravia, Strani, V.2002, M.Snizek leg. male bronze colour-26mm male blue colour-28mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi incompsus (as mendax Csiki,1906)
Male+Female adult
W.Ukraine,
Transcarpathian Reg., Rakhov Distr., Chornogora Mt Range, h=1900m. alpine zone, Panin R. leg/det. 28.VI-9.VIII.2008
female, 32 mm
photo endophallus Solodovnikov I.A
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi comptus Dejean, 1831
Male adult
Carabus (Morphocarabus) rothi comptus Dejean, 1831
Carpathians
male, 23mm
coll.SMNH Ukraine ,Lvi
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi alutensis Savulescu, 1972
Male adult
C Romania,
Olt River: Rimnica Vilcea VII.1971, Savulescu N. leg.
Copyright © Igor Solodovnikov
Carabus (Morphocarabus) regalis regalis Fischer, 1822
Male adult
Carabus (Morphocarabus) regalis regalis Fischer, 1822
S. Ural, Bashkiria, near Mesjagutovo vill., 09.07.2001 A. Menstschicov leg.
photo A. Menstschicov
Copyright © Anton Menstschikov
Carabus (Morphocarabus) regalis jenissoni Dejean, 1831
Male adult
Carabus (Morphocarabus) regalis jenissoni Dejean, 1831
Rusia, Tuva, Todzh, Azas lake, 990 m,
52°24'N 96°33'E, 8.VII.1998, D. Obydov leg.
Copyright © Dmitri Obydov
Carabus (Morphocarabus) regalis jenissoni Dejean, 1831
Female adult
Carabus (Morphocarabus) regalis jenissoni Dejean, 1831
SW Siberia, Altays Mts, Choya distr., Sukhoy Karasuk env.
23.07.2004, A.Safronov leg.
Copyright © Safronov
Carabus (Morphocarabus) regalis eltoni Obydov, 2006 Holotype
Male adult
Carabus (Morphocarabus) regalis eltoni Obydov, 2006 HT<br /> Rusia, Tver distr., near Ilyinskoe vill.,<br /> 29-30.VII.2006, D. Obydov leg.<br /> photo from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Morphocarabus) odoratus septentrionalis Breuning 1932
Female adult
N. Russia, arctic Urals, Paypudynya Riv., Pompeliz Mt.h=500-700 m.,10-15.07.2002
S. Andreev, A. Romanenko leg.
Copyright © Safronov
Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844: 100
Male adult
<b>Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844</b><br /> Buriatia, W Hamar-Daban Mts.,<br /> up riv. M.Zangisan, Baishint Ula Mt. 2300m<br /> 23.5.1999 Anichtchenko A. leg.<br /> male, 18 mm
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844: 100 Lectotype
Male adult
Carabus (Morphocarabus) odoratus odoratus LT<br /> Lectotype<br /> Hamar-Gaban Mts.<br /> photo from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844 (as melleus Lapouge, 1909)
Male+Female adult
Russia, S Siberia, East Sayans, Mondy circ., h~2300m., 5-8.VI.2001, R. Mishustin leg. 20mm.
Copyright © Ruslan Panin
Carabus (Morphocarabus) odoratus martjanovianus Obydov 1999 HT Holotype
Male adult
Carabus (Morphocarabus) odoratus martjanovianus Obydov 1999
Rusia, Sayan Mts., near Sayanogorsk, 7.VII.1993
photo from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Morphocarabus) odoratus magadanicus Obydov 1999 PT Paratype
Male adult
Carabus (Morphocarabus) odoratus magadanicus Obydov 1999 PT
Rusia, Far East, E of Magadan, W Nyuklia vill.
VII.1997, Yu. Marusik leg.
photo from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Morphocarabus) odoratus krugeri Obydov 1999 PT Paratype
Female adult
Carabus (Morphocarabus) odoratus krugeri Obydov 1999 PT
Rusia, Tuva, Akademika Obrucheva Mt.rng., 2100, 51°56'N 095°32'E,
Koptu pass, tundra 2.VII.1998, D. Obydov leg.
photo from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Morphocarabus) odoratus kamtchatensis Breuning 1942
Male adult
Carabus (Morphocarabus) odoratus kamtchatensis Breuning 1942
Rusia, Kamchatka peninsula, VII.21
photo from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Morphocarabus) odoratus dohrni Gebler, 1847
Male adult
Carabus (Morphocarabus) odoratus dohrni Gebler, 1847
Rusia, Kuznetskii Alatau, Taizhe-Su riv., 53°55'N 89°10'E,
1250-1650 m, 25.VI-30.VII.2002, V. Murzin and V. Sinyaev leg.
photo from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Morphocarabus) odoratus dabanensis Shilenkov 1996 loc.typ.
Male+Female adult
Carabus (Morphocarabus) odoratus dabanensis Shilenkov 1996 loc.typ.
Buryatia, Hamar-Daban mt. rng., Snezhnaya riv.,
4.6.2007 Anichtchenko A. leg.
male 17 mm, female 19 mm
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) odoratus czernyscheviellus Obydov, 2002 PT Paratype
Female adult
Carabus (Morphocarabus) odoratus czernyscheviellus PT
Rusia, SE Transbaikalia, Chita reg., Nerchinsko-Kugenskii Mt.rng.,
near Vershino-Darasunskii vill., Bogatui Mt. 1221 m.,
15.VII.2002, D. Obydov leg.
photo from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Morphocarabus) odoratus bargusinicus Shilenkov, 1996
Male+Female adult
Carabus (Morphocarabus) odoratus bargusinicus Shilenkov, 1996
Russia, N Burajatia,
Barguzinskiy Mts., the upper reaches of M.Chiverkuy riv., h=2000m, Vashchenko S. leg.
female, 21mm/male,19mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Morphocarabus) odoratus antropovi Shilenkov 1996
Male+Female adult
Russia, S Tuva, Khorumnug -Tayga mts. Range, h=1600-2300m, Buren riv. valley, 4-26.VI.2000 Vashchenko S & A. leg<br /> female, 20mm/male,18mm<br /> photo Panin R.
Copyright © Ruslan Panin
Carabus (Morphocarabus) odoratus ?antropovi Shilenkov 1996
Male adult
S Tuva, Horumnut-Taiga Mt.rng., Bureya riv., 2000 m, tundra, Vaschenko S. leg. 18 mm
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) monilis Fabricius, 1792
Male adult
France 39, Foncina le Haut 300m, 23.6.1979; 29 mm
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) monilis Fabricius, 1792 (as taunicus)
Male adult
W Germany, Vogelsberg Mt., 700m, 13.5.1985
male, 24 mm
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) monilis Fabricius, 1792 (as interpositus Géhin, 1880)
Male adult
NW France,
Basse-Normandie, Orne, Forкt d'Йcouves, V.2009, Pascal Stefani leg.
male, 29mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) monilis (as interpositus Gehin, 1880)
Male+Female adult
NW France,
Haute-Normandie reg., Bois Les Loges, Etretat, Seine-Maritime,10.V.1988
female, 28mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) monilis (as amoenus Baudet-Lafarge, 1836)
Female adult
C France,
Puy-de-Dome, Col de la Croix Saint Robert h=1300m, VII.2008, Pascal Stefani leg.
female, 24 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) monilis (as helveticus Heer, 1837 Holotype) Holotype
Male adult
Carabus (Morphocarabus) monilis (as helveticus Heer, 1837 Holotype)
?hybrid of monilis/cancellatus
Museum of Zurich
photo Anichtchenko
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) monilis (as dupeuxi Deuve, 1974)
Male+Female adult
France,
Charente, Foret de Braconne VII.1998, Laforgue leg.
famale, 27mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) michailovi Kabak, 1992
Male adult
NE Kazakhstan, S Altay Mts., Sarymsakty mt. rng.,
Sarymsakty riv., 2400 m, 22.6.2000 Nekroshevich S. leg.
male, 18 mm
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) michailovi Kabak, 1992
Male adult
NE Kazakhstan, S Altay Mts., Sarymsakty mt. rng.,
Sarymsakty riv., 2400 m, 22.6.2000 Nekroshevich S. leg.
male, 17 mm
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) michailovi Kabak, 1992
Male adult
NE-Kazakhstan,
S Altai Mts.,Sarym-Sakty Range, Sarym-Sakty riv., h=2400m. 21.VI.1999., A.Klimenko leg.
male,18mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) mestscherjakovi telezkianus Shilenkov, 1996
Male adult
Carabus (Morphocarabus) mestscherjakovi telezkianus Shilenkov, 1996
Rusia, Altai, S shore of Teletskoe lake, 5.VII.1968
Copyright © Dmitri Obydov
Carabus (Morphocarabus) mestscherjakovi telezkianus Shilenkov, 1996
Male adult
Carabus (Morphocarabus) mestscherjakovi telezkianus Shilenkov, 1996
SW Khakasia, Abakanskii Mt.rng., Choochek Mt., 20 km SSE Mrassu vill.,
1800m, 7-19.7.1999 Lomakin D. leg.
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
Male adult
W Sayan mts., Kopen mt., Dzhabash riv., 75 km
S from Arbaty vill., 2235m, 26-30.6.2003 Brinev A. leg.
long. 17 mm
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
Male adult
Russia, S Siberia,
S Khakassia, N Tuva reg., Sailyg-Khem-Taiga mts. range h=2400m, 10.VII.2004
male, 18mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
Male+Female adult
Russia, S Siberia,
S Khakassia, N Tuva reg., Sailyg-Khem-Taiga mts. range h=2000-2600m,10-30.VI.2005, S.Vashchenko & S.Alexeenko leg.
male,17.5mm/female,19mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Morphocarabus) mestscherjakovi pseudoodoratus Brinev, 2002 Paratype Paratype
Male adult
Krasnojarsk reg., Aradan distr., 5 km SW Aradan vill.,<br /> Us riv., forest, 52є32`N 93є27`E 9-12.VII.1998 A.Brinev leg<br /> male, 18 mm
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) mestscherjakovi pseudoodoratus Brinev, 2002 Paratype Paratype
Female adult
Krasnojarsk reg., Aradan distr., 5 km SW Aradan vill., vall. of Us riv., A.Brinev leg.<br /> female, 20 mm<br /> photo & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
Male adult
Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
W Sayan, Olenya rechka
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
Male adult
Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
W Sayan, Kulumys Mt.R., env. Oiskoje Lake, mountain tundra, 12-14. 07.1999 A.Semin leg.
male
L=17,0 mm
Copyright © Maksim Smirnov
Carabus (Morphocarabus) kozhantschikovi martjanovi Lutshnik, 1924
Female adult
Khakassia, 1800 m, near Cheremushki vill., 12.6.-20.7.2000<br /> female, 21 mm
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
Male adult
W Sayan Mts., Kazhyr-Suk riv., 80 km W Oleniya<br /> Rechka, W part of Oiskii mt. rng., 30.6.-7.7.1999<br /> 2100 m, Brinev A. leg.<br /> male, 19 mm
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
Male adult
W Sayan, low fl. Kazyr-Suk riv., 80km W meteostation Olenia Rechka, W part of Oiskii mt.rng., 30.VI-7.VII.1999 2100m, Brinev A. leg.<br /> male, 20mm<br /> det. Brinev A.<br /> photo Anichtchenko A.
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 18789 (as viridipennis Kraatz, 1886, nec Lewis,
Male adult
E Rusia, Khabarovsk reg.,
near Birobidzhan VI.1991
male, 26 mm
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
Male adult
Rusia, S Primorie, Vladivostok distr., Razdolnaya riv., near Kluchevoi vill., 10-14.VII.1988, S. Muhanov leg.; Obydov det. as tristiculus Kraatz, 1879
Copyright © Dmitri Obydov
Carabus (Morphocarabus) hummeli cf pusongensis Imura, 1993
Male adult
China, Beijing, Wulingshan mountain, 1985
Copyright © Ye Liu
Carabus (Morphocarabus) hummeli ssp.
Male adult
China, East Inner Mongolia, Chifen, 2006, Shi H.L. leg. Photo by liuye
Copyright © Ye Liu
Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
Male adult
China, Jinling pro.;
Photo by liuye
det. Anichtchenko A.
Copyright © Ye Liu
Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
Male adult
Russia, Far East,<br /> Primorskiy Kray, near Kamenushka vill., 2-3.VIII.1992, K.E.Dovgaylo leg.<br /> male, 27mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) hummeli pusongensis Imura, 1993
Male adult
S Primorie, Chernye gory, Poima riv.,
30.7-5.8.1999 Anichtchenko A. leg.
male, 26 mm
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) hummeli nevelskii Shilenkov, 1996
Female adult
Rusia, S Sakhalin isl., VII.1993, G. Vasiliev leg.
Copyright © Dmitri Obydov
Carabus (Morphocarabus) hummeli nevelskii Shilenkov, 1996
Male+Female adult
Russia, Sakhalin isl.,
Makarov distr., Lesnaja riv., VII.1992, M. Nesterov leg.
female,23mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) hummeli hummeli Fischer, 1823
Male adult
Carabus (Morphocarabus) hummeli hummeli Fischer, 1823
Rusia, Chita reg., Darasun, VII.1991
I. Avdeev leg.
male, 23 mm
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) hummeli biamensis Obydov, 2007 Paratypus Paratype
Male adult
Russia, North Primorie, B. Svetlovodnaya riv. (Biame riv.), 40 km from source, 30.07.1976, A. Klitin leg.
Copyright © Dmitri Obydov
Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
Female adult
E.RUSSIA, Primorsky ter., Chuguevsky distr., 30 km ESE from Chuguevka, 9-14.VIII.2007
Copyright © Nil Belonogov
Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
Male adult
E.RUSSIA, Primorsky ter., Chuguevsky distr., 30 km ESE from Chuguevka, 9-14.VIII.2007
Copyright © Nil Belonogov
Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 18789 (as viridipennis Kraatz, 1886, nec Lewis,
Female adult
Primorskii reg., near Kovalerovo<br /> VII.1990 Evstigneev A. leg.<br /> female, 25 mm
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) hummeli ? middendorfi Menetries, 1851
Male adult
Carabus (Morphocarabus) hummeli ? middendorfi Menetries, 1851
Rusia, Khabarovsk reg., Ohotsk sea, Port Ayan, 08.24
photo from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Morphocarabus) henningi pelevini Obydov 2006
Female adult
NE Kazakhstan, S Altai mt.rng., Sarymskiy mt. rng., Sarymsakty riv.,<br /> 2400m, 22.VI.2000 Nekroshevich S. leg. female 17mm det. Anichtchenko A.
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) henningi pelevini Obydov 2006 HT
Male adult
Carabus (Morphocarabus) henningi pelevini Obydov 2006 HT
Kazakhstan, S Altai, Sarym-Sakty Mt.rng., 3000-3400 m,
18-20.VI.1998, A. Klimenko leg.
Copyright © Dmitri Obydov
Carabus (Morphocarabus) henningi peczoricus Obydov 2000 Holotype
Male adult
Rusia, Komi republic, Kosiyu River mouth, (Pechora basin) 21.VII.1980. photo from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Morphocarabus) henningi oviformis Beheim & Breuning, 1943
Male adult
Carabus (Morphocarabus) henningi oviformis Beheim & Breuning, 1943
Rusia, Krasnoyarsk reg., Taimyr AO, near Norilsk,
07.2004, M. Shestopalov leg.
photo from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Morphocarabus) henningi melyachi Shilenkov & Menstshikov, 1996
Male adult
Carabus (Morphocarabus) henningi melyachi Shilenkov & Menstshikov, 1996
C Altai mts, Seminskii mt.rng., Katun riv., 3km S Ust-Sema vill.
4.VII.2002 A. & N. Menschikovy
male, 17mm
det. Menschikov A.
photo Anichtchenko
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) henningi henningi Fischer, 1817
Male adult
S. Ural, Bashkiria, near Mesjagutovo vill., 09.07.2001 A. Menstschicov leg.
Copyright © Anton Menstschikov
Carabus (Morphocarabus) henningi henningi Fischer, 1817
Male adult
SE Siberia, Russia, Irkutsk reg., near Pivovarikha vill., 7.V.07, D. Fominykh leg. female, 21mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) henningi hamarensis Shilenkov, 1996
Female adult
S Siberia, Hamar-Daban mts., Snezhnaya riv., 1-6.VI.2007, Anichtchenko A. leg. & det., 20 mm
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) henningi hamarensis Shilenkov, 1996
Female adult
Carabus (Morphocarabus) henningi hamarensis Shilenkov, 1996 loc.typ.
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) gebleri ultimus Obydov, 1999
Female adult
Carabus (Morphocarabus) gebleri ultimus Obydov, 1999 PT
E Kazakhstan, Altai, near Zyrjanovska, 6.VII.1995, D. Obydov leg.
female
photo from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Morphocarabus) gebleri ultimus Obydov, 1999 Holotype
Male adult
Carabus (Morphocarabus) gebleri ultimus Obydov, 1999 HT
E Kazakhstan, Altai, near Zyrjanovska, 6.VII.1995, D. Obydov leg.
male
photo from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Morphocarabus) gebleri gebleri Fischer, 1817
Female adult
Carabus (Morphocarabus) gebleri gebleri Fischer, 1817
Russia, Altai, Zmeinogorsk
female
photo from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Morphocarabus) gebleri gebleri Fischer, 1817
Male adult
Carabus (Morphocarabus) gebleri gebleri Fischer, 1817
Rusia, Altai, Zmeinogorsk
male
photo from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Morphocarabus) chaudoiri zaikai Obydov, 1999 Paratype
Female adult
Rusia, Tuva, Akademik Obruchev Mt. rng., 2100 m;, 52°00'N 95°34'E, Sainak pass, tundra, 2.VII.1998, D. Obydov leg.
Copyright © Dmitri Obydov
Carabus (Morphocarabus) chaudoiri juliae Obydov, 2002
Male adult
male, 18mm
Copyright © Boleslav Brezina
Carabus (Morphocarabus) chaudoiri czadanicus Obydov, 1997 Holotype
Male adult
Rusia, Tuva, near Hadyn vill., 28-30.VI.1994, R. Shmarzh leg.
Copyright © Dmitri Obydov
Carabus (Morphocarabus) chaudoiri beladici Obydov, 2001
Male adult
S-Tuva, East Tannu-Ola,Biche-Sailig r.h=2100m, 15.VII.1996, D.E.Lomakin leg.
Copyright © Anton Menstschikov
Carabus (Morphocarabus) chaudoiri beladici Obydov, 2001 Paratype
Female adult
Rusia, Tuva, E Tannu-Ola Mt.rng., near Kara-Hol lake, 29-31.V.2000, M. Beladzhik leg.
Copyright © Dmitri Obydov
Carabus (Morphocarabus) aeruginosus zyrjanovskianus Shilenkov & O.Berlov, 1996
Male+Female adult
NE KAZAKHSTAN, S-Altays, Ulbinsky Mt. h=900m., Bobrovka vill., 24.VI.01, A. Klimenko leg. female 25 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) aeruginosus verzhutzkii O.Berlov & Shilenkov, 1996
Male adult
Carabus (Morphocarabus) aeruginosus verzhutzkii O.Berlov & Shilenkov, 1996
C Altays, Katun r., 5 km. N Katanda vill., 21.06.2002, A. Menstschicov leg.
Copyright © Anton Menstschikov
Carabus (Morphocarabus) aeruginosus verzhutzkii O.Berlov & Shilenkov, 1996
Male adult
C Altai Mts., Seminskii Mt.rng., Katun riv., 3 km S Ust'-Sema riv.,
4.VII.2002 A. & N. Menschikovy leg.
det. Mentschikov A.
photo Anichtchenko A.
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) aeruginosus salechardensis Obydov, 1999 Paratype
Male adult
Rusia, Selehard, tundra, 30.VIII.1953, I. Telyshev leg
Copyright © Dmitri Obydov
Carabus (Morphocarabus) aeruginosus hermani Mannerheim, 1827 loc.typ.
Male adult
C Rusia, S Ural Mts., Zlatoust, 22-28.VI.1952, S. Nikulina leg.
Copyright © Dmitri Obydov
Carabus (Morphocarabus) aeruginosus eschscholtzi Mannerheim, 1827
Male adult
Russia, W Altai, Zmeinogorsk distr., near Cherepanovskii vill., 2-16.6.1990, D. Obydov leg.
Copyright © Dmitri Obydov
Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
Male adult
SE Siberia, E Sayan, Nizhneudinsk, h=500m, 21-26.VI.2006,
male, 26mm
Copyright © Ruslan Panin
Carabus (Rhigoidocarabus) zhubajie laojunensis Kleinfeld, 1999 Holotype
Male adult
Copyright © Frank Kleinfeld
Carabus (Morphocarabus) rothi rothi Dejean, 1829
Male+Female adult
Romania
Banat, Nemeseti h=250m. VI.1989 Lie P. leg.
photo Solodovnikov I.A.
Copyright © Igor Solodovnikov
Carabus (Morphocarabus) praecellens trans illigeri Dejean, 1826
Male+Female adult
W-Croatia, Karlovac, near Bosiljevo vill., h~300m, V.2000, I.Rapuzzi leg. female, 33 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) spasskianus aeruginosiformis Breuning, 1932
Male adult
Russia, Khakassia,
W-Sayan, Alan Mt range, valley Enisey riv., 5 km S Mayna vill., 15.VII.2000, A.& N. Menstschicovy leg.
L=24 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
Male+Female adult
Russia, S Siberia, W Sayan, Ergaki Mt range, Nizhnyaya Buyba river h=1700 m, 19.VII.1999, A.& N. Menstschikovy leg.
Copyright © Ruslan Panin
Carabus (Morphocarabus) spasskianus shoriensis Obydov, 1999
Male adult
Russia, Khakassia, Abakanskii Mt Range, Askiz river vall., 7 km E Kazanovka vill., 15.VI.2006, A.& N. Menstschikovy leg. L=24 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997
Male+Female adult
Russia,Tuva, Akademika Obrucheva Mt range, 15 km N Cherbi vill., Chailang ur. 7.VII.1999, A & N. Menstschikovy leg. L=23 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi hampei Kuster, 1846 (transition to rothi incompsus Kraatz, 1880)
Male adult
NW Romania , Bistrita Nasaud, Mts. Tibles, 6 km SE Bichigiu, 17.VI.2009, S.Budka leg.
L=35 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi hampei Kuster, 1846 (as dacicus Csiki, 1906)
Female adult
NW Romania , Bistrita Nasaud, Beclean env., 17.VI.2009, S.Budka leg.
female, 31 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) aeruginosus pitschugini Obydov, 2011 Holotype
Male adult
East Siberia, Tunkinskie Goltzy (East Sajans), Mondy env., 24.VI.1979, E. Ya. Berlov leg.
Copyright © Dmitri Obydov
Carabus (Morphocarabus) aeruginosus pitschugini Obydov, 2011 Paratype
Female adult
East Siberia, Tunkinskie Goltzy (East Sajans), Mondy env., 24.VI.1979, E. Ya. Berlov leg.
Copyright © Dmitri Obydov
Carabus (Morphocarabus) helleri Ganglbauer, 1893 (as pseudopreyssleri Breuning, 1932)
Male adult
Czech Rep., NE Moravia, Moravian-Silesian Beskids Mt Rng, PR Trojacka, Hodslavice vill., 29.V.12.VI.2010, Krause Rene leg./det.Panin R. L=30 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) kollari Palliardi, 1825 (as semetricus Kraatz, 1878)
Male adult
SW-Romania:Banat, Bocsa/Coltan, 21.V-5.VI.2010, Krause Rene leg.
L=29 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) karpinskii Kryzhanovskij & Matveev, 1993
Male adult
Copyright © Kirill Makarov
Carabus (Morphocarabus) spasskianus geblerianus Obydov, 1999
Male adult
Russia, NW Altay, Barnaul env.,23.07.1998, D.Lomakin leg.
Copyright © Ruslan Panin
Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859
Male adult
Russia, SE Siberia, Transbaikalia, N Chita reg., Kodar Mt. Range, Big Leprindo Lake env.h~1800m., 1.VII.2006, S. Rybalkin leg. L=20 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi hampei (as ormayi Reitter, 1896) loc.typ.
Male adult
W. UKRAINE,
Transcarpathian reg., near Beregovo, 27.IV-22.V..2011,, Panin R. & Chyzhevsky O. leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) henningi melyachi Shilenkov & Menstshikov, 1996 loc.typ.
Male adult
Russia, Altay, Katun Mt Range, Karakol lake,22.VI.1993, Melyach F.V. leg.<br /> L=18 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) chaudoiri marusiki Obydov, 1997 Holotype
Male adult
SE-Tuva: Sangilen Mts.: Kargy River valley: 50:31N/97:01E. image from zin.ru
Copyright © Dmitri Obydov
Carabus (Morphocarabus) excellens excellens (f.i. frivaldszkyi Kraatz, 1887)
Female adult
W-UKRAINE, NW Podolian Upland,<br /> Lviv Reg., Zolochiv Distr., Lysa Gora steppe massif, 14-30.V.2008, Panin. R. leg./det.<br /> L=26 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens excellens Fabricius, 1798
Male adult
NC Ukraine, Kiev, Lysa Gora, on the fringes forest, 5-24.VI.2007, Gerasimov R.P.. leg.<br /> L=25 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens excellens Fabricius, 1798
Male adult
C- UKRAINE, NW Odessa reg., Kodyma distr., near Aleksandrivka vill.,18.V.2003, Demidov G.N. leg.<br /> L=26 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) praecellens trans scheidleri
Male adult
NC Croatia, Ivanščica Mts , Ivanec,4-18.VI.2011, Krause Rene leg.<br /> L=31 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) praecellens praecellens Pillardi, 1825
Male adult
Serbia ,
Frushka Gora, Ruma VI.1985
Copyright © Igor Solodovnikov
Carabus (Morphocarabus) rothi hampei Kuster, 1846
Male adult
C Romania, Târgu Mureş, Ernei, 46°35′471″N 24°37’351″E, h=300 m,20.VIII.2004, Sl.Budka leg.
L=30 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi hampei (as zilahiensis Csiki, 1906)
Male+Female adult
NW Romania, Zalău env., 31.VII.2003, Sl.Budka leg.
L=24 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi comptus (as szoerenyensis Csiki, 1908)
Female adult
SW Romania, Tarcu Mts, vf. Matania Mt. h=2000 m, 12.VI.2008, S.Budka leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi hampei (as hunyadensis Csiki, 1926)
Male adult
CW Romania, Apuseni Mts, Muntii Metaliferi, Hunedoara reg., near Sacaramb vill., 6.VI.2011, S.Budka leg.
L=37 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878 (as viridipennis Kraatz, 1886, nec Lewis,18
Male+Female adult
Russia, Far East,Primorsky reg., S. Sikhote-Alin Mts. R., Kavalerovo vill. env.,
5-25.VII.1992, С. Kunitzyn leg.
male - 25 mm, female - 25 mm.
Copyright © Alexandr Safronov
Carabus (Morphocarabus) hummeli pusongensis Imura, 1993
Male+Female adult
Russia, Far East, Primorsky ter., Khasansky distr., Zanadvorovka vill.,
Gusevsky pit, S.Veriga leg.
male - 13.VII.2008, female 19-21.VI.2007
male - 28 мм, female - 28 мм.
Copyright © Alexandr Safronov
Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
Male+Female adult
Russia, Far East, Primorsky ter., Chuguevsky distr., 30 km ESE from Chuguevka, 9-14.VIII.2007 S. Shabalin leg. male - 25 mm, female - 26 mm.
Copyright ©
Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878 (as purpuripennis Kraatz, 1886)
Male adult
Russia, Khabarovsk reg., near Komsomolsk-on-Amur.
7-21.VI.2010, A. Kapkaev leg.
Male left - 24 mm, male right - 23 mm.
Copyright © Alexandr Safronov
Carabus (Morphocarabus) hummeli hummeli Fischer, 1823
Male+Female adult
Russia, Far East, Khabarovsk reg., Obluchye env.
M. Khingan Mts. R.
10-20.VII.1992 S. Andreev leg.
male - 26 mm, female - 27 mm.
Copyright © Alexandr Safronov
Carabus (Morphocarabus) regalis tschusovae Obydov, 2004
Male+Female adult
NE-Kazakhstan, W Altay, Ivanovsky Mt.Rng, near Ridder (Leninogorsk), h-1300-1500m, 4-6.VII.2003
female, 25 mm
det.Panin R.
Copyright © Ruslan Panin
Carabus (Morphocarabus) regalis as cyanescens Kraatz, 1882, nec Sturm, 1815
Male+Female adult
Kazakhstan, E. Kazakhstan area, Zyryanovsk env., near Putintzevo vill. 10-VI-2005 M. Danilevsky leg. male - 24 mm, female - 25 mm.
Copyright ©
Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
Male+Female adult
SW Siberia, Kemerovo reg., Krapivino distr.,<br /> 8 km SSW Saltymakovo vill., h=150 m.<br /> 1-20.VII.2010 D. Efimov leg.<br /> male - 25 mm, female - 26 mm.
Copyright ©
Carabus (Morphocarabus) spasskianus aeruginosiformis Breuning, 1932
Male+Female adult
Russia, S. Siberia, Krasnoyarsk reg., W. Sayan,<br /> Sayanogorsk distr., Cheremushki vill. env.,<br /> 07-VII-1994 М. Danilevsky leg.<br /> male - 22 mm, female - 25 mm.
Copyright ©
Carabus (Morphocarabus) helleri Ganglbauer, 1893
Male+Female adult
CZECH, SE Moravia, Velka nad Velickou. 20-V-2005, V. Slovak leg. male - 26 mm, female - 30 mm.
Copyright © Alexandr Safronov
Carabus (Morphocarabus) rothi hampei as telekii Csiki, 1937 (? hampei )
Female adult
CW Romania, Arad distr., Zarandulu Mt Range, near Ilteu vill., h=276 m, 46°66′487″N 22°20’455″E, 25.VI.2005, Sl.Budka leg.
L=30 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi rothi Dejean, 1829
Male+Female adult
C Romania, Sibiu Area, right bank Olt River, Bradu vill., h=380 m, 19.VIII.2001, S.Budka leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) venustus venustus Morawitz, 1862
Male adult
Russia, Far East, Primorskiy reg., Ussuriysk Distr., Kaymanovka vill., 6-15.VIII.2011, <br /> A. Korshunov leg.<br /> L= 21.5 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) praecellens praecellens Pillardi, 1825
Male adult
Serbia, Fruska Gora Mts, 20-31.V.2006, M.Sima leg.
L=29 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) illigeri versicolor Frivaldsky, 1835
Male+Female adult
C Bulgaria:Stara planina Mts, Gabrovo,12-15.VI.2007, M. Janalik leg. female, 31 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi comptus (as szoerenyensis Csiki, 1908)
Male adult
SW Romania, Tarcu Mts, vf. Nedeia h=2000m, 12.VI.2008, J.Kynkor leg.
L=22 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi hampei (as gutiiensis Takács & Lie, 1992)
Male adult
N Romania, E Carpathian, Maramureş Reg., Gutâi Mts, cabana Mogosa,4 km N Baia Sprie, 9-17.VI.2010, J.Kynkor leg.
L=33 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) wulffiusi dekraatzi Kraatz, 1881
Male adult
NE-China, Heilongjiang, Wuchang city, Mt Zheng-nan-gou, 1-11.VI.2011<br /> L=22 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi incompsus Kraatz, 1880
Male+Female adult
C Romania, Brașov County, Cristian vill., h=380 m,V-VI.2008, M.Pasek leg.
female, 27 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi (? incompsus as spectabilis Csiki, 1906)
Female adult
NC Romania, Bistrita Nasaud reg., Călimani Mts, Colibita vill., mixed forest, meadows h=800m, 15-21.V.2011, M.Pasek leg.<br /> L=34 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi incompsus (as mendax Csiki, 1906)
Female adult
W Ukraine, NE Carpathians,
Transcarpathian reg., Chornogora Mt Range, h=1100m,13.VI.2011, Panin R. leg./det.
L=33 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi alutensis Savulescu, 1972
Male+Female adult
C Romania, S Carpathians, Rimnicu Vilcea env., Calimaneşti, Cozia Mts., VII.2007, I. Marvan leg.
L=29 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi comptus (as borlovensis Lie, 1994)
Male adult
SW Romania, S Carpathians,
Banat , Tarcu Mts, Borlova env., 19-31.V.2005, P. Sima leg.
L=24 мм
Copyright © Ruslan Panin
Carabus (Morphocarabus) viridilimbatus panini I.Solodovnikov 2017 Paratype
Male adult
Russia, SE Yakutia, Neryungri distr., Tokinski Stanovik Mt Rng., Beder riv., (right tributary rivers Algama), 26.VI.1996, Krasnov M. leg. L=15 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) monilis Fabricius, 1792
Male adult
24 mm, England, Warwickshire
leg.det. D.J.Mann, 15.IV.1993
Photo: U.Schmidt, 2011
Copyright © Udo Schmidt
Carabus (Morphocarabus) rothi hampei (as diffinis Csiki, 1906)
Male adult
NW Romania, Apuseni Mts, Trascău Mt Rng, Petresti de Jos, 12 km E Turda,. 6.VIII.2008
L=27 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens (as lomnitzkii Reitter, 1896)
Female adult
W Ukraine, NW Podolian Upland, Lviv reg., Zolochiv distr., Steppe Reserve Lysa Gora, 19.V-7.VI.2007, Panin R. leg.<br /> L=26 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) illigeri versicolor Frivaldsky, 1835 (ab.)
Female adult
C Bulgaria: Stara Planina Mts, Schipka pass h=1450m, 12-15.VI.2007, M.Janalik leg. L=31 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) illigeri illigeri (as tarensis Toševski, 1990)
Male adult
WC-Serbia: National park Tara Mts.:NE Mitrovac h=1500m, 1-12.VII.2007, Pr.Sima leg. L=30 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi hampei (as hunyadensis Csiki, 1926)
Male adult
CW Romania, Apuseni Mts, Metalliferous Mt Rng, Hunedoara reg., near Săcărâmb vill., 45°97'472"N/23°02'630"E, h=680 m, mixed forest, meadows, 15-20.V.2011, B.Brezina leg.
L=36 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens excellens ( as jasilkowskii Born,1905)
Male adult
NE Romania, 5 km W Suceava, 47°63′940″N 26°18′795″E,h=340m, agricultural field, 15-20.V.2011, B. Brezina leg. L=24 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
Male adult
W-UKRAINE, Transcarpathian region, Rakhiv distr., Chornogora Mt Range, h=800 m, near Bogdan vill., 10.VI.2012, Panin R. leg.
L= 33 мм
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
Male adult
W Ukraine,Transcarpathian Reg., Rakhiv distr., Marmarosh Mts., near Lug (Lunca la Tisa) vill., h=400 m,12-15.VII.2011
L=32 mm
det.Panin R.
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
Female adult
W Ukraine, NE Carpathians,
Transcarpathian Reg., Marmarosh Mts., Rakhiv distr., 5 km W Dilowe vill., Site Kuziy, 5-15.VII.2010, Demidov G.N. leg./det.Panin R.
L= 35 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
Male adult
Ukrainian Carpathians,
Transcarpathian reg., Rakhov distr., Svidovec Mt Rng., Mt. Podpula h=1500-1600m. Lazutkin G. leg, 13-14.VII.1978, det. Panin R.
L=28mm.
Copyright © Ruslan Panin
Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878 (as vladobydovi Obydov, 2007) Paratype
Female adult
Russia, South Primorie, Popova isl., 12-14.07.1990, A. Aljakrinskii leg.
Copyright © Dmitri Obydov
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906 loc.typ.
Male+Female adult
W-UKRAINE,<br /> Transcarpathian region, Uzhgorod distr., h=250 m, 8.VII-25.VIII.2011<br /> Panin R. leg./det.<br /> female, 33 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906 loc.typ.
Male+Female adult
W-UKRAINE,<br /> Transcarpathian region, Uzhgorod distr., 28.IV-23.V.2011<br /> Panin R. leg./det.<br /> female, 33 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Male adult
SE Slovakia, Slanske vrchy Mts, Herlianske sedlo, 8-19.VI.2009, Krause Rene leg.
L=30 mm
det.Panin R.
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Female adult
NW Ukrainian Carpathians,
Lviv Region, Staryi Sambor Distr., near Dobromyl , h=450m, 9.VI-25.VII.07, Panin R. leg./det.Solodovnikov I. & Panin R.
female, 35mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Female adult
NW Ukrainian Carpathians
Lviv Region, Staryi Sambor Distr., near Dobromyl , h=500m 9.VI-25.VII.2007, Panin R. leg./det.Solodovnikov I. & Panin R.
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii zawadzkii, Kraatz 1854
Male adult
W.Ukraine, Ivano-Frankivsk reg.,
Yaremche distr., near Tatariv vill., Gorgany Mts. h=700 m, 16.VI.2011, Panin R. leg.
L=31 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii zawadzkii, Kraatz 1854
Male+Female adult
W.UKRAINE, Ukrainian Carpathians
Lviv Reg., Skole distr., Skolivski Beskidy Mts. Zelemyanka Mt. h=1000m. 9-20.VII.07 Panin R. leg.
female, 35mm.
Copyright © Ruslan Panin
Carabus (Morphocarabus) tarbagataicus birlikensis Kabak, 2013 Holotype
Male adult
China, Xinjiang, cresp of Birliktau Mt.R., WSW Toli, 45 49 23 N / 83 06 13 E, H=2565 m, 25.07.2012, Kabak I.I. leg. (ZIN)
Copyright © Ilya Kabak
Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Male adult
NE Serbia, Majdanpek; male, 33mm
Copyright © Aleš Sedláček
Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Female adult
NE Serbia, Kučevo; female, 33mm
Copyright © Aleš Sedláček
Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Female adult
NE Serbia, Majdanpek; 35mm
Copyright © Aleš Sedláček
Carabus (Morphocarabus) helleri Ganglbauer, 1893
Male adult
CZ; CE Moravia, Bílé Karpaty Mts.; 24mm
Copyright © Aleš Sedláček
Carabus (Morphocarabus) helleri Ganglbauer, 1893
Male adult
CZ; CE Moravia, Bílé Karpaty Mts.; 25mm
Copyright © Aleš Sedláček
Carabus (Morphocarabus) praecellens jucundus Csiki, 1906
Male adult
N Hungary, Pilis Mts., Piliszentlelek, 20-27.VII.2011, Krause Rene leg./det. R.Panin
L=30 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as pseudojucundus Retezár, 1974)
Male adult
NW-Hungary::Komarom distr., Tardos, Gerecse Mts., 20.-27.V.2011, Krause Rene leg.
L=26 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as vertesensis Retezár, 1974)
Male adult
NW Hungary: Fejer: Vértes Mt Rng, Tatabanya, 28.VII-2.VIII.1996
L=29 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens excellens (as moldaviensis Born, 1903)
Female adult
E-Romania, Vaslui reg.,15 km SE Vaslui, Crasna env., V-VI,2013, M,Pasek leg. L=26 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi rothi (as equistriatus Kraatz, 1878)
Female adult
C Romania, Sibiu reg., left bank Olt River, Avrig env., 12.VI.2013, V. Slovak leg.
L=30 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) odoratus magadanicus Obydov 1999: 614
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Carabus (Morphocarabus) odoratus magadanicus Obydov 1999: 614
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Carabus (Morphocarabus) brinevi Kabak, 2014 Paratype
Male adult
China, Xinjiang, S Altai Mts, Keshtau Mt., ENE Qinghe, 46°47'21''N / 90°44'42''E, 2730 m, 12.08.2013, I.I. Kabak leg.
Copyright © Ilya Kabak
Carabus (Morphocarabus) brinevi Kabak, 2014 Holotype
Male adult
China, Xinjiang, S Altai Mts, Keshtau Mt., ENE Qinghe, 46°47'21''N / 90°44'42''E, 2730 m, 12.08.2013, I.I. Kabak leg.
Copyright © Ilya Kabak
Carabus (Morphocarabus) brinevi Kabak, 2014 Paratype
Female adult
China, Xinjiang, S Altai Mts., Keshtau Mt., ENE of Qinghe, h~2730 49°47'21"N 90°44'42"E 12.VIII.2013 leg. I.Kabak L=19.5 mm
Copyright © Kirill Makarov
Carabus (Morphocarabus) chaudoiri mongolorum Csiki, 1927 Lectotype
Male adult
Muséum national d’Histoire naturelle, Paris (France); Collection : Insects: Coleoptera (EC); Specimen MNHN-EC- EC522; Credit: Antoine Mantilleri - 2010 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec522
Copyright ©
Carabus (Morphocarabus) odoratus chaffanjoni Lesne, 1898 Holotype
Male adult
"Bouloun, Env d'Jakoutsk, J. Chaffanjon" Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC157 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec157
Copyright © MNHN
Carabus (Morphocarabus) tarbagataicus var.motschulskyi Lapouge, 1909 Holotype
Male adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC525 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec525
Copyright © - - -
Carabus (Morphocarabus) henningi sahlbergi Dejean, 1826 Lectotype
Male adult
"Siberie, Slataoust" = [Златоуст, Zlatoust, Cheliabinsk prov.]. Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC1 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec1
Copyright © - - -
Carabus (Morphocarabus) odoratus melleus Lapouge, 1909 Paralectotypes
Male adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC521
Copyright © MNHN
Carabus (Morphocarabus) odoratus melleus Lapouge, 1909 Lectotype
Female adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC520
Copyright © MNHN
Carabus (Morphocarabus) hummeli stolidus Lapouge, 1924 Lectotype
Female adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC519
Copyright ©
Carabus (Morphocarabus) excellens paradoxus Lapouge, 1906 Lectotype
Female adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC539
Copyright © - - -
Carabus (Morphocarabus) odoratus dohrni Gebler, 1847: 300 Lectotype
Female adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC413
Copyright © MNHN
Carabus (Morphocarabus) mestscherjakovi zherichini Shilenkov, 1990 Paratype
Female adult
RUSSIA, Krasnoyarsk Terr., SW Tajmyr Pen., Nizhnyaya Agapa Riv., 40 km of source, south tundra, plakor, brooks holmme, on sandy soil, 7.VIII.1973 (V.V. Zherikhin).
