Carabus (Morphocarabus) aeruginosus zyrjanovskianus Shilenkov & O.Berlov, 1996
Male+Female adult
NE KAZAKHSTAN, S-Altays, Ulbinsky Mt. h=900m., Bobrovka vill., 24.VI.01, A. Klimenko leg. female 25 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) aeruginosus verzhutzkii O.Berlov & Shilenkov, 1996
Male adult
C Altai Mts., Seminskii Mt.rng., Katun riv., 3 km S Ust'-Sema riv.,
4.VII.2002 A. & N. Menschikovy leg.
det. Mentschikov A.
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) aeruginosus salechardensis Obydov, 1999 Paratype
Male adult
Rusia, Selehard, tundra, 30.VIII.1953, I. Telyshev leg
Copyright © Dmitri Obydov

Carabus (Morphocarabus) aeruginosus eschscholtzi Mannerheim, 1827
Male adult
Russia, W Altai, Zmeinogorsk distr., near Cherepanovskii vill., 2-16.6.1990, D. Obydov leg.
Copyright © Dmitri Obydov

Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822 (as pitschugini) Paratype
Female adult
East Siberia, Tunkinskie Goltzy (East Sajans), Mondy env., 24.VI.1979, E. Ya. Berlov leg.
Copyright © Dmitri Obydov

Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
Male+Female adult
SW Siberia, Kemerovo reg., Krapivino distr.,<br /> 8 km SSW Saltymakovo vill., h=150 m.<br /> 1-20.VII.2010 D. Efimov leg.<br /> male - 25 mm, female - 26 mm.
Copyright ©

Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
Endophallus
Russia, S Tuva, Khorumnug-Taiga Mts, Buren riv. Valley, Buren-Bay-Khaak vill. h=900 m
Copyright © Ruslan Panin

Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
Male adult
Russia, NE Altay, Turochak Distr., Lake Teletskoe N part, near Yaylyu vill. 23-25.VII.2019, Ivliev P. leg./ det. R. Panin, L=24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) aeruginosus eschscholtzi Mannerheim, 1827
Female adult
NE Kazakhstan, W Altay Mts, near Shemonaikha city., flood plain, 25-28.VII.2018, S.Vashchenko leg. L=25 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) aeruginosus eschscholtzi Mannerheim, 1827
Endophallus
NE Kazakhstan, W Altay Mts, near Shemonaikha city., flood plain,
Copyright © Ruslan Panin

Carabus (Trachycarabus) besseri Fischer, 1822
Male adult

Copyright © Safronov

Carabus (Trachycarabus) besseri Fischer, 1822
Male+Female adult
W Ukraine.NW Podolian,
Lviv reg., Zolochiv distr., Lysa Gora near Stinka vill., steppe sites, 25.IV-9.V.2009 Panin R. leg
female, 30mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) besseri Fischer, 1822
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) brinevi Kabak, 2014 Paratype
Male adult
China, Xinjiang, S Altai Mts, Keshtau Mt., ENE Qinghe, 46°47'21''N / 90°44'42''E, 2730 m, 12.08.2013, I.I. Kabak leg.
Copyright © Ilya Kabak

Carabus (Morphocarabus) brinevi Kabak, 2014 Holotype
Male adult
China, Xinjiang, S Altai Mts, Keshtau Mt., ENE Qinghe, 46°47'21''N / 90°44'42''E, 2730 m, 12.08.2013, I.I. Kabak leg.
Copyright © Ilya Kabak

Carabus (Morphocarabus) brinevi Kabak, 2014 Paratype
Endophallus

Copyright © Ilya Kabak

Carabus (Morphocarabus) brinevi Kabak, 2014 Paratype
Female adult
China, Xinjiang, S Altai Mts., Keshtau Mt., ENE of Qinghe, h~2730 49°47'21"N 90°44'42"E 12.VIII.2013 leg. I.Kabak L=19.5 mm
Copyright © Kirill Makarov

Carabus (Morphocarabus) tarbagataicus altaicola Deuve & Tian, 2019 Holotype
Female adult
Chine, Xinjiang, Préfecture d’Altaï, Monts Altaï, au nord de la ville d’Altaï, Parc forestier de Xiaodonggou (Chen Liusheng, 8.VIII. 2014), in coll. South China Agricultural University, Canton.
Copyright © Thierry Deuve

Carabus (Morphocarabus) brinevi altaicola Deuve & Tian, 2019
Female adult
«qing tong se», «28#» (ICXU)
Copyright © Ilya Kabak

Carabus (Morphocarabus) brinevi kanasensis Kabak, 2021 Holotype Paratype
Male adult
China, Xinjiang, Altai Mts, left bank of Kanas lake, 48°44ʹ35ʺN / 87°06ʹ03ʺE, H = 2710 m, alpine meadows, 15.07.2019 I.I. Kabak leg. (ZIN)
Copyright © Ilya Kabak

Carabus (Morphocarabus) chaudoiri zaikai Obydov, 1999 PT Paratype
Female adult
Carabus (Morphocarabus) chaudoiri zaikai Obydov, 1999 PT
Rusia, Tuva, Akademik Obruchev Mt. rng., 2100 m;, 52°00'N 95°34'E,
Sainak pass, tundra, 2.VII.1998, D. Obydov leg.
female,
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) chaudoiri juliae Obydov, 2002
Male adult
male, 18mm
Copyright © Boleslav Brezina

Carabus (Morphocarabus) chaudoiri czadanicus Obydov, 1997 HT Holotype
Male adult
Carabus (Morphocarabus) chaudoiri czadanicus Obydov, 1997 HT
Rusia, Tuva, near Hadyn vill., 28-30.VI.1994, R. Shmarzh leg.
male,
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) chaudoiri beladici Obydov, 2001
Male adult
S-Tuva
East Tannu-Ola,Biche-Sailig r.h=2100m, 15.VII.1996, D.E.Lomakin leg.
Copyright © Anton Menstschikov

Carabus (Morphocarabus) chaudoiri beladici Obydov, 2001 PT Paratype
Female adult
Carabus (Morphocarabus) chaudoiri beladici Obydov, 2001 PT
Rusia, Tuva, E Tannu-Ola Mt.rng., near Kara-Hol lake, 29-31.V.2000, M. Beladzhik leg
female,
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) chaudoiri marusiki Obydov, 1997 Holotype
Male adult
image from zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) chaudoiri mongolorum Csiki, 1927 Lectotype
Male adult
Muséum national d’Histoire naturelle, Paris (France); Collection : Insects: Coleoptera (EC); Specimen MNHN-EC- EC522; Credit: Antoine Mantilleri - 2010 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec522
Copyright © - - -

Carabus (Morphocarabus) chaudoiri juliae Obydov, 2002 Holotype Paratype
Male+Female adult
Russia, Tuva Republic, Mungun-Taiga, Khindiktig-Khol Lake, 2400 m, 22.-29.VI.2000, M. Beladič leg.
Copyright © - - -

Carabus (Morphocarabus) chaudoiri chaudoiri Gebler, 1847 Lectotype
Female adult
Altay
Copyright © Dmitri Obydov

