Carabus (Morphocarabus) aeruginosus zyrjanovskianus Shilenkov & O.Berlov, 1996
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) aeruginosus verzhutzkii O.Berlov & Shilenkov, 1996
Male adult
Carabus (Morphocarabus) aeruginosus verzhutzkii O.Berlov & Shilenkov, 1996
C Altays, Katun r., 5 km. N Katanda vill., 21.06.2002, A. Menstschicov leg.
Copyright © Anton Menstschikov

Carabus (Morphocarabus) aeruginosus verzhutzkii O.Berlov & Shilenkov, 1996
Male adult
C Altai Mts., Seminskii Mt.rng., Katun riv., 3 km S Ust'-Sema riv.,
4.VII.2002 A. & N. Menschikovy leg.
det. Mentschikov A.
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) aeruginosus eschscholtzi Mannerheim, 1827
Male adult
Russia, W Altai, Zmeinogorsk distr., near Cherepanovskii vill., 2-16.6.1990, D. Obydov leg.
Copyright © Dmitri Obydov

Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
Endophallus
endophallus
Copyright © Ruslan Panin

Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822 (as pitschugini) Holotype
Male adult
East Siberia, Tunkinskie Goltzy (East Sajans), Mondy env., 24.VI.1979, E. Ya. Berlov leg.
Copyright © Dmitri Obydov

Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822 (as pitschugini) Paratype
Female adult
East Siberia, Tunkinskie Goltzy (East Sajans), Mondy env., 24.VI.1979, E. Ya. Berlov leg.
Copyright © Dmitri Obydov

Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
Male adult
Russia, S Tuva, Khorumnug-Taiga Mts, Buren riv. Valley, Buren-Bay-Khaak vill. h=900 m, 17-24.VI.2002, Vashchenko A.& B.leg.,/ det. Panin R. L=25 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
Male adult
Russia, NE Altay, Turochak Distr., Lake Teletskoe N part, near Yaylyu vill. 23-25.VII.2019, Ivliev P. leg./ det. R. Panin, L=24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) aeruginosus eschscholtzi Mannerheim, 1827
Male adult
NE Kazakhstan, W Altay Mts, near Shemonaikha city., flood plain, 25-28.VII.2018, S.Vashchenko leg., L=22.5 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) brinevi Kabak, 2014 Paratype
Male adult
China, Xinjiang, S Altai Mts, Keshtau Mt., ENE Qinghe, 46°47'21''N / 90°44'42''E, 2730 m, 12.08.2013, I.I. Kabak leg.
Copyright © Ilya Kabak

Carabus (Morphocarabus) brinevi Kabak, 2014 Holotype
Male adult
China, Xinjiang, S Altai Mts, Keshtau Mt., ENE Qinghe, 46°47'21''N / 90°44'42''E, 2730 m, 12.08.2013, I.I. Kabak leg.
Copyright © Ilya Kabak

Carabus (Morphocarabus) brinevi Kabak, 2014 Paratype
Endophallus

Copyright © Ilya Kabak

Carabus (Morphocarabus) brinevi Kabak, 2014 Paratype
Female adult
China, Xinjiang, S Altai Mts., Keshtau Mt., ENE of Qinghe, h~2730 49°47'21"N 90°44'42"E 12.VIII.2013 leg. I.Kabak L=19.5 mm
Copyright © Kirill Makarov

Carabus (Morphocarabus) tarbagataicus altaicola Deuve & Tian, 2019 Holotype
Female adult
Chine, Xinjiang, Préfecture d’Altaï, Monts Altaï, au nord de la ville d’Altaï, Parc forestier de Xiaodonggou (Chen Liusheng, 8.VIII. 2014), in coll. South China Agricultural University, Canton.
Copyright © Thierry Deuve

Carabus (Morphocarabus) brinevi altaicola Deuve & Tian, 2019
Female adult
«qing tong se», «28#» (ICXU)
Copyright © Ilya Kabak

Carabus (Morphocarabus) brinevi kanasensis Kabak, 2021 Holotype Paratype
Male adult
China, Xinjiang, Altai Mts, left bank of Kanas lake, 48°44ʹ35ʺN / 87°06ʹ03ʺE, H = 2710 m, alpine meadows, 15.07.2019 I.I. Kabak leg. (ZIN)
Copyright © Ilya Kabak

Carabus (Morphocarabus) chaudoiri zaikai Obydov, 1999 PT Paratype
Female adult
Carabus (Morphocarabus) chaudoiri zaikai Obydov, 1999 PT
Rusia, Tuva, Akademik Obruchev Mt. rng., 2100 m;, 52°00'N 95°34'E,
Sainak pass, tundra, 2.VII.1998, D. Obydov leg.
female,
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) chaudoiri juliae Obydov, 2002
Male adult
male, 18mm
Copyright © Boleslav Brezina

Carabus (Morphocarabus) chaudoiri czadanicus Obydov, 1997 HT Holotype
Male adult
Carabus (Morphocarabus) chaudoiri czadanicus Obydov, 1997 HT
Rusia, Tuva, near Hadyn vill., 28-30.VI.1994, R. Shmarzh leg.
male,
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) chaudoiri beladici Obydov, 2001
Male adult
S-Tuva
East Tannu-Ola,Biche-Sailig r.h=2100m, 15.VII.1996, D.E.Lomakin leg.
Copyright © Anton Menstschikov

