Carabidae

Carabidae image gallery

Region

Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
Female adult

Copyright © Evgeniy Chibilov

Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
Male adult
Russia,
S Ural Mts, Mt.Iremel h=800m, 4.VI.2000 Filimonov V. leg.
male,21mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
Male adult
Carabus (Morphocarabus) karpinskii Rusia, S Ural mts., Jremel Mt., 1300 m 4.6.2000male, 20 mm
19.08.2008
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) karpinskii Kryzhanovskij & Matveev, 1993
Female adult

Copyright © Evgeniy Chibilov

Carabus (Morphocarabus) karpinskii Kryzhanovskij & Matveev, 1993
Female adult

Copyright © Evgeniy Chibilov

Carabus (Diocarabus) loschnikovi loschnikovi Fischer, 1823
Male adult
Komi Resp., Pecero-Ilych Nat. Res., Zajachij Kamen h~786 62°10'06" N 59°12'53" E 8.VIII.2004 M. Chernjakhovski leg.
L=15.0 mm
Copyright © Evgeniy Zinoviev

Aplothorax burchelli Waterhouse, 1841
Male adult
St. Helena island
Copyright © Boleslav Brezina

Calosoma (Calosoma) inquisitor cupreum Dejean, 1826: 196
Female adult
Armenia, Zangezur range, Meghri suburbs., Vahravar vill., Rivulet vall., h=1400 m Yuriy Shevnin leg.; 23.05.2007; 21 mm.
Copyright © Anton Menstschikov

Calosoma (Calosoma) inquisitor cupreum Dejean, 1826: 196
Male adult
Iran, Ardabil, 30 km NE Ardabil,
5 km SE Namin, 18-19.5.2007 Anichtchenko A. leg.
male, 20 mm
Copyright © Alexander Anichtchenko

Calosoma (Calosoma) schayeri schayeri Erichson, 1842
Male adult
Australia N.T. Alice Springs, Airport
19.5.2000 M. Langer leg.
male, 21 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisphaena) reticulatum reticulatum s.str.
Female adult
Germania, or. DDR, Bezirk Frankfurt, 24.V.1989
female, 25 mm
photo & det. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Callistenia) moniliatus Le Conte, 1851: 200
Male adult
SE Canada, Nueva Escocia
19.5.1970
male, 17 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) anthrax Semenov, 1900 Holotype
Male adult
collection of ZIN
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) breviusculus Mannerheim, 1830
Male adult
Armenia, near Ashtarik 15.6.1997 male, 22 mm photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) elegans s.str. (as declivis Dohrn, 1884b)
Male adult
SE Kazakhstan Alma-Ata reg., 30 km N Sary-Ozek 28.4.1998 Klimenko leg. male, 22 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) elegans s.str. (as declivis Dohrn, 1884b)
Female adult
SE. Kazakhstan, Sary-Olek vill., 26.IV.1997
Copyright © Anton Menstschikov

Calosoma (Callisthenes) elegans amethystinus Semenov & Redikorzev, 1928
Male adult
SE Kazakhstan, E Zailiyski mt. rng. Chilik riv.
Malybai vill. 28.4.1999 Klimenko A. leg.
male, 25 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) elegans amethystinus Semenov & Redikorzev, 1928
Male adult
26 mm.
Zailiyski Alatay.
06.1997 Chibilov E. leg.
Copyright © Sunvas

Calosoma (Callisthenes) elegans amethystinus Semenov & Redikorzev, 1928
Female adult
SE Kazakhstan, E Zailiyski mt. rng. Chilik riv.
Malybai vill. 28.4.1999 Klimenko A. leg.
female, 26 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) elegans elegans s.str.
Male adult
SE Kazakhstan, Tchu-Iliyskiy Mts., Kordai pass, h=1500; A. Klimenko leg.; 20.04.1997;
24 mm.
photo A. Menstschicov
Copyright © Anton Menstschikov

Calosoma (Callisthenes) elegans karagaicus Lapouge, 1924
Male adult
Inner Tian-Shan, 20 km N jf Naryn
27.5.1997 Klimenko A. leg.
male, 27 mm
photo, det. & coll. Anichtchenko A. 2009
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) elegans karagaicus Lapouge, 1924
Male adult
KIRGHIZSTAN, Inner Tian-Shan Mts., near Naryn tn., Naryn riv. Valley, 18-20.V.1999, Leg. Irtlatch A.
Copyright © Anton Menstschikov

Calosoma (Callisthenes) elegans manderstjernae Ballion, 1870
Female adult
Kirgizia, S Kirgizskii mt. rng., 2200m
Kichi-Kimin riv., Ak-Tuz vill. 05.1998 Klimenko leg.
female, 32 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) elegans semenovi Motschulsky, 1859
Female adult
SE Kazakhstan, Zailijsky Mt. rng.,
Talgar env., 1500 m, 21.5.2000
female, 27 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) eversmanni Chaudoir, 1850
Male adult
Syria, Antilibanon
1.3.1991
male, 22 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) fischeri Fischer, 1842
Male adult
Mongolia, Mandalgob, 20 km
N Amardalai vill. 11-12.6.2003 Anichtchenko A. leg
male, 19 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) grumi Semenov, 1900 Holotype
Female adult
collection ZIN
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) breuningi Mandl, 1954
Female adult
E Kazakhstan, S Alakol lake, Koktuma vill., 1-10.5.1999 Nekroshevich S. leg. female, 22 mm; det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) breuningi Mandl, 1954
Male adult
E.Kazakhstan, Alakol lake, Koktuma vill., 07.V.1999
Copyright © Anton Menstschikov

Calosoma (Callisthenes) breuningi iliense Kabak, 2022
Male adult
SE Kazakhstan, near Sary Ozek vill., A. Klimenko leg; 22.04.1997; 22 mm.; det. Anichtchenko A. 2022
Copyright © Anton Menstschikov

Calosoma (Callisthenes) kuschakewitschi kuschakewitschi s.str.
Male adult
Karzhantau mt. rng., Susungen riv.,
2300 m, 7.6.1998 Tretjakov leg.
male, 23 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) kuschakewitschi plasoni Born, 1917
Male adult
Kirgizia, 40 km S Bishkek
10.5.1990
male, 26 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) kuschakewitschi plasoni Born, 1917
Female adult
Kirgizia, Chon-Aryk. 23.V.1998 V. Dolin leg.
Copyright © Denis Potanin

Calosoma (Callisthenes) marginatus Gebler, 1829
Female adult
NE Kazakhstan, N Zaisan lake, 20.6.2000
Narymskii mt.rng. Klimenko A. leg.
female, 18 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) mirificus Casale, 1979
Female adult
Afganistan, S slope of Paropamiz Mt. rng., near Sabzan pass, 2400m, 22.VI.1967 M.Stoljarov leg.
Collection of ZIN
det. D. Obydov 1999
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) panderi Fischer, 1822
Male adult
NW Kazakhstan, Aktiubinsk reg., Baiganinsk distr.,
near Zharly vill. 10-12.05.1997 leg. M.& L. Smirnov
male, 20 mm
Photo from www.zin.ru
Copyright © Maksim Smirnov

Calosoma (Callisthenes) pavlovskii Kryzhanovskij, 1955
Male adult
Kirgizstan, Talas Mt. rng., Besh-Tash
riv., 1000m, 15.5.2003
male, 20 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) pseudocarabus Semenov et Redikorzev, 1928
Male adult
NE Kazakhstan, Saur mt.rng. 2000m
Sargar mt., 14.6.2001 Klimenko leg.
male, 16 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) pseudocarabus Semenov et Redikorzev, 1928
Female adult
E Kazakstan, NE Saur, 2 km. W m. Tas, h=2500; 10.06.2001, Ivanov A.V. leg.,
15 mm., female
photo, det. & coll. Menstschikov
Copyright © Anton Menstschikov

Calosoma (Callisthenes) pseudocarabus Semenov et Redikorzev, 1928
Male adult
E Kazakstan, NE Saur, 2 km. W m. Tas, h=2500; 10.06.2001, Ivanov A.V. leg., 14 mm., male
photo, det. & coll. Menstschikov
Copyright © Anton Menstschikov

Calosoma (Callisthenes) regelianus A. Morawitz, 1886
Female adult
Tadzikistan, Darvaz Ezgan 2.5.1971
female, 25 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) usgentensis usgentensis Solskyi, 1874
Male adult
Kirghizstan, near Jalal-Abad town, A. Klimenko leg., 05.04.1997; 30 mm.
photo, det. & coll. A. Menstschicov
Copyright © Anton Menstschikov

Calosoma (Callisthenes) usgentensis usgentensis Solskyi, 1874
Female adult
S Kirgizia, Dzhalal-Abad reg.,
Suzak vill., 1000m, 25.5.1999
male, 27 mm
photo, det. & coll. A. Anichtchenko
Copyright © Alexander Anichtchenko

Calosoma (Callitropa) externum Say, 1823: 149
Female adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Calosoma (Calosoma) scrutator Fabricius, 1775
Female adult
USA, Illinois, Peoria Co., Orange Prairie, 31.5.1990 32 mm coll. A.Montes photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Calosoma) scrutator Fabricius, 1775
Male adult
3 m SE Tishomingo , Tishomingo Co., Mississippi, USA; 24.VI.2005 Eric B. Lago leg.
Copyright © Denis Potanin

Calosoma (Calosoma) wilcoxi Le Conte, 1848
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Calosoma (Calosoma) wilcoxi Le Conte, 1848
Male adult
2 m. W Rawlings, Allegany Co., Maryland, USA; 26 May 1983 J.D. Glaser leg.
Copyright © Denis Potanin

Calosoma (Calosoma) cyanescens Motschulsky, 1859
Male adult
Primorskii reg., Kedrovaja Pad Nat.res., 11.VII.1994 Chichvarhin A. leg
male, 19 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Calosoma) frigidum Kirby, 1837: 19
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Calosoma (Calosoma) frigidum Kirby, 1837: 19
Male adult
2 m W Rawlings, Allegany Co., Maryland, USA; 26.V.1983 J.D. Glaser leg.
Copyright © Denis Potanin

Calosoma (Calosoma) maximowiczi A. Morawitz, 1863: 20
Female adult
China, Mt. Wangfoshan (1940 m.), Baokang Countri, Hubei; 20-28.06.2008, 30 mm.
Copyright © Anton Menstschikov

Calosoma (Calosoma) oceanicum klynstrai Breuning, 1927
Female adult
INDONESIA, Tanimbar Island, III.2009
photo & coll. N. Belonogov
Copyright © Nil Belonogov

Calosoma (Calosoma) oceanicum klynstrai Breuning, 1927
Male adult
INDONESIA, Tanimbar Island, III.2009
photo & coll. N. Belonogov
Copyright © Nil Belonogov

Calosoma (Calosoma) sycophanta Linne, 1758
Male adult
Spain, Sierra Nevada, Pico de Veleta
2800m
Copyright © Alexander Anichtchenko

Calosoma (Carabosoma) prominens LeConte, 1853
Male adult
USA Arizona, Cochise Co., Interstate 8 at Texas, Canyon Rest Area, 1500m, 32º00.3N 110º08.4W, 24.7.1996 Kavanaugh D. leg.; 30 mm
Copyright © Alexander Anichtchenko

Calosoma (Carabosoma) peregrinator Guerin-Meneville, 1844
Male adult
Fort Davis, Jeff Davis Co., Texas, USA; July 1998
Copyright © Denis Potanin

Calosoma (Caminara) denticolle Gebler, 1833: 274
Female adult
Burjatia, Selenga riv., Deben vill. 12.7.2003 Anichtchenko A. leg. female, 25 mm
Copyright © Alexander Anichtchenko

Calosoma (Caminara) imbricatum deserticola Semenov, 1897d: 242
Male adult
W Kazakstan, Ustjurtskij res., Kordon Kenderli; 01.05.2009, Gorbunov P.J. leg.,
25 mm., male
photo, det. & coll. Menstschikov
Copyright © Anton Menstschikov

Calosoma (Caminara) imbricatum deserticola Semenov, 1897d: 242
Male adult
W Kazakstan, Ustjurtskij res., Kordon Kenderli; 01.05.2009, Gorbunov P.J. leg., 24 mm., female photo, det. & coll. Menstschikov
Copyright © Anton Menstschikov

Calosoma (Campalita) chlorostictum Dejean, 1831a: 558
Male adult
RSA, Kwazulu/Natal, Badplaas, 1000m, 1.XII.2000 male, 24 mm det. P. Schüle photo &. coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Carabophanus) arrowi Breuning, 1928
Male adult
Ethiopia, Oromia reg., Arsi highlands, Wahoro env., Tiliku Lensho river valley10.-12.II.2010, J.Štastny lgt
photo Jaroslav Stastny
Copyright © Jaroslav Štastny

Calosoma (Castrida) alternans granulatum Perty, 1830
Male adult
Paraguay, Dep.Caaguazu, Tayao S25.28 W56.07 12-16.III.2003 leg. U.Drechsel
male, 28 mm
photo, det. & coll. Anichtchenko A. 2009
Copyright © Alexander Anichtchenko

Calosoma (Castrida) alternans granulatum Perty, 1830
Female adult
Argentina, Entre Rios,
Liebig, nov.1997
long. 32 mm
photo, det. & coll. Anichtchenko A. 2009
Copyright © Alexander Anichtchenko

Calosoma (Castrida) argentinensis Csiki, 1927
Male adult
Argentina, Prov. Santa Fe Coronda, 22.12.1965 leg. Mahunka, 23 mm
Copyright © Alexander Anichtchenko

Calosoma (Castrida) argentinensis Csiki, 1927
Male adult
ARGENTINA: Salta; November 1995
Copyright © Denis Potanin

Calosoma (Castrida) vagans Dejean,
Male adult
Chile, Santiago de Chile, Quilicura
19.9.1991 G. Aceituno leg.
male, 21 mm
Copyright © Alexander Anichtchenko

Calosoma (Charmosta) investigator Illiger, 1798
Male adult
Lake Baikal, Buriatia, Iliynka vill., 1.8.1997 Anichtchenko A. leg.; 20 mm
Copyright © Alexander Anichtchenko

Calosoma (Charmosta) investigator Illiger, 1798
Male adult
S. Ural, Bashkiria, near Mesjagutovo vill., 09.07.2001 A. Menstschicov leg.
Copyright © Anton Menstschikov

Calosoma (Charmosta) lugens Chaudoir, 1869f 372
Male adult
Wen-dong, North Fengcheng City, Liaoning, China; 1-5.VII.2006
Copyright © Denis Potanin

Calosoma (Chrysostigma) calidum Fabricius, 1775
Male adult
Canada Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Calosoma (Chrysostigma) semilaeve LeConte, 1851: 199
Male adult
Orange Co., California, USA; May 1973
Copyright ©

Calosoma (Ctenosta) bastardi Alluaud, 1925
Female adult

Copyright © Alexander Anichtchenko

Calosoma (Ctenosta) orientale Hope, 1838
Male adult
China, Yunnan pro.
Copyright © Ye Liu

Calosoma (Ctenosta) planicolle Chaudoir, 1869
Male adult
length: 30 mm. Kenia, Tsavo National Parc, leg. U.Schmidt, 1991; det. W.Lorenz, 2000. Foto: U.Schmidt, 2006
Copyright © U.Schmidt

Calosoma (Ctenosta) planicolle Chaudoir, 1869
Male adult
SOUTH AFRICA: Northern Transvaal
Copyright © Denis Potanin

Calosoma (Ctenosta) senegalense Dejean, 1831
Female adult
length: 22-27 mm
Kenia Tsavo National Parc
leg. U.Schmidt, 1991; det. W.Lorenz, 2000
Foto: U.Schmidt, 2006
Copyright © U.Schmidt

Calosoma (Microcallisthenes) relictum Apfelbeck, 1918c
Male adult
Macedonia, Shar mt., Peak Tserapashina
1850-2530m 9.7.1995
male, 13 mm
Copyright © Alexander Anichtchenko

Calosoma (Microcallisthenes) relictum Apfelbeck, 1918c
Male adult
Macedonia, Shar mt., Peak Titov Vrah
2747 m, 14.7.1995 V.Sakalian leg.
male, 13 mm
Copyright © Alexander Anichtchenko

Calosoma (Calosoma) inquisitor inquisitor Linne, 1758: 414
Male adult
length: 13-24 mm
England, Dendless Wood
leg.det. M.G.Telfer, 2003
Foto: U.Schmidt, 2009
Copyright © U.Schmidt

Calosoma (Calosoma) himalayanum Gestro, 1875b: 851
Female adult
China, Tibet
Copyright © Ye Liu

Calosoma (Carabops) abyssinicum Gestro, 1880
Male adult
Ethiopie, Shemene 08.2004
19 mm
coll. A.Montes
Copyright © Alexander Anichtchenko

Calosoma (Carabops) abyssinicum Gestro, 1880
Male adult
Ethiopia, Bale prov., Str. ZW. Sasheme u. Dodola, Mt. Mascall 3100-3300 m
21.VII.2005 Puchner leg.
male, 18 mm
det. Puchner F. 2005
photo & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Ceroglossus buqueti deuvei Jiroux, 1996
Male adult
Chile, VIII Reg., 1000-1250 m, P.N. Nahuelbuta, (Piedra del Aguila) 7-12.XII.1999 Heinz leg.
male, 22 mm
photo, det. & coll. Anichtchenko A. 2009
Copyright © Alexander Anichtchenko

Ceroglossus buqueti andestus Kraatz-Koschlau, 1887
Male adult
CHILE, Liquine; 05/12.02.2005; C.Saavedra leg.; Determination confirmed by Anichtchenko, 2019
Copyright © Anton Menstschikov

Ceroglossus buqueti cherquencoensis Dorsselaer, 1955
Male adult
Chile, IX reg., Cherquenco Vn. Llaima, 800-1000m, 29-30.I.1999 Heinz leg. 21 mm; det. & coll. Anichtchenko A. 2019
Copyright © Alexander Anichtchenko

Ceroglossus buqueti sybarita Gerstaecker, 1858
Female adult
Chile, Chiloe, 160m, 8 km N Butalcura 22-24.XII.1998 det. Heinz
female, 24 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Ceroglossus chilensis cyanicollis Kraatz, 1887
Male adult
Chile (VIII reg), Umg. Lag. de la Laja, 950 m, 19-20.12.1998 Heinz leg. 26 mm; det. Anichtchenko A., 2019
Copyright © Alexander Anichtchenko

Ceroglossus chilensis fallaciosus Kraatz, 1880
Male adult
CHILE, Talca prov., Constitucion tn. env., II.2008. det. Anichtchenko A., 2019
Copyright © Nil Belonogov

Ceroglossus chilensis ficheti Jiroux, 1996
Male adult
Chile (VIII reg), P.N. Nahuelbuta, (Piedra del Aguila) 700 m, 7-12.12.1999, Heinz leg.; 28 mm. Determination confirmed by Anichtchenko, 2019
Copyright © Alexander Anichtchenko

Ceroglossus chilensis gloriosus Gerstaecker, 1858
Male adult
Chile (IX reg.), Malalcahuello, 800 m, 9-10.12.1999 Heinz leg. 25 mm; Determination confirmed by Anichtchenko, 2019
Copyright © Alexander Anichtchenko

Ceroglossus chilensis latemarginatus Kraatz-Koschlau, 1889
Male adult
Chile (VIII. Reg.) Pata de Gallina, 650m, b.Contulmo 7-12.12.1999 Heinz leg. 27 mm; Determination confirmed by Anichtchenko, 2019
Copyright © Alexander Anichtchenko

Ceroglossus chilensis mochae Reed, 1874
Female adult
CHILE, Arauco prov., Isla Mocha. - II.2008. photo Belonogov N.; Determination confirmed by Anichtchenko, 2019
Copyright © Nil Belonogov

Ceroglossus chilensis resplendens Jaffrezic & Rataj, 2006
Male adult
CHILE, Huife, IX Region.
Copyright © Anton Menstschikov

Ceroglossus chilensis seladonicus Kraatz-Koschlau, 1887
Male adult
CHILE, Malleco prov., Malalcahuello Nat. Res. - I.2008
photo Belonogov N.
Copyright © Nil Belonogov

Ceroglossus chilensis seladonicus Kraatz-Koschlau, 1887
Female adult
Chile, Malleco prov., Madalcahuello
enero 2003 G. Aceituno leg.
28 mm
Copyright © Alexander Anichtchenko

Ceroglossus chilensis solieri Roeschke, 1900
Female adult
Chile, Chiloe, 160m, 8 km N Butalcura 22-24.XII.1998 det. Heinz
female, 28 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Ceroglossus darwini darwini s.str.
Male adult
Chile, Chiloe, ca. 25 km NW Queilen, 350 m, (n. Castro), 26-27.XII.1999 Leg. Heinz
male, 22 mm
photo, det. & coll. Anichtchenko A. 2009
Copyright © Alexander Anichtchenko

Ceroglossus resplendens Jaffrezic & Rataj, 2006
Female adult
CHILE. X Region, Termas de Carranco; 01/05 February. 2007; C.Saavedra leg.; det. Anichtchenko A. 2019
Copyright © Anton Menstschikov

Ceroglossus speciosus Gerstaecker, 1858
Female adult
Chile, Chiloe, S. Pedro de Piruquiña 350 m, (n. Castro), 24-25.XII.1999 Heinz leg.
female, 23 mm
photo, det. & coll. Anichtchenko A. 2009
Copyright © Alexander Anichtchenko

Calosoma (Teratexis) fabulosum Semenov & Znojko, 1933 Holotype
Male adult
collection of ZIN
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko

Calosoma (Teratexis) fabulosum Semenov & Znojko, 1933 Paratype
Male adult
collection of ZIN
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko

Pamborus alternans Latreille, 1817
Female adult
Australia, NSW. Kerewong S.F.
21.12.1978
30 mm
Copyright © Alexander Anichtchenko

Pamborus brisbanensis Castelnau, 1867
Male adult
Museum of Zurich
photo Anichtchenko
Copyright © Alexander Anichtchenko

Pamborus guerini Gory, 1830
Male adult
Museum of Zurich
photo Anichtchenko
Copyright © Alexander Anichtchenko

Pamborus opacus Gehin, 1885
Male adult
Museum of Zurich
photo Anichtchenko
Copyright © Alexander Anichtchenko

Pamborus pradieri darlingtoni Banninger, 1940 Paratype
Male adult
Museum of Zurich photo Anichtchenko
Copyright © Alexander Anichtchenko

Pamborus pradieri darlingtoni Banninger, 1940
Female adult
Australia, N.S.W. New England rng.,
Moonan Flats, 18.12.2000 T.Pollak leg.
21 mm
Copyright © Alexander Anichtchenko

Pamborus pradieri pradieri Chaudoir, 1869
Male adult
Museum of Zurich
photo Anichtchenko
Copyright © Alexander Anichtchenko

Pamborus punctatus Darlington, 1961 Paratype Paratype
Male adult
Museum of Zurich
photo Anichtchenko
Copyright © Alexander Anichtchenko

Pamborus tropicus Darlington, 1961 Paratype Paratype
Male adult
Museum of Zurich
photo Anichtchenko
Copyright © Alexander Anichtchenko

Carabus (Acathaicus) alexandrae alexandrae Semenov, 1887 Holotype Holotype
Male adult
Kwantin and Li-tschara-pu, 22.VI.1885, leg. Potanin Deposited in Zoolog. Museum St. Petersburg. male, long. 22 mm photo B. Brezina
Copyright © Boleslav Brezina

Carabus (Acathaicus) alexandrae alexandrae Semenov, 1887
Male adult
male, 25mm photo B. Brezina
Copyright © Boleslav Brezina

Carabus (Acathaicus) alexandrae alexandrae Semenov, 1887
Male adult
female, 28mm photo B. Brezina
Copyright © Boleslav Brezina

Carabus (Acathaicus) alexandrae alexandrae Semenov, 1887
Male adult
female, 24mm photo B. Brezina
Copyright © Boleslav Brezina

Carabus (Acathaicus) alexandrae cratocephaloides Semenov, 1887 Holotype Holotype
Male adult

Copyright © Boleslav Brezina

Carabus (Acathaicus) alexandrae cratocephaloides Semenov, 1887
Male adult
male, 26mm
Copyright © Boleslav Brezina

Carabus (Acathaicus) alexandrae cratocephaloides Semenov, 1887
Male adult
male, 24mm
Copyright © Boleslav Brezina

Carabus (Acathaicus) alexandrae cratocephaloides Semenov, 1887
Female adult
female, 28mm
Copyright © Boleslav Brezina

Carabus (Acathaicus) idolon Semenov, 1898 Holotype Holotype
Male adult
male
Copyright © Boleslav Brezina

Carabus (Acathaicus) interruptocostatus interruptocostatus s.str.
Female adult
female, 27mm
Copyright © Boleslav Brezina

Carabus (Acathaicus) interruptocostatus interruptocostatus s.str.
Male adult
male, 29mm
Copyright © Boleslav Brezina

Carabus (Acathaicus) interruptocostatus Semenov, 1887 Holotype Holotype
Female adult

Copyright © Boleslav Brezina

Carabus (Acoptolabrus) constricticollis constricticollis Kraatz, 1886
Female adult

Copyright © Alexander Anichtchenko

Carabus (Acoptolabrus) gehinii gehinii (as aereicollis Hauser, 1921)
Male adult

Copyright © Alexander Anichtchenko

Carabus (Acoptolabrus) gehinii gehinii Fairmaire, 1886
Male+Female adult
Komari, Chitose, Hokkaido, Japan.
20-27.05.2007
Copyright © Safronov

Carabus (Acoptolabrus) gehinii gehinii Fairmaire, 1886
Male+Female adult
Bibi, Chitose, Hokkaido, Japan.
03.05.2000
Copyright © Safronov

Carabus (Acoptolabrus) gehinii gehinii (as manoianus (Imura, 1989)
Male adult

Copyright © Alexander Anichtchenko

Carabus (Acoptolabrus) gehinii munakatai (Ishikawa, 1968)
Male+Female adult
Mt.Shiribetsudake, Kishibetsu, Hokkaido, Japan.
05.06.2005
Copyright © Safronov

Carabus (Acoptolabrus) leechi leechi Bates, 1888
Male adult

Copyright © Boleslav Brezina

Carabus (Acoptolabrus) leechi nangminicola Deuve & Li, 2001
Female adult
N Coree, Cha Gangdo, Nangnimsan Mt., 6.2006. 31 mm. (coll. A. Montes, Madrid)
Copyright © Alexander Anichtchenko

Carabus (Acoptolabrus) lopatini Morawitz, 1886
Male+Female adult
Russia,
Sakhalin isl., Tchekhov Mt, VI.1997
female, 40mm
photo & coll. Panin R.
Copyright © Ruslan Panin

Carabus (Acoptolabrus) mirabilissimus rigoutianus Deuve & Li, 1998
Female adult
N Koree, Pyonganbukto, mt. Miaoxiang 6.2006
28 mm
coll. A.Montes
Copyright © Alexander Anichtchenko

Carabus (Acoptolabrus) schrencki schrencki (as hauryi Gehin, 1885 )
Male adult

Copyright © Boleslav Brezina

Carabus (Acoptolabrus) schrencki schrencki (as hauryi Gehin, 1885 )
Male adult
det. Brezina B.
Copyright © Boleslav Brezina

Carabus (Acoptolabrus) schrencki schrencki (as hauryi Gehin, 1885 )
Male adult

Copyright © Boleslav Brezina

Carabus (Acoptolabrus) schrencki schrencki Motschulsky, 1860
Female adult
Primorie, Lazovskii distr., Chandalaz mt., L.Voita leg 19.07.2008
photo A.Menstschicov
Copyright © Anton Menstschikov

Carabus (Acoptolabrus) schrencki kimieae Kezuka & Imura, 1989
Male+Female adult
Mt. Wulongshan, Dandong City,
Liaoning Province, CHINE
25-30.VI.2008
Copyright © Safronov

Carabus (Acrocarabus) callisthenoides Semenov, 1888
Male adult
male, 22mm
Copyright © Boleslav Brezina

Carabus (Acrocarabus) callisthenoides Semenov, 1888
Male adult
female, 23mm
Copyright © Boleslav Brezina

Carabus (Acrocarabus) guerini aldarkose Kabak, 1992
Male adult
male, 22mm
Copyright © Boleslav Brezina

Carabus (Acrocarabus) guerini guerini Fischer, 1842
Male adult
male, 21mm
Copyright © Boleslav Brezina

Carabus (Acrocarabus) guerini guerini Fischer, 1842
Female adult
female, 25mm
Copyright © Boleslav Brezina

Carabus (Acrocarabus) guerini guerini Fischer, 1842
Male adult
SE Kazakhstan, W Dzhungar Mts, Lepsinsk env., 1600 m., 18.06.2000, A.Klimenko
Copyright © Anton Menstschikov

Carabus (Alipaster) barovskii bayanbulak Deuve, 1993 Paratype Paratype
Male adult
male, 15mm
Copyright © Boleslav Brezina

Carabus (Alipaster) barovskii bayanbulak Deuve, 1993 Paratype Paratype
Female adult
female, 17mm
Copyright © Boleslav Brezina

Carabus (Alipaster) infantulus infantulus Morawitz, 1886
Male adult
male, 15mm
Copyright © Boleslav Brezina

Carabus (Alipaster) pupulus pupulus Morawitz, 1889
Male adult
Inner Tien-Shan, W part of Terskei mnts., Ukek rng., 20-25 km E Kotshkorka, h=2800 m, 8.07.98 leg. Tretyakov male 13,6mm
Copyright © Maksim Smirnov

Carabus (Alipaster) pupulus pupulus Morawitz, 1889
Male adult
male, 13mm
Copyright © Boleslav Brezina

Carabus (Alipaster) pupulus pupulus Morawitz, 1889
Male adult
female, 16mm
Copyright © Boleslav Brezina

Carabus (Alipaster) pupulus pupulus Morawitz, 1889
Male adult
Kirgizstan, Terskei AlaToo Mt. rng, Kadzhi-Sai v., 2800 m.; A.Subankulov leg.; 30.05.2000
Copyright © Anton Menstschikov

Carabus (Apoplesius) deuvei hajdaji Hackel, 2007 Holotype Holotype
Male adult
Turkey- prov. Mus 60 km NW Mus 1630m
3856N 4108E Bulgan Gec. 23-24.06.03 E. & P. Hajdaj leg.
Copyright © Boleslav Brezina

Carabus (Apotomopterus) anxiensis Kleinfeld, 1998 Holotype & Paratype Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) arrowi hector Hauser, 1924
Female adult
China, Guangxi pro., Maoershan Mountain, Liang H.B. coll. Photo by Liuye det. Anichtchenko A.
Copyright © Ye Liu

Carabus (Apotomopterus) bousquetellus gengmaensis Kleinfeld, 2003 Holotype & Paratype Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) cantonensis pervarius Kleinfeld, 1997 Holotype
Male adult
Frank Kleinfeld
Copyright © - - -