Carabus (Morphocarabus) chaudoiri shilenkovi O.Berlov, 1989
Male adult
Россия, Якутия, Оймяконское нагорье, 12-18.VII.1998
Copyright © Dmitri Obydov

Carabus (Trachycarabus) estreicheri Fischer, 1822
Endophallus
Lviv Region, Brody distr., near Yasenov vill.,
Copyright © Ruslan Panin

Carabus (Trachycarabus) estreicheri Fischer, 1822
Male+Female adult
W UKRAINE, NW Podolian,
Lviv Region, Brody distr., near Yasenov vill., steppe sites, 2-28.V.2010, Panin R. leg.
female, 23mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) estreicheri Fischer, 1822
Male adult

Copyright © Safronov

Carabus (Trachycarabus) estreicheri Fischer, 1822
Female adult
W-UKRAINE, NW Podolian,
Lviv Region, Brody distr., near Yasenov vill., steppe sites, 12.V-2.VI.2007 Panin R. leg.
female, 22mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) estreicheri Fischer, 1822
Female adult
Volgograd Reg., Serafimovka Distr., Ust' -Khoperskaya Station 5.V.2009 leg.O.Sveshnikov
18.5 mm
Copyright © Kirill Makarov

Carabus (Trachycarabus) estreicheri Fischer, 1822 (as jaegeri Ménétriés, 1827)
Female adult
E UKAINE, Lugansk city, Site Luganskoe, 20-30.VII.2010, V.A.Landik leg.
L=21 mm
det.Panin.R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) estreicheri (ab.rufofemoratus Lomnicki, 1892)
Female adult
W Ukraine, NW Podolian, Lviv reg., Brody distr., Makitra Mt, steppe site, 15.VI - 24.IX.2017, Panin R. & Kanarsky Yu. leg.
Copyright © Ruslan Panin

Carabus (Trachycarabus) estreicheri Fischer, 1822 (ab.rufofemoratus Lomnicki, 1892)
Female adult
W Ukraina, Podolian Upland, Lviv reg., Brody distr., Mt. Makitra, steppe site, 19.V-5.VI.2020, Panin R. leg. L=23 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens Fabricius, 1798 (ab.erythromerus Dejean, 1826)
Male adult
Moldova, Anenii Noi distr., near Garbovets vill. 20.VI.08 Bodnar' I.I.leg. male, 26mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens Fabricius, 1798
Endophallus
Moldova,<br /> Anenii Noi distr., near Garbovets vill.
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens ( as jasilkowskii Born,1905)
Male adult
NE Romania, 5 km W Suceava, 47°63′940″N 26°18′795″E,h=340m, agricultural field, 15-20.V.2011, B. Brezina leg. L=24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Male adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 4-24.V.2010, Panin R. leg. L= 29 mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Male adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 18.V-12.VI.2014, L=28 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Male adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 27.IV-13.VI.2018, Panin R. leg. L= 28 mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Female adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 27.IV-13.VI.2018, Panin R. leg. L= 31 mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Male adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., Panin R. leg. L= 28 mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens (as lomnitzkii Reitter, 1896)
Male adult
W Ukraina, Podolian Upland, steppe reserve Kasova Gora near Burshtyn vill., 15.VI -28.VII.2020, R. Panin & D. Khrapov leg., L=24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Female adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky forest, near Pyatnychany vill., 28.V-9.VI.2013, Panin R. leg. L=30 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) gebleri ultimus Obydov, 1999
Female adult
Carabus (Morphocarabus) gebleri ultimus Obydov, 1999 PT
E Kazakhstan, Altai, near Zyrjanovska, 6.VII.1995, D. Obydov leg.
female
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) gebleri ultimus Obydov, 1999 Holotype
Male adult
Carabus (Morphocarabus) gebleri ultimus Obydov, 1999 HT
E Kazakhstan, Altai, near Zyrjanovska, 6.VII.1995, D. Obydov leg.
male
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) gebleri gebleri Fischer, 1817
Female adult
Carabus (Morphocarabus) gebleri gebleri Fischer, 1817
Russia, Altai, Zmeinogorsk
female
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) gebleri gebleri Fischer, 1817
Male adult
Carabus (Morphocarabus) gebleri gebleri Fischer, 1817
Rusia, Altai, Zmeinogorsk
male
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) helleri Ganglbauer, 1893
Male adult
Moravia Mer., Bile Karpaty 28.5.1993 Budka leg. male, 28 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) helleri Ganglbauer, 1893
Male+Female adult
SE Moravia, Strani, V.2002, M.Snizek leg. male bronze colour-26mm male blue colour-28mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) helleri Ganglbauer, 1893
Endophallus
SE Moravia, Strani, V.2002, M.Snizek leg. male bronze colour.
Copyright © Ruslan Panin

Carabus (Morphocarabus) helleri Ganglbauer, 1893
Endophallus
SE Moravia, Strani, V.2002, M.Snizek leg. male blue colour
Copyright © Ruslan Panin

Carabus (Morphocarabus) helleri Ganglbauer, 1893 (as pseudopreyssleri Breuning, 1932)
Male adult
Czech Rep., NE Moravia, Moravian-Silesian Beskids Mt Rng, PR Trojacka, Hodslavice vill., 29.V.12.VI.2010, Krause Rene leg./det.Panin R. L=30 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) helleri Ganglbauer, 1893 (as pseudopreyssleri Breuning, 1932)
Endophallus
Czech Rep., NE Moravia, Moravian-Silesian Beskids Mt Rng, PR Trojacka, Hodslavice vill.,
Copyright © Ruslan Panin

Carabus (Morphocarabus) helleri Ganglbauer, 1893
Male+Female adult
CZECH, SE Moravia, Velka nad Velickou. 20-V-2005, V. Slovak leg. male - 26 mm, female - 30 mm.
Copyright © Alexandr Safronov

Carabus (Morphocarabus) helleri Ganglbauer, 1893
Male adult
CZ; CE Moravia, Bílé Karpaty Mts.; 24mm
Copyright © Aleš Sedláček

Carabus (Morphocarabus) helleri Ganglbauer, 1893
Male adult
CZ; CE Moravia, Bílé Karpaty Mts.; 25mm
Copyright © Aleš Sedláček

Carabus (Morphocarabus) henningi pelevini Obydov 2006 HT
Male adult
Carabus (Morphocarabus) henningi pelevini Obydov 2006 HT
Kazakhstan, S Altai, Sarym-Sakty Mt.rng., 3000-3400 m,
18-20.VI.1998, A. Klimenko leg.
Copyright © Dmitri Obydov

Carabus (Morphocarabus) henningi peczoricus Obydov 2000 HT Holotype
Male adult
Carabus (Morphocarabus) henningi peczoricus Obydov 2000 HT
Rusia, Komi republic, Kosiyu River mouth, (Pechora basin) 21.VII.1980
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) henningi oviformis Beheim & Breuning, 1943
Male adult
Carabus (Morphocarabus) henningi oviformis Beheim & Breuning, 1943
Rusia, Krasnoyarsk reg., Taimyr AO, near Norilsk,
07.2004, M. Shestopalov leg.
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) henningi henningi Fischer, 1817
Male adult
S. Ural, Bashkiria, near Mesjagutovo vill., 09.07.2001 A. Menstschicov leg.
Copyright © Anton Menstschikov