Copyright © Roman Dudko
Carabus (Morphocarabus) odoratus baeri Menetries, 1851: 45
Male adult
Russia, SW Taymyr, near Talnakh, 25 km N Norilsk, hilly tundra, hillside overgrown with alder slate, dwarfish birches, blueberry, h = 150-250 м, 24-28.06.2003, leg. S.Saluk
Copyright © Igor Solodovnikov
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Male adult
NE-Hungary, 4 km S Sátoraljaújhely, right bank riv. Bodrog (Long-erdő), 8.VI.1992, leg./det. I.Retezár leg. L=28 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi distinguendus Csiki, 1906
Male adult
NC-Hungary: E- part of Budapest, Cinkota,28.VIII-17.IX.1980, I.Retezár leg. L=27 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi distinguendus (as bieneri Mandl, 1965)
Male adult
C-Slovakia, Detva Distr., Stara Huta env. h=700-800m, 3.VI.-5.VII.2012, V.Slovák lgt. L=29 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi distinguendus (as subparvulus Mandl, 1965)
Male+Female adult
NC-Hungary:Borzsony Mts, Nograd-Diosjeno,1-14.V.2004, Krause Rene leg. female, 32 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859
Female adult
Russia, SE Siberia, N Chita reg., Kodar Mt. Range, Big Leprindo Lake env., h~1800 m., L=21 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) odoratus kurmaevi I.Solodovnikov, 2017 Holotype
Male adult
Russia, Siberia, Krasnoyarsk Krai, the North Yenisei district, 55 km of CE of the vill. Bryanka, valley of the river Chirimba, stream Nakhodny, h = 350 m, the mixed forest (birch and pine); on a steep slope along an animal track on a hill brow, 23.VII-20.VIII.2015, leg. A.V. Kurmaev
Copyright © Igor Solodovnikov
Carabus (Morphocarabus) odoratus kurmaevi I.Solodovnikov, 2017 Paratype
Male adult
Russia, Siberia, Krasnoyarsk Krai, the North Yenisei district, 55 km of CE of the vill. Bryanka, valley of the river Chirimba, stream Nakhodny, h = 350 m, the mixed forest (birch and pine); on a steep slope along an animal track on a hill brow, 23.VII-20.VIII.2015, leg. A.V. Kurmaev
Copyright © Igor Solodovnikov
Carabus (Morphocarabus) odoratus kurmaevi I.Solodovnikov, 2017 Paratype
Female adult
Russia, Siberia, Krasnoyarsk Krai, the North Yenisei district, 55 km of CE of the vill. Bryanka, valley of the river Chirimba, stream Nakhodny, h = 350 m, the mixed forest (birch and pine); on a steep slope along an animal track on a hill brow, 23.VII-20.VIII.2015, leg. A.V. Kurmaev
Copyright © Igor Solodovnikov
Carabus (Morphocarabus) odoratus kurmaevi I.Solodovnikov, 2017 Paratype
Female adult
Russia, Siberia, Krasnoyarsk Krai, the North Yenisei district, 55 km of CE of the vill. Bryanka, valley of the river Chirimba, stream Nakhodny, h = 350 m, the mixed forest (birch and pine); on a steep slope along an animal track on a hill brow, 23.VII-20.VIII.2015, leg. A.V. Kurmaev
Copyright © Igor Solodovnikov
Carabus (Morphocarabus) viridilimbatus panini I.Solodovnikov 2017 Holotype
Male adult
Russia, SE Yakutia, Neryungri distr., Tokinski Stanovik Mt Rng., Beder riv., (right tributary rivers Algama), 26.VI.1996, Krasnov M. leg., 17 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) viridilimbatus panini I.Solodovnikov 2017 Paratype
Female adult
Russia, SE Yakutia, Neryungri distr., Tokinski Stanovik Mt Rng., Beder riv., (right tributary rivers Algama), taiga, h = 940 m, 55°55'27.78" N, 129°42'6.20" E, 26.VI.1996, leg. Krasnov M. 21,1 mm
Copyright © Igor Solodovnikov
Carabus (Morphocarabus) viridilimbatus saluki I.Solodovnikov 2017 Holotype
Male adult
Russia, SW Taymyr, near Talnakh, 25 km N Norilsk, hilly tundra, hillside overgrown with alder slate, dwarfish birches, blueberry, h = 150-250 м, 24-28.06.2003, leg. S.Saluk 20,5 mm
Copyright © Igor Solodovnikov
Carabus (Morphocarabus) viridilimbatus saluki I.Solodovnikov 2017 Paratype
Male adult
Russia, SW Taymyr, near Talnakh, 25 km N Norilsk, hilly tundra, hillside overgrown with alder slate, dwarfish birches, blueberry, h = 150-250 м, 24-28.06.2003, leg. S.Saluk
Copyright © Igor Solodovnikov
Carabus (Morphocarabus) viridilimbatus saluki I.Solodovnikov 2017 Paratype
Female adult
Russia, SW Taymyr, near Talnakh, 25 km N Norilsk, hilly tundra, hillside overgrown with alder slate, dwarfish birches, blueberry, h = 150-250 м, 24-28.06.2003, leg. S.Saluk
Copyright © Igor Solodovnikov
Carabus (Morphocarabus) viridilimbatus saluki I.Solodovnikov 2017 Paratype
Female adult
Russia, SW Taymyr, near Talnakh, 25 km N Norilsk, hilly tundra, hillside overgrown with alder slate, dwarfish birches, blueberry, h = 150-250 м, 24-28.06.2003, leg. S.Saluk
Copyright © Igor Solodovnikov
Carabus (Morphocarabus) monilis Fabricius, 1792
Female adult
France, Dröme, Ft. de Saou, 17.vii.1975-M. Grotz Leg.-Coll.RBINS, N. Mal as monilis saouensis nicodi
Copyright © Noël Mal
Carabus (Morphocarabus) wulffiusi liaoningensis Deuve & Imura, 1990
Female adult
NE-China:E-Liaoning: NP Kuandian Tianqiaogou , VI.2010, L=24
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
Female adult
NE-Hungary. (photo by Imre Retezár)
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
Male adult
NE-Hungary. (photo by Imre Retezár)
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Male adult
W Ukraine, Transcarpathian, Mts E of Uzgorod, Veliki Lazy h~270-300 m, 8.IV-23.V.2011, Panin R. leg. L=28 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) tarbagataicus altaicola Deuve & Tian, 2019 Holotype
Female adult
Chine, Xinjiang, Préfecture d’Altaï, Monts Altaï, au nord de la ville d’Altaï, Parc forestier de Xiaodonggou (Chen Liusheng, 8.VIII. 2014), in coll. South China Agricultural University, Canton.
Copyright © Thierry Deuve
Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
Female adult
W Ukraine, Transcarpathian lowland, near Chop, 11.VI-19.VII.2019, Panin R leg. L=29 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
Male adult
W Ukraine, Transcarpathian lowland, near Chop, 11.VI-19.VII.2019, Panin R leg. L=30 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
Female adult
W Ukraine, Transcarpathian lowland, near Chop, 11.VI-19.VII.2019, Panin R leg. L=31 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
Female adult
W Ukraine, Transcarpathian lowland, near Chop, 11.VI-19.VII.2019, Panin R leg. L=29 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
Male adult
W Ukraine, Transcarpathian lowland, near Chop, 11.VI-19.VII.2019, Panin R leg. L=28 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
Female adult
W Ukraine, Transcarpathian lowland, near Chop, 11.VI-19.VII.2019, Panin R leg. L=30 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Male adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 4-24.V.2010, Panin R. leg. L= 29 mm.
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens ciscarpathicusn (as rareulensis Born, 1907)
Male adult
NE-Romania, Rarau Mts. h-1300m, VI.2014, J. Kynkor leg. L=23 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens ciscarpathicus (as polonicus Lomnicki, 1886)
Male adult
SW Poland: Przemyśl env. photo by Mieczyslaw Stachowiak
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens ciscarpathicus (as polonicus Lomnicki, 1886)
Female adult
W Ukraine, Lviv Region, Zhydahciv distr., Borodchytsi vill.,the right bank of Dniester river, 11-16.VII.2012, Panin. R leg. L=27 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Holotype
Male adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 28.V-9.VI.2013, Panin R. leg. L=29 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Female adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 28.V-9.VI.2013, Panin R. leg. L= 28 mm.
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Male adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 18.V-12.VI.2014, L=28 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Male adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 27.IV-13.VI.2018, Panin R. leg. L= 28 mm.
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Male adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 27.IV-13.VI.2018, Panin R. leg. L= 29 mm.
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Female adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 20.V-25.VI.2008, Panin R. leg. L= 31 mm.
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Female adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 20.V-25.VI.2008, Panin R. leg. L= 30 mm.
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Male adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 12.V-2.VI.2009, Panin R. leg. L= 29 mm.
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Female adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 27.IV-13.VI.2018, Panin R. leg. L= 31 mm.
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Male adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., Panin R. leg. L= 28 mm.
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens excellens (ab.goldeggi Duftschmid, 1812)
Male adult
W-Ukraine, Podolian Tovtry, Ternopol reg., Pidvolochysk distr., Shevchenkove vill., steppe sites, 6.V-7.VI.2012, Panin R. leg. L=24 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) henningi hamarensis Shilenkov, 1996
Female adult
Lake Baikal, Maritui vill. 25.7.1994, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) henningi hamarensis Shilenkov, 1996
Female adult
Lake Baikal, Maritui vill. 25.7.1994, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
Female adult
W Ukraine, Transcarpathian, Marmarosh Mts., near Lug vill., h=400 m, 28.IV -26.VII.2017, Panin R. leg., L= 38 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
Male adult
Russia, S Tuva, Khorumnug-Taiga Mts, Buren riv. Valley, Buren-Bay-Khaak vill. h=900 m, 17-24.VI.2002, Vashchenko A.& B.leg.,/ det. Panin R. L=25 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) henningi cf. saldaitisi Obydov 1997
Female adult
Russia, S Tuva, E Tanu-Ola Mts., Cha-Ova-Art Pass, h=1700m, 28.VI-3.VII.2000, Vashchenko S. leg. L=20 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) henningi cf saldaitisi Obydov 1997
Male adult
Russia, S Tuva, E Tanu-Ola Mts., Cha-Ova-Art Pass, h=1700m, 28.VI-3.VII.2000, Vashchenko S. leg. L=19.5 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) henningi pelevini Obydov 2006
Female adult
E Kazakhstan, S Altay, Sarym-Sakty Mt Range, 12 km SE Ornek vill. h=1200m, 10.VII.1998, L=21 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) henningi pelevini Obydov 2006a: 99
Male adult
E Kazakhstan, S Altay, Sarym-Sakty Mt Range, 12 km SE Ornek vill. h=1200m, 10.VII.1998, L=20 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) henningi henningi Fischer, 1817
Male adult
Russia, S Siberia, NW Buryatia, E Sayans Mts., Tunka Range, near Mondy vill., h=2400 m, alpine tundra, 1-15.VI.2001, Mishustin R. leg./det. R.Panin
Copyright © Ruslan Panin
Carabus (Morphocarabus) hummeli putyatini Rapuzzi, 2012 Holotype
Male adult
Russia, South Pimorskiy region, Putyatin Island, 50-200 m, 14/22.VIII.2008, A. Plutenko leg. (in coll Rapuzzi). Image from Rapuzzi, 2012
Copyright © - - -
Carabus (Morphocarabus) brinevi altaicola Deuve & Tian, 2019
Female adult
«qing tong se», «28#» (ICXU)
Copyright © Ilya Kabak
Carabus (Morphocarabus) brinevi kanasensis Kabak, 2021 Holotype Paratype
Male adult
China, Xinjiang, Altai Mts, left bank of Kanas lake, 48°44ʹ35ʺN / 87°06ʹ03ʺE, H = 2710 m, alpine meadows, 15.07.2019 I.I. Kabak leg. (ZIN)
Copyright © Ilya Kabak
Carabus (Morphocarabus) excellens excellens (as lomnitzkii Reitter, 1896)
Male adult
W Ukraine, NW Podolian Upland, Lviv reg., Zolochiv distr., Lysa Gora steppe massif, 11.VII-14.VIII. 2021, Panin R. & D. Khrapov leg., L=25 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens excellens (as lomnitzkii Reitter, 1896)
Male adult
W Ukraine, NW Podolian Upland, Lviv reg., Zolochiv distr., Lysa Gora steppe massif, 11.VII-14.VIII. 2021, Panin R. & D. Khrapov leg., L=25 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens excellens (as lomnitzkii Reitter, 1896)
Female adult
W Ukraina, Podolian Upland, steppe reserve Kasova Gora near Burshtyn vill., 15.VI -28.VII.2020, R. Panin & D. Khrapov leg., L=27 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) excellens excellens (as lomnitzkii Reitter, 1896)
Male adult
W Ukraina, Podolian Upland, steppe reserve Kasova Gora near Burshtyn vill., 15.VI -28.VII.2020, R. Panin & D. Khrapov leg., L=24 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii zawadzkii, Kraatz 1854
Male adult
W Ukraine, NE Carpathians, Lviv region, Skole Beskids Mts, Zelemyanka Mt. Range h=915 m, 22.VI-17.VII.2022, Panin R. & Kanarsky Yu. leg., L= 32 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) chaudoiri juliae Obydov, 2002 Holotype Paratype
Male+Female adult
Russia, Tuva Republic, Mungun-Taiga, Khindiktig-Khol Lake, 2400 m, 22.-29.VI.2000, M. Beladič leg.
Copyright © - - -
Carabus (Morphocarabus) mestscherjakovi tuvinicus Obydov, 2002 Holotype
Female adult
Россия, Тува, хр. Таскыл, долина р. Уюк, 4.VI-8VI.1999, И. Белло leg.
Copyright © Dmitri Obydov
Carabus (Morphocarabus) odoratus chlebnikovae Obydov 2005 Paratype
Female adult
Россия, Северный Байкал, 80 км Ю Северобайкальска, 5 км Ю истока р. Куркула, 19.VII.2003, М. Хлебникова leg.
Copyright © Dmitri Obydov
Carabus (Morphocarabus) odoratus putoranicus Obydov, 2002 Paratype
Male adult
Россия, Север Вост. Сибири, Плато Путорана, окр. оз. Аян, 26.VI-10.VIII.1978, M. Сладков leg.
Copyright © Dmitri Obydov
Carabus (Morphocarabus) odoratus taskylensis Obydov, 2001 Holotype
Male adult
Россия, Тува, хр. Таскыл, р. Уюк, 4.VI-8.VI.1999, И. Белло и М. Беладжик leg.
Copyright © Dmitri Obydov
Carabus (Morphocarabus) odoratus jacuticus Obydov, 2002 Paratype
Male adult
Россия, Якутия, В. Хандыга, VII.1993, А. Наглис leg.
Copyright © Dmitri Obydov
Carabus (Morphocarabus) hummeli jurgae Obydov, 1995 Holotype
Male adult
Россия, Якутия, 300 км севернее хр. Сунта-Хаята, VII.1993, Н. Шивис leg.
Copyright © Dmitri Obydov
Carabus (Morphocarabus) chaudoiri chaudoiri Gebler, 1847 Lectotype
Female adult
Altay
Copyright © Dmitri Obydov
Carabus (Morphocarabus) chaudoiri shilenkovi O.Berlov, 1989
Male adult
Россия, Якутия, Оймяконское нагорье, 12-18.VII.1998
Copyright © Dmitri Obydov
Carabus (Morphocarabus) spasskianus cf putus Motschulsky, 1846
Female adult
Russia, SW Siberia, C-Altays Mts, Kuray Mt Rng NE Aktash vill., h=2100-3000 m, 8-13.VII.2022, E.Filippov leg./det.R Panin, L= 22 mm (alpine petrophilic population close to the form schestopalovi Plutenko, 1995)
Copyright © Ruslan Panin
Carabus (Morphocarabus) spasskianus cf putus Motschulsky, 1846
Male adult
Russia, SW Siberia, C-Altays Mts, Kuray Mt Rng NE Aktash vill., h=2100-3000 m, 8-13.VII.2022, E.Filippov leg./det.R Panin, L= 23 mm (alpine petrophilic population close to the form schestopalovi Plutenko, 1995)
Copyright © Ruslan Panin
Carabus (Morphocarabus) spasskianus putus Motschulsky, 1846 (loc.typ.)
Female adult
Russia, W Altay, 18 km N of Zmeinogorsk, 23.VI.1998, A.& N. Menstschikovy leg. L=24 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) spasskianus putus Motschulsky, 1846 (loc.typ.)
Male adult
Russia, W Altay, 18 km N of Zmeinogorsk, 23.VI.1998, A.& N. Menstschikovy leg. L=23 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) odoratus antropovi Shilenkov 1996
Female adult
Russia, S Siberia, Tuva, W Tannu-Ola Mts. 35 km NE Khandagayty vill., Teeli Pass 2200-2800 m, 23-25.VI.2021, A.& E.Filippovs leg. L=19 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) odoratus antropovi Shilenkov 1996
Male adult
Russia, S Siberia, Tuva, W Tannu-Ola Mts. 35 km NE Khandagayty vill., Teeli Pass 2200-2800 m, 23-25.VI.2021, A.& E.Filippovs leg. L=19 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Male adult
SE Poland, Subcarpathia Prov., Nadleśnictwo Kołaczyce N of Jasło vill. M. Kosibowicz leg./det. R Panin, VII.2016, L=29 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Male adult
SE Poland, N Carpathians, Subcarpathia Prov., Nadleśnictwo Baligród, (209a) 22.VII.2021, M.Kosibowicz leg./det. R.Panin, L=31 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Male adult
SE Poland, N Carpathians, Subcarpathia Prov., National Park Magura, VII.2016, M.Ranocha leg./ det. R. Panin, L=30 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Male adult
SE Poland, N Carpathians, Subcarpathia Prov., Nadleśnictwo Lesko, VII.2017 M.Ranocha leg./ det. R. Panin, L=31 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
Male adult
Russia, NE Altay, Turochak Distr., Lake Teletskoe N part, near Yaylyu vill. 23-25.VII.2019, Ivliev P. leg./ det. R. Panin, L=24 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) aeruginosus eschscholtzi Mannerheim, 1827
Female adult
NE Kazakhstan, W Altay Mts, near Shemonaikha city., flood plain, 25-28.VII.2018, S.Vashchenko leg. L=25 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) aeruginosus eschscholtzi Mannerheim, 1827
Male adult
NE Kazakhstan, W Altay Mts, near Shemonaikha city., flood plain, 25-28.VII.2018, S.Vashchenko leg., L=22.5 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
Male adult
Russia, W Sayan Mts., Kazyr-Suk river, W slope of Oyskiy Mt. rng., ?h=2200 m 7-15.VII.1999, Brinev A. leg., L=19 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) monilis tanarguensis Maguerre & Stefani, 2021 (syn.) Holotype Paratype
Male+Female adult
Holotype, m (21 mm), environs du Col de la Croix de Bauzon, 1300 m, communes de La Souche, Borne et Mayres, Ardèche, France (07/2003), in coll. D. Maguerre, à Mons-en-Barœul, France. Paratype f (23 mm), au dessus du Col de Meyrand, 1400 mètres, communes de Loubaresse et de Borne, Ardèche, France, (07/2004), in coll. P. Stéfani.
Copyright © - - -
Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Female adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky forest, near Pyatnychany vill., 28.V-9.VI.2013, Panin R. leg. L=30 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) hummeli hummeli Fischer, 1823
Male adult
E Siberia, Sokhondinskii NP, Ingoda riv., 49.50.207N 111.10.825E, 1700m, 23.7.2009 Shavrin A. leg.
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) hummeli bonoi Obydov, 2006 Holotype Paratype
Male+Female adult
Central Yakutia, 25.6.1958, Shubnikova leg.
Copyright © - - -
Carabus (Morphocarabus) zubatschiki Obydov & Saldaitis, 2008 Holotype Paratype
Male+Female adult
N-China: Nei Mongol: Greater Khingan Mt. R: Arxan 1500m 1-10.7.2008. (~47:10/119:58°)
Copyright © - - -
Carabus (Morphocarabus) wulffiusi liaoningensis Deuve & Imura, 1990
Female adult
China, Liaoning, Kuandian Co., Banlaing, Ba-che-chuan (cAA)
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) venustus Morawitz, 1862
Male adult
China, Heilongjiang, Wuchang city, Mt. Lao-Tu-Ding-Zi 1593m 1-18.6.2008 (cAA).
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) scheidleri Panzer, 1799
Male adult
E-Austria, Wienerwald, Hochstraß am Lengbachl, h=575 m, 25.VI-2.VII.2024, M. Fuchs leg. L= 30 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) aeruginosus aeruginosus (as salechardensis Obydov, 1999)
Female adult
N Russia, Polar Ural, Yamalo-Nenets reg., Harp station envir., 15.VII.1995, G. Chekura leg.
Copyright © Ruslan Panin
Carabus (Morphocarabus) henningi oviformis Beheim & Breuning, 1943
Male adult
Russia, Polar Ural, Yamalo-Nenets reg.,Harp station envir., 15.VII.1995, G. Chekura leg. L=19 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) scheidleri (as floriani Penecke, 1905)
Female adult
SE-Austria, Styria, Gratwein Kehrerwaldh=580 m, 47.130800, 15.270800, VI-VII.2025, P. Fritz leg., L=32 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) scheidleri (as floriani Penecke, 1905)
Male adult
SE-Austria, Styria, Gratwein Kehrerwaldh=580 m, 47.130800, 15.270800, VI-VII.2025, P. Fritz leg., L=31 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) scheidleri bergeri Peeters & I.Rapuzzi, 2025
Female adult
W-Austria, Tyrol, Ötztal Alps, Kaunertal, Feichten, h=1281 m, 47.035200, 10.748700, 1.VIII.2025, M.Egger leg., L=27 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) scheidleri bergeri Peeters & I.Rapuzzi, 2025
Male adult
W-Austria,Tyrol, Vergötschen (Kaunertal), Feichten, h= 1300 m 11.VIII.2024, M.Egger leg. L=26 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) scheidleri bergeri Peeters & I.Rapuzzi, 2025
Male adult
W-Austria, Tyrol, Ötztal Alps, Kaunertal, Feichten, h=1281 m, 47.035200, 10.748700, 1.VIII.2025, M.Egger leg., L=26 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Female adult
NE-Serbia, Majdanpek env., h=520 m, 10-18.VI.2015, M. Janalik leg., L=32 m
Copyright © Ruslan Panin
Carabus (Morphocarabus) regalis jenissoni Dejean, 1831
Male adult
C Altay Mts., Verhnyaya Karasuriv., 850m, 25.7.2000, A. Matalin leg. (DUBC)
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) regalis jenissoni Dejean, 1831
Male adult
NE Altay mts., Altaisky NP., S slope Korbu Mt rng., Kokshi, 650m, Teletskoe lake, 4-7.7.2002 Matalin A leg. (DUBC)
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878 (variability)
Male adult
Far East, Promorskij Krai, Ussuriisk reg., Kamenushka, 9-15.6.2008 (cAA)
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
Female adult
Russia, Far East, Primorskiy Kray, Chuguevsky distr., 30 km ESE from Chuguevka, 9-14.VIII.2007, S.Shabalin leg.
Copyright © Ruslan Panin -
Morvanicarabus taxon page (1 photos)
Carabus (Morvanicarabus) scheibei scheibei Eidam, 1937
Female adult
NE-Afghanistan, Hindu Kush, Nuristan Prov., near Parun, 35°19'N 70°52'E h=2900 m, Pine forest, VI.2022, local collector/det. Kleinfeld, L= 24 mm
Copyright © Ruslan Panin -
Neoplectes taxon page (286 photos)
Carabus (Neoplectes) starckianus ganglbaueri Reitter, 1888
Male adult
Russia, NW Caucasus, Krasnodar reg., Adygeia distr., Guzeripl env. 20.5.2003
male, 26 mm
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) starckianus ganglbaueri Reitter, 1888
Female adult
NW Caucasus, S Russia, Adygea Rep., Confluence of the river Belaya & Kisha h~700m, 24.V-7.VIII.2010, D.Fominyh leg.
female,32 mm
Copyright © Ruslan Panin
Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
Female adult
female, 49mm
Copyright © Boleslav Brezina
Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
Male adult
male, 43mm
Copyright © Boleslav Brezina
Carabus (Neoplectes) apollo apollo Zolotarev, 1913
Male adult
Caucasus, Abchasia
Copyright © Dmitri Obydov
Carabus (Neoplectes) apollo phoebus Kurnakov, 1962
Male adult
Abchasia, Kodori, A. Zolotarew leg.
Photo by D.V. Obydov
from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Neoplectes) daphnis askhicus Belousov, 1992 Paratype
Male adult
SC-Caucasus, W Georgia, valley of riv. Chkhorodukhu, h=1000-1200m, 14.V.-6.VII.1991, Zamotajlov A.S & Miroshnikov leg.
male, 29 mm
photo & collection Panin R.
Copyright © Ruslan Panin
Carabus (Neoplectes) daphnis askhicus Belousov, 1992 Paratype
Male+Female adult
W Georgia, valley of Chkhorodukhu riv., 1000-1200m
14.V-6.VII.1991
male 30mm, female 33mm
photo & collection Anichtchenko A.
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) daphnis askhicus Belousov, 1992 Paratype
Male adult
Georgia, env. Martvilli, Askhi plateau
10-15.7.2001 A.Rubenian leg.
27 mm
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) daphnis daphnis Kurnakov, 1962
Male adult
Georgia, Mingrelia, Lecheha vallis, Tehuri superior
Caucasus centr., 1200m, 27.6-7.7.1989 Retezar I. leg.
male, 28 mm
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) edithae exedithae Gottwald, 1985
Male adult
Karachaevo-Cherkessia, low cource of Daut river, 16-20.V.2009 Anichtchenko A. leg
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) edithae exedithae Gottwald, 1985
Female adult
Karachaevo-Cherkessia, Zelenchukskii distr., <br /> near Bukovo vill., 8-13.VII.2007, S.Dementiev leg.<br />
Copyright © Dmitri Obydov
Carabus (Neoplectes) edithae kabardensis Kurnakov, 1970
Female adult
Kabardino-Balkaria, Skalisty Mt Ridge, 1800 m, 20.VII.1996. Andreev leg. Photo by D.V. Obydov
Copyright © Dmitri Obydov
Carabus (Archiplectes) jason Semenov, 1898
Male adult
Caucasus, W Georgia, Svanetia, upper reaches of riv. Shoudidi, 1700-2100m, 20-21.V.1986
leg. A. Zamotajlov
Copyright © Sunvas
Carabus (Neoplectes) jason Semenov, 1898
Male adult
Abkhasia, up river Shaudidi, 21.V - 2.VII.1986, Zamotailov A. leg.
Photo by D.V. Obydov
Copyright © Dmitri Obydov
Carabus (Neoplectes) juenthneri atchibachi Zamotajlov, 1991 Paratype
Female adult
Abkhasia,
Valley of river Bzyb near Mt. Achibakhu. h=1500m., A.S. Zamotajlov leg.
female: 33mm, 31mm
coll. R.Panin
Copyright © Ruslan Panin
Carabus (Neoplectes) juenthneri atchibachi Zamotajlov, 1991 Paratype
Female adult
W Caucas, Abkhasia, Valley of river Bzyb near Mt. Achibakhu. h=1500m.,
24.IV.-6.VI.1991 A.S. Zamotajlov leg.
coll. Anichtchenko
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) juenthneri atchibachi Zamotajlov, 1991 Paratype
Male adult
W Caucas, Abkhasia, Valley of river Bzyb near Mt. Achibakhu. h=1500m.,
24.IV.-6.VI.1991 A.S. Zamotajlov leg.
coll. Anichtchenko
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) juenthneri atchibachi Zamotajlov, 1991 Paratype
Male adult
W Caucas, Abkhasia, Valley of river Bzyb near Mt. Achibakhu. h=1500m.,
24.IV.-6.VI.1991 A.S. Zamotajlov leg.
coll. Anichtchenko
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) juenthneri avadcharensis Kurnakov, 1970 (loc.typ.)
Male+Female adult
Abkhasia, near Avadhara, Site Pyv h~1850m, 43.490730 40.686433, 1.VI-12.VII.2009, Zamotajlov A.S., Bondarenko A.S. leg.
L=27mm
Copyright © Ruslan Panin
Carabus (Neoplectes) juenthneri avadcharensis Kurnakov, 1970
Male adult
Abkhasia, 7,4km NE from Ritza lake, 1,5 km N Kutykuh vill.,
1635 m, 19-25.VII.2008, A. Vlasenko leg.
Copyright © Dmitri Obydov
Carabus (Neoplectes) juenthneri dsychvensis Gottwald, 1985
Male adult
Abchasia, N Dsykhwa, coll. A. Zolotarew
Photo by D.V. Obydov
Copyright © Dmitri Obydov
Carabus (Neoplectes) juenthneri juenthneri Reitter, 1899
Female adult
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) koltzei faunulus Kurnakov, 1970
Female adult
Abchasia, Akarmara, 6500 f, A. Zolotarew leg.
Photo by D.V. Obydov
Copyright © Dmitri Obydov
Carabus (Neoplectes) koltzei faunus Kurnakov, 1970
Male adult
Abkhazia, Kadorsky Mt Ridge, 1000 m, Chlou, 22.V.1993, A. Evstigneev leg.
Photo by D.V. Obydov
Copyright © Dmitri Obydov
Carabus (Neoplectes) koltzei kobachidzei Kurnakov, 1970
Female adult
Caucasus, E Abkhasia, SW slopes of Akiba mt. rng., 1200m, 7.V.-2.VII.1991<br /> Zamotajlov & Miroshnikov leg.
Copyright © Sunvas
Carabus (Neoplectes) koltzei kobachidzei Kurnakov, 1970
Female adult
Caucasus, E Abkhasia, SW slopes of Akiba mt. rng., 1200m, 7.V.-2.VII.1991
Zamotajlov & Miroshnikov leg.
Copyright © Sunvas
Carabus (Neoplectes) koltzei kobachidzei Kurnakov, 1970
Male+Female adult
Caucasus, E Abkhasia, SW slope of Akiba mt.rng., 1200m, 7.V.-2.VII.1991 Zamotajlov & Miroshnikov leg.
male 37mm, female 38mm
photo & collection Anichtchenko A.
Copyright © Alexander Anichtchenko
Carabus (Archiplectes) koltzei Rost, 1889
Male adult
Abkhasia, Habyu 3.7.2002
Ivliev P. leg.
32 mm
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) koltzei koltzei Rost, 1889
Male adult
Abkhasia, Tsebeldy, 23.V.1986, A.Zamotailov leg.
Photo by D.V. Obydov
Copyright © Dmitri Obydov
Carabus (Neoplectes) koltzei lennoni Gottwald, 1985
Male adult
Abkhasia, Amtkel lake, 22.V - 4.VII.1986, A.Zamotailov leg.
Photo by D.V. Obydov
from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Neoplectes) komarowi komarowi Reitter, 1882
Female adult
Centr. Caucasus, Svanetia, coll. A. Zolotarew<br /> Photo by D.V. Obydov<br /> from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Neoplectes) kratkyi felicitanus Reitter, 1893
Male adult
Russia, N Caucasus, Karachaevo-Cherkessia 2200m, Bolshoi Pzizer mt. 8.8.2002
25 mm
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) kratkyi felicitanus Reitter, 1893 (as porcellus)
Male adult
NW Caucasus, Resp. Adygea, Mt.Tybga, 28.VI.1997
male,20mm
Copyright © Ruslan Panin
Carabus (Neoplectes) kratkyi felicitanus Reitter, 1893 (as porcellus)
Male adult
Prov. Kuban, mt. Tybga, 1300 m, VI.1913, Shaposhnikov leg.
Copyright © Dmitri Obydov
Carabus (Neoplectes) kratkyi felicitanus Reitter, 1893
Female adult
Rusia, Caucas, Dzhuga mt., 2200 m, 24.VI. - 22.VIII.1986, Zamotajlov A. leg.
Copyright © Dmitri Obydov
Carabus (Neoplectes) kratkyi Ganglbauer, 1890
Male+Female adult
W Caucas, 1900m, Khatsavitaya Mt., 4.VI-13.VII.1994
A.Miroshnikov leg.
male 25mm, female 26mm
photo & collection Anichtchenko A.
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) kratkyi Ganglbauer, 1890
Male adult
Russia, NW-Caucasus: <br /> Krasnodar reg., Mostovskoy distr., Khatsavitaja Mt. h=1900m, 4.VI.-13.VII.1994, A. Miroshnikov leg.<br /> male, 26mm<br /> photo Panin R./det.Solodovnikov I.A.
Copyright © Ruslan Panin
Carabus (Neoplectes) kratkyi Ganglbauer, 1890
Male+Female adult
W Caucas Mts., 2500m, Mt. Yatyrgvarta,
25.VI-12.VIII.1992 Shchurov V. leg.
male 20mm, female 21mm
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890
Female adult
NW-Caucasus, Russia,
Karacaj-Cherkessija, Abishira-Akhuba range h=2650m, upper Kyafar-Agur Riv., 4-9.VIII.2009, Solodovnikov I.A. leg.
female, 23 mm
Copyright © Ruslan Panin
Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890
Male adult
NW-Caucasus: Karacaj-Cherkessija,
Abishira–Akhuba Mnt. Range h=2000-2650m, Kyafar-Agur Riv., 14–17.VII.2009, Fominykh D. leg.
male, 22mm
Copyright © Ruslan Panin
Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890
Male adult
Caucasus, Kuban, Meyer-Darcis-Patria
Copyright © Dmitri Obydov
Carabus (Neoplectes) kratkyi solodovnikovi Zamotajlov, 1992
Male adult
W. Caucasus, M. Bambak Mt., VI.1931, Vvedensky leg.<br /> from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Archiplectes) plasoni dalensis Rost, 1893
Male adult
Abkhasia, near Yu. Ptysh vill., 14.7.1986 Zamotailov A. leg. 31 mm
Copyright © Alexander Anichtchenko
Carabus (Archiplectes) plasoni dalensis Rost, 1893
Male adult
Caucasus merid., Dal-Suchumi, N.S. Brianskyj leg.<br /> from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Archiplectes) plasoni plasoni Ganglbauer, 1886
Female adult
Gruzia, Centr. Caucasus, Svanetia, coll. A. Zolotarew<br /> det. D.V. Obydov<br /> from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Neoplectes) polychrous Rost, 1892
Male adult
Abkhasia, Gudaut distr., near Huap vill., 600 m, 9.VI.1990, M. Danilevskyi leg. det. D.V. Obydov
Copyright © Dmitri Obydov
Carabus (Neoplectes) rousianus rousianus Gottwald, 1985
Male adult
Abkhasia, 7,6 km NE Pitsunda, 2,1 km NNE Ryapsh riv mouth, Ryapsh canyon h=60 m, 43°10N 40°25E, 25-27.VII.2008, A.Vlasenko leg.<br /> det. D.V. Obydov
Copyright © Dmitri Obydov
Carabus (Archiplectes) prometheus miroshnikovi Zamotajlov, 1990
Female adult
NW-Caucasus, Krasnodar reg., Mostovskoy distr., Site Val , h=1800m. 07-08.VII.2008, D.Fominyh leg., 35mm; det. I. Solodovnikov 2023
Copyright © Ruslan Panin
Carabus (Archiplectes) prometheus zolotarevi Zamotajlov, 1988
Male adult
Paratypus, male Prov. Kuban distr., Maikop, Laba Min., Ch. Shaposhnikov leg.
from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Archiplectes) prometheus zolotarevi Zamotajlov, 1988 Paratype
Male adult
Paratypus, male - Fl. Min. Laba, VII.1918, Shaposhnikov leg.
from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Archiplectes) prometheus basilianus Starck, 1890
Male adult
Russia, W Caucas, Port-Artur Mt. 22.VI.1986, A.Zamotajlov leg.
det. D.V. Obydov
from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Neoplectes) prometheus dzhugensis Zamotajlov, 1988 Paratype
Female adult
Paratypus, female<br /> Prov. Kuban circ., mt. Dzhuga (Tigeni), 27.VI.1911.<br />
Copyright © Dmitri Obydov
Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990
Male+Female adult
W.Caucasus,
Psebai env., Nikitino, 2.V-3.VII.1993, leg. A.S.Zamotajlov
female 33mm
Copyright © Ruslan Panin
Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990 (as hephaestus)
Male adult
Paratypus Carabus (Archiplectes) hephaestus
Maikop, Ust. Urusht, 26.IV.1910
Copyright © Dmitri Obydov
Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990
Female adult
W Caucasus, Krasnodar reg., Maikop distr.
Nikel, 1.7.1989
38 mm
Copyright © Alexander Anichtchenko
Carabus (Archiplectes) prometheus pschadensis Zamotajlov, 1990
Female adult
NW Caucas, 5 km N from Pshada vill., 25.04.1989, Smorodinov leg.
specimen from collection of M.Shestopalov (Moscow).
det. D.V. Obydov
from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Neoplectes) prometheus prometheus Reitter, 1887
Male adult
NW-Caucasus,Krasnodar reg., Apsheron Distr., Lagonaki Mt., near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominykh leg.
L= 42mm.
Copyright © Ruslan Panin
Carabus (Neoplectes) prometheus prometheus Reitter, 1887 (as zugmayeriae)
Male adult
W. Caucasus, Maikop, 20.VI.1967., A. Bogachev leg.
det. D.V. Obydov
Copyright © Dmitri Obydov
Carabus (Neoplectes) prometheus prometheus Reitter, 1887
Male+Female adult
NW Caucasus, Russia,
Krasnodar reg.,Krasnoarmeisk distr., Krasny Les, near Podlesnyi vill.,17-22.IV.2008, D.D.Fominykh leg.
female, 40mm
Copyright © Ruslan Panin
Carabus (Archiplectes) prometheus prometheus Reitter, 1887 (as nubicola)
Male+Female adult
W Caucasus, Adygea,
h~1700m. upper res. Riv. Kurdzhips, 11-21.VIII.2007, I.A.Solodovnikov leg./det.
L=32mm.
Copyright © Ruslan Panin
Carabus (Neoplectes) prometheus prometheus Reitter, 1887
Male adult
Krasnodar distr., Mezmai
20.6.1989
38 mm
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) protensus olegi Belousov, 1992 Paratype
Male adult
Caucasus, W Georgia, Svanetia, right bank of riv. Magana, 800m, 12.V.-4.VII.1991, Zamotajlov & Miroshnikov leg.
male, 31mm
photo and collection Anichtchenko A.
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) protensus protensoides Novotny & Vorisek, 1988
Male adult
Georgia, Martvilli env., Akshi plateau 10-15.7.2001
Rubenian leg.
32 mm
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) reitteri gagrinus Starck, 1894
Female adult
Abkhasia, near Gagry, 16.V.1992, A. Evstigneev leg.
det. D.V. Obydov
Copyright © Dmitri Obydov
Carabus (Neoplectes) reitteri laevisternis Starck, 1890
Male adult
Abkhasia,W.slope of Gagrinsky Mt., Krepostnaya River source, h=1325m., A.S.Zamotajlov leg. 27.VII.2000
male, 29 mm
Copyright © Ruslan Panin
Carabus (Neoplectes) reitteri laevisternis Starck, 1890
Male adult
W-Caucasus:Sochi env., Aibga Mt. h=2000m., 06.VII.2000
male, 29mm
Copyright © Ruslan Panin
Carabus Neoplectes) reitteri reitteri Retowski, 1885
Female adult
NW Caucasus, Adygeya, Fischt-Oshten Mts. base, h=1800 m.<br /> 20.07.2006, D.Safronov leg.
Copyright © Safronov
Carabus (Neoplectes) reitteri reitteri Retowski, 1885 (as achunensis)
Male adult
W. Caucasus, Sochi Distr., Chernomoretz Vill. env.,
15.VII.1992, P. Gorbachev leg.
det. D.V. Obydov
Copyright © Dmitri Obydov
Carabus (Neoplectes) satyrus besleticus Kurnakov, 1972 (as gaskoi)
Male adult
USSR, Caucasus occid, Abchasia, Suchumi, vallis Kelasuri, 100m 25.6-17.7.1977 Retezar I. leg. 36 mm
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) satyrus mtsaranus Kurnakov, 1972
Male adult
Abkhasia, Bzhybskii Mt.rng., near Mtsara vill., 16.VI.1991, S.Gusev leg.
det. D.V. Obydov
Copyright © Dmitri Obydov
Carabus (Neoplectes) satyrus napraensis Belousov & Zamotajlov, 1993 Paratype
Male adult
Abkhasia, Bzybian Mt. Range near Mt. Chibzharga., h=1900-2000m., A.S. Zamotajlov leg.
male, 28mm
photo & coll.Panin R.
Copyright © Ruslan Panin
Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972
Male adult
Abkhasia, Habyu vill.,2.7.2002 Ibliev leg.<br /> 35 mm
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972
Male adult
Abkhasia, Habyu vill.<br /> 2.7.2002 Ibliev leg.<br /> 35 mm
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) starcki Heyden, 1885
Male+Female adult
NW-Caucasus,
Krasnodar reg., Mt. Aishkho, Malaya Laba River, h=2200m., 14.VII.2008, D.Fominyh leg.
female-21mm
Copyright © Ruslan Panin
Carabus (Neoplectes) starcki Heyden, 1885
Male adult
Caucasian Nat. Park, S slope
Pseashkho mt. 2500m 21.7.1975, Ovtsharenko leg.
20 mm
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) starcki Heyden, 1885
Male adult
NW Caucasus, Adygeya, Pshekho-Su Mt. env., h=1500 m., 21.07.2006, D.Safronov leg.
Copyright © Safronov
Carabus (Neoplectes) starcki Heyden, 1885 (as parvulus)
Male adult
Prov. Kuban, mt. Aischa, VI.1906, Shaposhnikov leg.
det. D.V. Obydov
Copyright © Dmitri Obydov
Carabus (Neoplectes) starckianus adelaidae Starck, 1889
Male adult
Russia, Caucasus, Chvizhepse River Val., 27.V.1994, P. Gorbachev leg.<br /> Photo from www.zin.ru<br /> det. D.V. Obydov
Copyright © Dmitri Obydov
Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988 (as shchessiensis) Paratype
Male+Female adult
NW Caucasus,
Krasnodar reg., Tuapsinskyi distr. Shessi Mt.
30.V.-25.VI.1993 Mt.Shessi, h=1800m., Imura & A.S.Zamotajlov, 1993 leg.
female, 29mm
Copyright © Ruslan Panin
Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988
Female adult
Russia,W Caucasus,
Krasnodar reg.,Chernogorie mts.,Chernogor mt. h=1700-1800m. 23.V.-30.VII.1999. A.S.Zamotajlov leg.
female 29mm.
Copyright © Ruslan Panin
Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988
Male adult
West. Caucasus, Shakhe Valley, 1.VI.1989, A. Shamaev leg.
det. D.V. Obydov
Copyright © Dmitri Obydov
Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
Male adult
NW-Caucasus, Russia,
Krasnodar reg., Apsheron Distr., Lagonaki Mt. ,near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominykh leg.
male, 28mm
Copyright © Ruslan Panin
Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
Male+Female adult
NW-Caucasus, Russia,
Krasnodar reg.,the right bank of Mzymta river, Engelmanovs glades, h=1200m., A.S.Zamotajlov leg.
female-30mm\male-28mm
Copyright © Ruslan Panin
Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
Male adult
Rusia, W Caucasus, Sochi distr., Aishkha mts.
900m, 15.7.2003 Gorbachev leg.
30 mm
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) starckianus starckianus Ganglbauer, 1886
Male adult
Russia, Krasnodar reg., 6 km NE Soloniki vill.,
Sochi Nat.Park, 28-30.V.2009 Anichtchenko A. leg.