Carabus (Apotomopterus) cantonensis pervarius Kleinfeld, 1997
Male adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) casaleianus jigongshanicus Kleinfeld, 1998 Holotype & Paratype Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) clermontianus Breuning, 1933
Female adult
N Viet-Nam, 100 km NW Hanoi 5.1995 Rapaltovskii E. leg.
Copyright © Alexander Anichtchenko

Carabus (Apotomopterus) datianshanicus Kleinfeld, 1997 Holotype & Paratype Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) davidis davidis Deyrolle & Fairmaire, 1878
Female adult

Copyright © Alexander Anichtchenko

Carabus (Apotomopterus) cantonensis metallaicus Kleinfeld, 1999 Holotype Holotype
Female adult
as pervarius metallaicus
Copyright © Frank Kleinfeld

Carabus (Apotomopterus) davidis neodavidis Kleinfeld & Schutze, 2001 Holotype & Paratype Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) davidis viridithoracis Kleinfeld & Schutze, 2001 Holotype & Paratype Holotype Paratype
Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) davidis yizhangensis Kleinfeld, 2002 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) delavayi s.str. (as medea Kleinfeld, 1999) Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) delavayi tenuimanus Deuve & Imura, 1990
Male adult
Lijiang Yunnan Chine
photo Daniel Prunier
Copyright © Daniel Prunier

Carabus (Apotomopterus) delavayi tenuimanus Deuve & Imura, 1990
Female adult
Lijiang, Yunnan, Chine
female
photo Daniel Prunier
Copyright © Daniel Prunier

Calosoma (Campalita) chlorostictum Dejean, 1831a: 558
Male adult
La Mandraka, Centre Est, Madagascar; December 1995
Copyright © Denis Potanin

Carabus (Morphocarabus) illigeri illigeri (as bjelasnicensis Apfelbeck, 1902)
Male+Female adult
Serbia, Debelo Brdo, 5.VI.2007, S. Sapaly leg. L=32 mm photo Panin R.
Copyright © Ruslan Panin

Carabus (Archicarabus) monticola fontanae Born, 1906
Male+Female adult
N-Italy,
Piemonte, Bielmonte, h1350 m, 14-15.VI.2010, Pascal Stefani leg.
female, 23 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Morphocarabus) kollari Palliardi, 1825
Female adult
female, 24 mm
Copyright © Boleslav Brezina

Carabus (Morphocarabus) kollari Palliardi, 1825
Male adult
male, 27 mm
Copyright © Boleslav Brezina

Carabus (Trachycarabus) estreicheri Fischer, 1822
Male+Female adult
W UKRAINE, NW Podolian,
Lviv Region, Brody distr., near Yasenov vill., steppe sites, 2-28.V.2010, Panin R. leg.
female, 23mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) estreicheri Fischer, 1822
Male adult

Copyright © Safronov

Carabus (Trachycarabus) estreicheri Fischer, 1822
Female adult
W-UKRAINE, NW Podolian,
Lviv Region, Brody distr., near Yasenov vill., steppe sites, 12.V-2.VI.2007 Panin R. leg.
female, 22mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei Kuster, 1846
Male+Female adult
W.Ukraine.
Transcarpathian region,Vinogradiv distr., near Malaya Kopanya vill., Chorna Gora Mt., h=180m. 7-18.VII.2008. Panin R. leg.
female 33 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei Kuster, 1846
Male adult
W.Ukraine.
Transcarpathian region,Vinogradov distr., near Malaya Kopanya vill., Chorna Gora Mt., h=180m. 7-18.VII.08. Panin R. leg.
male, 28 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei Kuster, 1846 (as ormayi Reitter, 1896) loc.typ.
Female adult
W.UKRAINE,
Transcarpathian reg., near Beregovo,15.V-16.VI.2009 Kanarsky Yu. & Geryak Yu. leg.
female, 30 mm
photo/det. Panin R.
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei Kuster, 1846
Female adult
Carabus (Morphocarabus) rothi hampei
W Ukraine, Zakarpatian reg., Vinogradov
1.7.1989 Avdeev I. leg.
female, 31 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) rothi hampei Kuster, 1846
Female adult
arabus (Morphocarabus) rothi hampei
W Ukraine, Zakarpatian reg., Vinogradov
1.7.1989 Avdeev I. leg.
female, 31 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) spasskianus schestopalovi Plutenko, 1995
Male adult
NE-Kazakhstan,
S-Altay, Sarym-Sakty Mt. h~2500m., Sarym-Sakty riv. 24.VI.1999
male, 23mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus schestopalovi Plutenko, 1995
Male adult
SE Kazakhstan,
S Altay Mts., Sarymsakty mt. rng., Sarymsakty riv.
22.VI.2000 2400m, Nekroshevich S. leg.
male, 20 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) kollari Palliardi, 1825
Male+Female adult
Romania, SW Carpathians,
Banat, Anina Mts. (Muntii Aninei) 8 km NE Oravita, 7.VI.2007, S.I. Budka leg.
female, 34 mm
photo Panin R
Copyright © Ruslan Panin

Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860
Male+Female adult
SW -France,
Pyrenees-Atlantiques, Foret de Sare, XII.2009, Pascal Stefani leg.
female, 31 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860
Female adult
Spain,
Navarra, Sierra de Aralar, Altxueta Mt., 5.V.2006
Anichtchenko A. leg.
L= 17mm
Copyright © Alexander Anichtchenko

Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860
Male adult
Spain,
Pais Vasco, Bera de Bidasoa, Larrun 450m,
9.9.2001 Anichtchenko A. leg.
male, 29mm
Copyright © Alexander Anichtchenko

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as delacrei Tarrier, 1975)
Male+Female adult
NE Spain, Catalonia, Sant Jaume de Frontanyа, h=1000 m, V-VI.2010, Pascal Stefani leg.
female, 38 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as delacrei Tarrier, 1975)
Male adult
Spain,
Catalunya, San Jaume de Frontana,
May 2001
Copyright © Ruslan Panin

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826
Female adult
Auriac alt.
500m. (France : Aude) 25.V.200
Copyright © Ruslan Panin

Carabus (Carabus) arvensis sylvaticus Dejean, 1826
Female adult
France, Centre reg., Eure-et-Loir, Forкt de Montecot, VI.2009, Pascal Stefani leg.
female, 20 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) arcensis sylvaticus Dejean, 1826 (as sugintensis Brunier, 1922)
Male adult
NW -France, Haut-Rhin Depart., Fulleren, XI.2004, Pascal Stefani leg.
Copyright © Ruslan Panin

Carabus (Carabus) arvensis sylvaticus Dejean, 1826 (as liebmanni Lengerken, 1911 )
Female adult
France 67, Weifersweller 350m, Vosges massif
25.8.91
long. 21mm
Copyright © Alexander Anichtchenko

Carabus (Carabus) arvensis sylvaticus Dejean, 1826
Male adult
France 61, Longny-au-Perche. 30.V.1991
male, 19 mm
Copyright © Alexander Anichtchenko

Carabus (Chrysocarabus) olympiae Sella, 1855
Male+Female adult
N-Italy,
Piemonte, Bielmonte, 14-15.VI.1981, Pascal Stefani leg.
female, 34 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Chrysocarabus) olympiae Sella, 1855
Male adult
Italy, Piemonte, Sessera, 2500m 23.III.1987
Copyright © Alexander Anichtchenko

Carabus (Coptolabrodes) haeckeli Brezina & Imura, 1997
Male adult
C China, Shaanxi,
Tsinling Mts., Taibaishan Mt., h=3000m, 33:55N/107:44 E, VI.2004, V.Siniaev & his team
female, 31 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus tscherkessicus Korge, 1964
Male adult
S Russia, NW-Caucasus,
Karachay-Cherkessia,Teberda, Mukhinsky (Mukhu) Mt. pass, h=2750-2800 m , 8.VIII.2010, I.A. Solodovnikov leg.
male, 23 mm

Copyright © Ruslan Panin

Carabus (Megodontus) imperialis Fischer, 1823
Female adult
Carabus (Megodontus) imperialis
photo M. Smirnov
from www.zin.ru
Copyright © Maksim Smirnov

Carabus (Eupachys) glyptopterus Fischer, 1827
Male+Female adult
Russia, SE-Siberia,
Buryat Republic, Ivolginsky Distr., 04.VII.2010, Filippov A.E. leg.
female, 32 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Eupachys) glyptopterus Fischer, 1827
Female adult
Mongolia, 15 km N Bayanteeg, 2000 m, 17.7.2006 Anichtchenko A. leg.
Copyright © Alexander Anichtchenko

Carabus (Megodontus) schoenherri sajanus Breuning, 1927
Male+Female adult
Carabus (Megodontus) schoenherri sajanus Breuning, 1927 Russia, SW Siberia, Krasnoyarsk reg., S of Abakan, Sayanogorsk distr., Cheryomushki vill. env. 07-VII-1994 M. Danilevsky leg. male 27mm, female 31 mm.
Copyright © Safronov

Carabus (Megodontus) schoenherri sajanus Breuning, 1927
Male adult
Carabus (Megodontus) schoenherri sajanus Breuning, 1927
W Sayan Mts, Borus mt. rng., near Cheremushki vill.,
29-31.7.2000 Brinev A. leg.
male, 30mm
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Coptolabrus) formosus brockmanni (as nili Deuve, 2005 PT) Paratype
Male+Female adult
C China, Shaanxi,
Liupan Mts., Guan Shan Mt. h=800 m, , 34:50N/106:48 E, Team V.Siniaev leg.
female, 38 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus fulgens Charpentier, 1825
Male adult
NC-Spain,
Navarra Reg., Sierra de Andнa, IX.2004, Pascal Stefani leg.
male, 30 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus fulgens Charpentier, 1825 (as gerundensis Born, 1916)
Male adult
Spain, Gerona, Garriguella, XI.1990 J. Soler leg.
male, 31mm
Copyright © Alexander Anichtchenko

Carabus (Procerus) gigas gigas Creutzer, 1799
Male adult
Croatia
Copyright © Stanislav Krejcik

Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792
Female adult
NE-FRANCE,
Lorraine Reg., Moselle, 15.II.2010, Pascal Stefani leg.
female, 26 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792
Male adult
Austria,
Weissenbach stt., III.1979, Kretschmer leg.
male, 24mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792 (as aureus)
Male adult
France 28, Saint-Eliph, V.1991
female, 25mm
Copyright © Alexander Anichtchenko

Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792
Female adult
France 28, Saint-Eliph, V.1991
female, 24 mm

Copyright © Alexander Anichtchenko

Carabus (Archicarabus) nemoralis lamadridae Born, 1895 (as layrei Tarrier, 1975)
Male adult
NW-Spain, León Province, Acebo, V.2007, Pascal Stefani leg.
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii zawadzkii Kraatz, 1854
Male adult
Ukrainian Carpathians
Lviv Reg., Skole Distr., near Grebeniv vill., h=1000m 9 - 20.VII.07, Panin R. leg.
male, 31 mm
Copyright © Safronov

Carabus (Acathaicus) alexandrae Semenov, 1887
Male adult
Carabus (Acathaicus) alexandrae alexandrae Semenov, 1897 Holotype
Kwantin and Li-tschara-pu, 22.VI.1885, leg. Potanin
Deposited in Zoolog. Museum St. Petersburg.
male, long. 22 mm
photo B. Brezina
Copyright © Boleslav Brezina

Carabus (Carpatophilus) linnaei linnaei Panzer, 1812
Male adult
Carabus (Carpatophilus) linnaei linnaei Panzer, 1812
Copyright © Safronov

Carabus (Isiocarabus) strandiellus dayongensis Kleinfeld, 1996 HT Holotype
Male adult
Carabus (Isiocarabus) strandiellus dayongensis HT
Copyright © Frank Kleinfeld

Carabus (Isiocarabus) dargei dargei Deuve, 1987 (as kulingensis Holotype) Holotype
Male adult
Carabus (Isiocarabus) dargei dargei (as kulingensis Holotype)
Copyright © Frank Kleinfeld

Carabus (Isiocarabus) dargei yulingensis (as dreuxi yulingensis) Kleinfeld, 2002 HT PT Holotype Paratype
Male+Female adult
Carabus (Isiocarabus) dargei yulingensis (as dreuxi yulingensis) HT PT
Copyright © Frank Kleinfeld

Carabus (Isiocarabus) fiduciarius fiduciarius (as fiduciarius tim) Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Isiocarabus) hunanicola jinxiuensis Imura, 1995 (as dreuxi xinyingensis) HT PT Holotype Paratype
Male+Female adult
Carabus (Isiocarabus) hunanicola jinxiuensis (as dreuxi xinyingensis) HT PT
Copyright © Frank Kleinfeld

Carabus (Isiocarabus) hunanicola liangshanicus (as dreuxi liangshanicus) HT PT Kleinfeld, 1998 Holotype Paratype
Male+Female adult
Carabus (Isiocarabus) hunanicola liangshanicus (as dreuxi liangshanicus) HT PT
Copyright © Frank Kleinfeld

Carabus (Isiocarabus) kiukiangensis orphniopterus Hauser, 1920
Male adult
Carabus (Isiocarabus) kuikiangensis orphniopterus Hauser, 1920
China, Fujian, Yun Ling Shan,
Fokien, 4.6.1998
Copyright © Alexander Anichtchenko

Carabus (Ulocarabus) theanus Reitter, 1895
Male adult
Vakhskiy Mt Range,Saryhosor, Mullonomi,
29.4.1990, S.Zonshteyn leg.
Copyright © Alexander Anichtchenko

Carabus (Tribax) titan titan Zolotarev, 1913
Male+Female adult
Russia,NW Caucasus,
Krasnodar reg.,Chernogorie Mnt. Range, Agulova Balka h=950m V-VII.1999, A.S.Zamotajlov leg.
female,32mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Tribax) titan titan Zolotarev, 1913
Male adult
Carabus (Tribax) titan titan Zolotarev, 1913
Tuapse distr., Anastasievka vill., Semiglavaya mt.,
Pshekaho riv., 20.7.1995
male, 30 mm
Copyright © Alexander Anichtchenko

Carabus (Tribax) titan abadzechus Zamotajlov, 1988
Male adult
Carabus (Tribax) titan abadzechus Zamotajlov, 1988
W Caucasus, Adygea, Asish-Tau Mt.R., vall riv. Bzykha, Fagus forest, h=1450m, 14-22.08.2008, I.A.Solodovnikov, O.Beljavskyi
male, 25 mm

Copyright © Igor Solodovnikov

Carabus (Tribax) steveni steveni Menetries, 1832
Male adult
Carabus (Tribax) steveni steveni Menetries, 1832
Karachaevo-Cherkessia,
Hudes riv., near Tsarskie vorota,
V.2009 Anichtchenko A. leg.
male, 25mm
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Tribax) steveni steveni Menetries, 1832
Male adult
Carabus (Tribax) steveni steveni Menetries, 1832
C Caucasus, Kabardino-Balkaria,
Elbrusskyi distr., riv. Malka, upper reaches riv. Sultan-Bashi, h=2400-2600 m, 26, VII, 2008. Leg. I.A.Solodovnikov.
male, 20,5 mm
Copyright © Igor Solodovnikov

Carabus (Tribax) steveni steveni Menetries, 1832
Female adult
Carabus (Tribax) steveni steveni Menetries, 1832
C Caucasus, Kabardino-Balkaria,
Elbrusskyi distr., riv. Malka, upper reaches riv. Sultan-Bashi, h=2400-2600 m, 26, VII, 2008. Leg. I.A.Solodovnikov.
female 24,5 mm
Copyright © Igor Solodovnikov

Carabus (Tribax) steveni schamyli Hampe, 1852
Female adult
S Russia, N Ossetia, Alagir distr., near Nizhniy Tsey vill.,VII.1981
female, 25mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Tribax) steveni schamyli Hampe, 1852
Male adult
C Caucasus, N Ossetia,
N slope of Pastbistshnyi Mt.R., Tagardon env., h=1100m, 7-9.08.1997, leg. A.G.Koval
male 25,5 mm
Copyright © Igor Solodovnikov

Carabus (Tribax) steveni schamyli Hampe, 1852
Male adult
C Caucasus, N Ossetia,
N slope of Pastbistshnyi Mt.R., Tagardon env., h=1100m, 7-9.08.1997, leg. A.G.Koval
female 27 mm
Copyright © Igor Solodovnikov

Carabus (Tribax) steveni schamyli Hampe, 1852
Male adult
N Osetia, B. Balkary, 1700 m, 2.9.85
Abdurahmanov leg.
male, 24 mm
Copyright © Alexander Anichtchenko

Carabus (Tribax) steveni satanas Semenov, 1896
Male adult
Carabus (Tribax) steveni satanas Semenov, 1896
S Russia, C-Caucasus:
Balkaria, Valley of riv. Cherek Balkarskyi, h=1750-1800m 10.VI-20.VII.1992 Leg. Zamotajlov A.
male, 23 mm
photo Panin R., det. Solodovnikov I.A
Copyright © Ruslan Panin

Carabus (Tribax) steveni satanas Semenov, 1896
Male+Female adult
Carabus (Tribax) steveni satanas Semenov, 1896
Russia, C.Caucasus, Kabardino-Balkaria,
valley of River Cherek Balkarsky, h=1800 m
10.VI-20.VII.1992
A. Zamotajlov leg.
Copyright © Safronov

Carabus (Tribax) puschkini puschkini Adams, 1817
Male adult
Armenia, Shirak Province, Ashotsk, 18.VI.2001, P.V.Kazaryan leg. male 25mm
Copyright © Ruslan Panin

Carabus (Tribax) puschkini ishikawaianus Breuning & Ruspoli, 1970
Male adult

Copyright © Alexander Anichtchenko

Carabus (Tribax) kasbekianus veselyi Gottwald, 1980
Female adult
Carabus (Tribax) kasbekianus veselyi Gottwald, 1980
Caucasus, Georgia, Egrissky Mt.R., riverheads Tekhuri river, h=2400, A.G.Koval leg.; 04.07.1989; 25 mm.; photo A. Menstschicov
Copyright © Anton Menstschikov

Carabus (Tribax) kasbekianus onerosus Belousov & Zamotajlov, 1993 Paratype
Male+Female adult
Carabus (Tribax) kasbekianus onerosus Belousov & Zamotajlov, 1993 Paratypes
Caucasus, W Georgia, Svanetia, right bank o Magana riv.,
800m, 12.V.-4.VII.1991 Zamotajlov & Miroshnikov leg.
male 27mm, female 33mm
photo & collection Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Tribax) edmundi Semenov, 1896
Male adult
Carabus (Tribax) edmundi Semenov, 1896
Copyright © Boleslav Brezina

Carabus (Tribax) circassicus teberdensis Zolotarev, 1913
Male+Female adult
W-Caucasus: Karacaj-Cherkessija,
Abishira–Akhuba Mnt. Range h=2000-2650m, Kyafar-Agur Riv., 14–17.VII.2009, Fominykh D. leg.
female, 25mm
Copyright © Ruslan Panin

Carabus (Tribax) circassicus kubaniensis Semenov, 1896
Male adult
N Caucasus, Krasnodar reg., 30 km S Maykop
august 1993
male, 26 mm
Copyright © Alexander Anichtchenko

Carabus (Tribax) circassicus kubaniensis Semenov, 1896
Male+Female adult
Russia, NW Caucasus,
Krasnodar reg., Chernogorie mts., 24.V-29.VII.1999. A.S. Zamotajlov leg.
female 29mm/male 25mm
Copyright © Ruslan Panin

Carabus (Tribax) circassicus circassicus Ganglbauer, 1886
Male+Female adult
Abkhasia,
W.slope of Gagrinsky Mt., Krepostnaya River source, h=930-980m., A.S.Zamotajlov leg. 27.VII.2000
female, 29 mm/male, 26mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Tribax) circassicus abagonensis Starck, 1894
Male+Female adult
S Russia, NW Caucasus,
Adygea Rep., Mt.Abago, 26.VI.1999
L=24mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Tribax) biebersteini ulgeni Gottwald, 1982
Male adult
S Russia, Karacaj-Cherkessija,
near Kart-Djurt vill., upper Kylan riv. h=1300m, 26-27.V.2009, Fominykh D. leg.
male, 28mm
Copyright © Ruslan Panin

Carabus (Tribax) biebersteini otcharensis Kurnakov, 1970
Male adult
Caucasus, W Georgia, Svanetia, upper reaches of riv. Shoudidi, 1700-2100m, 20-21.V.1986 leg. A. Zamotajlov
Copyright © Evgeniy Chibilov

Carabus (Tribax) biebersteini ingusch Zolotarev, 1913
Male+Female adult
Carabus (Tribax) biebersteini ingusch Zolotarev, 1913
Russia, C.Caucasus, N.Ossetia,
valley of River Khazniaush
10.VI-31.VII.1993
A. Zamotajlov leg.
Copyright © Safronov

Carabus (Tribax) biebersteini constantinowi Starck, 1894
Male adult
Carabus (Tribax) biebersteini constantinowi Starck, 1894
NW-Caucasus, S Russia,
Karacaj-Cherkessija, Nizhniy Arkhyz, 21-30.V.2001, Tikhonov V. leg.
male,30mm
Copyright © Ruslan Panin

Carabus (Tribax) apschuanus apschuanus Rost, 1893
Male+Female adult
Carabus (Tribax) apschuanus apschuanus Rost, 1893
Russia, Caucasus, Krasnodar reg.,
h=1200 m, 29.VII.2000
A. Zamotajlov leg.
Copyright © Safronov

Carabus (Tribax) agnatus agnatus Ganglbauer, 1889
Male adult
NW-Caucasus,
Krasnodar reg., Mostovskoj distr., Skirda Mts. N. slope Yatyrgvart Mt., h=2300-2500m. 09-10.VII.2008, D.Fominyh leg.
male, 23mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus sibiricus Fischer, 1822
Male adult

Copyright © Kirill Makarov

Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886
Male+Female adult
W Ukraine, Ternopol reg., Pidvolochisk distr., Supranivka vill. 3.VII.1908. Leg. Stobetskii. S. Photo R. Panin
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886
Male adult
W Ukraine,
Ternopil reg., Pidvolohcisk distr., Supranivka vill.
male, 26mm
The end of XIX the beginning of XX century
photo Panin R.
coll. SMNH Ukraine, Lviv
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus obliteratus Fischer, 1827
Male adult
Altai mts., Kalbinskii mt.rng.
Ognevka vill. VII.1994, Klimenko A. leg.
male, 28 mm
Copyright © Alexander Anichtchenko

Carabus (Trachycarabus) sibiricus obliteratus Fischer, 1827
Male adult
NE Kazakhstan,
S Altai Mts., Ulbinsky range h~500m.,Turgusun vill., 20.VI.2001
male 25mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus kolosovi Zinoviev, 1997 loc.typ.
Female adult
S. Ural, Bashkiria, near Mesjagutovo vill., 09.07.2001 A. Menstschicov leg.
photo A. Menstschicov
Copyright © Anton Menstschikov

Carabus (Trachycarabus) sibiricus kolosovi Zinoviev, 1997
Female adult
Russia, North-East Bashkortostan. Ai river, near Sikiyaz-Tamak vill.
12.07.1997, Zinoviev E.leg
length: 21,8 mm
Copyright © Evgeniy Zinoviev

Carabus (Trachycarabus) sibiricus haeres (as fossulatus Dejean, 1826)
Female adult
Carabus (Trachycarabus) sibiricus fossulatus Dejean, 1826 NE Ukraine, Kharkov reg.,1974 female, 27mm.
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus haeres (as fossulatus Dejean, 1826)
Male+Female adult
E.UKRAINE, Kharkov reg., Zmiev distr.,in steppe,10-20.V I.2007, D.Kovalchuk leg. female, 26mm/male, 24mm. photo/det. Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus errans Fischer, 1823
Female adult
Carabus (Trachycarabus) sibiricus errans Fischer, 1823
S.Russia, Rostov-Don reg., Avilo-Uspenskaya env.
V.2000
Copyright © Safronov

Carabus (Trachycarabus) sibiricus errans Fischer, 1823
Male adult
SW-Russia,
Rostov reg. env. stanitsa Razdorskaya, A. Ponomarev leg. 9-15.V.2001
male 25mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus errans Fischer, 1823
Male+Female adult
female:SE Ukraine,
Donetsk, 25.VII.2009, A.I.Gubin leg. L=30mm.
male:E Ukraine, Lugansk reg., near Slavyanoserbk, 5-15.VIII.2009, S.Konovalov leg. L=26mm.
photo/det.Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus bosphoranus Fischer, 1823
Female adult
Carabus (Trachycarabus) sibiricus bosphoranus Fischer, 1823
S Ukraine,
Krym Peninsula, Nikitskaya Jaila, S.Mosyanin leg. 10.VIII.2000
female 23mm.
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus bosphoranus Fischer, 1823
Male adult
Carabus (Trachycarabus) sibiricus bosphoranus Fisch. 1823 S Ukraine, Krym Peninsula, Dolgorukovskaya jaila, 25.VIII.1991 I.A.Solodovnikov leg. male, 23mm 
Copyright © Ruslan Panin

Carabus (Trachycarabus) scabriusculus scabriusculus (as erythropus Fischer, 1822)
Male+Female adult
W.Ukraine, NW Podolian Upland , Lviv Reg., Zolochiv distr., near Luky vill., steppe sites, 7.V-8.VI. 2010 leg. Panin R. female, 22mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) scabriusculus scabriusculus (as erythropus Fischer, 1822)
Male adult
W-UKRAINE NW Podolian Upland, Lviv Reg., Zolochiv Distr., near Luka vill., in steppe. Panin R. leg. 1.V - 16.VI.07
Copyright © Safronov

Carabus (Trachycarabus) scabriusculus inapertus Motschulsky, 1850 (loc.typ)
Male+Female adult
NE UKAINE,
Kharkov reg., Zmiev distr., in steppe, 15.VI-14.VII.2009, Kovalchuk D. leg.
female, 20mm
photo/det. Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus planus Gehin, 1885
Male+Female adult
S Russia, NW Caucasus, Karacaj-Cherkessija, near Kart-Djurt vill., upper Kylan riv. h=1300m, 26-27.V.2009 , Fominykh D. leg. L=24mm photo/det. Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus planus Gehin, 1885
Male adult
NW Caucasus, Rostov reg., Avilo-Uspenskaya vill. 22.5.2000 Arzanov Yu. leg. male, 25 mm
Copyright © Alexander Anichtchenko

Carabus (Trachycarabus) planus planus Gehin, 1885
Male adult
S Russia, Krasnodar reg., Anapa distr., B. Utrish vill., env., 11-20.VIII.2008, Khatchikov E. leg. male, 26mm photo Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus campicola Reitter, 1896
Male adult
Krym Peninsula, near Sudak. Leg.Vashchenko S. 1-5.V.2004 27 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus campicola Reitter, 1896
Male+Female adult
S Ukraine, Krym Peninsula, near Perevalnoe vill., in steppe, 15.IV.2009, Shaporinskiy V. leg. female, 28 mm photo/det.Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) mandibularis mandibularis Fischer, 1827
Male adult
NE Kazakhstan, S Altai Mt., Narymskii mt. rng.
Bolshenarymskoe vill. 26.7.2001, 1800m Klimenko leg.
male, 21 mm
Copyright © Alexander Anichtchenko

Carabus (Trachycarabus) mandibularis buchtarmensis Kryzhanovskij, 1953
Male adult
Kazakhstan, Kalbinsky mt. rng., 600m ,
Ognevka vill., 23.6.98 Klimenko A. leg.
male, 23 mm
Copyright © Alexander Anichtchenko

Carabus (Trachycarabus) mandibularis buchtarmensis Kryzhanovskij, 1953
Male adult
NE-Kazakhstan,
S-Altays, Kalbinky Mt. h=500m., near Ognevka vill., 20.VI.1998.A. Klimenko leg.
male, 23mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
Male adult
Burjatia, Dzhida distr., near Beloozersk vill. 4-9.VII.1997 Anichtchenko A. leg.
male, 23 mm
Copyright © Alexander Anichtchenko

Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
Male adult
Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
China, Beijing, Mentougou, 2005;
Photo & det. by liuye

Copyright © Ye Liu

Carabus (Trachycarabus) besseri Fischer, 1822
Male adult

Copyright © Safronov

Carabus (Trachycarabus) besseri Fischer, 1822
Male+Female adult
W Ukraine.NW Podolian,
Lviv reg., Zolochiv distr., Lysa Gora near Stinka vill., steppe sites, 25.IV-9.V.2009 Panin R. leg
female, 30mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) scabripennis scabripennis Chaudoir, 1850
Male adult
Carabus (Tomocarabus) scabripennis scabripennis Chaudoir, 1850
C. Armenia,
Kotayk reg., Yeghvard, Arailer Mt., 16.VI.1999, P.V.Kazaryan leg.
male, 20mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Tomocarabus) scabripennis marthaiellus Rapuzzi & Kleinfeld, 2000 Holotype Paratype
Male+Female adult
Carabus (Tomocarabus) scabripennis marthaiellus Rapuzzi & Kleinfeld, 2000
Copyright © Frank Kleinfeld

Carabus (Tomocarabus) marginalis Fabricius, 1794
Male adult
male, 26 mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) marginalis Fabricius, 1794
Male+Female adult
E Ukraine,
Kharkov reg.,near Vasishchevo vill.10-20.VI.2006 D.Kovalchuk leg.
female 24mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) decolor Fischer, 1823
Female adult
Carabus (Tomocarabus) decolor Fischer, 1823
Copyright © Alexander Anichtchenko

Carabus (Tomocarabus) decolor Fischer, 1823
Male adult
S Russia, NW-Caucasus,
Karacaj-Cherkessija,Skalisty Mnt. Range, Site Banditskoe h=1400m, 2-3.V.2009,
Fominykh D., Bondarenko A.S. leg.
male,17mm
Copyright © Ruslan Panin

Calosoma (Calosoma) cyanescens Motschulsky, 1859
Male adult
China,
Jilin prov, P'an-shih
42° 56' 32" N
126° 03' 22" E
20.06.2009
Local people leg.