Carabus (Morphocarabus) henningi melyachi Shilenkov & Menstshikov, 1996 loc.typ.
Male adult
Russia, Altay, Katun Mt Range, Karakol lake,22.VI.1993, Melyach F.V. leg.<br /> L=18 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) henningi henningi Fischer, 1817
Male adult

Copyright © Kirill Makarov

Carabus (Morphocarabus) henningi sahlbergi Dejean, 1826 Lectotype
Male adult
"Siberie, Slataoust" = [Златоуст, Zlatoust, Cheliabinsk prov.]. Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC1 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec1
Copyright © - - -

Carabus (Morphocarabus) henningi cf saldaitisi Obydov 1997
Male adult
Russia, S Tuva, E Tanu-Ola Mts., Cha-Ova-Art Pass, h=1700m, 28.VI-3.VII.2000, Vashchenko S. leg. L=19.5 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) henningi pelevini Obydov 2006
Female adult
E Kazakhstan, S Altay, Sarym-Sakty Mt Range, 12 km SE Ornek vill. h=1200m, 10.VII.1998, L=21 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) henningi henningi Fischer, 1817
Male adult
Russia, S Siberia, NW Buryatia, E Sayans Mts., Tunka Range, near Mondy vill., h=2400 m, alpine tundra, 1-15.VI.2001, Mishustin R. leg./det. R.Panin
Copyright © Ruslan Panin

Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 18789 (as viridipennis Kraatz, 1886, nec Lewis,
Male adult
E Rusia, Khabarovsk reg.,
near Birobidzhan VI.1991
male, 26 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) hummeli cf pusongensis Imura, 1993
Male adult
China, Beijing, Wulingshan mountain, 1985
Copyright © Ye Liu

Carabus (Morphocarabus) hummeli nevelskii Shilenkov, 1996
Female adult
Rusia, S Sakhalin isl., VII.1993, G. Vasiliev leg.

Copyright © Dmitri Obydov

Carabus (Morphocarabus) hummeli hummeli Fischer, 1823
Male adult
Carabus (Morphocarabus) hummeli hummeli Fischer, 1823
Rusia, Chita reg., Darasun, VII.1991
I. Avdeev leg.
male, 23 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 18789 (as viridipennis Kraatz, 1886, nec Lewis,
Female adult
Primorskii reg., near Kovalerovo
VII.1990 Evstigneev A. leg.
female, 25 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) hummeli ? middendorfi Menetries, 1851
Male adult
Carabus (Morphocarabus) hummeli ? middendorfi Menetries, 1851
Rusia, Khabarovsk reg., Ohotsk sea, Port Ayan, 08.24
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878 (as viridipennis Kraatz, 1886, nec Lewis,18
Male+Female adult
Russia, Far East,Primorsky reg., S. Sikhote-Alin Mts. R., Kavalerovo vill. env.,
5-25.VII.1992, С. Kunitzyn leg.
male - 25 mm, female - 25 mm.
Copyright © Alexandr Safronov

Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878 (as purpuripennis Kraatz, 1886)
Male adult
Russia, Khabarovsk reg., near Komsomolsk-on-Amur.
7-21.VI.2010, A. Kapkaev leg.
Male left - 24 mm, male right - 23 mm.
Copyright © Alexandr Safronov

Carabus (Morphocarabus) hummeli stolidus Lapouge, 1924
Label
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC519
Copyright © - - -

Carabus (Morphocarabus) hummeli hummeli Fischer, 1823
Male adult
E Siberia, Sokhondinskii NP, Ingoda riv., 49.50.207N 111.10.825E, 1700m, 23.7.2009 Shavrin A. leg.
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) illigeri illigeri (as bjelasnicensis Apfelbeck, 1902)
Male+Female adult
Serbia, Debelo Brdo, 5.VI.2007, S. Sapaly leg. L=32 mm photo Panin R.
Copyright © Ruslan Panin

Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Endophallus
NE-Serbia, Krs Mts., Majdanpek env.,h=300m.20-31.V.2006, Pr.Sima leg
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens trans illigeri Dejean, 1826
Male+Female adult
W-Croatia, Karlovac, near Bosiljevo vill., h~300m, V.2000, I.Rapuzzi leg. female, 33 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens trans illigeri Dejean, 1826
Endophallus
W-Croatia, Karlovac, near Bosiljevo vill., h~300m,
Copyright © Ruslan Panin

Carabus (Morphocarabus) illigeri versicolor Frivaldsky, 1835
Male+Female adult
C Bulgaria:Stara planina Mts, Gabrovo,12-15.VI.2007, M. Janalik leg. female, 31 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) illigeri versicolor Frivaldsky, 1835 (ab.)
Female adult
C Bulgaria: Stara Planina Mts, Schipka pass h=1450m, 12-15.VI.2007, M.Janalik leg. L=31 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) illigeri illigeri (as tarensis Toševski, 1990)
Male adult
WC-Serbia: National park Tara Mts.:NE Mitrovac h=1500m, 1-12.VII.2007, Pr.Sima leg. L=30 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) illigeri illigeri (as tarensis Toševski, 1990)
Endophallus
WC-Serbia: National park Tara Mts.:NE Mitrovac h=1500m,
Copyright © Ruslan Panin

Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Female adult
NE Serbia, Kučevo; female, 33mm
Copyright © Aleš Sedláček

Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Female adult
NE Serbia, Majdanpek; 35mm
Copyright © Aleš Sedláček

Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
Larva

Copyright © Evgeniy Chibilov

Endofallus Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
Male adult
Russia,
S Ural Mts, Mt.Iremel h=800m, 4.VI.2000 Filimonov V. leg.
male,21mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
Male adult
Carabus (Morphocarabus) karpinskii Rusia, S Ural mts., Jremel Mt., 1300 m 4.6.2000male, 20 mm
19.08.2008
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) karpinskii Kryzhanovskij & Matveev, 1993
Male adult

Copyright © Kirill Makarov

Carabus (Morphocarabus) kollari Palliardi, 1825
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) kollari Palliardi, 1825
Female adult
female, 24 mm
Copyright © Boleslav Brezina

Carabus (Morphocarabus) kollari Palliardi, 1825
Male adult
male, 27 mm
Copyright © Boleslav Brezina

Carabus (Morphocarabus) kollari Palliardi, 1825
Male+Female adult
Romania, SW Carpathians,
Banat, Anina Mts. (Muntii Aninei) 8 km NE Oravita, 7.VI.2007, S.I. Budka leg.
female, 34 mm
photo Panin R
Copyright © Ruslan Panin

Carabus (Morphocarabus) kollari Palliardi, 1825 (as semetricus Kraatz, 1878)
Male adult
SW-Romania:Banat, Bocsa/Coltan, 21.V-5.VI.2010, Krause Rene leg.
L=29 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) kollari Palliardi, 1825 (as semetricus Kraatz, 1878)
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) kozhantschikovi martjanovi Lutshnik, 1924
Female adult
Khakassia, 1800 m, near Cheremushki vill., 12.6.-20.7.2000<br /> female, 21 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
Male adult
W Sayan Mts., Kazhyr-Suk riv., 80 km W Oleniya<br /> Rechka, W part of Oiskii mt. rng., 30.6.-7.7.1999<br /> 2100 m, Brinev A. leg.<br /> male, 19 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
Male adult
W Sayan, low fl. Kazyr-Suk riv., 80km W meteostation Olenia Rechka, W part of Oiskii mt.rng., 30.VI-7.VII.1999 2100m, Brinev A. leg.<br /> male, 20mm<br /> det. Brinev A.<br /> photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
Endophallus

Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
Male adult
Russia, W Sayan Mts., Kazyr-Suk river, W slope of Oyskiy Mt. rng., ?h=2200 m 7-15.VII.1999, Brinev A. leg., L=19 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
Male adult
Burjatia, Dzhida distr., near Beloozersk vill. 4-9.VII.1997 Anichtchenko A. leg.
male, 23 mm
Copyright © Alexander Anichtchenko

Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
Male adult
Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
China, Beijing, Mentougou, 2005;
Photo & det. by liuye

Copyright © Ye Liu

Carabus (Trachycarabus) latreillei ?semicoriaceus Kraatz, 1881
Endophallus
NE-China: Liaodong Peninsula, Liaoning prov., Gaizhou distr., Mt. Buyun Shan.
Copyright © Ruslan Panin

Carabus (Trachycarabus) latreillei ? semicoriaceus Kraatz, 1881
Male adult
NE-China: Liaodong Peninsula, Liaoning prov., Gaizhou distr., Mt. Buyun Shan,VI.2007, Hao leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
Endophallus
N Mongolia, Khövsgöl reg., Lake Khovsgol, h=1640m, 51*:12’N; 100*:45’E, steppificated sites,
Copyright © Ruslan Panin

Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
Male adult
N Mongolia, Khövsgöl reg., Lake Khovsgol, h=1640m, 51*:12’N; 100*:45’E, steppificated sites,
20-25.VII.2010. A.Dergalev leg.
L=22 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) latreillei mengguicus Rapuzzi, 2023 Holotype
Male adult
China, Inner Mongolia Province, Tongliao, Ka’erqin, Jinbaotun, VIII.20. Image from Rapuzzi, 2023
Copyright © - - -

Carabus (Trachycarabus) latreillei shanxiensis Rapuzzi, 2024 Holotype
Male adult
China, Shanxi, Lu Liang, Kang Cheng Zhen, Jiao Kou Xian; Aug. 2022. Image from Rapuzzi, 2024
Copyright © - - -

Carabus (Trachycarabus) mandibularis mandibularis Fischer, 1827
Male adult
NE Kazakhstan, S Altai Mt., Narymskii mt. rng.
Bolshenarymskoe vill. 26.7.2001, 1800m Klimenko leg.
male, 21 mm
Copyright © Alexander Anichtchenko

Carabus (Trachycarabus) mandibularis buchtarmensis Kryzhanovskij, 1953
Male adult
Kazakhstan, Kalbinsky mt. rng., 600m ,
Ognevka vill., 23.6.98 Klimenko A. leg.
male, 23 mm
Copyright © Alexander Anichtchenko

Carabus (Trachycarabus) mandibularis buchtarmensis Kryzhanovskij, 1953
Male adult
NE-Kazakhstan,
S-Altays, Kalbinky Mt. h=500m., near Ognevka vill., 20.VI.1998.A. Klimenko leg.
male, 23mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) mandibularis buchtarmensis Kryzhanovskij, 1953
Endophallus

Copyright © Ruslan Panin

Carabus (Trachycarabus) mandibularis abakkereiorum Kabak & Hu, 2017 Holotype Paratype
Male+Female adult
(ZIN). China, Xinjiang, Altay Mt. R., Kran Valley, N of Altay Town, 47°55´37''N / 88°08´23´´E, h = 1015 m, 31.VII.2015 I.I. Kabak leg.
Copyright © Ilya Kabak

Carabus (Trachycarabus) mandibularis buchtarmensis Kryzhanovskij, 1953 Paralectotypes
Male genitalia
Aedeagus: from Tshernovaya; Endofallus: from Bolshenarymskoe.
Copyright © Ilya Kabak

Carabus (Trachycarabus) mandibularis abakkereiorum Kabak & Hu, 2017
Male genitalia

Copyright © Ilya Kabak

Carabus (Morphocarabus) mestscherjakovi telezkianus Shilenkov, 1996
Male adult
Carabus (Morphocarabus) mestscherjakovi telezkianus Shilenkov, 1996
Rusia, Altai, S shore of Teletskoe lake, 5.VII.1968
Copyright © Dmitri Obydov

Carabus (Morphocarabus) mestscherjakovi telezkianus Shilenkov, 1996
Male adult
Carabus (Morphocarabus) mestscherjakovi telezkianus Shilenkov, 1996
SW Khakasia, Abakanskii Mt.rng., Choochek Mt., 20 km SSE Mrassu vill.,
1800m, 7-19.7.1999 Lomakin D. leg.
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
Male adult
W Sayan mts., Kopen mt., Dzhabash riv., 75 km
S from Arbaty vill., 2235m, 26-30.6.2003 Brinev A. leg.
long. 17 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
Male+Female adult
Russia, S Siberia,
S Khakassia, N Tuva reg., Sailyg-Khem-Taiga mts. range h=2000-2600m,10-30.VI.2005, S.Vashchenko & S.Alexeenko leg.
male,17.5mm/female,19mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
Endophallus
endophallus
Copyright © Ruslan Panin

Carabus (Morphocarabus) mestscherjakovi pseudoodoratus Brinev, 2002 Paratype Paratype
Female adult
Krasnojarsk reg., Aradan distr., 5 km SW Aradan vill., vall. of Us riv., A.Brinev leg.<br /> female, 20 mm<br /> photo & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
Male adult
Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
W Sayan, Kulumys Mt.R., env. Oiskoje Lake, mountain tundra, 12-14. 07.1999 A.Semin leg.
male
L=17,0 mm
Copyright © Maksim Smirnov

Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
Male+Female adult
Russia, S Siberia, W Sayan, Ergaki Mt range, Nizhnyaya Buyba river h=1700 m, 19.VII.1999, A.& N. Menstschikovy leg.
Copyright © Ruslan Panin

Carabus (Morphocarabus) mestscherjakovi zherichini Shilenkov, 1990
Label

Copyright © Roman Dudko

Carabus (Morphocarabus) mestscherjakovi tuvinicus Obydov, 2002 Holotype
Female adult
Россия, Тува, хр. Таскыл, долина р. Уюк, 4.VI-8VI.1999, И. Белло leg.
Copyright © Dmitri Obydov