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) starckianus starckianus Ganglbauer, 1886
Female adult
W Caucas, Sochi distr., near Lazorevskoe vill., 9.06.1993, P.Gorbachev leg.
det. D.V. Obydov
Copyright © Dmitri Obydov
Carabus (Neoplectes) starckianus starckianus Ganglbauer, 1886 (as phlagochensis)
Female adult
Caucasus, Sotschi, Flagoch.
det. D.V. Obydov
from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Neoplectes) starckianus theseus Briansky, 1910
Male+Female adult
NW-Caucasus, Russia,
Krasnodar reg.,Tuapse distr., Mt.Semashkho 04.V.2008, D.Fominyh leg.
female, 32mm
Copyright © Ruslan Panin
Carabus (Neoplectes) starckianus theseus Briansky, 1910
Male adult
Tuapse distr., Anastasievka vill., Semiglavaya mt.
riv. Pshekaho 20.7.1995
30 mm
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) starckianus theseus Briansky, 1910
Female adult
Tuapse distr., Anastasievka vill., Semiglavaya mt.
bass. Pshekaho riv. 20.7.1995
female, 30 mm
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
Female adult
Copyright © Kirill Makarov
Carabus (Neoplectes) starckianus starckianus (as phlagochensis)
Female adult
Russia, Кrasnodar Reg., Dagomys, Flagokh, N43-44' / E39-40', 11.VII.2010, N. Pichuginleg.
Specimen from the collection N. U. Pichugin (Vladimir).
Copyright © Dmitri Obydov
Carabus (Neoplectes) satyrus duripshensis Kurnakov, 1972
Male+Female adult
W.Caucasus, Abkhasia, Gudauta distr.,
Duripsh village env., Khipsta valley, h=500-700 m.
07-12.VI.2009 A.Safronov leg.
Body lenght 30-33 mm.
Copyright © Alexandr Safronov
Carabus (Neoplectes) satyrus duripshensis Kurnakov, 1972
Female adult
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) satyrus duripshensis Kurnakov, 1972
Male adult
W.Caucasus, Abkhasia, Gudauta distr.,
Duripsh village env., Khipsta valley, h=500-700 m.
07-12.VI.2009 A.Safronov leg.
male - 30 mm.
Copyright © Alexandr Safronov
Carabus (Neoplectes) juenthneri avadcharensis Kurnakov, 1970
Male+Female adult
W. Caucasus, Abkhasia, 8 km NE of Ritza Lake,<br /> Avadkhara valley, h=1700 m<br /> 10-VI-2009 A.Safronov leg.
Copyright ©
Carabus (Neoplectes) polychrous polychrous Rost, 1892
Female adult
W.Caucasus, Abkhasia, Gudauta distr., Duripsh village env., Khipsta valley, h=500-700 m. 8-11.06.2009 A. Safronov leg. lenght 40-43 mm.
Copyright ©
Carabus (Neoplectes) reitteri gagrinus Starck, 1894 (as gegicus Gottwald, 1985)
Male+Female adult
W.Caucasus, Abkhasia, Gagra distr., Gega riv. valley,
13-15.07.2005 D. Kasatkin. leg.
male - 30 mm, female - 34 mm.
Copyright © Alexandr Safronov
Carabus (Neoplectes) edithae exedithae Gottwald, 1985
Male adult
NС-Caucasus, Karachaevo-Cherkessia, Abishir-Akhuba Mt.Range h=2250-2600 m, right bank Pkhiya river, 10-13.VI.2013, S. Pavluchuk leg.<br /> L=25 mm
Copyright © Ruslan Panin
Carabus (Neoplectes) reitteri reitteri Retowski, 1885
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Carabus (Neoplectes) satyrus satyrus Kurnakov, 1962
Male+Female adult
Abkhazia, Gulripsh Distr., NW slopes of Gurzul Mt. Range near Merkheul Village, 170-230 m, left tributary of Machara River, hornbeam, alder, rhododendron (Rhododendron ponticum) forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Neoplectes) satyrus satyrus Kurnakov, 1962
Female adult
Abkhazia, Gulripsh Distr., NW slopes of Gurzul Mt. Range near Merkheul Village, 170-230 m, left tributary of Machara River, hornbeam, alder, rhododendron (Rhododendron ponticum) forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Neoplectes) satyrus satyrus Kurnakov, 1962
Male adult
Abkhazia, Gulripsh Distr., NW slopes of Gurzul Mt. Range near Merkheul Village, 170-230 m, left tributary of Machara River, hornbeam, alder, rhododendron (Rhododendron ponticum) forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Neoplectes) satyrus satyrus Kurnakov, 1962
Male adult
Abkhazia, Gulripsh Distr., NW slopes of Gurzul Mt. Range near Merkheul Village, 170-230 m, left tributary of Machara River, hornbeam, alder, rhododendron (Rhododendron ponticum) forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014 Holotype Paratype
Male adult
Abkhazia, N Slopes of Bzybian Mt. Range, right bank of River Reshevie (= Reshava) (left tributary of River Bzyb), 700-1000 m, 13.V-02.VII.2010, leg. D. Fominykh, I. Solodovnikov
Copyright © Igor Solodovnikov
Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014 Paratype
Male adult
Abkhazia, N Slopes of Bzybian Mt. Range, right bank of River Reshevie (= Reshava) (left tributary of River Bzyb), 700-1000 m, 13.V-02.VII.2010, leg. D. Fominykh, I. Solodovnikov
Copyright © Igor Solodovnikov
Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014 Paratype
Male+Female adult
Abkhazia, N Slopes of Bzybian Mt. Range, right bank of River Reshevie (= Reshava) (left tributary of River Bzyb), 700-1000 m, 13.V-02.VII.2010, leg. D. Fominykh, I. Solodovnikov
Copyright © Igor Solodovnikov
Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014 Paratype
Male adult
Abkhazia, N Slopes of Bzybian Mt. Range, right bank of River Reshevie (= Reshava) (left tributary of River Bzyb), 700-1000 m, 13.V-02.VII.2010, leg. D. Fominykh, I. Solodovnikov
Copyright © Igor Solodovnikov
Carabus (Neoplectes) pseudopshuensis Zamotajlov, 1991
Male+Female adult
Abkhazia, Sukhum Distr., SE of Village Pskhu, hamlet Bitaga, Mt. Bzybskaya, 550-650 m, beech forest site, 12.V-30.VI.2010, leg. I. Solodovnikov, D. Fominykh
Copyright © Igor Solodovnikov
Carabus (Neoplectes) pseudopshuensis Zamotajlov, 1991
Male+Female adult
Abkhazia, Sukhum Distr., SE of Village Pskhu, hamlet Bitaga, Mt. Bzybskaya, 550-650 m, beech forest site, 12.V-30.VI.2010, leg. I. Solodovnikov, D. Fominykh
Copyright © Igor Solodovnikov
Carabus (Neoplectes) pseudopshuensis Zamotajlov, 1991
Male adult
1 male - Abkhazia, Sukhum Distr., vicinities of Village Sanchara, River Baul, beech forest site, 20.VII.1991, leg. M.N. Maksimenkov 2 male - Abkhazia, Sukhum Distr., SE of Village Pskhu, hamlet Bitaga, Mt. Bzybskaya, 550-650 m, beech forest site, 12.V-30.VI.2010, leg. I. Solodovnikov, D. Fominykh
Copyright © Igor Solodovnikov
Carabus (Neoplectes) pseudopshuensis Zamotajlov, 1991
Male+Female adult
Abkhazia, Sukhum Distr., N Slopes of Bzybian Mt. Range near Village Serebryanyi, 500-600 m, 25.IV-05.VII.2013, leg. D. Fominykh
Copyright © Igor Solodovnikov
Carabus (Neoplectes) pseudopshuensis Zamotajlov, 1991
Male+Female adult
Abkhazia, Sukhum Distr., N Slopes of Bzybian Mt. Range near Village Serebryanyi, 500-600 m, 25.IV-05.VII.2013, leg. D. Fominykh
Copyright © Igor Solodovnikov
Carabus (Neoplectes) pseudopshuensis Zamotajlov, 1991
Male+Female adult
Abkhazia, Sukhum Distr., N Slopes of Bzybian Mt. Range near Village Serebryanyi, 500-600 m, 25.IV-05.VII.2013, leg. D. Fominykh
Copyright © Igor Solodovnikov
Carabus (Neoplectes) besleticus besleticus Kurnakov, 1972
Male+Female adult
Abkhazia, Sukhum Distr., vicinities of Village Kaman, N slope of Mt. Birtskha, 160 m, beech, linden, box, elder forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Neoplectes) besleticus besleticus Kurnakov, 1972
Male+Female adult
Abkhazia, Sukhum Distr., vicinities of Village Kaman, N slope of Mt. Birtskha, 160 m, beech, linden, box, elder forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Neoplectes) besleticus besleticus Kurnakov, 1972
Male+Female adult
Abkhazia, Sukhum Distr., vicinities of Village Kaman, N slope of Mt. Birtskha, 160 m, beech, linden, box, elder forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Neoplectes) besleticus besleticus Kurnakov, 1972
Male+Female adult
Abkhazia, Sukhum Distr., vicinities of Village Kaman, N slope of Mt. Birtskha, 160 m, beech, linden, box, elder forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972
Male+Female adult
Abkhazia, Gudauta Distr, vicinities of Mtsara (Chiryuta), Zashirbara Mt. Range, source of River Mtsara, beech, blackberry, fern site, 575-690 m, 11.V-05.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972
Male+Female adult
Abkhazia, Gudauta Distr, vicinities of Mtsara (Chiryuta), Zashirbara Mt. Range, source of River Mtsara, beech, blackberry, fern site, 575-690 m, 11.V-05.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972
Male+Female adult
Abkhazia, Gudauta Distr, vicinities of Mtsara (Chiryuta), Zashirbara Mt. Range, source of River Mtsara, beech, blackberry, fern site, 575-690 m, 11.V-05.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972
Male+Female adult
Abkhazia, Gudauta Distr, vicinities of Mtsara (Chiryuta), Zashirbara Mt. Range, source of River Mtsara, beech, blackberry, fern site, 575-690 m, 11.V-05.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Neoplectes) besleticus dsychvensis Gottwald, 1985
Male+Female adult
Abkhazia, Bzybian Mt. Range, SW slopes of Mt. Dzykhva, 2000-2300 m, alpine zone, N 43º13’, E 041º08’, 29.VI-03.VII.2013, leg. D. Fominykh
Copyright © Igor Solodovnikov
Carabus (Neoplectes) besleticus dsychvensis Gottwald, 1985
Male+Female adult
Abkhazia, Bzybian Mt. Range, SW slopes of Mt. Dzykhva, 2000-2300 m, alpine zone, N 43º13’, E 041º08’, 29.VI-03.VII.2013, leg. D. Fominykh
Copyright © Igor Solodovnikov
Carabus (Neoplectes) besleticus duripshensis Kurnakov, 1972
Male+Female adult
Abkhazia, Gudauta Distr., valley of River Khipsta, vicinities of Village Tvanaarkhu, karst crater with cave, beech, blackberry site, 425 m, N 43º14’ E 040º39’9”, 08.V-05.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Neoplectes) besleticus napraensis Belousov & Zamotajlov, 1993 Paratype
Male+Female adult
Abkhazia, Gudauta Distr., Bzybian Mt. Range, vicinities of Mt. Chibzharga, 1900-2000 m, 20.VI-26.VII.1992, leg. I. Belousov, A. Zamotajlov, A. Miroshnikov
Copyright © Igor Solodovnikov
Carabus (Neoplectes) besleticus napraensis Belousov & Zamotajlov, 1993 Paratype
Female adult
Abkhazia, Gudauta Distr., Bzybian Mt. Range, vicinities of Mt. Chibzharga, 1900-2000 m, 20.VI-26.VII.1992, leg. I. Belousov, A. Zamotajlov, A. Miroshnikov
Copyright © Igor Solodovnikov
Carabus (Neoplectes) besleticus anastasiae Solodovnikov, 2014 Holotype Paratype
Female adult
Abkhasia, Gudauta distr., S slope Bzybian Mt.R., upper reach. Riv. Khipsta, 43°18' N / 40°41' E, h = 1907-1915 m, subalpine meadows with karst blocks, 07-12.08.2014, leg. I.A. Solodovnikov, E.V. Tatun
Copyright © Igor Solodovnikov
Carabus (Neoplectes) besleticus anastasiae Solodovnikov, 2014 Paratype
Female adult
Abkhasia, Gudaura distr., S slope Bzybian Mt.R., upper reach. Riv. Khipsta, 43°18' N / 40°41' E, h = 1907-1915 m, subalpine meadows with karst blocks, 07-12.08.2014, leg. I.A. Solodovnikov, E.V. Tatun
Copyright © Igor Solodovnikov
Carabus (Neoplectes) besleticus anastasiae Solodovnikov, 2014 Paratype
Male adult
Abkhasia, Gudauta distr., S slope Bzybian Mt.R., upper reach. Riv. Khipsta, 43°18' N / 40°41' E, h = 1984-1990 m, subalpine meadows with big karst blocks, 07-12.08.2014, leg. I.A. Solodovnikov, E.V. Tatun
Copyright © Igor Solodovnikov
Carabus (Neoplectes) besleticus anastasiae Solodovnikov, 2014 Paratype
Female adult
Abkhasia, Gudauta distr., S slope Bzybian Mt.R., upper reach. Riv. Khipsta, 43°18' N / 40°41' E, h = 1984-1990 m, subalpine meadows with big karst blocks, 07-12.08.2014, leg. I.A. Solodovnikov, E.V. Tatun
Copyright © Igor Solodovnikov
Carabus (Neoplectes) khipstaensis Solodovnikov, 2015 Paratype
Male adult
Абхазия, Гудаутский р-н, Ю скл. Бзыбского хр., водораздельный холм в истоках р. Хипста, 43°17'52.03" N / 40°40'8.72" E, h = 1435 m, Линия № 64-4, буковый спелый лес с папоротниками и ежевикой, 09-13.08.2014, leg. И.А. Солодовников, Е.В. Татун
Copyright © Igor Solodovnikov
Carabus (Neoplectes) khipstaensis Solodovnikov, 2015 Paratype
Male adult
Абхазия, Гудаутский р-н, Ю скл. Бзыбского хр., водораздельный холм в истоках р. Хипста, 43°17'52.03" N / 40°40'8.72" E, h = 1435 m, Линия № 64-4, буковый спелый лес с папоротниками и ежевикой, 09-13.08.2014, leg. И.А. Солодовников, Е.В. Татун
Copyright © Igor Solodovnikov
Carabus (Neoplectes) khipstaensis Solodovnikov, 2015 Paratype
Female adult
Абхазия, Гудаутский р-н, Ю скл. Бзыбского хр., водораздельный холм в истоках р. Хипста, 43°17'52.03" N / 40°40'8.72" E, h = 1435 m, Линия № 64-4, буковый спелый лес с папоротниками и ежевикой, 09-13.08.2014, leg. И.А. Солодовников, Е.В. Татун
Copyright © Igor Solodovnikov
Carabus (Neoplectes) khipstaensis Solodovnikov, 2015 Paratype
Female adult
Абхазия, Гудаутский р-н, Ю скл. Бзыбского хр., водораздельный холм в истоках р. Хипста, 43°17'52.03" N / 40°40'8.72" E, h = 1435 m, Линия № 64-4, буковый спелый лес с папоротниками и ежевикой, 09-13.08.2014, leg. И.А. Солодовников, Е.В. Татун
Copyright © Igor Solodovnikov
Carabus (Neoplectes) rousianus rousianus Gottwald, 1985
Male+Female adult
Абхазия, Гудаутский р-н, 5-8 км ЮВ от пос. Лдзаа (Ludzava) долина р. Риапши, левый борт; балки, поросшие орехом, каштаном, рододендроном понтийским и кленом, h = 58-70 m, 43º10’43” N / 40º25’37” E, 16-22.07.2011, leg. И.А. Солодовников, Н.Ю. Пичугин, С.M. Павлючук
Copyright © Igor Solodovnikov
Carabus (Neoplectes) rousianus rousianus Gottwald, 1985
Male+Female adult
Абхазия, Гудаутский р-н, 5-8 км ЮВ от пос. Лдзаа (Ludzava) долина р. Риапши, левый борт; балки, поросшие орехом, каштаном, рододендроном понтийским и кленом, h = 58-70 m, 43º10’43” N / 40º25’37” E, 16-22.07.2011, leg. И.А. Солодовников, Н.Ю. Пичугин, С.M. Павлючук
Copyright © Igor Solodovnikov
Carabus (Neoplectes) polychrous Rost, 1892
Female adult
Абхазия, Гудаутский р-н, окр. п. Тванаарху, дол. р. Хипсты, ущелье небольшого ручейка, поросшее буками, ежевикой и папоротником, Линия 25-3, 43°14’ N / 040°39’ E, h = 500 m,08.05-05.07.2013, leg. И.А. Солодовников, С.В. Солодовникова, С.M. Павлючук, Н.Ю. Пичугин, В.М. Коцур
Copyright © Igor Solodovnikov
Carabus (Neoplectes) polychrous Rost, 1892
Male+Female adult
Абхазия, Гудаутский р-н, окр. п. Тванаарху, дол. р. Хипсты, ущелье небольшого ручейка, поросшее буками, ежевикой и папоротником, Линия 25-3, 43°14’ N / 040°39’ E, h = 500 m,08.05-05.07.2013, leg. И.А. Солодовников, С.В. Солодовникова, С.M. Павлючук, Н.Ю. Пичугин, В.М. Коцур
Copyright © Igor Solodovnikov
Carabus (Neoplectes) polychrous Rost, 1892
Male+Female adult
Абхазия, Гудаутский р-н, окр. п. Тванаарху, дол. р. Хипсты, ущелье небольшого ручейка, поросшее буками, ежевикой и папоротником, Линия 25-3, 43°14’ N / 040°39’ E, h = 500 m,08.05-05.07.2013, leg. И.А. Солодовников, С.В. Солодовникова, С.M. Павлючук, Н.Ю. Пичугин, В.М. Коцур
Copyright © Igor Solodovnikov
Carabus (Neoplectes) rousianus svetlanae Solodovnikov, 2015 Paratype
Female adult
Абхазия, Гудаутский р-н, 12 км дороги на оз. Рица, лев. борт р. Бзыбь, буковый лес + каштан на щебнисто-глинистой почве, уроч. Авджуляр, Линия № 20, h = 210-230 m, 43º19’07” N / 040º25’06” E, 09.V-03.VII.2013, leg. И.А. Солодовников, С.В. Солодовникова, С.М. Павлючук, Н.Ю. Пичугин, В.М. Коцур
Copyright © Igor Solodovnikov
Carabus (Neoplectes) rousianus svetlanae Solodovnikov, 2015 Paratype
Male+Female adult
Абхазия, Гудаутский р-н, 12 км дороги на оз. Рица, лев. борт р. Бзыбь, буковый лес + каштан на щебнисто-глинистой почве, уроч. Авджуляр, Линия № 20, h = 210-230 m, 43º19’07” N / 040º25’06” E, 09.V-03.VII.2013, leg. И.А. Солодовников, С.В. Солодовникова, С.М. Павлючук, Н.Ю. Пичугин, В.М. Коцур
Copyright © Igor Solodovnikov
Carabus (Neoplectes) rousianus svetlanae Solodovnikov, 2015 Paratype
Male adult
Абхазия, Гудаутский р-н, 12 км дороги на оз. Рица, лев. борт р. Бзыбь, буковый лес + каштан на щебнисто-глинистой почве, уроч. Авджуляр, Линия № 20, h = 210-230 m, 43º19’07” N / 040º25’06” E, 09.V-03.VII.2013, leg. И.А. Солодовников, С.В. Солодовникова, С.М. Павлючук, Н.Ю. Пичугин, В.М. Коцур
Copyright © Igor Solodovnikov
Carabus (Neoplectes) rousianus svetlanae Solodovnikov, 2015 Paratype
Male adult
Абхазия, Гудаутский р-н, 12 км дороги на оз. Рица, лев. борт р. Бзыбь, буковый лес + каштан на щебнисто-глинистой почве, уроч. Авджуляр, Линия № 20, h = 210-230 m, 43º19’07” N / 040º25’06” E, 09.V-03.VII.2013, leg. И.А. Солодовников, С.В. Солодовникова, С.М. Павлючук, Н.Ю. Пичугин, В.М. Коцур
Copyright © Igor Solodovnikov
Carabus (Neoplectes) reitteri gagrinus Starck, 1894
Male adult
ABKHASIA, Gagra distr., Mamdsyshkha Mt., near Gagra, Fagus forest + karst, h = 1300-1330 m, biotope N 23, 43º18’186" N / 040º18’44" E, 12.05-05.07.2014, Leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Neoplectes) reitteri gagrinus Starck, 1894
Female adult
ABKHASIA, Gagra distr., Mamdsyshkha Mt., near Gagra, Fagus forest + karst, h = 1300-1330 m, biotope N 23, 43º18’186" N / 040º18’44" E, 12.05-05.07.2014, Leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Neoplectes) reitteri gagrinus Starck, 1894
Female adult
ABKHASIA, Gagra distr., Mamdsyshkha Mt., near Gagra, Fagus forest + karst, h = 1300-1330 m, biotope N 23, 43º18’186" N / 040º18’44" E, 12.05-05.07.2014, Leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov
Carabus (Neoplectes) starckianus olgae Semenov, 1898 Lectotype
Male adult
[Circassia. Rost. Car. imperatot St.] [44] [Plectes Olgae m. A. Semenov det., ♂ Typ. un. X.96] red label этикетка [LECTOTYPUS Gottwald, 1983 design], yellow label [Zoological Institute Russian Academy of Sciences St Peterburg]
Copyright © Igor Solodovnikov
Carabus (Neoplectes) starckianus olgae Semenov, 1898 Paralectotypes
Male adult
• [Circassien. Rost. [18]98 Pl. obtusus var. 5.] [5] [Plectes Olgae m. A. Semenov det., ♂ I.98] red label [PARALECTOTYPUS Gottwald, 1983 design], yellow label [Zoological Institute Russian Academy of Sciences St Peterburg]
Copyright © Igor Solodovnikov
Carabus (Neoplectes) starckianus starckianus Ganglbauer, 1886
Male adult
Russia, Krasnodar area, Sochi, Sochi distr., right bank of Riv. Persiyanka, NE slope Phlagokh Mt., 03.05-02.07.2013, Leg. I.A. Solodovnikov, S.V. Solodovnikova (TOPOTYPUS as phlagochensis Imura & Zamotajlov, 1993)
Copyright © Igor Solodovnikov
Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014 Paratype
Female adult
Abkhazia, Bzybsky Mt.Rng, right bank of river Reshevie (left tributary of river Bzyb) h~670-720 м, 13-14.V.2010, D.Fominykh & I.Solodovnikov leg. L=41 mm
Copyright © Ruslan Panin
Carabus (Neoplectes) ibericus s.str. (as djanoliensis Fominykh & Zamotajlov, 2016) Paratype
Male+Female adult
Image from Fominykh & Zamotajlov, 2016
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Carabus (Neoplectes) ibericus ibericus (as prunierianus Deuve, 2012)
Male+Female adult
Georgia, Magaroskari env. Image from Fominykh & Zamotajlov, 2016
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Carabus (Neoplectes) ibericus ibericus (as prunierianus Deuve, 2012)
Male adult
Georgia, Passanauri env. Image from Fominykh & Zamotajlov, 2016
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Carabus (Neoplectes) ibericus ibericus (as szekelyi Retezar, 2011)
Male+Female adult
Itrapula env. Image from Fominykh & Zamotajlov, 2016
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Carabus (Neoplectes) ibericus ibericus (as szekelyi Retezar, 2011)
Male+Female adult
Dzhalobet env.; Image from Fominykh & Zamotajlov, 2016
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Carabus (Neoplectes) ibericus ibericus (as szekelyi Retezar, 2011)
Male+Female adult
Nakerala env.; Image from Fominykh & Zamotajlov, 2016
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Carabus (Neoplectes) komarowi komarowi Reitter, 1882
Female adult
NW Georgia, Upper Svaneti, Mestia distr., Lakhamula env. h~1250 m, 19.VI.2014, V.Sima leg. L=33 mm
Copyright © Ruslan Panin
Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
Female adult
W Georgia, Martvili Distr., 5 km N T'aleri, 11-18.VI.2014, V. Sima leg., L=50 mm
Copyright © Ruslan Panin
Carabus (Neoplectes) ibericus ibericus (as imereticus Retezar, 2011) Paratype
Male adult
S-Caucasus: NC-Georgia, W slope Likhi Mt. Rng., 3 km W of Rikoti Pass h~900 m, 15.VI - 8.VII.2006, A.Puchkov leg., L=29.5 mm
Copyright © Ruslan Panin
Carabus (Neoplectes) compressus rebellis Reitter, 1895
Male adult
Georgia, Kvemo-Svaneti, S slopes of Svanetskiy Mt.R., S slope of Latpari pass, h=2225m, 17.VI -13.VII.2017 leg. D.Fominykh L=24.0
Copyright © Kirill Makarov
Carabus (Neoplectes) compressus rebellis Reitter, 1895
Female adult
Georgia, Kvemo-Svaneti, S slopes of Svanetskiy Mt.R., S slope of Latpari pass, h=2225m, 17.VI -13.VII.2017 leg. D.Fominykh L=26.0 mm
Copyright © Kirill Makarov
Carabus (Neoplectes) zubovi Fominykh, Zamotajlov et Titarenko, 2018 Holotype
Male adult
S slopes of Svaneti Mt. Range near Lentekhi Town, S slopes of Goldash Mt., 42°49'N, 42°45'E, 2300–2800 m, 21.VI.2016, D. Fominykh, A. Zherebilo (ZISP).
Copyright © Kirill Makarov
Carabus (Neoplectes) compressus lailensis Belousov, 1992
Male adult
Laila Mt.
Copyright © Kirill Makarov
Carabus (Neoplectes) daphnis tsekuricus Belousov, 1992
Male adult
LEFT: Lakhashuri River valley; RIGHT: Tzikuri Mt. Scale bar 10.0 mm.
Copyright © Kirill Makarov
Carabus (Tribax) lailensis stefaniianus Deuve, 2018 Holotype
Male adult
Géorgie, Basse-Svanétie, au nord de Lentekhi, versant sud de la chaîne de Svanétie, 1000 m., Th. Deuve et A. Matocq, juin 2016. (MNHN)
Copyright © Thierry Deuve
Carabus (Neoplectes) daphnis boulbenellus Deuve et Stéfani, 2017 Holotype
Male adult
Géorgie, Basse-Ratcha, à l’est de Nikortsminda, Skhartali, 1200-1300 m., Th. Deuve, P. Stéfani leg. (MNHN).
Copyright © Thierry Deuve
Carabus (Neoplectes) komarowi mestiaicus Deuve, 2016 Holotype
Female adult
Géorgie, Haute Svanétie, 10 km au nord de Mestia, vers les lacs de Koruldi, env. 2500 mètres, in coll. A. Wřzecionko, à Horní Sucha (République Tchèque).
Copyright © Thierry Deuve
Carabus (Neoplectes) martviliensis mzekalae Fominykh, Zamotajlov et Titarenko, 2019 Paratype
Male+Female adult
Imereti, left bank of river Semi near Melouri, 475 m, 19.V–12.VII.2019, D. Fominykh, A. Zubov, B. Dubinin & D. Safronov leg.
Copyright © Kirill Makarov
Carabus (Neoplectes) martviliensis mzekalae Fominykh, Zamotajlov et Titarenko, 2019 Paratype
Male+Female adult
Imereti, left bank of river Tskhenistskali near Dzedzileti, 620 m, 16.V–13.VII.2019, D. Fominykh, A. Zubov, B. Dubinin & D. Safronov leg.
Copyright © Kirill Makarov
Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
Male+Female adult
Imereti, S slopes of Askhi plateau near Zeda-Gordi, NE slope of Mt. Gulkuna, 980 m, 20.V–14.VII.2019, D. Fominykh & A. Zubov leg.
Copyright © Kirill Makarov
Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
Male+Female adult
Samegrelo, Doberazeni vic., left bank of river Tekhuri, 500 m, 16.IV–31.V.2017, D. Fominykh leg.
Copyright © Kirill Makarov
Carabus (Neoplectes) starckianus theseus Briansky, 1910
Male adult
Russia, Krasnodar Territory, Tuapse district, env. vill. Psebe, left bank of the Psebe River, hornbeam + oak + walnut, h = 146 m, 03.05-01.07.2014, leg. I.A. Solodovnikov, S.V. Solodovnikova, S.M. Pavlyuchuk
Copyright © Igor Solodovnikov
Carabus (Neoplectes) starckianus theseus Briansky, 1910
Female adult
Russia, Krasnodar Territory, Tuapse district, env. vill. Psebe, left bank of the Psebe River, hornbeam + oak + walnut, h = 146 m, 03.05-01.07.2014, leg. I.A. Solodovnikov, S.V. Solodovnikova, S.M. Pavlyuchuk
Copyright © Igor Solodovnikov
Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
Female adult
Russia, Krasnodar Territory, West-Caucasus, Apsheronsky district, Guam ridge, env. Mezmay village, by the road, beech, oak, blackberry, h = 1200-1250 m, 03.IV-14.VI.2020, leg. S.M. Pavlyuchuk, A.P. Tatulyan
Copyright © Igor Solodovnikov
Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
Female adult
Russia, Krasnodar Territory, West-Caucasus, Apsheronsky district, Guam ridge, env. Mezmay village, by the road, beech, oak, blackberry, h = 1200-1250 m, 03.IV-14.VI.2020, leg. S.M. Pavlyuchuk, A.P. Tatulyan
Copyright © Igor Solodovnikov
Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
Male adult
Russia, Krasnodar Territory, West-Caucasus, Apsheronsky district, Guam ridge, env. Mezmay village, by the road, beech, oak, blackberry, h = 1200-1250 m, 03.IV-14.VI.2020, leg. S.M. Pavlyuchuk, A.P. Tatulyan
Copyright © Igor Solodovnikov
Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
Male adult
Russia, Krasnodar Territory, West-Caucasus, Apsheronsky district, Guam ridge, env. Mezmay village, by the road, beech, oak, blackberry, h = 1200-1250 m, 03.IV-14.VI.2020, leg. S.M. Pavlyuchuk, A.P. Tatulyan
Copyright © Igor Solodovnikov
Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988 Paratype
Male adult
Krasnodar area, Babuk-Aul, 19.06.1983 , leg. A. Miroshnikov
Copyright © Igor Solodovnikov
Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
Female adult
W Caucasus, NW slope Aibga Mt.R. vall. riv. Galion, h = 650 m, Fagus, Alnus viridis forest, 18-24.VI.1999, I.A. Solodovnikov leg.
Copyright © Igor Solodovnikov
Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
Male adult
W Caucasus, Aibga Mt.R., 02.VIII.1994, A.S. Zamotailov leg.
Copyright © Igor Solodovnikov
Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
Male adult
Russia, W Caucasus, Krasnaya Polyana, Atchischkho Mt.R., h = 1200 m, 10.06.1995, N.Yu. Pichugin leg.
Copyright © Igor Solodovnikov
Carabus (Neoplectes) starckianus adelaidae Starck, 1889
Female adult
Russia, Krasnodar area, Adler, Ader distr., right bank vall. Riv. Msymta, near vill. Chvesipse, h = 800 m, 06.VII.1999, B. Saratovskikh leg.
Copyright © Igor Solodovnikov
Carabus (Neoplectes) starckianus adelaidae Starck, 1889
Female adult
Russia, Krasnodar area, Adler, Ader distr., right bank vall. Riv. Msymta, near vill. Chvesipse (Fagus+Quercus+Castanea forest) [Sochi national park], h = 720 m, 03.V-08.VII.2016, I.A. Solodovnikov, S.V.Solodovnikova, V.M.Kotsur leg.
Copyright © Igor Solodovnikov
Carabus (Neoplectes) starckianus adelaidae Starck, 1889
Female adult
Russia, Krasnodar area, Adler, Ader distr., right bank vall. Riv. Msymta, near vill. Chvesipse (Fagus+Quercus+Castanea forest) [Sochi national park], h = 720 m, 03.V-08.VII.2016, I.A. Solodovnikov, S.V.Solodovnikova, V.M.Kotsur leg.
Copyright © Igor Solodovnikov
Carabus (Neoplectes) starckianus adelaidae Starck, 1889
Male adult
Russia, Krasnodar area, Adler, Ader distr., right bank vall. Riv. Msymta, near vill. Chvesipse (Fagus+Quercus+Castanea forest) [Sochi national park], h = 720 m, 03.V-08.VII.2016, I.A. Solodovnikov, S.V.Solodovnikova, V.M.Kotsur leg.
Copyright © Igor Solodovnikov
Carabus (Neoplectes) ibericus lafertei Chaudoir, 1846
Female adult
SC-Georgia, Samtskhe–Javakheti, Meskheti Mt Range, Borjomi, Likani env., h=897 m, 41°50.125'N 43°19.905'E, VI.2018, D. Prunier leg., L=37 mm
Copyright © Ruslan Panin
Carabus (Neoplectes) edithae miloslavskii Obydov, 2010 Holotype Paratype
Male+Female adult
Holotype: Caucasus centr., Archyz, zona subalp., 15.VI.1967, A. Bogachev leg; Paratype: KarachaiCherkessian Region, Zagedan Mt. Ridge, 2500 m., 6-7.VII.1994, Melnik leg.
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Carabus (Neoplectes) jason Semenov, 1898
Male adult
Abhazia, Tchkhaltinskiy (Abkhazskiy) Mt.R., 20-26.VII.2022 leg. D. Fominykh
Copyright © Igor Solodovnikov
Carabus (Tribax) zubovi mananauriensis Deuve, 2022 Holotype
Male adult
Géorgie, Svanétie, 20 km à l’ouest de Lentekhi, 6 km à l’ouest de Mananaouri, 1350 m (A. Wřzecionko leg.), in coll. A. Wřzecionko, à Horni Sucha (République Tchèque).
Copyright © Thierry Deuve
Carabus (Neoplectes) zolotarevi Zamotajlov, 1988
Female adult
Carabus (Archiplectes) zolotarevi zolotarevi Zamotajlov, 1988 Russia: Krasnodar reg., SW slopes of Malye Balkany Mt.R., env. Allous pass, h = 1960 m, 08-18.VII.2023 leg. D. Fominykh
Copyright © Igor Solodovnikov
Carabus (Neoplectes) zolotarevi Zamotajlov, 1988
Male adult
Carabus (Archiplectes) zolotarevi zolotarevi Zamotajlov, 1988 Russia: Krasnodar reg., SW slopes of Malye Balkany Mt.R., env. Allous pass, h = 1960 m, 08-18.VII.2023 leg. D. Fominykh
Copyright © Igor Solodovnikov
Carabus (Neoplectes) komarowi komarowi Reitter, 1882 Lectotype Paralectotypes
Male+Female adult
LT - female «Kaukasus Swanetien Leder Reitter » , «Komarowi m.», «Roeschke C.», « MA.INS.COLE.194.1», «Lectotype», «Carabus komarowi Reitter, Lectotype, Th. Deuve des. 2024», ex coll. Vogt, in coll. Naturalis Biodiversity Center, Leyde (Pays-Bas). Image from Deuve, 2024
Copyright © Thierry Deuve
Carabus (Neoplectes) komarowi mutabilis Reitter, 1896 Lectotype Paralectotypes
Female adult
LT, left, female: «Kaukas Leder», «Komarowi v. mutabilis m.», «Roeschke C. », « Collectie C. & O. Vogt, Acq. 1960», «Plectes Komarowi mutabilis Reitter, 1896, ZMAN type COLE.0264.1», «Lectotype», «Carabus komarowi mutabilis Reitter, Lectotype, Th. Deuve des. 2024 » , ex coll. Vogt, in coll. Naturalis Biodiversity Center, Leyde (Pays-Bas). Image from Deuve, 2024
Copyright © Thierry Deuve
Carabus (Neoplectes) compressus rebellis Reitter, 1895 Lectotype
Male adult
Image from Deuve, 2024
Copyright © Thierry Deuve
Carabus (Tibax) komarowi antoni Deuve, 2024 Holotype
Male adult
«Kaukasus Swanetien Leder Reitter», ex coll. M. Maindron, ex coll. G. Babault, in coll. Muséum National d’Histoire Naturelle, Paris (France).
Copyright © Thierry Deuve
Carabus (Tribax) compressus pseudorebellis Deuve, 2024 Holotype
Male adult
Géorgie, Basse-Svanétie, 12 km au nord-est de Lentekhi, Mont Goldash, vallée de la Cholshoura, Durashi (« Dvashi »), env. 1150 mètres (W. Heinz, 1-4 juillet 2019), in coll. Muséum Stuttgart (Germany).
Copyright © Thierry Deuve
Carabus (Neoplectes) compressus zubovi Fominykh, Zamotajlov et Titarenko, 2018 Paratype
Female adult
S slopes of Svaneti Mt. Range near Lentekhi Town, S slopes of Goldash Mt., 42°50'N, 42°45'E, 2200–2600 m, 18.VI–13.VII.2017 (D. Fominykh). Image from Fominykh, Zamotajlov et Titarenko, 2018
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Carabus (Neoplectes) compressus zubovi Fominykh, Zamotajlov et Titarenko, 2018 Paratype
Male+Female adult
5-6: Paki Vill. environs; 7-8: Mannanauri Vill. environs. Scale bar 10.0 mm. Image from Fominykh, Zamotajlov et Titarenko, 2018
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Carabus (Neoplectes) ibericus lafertei Chaudoir, 1846
Male adult
Georgia, Lesser Caucasus, Imereti prov., Meskheti Mt Rng, N slope, Zekari Pass h~2150 -2250 m alpine meadows 25.VII.2021, E.Rujan leg./det. R. Panin, L=33 mm
Copyright © Ruslan Panin
Carabus (Neoplectes) zolotarevi dzhugensis Zamotajlov, 1988
Male adult
Russia, Adygea, E slopes of Dzuga Mt., left bank of Urushten Riv., near Meteostation “Dzhuga”, h = 1970 m, 21.VII-04.IX.2016, leg. Yu. Chumachenko
Copyright © Igor Solodovnikov
Carabus (Neoplectes) zolotarevi dzhugensis Zamotajlov, 1988
Female adult
Russia, Adygea, E slopes of Dzuga Mt., left bank of Urushten Riv., near Meteostation “Dzhuga”, h = 1970 m, 21.VII-04.IX.2016, leg. Yu. Chumachenko
Copyright © Igor Solodovnikov
Carabus (Neoplectes) compressus lailensis Belousov, 1992
Male adult
Carabus (Archiplectes) lailensis Belousov, 1992 (TOPOTYPUS) Georgia, Svaneti, S slopes of Svanetian Mts. rng., W slopes of Laila-lakheli Mt., upper of Skirili river, alpin zone, h = 3000 m, 27-30.VI.2014, leg. D. Fominykh, A. Zubov
Copyright © Igor Solodovnikov
Carabus (Neoplectes) compressus lailensis Belousov, 1992
Male adult
Carabus (Archiplectes) lailensis Belousov, 1992 (TOPOTYPUS) Georgia, Svaneti, S slopes of Svanetian Mts. rng., W slopes of Laila-lakheli Mt., upper of Skirili river, alpin zone, h = 3000 m, 27-30.VI.2014, leg. D. Fominykh, A. Zubov
Copyright © Igor Solodovnikov
Carabus (Neoplectes) protensus protensoides Novotny & Vorisek, 1988
Male adult
W Georgia, Martvili distr., W slope of Askhi Plateau h=1200-1800 m, 18-19.VI.2005, Shumov leg., L=33 mm
Copyright © Ruslan Panin
Carabus (Neoplectes) satyrus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014
Male adult
Abkhasia: N slope Bzybian Mt.R, vall riv. Reshevie (h =700-800 m) (Topotypus)
Copyright © Igor Solodovnikov
Carabus (Neoplectes) satyrus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014
Male adult
Abkhasia: N slope Bzybian Mt.R, vall riv. Reshevie (h =700-800 m) (Topotypus)
Copyright © Igor Solodovnikov
Carabus (Neoplectes) protensus protensus Schaum, 1864
Female adult
Georgia, Caucasus, Imereti reg., Dzedzileti env, h=500-600m, 18-24.VI.2024, I.Uspenskyi leg., L=36.5 mm
Copyright © Ruslan Panin
Carabus (Neoplectes) protensus protensus Schaum, 1864
Male adult
Georgia, Caucasus, Imereti reg., Dzedzileti env, h=500-600m, 18-24.VI.2024, I.Uspenskyi leg., L=32 mm
Copyright © Ruslan Panin
Carabus (Neoplectes) titarenkoi kozlovantoni Fominykh, Zamotajlov & Titarenko, 2025 Paratype
Female adult
Georgia, Ratcha, right bank of river Gamrula, near Gomi vill., 1385 m, 3.V–12.VII.2018, pitfall traps, D. Fominykh, B. Dubinin, D. Safronov leg. Image from Fominykh, Zamotajlov & Titarenko, 2025
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Carabus (Neoplectes) titarenkoi kozlovantoni Fominykh, Zamotajlov & Titarenko, 2025 Paratype
Male adult
Georgia, Ratcha, right bank of river Gamrula, near Gomi vill., 1385 m, 3.V–12.VII.2018, pitfall traps, D. Fominykh, B. Dubinin, D. Safronov leg. Image from Fominykh, Zamotajlov & Titarenko, 2025
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Carabus (Neoplectes) titarenkoi kozlovantoni Fominykh, Zamotajlov & Titarenko, 2025 Paratype
Male adult
Georgia, Racha, Oni town env., near Ghari vill., btw rivers Rioni and Gharula., 870 m, 13.V–22.VI.2024, pitfall traps, D. Fominykh, A. Tatulyan, Ya. Talibov leg. Image from Fominykh, Zamotajlov & Titarenko, 2025
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Carabus (Neoplectes) titarenkoi kozlovantoni Fominykh, Zamotajlov & Titarenko, 2025 Paratype
Female adult
Georgia, Racha, Oni town env., near Ghari vill., btw rivers Rioni and Gharula., 870 m, 13.V–22.VI.2024, pitfall traps, D. Fominykh, A. Tatulyan, Ya. Talibov leg. Image from Fominykh, Zamotajlov & Titarenko, 2025
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Carabus (Neoplectes) titarenkoi titarenkoi Zamotajlov & Fominykh, 2014
Male adult
Okureshi env. Image from Fominykh, Zamotajlov & Titarenko, 2025
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Carabus (Neoplectes) ibericus titarenkoi Zamotajlov & Fominykh, 2014
Female adult
Okureshi env. Image from Fominykh, Zamotajlov & Titarenko, 2025
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Carabus (Neoplectes) titarenkoi djanoliensis Fominykh & Zamotajlov, 2016
Male adult
Zubi env. Image from Fominykh, Zamotajlov & Titarenko, 2025
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Carabus (Neoplectes) titarenkoi djanoliensis Fominykh & Zamotajlov, 2016
Female adult
Zubi env. Image from Fominykh, Zamotajlov & Titarenko, 2025
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Carabus (Neoplectes) szekelyi tatulyani Fominykh, Zamotajlov & Titarenko, 2025 Paratype
Male adult
Kontanto env. Image from Fominykh, Zamotajlov & Titarenko, 2025
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Carabus (Neoplectes) szekelyi tatulyani Fominykh, Zamotajlov & Titarenko, 2025 Paratype
Female adult
Kontanto env. Image from Fominykh, Zamotajlov & Titarenko, 2025
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Carabus (Neoplectes) szekelyi tatulyani Fominykh, Zamotajlov & Titarenko, 2025 Paratype
Male adult
Tli env. Image from Fominykh, Zamotajlov & Titarenko, 2025
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Carabus (Neoplectes) szekelyi tatulyani Fominykh, Zamotajlov & Titarenko, 2025 Paratype
Female adult
Tli env. Image from Fominykh, Zamotajlov & Titarenko, 2025
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Carabus (Neoplectes) szekelyi szekelyi Retezar, 2011
Male adult
Shaori env. Image from Fominykh, Zamotajlov & Titarenko, 2025
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Carabus (Neoplectes) szekelyi szekelyi Retezar, 2011
Female adult
Shaori env. Image from Fominykh, Zamotajlov & Titarenko, 2025
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Carabus (Neoplectes) szekelyi szekelyi Retezar, 2011
Female adult
Kedisubani env. Image from Fominykh, Zamotajlov & Titarenko, 2025
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Carabus (Neoplectes) ibericus Fischer, 1823 (as szekelyi szekelyi)
Male adult
Kedisubani env. Image from Fominykh, Zamotajlov & Titarenko, 2025
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Carabus (Neoplectes) ibericus pseudoszekelyi Fominykh, Zamotajlov & Titarenko, 2025 Paratype
Female adult
Shkmeri env. Image from Fominykh, Zamotajlov & Titarenko, 2025
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Carabus (Neoplectes) ibericus pseudoszekelyi Fominykh, Zamotajlov & Titarenko, 2025 Paratype
Male adult
Shkmeri env. Image from Fominykh, Zamotajlov & Titarenko, 2025
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Carabus (Neoplectes) ibericus ibericus
Female adult
Ulumbo env. Image from Fominykh, Zamotajlov & Titarenko, 2025
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Carabus (Neoplectes) ibericus ibericus
Male adult
Ulumbo env. Image from Fominykh, Zamotajlov & Titarenko, 2025
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Carabus (Neoplectes) mellyi boulbenorum Deuve, 2015 Holotype
Male adult
Géorgie, au nord de Lentekhi, 1000 m. R. et A. Boulben (MNHN)
Copyright © Thierry Deuve
Carabus (Neoplectes) ibericus montreuilianus Deuve, 2014
Male adult
Haut-plateau Arméniano-Géorgien, entre Tsalka et le lac de Paravani : col de Tukmatash.