Copyright © Anton Menstschikov

Calosoma (Caminara) reitteri Roeschke, 1897
Male adult
W Kazakstan, Ustjurtskij res., Kordon Kenderli; 01.05.2009, Gorbunov P.J. leg.,
26 mm.; photo A. Menstschicov
Copyright © Anton Menstschikov

Calosoma (Caminara) reitteri Roeschke, 1897
Male adult
Turkmenia, Badkhyz,
Akar-Cheshme 9.4.1985
23 mm
Copyright © Alexander Anichtchenko

Calosoma (Caminara) olivieri Dejean, 1831
Male adult
З Казахстан, среднее течение р. Эмба, окр. п. Миялы, П.Ю.Горбунов leg. 02.07.2009; photo A. Menstschicov
Copyright © Anton Menstschikov

Carabus (Morphocarabus) wulffiusi wulffiusi Morawitz, 1862
Male+Female adult
Russia, Far East,
Primorskiy Kray, near Kamenushka vill., 2-3.VIII.1992, K.E.Dovgaylo leg.
female, 24mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) wulffiusi wulffiusi Morawitz, 1862
Female adult

Copyright © Safronov

Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Male adult
Serbia, Majdanpek, Ceremosja 15.6.2001 Z.Zlatic leg. male, 32 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Male+Female adult
NE-Serbia, Krs Mts., Majdanpek env.,h=300m.20-31.V.2006, Pr.Sima leg. female, 32mm/male, 30mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) venustus venustus Morawitz, 1862
Male adult
SE Primorskii reg., Lazovskii Nat.Res., 12 km S Staraya Kamenka vill., near Amerika kor. 43°15N, 134°1E, 19.VII.04 leg. M.& L. Smirnovy photo from www.zin.ru
Copyright © Maksim Smirnov

Carabus (Morphocarabus) tarbagataicus pietrorattii Deuve, 1991(as pseudogebleri Deuve, 1994)
Male adult
Carabus (Morphocarabus) tarbagataicus pietrorattii Deuve, 1991(as pseudogebleri Deuve, 1994)
S Tarbagatai mt.rng., Alexeevka vill.,
600m, 11.5.2001 Klimenko A. leg.
male, 30 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) tarbagataicus pietrorattii Deuve, 1991
Female adult

Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997 HT Holotype
Male adult
Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997 HT
Rusia, SE Tuva, Sangilen mt.rng., Kargy riv., 50°31'N 97°01'E,
1800 m, 29.07.1996, D. Obydov leg.

Copyright © Dmitri Obydov

Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997
Female adult
Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997
Tuva, Akademika Obrucheva mt.rng., 15 km N Cherbi vill.
ur. Chailag 7.VII.1999 A. & N. Menschikovy
female, 22 mm
det. Menschikov A.
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997
Male adult
Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997
Tuva, Akademika Obrucheva mt.rng., 15 km N Cherbi vill.
ur. Chailag 7.VII.1999 A. & N. Menschikovy
male, 21 mm
det. Menschikov A.
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) spasskianus tungus Obydov, 1999 HT Holotype
Male adult
Carabus (Morphocarabus) spasskianus tungus Obydov, 1999 HT
Rusia, N Krasnoyarsk reg., Ilimpeya riv and Dovogpokan riv.
VII.1974, Lunin leg.
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) spasskianus spasskianus Breuning, 1932
Male adult
Siberia, Lake Baikal, Listvianka vill. 25.8.90
Anichtchenko A. leg.
male, 23 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) spasskianus spasskianus Breuning, 1932
Male+Female adult
SE Siberia, Russia
Irkutsk reg., near Pivovarikha vill., 7.V.07, D. Fominyh leg.
male, 24mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus shoriensis Obydov, 1999 HT Holotype
Male adult
Gornaya Shoria, 8.VIII.1978, S. Nikireev leg.

Copyright © Dmitri Obydov

Carabus (Morphocarabus) spasskianus saragaschensis Obydov, 1999
Male adult
W Sayan Mts., Batenevskii Mt.rng., 12 km E Pervomaiskoe vill.,
Enisey riv., mountain steppe, 400m, 5.VII.2000 A. & N. Menschikovy leg.
female, 26mm

Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) spasskianus saragaschensis Obydov, 1999 Holotype
Male adult
Rusia, Khakassia, Saragash, 400 m, 6.VII.1994, D. Obydov leg.

Copyright © Dmitri Obydov

Carabus (Morphocarabus) spasskianus putus Motschulsky, 1846
Male adult
Carabus (Morphocarabus) spasskianus putus Motschulsky, 1846
SW Siberia, Altay Mts., Shebalino distr., Kumalyr env.
02.07.2003, A.Safronov leg.
Copyright © Safronov

Carabus (Morphocarabus) spasskianus microputus O.Berlov & Shilenkov, 1996
Female adult
Tuva, 25 km E Ak-Dovurak 23.VI.2006
A. & N. Menschikovy leg.
female, 24 mm
det. Menschikov A.

Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) spasskianus microputus O.Berlov & Shilenkov, 1996
Male adult
Tuva, 25 km E Ak-Dovurak 23.VI.2006
A. & N. Menschikovy leg
female, 24 mm
det. Menschikov A.

Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) spasskianus microputus O.Berlov & Shilenkov, 1996
Male adult
Rusia, SW Tuva, W Tannu-Ola mt.rng., bass. of Barlyk riv.,
50°29'N 90°44'E, 30.VII.2000, D. Obydov leg.
Copyright © Dmitri Obydov

Carabus (Morphocarabus) spasskianus korostelevi Obydov, 2004loc.typ.
Male adult
E Kazahstan, Ivanovskii Mts., Prohodnoy Belok, 03.06.2000, A. Ivanov leg.

Copyright © Anton Menstschikov

Carabus (Morphocarabus) spasskianus geblerianus Obydov, 1999 Holotype
Male adult
Rusia, Barnaul, 27.5.1965
Copyright © Dmitri Obydov

Carabus (Morphocarabus) spasskianus cracens Obydov, 1999
Male adult
Carabus (Morphocarabus) spasskianus cracens Obydov, 1999
NE Kazakhstan, S Altay Mts., Kalbinskii range,
Ognevka vill., 600m, 18.6.1999 A. Klimenko leg.
male, 22 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) spasskianus cracens Obydov, 1999 Paratype
Male adult
Museum of Institute of Systematic Biology (Daugavpils, Latvia)
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) spasskianus cracens Obydov, 1999
Female adult
Carabus (Morphocarabus) spasskianus cracens Obydov, 1999
NE-Kazakhstan:
S-Altay, Kalbinsky Mt., Ognevka vill., h~600m 18.VI.1999 A. Klimenko leg.
female, 24mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus cracens Obydov, 1999 Holotype
Male adult
Carabus (Morphocarabus) spasskianus cracens HT
E Kazakhstan, Kolbinskii Mt.rng., 600m, near Ognevka vill.,
13-18.07.1999, D. Obydov leg
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Cyclocarabus) belousovi Kabak, 1992
Female adult
Uzbekistan, Karzhantau Mt., Aktash vill.
1500 m, 12.4.1999
female, 23 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
Female adult
Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
Moravia mer.,Vranov, 26-29.06.1979, Dr. Resl leg.
photo A. Menstschicov
Copyright © Anton Menstschikov

Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
Male adult
Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
SE Moravia mer.,Vranov, 26-29.06.1979, Dr. Resl leg.
photo A. Menstschicov
Copyright © Anton Menstschikov

Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
Male adult
Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
S.Bohemia,
Hluboka nad Vltavou, 24.VI-18.VII.1990, S. Budka leg.
male, 27mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799 (as scheidleri floriani)
Male adult
Carabus (Morphocarabus) scheidleri scheidleri (as scheidleri floriani)
A.: Steiermark, Rein Umg. VIII.1983 Kenyery
female, 32 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) excellens excellens Fabricius, 1798
Male adult
NW Podolian
Copyright © Safronov

Carabus (Morphocarabus) excellens excellens Fabricius, 1798
Male adult
Moldova, 2003
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens (as lomnitzkii Reitter, 1896 )
Male adult
W-UKRAINE, NW Podolian, Lviv Reg., Zolochiv Distr., W Ukraine, Lysa Gora steppe massif, 19.V.-7.VI.2007, Panin R. leg. 25 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens Fabricius, 1798 (ab.erythromerus Dejean, 1826)
Male adult
Moldova, Anenii Noi distr., near Garbovets vill. 20.VI.08 Bodnar' I.I.leg. male, 26mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens (as lomnitzkii Reitter, 1896)
Male adult
W Ukraine, NW Podolian, Ternopil reg., Pidvilochisk distr., near Rosokhuvatets & Kamyanki vill., steppe sites, 24.V-15.VI.2008, Panin R. leg.
male, 24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) helleri Ganglbauer, 1893
Male adult
Moravia Mer., Bile Karpaty 28.5.1993 Budka leg. male, 28 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) helleri Ganglbauer, 1893
Male+Female adult
SE Moravia, Strani, V.2002, M.Snizek leg. male bronze colour-26mm male blue colour-28mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi incompsus (as mendax Csiki,1906)
Male+Female adult
W.Ukraine,
Transcarpathian Reg., Rakhov Distr., Chornogora Mt Range, h=1900m. alpine zone, Panin R. leg/det. 28.VI-9.VIII.2008
female, 32 mm
photo endophallus Solodovnikov I.A
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi comptus Dejean, 1831
Male adult
Carabus (Morphocarabus) rothi comptus Dejean, 1831
Carpathians
male, 23mm
coll.SMNH Ukraine ,Lvi
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi alutensis Savulescu, 1972
Male adult
C Romania,
Olt River: Rimnica Vilcea VII.1971, Savulescu N. leg.

Copyright © Igor Solodovnikov

Carabus (Eucarabus) obsoletus carpathicus Palliardi, 1825
Male+Female adult
female, 25 mm
SW-Romania, Banat, Resita - Valiug, 3-8.VI.2007, Krause R. leg.
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus tibiscinus Csiki, 1905
Male+Female adult
female,28 mm
SE Hungary, Csongrбd, Makу env., 1-8.VIII.2004, Krause R., Slovak, Pittner leg
Copyright © Ruslan Panin

Carabus (Tribax) apschuanus pseudoplatessa Gottwald, 1982
Male+Female adult
NW-Caucasus, Karachay-Cherkessia, Teberda: Dzhamagat gorge, the left bank of the riv. Dzhamagat, 10.VII-28.VIII.2010 S.Pavluchuk leg.
female, 27 mm
Copyright © Ruslan Panin

Carabus (Pachystus) hungaricus mingens Quensel, 1806
Male adult
male, 31 mm
SW-Russia: Stavropol Krai, near Stavropol city , Russkiy Les, 11.VI-28.IX.2010, Pavluchuk S. leg.
Copyright © Ruslan Panin

Carabus (Archicarabus) montivagus montivagus Palliardi, 1825
Male+Female adult
SW-Romania: Banat, Caras-Severin Reg., Sasca Montana – Stinapari, 21.V.-5.VI.2010, Krause Rene leg.
female, 26 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) montivagus blandus Frivaldsky, 1865
Female adult
Carabus (Archicarabus) montivagus blandus
Slovacchia, Plastovce,
16.7.1995 Budka leg.
female, 21 mm
Copyright © Alexander Anichtchenko

Carabus (Archicarabus) montivagus blandus Frivaldsky, 1865
Female adult
female, 23 mm
S-Slovakia, Region de Nitra, Plastovce viil., 1.V.2009, Rudolf Kmeco leg.
Copyright © Ruslan Panin

Carabus (Procechenochilus) kokujewi Semenov, 1898
Female adult
RUSSIA, Karachay-Cherkessia, NW Caucasus Mts., Daut Mt. Range, Iskrovtsev - W.Ullu-Kel Mt. pass circus, h=2800-2600m
Copyright © Alexander Anichtchenko

Carabus (Procechenochilus) ? kokujewi Semenov, 1898
Female adult
Russia, NW Caucasus, Karachay-Cherkessia, Karachaevsk distr., near Uchkula vill., valley Uchkulanichi riv. h=2800-2900m, 25-28.VII.2010, Fominykh D. leg.
female, 23 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) starckianus ganglbaueri Reitter, 1888
Male adult
Russia, NW Caucasus, Krasnodar reg., Adygeia distr., Guzeripl env. 20.5.2003
male, 26 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) starckianus ganglbaueri Reitter, 1888
Female adult
NW Caucasus, S Russia, Adygea Rep., Confluence of the river Belaya & Kisha h~700m, 24.V-7.VIII.2010, D.Fominyh leg.
female,32 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus andrzejuscii (as wolffii Dejean, 1826)
Male+Female adult
SW-Romania: Banat, Moldova Noua, Ravenska viil., 21.V.-5.VI.2010, Krause Rene leg. female, 32 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus andrzejuscii (as mehelyi Ganglbauer, 1896 loc. typ.)
Male+Female adult
N-Romania, Rodna Mts., Pasul Setref - Sacel 10km E, h=810 m, 8-19.VI.2009, Krause Rene leg. female, 31mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus andrzejuscii Dejean, 1826 (as pygmaeus Petri, 1912)
Male adult
W Ukraine, NE Carpathians, Transcarpathian Reg., Rakhov Distr., Chornogora Mt Range h=2030m, alpine zone, 13-24.VI.2010 Panin R. leg./det. male, 25 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) regalis regalis Fischer, 1822
Male adult
Carabus (Morphocarabus) regalis regalis Fischer, 1822
S. Ural, Bashkiria, near Mesjagutovo vill., 09.07.2001 A. Menstschicov leg.
photo A. Menstschicov
Copyright © Anton Menstschikov

Carabus (Morphocarabus) regalis jenissoni Dejean, 1831
Male adult
Carabus (Morphocarabus) regalis jenissoni Dejean, 1831
Rusia, Tuva, Todzh, Azas lake, 990 m,
52°24'N 96°33'E, 8.VII.1998, D. Obydov leg.

Copyright © Dmitri Obydov

Carabus (Morphocarabus) regalis jenissoni Dejean, 1831
Female adult
Carabus (Morphocarabus) regalis jenissoni Dejean, 1831
SW Siberia, Altays Mts, Choya distr., Sukhoy Karasuk env.
23.07.2004, A.Safronov leg.
Copyright © Safronov

Carabus (Morphocarabus) regalis eltoni Obydov, 2006 Holotype
Male adult
Carabus (Morphocarabus) regalis eltoni Obydov, 2006 HT
Rusia, Tver distr., near Ilyinskoe vill.,
29-30.VII.2006, D. Obydov leg.
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) odoratus septentrionalis Breuning 1932
Female adult
N. Russia, arctic Urals, Paypudynya Riv., Pompeliz Mt.h=500-700 m.,10-15.07.2002
S. Andreev, A. Romanenko leg.
Copyright © Safronov

Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844: 100
Male adult
Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844 Buriatia, W Hamar-Daban Mts., up riv. M.Zangisan, Baishint Ula Mt. 2300m 23.5.1999 Anichtchenko A. leg. male, 18 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844: 100 Lectotype
Male adult
Carabus (Morphocarabus) odoratus odoratus LT Lectotype Hamar-Gaban Mts. photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844 (as melleus Lapouge, 1909)
Male+Female adult
Russia, S Siberia, East Sayans, Mondy circ., h~2300m., 5-8.VI.2001, R. Mishustin leg. 20mm.
Copyright © Ruslan Panin

Carabus (Semnocarabus) rustemi Kabak, 2009 Paratype
Male adult
Paratype
Copyright © Ilya Kabak

Carabus (Eotribax) valikhanovi bagua Kabak, 2009 Holotype
Female adult
China, Xinjiang, N slope of Narat Mt. R., SSW of Tekes, Kokterek basin, Buryl Riv., 3145 m, 28.07.2009,
42°48'06»N / 81°41'20»E, I. Kabak leg. (ZIN)
Copyright © Ilya Kabak

Carabus (Alipaster) thianshanskii shatyensis Deuve et Tian, 2004 Paratype
Male adult

Copyright © Ilya Kabak

Carabus (Alipaster) thianshanskii sarkhobensis Kabak, 2009 Holotype
Female adult
CH, Xinjiang, N sl. of Khalyktau MR., N Kongurbulak R., H=3600 m, 42°35'50»N / 81°02'30»E, 31.07.2003, I. Kabak leg
(ZIN)
Copyright © Ilya Kabak

Carabus (Cechenotribax) petri alipastroides Kabak, 2009 Holotype
Male adult
E spur of Terskei Alatau Mt. R., 10 km S of Saryzhas Village, 2800-3000 m, 5.VII.1987, I. Kabak: 42°51’00”N / 79°37’00”E – 42°50’30”N / 79°39’00”E)
(ZIN)
Copyright © Ilya Kabak

Carabus (Tomocarabus) convexus s.str. (as pyrenaicus Born, 1907)
Male adult
Pirineos, Canfranc, 1300m, Barranco de Izas
15.6.2005 Anichtchenko A. leg.
male, 16 mm
Copyright © Alexander Anichtchenko

Carabus (Tomocarabus) convexus s.str. (as paganetti Born, 1905)
Male adult
Lazio, Roma, Bosco di Nettuno
22.5.1986 C. Iacovone leg.
male, 20 mm
Copyright © Alexander Anichtchenko

Carabus (Tomocarabus) convexus convexus Fabricius, 1775
Male adult
W.Ukraine,
Transcarpathian region,Vinogradov distr., near Malaya Kopanya vill., Chorna Gora Mt., h=180m. 7-18.VII.08. Panin R. leg.
male 18 mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) convexus s.str. (ab. niger)
Male adult
W Ukraine, NW Podolia, Lviv Region, Zolochiv distr., near Gologirky vill., in steppe 1-19.V.2007 Panin R. leg.
16mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) convexus erzeliki Schweiger, 1962
Male adult
Turkey, Sivas 1560 m. 19.7.1991
male, 19 mm
Copyright © Alexander Anichtchenko

Carabus (Tomocarabus) bessarabicus bessarabicus Fischer, 1823
Male+Female adult
UKRAINE,
Kherson Reg., Golaya Pristan distr. near Rybalche vill. 6.IV.2000
long (female) 20mm
Copyright © Ruslan Panin

Carabus (Tachypus) cristoforii Spence, 1823 (as nicolasi Reitter, 1888)
Male adult
S-France, Hautes-Pyrenees, Pic de Nere h=2200m,15.VII.1999,
male, 14mm
Copyright © Ruslan Panin

Carabus (Piocarabus) sui Imura & Zhou, 1998
Female adult
China, Beijing, Yunmengshan,2005, Liu ye coll.; 35mm
Copyright © Ye Liu

Carabus (Teratocarabus) azrael azrael Semenov & Znojko, 1932
Female adult

Copyright © Alexander Anichtchenko

Carabus (Teratocarabus) azrael mizunumaianus Imura, 1991
Male adult
China, Liaoning, Benxi, Gongchangung
20.VI.2004 Hong
Copyright © Boleslav Brezina

Carabus (Tanaocarabus) sylvosus Say, 1825
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Carabus (Tanaocarabus) sylvosus Say, 1825
Female adult
female, 26mm
Copyright © Boleslav Brezina

Carabus (Tanaocarabus) sylvosus Say, 1825
Male adult
male, 25mm
Copyright © Boleslav Brezina

Carabus (Tanaocarabus) finitimus Haldeman, 1852
Female adult

Copyright © Boleslav Brezina

Carabus (Tanaocarabus) hendrichsi Bolivar y Pieltain, Rotger & Coronado, 1967
Male adult
male, 23mm
Copyright © Boleslav Brezina

Carabus (Tanaocarabus) forreri Bates, 1882
Male adult
Mexico, Durango, Sierra Madre Occ.,
Buenos Aires w., 2700 m, La Ciudad, 14.6.90 Heinz leg.
Copyright © Alexander Anichtchenko

Carabus (Tanaocarabus) forreri forreri Bates, 1882
Female adult
female, 23mm
Copyright © Boleslav Brezina

Carabus (Tanaocarabus) forreri forreri Bates, 1882
Male adult
male, 22mm
Copyright © Boleslav Brezina

Carabus (Tachypus) cancellatus fusus Palliardi, 1825
Male+Female adult
France, Riveres Charente, leg. Laforgue VI.1996
female, 24mm
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus s.str. (as sulinensis Born, 1902)
Male+Female adult
S UKRAINE,
Krym Peninsula, Staryi Krym, 17.V.2002
L= 26 mm.
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus s.str. (as conspersus Lapouge, 1902)
Male+Female adult
E Byelorussia,
Mogilev Reg. Bykhov distr., near Voronino vill., 20.V-3.VI.2007, Komyagunova N. leg.
female, 26mm
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus s.str. (as tuberculatus Dejean, 1826)
Male adult
W Ukraine,NW Podolian,
Ternopil reg., Pidvolochisk distr., forest Bogdanivskiy, near Romanove Selo, 7-24.V.08, R.Yu.Panin & I.A.Solodovnikov leg.
male, 23mm
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus s.str. (as scythicus Schaum, 1857)
Male adult
Ukrainian Carpathians,
Transcarpathian reg., Rakhov Distr., Chornogora Mnt.Range h=1900m. Panin R. leg. 28.VI-9.VII.08
male, 24mm.
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus s.str. (as scythicus Schaum, 1857)
Female adult
Ukrainian Carpathians,
Transcarpathian reg., Rakhov Distr., Chornohora range h=1850m, 29.VI-28.VII.2007 Panin R. leg.
female, 27 mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) odoratus martjanovianus Obydov 1999 HT Holotype
Male adult
Carabus (Morphocarabus) odoratus martjanovianus Obydov 1999
Rusia, Sayan Mts., near Sayanogorsk, 7.VII.1993
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) odoratus magadanicus Obydov 1999 PT Paratype
Male adult
Carabus (Morphocarabus) odoratus magadanicus Obydov 1999 PT
Rusia, Far East, E of Magadan, W Nyuklia vill.
VII.1997, Yu. Marusik leg.
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) odoratus krugeri Obydov 1999 PT Paratype
Female adult
Carabus (Morphocarabus) odoratus krugeri Obydov 1999 PT
Rusia, Tuva, Akademika Obrucheva Mt.rng., 2100, 51°56'N 095°32'E,
Koptu pass, tundra 2.VII.1998, D. Obydov leg.
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) odoratus kamtchatensis Breuning 1942
Male adult
Carabus (Morphocarabus) odoratus kamtchatensis Breuning 1942
Rusia, Kamchatka peninsula, VII.21
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) odoratus dohrni Gebler, 1847
Male adult
Carabus (Morphocarabus) odoratus dohrni Gebler, 1847
Rusia, Kuznetskii Alatau, Taizhe-Su riv., 53°55'N 89°10'E,
1250-1650 m, 25.VI-30.VII.2002, V. Murzin and V. Sinyaev leg.
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) odoratus dabanensis Shilenkov 1996 loc.typ.
Male+Female adult
Carabus (Morphocarabus) odoratus dabanensis Shilenkov 1996 loc.typ.
Buryatia, Hamar-Daban mt. rng., Snezhnaya riv.,
4.6.2007 Anichtchenko A. leg.
male 17 mm, female 19 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) odoratus czernyscheviellus Obydov, 2002 PT Paratype
Female adult
Carabus (Morphocarabus) odoratus czernyscheviellus PT
Rusia, SE Transbaikalia, Chita reg., Nerchinsko-Kugenskii Mt.rng.,
near Vershino-Darasunskii vill., Bogatui Mt. 1221 m.,
15.VII.2002, D. Obydov leg.
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) odoratus bargusinicus Shilenkov, 1996
Male+Female adult

 Carabus (Morphocarabus) odoratus bargusinicus Shilenkov, 1996
 Russia, N Burajatia,
 Barguzinskiy Mts., the upper reaches of M.Chiverkuy riv., h=2000m, Vashchenko S. leg.
 female, 21mm/male,19mm
 photo Panin R.

Copyright © Ruslan Panin

Carabus (Morphocarabus) odoratus antropovi Shilenkov 1996
Male+Female adult
Russia, S Tuva, Khorumnug -Tayga mts. Range, h=1600-2300m, Buren riv. valley, 4-26.VI.2000 Vashchenko S & A. leg female, 20mm/male,18mm photo Panin R.
Copyright © Ruslan Panin

Carabus (Morphocarabus) odoratus ?antropovi Shilenkov 1996
Male adult
S Tuva, Horumnut-Taiga Mt.rng., Bureya riv., 2000 m, tundra, Vaschenko S. leg. 18 mm
Copyright © Alexander Anichtchenko

Carabus (Tachypus) auratus auratus Linne, 1761
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Carabus (Tachypus) auratus s.str. (as brunieri Barthe, 1921)
Male adult
SC-FRANCE, Aude, Rennes les Bains, 6.VII.1989, Sunard A. leg.
male, 26mm
Copyright © Ruslan Panin

Carabus (Tachypus) auratus auratus Linne, 1761
Female adult
30 mm
Germania, Oberfranken, Pegnitz
leg.det. U.Schmidt, 1972
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Carabus (Tachypus) auratus lasserrei Doue, 1855
Male adult
SC-FRANCE, Midi-Pyrуnуes reg. Tarn, Saint Paul cap de Joux, 24.V.2003, A. Laforgue leg.
male, 24mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) monilis Fabricius, 1792
Male adult
France 39, Foncina le Haut 300m, 23.6.1979; 29 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) monilis Fabricius, 1792 (as taunicus)
Male adult
W Germany, Vogelsberg Mt., 700m, 13.5.1985
male, 24 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) monilis Fabricius, 1792 (as interpositus Géhin, 1880)
Male adult
NW France,
Basse-Normandie, Orne, Forкt d'Йcouves, V.2009, Pascal Stefani leg.
male, 29mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) monilis (as interpositus Gehin, 1880)
Male+Female adult
NW France,
Haute-Normandie reg., Bois Les Loges, Etretat, Seine-Maritime,10.V.1988
female, 28mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) monilis (as amoenus Baudet-Lafarge, 1836)
Female adult
C France,
Puy-de-Dome, Col de la Croix Saint Robert h=1300m, VII.2008, Pascal Stefani leg.
female, 24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) monilis (as helveticus Heer, 1837 Holotype) Holotype
Male adult
Carabus (Morphocarabus) monilis (as helveticus Heer, 1837 Holotype)
?hybrid of monilis/cancellatus
Museum of Zurich
photo Anichtchenko
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) monilis (as dupeuxi Deuve, 1974)
Male+Female adult

France,
Charente, Foret de Braconne VII.1998, Laforgue leg.
famale, 27mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) michailovi Kabak, 1992
Male adult
NE Kazakhstan, S Altay Mts., Sarymsakty mt. rng.,
Sarymsakty riv., 2400 m, 22.6.2000 Nekroshevich S. leg.
male, 18 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) michailovi Kabak, 1992
Male adult
NE Kazakhstan, S Altay Mts., Sarymsakty mt. rng.,
Sarymsakty riv., 2400 m, 22.6.2000 Nekroshevich S. leg.
male, 17 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) michailovi Kabak, 1992
Male adult
NE-Kazakhstan,
S Altai Mts.,Sarym-Sakty Range, Sarym-Sakty riv., h=2400m. 21.VI.1999., A.Klimenko leg.
male,18mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) mestscherjakovi telezkianus Shilenkov, 1996
Male adult
Carabus (Morphocarabus) mestscherjakovi telezkianus Shilenkov, 1996
Rusia, Altai, S shore of Teletskoe lake, 5.VII.1968
Copyright © Dmitri Obydov

Carabus (Morphocarabus) mestscherjakovi telezkianus Shilenkov, 1996
Male adult
Carabus (Morphocarabus) mestscherjakovi telezkianus Shilenkov, 1996
SW Khakasia, Abakanskii Mt.rng., Choochek Mt., 20 km SSE Mrassu vill.,
1800m, 7-19.7.1999 Lomakin D. leg.
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
Male adult
W Sayan mts., Kopen mt., Dzhabash riv., 75 km
S from Arbaty vill., 2235m, 26-30.6.2003 Brinev A. leg.
long. 17 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
Male adult
Russia, S Siberia,
S Khakassia, N Tuva reg., Sailyg-Khem-Taiga mts. range h=2400m, 10.VII.2004
male, 18mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
Male+Female adult
Russia, S Siberia,
S Khakassia, N Tuva reg., Sailyg-Khem-Taiga mts. range h=2000-2600m,10-30.VI.2005, S.Vashchenko & S.Alexeenko leg.
male,17.5mm/female,19mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Morphocarabus) mestscherjakovi pseudoodoratus Brinev, 2002 Paratype Paratype
Male adult
Krasnojarsk reg., Aradan distr., 5 km SW Aradan vill., Us riv., forest, 52є32`N 93є27`E 9-12.VII.1998 A.Brinev leg male, 18 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) mestscherjakovi pseudoodoratus Brinev, 2002 Paratype Paratype
Female adult
Krasnojarsk reg., Aradan distr., 5 km SW Aradan vill., vall. of Us riv., A.Brinev leg. female, 20 mm photo & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
Male adult
Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
W Sayan, Olenya rechka
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
Male adult
Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
W Sayan, Kulumys Mt.R., env. Oiskoje Lake, mountain tundra, 12-14. 07.1999 A.Semin leg.
male
L=17,0 mm
Copyright © Maksim Smirnov

Carabus (Morphocarabus) kozhantschikovi martjanovi Lutshnik, 1924
Female adult
Khakassia, 1800 m, near Cheremushki vill., 12.6.-20.7.2000
female, 21 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
Male adult
W Sayan Mts., Kazhyr-Suk riv., 80 km W Oleniya
Rechka, W part of Oiskii mt. rng., 30.6.-7.7.1999
2100 m, Brinev A. leg.
male, 19 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
Male adult
W Sayan, low fl. Kazyr-Suk riv., 80km W meteostation Olenia Rechka, W part of Oiskii mt.rng., 30.VI-7.VII.1999 2100m, Brinev A. leg.
male, 20mm
det. Brinev A.
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 18789 (as viridipennis Kraatz, 1886, nec Lewis,
Male adult
E Rusia, Khabarovsk reg.,
near Birobidzhan VI.1991
male, 26 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
Male adult
Rusia, S Primorie, Vladivostok distr., Razdolnaya riv., near Kluchevoi vill., 10-14.VII.1988, S. Muhanov leg.; Obydov det. as tristiculus Kraatz, 1879
Copyright © Dmitri Obydov

Carabus (Morphocarabus) hummeli cf pusongensis Imura, 1993
Male adult
China, Beijing, Wulingshan mountain, 1985
Copyright © Ye Liu

Carabus (Morphocarabus) hummeli ssp.
Male adult
Carabus (Morphocarabus) hummeli ssp.
China, East Inner Mongolia, Chifen, 2006, Shi H.L. leg.
Photo & det. by liuye
Copyright © Ye Liu

Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
Male adult
China, Jinling pro.;
Photo by liuye
det. Anichtchenko A.
Copyright © Ye Liu

Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
Male adult
Russia, Far East,
Primorskiy Kray, near Kamenushka vill., 2-3.VIII.1992, K.E.Dovgaylo leg.
male, 27mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) hummeli pusongensis Imura, 1993
Male adult
S Primorie, Chernye gory, Poima riv.,
30.7-5.8.1999 Anichtchenko A. leg.
male, 26 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) hummeli nevelskii Shilenkov, 1996
Female adult
Rusia, S Sakhalin isl., VII.1993, G. Vasiliev leg.

