Carabidae image gallery
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Bembidarenini taxon page (14 photos)
Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Image from Sallenave, Erwin, and Roig-Juñent, 2008
Copyright © - - -
Andinodontis muellermotzfeldi Toledano & Erwin, 2010
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Tasmanitachoides erwini Maddison & Porch, 2021 Holotype
Male adult
Australia: Tasmania: River Forth at C136, 41.4712°S 146.1366°E, 126 m, 14.i.2019. DRM 19.012. D.R. Maddison & N.A. Porch. Scale bar 1mm
Copyright © David Maddison
Tasmanitachoides leai Sloane, 1896
Female adult
Australia: Victoria, Flat Rock Creek at highway B23. Scale bar 1mm
Copyright © David Maddison
Tasmanitachoides hobarti (Blackburn, 1901)
Male adult
TASMANIA: Ringarooma River at Derby, 148 m
Copyright © David Maddison
Tasmanitachoides wilsoni (Sloane, 1921)
Male adult
Australia: ACT: Murrumbidgee R. at Uriarra Crossing, 450 m, 35.2462°S, 148.9530°E , scale 1mm
Copyright © David Maddison
Tasmanitachoides baehri Maddison & Porch, 2021 Paratype
Male adult
Australia: ACT: Murrumbidgee River, 0.15 km u/s Uriarra Crossing (35°14.717'S, 148°57.135'E 440 m)
Copyright © David Maddison
Andinodontis maveetyae Erwin & Maddison, 2010 Holotype
Male adult
Perú: Cusco, Rocotal, 3.2 km E Tres Cruces, 13.1123° S, 71.5834° W, 2082 m, 27 June 2010 (S. MAVEETY) (MHN-UNSAAC: ADP124815)
Copyright © Terry Erwin
Andinodontis guzzettii Toledano & Erwin, 2010 Holotype
Female adult
Boliva: Cochabamba, Arque, 17.8232° S, 66.4022° W, 2714 m, 26 October 1995 (L. GUZZETTI) (MSNV: ADP116824)
Copyright © Terry Erwin
Andinodontis moreti Erwin & Toledano, 2010 Holotype
Female adult
Ecuador: Sucumbíos, La Bonita, 0.4600° N, 77.50533° W, 2000 m, 1 January 1999 (G. ONORE)
Copyright © Terry Erwin
Andinodontis muellermotzfeldi Toledano & Erwin, 2010 Paratype
Female adult
Sucumbios, La Bonita, 0.4600° N, 77.50533° W, 2000 m, 1 January 1994 (G. ONORE)
Copyright © Terry Erwin -
Bembidiini taxon page (942 photos)
Typhlocharis carmenae Zaballos & Ruiz-Tapiador, 1995
Male adult
Spain, Toledo, Los Navalucillos 23.I.1997
Zaballos & Ruiz-Tapiador leg.
long 1.3mm
Copyright © Alexander Anichtchenko
Asaphidion flavipes (Linne, 1761): 211 (Cicindela)
Male adult
lenght: 4,0-4,8 mm
Austria, Karawanken, Ferlach
leg.det. U.Schmidt, 1982
Copyright © U.Schmidt
Asaphidion stierlini (Heyden, 1880d): 304 (Tachypus)
Male adult
Spain, Sierra Nevada, El Castillejo 2000m
7.VII.2005 Anichtchenko A.
long. 4mm
Copyright © Alexander Anichtchenko
Asaphidion ussuriense Jedlicka, 1965e: 83
Male adult
Far East, Lazovskij Nat. Res., Proselochnaja Riv. near nouth 43°00'34"N 134°07'43"E 10.VII.2005 K.Makarov leg.
L=3.5 mm
Copyright © Kirill Makarov
Sakagutia umi Sasakawa, 2007
Male adult
Far East, Lazo Distr., 4 km VE Preobrazhenie Vill., Ezhovaja Bay 42°55'59N 26.VII.2005 133°59'29E K.Makarov leg.<br /> L=5.5 mm
Copyright © Kirill Makarov
Elaphropus (Tachyura) parvula (Dejean, 1831)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Elaphropus (Tachyura) parvula (Dejean, 1831)
Male adult
length: 1,7-2,4 mm
Germania, Oberbayern, Eching
leg.det. A.Skale, 2001 (as Elaphropus)
Foto: U.Schmidt, 2008
Copyright © U.Schmidt
Tachyta (Tachyta) nana (Gyllenhal, 1810)
Male adult
Far East, Lazovskij Nat. Res., Proselochnaja Riv. near nouth, N slope of Tumannaja Mt., h~350-500 43°00'08"N 134°06'56"E 9.VII.2005 K.Makarov leg.
L=2.7 mm
Copyright © Kirill Makarov
Tachys (Tachys) halophilus Lindroth, 1966
Female adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Tachys (Polyderis) laevis (Say, 1823)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Tachys (Paratachys) proximus Say, 1823
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Tachys (Paratachys) fulvicollis (Dejean, 1831)
Male adult
Volgograd Reg., Pallas Distr., Elton Vill., on UV-light 21.VII.2006 K.Makarov leg.
L=2.3 mm
Copyright © Kirill Makarov
Porotachys bisulcatus (Nicolai, 1822)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Pericompsus (Pericompsus) ephippiatus (Say, 1834)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Pericompsus (Pericompsus) elegantulus (Laferte, 1848)
Male adult
Puerto Rico<br /> photo I. M. Sokolov
Copyright © Igor Sokolov
Ocys harpaloides (Audinet-Serville, 1821)
Male adult
length: 4,2-5,2 mm
Korsika, Ghisonaccia
leg.det. U.Schmidt, 1973
Photo: U.Schmidt, 2008
Copyright © U.Schmidt
Mioptachys flavicauda (Say, 1823)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Erwiniana punctisculptilis (Erwin, 1994)
Male adult
Ecuador, Yasuni
Photo of Sokolov I. M.
Copyright © Igor Sokolov
Elaphropus (Barytachys) tripunctatus (Say, 1830)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Elaphropus (Tachyura) incurvus (Say, 1830)
Female adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Cillenus (Desarmatocillenus) sumaoi (Morita, 1981)
Male adult
Chiba, Honshu, Japan<br /> (c) Field Museum of Natural History <br /> (Maruyama Munetoshi)
Copyright © Munetoshi Maruyama
Bembidion (Limnaeoperyphus) quadriimpressum (Motschulsky, 1860)
Male adult
Kunashiri Isl., mouth of Medny Spring 44°16'31"N 145°56'29"E 27.VI.2008 leg. K.Makarov
L=4.3
Copyright © Kirill Makarov
Bembidion (Trepanes) articulatum (Panzer, 1796)
Female adult
length: 3,0-3,9 mm
Germania, Bayern, Oberfranken, Kulmbach
leg.det. U.Schmidt, 1992
Foto: U.Schmidt, 2010
Copyright © U.Schmidt
Bembidion (Trepanedoris) concretum Casey, 1918
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Bembidion (Plataphus) simplex Hayward, 1897
Male adult
Great Smoky Mountains National Park. photo I. M. Sokolov
Copyright © Igor Sokolov
Bembidion (Testedium) bipunctatum (Linne, 1761)
Male adult
length: 3,6-4,8 mm
Austria, Schladminger Tauern, Mariapfarr
leg.det. U.Schmidt, 1984
Foto: U.Schmidt, 2010
Copyright © U.Schmidt
Bembidion (Plataphus) gebleri (Gebler, 1833)
Male adult
Far East, Lazovskij Nat. Res., Formozova spring 43°00'06"N 134°05'13"E 10.VII.2005 K.Makarov leg.
L=4.5 mm
Copyright © Kirill Makarov
Bembidion (Plataphus) altaicum Gebler, 1833
Male adult
Primorie reg., Lazo distr., near Lazo vill. Lazovka riv. 20.06.04 leg. M.& L. Smirnov photo from www.zin.ru
Copyright © Maksim Smirnov
Bembidion (Phyla) obtusum Audinet-Serville, 1821: 83
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Bembidion (Philochthus) pallidiveste Carret, 1905
Male adult
Volgogrda Area, Pallasovka Distr., near Elton Vill. 4.V.2009 leg. K.Makarov
L = 3.8
Copyright © Kirill Makarov
Bembidion (Philochthus) guttula (Fabricius, 1792)
Male adult
length: 2,8-3,8 mm
Germania, Bayern, Oberfranken, Rudolfstein
leg.det. U.Schmidt, 2001
Foto: U.Schmidt, 2010
Copyright © U.Schmidt
Bembidion (Peryphus) tetracolum Say, 1823: 89
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Bembidion (Peryphus) petrosum Gebler, 1833: 275
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Bembidion (Peryphanes) lacunarium (Zimmermann, 1869)
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Bembidion (Odontium) confusum Hayward, 1897
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Bembidion (Odontium) chloropus Bates, 1883: 277
Male adult
Far East, Lazovskij Nat. Res., Proselochnaja Riv. near nouth, beach 43°00'34"N 134°07'43"E 17-21.VIII.2007 K.Makarov & A.Zaitsev leg. L=5.6 mm
Copyright © Kirill Makarov
Bembidion (Ochthedromus) bifossulatum (Leconte, 1852)
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Bembidion (Notaphus) patruele Dejean, 1831
Female adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Bembidion (Nepha) menetriesii (Kolenati, 1845)
Female adult
Russia, Orenburg reg., Sol'-Iletskii distr., 10 km W Troitsk vill. 6.05.09
leg. M. & L. Smirnov's
Female
photo M. Smirnov from www.zin.ru
Copyright © Maksim Smirnov
Bembidion (Nepha) genei illigeri Netolitzky, 1914
Male adult
length: 4,0-4,9 mm
Germania, Bayern, Oberfranken, Selbitz
leg.det. U.Schmidt, 2001
Foto: U.Schmidtz, 2010
Copyright © U.Schmidt
Bembidion (Nepha) genei illigeri Netolitzky, 1914: 54
Female adult
length: 4,6 mm
South Urals, Chelyabinsk area, Asha distr., the vicinities of Sim town, Sim river valley
14.05.2001.
Rudoiskatel' P.leg.
From the collection of A.Ivanov (Ekaterinburg)
Copyright © Evgeniy Zinoviev
Bembidion (Neoemphanes) shimoyamai Habu, 1978
Male adult
Far East, khanka Lake env., Bol.Usachi Riv. near Novokachalinsk 5.VII.2004 R.Filimonov leg.
L=3.5 mm
Copyright © Kirill Makarov
Bembidion (Metallina) properans (Stephens, 1828b): 26 (Tachypus)
Female adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Bembidion (Metallina) lampros (Herbst, 1784)
Male adult
length: 2,7-4,1 mm
Germania, Bayern, Oberfranken, Neuenmarkt
leg.det. U.Schmidt, 1972
Photo: U.Schmidt, 2009
Copyright © U.Schmidt
Bembidion (INCERTAE) bandotaro Morita, 1991
Female adult
Kuril Islands
photo Makarov K.
Copyright © Kirill Makarov
Bembidion (INCERTAE) amurense Motschulsky, 1859
Male adult
photo Makarov K.
Copyright © Kirill Makarov
Bembidion (Hirmoplataphus) nigrum Say, 1823
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Bembidion (Furcacampa) versicolor (Leconte, 1848)
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Bembidion (Furcacampa) impotens Casey, 1918
Female adult
Great Smoky Mountains National Park
photo I. M. Sokolov
Copyright © Igor Sokolov
Bembidion (Furcacampa) affine Say, 1823
Male adult
Great Smoky Mountains National Park
photo I. M. Sokolov
Copyright © Igor Sokolov
Bembidion (Eurytrachelus) pogonoides Bates, 1883
Female adult
Far East, Lazovsky Nat.Res., shore of Japan Sea, cordon Petrova, Petrovski spring 10.06.1995 leg. Yu.Sundukov
female
L=6,2 mm
Copyright © Maksim Smirnov
Bembidion (Eurytrachelus) nitidum (Kirby, 1837)
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Bembidion (Eurytrachelus) laticolle (Dufischmid, 1812)
Female adult
Orenburg Reg., Ilek Distr., Ilek env., valley of Ural Riv., beach 28.VII-8VIII.2005 S.Vastschenko leg.
L=6.2 mm
Copyright © Kirill Makarov
Bembidion (Asioperyphus) infuscatum Dejean, 1831
Female adult
length: 6,4 mm
West Siberia, KHMAO-Yugra, Khanty-Mansiysk distr., Ob river, Bolshoy Tchukhtinsiy island
4.08.2007.
Zinoviev E.leg.
Copyright © Evgeniy Zinoviev
Bembidion (Actedium) pallidipenne (Illiger, 1802b)
Male adult
length: 4,1-4,7 mm, Niederlande, Maasvlakte, leg. A.Teunisse, 1989; det. G.Rößler, 1996, Foto: U.Schmidt, 2010
Copyright © U.Schmidt
Amerizus (Amerizus) wingatei (Bland, 1863) (Bembidium)
Female adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Bembidion (Asioperyphus) postremum Say, 1830
Female adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Bembidion (Bembidion) quadrimaculatum (Linne, 1761)
Male adult
length: 2,5-3,6 mm
Germania, Oberfranken, Kulmbach
leg.det. U.Schmidt, 1972
Foto: U.Schmidt, 2008
Copyright © U.Schmidt
Bembidion (Bembidion) quadrimaculatum oppositum Say, 1823
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Bembidion (Bembidionetolitzkya) conforme Dejean, 1831
Male adult
length: 4,5-6 mm
Germania, Oberbayern, Oberstdorf
leg.det. U.Schmidt, 2005
Foto: U.Schmidt, 2006
Copyright © U.Schmidt
Bembidion (Bembidionetolitzkya) geniculatum Heer, 1837
Female adult
Ukrainian Carpathians,
Ivano-Frankivsk reg.,Yaremcha distr.,Gorgany Mts. ,h=750 m. 22.VI.08 Panin R. leg.
7mm.
Copyright © Ruslan Panin
Bembidion (Bracteon) conicolle Motschulsky, 1844: 273
Female adult
SE Siberia, Irkutsk Area, Khamar-Daban Mountains,6-8.VI.2007, Fominyh D. leg. long=7mm
Copyright © Ruslan Panin
Bembidion (Bracteon) foveum Motschulsky, 1844: 271
Male adult
Khabarovsk Reg., bank of Amur Rin. in Khabarovsk city 48°28'33"N 135°02'39"E 7.VIII.2003 I.Melnik leg.
L=6.0 mm
Copyright © Kirill Makarov
Bembidion (Bracteon) litorale (Olivier, 1790)
Female adult
length: 5,2-6,2 mm
Germania, Bayern, Forchheim, Hallerndorf
leg.det. A.Skale, 2001
Foto: U.Schmidt, 2008
Copyright © U.Schmidt
Bembidion (Bracteon) punctatostriatum Say, 1823
Female adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Bembidion (Diplocampa) transparens (Gebler, 1829)
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Bembidion (Eupetedromus) graciliforme Hayward, 1897
Female adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Bembidion (Eupetedromus) sibiricum Dejean, 1831a: 66
Female adult
S Primorie, Ussurijski Reserve, Mirinov Klyuch 43°38.644' N 132°26.693' E 25.VIII.2007 leg. K.Makarov, A.Zaitzev
L =4.7
Copyright © Kirill Makarov
Bembidion (Archaeophilochthus) christelae Ortuno & Arillo, 2010 Holotype
Male adult
Baltic Eocene amber
Copyright © Alexander Anichtchenko
Elaphropus (Elaphropus) latissimus (Motschulsky, 1851)
Female adult
Kunashiri Isl., Calder of Golovnina Volcano, S coast od Goryachee Lake 43°52'19"N 145°29'43"E 12.VIII.2008 leg.K.Makarov, A.Zaitzev
2.0 mm
Copyright © Kirill Makarov
Bembidion (Talanes) subfasciatum Chaudoir, 1850b: 187
Female adult
Krasnodar Reg., Taman Peninsula, Tuzlo Lake 6.V.2009
2.7 mm
Copyright © Kirill Makarov
Asaphidion pallipes (Duftschmid, 1812): 197 (Elaphrus)
Female adult
L=5,4 mm
RUSSIA, Tver reg., Likhoslavlskiy distr., Domanikha village, in the garden, under plank. 6.V.2000
leg., det. & photo - Vlasenko A. S.
Copyright © Andrey Vlasenko
Bembidion (Ocydromus) perspicuum (Leconte, 1848)
Male adult
USA: Colorado: Huerfano Co., Huerfano River at Badito, 37.7285°N 105.0167°W.
Scale bar is 1 mm.
Photo David R. Maddison & Alexander P. Swanson
Copyright © ---
Amerizus (Amerizus) teles Belousov & Dudko, 2010 Paratype
Male adult
NE Altay Mts., 35-40 km SSE Artybash, Evrechala Mt., 1850-2100 м, tundra, 51°28' N, 87°26' E, 20-22.05.2003, R.Yu.Dudko & I.I.Lubechanskii leg.