Copyright © Thierry Deuve
Carabus (Neoplectes) ibericus montreuilianus Deuve, 2014
Male adult
SC Georgia, Lesser Caucasus, Samsari Range, Mt Egoisar h=2100-2200m, 41.651528, 43.770083,12-16.VI.2025, I.Uspenskyi leg., 35 mm
Copyright © Ruslan Panin
Carabus (Neoplectes) compressus compressus Chaudoir, 1846 (? rebellis )
Male adult
Georgia, Caucasus, S of Sasashi vill., Lechkhumi Range N slope h=2430 m, alpine meadows, 242.759167, 42.973889, 20-21.VII.2025, I.Uspenskyi leg, L=26 mm
Copyright © Ruslan Panin -
Neoplesius taxon page (34 photos)
Carabus (Pagocarabus) wrzecionkoianus Deuve, 2007 Holotype Paratype
Male adult
Copyright © Boleslav Brezina
Carabus (Pagocarabus) wagae folwarcznyi Deuve, 1997 Paratype
Male adult
male, 21 mm
Copyright © Boleslav Brezina
Carabus (Pagocarabus) ? wagae Fairmaire, 1882
Male adult
China, E Tibet, pas Dongdola 5000 m, 4.7.1995
Copyright © Alexander Anichtchenko
Carabus (Pagocarabus) piceophilus piceophilus Deuve, 1994
Male adult
male, 22mm
Copyright © Boleslav Brezina
Carabus (Pagocarabus) paulusi Kalab, 1995
Male adult
China, E-Tibet, Chakla, 5000m, 100 km
N of Lhasa, 4.6.1997 Wrzecionko A. leg.
Copyright © Alexander Anichtchenko
Carabus (Pagocarabus) nanschanicus globulipennis Deuve, 2006 Holotype
Male adult
Copyright © Boleslav Brezina
Carabus (Pagocarabus) lama yajiangensis Deuve, 1994
Male adult
male, 23mm
Copyright © Boleslav Brezina
Carabus (Pagocarabus) lama daochengensis Deuve, 1994
Male adult
male, 21mm
Copyright © Boleslav Brezina
Carabus (Pagocarabus) borodini Heinz, 1996
Male adult
China, C Tibet, Mila 5000m, 150 km E of Lhasa 6.6.1997 Wrzecionko A. leg.
Copyright © Alexander Anichtchenko
Carabus (Eocechenus) takashimai Deuve & Imura, 1993
Female adult
female, 20mm
Copyright © Boleslav Brezina
Carabus (Eocechenus) takashimai Deuve & Imura, 1993
Male adult
male, 20mm
Copyright © Boleslav Brezina
Carabus (Eocechenus) mouthiezianoides Deuve & Kalab, 1992 Paratype
Male adult
male, 18mm
Copyright © Boleslav Brezina
Carabus (Eocechenus) leptoplesioides yanjinganus Imura & Mizusawa, 1998 Paratype
Female adult
female, 20mm
Copyright © Boleslav Brezina
Carabus (Eocechenus) leptoplesioides yanjinganus Imura & Mizusawa, 1998 Paratype
Male adult
male, 20mm
Copyright © Boleslav Brezina
Carabus (Eocechenus) kaznakovi kaznakovi (as zadoiensis Lassalle, 2003)
Female adult
female, 19mm
Copyright © Boleslav Brezina
Carabus (Eocechenus) kaznakovi nangqenicus Deuve, 1995 Paratype
Male adult
Copyright © Boleslav Brezina
Carabus (Eocechenus) kaznakovi kaznakovi Semenov & Znojko, 1932
Male adult
L=17 mm
Copyright © Boleslav Brezina
Carabus (Eocechenus) kaznakovi angenensis Deuve, 1996
Male adult
male, 17mm
Copyright © Boleslav Brezina
Carabus (Eocechenus) kaznakovi graziosii Deuve, 1995
Female adult
female, 16 mm
Copyright © Boleslav Brezina
Carabus (Pagocarabus) nanschanicus edacis Cavazzuti, 2006 Holotype
Male adult
China, Gansu, Str. Lintan - Diebu, 2. Pass & Bergmassiv, 3900m, 30.V.-19.VI.2009 (in coll. Puchner)<br /> 22,5 mm<br /> photo from carabus-online.de
Copyright ©
Carabus (Pagocarabus) nanschanicus edacis Cavazzuti, 2006 Paratype
Female adult
China, Gansu, Str. Lintan - Diebu, 2. Pass & Bergmassiv, 3900m, 30.V.-19.VI.2009 (in coll. Puchner) <br /> 26 mm<br /> photo from carabus-online.de
Copyright ©
Carabus (Pagocarabus) alexpuchneri Kleinfeld & Puchner, 2011 Holotype
Male adult
China, Sichuan, Straße Xinlong - Litang, Umgebung Heping (30°33'/100°20'), 2870m, 1.-16.VI.2010, leg. et in coll. Puchner
22,5 mm
Copyright © Frank Kleinfeld
Carabus (Pagocarabus) alexpuchneri Kleinfeld & Puchner, 2011 Paratype
Female adult
China, Sichuan, Straße Xinlong - Litang, Umgebung Heping (30°33'/100°20'), 2870m, 1.-16.VI.2010, leg. et in coll. Puchner
24 mm
Copyright © Frank Kleinfeld
Carabus (Pagocarabus) wagae wagae Fairmaire, 1882 Holotype
Female adult
Thibet MNHN) http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec146
Copyright © - - -
Carabus (Pagocarabus) lama achaicus Deuve & Kaláb, 2018 Holotype
Male adult
Chine, Ouest-Sichuan, col à 9 km au nord-ouest d’Acha, zone sud du col, 4700 mètres, 31º06’N/99º29’E (pelouses alpines, éboulis), in coll. J. Kaláb, à Jinačovice (République Tchèque).
Copyright © Thierry Deuve
Carabus (Neoplesius) nanschanicus heishuicus Deuve, 2016 Holotype
Male adult
Chine, Sichuan central, Heishui Xian, montagnes à 14 km au S-SSE de Heishui, 4200-4400m, 31º56´N/102º59´E (prairies alpines, éboulis), in coll. J. Kaláb, à Jinačovice (République Tchèque).
Copyright © Thierry Deuve
Carabus (Neoplesius) lama latitatus Deuve & Kalab, 2019 Holotype
Male adult
Chine, Sichuan occidental, montagnes à 17 km au nord-nord-ouest de Ganzi (= Garzê), 31º46’N/99º56’E, 4700-4800 mètres (prairies alpines et éboulis), in coll. J. Kaláb, à Jinačovice (République Tchèque).
Copyright © Thierry Deuve
Carabus (Neoplesius) lama lapidiphilus Deuve & Kalab, 2019 Holotype
Male adult
Chine, Sichuan occidental, montagnes à 15 km au nord-est de Baiyü, 31º19’N/98º55’E, 4750 mètres (prairies alpines et éboulis), in coll. J. Kaláb, à Jinačovice (République Tchèque).
Copyright © Thierry Deuve
Carabus (Neoplesius) lama paucipunctatus Deuve & Kalab, 2019 Holotype
Male adult
Chine, Sichuan occidental, montagnes à 20 km au sud-sud-est de Batang, 29º50’N/99º11’E, 4700-4800 mètres (prairies alpines et éboulis), in coll. J. Kaláb, à Jinačovice (République Tchèque).
Copyright © Thierry Deuve
Carabus (Neoplesius) nanschanicus nocticolor Deuve & Kalab, 1992
Female adult
Quinghai-Gansu border, Dalijia pass, 50km W Linxia town, 3800m 20.7.2019, V. Patrikeev leg.
Copyright © Alexander Anichtchenko
Carabus (Neoplesius) lama lama Semenov, 1898
Male adult
NW Sichuan, Chola mts, Maniganggo vill., 4600 m 15.6.2015, V. Patrikeev leg (cAA). det. Shi Hongliang, 2020
Copyright © Alexander Anichtchenko
Carabus (Neoplesius) zhangjieae Deuve & Li, 2020 Holotype
Male adult
Chine, Sichuan occidental, Danba Xian, env. 7 km à l’est de Danba, rive gauche de la rivière Xiaojin, Jikaiyi, env. 2300 mètres, 30°53’-101°55’E, Zhang Jie leg., 23 septembre 2019; 18 mm; Image from Deuve & Li, 2020
Copyright © - - -
Carabus (Neoplesius) piceophilus zheduo Cavazzuti, 2009 Holotype
Male adult
Passo a Ovest di Kangding, Mass. Zheduo Shan, 30°04'28"N, 101°48'16"E, 4400-4550 m, Sichuan occ., Cina.
Copyright © - - -
Carabus (Neoplesius) takashimai qamdoensis Deuve, Liu & Liang, 2024 Holotype
Female adult
Chine, Tibet, “Qamdo” (Chaodong Zhu leg., 8.VIII.1997), in IOZ. Image from Deuve et al 2024
Copyright © - - -
Carabus (Neoplesius) keluoensis Kleinfeld & Reuter, 2025 Holotype Paratype
Male+Female adult
China, W. Sichuan, Garzȇ pref., Dȇgȇ, 15 km west of Keluodong, 5 km nord Zama monastery, 31°58’N/98°33’E, 4600-4700 m, 30/VI-2/VII/2024, alpine valley, leg. et in coll. Reuter
Copyright © - - -
Carabus (Neoplesius) wagae degeicola Deuve, 1996
Male adult
China, W. Sichuan, pass 9 km W. Babang – S. env. (road to Dȇgȇ), 31°39’N/98°43’E, 4600-4800 m, 5-9/VII/2018, leg. Kalab (in coll. H. Schütze). 20 mm. Image from Kleinfeld & Reuter, 2025
Copyright © - - -
-
Nesaeocarabus taxon page (9 photos)
Carabus (Nesaeocarabus) coarctatus Brulle, 1838
Female adult
Canary isl., Las Palmas, Las Mesas 20.2.90
Copyright © Alexander Anichtchenko
Carabus (Nesaeocarabus) abbreviatus Brulle, 1835
Female adult
Spain, Canary Islands,
Tenerife, Ft de Esperanza 1360m, TF-24, 29.III.2007, D.Prunier leg.
female, 22mm
Copyright © Ruslan Panin
Carabus (Nesaeocarabus) abbreviatus Brulle, 1835
Male adult
Canarias Isle, NC Tenerife; 22mm
Copyright © Aleš Sedláček
Carabus (Nesaeocarabus) faustus cabrerai Enderlein, 1929
Male adult
Spain, Canarias Isl.,Teneriffe: NW-part:Teno Massif, Erjos env., h=1000m, 18-21.III.2013, V.Slovak leg.
L=23 mm
Copyright © Ruslan Panin -
Ohomopterus taxon page (38 photos)
Carabus (Ohomopterus) yamato Nakane, 1953
Female adult
Japan, Ohkubo, 740-770m, Shirakawa-mura
Ohno-gun (Gifu) 20-25.6.80 Ishikawa leg.
Copyright © Alexander Anichtchenko
Carabus (Ohomopterus) yaconinus yamaokai Ishikawa & Kubota, 1994
Female adult
Japan, Ehime pref., Taisanji, 14.5.2002, Matsuyama city, T. Kurihara leg. (cAA)
Copyright © Alexander Anichtchenko
Carabus (Ohomopterus) yaconinus yaconinus Bates, 1873
Male adult
Japan, Kagawa pref, Shin-ike
Awai town, Kannonji city, 4.3.1993
male, 28 mm
Copyright © Alexander Anichtchenko
Carabus (Ohomopterus) maiyasanus maiyasanus Bates, 1873
Female adult
Japan, Shiga pref.,
Lake Yogo 18.8.1995 Shimizu leg.
24 mm
Copyright © Alexander Anichtchenko
Carabus (Ohomopterus) japonicus tsushimae Breuning, 1932
Male adult
Japan, Tsushima isl., Nagasaki pref., Hiratake, Kamiagata town, 19.6.2002 T. Kurihara leg. 23 mm
Copyright © Alexander Anichtchenko
Carabus (Ohomopterus) insulicola insulicola Chaudoir, 1869
Male adult
Japan, Yamanashi pref., Kodachi, Kawaguchiko-cho, Minamitusru distr., 2.IX.2000 T. Kurihara leg. (cAA). 29 mm
Copyright © Alexander Anichtchenko
Carabus (Ohomopterus) insulicola insulicola Chaudoir, 1869
Female adult
15.VII.1998 JAPAN, Ichihara, env. Chiba
Copyright © Nil Belonogov
Carabus (Ohomopterus) dehaanii Chaudoir, 1848
Male adult
Japan, Hyogo pref., Ueda-rindo, Mihara, 4.3.93 N. Sano leg.
Copyright © Alexander Anichtchenko
Carabus (Ohomopterus) arrowianus murakii Ishikawa & Kubota, 1984
Male adult
Sho, alt 3m, Futami-T, Watarai-Co. Mie pref. Japan, 17.5.1987 Y. Imura leg. (cAA) 27 mm
Copyright © Alexander Anichtchenko
Carabus (Ohomopterus) arrowianus arrowianus Breuning, 1934
Male adult
Japan, Shizuoka pref., Mt. Mitake, Inasa town 18.10.1998 T. Kurihara leg.
Copyright © Alexander Anichtchenko
Carabus (Ohomopterus) albrechti esakianus Nakane, 1961
Male adult
Japan, Tokyo pref, Mt. Takao, Nachioji City, 11.8.1990 K. Kurihara leg. (cAA)
Copyright © Alexander Anichtchenko
Carabus (Ohomopterus) lewisianus lewisianus Breuning, 1932
Male adult
Japan, Yamanashi pref., Koganesawa-rindo, Otsuki city, 13.6.1990 K. Kurihara leg. (cAA). 20 mm
Copyright © Alexander Anichtchenko
Carabus (Ohomopterus) albrechti albrechti Morawitz, 1862
Female adult
Japan, S-Hokkaido:Oshima Peninsula,Pass btw Otoshibe - Assabu, h~250-300 m, 10-20.VII.2010, 42:01’53’Т; 140:22’09’E, I.Rapuzzi & L.Caldon leg.
Copyright © Ruslan Panin
Carabus (Ohomopterus) japonicus japonicus Motschulsky, 1857
Male adult
S Japan, Kyushu, Kumamoto Pref., Izumi viil., h=600 m,VII.2001, S. Remond leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Ohomopterus) yamato kitai (Imura, 2004)
Male adult
Japan, S Honshu, Mie Pref., Hakusan, VII.2001, S. Remond leg.<br /> L=21 mm
Copyright © Ruslan Panin
Carabus (Ohomopterus) kaempferi Thomson, 1875 Holotype
Male adult
Image courtesy of Biological Museum, Lund University
Copyright © - - -
Carabus (Ohomopterus) dehaanii dehaanii Chaudoir, 1848
Female adult
Japan, S Honshu, Hyogo Pref., Nishiwaki city, 20.VI.2010, Shinichi S. leg., L=37 mm
Copyright © Ruslan Panin
Carabus (Ohomopterus) dehaanii dehaanii Chaudoir, 1848
Male adult
Japan, S Honshu, Mie Pref., Inabe city, 21.II.2021, Shinichi S. leg., L=33 mm
Copyright © Ruslan Panin
Carabus (Ohomopterus) albrechti okumurai (Ishikawa, 1966)
Male+Female adult
Mt. Daibosatzu (Yamanashi, Honshu) JAPON
Copyright © Alain Mollard
Carabus (Ohomopterus) albrechti tohokuensis Ishikawa, 1984
Male+Female adult
Hashimori Pass (Iwate) JAPON
Copyright © Alain Mollard
Carabus (Ohomopterus) albrechti freyi van Emden, 1932
Male+Female adult
Sado Island (Niigata) JAPON
Copyright © Alain Mollard
Carabus (Ohomopterus) arrowianus nakamurai (Ishikawa, 1966)
Female adult
Mt.Takamoriyama (Tida-City, Nagano) JAPON VI-1987
Copyright © Alain Mollard
Carabus (Ohomopterus) chugokuensis chugokuensis (Nakane, 1961)
Male+Female adult
Mt.Takasho (Shiman) JAPON VI-1987
Copyright © Alain Mollard
Carabus (Ohomopterus) japonicus japonicus Motschulsky, 1857
Male adult
MT. Narabara-san, Ryuokakami Tamagawa town, Imabari city, Ehime pref. Japan 20.6.2015 Y. Amano (cAA). det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko
Carabus (Ohomopterus) japonicus awajiensis Imura, Dejima & Mizusawa, 1993
Female adult
Mt. Senzan, Sumoto city, Awajishima Is., Hyogo, Japan 28.4.2015 Y. Amano leg. (cAA).
Copyright © Alexander Anichtchenko
Carabus (Ohomopterus) japonicus japonicus Motschulsky, 1857
Female adult
Fukuoka pref., Omuta city, Hagio, 16.1.2012, S. Nishimoto leg. (cAA)
Copyright © Alexander Anichtchenko
Carabus (Ohomopterus) lewisianus awakazusanus Ishikawa, 1981
Male adult
Mt. Iyogatake 280m, Tomiyama town, Awa Co., Chiba pref, 4.2.1996 K. Watanabe leg. (cAA). det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko
Carabus (Ohomopterus) albrechti Morawitz, 1862
Male adult
Horoman, Samani town, Samani Co., Hokkaido, 16.6.2015 T. Nakamura leg. (cAA). det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko
Carabus (Ohomopterus) insulicola nishikawai (Ishikawa, 1966)
Male adult
Yamada, Isumi city, Chiba pref. Japan 20.5.2007 K. Osozawa leg. (cAA). 28mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko
Carabus (Ohomopterus) insulicola insulicola Chaudoir, 1869
Male adult
Japan, Kiyose, Tokyo 20.1.1961 (cAA). 29 mm
Copyright © Alexander Anichtchenko -
Ophiocarabus taxon page (34 photos)
Carabus (Ophiocarabus) regeli Morawitz, 1886
Male adult
Kirgizia, E Terskey Ala-Too, Chon-Akshuu pass,
2800 m, 13.6.1999
Copyright © Alexander Anichtchenko
Carabus (Ophiocarabus) aeneolus aeneolus Morawitz, 1886
Male adult
photo M. Smirnov from www.zin.ru
Copyright © Maksim Smirnov
Carabus (Ophiocarabus) aeneolus aeneolus Morawitz, 1886
Female adult
SE KAZAKHSTAN,
Alma-Ata reg., Zailiysky Mts., h=3000m, B. Almaatinka riv. valley,01.VI.2001
Copyright © Nil Belonogov
Carabus (Ophiocarabus) aeneolus aeneolus Morawitz, 1886
Male+Female adult
KAZAKHSTAN,
Bolshoe Almaatinskoe Lake,30.VI.1992
female, 17mm
Copyright © Ruslan Panin
Carabus (Ophiocarabus) regeli obscurior Semenov, 1888
Male+Female adult
Kyrgyzstan, NE Tian-Shan, E. Terskey Alatau Mts. R.,
Kok-kia River, h=2800 м.
7-8.VI.1997 М. Danilevsky leg.
Body lenght 15-18 mm.
Copyright © Alexandr Safronov
Carabus (Ophiocarabus) ballionis Kraatz, 1879
Male adult
NW-China, Xinjiang prov., 40 km N Narat, h=1500, 12-12.VI.2004, Benes & Kaiser leg.
L=22 mm
Copyright © Ruslan Panin
Carabus (Ophiocarabus) successor koeksuensis Kabak, 2013 Paratype
Male adult
China, Xinjiang, Koeksu Bas., Tshon-Aidak, SSE Tekes, 42* 32' 47'' N / 82* 05' 56'' E -- 42* 32' 51'' N / 82* 05' 36'' E, 3330-3550 m, 11.07.2012, Kabak I.I. leg.
Copyright © Ilya Kabak
Carabus (Ophiocarabus) nilkiensis Kabak, 2014 Paratype
Male adult
China, Xinjiang, Boro-Horo Mt. R., S slope, Nilki River, 44°01'25''N / 82°46'00''E, 2030 m, 5.08.2013, I.I. Kabak leg.
Copyright © Ilya Kabak
Carabus (Ophiocarabus) nilkiensis Kabak, 2014 Paratype
Female adult
China, Xinjiang, Boro-Horo Mt. R., S slope, Nilki River, 44°01'25''N / 82°46'00''E, 2030 m, 5.08.2013, I.I. Kabak leg.
Copyright © Ilya Kabak
Carabus (Ophiocarabus) successor koeksuensis Kabak, 2013 Paratype
Female adult
China, Xinjiang, Koeksu Bas. SSE Tekes h~3330-3550 11.VII.2012 leg. I.Kabak L=16.5 mm
Copyright © Kirill Makarov
Carabus (Ophiocarabus) successor koeksuensis Kabak, 2013 Paratype
Male adult
China, Xinjiang, Koeksu Bas. SSE Tekes h~3330-3550 11.VII.2012 leg. I.Kabak L=16.0 mm
Copyright © Kirill Makarov
Carabus (Ophiocarabus) tekesensis Deuve & Tian, 2004
Female adult
China, Xinjiang, Narat Mts. NE Tshon-Kushei Korbulak Riv., h~2630 42°56'25"N 82°15'27"E 13.VII.2014 leg. I.Kabak L=19.0 mm
Copyright © Kirill Makarov
Carabus (Ophiocarabus) kurdaiensis Kabak Paratype
Female adult
China, Xinjiang, Narat Mt. Range, Ekiasha River, near Kurdai Pass, 42°54ʹ21″N / 82°28ʹ30″E, 3640 m, 26.07.2013 I.I. Kabak leg. L=16.5 mm
Copyright © Kirill Makarov
Carabus (Ophiocarabus) latiballioni Deuve, 1993
Male adult
China, Xinjiang, Narat Mt.R., S of Kurdelin forest, h~1760 43°07'50"N 82°50'51"E 30.VII.2013 leg. I.Kabak L=17.0 mm
Copyright © Kirill Makarov
Carabus (Ophiocarabus) latiballioni Deuve, 1993
Female adult
China, Xinjiang, Narat Mt.R., S of Kurdelin forest, h~1760 43°07'50"N 82°50'51"E 30.VII.2013 leg. I.Kabak L=22.0 mm
Copyright © Kirill Makarov
Carabus (Ophiocarabus) successor successor Reitter, 1896
Male adult
SE-Kazakhstan, Narynkol distr., 4 km SW Bayankol, h~2400-2500 m, 2-14.VI.2007, Shumov leg. L=17.5 mm
Copyright © Ruslan Panin
Carabus (Ophiocarabus) successor successor Reitter, 1896
Female adult
SE-Kazakhstan, Narynkol distr., 4 km SW Bayankol, h~2400-2500 m, 2-14.VI.2007, Shumov leg. L=17 mm
Copyright © Ruslan Panin
Carabus (Ophiocarabus) ernsti ulastaiensis Kabak, 2019 Paratype
Male adult
China, Xinjiang, S slopes of Boro-Horo Mt. R., ridge between Kaptshik and Zekku rivers, NE of Ulastai (= Wulasitai) Vill., alpine meadows, 43°56'07"N / 83°18'33"E – 43°56'22"N / 83°18'30"E, 3135–3280 m, 17.07.2017 leg I.I. Kabak L=16.5 mm
Copyright © Kirill Makarov
Carabus (Ophiocarabus) ernsti ulastaiensis Kabak, 2019 Paratype
Female adult
China, Xinjiang, S slopes of Boro-Horo Mt. R., ridge between Kaptshik and Zekku rivers, NE of Ulastai (= Wulasitai) Vill., alpine meadows, 43°56'07"N / 83°18'33"E – 43°56'22"N / 83°18'30"E, 3135–3280 m, 17.07.2017 leg I.I. Kabak L=17.0 mm
Copyright © Kirill Makarov
Carabus (Ophiocarabus) successor eoketmenicus Kabak, 2016 Holotype
Male adult
China, Xinjiang, Wusunshan Mt. Range, E part, NE of Tekes Town, 43°20ʹ26″N / 81°59ʹ48″E, 2285 m, 27.07.2016 (I.I. Kabak leg). (ZIN)
Copyright © Ilya Kabak
Carabus (Ophiocarabus) kurdaiensis Kabak, 2016 Paratype
Female adult
China, Xinjiang, Narat Mt. Range, Ekiasha River, near Kurdai Pass, 42°54ʹ21″N / 82°28ʹ30″E, 3640 m, 26.07.2013 (I.I. Kabak leg.).
Copyright © Ilya Kabak
Carabus (Ophiocarabus) ernsti ernsti Kabak, 2001
Male adult
Left from the locality in right tributary of the Saryk River; Right – from the locality in mountains to the West of the Saryk River.
Copyright © Ilya Kabak
Carabus (Ophiocarabus) ernsti ulastaiensis Kabak, 2019 Paratype
Male+Female adult
China, Xinjiang, S slopes of Boro-Horo Mt. R., ridge between Kaptshik and Zekku rivers, NE of Ulastai (= Wulasitai) Vill., alpine meadows, 43°56ʹ07ʺN / 83°18ʹ33ʺE – 43°56ʹ22ʺN / 83°18ʹ30ʺE, 3135–3280 m, 17.07.2017 (I.I. Kabak leg.)
Copyright © Ilya Kabak -
Oreocarabus taxon page (61 photos)
Carabus (Pachystus) hortensis hortensis Linne, 1758
Male adult
Germania, Oberfranken, Langenau
leg.det. U.Schmidt, 2002
Foto: U.Schmidt, 2006
Copyright © U.Schmidt
Carabus (Pachystus) hortensis preslii Dejean, 1830
Female adult
S Greece, Peloponnesos, 14.7.1987
female, 28 mm
Copyright © Alexander Anichtchenko
Carabus (Pachystus) hortensis preslii Dejean, 1830 (as jonicus)
Male adult
Greece, Korfu 16.8.1987
male, 25 mm
Copyright © Alexander Anichtchenko
Carabus (Pachystus) hortensis neumeyeri Schaum, 1856
Female adult
Italy, Basilicata, Serra Calvello (PZ)
1400 m, 7.2003 F. Angelini leg.
female, 25 mm
Copyright © Alexander Anichtchenko
Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847 (as guadalupensis) Paratype
Male adult
ESP - Caceres, S. Guadalupe,
800 m XI.2002
Copyright © Alexander Anichtchenko
Carabus (Oreocarabus) ghiliani La Ferte-Senectere, 1847
Male adult
Spain, Segovia, Sierra de Somosierra,
La Pinilla, 1800m 4.4.2005
male, 19 mm
Copyright © Alexander Anichtchenko
Carabus (Oreocarabus) luetgensi luetgensi s.str.
Male adult
Spain, Zamora, San Martin de Castaneda
10.5.1997
male, 20 mm
Copyright © Alexander Anichtchenko
Carabus (Oreocarabus) taibaiensis Kleinfeld, 2001 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Oreocarabus) latro minshanensis Deuve, 1987
Male adult
China. N Sishuan, Juizhalgou, 12-16.6.1995 Benes leg., 22 mm
Copyright © Alexander Anichtchenko
Carabus (Oreocarabus) laotse paralaotse Kleinfeld, 2000 Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Oreocarabus) mikhaili Deuve & Mourzine, 1997
Male adult
China, S Gansu, Min Shan, 2100m
SW Wudu, 1.8.2000 V.Patrikeev leg.
male, 20 mm
Copyright © Alexander Anichtchenko
Carabus (Pachystus) hortensis chalkidikensis Müller, 2009 Holotype Paratype
Male+Female adult
Griechenland, Makedonien, Chalkidike, Holomondas Berge, zwischen den Orten Amea und Taxiarhis, 932 m, N 40°27'861", E0 23O32'661" leg. et coll. A. Muller (Wien).
Copyright ©
Carabus (Oreocarabus) luetgensi getschmanni Lapouge, 1924
Male adult
NW-Spain, León Province, Acebo, h=1000m, V.2007, Pascal Stefani leg.
L=22 mm
Copyright © Ruslan Panin
Carabus (Oreocarabus) carinthiacus Sturm, 1815
Male adult
NE-Italy, Bolzano, Brenner, Zirog h=2000 m, 26.VII.1992, Manfred Egger leg.
L=20 mm
Copyright © Ruslan Panin
Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847
Male adult
WC Spain, Cáceres, Hervás, h=1000m, V.2007, Pascal Stefani leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Oreocarabus) latro minshanensis Deuve, 1987
Male adult
China, N-Sichuan, Jiuzhaigou env., 13.VI.2001,E. Kučera leg. L=23 mm
Copyright © Ruslan Panin
Carabus (Oreocarabus) carinthiacus Sturm, 1815
Female adult
NW Slovenia, Trnovski Gozd; 24 mm
Copyright © Aleš Sedláček
Carabus (Oreocarabus) luetgensi luetgensi
Female adult
Maetzu, Alava, Spain. det. A. Mollard. Copyright © Magellanes
Copyright © - - -
Carabus (Oreocarabus) luetgensi getschmanni Lapouge, 1924
Male adult
Pto.de Opakua, Alava, Spain. det. A. Mollard. Copyright © Magellanes
Copyright © - - -
Carabus (Oreocarabus) luetgensi pseudosteuarti Lapouge, 1924
Male adult
Sierra de Estrela, Guarda, Portugal. det. A. Mollard. Copyright © Magellanes
Copyright © - - -
Carabus (Oreocarabus) ghiliani La Ferte-Senectere, 1847
Male adult
Pto.de Navafria, Madrid, Spain. det.A.Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Oreocarabus) ghiliani (as negrei Breuning, 1966)
Male adult
Sierra de Gredos, Avila, Spain. det.A.Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Oreocarabus) luetgensi luetgensi (f. typ.)
Female adult
Sierra de Marao, Vila Real, Portugal; det. A. Mollard. Copyright © Magellanes
Copyright © - - -
Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847
Female adult
los Santos, Salamanque, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847
Male adult
Villamadana, Palencia, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Oreocarabus) guadarramus (as duerensis Raynaud, 1974)
Female adult
Burgo de Osma, Soria, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Oreocarabus) guadarramus (as duerensis Raynaud, 1974)
Male adult
Sierra de Cucalon, Teruel, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Oreocarabus) guadarramus (as andiaensis Forel & Leplat, 1998)
Male adult
Sierra de Andia, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Oreocarabus) guadarramus (as guadalupensis Mollard, 2003) Paratype
Male adult
S.de Guadalupe, Extramadure, Spain. det.A.Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Oreocarabus) guadarramus (as sagranus)
Female adult
Sierra de la Sagra, Granada, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Oreocarabus) guadarramus (as hirschfelderi Branes, 1987)
Male adult
Puerto Viejo, Salamanque, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -
Carabus (Oreocarabus) ghiliani La Ferte-Senectere, 1847
Female adult
C-Spain: Sierra de Guadarrama, Puerto de Cotos (Madrid), 20.IX.2018, F.Rodriguez leg., L=22 mm
Copyright © Ruslan Panin
Carabus (Oreocarabus) luetgensi pseudosteuarti Lapouge, 1924
Female adult
Portugal, Guarda, Sierra de Estrela h=800 m, 26.VI-4.VIII.2015, L=26 mm
Copyright © Ruslan Panin
Carabus (Oreocarabus) luetgensi pseudosteuarti Lapouge, 1924
Male adult
Portugal, Guarda, Sierra de Estrela h=800 m, 26.VI-4.VIII.2015, L=24 mm
Copyright © Ruslan Panin
Carabus (Pachystus) hortensis preslii Dejean, 1830
Male adult
S-Greece:, N Peloponnes, Erymanthos Reg., Kalentzi, 26.VI.2007, Manfred Egger leg. L=26 mm
Copyright © Ruslan Panin
Carabus (Hypsocarabus) qinlingensis micangicola Deuve, 2017 Holotype
Female adult
Chine, Sichuan, Mts Micang Shan, env. de Guangwushan, 1700 m (in coll. Andreï Gorodinski, Moscow).
Copyright © Thierry Deuve
Carabus (Pachystus) hortensis hortensis Linne, 1758
Female adult
Natural hybrid Carabus hortensis X Carabus granulatus. Belarus, Minsk reg., 8,7 km of Borisov, near vill. Lyubatovstchina, pine forest (Pinetum pleuroziosum), h = 213 m, 54.31934° N, 28.511486° E, 20-27.09.2018, leg. Klychenok Snezhana
Copyright © Igor Solodovnikov
Carabus (Oreocarabus) gemellatus Menetries, 1832
Female adult
Col de Gascar (Mazaderan), IRAN, VI-2000
Copyright © Alain Mollard
Carabus (Oreocarabus) tewoensis Deuve, 1992
Female adult
Tewo, (Gansu) ,CHINE, VI-2012
Copyright © Alain Mollard
Carabus (Oreocarabus) mikhaili loustei Deuve & Mourzine, 1998
Female adult
Mts.Nanling (Sichuan) , CHINE , VI-2001
Copyright © Alain Mollard
Carabus (Oreocarabus) luetgensi pseudoguadarramus Mollard, 2006 Paratype
Female adult
Montes de Herrera (Valladolid) Espagne, XI-2005
Copyright © Alain Mollard
Carabus (Oreocarabus) gemellatus Menetries, 1832
Male adult
Iran, Mazandaran, pass b. Gach-i-Sar (N seite), 2700m, 17-21.vi.1998, Heinz leg. (cAA)
Copyright © Alexander Anichtchenko
Carabus (Hypsocarabus) mikhaili mikhaili Deuve & Mourzine, 1997
Female adult
L=23 mm
Copyright © Ruslan Panin
Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847
Female adult
Spain, Sierra de Guadarrama, Puerto de la Quesera, 1720m, 41.214640, -3.430896, Anichtchenko A. leg. 10.X.2025 (cAA).
Copyright © Alexander Anichtchenko -
Orinocarabus taxon page (68 photos)
Carabus (Orinocarabus) sylvestris transsylvanicus Dejean, 1826
Male adult
Ukrainian Carpathians,<br /> Transcarpathian reg., Rakhov Distr., Chornogora Mts. Petros Mt. h=1850-2000m. alpine zone, 29.VI-28.VII.2007<br /> Panin R. Leg.
Copyright © Safronov
Carabus (Orinocarabus) sylvestris transsylvanicus Dejean, 1826
Male+Female adult
Ukrainian Carpathians
Transcarpathian reg., Rakhov Distr., Chornogora Mts. Pietros Mt. h=1850-2000 m. alpine zone, 29.VI-28.VII.2007 Panin R. Leg.
long 22 mm.
Copyright © Ruslan Panin
Carabus (Orinocarabus) pedemontanus omensis Born, 1901
Female adult
Italy, Piemonte (CN), Col del Mulo
17.7.2001 leg- Grottolo
female, 22 mm
Copyright © Alexander Anichtchenko
Carabus (Orinocarabus) pedemontanus omensis Born, 1901
Male adult
Italy, Piemonte (CN), Col del Mulo
17.7.2001 leg. Grottolo
male, 22 mm
Copyright © Alexander Anichtchenko
Carabus (Orinocarabus) fairmairei Thomson, 1875
Female adult
SE France,
Grand Belvédère du refuge du Mont Viso, 5-20.VII.1991
female, 20mm.
photo Panin R.
Copyright © Ruslan Panin
Carabus (Orinocarabus) concolor concolor Fabricius, 1792
Male adult
Piemonte, Oropa (BI) , M. Camino,
2300 m, 20.7.2003 leg. F. Angelini
male, 18 mm
Copyright © Alexander Anichtchenko
Carabus (Orinocarabus) concolor concolor Fabricius, 1792
Female adult
Piemonte, Oropa (BI) , M. Camino,
2300 m, 20.7.2003 leg. F. Angelini
female, 19 mm
Copyright © Alexander Anichtchenko
Carabus (Orinocarabus) castanopterus A. & J.B. Villa, 1833
Male adult
Italy, Lombardia (BG), passo S. Simone 2000 m,
Mezzoldo 26.8.2006 leg. Grottolo
male, 17 mm
Copyright © Alexander Anichtchenko
Carabus (Orinocarabus) bertolinii Kraatz, 1878
Male adult
Italia, Bellamonte, 2200 m, VI.1982 Rubizzani leg.
male, 18 mm
Copyright © Alexander Anichtchenko
Carabus (Orinocarabus) pedemontanus maurinensis Deuve, 2002 Holotype
Male adult
Copyright © Frank Kleinfeld
Carabus (Orinocarabus) bremii Stierlin, 1881
Male adult
NW Italy, Piemonte, Mucrone Mt., 14-21.VI.1999
Copyright © Alexander Anichtchenko
Carabus (Orinocarabus) castanopterus A. & J.B. Villa, 1833
Female adult
NC-Italy - S-Switzerland, Val Colla – Gazzirola, 22.VI.1999, C. Sola leg.
L=21mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) sylvestris sylvestris Panzer, 1793 (as slabai Niedl, 1968)
Female adult
Czech, WS Bohemia,
Plzen Reg., Domazlice Distr., Babylon, h=500m, 30.VIII.2008, S.I. Budka leg.
L=24mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) alpestris hoppei Germar, 1824
Male adult
W Austria, Tirol, Innsbruck, Hafelekar Mts h=2200m, Manfred Egger leg.
L=19 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) alpestris alpestris Sturm, 1815
Female adult
C Austria, E-Alps, Windischgarsten, Mt Hoher Nock, VII.2004, Manfred Egger leg.
L=20 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) alpestris adamellicola Ganglbauer, 1904
Male adult
N-Italy, Brescia prov., Bagolino, Adamello Mts, Passo del Maniva env., h=2000m, VIII.2011, M.Monzini leg.
L=19 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) fairmairei fairmairei Thomson, 1875
Male adult
N Italy, Alpi Graie; 20 mm
Copyright © Aleš Sedláček
Carabus (Orinocarabus) fairmairei heteromorphus Daniel, 1896
Female adult
NW-Italy, Aosta region, Champorcher env., Mt.Dondenaz, VIII.2008, V.Monzini leg.
L=19 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) fairmairei fenestrellanus Beuthin, 1899
Male adult
NW-Italy, Piemonte prov., Cottian Alps, Mt Albergian h=2450 m.<br /> L=20 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) bertolinii Kraatz, 1878
Male adult
NE-Italy, Dolomites-Alps, Trentino prov., Rolle Pass env., Malga Fosse h=2000m, 8-28.VI.2013, Manfred Egger leg.
L=19 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) alpestris dolomitanus Mandl, 1956
Male adult
NE-Italy, South Tyrol, Dolomites-Alps, 500 m left of Würzjoch, Gömaalm - at the foot Mt.Peitlerkofel h=2100m, 28.VII-17.VIII.2013, Manfred Egger leg.
l=18 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) fairmairei stecki Born, 1902
Female adult
France, Hautes-Alpes, env. Lake Foreant, near Col de D'Agnel, ca 2700-2800 m 17.VIII.2013 leg. B.Kataev L=18.0 mm
Copyright © Kirill Makarov
Carabus (Orinocarabus) fairmairei stecki Born, 1902
Male adult
France, Hautes-Alpes, env. Lake Foreant, near Col de D'Agnel, ca 2700-2800 m 17.VIII.2013 leg. B.Kataev L=16.5 mm
Copyright © Kirill Makarov
Carabus (Orinocarabus) pedemontanus raynaudianus Deuve & Simard, 1977
Male adult
SE France, Alpes-Maritimes, Isola env., Colle Santa-Anna h=2309 m, 44*13',127" N, 07*06',242"E, 31-7.VIII..2012. L=21 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) concolor lepontinus Born, 1908
Male adult
NW-Italy, Piemonte (VB), Lepontine Alps, Mt. Zeda , 6.VII.1998, L Diotti leg., L=20 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) concolor concolor Fabricius, 1792
Female adult
NW-Italy, Breuil-Cervinia h~2500 m, 2.IX.1971, Georgeon leg. L=21 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) pedemontanus germanae Casale & Cavazzuti, 1977
Male adult
NW-Italy, Piemonte, Imperia prov., Mt.Saccarello h~2000m, 30.V.1986. L=21 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) fairmairei pelvicus Cavazzuti, 1973
Male adult
NW-Italy, Cuneo, Maira Vall., Acceglio env., La Colletta h~2800 m, 12.VII.2007, R. Monguzzi leg. L=19 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) latreilleanus Csiki, 1927
Female adult
NW-Italy, Piemonte, Biella, Biellese Alps Rng, Mt.Camino, 25.VI.1997, V.Monzini leg. L=18 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) sylvestris transsylvanicus Dejean, 1826
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=21 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) sylvestris transsylvanicus Dejean, 1826 (aberrant)
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg.
Copyright © Ruslan Panin
Carabus (Orinocarabus) bertolinii Kraatz, 1878
Male adult
NE-Italy, Trentino prov. Dolomites Alps, Plateau Pale di San Martino h=2500 m, 17-19.VIII.2019, M. Egger leg.
Copyright © Ruslan Panin
Carabus (Orinocarabus) sylvestris haberfelneri Ganglbauer, 1892
Female adult
Austria, Styria, Dietmannsdorf, Herrenwaldrücken, h=1550-1600m, 29.VII-15.VIII.2024, M.Fuchs leg., L=25 mm
Copyright © Ruslan Panin
Carabus (Orinocarabus) sylvestris sylvestris (as micklitzi Sokolar, 1910)
Male adult
W Austria, Styria, Mt Hochwechsel W slope, 47.531667, 15.906000 h=1640 m, 14.IV.2025, I. Uspenskyi leg.