Copyright © Dmitri Obydov

Carabus (Morphocarabus) hummeli nevelskii Shilenkov, 1996
Male+Female adult
Russia, Sakhalin isl.,
Makarov distr., Lesnaja riv., VII.1992, M. Nesterov leg.
female,23mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) hummeli hummeli Fischer, 1823
Male adult
Carabus (Morphocarabus) hummeli hummeli Fischer, 1823
Rusia, Chita reg., Darasun, VII.1991
I. Avdeev leg.
male, 23 mm
Copyright © Alexander Anichtchenko

Carabus (Sphodristocarabus) varians varians Fischer, 1827
Male+Female adult
female: Transcaucasia, Armenia, Amasia distr., Karakhachskiy mountain pass h=2270m, 3.VIII.1985 P.V.Kazaryan leg., 22mm male: Armenia, Amasia distr., Arpalach vill. VI.1999, P.V. Kazaryan leg.
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) varians rugatus Breuning, 1934
Female adult

Copyright © Ruslan Panin

Carabus (Sphodristocarabus) blumenthaliensis basilewskianus Breuning & Ruspoli, 1970
Male adult
SW Georgia, Adzharia 10 km Baku 2000m
18.6.98
25 mm
Copyright © Alexander Anichtchenko

Carabus (Sphodristocarabus) blumenthaliensis basilewskianus Breuning & Ruspoli, 1970
Male+Female adult
NE-Turkey,
Artvin Prov., Hopa vers Borcka, Cankutaran Gecidi h=800m, 17.VI.2007 leg. D. Prunier
female, 24mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) varians janthinus s.str.
Male+Female adult
NW-Caucasus, Russia, Krasnodar reg.,Tuapse distr., Mt.Semashkho 04.V.2008, D.Fominyh leg. female, 24mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) varians armeniacus Mannerheim, 1830
Female adult
NE Turkey, Artvin Prov., Borçka, Macahel Gecidi h=1450m, 18.VI.2007, D.Prunier leg. female, 26mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) tokatensis almushensis Mandl, 1975
Male adult
Turkey, Sivas, 20 km N Zara, N Arapca, 1750m; 11.VII.1993
Copyright © Alexander Anichtchenko

Carabus (Sphodristocarabus) scovitzi czipkai Heinz, 1980
Male adult
Turkey, Artvin, Kafkasor, 1600 m
4.7.1996 Staven and Skoupy leg.
male, 22 mm
Copyright © Alexander Anichtchenko

Carabus (Sphodristocarabus) macrogonus simonianus Breuning & Ruspoli, 1970
Female adult
Turkey, Ordu, Golkoy, 1220 m, 19.7.91
Copyright © Alexander Anichtchenko

Carabus (Sphodristocarabus) macrogonus resplendens Breuning, 1934
Female adult
Turkey; Cayirkent Samsun 12.VII.1996
Copyright © Alexander Anichtchenko

Carabus (Sphodristocarabus) georgia Cavazzuti, 1984
Male adult

Copyright © Boleslav Brezina

Carabus (Sphodristocarabus) bohemani bohemani Menetries, 1832
Female adult
S Azerbaydzan, Lenkoran distr., Vazha riv.,
2 km NW Dashdatuk vill. 38º40*09N 48º42*06E
17.4.2004 Makarov K. leg.
female, 22 mm
Copyright © Alexander Anichtchenko

Carabus (Sphodristocarabus) bohemani bohemani Menetries, 1832
Male adult
Azerbaijan,
Talysh Mts., Polikesh, h~1800m.,2-6.VI.2001 Dolin V.G. leg.
male, 20mm.
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) hollbergi Mannerheim, 1827
Male adult
N Azerbaijan,
4km W Ismayilli, h=700m, 9.VII.2008, Pljushch I.G. leg.
male,22mm
Copyright © Ruslan Panin

Carabus (Semnocarabus) transiliensis Semenov, 1896
Male adult
Kirgizstan, S slope Kungei Ala Too
Oi-Tal riv. 3000m 14.7.1999
13 mm
Copyright © Alexander Anichtchenko

Carabus (Semnocarabus) transiliensis Semenov, 1896
Male+Female adult
E KAZAKHSTAN,
Bolshoe Almaatinskoe Lake,30.VI.1992
15mm
Copyright © Ruslan Panin

Carabus (Semnocarabus) transiliensis Semenov, 1896
Male adult
SE-KAZAKHSTAN,
Ketmen Mts.,N-slopes h=2250-2500m.1-5.VI.1990
16mm
Copyright © Ruslan Panin

Carabus (Semnocarabus) regulus regulus Dohrn, 1882
Male adult
photo M. Smirnov from www.zin.ru
Copyright © Maksim Smirnov

Carabus (Semnocarabus) regulus regulus Dohrn, 1882
Male adult
E Kirgizia, Kaindy-Ketta mts. rng. Sary-Dzhas riv.
Maidadyr near Enelchek vill. 6-8.7.1999 Klimenko leg.
14 mm
Copyright © Alexander Anichtchenko

Carabus (Semnocarabus) erosus erosus Motschulsky, 1865
Male adult
Kirgizia, S Kungei Ala-Too, Kichi-Yuryukty
2800m 1.7.1999 Subankulov leg.
male, 21 mm
Copyright © Alexander Anichtchenko

Carabus (Semnocarabus) erosus corax Semenov, 1896
Male adult
SE Kazakhstan, Ili riv., Bakanas vill.,
1-5.5.1999 Klimenko A. leg.
Copyright © Alexander Anichtchenko

Carabus (Semnocarabus) carbonicolor carbonicolor Morawitz, 1886
Male adult
E Kirgizia, Kaindy-Ketta mts. rng., Sary-Dzas riv.,
Maidadyr near Enelchek vill., 6-8.7.99 Klimenko leg.
Copyright © Alexander Anichtchenko

Carabus (Semnocarabus) carbonicolor callosus Semenov, 1896
Male adult
Kazakhstan, Bol. Alma-Atinka riv. 1600m 1.6.1999 Klimenko A. leg.
male, 19 mm
Copyright © Alexander Anichtchenko

Carabus (Scambocarabus) sculptipennis Chaudoir, 1887
Male adult
China, Beijing, Xiaolongmeng, 2005, Liu ye coll.;
22mm; photo by Liuye
Copyright © Ye Liu

Carabus (Scambocarabus) kruberi kruberi Fischer, 1822
Male adult
Rusia, Chita reg., 23 km SW Borzia, Borzinskoe lake,
1.VI.2002 Anichtchenko A. leg.
Copyright © Alexander Anichtchenko

Carabus (Scambocarabus) kruberi Fischer, 1822
Male adult
China, Beijing, labagoumeng, 2004, Li fei coll.;
21mm; photo by Liuye
Copyright © Ye Liu

Carabus (Morphocarabus) hummeli biamensis Obydov, 2007 Paratypus Paratype
Male adult
Russia, North Primorie, B. Svetlovodnaya riv. (Biame riv.), 40 km from source, 30.07.1976, A. Klitin leg.
Copyright © Dmitri Obydov

Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
Female adult
E.RUSSIA, Primorsky ter., Chuguevsky distr., 30 km ESE from Chuguevka, 9-14.VIII.2007
Copyright © Nil Belonogov

Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
Male adult
E.RUSSIA, Primorsky ter., Chuguevsky distr., 30 km ESE from Chuguevka, 9-14.VIII.2007
Copyright © Nil Belonogov

Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 18789 (as viridipennis Kraatz, 1886, nec Lewis,
Female adult
Primorskii reg., near Kovalerovo
VII.1990 Evstigneev A. leg.
female, 25 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) hummeli ? middendorfi Menetries, 1851
Male adult
Carabus (Morphocarabus) hummeli ? middendorfi Menetries, 1851
Rusia, Khabarovsk reg., Ohotsk sea, Port Ayan, 08.24
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) henningi pelevini Obydov 2006
Female adult
NE Kazakhstan, S Altai mt.rng., Sarymskiy mt. rng., Sarymsakty riv., 2400m, 22.VI.2000 Nekroshevich S. leg. female 17mm det. Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) henningi pelevini Obydov 2006 HT
Male adult
Carabus (Morphocarabus) henningi pelevini Obydov 2006 HT
Kazakhstan, S Altai, Sarym-Sakty Mt.rng., 3000-3400 m,
18-20.VI.1998, A. Klimenko leg.
Copyright © Dmitri Obydov

Carabus (Morphocarabus) henningi peczoricus Obydov 2000 HT Holotype
Male adult
Carabus (Morphocarabus) henningi peczoricus Obydov 2000 HT
Rusia, Komi republic, Kosiyu River mouth, (Pechora basin) 21.VII.1980
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) henningi oviformis Beheim & Breuning, 1943
Male adult
Carabus (Morphocarabus) henningi oviformis Beheim & Breuning, 1943
Rusia, Krasnoyarsk reg., Taimyr AO, near Norilsk,
07.2004, M. Shestopalov leg.
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) henningi melyachi Shilenkov & Menstshikov, 1996
Male adult
Carabus (Morphocarabus) henningi melyachi Shilenkov & Menstshikov, 1996
C Altai mts, Seminskii mt.rng., Katun riv., 3km S Ust-Sema vill.
4.VII.2002 A. & N. Menschikovy
male, 17mm
det. Menschikov A.
photo Anichtchenko
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) henningi henningi Fischer, 1817
Male adult
S. Ural, Bashkiria, near Mesjagutovo vill., 09.07.2001 A. Menstschicov leg.
Copyright © Anton Menstschikov

Carabus (Morphocarabus) henningi henningi Fischer, 1817
Male adult
SE Siberia, Russia, Irkutsk reg., near Pivovarikha vill., 7.V.07, D. Fominykh leg. female, 21mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) henningi hamarensis Shilenkov, 1996
Female adult
S Siberia, Hamar-Daban mts., Snezhnaya riv., 1-6.VI.2007, Anichtchenko A. leg. & det., 20 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) henningi hamarensis Shilenkov, 1996
Female adult
Carabus (Morphocarabus) henningi hamarensis Shilenkov, 1996 loc.typ.
Copyright © Alexander Anichtchenko

Carabus (Rhigocarabus) xiei xiei Deuve, 1992
Male adult
male, 18mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) xiei xiei Deuve, 1992
Female adult
female, 19mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) xiei xiei Deuve, 1992
Male adult
male, 20mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) xiei xiei Deuve, 1992
Male adult
male, 17mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) xiei sanggarparicus Deuve, 1996 Holotype Paratype
Male adult
Holotype
male, 19mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) xiei sanggarparicus Deuve, 1996
Female adult
Paratype
female, 20mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) xiei barkamensis Deuve, 1994 Holotype Paratype
Male adult
Holotype, male, 19mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) xiei barkamensis Deuve, 1994
Male adult
Paratype, female, 19mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) xiangchengicus qingmaicus Deuve, 2007
Male adult
male, 15mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) xiangchengicus qingmaicus Deuve, 2007
Female adult
female, 18mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) xiangchengicus qingmaicus Deuve, 2007
Female adult
female, 16mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) xiangchengicus sangpiensis Deuve, 2007 Paratype
Male adult
male, 15 mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) thais thais Heinz, 1997
Male adult
male, 16mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) thais thais Heinz, 1997
Male adult
male, 19mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) thais thais Heinz, 1997
Male adult
male, 20mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) subindigestus subindigestus Deuve, 1994
Female adult
female, 15mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) subindigestus subindigestus Deuve, 1994
Male adult
male, 16mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) subindigestus subindigestus Deuve, 1994
Male adult
male, 15mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) subindigestus subindigestus Deuve, 1994
Male adult
male, 16mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) sehnali Brezina & Hackel, 2006 Holotype Paratype
Male adult
male, 20mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) sehnali Brezina & Hackel, 2006
Female adult
female, 23mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) roborowskii roborowskii Semenov, 1887
Male adult
China, E Tibet, Chegarla 4800m, 20 km E
of Rioche, 26.6.97 A. Wrzecionko leg.
Copyright © Alexander Anichtchenko

Carabus (Rhigocarabus) rhododendron Deuve & Imura, 1991
Male adult
male, 18mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) rhododendron Deuve & Imura, 1991
Female adult
female, 19mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) rhododendron Deuve & Imura, 1991
Male adult
male, 17mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) quindecim Cavazzuti, 2002 Holotype
Male adult
Holotype, male, 15mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) pseudopusio Deuve, 1996 Paratype
Male adult
male, 15mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) pseudopusio Deuve, 1996
Female adult
female, 16mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) poeta Semenov, 1898 Holotype Paratype Lectotype
Male adult
male, 17mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) poeta Semenov, 1898
Male adult
male, 18mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) poeta (as korsakowi Semenov, 1898) Holotype
Female adult
Holotype, female 18 mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) poeta (as korsakowi Semenov, 1898) Syntypes
Female adult
female, 16mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) poeta (as korsakowi Semenov, 1898) Lectotype
Male adult
male, 17mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) pepek Imura & Kalab, 2006 Paratype
Male adult
male, 17mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) pepek Imura & Kalab, 2006 Paratype
Female adult
female, 18mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) morawitzianus breviculus Cavazzuti & Ratti, 1998 Paratype
Male adult
male, 15 mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) maleki Deuve, 1991 Paratype
Male adult
male, 14mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) maleki Deuve, 1991
Female adult
female, 15mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) maleki Deuve, 1991 Paratype
Male adult
male, 15mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) itzingeri rugulosior Deuve, 1996 Paratype Paratype
Male adult
male, 18mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) itzingeri itzingeri Breuning, 1934
Male adult
male, 19mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) itzingeri itzingeri Breuning, 1934
Female adult
female, 20mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) itzingeri kozaburoi Imura, 1999 Paratype
Male adult
male, 16mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) itzingeri kozaburoi Imura, 1999 Paratype
Female adult
female, 18mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) handelmazzettii handelmazzettii Mandl, 1955
Male adult
male, 18mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) handelmazzettii handelmazzettii Mandl, 1955
Female adult
female, 20mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) handelmazzettii handelmazzettii Mandl, 1955
Female adult
female, 21mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) handelmazzettii habaicus Deuve, 2005 Paratype
Male adult
male, 16mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) pusio pusio s.str.
Male adult
male, 16mm
(as hylonomus)
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) pusio pusio s.str.
Female adult
female, 17mm
(as hylonomus)
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) gracilicollis gracilicollis Semenov, 1887
Male adult
male, 15mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) gracilicollis gracilicollis Semenov, 1887
Female adult
female, 16mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) gigoloides priapeius Cavazzuti, 2006
Male adult
male, 20mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) gigoloides jone Brezina & Hackel, 2004 Holotype Paratype
Male adult
male, 19mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) gigoloides jone Brezina & Hackel, 2004
Female adult
female, 20mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) gigolo (as qilingianus Cavazzuti, 2006)
Male adult
18mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) gigolo Heinz & Brezina, 1996 Paratype
Male adult
20mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) gigolo Heinz & Brezina, 1996 Paratype
Male adult
20 mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) gigolo Heinz & Brezina, 1996 Paratype
Female adult
21 mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) gentleman Brezina & Hackel, 2004 Holotype
Male adult
male, 19mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) gentleman Brezina & Hackel, 2004 Paratype
Female adult
female, 20mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) dacatraianus Deuve, 1995
Male adult
China, E Tibet, Angenla 5000-5100m
45 km W of Bachen 29.6-2.7.1997 Wrzecionko leg.
16 mm
Copyright © Alexander Anichtchenko

Carabus (Rhigocarabus) choui Deuve, 1989
Male adult
male, 16mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) choui Deuve, 1989
Female adult
female, 17mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) cateniger cateniger s.str. (as sommitalis)
Male adult
male, 16 mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) cateniger cateniger Morawitz, 1886
Female adult
female, 19mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) cateniger cateniger Morawitz, 1886
Male adult
male, 18mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) cateniger saltuensis Cavazzuti, 2006 Paratype
Female adult
female, 19mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) cateniger dabanicola Deuve, 1998
Male adult
male, 16mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) cateniger dabanicola Deuve, 1998
Female adult
female, 18mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) buddaicus bocsiki Deuve, 2007 Holotype
Male adult
male, 20 mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) buddaicus angenlaicus Deuve, 2007 Paratype
Male adult
male, 18 mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) broukpytlik Brezina & Hackel, 2004 Holotype Paratype
Male adult
male, 19mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) broukpytlik Brezina & Hackel, 2004
Female adult
female, 20mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) broukpytlik Brezina & Hackel, 2004
Male adult
male, 16mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) morawitzianus breviculus Cavazzuti & Ratti, 1998 Paratype
Male adult
China, N Tibet, Tangula Shan Mts., Tangula
pass 5100-5300m, 8-11.7.1997 A. Wrzecionko leg.
Copyright © Alexander Anichtchenko

Carabus (Rhigocarabus) batangicus fickleri Korell & Richter, 1996 Paratype
Male adult
male, 16 mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) poeta Semenov, 1898
Male adult
male, 17mm
Copyright © Boleslav Brezina

Carabus (Piocarabus) reitterianus Breuning, 1934
Male adult
China, Gansu, ca 25 km E Xiahe 2700-2900m, Wagatan Timber 30.7.-2.8.1993 Heinz leg. male, 23 mm
Copyright © Alexander Anichtchenko

Carabus (Piocarabus) nanwutai Kleinfeld, Korell & Wrase, 1996 Holotype Paratype
Male adult
China, Shaanxi, Qing Ling Shan, track Hou Zen Zi vill. to Taiban Shan, 2500m, 27-29.6.1998 mixed forest male, long. 19 mm
Copyright © Alexander Anichtchenko

Carabus (Piocarabus) nanwutai Kleinfeld, Korell & Wrase, 1996
Male+Female adult

Copyright © Frank Kleinfeld

Carabus kitawakianus kitawakianus Imura, 1993
Male adult
China, Shaanxi, Tschinling mts.,
Taibeichan Nat. res. 2000m 5-15.8.2005
male, 27 mm
Copyright © Alexander Anichtchenko

Carabus (Pseudotribax) ferghanicus Breuning, 1933
Male adult
male, 29mm
Copyright © Boleslav Brezina

Carabus (Pseudotribax) validus Kraatz, 1884
Male adult
male, 31mm
Copyright © Boleslav Brezina

Carabus (Pseudotribax) validus Kraatz, 1884
Female adult
female, 32mm
Copyright © Boleslav Brezina

Carabus (Pseudocranion) sackeni sackeni Semenov, 1898
Male adult
male, 23mm
Copyright © Boleslav Brezina

Carabus (Pseudocranion) sackeni remondianus Deuve, 1996 Paratype
Female adult
Museum of Institute of Systematic Biology (Daugavpils, Latvia)
Copyright © Alexander Anichtchenko

Carabus (Pseudocranion) labrangicus macilentior Deuve, 1996
Female adult
female, 21mm
Copyright © Boleslav Brezina

Carabus (Pseudocranion) gansuensis jedompansis Deuve, 1989
Male adult
China, Sichuan, Langmusi 3200m 7.1999
Copyright © Alexander Anichtchenko

Carabus (Pseudocranion) fumigatus fumigatus Semenov, 1898 Lectotype
Male adult
male, 22mm
Copyright © Boleslav Brezina

Carabus (Pseudocranion) fumigatus fumigatus Semenov, 1898 Syntypes
Female adult
female, 22mm
Copyright © Boleslav Brezina

Carabus (Morphocarabus) gebleri ultimus Obydov, 1999
Female adult
Carabus (Morphocarabus) gebleri ultimus Obydov, 1999 PT
E Kazakhstan, Altai, near Zyrjanovska, 6.VII.1995, D. Obydov leg.
female
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) gebleri ultimus Obydov, 1999 Holotype
Male adult
Carabus (Morphocarabus) gebleri ultimus Obydov, 1999 HT
E Kazakhstan, Altai, near Zyrjanovska, 6.VII.1995, D. Obydov leg.
male
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) gebleri gebleri Fischer, 1817
Female adult
Carabus (Morphocarabus) gebleri gebleri Fischer, 1817
Russia, Altai, Zmeinogorsk
female
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) gebleri gebleri Fischer, 1817
Male adult
Carabus (Morphocarabus) gebleri gebleri Fischer, 1817
Rusia, Altai, Zmeinogorsk
male
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) chaudoiri zaikai Obydov, 1999 PT Paratype
Female adult
Carabus (Morphocarabus) chaudoiri zaikai Obydov, 1999 PT
Rusia, Tuva, Akademik Obruchev Mt. rng., 2100 m;, 52°00'N 95°34'E,
Sainak pass, tundra, 2.VII.1998, D. Obydov leg.
female,
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) chaudoiri juliae Obydov, 2002
Male adult
male, 18mm
Copyright © Boleslav Brezina

Carabus (Morphocarabus) chaudoiri czadanicus Obydov, 1997 HT Holotype
Male adult
Carabus (Morphocarabus) chaudoiri czadanicus Obydov, 1997 HT
Rusia, Tuva, near Hadyn vill., 28-30.VI.1994, R. Shmarzh leg.
male,
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) chaudoiri beladici Obydov, 2001
Male adult
S-Tuva
East Tannu-Ola,Biche-Sailig r.h=2100m, 15.VII.1996, D.E.Lomakin leg.
Copyright © Anton Menstschikov

Carabus (Morphocarabus) chaudoiri beladici Obydov, 2001 PT Paratype
Female adult
Carabus (Morphocarabus) chaudoiri beladici Obydov, 2001 PT
Rusia, Tuva, E Tannu-Ola Mt.rng., near Kara-Hol lake, 29-31.V.2000, M. Beladzhik leg
female,
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Morphocarabus) aeruginosus zyrjanovskianus Shilenkov & O.Berlov, 1996
Male+Female adult
NE KAZAKHSTAN,
S-Altays, Ulbinsky Mt. h=900m., Bobrovka vill., 24.VI.01, A. Klimenko leg.
female,25 mm
Copyright © Ruslan Panin

Carabus (Pseudocoptolabrus) taliensis taliensis (Fairmaire, 1886)
Male adult
China, Yunnan, Lijiang env., 2700 m, 10.6.1998
Copyright © Alexander Anichtchenko

Carabus (Pseudocoptolabrus) taliensis s.str. (as yulongxuensis)
Male adult
male, 26mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) taliensis s.str. (as yulongxuensis)
Male adult
male, 24mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) taliensis s.str. (as yulongxuensis)
Female adult
female, 29mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) taliensis yanmenensis Deuve, 1996
Male adult
male, 27mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) taliensis yanmenensis Deuve, 1996
Female adult
female, 28mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) taliensis xueshanicola Deuve, 1992
Female adult
female, 30mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) taliensis xueshanicola Deuve, 1992
Male adult
male, 26mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) taliensis xueshanicola Deuve, 1992
Male adult
male, 27mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) taliensis xueshanicola Deuve, 1992
Male adult
male, 28mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) taliensis peterheisei Holotype
Male adult
Holotype
Copyright © Frank Kleinfeld

Carabus (Pseudocoptolabrus) taliensis myanmarensis Imura, 1999
Female adult
female, 36 mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) taliensis mulianus Deuve, 1995
Male adult
male, 27mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) taliensis liqiuensis Imura, 1999
Male adult
male, 32mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) taliensis lijiangensis Paratype
Female adult
female, 33mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) taliensis lijiangensis
Male adult
male, 31mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) taliensis jingpingensis Deuve, 2005 Paratype
Male adult
male, 26mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) taliensis jingpingensis Deuve, 2005 Paratype
Female adult
female, 30mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) nosei taronensis Imura, 2001 Paratype
Female adult
female, 28mm
Copyright © Boleslav Brezina

Carabus (Pseudocoptolabrus) burmanensis Breuning, 1932
Male adult
male, 23mm
Copyright © Boleslav Brezina

Carabus (Morphocarabus) aeruginosus verzhutzkii O.Berlov & Shilenkov, 1996
Male adult
Carabus (Morphocarabus) aeruginosus verzhutzkii O.Berlov & Shilenkov, 1996
C Altays, Katun r., 5 km. N Katanda vill., 21.06.2002, A. Menstschicov leg.
Copyright © Anton Menstschikov

Carabus (Pseudocoptolabrus) armiger armiger Imura, 1997
Female adult
female, 30mm
Copyright © Boleslav Brezina

Carabus (Morphocarabus) aeruginosus verzhutzkii O.Berlov & Shilenkov, 1996
Male adult
C Altai Mts., Seminskii Mt.rng., Katun riv., 3 km S Ust'-Sema riv.,
4.VII.2002 A. & N. Menschikovy leg.
det. Mentschikov A.
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) aeruginosus salechardensis Obydov, 1999 Paratype
Male adult
Rusia, Selehard, tundra, 30.VIII.1953, I. Telyshev leg

Copyright © Dmitri Obydov

Carabus (Morphocarabus) aeruginosus hermani Mannerheim, 1827 loc.typ.
Male adult
C Rusia, S Ural Mts., Zlatoust, 22-28.VI.1952, S. Nikulina leg.
male,

Copyright © Dmitri Obydov

Carabus (Procrustes) paiafa acuticollis Motschulsky, 1858
Male adult

Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) aeruginosus eschscholtzi Mannerheim, 1827
Male adult
Russia, W Altai, Zmeinogorsk distr., near Cherepanovskii vill., 2-16.6.1990, D. Obydov leg.
Copyright © Dmitri Obydov

Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
Male adult
SE Siberia, E Sayan, Nizhneudinsk, h=500m, 21-26.VI.2006,
male, 26mm
Copyright © Ruslan Panin

Carabus (Mimocarabus) maurus osculatii Osculati, 1844
Male adult
Carabus (Mimocarabus) maurus osculatii Osculati, 1844
Copyright © Alexander Anichtchenko

Carabus (Mimocarabus) maurus maurus Adams, 1817
Male+Female adult
Armenia,
Ashotsk distr., Tsoghamarg vill., 24-27.V.1999 P.V.Kazaryan leg.
female, 20mm/male,18mm
Copyright © Ruslan Panin

Carabus (Procrustes) impressus Klug, 1832 (as carmelita)
Male adult

Copyright © Safronov

Carabus (Procrustes) iimpressus Klug, 1832 (as drusum) Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Procrustes) impressus Klug, 1832 (as desertum) Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Procrustes) coriaceus rugifer Kraatz, 1877
Male adult

Copyright © Safronov

Carabus (Procrustes) coriaceus kindermanni Waltl, 1838
Male+Female adult
NW TURKEY,
Stambul Avdzhilar, (Eropean part), h~20 m, 10-29.X.2007 M.V.Cheriavskaya leg./det. I.A.Solodovnikov
female,31mm
Copyright © Ruslan Panin

Carabus (Procrustes) coriaceus hopffgarteni Kraatz, 1878
Male adult

Copyright © Safronov

Carabus (Procrustes) coriaceus cerisyi Dejean, 1826 (as foudrasi)
Male adult

Copyright © Boleslav Brezina

Carabus (Procrustes) coriaceus coriaceus Linne, 1758
Female adult
The neighborhood Nicolo-Berezovskoe republic of
Bashkortostan 5.07
Copyright © Sunvas

Carabus (Procrustes) coriaceus coriaceus Linne, 1758
Male adult
length: 32.42 mm
Germania, Oberfranken, Weismain
leg.det. U.Schmidt, 1983
Foto: U.Schmidt, 2005
Copyright © U.Schmidt

Carabus (Procrustes) coriaceus cerisyi Dejean, 1826
Male adult
male, 30mm
Copyright © Boleslav Brezina

Carabus (Procrustes) coriaceus cerisyi Dejean, 1826
Female adult
female, 31mm
Copyright © Boleslav Brezina

Carabus (Procrustes) clypeatus clypeatus Adams, 1817
Female adult
C-Caucasus, N Ossetia, vall. Mamyhdon riv., h=2100
Koval A.G. leg; 08.08.2001
Copyright © Anton Menstschikov

Carabus (Procrustes) chevrolati korbi Breuning, 1928
Male adult
length: 28 mm
Türkei, Tülü/Sathan, Erzincan
leg. Staven & Willmer; det. K.Staven, 2003
Foto: U.Schmidt, 2009
Copyright © U.Schmidt

Carabus (Procrustes) anatolicus lycicus Breuning, 1934
Male adult

Copyright © Safronov

Carabus (Procrustes) chevrolati chevrolati Cristoforis & Jan, 1837
Male+Female adult
female, 31mm: Ashotsk,Vardaghbyur vill., 30.V.1997
male, 30 mm: Amasia, Chaldash vill.,8-11.VIII.1997
P.V. Kazaryan leg.
Copyright © Ruslan Panin

Carabus (Procerus) scabrosus tauricus Bonelli, 1811
Female adult

Copyright © Safronov

Carabus (Cechenochilus) heydenianus Starck, 1889
Male adult
Caucasian nat. park., S slope Pseashkho mt., 2500 m
21.7.1975 Ovtsharenko V. leg.
Copyright © Alexander Anichtchenko

Carabus (Cechenochilus) heydenianus Starck, 1889
Male+Female adult
NW-Caucasus,
Krasnodar reg., Mostovskoy distr., Skirda Mt Range, N slopes Yatyrgvart Mt. h=2300-2500m., 09-10.VII.2008, D. Fominykh leg.
female,19mm/male,18mm
Copyright © Ruslan Panin

Carabus (Cechenochilus) adangensis Gottwald, 1983
Male adult
male, 19mm
Copyright © Boleslav Brezina

Carabus (Cechenochilus) adangensis Gottwald, 1983
Female adult
female, 18mm
Copyright © Boleslav Brezina

Carabus (Cechenochilus) adangensis Gottwald, 1983
Female adult
Russia, NW-Caucasus:
Karacaj-Cherkessija, Abishira–Akhuba Mt. Range h=2000-2650m, Kyafar-Agur Riv. 14–17.VII.2009, Fominykh D. leg.
female,21mm
Copyright © Ruslan Panin

Carabus (Cechenochilus) adangensis (as gusevi)
Male adult
male, 19mm
Copyright © Boleslav Brezina

Carabus (Cechenochilus) adangensis (as gusevi)
Male adult
male, 19mm
Copyright © Boleslav Brezina

Carabus (Cechenochilus) lazorum Belousov & Zamotajlov, 1999
Male adult
male, 20mm
Copyright © Boleslav Brezina

Carabus (Cechenochilus) boeberi longiceps Chaudoir, 1846
Male adult
C-Caucasus, N Ossetia, near Kozskii, h=2800,
A.G. Koval leg. 10.08.2001; 12 mm
Copyright © Anton Menstschikov

Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Holotype
Male adult
Russia, Karachai-Circassian Republic, Abishira-Akhuba Mount Range between passes Agur and Mylgvar, 2785–2830 m.
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009
Male adult
male, 17mm
Copyright © Boleslav Brezina

Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Paratype
Male adult
Russia, W Caucasus,
Karacaj-Cherkessija, Abishira-Akhuba Mnt Range, Area of pass Turii h~2650-2850m, 17.VII.2009, Fominykh D. leg.
male,17 mm
photo & coll. Panin R.
Copyright © Ruslan Panin

Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Paratype
Female adult
Russia, Karachai-Circassian Republic, Abishira-Akhuba Mount Range between passes Agur and Mylgvar, 2785–2830 m. PARATYPUS
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009
Female adult
Russia, Karachai-Circassian Republic, Abishira-Akhuba Mount Range between passes Agur and Mylgvar, 2785–2830 m. PARATYPUS
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Paratype
Female adult
Russia, Karachai-Circassian Republic, Abishira-Akhuba Mount Range between passes Agur and Mylgvar, 2785–2830 m.
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Paratype
Male adult
Russia, Karachai-Circassian Republic, Abishira-Akhuba Mount Range between passes Agur and Mylgvar, 2785–2830 m.
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Paratype
Female adult
female, 21mm
Copyright © Boleslav Brezina