Carabus (Morphocarabus) chaudoiri beladici Obydov, 2001 PT Paratype
Female adult
Carabus (Morphocarabus) chaudoiri beladici Obydov, 2001 PT
Rusia, Tuva, E Tannu-Ola Mt.rng., near Kara-Hol lake, 29-31.V.2000, M. Beladzhik leg
female,
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) chaudoiri marusiki Obydov, 1997 Holotype
Male adult
image from zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) chaudoiri mongolorum Csiki, 1927 Lectotype
Male adult
Muséum national d’Histoire naturelle, Paris (France); Collection : Insects: Coleoptera (EC); Specimen MNHN-EC- EC522; Credit: Antoine Mantilleri - 2010 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec522
Copyright © - - -

Carabus (Morphocarabus) chaudoiri juliae Obydov, 2002 Holotype Paratype
Male+Female adult
Russia, Tuva Republic, Mungun-Taiga, Khindiktig-Khol Lake, 2400 m, 22.-29.VI.2000, M. Beladič leg.
Copyright © - - -

Carabus (Morphocarabus) chaudoiri chaudoiri Gebler, 1847 Lectotype
Female adult
Altay
Copyright © Dmitri Obydov

Carabus (Morphocarabus) chaudoiri shilenkovi O.Berlov, 1989
Male adult
Россия, Якутия, Оймяконское нагорье, 12-18.VII.1998
Copyright © Dmitri Obydov

Carabus (Morphocarabus) excellens excellens Fabricius, 1798
Male adult
Moldova, 2003
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens ( as jasilkowskii Born,1905)
Male adult
NE Romania, 5 km W Suceava, 47°63′940″N 26°18′795″E,h=340m, agricultural field, 15-20.V.2011, B. Brezina leg. L=24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens paradoxus
Label
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC539
Copyright © - - -

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Male adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 4-24.V.2010, Panin R. leg. L= 29 mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus (as polonicus Lomnicki, 1886)
Male adult
SW Poland: Przemyśl env. photo by Mieczyslaw Stachowiak
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Holotype
Male adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 28.V-9.VI.2013, Panin R. leg. L=29 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Female adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 28.V-9.VI.2013, Panin R. leg. L= 28 mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Male adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 12.V-2.VI.2009, Panin R. leg. L= 29 mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens (as lomnitzkii Reitter, 1896)
Female adult
W Ukraina, Podolian Upland, steppe reserve Kasova Gora near Burshtyn vill., 15.VI -28.VII.2020, R. Panin & D. Khrapov leg., L=27 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens (as lomnitzkii Reitter, 1896)
Male adult
W Ukraina, Podolian Upland, steppe reserve Kasova Gora near Burshtyn vill., 15.VI -28.VII.2020, R. Panin & D. Khrapov leg., L=24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) gebleri ultimus Obydov, 1999
Female adult
Carabus (Morphocarabus) gebleri ultimus Obydov, 1999 PT
E Kazakhstan, Altai, near Zyrjanovska, 6.VII.1995, D. Obydov leg.
female
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) gebleri ultimus Obydov, 1999 Holotype
Male adult
Carabus (Morphocarabus) gebleri ultimus Obydov, 1999 HT
E Kazakhstan, Altai, near Zyrjanovska, 6.VII.1995, D. Obydov leg.
male
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) gebleri gebleri Fischer, 1817
Female adult
Carabus (Morphocarabus) gebleri gebleri Fischer, 1817
Russia, Altai, Zmeinogorsk
female
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) gebleri gebleri Fischer, 1817
Male adult
Carabus (Morphocarabus) gebleri gebleri Fischer, 1817
Rusia, Altai, Zmeinogorsk
male
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) helleri Ganglbauer, 1893
Male adult
Moravia Mer., Bile Karpaty 28.5.1993 Budka leg. male, 28 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) helleri Ganglbauer, 1893
Male+Female adult
SE Moravia, Strani, V.2002, M.Snizek leg. male bronze colour-26mm male blue colour-28mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) helleri Ganglbauer, 1893
Endophallus
SE Moravia, Strani, V.2002, M.Snizek leg. male bronze colour.
Copyright © Ruslan Panin

Carabus (Morphocarabus) helleri Ganglbauer, 1893
Endophallus
SE Moravia, Strani, V.2002, M.Snizek leg. male blue colour
Copyright © Ruslan Panin

Carabus (Morphocarabus) helleri Ganglbauer, 1893 (as pseudopreyssleri Breuning, 1932)
Male adult
Czech Rep., NE Moravia, Moravian-Silesian Beskids Mt Rng, PR Trojacka, Hodslavice vill., 29.V.12.VI.2010, Krause Rene leg./det.Panin R. L=30 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) helleri Ganglbauer, 1893 (as pseudopreyssleri Breuning, 1932)
Endophallus
Czech Rep., NE Moravia, Moravian-Silesian Beskids Mt Rng, PR Trojacka, Hodslavice vill.,
Copyright © Ruslan Panin

Carabus (Morphocarabus) helleri Ganglbauer, 1893
Male+Female adult
CZECH, SE Moravia, Velka nad Velickou. 20-V-2005, V. Slovak leg. male - 26 mm, female - 30 mm.
Copyright © Alexandr Safronov

Carabus (Morphocarabus) helleri Ganglbauer, 1893
Male adult
CZ; CE Moravia, Bílé Karpaty Mts.; 24mm
Copyright © Aleš Sedláček