Copyright © Ruslan Panin -
Oxycarabus taxon page (8 photos)
Carabus (Oxycarabus) saphyrinus pseudosaphyrinus Schweiger, 1964 (as boluensis)
Male adult
Turkei, Bolu, Seben, Aladag,
1500m, 17.7.1988 Staven K. leg.
male, 23 mm
Copyright © Alexander Anichtchenko
Carabus (Oxycarabus) saphyrinus pseudosaphyrinus Schweiger, 1964 (as rabaronianus)
Male adult
Anatolia, Kastamonu, Yaraligoz verso Catalzeitin,
VI.2001 Sola Claudio leg.
male, 28 mm
Copyright © Alexander Anichtchenko
Carabus (Oxycarabus) saphyrinus pseudosaphyrinus Schweiger, 1964 (as rabaronianus)
Female adult
Anatolia, Kastamonu, Yaraligoz
verso Catalzeytin, VI.2001 Sola C. leg.
female, 30 mm
Copyright © Alexander Anichtchenko
Carabus (Oxycarabus) saphyrinus Cristoforis & Jan, 1837
Female adult
Turkey, NE Giresun, Dereli env., h~400 40°43'01"N 38°28'20"E 12.IX.2013 leg. B.Saratovskikh L=28.0 mm
Copyright © Kirill Makarov
Carabus (Oxycarabus) saphyrinus Cristoforis & Jan, 1837
Male adult
Turkey, NE Giresun, Dereli env., h~400 40°43'01"N 38°28'20"E 12.IX.2013 leg. B.Saratovskikh L=27.0 mm
Copyright © Kirill Makarov
Carabus (Oxycarabus) saphyrinus pirithous Reitter, 1896
Male adult
NW-Turkey,Istanbul (Asian part),Polonezköy env., 2.VI.2009. S. Dacatra leg. L=27 mm
Copyright © Ruslan Panin
Carabus (Oxycarabus) saphyrinus notabilis Roeschke, 1898
Male adult
Ulu Dag (Bursa) TURQUIE VI-1990
Copyright © Alain Mollard
Carabus (Oxycarabus) saphyrinus saphyrinus Cristoforis & Jan, 1837
Male adult
Gulug (Kumbet, Giresun) TURQUIE VII-1981
Copyright © Alain Mollard -
Pachycarabus taxon page (27 photos)
Carabus (Pachycarabus) staehlini Adams, 1817
Male adult
N Osetia, Skalistyi mt. rng.
Hamarat-Hoh mt. 14.7.1996 Arzanov leg.
male, 21 mm
Copyright © Alexander Anichtchenko
Carabus (Pachycarabus) roseri ciftekopruleriensis Machard, 1975
Female adult
NE Turkey,
Artvin Prov., Borcka, Macahel Gecidi h=1450m, 18.VI.2007, D.Prunier leg.
female, 22mm
Copyright © Ruslan Panin
Carabus (Pachycarabus) imitator katherinae Reitter, 1898
Male adult
Abkhasia, Anchkho mnt.Range, near Lake Chkho, h~2450-2550m, 13-14.VII.2009, Bondarenko A.S. leg., 20mm; det. I. Solodovnikov, 2025
Copyright © Ruslan Panin
Carabus (Pachycarabus) koenigi Ganglbauer, 1886
Male adult
NW Caucasus, S Russia, Adygea Rep., Lagonaki plateau, Site Murzikal h=1800-2000m, 24.V-18.IX.2010, D.Fominykh leg.
L= 22 mm
Copyright © Ruslan Panin
Carabus (Pachycarabus) roseri soganliensis Heinz & Korge, 1964
Male+Female adult
Turkey, Rize prov., Ikizdere distr., Rize Daglari,
47 km to SE from lyidere, Sivrikaya Koyu, h=2025 m.
30.VI-5.VII.2011, A. Vlasenko leg.
male - 23 mm, female - 24 mm.
Copyright © Alexandr Safronov
Carabus (Pachycarabus) roseri soganliensis Heinz & Korge, 1964
Female adult
NE-Turkey: Rize Prov.Ovit Pass (40.37;40.46) h-2700m,15.VII.2008, I.Rapuzzi & Caldon leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Pachycarabus) imitator imitator Reitter, 1883
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Carabus (Pachycarabus) staehlini Adams, 1817
Male adult
S Russia, E-Caucasus, Dagestan, Nukatl Mt Rng h=2600m, 25.VII.1997, V. Savizky leg. L=21 mm
Copyright © Ruslan Panin
Carabus (Pachycarabus) swaneticus kvirensis Belousov, 1992
Male adult
SC-Caucasus: NW-Georgia, Mingrelia, Egrissky Mt.Rng., h~1800-2000 m, 23-26.VII.1990, I.Retezár leg. L=20 mm
Copyright © Ruslan Panin
Carabus (Pachycarabus) imitator katherinae Reitter, 1898
Male adult
Abkhasia, Sukhumi distr., Anchkho Mt. Pass E- part h=1870-1920 m, 15-20.VII.2011, Pavluchuk S. leg., L= 21 mm; det. I. Solodovnikov, 2025
Copyright © Ruslan Panin
Carabus (Pachycarabus) imitator imitator Reitter, 1883
Male adult
Caucasus Major, NW Georgia, Utviri Pass env. h=2630 m, 43.083111, 42.255944, 13-14.VI.2023, I. Uspenskyi leg., L=19 mm
Copyright © Ruslan Panin
Carabus (Pachycarabus) roseri roseri Faldermann, 1835
Female adult
Georgia, Lesser Caucasus, Imereti prov., N slope Meskheti Mt Rng, Zekari Pass, h~1950 m, 24.VII.2021, E.Rujan leg., L=22 mm
Copyright © Ruslan Panin
Carabus (Pachycarabus) swaneticus swaneticus Reitter, 1883
Male adult
Georgia, Caucasus, Svaneti Range E part, Latpari Pass h=2700-2800 m, 23.VI.2024, I.Uspenskyi leg./ det. R. Panin, L=18 mm
Copyright © Ruslan Panin
Carabus (Pachycarabus) swaneticus swaneticus Reitter, 1883
Female adult
Georgia, Caucasus, Svaneti Range E part, Latpari Pass h=2700-2800 m, 23.VI.2024, I.Uspenskyi leg., L=19 mm
Copyright © Ruslan Panin
Carabus (Pachycarabus) swaneticus swaneticus Reitter, 1883
Male adult
Georgia , Caucasus, E part Egrisi Mt Range, Mt. Tsekuri N slope h=2370 m, 18-20.VI..2025, I.Uspenskyi leg. L=19 mm
Copyright © Ruslan Panin
Carabus (Pachycarabus) staehlini rionicus Belousov, 1992
Male adult
Georgia, Caucasus, S of Sasashi, Lechkhumi Range N sope h=2430m, alpine meadows 242.759167, 42.973889, 20-21.VII.2025, I.Uspenskyi leg./det. R,Panin, L=20 mm
Copyright © Ruslan Panin
Carabus (Pachycarabus) staehlini rionicus Belousov, 1992
Male adult
Georgia, Caucasus, S of Sasashi, Lechkhumi Range N sope h=2430m, alpine meadows 242.759167, 42.973889, 20-21.VII.2025, I.Uspenskyi leg./det. R. Panin, L=19.5 mm
Copyright © Ruslan Panin
Carabus (Pachycarabus) staehlini rionicus Belousov, 1992
Female adult
Georgia, Caucasus, S of Sasashi, Lechkhumi Range N sope h=2430m, alpine meadows 242.759167, 42.973889, 20-21.VII.2025, I.Uspenskyi leg./det. R. Panin, L=20.5 mm
Copyright © Ruslan Panin
Carabus (Pachycarabus) koenigi Ganglbauer, 1886
Male adult
NW Caucasus, Republic Adygea, Mt.Tybga h=2100 m, 28.VI.2006, Tret’yakov Yu. leg., L=21 mm
Copyright © Ruslan Panin -
Pachystus taxon page (45 photos)
Carabus (Pachystus) hungaricus mingens Quensel, 1806
Male adult
male, 31 mm
SW-Russia: Stavropol Krai, near Stavropol city , Russkiy Les, 11.VI-28.IX.2010, Pavluchuk S. leg.
Copyright © Ruslan Panin
Carabus (Pachystus) tamsi tamsi Menetries, 1832
Male+Female adult
Transcaucasia, Armenia, <br /> Ashotsk, 18.VI.2001, P.V.Kazaryan leg.<br /> female, 26mm//male, 24mm
Copyright © Ruslan Panin
Carabus (Pachystus) tamsi tamsi Menetries, 1832
Female adult
Azerbayjan, Talysh mt, Zuvant<br /> Pirasora, 1900-2000 m 9.7.1983 I. Belousov leg.<br /> female, 25 mm
Copyright © Alexander Anichtchenko
Carabus (Pachystus) tamsi tamsi Menetries, 1832
Male adult
N Iran, Azarbaygan-e-Sharqi, 21 km SE<br /> Halhal, 3734N 4841E, 1633m<br /> 3.6.2003 Kabatek P. leg<br /> male, 23 mm
Copyright © Alexander Anichtchenko
Carabus (Pachystus) hungaricus scythus Motschulsky, 1847
Male adult
S Ukraine,
Zaporizhia reg., Khortytsa island, 19.VIII.2008, M. Mulenko leg., det.Panin R.
male, 27mm
Copyright © Ruslan Panin
Carabus (Pachystus) hungaricus cribellatus Adams, 1812
Female adult
Kazakhstan,
Astana reg. Enbekshil-der distr., Burevestnik vill., 9.VIII.2002
female,26 mm
Copyright © Ruslan Panin
Carabus (Pachystus) graecus morio Mannerheim, 1830
Male adult
Turchia, Sehtavul Geg 1700m
5.1996 Rabizzani leg.
male, 26 mm
Copyright © Alexander Anichtchenko
Carabus (Pachystus) graecus kykladensis Breuning, 1961
Male adult
male, 24mm
Copyright © Boleslav Brezina
Carabus (Pachystus) graecus kykladensis Breuning, 1961
Male adult
female, 24mm
Copyright © Boleslav Brezina
Carabus (Pachystus) glabratus glabratus Paykull, 1790
Female adult
W UKRAINE<br /> Lviv region, Stryi distr., near Bilche vill., 9-20.VI.2005, Panin R. leg.<br /> female 30 mm
Copyright © Ruslan Panin
Carabus (Pachystus) glabratus glabratus Paykull, 1790
Male adult
W Ukraine, NW Podolian, Lviv reg., Zolochiv distr., near Stinka vill., Lysa Gora, 26.IV.2009 R.Panin leg., 29mm
Copyright © Ruslan Panin
Carabus (Pachystus) glabratus glabratus Paykull, 1790
Female adult
Germania, Oberfranken, Kulmbach leg.det. U.Schmidt, 1974 Foto: U.Schmidt, 2008
Copyright © U.Schmidt
Carabus (Pachystus) cavernosus cavernosus Frivaldsky, 1837
Male adult
Bulgaria, Middle Stara Planina Mts.,
feeder of Byala Reka riv., below Botev Peak
1900-2000m 25.7.1996 B. Gueorguiev leg.
male, 25 mm
Copyright © Alexander Anichtchenko
Carabus (Pachystus) hungaricus gastridulus Fischer, 1823
Male adult
N. Krym Peninsula,
near Armyansk, Demidov G. leg.5.V.2006
male, 28mm
Copyright © Ruslan Panin
Carabus (Pachystus) cribratus cribratus Quensel, 1806
Male adult
Turkey, Gьmьshane<br /> leg. det. K.Staven, 1992<br /> length: 26 mm
Copyright © U.Schmidt
Carabus (Pachystus) cribratus cribratus Quensel, 1806
Male adult
near Tbilisi, Bakuriani<br /> 7.VI.1973, Shilenkov V. leg.<br /> male, 25 mm
Copyright © Alexander Anichtchenko
Carabus (Pachystus) hungaricus hungaricus Fabricius, 1792
Female adult
C Hungary, Pest reg., Örkény, 9-10.V.1997, S.Budka leg.
Copyright © Ruslan Panin
Carabus (Pachystus) hungaricus cribellatus Adams, 1812
Male adult
E Siberia, Russia, Krasnoyarsk Reg., near Krasnoyarsk, forest-steppe,1.VII-20.VIII,2011,M.Ivanov leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Pachystus) hungaricus scythus Motschulsky, 1847
Female adult
S Ukraine, Zaporizhzhya reg., Khortytsa island, 16-20.VII.2011, M. Mulenko leg., det. Panin R.
L=28 mm
Copyright © Ruslan Panin
Carabus (Pachystus) hungaricus hungaricus Fabricius, 1792
Male adult
Czech, S Moravia, Pouzdrany, 8,VIII.2005, V. Slovak lgt.
L=27 mm
Copyright © Ruslan Panin
Carabus (Pachystus) cribratus staneki Sterba, 1931
Male adult
NC Turkey, Küre Mts., 60km N Kastamonu; 25mm
Copyright © Aleš Sedláček
Carabus (Pachystus) cribratus staneki Sterba, 1931
Female adult
NC Turkey, Küre Mts., 60km N Kastamonu; 27mm
Copyright © Aleš Sedláček
Carabus (Pachystus) cavernosus cavernosus Frivaldsky, 1837
Female adult
W Bulgaria, Cerovo, 31 mm
Copyright © Aleš Sedláček
Carabus (Pachystus) hungaricus scythus Motschulsky, 1847 (loc.typ.)
Male adult
S-Ukraine, near Dnipropetrovsk, 6-15.IX.2013, A. Sukhenko leg.<br /> L=28 mm
Copyright © Ruslan Panin
Carabus (Pachystus) cribratus echarouxi Lassalle, 1996 Holotype
Male adult
in coll Lassalle
Copyright © Bernard Lassalle
Carabus (Pachystus) cavernosus variolatus Costa, 1839
Male adult
C-Italy:Abruzzo reg., Gran Sasso d'Italia Massif, h~1600 m, VIII.1997, Seichert leg. L=26 mm
Copyright © Ruslan Panin
Carabus (Pachystus) cavernosus rozneri Retezár & Gy. Szél, 2013 Paratype
Male adult
Romania, Apuseni : Trascău Mts. L=27 mm
Copyright © Ruslan Panin
Carabus (Pachystus) pisidicus pisidicus Peyron, 1854
Male adult
Col Yellibel (Konya), TURQUIE VII-1984
Copyright © Alain Mollard
Carabus (Pachystus) cribratus pulumuricus Cavazzuti, 1994
Female adult
Pulumur (Tunceli) TURQUIE, VI-1991
Copyright © Alain Mollard
Carabus (Pachystus) cribratus porrectangulus Gehin, 1885
Female adult
Col d’Egribel (Bolu)), TURQUIE, VII-1999
Copyright © Alain Mollard
Carabus (Pachystus) cribratus yaraligozi Battoni, 1982
Male adult
Yaraligoz (Kastamonu), TURQUIE VI-2006
Copyright © Alain Mollard
Carabus (Pachystus) glabratus gibbosus Heyden, 1866
Male adult
N Croatia, Ivanščica Mts, Ivanec, 4-18.VI.2011, Krause Rene leg., L=32 mm
Copyright © Ruslan Panin
Carabus (Pachystus) cavernosus variolatus Costa, 1839
Female adult
C-Italy: Abruzzo reg., Gran Sasso massif, Campo Imperatore, h=1900 m, VIII.2021, G.Proscia leg., L=27 mm
Copyright © Ruslan Panin
Carabus (Pachystus) cribratus cribratus Quensel, 1806
Male adult
NW Armenia, Amasia, (Shirak prov.), 20-24.VIII.1998, Kazaryan P.V. leg.
Copyright © Ruslan Panin -
Pagocarabus taxon page (49 photos)
Carabus (Pagocarabus) crassesculptus eleganticolor Deuve, 1997 Paratype
Male adult
China, Gansu, Min Shan mts., 70 km NW Wudu
Copyright © Alexander Anichtchenko
Carabus (Pagocarabus) crassesculptus diruptus Morawitz, 1886
Male adult
China, Gansu, Umg. Xiahe 3000-3200m
24-26.6.1993 Heinz leg.
20 mm
Copyright © Alexander Anichtchenko
Carabus (Pagocarabus) crassesculptus crassesculptus Kraatz, 1881
Male adult
China, Beijing, Wulingshan, 2007, Liu ye leg.;
Photo & det. by liuye
Copyright © Ye Liu
Carabus (Pagocarabus) crassesculptus crassesculptus Kraatz, 1881
Male adult
China, Hebei, Xianhua (=Kalgan)
10-15.5.1995
21 mm
Copyright © Alexander Anichtchenko
Carabus (Pagocarabus) crassesculptus tonghua Deuve & Mourzine, 2001
Male adult
China, NW Sishuan, 3500m, Hongyuan, Somang Qu riv. grassland, 9-29.6.2004 (cAA). 20 mm
Copyright © Alexander Anichtchenko
Carabus (Pagocarabus) crassesculptus eleganticolor Deuve, 1997
Female adult
S Gansu, Min Shan, 5-20 km S Cagan, 2300-3200m 7-20.7.2019 V. Patrikeev leg. (cAA). det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko -
Pantophyrtus taxon page (20 photos)
Carabus (Pantophyrtus) turcomanorum karaalmicus Gottwald, 1994
Male adult
male, 27mm
Copyright © Boleslav Brezina
Carabus (Pantophyrtus) turcomanorum turcomanorum Thieme, 1881
Female adult
female, 32mm
Copyright © Boleslav Brezina
Carabus (Pantophyrtus) turcomanorum turcomanorum Thieme, 1881
Male adult
male, 28mm
Copyright © Boleslav Brezina
Carabus (Pantophyrtus) turcomanorum s.str. (as orbitosus)
Male adult
male, 27mm
Copyright © Boleslav Brezina
Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993
Male adult
Kirgizstan, S Chatkal mt. rng., Sary-Chelek lake 2000 m, 25.6.2000
Copyright © Alexander Anichtchenko
Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993
Male adult
male, 29mm
Copyright © Boleslav Brezina
Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993 Paratype
Female adult
female, 28mm
Copyright © Boleslav Brezina
Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993 Paratype
Female adult
female, 34mm
Copyright © Boleslav Brezina
Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993 Paratype
Male adult
male, 27mm
Copyright © Boleslav Brezina
Carabus (Pantophyrtus) brachypedilus brachypedilus s.str.
Male adult
N Kirgizskii mt rng., Ak-Suu riv., 1400 m,
Belovodskoe, 16.4.2000 Klimenko A. leg.
Copyright © Alexander Anichtchenko
Carabus (Pantophyrtus) brachypedilus brachypedilus s.str.
Female adult
female, 27mm
Copyright © Boleslav Brezina
Carabus (Pantophyrtus) brachypedilus brachypedilus s.str.
Male adult
male, 26mm
Copyright © Boleslav Brezina
Carabus (Pantophyrtus) brachypedilus brachypedilus s.str.
Female adult
female, 28mm
Copyright © Boleslav Brezina
Carabus (Pantophyrtus) brachypedilus absentivus Belousov & Kabak 1993 Paratype
Male adult
male, 24mm
Copyright © Boleslav Brezina
Carabus (Pantophyrtus) brachypedilus absentivus Belousov & Kabak 1993 Paratype
Male adult
male, 24mm
Copyright © Boleslav Brezina
Carabus (Pantophyrtus) brachypedilus absentivus Belousov & Kabak 1993 Paratype
Female adult
female, 26mm
Copyright © Boleslav Brezina
Carabus (Pantophyrtus) brachypedilus absentivus Belousov & Kabak 1993 Paratype
Female adult
female, 30mm
Copyright © Boleslav Brezina
Carabus (Pantophyrtus) turcomanorum turcomanorum Thieme, 1881
Male+Female adult
Kyrgyzstan, Kyrgyzskyi Mts. R., Uzun Bulak Gorge, h=1800 ms.
01-09.VI.2002 loc. coll.
male - 28 mm, female - 33 mm.
Copyright © Alexandr Safronov
Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993
Male adult
Uzbekistan, Chatkal Mt. rng., Irtash, 41°10'25.7"N 70°16'20.4"E, 5-7.8.2021, 2400m, A. Anichtchenko leg. (cAA). 27mm
Copyright © Alexander Anichtchenko
Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993
Female adult
Uzbekistan, Chatkal Mt. rng., Irtash, 41°10'25.7"N 70°16'20.4"E, 5-7.8.2021, 2400m, A. Anichtchenko leg. (cAA). 30 mm
Copyright © Alexander Anichtchenko -
Piocarabus taxon page (5 photos)
Carabus (Piocarabus) sui Imura & Zhou, 1998
Female adult
China, Beijing, Yunmengshan,2005, Liu ye coll.;<br /> 35mm
Copyright © Ye Liu
Carabus (Piocarabus) reitterianus Breuning, 1934
Male adult
China, Gansu, ca 25 km E Xiahe 2700-2900m, Wagatan Timber<br /> 30.7.-2.8.1993 Heinz leg.<br /> male, 23 mm
Copyright © Alexander Anichtchenko
Carabus (Piocarabus) nanwutai Kleinfeld, Korell & Wrase, 1996
Male adult
China, Shaanxi, Qing Ling Shan, track Hou Zen Zi vill.<br /> to Taiban Shan, 2500m, 27-29.6.1998 mixed forest<br /> male, long. 19 mm
Copyright © Alexander Anichtchenko
Carabus (Piocarabus) nanwutai Kleinfeld, Korell & Wrase, 1996
Male+Female adult
Copyright © Frank Kleinfeld
Carabus kitawakianus kitawakianus Imura, 1993
Male adult
China, Shaanxi, Tschinling mts., Taibeichan Nat. res. 2000m 5-15.8.2005 (cAA). 27 mm
Copyright © Alexander Anichtchenko
Carabus (Piocarabus) vladimirskyi Dejean & Boisduval, 1830
Male adult
China, Beijing, Jiufeng mountain, 2007, Liu ye leg.;<br /> Photo & det. by liuye
Copyright © Ye Liu
Carabus (Piocarabus) titanus Breuning, 1933
Male adult
C-China Sichuan prov., Nanping, h=1500m, VI.2000, S. Remond leg.<br /> L=34 mm
Copyright © Ruslan Panin
Carabus (Piocarabus) nanwutai Kleinfeld, Korell & Wrase, 1996
Male adult
China, W-Shaanxi, Taibai Shan Mts, btw. Houzhenzi/Ba Xian Tai, nr.Nan Tian Men h=3200m, 33°55'17"N/107°46'53"E, 25-30.VII.2011, Li Wuyang leg.<br /> L=19 mm
Copyright © Ruslan Panin
Carabus (Piocarabus) sui Imura & Zhou, 1998
Female adult
CE-China, Henan, Mt Wangwu, V.2007, local collector. L= 33 mm
Copyright © Ruslan Panin
Carabus (Piocarabus) vladimirskyi Dejean & Boisduval, 1830
Male adult
Buryatia, Kyahta distr. [Khyagh], 5 km SE Chikoi vill., 16.8.1998, Anichtchenko A. leg.
Copyright © Alexander Anichtchenko
Carabus (Piocarabus) titanus Breuning, 1933
Female adult
China Sichuan prov., Nanping, h=1500m, VI.2000, S. Remond leg.
L=38 mm
Copyright © Ruslan Panin -
Platycarabus taxon page (37 photos)
Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Male adult
W.Ukraine, Transcarpathian Reg., Rakhov Distr., Chornogora range., h=2020m. Panin R. leg. 28.VI-9.VIII.2008, male, 19 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Female adult
W Ukraine, NE Carpathians, Transcarpathian Reg., Rakhov Distr., Chornogora Mts Range h=2030m, alpin zone, 13-24.VIII.2010, Panin R. leg., female, 22 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Female adult
Ukrainian Carpathians, Transcarpathian reg., Rakhov Distr., Chornogora range, h=2010m, 29.VI-28.VII.2007, Panin R. leg., female, 23 mm., photo/det.Panin R
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Female adult
Ukrainian Carpathians, Transcarpathian reg., Rakhov Distr., Chornogora range, h=2010 m., alpine zone 29.VI-28.VII.2007 Panin R. leg., female,23 mm.
Copyright © Ruslan Panin
Carabus (Platycarabus) irregularis montandoni Buysson, 1882
Male adult
W-Ukraine,
Lviv Reg., Zhydachiv distr., near Vybranivka, under the bark of oak tree, in the forest,12.X.2007 Panin R. leg.
male,22 mm.
Copyright © Ruslan Panin
Carabus (Platycarabus) irregularis montandoni Buysson, 1882
Male adult
W-Ukraine, NW Podolian,
Lviv Reg., Zhydachiv distr., near Vybranivka, under the bark of oak tree, in the forest,12.X.2007 Panin R. leg
L= 22 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) depressus lucens Schaum, 1856
Female adult
Italia, Alpi Cozie, Col. delle Finestre,
2150m, VII.1986 Ratti leg.
female, 24 mm
Copyright © Alexander Anichtchenko
Carabus (Platycarabus) depressus depressus Bonelli, 1811
Female adult
Italia, Bellamonte, 2200m
VI.1992 Rubizzani leg.
female, 20 mm
Copyright © Alexander Anichtchenko
Carabus (Platycarabus) cychroides Baudi, 1860
Male+Female adult
female: NW-Italy, Provincia Torino, Mt.Albergian h~2600 m, 9.VI.2007, V.Monzini leg.
L=20 mm
male:, N slope Mt.Albergian h~2000 m, 9.VI.2003, P.Zorgati leg.
photo Panin R.
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Male adult
W.Ukraine, Transcarpathian Reg., Rakhov Distr., Chornogora Mt range h=2020m alpine zone, Panin R. leg. 28.VI-9.VIII.2008, male 19 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii fabricii Panzer, 1812
Male adult
NE-Italy,
Trentino-Alto Adige Reg,. Bolzano-Bozen Prov.,Val Aurina – Rottal 2050-2150m, 28.VI.1997, I.Rapuzzi leg.
male, 19mm
Copyright © Ruslan Panin
Carabus (Platycarabus) irregularis irregularis Fabricius, 1792
Male adult
NE-Italy,
Friuli Reg., Udine, Carnia, Pierabech h~1100m, I.Rapuzzi leg.
male, 23 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) irregularis ramanus Sokolar, 1909 (as hayeki Kobmann, 1925)
Male adult
N Slovenia, Maribor – Dravograd, Pomorje Mts.,13.XI.2005, I.Rapuzzi leg.
Copyright © Ruslan Panin
Carabus (Platycarabus) irregularis ramanus Sokolar, 1909 (as bucephalus Kraatz, 1879)
Male+Female adult
W Slovenia, Goriška region Trnovo, h~1000 m, IX.2002, I.Rapuzzi leg.
female, 29 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) creutzeri creutzeri Fabricius, 1801 (as trnovensis Bernau, 1914)
Male adult
W Slovenia,
Lokve, Trnovski Gozd h=1100m, 17.IV.2006, leg. I.Rapuzzi
male, 26mm
Copyright © Ruslan Panin
Carabus (Platycarabus) creutzeri creutzeri Fabricius, 1801
Male adult
Italy, Fruili - UD - Carnia, N. Paulano 29.III.2002 leg. I.Rapuzzi
L=22.0 mm
Copyright © Kirill Makarov
Carabus (Platycarabus) creutzeri creutzeri Fabricius, 1801(as posarinii Rapuzzi, 1999 PT) Paratype
Male+Female adult
Slovenia:Koper:Kras Mt.R.:Brkini Mts.:Suhorije env., VIII.1998, I.Rapuzzi leg.
photo & koll. Panin R.
female, 32 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) creutzeri kircheri Germar, 1824
Male+Female adult
NE-Italy, Veneto Reg., Belluno, Forcella Lavardet h=2000m, VII.2002, I.Rapuzzi leg.
female, 24 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii fabricii (as fassatii Smetana, 1955)
Male adult
C-Slovakia:Nízke Tatry Mts., 4.VI.2011, Krause Rene leg.
L=19 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) depressus lucens Schaum, 1856
Female adult
SE-France: Alpi Marittime, Monte Viso, 14.VII.2010
L=23 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii fabricii (as fassatii Smetana, 1955)
Male adult
C-Slovakia:Nizke Tatry Mts., h=1750 m,.9.VI.2007, V.Slovak leg.
L=20 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii fabricii (as feichtauensis Müller & Sach, 1999)
Male adult
NC Austria, E-Alps, Windischgarsten, Mt Hoher Nock, VII.2004, Manfred Egger leg.
L=18 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii fabricii (as koralpicus Sokolar, 1910)
Female adult
S Austria, Carinthia reg.(Kärnten), Wolfsberg, Koralpe Mt Range,18.VI.2005, Manfred Egger leg.
L=22 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii fabricii (as poloniensis Géhin, 1876)
Female adult
N-Slovakia, Orava Beskids Mts, Mt Babia Góra h=1720 m, 5.VII.20.VIII.2010, Rudolf Kmeco leg.
L=22 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii ucrainicus Lazorko, 1951
Male adult
Ukrainian Carpathians, Ivano-Frankivsk reg., Yaremcha distr., Gorgany Mts., h=1590 m, 22.VII-3.VIII.08, Panin R. leg., male, 23mm
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii ucrainicus Lazorko, 1951
Male adult
Ukrainian Carpathians, Ivano-Frankivsk reg., Gorgany Mts., h=1590 m, 17.VI-26.VII.2011, Panin R. leg.<br /> male, 23mm
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii ucrainicus Lazorko, 1951
Female adult
W-UKRAINE, Ivano-Frankivsk reg., Gorgany Mts, Syvula Mt., h =1820 м., 25.VII.2011,Panin R. leg., L=27 mm<br />
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Female adult
N- Romania, E-Carpathians, Rodna Mts, h=2000m, 7-8.VI.2012, Cartuz Z.leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) creutzeri kircheri Germar, 1824
Male adult
S Austria, Karnische Alpen; male, 24mm
Copyright © Aleš Sedláček
Carabus (Platycarabus) creutzeri kircheri Germar, 1824
Female adult
S Austria, Karnische Alpen; 25mm
Copyright © Aleš Sedláček
Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Female adult
W Ukraine, NE Carpathians, Transcarpathian Region, Chornogora Mt Range, h=2000 m, alpin zone, 28.VII-11.IX.2014, Panin R. leg. L=22 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) creutzeri baldensis Schaum, 1857
Male adult
NC-Italy: Trento, Sega di Ala, IX.1999, A. Vicini leg. , L=24 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) irregularis irregularis (as jurassicus Born, 1911)
Male+Female adult
France, Ain (01), Jura Mts, Grand Colombier, Forêt d' Arvières h~1250 m, 25.X.2014, Ya. Vasseur leg., L=23 mm
Copyright © Ruslan Panin
Carabus (Platycarabus) irregularis ramanus Sokolar, 1909
Female adult
NW Croatia, Ivanščica Mts , Ivanec, 4-18.VI.2011, Krause Rene leg., L=33 mm
Copyright © Ruslan Panin -
Procerus taxon page (54 photos)
Carabus (Procerus) gigas gigas Creutzer, 1799
Male+Female adult
N Croatia, Ivanščica Mts , Ivanec, 4-18.VI.2011, Krause Rene leg.
female, 54 mm
Copyright © Ruslan Panin
Carabus (Procerus) scabrosus caucasicus Adams, 1817
Male adult
N Caucasus,
Stavropol Krai, Pyatigorsk, Mt.Mashuk, VI.2007 Tikhonov Yu. leg.
44mm
Copyright © Ruslan Panin
Carabus (Procerus) gigas duponcheli Dejean, 1831 (as kusdasi Mandl, 1966)
Male adult
S-Greece: Peloponnes, Mt.Chelmos, 20.V.2007.
L=46 mm
Copyright © Ruslan Panin
Carabus (Procerus) scabrosus bureschianus Breuning, 1928
Male adult
Bulgaria, W Rodopi, Dobrostan masive, backovo env. M.Janalik leg.
Copyright © Igor Solodovnikov
Carabus (Procerus) scabrosus colchicus Motschulsky, 1844
Male adult
Abkhasia: S part of Bzybian Mt.R. vall. riv. Khipsta, h= 600 m, VIII.2014, leg. I.A. Solodovnikov
Copyright © Igor Solodovnikov
Carabus (Procerus) scabrosus colchicus Motschulsky, 1844
Male adult
Abkhasia: S part of Bzybian Mt.R. vall. riv. Khipsta, h= 600 m, VIII.2014, leg. I.A. Solodovnikov, 58 mm
Copyright © Igor Solodovnikov
Carabus (Procerus) scabrosus schuberti Breuning, 1968
Male adult
NC-Turkey: 10 km E of Sinop, h = 0 m, VI.1999, leg. St. Remond, 42 mm
Copyright © Igor Solodovnikov
Carabus (Procerus) scabrosus tauricus Bonelli, 1811
Male adult
f. virens Crimea, Kazu-Kau Mt. (South part Chatyr-Dagh Mt.) 14.06.2009. Leg V.V.Savchuk, male. 41 mm
Copyright © Igor Solodovnikov
Carabus (Procerus) scabrosus tauricus Bonelli, 1811
Male adult
f. purpureus Kraatz Crimea, Inkerman, steppe, 26.03.1987. Leg I.M. Lutzky. 42 mm
Copyright © Igor Solodovnikov
Carabus (Procerus) scabrosus bulgharmaadensis Bodemeyer, 1915 (as ludademae)
Female adult
Turkey, Karaman, 1700 m., 03 March 2014
Copyright © Özgür KOÇAK
Carabus (Procerus) syriacus galilaeus Schweiger, 1970
Male adult
Elon, North Galilea (Israel)
Copyright © Alain Mollard
Carabus (Procerus) syriacus syriacus Kollar, 1843
Female adult
East Saida, Montes de Lebanon (SC. Lebanon)
Copyright © Alain Mollard
Carabus (Procerus) syriacus bulgharmaadensis
Female adult
Etekli (Adana, S.Turquey,)
Copyright © Alain Mollard
Carabus (Procerus) scabrosus akbesianus Breuning, 1975
Male adult
Akbés (Antakya, Turquey)
Copyright © Alain Mollard
Carabus (Procerus) scabrosus amasicus Csiki, 1927
Male adult
Almus (Tokat, Turquey)
Copyright © Alain Mollard
Carabus (Procerus) scabrosus audouini Brulle, 1835
Male adult
Isiklar (Trabzon,Turquey)
Copyright © Alain Mollard
Carabus (Procerus) scabrosus callipygius Cavazzuti, 1986
Male adult
Devrekani (Kastamonu, Turquey)
Copyright © Alain Mollard
Carabus (Procerus) gigas parnassicus Kraatz, 1884
Female adult
N. Greece, Phalakron Mt., 750 m, 09.05.2004. Image of В.И. Гуменюк from www.zin.ru
Copyright © - - -
Carabus (Procerus) scabrosus audouini Brulle, 1835
Female adult
Turkey. Rize Prov., South of Ikizdere, 800 m, 17.08.2009. Image of В.И. Гуменюк from www.zin.ru
Copyright © - - -
Carabus (Procerus) scabrosus armenius Zaitzev, 1930
Male adult
Balkaya (Gumushane, Turquey)
Copyright © Alain Mollard
Carabus (Procerus) scabrosus dardanellicus Kraatz, 1886
Male adult
Sancaeli (Canakkale, Turquey)
Copyright © Alain Mollard
Carabus (Procerus) scabrosus fallettianus (Cavazzuti, 1997)
Male adult
Khndzorek (Goris, Arménie)
Copyright © Alain Mollard
Carabus (Procerus) scabrosus montisabanti Schweiger, 1962
Male adult
Abant (Bolu, Turquey)
Copyright © Alain Mollard
Carabus (Procerus) scabrosus scabrosus Olivier, 1795
Male adult
Dermirkoy (Kirklareli, Turquey). L = 54mm
Copyright © Alain Mollard
Carabus (Procerus) scabrosus propinquus Csiki, 1927
Male adult
Alem Dag (Istambul, Turquey) L=57 mm
Copyright © Alain Mollard
Carabus (Procerus) scabrosus mentor Blumenthal & Breuning, 1967
Male adult
Catalzeytin,(Katsamonu,Turquey)
Copyright © Alain Mollard
Carabus (Procerus) scabrosus munzurensis Cavazzuti & Lassalle, 1987 Paratype
Male adult
leg. B.Lassalle, from Pulumur (Tunceli,Turquey)
Copyright © Alain Mollard
Carabus (Procerus) caucasicus var.tuberculosus Lectotype
Female adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC468 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec468
Copyright © - - -
Carabus (Procerus) gigas duponcheli Dejean, 1831 Holotype
Male adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC54 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec54
Copyright © - - -
Carabus (Procerus) syriacus kozloviorum Cavazzuti, 2014 Holotype
Male adult
Turkey. Nigde. Ulukisla
Copyright © Anton Kozlov
Carabus (Procerus) syriacus kozloviorum Cavazzuti, 2014 Paratype
Female adult
Turkey. Nigde. Ulukisla
Copyright © Anton Kozlov
Carabus (Procerus) syriacus ludademae Cavazzuti & Kozlov, 2014 Holotype
Male adult
Prov. Karaman : circa 10 km a Ovest di Karaman IV.2014. (in coll A. Kozlov)
Copyright © Anton Kozlov
Carabus (Procerus) syriacus ludademae Cavazzuti & Kozlov, 2014 Paratype
Female adult
Prov. Karaman : circa 10 km a Ovest di Karaman IV.2014. (in coll A. Kozlov)
Copyright © Anton Kozlov
Carabus (Procerus) scabrosus antonkozlovi Cavazzuti, 2014 Paratype
Female adult
Tavush prov., Ditavan vill. 25-29.VII.2013 (in coll Kozlov)
Copyright © Anton Kozlov
Carabus (Procerus) scabrosus antonkozlovi Cavazzuti, 2014 Holotype
Male adult
Tavush prov., Mehrab vill. 950-1005m (in coll Kozlov). 44 mm
Copyright © Anton Kozlov
Carabus (Procerus) scabrosus tatianagorokhovae Cavazzuti & Kozlov, 2014 Holotype
Male adult
Armenia. Lori prov. Akhtala
Copyright © Anton Kozlov
Carabus (Procerus) scabrosus tauricus Bonelli, 1811
Male adult
Crimea, Sevastopol, leg.det. A.Chuvilin, 12.VIII.1999. Photo: U.Schmidt, 2016. 40mm
Copyright © Udo Schmidt
Carabus (Procerus) scabrosus nakagomei Cavazzuti & Kozlov, 2016 Paratype
Female adult
Copyright © Anton Kozlov
Carabus (Procerus) scabrosus nakagomei Cavazzuti & Kozlov, 2016 Holotype
Male adult
Copyright © Anton Kozlov
Carabus (Procerus) syriacus bulgharmaadensis Bodemeyer, 1915 (as toulgoetianus)
Female adult
North Gulek (Prov.Adana), South Turquey.
Copyright © Alain Mollard
Carabus (Procerus) syriacus limitaneus (Korell, 1989)
Male adult
Daret’Azze; North-West Syria
Copyright © Alain Mollard
Carabus (Procerus) scabrosus akbesianus Breuning, 1975 Holotype
Male adult
Syrie, Akbes (MNHN-EC-EC131)
Copyright © - - -
Carabus (Procerus) scabrosus kahramanicus Cavazzuti & Myška, 2004 Holotype
Female adult
Anatolia Mer., Kahramanmaras, 20/08/2000, P. Smrz leg., in coli. Myska.
Copyright © - - -
-
Procrustes taxon page (48 photos)
Carabus (Procrustes) paiafa acuticollis Motschulsky, 1858
Male adult
Copyright © Alexander Anichtchenko
Carabus (Procrustes) iimpressus Klug, 1832 (as drusum) Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Procrustes) impressus Klug, 1832 (as desertum) Holotype Paratype
Male+Female adult
Copyright © Frank Kleinfeld
Carabus (Procrustes) coriaceus kindermanni Waltl, 1838
Male+Female adult
NW TURKEY,
Stambul Avdzhilar, (Eropean part), h~20 m, 10-29.X.2007 M.V.Cheriavskaya leg./det. I.A.Solodovnikov
female,31mm
Copyright © Ruslan Panin
Carabus (Procrustes) coriaceus cerisyi Dejean, 1826 (as foudrasi)
Male adult
Copyright © Boleslav Brezina
Carabus (Procrustes) coriaceus coriaceus Linne, 1758
Female adult
The neighborhood Nicolo-Berezovskoe republic of
Bashkortostan 5.07
Copyright © Sunvas
Carabus (Procrustes) coriaceus coriaceus Linne, 1758
Male adult
length: 32.42 mm
Germania, Oberfranken, Weismain
leg.det. U.Schmidt, 1983
Foto: U.Schmidt, 2005
Copyright © U.Schmidt
Carabus (Procrustes) coriaceus cerisyi Dejean, 1826
Male adult
male, 30mm
Copyright © Boleslav Brezina
Carabus (Procrustes) coriaceus cerisyi Dejean, 1826
Female adult
female, 31mm
Copyright © Boleslav Brezina
Carabus (Procrustes) clypeatus clypeatus Adams, 1817
Female adult
C-Caucasus, N Ossetia, vall. Mamyhdon riv., h=2100
Koval A.G. leg; 08.08.2001
Copyright © Anton Menstschikov
Carabus (Procrustes) chevrolati korbi Breuning, 1928
Male adult
length: 28 mm
Türkei, Tülü/Sathan, Erzincan
leg. Staven & Willmer; det. K.Staven, 2003
Foto: U.Schmidt, 2009
Copyright © U.Schmidt
Carabus (Procrustes) chevrolati chevrolati Cristoforis & Jan, 1837
Male+Female adult
female, 31mm: Ashotsk,Vardaghbyur vill., 30.V.1997
male, 30 mm: Amasia, Chaldash vill.,8-11.VIII.1997
P.V. Kazaryan leg.
Copyright © Ruslan Panin
Carabus (Procrustes) coriaceus banaticus Redtenbacher, 1849
Male adult
SW Romania, Banat, Băile Herculane env., Domogled valley, 24-30.V.2008, Krause Rene leg.
L=35 mm
Copyright © Ruslan Panin
Carabus (Procrustes) chevrolati thirki Chaudoir, 1857
Male adult
NE Turkey, Trabzon, Zigana Pass, h~2000m, V. Slovak leg.
L=27 mm
Copyright © Ruslan Panin
Carabus (Procrustes) coriaceus hopffgarteni (as emgei Ganglbauer, 1888)
Male adult
CE-Greece: Larissa reg. Stomio, Ossa Mt. 6-17.VII.2011, Krause Rene leg.
L=32 mm
Copyright © Ruslan Panin
Carabus (Procrustes) chevrolati thirki Chaudoir, 1857
Male+Female adult
Turkey, Rize prov., Ikizdere distr., Rize Daglari,
47 km to SE from lyidere, Sivrikaya Koyu, h=2025 m.
30.VI-5.VII.2011, A. Vlasenko leg.
male - 28 mm, female - 31 mm.
Copyright © Alexandr Safronov
Carabus (Procrustes) anatolicus anatolicus Chaudoir, 1857
Male adult
Cyprus, Troodos, 17.2.1V.2010,. Manfred Egger leg.