Carabus (Cechenochilus) boeberi buschi Semenov & Znojko, 1931
Female adult
female, 19mm
Copyright © Boleslav Brezina

Carabus (Cechenochilus) boeberi buschi Semenov & Znojko, 1931
Female adult
female, 23mm
Copyright © Boleslav Brezina

Carabus (Cechenochilus) boeberi boeberi Adams, 1817
Male adult
male, 18mm
Copyright © Boleslav Brezina

Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Male adult
W.Ukraine, Transcarpathian Reg., Rakhov Distr., Chornogora range., h=2020m. Panin R. leg. 28.VI-9.VIII.2008, male, 19 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Female adult
W Ukraine, NE Carpathians, Transcarpathian Reg., Rakhov Distr., Chornogora Mts Range h=2030m, alpin zone, 13-24.VIII.2010, Panin R. leg., female, 22 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Female adult
Ukrainian Carpathians, Transcarpathian reg., Rakhov Distr., Chornogora range, h=2010m, 29.VI-28.VII.2007, Panin R. leg., female, 23 mm., photo/det.Panin R
Copyright © Ruslan Panin

Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Female adult
Ukrainian Carpathians, Transcarpathian reg., Rakhov Distr., Chornogora range, h=2010 m., alpine zone 29.VI-28.VII.2007 Panin R. leg., female,23 mm.
Copyright © Ruslan Panin

Carabus (Platycarabus) irregularis montandoni Buysson, 1882
Male adult
W-Ukraine,
Lviv Reg., Zhydachiv distr., near Vybranivka, under the bark of oak tree, in the forest,12.X.2007 Panin R. leg.
male,22 mm.
Copyright © Ruslan Panin

Carabus (Platycarabus) irregularis montandoni Buysson, 1882
Male adult
W-Ukraine, NW Podolian,
Lviv Reg., Zhydachiv distr., near Vybranivka, under the bark of oak tree, in the forest,12.X.2007 Panin R. leg
L= 22 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) depressus lucens Schaum, 1856
Female adult
Italia, Alpi Cozie, Col. delle Finestre,
2150m, VII.1986 Ratti leg.
female, 24 mm
Copyright © Alexander Anichtchenko

Carabus (Platycarabus) depressus depressus Bonelli, 1811
Female adult
Italia, Bellamonte, 2200m
VI.1992 Rubizzani leg.
female, 20 mm
Copyright © Alexander Anichtchenko

Carabus (Platycarabus) cychroides Baudi, 1860
Male+Female adult
female: NW-Italy, Provincia Torino, Mt.Albergian h~2600 m, 9.VI.2007, V.Monzini leg.
L=20 mm
male:, N slope Mt.Albergian h~2000 m, 9.VI.2003, P.Zorgati leg.
photo Panin R.
Copyright © Ruslan Panin

Carabus (Piocarabus) vladimirskyi Dejean & Boisduval, 1830
Male adult
China, Beijing, Jiufeng mountain, 2007, Liu ye leg.; Photo & det. by liuye
Copyright © Ye Liu

Carabus (Eucarabus) sternbergi Roeschke, 1898
Female adult
Korea, Gyeongi Do,
Bokwang Sa 30.2.1976
female, 28 mm
Copyright © Alexander Anichtchenko

Carabus (Eucarabus) pawlowskianus Deuve, 1989
Female adult
N Korea, Pong-San Mt.
12.7.1997
female, 25 mm
Copyright © Alexander Anichtchenko

Carabus (Eucarabus) nitididorsus Ishikawa & Kim, 1983
Male adult
S Korea, Deagu, 25.8.2003
Copyright © Alexander Anichtchenko

Carabus (Eucarabus) nitididorsus Ishikawa & Kim, 1983
Female adult
Korea, Peak Chonwongbong, 1600-1900m
Mt. Chiri-San, Kyongsangnam-Do, 18.7.87 Y. Imura leg.
Copyright © Alexander Anichtchenko

Carabus (Eucarabus) manifestus guanmenshanus Imura, 1991
Female adult
China, Liaoning, Mt. Lao Gu Go, Dandong, 29.5.97
Copyright © Alexander Anichtchenko

Carabus (Eucarabus) billbergi billbergi Mannerheim, 1827
Male adult
Russia, E Siberia, Buryatia, lake Baikal, near Il'inka vill. Anichtchenko A. leg.
Copyright © Alexander Anichtchenko

Carabus (Eucarabus) billbergi billbergi Mannerheim, 1827
Male adult
Tian tou, Yueshi Counti, Anhui, China
1-10.08.2004
Copyright © Safronov

Carabus (Pagocarabus) crassesculptus eleganticolor Deuve, 1997 Paratype
Male adult
China, Gansu, Min Shan mts., 70 km NW Wudu
Copyright © Alexander Anichtchenko

Carabus (Eucarabus) ulrichi s.str. (as volhynicus Lutshnik, 1927)
Male+Female adult
W Ukraine, NW Podolian,
Ternopil reg., Pidvolochisk distr., forest Bogdanivskiy, near Romanove Selo, 7-24.V.2008, Panin R. & Solodovnikov I. leg.
Copyright © Ruslan Panin

Carabus (Pantophyrtus) turcomanorum karaalmicus Gottwald, 1994
Male adult
male, 27mm
Copyright © Boleslav Brezina

Carabus (Pantophyrtus) turcomanorum turcomanorum Thieme, 1881
Female adult
female, 32mm
Copyright © Boleslav Brezina

Carabus (Pantophyrtus) turcomanorum turcomanorum Thieme, 1881
Male adult
male, 28mm
Copyright © Boleslav Brezina

Carabus (Pantophyrtus) turcomanorum s.str. (as orbitosus)
Male adult
male, 27mm
Copyright © Boleslav Brezina

Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993
Male adult
Kirgizstan, S Chatkal mt. rng., Sary-Chelek lake 2000 m, 25.6.2000
Copyright © Alexander Anichtchenko

Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993
Male adult
male, 29mm
Copyright © Boleslav Brezina

Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993 Paratype
Female adult
female, 28mm
Copyright © Boleslav Brezina

Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993 Paratype
Female adult
female, 34mm
Copyright © Boleslav Brezina

Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993 Paratype
Male adult
male, 27mm
Copyright © Boleslav Brezina

Carabus (Pantophyrtus) brachypedilus brachypedilus s.str.
Male adult
N Kirgizskii mt rng., Ak-Suu riv., 1400 m,
Belovodskoe, 16.4.2000 Klimenko A. leg.
Copyright © Alexander Anichtchenko

Carabus (Pantophyrtus) brachypedilus brachypedilus s.str.
Female adult
female, 27mm
Copyright © Boleslav Brezina

Carabus (Pantophyrtus) brachypedilus brachypedilus s.str.
Male adult
male, 26mm
Copyright © Boleslav Brezina

Carabus (Pantophyrtus) brachypedilus brachypedilus s.str.
Female adult
female, 28mm
Copyright © Boleslav Brezina

Carabus (Pantophyrtus) brachypedilus absentivus Belousov & Kabak 1993 Paratype
Male adult
male, 24mm
Copyright © Boleslav Brezina

Carabus (Pantophyrtus) brachypedilus absentivus Belousov & Kabak 1993 Paratype
Male adult
male, 24mm
Copyright © Boleslav Brezina

Carabus (Pantophyrtus) brachypedilus absentivus Belousov & Kabak 1993 Paratype
Female adult
female, 26mm
Copyright © Boleslav Brezina

Carabus (Pantophyrtus) brachypedilus absentivus Belousov & Kabak 1993 Paratype
Female adult
female, 30mm
Copyright © Boleslav Brezina

Carabus (Pagocarabus) crassesculptus diruptus Morawitz, 1886
Male adult
China, Gansu, Umg. Xiahe 3000-3200m
24-26.6.1993 Heinz leg.
20 mm
Copyright © Alexander Anichtchenko

Carabus (Pagocarabus) crassethoracis Kleinfeld, 1999 Holotype
Male adult

Copyright © Frank Kleinfeld

Carabus (Pagocarabus) crassesculptus crassesculptus Kraatz, 1881
Male adult
China, Beijing, Wulingshan, 2007, Liu ye leg.;
Photo & det. by liuye
Copyright © Ye Liu

Carabus (Pagocarabus) crassesculptus crassesculptus Kraatz, 1881
Male adult
China, Hebei, Xianhua (=Kalgan)
10-15.5.1995
21 mm
Copyright © Alexander Anichtchenko

Carabus (Pagocarabus) crassesculptus tonghua Deuve & Mourzine, 2001
Male adult
China, NW Sishuan, 3500m, Hongyuan
Somang Qu riv. grassland, 9-29.6.2004
male, 20 mm
Copyright © Alexander Anichtchenko

Carabus (Eucarabus) ulrichi ulrichi Germar, 1824 (as podolicus Semenov, 1898 loc.typ)
Female adult
C-UKRAINE, Vinnitsa Reg., near Khmilnyk, forest, 10-20.VIII.2002, Demidov G.N. leg., det. Panin R female, 30mm
Copyright © Ruslan Panin

Carabus (Eucarabus) ulrichi ulrichi Germar, 1824
Male adult
NW Ukrainian Carpathians
Lviv Reg., Staryi Sambir Distr., near Dobromyl , h=500m 9.VI-25.VII.2007, Panin R. Leg.
Copyright © Safronov

Carabus (Tribax) nothus planipennis Chaudoir, 1846
Female adult
Dagestan, Huizah., near Motchoh, h=1800, 27.07.1993, Ilina leg.
photo A. Menstschicov
Copyright © Anton Menstschikov

Carabus (Tribax) nothus planipennis Chaudoir, 1846
Male adult
male, 22mm
Copyright © Boleslav Brezina

Carabus (Tribax) nothus planipennis Chaudoir, 1846
Male adult
male, 21mm
Copyright © Boleslav Brezina

Carabus (Tribax) nothus planipennis Chaudoir, 1846
Male adult
Dagestan, Bogosskii mt. rng, Inhokvari-Honoh
3500 m, 15-25.6.85 Abdurakhmanov leg.
Copyright © Alexander Anichtchenko

Carabus (Tribax) nothus nothus Adams, 1817
Male adult
male, 24mm
Copyright © Boleslav Brezina

Carabus (Tribax) nothus nothus Adams, 1817
Female adult
female, 24mm
Copyright © Boleslav Brezina

Carabus (Tribax) nothus nothus Adams, 1817
Female adult
female, 25mm
Copyright © Boleslav Brezina

Carabus (Tribax) nothus fausti Dohrn, 1873
Male adult
male, 24mm
Copyright © Boleslav Brezina

Carabus (Eucarabus) billbergi billbergi Mannerheim, 1827
Male adult
Rusia, Primorskii reg., Kiparisovo vill., 5.8.1993
Chichvarhin A. leg.
male, 20 mm
Copyright © Alexander Anichtchenko

Carabus (Eucarabus) billbergi billbergi Mannerheim, 1827
Male adult
Rusia, Primorskii reg., Anuchinskii distr.,
near LZU-3, 7.8.1979 Bakurov leg.
male, 21 mm
Copyright © Alexander Anichtchenko

Carabus (Tribax) nothus (as abdurachmanovi)
Male adult
Carabus (Tribax) nothus (as abdurachmanovi)
male, 22mm
Copyright © Boleslav Brezina

Carabus (Tribax) nothus (as abdurachmanovi)
Female adult
Carabus (Tribax) nothus (as abdurachmanovi)
female, 23mm
Copyright © Boleslav Brezina

Carabus (Tribax) kasakorum kasakorum Semenov, 1896
Male adult
male, 26mm
Copyright © Boleslav Brezina

Carabus (Tribax) kasakorum kasakorum Semenov, 1896
Male adult
male, 27mm
Copyright © Boleslav Brezina

Carabus (Tribax) kasakorum kasakorum Semenov, 1896
Female adult
female, 29mm
Copyright © Boleslav Brezina

Carabus (Tribax) kasakorum kasakorum Semenov, 1896
Female adult
female, 29mm
Copyright © Boleslav Brezina

Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
Male adult
male, 23mm
Copyright © Boleslav Brezina

Carabus (Microplectes) riedeli Menetries, 1832
Female adult
female, 19mm
Copyright © Boleslav Brezina

Carabus (Microplectes) riedeli Menetries, 1832
Male adult
male, 17mm
Copyright © Boleslav Brezina

Carabus (Microplectes) riedeli Menetries, 1832
Male adult
male, 17mm
Copyright © Boleslav Brezina

Carabus (Microplectes) riedeli Menetries, 1832
Male adult
N-Ossetia,
Khazniaush Pass, h=2500-2700m, 9.VI-31.VII.1994, Zamotajlov A.leg.
male, 17mm
Copyright © Ruslan Panin

Carabus (Microplectes) convallium convallium Starck, 1889
Male adult
male, 19mm
Copyright © Boleslav Brezina

Carabus (Microplectes) convallium convallium Starck, 1889
Male adult
male, 19mm
Copyright © Boleslav Brezina

Carabus (Microplectes) convallium convallium Starck, 1889
Male+Female adult
female:NW-Caucasus, Russia,
Krasnodar reg., Apsheron Distr., Lagonaki Mt., near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominyh leg.
male:Karachaevo-Cherkessia,the right source of the river Beskess, h=2100m, 10.VIII.1997, I.A. Solodovnikov leg.
female, 21mm/male, 19mm
Copyright © Ruslan Panin

Carabus (Microplectes) convallium argonautarum Semenov, 1898
Male adult
male, 18mm
Copyright © Boleslav Brezina

Carabus (Microplectes) convallium argonautarum Semenov, 1898
Male adult
male, 19mm
Copyright © Boleslav Brezina

Carabus (Microplectes) convallium argonautarum Semenov, 1898
Female adult
female, 20mm
Copyright © Boleslav Brezina

Carabus (Microplectes) convallium argonautarum Semenov, 1898
Male+Female adult
Abkhasia,
W.slope of Gagrinsky Mt., Krepostnaya River source, h=930-980m., A.S.Zamotajlov leg. 27.VII.2000
female, 19mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Pachystus) tamsi tamsi Menetries, 1832
Male+Female adult
Transcaucasia, Armenia,
Ashotsk, 18.VI.2001, P.V.Kazaryan leg.
female, 26mm//male, 24mm
Copyright © Ruslan Panin

Carabus (Pachystus) tamsi tamsi Menetries, 1832
Female adult
Azerbayjan, Talysh mt, Zuvant
Pirasora, 1900-2000 m 9.7.1983 I. Belousov leg.
female, 25 mm
Copyright © Alexander Anichtchenko

Carabus (Mesocarabus) macrocephalus Dejean, 1826
Male adult
Spain, Picos de Europa, Macizo Central,
Llagu Bajero 1890m 6.8.2003
Copyright © Alexander Anichtchenko

Carabus (Pachystus) tamsi tamsi Menetries, 1832
Male adult
N Iran, Azarbaygan-e-Sharqi, 21 km SE
Halhal, 3734N 4841E, 1633m
3.6.2003 Kabatek P. leg
male, 23 mm
Copyright © Alexander Anichtchenko

Carabus (Mesocarabus) lusitanicus lusitanicus Fabricius, 1801
Male adult
Portugal, Vale de Estrela, Guarda, VI.1976
Copyright © Alexander Anichtchenko

Carabus (Mesocarabus) lusitanicus lusitanicus Fabricius, 1801(as schaumi Gaubil, 1849)
Male+Female adult
Portugal,
Viseu distr., Gosende, h~1010m, 15.X.2009, D.Prunier leg.
female,26 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus brevis Dejean, 1826
Female adult
Spain, Sierra Guadarrama, Montes Carpetinos,
Pto. de Somosierra 1600-1700m, 3.VI.2006
Copyright © Alexander Anichtchenko

Carabus (Mesocarabus) breuningi Csiki, 1927
Male adult
Spain, San Martin de Castaneda, Zamora 11 oct 1984,h~1500 m
Copyright © Alexander Anichtchenko

Carabus (Mesocarabus) lusitanicus gougeleti Reiche, 1863 (as alcaracinus Kraatz, 1886)
Male adult
Spain, Riopa (Albacete)
Copyright © Alexander Anichtchenko

Carabus (Pachystus) hungaricus scythus Motschulsky, 1847
Male adult
S Ukraine,
Zaporizhia reg., Khortytsa island, 19.VIII.2008, M. Mulenko leg., det.Panin R.
male, 27mm
Copyright © Ruslan Panin

Carabus (Pachystus) hungaricus cribellatus Adams, 1812
Female adult
Kazakhstan,
Astana reg. Enbekshil-der distr., Burevestnik vill., 9.VIII.2002
female,26 mm
Copyright © Ruslan Panin

Carabus (Pachystus) graecus morio Mannerheim, 1830
Male adult
Turchia, Sehtavul Geg 1700m
5.1996 Rabizzani leg.
male, 26 mm
Copyright © Alexander Anichtchenko

Carabus (Pachystus) graecus kykladensis Breuning, 1961
Male adult
male, 24mm
Copyright © Boleslav Brezina

Carabus (Pachystus) graecus kykladensis Breuning, 1961
Male adult
female, 24mm
Copyright © Boleslav Brezina

Carabus (Mesocarabus) dufouri dufouri Dejean, 1829 (as papillatus Lapouge, 1902)
Male adult
Spain, Algeciras, Molino de las Cuevas
5-6.4.2003 Anichtchenko A. leg.
Copyright © Alexander Anichtchenko

Carabus (Pachystus) glabratus glabratus Paykull, 1790
Female adult
W UKRAINE Lviv region, Stryi distr., near Bilche vill., 9-20.VI.2005, Panin R. leg. female 30 mm
Copyright © Ruslan Panin

Carabus (Pachystus) glabratus glabratus Paykull, 1790
Male adult
W Ukraine, NW Podolian, Lviv reg., Zolochiv distr., near Stinka vill., Lysa Gora, 26.IV.2009 R.Panin leg., 29mm
Copyright © Ruslan Panin

Carabus (Pachystus) glabratus glabratus Paykull, 1790
Female adult
Germania, Oberfranken, Kulmbach leg.det. U.Schmidt, 1974 Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Carabus (Pachystus) cavernosus cavernosus Frivaldsky, 1837
Male adult
Bulgaria, Middle Stara Planina Mts.,
feeder of Byala Reka riv., below Botev Peak
1900-2000m 25.7.1996 B. Gueorguiev leg.
male, 25 mm
Copyright © Alexander Anichtchenko

Carabus (Pachycarabus) staehlini Adams, 1817
Male adult
N Osetia, Skalistyi mt. rng.
Hamarat-Hoh mt. 14.7.1996 Arzanov leg.
male, 21 mm
Copyright © Alexander Anichtchenko

Carabus (Pachycarabus) roseri ciftekopruleriensis Machard, 1975
Female adult
NE Turkey,
Artvin Prov., Borcka, Macahel Gecidi h=1450m, 18.VI.2007, D.Prunier leg.
female, 22mm
Copyright © Ruslan Panin

Carabus (Pachycarabus) imitator imitator Reitter, 1883
Male adult
Abkhasia,
Anchkho mnt.Range, near Lake Chkho, h~2450-2550m, 13-14.VII.2009, Bondarenko A.S. leg.
male, 20mm
Copyright © Ruslan Panin

Carabus (Oxycarabus) saphyrinus pseudosaphyrinus Schweiger, 1964 (as boluensis)
Male adult
Turkei, Bolu, Seben, Aladag,
1500m, 17.7.1988 Staven K. leg.
male, 23 mm
Copyright © Alexander Anichtchenko

Carabus (Oxycarabus) saphyrinus pseudosaphyrinus Schweiger, 1964 (as rabaronianus)
Male adult
Anatolia, Kastamonu, Yaraligoz verso Catalzeitin,
VI.2001 Sola Claudio leg.
male, 28 mm
Copyright © Alexander Anichtchenko

Carabus (Oxycarabus) saphyrinus pseudosaphyrinus Schweiger, 1964 (as rabaronianus)
Female adult
Anatolia, Kastamonu, Yaraligoz
verso Catalzeytin, VI.2001 Sola C. leg.
female, 30 mm
Copyright © Alexander Anichtchenko

Carabus (Orinocarabus) sylvestris transsylvanicus Dejean, 1826
Male adult
Ukrainian Carpathians, Transcarpathian reg., Rakhov Distr., Chornogora Mts. Petros Mt. h=1850-2000m. alpine zone, 29.VI-28.VII.2007 Panin R. Leg.
Copyright © Safronov

Carabus (Orinocarabus) sylvestris transsylvanicus Dejean, 1826
Male+Female adult
Ukrainian Carpathians
Transcarpathian reg., Rakhov Distr., Chornogora Mts. Pietros Mt. h=1850-2000 m. alpine zone, 29.VI-28.VII.2007 Panin R. Leg.
long 22 mm.
Copyright © Ruslan Panin

Carabus (Orinocarabus) pedemontanus omensis Born, 1901
Female adult
Italy, Piemonte (CN), Col del Mulo
17.7.2001 leg- Grottolo
female, 22 mm
Copyright © Alexander Anichtchenko

Carabus (Orinocarabus) pedemontanus omensis Born, 1901
Male adult
Italy, Piemonte (CN), Col del Mulo
17.7.2001 leg. Grottolo
male, 22 mm
Copyright © Alexander Anichtchenko

Carabus (Orinocarabus) fairmairei Thomson, 1875
Female adult
SE France,
Grand Belvédère du refuge du Mont Viso, 5-20.VII.1991
female, 20mm.
photo Panin R.
Copyright © Ruslan Panin

Carabus (Orinocarabus) concolor concolor Fabricius, 1792
Male adult
Piemonte, Oropa (BI) , M. Camino,
2300 m, 20.7.2003 leg. F. Angelini
male, 18 mm
Copyright © Alexander Anichtchenko

Carabus (Orinocarabus) concolor concolor Fabricius, 1792
Female adult
Piemonte, Oropa (BI) , M. Camino,
2300 m, 20.7.2003 leg. F. Angelini
female, 19 mm
Copyright © Alexander Anichtchenko

Carabus (Orinocarabus) castanopterus A. & J.B. Villa, 1833
Male adult
Italy, Lombardia (BG), passo S. Simone 2000 m,
Mezzoldo 26.8.2006 leg. Grottolo
male, 17 mm
Copyright © Alexander Anichtchenko

Carabus (Orinocarabus) bertolinii Kraatz, 1878
Male adult
Italia, Bellamonte, 2200 m, VI.1982 Rubizzani leg.
male, 18 mm
Copyright © Alexander Anichtchenko

Carabus (Pachystus) hortensis hortensis Linne, 1758
Male adult
Germania, Oberfranken, Langenau
leg.det. U.Schmidt, 2002
Foto: U.Schmidt, 2006
Copyright © U.Schmidt

Carabus (Pachystus) hortensis preslii Dejean, 1830
Female adult
S Greece, Peloponnesos, 14.7.1987
female, 28 mm
Copyright © Alexander Anichtchenko

Carabus (Pachystus) hortensis preslii Dejean, 1830 (as jonicus)
Male adult
Greece, Korfu 16.8.1987
male, 25 mm
Copyright © Alexander Anichtchenko

Carabus (Pachystus) hortensis neumeyeri Schaum, 1856
Female adult
Italy, Basilicata, Serra Calvello (PZ)
1400 m, 7.2003 F. Angelini leg.
female, 25 mm
Copyright © Alexander Anichtchenko

Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847 (as guadalupensis) Paratype
Male adult
ESP - Caceres, S. Guadalupe,
800 m XI.2002
Copyright © Alexander Anichtchenko

Carabus (Oreocarabus) ghiliani La Ferte-Senectere, 1847
Male adult
Spain, Segovia, Sierra de Somosierra,
La Pinilla, 1800m 4.4.2005
male, 19 mm
Copyright © Alexander Anichtchenko

Carabus (Oreocarabus) luetgensi luetgensi s.str.
Male adult
Spain, Zamora, San Martin de Castaneda
10.5.1997
male, 20 mm
Copyright © Alexander Anichtchenko

Carabus (Ophiocarabus) regeli Morawitz, 1886
Male adult
Kirgizia, E Terskey Ala-Too, Chon-Akshuu pass,
2800 m, 13.6.1999
Copyright © Alexander Anichtchenko

Carabus (Ophiocarabus) aeneolus aeneolus Morawitz, 1886
Male adult
photo M. Smirnov from www.zin.ru
Copyright © Maksim Smirnov

Carabus (Ophiocarabus) aeneolus aeneolus Morawitz, 1886
Female adult
SE KAZAKHSTAN,
Alma-Ata reg., Zailiysky Mts., h=3000m, B. Almaatinka riv. valley,01.VI.2001
Copyright © Nil Belonogov

Carabus (Ophiocarabus) aeneolus aeneolus Morawitz, 1886
Male+Female adult
KAZAKHSTAN,
Bolshoe Almaatinskoe Lake,30.VI.1992
female, 17mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) baguenai Breuning, 1926
Male adult
Spain, Almeria, Sierra de Baza 10.III.1995 male photo Anichtchenko A. 
Copyright © Alexander Anichtchenko

Carabus (Ohomopterus) yamato Nakane, 1953
Female adult
Japan, Ohkubo, 740-770m, Shirakawa-mura
Ohno-gun (Gifu) 20-25.6.80 Ishikawa leg.
Copyright © Alexander Anichtchenko

Carabus (Ohomopterus) yaconinus yamaokai Ishikawa & Kubota, 1994
Female adult
Japan, Ehime pref., Taisanji, 14.5.2002
Matsuyama city, T. Kurihara leg.
Copyright © Alexander Anichtchenko

Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
Male adult
SW-France,
Foret d'Iraty Pyrenees-Atlantiques, d'Occabe, 15.VIII.1988
male,30 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Ohomopterus) yaconinus yaconinus Bates, 1873
Male adult
Japan, Kagawa pref, Shin-ike
Awai town, Kannonji city, 4.3.1993
male, 28 mm
Copyright © Alexander Anichtchenko

Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876 Loc.typ.
Female adult
Spain, Navarra, Sierra de Aralar,
Altxueta 1200-1300m 6.1.2001
Copyright © Alexander Anichtchenko

Carabus (Ohomopterus) maiyasanus maiyasanus Bates, 1873
Female adult
Japan, Shiga pref.,
Lake Yogo 18.8.1995 Shimizu leg.
24 mm
Copyright © Alexander Anichtchenko

Carabus (Ohomopterus) japonicus tsushimae Breuning, 1932
Male adult
Japan, Tsushima isl., Nagasaki pref., Hiratake, Kamiagata town, 19.6.2002 T. Kurihara leg.
male, 23 mm
Copyright © Alexander Anichtchenko

Carabus (Ohomopterus) insulicola insulicola Chaudoir, 1869
Male adult
Japan, Yamanashi pref., Kodachi, Kawaguchiko-cho, Minamitusru distr.
2.IX.2000 T. Kurihara leg.
male, 29 mm
Copyright © Alexander Anichtchenko

Carabus (Ohomopterus) insulicola insulicola Chaudoir, 1869
Female adult
15.VII.1998 JAPAN, Ichihara, env. Chiba
Copyright © Nil Belonogov

Carabus (Megodontus) violaceus rilvensis Kolbe, 1887
Female adult
Bulgaria, Stara Zagora distr. Stara Planina Mts., Shipka Mt., 16.08.2005
A. Safronov leg.