Copyright © Igor Belousov
Tachys (Paratachys) kaorutanakai Habu, 1977 Holotype
Male adult
Kamiube, Ube, YAMAGUCHI, 09. V. 1976, K. Tanaka
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-019
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Tachys (Paratachys) kaorutanakai Habu, 1977 Holotype
Male adult
Kamiube, Ube, YAMAGUCHI, 09. V. 1976, K. Tanaka
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-019
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Tachys (Paratachys) kaorutanakai Habu, 1977 Holotype
Male adult
Kamiube, Ube, YAMAGUCHI, 09. V. 1976, K. Tanaka
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-019
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Tachys (Paratachys) kaorutanakai Habu, 1977 (Label) Holotype
Male adult
Kamiube, Ube, YAMAGUCHI, 09. V. 1976, K. Tanaka
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-019
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Elaphropus (Elaphropus) nipponicus (Habu & Baba, 1967) Holotype
Male adult
Kurokawa, Niigata Pref., Japan, 24. VI. 1963, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-020
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Elaphropus (Elaphropus) nipponicus (Habu & Baba, 1967) Holotype
Male adult
Kurokawa, Niigata Pref., Japan, 24. VI. 1963, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-020
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Elaphropus (Elaphropus) nipponicus (Habu & Baba, 1967) Holotype
Male adult
Kurokawa, Niigata Pref., Japan, 24. VI. 1963, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-020
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Elaphropus (Elaphropus) nipponicus (Habu & Baba, 1967) (Label) Holotype
Male adult
Kurokawa, Niigata Pref., Japan, 24. VI. 1963, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-020
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Tachys (Paratachys) koizumii (Habu, 1961) Holotype
Male adult
Mt. Nachi, WAKAYAMA, 27. VIII. 1957, K. Koizumi
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-021
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Tachys (Paratachys) koizumii (Habu, 1961) Holotype
Male adult
Mt. Nachi, WAKAYAMA, 27. VIII. 1957, K. Koizumi
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-021
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Tachys (Paratachys) koizumii (Habu, 1961) Holotype
Male adult
Mt. Nachi, WAKAYAMA, 27. VIII. 1957, K. Koizumi
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-021
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Tachys (Paratachys) koizumii (Habu, 1961) Holotype
Male adult
Mt. Nachi, WAKAYAMA, 27. VIII. 1957, K. Koizumi
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-021
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Tachys (Paratachys) fasciatus uenoi Tanaka, 1960 Holotype
Male adult
Chitose-funabashi, Tokyo, Japan, 3. VII. 1957, K. Tanaka
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-240
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Tachys (Paratachys) fasciatus uenoi Tanaka, 1960 Holotype
Male adult
Chitose-funabashi, Tokyo, Japan, 3. VII. 1957, K. Tanaka
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-240
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Tachys (Paratachys) fasciatus uenoi Tanaka, 1960 (Label) Holotype
Male adult
Chitose-funabashi, Tokyo, Japan, 3. VII. 1957, K. Tanaka
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-240
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Tachys (Paratachys) gyotokuensis Tanaka, 1956 Holotype
Male adult
Gyôtoku, Chiba Pref., Japan, 3. VI. 1952, K. Tanaka
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-239
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Tachys (Paratachys) gyotokuensis Tanaka, 1956 Holotype
Male adult
Gyôtoku, Chiba Pref., Japan, 3. VI. 1952, K. Tanaka
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-239
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Tachys (Paratachys) gyotokuensis Tanaka, 1956 (Label) Holotype
Male adult
Gyôtoku, Chiba Pref., Japan, 3. VI. 1952, K. Tanaka
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-239
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Tachys (Paratachys) plagiatus shimosae Tanaka, 1956 Holotype
Male adult
Gyôtoku, at the mouth of the Edogawa River, Chiba Pref., Japan, 3. VI. 1952, K. Tanaka
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-238
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Tachys (Paratachys) plagiatus shimosae Tanaka, 1956 (Label) Holotype
Female adult
Gyôtoku, at the mouth of the Edogawa River, Chiba Pref., Japan, 3. VI. 1952, K. Tanaka
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-238
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Lymnastis sugimotoi Habu, 1975 Holotype
Male adult
Ishigaki Is., RYUKYUS, 29. VI. 1973, A. Sugimoto
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-022
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Lymnastis sugimotoi Habu, 1975 Holotype
Male adult
Ishigaki Is., RYUKYUS, 29. VI. 1973, A. Sugimoto
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-022
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Lymnastis sugimotoi Habu, 1975 Holotype
Male adult
Ishigaki Is., RYUKYUS, 29. VI. 1973, A. Sugimoto
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-022
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Lymnastis sugimotoi Habu, 1975 (Label) Holotype
Male adult
Ishigaki Is., RYUKYUS, 29. VI. 1973, A. Sugimoto
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-022
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Asaphidion tenryuense tenryuense Habu, 1954 Holotype
Male adult
Tenryu River-side, Iijima, Kamiina, NAGANO, 6. X.1945, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-023
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Asaphidion tenryuense tenryuense Habu, 1954 Holotype
Male adult
Tenryu River-side, Iijima, Kamiina, NAGANO, 6. X.1945, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-023
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Asaphidion tenryuense tenryuense Habu, 1954 Holotype
Male adult
Tenryu River-side, Iijima, Kamiina, NAGANO, 6. X.1945, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-023
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Asaphidion tenryuense tenryuense Habu, 1954 (Label) Holotype
Male adult
Tenryu River-side, Iijima, Kamiina, NAGANO, 6. X.1945, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-023
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Cillenus (Desarmatocillenus) kasaharai Habu, 1978 Holotype
Male adult
Oohara, Iriomote Is., OKINAWA, 11. VI. 1976, S. Kasahara
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-024
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Cillenus (Desarmatocillenus) kasaharai Habu, 1978 Holotype
Male adult
Oohara, Iriomote Is., OKINAWA, 11. VI. 1976, S. Kasahara
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-024
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Cillenus (Desarmatocillenus) kasaharai Habu, 1978 Holotype
Male adult
Oohara, Iriomote Is., OKINAWA, 11. VI. 1976, S. Kasahara
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-024
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Cillenus (Desarmatocillenus) kasaharai Habu, 1978 (Label) Holotype
Male adult
Oohara, Iriomote Is., OKINAWA, 11. VI. 1976, S. Kasahara
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-024
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Plataphodes) tetraporum kurokawense Habu & Baba, 1958 Holotype
Male adult
Kurokawa, NIIGATA, 27. VII. 1957, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-025
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Plataphodes) tetraporum kurokawense Habu & Baba, 1958 Holotype
Male adult
Kurokawa, NIIGATA, 27. VII. 1957, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-025
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Plataphodes) tetraporum kurokawense Habu & Baba, 1958 Holotype
Male adult
Kurokawa, NIIGATA, 27. VII. 1957, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-025
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Plataphodes) tetraporum kurokawense Habu & Baba, 1958 (Label) Holotype
Male adult
Kurokawa, NIIGATA, 27. VII. 1957, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-025
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Plataphodes) fujiyamai Habu, 1958 Holotype
Male adult
Mt. Shozu-dake, Hakuba, NAGANO, 31. VIII. 1949, I. Fujiyama
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-026
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Plataphodes) fujiyamai Habu, 1958 Holotype
Male adult
Mt. Shozu-dake, Hakuba, NAGANO, 31. VIII. 1949, I. Fujiyama
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-026
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Plataphodes) fujiyamai Habu, 1958 Holotype
Male adult
Mt. Shozu-dake, Hakuba, NAGANO, 31. VIII. 1949, I. Fujiyama
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-026
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Plataphodes) fujiyamai Habu, 1958 (Label) Holotype
Male adult
Mt. Shozu-dake, Hakuba, NAGANO, 31. VIII. 1949, I. Fujiyama
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-026
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Alexander Anichtchenko
Bembidion (Plataphodes) daisetsuzanum Habu, 1958 Holotype
Male adult
Mt. Daisetsu, HOKKAIIDO, 15. VII. 1952, T. Shirozu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-027
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Plataphodes) daisetsuzanum Habu, 1958 Holotype
Male adult
Mt. Daisetsu, HOKKAIIDO, 15. VII. 1952, T. Shirozu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-027
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Plataphodes) daisetsuzanum Habu, 1958 Holotype
Male adult
Mt. Daisetsu, HOKKAIIDO, 15. VII. 1952, T. Shirozu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-027
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Plataphodes) daisetsuzanum Habu, 1958 Holotype
Male adult
Mt. Daisetsu, HOKKAIIDO, 15. VII. 1952, T. Shirozu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-027
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Trichoplataphus) inaense Habu, 1956 Holotype
Male adult
Tenryu River-side, Iijima, Kamiina, NAGANO, 06. X. 1945, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-028
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Trichoplataphus) inaense Habu, 1956 Holotype
Male adult
Tenryu River-side, Iijima, Kamiina, NAGANO, 06. X. 1945, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-028
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Trichoplataphus) inaense Habu, 1956 Holotype
Male adult
Tenryu River-side, Iijima, Kamiina, NAGANO, 06. X. 1945, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-028
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Trichoplataphus) inaense Habu, 1956 (Label) Holotype
Male adult
Tenryu River-side, Iijima, Kamiina, NAGANO, 06. X. 1945, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-028
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Peryphus) koikei Habu & Baba, 1960 Holotype
Male adult
Kurokawa, NIIGATA, 27. VII. 1957, K. Baba<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-029<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Peryphus) koikei Habu & Baba, 1960 Holotype
Male adult
Kurokawa, NIIGATA, 27. VII. 1957, K. Baba<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-029<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Peryphus) koikei Habu & Baba, 1960 Holotype
Male adult
Kurokawa, NIIGATA, 27. VII. 1957, K. Baba<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-029<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Ocydromus) echigonum Habu & Baba, 1957 Holotype
Male adult
Mt. Amakazari (alt. 800m), NIIGATA, 25. VI. 1955, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-030
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Ocydromus) echigonum Habu & Baba, 1957 Holotype
Male adult
Mt. Amakazari (alt. 800m), NIIGATA, 25. VI. 1955, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-030
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Ocydromus) echigonum Habu & Baba, 1957 Holotype
Male adult
Mt. Amakazari (alt. 800m), NIIGATA, 25. VI. 1955, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-030
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Ocydromus) echigonum Habu & Baba, 1957 (Label) Holotype
Male adult
Mt. Amakazari (alt. 800m), NIIGATA, 25. VI. 1955, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-030
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (INCERTAE) umeyai Habu, 1959 Holotype
Male adult
Jozankei, HOKKAIDO, 18. IX. 1954, K. Umeya
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-031
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (INCERTAE) umeyai Habu, 1959 Holotype
Male adult
Jozankei, HOKKAIDO, 18. IX. 1954, K. Umeya
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-031
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (INCERTAE) umeyai Habu, 1959 Holotype
Male adult
Jozankei, HOKKAIDO, 18. IX. 1954, K. Umeya
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-031
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (INCERTAE) umeyai Habu, 1959 (Label) Holotype
Male adult
Jozankei, HOKKAIDO, 18. IX. 1954, K. Umeya
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-031
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Sinechostictus (Sinechostictus) kyushuensis (Habu, 1957) Holotype
Male adult
Mt. Hiko, FUKUOKA, 03. XI. 1952, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-032
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Sinechostictus (Sinechostictus) kyushuensis (Habu, 1957) Holotype
Male adult
Mt. Hiko, FUKUOKA, 03. XI. 1952, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-032
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Sinechostictus (Sinechostictus) kyushuensis (Habu, 1957) Holotype
Male adult
Mt. Hiko, FUKUOKA, 03. XI. 1952, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-032
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Ocydromus) negrei Habu, 1958 Holotype
Male adult
Mt. Osore, AOMORI, 01. VIII. 1953, A. Habu Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-033. (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html). Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Ocydromus) negrei Habu, 1958 Holotype
Male adult
Mt. Osore, AOMORI, 01. VIII. 1953, A. Habu<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-033<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Ocydromus) negrei Habu, 1958 Holotype
Male adult
Mt. Osore, AOMORI, 01. VIII. 1953, A. Habu. Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-033 (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html) Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Nipponobembidion) ainu Habu & Baba, 1968a: 145 Holotype
Male adult
Mt. Daisetsu, HOKKAIDO, 26. VI. 1958, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-034
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Nipponobembidion) ainu Habu & Baba, 1968a: 145 Holotype
Male adult
Mt. Daisetsu, HOKKAIDO, 26. VI. 1958, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-034
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Nipponobembidion) ainu Habu & Baba, 1968a: 145 Holotype
Male adult
Mt. Daisetsu, HOKKAIDO, 26. VI. 1958, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-034
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Nipponobembidion) ainu Habu & Baba, 1968a: 145 Holotype
Male adult
Mt. Daisetsu, HOKKAIDO, 26. VI. 1958, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-034
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (INCERTAE) ehikoense Habu, 1984 Holotype
Male adult
Mt. Hiko, FUKUOKA, 03. VIII. 1948, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-035
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (INCERTAE) ehikoense Habu, 1984 Holotype
Male adult
Mt. Hiko, FUKUOKA, 03. VIII. 1948, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-035
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (INCERTAE) ehikoense Habu, 1984 Holotype
Male adult
Mt. Hiko, FUKUOKA, 03. VIII. 1948, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-035
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (INCERTAE) ehikoense Habu, 1984 (Label) Holotype
Male adult
Mt. Hiko, FUKUOKA, 03. VIII. 1948, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-035
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Neoemphanes) shimoyamai Habu, 1978 Holotype
Male adult
Oirase, Nishitsugaru-gun, AOMORI, 10. VIII. 1973, K. Shimoyama
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-036
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Neoemphanes) shimoyamai Habu, 1978 Holotype
Male adult
Oirase, Nishitsugaru-gun, AOMORI, 10. VIII. 1973, K. Shimoyama
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-036
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Neoemphanes) shimoyamai Habu, 1978 Holotype
Male adult
Oirase, Nishitsugaru-gun, AOMORI, 10. VIII. 1973, K. Shimoyama
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-036
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Neoemphanes) shimoyamai Habu, 1978 Holotype
Male adult
Oirase, Nishitsugaru-gun, AOMORI, 10. VIII. 1973, K. Shimoyama
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-036
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Neoemphanes) shimoyamai Habu, 1978 (Label) Holotype
Male adult
Oirase, Nishitsugaru-gun, AOMORI, 10. VIII. 1973, K. Shimoyama
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-036
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Asioperyphus) gotoense Habu, 1973 Holotype
Male adult
Goto, Tomie, NAGASAKI, 02. V. 1970, M. Ezima
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-037
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Asioperyphus) gotoense Habu, 1973 Holotype
Male adult
Goto, Tomie, NAGASAKI, 02. V. 1970, M. Ezima
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-037
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Asioperyphus) gotoense Habu, 1973 Holotype
Male adult
Goto, Tomie, NAGASAKI, 02. V. 1970, M. Ezima
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-037
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Asioperyphus) gotoense Habu, 1973 (Label) Holotype
Male adult
Goto, Tomie, NAGASAKI, 02. V. 1970, M. Ezima
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-037
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Bembidion (Trichoplataphus) ozarkense Maddison & Hildebrandt, 2011 Paratype
Male adult
USA: Missouri: Carter Co., Current River at Van Buren, voucher DRMV100311
Copyright © - - -
Bembidion (Trichoplataphus) rolandi Fall, 1922
Male adult
USA: Virginia: Rockbridge Co., Maury River at Glasgow, voucher DRMV100193
Copyright © ---
Bembidion (Hydrium) laevigatum Say, 1830
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -
Bembidion (Peryphanes) geberti Marggi, 2011 Holotype
Male adult
Türkei, Provinz Nevşehir, NE Avanos Kızılırmak, temporärer Flusslauf, 24.5.2001, leg. J. Gebert [38°46.070 N, 34°53.755 E], coll.
Naturhistorisches Museum Bern (NMBE).
Foto: Ortwin Bleich
Copyright © ---
Bembidion (Plataphodes) tetraporum Bates, 1883: 270
Female adult
Kuriles isl., Kunashir
Copyright © Kirill Makarov
Bembidion (Testediolum) armeniacum kokandicum Solsky, 1874: 120
Female adult
Copyright © Kirill Makarov
Bembidion (Bembidion) quadrimaculatum mandli Netolitzky, 1932
Female adult
Copyright © Kirill Makarov
Asaphidion substriatum Andrewes, 1925a: 56
Male adult
India, Uttarakhand, 2-3 km SW of Sonprayag, 2000-2200 m asl, 30°37' N, 78°59' E, 8-9.04.2012, R. Dudko, I. Lyubechanskii
R. Dudko det. 2012
Copyright © Roman Dudko
Bembidion (Princidium) compactum Andrewes, 1922c: 240
Female adult
India, Uttarakhand, 10 km NNE of Ramnagar, gorge of left trib. of Kosi River, 400-450 m,
29°29' N, 79°09,5' E, R. Dudko, I. Lyubechanskii, 19-20.04.2012
det. R. Dudko 2012
Copyright © Roman Dudko
Bembidion (Lymnaeum) nigropiceum (Marsham, 1802)
Male adult
Thompson Island, 7.5.2008, Massachusetts
Images J. Rawlins
Copyright © ---
Elaphropus (Tachyura) ocellatus (Bates, 1892)
Female adult
India, Uttarakhand, 5 km NNE of Ramnagar, valley of Kosi River, 400-420 m, 29°26' N, 79°08,5' E, 21.04.2012, R. Dudko, I. Lyubechanskii
det. R. Dudko 2012
Copyright © Roman Dudko
Elaphropus (Tachyura) bombycinus (Andrewes, 1925)
Male adult
India, Uttarakhand, 5 km NNE of Ramnagar, valley of Kosi River, 400-420 m, 29°26' N, 79°08,5' E, 21.04.2012, R. Dudko, I. Lyubechanskii
det. R. Dudko 2012
Copyright © Roman Dudko
Tachyura (Amaurotachys) rhombophorus (Andrewes, 1925)
Male adult
India, Uttarakhand, 8 km NNE of Ramnagar, valley of Pipaliasot River, 420 m, 29°28,5' N, 79°10' E, 20.04.2012, R. Dudko, I. Lyubechanskii<br /> det. R. Dudko 2012
Copyright © Roman Dudko
Elaphropus (Tachyura) klugii (Nietner, 1858)
Female adult
India, Uttarakhand, 10 km NNE of Ramnagar, gorge of left trib. of Kosi River, 400-450 m,
29°29' N, 79°09,5' E, 19-20.04.2012, R. Dudko, I. Lyubechanskii
det. R. Dudko 2012
Copyright © Roman Dudko
Tachyura (Amaurotachys) notaphoida (Bates, 1886)
Male adult
India, Uttarakhand, 5 km NNE of Ramnagar, valley of Kosi River, 400-420 m, 29°26' N, 79°08,5' E, 21.04.2012, R. Dudko, I. Lyubechanskii
Copyright © Roman Dudko
Bembidion (Asioperyphus) altestriatum Netolitzky, 1934a: 68
Male adult
Russia, SE Primorie, Lazo Nat. res., 10 km SW Sokol'chi vill.
(Приморский кр., Лазовский зап., 10 км SW пос. Сокольчи, окр. кордона Корпадь 43°15' N, 134°7' E, 23-27.06.10, leg. М.Смирнов)
Image of M. Smirnov (М.Э. Смирнов) from www.zin.ru\Animalia\Coleoptera
Copyright © Maksim Smirnov
Bembidion (Ocydromus) zolotarewi Reitter, 1910b: 313
Male adult
Russia, NW Caucas, Krasnodar reg., Tuapse distr., Podchrebetovoe vicinity
(NW Кавказ, Краснодарский край, Туапсинский р-н, окр пос. Подхребeтовое 19.V.2012 leg. М. Смирнов)
Image of M. Smirnov (М.Э. Смирнов) from www.zin.ru\Animalia\Coleoptera
Copyright © Maksim Smirnov
Elaphropus (Elaphropus) zouhari (Jedlicka, 1961)
Female adult
SE Приморский кр., Лазовский зап., ~10 км SW пос. Сокольчи, окр. кордона Корпадь, 43°15' N, 134°7' E, 27.06.06 leg. М.& Л.Смирновы
Russia, SE Primorie, Lazo Nat. res., 10 km SW from Sokol'chi vill.
Image of M. Smirnov (М.Э. Смирнов) from www.zin.ru\Animalia\Coleoptera
Copyright © Maksim Smirnov
Bembidion (Plataphus) gebleri persuasum Netolitzky, 1938a: 38
Male adult
SE Приморский кр., Лазовский зап., побережье Японского моря, бухта Тачингоуза, 27.06.04 leg. М.& Л.Смирновы
Russia, SE Primorie, Lazo Nat. res., seashore, Tachingouz bay
Image of M. Smirnov (М.Э. Смирнов) from www.zin.ru\Animalia\Coleoptera
Copyright © Maksim Smirnov
Bembidion (Bembidionetolitzkya) cyaneum Chaudoir, 1846a: 203
Male adult
N Кавказ, Карачаево-Черкесия, окр. г.Теберда 1.VII.1994 leg. А. Гусаков
N Caucas, Karachaevo-Cherkessia, Teberda vicinity
Image of M. Smirnov (М.Э. Смирнов) from www.zin.ru\Animalia\Coleoptera
Copyright © Maksim Smirnov
Bembidion (Bembidionetolitzkya) tibiale (Duftschmid, 1812)
Male adult
NW Кавказ, 20 км S г. Горячий Ключ, бер. горной речки 3.04.1996 leg. М. Смирнов
NW Caucas, 20 km S from Goriachii Kluch
Image of M. Smirnov (М.Э. Смирнов) from www.zin.ru\Animalia\Coleoptera
Copyright © Maksim Smirnov
Bembidion (Ecuadion) ricei Maddison & Toledano, 2012 Paratype
Male adult
ECUADOR: Napo: Rio Chalpi Grande, 2800m, 0.3645°S, 78.0852°W, D.R. Maddison voucher V100622
Scale bar is 1 mm
image from (Maddison & Toledano, 2012)
Copyright © ---
Bembidion (Ecuadion) georgeballi Toledano, 2008
Male adult
ECUADOR: Pichincha: Quebrada Lozada, on road to Res. Yanacocha, 3460m, 0.1105°S, 78.5642°W, voucher V100658<br /> Scale bar is 1 mm.<br /> Image from Maddison & Toledano, 2012
Copyright © - - -
Asaphidion transcaspicum (Semenov, 1889)
Female adult
Kazakhstan, Alma-Ata, 2.06.1990
Copyright © Alexander Anichtchenko
Asaphidion championi Andrewes, 1924
Female adult
N INDIA, Uttarakhand, 17-18.04.2012, Uttarkashi distr., N30°45'15.72", E78°34'32.86", 1374 m, 14 km E Uttarkashi
Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko
Asaphidion stierlini (Heyden, 1880d)
Male adult
Italy, Basilicata, Laurenzana (PZ), 5.X.1997, 1100m, F. Angelini leg.
Copyright © Alexander Anichtchenko
Asaphidion caraboides (Schrank, 1781)
Male adult
France, B. Alp., Digne, 1000 m, 23.VII.1974, P. Poot leg.
Copyright © Alexander Anichtchenko
Bembidion (Peryphanes) adygorum Belousov & Sokolov, 1996
Male adult
NW Caucas, Krasnodar reg., Tuapse distr., Podkhrebetovoe vill. vicinity, 19.V.2012 leg. M. Smirnov
Фотография с сайта zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov
Bembidion (Peryphanes) grapii Gyllenhal, 1827
Male adult
NW Caucas, Krasnodar reg., Tuapse distr., Podkhrebetovoe vill. vicinity, 19.V.2012 leg. M. Smirnov <br /> Фотография с сайта zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov
Bembidion (Nepha) tetrasemum Chaudoir, 1846a: 207
Male adult
NW Caucas, Krasnodar reg., Tuapse distr., Podkhrebetovoe vill. vicinity, 19.V.2012 leg. M. Smirnov
Фотография с сайта zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov
Bembidion (Plataphus) infuscatipenne Netolitzky, 1938
Male adult
Russia, SE Primorie, Lazo Nat. res., ~10 км SW Sokolchi vill., Korpad' vicinity 43°15'N / 134°7'E, 23-27.06.10, leg. M. Smirnov
Фотография с сайта zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov
Bembidion (Microserrullula) luniferum Andrewes, 1924
Female adult
N INDIA, Uttarakhand, 22-23.04.2012, 8 km NW Jaspur, lake shore, 29°21'10.54", 78°48' 30.99", Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko
Bembidion (Microserrullula) xanthacrum Chaudoir, 1850b: 175
Male adult
N India, Uttarakhand, left side of Kosi riv., 5 km N Ramnagar, N29*432 E79*140, 7-11.06.2011 Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko
Bembidion (Microserrullula) splendens Andrewes, 1923
Male adult
N India, Uttarakhand, left side of Kosi riv., 5 km N Ramnagar, N29*432 E79*140, 23-26.04.2012 Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko
Bembidion (Diplocampa) fumigatum (Duftschmid, 1812)
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Bembidion (Notaphemphanes) ephippium (Marsham, 1802)
Male adult
S Hungary, Kiskunsági Nemzeti Park; 3,2mm
Copyright © Aleš Sedláček
Bembidion (Bracteon) argenteolum Ahrens, 1812
Male adult
E Slovakia, Tisa rive; 6,5mm
Copyright © Aleš Sedláček
Bembidion (Bracteon) litorale (Olivier, 1790)
Female adult
E Slovakia, Tisa rive; 6mm
Copyright © Aleš Sedláček
Bembidion (Chlorodium) pygmaeum ab. bilunulatum Bielz, 1852
Male adult
N Moravia, Týn nad Bečvou; 2,5mm. det. J. Coulon, 2015
Copyright © Aleš Sedláček
Cillenus (Cillenus) lateralis Samouelle, 1819
Male adult
Netherlands, Bruinisse, leg. Krift, 1976; det. U.Schmidt, 1982
Photo: U.Schmidt, 2008
Copyright © Udo Schmidt
Bembidion (Blepharoplataphus) hastii C.R. Sahlberg, 1827
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Bembidion (Blepharoplataphus) hastii C.R. Sahlberg, 1827
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Bembidion (Bembidionetolitzkya) cyaneum Chaudoir, 1846
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Bembidion (Bracteon) velox (Linne, 1761)
Male adult
E Poland, Siemiatycze env., Bug rive; 5,5mm
Copyright © Aleš Sedláček
Bembidion (Chlorodium) pygmaeum (Fabricius, 1792)
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Bembidion (Diplocampa) transparens (Gebler, 1829)
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Bembidion (Peryphus) captivorum Netolitzky, 1943
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Bembidion (Philochthus) guttula (Fabricius, 1792)
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Bembidion (Philochthus) inoptatum Schaum, 1857a: 149
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Bembidion (Plataphus) lucillum lucillun Bates, 1883
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Bembidion (Semicampa) schueppelii Dejean, 1831
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Sinechostictus (Sinechostictus) lederi (Reitter, 1888)
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Bembidion (Metallina) elevatum (Motschulsky, 1844)
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Bembidion (Peryphanes) olegleonidovici Fassati, 1990: 169
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Sinechostictus (Sinechostictus) nordmanni (Chaudoir, 1844)
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Tachys (Paratachys) micros (Fischer von Waldheim, 1828)
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Geocharidius phineus Erwin, 1982
Male adult
Guatemala
image from Sokolov I., 2013. A new genus and eight new species of the subtribe Anillina (Carabidae, Trechinae, Bembidiini) from Mexico, with a cladistic analysis and some notes on the evolution of the genus.
Copyright © Igor Sokolov
Nesamblyops sp.
Male adult
New Zealand
Scale bars = 0.5mm
image from Sokolov I., 2013. A new genus and eight new species of the subtribe Anillina (Carabidae, Trechinae, Bembidiini) from Mexico, with a cladistic analysis and some notes on the evolution of the genus.
Copyright © Igor Sokolov
Zapotecanillus montanus Sokolov, 2013 Paratype
Male adult
MEXICO, Oaxaca, 52miles N of Oaxaca, Ber.202, sink litter, 17 May 1971 S.B.Peck collector (CMNH)
Scale bar = 0.5mm
Copyright © Igor Sokolov
Zapotecanillus oaxacanus Sokolov, 2013 Paratype
Male adult
MEXICO. Oaxaca 18.7mi S Valle Nacional 5200’ 17 Aug.1973\ A.Newton Collector
Scale bar = 0.5mm
Copyright © Igor Sokolov
Bembidion (Notaphocampa) niloticum Dejean, 1831
Female adult
4,4 mm
Namibia, Etosha National Park, Okaukuejo, 1400 mб leg. U.Schmidt, 18.-20.III.1994;
det. M.Baehr, 2012
Photo: U.Schmidt, 2014
Copyright © Udo Schmidt
Zapotecanillus nanus Sokolov, 2013 Paratype
Female adult
MEXICO, Oaxaca, 18.7mi N Valle National, 5200’ 17 Aug.1973\ A.Newton Collector
Copyright © Igor Sokolov
Zapotecanillus ixtlanus Sokolov, 2013 Paratype
Male adult
MEXICO, Oaxaca, 37mi S Valle National
Copyright © Igor Sokolov
Zapotecanillus iviei Sokolov, 2013 Paratype
Male adult
OAXACA, 2 mi S. Cerro Pelon, 8–9000 ft. 03 JUL 1982 M.A. Ivie colr.
Copyright © Igor Sokolov
Zapotecanillus kavanaughi Sokolov, 2013 Paratype
Male adult
MEXICO, Oaxaca, 14km N San Juan
Copyright © Igor Sokolov
Zapotecanillus pecki Sokolov, 2013 Paratype
Male adult
MEXICO, Oaxaca, 3.5mi S Suchixtepec
Copyright © Igor Sokolov
Zapotecanillus longinoi Sokolov, 2013 Paratype
Male adult
MEXICO, Chiapas, Sierra Morena
Copyright © Igor Sokolov
Bembidion (Bembidionetolitzkya) varicolor agurense Lutshnik, 1938
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Bembidion (Princidium) punctulatum punctulatum s.str.