Carabus (Morphocarabus) helleri Ganglbauer, 1893
Male adult
CZ; CE Moravia, Bílé Karpaty Mts.; 25mm
Copyright © Aleš Sedláček

Carabus (Morphocarabus) henningi pelevini Obydov 2006 HT
Male adult
Carabus (Morphocarabus) henningi pelevini Obydov 2006 HT
Kazakhstan, S Altai, Sarym-Sakty Mt.rng., 3000-3400 m,
18-20.VI.1998, A. Klimenko leg.
Copyright © Dmitri Obydov

Carabus (Morphocarabus) henningi henningi Fischer, 1817
Male adult
S. Ural, Bashkiria, near Mesjagutovo vill., 09.07.2001 A. Menstschicov leg.
Copyright © Anton Menstschikov

Carabus (Morphocarabus) henningi henningi Fischer, 1817
Male adult
SE Siberia, Russia, Irkutsk reg., near Pivovarikha vill., 7.V.07, D. Fominykh leg. female, 21mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) henningi hamarensis Shilenkov, 1996
Female adult
S Siberia, Hamar-Daban mts., Snezhnaya riv., 1-6.VI.2007, Anichtchenko A. leg. & det., 20 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) henningi hamarensis Shilenkov, 1996
Female adult
Carabus (Morphocarabus) henningi hamarensis Shilenkov, 1996 loc.typ.
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) henningi melyachi Shilenkov & Menstshikov, 1996 loc.typ.
Male adult
Russia, Altay, Katun Mt Range, Karakol lake,22.VI.1993, Melyach F.V. leg.<br /> L=18 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) henningi sahlbergi Dejean, 1826
Label
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC1 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec1
Copyright © - - -

Carabus (Morphocarabus) henningi cf saldaitisi Obydov 1997
Male adult
Russia, S Tuva, E Tanu-Ola Mts., Cha-Ova-Art Pass, h=1700m, 28.VI-3.VII.2000, Vashchenko S. leg. L=19.5 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) henningi pelevini Obydov 2006
Female adult
E Kazakhstan, S Altay, Sarym-Sakty Mt Range, 12 km SE Ornek vill. h=1200m, 10.VII.1998, L=21 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) henningi henningi Fischer, 1817
Male adult
Russia, S Siberia, NW Buryatia, E Sayans Mts., Tunka Range, near Mondy vill., h=2400 m, alpine tundra, 1-15.VI.2001, Mishustin R. leg./det. R.Panin
Copyright © Ruslan Panin

Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
Male adult
China, Jinling pro.;
Photo by liuye
det. Anichtchenko A.
Copyright © Ye Liu

Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
Male adult
Russia, Far East,
Primorskiy Kray, near Kamenushka vill., 2-3.VIII.1992, K.E.Dovgaylo leg.
male, 27mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) hummeli pusongensis Imura, 1993
Male adult
S Primorie, Chernye gory, Poima riv.,
30.7-5.8.1999 Anichtchenko A. leg.
male, 26 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) hummeli nevelskii Shilenkov, 1996
Male+Female adult
Russia, Sakhalin isl.,
Makarov distr., Lesnaja riv., VII.1992, M. Nesterov leg.
female,23mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) hummeli hummeli Fischer, 1823
Male adult
Carabus (Morphocarabus) hummeli hummeli Fischer, 1823
Rusia, Chita reg., Darasun, VII.1991
I. Avdeev leg.
male, 23 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
Male adult
E.RUSSIA, Primorsky ter., Chuguevsky distr., 30 km ESE from Chuguevka, 9-14.VIII.2007
Copyright © Nil Belonogov

Carabus (Morphocarabus) hummeli ? middendorfi Menetries, 1851
Male adult
Carabus (Morphocarabus) hummeli ? middendorfi Menetries, 1851
Rusia, Khabarovsk reg., Ohotsk sea, Port Ayan, 08.24
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878 (as viridipennis Kraatz, 1886, nec Lewis,18
Male+Female adult
Russia, Far East,Primorsky reg., S. Sikhote-Alin Mts. R., Kavalerovo vill. env.,
5-25.VII.1992, С. Kunitzyn leg.
male - 25 mm, female - 25 mm.
Copyright © Alexandr Safronov

Carabus (Morphocarabus) hummeli stolidus Lapouge, 1924
Label
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC519
Copyright © - - -

Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
Male adult

Copyright © Kirill Makarov

Carabus (Morphocarabus) illigeri illigeri Dejean, 1826 (as bjelasnicensis Apfelbeck, 1902)
Endophallus
endophallus as bjelasnicensis Apfelbeck, 1902
Copyright © Ruslan Panin

Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Male adult
Serbia, Majdanpek, Ceremosja 15.6.2001 Z.Zlatic leg. male, 32 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Endophallus
NE-Serbia, Krs Mts., Majdanpek env.,h=300m.20-31.V.2006, Pr.Sima leg
Copyright © Ruslan Panin

Carabus (Morphocarabus) illigeri versicolor Frivaldsky, 1835
Male+Female adult
C Bulgaria:Stara planina Mts, Gabrovo,12-15.VI.2007, M. Janalik leg. female, 31 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) illigeri versicolor Frivaldsky, 1835 (ab.)
Female adult
C Bulgaria: Stara Planina Mts, Schipka pass h=1450m, 12-15.VI.2007, M.Janalik leg. L=31 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) illigeri illigeri (as tarensis Toševski, 1990)
Male adult
WC-Serbia: National park Tara Mts.:NE Mitrovac h=1500m, 1-12.VII.2007, Pr.Sima leg. L=30 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) illigeri illigeri (as tarensis Toševski, 1990)
Endophallus
WC-Serbia: National park Tara Mts.:NE Mitrovac h=1500m,
Copyright © Ruslan Panin

Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Male adult
NE Serbia, Majdanpek; male, 33mm
Copyright © Aleš Sedláček

Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Female adult
NE Serbia, Kučevo; female, 33mm
Copyright © Aleš Sedláček

Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Female adult
NE Serbia, Majdanpek; 35mm
Copyright © Aleš Sedláček

Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
Larva

Copyright © Evgeniy Chibilov

Endofallus Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
Male adult
Russia,
S Ural Mts, Mt.Iremel h=800m, 4.VI.2000 Filimonov V. leg.
male,21mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
Male adult
Carabus (Morphocarabus) karpinskii Rusia, S Ural mts., Jremel Mt., 1300 m 4.6.2000male, 20 mm
19.08.2008
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) karpinskii Kryzhanovskij & Matveev, 1993
Male adult

Copyright © Kirill Makarov

Carabus (Morphocarabus) kollari Palliardi, 1825
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) kollari Palliardi, 1825
Female adult
female, 24 mm
Copyright © Boleslav Brezina

Carabus (Morphocarabus) kollari Palliardi, 1825
Male adult
male, 27 mm
Copyright © Boleslav Brezina

Carabus (Morphocarabus) kollari Palliardi, 1825
Male+Female adult
Romania, SW Carpathians,
Banat, Anina Mts. (Muntii Aninei) 8 km NE Oravita, 7.VI.2007, S.I. Budka leg.
female, 34 mm
photo Panin R
Copyright © Ruslan Panin

Carabus (Morphocarabus) kollari Palliardi, 1825 (as semetricus Kraatz, 1878)
Male adult
SW-Romania:Banat, Bocsa/Coltan, 21.V-5.VI.2010, Krause Rene leg.
L=29 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) kollari Palliardi, 1825 (as semetricus Kraatz, 1878)
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) kozhantschikovi martjanovi Lutshnik, 1924
Female adult
Khakassia, 1800 m, near Cheremushki vill., 12.6.-20.7.2000
female, 21 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
Male adult
W Sayan Mts., Kazhyr-Suk riv., 80 km W Oleniya
Rechka, W part of Oiskii mt. rng., 30.6.-7.7.1999
2100 m, Brinev A. leg.
male, 19 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
Male adult
W Sayan, low fl. Kazyr-Suk riv., 80km W meteostation Olenia Rechka, W part of Oiskii mt.rng., 30.VI-7.VII.1999 2100m, Brinev A. leg.
male, 20mm
det. Brinev A.
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
Endophallus

Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
Male adult
Russia, W Sayan Mts., Kazyr-Suk river, W slope of Oyskiy Mt. rng., ?h=2200 m 7-15.VII.1999, Brinev A. leg., L=19 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) mestscherjakovi telezkianus Shilenkov, 1996
Male adult
Carabus (Morphocarabus) mestscherjakovi telezkianus Shilenkov, 1996
Rusia, Altai, S shore of Teletskoe lake, 5.VII.1968
Copyright © Dmitri Obydov

Carabus (Morphocarabus) mestscherjakovi telezkianus Shilenkov, 1996
Male adult
Carabus (Morphocarabus) mestscherjakovi telezkianus Shilenkov, 1996
SW Khakasia, Abakanskii Mt.rng., Choochek Mt., 20 km SSE Mrassu vill.,
1800m, 7-19.7.1999 Lomakin D. leg.
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
Male+Female adult
Russia, S Siberia,
S Khakassia, N Tuva reg., Sailyg-Khem-Taiga mts. range h=2000-2600m,10-30.VI.2005, S.Vashchenko & S.Alexeenko leg.
male,17.5mm/female,19mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
Endophallus
endophallus
Copyright © Ruslan Panin

Carabus (Morphocarabus) mestscherjakovi pseudoodoratus Brinev, 2002 Paratype Paratype
Male adult
Krasnojarsk reg., Aradan distr., 5 km SW Aradan vill.,<br /> Us riv., forest, 52є32`N 93є27`E 9-12.VII.1998 A.Brinev leg<br /> male, 18 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
Male adult
Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
W Sayan, Olenya rechka
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
Male adult
Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
W Sayan, Kulumys Mt.R., env. Oiskoje Lake, mountain tundra, 12-14. 07.1999 A.Semin leg.
male
L=17,0 mm
Copyright © Maksim Smirnov

Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
Male+Female adult
Russia, S Siberia, W Sayan, Ergaki Mt range, Nizhnyaya Buyba river h=1700 m, 19.VII.1999, A.& N. Menstschikovy leg.
Copyright © Ruslan Panin

Carabus (Morphocarabus) mestscherjakovi zherichini Shilenkov, 1990 Paratype
Female adult
RUSSIA, Krasnoyarsk Terr., SW Tajmyr Pen., Nizhnyaya Agapa Riv., 40 km of source, south tundra, plakor, brooks holmme, on sandy soil, 7.VIII.1973 (V.V. Zherikhin).
Copyright © Roman Dudko