Carabus (Morphocarabus) michailovi Kabak, 1992
Male adult
NE Kazakhstan, S Altay Mts., Sarymsakty mt. rng.,
Sarymsakty riv., 2400 m, 22.6.2000 Nekroshevich S. leg.
male, 18 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) michailovi Kabak, 1992
Male adult
NE Kazakhstan, S Altay Mts., Sarymsakty mt. rng.,
Sarymsakty riv., 2400 m, 22.6.2000 Nekroshevich S. leg.
male, 17 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) michailovi Kabak, 1992
Male adult
NE-Kazakhstan,
S Altai Mts.,Sarym-Sakty Range, Sarym-Sakty riv., h=2400m. 21.VI.1999., A.Klimenko leg.
male,18mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) monilis Fabricius, 1792
Male adult
France 39, Foncina le Haut 300m, 23.6.1979; 29 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) monilis Fabricius, 1792 (as taunicus)
Male adult
W Germany, Vogelsberg Mt., 700m, 13.5.1985
male, 24 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) monilis Fabricius, 1792 (as interpositus Géhin, 1880)
Male adult
NW France,
Basse-Normandie, Orne, Forкt d'Йcouves, V.2009, Pascal Stefani leg.
male, 29mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) monilis Fabricius, 1792 (as interpositus Géhin, 1880)
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) monilis (as amoenus Baudet-Lafarge, 1836)
Female adult
C France,
Puy-de-Dome, Col de la Croix Saint Robert h=1300m, VII.2008, Pascal Stefani leg.
female, 24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) monilis (as helveticus Heer, 1837 Holotype) Holotype
Male adult
Carabus (Morphocarabus) monilis (as helveticus Heer, 1837 Holotype)
?hybrid of monilis/cancellatus
Museum of Zurich
photo Anichtchenko
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) monilis (as dupeuxi Deuve, 1974)
Male+Female adult

France,
Charente, Foret de Braconne VII.1998, Laforgue leg.
famale, 27mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) monilis Fabricius, 1792
Male adult
24 mm, England, Warwickshire
leg.det. D.J.Mann, 15.IV.1993
Photo: U.Schmidt, 2011
Copyright © Udo Schmidt

Carabus (Morphocarabus) monilis Fabricius, 1792
Female adult
France, Dröme, Ft. de Saou, 17.vii.1975-M. Grotz Leg.-Coll.RBINS, N. Mal as monilis saouensis nicodi
Copyright © Noël Mal

Carabus (Morphocarabus) monilis tanarguensis Maguerre & Stefani, 2021 (syn.) Holotype Paratype
Male+Female adult
Holotype, m (21 mm), environs du Col de la Croix de Bauzon, 1300 m, communes de La Souche, Borne et Mayres, Ardèche, France (07/2003), in coll. D. Maguerre, à Mons-en-Barœul, France. Paratype f (23 mm), au dessus du Col de Meyrand, 1400 mètres, communes de Loubaresse et de Borne, Ardèche, France, (07/2004), in coll. P. Stéfani.
Copyright © - - -

Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844: 100 Lectotype
Male adult
Carabus (Morphocarabus) odoratus odoratus LT<br /> Lectotype<br /> Hamar-Gaban Mts.<br /> photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844 (as melleus Lapouge, 1909)
Male+Female adult
Russia, S Siberia, East Sayans, Mondy circ., h~2300m., 5-8.VI.2001, R. Mishustin leg. 20mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) odoratus kamtchatensis Breuning 1942
Male adult
Carabus (Morphocarabus) odoratus kamtchatensis Breuning 1942
Rusia, Kamchatka peninsula, VII.21
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) odoratus melleus Lapouge, 1909 Paralectotypes
Male adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC521
Copyright © MNHN

Carabus (Morphocarabus) odoratus kurmaevi I.Solodovnikov, 2017 Paratype
Female adult
Russia, Siberia, Krasnoyarsk Krai, the North Yenisei district, 55 km of CE of the vill. Bryanka, valley of the river Chirimba, stream Nakhodny, h = 350 m, the mixed forest (birch and pine); on a steep slope along an animal track on a hill brow, 23.VII-20.VIII.2015, leg. A.V. Kurmaev
Copyright © Igor Solodovnikov

Carabus (Morphocarabus) odoratus kurmaevi I.Solodovnikov, 2017 Holotype
Endophallus
Russia, Siberia, Krasnoyarsk Krai, the North Yenisei district, 55 km of CE of the vill. Bryanka, valley of the river Chirimba, stream Nakhodny, h = 350 m, the mixed forest (birch and pine); on a steep slope along an animal track on a hill brow, 23.VII-20.VIII.2015, leg. A.V. Kurmaev
Copyright © Igor Solodovnikov

Carabus (Morphocarabus) odoratus chlebnikovae Obydov 2005 Paratype
Female adult
Россия, Северный Байкал, 80 км Ю Северобайкальска, 5 км Ю истока р. Куркула, 19.VII.2003, М. Хлебникова leg.
Copyright © Dmitri Obydov

Carabus (Morphocarabus) odoratus jacuticus Obydov, 2002 Paratype
Male adult
Россия, Якутия, В. Хандыга, VII.1993, А. Наглис leg.
Copyright © Dmitri Obydov

Carabus (Morphocarabus) odoratus antropovi Shilenkov 1996
Female adult
Russia, S Siberia, Tuva, W Tannu-Ola Mts. 35 km NE Khandagayty vill., Teeli Pass 2200-2800 m, 23-25.VI.2021, A.& E.Filippovs leg. L=19 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) odoratus antropovi Shilenkov 1996
Endophallus
Russia, S Siberia, Tuva, W Tannu-Ola Mts. 35 km NE Khandagayty vill., Teeli Pass 2200-2800 m
Copyright © Ruslan Panin

Carabus (Trachycarabus) planarius Obydov, 1994 Paratype
Female adult
Казахстан, Семипалатинский р-н, окр. пос. Шолпан, 500 м, 20 км. южнее пос. Таскескен, 2.06.1993, М. Данилевский leg.
Copyright © Dmitri Obydov

Carabus (Trachycarabus) planus planus Gehin, 1885
Male+Female adult
S Russia, NW Caucasus,<br /> Karacaj-Cherkessija, near Kart-Djurt vill., upper Kylan riv. h=1300m, 26-27.V.2009 , Fominykh D. leg.<br /> L=24mm<br /> photo/det. Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus planus Gehin, 1885
Male adult
NW Caucasus, Rostov reg.,<br /> Avilo-Uspenskaya vill. 22.5.2000 Arzanov Yu. leg.<br /> male, 25 mm
Copyright © Alexander Anichtchenko

Carabus (Trachycarabus) planus planus Gehin, 1885
Male adult
S Russia,<br /> Krasnodar reg., Anapa distr., B. Utrish vill., env., 11-20.VIII.2008, Khatchikov E. leg.<br /> male, 26mm<br /> photo Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus planus Gehin, 1885
Endophallus
<b>endophalus </b> S Russia,Krasnodar reg., Anapa distr., B. Utrish vill., env.,
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus planus Gehin, 1885
Endophallus
<b>endophalus</b> Karacaj-Cherkessija, near Kart-Djurt vill., upper Kylan riv. h=1300m
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus campicola Reitter, 1896
Male adult
Krym Peninsula, near Sudak. Leg.Vashchenko S. 1-5.V.2004<br /> 27 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus campicola Reitter, 1896
Male+Female adult
S Ukraine, Krym Peninsula,<br /> near Perevalnoe vill., in steppe, 15.IV.2009, Shaporinskiy V. leg.<br /> female, 28 mm<br /> photo/det.Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus campicola Reitter, 1896
Endophallus