Male adult
Germany, Bavaria, Ascholding, Isar, leg. det. U.Schmidt, 3.V.1975 Photo: U.Schmidt, 2012
Copyright © Udo Schmidt
Bembidion (Bracteon) stenoderum stenoderum Bates, 1873a: 300
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Bembidion (Diplocampa) transparens prostratum (Motschulsky, 1844)
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Bembidion (Eupetedromus) dentellum (Thunberg, 1787)
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Bembidion (Peryphus) subcostatum subcostatum (Motschulsky, 1850)
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Bembidion (Philochthus) mannerheimii C.R. Sahlberg, 1827
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Sinechostictus (Sinechostictus) ruficornis Sturm, 1825
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Bembidion (Metallina) properans (Stephens, 1828)
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Anillinus langdoni Sokolov & Carlton, 2004
Male adult
USA-TN, Sevier Co, GSMNP, secondary site, 490 m, ex leaf et log litter, 35°43.157' N, 83°23.309' W, 19.04.2011, J.-S. Park det. I. Sokolov, 2013
Copyright © Roman Dudko
Bembidion (Peryphanes) milleri Jacquelin du Val, 1852
Male adult
4,5 mm; Germany, Rheinland-Pfalz, Eifel, Gerolstein, leg. J.Scheuern, 30.IV.1983; det. F.Koehler, 1994; Photo: U.Schmidt, 2014
Copyright © Udo Schmidt
Bembidion (Ocydromus) scopulinum (Kirby, 1837)
Male adult
Russia, Kunashir isl., Severjanka riv., 27.VIII.2014 (Кунашир, пойма р. Северянка)
Copyright © Kirill Makarov
Bembidion (Ocydromus) scopulinum (Kirby, 1837)
Male adult
Russia, Kunashir isl., Severjanka riv., 27.VIII.2014 (Кунашир, пойма р. Северянка)
Copyright © Kirill Makarov
Elaphropus (Tachyura) sexstriata (Dufischmid, 1812)
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Sinechostictus (Sinechostictus) frederici (G. Muller, 1918)
Female adult
Spain, Jaen prov., Arroyo de la Piedra del Aguila; 3-4.05.2014, 37.683885, -3.485062, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko
Bembidion (Notholopha) scitulum (Erichson, 1834)
Male adult
CHILE: Reg. VII: Los Niches E of Curicó, DRM voucher V100790; Image from Maddison D.R., 2014. Scale bar 1 mm.
Copyright © - - -
Bembidion (Antiperyphus) philippii Germain, 1906
Male adult
Argentina: Neuquén: Rio Collón Curá ca 13 km S La Rinconada, DRM voucher V100787. Image from Maddison D.R., 2014. Scale bar 1 mm.
Copyright © - - -
Bembidion (Notholopha) sexfoveatum Germain, 1906
Male adult
CHILE: Reg. IX: 16.3 km E Malalcahuello, Cuesta Las Raices, DRM voucher V100598.. Image from Maddison D.R., 2014. Scale bar 1 mm.
Copyright © - - -
Bembidion (Antiperyphanes) mandibulare (Solier, 1849)
Male adult
Chile: Reg. X, Chiloé: Cucao, DRM voucher V100789. Image from Maddison D.R., 2014. Scale bar 1 mm.
Copyright © - - -
Bembidion (Ecuadion) andersoni Toledano, 2008
Male adult
ECUADOR: Pichincha: Reserva Yanacocha, start Andean Snipe Trail, DRM voucher V100655. Image from Maddison D.R., 2014. Scale bar 1 mm.
Copyright © - - -
Bembidion (Nothonepha) eburneonigrum Germain, 1906
Male adult
Argentina: Neuquén: Rio Neuquén at Chos Malal, DRM voucher V100785. Image from Maddison D.R., 2014.
Copyright © - - -
Anillinus affabilis (Brues, 1902) Lectotype
Male adult
Image from Sokolov I.M., Reddell J.R. & Kavanaugh D.H., 2014 Scale = 0.5 mm.
Copyright © Igor Sokolov
Anillinus sinuatus (Jeannel, 1963) Holotype
Female adult
Image from Sokolov I.M., Reddell J.R. & Kavanaugh D.H., 2014 Scale = 0.5 mm. (Deposited in NMNH)
Copyright © Igor Sokolov
Anillinus depressus (Jeannel, 1963) Holotype
Female adult
Image from Sokolov I.M., Reddell J.R. & Kavanaugh D.H., 2014 Scale = 0.5 mm. (NMNH)
Copyright © Igor Sokolov
Bembidion (Ecuadion) chimborazonum (Bates, 1891)
Male adult
ECUADOR: Pichincha: Paso de la Virgen, DRM voucher V100793. Image from Maddison D.R., 2014.
Copyright © - - -
Bembidion (Ecuadion) sanctaemarthae Darlington, 1934
Male adult
ECUADOR: Napo: Rio Chalpi Grande. Image from Maddison D.R., 2014.
Copyright © - - -
Bembidion (Ecuadion) ricei Maddison & Toledano, 2012
Male adult
ECUADOR: Napo: Rio Chalpi Grande. Scale bar 1 mm. Image from Maddison D.R., 2014.
Copyright © - - -
Bembidion (Ecuadion) cotopaxi Moret & Toledano, 2002
Male adult
ECUADOR: Pichincha: km 17 on route 28 W of Papallacta. Scale bar 1 mm. Image from Maddison D.R., 2014.
Copyright © - - -
Bembidion (Ecuadion) walterrossii Toledano, 2008
Male adult
ECUADOR: Napo: Vinillos, 4.1 km S Cosanga. Scale bar 1 mm. Image from Maddison D.R., 2014.
Copyright © - - -
Bembidion (Nothonepha) tetrapholeon Maddison, 2014 Paratype
Male adult
Argentina: Neuquén: Arroyo Queñi at Lago Queñi, 830m, 40.1575°S, 71.721°W, / 10–11.ii.2007. DRM 07.035. D.R. Maddison, S.A.Roig. Scale bar 1 mm. Image from Maddison D.R., 2014.
Copyright © - - -
Bembidion (Nothonepha) tetrapholeon Maddison, 2014 Paratype
Female adult
Argentina: Chubut: Rio Azul at Lago Puelo. Scale bar 1 mm. Image from Maddison D.R., 2014.
Copyright © - - -
Bembidion (Nothonepha) lonae Jensen-Haarup, 1910
Male adult
Argentina: Mendoza: Salinas del Diamante. Scale bar 1 mm. Image from Maddison D.R., 2014.
Copyright © - - -
Bembidion (Nothonepha) engelhardti engelhardti
Male adult
Argentina: Neuquén: Rio Salado at route 40. Scale bar 1 mm. Image from Maddison D.R., 2014.
Copyright © - - -
Bembidion (Nothonepha) germainianum (Toledano, 2002)
Male adult
Argentina: Neuquén: Rio Salado at route 40. Scale bar 1 mm. Image from Maddison D.R., 2014.
Copyright © - - -
Bembidion (Nothonepha) tucumanum (Jeannel, 1962)
Male adult
Argentina: Mendoza: Salinas del Diamante. Scale bar 1 mm. Image from Maddison D.R., 2014.
Copyright © - - -
Bembidion (Chilioperyphus) orregoi Germain, 1906
Male adult
Argentina: Chubut: Rio Azul at Lago Puelo. Scale bar 1 mm. Image from Maddison D.R., 2014.
Copyright © - - -
Bembidion (Antiperyphanes) hirtipes (Jeannel, 1962)
Male adult
Argentina, Mendoza, Pampa Palauco. Image from Maddison D.R., 2014.
Copyright © - - -
Bembidion (Antiperyphanes) rufoplagiatum Germain, 1906
Male adult
Argentina, Neuquen, Arroyo Queni at Lago Queni. Image from Maddison D.R., 2014.
Copyright © - - -
Anillinus forthoodensis Sokolov & Reddell, 2014 Holotype
Male adult
TX: Bell Co., Talking Crows Cave, Fort Hood, 4.V.2006, J. Fant, M. Reyes \ Texas Memorial Museum Invertebrate Zool Coll #45.781 \ Holotype Anillinus forthoodensis Sokolov & Reddell 2014 [red label] \ CAS Type No. 18872
Copyright © Igor Sokolov
Anillinus affabilis (Brues, 1902)
Male adult
TEXAS, Travis County, Tooth Cave
Copyright © Igor Sokolov
Anillinus acutipennis Sokolov & Reddell, 2014 Paratype
Female adult
TX: Bell Co., Hidden Pit Cave, Fort Hood, 27.X.2007, J. Reddell, M. Reyes \ Texas Memorial Museum Invertebrate Zool Coll #60.107
Copyright © Igor Sokolov
Anillinus comalensis Sokolov & Kavanaugh, 2014 Paratype
Female adult
TX: Comal Co., 7mi W New Braunfels, 27.I.1995, A. G. Grubbs
Copyright © Igor Sokolov
Anillinus wisemanensis Sokolov & Kavanaugh, 2014 Holotype
Male adult
TX: Hays Co., Wiseman Sink, 28.IV.1995, A. G. Grubbs \ Texas Memorial Museum Invertebrate Zool Coll #27.149 \ Holotype Anillinus wisemanensis Sokolov & Kavanaugh 2014 [red label] \ CAS Type No. 18873
Copyright © Igor Sokolov
Bembidion (Trepanes) octomaculatum (Goeze, 1777)
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Bembidion (Nepha) caucasicum (Motschulsky, 1844)
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Bembidion (Talanes) aspericolle (Germar, 1829)
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Sakagutia umi Sasakawa, 2007
Male adult
SE Приморье, Лазовский р-н., побережье Японского моря, 3 км NE пос. Преображение бухта Ежовая (Тасовая), литораль, 12.07.2010, leg. М. Смирнов. Image from www.zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov
Bembidion (Semicampa) chaudoirii Chaudoir, 1850b: 179
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Bembidion (Peryphanes) dalmatinum dalmatinum Dejean, 1831
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Bembidion (Peryphanes) adygorum Belousov & Sokolov, 1996: 14
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Bembidion (Peryphus) obscurellum obscurellum (Motschulsky, 1845)
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Bembidion (Eupetedromus) ruthenum Tschitscherine, 1895c: 298
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Bembidion (Asioperyphus) pamiricola pamiricola Lutshnik, 1930: 34
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Bembidion (Asioperyphus) bandotaro Morita, 1991a: 119
Female adult
Kuriles isl. det. K. Makarov
Copyright © Kirill Makarov
Bembidion (Antiperyphanes) zanettii Toledano, 2008
Male adult
Ecuador: Napo: Rio Quijos W of Baeza. Image from Maddison D.R., 2014.
Copyright © - - -
Bembidion (Antiperyphanes) spinolai Solier, 1849
Male adult
Argentina: Chubut: Rio Azul at Lago Puelo. Image from Maddison D.R., 2014.
Copyright © - - -
Hypotyphlus degiovannii Magrini, 2013 Holotype
Male adult
Italia, Lazio, Montagna di Scauri (Latina), 50 m s.l.m., 1.V.2008, P. Magrini leg. (CM).
Copyright © Paolo Magrini
Cryptocharidius mandibularis M.Etonti & Mateu, 1992
Male adult
Peru, Dept. Ancash, Cordillera Blanca, Punta Olimpica Pass, m 4400 a.s.l., G. Allegro legit
Copyright © Gianni Allegro
Bembidion (Philochthus) cf. neresheimeri J. Muller, 1929
Female adult
Ивановская обл., г. Иваново, парк, 1905 г., 29.05.2014, М. Смирнов leg.; det. K.Makarov. Image from www.zin.ru\Animalia\Coleoptera
Copyright © Maksim Smirnov
Bembidion (Nepha) schmidti subcallosum Wollaston, 1864: 71
Male adult
3,6 mm; Spain, Canary Islands, Tenerife, Teno, Erjos, 1000 m, leg. J.Schoenfeld, 11.VII.2010; det. Lompe, 2012. Photo: U.Schmidt, 2014
Copyright © Udo Schmidt
Bembidion (Philochthus) lunulatum (Geoffroy, 1785)
Male adult
Turkey, Marmaris, Hisarönü, leg. U.Schmidt, 18.V.1997; det. Marggi, 2014; Photo: U.Schmidt, 2014. 3,8 mm
Copyright © Udo Schmidt
Tachys (Paratachys) oblitus Casey, 1918
Male adult
Copyright © Ben Smith
Copyright © - - -
Bembidion (Nipponobembidion) ruruy Makarov & Sundukov, 2014 Holotype
Male adult
Northern Kunashir, 2.5 km NW of Cape Nelyudimyi, N44°29.433', E146°11.783', 4.VIII.2013, leg. K. Makarov & Y. Sundukov” and “HOLOTYPE Bembidion (Nipponobembidion) ruruy Makarov & Sundukov 2014 [printed on red paper]
Copyright © Kirill Makarov
Bembidion (Nipponobembidion) ruruy Makarov & Sundukov, 2014 Paratype
Female adult
Copyright © Kirill Makarov
Bembidion (Ocydromus) transsylvanicum Bielz, 1852
Male adult
W Ukraine, NE Carpathians, Transcarpathian Reg., Marmarosh Mts., near Lug vill., bank of Kuziy (Kizi) stream, 400m, 26.IV.2014, Panin R. & Kanarsky Yu. leg. L= 5 mm
Copyright © Ruslan Panin
Amerizus (Tiruka) gaoligongensis Gueorguiev, 2015 Paratype
Female adult
CHINA, Yunnan, Tengchong County, cave Wu Shi Shan 1-2-3, near vil. Kongshuhe, ca. 2580 m, N25.73111 E98.66459, 11.11.2013, B. Petrov leg.; Scale bar = 1.5 mm.
Copyright © Borislav Gueorguiev
Bembidion (Zeperyphodes) nesophilum Broun, 1886
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
Copyright © - - -
Zecillenus karikari Larochelle & Larivière, 2015
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
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Bembidion (Zecillenus) alacre (Broun, 1921)
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
Copyright © - - -
Bembidion (Zecillenus) albescens (Bates 1878)
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
Copyright © - - -
Bembidion (Zecillenus) tepaki Larochelle & Larivière, 2015
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
Copyright © - - -
Bembidion (Zecillenus) tillyardi (Brookes, 1927)
Female adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
Copyright © - - -
Bembidion (Zecillenus) waimarama Larochelle & Larivière, 2015
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
Copyright © - - -
Bembidion (Zecillenus) puponga Larochelle & Larivière, 2015
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
Copyright © - - -
Bembidion (Zecillenus) chalmeri (Broun, 1886)
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
Copyright © - - -
Bembidion (Zeplataphus) tairuense Bates, 1878
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
Copyright © - - -
Bembidion (Zeactedium) giachinoi Toledano, 2005
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
Copyright © - - -
Bembidion (Zeperyphodes) callipeplum Bates, 1878
Female adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
Copyright © - - -
Bembidion (Zemetallina) wanakense Lindroth, 1976
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78.
Copyright © - - -
Bembidion (Zemetallina) chalceipes Bates, 1878
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78.
Copyright © - - -
Bembidion (Zemetallina) bullerense Larochelle & Larivière, 2015
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78.
Copyright © - - -
Bembidion (Zemetallina) hokitikense Bates, 1878
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78.
Copyright © - - -
Elaphropus (Tachyura) cf. fuscicauda (Bates, 1873)
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Elaphropus (Tachyura) diabrachys (Kolenati, 1845)
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Elaphropus (Tachyuropsis) exaratus (Bates, 1873)
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Elaphropus (Tachyura) tetraspilus (Solsky, 1874)
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Bembidion (Peryphus) captivorum Netolitzky, 1943a: 29
Male adult
Россия, SE Приморский кр., Лазовский зап., ~12 км S пос. Старая Каменка, окр. кордона Америка, 43°15'N / 134°01'E, 10.07.04, leg. М. & Л. Смирновы. image from www.zin.ru
Copyright © Maksim Smirnov
Bembidion (Terminophanes) pulcherrimum (Motschulsky, 1850)
Male adult
Caucasus, Elbrus, Itkol (CTVR). Image from Toledano L. & Gueorguiev B., 2015
Copyright © - - -
Bembidion (Ochthedromus) americanum Dejean, 1831
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -
Asaphidion indicum (Chaudoir, 1850)
Male adult
India, Uttarakhand, Nainital distr., Peora vill. env., 1950m, 20.X-6.XI.2003, S. Saluk leg. det. Anichtchenko A. 2015 3.8 mm
Copyright © Alexander Anichtchenko
Bembidion (Philochthus) biguttatum (Fabricius, 1779)
Male adult
Russia, Ivanovo, "Kharinka" city park, 30.V.2015, M. Smirnov leg.
Copyright © Maksim Smirnov
Bembidion (Testedium) laetum Brulle, 1836: pl.2
Female adult
Morocco, 65 km N Essaouria, 5 km N Jamaa Shoim, 23-25.III.2011, A. Gusakov. Фотография с сайта zin.ru\Animalia\Coleoptera.
Copyright © Maksim Smirnov
Elaphropus (Sphaerotachys) fumigatus (Motschulsky, 1851)
Female adult
2,2 mm; Senegal, M`Bour, environment Club Aldiana leg. U.Schmidt, 8.-20.X.1992; det. M.Baehr, 2014; Photo: U.Schmidt, 2016
Copyright © Udo Schmidt
Elaphropus (Tachyura) germanus (Chaudoir, 1876)
Female adult
2,2 mm; Gambia, Serrekunda, envir. Senegambia Beach Hotel leg. U.Schmidt, 16.-22.XI.1990; det. M.Baehr, 2014; Photo: U.Schmidt, 2016
Copyright © Udo Schmidt
Bembidion (Ocydromus) decorum caraffai Sainte-Claire Deville, 1906a: 15
Female adult
France Corse
Copyright © Pascal Deschamps
Bembidion (Bembidionetolitzkya) longipes K. Daniel, 1902a: 29
Male adult
France, 05380 Chateauroux les Alpes Le Rabioux, 02-VII-2014
Copyright © Pascal Deschamps
Bembidion (Notaphus) obliquum Sturm 1825: 160
Male adult
Russia, Ивановская обл., Пестяковский р-н, окр. дер. Пурежка, бер. р. Лух 24-25.VI.2011 М. Смирнов. Фотография с сайта www.zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov
Asaphidion ganglbaueri J. Muller, 1921
Female adult
S Turkey, Antalya prov., Kemer reg., Göynük env. 16-20.IV.2015 M. Smirnov. Фотография с сайта www.zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov
Bembidion (Eupetedromus) immaturum Lindroth, 1954 Paratype
Male adult
W Newfoundland, Deer Lake, 30.V.51, Lindroth. 4254:2 (MZLU)
Copyright © Lund University
Bembidion (Notaphocampa) foveolatum Dejean, 1831
Female adult
Bembidion sobrinum Boheman, 1848 det. M.Baehr, 2014; 4,2 mm. Namibia, Otjiwarongo, Kalkfeld, 30 km East, Mt. Etjo Safari Lodge, 1500 m, leg. U.Schmidt, 15.III.1994; Photo: U.Schmidt, 2016
Copyright © Udo Schmidt
Tachyura ascendens (Alluaud, 1917)
Female adult
TANZANIA, Kilimanjaro, Nkuansa, grassland,1660m, 3.7.2011, R. Felix det. as Elaphropus (Tachyura) exiguus (BOHEMAN 1848)
Copyright © Peter Schüle
Elaphropus (Sphaerotachys) hoemorroidalis (Ponza, 1805)
Female adult
2,5 mm. Namibia, Etosha National Park, Okaukuejo, 1400 m, leg. U.Schmidt, 19.III.1994; det. M.Baehr, 2014. Photo: U.Schmidt, 2016
Copyright © Udo Schmidt
Bembidion (Chlorodium) leucolenum Bates, 1873a: 275
Female adult
Kunashir Island, lower reaches of Krivonozhka Stream, 20-22.VI.2013, Yu. & L. Sundukov leg.
Copyright © Kirill Makarov
Bembidion (Chlorodium) leucolenum Bates, 1873a: 275
Male adult
Kunashir Island, watershed of Golovnina and Khlebnikova rivers west of Dubovoe, 28.IX.2015, Yu. Sundukov
Copyright © Kirill Makarov
Bembidion () pseudolucillum Netolitzky, 1938a: 37 EA: JA
Male adult
Kunashir Island, creek mouth near Moristyi Island, 17.VIII. 2013, K. Makarov
Copyright © Kirill Makarov
Bembidion () pseudolucillum Netolitzky, 1938a: 37 EA: JA
Female adult
Kunashir Island, creek mouth near Moristyi Island, 17.VIII. 2013, K. Makarov
Copyright © Kirill Makarov
Bembidion () pseudolucillum Netolitzky, 1938a: 37 EA: JA
Female adult
Kunashir Island, creek mouth near Moristyi Island, 17.VIII. 2013, K. Makarov
Copyright © Kirill Makarov
Bembidion () pseudolucillum Netolitzky, 1938a: 37 EA: JA
Female adult
Kunashir Island, creek mouth near Moristyi Island, 17.VIII. 2013, K. Makarov
Copyright © Kirill Makarov
Bembidion (Trichoplataphus) parconaturaviva Toledano, 2010: 394
Male adult
SE Maritime Prov., Lazovsky dist., ~10 кm SW Sokoltchi vill., 23-27.VI.2010 leg.М. Smirnov. Фотография с сайта www.zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов. det. K. Makarov 2016
Copyright © Maksim Smirnov
Bembidion (Neja) nigricorne Gyllenhal, 1827: 402
Female adult
Czech Republic, Bohemia occ., Kraslice, Tisovec hill, 750-755 m a.s.l., 24.iv.1999, lgt. et det. J. Hejkal
Copyright ©
Micratopus sp.
Male adult
Panama, Darien Nat. Park, Rancho Frio, 10.I.2017, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko
Bembidion (Peryphus) tetracolum tetracolum Say, 1823: 89
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Bembidion (Ocydromus) decorum subconvexum K. Daniel & J. Daniel, 1902: 102
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Tachys doddi Sloane, 1903 Syntypes
Male adult
Tachys (Paratachys) plagiatus plagiatus
Copyright © Museums Victoria
Pelodiaetodes (Pelodiaetodes) constricticollis Sokolov, 2015 Paratype
Male adult
NZ, Northland, Radar Bush. Image from Sokolov, 2015
Copyright © - - -
Bembidion (Jammuphanes) sinuosum Schmidt & Marggi, 2014 Paratype
Male adult
Pakistan, Kaghan valley, Shogran, 3050m, 19.6.2001. Imagr from Schmidt & Marggi, 2014
Copyright © - - -
Bembidion (Paraprincidium) karinae Schmidt, 2009 Holotype
Female adult
NEPAL, Prov. Koshi, Tumlingtar, Arun-Ufer [Arun river bank], 300 mNN, 20.X.2002 leg. J. Weipert (Naturkundemuseum Erfurt). Image from Schmidt, 2009
Copyright © - - -
Bembidion (Andrewesa) patris Schmidt, 2010 Paratype
Male adult
NEPAL Annapurna Mts. / Madi Khola below Sikles / 1400 m, water power st. [station, 28°22N 84°08E] / 10.5.1996 leg. J. Schmidt. Image from Schmidt, 2010
Copyright © - - -
Setitachys macrops Baehr, 2016 Paratype
Female adult
Qld. Bentinck Is. Rokoti, 16 June 1963. P. Aitken, N. B. Tindale
Copyright © Martin Baehr
Tachyxysta howdenorum Boyd & Erwin, 2016 Holotype
Male adult
México, Chiapas, El Aguacero.