Carabus (Morphocarabus) michailovi Kabak, 1992
Male adult
NE Kazakhstan, S Altay Mts., Sarymsakty mt. rng.,
Sarymsakty riv., 2400 m, 22.6.2000 Nekroshevich S. leg.
male, 18 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) michailovi Kabak, 1992
Male adult
NE Kazakhstan, S Altay Mts., Sarymsakty mt. rng.,
Sarymsakty riv., 2400 m, 22.6.2000 Nekroshevich S. leg.
male, 17 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) michailovi Kabak, 1992
Male adult
NE-Kazakhstan,
S Altai Mts.,Sarym-Sakty Range, Sarym-Sakty riv., h=2400m. 21.VI.1999., A.Klimenko leg.
male,18mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) monilis Fabricius, 1792
Male adult
France 39, Foncina le Haut 300m, 23.6.1979; 29 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) monilis Fabricius, 1792
Endophallus
France 39, Foncina le Haut 300m
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) monilis Fabricius, 1792 (as taunicus)
Male adult
W Germany, Vogelsberg Mt., 700m, 13.5.1985
male, 24 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) monilis Fabricius, 1792 (as interpositus Géhin, 1880)
Male adult
NW France,
Basse-Normandie, Orne, Forкt d'Йcouves, V.2009, Pascal Stefani leg.
male, 29mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) monilis Fabricius, 1792 (as interpositus Géhin, 1880)
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) monilis (as interpositus Gehin, 1880)
Male+Female adult
NW France,
Haute-Normandie reg., Bois Les Loges, Etretat, Seine-Maritime,10.V.1988
female, 28mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) monilis (as helveticus Heer, 1837 Holotype) Holotype
Male adult
Carabus (Morphocarabus) monilis (as helveticus Heer, 1837 Holotype)
?hybrid of monilis/cancellatus
Museum of Zurich
photo Anichtchenko
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) monilis (as dupeuxi Deuve, 1974)
Male+Female adult

France,
Charente, Foret de Braconne VII.1998, Laforgue leg.
famale, 27mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) monilis Fabricius, 1792
Female adult
France, Dröme, Ft. de Saou, 17.vii.1975-M. Grotz Leg.-Coll.RBINS, N. Mal as monilis saouensis nicodi
Copyright © Noël Mal

Carabus (Morphocarabus) monilis tanarguensis Maguerre & Stefani, 2021 (syn.) Holotype Paratype
Male+Female adult
Holotype, m (21 mm), environs du Col de la Croix de Bauzon, 1300 m, communes de La Souche, Borne et Mayres, Ardèche, France (07/2003), in coll. D. Maguerre, à Mons-en-Barœul, France. Paratype f (23 mm), au dessus du Col de Meyrand, 1400 mètres, communes de Loubaresse et de Borne, Ardèche, France, (07/2004), in coll. P. Stéfani.
Copyright © - - -

Carabus (Morphocarabus) odoratus septentrionalis Breuning 1932
Female adult
N. Russia, arctic Urals, Paypudynya Riv., Pompeliz Mt.h=500-700 m.,10-15.07.2002
S. Andreev, A. Romanenko leg.
Copyright © Safronov

Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844: 100
Endophallus
endophallus
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) odoratus martjanovianus Obydov 1999 HT Holotype
Male adult
Carabus (Morphocarabus) odoratus martjanovianus Obydov 1999
Rusia, Sayan Mts., near Sayanogorsk, 7.VII.1993
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) odoratus krugeri Obydov 1999 PT Paratype
Female adult
Carabus (Morphocarabus) odoratus krugeri Obydov 1999 PT
Rusia, Tuva, Akademika Obrucheva Mt.rng., 2100, 51°56'N 095°32'E,
Koptu pass, tundra 2.VII.1998, D. Obydov leg.
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) odoratus dohrni Gebler, 1847
Male adult
Carabus (Morphocarabus) odoratus dohrni Gebler, 1847
Rusia, Kuznetskii Alatau, Taizhe-Su riv., 53°55'N 89°10'E,
1250-1650 m, 25.VI-30.VII.2002, V. Murzin and V. Sinyaev leg.
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) odoratus dabanensis Shilenkov 1996 loc.typ.
Male+Female adult
Carabus (Morphocarabus) odoratus dabanensis Shilenkov 1996 loc.typ.
Buryatia, Hamar-Daban mt. rng., Snezhnaya riv.,
4.6.2007 Anichtchenko A. leg.
male 17 mm, female 19 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) odoratus ?antropovi Shilenkov 1996
Male adult
S Tuva, Horumnut-Taiga Mt.rng., Bureya riv., 2000 m, tundra, Vaschenko S. leg. 18 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) odoratus melleus Lapouge, 1909 Paralectotypes
Male adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC521
Copyright © MNHN

Carabus (Morphocarabus) odoratus melleus Lapouge, 1909 Lectotype
Female adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC520
Copyright © MNHN

Carabus (Morphocarabus) odoratus kurmaevi I.Solodovnikov, 2017 Holotype
Male adult
Russia, Siberia, Krasnoyarsk Krai, the North Yenisei district, 55 km of CE of the vill. Bryanka, valley of the river Chirimba, stream Nakhodny, h = 350 m, the mixed forest (birch and pine); on a steep slope along an animal track on a hill brow, 23.VII-20.VIII.2015, leg. A.V. Kurmaev
Copyright © Igor Solodovnikov