Copyright © Safronov

Carabus (Ohomopterus) dehaanii Chaudoir, 1848
Male adult
Japan, Hyogo pref., Ueda-rindo, Mihara, 4.3.93 N. Sano leg.
Copyright © Alexander Anichtchenko

Carabus (Ohomopterus) arrowianus murakii Ishikawa & Kubota, 1984
Male adult
Sho, alt 3m, Futami-T, Watarai-Co. Mie pref. Japan, 17.5.1987 Y. Imura leg.
male, 27 mm
Copyright © Alexander Anichtchenko

Carabus (Ohomopterus) arrowianus arrowianus Breuning, 1934
Male adult
Japan, Shizuoka pref., Mt. Mitake, Inasa town 18.10.1998 T. Kurihara leg.
Copyright © Alexander Anichtchenko

Carabus (Ohomopterus) albrechti esakianus Nakane, 1961
Male adult
Japan, Tokyo pref, Mt. Takao, Nachioji City, 11.8.1990 K. Kurihara leg.
Copyright © Alexander Anichtchenko

Carabus (Megodontus) violaceus picenus A.& J.B.Villa, 1838
Male adult
Italia, Marche Pesaro
leg.det. K.Staven, 1998
Foto: U.Schmidt, 2008
length: 26-30 mm
Copyright © U.Schmidt

Carabus (Ohomopterus) lewisianus lewisianus Breuning, 1932
Male adult
Japan, Yamanashi pref., Koganesawa-rindo, Otsuki city,
13.6.1990 K. Kurihara leg.
male, 20 mm
Copyright © Alexander Anichtchenko

Carabus (Nesaeocarabus) gomerae A.Muller, 2004 Paratype
Female adult

Copyright © Boleslav Brezina

Carabus (Megodontus) violaceus mixtus Gehin, 1876
Female adult
France, Eloise 74, pont de Gresin 14.07.1991
female, 29mm
Copyright © Alexander Anichtchenko

Carabus (Megodontus) violaceus fiorii Born, 1901
Male+Female adult
NW-Italy,
Reggio Emilia, Regnano, 5.IX.1996 leg. Cartini A.
female, 30 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus baeterrensis Lapouge, 1901
Male+Female adult
S France,
Midi-Pyrénées Region, Ariège, Col du Chioula h=1500m, V-VI.2005, Pascal Stefani leg.
photo Panin R.
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus aurolimbatus Dejean, 1829
Female adult
Kazakhstan,
North Kazakhstan Province (Kokchetavski region)1.VII.1986
female, 24mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus aurolimbatus Dejean, 1829
Male adult
C.UKRAINE,
Poltava reg., near Novi Sanzhary, bank of Vorskla River, 1-2.VII.07, D.Kovalchuk leg.
male 25mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus aurichalceus Kraatz, 1879
Male adult
Spain, Pais Vasco, Aranzazu, 6.I.2005
Anichtchenko A. leg.
male, 27 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823
Male adult
W Ukraine, NW Podolian, Ternopol reg., Kremenets distr., near Chornyi Lis vill., steppe sites, 4-22.V.2009, Panin R. leg. female, 29 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823
Male adult
W-UKRAINE
NW Podolian, Lviv Reg., Zolochiv Distr., near Osovitsa, steppe sites, Panin R. leg. 1.V-6.VI.07
male, 26mm
Copyright © Safronov

Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823
Male+Female adult
W. Ukraine, W. Podolian, Lviv reg., Nikolaev distr., near Krasiv vill., under the bark of oak tree, 8.II.2009, Panin R. leg. female, 29mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823 (as carbonatus Schaufuss, 1882)
Male adult
W.UKRAINE, Ukrainian Carpathians, Lviv Reg., Skolivski Beskidy Mts., near Skole, h=700m, 20.IX.2009, Panin R. leg. male, 33mm
Copyright © Ruslan Panin

Carabus (Megodontus) vietinghoffi vietinghoffi Adams, 1812
Male adult
Yakutia, near Kular vill., 27.7.2000, 20 mm det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Megodontus) vietinghoffi vietinghoffi Adams, 1812
Female adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada

Copyright © Henri Goulet

Carabus (Megodontus) vietinghoffi fulgidus Fischer, 1827
Male adult
Russia, Evreiskaya AO., near Obluchie 14.6.1993
Chichvarhin A. leg.
male, 20 mm
photo, det. & coll. Anichtchenko A
Copyright © Alexander Anichtchenko

Carabus (Megodontus) tuvensis Shilenkov, 1996
Male adult
Photo A. Menstschicov
Copyright © Anton Menstschikov

Carabus (Megodontus) tuvensis Shilenkov, 1996
Female adult
Photo A. Mentschikov
Copyright © Anton Menstschikov

Carabus (Megodontus) stroganowi tiedemanni Breuning, 1934
Male adult
IRAN, Gilan, Masuleh, 1550 m., 17.06.2002., O.Montrieull leg.
photo A. Menstschicov
Copyright © Anton Menstschikov

Carabus (Megodontus) stroganowi stroganowi Zoubkoff, 1837
Male adult
Iran,
Mazanderan, Naharkoran, b. Gorgan
500m, 4-11.VII.1975 Heinz leg.
male, 30 mm
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Megodontus) septemcarinatus Motschulsky, 1840
Male adult

Copyright © Safronov

Carabus (Megodontus) schoenherri schoenherri Fischer, 1822
Male adult

Copyright © Anton Menstschikov

Carabus (Megodontus) schoenherri schoenherri Fischer, 1822
Male adult
enza Reg., Kuznetsk Distr., env. Tikhmenevo Vill., upper course of Sura River, pine forest VI.2005 I.Lebezhinskaja leg.
L=28.0 mm
Copyright © Kirill Makarov

Carabus (Megodontus) schoenherri schoenherri Fischer, 1822 (as sangilenus Obydov, 1997)
Male adult
Tyva, Akademika Obrucheva mt.rng., 15km N Cherbi vill.,
Chailag ur., 7.7.1999 A. & N. Menshikovy leg.
male, 26mm
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Megodontus) schoenherri schoenherri Fischer, 1822
Female adult

Copyright © Safronov

Carabus (Megodontus) persianus persianus Roeschke, 1896
Male adult
Iran, Gilan, pas. N.Dailiman, 1100m, 12-19.VII.1999 Heinz leg.
male, 32 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Megodontus) obovatus Fischer, 1827
Female adult
Kemerovo reg., Kuznetskii Mt. rng., Berikul, 1.6.1982
female, 16mm
Copyright © Alexander Anichtchenko

Carabus (Megodontus) obovatus Fischer, 1827
Male adult
Kemerovo reg., Kuznetskii Mt. rng., Berikul, 1.6.1982
male, 17mm
Copyright © Alexander Anichtchenko

Carabus (Megodontus) obovatus Fischer, 1827
Female adult
Kuznetskii Mt., Gluhari vill, A. Menstschicov leg., 07.06.2006
female
Copyright © Anton Menstschikov

Carabus (Megodontus) leachi surakovi Obydov, 1995
Male adult

Copyright © Kirill Makarov

Carabus (Megodontus) leachi klimenkoi Obydov, 1999 Holotype
Male adult
Museum of Institute of Systematic Biology (Daugavpils, Latvia)
Copyright © Alexander Anichtchenko

Carabus (Megodontus) leachi panzeri Dejean, 1829 (as klimenkoi Obydov, 1999)
Male adult
Kazakhstan, S Altay mts., Ognevka vill., Kalbinskii mt. rng., 16.6.1999 Klimenko A. leg.
Copyright © Alexander Anichtchenko

Carabus (Megodontus) gyllenhali Fischer, 1827
Male adult
Krym Peninsula
male, 29 mm
Copyright © Ruslan Panin

Carabus (Megodontus) gyllenhali Fischer, 1827
Male adult
Krym Peninsula, near Sevastopol, 05.05.2000, A.V. Konovalov leg.
Copyright © Anton Menstschikov

Carabus (Megodontus) exaratus Quensel, 1806
Male adult
Russia, C.Caucasus, N.Ossetia,
valley of River Khazniaush,
h=1400 m, 10.VI-31.VII.1993
A. Zamotajlov leg.
Copyright © Safronov

Carabus (Megodontus) exaratus Quensel, 1806
Male+Female adult
NW Caucasus, Russia,
Krasnodar reg., Krasnodar City, 27.IX-07.X.07, D.D.Fominykh leg.
female,29mm/male, 27mm
Copyright © Ruslan Panin

Carabus (Megodontus) kantaikensis Gehin, 1885
Female adult
Russia, Lake Baikal, vall. of Goloustnaya riv., Nizhnii Kochergat vill.,
27.VII.1994 Anichtchenko A. leg.
female, 18 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Megodontus) kantaikensis Gehin, 1885
Male adult
E Sayan Mts., Sentsa riv. (Oka riv. confluence), near Shasnur vill.
7-16.7.1998 Anichtchenko A. leg.
male, 18 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Megodontus) kantaikensis Gehin, 1885
Male adult
Krasnoyarsk Area, Sayanogorsk Distr, env. Cheremushki Vill., right bank of Enisej Riv., h= 500 - 800 m, 23.07.1999 A. Semin leg. male L=17,0 mm
Copyright © Maksim Smirnov

Carabus (Megodontus) viridicans Obydov, 1993
Male adult
W Sayan, Cheremushki vill., Borus Mt. rng., h= 500m, 29-30.7.2000, Brinev A. leg.; 16 mm
Copyright © Alexander Anichtchenko

Carabus (Megodontus) viridicans Obydov, 1993
Female adult
Photo A. Menstschicov
Copyright © Anton Menstschikov

Carabus (Megodontus) viridicans Obydov, 1993
Male adult

Copyright © Anton Menstschikov

Carabus (Megodontus) danilevskii Obydov, 1993
Male adult
Krasnoyarsk Area, Sayanogorsk Distr, env. Cheremushki Vill., right bank of Enisej Riv., h= 500 - 800 m, 23.07.1999 A. Semin leg.
L=17,0 mm
Copyright © Maksim Smirnov

Carabus (Nesaeocarabus) faustus faustus Brulle, 1838
Male adult

Copyright © Boleslav Brezina

Carabus (Nesaeocarabus) faustus faustus Brulle, 1838
Male adult

Copyright © Boleslav Brezina

Carabus (Nesaeocarabus) faustus cabrerai Enderlein, 1929
Male adult

Copyright © Boleslav Brezina

Carabus (Nesaeocarabus) coarctatus Brulle, 1838
Female adult
Canary isl., Las Palmas, Las Mesas 20.2.90
Copyright © Alexander Anichtchenko

Carabus (Nesaeocarabus) abbreviatus Brulle, 1835
Male adult

Copyright © Boleslav Brezina

Carabus (Nesaeocarabus) abbreviatus Brulle, 1835
Female adult
Spain, Canary Islands,
Tenerife, Ft de Esperanza 1360m, TF-24, 29.III.2007, D.Prunier leg.
female, 22mm
Copyright © Ruslan Panin

Carabus (Pagocarabus) wrzecionkoianus Deuve, 2007 Holotype Paratype
Male adult

Copyright © Boleslav Brezina

Carabus (Pagocarabus) wrzecionkoianus Deuve, 2007
Female adult

Copyright © Boleslav Brezina

Carabus (Pagocarabus) wagae folwarcznyi Deuve, 1997 Paratype
Male adult
male, 21 mm
Copyright © Boleslav Brezina

Carabus (Pagocarabus) ? wagae Fairmaire, 1882
Male adult
China, E Tibet, pas Dongdola 5000 m, 4.7.1995
Copyright © Alexander Anichtchenko

Carabus (Pagocarabus) wagae degeicola Deuve, 1996
Male adult

Copyright © Boleslav Brezina

Carabus (Pagocarabus) tsogoensis Deuve, 1997
Male adult
male, 20 mm
Copyright © Boleslav Brezina

Carabus (Pagocarabus) tryznai Brezina & Hackel, 2006 Holotype
Male adult

Copyright © Boleslav Brezina

Carabus (Pagocarabus) tryznai Brezina & Hackel, 2006 Paratype
Female adult

Copyright © Boleslav Brezina

Carabus (Pagocarabus) sinotibeticola xiaodongi (Imura, Zhou & Su, 2004)
Male adult

Copyright © Boleslav Brezina

Carabus (Pagocarabus) sinotibeticola sinotibeticola Mandl, 1975
Male adult

Copyright © Boleslav Brezina

Carabus (Pagocarabus) sinotibeticola sinotibeticola Mandl, 1975
Female adult

Copyright © Boleslav Brezina

Carabus (Pagocarabus) sinotibeticola pantholops Brezina, Hackel & Sehnal, 2007 Holotype Paratype
Male adult
male, 23mm
Copyright © Boleslav Brezina

Carabus (Pagocarabus) sinotibeticola pantholops Brezina, Hackel & Sehnal, 2007
Female adult
female, 24mm
Copyright © Boleslav Brezina

Carabus (Pagocarabus) sichuanicola sigena Kleinfeld, 2000 Holotype
Male adult

Copyright © Frank Kleinfeld

Carabus (Pagocarabus) sichuanicola sigena Kleinfeld, 2000
Male adult
male, 26mm
Copyright © Boleslav Brezina

Carabus (Pagocarabus) sichuanicola morvanellus Deuve, 1998 Holotype
Female adult

Copyright © Boleslav Brezina

Carabus (Pagocarabus) sichuanicola sichuanicola Deuve, 1989 (as dromlae) Paratype
Male adult

Copyright © Boleslav Brezina

Carabus (Pagocarabus) puetzi Kleinfeld, 2000 Holotype
Male adult

Copyright © Frank Kleinfeld

Carabus (Pagocarabus) piceophilus piceophilus Deuve, 1994
Male adult

Copyright © Boleslav Brezina

Carabus (Pagocarabus) piceophilus piceophilus Deuve, 1994
Male adult
male, 22mm
Copyright © Boleslav Brezina

Carabus (Pagocarabus) paulusi Kalab, 1995
Male adult
China, E-Tibet, Chakla, 5000m, 100 km
N of Lhasa, 4.6.1997 Wrzecionko A. leg.
Copyright © Alexander Anichtchenko

Carabus (Pagocarabus) nanschanicus globulipennis Deuve, 2006 Holotype
Male adult

Copyright © Boleslav Brezina

Carabus (Rhigoidocarabus) zhubajie laojunensis Kleinfeld, 1999 Holotype
Male adult

Copyright © Frank Kleinfeld

Carabus (Oreocarabus) taedatus agassi Le Conte, 1850
Female adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Carabus (Oreocarabus) taedatus agassi Le Conte, 1850
Male adult
USA, Arizona, Coconino Co., Humphreys Peak,
bei Flagstaff 14.6.1990 Heinz leg.
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
Female adult
female, 49mm
Copyright © Boleslav Brezina

Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
Male adult
male, 43mm
Copyright © Boleslav Brezina

Carabus (Pagocarabus) lama yajiangensis Deuve, 1994
Male adult
male, 23mm
Copyright © Boleslav Brezina

Carabus (Archicarabus) pseudomonticola Lapouge, 1908 ? nemoralis
Male adult
NE Spain, Catalonia, Sant Jaume de Frontanyà, h=1000 m, V-VI.2010, Pascal Stefani leg.
L=26 мм
Copyright © Ruslan Panin

Carabus (Pagocarabus) lama daochengensis Deuve, 1994
Male adult
male, 21mm
Copyright © Boleslav Brezina

Carabus (Archicarabus) pseudomonticola Lapouge, 1908
Male adult
Spain, Montseny, Barcelona
19.VI.1986 M. Tomas leg.
male, 21 mm
Copyright © Alexander Anichtchenko

Carabus (Pagocarabus) kamensis tatsienlui Breuning, 1934
Female adult

Copyright © Boleslav Brezina

Carabus (Pagocarabus) jiudingensis jiudingensis Deuve, 1994
Male adult

Copyright © Boleslav Brezina

Carabus (Pagocarabus) jiudingensis jiudingensis Deuve, 1994
Female adult

Copyright © Boleslav Brezina

Carabus (Pagocarabus) draco Brezina, 1999
Male adult

Copyright © Boleslav Brezina

Carabus (Pagocarabus) chomae (Imura, 2002)
Male adult

Copyright © Boleslav Brezina

Carabus (Pagocarabus) chomae (Imura, 2002)
Female adult

Copyright © Boleslav Brezina

Carabus (Pagocarabus) chomae (Imura, 2002)
Male adult
male, 24mm
Copyright © Boleslav Brezina

Carabus (Pagocarabus) caelestinus garpen Kalab, 2007 Holotype Paratype
Male adult

Copyright © Boleslav Brezina

Carabus (Pagocarabus) caelestinus garpen Kalab, 2007
Female adult
female, 21 mm
Copyright © Boleslav Brezina

Carabus (Pagocarabus) caelestinus caelestinus (Imura, 2007) Paratype
Male adult

Copyright © Boleslav Brezina

Carabus (Pagocarabus) borodini Heinz, 1996
Male adult
China, C Tibet, Mila 5000m, 150 km E of Lhasa 6.6.1997 Wrzecionko A. leg.
Copyright © Alexander Anichtchenko

Carabus (Pagocarabus) alpherakii takin Hackel & Sehnal, 2007 Holotype Paratype
Male adult
male, 19mm
Copyright © Boleslav Brezina

Carabus (Pagocarabus) alpherakii takin Hackel & Sehnal, 2007
Male adult
male, 17mm
Copyright © Boleslav Brezina

Carabus (Pagocarabus) alpherakii takin Hackel & Sehnal, 2007 Paratype
Female adult
female, 22mm
Copyright © Boleslav Brezina

Carabus (Pagocarabus) alpherakii takin Hackel & Sehnal, 2007 Paratype
Female adult
female, 23mm
Copyright © Boleslav Brezina

Carabus (Pagocarabus) alpherakii takin Hackel & Sehnal, 2007 Paratype
Male adult
male, 18 mm
Copyright © Boleslav Brezina

Carabus (Pagocarabus) alpherakii pignataroi Cavazzuti, 1996 Paratype
Female adult
female, 20mm
Copyright © Boleslav Brezina

Carabus (Pagocarabus) alpherakii pignataroi Cavazzuti, 1996
Male adult
male, 20mm
Copyright © Boleslav Brezina

Carabus (Pagocarabus) alpherakii pignataroi Cavazzuti, 1996
Male adult

Copyright © Boleslav Brezina

Carabus (Pagocarabus) alpherakii pignataroi Cavazzuti, 1996 Paratype
Female adult

Copyright © Boleslav Brezina

Carabus (Cupreocarabus) alpherakii alpherakii Semenov, 1898
Male adult
male, 23mm
Copyright © Boleslav Brezina

Carabus (Cupreocarabus) alpherakii alpherakii Semenov, 1898
Female adult
female, 24mm
Copyright © Boleslav Brezina

Carabus (Cupreocarabus) alpherakii alpherakii Semenov, 1898
Male adult
Chine, W Sichuan, Litang env., 3-6.7.99 Dr. V. Banes leg.
Copyright © Alexander Anichtchenko

Carabus (Pagocarabus) alpherakii budha Breuning, 1950
Male adult

Copyright © Boleslav Brezina

Carabus (Pagocarabus) alpherakii budha Breuning, 1950
Female adult
female, 20mm
Copyright © Boleslav Brezina

Carabus (Pagocarabus) alpherakii budha Breuning, 1950
Female adult
female, 24mm
Copyright © Boleslav Brezina

Carabus (Pagocarabus) alpherakii budha Breuning, 1950
Male adult
male, 20 mm
Copyright © Boleslav Brezina

Carabus (Pagocarabus) alpherakii budha Breuning, 1950
Male adult
male, 19mm
Copyright © Boleslav Brezina

Carabus (Apotomopterus) leda Kleinfeld, 2000 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) korellianus Kleinfeld, 2002 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) infirmior homodynamus Deuve, 1994 (as jiucaishan) Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) hera Kleinfeld, 2000 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) guzhangensis Kleinfeld, 1998 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) guizhouensis yunwuensis Kleinfeld, 1996 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) guangxicus Deuve, 1989
Male adult
Guposhan, Hezhou, Guangxi prov. China
male,
photo Daniel Prunier
Copyright © Daniel Prunier

Carabus (Apotomopterus) grossefoveatus grossefoveatus Hauser, 1913 (as zhongnanshan) Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) grossefoveatus annikae Kleinfeld, 1996 Holotype
Male adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) delavayi yunlongensis Kleinfeld, 2000 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) longeantennatus angulihabitus Kleinfeld, 1998 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) longeantennatus changyangensis Kleinfeld, 1998 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) longeantennatus yongshan Kleinfeld, 1999 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) mecynodes helena Kleinfeld, 1999 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) penelope penelope Kleinfeld, 1997 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) rapuzzi rapuzzi Kleinfeld, 2000 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) roxane roxane Kleinfeld, 2002 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) yunanensis yunanensis Born, 1905 Paralectotypes
Female adult

Copyright © Alexander Anichtchenko

Carabus (Apotomopterus) sauteri fujianensis Kleinfeld, 1997 Holotype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) sauteri sanjiangensis Kleinfeld, 2001 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) sauteri tianpingensis Deuve & Yu, 1992 (as eleganticollis chalingensis) Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) sauteri tianpingensis Deuve & Yu, 1992 (as longhuiensis) Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) sauteri tianpingensis Deuve & Yu, 1992 (as peterfrankei) Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) schuetzei Kleinfeld, 1998 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) shun gigantothoracis Kleinfeld, 1997 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) shun paraschuetzei Kleinfeld, 2002 Holotype
Male+Female adult

Copyright © Kirschenhofer

Carabus (Apotomopterus) solidior perangustus Kleinfeld & Schutze, 2003 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) songshanicus Kleinfeld, 1994 Holotype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) tonkinensis pandora Kleinfeld, 2000 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) tonkinensis tonkinensis Deuve, 1990
Female adult
N Viet Nam, Fan-si-pan mt., N seite Cha-pa (=Sapa)
1600 m, 20-30.4.95; 22.17N 103.44E Siniaev leg.
Copyright © Alexander Anichtchenko

Carabus (Apotomopterus) torquatus lushanicus Kleinfeld, 1996 Holotype
Male adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) vitalisi bernardlassallei A.Muller, 2005 Paratype
Female adult

Copyright © Boleslav Brezina

Carabus (Apotomopterus) vogtae artemis Kleinfeld, 2000 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) vogtae menghaiensis Kleinfeld, 2000 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) vogtae mojiangensis Kleinfeld, 2000 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) vogtae paravogtae Kleinfeld, 2000 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) vogtae thetis Kleinfeld, 2000 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) yuanshanensis Kleinfeld, 1996 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) yunanensis kassandra Kleinfeld, 1999 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) yunanensis leiboensis Kleinfeld, 1999 Holotype Paratype
Male+Female adult

Copyright © Kirschenhofer

Carabus (Carabus) bornianus bornianus Hauser, 1922 Lectotype
Male adult

Copyright © Frank Kleinfeld

Carabus (Carabus) dreuxioides wushanling Kleinfeld, 1998 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Carabus) emanuelei grabneri Kleinfeld, 2003 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Carabus) gressittianus daiyunshan Kleinfeld, 1998 Holotype
Male adult

Copyright © Frank Kleinfeld

Carabus (Carabus) gressittianus fuscomarginatus Kleinfeld, 1997 Holotype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Carabus) kucerai wuangxianus Kleinfeld, 1998 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Carabus) mianningensis Kleinfeld, 2000 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Carabus) nanosomus wufengensis Kleinfeld, 1998 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Carabus) oblongior yunlingicus Deuve, 1994
Male adult
China, NW Yunnan, Zhongdian env.
3400m, 5.7.2000
male, 22 mm
Copyright © Alexander Anichtchenko

Carabus (Carabus) pseudolatipennis wenxianensis Deuve, 1992
Male adult
China, S Gansu, Min Shan 2100 m,
SW Wudu 1.8.2000, V. Patrikeev leg.
Copyright © Alexander Anichtchenko

Carabus (Carabus) pseudolatipennis wenxianensis Deuve, 1992
Male adult
male, 20 mm
Copyright © Boleslav Brezina

Carabus (Carabus) tianbaoshan Kleinfeld, 1998 Holotype
Female adult

Copyright © Frank Kleinfeld

Carabus (Carabus) xupuensis Kleinfeld, 1998 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Archicarabus) gotschi gotschi Chaudoir, 1846
Male+Female adult
Transcaucasia, Armenia,
Ashotsk distr., Bandivan, 8-21.X.1991 P.V.Kazaryan leg.
female, 20mm
Copyright © Ruslan Panin

Carabus (Archicarabus) nemoralis nemoralis Muller, 1764
Male+Female adult
W Ukraine, NW Podolian,
Lviv reg., Zolochiv distr., near Vilshanitsya vill., Lysa Gora, 16-30.V.2009 R.Panin leg.
female, 26 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) nemoralis nemoralis Muller, 1764
Female adult

Copyright © Ruslan Panin

Carabus (Archicarabus) nemoralis prasinotinctus Heyden, 1880
Male+Female adult
AUDE.Pic de Nore, h=1200m.IX.2005, A. Laforgue leg.
female,25mm/male, 24mm
Copyright © Ruslan Panin

Carabus (Archicarabus) nemoralis prasinotinctus Heyden, 1880
Male adult
N Spain, 3 km N Bera de Bidasoa 14.9.2002
Anichtchenko A. leg.
long. 25mm
Copyright © Alexander Anichtchenko

Carabus (Archicarabus) rossii Dejean, 1826
Male adult
Lazio (Roma), Bosco di
Nettuno 22.5.1986 C. Iacovone leg.
male, 23 mm
Copyright © Alexander Anichtchenko

Carabus (Archicarabus) steuartii Deyrolle, 1852
Male adult
Portugal, S. Martinho, 27.5.1987
male, 18 mm
Copyright © Alexander Anichtchenko

Carabus (Archicarabus) victor victor Fischer, 1836
Male adult
Anatolia Or.
Artvin Heinz leg.
male, 18 mm
Copyright © Alexander Anichtchenko

Carabus (Archicarabus) victor wiedemanni Menetries, 1836
Male adult
Anatolia, Kocaeli, tra Kandira e Sile VII.2001
Solo Claudio leg.
long. 24mm
Copyright © Alexander Anichtchenko

Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Male adult
W.Ukraine, Transcarpathian Reg., Rakhov Distr., Chornogora Mt range h=2020m alpine zone, Panin R. leg. 28.VI-9.VIII.2008, male 19 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi rothi Dejean, 1829
Male+Female adult
Romania
Banat, Nemeseti h=250m. VI.1989 Lie P. leg.

photo Solodovnikov I.A.
Copyright © Igor Solodovnikov

Carabus (Neoplectes) apollo apollo Zolotarev, 1913
Male adult
Caucasus, Abchasia

Copyright © Dmitri Obydov

Carabus (Neoplectes) apollo phoebus Kurnakov, 1962
Male adult
Abchasia, Kodori, A. Zolotarew leg.
Photo by D.V. Obydov
from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Neoplectes) apollo tenebricosus Kurnakov, 1962
Male adult

Copyright © Safronov

Carabus (Pachystus) hungaricus gastridulus Fischer, 1823
Male adult
N. Krym Peninsula,
near Armyansk, Demidov G. leg.5.V.2006
male, 28mm
Copyright © Ruslan Panin

Carabus (Neoplectes) daphnis askhicus Belousov, 1992 Paratype
Male adult
SC-Caucasus, W Georgia, valley of riv. Chkhorodukhu, h=1000-1200m, 14.V.-6.VII.1991, Zamotajlov A.S & Miroshnikov leg.
male, 29 mm
photo & collection Panin R.
Copyright © Ruslan Panin

Carabus (Neoplectes) daphnis askhicus Belousov, 1992 Paratype
Male+Female adult
W Georgia, valley of Chkhorodukhu riv., 1000-1200m
14.V-6.VII.1991
male 30mm, female 33mm
photo & collection Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) daphnis askhicus Belousov, 1992 Paratype
Male adult
Georgia, env. Martvilli, Askhi plateau
10-15.7.2001 A.Rubenian leg.
27 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) daphnis daphnis Kurnakov, 1962
Male adult
Georgia, Mingrelia, Lecheha vallis, Tehuri superior
Caucasus centr., 1200m, 27.6-7.7.1989 Retezar I. leg.
male, 28 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) edithae exedithae Gottwald, 1985
Male adult
Karachaevo-Cherkessia, low cource of Daut river,
16-20.V.2009 Anichtchenko A. leg

Copyright © Alexander Anichtchenko

Carabus (Neoplectes) edithae exedithae Gottwald, 1985
Female adult
Karachaevo-Cherkessia, Zelenchukskii distr.,
near Bukovo vill., 8-13.VII.2007, S.Dementiev leg.

Copyright © Dmitri Obydov

Carabus (Neoplectes) edithae kabardensis Kurnakov, 1970
Female adult
Kabardino-Balkaria, Skalisty Mt Ridge, 1800 m,
20.VII.1996. Andreev leg.
Photo by D.V. Obydov

Copyright © Dmitri Obydov

Carabus (Archiplectes) jason Semenov, 1898
Male adult
Caucasus, W Georgia, Svanetia, upper reaches of riv. Shoudidi, 1700-2100m, 20-21.V.1986
leg. A. Zamotajlov
Copyright © Sunvas

Carabus (Neoplectes) jason Semenov, 1898
Male adult
Abkhasia, up river Shaudidi, 21.V - 2.VII.1986, Zamotailov A. leg.
Photo by D.V. Obydov

Copyright © Dmitri Obydov

Carabus (Neoplectes) juenthneri atchibachi Zamotajlov, 1991 Paratype
Female adult
Abkhasia,
Valley of river Bzyb near Mt. Achibakhu. h=1500m., A.S. Zamotajlov leg.
female: 33mm, 31mm
coll. R.Panin
Copyright © Ruslan Panin

Carabus (Neoplectes) juenthneri atchibachi Zamotajlov, 1991 Paratype
Female adult
W Caucas, Abkhasia, Valley of river Bzyb near Mt. Achibakhu. h=1500m.,
24.IV.-6.VI.1991 A.S. Zamotajlov leg.
coll. Anichtchenko
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) juenthneri atchibachi Zamotajlov, 1991 Paratype
Male adult
W Caucas, Abkhasia, Valley of river Bzyb near Mt. Achibakhu. h=1500m.,
24.IV.-6.VI.1991 A.S. Zamotajlov leg.
coll. Anichtchenko
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) juenthneri atchibachi Zamotajlov, 1991 Paratype
Male adult
W Caucas, Abkhasia, Valley of river Bzyb near Mt. Achibakhu. h=1500m.,
24.IV.-6.VI.1991 A.S. Zamotajlov leg.
coll. Anichtchenko
Copyright © Alexander Anichtchenko

Carabus (Megodontus) croaticus croaticus Dejean, 1826
Male adult
YU- Slovenija, Krim 1000 m, 30.10.1975 Drovenik leg.
Copyright © Alexander Anichtchenko

Carabus (Megodontus) caelatus caelatus Fabricius, 1801
Male adult
Bosnia, Trebevic, Vidikovac-Brus, 1200m
21.11.1985 Mihljevic B. leg.
Copyright © Alexander Anichtchenko

Carabus (Megodontus) bonvouloiri bonvouloiri Chaudoir, 1863
Female adult
Anatolia Orientale, Macka-Zigana VIII.1981
Copyright © Alexander Anichtchenko

Carabus (Megodontus) avinovi Semenov & Znojko, 1932
Male adult
Russia, Sakhalin isl., Susunaiskyi rng.,
Chehov Mt., 400m, 19.6.1990 Berlov E. leg.
female, 22 mm
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) juenthneri avadcharensis Kurnakov, 1970 (loc.typ.)
Male+Female adult
Abkhasia, near Avadhara, Site Pyv h~1850m, 43.490730 40.686433, 1.VI-12.VII.2009, Zamotajlov A.S., Bondarenko A.S. leg.
L=27mm

Copyright © Ruslan Panin

Carabus (Neoplectes) juenthneri avadcharensis Kurnakov, 1970
Male adult
Abkhasia, 7,4km NE from Ritza lake, 1,5 km N Kutykuh vill.,
1635 m, 19-25.VII.2008, A. Vlasenko leg.
Copyright © Dmitri Obydov

Carabus (Neoplectes) juenthneri dsychvensis Gottwald, 1985
Male adult
Abchasia, N Dsykhwa, coll. A. Zolotarew
Photo by D.V. Obydov
Copyright © Dmitri Obydov

Carabus (Neoplectes) juenthneri juenthneri Reitter, 1899
Female adult

Copyright © Alexander Anichtchenko

Carabus (Neoplectes) koltzei faunulus Kurnakov, 1970
Female adult
Abchasia, Akarmara, 6500 f, A. Zolotarew leg.
Photo by D.V. Obydov

Copyright © Dmitri Obydov

Carabus (Neoplectes) koltzei faunus Kurnakov, 1970
Male adult
Abkhazia, Kadorsky Mt Ridge, 1000 m, Chlou, 22.V.1993, A. Evstigneev leg.
Photo by D.V. Obydov

Copyright © Dmitri Obydov

Carabus (Neoplectes) koltzei kobachidzei Kurnakov, 1970
Female adult
Caucasus, E Abkhasia, SW slopes of Akiba mt. rng., 1200m, 7.V.-2.VII.1991 Zamotajlov & Miroshnikov leg.
Copyright © Sunvas

Carabus (Neoplectes) koltzei kobachidzei Kurnakov, 1970
Female adult
Caucasus, E Abkhasia, SW slopes of Akiba mt. rng., 1200m, 7.V.-2.VII.1991
Zamotajlov & Miroshnikov leg.
Copyright © Sunvas

Carabus (Neoplectes) koltzei kobachidzei Kurnakov, 1970
Male+Female adult
Caucasus, E Abkhasia, SW slope of Akiba mt.rng., 1200m, 7.V.-2.VII.1991 Zamotajlov & Miroshnikov leg.
male 37mm, female 38mm
photo & collection Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Archiplectes) koltzei Rost, 1889
Male adult
Abkhasia, Habyu 3.7.2002
Ivliev P. leg.
32 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) koltzei koltzei Rost, 1889
Male adult
Abkhasia, Tsebeldy, 23.V.1986, A.Zamotailov leg.
Photo by D.V. Obydov

Copyright © Dmitri Obydov

Carabus (Neoplectes) koltzei lennoni Gottwald, 1985
Male adult
Abkhasia, Amtkel lake, 22.V - 4.VII.1986, A.Zamotailov leg.
Photo by D.V. Obydov
from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Cratophyrtus) turcosinensis medvedevi Kryzhanovskij, 1968
Female adult
S Kazakhstan, NW slope of Ugamskii mt.rng., 3 km W Sairamskii peak, 2200m
tributary of Karzhantau riv., 15.6.2004 Matalin A. leg.
20 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) komarowi komarowi Reitter, 1882
Female adult
Centr. Caucasus, Svanetia, coll. A. Zolotarew
Photo by D.V. Obydov
from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Neoplectes) kratkyi felicitanus Reitter, 1893
Male adult
Russia, N Caucasus, Karachaevo-Cherkessia 2200m, Bolshoi Pzizer mt. 8.8.2002
25 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) kratkyi felicitanus Reitter, 1893 (as porcellus)
Male adult
NW Caucasus, Resp. Adygea, Mt.Tybga, 28.VI.1997
male,20mm
Copyright © Ruslan Panin

Carabus (Neoplectes) kratkyi felicitanus Reitter, 1893 (as porcellus)
Male adult
Prov. Kuban, mt. Tybga, 1300 m, VI.1913, Shaposhnikov leg.