Copyright © Ruslan Panin

Carabus (Trachycarabus) planus retowskianus Mandl, 1955
Male adult
S Ukraine, Zaporizhia reg., Melitopol distr., near Troitskoe vill., 1.VI.2010, I.V. Kovalev leg., L=27 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus campicola Dejean, 1831
Female adult
S Ukraine, Crimea, S slope Mt Demerdzhi h=650 m, sibljak, VII.2004, R. Panin leg., L=29 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens trans scheidleri
Endophallus
NC Croatia, Ivanščica Mts , Ivanec,4-18.VI.2011, Krause Rene leg.
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens trans scheidleri
Male adult
NC Croatia, Ivanščica Mts , Ivanec,4-18.VI.2011, Krause Rene leg.<br /> L=31 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens praecellens Pillardi, 1825
Endophallus
Serbia, Fruska Gora Mts, 20-31.V.2006, M.Sima leg.
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens praecellens Pillardi, 1825
Male adult
Serbia, Fruska Gora Mts, 20-31.V.2006, M.Sima leg.
L=29 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens jucundus Csiki, 1906
Male adult
N Hungary, Pilis Mts., Piliszentlelek, 20-27.VII.2011, Krause Rene leg./det. R.Panin
L=30 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens jucundus Csiki, 1906
Endophallus
N Hungary, Pilis Mts., Piliszentlelek,20-27.VII.2011, Krause Rene leg.
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as pseudojucundus Retezár, 1974)
Male adult
NW-Hungary::Komarom distr., Tardos, Gerecse Mts., 20.-27.V.2011, Krause Rene leg.
L=26 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as pseudojucundus Retezár, 1974)
Endophallus
NW-Hungary::Komarom distr., Tardos, Gerecse Mts.
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as vertesensis Retezár, 1974)
Male adult
NW Hungary: Fejer: Vértes Mt Rng, Tatabanya, 28.VII-2.VIII.1996
L=29 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as vertesensis Retezár, 1974)
Endophallus
NW Hungary: Fejer: Vertes Mt Rng, Tatabanya
Copyright © Ruslan Panin

Carabus (Morphocarabus) regalis regalis Fischer, 1822
Male adult
Carabus (Morphocarabus) regalis regalis Fischer, 1822
S. Ural, Bashkiria, near Mesjagutovo vill., 09.07.2001 A. Menstschicov leg.
photo A. Menstschicov
Copyright © Anton Menstschikov

Carabus (Morphocarabus) regalis jenissoni Dejean, 1831
Male adult
Carabus (Morphocarabus) regalis jenissoni Dejean, 1831
Rusia, Tuva, Todzh, Azas lake, 990 m,
52°24'N 96°33'E, 8.VII.1998, D. Obydov leg.

Copyright © Dmitri Obydov

Carabus (Morphocarabus) regalis jenissoni Dejean, 1831
Female adult
Carabus (Morphocarabus) regalis jenissoni Dejean, 1831
SW Siberia, Altays Mts, Choya distr., Sukhoy Karasuk env.
23.07.2004, A.Safronov leg.
Copyright © Safronov

Carabus (Morphocarabus) regalis eltoni Obydov, 2006 Holotype
Male adult
Carabus (Morphocarabus) regalis eltoni Obydov, 2006 HT
Rusia, Tver distr., near Ilyinskoe vill.,
29-30.VII.2006, D. Obydov leg.
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) regalis tschusovae Obydov, 2004
Male+Female adult
NE-Kazakhstan, W Altay, Ivanovsky Mt.Rng, near Ridder (Leninogorsk), h-1300-1500m, 4-6.VII.2003
female, 25 mm
det.Panin R.
Copyright © Ruslan Panin

Carabus (Morphocarabus) regalis tschusovae Obydov, 2004
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) regalis jenissoni Dejean, 1831 (as cyanescens Kraatz, 1882, nec Sturm, 1815)
Male+Female adult
Kazakhstan, E. Kazakhstan area,
Zyryanovsk env., near Putintzevo vill.
10-VI-2005 M. Danilevsky leg.
male - 24 mm, female - 25 mm.
Copyright © Alexandr Safronov

Carabus (Morphocarabus) rothi hampei Kuster, 1846
Male adult
W.Ukraine.
Transcarpathian region,Vinogradov distr., near Malaya Kopanya vill., Chorna Gora Mt., h=180m. 7-18.VII.08. Panin R. leg.
male, 28 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei Kuster, 1846
Female adult
Carabus (Morphocarabus) rothi hampei
W Ukraine, Zakarpatian reg., Vinogradov
1.7.1989 Avdeev I. leg.
female, 31 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) rothi hampei Kuster, 1846 (transition to rothi incompsus Kraatz, 1880)
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei Kuster, 1846 (as dacicus Csiki, 1906)
Female adult
NW Romania , Bistrita Nasaud, Beclean env., 17.VI.2009, S.Budka leg.
female, 31 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei (as zilahiensis Csiki, 1906)
Male+Female adult
NW Romania, Zalău env., 31.VII.2003, Sl.Budka leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi comptus (as szoerenyensis Csiki, 1908)
Male adult
SW Romania, Tarcu Mts, vf. Nedeia h=2000m, 12.VI.2008, J.Kynkor leg.
L=22 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi alutensis Savulescu, 1972
Endophallus
C Romania, S Carpathians, Rimnicu Vilcea env., Calimaneşti, Cozia Mts.,
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei (as diffinis Csiki, 1906)
Endophallus
NW Romania, Apuseni Mts, Trascău Mt Rng, Petresti de Jos, 12 km E Turda,.
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi distinguendus (as bieneri Mandl, 1965)
Endophallus
C-Slovakia, Detva Distr., Stara Huta env. h=700-800m,
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi distinguendus (as bieneri Mandl, 1965)
Male adult
C-Slovakia, Detva Distr., Stara Huta env. h=700-800m, 3.VI.-5.VII.2012, V.Slovák lgt. L=29 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) scabriusculus scabriusculus (as erythropus Fischer, 1822)
Male+Female adult
W.Ukraine,<br /> NW Podolian Upland , Lviv Reg., Zolochiv distr., near Luky vill., steppe sites, 7.V-8.VI. 2010<br /> leg. Panin R.<br /> female, 22mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) scabriusculus scabriusculus (as erythropus Fischer, 1822)
Male adult
W-UKRAINE<br /> NW Podolian Upland, Lviv Reg., Zolochiv Distr., near Luka vill., in steppe. Panin R. leg. 1.V - 16.VI.07
Copyright © Safronov

Carabus (Trachycarabus) scabriusculus scabriusculus Olivier, 1795
Endophallus

Copyright © Ruslan Panin

Carabus (Trachycarabus) scabriusculus inapertus Motschulsky, 1850 (loc.typ)
Male+Female adult
NE UKAINE,
Kharkov reg., Zmiev distr., in steppe, 15.VI-14.VII.2009, Kovalchuk D. leg.
female, 20mm
photo/det. Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) scabriusculus inapertus Motschulsky, 1850 (loc.typ)
Endophallus