L=27 mm
Copyright © Ruslan Panin
Carabus (Procrustes) banoni Dejean, 1829
Male adult
Greece,W Crete Island, Mili gorge ,12.V.2009, Manfred Egger leg.
L=26 mm
Copyright © Ruslan Panin
Carabus (Procrustes) coriaceus mopsucrenae Peyron, 1858
Male adult
SC-Turkey, Adana Prov., Doganbeyli, 8.V.2001, Snizhek M. leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Procrustes) talyschensis talyschensis Menetries, 1832
Female adult
Azerbaijan, Talysh Mts., Zuvand, Kalvaz vill., h=1650 m, 15.VII.2003, Dolin V.G. leg.
L=32 mm
Copyright © Ruslan Panin
Carabus (Procrustes) piochardi piochardi Gehin, 1883
Male adult
C-Lebanon, Faraya vill., Mt. Djebel Sannin h=1800m, X.2012, C. Reuter leg. <br /> L=31 mm
Copyright © Ruslan Panin
Carabus (Procrustes) banoni Dejean, 1829
Male adult
Crete, Agia Irini (Sélinos); 31mm
Copyright © Aleš Sedláček
Carabus (Procrustes) banoni Dejean, 1829
Female adult
Crete, Agia Irini (Sélinos); 34mm
Copyright © Aleš Sedláček
Carabus (Procrustes) talyschensis Menetries, 1832
Female adult
det. K. Makarov
Copyright © Alexander Anichtchenko
Carabus (Procrustes) paiafa acuticollis Motschulsky, 1858
Male adult
SC-Turkey: Alania (Irmasan), rte 695 au km 25 Akseki, h= 1225 m, 02.X.2003
Copyright © Igor Solodovnikov
Carabus (Procrustes) piochardi pierpaolo Kleinfeld & Rapuzzi, 2004
Female adult
29 mm
Copyright © Frank Kleinfeld
Carabus (Procrustes) piochardi praestigiator Morawitz, 1886
Male adult
C-Lebanon, Byblos /Jbeil Distr., Kfardebian env., h=1200m, XI.2014, C. Reuter leg. L=31 mm
Copyright © Ruslan Panin
Carabus (Procrustes) coriaceus rugifer Kraatz, 1877
Female adult
W.UKRAINE, NE Carpathians, Lviv Reg., Skole distr., Zelemyanka Mt.Range h=700m, 26.V-1.VII.2010, Panin R. leg. L=43 mm
Copyright © Ruslan Panin
Carabus (Procrustes) paiafa paiafa White, 1845
Male adult
Turkey, Antalya Prov., Finikie env., h~200 20.V.2015 L=41 mm
Copyright © Kirill Makarov
Carabus (Procrustes) coriaceus coriaceus Linne, 1758
Female adult
Belgique : Prov. Luxembourg, Harnoncourt 31.vii.1975 - M. Grotz Leg. (Size : 36 mm)
Copyright © Noël Mal
Carabus (Procrustes) piochardi piochardi Gehin, 1883 Lectotype
Male adult
Liban, Jebel Sannin. http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec511
Copyright © MNHN
Carabus (Procrustes) piochardi labruleriei Gehin, 1883
Male adult
Lebanon. det. Mingyi Tian
Copyright © Mingyi Tian
Carabus (Procrustes) coriaceus ressli Mandl, 1961
Male adult
SC-Turkey: Hatay Prov, Dörtyol, Nur Mts, Topaktaş h=900-1200 m, 12.V.2005, D. Prunier leg.
Copyright © Ruslan Panin
Carabus (Procrustes) coriaceus ressli Mandl, 1961
Female adult
SC-Turkey: Hatay Prov, Dörtyol, Nur Mts, Topaktaş h=900-1200 m, 12.V.2005, D. Prunier leg. L=35 mm
Copyright © Ruslan Panin
Carabus (Procrustes) chevrolati internatus Heinz & Korge, 1964
Female adult
Turkey, Bolu, Gerede, Esentepe 1350m, 1.7.1994, leg. K. Staven
Copyright © Alexander Anichtchenko
Carabus (Procrustes) coriaceus excavatus Charpentier, 1825
Male adult
W Croatia, Velebit Mts, Jablanac Stirovaca h=998 m, 44.71124, 14.95673,3.VI-7.VII.2017, G. Proscia leg., L= 37 mm
Copyright © Ruslan Panin
Carabus (Procrustes) coriaceus excavatus Charpentier, 1825
Female adult
W Croatia, Velebit Mts, Jablanac Stirovaca h=998 m, 44.71124, 14.95673,3.VI-7.VII.2017, G. Proscia leg., L= 38 mm
Copyright © Ruslan Panin
Carabus (Procrustes) mulsantianus mulsantianus Morawitz, 1886 (as akbesianus)
Male adult
Anatolia Mer., Martavan, ca. 600m, 32 km NW Kilis, 15.4.1989, Heinz leg.(cAA)
Copyright © Alexander Anichtchenko -
Pseudocoptolabrus taxon page (42 photos)
Carabus (Pseudocoptolabrus) taliensis taliensis (Fairmaire, 1886)
Male adult
China, Yunnan, Lijiang env., 2700 m, 10.6.1998
Copyright © Alexander Anichtchenko
Carabus (Pseudocoptolabrus) taliensis s.str. (as yulongxuensis)
Male adult
male, 26mm
Copyright © Boleslav Brezina
Carabus (Pseudocoptolabrus) taliensis s.str. (as yulongxuensis)
Male adult
male, 24mm
Copyright © Boleslav Brezina
Carabus (Pseudocoptolabrus) taliensis s.str. (as yulongxuensis)
Female adult
female, 29mm
Copyright © Boleslav Brezina
Carabus (Pseudocoptolabrus) taliensis yanmenensis Deuve, 1996
Male adult
male, 27mm
Copyright © Boleslav Brezina
Carabus (Pseudocoptolabrus) taliensis yanmenensis Deuve, 1996
Female adult
female, 28mm
Copyright © Boleslav Brezina
Carabus (Pseudocoptolabrus) taliensis xueshanicola Deuve, 1992
Female adult
female, 30mm
Copyright © Boleslav Brezina
Carabus (Pseudocoptolabrus) taliensis xueshanicola Deuve, 1992
Male adult
male, 26mm
Copyright © Boleslav Brezina
Carabus (Pseudocoptolabrus) taliensis xueshanicola Deuve, 1992
Male adult
male, 27mm
Copyright © Boleslav Brezina
Carabus (Pseudocoptolabrus) taliensis xueshanicola Deuve, 1992
Male adult
male, 28mm
Copyright © Boleslav Brezina
Carabus (Pseudocoptolabrus) taliensis peterheisei Holotype
Male adult
Holotype
Copyright © Frank Kleinfeld
Carabus (Pseudocoptolabrus) taliensis myanmarensis Imura, 1999
Female adult
female, 36 mm
Copyright © Boleslav Brezina
Carabus (Pseudocoptolabrus) taliensis mulianus Deuve, 1995
Male adult
male, 27mm
Copyright © Boleslav Brezina
Carabus (Pseudocoptolabrus) taliensis liqiuensis Imura, 1999
Male adult
male, 32mm
Copyright © Boleslav Brezina
Carabus (Pseudocoptolabrus) taliensis lijiangensis Paratype
Female adult
female, 33mm
Copyright © Boleslav Brezina
Carabus (Pseudocoptolabrus) taliensis jingpingensis Deuve, 2005 Paratype
Male adult
male, 26mm
Copyright © Boleslav Brezina
Carabus (Pseudocoptolabrus) taliensis jingpingensis Deuve, 2005 Paratype
Female adult
female, 30mm
Copyright © Boleslav Brezina
Carabus (Pseudocoptolabrus) nosei taronensis Imura, 2001 Paratype
Female adult
female, 28mm
Copyright © Boleslav Brezina
Carabus (Pseudocoptolabrus) burmanensis Breuning, 1932
Male adult
male, 23mm
Copyright © Boleslav Brezina
Carabus (Pseudocoptolabrus) armiger armiger Imura, 1997
Female adult
female, 30mm
Copyright © Boleslav Brezina
Carabus (Pseudocoptolabrus) businskyi Deuve, 1990
Male adult
CH, NW-Yunnan (Lijiang) Yulongxue Shan, E-slope
27:10:50N/100:12:26E/ 4400m, alpine meadows 12.VII.2000
K. & B. Brezina, M.Brychcin, M. Hackel
Copyright © Boleslav Brezina
Carabus (Pseudocoptolabrus) taliensis (Fairmaire, 1886) Holotype
Female adult
Yunnan; http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec147
Copyright ©
Carabus (Pseudocoptolabrus) taliensis kawagarbo Deuve, 2018 Holotype
Male adult
Chine, Nord-Ouest du Yunnan, Monts Hengduan Shan, au nord du Mont Meili Xueshan (Kawagarbo), à l’ouest du village de Shuaqu, 4268 mètres, 28°35’12’’N-98°40’36’’E, in coll. R. Sehnal, à Unhošt (République Tchèque).
Copyright © Thierry Deuve
Carabus (Pseudocoptolabrus) watanabei valdemar Deuve, 2016 Holotype
Male adult
Chine, Yunnan, Dêqên, monts Hengduan Shan, près du col de Doker La, 3600 m. (in coll. I. Brunk)
Copyright © Thierry Deuve
Carabus (Pseudocoptolabrus) taliensis lijiangensis Deuve, 1990
Male adult
N Yunnan, S slope Yulong mts., near Lijiang, 3900m, 15-25.6.2016, leg. V. Patrikeev. det. A. Anichtchenko, 2020
Copyright © Alexander Anichtchenko
Carabus (Pseudocoptolabrus) chortenensis Cavazzuti, 2005
Male adult
specimen from type locality. Image from Rapuzzi, 2020
Copyright © - - -
Carabus (Pseudocoptolabrus) taliensis xerophilous Rapuzzi, 2020 Holotype
Female adult
nE Yunnan, dahai xiang, Vii.2019, Wang legit. Image from Rapuzzi, 2020
Copyright ©
Carabus (Pseudocoptolabrus) taliensis ridilioides Deuve & Tian, 2020 Holotype
Male adult
Chine, Yunnan, col à env. 17 km au nord-ouest de Shangri-La (Zhongdian), 3684 mètres, 27°55’33’’N-99°36’42’’E, in coll. R. Sehnal (Unhošt, République Tchèque). Image from Deuve & Tian, 2020
Copyright ©
Carabus (Pseudocoptolabrus) pseudochortenensis Deuve & Tian, 2020 Paratype
Male adult
Chine, Sichuan, entre Yanyuan et Muli, col de Mianbu, 3244 mètres, 27,68638°N-101,22335°E (Shi Hongliang et Liu Ye), in coll. Institut Zoologique, Academia Sinica, Pékin (Chine). Image from Deuve & Tian, 2020
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Carabus (Pseudocoptolabrus) zuzkae Imura & Březina, 2012 Holotype Paratype
Male+Female adult
Holotype: m, Northeast of Fugong, 3,440 m in altitude, 26.57.32N/ 98.56.44E, on the Nu Shan Mountain Range, in Fugong Xian, of Nujianglilizu- zizhizhou, northwestern Yunnan, Southwest China, 1.VI.2006, preserved in the Zoological Institute of Academy of Sciences, St. Petersburg, Russia. Paratype: female, same locality, 31.V.2006, preserved in the collection of the first author, Y. IBJG6 (Yokohama).Image from Imura & Březina, 2012
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Carabus (Pseudocoptolabrus) watanabei watanabei (Imura, 2003) Holotype
Male adult
Imau Bum of northern Myanmar. Image from Imura & Březina, 2012
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Carabus (Pseudocoptolabrus) belousovianus Imura, 2011 Holotype Paratype
Male+Female adult
uppermost zone of the Chuandongzi Massif. '27・20' 21-34" NI IOI・ 32' 42-52" E. 4070・4135 m in alt . south of Yanyuan, Yanyuan Xian. Liangshan-yizu-Zizhizhou. southern Sichuan, Southwest China, 4-VII-2010. preserved in the Zoological Institute of Academy of Sciences. St. Petersburg. Russia. Image from Imura, 2011
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Carabus (Pseudocoptolabrus) masahiroi (Imura, 2006) Holotype
Male adult
Chudu Razi Massif, E. Kachin, N. Myanmar, VIII-2005, to be preserved in the National Science Museum (Nat. Hist), Tokyo. Image from Imura, 2011
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Carabus (Pseudocoptolabrus) branaungi liukuensis (Imura, 2007) Holotype
Male adult
south-southwest of Liuku , (25°41'31"N/98°45 '47"E), 3,545 m in altitude, in Lushui Xian, of western Yunnan, China, 1 9-V-2006, preserved in the collection of the Department of Zoology, National Science Museum (Nat. Hist.), Tokyo. Image from Imura, 2011
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Carabus (Pseudocoptolabrus) armiger legionarius Cavazzuti & Rapuzzi, 2005 Holotype
Female adult
Cina, Sichuan meridionale, Passo a sud-ovest di Mianning, 2650 m. (in coll. Rapuzzi). Image from Cavazzuti & Rapuzzi, 2005
Copyright © - - -
Carabus (Pseudocoptolabrus) taliensis jiaozicola Deuve, 2019 Holotype
Male adult
Chine, Yunnan septentrional, Mont Jiaozi Xueshan, 3640 m, 26°05’N-102°51’E, in coll. Jaroslav Turna, Kostelec na Hane (République Tchèque)
Copyright © Thierry Deuve
Carabus (Pseudocoptolabrus) taliensis huiliensis Deuve, Liu & Liang, 2024 Holotype
Male adult
Chine, province du Yunnan, canton de Huili, Mont Longzhou Shan, chemin vers le sommet, 3024 m, 26,8117°N 102,2238°E, forêt de Rhododendron (Hongliang Shi & Bo Liu leg., 19.VII.2015), in IOZ. Image from Deuve 2024
Copyright © - - -
Carabus (Pseudocoptolabrus) taliensis shanlenggang Rapuzzi, 2024 Holotype
Male adult
Sichuan, 30 km behind Meigu to Shanglenggang; 2700 m; Jul. 2022
Copyright © Alexander Anichtchenko -
Pseudocranion taxon page (14 photos)
Carabus (Pseudocranion) sackeni sackeni Semenov, 1898
Male adult
male, 23mm
Copyright © Boleslav Brezina
Carabus (Pseudocranion) sackeni remondianus Deuve, 1996 Paratype
Female adult
Museum of Institute of Systematic Biology (Daugavpils, Latvia)
Copyright © Alexander Anichtchenko
Carabus (Pseudocranion) labrangicus macilentior Deuve, 1996
Female adult
female, 21mm
Copyright © Boleslav Brezina
Carabus (Pseudocranion) gansuensis jedompansis Deuve, 1989
Male adult
China, Sichuan, Langmusi 3200m 7.1999
Copyright © Alexander Anichtchenko
Carabus (Pagocarabus) jiudingensis jiudingensis Deuve, 1994
Female adult
Copyright © Boleslav Brezina
Carabus (Pseudocranion) lazikouensis lazikouensis (as kundaensis Deuve, 2003) Paratype
Female adult
C-China, S-Gansu: SSW of Minxian: Qilian Shan: W Kunda, 34:10N/103:47E, h=2800-3000m, forest, 7.VII.2002, Belousov & Kabak leg.
L=24 mm
Copyright © Ruslan Panin
Carabus (Pseudocranion) taibaishanicus taibaishanicus Deuve, 1989
Female adult
China, W-Shaanxi, Taibai Shan Mts, btw. Houzhenzi/Ba Xian Tai, nr. Nan Tian Men h=3200m, 33°55'17"N/ 107°46'53"E, 25-30.VII.2011, Li Wuyang leg. L=29 mm
Copyright © Ruslan Panin
Carabus (Pseudocranion) zhanglaensis potrikeevi Deuve, 2015 Holotype
Male adult
Chine, Gansu, env. 60 km à l’est de Têwo, Palanama, 4000 mètres, in coll. Andreï Gorodinski, Moscou (Russie).
Copyright © Thierry Deuve
Carabus (Pseudocranion) gansuensis tiro Semenov, 1898 Lectotype
Male adult
Sun-pan
Copyright © Thierry Deuve
Carabus (Pseudocranion) gansuensis altiphilus Deuve, 2010 Holotype
Male adult
Chine, Nord Sichuan, alentours meridionaux du col situe a 12 km a l'est-sud-est de Zhangla (pelouses alpines, eboulis) 32°43N 103°45E (in coll. J. Kalab)
Copyright © Thierry Deuve
Carabus (Pseudocranion) shuamaluko luhuanus Cavazzuti & Rapuzzi, 2005 Holotype
Male adult
Cina, Sichuan, Heishui Xian (Luhua), Dakog Bing Chuan 2700 m. (Holotypus in coll. Cavazzuti)
Copyright © - - -
Carabus (Pseudocranion) absonus Cavazzuti & Rapuzzi, 2005 Holotype
Male adult
Cina, Shaanxi meridionale, Zhiyang, Dabashan 1600 m i meridionale, Zhiyang, Dabashan 1600 m. VII-2003 (in coll. Rapuzzi). Image from Cavazzuti & Rapuzzi, 2005
Copyright © - - -
Carabus (Pseudocranion) kitawakiellus Imura, 1995
Female adult
Image courtesy A. Gorodinsky
Copyright © - - - -
Relictocarabus taxon page (1 photos)
Carabus (Relictocarabus) meurguesianus Ledoux, 1990
Male adult
Image from molbiol.ru
Copyright © - - - -
Rhigocarabus taxon page (163 photos)
Carabus (Rhigocarabus) xiei sanggarparicus Deuve, 1996 Holotype
Male adult
19mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) xiei sanggarparicus Deuve, 1996 Paratype
Female adult
20mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) xiei barkamensis Deuve, 1994 Holotype
Male adult
Holotype, male, 19mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) xiei barkamensis Deuve, 1994 Paratype
Male adult
Paratype, female, 19mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) xiangchengicus qingmaicus Deuve, 2007
Male adult
male, 15mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) xiangchengicus qingmaicus Deuve, 2007
Female adult
female, 18mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) xiangchengicus qingmaicus Deuve, 2007
Female adult
female, 16mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) xiangchengicus sangpiensis Deuve, 2007 Paratype
Male adult
male, 15 mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) subindigestus subindigestus Deuve, 1994
Female adult
female, 15mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) subindigestus subindigestus Deuve, 1994
Male adult
male, 16mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) subindigestus subindigestus Deuve, 1994
Male adult
male, 15mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) subindigestus subindigestus Deuve, 1994
Male adult
male, 16mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) sehnali Brezina & Hackel, 2006 Holotype Paratype
Male adult
male, 20mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) sehnali Brezina & Hackel, 2006
Female adult
female, 23mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) roborowskii roborowskii Semenov, 1887
Male adult
China, E Tibet, Chegarla 4800m, 20 km E
of Rioche, 26.6.97 A. Wrzecionko leg.
Copyright © Alexander Anichtchenko
Carabus (Rhigocarabus) rhododendron Deuve & Imura, 1991
Male adult
male, 18mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) rhododendron Deuve & Imura, 1991
Female adult
female, 19mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) rhododendron Deuve & Imura, 1991
Male adult
male, 17mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) quindecim Cavazzuti, 2002 Holotype
Male adult
Holotype, 15mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) pseudopusio Deuve, 1996 Paratype
Male adult
male, 15mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) poeta Semenov, 1898 Holotype Paratype Lectotype
Male adult
male, 17mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) poeta (as korsakowi Semenov, 1898) Holotype
Female adult
Holotype, female 18 mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) poeta (as korsakowi Semenov, 1898) Syntypes
Female adult
female, 16mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) poeta (as korsakowi Semenov, 1898) Lectotype
Male adult
male, 17mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) pepek Imura & Kalab, 2006 Paratype
Male adult
male, 17mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) pepek Imura & Kalab, 2006 Paratype
Female adult
female, 18mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) morawitzianus breviculus Cavazzuti & Ratti, 1998 Paratype
Male adult
male, 15 mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) itzingeri rugulosior Deuve, 1996 Paratype Paratype
Male adult
male, 18mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) itzingeri itzingeri Breuning, 1934
Male adult
male, 19mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) itzingeri itzingeri Breuning, 1934
Female adult
female, 20mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) itzingeri kozaburoi Imura, 1999 Paratype
Male adult
male, 16mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) itzingeri kozaburoi Imura, 1999 Paratype
Female adult
female, 18mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) handelmazzettii handelmazzettii Mandl, 1955
Male adult
male, 18mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) handelmazzettii handelmazzettii Mandl, 1955
Female adult
female, 20mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) handelmazzettii handelmazzettii Mandl, 1955
Female adult
female, 21mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) handelmazzettii habaicus Deuve, 2005 Paratype
Male adult
male, 16mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) pusio pusio s.str.
Male adult
male, 16mm
(as hylonomus)
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) pusio pusio s.str.
Female adult
female, 17mm
(as hylonomus)
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) gracilicollis gracilicollis Semenov, 1887
Male adult
male, 15mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) gracilicollis gracilicollis Semenov, 1887
Female adult
female, 16mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) gigoloides priapeius Cavazzuti, 2006
Male adult
male, 20mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) gigoloides jone Brezina & Hackel, 2004 Holotype Paratype
Male adult
male, 19mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) gigoloides jone Brezina & Hackel, 2004
Female adult
female, 20mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) gigolo (as qilingianus Cavazzuti, 2006)
Male adult
18mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) gigolo Heinz & Brezina, 1996 Paratype
Male adult
20mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) gigolo Heinz & Brezina, 1996 Paratype
Male adult
20 mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) gigolo Heinz & Brezina, 1996 Paratype
Female adult
21 mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) gentleman Brezina & Hackel, 2004 Holotype
Male adult
male, 19mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) gentleman Brezina & Hackel, 2004 Paratype
Female adult
female, 20mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) dacatraianus Deuve, 1995
Male adult
China, E Tibet, Angenla 5000-5100m
45 km W of Bachen 29.6-2.7.1997 Wrzecionko leg.
16 mm
Copyright © Alexander Anichtchenko
Carabus (Rhigocarabus) cateniger cateniger s.str. (as sommitalis)
Male adult
male, 16 mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) cateniger cateniger Morawitz, 1886
Female adult
female, 19mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) cateniger cateniger Morawitz, 1886
Male adult
male, 18mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) cateniger saltuensis Cavazzuti, 2006 Paratype
Female adult
female, 19mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) cateniger dabanicola Deuve, 1998
Male adult
male, 16mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) cateniger dabanicola Deuve, 1998
Female adult
female, 18mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) buddaicus bocsiki Deuve, 2007 Holotype
Male adult
male, 20 mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) buddaicus angenlaicus Deuve, 2007 Paratype
Male adult
male, 18 mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) broukpytlik Brezina & Hackel, 2004 Holotype Paratype
Male adult
male, 19mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) broukpytlik Brezina & Hackel, 2004
Female adult
female, 20mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) broukpytlik Brezina & Hackel, 2004
Male adult
male, 16mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) morawitzianus breviculus Cavazzuti & Ratti, 1998 Paratype
Male adult
China, N Tibet, Tangula Shan Mts., Tangula, pass 5100-5300m, 8-11.7.1997 A. Wrzecionko leg.
Copyright © Alexander Anichtchenko
Carabus (Rhigocarabus) batangicus fickleri Korell & Richter, 1996 Paratype
Male adult
male, 16 mm
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) wengdaensis Kleinfeld & Puchner, 2009 Holotype
Male adult
HOLOTYPUS Sichuan, Str. Wengda-Luhuo, Pass 20 km südl. Wengda, 4100m (ca. 31°43'N/100°44'E), VIII.2007, leg. Ding - coll. Puchner 20mm lang / 7 mm breit
Copyright © Frank Kleinfeld
Carabus (Rhigocarabus) buddaicus gansuicus Deuve, 1991
Male adult
China, NC-Sichuan (ZOIGE), LANGMUSI (=DOGCANGLHAMO), 34:03:48N/102:38:08E, h=4100m, alpine meadows, screes, 16-18.VII.1999, B.Brezina leg.
L=20 mm
Copyright © Ruslan Panin
Carabus (Rhigocarabus) indigestus confucius Breuning, 1933
Female adult
C-China, E Tibet,W-Sichuan prov., Kangding, Zheduo Pass h=4300m 30.VI.2005 , V.Benes leg,
L=17 mm
Copyright © Ruslan Panin
Carabus (Rhigocarabus) ladygini degensis Deuve, 1989
Male adult
C-China, NW- Sichuan prov., Garzê Tibetan AP, Pass Garzê – Sêrtar, 19.VII.2007, V.Benes leg.
L=17 mm
Copyright © Ruslan Panin
Carabus (Rhigocarabus) buddaicus linxiaicus Deuve, 1995
Male adult
China, S-Gansu (LABRANG):, valley.E of Ponggartang, 35:12-13N/102:54-55E, h=3000-3700m, alpine meadows, coniferous forests,10.-14.VII.1999, B.Brezina leg. L=20 mm
Copyright © Ruslan Panin
Carabus (Rhigocarabus) shaluishan Kleinfeld & Puchner, 2011 Holotype
Male adult
CHINA, Prov. Sichuan, zwischen Garzê und Baiyü, Berg 25 km südlich von Garzê, Nordseite, 3820m, - 31°31.985 N / /99°58.670´ E, 30.V-14.VI.2010, leg. et in coll. Puchner.
19 mm
Foto Kleinfeld XII.2010
Copyright © Frank Kleinfeld
Carabus (Rhigocarabus) shaluishan Kleinfeld & Puchner, 2011 Paratype
Female adult
CHINA, Prov. Sichuan, zwischen Garzê und Baiyü, Berg 25 km südlich von Garzê, Nordseite, 3820m, - 31°31.985 N / /99°58.670´ E, 30.V-14.VI.2010, leg. et in coll. Puchner.
18 mm Foto Kleinfeld XII.2010
Copyright © Frank Kleinfeld
Carabus (Rhigocarabus) allegroi Cavazzuti & Rapuzzi, 2007
Male adult
CINA S. Gansu, Qiling Shan, Pass N. Tewo (N 34°16’-E 103°03’), 4.100 m, 17.VI-1.VII.2010 (Colleccion Gianni Allegro)
Copyright © Gianni Allegro
Carabus (Rhigocarabus) morawitzianus morawitzianus Semenov, 1887
Male adult
C China, Qinghai, Zadoi County, Toramarkog h~4600 m, VI.2001, P. Cavazzuti leg. L=17 mm
Copyright © Ruslan Panin
Carabus (Rhigocarabus) turnai turnai Deuve, 1994
Male adult
China, Tibet, W-Sichuan, pass btw Sêrtar / Luhuo h=4200 m, 28.VI.2011, V.Patrikeev leg.L=19 mm
Copyright © Ruslan Panin
Carabus (Rhigocarabus) turnai turnai Deuve, 1994
Female adult
China, Tibet, W-Sichuan, pass btw Sêrtar / Luhuo h=4200 m, 28.VI.2011, V.Patrikeev leg. L=21 mm
Copyright © Ruslan Panin
Carabus (Rhigocarabus) indigestus dilbu Deuve & Kaláb, 2018 Holotype
Male adult
Chine, Ouest-Sichuan, col à 9 km à l’ouest de Babang, (route de Dêgê), zone sud du col, 4600-4800 mètres, 31º39’N/98º43’E (pelouses alpines, éboulis), in coll. J. Kaláb, à Jinačovice (République Tchèque).
Copyright © Thierry Deuve
Carabus (Rhigocarabus) indigestus ronggaicus Deuve & Kaláb, 2018 Holotype
Male adult
Chine, Ouest-Sichuan, col à 17 km au SE-SSE de Ronggai, 4800 mètres, 30º59’N/99º00’E (pelouses alpines, éboulis), in coll. J. Kaláb, à Jinačovice (République Tchèque).
Copyright © Thierry Deuve
Carabus (Rhigocarabus) ladygini babangensis Deuve & Kaláb, 2018 Holotype
Male adult
Chine, Ouest-Sichuan, col à 9 km à l’ouest de Babang, (route de Dêgê), zone sud du col, 4600-4800 mètres, 31º39’N/98º43’E (pelouses alpines, éboulis), in coll. J. Kaláb, à Jinačovice (République Tchèque).
Copyright © Thierry Deuve
C. (Rhigocarabus) itzingeri gongshanensis Deuve, 2016 Holotype
Male adult
Chine, Yunnan, Monts Gaoligong Shan, Préfecture de Nujiang, Gongshan Xian, 21 km à vol d’oiseau à l’ouest de Gongshan, col entre Gongshan et Dulong, 3300-3680 mètres, 27,69655°N-98,45349°E (D. H. Kavanaugh, C. E. Griswold, H. B. Liang, D. Ubick, D. Z. Dobg leg., 16-17 juillet 2000, in coll. Institut Zoologique, Pékin.
Copyright © Thierry Deuve
Carabus (Rhigocarabus) dungchen Deuve & Kalab, 2019 Paratype
Male adult
Chine, Sichuan occidental, montagnes à 13 km au S-SSW de Cuo’a, 4700 mètres, 31º42’N/99º24’E (prairies alpines et éboulis).
Copyright © Thierry Deuve
Carabus (Rhigocarabus) ladygini ganzicola Deuve & Kalab, 2019 Holotype
Male adult
Chine, Sichuan occidental, montagnes à 17 km au nord-nord-ouest de Ganzi (= Garzê), 31º46’N/99º56’E, 4700-4800 mètres (prairies alpines et éboulis)
Copyright © Thierry Deuve
Carabus (Rhigocarabus) morawitzianus morawitzianus Semenov, 1887
Male adult
Qinghai, Anemaqen mts pass, 20 km N of Maduo town, 4500 m, 6.7.2012, V. Patrikeev leg.; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko
Carabus (Rhigocarabus) buddaicus gansuicus Deuve, 1991
Male adult
N Sichuan, Langmusi vill. 3700 m, 1-25.7.2018, Leg. V. Patrikeev. det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko
Carabus (Rhigocarabus) gigolo Heinz & Brezina, 1996
Male adult
S Gansu, near Tewo, 8.8.2014, V. Patrikeev leg.
Copyright © Alexander Anichtchenko
Carabus (Rhigocarabus) roborowskii langmusiensis Deuve, 1996
Male adult
N Sichuan, near Langmusi, 3700m 1-25.7.2018, V. Patrikeev leg. det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko
Carabus (Rhigocarabus) pseudopusio Deuve, 1996
Male adult
NW Sichuan, Zonggrannkow pass, 60km W Heishue city, 4000m, 22.7.2017 V.Patrikeev leg. (cAA). det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko
Carabus (Rhigocarabus) gracilicollis pusio Semenov, 1898
Male adult
N Sichuan, pass between Songpan and Juzaigow, W side, 25-30.6.2015, 4200 m V. Patrikeev leg. (cAA). det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko
Carabus (Rhigocarabus) ladygini degensis Deuve, 1989
Male adult
NW Sichuan, Chola Mts, Maniganggo vill., 4600m, 15.6.2015 V. Patrikeev leg. (cAA). det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko
Carabus (Rhigocarabus) thais thais Heinz, 1997
Male adult
E Qinghai-Gansu border, Dalia pass 50km W Lixia town 3800m 20.7.2019 (cAA) det. Anichtchenko 2020
Copyright © Alexander Anichtchenko
Carabus (Rhigocarabus) cateniger copulans Semenov, 1898
Male adult
E Qinghai, Laji Shan, near Garang, 3900m, 19.6.2011, Patrikeev V. leg (cAA) det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko
Carabus (Rhigocarabus) cateniger papaoensis Cavazzuti, 2006
Male adult
E Qinghai, near Babao town, 3800m 5.8.12 Patrikeev V. leg. (cAA) det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko
Carabus (Rhigocarabus) cateniger cateniger Morawitz, 1886
Male adult
NW Gansu, 80 km S Jayuguan, Tulai shan, Qionglai mts, 10.7.2014, 4200 m, Patrikeev leg (cAA)
Copyright © Alexander Anichtchenko
Carabus (Rhigocarabus) indigestus dongziensis Deuve & Kaláb, 2022 Holotype
Male adult
Chine, Sud-Sichuan, montagnes à 10 km au nordouest de Dongzi, 4500 m, 28°34’N/100°46’E, pelouses alpines et éboulis, in coll. J. Kaláb, à Jinačovice (République Tchèque).
Copyright © Thierry Deuve
Carabus (Rhigocarabus) ladygini shiliensis Deuve et Müller, 2022 Holotype
Male adult
Chine, Sichuan nord-occidental, environs de Shili, Zango (31,85°N-101,05°E), 4000 m (V. Benes leg.), in coll. A. Müller, Vienne (Autriche).
Copyright © Thierry Deuve
Carabus (Rhigocarabus) casanova Imura & Brezina, 2008 Holotype Paratype
Male+Female adult
north-northwest of Kaba, 34°11'09-14"N I 103°23' 10-21 "E, 3,800-4,100 m in altitude, on the Die Shan Mountain Range in Tewo Xian, of Gannan Zangzu Zizhizhou, in southern Gansu, China, 14-VI-2006. HT to be deposited in the Department of Zoology, National Museum of Nature and Science, Tokyo
Copyright © Boleslav Brezina
Carabus (Rhigocarabus) nianjuaensis Imura, 2016 Holotype Paratype
Male adult
China, NW. Yunnan, NE. of Weixi-lisuzu Zizhixian ［维西傈僳族自治县］, 8.65 km west of Laoantong ［老安 统］, 7.35 km east of Mt. Nianjua ［念菊阿］, 27°44ʼ41” N / 98°56ʼ58” E, 3950 m in altitude, 16-VI-2015, to be preserved in the Zoological Institute of Academy of Sciences, St. Petersburg, Russia. Paratype: 1♂, same data as for the holotype, preserved in the collections of Y. Imura (Yokohama). Image from Imura, 2016
Copyright © - - -
Carabus (Rhigocarabus) kozaburoi Imura, 1999 Paratype
Male adult
Gaoligong Shan, N. Myanmar. Image from Imura, 2016
Copyright © - - -
Carabus (Rhigocarabus) itzingeri rugulosior Deuve, 1996 Holotype
Male adult
NW-Yunnan: Hengduan Shan: Yanmen env.: 28:06N/98:52E. Image from Imura, 2016
Copyright © - - -
Carabus (Rhigocarabus) itzingeri Breuning, 1934 Holotype
Female adult
Batang, W. Schuan. Image from Imura, 2016
Copyright © - - -
Carabus (Rhigocarabus) itzingeri tanakai Imura, 1997
Male adult
NE. Putao, N. Myanmar. Image from Imura, 2016
Copyright © - - -
Carabus (Rhigocarabus) itzingeri choguy Imura, 2001 Holotype
Male adult
Chayu, E. Xizang. Image from Imura, 2016
Copyright © - - -
Carabus (Rhigocarabus) rhododendron Deuve & Imura, 1991 Holotype
Male adult
Mt. Bamaxue Shan, NW. Yunnan. Image from Imura, 2016
Copyright © - - -
Carabus (Rhigocarabus) rhododendron lidimaensis Imura, 2016 Holotype
Male adult
China, NW. Yunnan, near the borders between Weixi-lisuzu Zizhixian ［维西傈僳族自治县］ and Deqin Xian ［德钦县］, 4.3 km east of Lidima [里底 玛], 27°40ʼ43” N / 99 °19ʼ20” E, 4110 m in altitude, 23-VI-2015, to be preserved in the Zoological Institute of Academy of Sciences, St. Petersburg, Russia. Image from Imura, 2016
Copyright © - - -
Carabus (Rhigocarabus) rhododendron xuebangshanensis Imura, 2016 Holotype
Male adult
China, NW. Yunnan, near the borders between Lanping-baizu-pumizu Zizhixian [兰坪白族普米族自治县] and Jianchuan Xian [剑川县], NE. of Lanping [兰 坪], 0.95 km NNE. of Mt. Xuebang Shan [雪 邦 山], 26 °29ʼ14” N / 99 °30ʼ08” E, 4035 m in altitude, 29-V-2015, to be preserved in the Zoological Institute of Academy of Sciences, St. Petersburg, Russia. Image from Imura, 2016
Copyright © - - -
Carabus (Rhigocarabus) rhododendron xinshanchangensis Imura, 2016 Holotype
Male adult
China, NW. Yunnan, Lanping-baizu-pumizu Zizhixian [兰坪白族普米族自治县], ca. 10km NNE. of Lanping [兰坪], SE. of Xinshanchang [新山 场], 26 °30ʼ37” N / 99 °29ʼ26” E, 3780 m in altitude, 19-VI-2014, to be preserved in the Zoological Institute of Academy of Sciences, St. Petersburg, Russia. Image from Imura, 2016
Copyright © - - -
Carabus (Rhigocarabus) pseudopusio metallicior Deuve, 1996 Holotype
Male adult
NW-Sichuan: 23km SSW of Shangbazhai, 32.25N 102.57E 4300m
Copyright © Thierry Deuve
Carabus (Zheduocarabus) jiulongensis paolorapuzzii Cavazzuti & Rapuzzi, 2005 Holotype
Male adult
Cina, Sichuan meridionale, Sabde. 24-Vl/28-VI-2001, Kucera leg, in coll. Rapuzzi.
Copyright © - - -
Carabus (Rhigocarabus) cateniger cateniger (as sommitalis Cavazzuti, 2006)
Female adult
China, C-Gansu, Mts Qilian h=3378 m, road G227, Yangxiongzi env. 38.050917, 100.887267, 18-23.VI.2011, G.Proscia leg., L=19 mm
Copyright © Ruslan Panin
Carabus (Rhigocarabus) cateniger cateniger (as sommitalis Cavazzuti, 2006)
Male adult
China, C-Gansu, Mts Qilian h=3378 m, road G227, Yangxiongzi env. 38.050917, 100.887267, 18-23.VI.2011, G.Proscia leg., L=17 mm
Copyright © Ruslan Panin
Carabus (Rhigocarabus) cateniger dabanicola Deuve, 1998
Male adult
China, Qinghai, Xining city, 55km NE of Datong, Daban Shan h=3936 m, 37.348450, 101.397600, 17-23.VI.2011, G.Proscia leg., L= 16 mm
Copyright © Ruslan Panin
Carabus (Rhigocarabus) cateniger dabanicola Deuve, 1998
Female adult
China, Qinghai, Xining city, 55km NE of Datong, Daban Shan h=3936 m, 37.348450, 101.397600, 17-23.VI.2011, G.Proscia leg., L= 18.5 mm
Copyright © Ruslan Panin
Carabus (Rhigocarabus) roborowskii roborowskii Semenov, 1887
Male adult
China, E-Qinghai, Mts NW of Tongren city, 35.562778, 101.843667, h=3870 m, 15-26.VI.2011, G. Proscia leg./det.R.Panin, L=20 mm
Copyright © Ruslan Panin
Carabus (Rhigocarabus) roborowskii roborowskii Semenov, 1887
Female adult
China, E-Qinghai, Mts NW of Tongren city, 35.562778, 101.843667, h=3870 m, 15-26.VI.2011, G. Pros cia leg./det.R.Panin, L=20 mm
Copyright © Ruslan Panin
Carabus (Rhigocarabus) degensis ladyginioides Deuve, Liu & Liang, 2024 Holotype
Male adult
Chine, province de Qinghai, canton de Madoi, col de Bayan Har, 4827 m, 34,1267°N 97,6575°E, pelouses alpines et éboulis (Hongliang Shi, Zhongbao Lu, Weifeng Yan & Pingzhou Zhu leg., 30.VII.2017, dans des pièges), in IOZ. Image from Deuve et al, 2024
Copyright © - - -
Carabus (Rhigocarabus) rhododendron Deuve & Imura, 1991
Male adult
China, N Yunnan: Baimaxue Shan, S of Deqen, h=4300m, 28°20'N 99°03'E, 16-20.VI.2016, C.Reuter leg., L=18 mm
Copyright © Ruslan Panin
Carabus (Rhigocarabus) pepek Imura & Kalab, 2006 Holotype Paratype
Male+Female adult
SE of Zhenjiangguan (32°17'54-57"N/ 103°47'56"-47'56"E), 4,100-4,185 m, southeastern Songpan Xian, northern Sichuan, Southwest China, 17-VII-2005, deposited in the Department of Zoology, National Science Museum (Nat. Hist., Tokyo). Image from Imura & Kalab, 2006
Copyright © - - -
Carabus (Rhigocarabus) roborowskii sininensis Semenov, 1898 (as lajiensis Deuve & Imura, 1991)
Male adult
E Qinghai, E Laji Shan, N of Hualun, 3900m, 28.7.2019 Patrikeev leg (cAA). det. Anichtchenko A.
Copyright © Alexander Anichtchenko
Carabus (Rhigocarabus) roborowskii generalovi Deuve, 2014 Paratype
Male adult
S-Gansu, Dalijia Pass, 4000-4100m
Copyright © - - -
Carabus (Rhigocarabus) ladygini jigzhianus Cavazzuti, 2006
Female adult
S Quinghai, Guoluo mts., NW part, 4200-4500m, 11.7.17 Patrikeev V. leg (cAA). det. Anichtchenko A.
Copyright © Alexander Anichtchenko
Carabus (Rhigocarabus) buddaicus cholaensis Deuve, 2008 Holotype
Male adult
W-Sichuan: Chola Shan Pass, 4500m (in coll Benes, Prague)
Copyright © Thierry Deuve
Carabus (Rhigocarabus) buddaicus angenlaicus Deuve, 2008 Holotype
Male adult
E-Tibet: 30km E of Yanga, Angen La Pass 4800m (in coll Wrzecionko)
Copyright © Thierry Deuve
Carabus (Rhigocarabus) buddaicus obenbergeri Semenov & Znojko, 1932
Male adult
Qinghai, S of Kukunor lake, pass near Heka, 25.7.2010 (cAA)
Copyright © Alexander Anichtchenko
Carabus (Rhigocarabus) impavidus Cavazzuti, 2008 Holotype
Male adult
Cina Sichuan occidentale, Xiaojin, Passo a Nord di Fubian, 4550 m, Buma Shan, praterie alpine, 31°33159N, 102°24679E.
Copyright © - - - -
Scambocarabus taxon page (6 photos)
Carabus (Scambocarabus) sculptipennis Chaudoir, 1887
Male adult
China, Beijing, Xiaolongmeng, 2005, Liu ye coll.;
22mm; photo by Liuye
Copyright © Ye Liu
Carabus (Scambocarabus) kruberi kruberi Fischer, 1822
Male adult
Rusia, Chita reg., 23 km SW Borzia, Borzinskoe lake,
1.VI.2002 Anichtchenko A. leg.