Carabus (Morphocarabus) praecellens trans scheidleri
Endophallus
NC Croatia, Ivanščica Mts , Ivanec,4-18.VI.2011, Krause Rene leg.
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens praecellens Pillardi, 1825
Male adult
Serbia ,
Frushka Gora, Ruma VI.1985
Copyright © Igor Solodovnikov

Carabus (Morphocarabus) praecellens praecellens Pillardi, 1825
Endophallus
Serbia, Fruska Gora Mts, 20-31.V.2006, M.Sima leg.
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens praecellens Pillardi, 1825
Male adult
Serbia, Fruska Gora Mts, 20-31.V.2006, M.Sima leg.
L=29 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens jucundus Csiki, 1906
Male adult
N Hungary, Pilis Mts., Piliszentlelek, 20-27.VII.2011, Krause Rene leg./det. R.Panin
L=30 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens jucundus Csiki, 1906
Endophallus
N Hungary, Pilis Mts., Piliszentlelek,20-27.VII.2011, Krause Rene leg.
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as pseudojucundus Retezár, 1974)
Male adult
NW-Hungary::Komarom distr., Tardos, Gerecse Mts., 20.-27.V.2011, Krause Rene leg.
L=26 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as pseudojucundus Retezár, 1974)
Endophallus
NW-Hungary::Komarom distr., Tardos, Gerecse Mts.
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as vertesensis Retezár, 1974)
Male adult
NW Hungary: Fejer: Vértes Mt Rng, Tatabanya, 28.VII-2.VIII.1996
L=29 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as vertesensis Retezár, 1974)
Endophallus
NW Hungary: Fejer: Vertes Mt Rng, Tatabanya
Copyright © Ruslan Panin

Carabus (Morphocarabus) regalis regalis Fischer, 1822
Male adult
Carabus (Morphocarabus) regalis regalis Fischer, 1822
S. Ural, Bashkiria, near Mesjagutovo vill., 09.07.2001 A. Menstschicov leg.
photo A. Menstschicov
Copyright © Anton Menstschikov

Carabus (Morphocarabus) regalis jenissoni Dejean, 1831
Male adult
Carabus (Morphocarabus) regalis jenissoni Dejean, 1831
Rusia, Tuva, Todzh, Azas lake, 990 m,
52°24'N 96°33'E, 8.VII.1998, D. Obydov leg.

Copyright © Dmitri Obydov

Carabus (Morphocarabus) regalis jenissoni Dejean, 1831
Female adult
Carabus (Morphocarabus) regalis jenissoni Dejean, 1831
SW Siberia, Altays Mts, Choya distr., Sukhoy Karasuk env.
23.07.2004, A.Safronov leg.
Copyright © Safronov

Carabus (Morphocarabus) regalis eltoni Obydov, 2006 Holotype
Male adult
Carabus (Morphocarabus) regalis eltoni Obydov, 2006 HT
Rusia, Tver distr., near Ilyinskoe vill.,
29-30.VII.2006, D. Obydov leg.
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) regalis tschusovae Obydov, 2004
Male+Female adult
NE-Kazakhstan, W Altay, Ivanovsky Mt.Rng, near Ridder (Leninogorsk), h-1300-1500m, 4-6.VII.2003
female, 25 mm
det.Panin R.
Copyright © Ruslan Panin

Carabus (Morphocarabus) regalis tschusovae Obydov, 2004
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) regalis jenissoni Dejean, 1831 (as cyanescens Kraatz, 1882, nec Sturm, 1815)
Male+Female adult
Kazakhstan, E. Kazakhstan area,
Zyryanovsk env., near Putintzevo vill.
10-VI-2005 M. Danilevsky leg.
male - 24 mm, female - 25 mm.
Copyright © Alexandr Safronov

Carabus (Morphocarabus) rothi incompsus (as mendax Csiki,1906)
Male+Female adult
W.Ukraine,
Transcarpathian Reg., Rakhov Distr., Chornogora Mt Range, h=1900m. alpine zone, Panin R. leg/det. 28.VI-9.VIII.2008
female, 32 mm
photo endophallus Solodovnikov I.A
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi alutensis Savulescu, 1972
Male adult
C Romania,
Olt River: Rimnica Vilcea VII.1971, Savulescu N. leg.

Copyright © Igor Solodovnikov

Carabus (Morphocarabus) rothi hampei Kuster, 1846 (transition to rothi incompsus Kraatz, 1880)
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi rothi Dejean, 1829
Endophallus
C Romania, Sibiu Area, right bank Olt River, Bradu vill., h=380 m,
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi incompsus Kraatz, 1880
Endophallus
C Romania, Brașov County, Cristian vill., h=380 m
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi incompsus Kraatz, 1880
Male+Female adult
C Romania, Brașov County, Cristian vill., h=380 m,V-VI.2008, M.Pasek leg.
female, 27 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi alutensis Savulescu, 1972
Male+Female adult
C Romania, S Carpathians, Rimnicu Vilcea env., Calimaneşti, Cozia Mts., VII.2007, I. Marvan leg.
L=29 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei (as hunyadensis Csiki, 1926)
Male adult
CW Romania, Apuseni Mts, Metalliferous Mt Rng, Hunedoara reg., near Săcărâmb vill., 45°97'472"N/23°02'630"E, h=680 m, mixed forest, meadows, 15-20.V.2011, B.Brezina leg.
L=36 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi rothi (as equistriatus Kraatz, 1878)
Female adult
C Romania, Sibiu reg., left bank Olt River, Avrig env., 12.VI.2013, V. Slovak leg.
L=30 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi distinguendus (as subparvulus Mandl, 1965)
Male+Female adult
NC-Hungary:Borzsony Mts, Nograd-Diosjeno,1-14.V.2004, Krause Rene leg. female, 32 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
Female adult
Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
Moravia mer.,Vranov, 26-29.06.1979, Dr. Resl leg.
photo A. Menstschicov
Copyright © Anton Menstschikov

Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
Male adult
Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
SE Moravia mer.,Vranov, 26-29.06.1979, Dr. Resl leg.
photo A. Menstschicov
Copyright © Anton Menstschikov

Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
Male adult
Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
S.Bohemia,
Hluboka nad Vltavou, 24.VI-18.VII.1990, S. Budka leg.
male, 27mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799 (as scheidleri floriani)
Male adult
Carabus (Morphocarabus) scheidleri scheidleri (as scheidleri floriani)
A.: Steiermark, Rein Umg. VIII.1983 Kenyery
female, 32 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997
Male adult
Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997
Tuva, Akademika Obrucheva mt.rng., 15 km N Cherbi vill.
ur. Chailag 7.VII.1999 A. & N. Menschikovy
male, 21 mm
det. Menschikov A.
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) spasskianus shoriensis Obydov, 1999 HT Holotype
Male adult
Gornaya Shoria, 8.VIII.1978, S. Nikireev leg.

Copyright © Dmitri Obydov

Carabus (Morphocarabus) spasskianus saragaschensis Obydov, 1999
Male adult
W Sayan Mts., Batenevskii Mt.rng., 12 km E Pervomaiskoe vill.,
Enisey riv., mountain steppe, 400m, 5.VII.2000 A. & N. Menschikovy leg.
female, 26mm

Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) spasskianus saragaschensis Obydov, 1999 Holotype
Male adult
Rusia, Khakassia, Saragash, 400 m, 6.VII.1994, D. Obydov leg.

Copyright © Dmitri Obydov

Carabus (Morphocarabus) spasskianus microputus O.Berlov & Shilenkov, 1996
Male adult
Rusia, SW Tuva, W Tannu-Ola mt.rng., bass. of Barlyk riv.,
50°29'N 90°44'E, 30.VII.2000, D. Obydov leg.
Copyright © Dmitri Obydov

Carabus (Morphocarabus) spasskianus shoriensis Obydov, 1999
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997
Male+Female adult
Russia,Tuva,
Akademika Obrucheva Mt range, 15 km N Cherbi vill., Chailang ur. 7.VII.1999, A & N. Menstschikovy leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus geblerianus Obydov, 1999
Male adult
Russia, NW Altay, Barnaul env.,23.07.1998, D.Lomakin leg.

Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus cf putus Motschulsky, 1846
Female adult
Russia, SW Siberia, C-Altays Mts, Kuray Mt Rng NE Aktash vill., h=2100-3000 m, 8-13.VII.2022, E.Filippov leg./det.R Panin, L= 22 mm (alpine petrophilic population close to the form schestopalovi Plutenko, 1995)
Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus putus Motschulsky, 1846 (loc.typ.)
Male adult
Russia, W Altay, 18 km N of Zmeinogorsk, 23.VI.1998, A.& N. Menstschikovy leg. L=23 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) tarbagataicus pietrorattii Deuve, 1991(as pseudogebleri Deuve, 1994)
Male adult
Carabus (Morphocarabus) tarbagataicus pietrorattii Deuve, 1991(as pseudogebleri Deuve, 1994)
S Tarbagatai mt.rng., Alexeevka vill.,
600m, 11.5.2001 Klimenko A. leg.
male, 30 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) tarbagataicus pietrorattii Deuve, 1991
Female adult

Copyright © Ruslan Panin

Carabus (Morphocarabus) tarbagataicus birlikensis Kabak, 2013 Holotype
Male adult
China, Xinjiang, cresp of Birliktau Mt.R., WSW Toli, 45 49 23 N / 83 06 13 E, H=2565 m, 25.07.2012, Kabak I.I. leg. (ZIN)
Copyright © Ilya Kabak

Carabus (Morphocarabus) tarbagataicus var.motschulskyi Lapouge, 1909 Holotype
Male adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC525 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec525
Copyright © - - -

Carabus (Morphocarabus) monilis var.motschulskyi Lapouge, 1909
Label
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC525 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec525
Copyright © - - -

Carabus (Morphocarabus) venustus venustus Morawitz, 1862
Male adult
SE Primorskii reg., Lazovskii Nat.Res., 12 km S Staraya Kamenka vill., near Amerika kor. 43°15N, 134°1E, 19.VII.04 leg. M.& L. Smirnovy photo from www.zin.ru
Copyright © Maksim Smirnov

Carabus (Morphocarabus) venustus venustus Morawitz, 1862
Male adult
Russia, Far East, Primorskiy reg., Ussuriysk Distr., Kaymanovka vill., 6-15.VIII.2011,
A. Korshunov leg.
L= 21.5 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) venustus venustus Morawitz, 1862
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) venustus venustus Morawitz, 1862
Male adult