Copyright © Ruslan Panin

Carabus (Trachycarabus) scabriusculus lippii Dejean, 1826
Female adult
SW Ukraine, Odessa reg., Ivanovka distr., steppe ravine near Gudevichevo vill., 22.IV-5.V.2022, A.Gontarenko leg., L=22 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) scabriusculus lippii Dejean, 1826
Male adult
SW Ukraine, Odessa reg., Ivanovka distr., steppe ravine near Gudevichevo vill., 22.IV-5.V.2022, A.Gontarenko leg., L=21.5 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) scabriusculus inapertus Motschulsky, 1850
Male adult
NC Ukraine, Kiev city, Lysa Gora, 1-15.VI.2012, Gerasimov R.leg./det. R.Panin, L=19 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
Female adult
Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
Moravia mer.,Vranov, 26-29.06.1979, Dr. Resl leg.
photo A. Menstschicov
Copyright © Anton Menstschikov

Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
Male adult
Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
SE Moravia mer.,Vranov, 26-29.06.1979, Dr. Resl leg.
photo A. Menstschicov
Copyright © Anton Menstschikov

Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
Male adult
Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
S.Bohemia,
Hluboka nad Vltavou, 24.VI-18.VII.1990, S. Budka leg.
male, 27mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799 (as scheidleri floriani)
Male adult
Carabus (Morphocarabus) scheidleri scheidleri (as scheidleri floriani)
A.: Steiermark, Rein Umg. VIII.1983 Kenyery
female, 32 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) scheidleri Panzer, 1799
Endophallus
W-Austria, Wienerwald, Hochstraß am Lengbachl, h=575 m,
Copyright © Ruslan Panin

Carabus (Morphocarabus) scheidleri Panzer, 1799
Male adult
W-Austria, Wienerwald, Hochstraß am Lengbachl, h=575 m, 25.VI-2.VII.2024, M. Fuchs leg. L= 30 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886
Male+Female adult
W Ukraine,<br /> Ternopol reg., Pidvolochisk distr., Supranivka vill.<br /> 3.VII.1908. Leg. Stobetskii. S.<br /> Photo R. Panin
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886
Endophallus

Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886
Endophallus
aedeagus
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus obliteratus Fischer, 1827
Endophallus
endophallus sibiricus obliteratus Fischer, 1827
NE Kazakhstan,
S Altai mts.,S Ulbinsky mt.h=500m.,Turgusun vill., 20.VI.2001
male 25mm.
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus kolosovi Zinoviev, 1997 loc.typ.
Female adult
S. Ural, Bashkiria, near Mesjagutovo vill., 09.07.2001 A. Menstschicov leg.
photo A. Menstschicov
Copyright © Anton Menstschikov

Carabus (Trachycarabus) sibiricus kolosovi Zinoviev, 1997
Female adult
Russia, North-East Bashkortostan. Ai river, near Sikiyaz-Tamak vill.
12.07.1997, Zinoviev E.leg
length: 21,8 mm
Copyright © Evgeniy Zinoviev

Carabus (Trachycarabus) sibiricus haeres (as fossulatus Dejean, 1826)
Male+Female adult
E.UKRAINE,<br /> Kharkov reg., Zmiev distr.,in steppe,10-20.V I.2007, D.Kovalchuk leg.<br /> female, 26mm/male, 24mm.<br /> photo/det. Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus haeres (as fossulatus Dejean, 1826)
Endophallus
endophallus
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus gvozdevae Obydov, 1997
Male adult
Russia, Khakassia,
W Sayan, Enisey riv., Batenevskiy Mt Range, 12 km E Pervomayskiy vill., 11.VII.2000, A.& N. Menstschicovy leg.
L=21 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus seriatoporus Fischer von Waldheim, 1828
Male adult
окр. Якутска, пос. Бестях на р. Лена, 30.VI.1962, Сладков leg.
Copyright © Dmitri Obydov

Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997 HT Holotype
Male adult
Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997 HT
Rusia, SE Tuva, Sangilen mt.rng., Kargy riv., 50°31'N 97°01'E,
1800 m, 29.07.1996, D. Obydov leg.

Copyright © Dmitri Obydov

Carabus (Morphocarabus) spasskianus spasskianus Breuning, 1932
Male adult
Siberia, Lake Baikal, Listvianka vill. 25.8.90
Anichtchenko A. leg.
male, 23 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) spasskianus spasskianus Breuning, 1932
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus schestopalovi Plutenko, 1995
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus aeruginosiformis Breuning, 1932
Male adult
Russia, Khakassia,
W-Sayan, Alan Mt range, valley Enisey riv., 5 km S Mayna vill., 15.VII.2000, A.& N. Menstschicovy leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus aeruginosiformis Breuning, 1932
Endophallus
Russia, Khakassia,
W-Sayan, Alan Mt range, valley Enisey riv., 5 km S Mayna vill.,
Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus shoriensis Obydov, 1999
Male adult
Russia, Khakassia,
Abakanskii Mt Range, Askiz river vall., 7 km E Kazanovka vill., 15.VI.2006, A.& N. Menstschikovy leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus aeruginosiformis Breuning, 1932
Male+Female adult
Russia, S. Siberia, Krasnoyarsk reg., W. Sayan,<br /> Sayanogorsk distr., Cheremushki vill. env.,<br /> 07-VII-1994 М. Danilevsky leg.<br /> male - 22 mm, female - 25 mm.
Copyright ©

Carabus (Morphocarabus) spasskianus cf putus Motschulsky, 1846
Male adult
Russia, SW Siberia, C-Altays Mts, Kuray Mt Rng NE Aktash vill., h=2100-3000 m, 8-13.VII.2022, E.Filippov leg./det.R Panin, L= 23 mm (alpine petrophilic population close to the form schestopalovi Plutenko, 1995)
Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus putus Motschulsky, 1846 (loc.typ.)
Male adult
Russia, W Altay, 18 km N of Zmeinogorsk, 23.VI.1998, A.& N. Menstschikovy leg. L=23 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) tarbagataicus pietrorattii Deuve, 1991(as pseudogebleri Deuve, 1994)
Male adult
Carabus (Morphocarabus) tarbagataicus pietrorattii Deuve, 1991(as pseudogebleri Deuve, 1994)
S Tarbagatai mt.rng., Alexeevka vill.,
600m, 11.5.2001 Klimenko A. leg.
male, 30 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) tarbagataicus pietrorattii Deuve, 1991
Female adult

Copyright © Ruslan Panin

Carabus (Morphocarabus) tarbagataicus birlikensis Kabak, 2013 Holotype
Male adult
China, Xinjiang, cresp of Birliktau Mt.R., WSW Toli, 45 49 23 N / 83 06 13 E, H=2565 m, 25.07.2012, Kabak I.I. leg. (ZIN)
Copyright © Ilya Kabak

Carabus (Morphocarabus) tarbagataicus var.motschulskyi Lapouge, 1909 Holotype
Male adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC525 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec525
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Carabus (Morphocarabus) monilis var.motschulskyi Lapouge, 1909
Label
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC525 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec525
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Carabus (Morphocarabus) venustus venustus Morawitz, 1862
Male adult
SE Primorskii reg., Lazovskii Nat.Res., 12 km S Staraya Kamenka vill., near Amerika kor. 43°15N, 134°1E, 19.VII.04 leg. M.& L. Smirnovy photo from www.zin.ru
Copyright © Maksim Smirnov