Copyright © Terry Erwin
Elaphropus (Nototachys) occidentalis Boyd & Erwin, 2016 Paratype
Male adult
Perú, Madre de Dios, Pakitza
Copyright © Terry Erwin
Elaphropus (Idiotachys) acutifrons Boyd & Erwin, 2016 Holotype
Female adult
Brazil, Santarem
Copyright © Terry Erwin
Meotachys (Scolistichus) riparius Boyd & Erwin, 2016 Paratype
Female adult
Brazil, Amazonas, Rio Solimões, Ilha de Marchantaria
Copyright © Terry Erwin
Elaphropus (Ammotachys) marchantarius Boyd & Erwin, 2016 Paratype
Male adult
Venezuela, San Carlos de Rio Negro
Copyright © Terry Erwin
Meotachys (Hylotachys) ballorum Boyd & Erwin, 2016 Paratype
Female adult
Brazil, Amazonas, Tapurucuara
Copyright © Terry Erwin
Stigmatachys uvea Boyd & Erwin, 2016 Holotype
Female adult
Perú, Loreto, Campamento San Jacinto
Copyright © Terry Erwin
Xystosomus laevis Erwin, 1973 Holotype
Male adult
Brasilien, Nova Teutonia
Copyright © - - -
Bembidion (Terminophanes) kaschmirense Netolitzky, 1920
Female adult
India, Uttarakhand, 5 km NE Govind Ghat, road to Gangaria, 9-10.6.2011, Anichtchenko A. leg. det. J. Schmidt 2015 8 mm
Copyright © Alexander Anichtchenko
Bembidion (Asioperyphus) xanthochiton Andrewes, 1922b: 176
Female adult
India, Uttarakhand, 14 km NW New Tehri, 950m, N30*28'25.3" E 078*20'56.2", 10.4.2012, A. Anichtchenko leg. det. J. Schmidt 2015
Copyright © Alexander Anichtchenko
Bembidion (Testediolum) gagates gagates Andrewes, 1924a: 72
Male adult
India, Uttarakhand, 13-15.4.2012, Uttarkashi distr., left tributary of Bhagirathi riv., 2300m, N30*57'41.57" E78*41'54.75", Anichtchenko A. leg. det. J. Schmidt 2015
Copyright © Alexander Anichtchenko
Bembidion (Bembidionetolitzkya) cimmerium Andrewes, 1922c: 238
Male adult
India, Uttarakhand, 5km NW Ghangaria, Valley of Flowers, N30*43'356" E079*85.309 , 3417 m, 06.2011, Anichtchenko A. leg. 5.5 mm. det. J. Schmidt 2015
Copyright © Alexander Anichtchenko
Bembidion (Omoperyphus) kempi Andrewes, 1922b: 178
Male adult
India, Uttarakhand, 13-15.4.2012, Uttarkashi distr., left tributary of Bhagirathi riv., 2150m, N30*57'43" E78*41'54.75", Anichtchenko A. leg. det. J. Schmidt 2015
Copyright © Alexander Anichtchenko
Bembidion (Peryphus) bracculatum Bates, 1889b: 212
Female adult
India, Uttarakhand, 5km NW Ghangaria, Valley of Flowers, N30*43'356" E079*85.309 , 3417 m, 06.2011, Anichtchenko A.; 5.5 mm, det. J. Schmidt, 2015
Copyright © Alexander Anichtchenko
Bembidion (Trichoplataphus) kara Andrewes, 1921d: 250
Female adult
India, Uttarakhand, 15 km SW New Tehri, 869 m, N30*15'874" E 078*21'587", 18-20.4.2012, A. Anichtchenko leg.; 6 mm; det. J. Schmidt, 2015
Copyright © Alexander Anichtchenko
Bembidion (Ocydromus) phaedrum Andrewes, 1923b: 97
Female adult
India, Uttarakhand, 15 km SW New Tehri, 869 m, N30*15'874" E 078*21'587", 18-20.4.2012, A. Anichtchenko leg.; 5 mm; det. J. Schmidt 2015
Copyright © Alexander Anichtchenko
Bembidion () eutherum Andrewes, 1923b: 96
Male adult
India, Uttarakhand, 15 km SW New Tehri, 869m, 18-20.4.2012, Anichtchenko A. leg.; det. J. Schmidt 2015
Copyright © Alexander Anichtchenko
Bembidion (Ocydromus) hasurada Andrewes, 1924a: 77
Male adult
India, Uttarakhand, 10-12.iV.2012, 14km E Uttarkashi, Anichtchenko A. leg.; 5 mm. det. J. Schmidt, 2015
Copyright © Alexander Anichtchenko
Bembidion (Pamirium) zlotini Mikhailov, 1996: 71
Female adult
det. I.Kabak et B.Kataev SE Kazakhstan, Zailiysky Alatau mts. Kok-Zhaylau. N 43°08'15" / E 77°00'08". h=2270 m. 24-26.06.2013. S.Kolov leg. Image of S. Kolov, from www.zin.ru
Copyright © - - -
Bembidion (Lionepha) disjunctum Lindroth, 1963 Holotype
Male adult
Copyright © - - -
Bembidion (Gnatholymnaeum) spurcum Blackburn, 1881
Male adult
Hawaii. Image from J.K. Liebherr, 2008
Copyright © - - -
Bembidion (Peryphus) transversale Dejean, 1831
Male adult
Images from David R. Maddison & Alexander P. Swanson, 2010
Copyright © - - -
Bembidion (Hirmoplataphus) hirmocaelum Chaudoir, 1850
Female adult
Siberia, Irkutsk, Irkut riv., 3.6.1997, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko
Bembidion (Trichoplataphus) deplanatum A. Morawitz, 1862a: 262
Male adult
E Siberia, Irkutsk, Irkut riv., 3.VI.1997, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko
Bembidion (Pamirium) platypterum Solsky, 1874: 120
Female adult
Kazakhstan, Alma-Ata, 2.06.1990. det. Anichtchenko A. 2016
Copyright © Alexander Anichtchenko
Bembidion (Terminophanes) mckinleyi scandicum Lindroth, 1943
Female adult
E Siberia, Lake Baikal, Malyi Kochergat vill., Goloustnaya riv., 16.VII.1991, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko
Bembidion (Asioperyphus) ovale Motschulsky, 1844: 239
Male adult
Siberia, E Sajan Mts., Sentsa riv, trib. of Oka riv., 1700m, 9-10.7.1998, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko
Bembidion (Peryphus) jedlickai jedlickai Fassati, 1945: 22
Female adult
Siberia, E Sajan Mts., Sentsa riv, trib. of Oka riv., 1700m, 9-10.7.1998, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko
Bembidion (Peryphus) bualei bualei Jacquelin du Val, 1852
Male adult
Italy, Lombardia, Brescla, Sabbio Chiese, 20.5.2011, E. Ruzzier leg.
Copyright © Alexander Anichtchenko
Bembidion (Asioperyphus) lunatum Duftschmid, 1812: 211
Male adult
Latvia, Daugavpils distr., Silene, Ilgas, 18.5.1991, A. Barsevskis leg.
Copyright © Alexander Anichtchenko
Bembidion (Asioperyphus) sajanum Shilenkov in Kryzhanovskij & al., 1995
Male adult
Siberia, E Sajan Mts., Sentsa riv, trib. of Oka riv., 1700m, 9-10.7.1998, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko
Scotodipnus alpinus Baudi di Selve, 1871
Female adult
Piemonte, Valle Po, Crissolo (CN), 27.V.2000, 1250m, fagus, leg. F. Angelini
Copyright © Alexander Anichtchenko
Scotodipnus (Binaghites) affinis affinis (Baudi di Selva, 1871)
Male adult
Italy, Emilia, M. Penna (PR), 1400m, 24.5.1999, leg. F. Angelini
Copyright © Alexander Anichtchenko
Bembidion (Nepha) hauserianum Netolitzky, 1918
Female adult
Iran, 20 km SW Yasuj, 5-6.V.2007, Anichtchenko A. leg.
Copyright © Alexander Anichtchenko
Sinechostictus (Sinechostictus) moschatus (Peyron, 1858)
Male adult
Iran, Lorestan, 50km N Ardimeshk, 2000m, Sar Takht 3-4.V.2007, Anichtchenko A. leg.; det. L. Toledano 2008
Copyright © Alexander Anichtchenko
Bembidion (Trichoplataphus) oxyglymma Bates, 1883
Male adult
Japan, Honshu Pref., Gifu, Tohuyama Dam, Ibigawa river, 16.VII.1992, leg. B. Kuznetsov. det. L. Toledano
Copyright © Alexander Anichtchenko
Bembidion (Ocyturanes) parsorum Netolitzky, 1934
Female adult
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin L=4.7 mm
Copyright © Kirill Makarov
Bembidion (Ocyturanes) parsorum Netolitzky, 1934
Male adult
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin L=4.9 mm
Copyright © Kirill Makarov
Asaphidion flavicorne (Solsky, 1874)
Male adult
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin L=4.0
Copyright © Kirill Makarov
Bembidion (Emphanes) latiplaga flavibase De Monte, 1956a: 179
Male adult
Morocco, Souss-Massa Draa, Zagora, OuedDraa riv., 1.2.2013. det. M. Toribio, 2017. 2.5 mm
Copyright © Alexander Anichtchenko
Bembidion (Euperyphus) combustum combustum Menetries, 1832: 138
Female adult
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin L=7.0 mm
Copyright © Kirill Makarov
Bembidion (Peryphanes) phryganobium Belousov et Sokolov, 1996:24
Female adult
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin L=6.5 mm
Copyright © Kirill Makarov
Bembidion (Peryphus) hummleri hummleri J. Muller, 1918
Male adult
Morocco, Quarzazate prov., Quwd Drag N 30*30'26" W006*06'01", near Tin-Zoulin vill., 27.6.2010, A. Anichtchenko leg. det. M. Toribio, 2017
Copyright © Alexander Anichtchenko
Tachyura (Tachyura) nitens (Andrewes, 1925)
Female adult
Indonesia E, W New Guinea, Doberai Peninsula, Ayamaru vill. 27 km SE Uter (Aitinyo) lakeside, 1*23'01S 132*23'09E, 175-177m, 4.IX.2015, muddy bank, between lakeside vegetation, leg. D. Telnov. 2.8 mm. det. M. Baehr, 2016
Copyright © Alexander Anichtchenko
Tachyura (Sphaerotachys) curticollis (Sloane, 1896)
Male adult
Indonesia E, W New Guinea, Doberai Peninsula, Ayamaru vill. 27 km SE Uter (Aitinyo) lakeside, 1*23'01S 132*23'09E, 175-177m, 4.IX.2015, muddy bank, between lakeside vegetation, leg. D. Telnov. 2.2 mm. det. M. Baehr, 2016
Copyright © Alexander Anichtchenko
Bembidion (Peryphanes) dalmatinum haupti Reitter, 1908a: 110
Male adult
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin L=5.7 mm
Copyright © Kirill Makarov
Bembidion (Peryphanes) dalmatinum haupti Reitter, 1908a: 110
Female adult
Iran, Lorestan prov., Dorud distr., near Cham Chit vill. 13.V.2017 leg. D.Kasatkin L=5.6 mm
Copyright © Kirill Makarov
Bembidion (Diplocampa) blandulum Netolitzky, 1910a: 221
Male adult
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin L=3.6 mm
Copyright © Kirill Makarov
Bembidion (Diplocampa) blandulum Netolitzky, 1910a: 221
Female adult
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin L=4.0 mm
Copyright © Kirill Makarov
Bembidion (Nepha) hauserianum Netolitzky, 1918a: 23
Male adult
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin L=4.1 mm
Copyright © Kirill Makarov
Bembidion (Nepha) hauserianum Netolitzky, 1918a: 23
Female adult
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin L=4.2 mm
Copyright © Kirill Makarov
Bembidion (Bembidionetolitzkya) penninum Netolitzky, 1918a: 20
Male adult
Val Cairasca Sempione VI-1915 Mancini
Copyright © Jan Muilwijk
Bembidion (Peryphus) irregulare Netolitzky, 1934b: 90
Male adult
Shyock valley betw. Karghalik & Saser Brangsa, 4150-4500 m, 24.VI-11.VIII.1930
Copyright © Jan Muilwijk
Eotachys (Eotachys) enormis Dostal, 2015 Holotype
Male adult
Madagascar, Andasibe National Park, Lokato, near Andasibe, 10.X.2007, leg. Zd. Mracek (in coll. Dostal, Vienna). Scale bar 1 mm
Copyright © Alexander Dostal
Orzolina thalassophila Machado, 1987
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison
Bembidion (Desarmatocillenus) aestuarii (Habu & Ueno, 1954)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison
Bembidion (Cillenus) palosverdes Kavanaugh and Erwin, 1992
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison
Bembidion (Australoemphanes) ateradustum Liebherr, 2008
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison
Bembidion (Notaphus) contractum Say, 1823
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison
Bembidion (Peryphodes) ephippigerum (Leconte, 1852)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison
Bembidion (Notaphus) indistinctum Dejean, 1831
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison
Bembidion (Desarmatocillenus) kasaharai Habu, 1978
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison
Bembidion (Cillenus) lateralis Samouelle, 1819
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison
Bembidion (Lymnaeum) laticeps (Leconte, 1858)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison
Bembidion (Lymnaeum) nigropiceum (Marsham, 1802)
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Bembidion (Ocyturanes) praeustum Dejean, 1831
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Bembidion (Limnaeoperyphus) quadriimpressum (Motschulsky, 1860)
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Bembidion (Peryphus) sejunctum Casey, 1918
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Bembidion (Leuchydrium) tigrinum Leconte, 1879
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Bembidion (Desarmatocillenus) yakushimanum Sasakawa, 2007
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Bembidion (Bracteon) zephyrum Fall, 1910
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Amerizus (Amerizus) oblongulus (Mannerheim, 1852)
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Anomotachys acaroides (Motschulsky, 1859)
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Elaphropus (Elaphropus) cf. haliploides Bates, 1892
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Kiwitachys cf. antarcticus Bates, 1874
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Nothoderis rufotestacea Hayward, 1900
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Tachys (Paratachys) scitulus Leconte, 1848
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Pericompsus (Pericompsus) laetulus LeConte, 1851
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Sinechostictus (Pseudolimnaeum) alesmetana Toledano, 2008
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Tachys (incertae) mulwalensis Sloane, 1900
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Tachyura (Tachylopha) leleupi (Basilewsky In Basilewsky & Straneo, 1950)
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Caecidium trechomorphum Ueno, 1971b: 150 Paratype
Male adult
(MNHN) Image from Toledano, 2011
Copyright © - - -
Neolovricia ozimeci Lakota, Jazlic & Moravec, 2009 Holotype
Female adult
Spilja u Radinovcima, Dedici, Rastovac, Biokovo, 25.10.2006 Coleoptera (1) Leg: Roman Ozimec. Deposited in the Croatian Natural History Museum, Zagreb. Image from Lakota, Jazlic & Moravec, 2009
Copyright © - - -
Winklerites stevanovici Hlavac & Margini, 2017 Paratype
Male adult
Srbija [p], Dimitrovgrad, Petrlaš, Džemanska prop[ast], 02.05.2002 [h], M. Stevanović leg. Image from HLAVÁČ & MAGRINI, 2017
Copyright © - - -
Scotodipnus (Binaghites) armellinii (Ganglbauer, 1900)
Male adult
Veneto Italia
Copyright © Erminio Piva
Bembidion (Sloanephila) tahitiense Liebherr & Maddison, 2013 Holotype
Male adult
(MNHN): French Polynesia: Tahiti Nui / Mt. Mauru lava tube / el. 705 m 18-IX-2006 lot 01 / 17˚38.017´S 149˚21.284´W / pyr. fog moss/liverworts / along stream J.K. Liebherr // HOLOTYPE / Bembidion / tahitiense / Liebherr & / Maddison 2012 (black-bordered red label).
Copyright © David Maddison
Bembidion (Sloanephila) jacksoniense Guerin Meneville, 1830
Female adult
Wiluna W.A. / ix 25 1931 //Australia / Harvard Exp. / Darlington (MCZ). Scale bars 0.5 mm.
Copyright © David Maddison
Bembidion (Odontium) subfusum Darlington, 1959 Holotype
Male adult
Central plains of Luzon Is., P. J. Feb.-Sept. 1945 Darlington; Bembidion (Bracteon) subfusum Darl. (MCZ).
Copyright © - - -
Sinechostictus (Pseudolimnaeum) cf. pendleburyi (Andrewes, 1931)
Female adult
Philippines, N Luzon, Mt. Province, Mt. Polis 7.2014. det. Anichtchenko A. 2018
Copyright © Alexander Anichtchenko
Bembidion (Bembidionetolitzkya) chaklaense Schmidt, 2018 Paratype
Male adult
“TIBET South Centr. 16.VII.07 / 50 km NNE Lhasa, SW of / Chak La pass 4100-4550m / Chakla river valley, lg. Schmidt”, “30°03’36.6N 91°16’51.7E / to / 30°06’10.1N 91°16’38.4E” (CSCHM).
Copyright © Joachim Schmidt
Bembidion (Bembidionetolitzkya) lhai Schmidt, 2018 Paratype
Male adult
“TIBET South Centr. 17-20.VI.07 / Budha Vall. N of Yangpachem / ca. 30°10’56N 90°29’21E / 5000-5200m, leg. J. Schmidt” (CSCHM).
Copyright © Alexander Anichtchenko
Bembidion (Bembidionetolitzkya) pinnigerum Schmidt, 2018 Holotype
Male adult
“TIBET South Centr. 17/18.VII.07 / Reting Tsangpo Vall. E Reting / 4230-4800 m, Kiykiy place / ca. 30°24’15N 91°42’59E lg.J.S.” (CSCHM).
Copyright © Joachim Schmidt
Bembidion (Bembidionetolitzkya) lunepabomaguvi lunepabomaguvi Paratype
Male adult
“INDIA Arunachal Pradesh / 10-15 km W Sela Pass / 34-3500 m, 24.VI.2008 / 27°30N 92°03E lg. Reuter” (CSCHM).
Copyright © Joachim Schmidt
Bembidion (Bembidionetolitzkya) lunepabomaguvi bhutanicum Schmidt, 2018 Paratype
Male adult
“BT Wangdue Phodrang distr. / Phobjikha Vall., 2800 m / 28.IX.2017, leg. J. Schmidt / 27°24’40”N 90°12’12”E” (NBCB).
Copyright © Joachim Schmidt
Bembidion (Bembidionetolitzkya) namtso Schmidt, 2018 Paratype
Male adult
“TIBET (South Central) 12.VII.07 / S Namtso, Langma Valley / 5100-5150 m, leg. Schmidt / 30°37’39,1”N90°51’56,5”E” (CSCHM).
Copyright © Joachim Schmidt
Bembidion (Bembidionetolitzkya) retingense Schmidt, 2018 Paratype
Male adult
“CHINA Tibet 1.VII.2015 / side valley S Reting / 4400 m leg. J. Schmidt / 30°16’44”N 91°31’55”E” (CSCHM).
Copyright © Joachim Schmidt
Bembidion (Bembidionetolitzkya) smetanai qingmaiense Schmidt, 2018 Paratype
Male adult
“China, S.W. Sichuan / Qingmai 3000 m / N 28°48’46.7”, E 099°52’09.2” / 3.7.2006, lgt. Janata, M.” (CSCHM).
Copyright © Joachim Schmidt
Bembidion (Bembidionetolitzkya) transhimalayanum Schmidt, 2018 Paratype
Female adult
“S TIBET, ca. 40km SW of / Yangpachem, 13.VII.2010 / Pijin moraine 5000-5500 m / 29°45’36N 90°17’54E” (CSCHM).
Copyright © Joachim Schmidt
Bembidion (Bembidionetolitzkya) wrzecionkoi Toledano & Schmidt, 2008
Male adult
south central Tibet, NE of Gyantse, Yung La pass and above, 4850-5400 m, 28°57’14”N 89°50’04”E, 17.VII.2010
Copyright © Joachim Schmidt
Bembidion (Bembidionetolitzkya) yamtso Schmidt, 2018 Holotype
Male adult
“S TIBET 14/15.VI.2011 / E-slope Karo La pass / 4900-5300 m, leg. Schmidt / 28°55’N 90°18’E” (CSCHM).
Copyright © Joachim Schmidt
Bembidion (Bembidionetolitzkya) zugmayeri Schmidt, 2018 Holotype
Male adult
“Sumdschiling / Ebene [plateau] W Tibet”, “Süssw. Tümp. [fresh water pond] / VII/06. Zugm.” (ex coll. W. Marggi, Thun, in CSCHM).
Copyright © Joachim Schmidt
Bembidion (Josefia) praeceptor Schmidt, 2018 Paratype
Male adult
“Tibet (South Central) 26.VI.07 / Gyama vall. ca. 60km E Lhasa / 4230-4620 m leg. J. Schmidt / ca. 29°41’48N 91°38’29E”, “south ascent of / Chungenpo Shong side vall” (CSCHM).
Copyright © Joachim Schmidt
Bembidion (Ocyturanes) christophi Schmidt, 2018 Paratype
Male adult
“S TIBET 15.VII.2010 / Tibetan Himalaya / above Kampa La 48-4850 m” (CSCHM).
Copyright © Joachim Schmidt
Bembidion (Ocyturanes) gracilipenne Schmidt, 2018 Paratype
Male adult
“S TIBET, N Lhasa Valley / above Chupsang Monast. / 19.IX.2010, 4900m / 29°44’01N 91°07’47E ” (CSCHM).
Copyright © Joachim Schmidt
Bembidion (Ocyturanes) lhatseense Schmidt, 2018 Paratype
Male adult
“S TIBET 17.VI.2011 / above pass Tsuo La / 47-4940 m, leg. Schmidt / 29°03’37N 87°58’52E” (CSCHM).
Copyright © Joachim Schmidt
Bembidion (Peryphophila) problematicum Schmidt, 2018 Holotype
Male adult
“NEPAL, Ghatte / Kirola – Khola. Camp / 3500m. Pasang 12.5.88” (CSCHM).
Copyright © Joachim Schmidt
Bembidion (Peryphophila) pseudoclarum Schmidt, 2018 Holotype
Male adult
“NEPAL Manaslu Mts. / Marsyangdi Khola vall. / Bhachok Khola 1000m / 13.4.1999 leg. Schmidt” (CSCHM).
Copyright © Joachim Schmidt
Bembidion (Peryphus) ahrensi Schmidt, 2018 Paratype
Male adult
“Uttar Ganga Ebene / bei Dhorpathan / 3000 m, 6.-8.3.”, “C-NEPAL-Himalaya / Dhaulagiri-Mts. / leg. Ahrens 1994” (CSCHM).
Copyright © Joachim Schmidt
Bembidion (Peryphus) aparupa Schmidt, 2018 Paratype
Male adult
“W-NEPAL Baglung Lekh / W Pokhara, 10-15km NW / Baglung, 2350-2550m / leg. Schmidt 10-12.5.04 / 28°18‘36”N 83°31‘56“E” (CSCHM).