Copyright © Alexander Anichtchenko
Carabus (Scambocarabus) kruberi Fischer, 1822
Male adult
China, Beijing, labagoumeng, 2004, Li fei coll.;
21mm; photo by Liuye
Copyright © Ye Liu
Carabus (Scambocarabus) kruberi laobeiensis Deuve & Li, 2000
Male adult
NE-China: Liaoning prov., Fengcheng City, Miaoyang, 12-17.VI.2011
L=27 mm
Copyright © Ruslan Panin
Carabus (Scambocarabus) kruberi bungii Gebler, 1830 Lectotype
Male adult
Not TYPE, because type was described from Altay. Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC416 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec416
Copyright © MNHN -
Semnocarabus taxon page (29 photos)
Carabus (Semnocarabus) transiliensis Semenov, 1896
Male adult
Kirgizstan, S slope Kungei Ala Too
Oi-Tal riv. 3000m 14.7.1999
13 mm
Copyright © Alexander Anichtchenko
Carabus (Semnocarabus) transiliensis Semenov, 1896
Male+Female adult
E KAZAKHSTAN,
Bolshoe Almaatinskoe Lake,30.VI.1992
15mm
Copyright © Ruslan Panin
Carabus (Semnocarabus) transiliensis Semenov, 1896
Male adult
SE-KAZAKHSTAN,
Ketmen Mts.,N-slopes h=2250-2500m.1-5.VI.1990
16mm
Copyright © Ruslan Panin
Carabus (Semnocarabus) regulus regulus Dohrn, 1882
Male adult
photo M. Smirnov from www.zin.ru
Copyright © Maksim Smirnov
Carabus (Semnocarabus) regulus regulus Dohrn, 1882
Male adult
E Kirgizia, Kaindy-Ketta mts. rng. Sary-Dzhas riv.
Maidadyr near Enelchek vill. 6-8.7.1999 Klimenko leg.
14 mm
Copyright © Alexander Anichtchenko
Carabus (Semnocarabus) erosus erosus Motschulsky, 1865
Male adult
Kirgizia, S Kungei Ala-Too, Kichi-Yuryukty
2800m 1.7.1999 Subankulov leg.
male, 21 mm
Copyright © Alexander Anichtchenko
Carabus (Semnocarabus) erosus kuldshaensis Ballion, 1878
Male adult
SE Kazakhstan, Ili riv., Bakanas vill.,<br /> 1-5.5.1999 Klimenko A. leg.
Copyright © Alexander Anichtchenko
Carabus (Semnocarabus) carbonicolor carbonicolor Morawitz, 1886
Male adult
E Kirgizia, Kaindy-Ketta mts. rng., Sary-Dzas riv.,
Maidadyr near Enelchek vill., 6-8.7.99 Klimenko leg.
Copyright © Alexander Anichtchenko
Carabus (Semnocarabus) carbonicolor callosus Semenov, 1896
Male adult
Kazakhstan, Bol. Alma-Atinka riv. 1600m 1.6.1999 Klimenko A. leg.
male, 19 mm
Copyright © Alexander Anichtchenko
Carabus (Semnocarabus) carbonicolor carbonicolor Morawitz, 1886
Male+Female adult
Kyrgyzstan, NC Tianshan, Kyrgyz Mt. R., Tuyuk riv., h=2000 м.
25-VI-1997 М. Danilevsky leg.
male - 20 mm, female - 21 mm.
Copyright © Alexandr Safronov
Carabus (Semnocarabus) erosus kuldshaensis Ballion, 1878
Male adult
S-Kazakhstan:Ili River valley,:near Bakanas,3-5.V.2002
L=21 mm
Copyright © Ruslan Panin
Carabus (Semnocarabus) carbonicolor callosus Semenov, 1896
Male+Female adult
KAZAKHSTAN,
Bolshoe Almaatinskoe Lake,30.VI.1992/ det.Panin R.
female 19mm
Copyright © Ruslan Panin
Carabus (Semnocarabus) carbonicolor korolkowi Semenov, 1896
Male adult
NC-Kyrgyzstan, Kyrgyz Mt.Rng , 10 km SW of Bishkek, Chon-Aryk env, h=1100m, 24.V.1998, /det. Panin R.
22 mm
Copyright © Ruslan Panin
Carabus (Semnocarabus) erosus karascharensis Eidam, 1931 Lectotype Paralectotypes
Male adult
Lectotype, left: “Ost Turkestan, Bagratsch-kul, Kurla 5.1902. Coll. Hsr,” “Syntypus,” “O. Leonhard,” “erosus subsp. karascharensis type Eidam, det. P. Eidam;” “coll DEI, Eberswalde;” “coll DEI, Müncheberg Col.–07059”
Copyright © Kirill Makarov
Carabus (Semnocarabus) erosus corax Semenov, 1896
Male adult
SE-Kazakhstan, Almaty reg., C Trans-Ili Alatau N-slopes, Kaskalen river Valley, Izvestkovaya vill., h=1700 m, 6.V.2015, D.Kovalchuk leg./det. R.Panin,
Copyright © Ruslan Panin
Carabus (Semnocarabus) erosus corax Semenov, 1896
Female adult
SE-Kazakhstan, Almaty reg., C Trans-Ili Alatau N-slopes, Kaskalen river Valley, Izvestkovaya vill., h=1700 m, 6.V.2015, D.Kovalchuk leg./det. R.Panin, L=24 mm
Copyright © Ruslan Panin
Carabus (Semnocarabus) carbonicolor korolkowi Semenov, 1896
Female adult
SE-Kazakhstan, Almaty, E Zailisky Alatau, Bokaydin-tau, Belshabdar valley h=1300 m, 5-27.VI.2002, A.Puchkov leg.L=21 mm
Copyright © Ruslan Panin
Carabus (Semnocarabus) carbonicolor carbonicolor Morawitz, 1886
Female adult
SE-Kazakhstan, Almaty reg., Khan Tengri Massif, Narynkol env.h=1800 m, 30-31.-V.2007, Shumov leg., L=21 mm
Copyright © Ruslan Panin
Carabus (Semnocarabus) erosus prope erosus Motschulsky, 1865
Male adult
SE-Kazakhstan, 70 km E of Almaty, E Trans-Ili Alatau N-slopes, valley Assy (Asy) riv. Assy plateau h=1700 m, 12.V.2015, L=21 mm,/ det. R. Panin
Copyright © Ruslan Panin
Carabus (Semnocarabus) rustemi Kabak, 2009
Male adult
SE-Kazakhstan, Almaty reg., near Bodokhudzir 43.916667, 79.716667 h=500 m, 12.VIII.2019, det. R.Panin, L=22 mm
Copyright © Ruslan Panin -
Shunichiocarabus taxon page (1 photos)
Carabus (Shunichiocarabus) uenoianus uenoianus Imura, 1995
Male adult
Copyright © - - - -
Sigenacarabus taxon page (6 photos)
Carabus (Pagocarabus) tryznai Brezina & Hackel, 2006 Holotype
Male adult
Copyright © Boleslav Brezina
Carabus (Pagocarabus) tryznai Brezina & Hackel, 2006 Paratype
Female adult
Copyright © Boleslav Brezina
Carabus (Pagocarabus) sichuanicola sigena Kleinfeld, 2000 Holotype
Male adult
Copyright © Frank Kleinfeld
Carabus (Pagocarabus) sichuanicola sigena Kleinfeld, 2000
Male adult
male, 26mm
Copyright © Boleslav Brezina
Carabus (Pagocarabus) sichuanicola morvanellus Deuve, 1998 Holotype
Female adult
Copyright © Boleslav Brezina
Carabus (Pagocarabus) sichuanicola sichuanicola Deuve, 1989 (as dromlae) Paratype
Male adult
Copyright © Boleslav Brezina
Carabus (Pagocarabus) caelestinus garpen Kalab, 2007 Holotype
Male adult
Copyright © Boleslav Brezina
Carabus (Pagocarabus) caelestinus garpen Kalab, 2007 Paratype
Female adult
female, 21 mm
Copyright © Boleslav Brezina
Carabus (Pagocarabus) caelestinus caelestinus (Imura, 2007) Paratype
Male adult
Copyright © Boleslav Brezina
Carabus (Pagocarabus) trachynodes trachynodes Bates, 1891 Lectotype
Male adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC89 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec89
Copyright © - - -
Carabus (Pagocarabus) sichuanicola sichuanicola (as drolmae Lassalle, 2003)
Female adult
China, Sichuan, NW Chengdu, Mt. Jiajin Shan h~4000 m,VI. 2016, local collector, L= 24 mm
Copyright © Ruslan Panin
Carabus (Sigenacarabus) draco lubasha Deuve, 2012
Male adult
China, C Sichuan, Jiyuding shan, 30 km N of Wenchuan, 3250 m 2.7.2017, leg. V. Patrikeev (cAA). det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko
Carabus (Sigenacarabus) trachynodes moxiensis Deuve, 1992
Male adult
China, W-Sichuan, W of Moxi, Daxue Range, Mt.Gongga Shan h=3000-3100m, 29°34′29.6″N /101°59′44.0″E, 10-12.VI.2018, Ch. Reuter leg. L=21 mm
Copyright © Ruslan Panin
Carabus (Sigenacarabus) trachynodes moxiensis Deuve, 1992
Male adult
China, W-Sichuan, Daxue Range, Mt.Gongga Shan h=3000-3100m, Hailuogou Glacier Forest Park, 29.574889, 101.995556, 10-12.VI.2018, Ch. Reuter leg., L=21 mm
Copyright © Ruslan Panin
Carabus (Sigenacarabus) huangcaoshanicus Rapuzzi, 2013 Holotype Paratype
Male+Female adult
China, Sichuan Province, Yaan, Shimian, Mt. Huangcao Shan, VII.2012;
Copyright © - - - -
Sphodristocarabus taxon page (73 photos)
Carabus (Sphodristocarabus) varians varians Fischer, 1827
Male+Female adult
female: Transcaucasia, Armenia, Amasia distr., Karakhachskiy mountain pass h=2270m, 3.VIII.1985 P.V.Kazaryan leg., 22mm<br /> male: Armenia, Amasia distr., Arpalach vill. VI.1999, P.V. Kazaryan leg.
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) blumenthaliensis basilewskianus Breuning & Ruspoli, 1970
Male adult
SW Georgia, Adzharia 10 km Baku 2000m, 18.6.98 25 mm
Copyright © Alexander Anichtchenko
Carabus (Sphodristocarabus) blumenthaliensis basilewskianus Breuning & Ruspoli, 1970
Male+Female adult
NE-Turkey, Artvin Prov., Hopa vers Borcka, Cankutaran Gecidi h=800m, 17.VI.2007 leg. D. Prunier. female, 24mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) varians janthinus s.str.
Male+Female adult
NW-Caucasus, Russia, Krasnodar reg.,Tuapse distr., Mt.Semashkho 04.V.2008, D.Fominyh leg.<br /> female, 24mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) varians armeniacus Mannerheim, 1830
Female adult
NE Turkey,<br /> Artvin Prov., Borçka, Macahel Gecidi h=1450m, 18.VI.2007, D.Prunier leg.<br /> female, 26mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) tokatensis almushensis Mandl, 1975
Male adult
Turkey, Sivas, 20 km N Zara, N Arapca, 1750m; 11.VII.1993
Copyright © Alexander Anichtchenko
Carabus (Sphodristocarabus) scovitzi czipkai Heinz, 1980
Male adult
Turkey, Artvin, Kafkasor, 1600 m
4.7.1996 Staven and Skoupy leg.
male, 22 mm
Copyright © Alexander Anichtchenko
Carabus (Sphodristocarabus) macrogonus pseudosimonianus Mandl, 1975
Female adult
Turkey, Ordu, Golkoy, 1220 m, 19.7.91
Copyright © Alexander Anichtchenko
Carabus (Sphodristocarabus) macrogonus resplendens Breuning, 1934
Female adult
Turkey; Cayirkent Samsun 12.VII.1996
Copyright © Alexander Anichtchenko
Carabus (Sphodristocarabus) bohemani bohemani Menetries, 1832
Female adult
S Azerbaydzan, Lenkoran distr., Vazha riv.,
2 km NW Dashdatuk vill. 38º40*09N 48º42*06E
17.4.2004 Makarov K. leg.
female, 22 mm
Copyright © Alexander Anichtchenko
Carabus (Sphodristocarabus) bohemani bohemani Menetries, 1832
Male adult
Azerbaijan,
Talysh Mts., Polikesh, h~1800m.,2-6.VI.2001 Dolin V.G. leg.
male, 20mm.
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) hollbergi Mannerheim, 1827
Male adult
N Azerbaijan,
4km W Ismayilli, h=700m, 9.VII.2008, Pljushch I.G. leg.
male,22mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) adamsi Adams, 1817 (as chiragricus)
Male adult
Russia, Stavropol reg., Kislovodsk vic., 1,5 км ESE Podkumok 905 м, N43*57E42*47, 13-18.05.2006, A.Vlasenko leg.
det. Obydov
Copyright © Dmitri Obydov
Carabus (Sphodristocarabus) varians janthinus (as dvorshaki)
Male adult
Abkhasia, Gagra, 12.VI.1986<br /> photo from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Sphodristocarabus) adamsi Adams, 1817 (as chiragricus Fischer, 1828)
Male+Female adult
Russia, N. Caucasus, Stavropol reg., Zheleznovodsk, Zheleznaya Mt., h=600 m. 5-11.VI.2011, Е. Shankhiza leg. male - 24 mm, female - 25 mm.
Copyright ©
Carabus (Sphodristocarabus) varians varians Fischer, 1827
Male+Female adult
N. Armenia, Idzhevan distr., Acharkut vill. env., h=800 m.<br /> 26-28.VII.2008 S. Andreev leg.<br /> male - 22 mm, female - 23 mm.
Copyright ©
Carabus (Sphodristocarabus) varians janthinus s.str. (as dvorshaki Mandl, 1975)
Male+Female adult
W. Caucasus, Abkhazia, Gudauta distr..,<br /> Duripsh vill. env., Khipsta riv., h=400-500 м.<br /> 03-13.VI.2009 A. Safronov leg.<br /> body lenght 20-25 мм.
Copyright ©
Carabus (Sphodristocarabus) macrogonus macrogonus Chaudoir, 1847
Male adult
NE-Turkey:Trabzon env., Sumela, 17-21.VI.2010, M.Janalik leg.
L=20 mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) macrogonus pseudokolatensis Heinz & Korge, 1967
Male adult
NE-Turkey::Kackar Dagi Mt, Ayder S of Ardesen h=1600m, 17-21.VI.2010, M.Janalik leg.
L=20 mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) varians armeniacus Mannerheim, 1830
Male adult
NE Turkey, Artvin Prov., Şavşat, Çamlıca,18.VI.1993, M. Janata leg.<br /> L=22 mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) separatus persa Lapouge, 1924
Male adult
N-Iran: Mazandaran Province, Elborz Mt.Rng, Vandarbon env., 10.VII.2002 L=23 mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) blumenthaliensis aequicostalis Staven & Heinz, 1995
Male adult
NE-Turkey:NE of Artvin:Dalis Dağ ,h=1600-1800m, 8-18.VII.2014, C. Reuter leg. L= 25 mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) adamsi Adams, 1817
Male adult
CN-Georgia, N of Tbilisi, NW of Tianeti, Tvalivi env., 20.VI.2015, F. Grycz leg. L=24 mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) separatus bararii Deuve, 2000
Male adult
N-Iran: Mazendaran: Ramsar env., deciduous forest, h~305 m, 16-24.VI.2017, Václav Čutka leg. L=22 mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) separatus bararii Deuve, 2000
Female adult
N-Iran: Mazendaran: Ramsar env., deciduous forest, h~305 m, 16-24.VI.2017, Václav Čutka leg. L=23,5 mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) separatus separatus Gehin, 1885
Male adult
N-Iran:Gilan Prov., Asalem - Khalkal road, 25-30.V.2018, Václav Čutka leg. L= 21 mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) separatus separatus Gehin, 1885
Female adult
N-Iran: Gilan Prov., Asalem - Khalkal road, 25-30.V.2018, Václav Čutka leg. L= 23 mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) separatus separatus Gehin, 1885
Male adult
N-Iran:Gilan Prov., Asalem - Khalkal road, 25-30.V.2018, Václav Čutka leg. L= 22 mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) separatus haerii Heinz, 1973
Female adult
N-Iran:Gilan Prov., Rudbar, Harzevil env.,23-24.V.2018, Václav Čutka leg. L=24 mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) separatus haerii Heinz, 1973
Male adult
N-Iran: Gilan Prov., E of Rudbar, Barehsar env., 19-23.VI.2017, Václav Čutka leg. L=22 mm
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) biroi biroi Menetries, 1832
Male+Female adult
Amasya (Samsun), TURQUIE VI-1987
Copyright © Alain Mollard
Carabus (Sphodristocarabus) bohemani esfandiarii Heinz, 1973
Female adult
Hasan-Kif (Mazandaran) IRAN VII-2000
Copyright © Alain Mollard
Carabus (Sphodristocarabus) bohemani esperanzae Heinz, 1970
Male+Female adult
Rasht, (Masuleh) IRAN VI-2000
Copyright © Alain Mollard
Carabus (Sphodristocarabus) elegantulus Motschulsky, 1850 (as astrabadensis Semenov, 1903)
Male+Female adult
Ft.Pashmaky, (Golestan) IRAN VI-2000
Copyright © Alain Mollard
Carabus (Sphodristocarabus) giachinoi Cavazzuti, 1991
Female adult
Ulukoy, (Gumushane) TURQUIE VI-1991
Copyright © Alain Mollard
Carabus (Sphodristocarabus) gilnickii brunneicupreus Deuve, 1994
Male+Female adult
Camlik, (Rize), TURQUIE VI-1997
Copyright © Alain Mollard
Carabus (Sphodristocarabus) gilnickii hasandaghensis Darge & Basquin, 1990
Male+Female adult
Ispir (Erzurum), TURQUIE VI-1997
Copyright © Alain Mollard
Carabus (Sphodristocarabus) heinzi heinzi Breuning, 1964
Male+Female adult
Egribel (Gerisum), TURQUIE VI-1992
Copyright © Alain Mollard
Carabus (Sphodristocarabus) heinzi mateuellus Deuve, 2003
Male adult
Col Harçebeli,(Ordu) TURQUIE V-2001
Copyright © Alain Mollard
Carabus (Sphodristocarabus) kurdicus kurdicus Heinz, 1975
Female adult
Resadiye,(Bitlis) TURQUIE V-1991
Copyright © Alain Mollard
Carabus (Sphodristocarabus) macrogonus akkusensis Breuning & Ruspoli, 1970 Paratype
Male+Female adult
Akkus,(Ordu) TURQUIE X-1962
Copyright © Alain Mollard
Carabus (Sphodristocarabus) macrogonus diogenei Cavazzuti, 1991
Female adult
Gokçebel,(Sinop) TURQUIE VI-1988
Copyright © Alain Mollard
Carabus (Sphodristocarabus) macrogonus keltepensis Vigna-Taglianti & Bruschi, 1976
Female adult
Mt. Keltepe (Izmit) TURQUIE VI-2005
Copyright © Alain Mollard
Carabus (Sphodristocarabus) macrogonus drannadaghensis Breuning, 1964
Male+Female adult
Drannaz Dag,(Sinop) TURQUIE VII-1993
Copyright © Alain Mollard
Carabus (Sphodristocarabus) macrogonus schweigerinae Schweiger, 1962
Male+Female adult
Lac d’Abant, (Bolu) TURQUIE VIII-1993
Copyright © Alain Mollard
Carabus (Sphodristocarabus) tokatensis tokatensis Roeschke, 1898
Male+Female adult
Yilidiz Dag, (Tokat) TURQUIE VI-1985
Copyright © Alain Mollard
Carabus (Sphodristocarabus) tokatensis simonae Cavazzuti, 1984
Male+Female adult
Pulumur, (Tunceli) TURQUIE VI-1988
Copyright © Alain Mollard
Carabus (Sphodristocarabus) tokatensis kosensis Cavazzuti, 1984
Female adult
Kosedagi-Geç (Gumushane) TURQUIE VII-2000
Copyright © Alain Mollard
Carabus (Sphodristocarabus) scovitzi hedini Heinz, 1973
Male+Female adult
Borçka (Artvin) TURQUIE V-2005
Copyright © Alain Mollard
Carabus (Sphodristocarabus) scovitzi sarigoelensis Deuve & Charet, 1992
Male+Female adult
Kaçkar (Artvin) TURQUIE VII-1992
Copyright © Alain Mollard
Carabus (Sphodristocarabus) adamsi adamsi (as adamsianus Mandl, 1975)
Male adult
Dagestan. det & Photo I. Belousov
Copyright © Creative Commons
Carabus (Sphodristocarabus) adamsi adamsi (as adamsianus Mandl, 1975)
Female adult
Dagestan. det & Photo I. Belousov
Copyright © Creative Commons
Carabus (Sphodristocarabus) adamsi hollbergi Mannerheim, 1827
Male adult
det & Photo I. Belousov
Copyright © Creative Commons
Carabus (Sphodristocarabus) enigmaticus enigmaticus Heinz, 1980
Male+Female adult
Ispir (Erzurum) TURQUIE VI-1991
Copyright © Alain Mollard
Carabus (Sphodristocarabus) enigmaticus citamgineus Lassalle & Charet, 1992
Male+Female adult
Gole (Erzurum) TURQUIE VII-1991
Copyright © Alain Mollard
Carabus (Sphodristocarabus) varians janthinus Ganglbauer, 1887
Female adult
Carabus (Sphrodistocarabus) janthinus dvorschaki Mandl, 1975 Abkhazia, Gudautsky district, S- Bzybsky Ridge, a watershed hill at the head of the river Khipsta, 43°17'56.63" N, 40°40'16.74" E, h= 1490 m, mature beech-maple forest with butterbur, moist soil, 05.07-17.08.2017, leg. I.A. Solodovnikov, S.V. Solodovnikova, E.V. Tatun
Copyright © Igor Solodovnikov
Carabus (Sphodristocarabus) scovitzi scovitzi Faldermann, 1835
Male adult
Georgia, Adzharia, Adzharo-Imeretinskiy Mt.R., near Tchidila pass, alpin zone, h=2350 m, 05.VI.2016 leg. D. Fominykh
Copyright © Igor Solodovnikov
Carabus (Sphodristocarabus) tokatensis kosensis Cavazzuti, 1984
Male adult
CE-Turkey: Gümüshane: Köse Dag(pass N of Köse), h = 1700 m, VII.2000, leg. St. Il. Remond (TOPOTYPUS)
Copyright © Igor Solodovnikov
Carabus (Sphodristocarabus) varians armeniacus Mannerheim, 1830
Male adult
C Georgia, Imereti, Caucasus, Likhi Mt Range, Rikoti Pass h=900-1000 m, 14-16.V.2017, E. Rutjan E. leg., L=22 mm
Copyright © Ruslan Panin -
Stephanocarabus taxon page
-
Tachypus taxon page (50 photos)
Carabus (Tachypus) cancellatus tibiscinus Csiki, 1905
Male+Female adult
female,28 mm<br /> SE Hungary, Csongrбd, Makу env., 1-8.VIII.2004, Krause R., Slovak, Pittner leg
Copyright © Ruslan Panin
Carabus (Tachypus) cristoforii Spence, 1823 (as nicolasi Reitter, 1888)
Male adult
S-France, Hautes-Pyrenees, Pic de Nere h=2200m,15.VII.1999,
male, 14mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus carinatus Charpentier, 1825
Male+Female adult
France, Riveres Charente, leg. Laforgue VI.1996<br /> female, 24mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus s.str. (as sulinensis Born, 1902)
Male+Female adult
S UKRAINE, <br /> Krym Peninsula, Staryi Krym, 17.V.2002<br /> L= 26 mm.
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus s.str. (as conspersus Lapouge, 1902)
Male+Female adult
E Byelorussia,
Mogilev Reg. Bykhov distr., near Voronino vill., 20.V-3.VI.2007, Komyagunova N. leg.
female, 26mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus s.str. (as tuberculatus Dejean, 1826)
Male adult
W Ukraine,NW Podolian,
Ternopil reg., Pidvolochisk distr., forest Bogdanivskiy, near Romanove Selo, 7-24.V.08, R.Yu.Panin & I.A.Solodovnikov leg.
male, 23mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus durus (as strictus Kolbe, 1913)
Female adult
Ukrainian Carpathians,<br /> Transcarpathian reg., Rakhov Distr., Chornohora range h=1850m, 29.VI-28.VII.2007 Panin R. leg.<br /> female, 27 mm.
Copyright © Ruslan Panin
Carabus (Tachypus) auratus auratus Linne, 1761
Male adult
Canada<br /> Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Carabus (Tachypus) auratus brunieri Barthe, 1921
Male adult
SC-FRANCE, Aude, Rennes les Bains, 6.VII.1989, Sunard A. leg., 26mm
Copyright © Ruslan Panin
Carabus (Tachypus) auratus auratus Linne, 1761
Female adult
30 mm. Germania, Oberfranken, Pegnitz, eg.det. U.Schmidt, 1972
Copyright © U.Schmidt
Carabus (Tachypus) auratus lasserrei Doue, 1855
Male adult
SC-FRANCE, Midi-Pyrуnуes reg. Tarn, Saint Paul cap de Joux, 24.V.2003, A. Laforgue leg., 24mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
Male adult
Italy-Switzerland, Riviera – Preonzo, 18.II.1998, C.Sola leg.
L=26 mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus graniger Palliardi, 1825
Male adult
SW-Romania, Banatului Montan, Resita - Valiug, 29.V-9.VI.2005. Krause R. leg.
L=25 mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
Male adult
relict lowland populations of blue-purple color, of the the river Ticino park - Italy - Lombardy
Copyright © Monzini
Carabus (Tachypus) auratus lotharingus Dejean, 1826
Male adult
SC France: Herault departament, Foret de Saint Gervais, V. 2009 leg. J. Nicolas<br /> L=27mm
Copyright © Ruslan Panin
Carabus (Tachypus) cristoforii Spence, 1823
Female adult
S France, Pyrénés Mts., Tarascon; 15mm
Copyright © Aleš Sedláček
Carabus (Tachypus) auratus honnoratii Dejean, 1826
Male adult
SE France, Carpentras; 23mm
Copyright © Aleš Sedláček
Carabus (Tachypus) cancellatus durus (as carpathicus Kolbe, 1913)
Male+Female adult
Ukrainian Carpathians, Ivano-Frankivsk reg., Gorgany Mts., 20km SW Nadvorna, 22.VI-3.VIII.20, Panin R. leg.<br /> female, 26mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus durus Reitter, 1896
Male+Female adult
W Ukraine, Transcarpathian reg., Vinogradov distr., near Malaya Kopanya vill.,7-18.VII.2008, Panin R. leg. <br /> female, 32mm.
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus durus Reitter, 1896
Female adult
W. UKRAINE, NE Carpathians, Marmarosh Mts h=450m, Rakhiv distr., near Dilove vill.,19-23.VII.2010, Panin R. leg.<br /> L=29 mm
Copyright © Ruslan Panin
Carabus (Tachypus) vagans Olivier, 1795
Female adult
SE-France, Var: Le Muy env., 30.VII.2008., A. Paulian leg.
L=24 mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus intermedius Dejean, 1826
Female adult
NW -Serbia,Fruška Gora,Vrdnik env. h=380m, 9-10.VII.2013, M. Janalik leg. L=29 mm
Copyright © Ruslan Panin
Carabus (Tachypus) auratus honnoratii Dejean, 1826
Male+Female adult
France : Vaucluse, Mt.Ventoux-Mt. Serein, vii.1969, N. Mal Leg., Coll.RBINS-N. Mal incl. ventouxensis and Nicolasi
Copyright © Noël Mal
Carabus (Tachypus) auratus honnoratii Dejean, 1826
Male+Female adult
France : Vaucluse, Mt. Ventoux-Mt. Serein, 1400-1800m, vii.1969-N. Mal Leg.-Coll.RBINS, N. Mal - including f.i. ventouxensis and nicolasi
Copyright © Noël Mal
Carabus (Tachypus) cancellatus cancellatus Illiger, 1798
Male adult
Carabus (Tachypus) cancellatus tuberculatus f.conspersus Lapouge, 1902 (Blaс form) ugly specimens N-Belarus, Vitebsk reg., Senno distr., 37 km SSE Vitebsk, near Lake Streshno, 01-10.VII.2006, I.A. Solodovnikov
Copyright © Igor Solodovnikov
Carabus (Tachypus) cancellatus cancellatus Illiger, 1798
Male adult
Carabus (Tachypus) cancellatus tuberculatus f.conspersus Lapouge, 1902 (Blaс form) E-Belarus, Mogilev reg., Bykhov distr., 5 km N Voronino Vill., mixed-pine forest, 20.V-03.VI.2007, Komyagunova N.P.
Copyright © Igor Solodovnikov
Carabus (Tachypus) cancellatus cancellatus Illiger, 1798
Male adult
Carabus (Tachypus) cancellatus tuberculatus f. tenebrosus Ball. (Blue form) S-Belarus, Brest reg., env. Tomashovka, pine forest, 26.VI-13.VII.2022, Kuznetzov V.A.
Copyright © Igor Solodovnikov
Carabus (Tachypus) cancellatus cancellatus Illiger, 1798
Female adult
Carabus (Tachypus) cancellatus tuberculatus f.conspersus Lapouge, 1902 (Blaс form) E-Belarus, Mogilev reg., Bykhov distr., 5 km N Voronino Vill., mixed-pine forest, 20.V-03.VI.2007, Komyagunova N.P.
Copyright © Igor Solodovnikov
Carabus (Tachypus) cancellatus cancellatus Illiger, 1798
Female adult
Carabus (Tachypus) cancellatus tuberculatus f. tenebrosus Ball. (Blue form) S-Belarus, Brest reg., env. Tomashovka, pine forest, 26.VI-13.VII.2022, Kuznetzov V.A.
Copyright © Igor Solodovnikov
Carabus (Tachypus) cancellatus cancellatus Illiger, 1798
Female adult
Carabus (Tachypus) cancellatus tuberculatus f. tenebrosus Ball. (Blue form) S-Belarus, Brest reg., env. Tomashovka, pine forest, 26.VI-13.VII.2022, Kuznetzov V.A.
Copyright © Igor Solodovnikov
Carabus (Tachypus) cancellatus cancellatus Illiger, 1798
Male adult
Carabus (s.str.) cancellatus cancellatus f.balticus Bernau, 1915 N-Belarus, Vitebsk reg. 21 mm
Copyright © Igor Solodovnikov
Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
Male adult
Carabus (s.str.) cancellatus emarginatus n.monzinii Monzini, 1983 25,5 mm
Copyright © Igor Solodovnikov
Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
Male adult
Carabus (Tachypus) cancellatus emarginatus n. dahli Heer, 1841
Copyright © Igor Solodovnikov
Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
Female adult
Carabus (Tachypus) cancellatus emarginatus n. dahli Heer, 1841
Copyright © Igor Solodovnikov
Carabus (Tachypus) cancellatus durus (as strictus Kolbe, 1913)
Male adult
W Ukraine, NE Carpathians, Maramaros Mts (Hutsul Alps) Gorge of Kuziy (Kizi) stream, h = 400m, 20-23.V.2021, Panin R. leg., L=30 mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus durus (as strictus Kolbe, 1913)
Female adult
W Ukraine, NE Carpathians, Maramaros Mts (Hutsul Alps), Gorge of Kuziy (Kizi) stream, h = 400m, 20-23.V.2021, Panin R. leg., L=32 mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus durus Reitter, 1896
Female adult
W Ukraine, Transcarpathian, Velykyi Bereznyi distr., Stuzhytsya (Stužica), 2-29.VI.2011, Yu. Kanarsky leg. L=33.5 mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus carinatus (misellus Lapouge, 1898)
Male adult
SC-France, Massif central, Mt.Cantal, Plateau du Limon h=1300m, VIII.2013, Pascal Stefani leg., L=23 mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus excisus Dejean, 1826
Female adult
W-Czech, Plzeň Reg, Újezd nade Mží vill.,10.VI.2014, L=25.5 mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus excisus Dejean, 1826
Male adult
W Austria, Tyrol, Vomp, forest path to Sigmundslust Castle, 18.V-3.VI.2020, M. Egger leg., L=23 mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus graniger (as biharicus Reitter, 1896)
Female adult
CW-Romania, Arad distr., Săvârşin - Ilteu, 24-30.V.2017, V. Šima leg., l=30 mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus graniger (as szobroniensis Gehin, 1885)
Female adult
SE-Romania, Bucharest, Padurea (Forest) Cernica, h=150 m, VI.1989, Stela Lie leg., L=29 mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus intermedius (as maximus Haury, 1880)
Male adult
SW Hungary, Bátaapáti, Com. Baranya, 6-9.VI.2003, Gy. Szel leg/det. I.Retezar 2010, L=25 mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus cancellatus Illiger, 1798
Male adult
W-Ukraine, Podolian Upland (Lviv Opillia) W of Vybranivka vill., under the bark, 8.II.2008, R.Panin leg., L=22 mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus durus (as carpathicus Kolbe, 1913)
Male adult
W-UKRAINE, Ivano-Frankivsk reg., Gorgany Mts, Mt.Synyak h =1530м., 25.V-17.VII.2014, Panin R. leg., L=28 mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus cancellatus (as charcoviensis Bernau, 1915)
Female adult
NE-Ukraine, Kharkov reg., Zmiev distr., 1-7.VI.2007, Kovalchuk I. leg., L=27 mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus durus Reitter, 1896
Female adult
W-Ukraine, Transcarpathian Lowland, Mukachevo distr., near Znayatsevo vill., 12.VI-8.VII.2009, R. Panin leg., L=30 mm
Copyright © Ruslan Panin
Carabus (Tachypus) cancellatus carinatus (as subcarinatus Kolbe, 1912)
Male adult
E-Netherlands, Gelderland prov., Nijmegen., Berg en Dal vill., 6.XII.2014, H.de Haan leg., L=25 mm
Copyright © Ruslan Panin -
Tanaocarabus taxon page (14 photos)
Carabus (Tanaocarabus) sylvosus Say, 1825
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Carabus (Tanaocarabus) hendrichsi Bolivar y Pieltain, Rotger & Coronado, 1967
Male adult
male, 23mm
Copyright © Boleslav Brezina
Carabus (Tanaocarabus) forreri Bates, 1882
Male adult
Mexico, Durango, Sierra Madre Occ.,
Buenos Aires w., 2700 m, La Ciudad, 14.6.90 Heinz leg.
Copyright © Alexander Anichtchenko
Carabus (Tanaocarabus) forreri forreri Bates, 1882
Female adult
female, 23mm
Copyright © Boleslav Brezina
Carabus (Oreocarabus) taedatus agassi Le Conte, 1850
Female adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Carabus (Oreocarabus) taedatus agassi Le Conte, 1850
Male adult
USA, Arizona, Coconino Co., Humphreys Peak,
bei Flagstaff 14.6.1990 Heinz leg.
Copyright © Alexander Anichtchenko
Carabus (Tanaocarabus) sylvosus Say, 1825
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -
Carabus (Oreocarabus) taedatus agassi Le Conte, 1850
Male adult
SC-USA, New Mexico, Santa Fe,VIII.1999, Sehnal R. leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Oreocarabus) taedatus bicanaliceps Casey, 1920
Male adult
NW-USA, Washington, Thurston County, Mima Mound Prairie, 15.IX.2005, A. Laforgue leg. L=23 мм
Copyright © Ruslan Panin
Carabus (Tanaocarabus) sylvosus Say, 1825 Lectotype
Male adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC63 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec63
Copyright © - - -
Carabus (Tanaocarabus) caseyi Angell, 1914 Holotype
Female adult
Texas (Smithsonian Institute, Washington)
Copyright © Terry Erwin
Carabus (Tanaocarabus) forreri chiricahuaensis Lassalle, 2016 Holotype
Male adult
S-USA: Arizona: Chiricahua Mts.: Rustler Camp, 31:54.828N/109:16.185W/ 2514m, Van Den Berghe leg. (Carnegie Mus. Pittsburg)
Copyright © Bernard Lassalle -
Teratocarabus taxon page (2 photos)
Carabus (Teratocarabus) azrael azrael Semenov & Znojko, 1932
Female adult
Copyright © Alexander Anichtchenko
Carabus (Teratocarabus) azrael mizunumaianus Imura, 1991
Male adult
China, Liaoning, Benxi, Gongchangung
20.VI.2004 Hong
Copyright © Boleslav Brezina -
Tomocarabus taxon page (30 photos)
Carabus (Tomocarabus) scabripennis scabripennis Chaudoir, 1850
Male adult
Carabus (Tomocarabus) scabripennis scabripennis Chaudoir, 1850
C. Armenia,
Kotayk reg., Yeghvard, Arailer Mt., 16.VI.1999, P.V.Kazaryan leg.
male, 20mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Tomocarabus) scabripennis marthaiellus Rapuzzi & Kleinfeld, 2000 Holotype Paratype
Male+Female adult
<b>Carabus (Tomocarabus) scabripennis marthaiellus Rapuzzi & Kleinfeld, 2000</b>
Copyright © Frank Kleinfeld
Carabus (Tomocarabus) marginalis Fabricius, 1794
Male+Female adult
E Ukraine,
Kharkov reg.,near Vasishchevo vill.10-20.VI.2006 D.Kovalchuk leg.
female 24mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) decolor Fischer, 1823
Female adult
Carabus (Tomocarabus) decolor Fischer, 1823
Copyright © Alexander Anichtchenko
Carabus (Tomocarabus) decolor Fischer, 1823
Male adult
S Russia, NW-Caucasus,
Karacaj-Cherkessija,Skalisty Mnt. Range, Site Banditskoe h=1400m, 2-3.V.2009,
Fominykh D., Bondarenko A.S. leg.
male,17mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) convexus s.str. (as pyrenaicus Born, 1907)
Male adult
Pirineos, Canfranc, 1300m, Barranco de Izas
15.6.2005 Anichtchenko A. leg.
male, 16 mm
Copyright © Alexander Anichtchenko
Carabus (Tomocarabus) convexus s.str. (as paganetti Born, 1905)
Male adult
Lazio, Roma, Bosco di Nettuno
22.5.1986 C. Iacovone leg.
male, 20 mm
Copyright © Alexander Anichtchenko
Carabus (Tomocarabus) convexus convexus Fabricius, 1775
Male adult
W.Ukraine,
Transcarpathian region,Vinogradov distr., near Malaya Kopanya vill., Chorna Gora Mt., h=180m. 7-18.VII.08. Panin R. leg.
male 18 mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) convexus s.str. (ab. niger)
Male adult
W Ukraine, NW Podolia, Lviv Region, Zolochiv distr., near Gologirky vill., in steppe 1-19.V.2007 Panin R. leg.
16mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) convexus erzeliki Schweiger, 1962
Male adult
Turkey, Sivas 1560 m. 19.7.1991
male, 19 mm
Copyright © Alexander Anichtchenko
Carabus (Tomocarabus) bessarabicus bessarabicus Fischer, 1823
Male+Female adult
UKRAINE,
Kherson Reg., Golaya Pristan distr. near Rybalche vill. 6.IV.2000
long (female) 20mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) convexus gracilior Gehin, 1885
Male adult
SE-Bulgaria, Burgas Prov., Primorsko env., 18-30.VI.2000, Krause Rene leg.
L=18 mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) bessarabicus ? concretus Fischer, 1823
Male adult
E Ukraine, Lugannsk Reg., Sverdlovsk distr., Steppe Reserve “Provalsky", 3.V-3.VI.2011, leg.Glotov S./det.Panin R.<br /> female, 22 mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) convexus moreanus Reitter, 1896
Female adult
Greece:, N Peloponnes, Xylokastro, Zemeno,h=1000m, 6.VI.1997, Manfred Egger leg. L=21 mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) convexus dilatatus (as perplexus Schaum, 1857)
Male adult
CE-Greece: Larissa reg., Stomio, Ossa Mt., 6-17.VII.2011, Krause Rene leg.
L=19 mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) rumelicus syriensis Breuning, 1943
Male adult
N Lebanon, Fnaпdek, Djebel Qammoua, h=1100-1200m, C. Reuter leg.
L=17.5 mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) convexus moreanus Reitter, 1896
Male adult
S Greece, Taygetos Mts.; 13mm
Copyright © Aleš Sedláček
Carabus (Tomocarabus) convexus dilatatus Dejean, 1826
Male adult
NE Serbia, Majdanpek; 13mm
Copyright © Aleš Sedláček
Carabus (Tomocarabus) convexus gracilior Gehin, 1885
Male adult
SW Romania, Banat; 14mm
Copyright © Aleš Sedláček
Carabus (Tomocarabus) convexus gracilior Gehin, 1885
Female adult
SW Romania, Banat; 15mm
Copyright © Aleš Sedláček
Carabus (Tomocarabus) convexus erzeliki Schweiger, 1962
Female adult
NC-Turkey, Sinop province, Erfelek env., h=200m, 10-27.VII.2014, C. Reuter leg. L=20 mm
Copyright © Ruslan Panin
Carabus (Tomocarabus) convexus rhinopterus Hampe, 1852 (as salax Motschulsky, 1865)
Male adult
Turkey, Sivas, 20 km N Zara, 1750 m 10 km S Serefiye 17.8.1992 K. Staven leg
Copyright © Alexander Anichtchenko
Carabus (Tomocarabus) rumelicus syriensis Breuning, 1943 (as libanoticus)
Male adult
Lebanon
Copyright © Mingyi Tian
Carabus (Tomocarabus) rumelicus rumelicus Chaudoir, 1867
Male adult
Turkey, Yavsan, K. Maras, 7.7.2004 (cAA)
Copyright © Alexander Anichtchenko
Carabus (Tomocarabus) marginalis Fabricius, 1794
Male adult
N Ukraine, Kiev reg., Vyshgorod disrt., near Khotyanivka vill., 1.VII-7.IX..2025, E. Rutjan leg.l=22 mm
Copyright © Ruslan Panin -
Trachycarabus taxon page (9 photos)
Carabus (Trachycarabus) estreicheri Fischer, 1822
Male+Female adult
W UKRAINE, NW Podolian,
Lviv Region, Brody distr., near Yasenov vill., steppe sites, 2-28.V.2010, Panin R. leg.
female, 23mm
Copyright © Ruslan Panin
Carabus (Trachycarabus) estreicheri Fischer, 1822
Female adult
W-UKRAINE, NW Podolian,
Lviv Region, Brody distr., near Yasenov vill., steppe sites, 12.V-2.VI.2007 Panin R. leg.
female, 22mm
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus tscherkessicus Korge, 1964
Male adult
S Russia, NW-Caucasus,<br /> Karachay-Cherkessia,Teberda, Mukhinsky (Mukhu) Mt. pass, h=2750-2800 m , 8.VIII.2010, I.A. Solodovnikov leg.<br /> male, 23 mm<br />
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886
Male+Female adult
W Ukraine,<br /> Ternopol reg., Pidvolochisk distr., Supranivka vill.<br /> 3.VII.1908. Leg. Stobetskii. S.<br /> Photo R. Panin
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886
Male adult
W Ukraine,
Ternopil reg., Pidvolohcisk distr., Supranivka vill.
male, 26mm
The end of XIX the beginning of XX century
photo Panin R.
coll. SMNH Ukraine, Lviv
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus obliteratus Fischer, 1827
Male adult
Altai mts., Kalbinskii mt.rng.
Ognevka vill. VII.1994, Klimenko A. leg.
male, 28 mm
Copyright © Alexander Anichtchenko
Carabus (Trachycarabus) sibiricus obliteratus Fischer, 1827
Male adult
NE Kazakhstan,
S Altai Mts., Ulbinsky range h~500m.,Turgusun vill., 20.VI.2001
male 25mm
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus kolosovi Zinoviev, 1997 loc.typ.