Carabus (Morphocarabus) venustus venustus Morawitz, 1862
Male adult
Russia, Far East, Primorskiy reg., Ussuriysk Distr., Kaymanovka vill., 6-15.VIII.2011, <br /> A. Korshunov leg.<br /> L= 21.5 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) venustus venustus Morawitz, 1862
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) venustus venustus Morawitz, 1862
Male adult

Copyright © Kirill Makarov

Carabus (Morphocarabus) venustus Morawitz, 1862
Male adult
China, Heilongjiang, Wuchang city, Mt. Lao-Tu-Ding-Zi 1593m 1-18.6.2008 (cAA).
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) odoratus czernyscheviellus Obydov, 2002 PT Paratype
Female adult
Carabus (Morphocarabus) odoratus czernyscheviellus PT
Rusia, SE Transbaikalia, Chita reg., Nerchinsko-Kugenskii Mt.rng.,
near Vershino-Darasunskii vill., Bogatui Mt. 1221 m.,
15.VII.2002, D. Obydov leg.
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859
Male adult
Russia, SE Siberia, Transbaikalia, N Chita reg., Kodar Mt. Range, Big Leprindo Lake env.h~1800m., 1.VII.2006, S. Rybalkin leg. L=20 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) viridilimbatus panini I.Solodovnikov 2017 Paratype
Male adult
Russia, SE Yakutia, Neryungri distr., Tokinski Stanovik Mt Rng., Beder riv., (right tributary rivers Algama), 26.VI.1996, Krasnov M. leg. L=15 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) viridilimbatus panini I.Solodovnikov 2017 Holotype
Male adult
Russia, SE Yakutia, Neryungri distr., Tokinski Stanovik Mt Rng., Beder riv., (right tributary rivers Algama), 26.VI.1996, Krasnov M. leg., 17 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) viridilimbatus panini I.Solodovnikov 2017 Paratype
Female adult
Russia, SE Yakutia, Neryungri distr., Tokinski Stanovik Mt Rng., Beder riv., (right tributary rivers Algama), taiga, h = 940 m, 55°55'27.78" N, 129°42'6.20" E, 26.VI.1996, leg. Krasnov M. 21,1 mm
Copyright © Igor Solodovnikov

Carabus (Morphocarabus) viridilimbatus saluki I.Solodovnikov 2017 Holotype
Male adult
Russia, SW Taymyr, near Talnakh, 25 km N Norilsk, hilly tundra, hillside overgrown with alder slate, dwarfish birches, blueberry, h = 150-250 м, 24-28.06.2003, leg. S.Saluk 20,5 mm
Copyright © Igor Solodovnikov

Carabus (Morphocarabus) viridilimbatus saluki I.Solodovnikov 2017 Paratype
Male adult
Russia, SW Taymyr, near Talnakh, 25 km N Norilsk, hilly tundra, hillside overgrown with alder slate, dwarfish birches, blueberry, h = 150-250 м, 24-28.06.2003, leg. S.Saluk
Copyright © Igor Solodovnikov

Carabus (Morphocarabus) viridilimbatus saluki I.Solodovnikov 2017 Paratype
Female adult
Russia, SW Taymyr, near Talnakh, 25 km N Norilsk, hilly tundra, hillside overgrown with alder slate, dwarfish birches, blueberry, h = 150-250 м, 24-28.06.2003, leg. S.Saluk
Copyright © Igor Solodovnikov

Carabus (Morphocarabus) viridilimbatus saluki I.Solodovnikov 2017 Paratype
Endophallus
Russia, SW Taymyr, near Talnakh, 25 km N Norilsk, hilly tundra, hillside overgrown with alder slate, dwarfish birches, blueberry, h = 150-250 м, 24-28.06.2003, leg. S.Saluk
Copyright © Igor Solodovnikov

Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859
Endophallus
Russia, Chita Region, Udokansky district. Kodar Range, 56 °34'42,9 " N, 118 °29'34,95" E, 6/13/2002, leg. A. Pokhorov
Copyright © Igor Solodovnikov

Carabus (Morphocarabus) wulffiusi wulffiusi Morawitz, 1862
Male+Female adult
Russia, Far East,
Primorskiy Kray, near Kamenushka vill., 2-3.VIII.1992, K.E.Dovgaylo leg.
female, 24mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) wulffiusi wulffiusi Morawitz, 1862
Female adult

Copyright © Safronov

Carabus (Morphocarabus) wulffiusi wulffiusi Morawitz, 1862
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) wulffiusi dekraatzi Kraatz, 1881
Male adult
NE-China, Heilongjiang, Wuchang city, Mt Zheng-nan-gou, 1-11.VI.2011<br /> L=22 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) wulffiusi liaoningensis Deuve & Imura, 1990
Female adult
NE-China:E-Liaoning: NP Kuandian Tianqiaogou , VI.2010, L=24
Copyright © Ruslan Panin

Carabus (Morphocarabus) wulffiusi liaoningensis Deuve & Imura, 1990
Female adult
China, Liaoning, Kuandian Co., Banlaing, Ba-che-chuan (cAA)
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
Male adult
W-UKRAINE, Transcarpathian region, Rakhiv distr., Chornogora Mt Range, h=800 m, near Bogdan vill., 10.VI.2012, Panin R. leg.
L= 33 мм
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
Female adult
W Ukraine, NE Carpathians,
Transcarpathian Reg., Marmarosh Mts., Rakhiv distr., 5 km W Dilowe vill., Site Kuziy, 5-15.VII.2010, Demidov G.N. leg./det.Panin R.
L= 35 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Endophallus
Transcarpathian region, Uzhgorod distr., 28.IV-23.V.2011
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Endophallus
SE Slovakia, Slanske vrchy Mts, Herlianske sedlo, 8-19.VI.2009, Krause Rene leg.
det.Panin R.
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii zawadzkii, Kraatz 1854
Endophallus
Gorgany Mts. h=700 m, 16.VI.2011,
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii zawadzkii, Kraatz 1854
Male+Female adult
W.UKRAINE, Ukrainian Carpathians
Lviv Reg., Skole distr., Skolivski Beskidy Mts. Zelemyanka Mt. h=1000m. 9-20.VII.07 Panin R. leg.
female, 35mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Male adult
NE-Hungary, 4 km S Sátoraljaújhely, right bank riv. Bodrog (Long-erdő), 8.VI.1992, leg./det. I.Retezár leg. L=28 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
Male adult
W Ukraine, Transcarpathian lowland, near Chop, 11.VI-19.VII.2019, Panin R leg. L=28 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii zawadzkii, Kraatz 1854
Male adult
W Ukraine, NE Carpathians, Lviv region, Skole Beskids Mts, Zelemyanka Mt. Range h=915 m, 22.VI-17.VII.2022, Panin R. & Kanarsky Yu. leg., L= 32 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Endophallus
SE Poland, Subcarpathia Prov., Nadleśnictwo Kołaczyce N of Jasło vill., M. Kosibowicz leg./det. R Panin
Copyright © Ruslan Panin

Carabus (Rhigoidocarabus) zhubajie laojunensis Kleinfeld, 1999 Holotype
Male adult

Copyright © Frank Kleinfeld

Carabus (Morphocarabus) zhubajie zhubajie Imura, 1993
Male adult

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