Copyright © Dmitri Obydov

Carabus (Neoplectes) kratkyi felicitanus Reitter, 1893
Female adult
Rusia, Caucas, Dzhuga mt., 2200 m, 24.VI. - 22.VIII.1986, Zamotajlov A. leg.

Copyright © Dmitri Obydov

Carabus (Neoplectes) kratkyi Ganglbauer, 1890
Male+Female adult
W Caucas, 1900m, Khatsavitaya Mt., 4.VI-13.VII.1994
A.Miroshnikov leg.
male 25mm, female 26mm
photo & collection Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) kratkyi Ganglbauer, 1890
Male adult
Russia, NW-Caucasus: Krasnodar reg., Mostovskoy distr., Khatsavitaja Mt. h=1900m, 4.VI.-13.VII.1994, A. Miroshnikov leg. male, 26mm photo Panin R./det.Solodovnikov I.A.
Copyright © Ruslan Panin

Carabus (Neoplectes) kratkyi Ganglbauer, 1890
Male+Female adult
W Caucas Mts., 2500m, Mt. Yatyrgvarta,
25.VI-12.VIII.1992 Shchurov V. leg.
male 20mm, female 21mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890
Female adult
NW-Caucasus, Russia,
Karacaj-Cherkessija, Abishira-Akhuba range h=2650m, upper Kyafar-Agur Riv., 4-9.VIII.2009, Solodovnikov I.A. leg.
female, 23 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890
Male adult
NW-Caucasus: Karacaj-Cherkessija,
Abishira–Akhuba Mnt. Range h=2000-2650m, Kyafar-Agur Riv., 14–17.VII.2009, Fominykh D. leg.
male, 22mm
Copyright © Ruslan Panin

Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890
Male adult
Caucasus, Kuban, Meyer-Darcis-Patria

Copyright © Dmitri Obydov

Carabus (Neoplectes) kratkyi solodovnikovi Zamotajlov, 1992
Male adult
W. Caucasus, M. Bambak Mt., VI.1931, Vvedensky leg.
from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Archiplectes) plasoni dalensis Rost, 1893
Male adult
Abkhasia, near Yu. Ptysh vill.
14.7.1986 Zamotailov A. leg.
31 mm
Copyright © Alexander Anichtchenko

Carabus (Archiplectes) plasoni dalensis Rost, 1893
Male adult
Caucasus merid., Dal-Suchumi, N.S. Brianskyj leg.
from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Archiplectes) plasoni plasoni Ganglbauer, 1886
Female adult
Gruzia, Centr. Caucasus, Svanetia, coll. A. Zolotarew
det. D.V. Obydov
from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Neoplectes) polychrous Rost, 1892
Male adult
Abkhasia, Gudaut distr., near Huap vill., 600 m, 9.VI.1990, M. Danilevskyi leg. det. D.V. Obydov
Copyright © Dmitri Obydov

Carabus (Neoplectes) rousianus rousianus Gottwald, 1985
Male adult
Abkhasia, 7,6 km NE Pitsunda, 2,1 km NNE Ryapsh riv mouth, Ryapsh canyon h=60 m, 43°10N 40°25E, 25-27.VII.2008, A.Vlasenko leg. det. D.V. Obydov
Copyright © Dmitri Obydov

Carabus (Archiplectes) prometheus miroshnikovi Zamotajlov, 1990
Female adult
NW-Caucasus, Krasnodar reg., Mostovskoy distr., Site Val , h=1800m. 07-08.VII.2008, D.Fominyh leg., 35mm; det. I. Solodovnikov 2023
Copyright © Ruslan Panin

Carabus (Archiplectes) prometheus zolotarevi Zamotajlov, 1988
Male adult
Paratypus, male Prov. Kuban distr., Maikop, Laba Min., Ch. Shaposhnikov leg.
from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Archiplectes) prometheus zolotarevi Zamotajlov, 1988 Paratype
Male adult
Paratypus, male - Fl. Min. Laba, VII.1918, Shaposhnikov leg.
from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Archiplectes) prometheus basilianus Starck, 1890
Male adult
Russia, W Caucas, Port-Artur Mt. 22.VI.1986, A.Zamotajlov leg.
det. D.V. Obydov
from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Neoplectes) prometheus dzhugensis Zamotajlov, 1988 Paratype
Female adult
Paratypus, female Prov. Kuban circ., mt. Dzhuga (Tigeni), 27.VI.1911. 
Copyright © Dmitri Obydov

Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990
Male+Female adult
W.Caucasus,
Psebai env., Nikitino, 2.V-3.VII.1993, leg. A.S.Zamotajlov
female 33mm
Copyright © Ruslan Panin

Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990 (as hephaestus)
Male adult
Paratypus Carabus (Archiplectes) hephaestus
Maikop, Ust. Urusht, 26.IV.1910

Copyright © Dmitri Obydov

Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990
Female adult
W Caucasus, Krasnodar reg., Maikop distr.
Nikel, 1.7.1989
38 mm
Copyright © Alexander Anichtchenko

Carabus (Archiplectes) prometheus pschadensis Zamotajlov, 1990
Female adult
NW Caucas, 5 km N from Pshada vill., 25.04.1989, Smorodinov leg.
specimen from collection of M.Shestopalov (Moscow).
det. D.V. Obydov
from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Neoplectes) prometheus prometheus Reitter, 1887
Male adult
NW-Caucasus,Krasnodar reg., Apsheron Distr., Lagonaki Mt., near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominykh leg.
L= 42mm.
Copyright © Ruslan Panin

Carabus (Neoplectes) prometheus prometheus Reitter, 1887 (as zugmayeriae)
Male adult
W. Caucasus, Maikop, 20.VI.1967., A. Bogachev leg.
det. D.V. Obydov

Copyright © Dmitri Obydov

Carabus (Neoplectes) prometheus prometheus Reitter, 1887
Male+Female adult
NW Caucasus, Russia,
Krasnodar reg.,Krasnoarmeisk distr., Krasny Les, near Podlesnyi vill.,17-22.IV.2008, D.D.Fominykh leg.
female, 40mm
Copyright © Ruslan Panin

Carabus (Archiplectes) prometheus prometheus Reitter, 1887 (as nubicola)
Male+Female adult
W Caucasus, Adygea,
h~1700m. upper res. Riv. Kurdzhips, 11-21.VIII.2007, I.A.Solodovnikov leg./det.
L=32mm.
Copyright © Ruslan Panin

Carabus (Neoplectes) prometheus prometheus Reitter, 1887
Male adult
Krasnodar distr., Mezmai
20.6.1989
38 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) protensus olegi Belousov, 1992 Paratype
Male adult
Caucasus, W Georgia, Svanetia, right bank of riv. Magana, 800m, 12.V.-4.VII.1991, Zamotajlov & Miroshnikov leg.
male, 31mm
photo and collection Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) protensus protensoides Novotny & Vorisek, 1988
Male adult
Georgia, Martvilli env., Akshi plateau 10-15.7.2001
Rubenian leg.
32 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) reitteri gagrinus Starck, 1894
Female adult
Abkhasia, near Gagry, 16.V.1992, A. Evstigneev leg.
det. D.V. Obydov
Copyright © Dmitri Obydov

Carabus (Neoplectes) reitteri laevisternis Starck, 1890
Male adult
Abkhasia,W.slope of Gagrinsky Mt., Krepostnaya River source, h=1325m., A.S.Zamotajlov leg. 27.VII.2000
male, 29 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) reitteri laevisternis Starck, 1890
Male adult
W-Caucasus:Sochi env., Aibga Mt. h=2000m., 06.VII.2000
male, 29mm
Copyright © Ruslan Panin

Carabus Neoplectes) reitteri reitteri Retowski, 1885
Female adult
NW Caucasus, Adygeya, Fischt-Oshten Mts. base, h=1800 m. 20.07.2006, D.Safronov leg.
Copyright © Safronov

Carabus (Neoplectes) reitteri reitteri Retowski, 1885 (as achunensis)
Male adult
W. Caucasus, Sochi Distr., Chernomoretz Vill. env.,
15.VII.1992, P. Gorbachev leg.
det. D.V. Obydov

Copyright © Dmitri Obydov

Carabus (Neoplectes) satyrus besleticus Kurnakov, 1972 (as gaskoi)
Male adult
USSR, Caucasus occid, Abchasia, Suchumi, vallis Kelasuri, 100m 25.6-17.7.1977 Retezar I. leg. 36 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) satyrus mtsaranus Kurnakov, 1972
Male adult
Abkhasia, Bzhybskii Mt.rng., near Mtsara vill., 16.VI.1991, S.Gusev leg.
det. D.V. Obydov

Copyright © Dmitri Obydov

Carabus (Neoplectes) satyrus napraensis Belousov & Zamotajlov, 1993 Paratype
Male adult
Abkhasia, Bzybian Mt. Range near Mt. Chibzharga., h=1900-2000m., A.S. Zamotajlov leg.
male, 28mm
photo & coll.Panin R.
Copyright © Ruslan Panin

Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972
Male adult
Abkhasia, Habyu vill.,2.7.2002 Ibliev leg. 35 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972
Male adult
Abkhasia, Habyu vill. 2.7.2002 Ibliev leg. 35 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) starcki Heyden, 1885
Male+Female adult
NW-Caucasus,
Krasnodar reg., Mt. Aishkho, Malaya Laba River, h=2200m., 14.VII.2008, D.Fominyh leg.
female-21mm
Copyright © Ruslan Panin

Carabus (Neoplectes) starcki Heyden, 1885
Male adult
Caucasian Nat. Park, S slope
Pseashkho mt. 2500m 21.7.1975, Ovtsharenko leg.
20 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) starcki Heyden, 1885
Male adult
NW Caucasus, Adygeya, Pshekho-Su Mt. env., h=1500 m., 21.07.2006, D.Safronov leg.
Copyright © Safronov

Carabus (Neoplectes) starcki Heyden, 1885 (as parvulus)
Male adult
Prov. Kuban, mt. Aischa, VI.1906, Shaposhnikov leg.
det. D.V. Obydov

Copyright © Dmitri Obydov

Carabus (Neoplectes) starckianus adelaidae Starck, 1889
Male adult
Russia, Caucasus, Chvizhepse River Val., 27.V.1994, P. Gorbachev leg.
Photo from www.zin.ru
det. D.V. Obydov
Copyright © Dmitri Obydov

Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988 (as shchessiensis) Paratype
Male+Female adult
NW Caucasus,
Krasnodar reg., Tuapsinskyi distr. Shessi Mt.
30.V.-25.VI.1993 Mt.Shessi, h=1800m., Imura & A.S.Zamotajlov, 1993 leg.
female, 29mm
Copyright © Ruslan Panin

Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988
Female adult
Russia,W Caucasus,
Krasnodar reg.,Chernogorie mts.,Chernogor mt. h=1700-1800m. 23.V.-30.VII.1999. A.S.Zamotajlov leg.
female 29mm.
Copyright © Ruslan Panin

Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988
Male adult
West. Caucasus, Shakhe Valley, 1.VI.1989, A. Shamaev leg.
det. D.V. Obydov

Copyright © Dmitri Obydov

Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
Male adult
NW-Caucasus, Russia,
Krasnodar reg., Apsheron Distr., Lagonaki Mt. ,near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominykh leg.
male, 28mm
Copyright © Ruslan Panin

Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
Male+Female adult
NW-Caucasus, Russia,
Krasnodar reg.,the right bank of Mzymta river, Engelmanovs glades, h=1200m., A.S.Zamotajlov leg.
female-30mm\male-28mm
Copyright © Ruslan Panin

Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
Female adult

Copyright © Safronov

Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
Male adult
Rusia, W Caucasus, Sochi distr., Aishkha mts.
900m, 15.7.2003 Gorbachev leg.
30 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) starckianus starckianus Ganglbauer, 1886
Male adult
Russia, Krasnodar reg., 6 km NE Soloniki vill.,
Sochi Nat.Park, 28-30.V.2009 Anichtchenko A. leg.

Copyright © Alexander Anichtchenko

Carabus (Neoplectes) starckianus starckianus Ganglbauer, 1886
Female adult
W Caucas, Sochi distr., near Lazorevskoe vill., 9.06.1993, P.Gorbachev leg.
det. D.V. Obydov

Copyright © Dmitri Obydov

Carabus (Neoplectes) starckianus starckianus Ganglbauer, 1886 (as phlagochensis)
Female adult
Caucasus, Sotschi, Flagoch.
det. D.V. Obydov
from www.zin.ru

Copyright © Dmitri Obydov

Carabus (Neoplectes) starckianus theseus Briansky, 1910
Male+Female adult
NW-Caucasus, Russia,
Krasnodar reg.,Tuapse distr., Mt.Semashkho 04.V.2008, D.Fominyh leg.
female, 32mm
Copyright © Ruslan Panin

Carabus (Neoplectes) starckianus theseus Briansky, 1910
Male adult
Tuapse distr., Anastasievka vill., Semiglavaya mt.
riv. Pshekaho 20.7.1995
30 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) starckianus theseus Briansky, 1910
Female adult
Tuapse distr., Anastasievka vill., Semiglavaya mt.
bass. Pshekaho riv. 20.7.1995
female, 30 mm
Copyright © Alexander Anichtchenko

Carabus (Aristocarabus) viridifossulatus viridifossulatus Fairmaire, 1887
Female adult
35mm
Copyright © Boleslav Brezina

Carabus (Aristocarabus) viridifossulatus viridifossulatus Fairmaire, 1887
Male adult
29 mm
Copyright © Boleslav Brezina

Carabus (Aristocarabus) viridifossulatus aleatus Cavazzuti, 2003
Male adult
26 mm
Copyright © Boleslav Brezina

Carabus (Aristocarabus) viridifossulatus gonggashanensis Deuve, 1992
Male adult
32 mm
Copyright © Boleslav Brezina

Carabus (Aristocarabus) viridifossulatus jiajiensis Deuve, 1998
Male adult
27 mm
Copyright © Boleslav Brezina

Carabus (Aristocarabus) viridifossulatus kvetoslavae Sehnal, Farkac & Hackel, 2006 Holotype Paratype
Female adult
36 mm
Copyright © Boleslav Brezina

Carabus (Aristocarabus) viridifossulatus kvetoslavae Sehnal, Farkac & Hackel, 2006
Male adult
30 mm
Copyright © Boleslav Brezina

Carabus (Aristocarabus) viridifossulatus kvetoslavae Sehnal, Farkac & Hackel, 2006 Paratype
Female adult
35 mm
Copyright © Boleslav Brezina

Carabus (Aristocarabus) viridifossulatus phamianus Deuve, 1990
Male adult
27 mm
Copyright © Boleslav Brezina

Carabus (Aristocarabus) viridifossulatus seticollis Deuve & Mourzine, 2000
Male adult
28 mm
Copyright © Boleslav Brezina

Carabus (Aulonocarabus) canaliculatus rufinus Beheim & Breuning, 1943
Female adult

Copyright © Safronov

Carabus (Aulonocarabus) canaliculatus canaliculatus Adams, 1812
Male adult

Copyright © Safronov

Carabus (Aulonocarabus) canaliculatus jankowskiellus Deuve, 1991
Female adult
CHINA, Jilin prov., Hongshi
forest plant
Copyright © Anton Menstschikov

Carabus (Aulonocarabus) careniger careniger Chaudoir, 1863
Male adult
S Primorie, Chernye mts.,
Poima riv., 30.07.- 6.08.1999 Anichtchenko A. leg
male, 25 mm
Copyright © Alexander Anichtchenko

Carabus (Aulonocarabus) gossarei imanensis Lafer, 1989
Male adult
Primorskii reg., Chug. distr., Oblachnaya mt.,
Zabytyi, 11.7.1977 Kuznetsov V. leg
male, 19 mm
Copyright © Alexander Anichtchenko

Carabus (Aulonocarabus) kolymensis kodarensis Brinev, 1997
Male adult
NE Chita reg., Kodar mt. rng., up of Chara riv.,
15 km N Leprindo, 2200m 5-15.7.2001 Brinev A. leg.
Copyright © Alexander Anichtchenko

Carabus (Aulonocarabus) kolymensis viridicupreior Deuve & Mourzine, 1993
Male adult
SE Yakutia, Cherskii Mt.rng., Garmychan Mts., 20 km NNE Sasyr,
Kalgar riv., 7.VIII.1998 Nogovitsyna leg.
Copyright © Alexander Anichtchenko

Carabus (Aulonocarabus) kolymensis viridicupreior Deuve & Mourzine, 1993
Female adult
10.VII.2001 NE RUSSIA, Magadan, near Karmaken vill
Copyright © Nil Belonogov

Carabus (Aulonocarabus) kurilensis kurilensis Lapouge, 1913
Male adult
Rusia, Sakhalin isl., 44 kordon
6.08.1991
male, 20mm
Copyright © Alexander Anichtchenko

Carabus (Aulonocarabus) kurilensis pseudodiamesus Ivanovs, 1994
Male adult
Russia, Sakhalin isl., Susunaiskii mg.
Chehov mt., 19.6.1990 Berlov E. leg.
male, 22 mm
Copyright © Alexander Anichtchenko

Carabus (Aulonocarabus) truncaticollis polaris Poppius, 1905
Male adult
Arctic Russia, NW Siberia,
Yamal, 230 N from Labytnangi, Panta-Yakha Riv., 15-29.VI.2002
male,18mm.
Copyright © Ruslan Panin

Carabus (Aulonocarabus) truncaticollis polaris Poppius, 1905
Female adult
N.Russia, Polar Ural, Paypudynya River, Pompeliz Mt., h=500-700 m.
10-15.VII.2002 S. Andreev, A. Romanenko leg.
Copyright © Safronov

Carabus (Axinocarabus) fedtschenkoi fedtschenkoi Solsky, 1874
Male adult
27 mm
Copyright © Boleslav Brezina

Carabus (Axinocarabus) fedtschenkoi fedtschenkoi Solsky, 1874
Female adult
30 mm
Copyright © Boleslav Brezina

Carabus (Axinocarabus) fedtschenkoi kondarensis Kryzhanovskij, 1953
Male adult
Tadzhikistan, Varzhobskoe usch., kishlak Navobob
4.1994 Avdeev I. leg.
Copyright © Alexander Anichtchenko

Carabus (Axinocarabus) fedtschenkoi kondarensis Kryzhanovskij, 1953
Female adult
31 mm
Copyright © Boleslav Brezina

Carabus (Axinocarabus) fedtschenkoi miles Semenov, 1886
Male adult
27 mm
Copyright © Boleslav Brezina

Carabus (Axinocarabus) fedtschenkoi miles Semenov, 1886
Female adult
29 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) aristochroides aristochroides Deuve, 1992
Male adult
China Sichuan N-Occ. 25 km Nord Maniganggo
4050 m, VI.2001 Cavazzuti leg.
Copyright © Pierfranco Cavazzuti

Carabus (Calocarabus) aristochroides qagcaensis Deuve, 1995
Male adult
17 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) aristochroides qagcaensis Deuve, 1995
Female adult
19 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) deuveianus Cavazzuti & Casale, 2006 Holotype
Male adult
Cina, Gansu mer., Qiling Shan, Passo a Nord di Tewo 4000m
Guan Gai Shan. Colle mer. 18.VII.2005 Cavazzuti leg.
Copyright © Pierfranco Cavazzuti

Carabus (Calocarabus) deuveianus Cavazzuti & Casale, 2006
Male adult
17 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) dietererberi Heinz, 2001
Female adult
20 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) gologensis Deuve e Tian 2004
Male adult
18 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) gratus gratus Semenov, 1887
Female adult
17 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) gratus nishidai Deuve & Imura, 1993 Paratype
Male adult

Copyright © Boleslav Brezina

Carabus (Calocarabus) gratus nishidai Deuve & Imura, 1993 Paratype
Female adult
19 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) gratus serxuensis Deuve, 1995
Male adult
17 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) guinanensis guinanensis Deuve, 1991
Male adult
16 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) guinanensis guinanensis Deuve, 1991
Female adult
17 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) guinanensis maqenicola Kalab, 2005 Paratype
Male adult
China, SE Qinghai mts. 15 km E Maqen, 34.27N/100.24E 4300m
J.Kalab leg., 22-24.7.2004 alpine meadows, screes
Copyright © Pierfranco Cavazzuti

Carabus (Calocarabus) janatai Brezina, 1996 Paratype
Female adult
21 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) janatai Brezina, 1996
Male adult
18 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) janatai Brezina, 1996
Male adult
19 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) juengerianus Kleinfeld, 1995 Paratype
Male adult
18 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) juengerianus Kleinfeld, 1995
Female adult

Copyright © Boleslav Brezina

Carabus (Calocarabus) juengerianus Kleinfeld, 1995
Female adult
20 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) juengerianus Kleinfeld, 1995 (as yangguifei)
Male adult
17 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) juengerianus Kleinfeld, 1995
Male adult

Copyright © Boleslav Brezina

Carabus (Calocarabus) juengerianus Kleinfeld, 1995 (as yangguifei)
Male adult
18 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) juengerianus Kleinfeld, 1995 (as yangguifei) Paratype
Female adult
20 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) linxiaensis darcangicus Deuve, 1995 Paratype
Female adult
20 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) linxiaensis darcangicus Deuve, 1995 Paratype
Male adult
18 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) linxiaensis linxiaensis Deuve, 1992
Male adult
17 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) linxiaensis linxiaensis Deuve, 1992
Female adult
18 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) linxiaensis linxiaensis Deuve, 1992
Male adult

Copyright © Boleslav Brezina

Carabus (Calocarabus) linxiaensis ponggartangensis Deuve, 1995 Holotype
Male adult
17 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) linxiaensis pseudogratus Deuve, 1994 Paratype
Male adult
19 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) linxiaensis pseudogratus Deuve, 1994
Male adult
17 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) linxiaensis pseudogratus Deuve, 1994
Male adult
17 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) linxiaensis pseudogratus Deuve, 1994
Male adult
17 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) linxiaensis pseudogratus Deuve, 1994 Paratype
Male adult
China, NW Sichuan, 45 km S Zamtang 31.52N 100.59E,
4600 m, 15-19.7.2001 Rejsek leg.
Copyright © Alexander Anichtchenko

Carabus (Calocarabus) linxiaensis subacidus Cavazzuti, 2002 Holotype
Male adult
China, Sichuan N-Occ., Sertar-Garze, 50 km a Nord del bivio Luhuo-Garze,
4400.VI.2001 Cavazzuti leg.
Copyright © Pierfranco Cavazzuti

Carabus (Calocarabus) mandarin Kalab, 2002 Paratype
Male adult
17 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) turnaianus Deuve, 1995
Male adult
China, Sichuan N-Occ., 20 km Sud di Qagca, Lala Shan 42-4500m
VI.2001 Cavazzuti leg.
Copyright © Pierfranco Cavazzuti

Carabus (Calocarabus) trichothorax trichothorax Brezina & Imura, 1997
Male adult
22 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) trichothorax trichothorax Brezina & Imura, 1997
Female adult
22 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) pingpong Brezina & Hackel, 2004 Holotype Paratype
Male adult
CH, S Gansu, SSW Minxian (12) W Qilian Shan
11 km W of Kunda 34:08N/103:45E/ 4000-4300m
alpine zone 9-10.VII.2002 Belousov & Kabak
Copyright © Boleslav Brezina

Carabus (Calocarabus) pingpong Brezina & Hackel, 2004
Male adult
CH, S Gansu, SSW Minxian (12) W Qilian Shan
11 km W of Kunda 34:08N/103:45E/ 4000-4300m
alpine zone 9-10.VII.2002 Belousov & Kabak
Copyright © Boleslav Brezina

Carabus (Calocarabus) pingpong Brezina & Hackel, 2004 Paratype
Female adult
20 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) pingpong Brezina & Hackel, 2004 Paratype
Male adult
18 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) przewalskii somoli Deuve, 2007 Holotype
Male adult
16 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) przewalskii somoli Deuve, 2007
Female adult
19 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) przewalskii przewalskii Morawitz, 1886 (as elnarae)
Male adult
CINA, Qinghai Nord-Or. Passo fre Daliang e Obo
3600 m, 11.VII.2005 Cavazzuti leg.
Copyright © Pierfranco Cavazzuti

Carabus (Calocarabus) przewalskii przewalskii Morawitz, 1886
Male adult
17 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) przewalskii przewalskii Morawitz, 1886
Female adult

Copyright © Boleslav Brezina

Carabus (Calocarabus) przewalskii przewalskii Morawitz, 1886
Male adult

Copyright © Boleslav Brezina

Carabus (Calocarabus) przewalskii przewalskii Morawitz, 1886
Female adult
19 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) przewalskii sewertzowi Semenov, 1898 Holotype
Male adult
17 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) przewalskii sewertzowi Semenov, 1898
Male adult
19 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) przewalskii sewertzowi Semenov, 1898
Female adult
20 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) przewalskii paulusianus Deuve, 2007
Male adult
19 mm
Copyright © Boleslav Brezina

Carabus (Carabus) arvensis arvensis Herbst, 1784
Male+Female adult
W Ukraine, NW Podolian, Lviv reg., Javoriv distr., near Stradch vill., 20-30.IV.2009, Panin R. leg.
Copyright © Ruslan Panin

Carabus (Carabus) arvensis arvensis Herbst, 1784
Male adult
W Ukraine, Male Polissya, Lviv reg., near Kamenka-Bugzkaya, under the bark of oak tree, in the forest, 29.III.08, Panin R. leg. male,17mm
Copyright © Ruslan Panin

Carabus (Carabus) arvensis arvensis Herbst, 1784
Male adult
W. Ukraine , NW. Podolian, Lviv reg., Nikolaev distr., near Krasiv vill., 9.II.08, Panin R. leg. L=17mm
Copyright © Ruslan Panin

Carabus (Carabus) arvensis conciliator Fischer, 1822 (as jurgitae)
Male adult
Russland, Altai, Aktasch
leg. Nikolaevsky, 1991; det. U.Schmidt, 1995
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Carabus (Carabus) arvensis baschkiricus Breuning, 1936
Female adult
Chebarkul, Chelyabinsk area 2.08.2008
Copyright © Evgeniy Chibilov

Carabus (Carabus) arvensis carpathus Born, 1902
Male+Female adult
Ukrainian Carpathians,
Transcarpathian reg., Rakhov Distr., Chornohora range, h=1850 -2000m., 29.VI-28.VII.2007 Panin R. leg.
18mm.
Copyright © Ruslan Panin

Carabus (Carabus) arvensis deyrollei Gory, 1839
Male adult
Spain, Asturias, 1-8.7.1999
long. 17mm
Copyright © Alexander Anichtchenko

Carabus (Carabus) arvensis faldermanni Dejean, 1829
Female adult
Rusia, S Primorskii reg., Poima riv., 30.VII-6.VIII.1999
Anichtchenko A. leg.
long. 22mm
Copyright © Alexander Anichtchenko

Carabus (Carabus) arvensis faldermanni Dejean, 1829
Female adult
Rusia, Evreiskaja AO., near Obluchie 20.6.93
Chichvarhin A. leg.
long. 22mm
Copyright © Alexander Anichtchenko

Carabus (Carabus) granulatus leander Kraatz, 1878
Male+Female adult
Russia, C-Caucasus: Balkaria, Valley of riv. Cherek Balkarskyi, h=1750-1800m 10.VI-20.VII.1992, Zamotajlov A. leg.
female,22mm
Copyright © Ruslan Panin

Carabus (Carabus) granulatus calabricus Spettoli & Vigna Taglianti, 2001
Male adult
Italia, Pollino, San Sever. Luc. Bosco Magnano,
(PZ) 13.X.2002 800m, F.Angelini leg.
long.18mm
Copyright © Alexander Anichtchenko

Carabus (Carabus) granulatus duarius Fischer, 1844
Male+Female adult
SE Siberia, Russia
Irkutsk reg., near Pivovarikha vill., 7.V.07, D.Fominyh leg.
female, 22mm
Copyright © Ruslan Panin

Carabus (Carabus) granulatus granulatus Linne, 1758
Male adult
W Ukraine,
Lviv Reg., Javoriv distr., near Ivano-Frankovo vill., 30.IV-18.V.2009, Panin R. leg.
male, 21mm
Copyright © Ruslan Panin

Carabus (Carabus) granulatus granulatus Linne, 1758
Male adult
23 mm
Germania, Oberfranken, Bad Steben
leg.det. U.Schmidt, 2006
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Carabus (Carabus) granulatus granulatus Linne, 1758
Female adult
Polonia, Verh. Zalesie, V.1994 Strelnikov D. leg.
long. 21mm
Copyright © Alexander Anichtchenko

Carabus (Carabus) granulatus granulatus Linne, 1758
Female adult
long. 22mm
Copyright © Ruslan Panin

Carabus (Carabus) granulatus granulatus Linne, 1758
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Carabus (Carabus) granulatus interstitialis Duftschmid, 1812
Male adult
Austria, Murek, X.1971 leg. Orselli
long. 22mm
Copyright © Alexander Anichtchenko

Carabus (Carabus) granulatus yezoensis Bates, 1883
Male adult
S Sakhalin isl., Novoaleksandrovka, Susun riv., 29.V.90 long. 19mm
Copyright © Alexander Anichtchenko

Carabus (Carabus) menetriesi menetriesi Hummel, 1827
Male adult
W-Ukraine, Roztochia, Lviv Reg., Yavorov distr., near Ivano-Frankovo vill., Site Zalyvki, Panin R. leg., 1-20.V.2005, male, 21 mm.
Copyright © Ruslan Panin