Copyright © Joachim Schmidt
Bembidion (Peryphus) dhaulagiricum Schmidt, 2018 Holotype
Male adult
“NEPAL W Dhaulagiri / below Dhule 26-2700m / 7.IX.12 leg. J. Schmidt / 28°41‘07N 82°56‘31E” (CSCHM).
Copyright © Joachim Schmidt
Bembidion (Peryphus) glabripenne Schmidt, 2018 Paratype
Female adult
“Nepal, District Karnali, / Nähe Pohada 02.06.1997 / 3010-3400 mNN, leg. Grill, / Flußufer / HF, 29°04’N, 82°42’E” (CSCHM)
Copyright © Joachim Schmidt
Bembidion (Politophanes) inventor Schmidt, 2018 Holotype
Male adult
“E. NEPAL: KOSI / Val. Induwa Kola / 2000 m, 18.IV.84 / Löbl - Smetana” (MHNG).
Copyright © Joachim Schmidt
Bembidion (Terminophanes) reibnitzi Schmidt, 2018 Paratype
Male adult
“S TIBET Kyi Chu river / betw. Lhasa and airport / ca. 3650 m, 22.IX.2010 / leg. J. Schmidt” (CSCHM).
Copyright © Joachim Schmidt
Bembidion (Emphanes) bulgani Jedlicka, 1968
Female adult
Russia: Primorskii krai: 20 km NWW Spassk-Dal'ny, Khanka Lake, Khankaysky Nat. Res., kordon Vostochnyi, 3-8.VI 2008, A. Napolov leg.
Copyright © Kirill Makarov
Bembidion (Emphanes) tenellum tenellum Erichson, 1837
Male adult
C Russia, Ivanovo ar., Pestyaki distr., Demidovo vill. env., Lukh riv. 4-5.VII.2014 leg. M. Smirnov ; Фотография с сайта www.zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov
Tachys (Tachys) lenkoranus Csiki, 1928
Male adult
8 км З гю Карасук, Ю бер оз. Кусган, 53°44.2' N, 77°52.5' E, 4.05.2014, Р.Ю. и Е.Р. Дудко leg. Image from Dudko, Bespalov, Zinovyev & Lyubechanskii, 2018
Copyright © - - -
Bembidion (Nepha) glabrum (Motschulsky, 1850)
Male adult
Новосибирская обл., 9 км З г. Карасук, оз Кротовое. Image from Dudko, Bespalov, Zinovyev & Lyubechanskii, 2018
Copyright © - - -
Pelodiaetus sulcatipennis Jeannel, 1937
Male adult
New Zealand, Otago, Outram
Copyright © Igor Sokolov
Pelodiaetus nunni Sokolov, 2019 Paratype
Male adult
NZ, Canterbury, Ahuriri Scenic Reserve. 1.27 mm
Copyright © Igor Sokolov
Bembidion (Omotaphus) alsium Coquerel, 1866
Male adult
St. Helena Island. Image courtesy of David Maddison
Copyright © CC-BY 4.0
Bembidion (Endosomatium) megalops (Wollaston, 1877)
Female adult
St. Helena Island. Image courtesy of David Maddison
Copyright © CC-BY 4.0
Bembidion (Pseudophilochthus) nubigena (Wollaston, 1877)
Male adult
St. Helena Island. Image courtesy of David Maddison
Copyright © CC-BY 4.0
Bembidion (Apteromimus) platyderoides (Wollaston, 1877)
Male adult
St. Helena Island. Image courtesy of David Maddison
Copyright © CC-BY 4.0
Bembidion (Ecuadion) cotopaxi Moret & Toledano, 2002 Paratype
Female adult
Ecuador, Cotopaxi, Laguna Quilotoa, 3500m, 22.7.2001 P. Moret leg.
Copyright © Alexander Anichtchenko
Bembidion (Peryphus) cordicolle Jacquelin da Val, 1852
Male adult
Turkey, Kemer distr., Mts.Taurus, Ulupinar riv.,36*27'01N 30*26'14E 21.9.2014, I. Enuschenko leg. (DUBC) det. L. Toledano, 2019
Copyright © Alexander Anichtchenko
Bembidion (Philochthus) ellipticocurtum Netolitzky, 1935c: 167
Female adult
Lebanon, Jbail prov., Laqlouq, 8.5.2000. det. L. Toledano, 2019
Copyright © Alexander Anichtchenko
Bembidion (Ocyturanes) eucheres Netolitzky, 1943
Male adult
Tajikistan, Khoja Obi, Garm, 15.9.2011, Barsevskis A. leg. (DUBC) det. L. Toledano, 2019
Copyright © Alexander Anichtchenko
Bembidion (Terminophanes) terminale kirgisorum Netolitzky, 1934
Male adult
Tajikistan, Ayni distr., Sughd prov., Sarytag, 9km S lake Iskanderkul, 2380m, 6.7.2012, A. Barsevskis leg. (DUBC) det. L. Toledano, 2019
Copyright © Alexander Anichtchenko
Bembidion (Peryphanes) lirykense Reitter, 1908
Male adult
Iran, 40km S Ardabil, Goli, 20.5.2005, Anichtchenko A. leg. (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko
Sinechostictus (Sinechostictus) dahlii (Dejean, 1831)
Female adult
Spain, Jaen prov., Arroyo de la Piedra del Aguila, 3-4.5.2014, Anichtchenko A. leg. (DUBC)
Copyright © Alexander Anichtchenko
Sinechostictus (Sinechostictus) elongatus (Dejean, 1831)
Female adult
Spain, Madrid, Tres Cantos, Arroyo de Tejada, 10.4.2010, Anichtchenko A. leg. (DUBC)
Copyright © Alexander Anichtchenko
Sinechostictus (Sinechostictus) effluviorus (Peyron, 1858)
Male adult
Turkey, Kemer distr., Mts.Taurus, Ulupinar riv.,36*27'01N 30*26'14E 21.9.2014, I. Enuschenko leg. (DUBC) det. L. Toledano, 2019
Copyright © Alexander Anichtchenko
Bembidion (Phyla) phoeniceum Huber & Marggi, 1997: 778
Female adult
Turkey, Kirish distr., Camuyuva riv. 28.9.2014, I. Enischenko leg. (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko
Bembidion (Peryphus) persicum Menetries, 1832
Male adult
Tajikistan, Khoja Obi, Garm, 15.9.2011, Barsevskis A. leg. (DUBC) det. L. Toledano, 2019
Copyright © Alexander Anichtchenko
Bembidion (Ocyturanes) samai Neri & Toledano, 2017 Paratype
Male adult
Afghanistan, Vt. N Salang 1450 m, Khenjan, bord torrent, 11.8.1975, leg. Ledoux (DUBC)
Copyright © Alexander Anichtchenko
Bembidion (Bembidionetolitzkya) bactrianum K. Daniel, 1902
Male adult
Tadzhikistan, Varzob, Khodja obi Garm, 2000m 21-24.3.2015 (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko
Bembidion (Peryphus) abbreviatum pulpani Fassati, 1955
Male adult
Tadzhikistan, Ayni distr., Sughd prov., Iskanderkul lake, 2300m 5.7.2012, A. Barsevskis leg. (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko
Bembidion (Asioperyphus) notatum Andrewes, 1922b: 175
Female adult
India, Uttarakhand, 8 km NW Jaspur, lake shore, 22-23.4.2012, Anichtchenko A. leg. (DUBC). det. L. Toledano
Copyright © Alexander Anichtchenko
Bembidion (Plataphus) coelestinum (Motschulsky, 1844)
Female adult
N Mongolia, Tuul-Gol R., Engin Aimak Tev, 14.8.2013, I. Enuschenko leg. (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko
Bembidion (Plataphus) difficile (Motschulsky, 1844)
Male adult
Russia, Tuva rep.,Todzhynsk distr., E Sayan mts., Arzhkhan-Khem, 2200 m, 21.6.2017, I. Enuschenko leg. (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko
Bembidion (Ocyturanes) muilwijki Neri & Toledano, 2017 Paratype
Male adult
S Iran, Sisakht Dena, 2500-3000 m, 13-14.6.1973
Copyright © Alexander Anichtchenko
Bembidion (Peryphanes) fraxator Menetries, 1832
Male adult
Russia, Karachaevo-Cherkessia, urochishe Dzhalpak, 17-19.5.2009, A. Anichtchenko leg. (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko
Bembidion (Euperyphus) giganteum giganteum J.R. Sahlberg, 1900
Female adult
S Kazakhstan, N slope Talas Mt.rng., Dzhabagly riv., 1100m 28.7.2004, A.Matalin leg. (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko
Bembidion (Peryphus) morawitzi Csiki, 1928
Female adult
China, Liaoning, Kuandian co., Banlaing, Ba-che-chuan (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko
Bembidion (Ocyturanes) dyscheres Netolitzky, 1943
Female adult
N Iran, C Elburz, Val. Darband Sar, 2500-3000m 18.7.1970 (DUBC). det. L. Toledano
Copyright © Alexander Anichtchenko
Bembidion (Phyla) incommodum Netolitzky, 1926
Male adult
Iran, 40 km S Ardabil, Goli, 20.5.2005, Anichtchenko A. leg. (DUBC). det. L. Toledano
Copyright © Alexander Anichtchenko
Bembidion (Chlorodium) difforme (Motschulsky, 1844)
Female adult
Russia, Burjatia, Muhor-Shibirsk distr., 5km N Nikolsk vill., salt marsh, 27.8.1998, Anichtchenko A. leg & det.
Copyright © Alexander Anichtchenko
Bembidion (Ocydromus) saxatile morsum Netolitzky, 1930
Male adult
Tajikistan, Khoja Obi Garm, 15.9.2011, A. Barsevskis leg. (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko
Bembidion (Nepha) callosum subconnexum De Monte, 1953
Male adult
Spain, Gredos Mts., 2km E from La Mira, 1800m, 4-7.7.2007, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko
Bembidion (Nepha) schmidti alluaudi (Antoine, 1925)
Male adult
Morocco, Grand Atlas, 2200m Oukaimedene env., 16-17..6.2010, Anichtchenko A. leg. (DUBC). det. M. Toribio
Copyright © Alexander Anichtchenko
Bembidion (Eurytrachelus) vitiosum Gemminger & Harold, 1868
Female adult
Russia, Irkutsk reg., Nizhneudinsk distr., Uda riv., Bolshoy Uk waterfall, 22.7.1898, A.Anichtchenko leg. (DUBC).
Copyright © Alexander Anichtchenko
Bembidion (Trepanedoris) faraticolle (Jeanne, 1995) Holotype
Female adult
Image courtesy Biological Museum, Lund University
Copyright © - - -
Anillus angelae Magrini & Vanni, 1989
Male adult
Arnale cave, Coreno Ausonio, Italia
Copyright © Vittorino Monzini
Bembidion (Lionepha) australerasa Maddison, 2020 Holotype
Male adult
USA: California: Amador Co., Oyster Lake, Silver Lake Campground
Copyright © David Maddison
Bembidion (Lionepha) tuulukwa Maddison, 2020
Male adult
USA: Oregon: Benton Co., Marys Peak Rd, nr Alder Creek Falls
Copyright © David Maddison
Bembidion (Lionepha) lindrothi Maddison & Sproul, 2020 Holotype
Male adult
USA: California: Tulare Co., outlet of Emerald Lake
Copyright © David Maddison
Bembidion (Lionepha) kavanaughi Maddison, 2020 Holotype
Male adult
USA: Oregon: Wallowa Co., Lostine River Valley
Copyright © David Maddison
Bembidion (Lionepha) probata (Casey, 1918)
Male adult
USA: Oregon: Klamath Co., Odell Creek near Davis Lake
Copyright © David Maddison
Bembidion (Lionepha) pseudoerasum Lindroth, 1963
Male adult
USA: California: El Dorado Co., Strawberry Creek at Sciots Camp
Copyright © David Maddison
Bembidion (Lionepha) osculans Casey, 1918
Male adult
USA: California: Tehama Co., Nanny Creek
Copyright © David Maddison
Bembidion (Lionepha) erasa Leconte, 1859
Male adult
USA: Oregon: Benton Co., Marys Peak
Copyright © David Maddison
Bembidion (Lionepha) casta Casey, 1918
Male adult
USA: Oregon: Benton Co., Marys Peak, Alder Creek Falls
Copyright © David Maddison
Bembidion (Lionepha) sequoiae Lindroth, 1963
Male adult
USA: Oregon: Lane Co., School Creek near Lookout Point Reservoir
Copyright © David Maddison
Bembidion (Omotaphus) gorilla Bonavita, Toledano & Vigna Taglianti, 2016
Male adult
Copyright © David Maddison
Mioptachys flavicauda (Say, 1823)
Male adult
Ukraine, Vic. Odessa, forest Lusanovskiy (Liski) 29.4.2019 Gontarenko A.V. leg
Copyright © Alexander Anichtchenko
Tachyura (Tachyphanes) patruelis Baehr, 2016 Holotype
Male adult
Turkey, Pr. Siirt Botan Cayi-Tal 500 m, 23.4.1983 Leg. W. Schacht (Zoologische Staatssammlung München).
Copyright © Martin Baehr
Ocys monzinii Magrini & Vigna Taglianti, 2006
Male adult
1, Lombardy (Lecco), Monte Resegone, m 1750 a.s.l. (Brumano), surr. of Pizzo Brumano SW slope, 4.XI.2017, leg. D. Pedersoli (CP); 2, Lombardy (Bergamo), Monte Alben, m 1750 a.s.l. (Colle dei Brassamonti, Oneta), 14.V.2017, leg. M. Grottolo (CM). Image from Magrini et al, 2020
Copyright © - - -
Tachysbembix sirena Erwin, 2004 Holotype
Male adult
COSTA RICA: Osa, P.N. Corcovado, Llorona, 20km N Estación Sirena, 1–100 m, 08º 37’ N 083º 42’ W, LS 270500,508300, January (R.W. Matthews & J.R. Matthews)(INBio: ADP103930).
Copyright © Terry Erwin
Tachysbembix wendyporrasae Erwin, 2004 Holotype
Male adult
COSTA RICA: Guanacaste, Playa del Coco, 35 km W Liberia, sea level, 10° 33' N, 85° 40' W, July (R.T. Allen)
Copyright © Terry Erwin
Bembidion (Hydrium) laevigatum Say, 1830
Male adult
USA: Utah: Grand Co., Moab, Colorado River, 1190 m, 38.5739°N, 109.5769°W
Copyright © David Maddison
Bembidion (Hydrium) mimbres Maddison, 2020 Paratype
Male adult
USA: New Mexico: Grant Co., Gila River, Billings Vista, 1320 m, 32.8163°N, 108.6032°W, 11.viii.2005. DRM 05.043. D.R. & J.H.A. Maddison
Copyright © David Maddison
Bembidion (Ocydromus) corgenoma Maddison, 2020 Paratype
Male adult
USA: Oregon: Benton Co., Corvallis, Willamette River, 44.5491°N, 123.2449°W, 60 m
Copyright © David Maddison
Bembidion (Ocydromus) transversale Dejean, 1831
Male adult
Canada: Alberta: Burbank, junction of Red Deer and Blindman Rivers, 52.3542°N, 113.7556°W (voucher V101454)
Copyright © David Maddison
Bembidion (Ocydromus) erosum (Motschulsky, 1850)
Male adult
USA: California: Del Norte Co., Wilson Creek, 3 m, 41.6051°N, 124.1005°W (V101453)
Copyright © David Maddison
Bembidion (Ocydromus) perspicuum (Leconte, 1848)
Male adult
USA: Arizona: Cochise Co., San Pedro R at Charleston, 31.6239°N, 110.1722°W (voucher V101461)
Copyright © David Maddison
Bembidium haplogonum Chaudoir, 1868 (NEOTYPE)
Male adult
USA: California: Lake Co., North Branch Cache Creek at hwy 20, 305 m 38.9881°N, 122.5400°W
Copyright © David Maddison
Bembidion (Ocydromus) sarpedon Casey, 1918
Male adult
USA: Colorado: Las Animas Co., Cokedale, Reilly Canyon, 37.1346°N, 104.6114°W (V101459)
Copyright © David Maddison
Bembidion (Peryphiolus) caligatum Jeanne & Muller-Motzfeld, 1982
Male adult
Spain, Guadarrama, Cotos, 25.8.2001, Anichtchenko A. leg & det.
Copyright © Alexander Anichtchenko
Bembidion (Terminophanes) consummatum Bates, 1873
Male adult
Russia, E Siberia, Irkutsk reg., Bolshaya Rechka vill, 3.8.1990, Anichtchenko A. leg. & det
Copyright © Alexander Anichtchenko
Anillinus felicianus Sokolov, 2021 Holotype
Male adult
(NMNH): \ USA-LA: West Feliciana Par., ~4mi SW Jackson at 30.794°N, 91.254°W, mixed pine-hardwood forest, soil washing/berlese Sokolov I.M. 19–25 Apr 2018
Copyright © Igor Sokolov
Anillinus relictus Sokolov, 2021 Holotype
Male adult
(CMNH): ALABAMA: Blount Co., Tidwell Hollow Nature Trail east of Oneonta. T. N. King April 1 1972 \ 4/1/72 o [handwritten] \ THOMAS C. BARR COLLECTION 2011 Acc. No. 38,014
Copyright © Igor Sokolov
Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Sakhalin Isl., valley of the Rogatka River, 46°58'03"N 142°46'24"E, 18.vii.2011, leg. K.Makarov L=5,7
Copyright © Kirill Makarov
Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Sakhalin Isl., valley of the Rogatka River, 46°58'03"N 142°46'24"E, 18.vii.2011, leg. K.Makarov L=5,8
Copyright © Kirill Makarov
Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Kinashir Isl., 1.5 km south-west mouth of the Ozernaya River, 43°52'27"N 145°27'16"E, 26.vii.2011, leg. K.Makarov L=5,2
Copyright © Kirill Makarov
Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., 1.5 km south-west mouth of the Ozernaya River, 43°52'27"N 145°27'16"E, 26.vii.2011, leg. K.Makarov L=5.0
Copyright © Kirill Makarov
Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Kinashir Isl., Zmeinyy (Stolbovskoy) Stream, 44°00'31"N 145°40'49"E, 26.viii.2017, leg. K.Makarov L=5,75 mm
Copyright © Kirill Makarov
Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., Zmeinyy (Stolbovskoy) Stream, 44°00'31"N 145°40'49"E, 26.viii.2017, leg. K.Makarov L=5,6 mm
Copyright © Kirill Makarov
Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Kinashir Isl., mouth of the Medny Stream, 44°16'34"N 145°56'27"E, 16.viii.2017, leg. K.Makarov L=5,85 mm
Copyright © Kirill Makarov
Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., mouth of the Medny Stream, 44°16'34"N 145°56'27"E, 16.viii.2017, leg. K.Makarov L=5,45
Copyright © Kirill Makarov
Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Kinashir Isl., Dokuchaeva Ridge, solfatara field “Bolshoye”, h~400, 44°26'57"N 146°5'28"E, 8.viii.2017, leg. K.Makarov L=5.0 mm
Copyright © Kirill Makarov
Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., Dokuchaeva Ridge, solfatara field “Bolshoye”, h~400, 44°26'57"N 146°5'28"E, 8.viii.2017, leg. K.Makarov L=4,7 mm
Copyright © Kirill Makarov
Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Kinashir Isl., caldera of Golovnin Volcano, shore of Kipyashcheye Lake, 43°51'47"N 145°30'04"E, 24.vi.2008, leg. K.Makarov L=4,55 mm
Copyright © Kirill Makarov
Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., caldera of Golovnin Volcano, shore of Kipyashcheye Lake, 43°51'47"N 145°30'04"E, 24.vi.2008, leg. K.Makarov L=4,75
Copyright © Kirill Makarov
Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Kinashir Isl., caldera of Golovnin Volcano, solfatara field "Cherepakhovoye", 43°52'51"N 145°30'01"E, 28.vii.2011, leg. K.Makarov L=4,8 mm
Copyright © Kirill Makarov
Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., caldera of Golovnin Volcano, solfatara field "Cherepakhovoye", 43°52'51"N 145°30'01"E, 28.vii.2011, leg. K.Makarov L=4,8 mm
Copyright © Kirill Makarov
Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., Mendeleev Volcano, source of Kislyi Stream below the solfatara field, 43°59'46"N 145°45'54"E, 22.ix.2016, leg. Yu.Sundukov L=4,9 mm
Copyright © Kirill Makarov
Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., Mendeleev Volcano, source of Kislyi Stream below the solfatara field, 43°59'46"N 145°45'54"E, 22.ix.2016, leg. Yu.Sundukov L=4,8
Copyright © Kirill Makarov
Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Shikotan Isl., the Tserkovnaya Bay, 43°44'59"N 146°41'40"E, 29.vi-10.vii.2012, leg. Yu.Sundukov L=5,3 mm
Copyright © Kirill Makarov
Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Shikotan Isl., the Tserkovnaya Bay, 43°44'59"N 146°41'40"E, 29.vi-10.vii.2012, leg. Yu.Sundukov L=5,6 mm
Copyright © Kirill Makarov
Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Yuriy Isl., Shirokaya Bay, 43°25'33"N 146°04'08"E, 2.ix.2016, leg. Yu.Sundukov L=4,8 mm
Copyright © Kirill Makarov
Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Yuriy Isl., Shirokaya Bay, 43°25'33"N 146°04'08"E, 2.ix.2016, leg. Yu.Sundukov L=5 mm
Copyright © Kirill Makarov
Bembidion (Ocydromus) fontinale (Raffray, 1885)
Male adult
Ethiopia Amhara, Mt. Abuna yosef, n-slope, 3800-3950 m, 12°07’52”N 39°11’39”E, 4.iii.2019, leg. D. Hauth, J. Schmidt, Yeshitla M., Yitbarek, W. (CSCHM)
Copyright © Joachim Schmidt
Bembidion (Peryphus) baleense Toledano, Bonavita & Schmidt, 2021 Holotype
Male adult
Ethiopia, Oromia, Bale Mts., Sebsebe Washia Forest, Salgen Vall. 3130 m, 07°02’08”n 39°36’06”e, 4.ii.2019, leg. R.Emmerich, J. Schmidt, Yeshitla M. (CSCHM).
Copyright © Joachim Schmidt
Bembidion (Ocydromus) basistriatum Fairmaire, 1893
Male adult
Ethiopia, Oromia, Mt Enkuolo, NE-slope, 3100-3200m, 07°24’41”N 39°22’26”E, 5-7.Xii.16, leg. J. Schmidt (CSChM)
Copyright © Joachim Schmidt
Bembidion (Peryphus) damota Toledano, Bonavita & Schmidt, 2021 Holotype
Male adult
Ethiopia, Sodo, Mt. Damot, 2380 m, 06°53’26”N 37°46’51”E, 3.Xii.2017, leg. D. Hauth, J. Schmidt, Yeshitla M.” (CSCHM).
Copyright © Joachim Schmidt
Bembidion (Ocydromus) melanoceroides Toledano, Bonavita & Schmidt, 2021 Paratype
Male adult
Ethiopia, Oromia, Mt Enkuolo, N-slope, 3350-3600m, 07°22’48”N 39°21’07”E, 6-9.Xii.16, leg. J. Schmidt (CSCHM).