Female adult
S. Ural, Bashkiria, near Mesjagutovo vill., 09.07.2001 A. Menstschicov leg.
photo A. Menstschicov
Copyright © Anton Menstschikov
Carabus (Trachycarabus) sibiricus kolosovi Zinoviev, 1997
Female adult
Russia, North-East Bashkortostan. Ai river, near Sikiyaz-Tamak vill.
12.07.1997, Zinoviev E.leg
length: 21,8 mm
Copyright © Evgeniy Zinoviev
Carabus (Trachycarabus) sibiricus haeres (as fossulatus Dejean, 1826)
Female adult
<b>Carabus (Trachycarabus) sibiricus fossulatus Dejean, 1826</b><br /> NE Ukraine,<br /> Kharkov reg.,1974<br /> female, 27mm.
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus haeres (as fossulatus Dejean, 1826)
Male+Female adult
E.UKRAINE,<br /> Kharkov reg., Zmiev distr.,in steppe,10-20.V I.2007, D.Kovalchuk leg.<br /> female, 26mm/male, 24mm.<br /> photo/det. Panin R.
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus errans Fischer, 1823
Female adult
Carabus (Trachycarabus) sibiricus errans Fischer, 1823
S.Russia, Rostov-Don reg., Avilo-Uspenskaya env.
V.2000
Copyright © Safronov
Carabus (Trachycarabus) sibiricus errans Fischer, 1823
Male adult
SW-Russia,
Rostov reg. env. stanitsa Razdorskaya, A. Ponomarev leg. 9-15.V.2001
male 25mm
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus errans Fischer, 1823
Male+Female adult
female:SE Ukraine,
Donetsk, 25.VII.2009, A.I.Gubin leg. L=30mm.
male:E Ukraine, Lugansk reg., near Slavyanoserbk, 5-15.VIII.2009, S.Konovalov leg. L=26mm.
photo/det.Panin R.
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus bosphoranus Fischer, 1823
Female adult
Carabus (Trachycarabus) sibiricus bosphoranus Fischer, 1823
S Ukraine,
Krym Peninsula, Nikitskaya Jaila, S.Mosyanin leg. 10.VIII.2000
female 23mm.
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus bosphoranus Fischer, 1823
Male adult
Carabus (Trachycarabus) sibiricus bosphoranus Fisch. 1823<br /> S Ukraine, Krym Peninsula,<br /> Dolgorukovskaya jaila, 25.VIII.1991 I.A.Solodovnikov leg.<br /> male, 23mm<br />
Copyright © Ruslan Panin
Carabus (Trachycarabus) scabriusculus scabriusculus (as erythropus Fischer, 1822)
Male+Female adult
W.Ukraine,<br /> NW Podolian Upland , Lviv Reg., Zolochiv distr., near Luky vill., steppe sites, 7.V-8.VI. 2010<br /> leg. Panin R.<br /> female, 22mm
Copyright © Ruslan Panin
Carabus (Trachycarabus) scabriusculus scabriusculus (as erythropus Fischer, 1822)
Male adult
W-UKRAINE<br /> NW Podolian Upland, Lviv Reg., Zolochiv Distr., near Luka vill., in steppe. Panin R. leg. 1.V - 16.VI.07
Copyright © Safronov
Carabus (Trachycarabus) scabriusculus inapertus Motschulsky, 1850 (loc.typ)
Male+Female adult
NE UKAINE,
Kharkov reg., Zmiev distr., in steppe, 15.VI-14.VII.2009, Kovalchuk D. leg.
female, 20mm
photo/det. Panin R.
Copyright © Ruslan Panin
Carabus (Trachycarabus) planus planus Gehin, 1885
Male+Female adult
S Russia, NW Caucasus,<br /> Karacaj-Cherkessija, near Kart-Djurt vill., upper Kylan riv. h=1300m, 26-27.V.2009 , Fominykh D. leg.<br /> L=24mm<br /> photo/det. Panin R.
Copyright © Ruslan Panin
Carabus (Trachycarabus) planus planus Gehin, 1885
Male adult
NW Caucasus, Rostov reg.,<br /> Avilo-Uspenskaya vill. 22.5.2000 Arzanov Yu. leg.<br /> male, 25 mm
Copyright © Alexander Anichtchenko
Carabus (Trachycarabus) planus planus Gehin, 1885
Male adult
S Russia,<br /> Krasnodar reg., Anapa distr., B. Utrish vill., env., 11-20.VIII.2008, Khatchikov E. leg.<br /> male, 26mm<br /> photo Panin R.
Copyright © Ruslan Panin
Carabus (Trachycarabus) planus campicola Reitter, 1896
Male adult
Krym Peninsula, near Sudak. Leg.Vashchenko S. 1-5.V.2004<br /> 27 mm
Copyright © Ruslan Panin
Carabus (Trachycarabus) planus campicola Reitter, 1896
Male+Female adult
S Ukraine, Krym Peninsula,<br /> near Perevalnoe vill., in steppe, 15.IV.2009, Shaporinskiy V. leg.<br /> female, 28 mm<br /> photo/det.Panin R.
Copyright © Ruslan Panin
Carabus (Trachycarabus) mandibularis mandibularis Fischer, 1827
Male adult
NE Kazakhstan, S Altai Mt., Narymskii mt. rng.
Bolshenarymskoe vill. 26.7.2001, 1800m Klimenko leg.
male, 21 mm
Copyright © Alexander Anichtchenko
Carabus (Trachycarabus) mandibularis buchtarmensis Kryzhanovskij, 1953
Male adult
Kazakhstan, Kalbinsky mt. rng., 600m ,
Ognevka vill., 23.6.98 Klimenko A. leg.
male, 23 mm
Copyright © Alexander Anichtchenko
Carabus (Trachycarabus) mandibularis buchtarmensis Kryzhanovskij, 1953
Male adult
NE-Kazakhstan,
S-Altays, Kalbinky Mt. h=500m., near Ognevka vill., 20.VI.1998.A. Klimenko leg.
male, 23mm
Copyright © Ruslan Panin
Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
Male adult
Burjatia, Dzhida distr., near Beloozersk vill. 4-9.VII.1997 Anichtchenko A. leg.
male, 23 mm
Copyright © Alexander Anichtchenko
Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
Male adult
Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
China, Beijing, Mentougou, 2005;
Photo & det. by liuye
Copyright © Ye Liu
Carabus (Trachycarabus) besseri Fischer, 1822
Male+Female adult
W Ukraine.NW Podolian,
Lviv reg., Zolochiv distr., Lysa Gora near Stinka vill., steppe sites, 25.IV-9.V.2009 Panin R. leg
female, 30mm
Copyright © Ruslan Panin
Carabus (Trachycarabus) estreicheri Fischer, 1822
Female adult
Volgograd Reg., Serafimovka Distr., Ust' -Khoperskaya Station 5.V.2009 leg.O.Sveshnikov
18.5 mm
Copyright © Kirill Makarov
Carabus (Trachycarabus) sibiricus (as gvozdevae Obydov, 1997)
Male adult
Russia, Khakassia, <br /> W Sayan, Enisey riv., Batenevskiy Mt Range, 12 km E Pervomayskiy vill., 11.VII.2000, A.& N. Menstschicovy leg.<br /> L=21 mm
Copyright © Ruslan Panin
Carabus (Trachycarabus) estreicheri Fischer, 1822 (as jaegeri Ménétriés, 1827)
Female adult
E UKAINE, Lugansk city, Site Luganskoe, 20-30.VII.2010, V.A.Landik leg.
L=21 mm
det.Panin.R.
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus sibiricus Fischer, 1822
Male adult
E Siberia, Russia, Krasnoyarsk Reg., near Krasnoyarsk, on dry steppe slopes, 7.VIII,2011, M.Ivanov leg.
L=21 mm
Copyright © Ruslan Panin
Carabus (Trachycarabus) latreillei ? semicoriaceus Kraatz, 1881
Male adult
NE-China: Liaodong Peninsula, Liaoning prov., Gaizhou distr., Mt. Buyun Shan,VI.2007, Hao leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
Male adult
N Mongolia, Khövsgöl reg., Lake Khovsgol, h=1640m, 51*:12’N; 100*:45’E, steppificated sites,
20-25.VII.2010. A.Dergalev leg.
L=22 mm
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus haeres Fischer, 1823
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Carabus (Trachycarabus) estreicheri (ab.rufofemoratus Lomnicki, 1892)
Female adult
W Ukraine, NW Podolian, Lviv reg., Brody distr., Makitra Mt, steppe site, 15.VI - 24.IX.2017, Panin R. & Kanarsky Yu. leg.
Copyright © Ruslan Panin
Carabus (Trachycarabus) mandibularis abakkereiorum Kabak & Hu, 2017 Holotype Paratype
Male+Female adult
(ZIN). China, Xinjiang, Altay Mt. R., Kran Valley, N of Altay Town, 47°55´37''N / 88°08´23´´E, h = 1015 m, 31.VII.2015 I.I. Kabak leg.
Copyright © Ilya Kabak
Carabus (Trachycarabus) sibiricus obliteratus Fischer, 1827 Paratype
Female adult
China, Xinjiang, “Prov. Xinjiang: Ake Tei A Zhu Pai 1750 m, 5.VIII.1960, leg. Wan Su Yung, China Naturwiss. Institut”; “Paratypus Carabus (Trachycarabus) ♀ sibiricus pseudobliteratus nov. ssp. det. Korell et Kleinfeld, 1982”. (MFNB)
Copyright © Ilya Kabak
Carabus (Trachycarabus) sibiricus haeres (trans s. errans)
Female adult
C Ukraine, S of Poltava city, edge forest-lane, 2-26.IX.2010, O. Pozharov leg. /det. R.Panin. L=30 mm
Copyright © Ruslan Panin
Carabus (Trachycarabus) scabriusculus lippii Dejean, 1826
Female adult
SW Ukraine, Odessa reg., Ivanovka distr., steppe ravine near Gudevichevo vill., 22.IV-5.V.2022, A.Gontarenko leg./det. R.Panin, L=22 mm
Copyright © Ruslan Panin
Carabus (Trachycarabus) scabriusculus lippii Dejean, 1826
Male adult
SW Ukraine, Odessa reg., Ivanovka distr., steppe ravine near Gudevichevo vill., 22.IV-5.V.2022, A.Gontarenko leg./det. R.Panin, L=21.5 mm
Copyright © Ruslan Panin
Carabus (Trachycarabus) planarius Obydov, 1994 Paratype
Female adult
Казахстан, Семипалатинский р-н, окр. пос. Шолпан, 500 м, 20 км. южнее пос. Таскескен, 2.06.1993, М. Данилевский leg.
Copyright © Dmitri Obydov
Carabus (Trachycarabus) sibiricus seriatoporus Fischer von Waldheim, 1828
Male adult
окр. Якутска, пос. Бестях на р. Лена, 30.VI.1962, Сладков leg.
Copyright © Dmitri Obydov
Carabus (Trachycarabus) planus retowskianus Mandl, 1955
Male adult
S Ukraine, Zaporizhia reg., Melitopol distr., near Troitskoe vill., 1.VI.2010, I.V. Kovalev leg., L=27 mm
Copyright © Ruslan Panin
Carabus (Trachycarabus) estreicheri Fischer, 1822 (ab.rufofemoratus Lomnicki, 1892)
Female adult
W Ukraina, Podolian Upland, Lviv reg., Brody distr., Mt. Makitra, steppe site, 19.V-5.VI.2020, Panin R. leg. L=23 mm
Copyright © Ruslan Panin
Carabus (Trachycarabus) planus campicola Dejean, 1831
Female adult
S Ukraine, Crimea, S slope Mt Demerdzhi h=650 m, sibljak, VII.2004, R. Panin leg., L=29 mm
Copyright © Ruslan Panin
Carabus (Morphocarabus) latreillei mengguicus Rapuzzi, 2023 Holotype
Male adult
China, Inner Mongolia Province, Tongliao, Ka’erqin, Jinbaotun, VIII.20. Image from Rapuzzi, 2023
Copyright © - - -
Carabus (Trachycarabus) scabriusculus inapertus Motschulsky, 1850
Male adult
NC Ukraine, Kiev city, Lysa Gora, 1-15.VI.2012, Gerasimov R.leg./det. R.Panin, L=19 mm
Copyright © Ruslan Panin
Carabus (Trachycarabus) latreillei shanxiensis Rapuzzi, 2024 Holotype
Male adult
China, Shanxi, Lu Liang, Kang Cheng Zhen, Jiao Kou Xian; Aug. 2022. Image from Rapuzzi, 2024
Copyright © - - - -
Tribax taxon page (145 photos)
Carabus (Tribax) titan titan Zolotarev, 1913
Male+Female adult
Russia,NW Caucasus,
Krasnodar reg.,Chernogorie Mnt. Range, Agulova Balka h=950m V-VII.1999, A.S.Zamotajlov leg.
female,32mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Tribax) titan titan Zolotarev, 1913
Male adult
Carabus (Tribax) titan titan Zolotarev, 1913
Tuapse distr., Anastasievka vill., Semiglavaya mt.,
Pshekaho riv., 20.7.1995
male, 30 mm
Copyright © Alexander Anichtchenko
Carabus (Tribax) titan abadzechus Zamotajlov, 1988
Male adult
Carabus (Tribax) titan abadzechus Zamotajlov, 1988
W Caucasus, Adygea, Asish-Tau Mt.R., vall riv. Bzykha, Fagus forest, h=1450m, 14-22.08.2008, I.A.Solodovnikov, O.Beljavskyi
male, 25 mm
Copyright © Igor Solodovnikov
Carabus (Tribax) steveni steveni Menetries, 1832
Male adult
Carabus (Tribax) steveni steveni Menetries, 1832
Karachaevo-Cherkessia,
Hudes riv., near Tsarskie vorota,
V.2009 Anichtchenko A. leg.
male, 25mm
photo Anichtchenko A.
Copyright © Alexander Anichtchenko
Carabus (Tribax) steveni steveni Menetries, 1832
Male adult
Carabus (Tribax) steveni steveni Menetries, 1832
C Caucasus, Kabardino-Balkaria,
Elbrusskyi distr., riv. Malka, upper reaches riv. Sultan-Bashi, h=2400-2600 m, 26, VII, 2008. Leg. I.A.Solodovnikov.
male, 20,5 mm
Copyright © Igor Solodovnikov
Carabus (Tribax) steveni steveni Menetries, 1832
Female adult
Carabus (Tribax) steveni steveni Menetries, 1832
C Caucasus, Kabardino-Balkaria,
Elbrusskyi distr., riv. Malka, upper reaches riv. Sultan-Bashi, h=2400-2600 m, 26, VII, 2008. Leg. I.A.Solodovnikov.
female 24,5 mm
Copyright © Igor Solodovnikov
Carabus (Tribax) steveni schamyli Hampe, 1852
Female adult
S Russia, N Ossetia, Alagir distr., near Nizhniy Tsey vill.,VII.1981
female, 25mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Tribax) steveni schamyli Hampe, 1852
Male adult
C Caucasus, N Ossetia,
N slope of Pastbistshnyi Mt.R., Tagardon env., h=1100m, 7-9.08.1997, leg. A.G.Koval
male 25,5 mm
Copyright © Igor Solodovnikov
Carabus (Tribax) steveni schamyli Hampe, 1852
Male adult
C Caucasus, N Ossetia,
N slope of Pastbistshnyi Mt.R., Tagardon env., h=1100m, 7-9.08.1997, leg. A.G.Koval
female 27 mm
Copyright © Igor Solodovnikov
Carabus (Tribax) steveni schamyli Hampe, 1852
Male adult
N Osetia, B. Balkary, 1700 m, 2.9.85
Abdurahmanov leg.
male, 24 mm
Copyright © Alexander Anichtchenko
Carabus (Tribax) steveni satanas Semenov, 1896
Male adult
Carabus (Tribax) steveni satanas Semenov, 1896
S Russia, C-Caucasus:
Balkaria, Valley of riv. Cherek Balkarskyi, h=1750-1800m 10.VI-20.VII.1992 Leg. Zamotajlov A.
male, 23 mm
photo Panin R., det. Solodovnikov I.A
Copyright © Ruslan Panin
Carabus (Tribax) steveni satanas Semenov, 1896
Male+Female adult
Carabus (Tribax) steveni satanas Semenov, 1896
Russia, C.Caucasus, Kabardino-Balkaria,
valley of River Cherek Balkarsky, h=1800 m
10.VI-20.VII.1992
A. Zamotajlov leg.
Copyright © Safronov
Carabus (Tribax) puschkini puschkini Adams, 1817
Male adult
Armenia,<br /> Shirak Province, Ashotsk, 18.VI.2001, P.V.Kazaryan leg.<br /> male 25mm
Copyright © Ruslan Panin
Carabus (Tribax) puschkini ishikawaianus Breuning & Ruspoli, 1970
Male adult
Copyright © Alexander Anichtchenko
Carabus (Tribax) kasbekianus veselyi Gottwald, 1980
Female adult
<b>Carabus (Tribax) kasbekianus veselyi Gottwald, 1980</b><br /> Caucasus, Georgia, Egrissky Mt.R., riverheads Tekhuri river, h=2400, A.G.Koval leg.; 04.07.1989; 25 mm.; photo A. Menstschicov
Copyright © Anton Menstschikov
Carabus (Tribax) kasbekianus onerosus Belousov & Zamotajlov, 1993 Paratype
Male+Female adult
Carabus (Tribax) kasbekianus onerosus Belousov & Zamotajlov, 1993 Paratypes
Caucasus, W Georgia, Svanetia, right bank o Magana riv.,
800m, 12.V.-4.VII.1991 Zamotajlov & Miroshnikov leg.
male 27mm, female 33mm
photo & collection Anichtchenko A.
Copyright © Alexander Anichtchenko
Carabus (Tribax) edmundi Semenov, 1896
Male adult
Carabus (Tribax) edmundi Semenov, 1896
Copyright © Boleslav Brezina
Carabus (Tribax) circassicus teberdensis Zolotarev, 1913
Male+Female adult
W-Caucasus: Karacaj-Cherkessija,
Abishira–Akhuba Mnt. Range h=2000-2650m, Kyafar-Agur Riv., 14–17.VII.2009, Fominykh D. leg.
female, 25mm
Copyright © Ruslan Panin
Carabus (Tribax) circassicus kubaniensis Semenov, 1896
Male adult
N Caucasus, Krasnodar reg., 30 km S Maykop
august 1993
male, 26 mm
Copyright © Alexander Anichtchenko
Carabus (Tribax) circassicus kubaniensis Semenov, 1896
Male+Female adult
Russia, NW Caucasus,
Krasnodar reg., Chernogorie mts., 24.V-29.VII.1999. A.S. Zamotajlov leg.
female 29mm/male 25mm
Copyright © Ruslan Panin
Carabus (Tribax) circassicus circassicus Ganglbauer, 1886
Male+Female adult
Abkhasia,
W.slope of Gagrinsky Mt., Krepostnaya River source, h=930-980m., A.S.Zamotajlov leg. 27.VII.2000
female, 29 mm/male, 26mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Tribax) circassicus abagonensis Starck, 1894
Male+Female adult
S Russia, NW Caucasus,
Adygea Rep., Mt.Abago, 26.VI.1999
L=24mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Tribax) biebersteini ulgeni Gottwald, 1982
Male adult
S Russia, Karacaj-Cherkessija,
near Kart-Djurt vill., upper Kylan riv. h=1300m, 26-27.V.2009, Fominykh D. leg.
male, 28mm
Copyright © Ruslan Panin
Carabus (Tribax) biebersteini otcharensis Kurnakov, 1970
Male adult
Caucasus, W Georgia, Svanetia, upper reaches of riv. Shoudidi, 1700-2100m, 20-21.V.1986 leg. A. Zamotajlov
Copyright © Evgeniy Chibilov
Carabus (Tribax) biebersteini ingusch Zolotarev, 1913
Male+Female adult
Carabus (Tribax) biebersteini ingusch Zolotarev, 1913
Russia, C.Caucasus, N.Ossetia,
valley of River Khazniaush
10.VI-31.VII.1993
A. Zamotajlov leg.
Copyright © Safronov
Carabus (Tribax) biebersteini constantinowi Starck, 1894
Male adult
Carabus (Tribax) biebersteini constantinowi Starck, 1894
NW-Caucasus, S Russia,
Karacaj-Cherkessija, Nizhniy Arkhyz, 21-30.V.2001, Tikhonov V. leg.
male,30mm
Copyright © Ruslan Panin
Carabus (Tribax) apschuanus apschuanus Rost, 1893
Male+Female adult
Carabus (Tribax) apschuanus apschuanus Rost, 1893
Russia, Caucasus, Krasnodar reg.,
h=1200 m, 29.VII.2000
A. Zamotajlov leg.
Copyright © Safronov
Carabus (Tribax) agnatus agnatus Ganglbauer, 1889
Male adult
NW-Caucasus,
Krasnodar reg., Mostovskoj distr., Skirda Mts. N. slope Yatyrgvart Mt., h=2300-2500m. 09-10.VII.2008, D.Fominyh leg.
male, 23mm
photo Panin R.
Copyright © Ruslan Panin
Carabus (Tribax) apschuanus pseudoplatessa Gottwald, 1982
Male+Female adult
NW-Caucasus, Karachay-Cherkessia, Teberda: Dzhamagat gorge, the left bank of the riv. Dzhamagat, 10.VII-28.VIII.2010 S.Pavluchuk leg.
female, 27 mm
Copyright © Ruslan Panin
Carabus (Tribax) nothus planipennis Chaudoir, 1846
Female adult
Dagestan, Huizah., near Motchoh, h=1800, 27.07.1993, Ilina leg.
photo A. Menstschicov
Copyright © Anton Menstschikov
Carabus (Tribax) nothus planipennis Chaudoir, 1846
Male adult
Dagestan, Bogosskii mt. rng, Inhokvari-Honoh
3500 m, 15-25.6.85 Abdurakhmanov leg.
Copyright © Alexander Anichtchenko
Carabus (Tribax) nothus (as abdurachmanovi)
Male adult
Carabus (Tribax) nothus (as abdurachmanovi)
male, 22mm
Copyright © Boleslav Brezina
Carabus (Tribax) nothus (as abdurachmanovi)
Female adult
Carabus (Tribax) nothus (as abdurachmanovi)
female, 23mm
Copyright © Boleslav Brezina
Carabus (Tribax) kasakorum kasakorum Semenov, 1896
Female adult
female, 29mm
Copyright © Boleslav Brezina
Carabus (Tribax) kasakorum kasakorum Semenov, 1896
Female adult
female, 29mm
Copyright © Boleslav Brezina
Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
Male adult
male, 23mm
Copyright © Boleslav Brezina
Carabus (Tribax) biebersteini otcharensis Kurnakov, 1970 (as retezari)
Female adult
Abkhazia, Lake Ritsa, 43-31'N / 40-37'E, 19-25.VII.2008, А. Vlasenko leg.
photo from zin.ru
Copyright © Dmitri Obydov
Carabus (Tribax) kraatzi Chaudoir, 1877
Male adult
Georgia: Achalzich (Chambobel), 1910, Korb leg.
photo from zin.ru
Copyright © Dmitri Obydov
Carabus (Tribax) macropus Chaudoir, 1877
Female adult
Georgia, Khoc(aldagh, prope Lagodechi, Kachetia, VI.1907.
photo from zin.ru
Copyright © Dmitri Obydov
Carabus (Tribax) kasakorum kasakorum Semenov, 1896
Male adult
Pr. Wladikavkas, montes, coll. A. Zolotarew
photo from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Tribax) kasakorum kasakorum Semenov, 1896
Male adult
Montes prope Wladikavkas, 6000 f., coll. A. Zolotarew
photo from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Female adult
Caucasus, N. Osetia, 60 km S Alagir, 1850 m, 12.IX.1990, A. Klimenko leg.<br /> photo from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Female adult
Cuaucas, N Osetia, Ardon riv., Alagir, Duravtuan, 28.IV.1982, A.Sokolov leg.<br /> photo from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Tribax) nothus planipennis Chaudoir, 1846
Male adult
Georgia, Lagodehi, Ninigora, 2750 м, near snow 24.V.1979, I.Belousov leg.
photo from www.zin.ru
Copyright © Dmitri Obydov
Carabus (Tribax) nothus formaneki Reitter, 1896
Male adult
Topotypus: Azerbaijan, Caucas. or., distr. Nukha, 3000 m, 20.VII.1934, A. Bogachev leg.
Copyright © Dmitri Obydov
Carabus (Tribax) biebersteini otcharensis Kurnakov, 1970
Male adult
W. Caucasus, Abkhazia, Gudauta distr.,
Duripsh vill. env., Khipsta River, h=700 m,
11-06-2009 A. Safronov leg.
male - 40 mm
Copyright © Alexandr Safronov
Carabus (Tribax) puschkini ishikawaianus Breuning & Ruspoli, 1970
Male adult
NE-Turkey: Artvin:Hopa, Borçka env., h=450 m, 29.VI-7.VII.2011, M.Janalik leg.
L=33 mm
Copyright © Ruslan Panin
Carabus (Tribax) puschkini ponticus Deyrolle, 1869
Male adult
NE-Turkey:Trabzon, Zigana Pass h=1700m,14-24.VI.2010, M.Janalik leg.
L=28 mm
Copyright © Ruslan Panin
Carabus (Tribax) circassicus circassicus (as tshchaltensis Novotný & Voříšek
Male adult
W-Caucasus: Abkhasia, Gudauta distr., E slope Mt Anchkho h=2200-2350 m, 14-20.VII.2011, S.Pavluchuk, I. Solodovnikov leg./det. I. A. Solodovnikov.
25 mm
Copyright © Ruslan Panin
Carabus (Tribax) agnatus pseudagnatus Novotný & Voříšek, 1988
Female adult
NW-Caucasus, Karachay-Cherkessia, Bol'shaya Marka River h=2500-2600m, 12.VIII.2010, I.A. Solodovnikov leg./det.
Copyright © Ruslan Panin
Carabus (Tribax) agnatus pseudagnatus Novotný & Voříšek, 1988
Male adult
W-Caucasus: Abkhasia, Gudauta distr., Mt Anchkho h=2200-2350 m, 14-20.VII.2011, S.Pavluchuk, I. Solodovnikov leg./det. I. A. Solodovnikov.
L=25 mm
Copyright © Ruslan Panin
Carabus (Tribax) apschuanus pseudoplatessa Gottwald, 1982
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Carabus (Tribax) biebersteini fossiger Chaudoir, 1877
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Carabus (Tribax) puschkini platypterus Ganglbauer, 1886
Male adult
SW Georgia, Lesser Caucasus, Adzhar-Imereti (Meskheti) Mt Range, 10 km S Sairme h~1850 m, 27.VI-2.VII.2013, A.Puchkov & Martynov leg. L=27 mm
Copyright © Ruslan Panin
Carabus (Tribax) puschkini kolenatii (as suramensis Kraatz, 1887)
Male adult
S-Caucasus: NC-Georgia, W slope Likhi Mt.Range, 3 km W Rikoti Pass h~900m 11-14.VI -2006, A.Puchkov leg. L=27 mm
Copyright © Ruslan Panin
Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
Male adult
Russia, C-Caucasus: N Ossetia, Chekhatsirtit Mt range, Mt Tepli h=3000 m, 21.VII.2009. /det. Solodovnikov I.A, L=20.5 mm
Copyright © Ruslan Panin
Carabus (Tribax) adelphus shtavlerensis Fominykh, Zamotajlov et Titarenko, 2018
Male adult
Georgia, Zemo Svaneti, left bank of Nenskra River near Zemo Margi Vill., 1490 m, 43°04'23.5''N, 42°11'55.2''E, 10.V.2018 (D. Fominykh) PARATYPE
Copyright © Igor Solodovnikov
Carabus (Tribax) adelphus shtavlerensis Fominykh, Zamotajlov et Titarenko, 2018
Male adult
Abkhazia, Tkuarchalsky district, Kodorsky Mt.R., a source of the river Mokvi, a talus under the pass, h = 2635 m, 17-23.07.2018, leg. I.A. Solodovnikov
Copyright © Igor Solodovnikov
Carabus (Tribax) macropus Chaudoir, 1877
Male adult
Daghestan: Bogosskyi Mts., Inkhokvari
Copyright © Igor Solodovnikov
Carabus (Tribax) macropus Chaudoir, 1877
Male adult
Daghestan: Bogosskyi Mts., Inkhokvari
Copyright © Igor Solodovnikov
Carabus (Tribax) kasakorum daghestanicola Rapuzzi, 2018 Holotype
Female adult
Russia, East Caucasus, Daghestan, Andiysky Mountain Range, Kharami pass, 2,500 m, 5.VII.2010, S. Rybalkin legit, Ivan Rapuzzi collection. Image from Rapuzzi, 2018
Copyright © - - -
Carabus (Tribax) kasakorum scintillans Rapuzzi, 2018 Holotype
Male adult
Russia, Caucasus, Ingushetia, South slopes of Skalistyi Mountain, Range, Beyni village environs, 2,800 m, 21.V/19.VII.2018, Ivan Rapuzzi collection. Image from Rapuzzi, 2018
Copyright © - - -
Carabus (Tribax) macropus Chaudoir, 1877
Male adult
Russia, Ingushetia, S slopes of Skalistyi Mt.R., Khahalgi Mt., alpin zone, h = 2500 m, 25.V-19.VII.2018 leg. D.Fominykh
Copyright © Igor Solodovnikov
Carabus (Tribax) macropus Chaudoir, 1877
Female adult
Russia, Ingushetia, S slopes of Skalistyi Mt.R., Khahalgi Mt., alpin zone, h = 2500 m, 25.V-19.VII.2018 leg. D.Fominykh
Copyright © Igor Solodovnikov
Carabus (Tribax) steveni schamyli Hampe, 1852
Female adult
C Caucasus, North Ossetia, the road on the village of Coban (Koban), the left river bank Gizeldon, the deciduous wood, h = 895-905 m, N 42°55'32.91", E 44°31'53.53", 20.05-03.07.2017, leg. A.P. Tatulyan
Copyright © Igor Solodovnikov
Carabus (Tribax) steveni schamyli Hampe, 1852
Female adult
C Caucasus, North Ossetia, the road on the village of Coban (Koban), the left river bank Gizeldon, the deciduous wood, h = 895-905 m, N 42°55'32.91", E 44°31'53.53", 20.05-03.07.2017, leg. A.P. Tatulyan
Copyright © Igor Solodovnikov
Carabus (Tribax) steveni schamyli Hampe, 1852
Male adult
C Caucasus, North Ossetia, the road on the village of Coban (Koban), the left river bank Gizeldon, the deciduous wood, h = 895-905 m, N 42°55'32.91", E 44°31'53.53", 20.05-03.07.2017, leg. A.P. Tatulyan
Copyright © Igor Solodovnikov
Carabus (Tribax) steveni schamyli Hampe, 1852
Male adult
C Caucasus, North Ossetia, the road on the village of Coban (Koban), the left river bank Gizeldon, the deciduous wood, h = 895-905 m, N 42°55'32.91", E 44°31'53.53", 20.05-03.07.2017, leg. A.P. Tatulyan
Copyright © Igor Solodovnikov
Carabus (Tribax) nothus sharoiensis Fominykh, Zamotajlov, Avtaeva et Titarenko, 2019 Paratype
Male adult
Copyright © Igor Solodovnikov
Carabus (Tribax) nothus fausti Dohrn, 1873
Male adult
Carabus (Microtribax) fausti fausti Dohrn, 1873 Russia: Daghestan, Kyabyak Mt.R.
Copyright © Igor Solodovnikov
Carabus (Tribax) nothus fausti Dohrn, 1873
Female adult
Carabus (Microtribax) fausti fausti Dohrn, 1873 Russia: Daghestan, Kyabyak Mt.R.
Copyright © Igor Solodovnikov
Carabus (Tribax) lederi lederi Reitter, 1882
Male adult
Georgia, Kvemo-Svaneti, Svanetian Mt.R., near Latpari pass., alpin zone, 2700-3000m (TOPOTYPUS)
Copyright © Igor Solodovnikov
Carabus (Tribax) lederi lederi Reitter, 1882
Female adult
Georgia, Kvemo-Svaneti, Svanetian Mt.R., near Latpari pass., alpin zone, 2700-3000m (TOPOTYPUS)
Copyright © Igor Solodovnikov
Carabus (Tribax) kasakorum kasakorum Semenov, 1896
Male adult
Russia: N.Osetia: Pastbyshchniy Mt.R., S slopes of Khoshkharanrag Mt.R., right bank of Amusadon riv., h=1500 m
Copyright © Igor Solodovnikov
Carabus (Tribax) kasakorum kasakorum Semenov, 1896
Female adult
Russia: N.Osetia: Pastbyshchniy Mt.R., S slopes of Khoshkharanrag Mt.R., right bank of Amusadon riv., h=1500 m
Copyright © Igor Solodovnikov
Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Male adult
Russia, Caucasus, Ingushetia, South slopes of Skalistyi Mt Range, Stolovaya Mt., near Beyni, alpin zone, h = 2800 m, N 42°51’35.30",E 44°42’13.70", h = 2800m, 21.V–19.VII.2018, D. Fominykh leg.
Copyright © Igor Solodovnikov
Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Female adult
Russia, Caucasus, Ingushetia, South slopes of Skalistyi Mt Range, Stolovaya Mt., near Beyni, alpin zone, h = 2800 m, N 42°51’35.30",E 44°42’13.70", h = 2800m, 21.V–19.VII.2018, D. Fominykh leg.
Copyright © Igor Solodovnikov
Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Male adult
Russia, Caucasus, Ingushetia, Skalistyi Mt. Range, S slopes of Tsoreylam Mt. Range, 5 km W of Mt. Khakhalgi, N 42°50’07.60", E 45°05’02.20", h=2500 m, 25.V–21.VII.2017, D. Fominykh leg.
Copyright © Igor Solodovnikov
Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Female adult
Russia, Chechnya, Caucasus, Skalystyi Mt. Range upper river Gekhi, Mt. Ekkyrkort, N 42°52', E 45°26', h=2230 m, 26.V–21.VII.2018, D.Fominykh leg.
Copyright © Igor Solodovnikov
Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
Female adult
Russia: N.Osetia, Bokovoy Mt.R., near Arkhonskiy pass., alpin zone, h = 2700 m, 23.V-21VII.2018, D. Fominykh leg.
Copyright © Igor Solodovnikov
Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
Male adult
Russia: N.Osetia, Bokovoy Mt.R., near Arkhonskiy pass., alpin zone, h = 2700 m, 23.V-21VII.2018, D. Fominykh leg.
Copyright © Igor Solodovnikov
Carabus (Tribax) puschkini kolenatii Chaudoir, 1846
Female adult
C-Georgia, Trialeti Mt Range, 2 km SE of Borjomi h=916 m, 41°48'35.3"N 43°26'11.8"E, 17.VI.2013, D. Prunier leg./det. Panin R., L=29 mm
Copyright © Ruslan Panin
Carabus (Tribax) puschkini kolenatii Chaudoir, 1846
Male adult
C-Georgia, Trialeti Mt Range, 2 km SE of Borjomi h=916 m, 41°48'35.3"N 43°26'11.8"E, 17.VI.2013, D. Prunier leg./det. Panin R., L=28 mm
Copyright © Ruslan Panin
Carabus (Tribax) boulbenicus Deuve, 2022 Holotype
Male adult
Géorgie, Svanétie, à l’est de Khaishi, vallée de la Kasleti, 950 m. R. Boulben et P. Stéfani leg.
Copyright © Thierry Deuve
Carabus (Tribax) adelphus shtavlerensis Fominykh, Zamotajlov et Titarenko, 2018 Holotype Paratype
Male+Female adult
Georgia, left bank of Nenskra River near Tita Vill., Shtavler Mt. Range, NW slopes of Utur Mt., 43°04'N, 42°11'E, 1048 m, 14.IV–13.VI.2016 D. Fominykh.
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Carabus (Tribax) adelphus adelphus Rost, 1892
Male+Female adult
Muzhava Vill. environs
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Carabus (Tribax) apschuanus galianus Belousov & Zamotajlov, 1993 Paratype
Male+Female adult
Mt.Okhachkue (L’Achake) GEORGIE, VII-1991
Copyright © Alain Mollard
Carabus (Tribax) biebersteini hurkai Gottwald, 1980
Male+Female adult
Chkumi (Racha-Lechkumi) GEORGIE, VII-2018
Copyright © Alain Mollard
Carabus (Tribax) kasbekianus mingrelicus Reitter, 1889
Male+Female adult
Chkhorodukhu GEORGIE, VII-2018
Copyright © Alain Mollard
Carabus (Tribax) osseticus Adams, 1817
Male+Female adult
Kzabegi , GEORGIE, VIII-1991
Copyright © Alain Mollard
Carabus (Tribax) biebersteini vlasenkoi Obydov, 2011 Holotype Paratype
Male+Female adult
Russia, North Caucasus,Stavropol Region, 33 km from Kislovodsk City to the north-west, Predgornyi Dist.,8 km from st. Bekeshevskaya to the south-west, Forestry “Ugolnaya Dacha”,Pastbischnyi Mt. Ridge, left coast of Kuma River, 800 m, N44º02’ E42°21’, 11-17.05.2006, A. Vlasenko leg.
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Carabus (Tribax) apschuanus galianus Belousov & Zamotajlov, 1993 Paratype
Male adult
Eastern Abkhazia, south slope of Mt. Okhachku, 4.V.1989, I. Belousov leg.
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Carabus (Tribax) apschuanus galianus Belousov & Zamotajlov, 1993
Male adult
Copyright © Igor Solodovnikov
Carabus (Tribax) apschuanus galianus Belousov & Zamotajlov, 1993
Female adult
Copyright © Igor Solodovnikov
Carabus (Tribax) edmundi Semenov, 1896
Female adult
Russia, S-Dagestan, 18.IV-19.VI.2022 leg. D. Fominykh
Copyright © Igor Solodovnikov
Carabus (Tribax) edmundi Semenov, 1896
Male adult
Russia, S-Dagestan, 18.IV-19.VI.2022 leg. D. Fominykh
Copyright © Igor Solodovnikov
Carabus (Tribax) puschkini kraatzi Chaudoir, 1877
Female adult
Georgia, Samtskhe-Javakheti, env. Uraveli vill., 14.VI.2024 leg. D. Fominykh
Copyright © Igor Solodovnikov
Carabus (Tribax) puschkini kraatzi Chaudoir, 1877
Male adult
Georgia, Samtskhe-Javakheti, env. Uraveli vill., 14.VI.2024 leg. D. Fominykh
Copyright © Igor Solodovnikov
Carabus (Tribax) apschuanus apschuanus Rost, 1893
Male adult
ABKHASIA, Sukhum distr., N slope of Dou Mt.R., left board of the river Bzyb, (Fagus + ferns + stony taluses (the beginning of a track on Dow's (Dou) pass), H = 740 m, 43°19'3.64" N, 40°52'37.55" E, 10.V-19.VI.2015, Leg. I.A. Solodovnikov, E.V. Tatun
Copyright © Igor Solodovnikov
Carabus (Tribax) kasbekianus mingrelicus Reitter, 1889
Female adult
Georgia, Caucasus, 3 km W Nakerala Pass 1450-1500 m, 42.386528, 42.977944, 22-25.VI.2024, I. Uspenskyi leg., L=27 mm
Copyright © Ruslan Panin
Carabus (Tribax) kasbekianus mingrelicus Reitter, 1889
Male adult
Georgia, Caucasus, 3 km W Nakerala Pass 1450-1500 m, 42.386528, 42.977944, 22-25.VI.2024, I. Uspenskyi leg., L=26 mm
Copyright © Ruslan Panin
Carabus (Tribax) kasbekianus veselyi Gottwald, 1980
Female adult
Georgia, Caucasus, Svaneti Range E part, Latpari Pass h=2700-2800 m, 23.VI.2024, I.Uspenskyi leg., L= 24 mm
Copyright © Ruslan Panin
Carabus (Tribax) kasbekianus veselyi Gottwald, 1980
Male adult
Georgia, Caucasus, Svaneti Range E part, Latpari Pass h=2700-2800 m, 23.VI.2024, I.Uspenskyi leg., L= 22 mm
Copyright © Ruslan Panin
Carabus (Tribax) kasbekianus veselyi Gottwald, 1980
Male adult
Georgia, Caucasus, S of Sasashi vill., Lechkhumi Range N slope h=2430 m, alpine meadows, 242.759167, 42.973889, 20-21.VII.2025, I.Uspenskyi leg./det. R.Panin, L=24 mm
Copyright © Ruslan Panin -
Ulocarabus taxon page (14 photos)
Carabus (Ulocarabus) theanus Reitter, 1895
Male adult
Vakhskiy Mt Range,Saryhosor, Mullonomi,
29.4.1990, S.Zonshteyn leg.
Copyright © Alexander Anichtchenko
Carabus (Ulocarabus) ? theanus Reitter, 1895
Male adult
NC Tadzhikistan, Pamir, N-slope Peter I.Mt Rng, h=2100m, Ganishou vill., 10-12.VI.2006, Ivanova E. leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Ulocarabus) stschurovskii Solsky, 1874
Male adult
Tadzhikistan, Pamir-Alay, S slope Gissar Mt.Range, Takob, h-1700m, 10.V.1996, Dolin V.G. leg.
det. Panin R.Yu.
L=22 mm
Copyright © Ruslan Panin
Carabus (Ulocarabus) theanus Reitter, 1895
Male adult
Tadzhikistan, S slope Gissar Mt.Rng, left bank of the river Sorbo,Haidar gorge,1.VI.1984,Dolin V.G. leg.
det.Panin R.Yu.
L=21 mm
Copyright © Ruslan Panin
Carabus (Ulocarabus) stschurovskii Solsky, 1874
Female adult
Tadzhikistan, Pamir, N-slope Peter I.Mt Rng,h=2100m,Ganishou vill., Dolin V.G. leg.
det.Panin R.Yu.
L=24 mm
Copyright © Ruslan Panin
Carabus (Ulocarabus) theanus delerei Mandl, 1955
Male adult
N-Afghanistan, Hindukush Mt.Rng, NE Salang Pass h=2300 m, 35°27'494"N, 68°58'399"E, IV.2011, C. Reuter leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Ulocarabus) theanus theanus Reitter, 1895
Male adult
Vakhskiy Mt Range, Sary Khosor, Mullokoni, 29.4.1990, S.Zonshteyn leg. (cAA)
Copyright © Alexander Anichtchenko
Carabus (Ulocarabus) theanus theanus Reitter, 1895
Male adult
Tadzhikistan, S slope Gissar Mt.Rng, left bank of the river Sorbo, Haidar gorge,1.VI.1984, Dolin V.G. leg./det.Panin R.Yu.
L=21 mm
Copyright © Ruslan Panin
Carabus (Ulocarabus)? theanus theanus Reitter, 1895
Male adult
NC Tadzhikistan, Pamir, N-slope Peter I.Mt Rng, h=2100m, Ganishou vill., 10-12.VI.2006, Ivanova E. leg.
L=23 mm
Copyright © Ruslan Panin
Carabus (Ulocarabus) theanus xerophiloides Deuve, 2019 Holotype
Female adult
NO LABEL DATA! ex coll. M. Maindron, ex coll. G. Babault (MNHN).
Copyright © Thierry Deuve