Carabus (Carabus) stscheglowi Mannerheim, 1827
Female adult
E Ukraine,
Dniepropetrovsk reg.,Novomoskovskiy distr. near Andriivka vill.16.VII.06 Kanarsky Yu.leg.
female 23mm
Copyright © Ruslan Panin

Carabus (Carabus) stscheglowi Mannerheim, 1827
Male+Female adult
Central Russia , Voronezh region, near Grafskoe, 12.VIII.1984 long=20mm
Copyright © Ruslan Panin

Carabus (Carabus) vanvolxemi Putzeys, 1875
Male adult

Copyright © Alexander Anichtchenko

Carabus (Cathaicus) brandti aishanensis Kleinfeld, 1999 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Cathaicus) brandti brandti Faldermann, 1835
Male adult
China, Beijing, Zhongguanchun, in the flower bed, 2007, Liu ye leg.; Photo & det. by liuye
Copyright © Ye Liu

Carabus (Cathaicus) brandti brandti Faldermann, 1835
Male adult
28 mm
Copyright © Boleslav Brezina

Carabus (Cathaicus) brandti brandti Faldermann, 1835
Female adult
27 mm
Copyright © Boleslav Brezina

Carabus (Acathaicus) interruptocostatus fantingi Imura & Su, 1998 Holotype
Male adult
China, N.-Sichuan, above Jiuzhaigou, 2100-2200m, N bank of Riv. Bai He, Jiuzhaigou Xian" - 24 mm lang - in coll. Imura.
Copyright © Frank Kleinfeld

Carabus (Rhigocarabus) wengdaensis Kleinfeld & Puchner, 2009 Holotype
Male adult
HOLOTYPUS Sichuan, Str. Wengda-Luhuo, Pass 20 km südl. Wengda, 4100m (ca. 31°43'N/100°44'E), VIII.2007, leg. Ding - coll. Puchner 20mm lang / 7 mm breit
Copyright © Frank Kleinfeld

Carabus (Acathaicus) interruptocostatus nanpingicus Deuve, 1995 Holotype
Male adult
Chine, Sichuan, Nanping, VII.1995, S. J. L. Remond" - Länge 25 mm - in MNHN Paris.
Copyright © Frank Kleinfeld

Carabus (Acathaicus) interruptocostatus amplior Deuve, 1994 Holotype
Male adult
Chine, Gansu, Min Shan, 2000m, J. M. Bousquet, 1994" - Länge 29 mm - in MNHN Paris
Copyright © Frank Kleinfeld

Carabus (Acathaicus) interruptocostatus tewoicus Deuve, 1996 Holotype
Male adult
Chine, Gansu, Tewo, 2000m, J. L. Remond, SR. 24,5 mm, in MNHN Paris
Copyright © Frank Kleinfeld

Carabus (Imaibius) reuteri Kleinfeld, 2008 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Platycarabus) fabricii fabricii Panzer, 1812
Male adult
NE-Italy,
Trentino-Alto Adige Reg,. Bolzano-Bozen Prov.,Val Aurina – Rottal 2050-2150m, 28.VI.1997, I.Rapuzzi leg.
male, 19mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) pedemontanus maurinensis Deuve, 2002 Holotype
Male adult

Copyright © Frank Kleinfeld

Carabus (Imaibius) pachtoun kabakovianus Gottwald, 1992 Holotype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Imaibius) pachtoun kabakovianus Gottwald, 1992
Male adult

Copyright © Frank Kleinfeld

Carabus (Imaibius) nouristani nouristani (Ledoux, 1977)
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Imaibius) anami Ledoux, 1977
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Imaibius) pachtoun pachtoun (Ledoux, 1975) Paratype
Male adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) angermaieri Kleinfeld, 2008 Holotype Paratype
Male+Female adult
China, Guangxi, Rong'an (=Chang'an: 25°13'109°23'), Durongshan, 1500m, V.2007, leg. Li et al. in coll. Kleinfeld
male 30mm, female 32 mm
Copyright © Frank Kleinfeld

Carabus (Pagocarabus) qiangding Kleinfeld & Puchner, 2007 Holotype Paratype
Male+Female adult
China. Sichuan, Str. zwischen Yajiang und Kangding, Tagong Grasslands (30°02'59,3''N/101°34'14,3''E)
28 mm
Copyright © Frank Kleinfeld

Carabus (Pagocarabus) thibetanus Breuning, 1950 Holotype
Male adult
Tong-ho River, Szetschuan, VII.1947, 4.000m / C. Plesius thibetanus mihi, Typ / HOLOTYPE Coll. Frey, NHM Basel ? 27 mm
Copyright © Frank Kleinfeld

Carabus (Coptolabrus) formosus elytraviridis Kleinfeld, 2007 Holotype Paratype
Male+Female adult
China, Sichuan, Aba pref., Nanping Co., Jiuzhaigou (ca. 33.17N/103.54E), 2000 m, 30.5.-6.6.1990, coll. Kleinfeld.
HT 32 mm, PT 36 mm
Copyright © Frank Kleinfeld

Carabus (Carabus) kucerai puchneri Kleinfeld, 2007
Male adult
China, Sichuan, road Pingwu-Nanping, Umg. Mupi Tibetan (Mupi 32.33N/104.33, 1250 m, 28.5.-4.6.2006),
27 mm, coll. Puchner
Copyright © Frank Kleinfeld

Carabus (Cathaicus) brandti brandti Faldermann, 1835 (as grossegranulatus) Syntypes Lectotype
Male adult
C. (Cathaicus) brandti grossegranulatus Breuning 1943 Lectotypus/SyntypusLectotypus (25 mm) von "South Mandschou". Syntypus (25 mm) "South Mandschou, 30. V. 1937"
Copyright © Frank Kleinfeld

Carabus (Cathaicus) brandti brandti Faldermann, 1835 Syntypes Lectotype
Male adult
C. (Cathaicus) brandti grossegranulatus Breuning 1943
Lectotypus/SyntypusLectotypus (25 mm) von "South Mandschou". Syntypus (25 mm) "South Mandschou, 30. V. 1937"
Copyright © Frank Kleinfeld

Carabus (Cathaicus) brandti reginae Deuve, 1997 Holotype
Male adult
Ex Ningxia, Suyuko, Helan Shan 2800 m, 2.VII.1996.
MNHN Paris. 30 mm
Copyright © Frank Kleinfeld

Carabus (Carabus) sculpturatus Menetries, 1832
Male+Female adult
Azerbaijan,
Talysh Mts. Hirkan National Park, Moscow Forest, 8-9.VII.1988, P.V.Kazaryan leg.
female, 22 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) estreicheri Fischer, 1822
Female adult
Volgograd Reg., Serafimovka Distr., Ust' -Khoperskaya Station 5.V.2009 leg.O.Sveshnikov
18.5 mm
Copyright © Kirill Makarov

Carabus (Hygrocarabus) nodulosus Creutzer, 1799
Male+Female adult
Austria, Styria Region (Steiermark), Leibnitz, 31.X.1999, I.Rapuzzi leg. female, 29 mm
Copyright © Ruslan Panin

Carabus (Hygrocarabus) variolosus Fabricius, 1787
Male adult
W-Ukraine, NW Podolian, Lviv reg., Zhydachiv distr.,near Vybranivka vill., 15.V-8.VI.2008, Panin R. leg. male, 27mm
Copyright © Ruslan Panin

Carabus (Hygrocarabus) variolosus Fabricius, 1787
Male+Female adult
W-Ukraine, Lviv Reg., Zhydachiv distr., near Vybranivka vill., Panin R. leg. female, 30mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) lambrechti Kleinfeld, 2006 Holotype
Male adult
26 mm
Copyright © Frank Kleinfeld

Carabus (Apotomopterus) pseudoguizhouensis Kleinfeld, 2006 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Platycarabus) irregularis irregularis Fabricius, 1792
Male adult
NE-Italy,
Friuli Reg., Udine, Carnia, Pierabech h~1100m, I.Rapuzzi leg.
male, 23 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) shun emeifeng Kleinfeld, 2006 Holotype
Male adult
China, Fujian, Taining Co., SW Xinqiao: Emeifeng Shan (ca. 26.59N/117.05E), 1500 m, 34 mm, coll. Kleinfeld
Copyright © Frank Kleinfeld

Carabus (Apotomopterus) delavayi planithoracis Kleinfeld, 2006 Holotype
Male adult
30 mm,
China, Süd-Yunnan, Lüchün (23.00N/102.20E): Shanbashan, 2200 m, 9/2005, coll. Kleinfeld
Copyright © Frank Kleinfeld

Carabus (Imaibius) nouristani bandaensis Kleinfeld & Reuter, 2007 Holotype
Male adult
Afghanistan, prov. Konar, NE Jalalabad, Schamschehr - Banda-Wald (ca. 34.44N/70.50E), 2800 m
23 mm
Copyright © Frank Kleinfeld

Carabus (Eucarabus) manifestus pyonganicola Deuve & Li, 1998
Male+Female adult
N-Korea: Pyonganam-Do, YangTok County,(39;127), 12-19,VI.2006
Copyright © Ruslan Panin

Carabus (Platycarabus) irregularis ramanus Sokolar, 1909 (as hayeki Kobmann, 1925)
Male adult
N Slovenia, Maribor – Dravograd, Pomorje Mts.,13.XI.2005, I.Rapuzzi leg.
Copyright © Ruslan Panin

Carabus (Platycarabus) irregularis ramanus Sokolar, 1909 (as bucephalus Kraatz, 1879)
Male+Female adult
W Slovenia, Goriška region Trnovo, h~1000 m, IX.2002, I.Rapuzzi leg.
female, 29 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens trans illigeri Dejean, 1826
Male+Female adult
W-Croatia, Karlovac, near Bosiljevo vill., h~300m, V.2000, I.Rapuzzi leg. female, 33 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) creutzeri creutzeri Fabricius, 1801 (as trnovensis Bernau, 1914)
Male adult
W Slovenia,
Lokve, Trnovski Gozd h=1100m, 17.IV.2006, leg. I.Rapuzzi
male, 26mm
Copyright © Ruslan Panin

Carabus (Platycarabus) creutzeri creutzeri Fabricius, 1801
Male adult
Italy, Fruili - UD - Carnia, N. Paulano 29.III.2002 leg. I.Rapuzzi
L=22.0 mm
Copyright © Kirill Makarov

Carabus (Platycarabus) creutzeri creutzeri Fabricius, 1801(as posarinii Rapuzzi, 1999 PT) Paratype
Male+Female adult
Slovenia:Koper:Kras Mt.R.:Brkini Mts.:Suhorije env., VIII.1998, I.Rapuzzi leg.
photo & koll. Panin R.
female, 32 mm
Copyright © Ruslan Panin

Carabus (Diocarabus) opaculus opaculus Putzeys, 1875
Male adult
Rusia, Kunashir isl., 8.1991 Shestopalov M. leg. male, 15 mm
Copyright © Alexander Anichtchenko

Carabus (Diocarabus) opaculus opaculus Putzeys, 1875
Male adult
Japan, Hokkaido, Hokuryu-cho 260 m., Museushi (West), 43:48’37’N;141:40’13’E, 6-26.VII.2010, I.Rapuzzi & L.Caldon leg.
Copyright © Ruslan Panin

Carabus (Platycarabus) creutzeri kircheri Germar, 1824
Male+Female adult
NE-Italy, Veneto Reg., Belluno, Forcella Lavardet h=2000m, VII.2002, I.Rapuzzi leg.
female, 24 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) casaleianus casaleianus Deuve, 1994
Female adult
China, SW Hunan Prov., Chengbu Co, Nan-Shan-Ding (26.34;110,08)1-10.VIII,2006, I.Rapuzzi leg.
L=33 mm
Copyright © Ruslan Panin

Carabus (Diocarabus) lucepunctus ignipunctatus Rapuzzi, 2009 PT Paratype
Male+Female adult
China, SW Hunan Prov., Chengbu Co, Nan-Shan-Ding (26.34;110,08)1-10.VIII,2006, I.Rapuzzi leg.
femele, 18 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) morbillosus alternans Palliardi, 1825
Male adult
S Italy, Sicily, Province Palermo, Piana degli Albanesi h~700m, XI.2010, I.Rapuzzi leg.
L=32 mm
Copyright © Ruslan Panin

Carabus (Aulonocarabus) canaliculatus pseudocareniger Deuve, 1991
Male adult
N Korea, Ryanggang Prov., Mt. Tongjom Ridge h~1861m, (128.60E;41.20N) I. Rapuzzi leg.
L=27 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) manifestus guanmenshanus Imura, 1991
Female adult
NE China, Liaoning, Anshan City, Mt.Qian Shan, 4-16.VII.2005, I.Rapuzzi leg.
L=26 mm
Copyright © Ruslan Panin

Carabus (Ohomopterus) albrechti albrechti Morawitz, 1862
Female adult
Japan, S-Hokkaido:Oshima Peninsula,Pass btw Otoshibe - Assabu, h~250-300 m, 10-20.VII.2010, 42:01’53’Т; 140:22’09’E, I.Rapuzzi & L.Caldon leg.
Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus aeruginosiformis Breuning, 1932
Male adult
Russia, Khakassia,
W-Sayan, Alan Mt range, valley Enisey riv., 5 km S Mayna vill., 15.VII.2000, A.& N. Menstschicovy leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Pagocarabus) promachus promachus s.str.
Male adult

Copyright © Boleslav Brezina

Carabus (Pagocarabus) promachus danbaensis Kleinfeld, 1999
Male adult

Copyright © Boleslav Brezina

Carabus (Pagocarabus) promachus danbaensis Kleinfeld, 1999
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Pagocarabus) promachus promachus s.str. (as konkerianus)
Female adult

Copyright © Boleslav Brezina

Carabus (Pagocarabus) poschingerianus Korell & Kleinfeld, 1989 Holotype
Male adult

Copyright © Frank Kleinfeld

Carabus (Pagocarabus) erwini plesiolabrus Deuve, 1991
Male adult

Copyright © Boleslav Brezina

Carabus (Pagocarabus) erwini erwini s.str.
Male adult

Copyright © Boleslav Brezina

Carabus (Meganebrius) wallichi wittmeri Mandl, 1974
Female adult
Nepal, Lantang, near Dunche, Songta vill.
3500 m 15.5.2000
Copyright © Alexander Anichtchenko

Carabus (Meganebrius) arunensis Heinertz, 1980
Female adult

Copyright © Boleslav Brezina

Carabus (Meganebrius) franzi Mandl, 1974
Female adult
Nepal, Lantang, near Dunche, Songta vill,
3500 m, 15.5.2000
Copyright © Alexander Anichtchenko

Carabus (Meganebrius) tamang gurkhaorum Lassalle, 1985
Female adult
CW Nepal, Manaslu massif 2300 m,
Bara Pokhari Lekh, 5.2001 M.Hackel leg.
Copyright © Alexander Anichtchenko

Carabus (Macrothorax) aumonti Lucas, 1849
Male adult

Copyright © Alexander Anichtchenko

Carabus (Macrothorax) planatus Chaudoir, 1843
Male adult
Sicilia, Madonie, Piano Battaglia (PA) 1500m
4.6.1992 Aliquo leg.
male, 31 mm
Copyright © Alexander Anichtchenko

Carabus (Macrothorax) morbillosus macilentus Lapouge, 1899
Male adult
Spain, Alicante, Santa Pola, 20.2.1988 M. Godoy leg.
male, 29 mm
Copyright © Alexander Anichtchenko

Carabus (Macrothorax) morbillosus constantinus Kraatz, 1899
Female adult

Copyright © Safronov

Carabus (Macrothorax) rugosus boeticus Deyrolle, 1852
Female adult
Spain, Cadiz, Novo Sancti Petri, 10.6.2003
female, 32 mm
Copyright © Alexander Anichtchenko

Carabus (Macrothorax) rugosus rugosus Fabricius, 1792
Male adult
N Africa, 2 km W Ceuta, Allabacal,
17-18.2.2004 Anichtchenko A. leg.
male, 26 mm
Copyright © Alexander Anichtchenko

Carabus (Macrothorax) rugosus laufferi Breuning, 1927
Female adult
Spain, Gredos Mts., 2 km E from La Mira Mt.
4-7.7.2007 Anichtchenko A. leg.
female, 29 mm
Copyright © Alexander Anichtchenko

Carabus (Macrothorax) rugosus celtibericus Germar, 1824
Male adult
Portugal, Vila Fresca de Azeitas,
Santana, 13.12.2001
male, 30 mm
Copyright © Alexander Anichtchenko

Carabus (Lipaster) stjernvalli stjernvalli Mannerheim, 1830
Male+Female adult
Armenia,
Ashotsk, 22.VI.1990, P.V.Kazaryan leg.
female 34mm
Copyright © Ruslan Panin

Carabus (Lipaster) stjernvalli (as tabackuriensis Novotny & Vorisek, 1988)
Male adult

Copyright © Alexander Anichtchenko

Carabus (Limnocarabus) clathratus clathratus Linne, 1761
Female adult
W-Ukraine, Maloe Polesie, Lviv Region, near Kamenka-Bugzkaya, 28.III.2008, Panin R. leg. L=31 mm
Copyright © Ruslan Panin

Carabus (Limnocarabus) clathratus clathratus Linne, 1761
Male adult
length: 20-36 mm, Germania, Mecklenburg, Wismar, leg. F.Wolf, 1998; det. A.Skale, Photo: U.Schmidt, 2009
Copyright © U.Schmidt

Carabus (Limnocarabus) clathratus clathratus (as clathratus foveolatoseriatus)
Female adult
The Chelyabinsk area, Kizilskyi township, Cheka mountain (steppe). 14-15/07/2008
Copyright © Sunvas

Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
Male adult
Transcaucasia, Armenia,
Amasia distr., Arpalach vill., VI-VII.1999, P.V.Kazaryan leg.
male, 25mm
Copyright © Ruslan Panin

Carabus (Carabus) limbatus Say, 1825 (as goryi)
Female adult

Copyright © Henri Goulet

Carabus (Carabus) limbatus Say, 1825
Male adult

Copyright © Denis Potanin

Carabus (Leptoplesius) merzbacheri kirgisiensis Mandl, 1967
Female adult
22mm
Copyright © Boleslav Brezina

Carabus (Leptoplesius) merzbacheri kirgisiensis Mandl, 1967
Female adult
female, 27mm
Copyright © Boleslav Brezina

Carabus (Leptoplesius) merzbacheri kirgisiensis Mandl, 1967
Female adult
female, 24mm
Copyright © Boleslav Brezina

Carabus (Leptoplesius) shokalskii kuqaensis Deuve, 1994 Paratype
Female adult
female, 24mm
Copyright © Boleslav Brezina

Carabus (Leptocarabus) yokoae marcilhaci Deuve, 1992
Male adult
CHINA, Shanxi, Tschingling Mts., Taibeichan Nat. reserv.;
Copyright © Anton Menstschikov

Carabus (Leptocarabus) yokoae marcilhaci Deuve, 1992
Male adult
China, Shanxi, Taschingling Mts., Taibeishan Nat. res.
2000 m., 5-15.08.2005
Copyright © Alexander Anichtchenko

Carabus (Leptocarabus) semiopacus Reitter, 1895
Female adult
N Korea, Nangnim, 14.VII.1981
female, long. 24 mm
Copyright © Alexander Anichtchenko

Carabus (Leptocarabus) procerulus Chaudoir, 1862
Male adult
Japan, Yamanashi pref., Otawa-rindo,
Narusawa vill., 9.IX.2002 T. Kurihara leg
male, 31 mm
Copyright © Alexander Anichtchenko

Carabus (Leptocarabus) harmandi fujisan Ishikawa, 1972
Female adult
Japan, Yamanashi pref., Fuji-Rinds 1600m, Narusawa vill. 3.V.2005 Kurihara K. leg. 25 mm
Copyright © Alexander Anichtchenko

Carabus (Leptocarabus) arboreus exarboreus (Ishikawa, 1992)
Male adult
E.Russia, Far East, Sakhalin Isl., forest of M.Tym river 11-IX-2000 . Egorov leg. male 27mm, female 30mm.
Copyright © Safronov

Carabus (Lasiocoptolabrus) sunwukong sunwukong Imura, 1993 Paratype
Male adult

Copyright © Boleslav Brezina

Carabus (Lasiocoptolabrus) sunwukong baotianmanus (Imura, 2005) Paratype
Male adult

Copyright © Boleslav Brezina

Carabus (Lamprostus) calleyi pseudoprasinus Lapouge, 1912
Male+Female adult
Transcaucasia, Armenia,
Ashotsk distr. Tsoghamarg vill, .21-22.V.1997, P.V.Kazaryan leg
female, 24 mm
Copyright © Ruslan Panin

Carabus (Lamprostus) calleyi nigrinus Motschulsky, 1865
Male adult
Azerbaijan,
Talysh Mts., Palikesh, Zuvand, 08.VI.2001, Dolin V.G. leg.
male, 24mm
Copyright © Ruslan Panin

Carabus (Lamprostus) torosus spinolae Cristoforis & Jan, 1837
Male adult
length: 29 mm
Türkei, Cankiri, Cerkes, Kurtcimeni
leg.det. K.Staven, 1989
Foto: U.Schmidt, 2009
Copyright © U.Schmidt

Carabus (Ischnocarabus) tenuitarsis tenuitarsis Kraatz, 1877
Male adult
male, 24mm
Copyright © Boleslav Brezina

Carabus (Ischnocarabus) tenuitarsis tenuitarsis Kraatz, 1877
Female adult
female, 26mm
Copyright © Boleslav Brezina

Carabus (Ischnocarabus) cychropalpus Peyron, 1858
Male adult
male, 27mm
Copyright © Boleslav Brezina

Carabus (Ischnocarabus) cychropalpus Peyron, 1858
Female adult
female, 28mm
Copyright © Boleslav Brezina

Carabus (Iniopachus) pyrenaeus jeanneli Deuve, 1994
Male adult
1-2: Pirineos, Canfranc, Ibon de Iserias 2200 m 16-17.7.05 3: Pirineos, Monte Perdido, Collata Anisolo 2400m 25.8.05
Copyright © Alexander Anichtchenko

Carabus (Iniopachus) auriculatus ubinensis (Puissegur, 1955)
Female adult
Spain, Montes Cantabricos, Peña Ubiña, 2400m
5-6.VII.2009 Anichtchenko A. leg.
female, 19 mm
Copyright © Alexander Anichtchenko

Carabus (Iniopachus) auriculatus auriculatus Putzeys, 1872
Female adult
Spain, Picos de Europa, Macizo Central, Las Colladinas,
ref. Hermoso 2100m, 8.8.2003
Copyright © Alexander Anichtchenko

Carabus (Imaibius) wittmerorum Heinertz, 1978
Male adult

Copyright © Alexander Anichtchenko

Carabus (Imaibius) stoliczkanus Bates, 1878
Male adult

Copyright © Alexander Anichtchenko

Carabus (Imaibius) pachtoun marysae Letellier, 1979
Male adult

Copyright © Alexander Anichtchenko

Carabus (Imaibius) isabellae Lassalle, 1985
Male adult
male, 27mm
Copyright © Boleslav Brezina

Carabus (Imaibius) isabellae Lassalle, 1985
Female adult
female, 30mm
Copyright © Boleslav Brezina

Carabus (Imaibius) gandharae Heinertz, 1978
Male adult

Copyright © Alexander Anichtchenko

Carabus (Imaibius) dardiellus polymorphus Heinz, 1993 Paratype
Female adult
Museum of Institute of Systematic Biology (Daugavpils, Latvia)

Copyright © Alexander Anichtchenko

Carabus (Imaibius) caschmirensis caschmirensis Redtenbacher, 1844
Female adult
length: 32 mm Pakistan, Mansehra leg. Heinz & Staven, 2001; det. K.Staven, 2001 Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Carabus (Imaibius) boysi Tatum, 1851
Male adult

Copyright © Alexander Anichtchenko

Carabus (Imaibius) barysomus hazarensis Deuve, 1982
Male adult
length: 35 mm Pakistan, Mansehra leg.det. K.Staven, 2001 Topotypus Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Carabus (Imaibius) baronii sharanicus Deuve & Letellier, 1980 Paratype
Male adult

Copyright © Alexander Anichtchenko

Carabus (Imaibius) baronii sharanicus Deuve & Letellier, 1980
Male adult

Copyright © Alexander Anichtchenko

Carabus (Imaibius) baronii baronii Heinertz, 1977
Male adult
length: 23 mm
Pakistan, Mansehra
leg.det. K.Staven, 1997
Topotypus
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Carabus (Imaibius) baronii battalensis Heinz & Staven, 1998 Paratype
Male adult

Copyright © Alexander Anichtchenko

Carabus (Pseudocoptolabrus) businskyi Deuve, 1990
Male adult
CH, NW-Yunnan (Lijiang) Yulongxue Shan, E-slope
27:10:50N/100:12:26E/ 4400m, alpine meadows 12.VII.2000
K. & B. Brezina, M.Brychcin, M. Hackel
Copyright © Boleslav Brezina

Carabus (Oreocarabus) taibaiensis Kleinfeld, 2001 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Oreocarabus) latro minshanensis Deuve, 1987
Male adult
China. N Sishuan, Juizhalgou
12-16.6.1995 Benes leg.
male, 22 mm
Copyright © Alexander Anichtchenko

Carabus (Oreocarabus) laotse paralaotse Kleinfeld, 2000 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Oreocarabus) mikhaili Deuve & Mourzine, 1997
Male adult
China, S Gansu, Min Shan, 2100m
SW Wudu, 1.8.2000 V.Patrikeev leg.
male, 20 mm
Copyright © Alexander Anichtchenko

Carabus (Trachycarabus) sibiricus gvozdevae Obydov, 1997
Male adult
Russia, Khakassia,
W Sayan, Enisey riv., Batenevskiy Mt Range, 12 km E Pervomayskiy vill., 11.VII.2000, A.& N. Menstschicovy leg.
L=21 mm
Copyright © Ruslan Panin

Carabus (Homoeocarabus) maeander paludis Gehin, 1885
Female adult
female, 21mm
Copyright © Boleslav Brezina

Carabus (Homoeocarabus) maeander maeander s.str.
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Carabus (Homoeocarabus) maeander maeander s.str.
Male adult
male, 18mm
Copyright © Boleslav Brezina

Carabus (Homoeocarabus) maeander maeander s.str.
Male adult
male, 17mm
Copyright © Boleslav Brezina

Carabus (Cathoplius) aliai Escalera, 1944
Male adult
male, 27mm
Copyright © Boleslav Brezina

Carabus (Cathoplius) asperatus asperatus (Dejean, 1826)
Male adult
male, 24mm
Copyright © Boleslav Brezina

Carabus (Cathoplius) asperatus asperatus (Dejean, 1826)
Female adult
female, 26mm
Copyright © Boleslav Brezina

Carabus (Cathoplius) asperatus ifniensis (Zarco, 1941)
Male adult
male, 30mm
Copyright © Boleslav Brezina

Carabus (Cathoplius) asperatus ifniensis (Zarco, 1941)
Male adult
male, 34mm
Copyright © Boleslav Brezina

Carabus (Cathoplius) stenocephalus Lucas, 1866 (as escalerae)
Male adult
male, 29mm
Copyright © Boleslav Brezina

Carabus (Cathoplius) stenocephalus Lucas, 1866 (as escalerae)
Female adult
female, 33mm
Copyright © Boleslav Brezina

Carabus (Cathoplius) stenocephalus Lucas, 1866
Female adult
female, 28mm
Copyright © Boleslav Brezina

Carabus (Cathoplius) stenocephalus Lucas, 1866
Male adult
male, 25 mm
Copyright © Boleslav Brezina

Carabus (Cathoplius) asperatus susicus Antoine, 1941
Male adult
male, 27mm
Copyright © Boleslav Brezina

Carabus (Cathoplius) asperatus susicus Antoine, 1941
Female adult
female, 29mm
Copyright © Boleslav Brezina

Carabus (Aulonocarabus) mouthiezianus Deuve, 1991
Male adult
Yakutia, Verhoyanskii Mt.rng., âåðõîâüÿ Êåëå, ð. Íÿìíè, ëåâ. èñòîê. ð. Êîê÷èí 29.7.1994
Copyright © Alexander Anichtchenko

Carabus (Orinocarabus) bremii Stierlin, 1881
Male adult
NW Italy, Piemonte, Mucrone Mt., 14-21.VI.1999
Copyright © Alexander Anichtchenko

Carabus (Cechenotribax) petri Semenov & Znojko, 1932
Female adult
female, 18mm
Copyright © Boleslav Brezina

Carabus (Cechenotribax) petri Semenov & Znojko, 1932
Male adult
male, 14mm
Copyright © Boleslav Brezina

Carabus (Cephalornis) potanini (Semenov, 1887)
Male adult
China, S Gansu, 3400m, Minshan mts.,
Wenxian 8.92, J. M. Bousquet
Copyright © Boleslav Brezina

Carabus (Chaetocarabus) arcadicus arcadicus Gistl, 1850
Female adult
female, 35mm
Copyright © Boleslav Brezina

Carabus (Chaetocarabus) arcadicus arcadicus Gistl, 1850
Male adult
male, 29mm
Copyright © Boleslav Brezina

Carabus (Chaetocarabus) arcadicus merlini Schaum, 1861
Male adult
male, 30mm
Copyright © Boleslav Brezina

Carabus (Chaetocarabus) arcadicus merlini Schaum, 1861
Female adult
female, 36mm
Copyright © Boleslav Brezina

Carabus (Chaetocarabus) intricatus bayardi Solier, 1835
Male+Female adult
S-Italy,
Basilicata Reg., Potenza Prov., Latronico, F. Angelini leg. 14.VI.2002
female, 26mm
Copyright © Ruslan Panin

Carabus (Chaetocarabus) intricatus bayardi Solier, 1835
Male adult
male, 25mm
Copyright © Boleslav Brezina

Carabus (Chaetocarabus) intricatus bayardi Solier, 1835
Female adult
female, 27mm
Copyright © Boleslav Brezina

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761 (as bohemicus)
Female adult
female, 33mm
Copyright © Boleslav Brezina

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Male adult
male, 26mm
Copyright © Boleslav Brezina

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Male adult
W-Ukraine,
Lviv Reg., Zhydachiv distr., near Vybranivka Vill. Panin R.leg. 12.X.2007
31mm.
Copyright © Ruslan Panin

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Female adult
36 mm
Fundort: Germania, Oberfranken, Kulmbach
leg.det. U.Schmidt, 1977
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Carabus (Chaetocarabus) intricatus krueperi Reitter, 1896
Male adult
male, 35mm
Copyright © Boleslav Brezina

Carabus (Chaetocarabus) intricatus krueperi Reitter, 1896
Female adult
female, 38mm
Copyright © Boleslav Brezina

Carabus (Chaetocarabus) intricatus lefebvrei Dejean, 1826
Male adult
male, 28mm
Copyright © Boleslav Brezina

Carabus (Chaetocarabus) intricatus lefebvrei Dejean, 1826
Female adult
female, 30mm
Copyright © Boleslav Brezina
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