Copyright © Joachim Schmidt
Bembidion (Ocydromus) melanocerum Chaudoir, 1876
Male adult
Ethiopia, Amhara, W-slope Mt. Choke, 2750-2900 m, 10°38’N 37°45’E, 1.iii.2019, leg. D. Hauth, J. Schmidt, Yeshitla M., Yitbarek, W. (CSCHM)
Copyright © Joachim Schmidt
Bembidion (Peryphus) scottustulatum Netolitzky, 1937
Male adult
Ethiopia, Amhara, n-slope Mt. Choke, 3800-3950 m, 10°43’16”N 37°51’15”E, 26.ii.2019, leg. D. Hauth, J. Schmidt, Yeshitla M., Yitbarek, W. (CSChM)
Copyright © Joachim Schmidt
Odontium (Microserrullula) australe Bonavita & Vigna Taglianti, 2021 Paratype
Male adult
Kenya, Bushwackers Camp, Athi river Beach, 11-13 ii 1977, J C A van Etten. (Museum Leiden)
Copyright © Paolo Bonavita
Odontium (Microserrullula) clarkei Bonavita & Vigna Taglianti, 2021 Paratype
Male adult
Blue nile Gorge, 1100 m, 10.Xi.1971 / Coll. Mus. Tervuren, Ethiopie, r.o.S. Clarke / Musée royal de l’Afrique centrale Tervuren
Copyright © Paolo Bonavita
Odontium (Microserrullula) pogonopsis Alluaud, 1933
Male adult
Musee Du Congo, prés Bombesa [= Bambesa], Alipago – ii-1935, J. Leroy (MRAC)
Copyright © Paolo Bonavita
Odontium (Microserrullula) okavangum Bonavita & Vigna Taglianti, 2021 Paratype
Male adult
Angola, riv. Cuilo, Cacolo, Alto Cuilo, Vi-1954, 3694.3, E. Luna de Carvalho
Copyright © Paolo Bonavita
Odontium (Microserrullula) basilewskyi Bonavita & Vigna Taglianti, 2021 Paratype
Male adult
Congo Belge, Libenge, Zambi 6.1.1948, r. Cremer-M. neumann / r. i. Sc. nat. Belg. , i.G. 16.655 / Institut Royal des Sciences naturelles de Belgique, Bruxelles
Copyright © Paolo Bonavita
Odontium (Microserrullula) aegyptiacum Dejean, 1831
Male adult
Sudan, Wadi Hatta
Copyright © Paolo Bonavita
Odontium (Microserrullula) icterodes Alluaud, 1933
Male adult
Gantchou, 19.Vi.1927, A. Collart (RBINS)
Copyright © Paolo Bonavita
Anillinus albrittonorum Sokolov & Schnepp, 2021 Paratype
Female adult
6 mi NW High Springs, Columbia County, Florida. Image from Sokolov & Schnepp, 2021
Copyright © Igor Sokolov
Bembidion (Peryphanes) sanatum Bates, 1883: 274 Holotype
Male adult
“Type [round, bordered in red] // Japan / G. Lewis / 1910 – 310 [printed] // B. / sanatum / Bates [handwritten]” (NHMUK).
Copyright © Luca Toledano
Bembidion (Plataphus) pseudolucillum Netolitzky, 1938a: 37
Male adult
Kibune, Kyoto (NHMW)
Copyright © Luca Toledano
Bembidion (Peryphus) psuchrum Andrewes, 1922c: 241
Male adult
Inde, Himachal Pradesh, Mahari, 3500 m (CTVR)
Copyright © Luca Toledano
Tachys (Paratachys) terryli Liebherr, 2021 Paratype
Female adult
Kauai: Halelea F. R.: Namolokama Mtn., Waioli Str., litter, sift, ohia/ferns, 22°08.00'N, 159°29.85'W, 1340 m el., 21-v-2005, lot 1 Liebherr
Copyright © James K. Liebherr
Tachys (Paratachys) arcanicola Blackburn, 1878 Paralectotypes
Female adult
Oahu: Honolulu (Bates collection, box 389). Image from Liebherr, 2021
Copyright © James K. Liebherr
Tachys (Paratachys) perkinsi Liebherr, 2021 Holotype
Male adult
(NHMUK): platen mount / 170 on reverse // Hawaiian Is. / R.C.L. Perkins // Kaunakakai / sea level / vii-93 // atomarium (label upside down indicating misidentification) // HOLOTYPE ♀ / Paratachys / perkinsi / J. K. Liebherr 2020 (black-margined red label)
Copyright © James K. Liebherr
Tachys (Paratachys) haleakalae Liebherr, 2021 Holotype
Female adult
(CUIC): HI:Maui Haleakala N.P. / Kipahulu Vy. Central / Pali Tr. 910 m el. / 30-IV-1991 under logs // J.K. Liebherr / A.C. Medeiros / Jr. collectors // HOLOTYPE ♀ / Paratachys / haleakalae / J. K. Liebherr 2020.
Copyright © James K. Liebherr
Tachys (Paratachys) aaa Liebherr, 2021 Paratype
Male adult
Kau, Kona Mala, 21–xi–2017, lot HI00179
Copyright © James K. Liebherr
Bembidion (Ocydromus) terryerwini Neri & Toledano, 2021 Paratype
Male adult
N 29°21'14.7" E 57°20'31.7" / Iran, Prov. Kerman, Rāyen, Goroah, / Babtahoune, Bachufer-Schotterbank / 06.06.2014 2700 m / leg. Schnitter
Copyright © Luca Toledano
Bembidion (Ocydromus) hiekei Muller-Motzfeld, 1986b: 171 Paratype
Male adult
Copyright © Luca Toledano
Bembidion (Ocydromus) sokolowskii Fassati, 1957b: 154 Holotype
Male adult
MHNG
Copyright © Luca Toledano
Tachyta (Tachyta) umbrosa (Motschulsky, 1851)
Male adult
Thailand, Prov. Phang Nga, Khao Lak, Similana Resort, leg. U.Schmidt, 14.XI.2007; det. M.Baehr, 2008; Photo: U.Schmidt, 2021
Copyright © Udo Schmidt
Tachyta (Paratachyta) coracina (Putzeys, 1875)
Male adult
Malaysia, Langkawi, environment of Hotel Berjaya, leg. U.Schmidt, 7.-22.XI.2009; det. M.Baehr, 2010; Photo: U.Schmidt, 2021; 2.1mm
Copyright © Udo Schmidt
Tachyura (Tachyura) charactus (Andrewes, 1925)
Male adult
Nepal, Prov. Narayani, Sauraha, Rapti River, 180 m, leg. A. Skale, 18.IV.2000, det. M. Hartmann, 2000; Photo: U.Schmidt, 2021. 3.1 mm
Copyright © Udo Schmidt
Tachyura (Tachyura) politus (Motschulsky, 1851)
Female adult
Thailand, Prov. Phang Nga, Khao Lak, Similana Resort, leg. U.Schmidt, 14.XI.2007; det. M.Baehr, 2008; Photo: U.Schmidt, 2021; 2.5mm
Copyright © Udo Schmidt
Tachyura (Tachyura) ceylanicus (Nietner, 1858)
Female adult
Malaysia, Langkawi, environment of Hotel Berjaya, 7.-22.XI.2009 leg. U.Schmidt; det. M.Baehr, 2010; Photo: U.Schmidt, 2021; 2.1mm
Copyright © Udo Schmidt
Tachyura (Tachyura) poecilopterus (Bates, 1873)
Male adult
Thailand, Prov. Phang Nga, Khao Lak, Similana Resort 14.XI.2007 leg. U.Schmidt; det. M.Baehr, 2008; Photo: U.Schmidt, 2021; 2.4mm
Copyright © Udo Schmidt
Tachyura (Amaurotachys) notaphoides (Bates, 1886)
Female adult
Thailand, Prov. Phang Nga, Khao Lak, Similana Resort leg. U.Schmidt, 14.XI.2007; det. M.Baehr, 2008 as Tachys callispilotus. Photo: U.Schmidt, 2021; 2.3mm
Copyright © Udo Schmidt
Tachyura (Amaurotachys) holzschuhi Baehr, 2015 Holotype
Male adult
INDIA; Himalaya; U.P. Uttarkashi Distr., Gangani 14.-18.VI.1981, 1300 m leg. C. Holzschuh (NME).
Copyright © Martin Baehr
Tachyta (Eurytachyta) laticollis Baehr, 2016 Holotype
Female adult
Borneo, Sabah, Kinabalu NP, HQ vic., 1550 m, 22.–25.V.2005, R. GRIMM (CBM).
Copyright © Martin Baehr
Elaphropus (Tachyura) serrulipennis Baehr, 2016 Holotype
Male adult
Philippines, Leyte, visca N Baybay, cultiv. land, 1991, leg. SCHAWALLER & al. / 4.III.1991 (SMNS).
Copyright © Martin Baehr
Elaphropus (Tachyura) schawalleri Baehr, 2016 Holotype
Male adult
Philippines, Leyte, SW Abuyog, 28.II.1991, river bank, leg. SCHAWALLER & al. (SMNS).
Copyright © Martin Baehr
Tachyura (Tachyura) rufinus Baehr, 2016 Holotype
Male adult
Borneo, Brunei, Temburong Kuala Belalong, BORCHERDING leg. / 10.II.1995 HW KBFSC (CWB).
Copyright © Martin Baehr
Elaphropus (Tachyura) martensi Baehr, 2016 Paratype
Female adult
321 Ilam Distr., Gitang Khola Valley, Alnus forest along river, 1750 m, 11–13 Apr 1988, J. MARTENS & W. SCHAWALLER leg. / Nepal-Expeditionen JOCHEN MARTENS (CBM).
Copyright © Martin Baehr
Elaphropus (Tachyura) grimmi Baehr, 2016 Holotype
Male adult
Borneo, Malaysia, Sabah NE, Keningau, Bingkor, 380 m, 22.III.2013, R. GRIMM (CBM).
Copyright © Martin Baehr
Tachyta (Paratachyta) quadrinotata Baehr, 2016 Holotype
Female adult
320 Ilam Distr., betw. Hodia Khola Valley to Sktia, 250–500 m, Shorea forest, dry, 7 Apr 1988, MARTENS & SCHAWALLER leg. / Nepal-Expeditionen JOCHEN MARTENS (SMNS).
Copyright © Martin Baehr
Paratachys leytensis Baehr, 2016 Holotype
Female adult
Philippines, Leyte, visca N Baybay, 1991, sec. forest, 100–200 m, leg. SCHAWALLER & al. / 11.III.1991 (SMNS).
Copyright © Alexander Anichtchenko
Tachyta (Tachyta) quadriplagiata Baehr, 2014 Holotype
Male adult
S-VIETNAM 40km NW An Khe Buoan Luoi, 620-750m / 14°10’N 106°30’E 28.3.-12.4.1995 Pacholatko & Dembicky (NMW).
Copyright © Martin Baehr
Tachys (Tachys) federicae Baehr, 2016 Holotype
Female adult
QUEENSLAND: 27.295°S 152.749°E, Mt.Tenison Woods, 631 m, 29 Oct 2009, 19171, G.B. Monteith & F. Turco, sieved litter (QM Reg. No. QMT224214).
Copyright © Martin Baehr
Aenigmatachys rubens Baehr, 2020 Holotype
Male adult
“Australia Black B . . . (unreadable) / G.C. Champion Coll. B.M. 1927-409” (NHM).
Copyright © Martin Baehr
Bembidion (Bembidionetolitzkya) conforme Dejean, 1831
Male adult
W Ukraine, NE Carpathians, Tiachiv distr., h~400 m, bank of stream Mala Ugolka, 23.VI.2016, Panin R. leg.
Copyright © Ruslan Panin
Bembidion (Philochthus) neresheimeri J. Muller, 1929: 101
Male adult
W Ukraine, Lviv city, Sykhiv, waste-ground, 49°47'29"N 24°3'18"E, 7.IV.2018, Panin R. leg., L=3.1mm
Copyright © Denis Khrapov
Bembidion (Peryphanes) sanatum sanatum
Male adult
Japan, Honshu Island, Mount Hikageyama Jyoushu, Gunma Pref., 18.IX.1993 leg. S. Morita
Copyright © Kirill Makarov
Bembidion (Peryphanes) sanatum sanatum
Female adult
Japan, Honshu Island, Mount Hikageyama Jyoushu, Gunma Pref., 18.IX.1993 leg. S. Morita
Copyright © Kirill Makarov
Bembidion (Peryphanes) sanatum iwanai Makarov & Sundukov, 2021 Holotype
Male adult
Kunashir Island, valley of Kislaya River, below hot springs, 18.VI.2011, leg. A. Matalin
Copyright © Kirill Makarov
Bembidion (Peryphanes) sanatum iwanai Makarov & Sundukov, 2021 Paratype
Female adult
Kunashir Island, valley of Kislaya River, below hot springs, 18.VI.2011, leg. A. Matalin
Copyright © Kirill Makarov
Tachys (Paratachys) frischi Coulon & Felix, 2009
Female adult
Iran, Khuzestan, 30 km E Shushtar, 5.5.2007, A. Anichtchenko leg. (cAA). det. J. Muilwijk, 2021
Copyright © Alexander Anichtchenko
Tachys (Polyderis) algiricus (Lucas, 1846)
Male adult
Iran, Chaharmahal and Bakhtiari, 10 km W Naghan, 3.iv.2007; Muilwijk leg. & det. confirmed by J. Coulon.
Copyright © Jan Muilwijk
Tachys (Polyderis) impressipennis Motschulsky, 1859
Male adult
Iran, Iran, Hormozgan, Minab, 19–20.v.1973, Loc. no 203; Exp. Nat. Mus. Praha. Confirmed by J. Coulon.
Copyright © Jan Muilwijk
Porotachys ottomanus ottomanus Schweiger, 1968
Male adult
Iran, Kohgiluyeh and Boyer-Ahmad, Suq cave, 812 m, 30°49ˊ25.9´´ N, 50° 25ˊ37.9´´ E, 16.vi.2018, S. Rahideh & J. Muilwijk leg
Copyright © Jan Muilwijk
Tachyura (Tachyura) conspicua (Schaum, 1863)
Male adult
Hormozgan, Kuh-e- Geno, 600–1000 m, 27–28.iv.1977, Loc. no. 318, Exp. Nat. Mus. Praha. det. Muilwijk; confirmed by J. Coulon
Copyright © Jan Muilwijk
Tachyura (Tachyura) shahinei (Schatzmayr & Koch, 1934)
Male adult
Iran, Sistan and Baluchestan, 13 km SSE Nikshahr riv., 8–9.iv.1973, Loc. no. 152, Exp. Nat. Mus. Praha. Muilwijk det.
Copyright © Jan Muilwijk
Bembidion (Emphanes) latiplaga latiplaga Chaudoir, 1850
Female adult
NW-Ukraine, Volhynia, Turiysk env., 17-26.VI.2016, to light, Voytko P. leg./ Panin R. det., L=3.1 mm
Copyright © Denis Khrapov
Bembidion (Bembidion) humerale humerale Sturm, 1825: 176
Female adult
W Ukraine, Lviv city, Landscape Reserve “Peatbog Bilogorshcha” NW part, 17.IV-10.VII.2018, Panin R. leg., L =2.95 mm
Copyright © Denis Khrapov
Tachys (Paratachys) fulvicollis (Dejean, 1831)
Female adult
NW-Ukraine, Volhynia, Turiysk env ., into the light, 6.VII.2015, Voytko P. leg./ R.Panin det., L=3,3 мм
Copyright © Denis Khrapov
Sinechostictus (Pseudolimnaeum) inustus (Jacquelin du Val, 1857)
Female adult
Fiume Imagna, Bg, Lombardia, Italy
Copyright © Vittorino Monzini
Bembidion (Peryphanes) milleri carpathicum J. Muller, 1918b: 88
Male adult
W Ukraine, W Podolian Upland, Rogatyn Opillia, Zaluzhzhya vill, 49°24'33"N 24°35'36"E, clay quarry, 30-31.VIII.2018, Panin R leg., L=4.6mm
Copyright © Denis Khrapov
Bembidion (Euperyphus) scapulare lomnickii Netolitzky, 1916: 260
Female adult
W Ukraine, NE Carpathians, Lviv reg., near Rozgirche vill., bank of Stryi river, 11.V.2014, Khrapov D. leg./det.Panin R., L= 5.4 mm
Copyright © Denis Khrapov
Bembidion (Peryphanes) cf brunnicorne Dejean, 1831
Male adult
W.Ukraine, Transcarpathian, Polonynian Beskids, Yavirnyk Rng, h=600 m, near Ruskyi Mochar vill.,15.V.2021, V. Khrapova leg./R. Panin det., L=4.7 mm
Copyright © Denis Khrapov
Scotodipnus (Scotodipnus) glaber (Baudi di Selve, 1869)
Male adult
Monte Chiappo, PV, Italia
Copyright © Vittorino Monzini
Tachyura (Tachyura) yuezhi Kabak, 2021 Holotype
Male adult
China, Xinjiang, Ruoqiang River, Ruoqiang (= Charklik) Town, 39°00ʹ59ʺN / 88°09ʹ24ʺE, H = 820 m, 19.07.2015, Kabak I.I. leg.
Copyright © Ilya Kabak
Bembidion (Euperyphus) scapulare lomnickii Netolitzky, 1916: 260
Male adult
W-Ukraine, Transcarpathian region, Vynogradiv distr., bank of Tisza river near Mala Kopanya vill., 5-7.VI.2018, Panin.R leg., L=5,8 mm
Copyright © Denis Khrapov
Elaphropus (Elaphropus) globulus (Dejean, 1831)
Male adult
Spain, Granada, rio Darro, barranco de Teatino, 20.V.2005, Anichtchenko A. leg & det. (cAA). 1.8 mm
Copyright © Alexander Anichtchenko
Tachys (Polyderis) truncatus (Nietner, 1858)
Male adult
India, South Goa, Distr. Canacona, Raja Baga Beach, on light leg. U.Schmidt, 16.-29.XI.2010; det. M.Hartmann. Photo: U.Schmidt, 2022 1.3 mm
Copyright © Udo Schmidt
Costitachys inusitatus Erwin, 1974 Holotype
Male adult
Santarem, Para, Brazil, IV.27.1963 FG Werner (MCZ)
Copyright © - - -
Bembidion (Ocyturanes) montislunae Neri & Toledano, 2022 Holotype
Male adult
Oman, Dhofer province / Jabal al Qamar / 20 km N Dhalqut / N 16.71092° E 53.15350° / (1100 m.n.m.) 28.8.2012 / lgt. P. Kučera (CTVR). Image from Paolo Neri & Luca Toledano, 2022
Copyright © - - -
Microtyphlus saxatilis Magrini, Leo & Fancello, 2004
Male adult
Italia, Sardegna, Perda Longa (NU) Baunei.
Copyright © Augusto Degiovanni
Rhegmatobius bastianinii Magrini & Casale, 2015 Paratype
Male adult
Italia, Lazio (LT) M.te S. Angelo, Formia, 1200m.
Copyright © Augusto Degiovanni
Bembidion (Princidium) punctulatum Drapiez, 1820
Male adult
Pechoro-Ilychskii N.P. (Печеро-Илычский зап)
Copyright © Kirill Makarov
Pericompsus (Pericompsus) anassa Erwin, 1974
Female adult
Polvaredas, partido de Saladillo, provincia de Buenos Aires I-2000
Copyright © Armando Cicchino
Tachys (Tachys) argentinicus Csiki, 1928
Female adult
Mar Chiquita, Partido de Mar Chiquita, provincia de Buenos Aires, XII-2010
Copyright © Armando Cicchino
Tachys (Paratachys) bonariensis Steinheil, 1869
Female adult
San Fernando, provincia de Buenos Aires XI-2015
Copyright © Armando Cicchino
Tachys (Paratachys) laevigatus (Boheman, 1858)
Female adult
San Fernando, provincia de Buenos Aires XI-2015
Copyright © Armando Cicchino
Bembidion (Notaphus) brownorum Maddison, Sproul & Will, 2023 Holotype
Male adult
EMEC: 39.13841/−122.34621 USA: California: Colusa Co. Antelope Valley, Freshwater Creek uv light pan trap 133 m. 1.vii.2021 K.Will
Copyright © David Maddison
Bembidion (Notaphus) callens Casey, 1918 Paralectotypes
Female adult
Tuçson [Pima County], Arizona.
Copyright © David Maddison
Bembidion (Liocosmius) cachagua Maddison, 2023 Holotype
Male adult
CAS: “U.S.A.: CALIFORNIA: Monterey Co., Hastings Nat. History Reserve. J.M. Linsdale Colln.”, “C.A.S. Accession 29-V-48 Cachagua Creek”, “David R. Maddison DNA3842 DNA Voucher”
Copyright © David Maddison
Nesamblyops townsendi Sokolov, 2023
Male adult
Tennyson Inlet, Marlborough Sounds, SO
Copyright © Igor Sokolov
Nesamblyops tararua Sokolov, 2023
Male adult
Judd Ridge, Tararua Range, Wellington, NO
Copyright © Igor Sokolov
Nesamblyops parvulus Sokolov, 2023 Paratype
Female adult
Mount Stokes, Marlborough Sounds, SO
Copyright © Igor Sokolov
Odontium (Odontium) subfusum Darlington, 1959
Female adult
Philippines, Mindanao, Mati, Surigao-Davao coastal rd., 6.96N 126.29E, river bank, 28.4.2023, Anichtchenko A. leg & det.
Copyright © Alexander Anichtchenko
Tachyura (Tachyura) lucasi (Jacquelin da Val, 1852)
Female adult
Gambia, Serrekunda, environment Senegambia Beach Hotel, leg. U.Schmidt, 16.-22.XI.1990; det. M. Baehr, 2017; 2.5 mm.
Copyright © Udo Schmidt
Pericompsus (Pericompsus) catamarcensis Roig-Juñent & Scheibler, 2004 Paratype
Male adult
Image from: Córdoba, Silvia Patricia, Sánchez, Francisco Rolando & Bezdjian, Laura Patricia, 2023, Types of Carabidae (Insecta: Coleoptera) deposited at the entomological collection of Instituto- Fundación Miguel Lillo, Tucumán, Argentina
Copyright © - - -
Ocys andreae (Jeannel, 1937)
Female adult
Spain, Andalusia, Sierra Nevada, pico Veleta, 2500-2800m, 29-30.9.2001 Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko
Anillinus bexarensis Sokolov, 2022 Paratype
Female adult
Up the Creek Cave, Bexar County, Texas ; 1.7 mm
Copyright © Igor Sokolov
Anillinus reddelli Sokolov, 2022 Paratype
Female adult
Beck Horse Cave, Williamson County, Texas
Copyright © Igor Sokolov
Bembidion (Hirmoplataphus) pliculatum Bates, 1883
Male adult
S Sakhalin isl., Susuya riv. leg & det. Sundukov
Copyright © Kirill Makarov
Bembidion (Hirmoplataphus) hirmocaelum Chaudoir, 1850
Male adult
Primorski reg., Kedrovaya riv. det. Sundukov, 2023
Copyright © Kirill Makarov
Medusapyga hehalis LaBonte, 2023 Paratype
Female adult
WASHINGTON, Thurston Co., 11.3 km NW Littlerock, elev. ~400 m, D.J. Cox and J.R. LaBonte, 6 May 2007, 46.9600°N 123.13450°W. Image from LaBonte & Maddison, 2023
Copyright © - - -
Medusapyga alsea LaBonte, 2023 Paratype
Female adult
“OREGON, Benton Co., Prairie Peak, 11 km S Alsea, J.R. LaBonte, 9 V 1993, elev. 810 m, 44.2844°N 123.5858°W, under stones". Image from LaBonte & Maddison, 2023
Copyright © - - -
Anillaspis explanata (Horn, 1888)
Female adult
USA: California: El Dorado Co., Alabaster Cave (CMNH). Scale bar 1 mm; Image from LaBonte & Maddison, 2023
Copyright © - - -
Amerizus (Tiruka) janatai Deuve, 2023 Holotype
Female adult
Chine, Sichuan, Maoxian, Jiuding Shan, 3000 m, 31°40’03”N/103°42’33”E (M. Janata), in coll. M. Janata, Prague (République Tchèque).