Copyright © Kirill Makarov

Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859
Male adult
Russia, SE Siberia, Transbaikalia, N Chita reg., Kodar Mt. Range, Big Leprindo Lake env.h~1800m., 1.VII.2006, S. Rybalkin leg. L=20 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) viridilimbatus panini I.Solodovnikov 2017 Holotype
Endophallus
Russia, SE Yakutia, Neryungri distr., Tokinski Stanovik Mt Rng., Beder riv., (right tributary rivers Algama), 26.VI.1996
Copyright © Ruslan Panin

Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859
Female adult
Russia, SE Siberia, N Chita reg., Kodar Mt. Range, Big Leprindo Lake env., h~1800 m., L=21 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) viridilimbatus panini I.Solodovnikov 2017 Holotype
Male adult
Russia, SE Yakutia, Neryungri distr., Tokinski Stanovik Mt Rng., Beder riv., (right tributary rivers Algama), 26.VI.1996, Krasnov M. leg., 17 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) viridilimbatus saluki I.Solodovnikov 2017 Paratype
Male adult
Russia, SW Taymyr, near Talnakh, 25 km N Norilsk, hilly tundra, hillside overgrown with alder slate, dwarfish birches, blueberry, h = 150-250 м, 24-28.06.2003, leg. S.Saluk
Copyright © Igor Solodovnikov

Carabus (Morphocarabus) viridilimbatus saluki I.Solodovnikov 2017 Paratype
Female adult
Russia, SW Taymyr, near Talnakh, 25 km N Norilsk, hilly tundra, hillside overgrown with alder slate, dwarfish birches, blueberry, h = 150-250 м, 24-28.06.2003, leg. S.Saluk
Copyright © Igor Solodovnikov

Carabus (Morphocarabus) viridilimbatus saluki I.Solodovnikov 2017 Paratype
Female adult
Russia, SW Taymyr, near Talnakh, 25 km N Norilsk, hilly tundra, hillside overgrown with alder slate, dwarfish birches, blueberry, h = 150-250 м, 24-28.06.2003, leg. S.Saluk
Copyright © Igor Solodovnikov

Carabus (Morphocarabus) viridilimbatus saluki I.Solodovnikov 2017 Paratype
Endophallus
Russia, SW Taymyr, near Talnakh, 25 km N Norilsk, hilly tundra, hillside overgrown with alder slate, dwarfish birches, blueberry, h = 150-250 м, 24-28.06.2003, leg. S.Saluk
Copyright © Igor Solodovnikov

Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859
Endophallus
Russia, Chita Region, Udokansky district. Kodar Range, 56 °34'42,9 " N, 118 °29'34,95" E, 6/13/2002, leg. A. Pokhorov
Copyright © Igor Solodovnikov

Carabus (Morphocarabus) wulffiusi wulffiusi Morawitz, 1862
Male+Female adult
Russia, Far East,
Primorskiy Kray, near Kamenushka vill., 2-3.VIII.1992, K.E.Dovgaylo leg.
female, 24mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) wulffiusi wulffiusi Morawitz, 1862
Female adult

Copyright © Safronov

Carabus (Morphocarabus) wulffiusi wulffiusi Morawitz, 1862
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) wulffiusi dekraatzi Kraatz, 1881
Male adult
NE-China, Heilongjiang, Wuchang city, Mt Zheng-nan-gou, 1-11.VI.2011
L=22 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) wulffiusi liaoningensis Deuve & Imura, 1990
Female adult
NE-China:E-Liaoning: NP Kuandian Tianqiaogou , VI.2010, L=24
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii zawadzkii Kraatz, 1854
Male adult
Ukrainian Carpathians
Lviv Reg., Skole Distr., near Grebeniv vill., h=1000m 9 - 20.VII.07, Panin R. leg.
male, 31 mm
Copyright © Safronov

Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
Male adult
Ukrainian Carpathians,
Transcarpathian reg., Rakhov distr., Svidovec Mt Rng., Mt. Podpula h=1500-1600m. Lazutkin G. leg, 13-14.VII.1978, det. Panin R.
L=28mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906 loc.typ.
Male+Female adult
W-UKRAINE,<br /> Transcarpathian region, Uzhgorod distr., 28.IV-23.V.2011<br /> Panin R. leg./det.<br /> female, 33 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Female adult
NW Ukrainian Carpathians
Lviv Region, Staryi Sambor Distr., near Dobromyl , h=500m 9.VI-25.VII.2007, Panin R. leg./det.Solodovnikov I. & Panin R.
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii zawadzkii, Kraatz 1854
Endophallus
Gorgany Mts. h=700 m, 16.VI.2011,
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Male adult
NE-Hungary, 4 km S Sátoraljaújhely, right bank riv. Bodrog (Long-erdő), 8.VI.1992, leg./det. I.Retezár leg. L=28 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
Female adult
W Ukraine, Transcarpathian lowland, near Chop, 11.VI-19.VII.2019, Panin R leg. L=29 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
Male adult
W Ukraine, Transcarpathian lowland, near Chop, 11.VI-19.VII.2019, Panin R leg. L=28 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Endophallus
SE Poland, Subcarpathia Prov., Nadleśnictwo Kołaczyce N of Jasło vill., M. Kosibowicz leg./det. R Panin
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Male adult
SE Poland, Subcarpathia Prov., Nadleśnictwo Kołaczyce N of Jasło vill. M. Kosibowicz leg./det. R Panin, VII.2016, L=29 mm
Copyright © Ruslan Panin

Carabus (Rhigoidocarabus) zhubajie laojunensis Kleinfeld, 1999 Holotype
Male adult

Copyright © Frank Kleinfeld