Copyright © Thierry Deuve
Bembidion (Euperyphus) eques Sturm, 1825
Female adult
Italy, Moggio Udinese, Torrente Aupa, 6.9.2023, 13.2007° E, 46.4084° N, J. Gebert leg.
Copyright © Jörg Gebert
Bembidion (Omoperyphus) hypocrita Dejean, 1831
Male adult
Spain, Jaen prov., Arroyo de la piedra del Aguila, 37.683885, -3.485062, 3-4.5.2014, Anichtchenko A. leg & det. (cAA)
Copyright © Alexander Anichtchenko
Horologion speokoites Valentine, 1932 Holotype
Male adult
"Arbuckle's Cave Maxwelton, W.Va. July 12 1931 J.M. Valentine / On bit of wet decaying wood, muddy floor of lower level / Type gen. Type sp. / U.S.N.M. Type 44255"
Copyright © Curt Harden
Horologion hubbardi Harden & Davidson, 2024 Paratype
Female adult
CWH-452 (VMNH).
Copyright © Curt Harden
Elaphropus (Barytachys) ferrugineus (Dejean, 1831)
Female adult
Alabama, Blount Co., Highland Lake. 9.iv.2019. T. King leg. Soil berlese
Copyright © Curt Harden
Tachys (Paratachys) austinicus Casey, 1918
Male adult
USA: Alabama, Blount Co., Highland Lake, 11.ix.2020, T. King leg. Window trap
Copyright © Curt Harden
Tachys (Paratachys) albipes Leconte, 1863
Male adult
"USA: Florida, Columbia Co., Osceola National Forest, Range Rd., 1.ii.2023, C.W. Harden leg., sifted from litter." ca. 3mm
Copyright © Curt Harden
Tachys (Paratachys) sagax Casey, 1918
Male adult
"USA: Alabama, Blount Co., Highland Lake, 24.viii.2020, T. King leg., window trap"
Copyright © Curt Harden
Tachys (Paratachys) ventricosus Leconte, 1863
Female adult
"USA: Virginia, Virginia Beach City, First Landing State Park, 18.iv.2018, C.W. Harden leg., sifting litter"
Copyright © Curt Harden
Micratopus anaisae Lemaire, 2021 Holotype
Female adult
Saint-Félix, commune du Gosier, Guadeloupe, 16.210 N, 61.461 O, J. M. Lemaire leg. 15/04/2016. (MNHN). Image from Lemaire, 2021
Copyright © - - -
Tachyura (Tachyura) lucasi (Jacquelin da Val, 1852)
Female adult
Senegal, M`Bour, 14 km south, Club Aldiana, leg. U.Schmidt, 8.-20.X.1992; det. R.F.F.L. Felix; 2.5 mm
Copyright © Udo Schmidt
Tachyura (Tachyura) imitans (Peringuey, 1896)
Female adult
South Africa, Mpumalanga, White River, leg. A. Skale, 31.V.2001; det. R.F.F.L. Felix; 3.2 mm
Copyright © Udo Schmidt
Bembidion (Armatocillenus) sauteri (Jedlicka, 1958) Holotype
Female adult
Formosa, Sauter // Anping, 1909.X. // Typus [red] // Armatocillenus sauteri n.sp. [handwritten], det. ING. JEDLIČKA [pink] (HNHM).
Copyright © - - -
Bembidion (Desarmatocillenus) foochowense Lindroth, 1980
Male adult
TAIWAN: Lunglungchi, Fulung, Taipei Hsien, 31.vii.2001, Terada lgt.
Copyright © - - -
Odontium (Odontium) striatum (Fabricius, 1792)
Female adult
Latvia, Dunava
Copyright © Alexander Anichtchenko
Odontium (Bracteon) lapponicum Zetterstedt, 1828
Female adult
Irkutsk reg., Usolskii distr., Malaya Belaya riv., Mutovka vill., 12-15.7.1996, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko
Odontium (Bracteon) conicolle Motschulsky, 1844: 273
Female adult
Burjatia, Barguzin depr., Alla riv., 8.8.1997, Anichtchenko A. leg & det.
Copyright © Alexander Anichtchenko
Megastylulus pivai Giachino & Sciaky, 2003 Holotype
Male adult
leg. E. Piva - Isl. St.Lucia Caraibi
Copyright © Vittorino Monzini
Bembidion (Nepha) ibericum Piochard de la Brulerie, 1868
Male adult
Spain, Sierra de Guadarrama, Laguna de los Cleveles, 6.VIII.2024, 2000m, Anichtchenko A. leg & det. (cAA).
Copyright © Alexander Anichtchenko
Bembidion (Nepha) genei trinacriae (Bonavita & Vigna-Taglianti, 2010)
Female adult
S Spain, Sierra de las Nieves, Los Quejigales 6-8.4.2010, Anichtchenko A. leg & det. (cAA)
Copyright © Alexander Anichtchenko
Ocys reuteri Toledano & Wrase, 2016 Holotype
Female adult
“N-Iraq, S Rawanduz/ XI.2007 – III.2008 / 36°30’N, 44°36’E / 1400m, leg. C. Reuter” // Ocys reuteri n.sp./ Toledano & Wrase des. 2016” (CWBE).
Copyright © Luca Toledano
Ocys hiekei Toledano & Wrasei, 2016 Paratype
Male adult
“CHINA (Nei Mongol Aut. Reg.) / Helan Shan / above Guangzong temple / 38°39’14.8”N/105°50’00.9”E / 2575m (Picea forest, litter sifted) / 22.VI.2011 D.W. Wrase
Copyright © Luca Toledano
Stylulus (Stylulus) isabelae Giachino & Sciaky, 2003 Holotype
Male adult
Copyright © Vittorino Monzini
Bembidion (Ocyturanes) ivanloebli Neri & Toledano, 2018 Holotype
Male adult
“Iran, Fars, Sepidan, 21-9-2005, Muilwijk leg.” (CTVR)
Copyright © Luca Toledano
Bembidion (Ocyturanes) waziristanum Andrewes, 1932c: 863
Male adult
Pakistan, W. Himalai mts., 20 km SW Chilez (CTVR)
Copyright © Luca Toledano
Bembidion (Ocyturanes) lobanovi Mikhailov, 1984b: 297
Female adult
N Tajikistan, Fanskie Mts. Khurdak (CTVR). Scale: 1 mm
Copyright © Luca Toledano
Bembidion (Ocyturanes) signatipenne Jacquelin du Val, 1852
Male adult
Imago: NW Turkey, Edremit, Kaz Dağ, Ay i Deresi (PN); Aedeagus specimen from Crete, Rethimno, Plakias, river Megalopotamos (PN).
Copyright © Luca Toledano
Bembidion (Ocyturanes) viduum Netolitzky, 1910a: 215 Holotype
Male adult
Kilikisch. Taurus (NHMW). Scale: 1 mm.
Copyright © Luca Toledano
Bembidion (Ocyturanes) kurdistanicum Netolitzky, 1920
Male adult
Imago: Iran, Mazandaran Prov., Karaj-Chalus road (MNB); Aedeagus: Holotype, Persia Septentr., Kerim (NHMW).
Copyright © Luca Toledano
Bembidion (Ocyturanes) heinzi Korge, 1971: 16 Paratype
Male adult
Anatolia bor., Köse Flußufer (MNB). Aedeagus of Holotype, Anatolia bor., Köse Flußufer (MNB).
Copyright © Luca Toledano
Bembidion (Ocyturanes) antennarium (Morvan, 1972) Holotype
Male adult
Iran, Zagros, Kurang (PM). Scale 1mm
Copyright © Luca Toledano
Bembidion (Ocyturanes) mirzayani (Morvan, 1973) Paratype
Female adult
Iran, Zagros Mts., Sisakht (PM). Scale: 1 mm.
Copyright © Luca Toledano
Bembidion (Peryphus) tricuspis Neri & Toledano, 2018 Paratype
Male adult
“SW Iran, Hoseiniyeh / 28 km NNW Andimeshk / 12-13. 4. 1977, 360 m // Loc. no. 286 / Exped Nat Mus / Praha” (NMPC).
Copyright © Luca Toledano
Bembidion (Peryphidium) validum salangense Neri & Toledano, 2021 Paratype
Male adult
“Afghanistan / Lac du Salang 3700 m / 4.8.74 Leg. Ledoux”
Copyright © Luca Toledano
Bembidion (Omotaphus) mixtum Schaum, 1863b: 90 Lectotype
Male adult
43101 [handwritten] // mixtum Sch. [handwritten] / dt. Netolitzky [printed] // Hist. -Coll. (Coleoptera) / Nr. 43101 (=55133) / Bembidion mixtum Klug / Aegypt. Ehrenberg / Zool. Mus. Berlin [blue, printed] // SYNTYPE / Bembidium mixtum / Schaum, 1863 / labelled by MFNB 2015 [red, printed] (MNB).
Copyright © Luca Toledano
Bembidion (Sakagutia) umi Sasakawa, 2007
Male adult
S. Korea, Gyeongsanbuk-do, Pohang-si, collected by digging the pebbles near the estuary
Copyright © Dooyoung Kim
Bembidion (Omotaphus) kivuanum (Basilewsky, 1951) Holotype
Male adult
Imago - Holotype: “HOLOTYPUS [red, printed] // édéage en preparation microscop. [hand�written] // I.R.S.A.C. – MUS. Congo, Uvira: Bassin Ulindi / 2850m. 29-IX-1950 / N. Leleup [printed] / (Bord d’un marais) [handwritten] // RMCA ENT / 000019230 “ (MRAC); Aedeagus - paratype.
Copyright © Paolo Bonavita
Bembidion (Omotaphus) kilimanum Alluaud, 1908 Holotype
Male adult
“AFRIQUE ORle ALLEMANDE / KILIMANDJARO / (ZONE DES PRAIRIES) / CH. ALLUAUD I-IV 1904 [white, printed] // TYPE [red, printed] // MUSEUM PARIS / Coll. Ch. ALLUAUD [blue, printed] // Bembid. / (Testediolum / kilimanum / n.sp. Alluaud / [white, handwritten]” (MNHN).
Copyright © Paolo Bonavita
Bembidion (Omotaphus) demeyeri Toledano & Bonavita, 2016 Holotype
Male adult
“Uganda Occidental / Province D’UNYORO / Rég. de l’Albert-Nyanza / CH. ALLUAUD II 1909 [white, printed] // MUSEUM PARIS / Coll. Ch. ALLUAUD [blue, printed] // Bembid. (Notaphus) / Bakeri / type. Alluaud / Alluaud détérm [white, handwritten, with a blue strip on top]” (MNHN)
Copyright © Paolo Bonavita
Bembidion (Omotaphus) gorilla Bonavita, Toledano & Vigna Taglianti, 2016 Holotype
Male adult
? Holotype. “RWERU: 2800/2900M, 20 - VI – 1964, Tamisage, Bambousaise // Coll. Mus. Tervuren, Zaire: P. N. Virunga, Volcan Mikeno, R. P. Celis VI-64” (MRAC)
Copyright © Paolo Bonavita
Bembidion (Omotaphus) mellissii Wollaston, 1869
Male adult
Image from Bonavita et al. 2016, with modifications
Copyright © Paolo Bonavita
Bembidion (Omotaphus) uhligi Toledano & Maddison, 2016 Holotype
Male adult
Holotype? RSA-Western Cape, NE of Botrivier env., 34°13’S, 19°12’E, 24.II.2014 R. Kmeco leg.
Copyright © Paolo Bonavita
Tachys (Tachys) shirazi Jedlicka, 1968 Holotype
Male adult
Iran, Fars Province, Maharlou Lake, a salt lake in south of Shiraz, 18.III.1965 (MNHM)
Copyright © Jan Muilwijk
Tachys (Tachys) centralis J.Sahlberg, 1900
Male adult
Iran, Queshm Island. Image from Muilwijk et al, 2019
Copyright © - - -
Tachys (Tachys) torretassoi Schatzmayr & Koch, 1934
Male adult
Iran, Hormozgan Province, Queshm Isl., Tola, W. Qeshm, 5.III.2015, Wrase & Laser. Image from Muilwijk et al, 2019
Copyright © - - -
Tachyura (Tachyphanes) sinepuncta (Coulon, 2016) Holotype
Male adult
“N.O. Thailand, Khon Kaen, lux, 25-2-1981, leg. Dr. S. Saowakontha / Tachyura (Tachyphanes) lata (Peyron, 1858), Det. T. Kopecký 2011 / Sphaerotachys sinepunctis n. sp., J. Coulon det. 2016 / Holotype [étiquette rouge]” (MNHUB).
Copyright © - - -
Tachyura (Tachyphanes) lata (Peyron, 1858)
Male adult
N.O. Thailand, Khon Kaen. Image from Coulon, 2016
Copyright © - - -
Sinechostictus (Sinechostictus) millerianus (Heyden, 1883)
Male adult
CZECH REPUBLIC - Bystřice nad Olší, 26.VIII.2021, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar
Bembidion (Bembidionetolitzkya) varicolor varicolor
Male adult
CZECH REPUBLIC - Bystřice nad Olší, 15.III.2020, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar
Bembidion (Euperyphus) fluviatile fluviatile Dejean, 1831: 113
Male adult
CZECH REPUBLIC - Polanka nad Odrou, 6.IV.2017, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar
Bembidion (Notaphus) varium (Olivier, 1795)
Female adult
CZECH REPUBLIC - Olomouc dist., Skalička, 8.IV.2020, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar
Bembidion (Ocydromus) decorum decorum (Panzer, 1799b)
Male adult
CZECH REPUBLIC - Staré Hamry, 22.IX.2020, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar
Bembidion (Ocydromus) modestum (Fabricius, 1801)
Male adult
CZECH REPUBLIC - Olomouc dist., Skalička, 11.VI.2020, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar
Bembidion (Peryphanes) deletum deletum Audinet-Serville, 1821: 77
Male adult
CZECH REPUBLIC - Beskydy Mts., Kněhyně, 27.VI.2020, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar
Bembidion (Peryphanes) stephensii Crotch, 1866
Female adult
CZECH REPUBLIC - Moravian-Silesian reg., Opava dist., Závada, 25.VI.2020, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar
Bembidion (Peryphiolus) monticola monticola Sturm, 1825: 135
Male adult
CZECH REPUBLIC - Frýdek-Místek dist., Staré Hamry, 22.IX.2020, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar
Bembidion (Peryphus) subcostatum vau Netolitzky, 1913: 213
Male adult
CZECH REPUBLIC - Olomouc reg., Přerov dist., Skalička, 8.IV.2020, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar
Tachyura (Tachyura) stevensi (Andrewes, 1925)
Female adult
India, Uttarakhand, 5 km SW Rishikesh, left bank of Ganga riv., 9.4.2012, Anichtchenko A. leg. (cAA). det. M. Baehr
Copyright © Alexander Anichtchenko
Tachyura (Tachyura) pulcher pulcher
Male adult
Malaysia, Borneo, Sarawak, Miri div., Bario 10km NE, Pa Lungan, 7.3.2014 (cAA). det. M. Baehr
Copyright © Alexander Anichtchenko
Tachyura (Tachyura) thoracicus (Kolenati, 1845)
Female adult
India, Uttarakhand, 14 km E Uttarkashi, 1500m, 10-12.4.2012, Anichtchenko A. leg. (cAA). det. M. Baehr as octostriata (Netolitzky, 1929)
Copyright © Alexander Anichtchenko
Tachyura (Tachyura) aeneus (Putzeys, 1875)
Female adult
Indonesia, W New Guinea, Doberai Peninsula, Ayamaru vill., Ayamaru lakeside, 3.9.2015 (cAA). det. M. Baehr
Copyright © Alexander Anichtchenko
Tachyura (Tachyura) championi (Andrewes, 1925)
Female adult
India, Uttarakhand, 5km SW Rishikesh, left bank of Ganga riv., 9.4.2012 (cAA). det. M. Baehr
Copyright © Alexander Anichtchenko
Tachyura (Tachyura) vixmaculatus (Andrewes, 1925)
Female adult
Laos, Vientiane prov., Vang Vieng, 6km N Pakpo, Nam Song riv., 27.2.2014 (cAA). det. M. Baehr
Copyright © Alexander Anichtchenko
Tachyura (Tachyura) tagax (Andrewes, 1925)
Female adult
India, Uttarakhand, 18-20.4.2012, 15km SW New Tehri, 869m, Anichtchenko A. leg. (cAA). det. M. Baehr
Copyright © Alexander Anichtchenko
Bembidion (Ocyturanes) dudichi Csiki, 1928
Male adult
Morocco, Great Atlas mts, 2200-2300m Oukaimeden env., 16.6.2010, Anichtchenko A. leg. (cAA). det. M. Toribio
Copyright © Alexander Anichtchenko
Bembidion (Bembidionetolitzkya) geniculatum claudejeannei Marggi in Löbl & Smetana, 2003
Male adult
Spain, Sierra Nevada, rio Lanjaron, 1200m, 29.3.2010, Anichtchenko A. leg. (cAA). 5 mm; det. M. Toribio
Copyright © Alexander Anichtchenko
Bembidion (Paraprincidium) ruficolle (Panzer, 1796)
Female adult
Latvia, Jekabpils, Dunava 1.7.2006 (DUBC)
Copyright © Alexander Anichtchenko
Liotachys antennatus Bates, 1871
Female adult
Bolivia: Santa Cruz Prov. Andrez Ibanez Potrerillo de Guenda, Terebinto. / 3-4 October 1994 / leg. P. Bettella / det. R. Davidson. Carnegie Museum of Natural History, Pittsburg, PA, USA
Copyright © Curt Harden
Anillinus alleni Sokolov & Carlton, 2017
Male adult
USA: Arkansas, Stone Co., Ozark National Forest, Blanchard Springs Rec. Area. 27 September 2024. C.W. Harden & K.D. Karns leg. Under rock during rain. / Det. C.W. Harden 2024.
Copyright © Curt Harden
Bembidion (Testedium) laetum Brulle, 1836
Female adult
Morocco 65 km N Essaouria, 5 km N Jamaa Shoim 31°57'02"N 009°25'25"E 22-24.III.2011 leg. A.Gusakov. L= 4.0mm
Copyright © Kirill Makarov
Bembidion (Testedium) laetum Brulle, 1836
Male adult
Morocco 65 km N Essaouria, 5 km N Jamaa Shoim 31°57'02"N 009°25'25"E 22-24.III.2011 leg. A.Gusakov. L= 3.9 mm
Copyright © Kirill Makarov
Bembidion (Philochthus) vicinum Lucas, 1846
Male adult
Morocco 65 km N Essaouria, 5 km N Jamaa Shoim 31°57'02"N 009°25'25"E 22-24.III.2011 leg. A.Gusakov. L= 3.7 mm
Copyright © Kirill Makarov
Bembidion (Philochthus) vicinum Lucas, 1846: 86
Female adult
Morocco 65 km N Essaouria, 5 km N Jamaa Shoim 31°57'02"N 009°25'25"E 22-24.III.2011 leg. A.Gusakov. L= 4.0 mm
Copyright © Kirill Makarov
Rhegmatobius strictus (Baudi di Selva, 1891)
Male adult
Sardegna, Gennargentu Mte., Spada Mt., 1000, 27.4.2003 A. Degiovanni leg. (cAA).
Copyright © Alexander Anichtchenko
Bembidion (Ecuadion) agonoides Vigna-Taglianti & Toledano, 2008
Female adult
Ecuador, Provincia Cotopaxi, Otonga Nature Reserve, 0°25’15” S, 79°00’23” W, 1800-2000 m
Copyright © Pierre Moret
Bembidion (Ecuadion) walterrossii Toledano, 2008: 94
Female adult
Ecuador, Provincia Cotopaxi, Otonga Nature Reserve, 0°25’15” S, 79°00’23” W, 1800-2000 m
Copyright © Pierre Moret
Bembidion (Peryphus) andreae (Fabricius, 1787)
Female adult
Italy, Tagliamento, Ronchis, iv.2024, leg. Jörg Gebert
Copyright © Jörg Gebert
Bembidion (Euperyphus) scapulare tergluense Netolitzky, 1918
Male adult
Italy, Fiume Tagliamento, Cornino, iv.2024, leg. Jörg Gebert
Copyright © Jörg Gebert
Bembidion (Bembidionetolitzkya) coeruleum Audinet-Serville, 1821
Male adult
Marocco c., Asni, National Park Toubkal
Copyright © Paolo Bonavita
Bembidion (Bembidionetolitzkya) toubkalense Bonavita & Vigna Taglianti, 2018 Paratype
Male adult
Morocco, Toubkal
Copyright © Paolo Bonavita
Bembidion (Bembidionetolitzkya) scelio (Antoine, 1945)
Male adult
Maroc, Haut-Atlas, Gorges du Todra 1500 m
Copyright © Paolo Bonavita
Bembidion (Bembidionetolitzkya) cassolai (Bonavita & Vigna Taglianti, 1993)
Male adult
Italy, Sardegna, Medio Campedano, Gonnosfanadiga, 1.5km SW Riu Piras. leg & det. J. Gebert
Copyright © Jörg Gebert
Bembidion (Asioperyphus) jorgeberti Neri & Toledano, 2024 Paratype
Male adult
Burjatia, East Sajan Mts, Black Irkut valley, mouth of Susar riv., 1860m, 30.6.2012, leg J. Gebert
Copyright © Jörg Gebert
Peryphus conformis Motschulsky, 1844 Lectotype
Male adult
Mt. Hamar-Dab. (ZMU)
Copyright © Kirill Makarov
Bembidion (Peryphus) bualei polonicum J. Muller, 1930
Male adult
Ahrenshoop, Steilufer s. Ort 12.4047E, 54.3726N, 5.2025, leg. & det. J. Gebert
Copyright © Jörg Gebert
Bembidion (Peryphus) subflavescens Antoine, 1944
Male adult
Morocco, Souss-Massa, Draa, Zagora, Oued Draa riv., 1.2.2013 (cAA). det. M. Toribio 2017
Copyright © Alexander Anichtchenko
Bembidion (Peryphus) maritimum (Stephens, 1839)
Female adult
Spain, Guipuzcoa, Playaundi, 20.3.2006, Anichtchenko A. leg & det. (cAA)
Copyright © Alexander Anichtchenko
Bembidion (Peryphus) femoratum femoratum Sturm, 1825
Female adult
Latvia, Jekabpils distr, Dunava, 23.7.2008 (DUBC)
Copyright © Alexander Anichtchenko
Bembidion (Peryphus) petrosum petrosum
Male adult
E Siberia, Lake Baikal, Baikalo-Lenskii NP., Bereg Buryh Medvedei, Pokoiniki, 27.6.1995, Anichtchenko A. leg & det. (cAA)
Copyright © Alexander Anichtchenko
Bembidion (Semicampa) gilvipes Sturm, 1825
Female adult
Latvia, Daugavpils loki, Daugava 29.4.2008 (DUBC)
Copyright © Alexander Anichtchenko
Bembidion (Emphanes) normannum normannum Dejean, 1831
Male adult
Spain, Guipuzcoa, Playaundi, 20.3.2006, Anichtchenko A. leg. (cAA)
Copyright © Alexander Anichtchenko
Bembidion (Ocydromus) loebli Schmidt, 2014 Holotype
Male adult
Holotype male, with label data „INDIA Madras / Nilgiri 2100 m / Ootacamund - Pykara / 21.XI.72 / Besuchet Löbl Mussard“ in MHNG
Copyright © Qin Yuyao
Bembidion (Ocydromus) salweenum Schmidt, 2014 Holotype
Male adult
Holotype male, with label data “STIBET 26-29.IX.2010 / Kema SSE Naqu 4450 m / Salween River, banks, alpine / steppe, 31°16’34N 92°06’20E” in ZSM
Copyright © Qin Yuyao
Bembidion (Ocydromus) saxatile himalotibeticum Schmidt, 2014 Holotype
Male adult
Holotype male, with label data “S TIBET 15.VII.2010 / YamTso lake shore near / Nagartse, near Samding / Monastery, 4420 m” in ZSM.
Copyright © Qin Yuyao
Bembidion (Ocydromus) primorjense Neri & Toledano, 2023 Holotype
Male adult
Holotype, ♂, “Russia N Primorskyi reg. / 70 km NW Plastun 18-26 VII / Podnebesnyi pass 600-800 m / A. Plutenko leg. 1996” [printed]” (CTVR)
Copyright © Qin Yuyao
Bembidion (Asioperyphus) pseudoinfuscatum Neri, Toledano & Rébl, 2023 Holotype
Male adult
Holotype, ♂, “N 49°38’50.3” E 114°41’27.1 / Mongolei Dornod Aimak / Dashbalbar NE, / Salzsee Steppe / 27.07.2012 670m / leg. Schnitter” [printed]” (CTVR).
Copyright © Qin Yuyao
Bembidion (Diplocampa) osellai Neri & Toledano, 2023 Holotype
Male adult
1 ♂, Holotype, (3.40 mm), Median lobe of the aedeagus, holotype (0.64 mm) “Turchia – vil. Gümüshane / Bayburt / m 1200, 1.VII.75 / Osella leg. [printed]” (PN); the specimen is not in good conditions, lacking the two terminal right antennomeres, fore right leg, median left leg and left hind tarsomeres. We added to the specimen the following label [red, printed]: “Bembidion (Diplocampa) osellai P. Neri & L. Toledano, 2022 – Holotypus”
Copyright © Qin Yuyao
Bembidion (Diplocampa) acutangulum Neri & Toledano, 2023 Holotype Paratype
Female adult
1 ♀, paratype, (3.50 mm); “Armenia: ca. 10 km N Sevan / 40°36’48”N 44°57’29”E / nr. Tsovagyugh village / 22.5.2001, ca. 1940m / Shaverdo & Schillhammer [printed]” (NHMW) Median lobe of the aedeagus, holotype (0.68 mm) (NHMW)
Copyright © Qin Yuyao
Bembidion (Diplocampa) loeffleri Jedlicka, 1963c: 169 Holotype Paratype
Male adult
1 ♂, holotype, (3.80 mm); “Persien / Dr. Löffler // Holotype (pink) // Montage / Morvan / 1974 n. 474 (handwritten) // Mus. Nat. Pragae (red, printed) / inv. 66303 (handwritten) // Diplocampa / löffleri sp. n. (handwritten) / det. Ing. Jedlička (pink, printed)” (NMPC) Median lobe of the aedeagus, ♂ paratype (0.73 mm), “Persien / Dr. Löffler // Paratypus (pink) // Mus. Nat. Pragae (red, printed) / inv. 66304 (handwritten) // löffleri sp. n. (handwritten) / det. Ing. Jedlička (pink, printed)” (NMPC).
Copyright © Qin Yuyao
Bembidion (Diplocampa) blandulum Netolitzky, 1910a: 221 Holotype
Male adult
1 ♂, holotype, (3.80 mm); “Persia sett. / Doria 62 [handwritten] // Mus. Civ. / Genova [printed] // Diplocampa / blandulum Net. [handwritten] / det. Netolitzky [printed] // Holotypus [printed] / Bembidion (Diplo / campa) blandulum / Netolitzky, 1910 [pink, handwritten]” (MCSNG) the specimen is in rather good condition, missing only the left antenna, two right antennomeres and a few tarsomeres of the right median leg.
Copyright © Qin Yuyao -
Lissopogonini taxon page (6 photos)
Lissopogonus tonkinensis Zamotajlov & Sciaky, 1996: 40
Male adult
N Vietnam, Lao Cai Province 250 km from Hanoi bearing 310°, SaPa vill env., h~1250 Hoang Lien Son Nat. Park 27.VI-2.VII.1998 leg.A.Napolov
L= 4,8
Copyright © Kirill Makarov
Lissopogonus cf. borneensis Baehr, 2001
Male adult
Malaysia, Borneo, Sarawak, Miri div., Bario 10 km NE, 3 km N Pa Lungan, 7-8.III.2014, R. Cibulskis leg. det. A. Anichtchenko 2015
Copyright © Alexander Anichtchenko
Lissopogonus glabellus Andrewes, 1923
Male adult
Nepal: Annapurna Mts, Madi Khola river below Khilang ca. 1100 m, 14.5.1996 leg. J. Schmidt. Creative Commons CC-BY 3.0
Copyright © David Maddison
Lissopogonus morvani Deuve, 2009
Female adult
Nepal, Kosi Mure, 2000-2100m, 27.30N 87.16E, 6-12.6.2001. 7 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko
Lissopogonus glabellus Andrewes, 1923
Male adult
Nepal, 12km NE Chainpur, 1400m, 29.36.17N, 81.17.54E, 28.6.2009. 5.8 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko -
Patrobini taxon page (75 photos)
Platypatrobus lacustris Darlington, 1938
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Platidiolus vandykei Kurnakov, 1960
Female adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Patrobus quadricollis L.Miller, 1868
Male adult
W Ukraine, NE Carpathians Mts., Transcarpathian reg., Rakhov Distr., Chornogora range, N slope of Sheshul Mt., h~1450m, 9.VI.2009, Panin R. leg./det. length: 9mm
Copyright © Ruslan Panin
Patrobus longicornis (Say, 1823)
Male adult
Great Smoky Mountains National Park
photo I. M. Sokolov
Copyright © Igor Sokolov
Patrobus longicornis (Say, 1823)
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Patrobus foveocollis (Eschscholtz, 1823)
Female adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Patrobus cinctus Motschulsky, 1860
Female adult
Jewish Aut. Reg., Obluchie Distr., 3-5 km upper Radde Vill. on Lagar Riv., forest 24.07.2004 I.Melnik leg.
L=9.5 mm
Copyright © Kirill Makarov
Patrobus atrorufus (Stroem, 1768)
Male adult
length: 7-10 mm Germania, Niedersachsen, Hildesheim leg.det. Kuntze, 1959 Photo: U.Schmidt, 2008
Copyright © U.Schmidt
Patrobus atrorufus (Stroem, 1768)
Female adult
W-Ukraine, Lviv Region, near Kamenka-Buzkaya 20-25.V.07 Panin R. leg. long 8mm
Copyright © Ruslan Panin
Diplous (Platidius) rugicollis (Randall, 1838)
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Diplous (Platidius) dolini Zamotajlov, 2005
Female adult
S Primorie, Lazovsky Reserve, Formosova Spring 43°00'06"N 134°05'13"E 10.VII.2005 leg. K.Makarov
L=9.3
Copyright © Kirill Makarov
Diplous (Diplous) sibiricus sibiricus (Motschulsky, 1844)
Female adult
Transbaikalia, Chita area, Krasnochikoyskyi distr., Chikoy river valley near Nizhnyi Naryn village
18.07.1999
Zinoviev E.leg.
Copyright © Evgeniy Zinoviev
Deltomerus (Deltomerus) circassicus Reitter, 1890
Male adult
Russia, NW-Caucasus,
Karacaj-Cherkessija, upper Pkhiya riv., Arkasara Mt.Range, Zakzan-Syrt Mt., h=2600m, 14.VIII.2009
Solodovnikov I.A. leg.
L=12.5mm
photo Panin R.
Copyright © Ruslan Panin
Deltomerus (Deltomerus) carpathicus L. Miller, 1868:
Female adult
Ukrainian Carpathians, Transcarpathian reg., Rakhov Distr., Chornogora range, h=1650m., 29.VI-28.VII.2007 photo/leg.Panin R. leg. L= 11 mm.
Copyright © Ruslan Panin
Deltomerus (Deltomerus) pseudoplatynus pseudoplatynus s.str.
Female adult
NW-Caucasus,
Krasnodar reg., Apsheron Distr., Lagonaki Mt., near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominykh leg.
long - 11mm.
Copyright © Ruslan Panin
Deltomerus (Deltomerus) pseudoplatynus inferior Zamotajlov,1988
Male adult
S Russia, Krasnodar Reg.,
Near Sochi , Solokh-Aul h=500m, 5.VIII.2007
I.A.Solodovnikov leg.
L=10mm
photo Panin R.
Copyright © Ruslan Panin
Apatrobus (Apatrobus) echigonus (Habu et Baba, 1962) Holotype
Male adult
Mt. Asahidake (alt. 2,000m), NIIGATA, 27. VII. 1961, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Apatrobus (Apatrobus) echigonus (Habu et Baba, 1962) Holotype
Male adult
Mt. Asahidake (alt. 2,000m), NIIGATA, 27. VII. 1961, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Apatrobus (Apatrobus) echigonus (Habu et Baba, 1962) Holotype
Male adult
Mt. Asahidake (alt. 2,000m), NIIGATA, 27. VII. 1961, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Apatrobus (Apatrobus) echigonus (Habu et Baba, 1962) (Label) Holotype
Male adult
Mt. Asahidake (alt. 2,000m), NIIGATA, 27. VII. 1961, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Apatrobus (Apenetretus) shirahatai (Habu et Baba, 1960) Holotype
Male adult
Mt. Kitamata, NIIGATA, 28. VII. 1957, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Apatrobus (Apenetretus) shirahatai (Habu et Baba, 1960) Holotype
Male adult
Mt. Kitamata, NIIGATA, 28. VII. 1957, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Apatrobus (Apenetretus) shirahatai (Habu et Baba, 1960) Holotype
Male adult
Mt. Kitamata, NIIGATA, 28. VII. 1957, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Apatrobus (Apenetretus) shirahatai (Habu et Baba, 1960) (Label) Holotype
Male adult
Mt. Kitamata, NIIGATA, 28. VII. 1957, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Apatrobus (Apenetretus) shoorengensis (Habu et Baba, 1962) Holotype
Male adult
Mt. Shoorenge, NIIGATA, 25. VII. 1961, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Apatrobus (Apenetretus) shoorengensis (Habu et Baba, 1962) Holotype
Male adult
Mt. Shoorenge, NIIGATA, 25. VII. 1961, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Apatrobus (Apenetretus) shoorengensis (Habu et Baba, 1962) Holotype
Male adult
Mt. Shoorenge, NIIGATA, 25. VII. 1961, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Apatrobus (Apenetretus) shoorengensis (Habu et Baba, 1962) (Label) Holotype
Male adult
Mt. Shoorenge, NIIGATA, 25. VII. 1961, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Apatrobus (Apatrobus) jakuchiensis (Habu, 1977) Holotype
Male adult
Mt. Jakuchi, YAMAGUCHI, 27. VI. 1976, K. Tanaka
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-014
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Apatrobus (Apatrobus) jakuchiensis (Habu, 1977) Holotype
Male adult
Mt. Jakuchi, YAMAGUCHI, 27. VI. 1976, K. Tanaka
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Apatrobus (Apatrobus) jakuchiensis (Habu, 1977) Holotype
Male adult
Mt. Jakuchi, YAMAGUCHI, 27. VI. 1976, K. Tanaka
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Apatrobus (Apatrobus) jakuchiensis (Habu, 1977) (Label) Holotype
Male adult
Mt. Jakuchi, YAMAGUCHI, 27. VI. 1976, K. Tanaka
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Apatrobus (Apatrobus) satoui (Habu, 1976) Holotype
Male adult
Mt. Okawa (alt. 1,000m), Kotonami, KAGAWA, 10. X. 1973, M. Sato
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-015
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Apatrobus (Apatrobus) satoui (Habu, 1976) Holotype
Male adult
Mt. Okawa (alt. 1,000m), Kotonami, KAGAWA, 10. X. 1973, M. Sato
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-015
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Apatrobus (Apatrobus) satoui (Habu, 1976) Holotype
Male adult
Mt. Okawa (alt. 1,000m), Kotonami, KAGAWA, 10. X. 1973, M. Sato
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-015
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Apatrobus (Apatrobus) satoui (Habu, 1976) (Label) Holotype
Male adult
Mt. Okawa (alt. 1,000m), Kotonami, KAGAWA, 10. X. 1973, M. Sato
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-015
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Apatrobus (Apatrobus) tsurugiensis (Habu, 1976) Holotype
Male adult
Mt. Tsurugi (alt. 1,400m), TOKUSHIMA, 17. VII. 1970, M. Sato
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-016
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Apatrobus (Apatrobus) tsurugiensis (Habu, 1976) Holotype
Male adult
Mt. Tsurugi (alt. 1,400m), TOKUSHIMA, 17. VII. 1970, M. Sato
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-016
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Apatrobus (Apatrobus) tsurugiensis (Habu, 1976) Holotype
Male adult
Mt. Tsurugi (alt. 1,400m), TOKUSHIMA, 17. VII. 1970, M. Sato
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-016
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Apatrobus (Apatrobus) tsurugiensis (Habu, 1976) Holotype
Male adult
Mt. Tsurugi (alt. 1,400m), TOKUSHIMA, 17. VII. 1970, M. Sato
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-016
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Apatrobus (Apatrobus) tsurugiensis (Habu, 1976) (Label) Holotype
Male adult
Mt. Tsurugi (alt. 1,400m), TOKUSHIMA, 17. VII. 1970, M. Sato
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-016
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Archipatrobus flavipes (Motschulsky, 1864)
Male adult
Collection ground Kobe city, Hyogo Pref. 33645<br /> (C) Working Group for Biological Indicator Ground Beetles Database, Japan<br /> Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Diplous (Diplous) sibiricus caligatus Bates, 1873a: 294
Male adult
Collection ground Kobe city, Hyogo Pref. 34804
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Penetretus rufipennis (Dejean, 1828)
Female adult
Spain, Gredos mts., 2 km E La Mira mt., 4-7.07.2007, 1800m, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko
Deltomerus (Deltomerus) tatricus L. Miller, 1859: 306
Male adult
N Slovakia; Vysoké Tatry Mts., 10mm
Copyright © Aleš Sedláček
Patrobus assimilis Chaudoir, 1844
Female adult
CZ, N Bohemia; Krušné Hory Mts., Výsluní; 8mm
Copyright © Aleš Sedláček
Patrobus septentrionis Dejean, 1828 (? australis)
Male adult
CZ, NE Moravia; Háj ve Slezsku; 8mm
Copyright © Aleš Sedláček
Penetretus temporalis Bedel, 1909
Female adult
Spain, Cadiz prov., Algeciras vic., h=50m, 36.080429, -5.428613, 11-12.05.2014, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko
Patrobus quadricollis L.Miller, 1868
Male adult
W Ukraine, NE Carpathians Mts., Transcarpathian reg., Rakhov Distr., Chornogora range, N slope of Sheshul Mt., h=1450m, 9.VI.2009, Panin R.leg. length: 9mm
Copyright © Ruslan Panin
Dimorphopatrobus ludmilae Casale et Sciaky, 1994: 47
Female adult
Tibet, Motuo, Hanmi, 8.2013. det. A. Anichtchenko, 2015 17 mm
Copyright © Alexander Anichtchenko
Deltomerus (Deltomerus) jelineki Zamotajlov, 2001
Female adult
Iran Alburz prov Shemshak 2750m 12.06.17
Copyright © Jan Muilwijk
Diplous (Platidius) californicus (Motschulsky, 1859)
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Platidiolus cf. borealis Zamotajlov, 2001
Female adult
Russia, Khabarovsk reg., Verhnebureinski distr. Bureya river valley, Ussomah river, 22.7.2006 (anal sternite with 3 pairs of setae). det. Anichtchenko A. 2018
Copyright © Alexander Anichtchenko
Platidiolus rufus Chaudoir, 1878
Male adult
Russia, Hamar-Daban Mt. rng., Komarinskiy Mt., upper flow of Slyudyanka riv. 15.7.2013, A. Shavrin leg. det. Anichtchenko A. 2018
Copyright © Alexander Anichtchenko
Deltomerus (Deltomerus) veldkampi Muilwijk, 2015 Holotype
Male adult
Iran Azarbayjan-e Gharbi/18 km S. Ziveh (Ulugh Dagh)/N 37’07’14” E 044’52’26”/01.09.2008, lg. Frisch & Serri
Copyright © Jan Muilwijk
Penetretus nebrioides (Vuillefroy, 1866)
Male adult
SPAIN, Cantabria, 2 km E Puerto de San Glorio, near Vejo riv., 43.072369, -4.733061, 1300 m., 23.07.2019, A. Anichtchenko & R. Sciaky leg.; det. Anichtchenko A. 2019
Copyright © Alexander Anichtchenko
Himalopenetretus franzi (Zamotajlov & Sciaky, 1998) Holotype
Male adult
India, Uttar Pradesh; 17 km SE of Gangotri, Tapovan, 4500 m, 26.VI.1996 (coll. R. Sciaky)
Copyright © Riccardo Sciaky
Grandipenetretus major (Zamotajlov et Wrase, 1997) Paratype
Male adult
China, Shaanxi, Qin Ling Shan, 47 km S Xian, 26-29.VIII.1995 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky
Parapenetretus (Robustopenetretus) saueri Zamotajlov & Sciaky, 1996 Holotype
Male adult
China – W Sichuan, Emei Shan. 2800 25-31.VI.1992 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky
Prodiplous businskyi (Casale & Sciaky, 2003) Paratype
Male adult
China, SE Tibet, Zayu co., Irawaddy-Salween divide East of Tsema-la pass, 4300 m, 18.VI.1999 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky
Himalopenetretus burangensis Yan, Shi & Liang, 2020 Holotype
Male adult
“Xizang, Ngari Pref., Burang County, Burang Town. N of Tinkar pass, alpine meadow. 30.2085N, 81.0661E, 4800m.”; “under stone, 2019. VII. 15, Shi HL lgt., Exp. BJFU 2019”; “DNA sample series number SHL2019--Xiz006”; “HOLOTYPE♂ Himalopenetretus burangensis sp. nov., des. YAN & SHI, 2020” [red label]. (IZAS). Image from Yan, Shi & Liang, 2020
Copyright © - - -
Troglopatrobus zhouchaoi Deuve, He & Tian, 2020 Holotype
Male adult
cave Lianhua Dong, Tangba Cun, Bailu Zhen, Pengzhou Shi, Chengdu, Sichuan, 31°10ʹ5852ʺN 103°56ʹ09.13ʺE, 1064 m in altitude, 20.VI.2019, leg. Li He, Yuan Li & Chao Zhou (SCAU). Image from Deuve, He & Tian, 2020
Copyright © - - -
Deltomerodes ovicollis Yan, H. Shi, Liang & J. Shi, 2021 Holotype
Male adult
Image from Yan, H. Shi, Liang & J. Shi, 2021
Copyright © - - -
Deltomerodes conaensis Yan, H. Shi, Liang & J. Shi, 2021 Holotype
Male adult
Image from Yan, H. Shi, Liang & J. Shi, 2021
Copyright © - - -
Deltomerus (Deltomerus) corax Peyerimhoff, 1922
Male adult
Morocco, Grand Atlas, 2200-2300m, N31.13.09.5 W07.49.29.7 16-17.6.2010 Anichtchenko A. leg. & det. (cAA) 12 mm
Copyright © Alexander Anichtchenko
Penetretus andalusicus (Reitter, 1897)
Male adult
Spain, Sierra Nevada, Lagunillos de la Virgen, 2850m, 18.5.2005, A. Anichtchenko leg. & det (cAA)
Copyright © Alexander Anichtchenko -
Pogonini taxon page (44 photos)
Cardiaderus chloroticus (Fischer von Waldheim, 1823)
Male adult
Volgogrda Area, Pallasovka Distr., N coast of Elton Lake, right bank of Khara River, 3 km upstream of the mouth, Salina on fl oodplain terrace 20.IX.2006 leg. K.Makarov
L=7.0
Copyright © Kirill Makarov
Pogonistes convexicollis Chaudoir, 1871a: 33
Male adult
Volgogrda Area, Pallasovka Distr., N coast of Elton Lake, Lakeside near the mouth of Khara River, Salt-marsh 20.IX.2006 leg. K.Makarov
L=4.0
Copyright © Kirill Makarov
Pogonus (Pogonoidius) punctulatus Dejean, 1828
Male adult
Volgograd Reg., Pallas Distr., Elton Vill., on UV-light 16-18.VII.2006 K.Makarov leg.
L=6.4 mm
Copyright © Kirill Makarov
Pogonus (Pogonus) iridipennis Nicolai, 1822
Male adult
Volgograd Reg., Pallas Distr., N shore of Elton Lake, right bank of Khara Riv., mouth 13.V.2006 A.Matalin leg.
L=6.7 mm
Copyright © Kirill Makarov
Pogonus (Pogonus) japonicus Putzeys, 1875
Male adult
N. Korea, prov. Phjongan-namdo, Nampho, 24 V 1965, leg. M. Mroczkowski et A. Riedel
Copyright © Mieczyslaw Stachowiak
Pogonus (Pogonus) chalceus chalceus (Marsham, 1802)
Male adult
length: 5,3-7,7 mm
Holland, Goes
leg. W.Veldkamp, 1972; det. U.Schmidt, 1974
Photo: U.Schmidt, 2008
Copyright © U.Schmidt
Pogonus (Pogonus) littoralis Duftschmid, 1812: 183
Male adult
length: 6,5-8 mm
Frankreich, Bretagne, Coulomb
leg. A.Skale, 2006; det. Wrase, 2007
Photo: U.Schmidt, 2008
Copyright © U.Schmidt
Pogonus (Pogonus) smaragdinus Waltl, 1835: 53
Male adult
Oran, Sidi bel Abbes, coll. Plason
Copyright © Mieczyslaw Stachowiak
Pogonus (Pogonus) texanus Chaudoir, 1868
Female adult
Louisiana, Cameron Parish
photo I. M. Sokolov
Copyright © Igor Sokolov
Pogonus (Pogonus) itoshimaensis Habu, 1954 Holotype
Female adult
Maebaru, FUKUOKA, 06. VI. 1952, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-017
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Pogonus (Pogonus) itoshimaensis Habu, 1954 Holotype
Female adult
Maebaru, FUKUOKA, 06. VI. 1952, A. Habu Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-017 (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html) Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Pogonus (Pogonus) itoshimaensis Habu, 1954 Holotype
Female adult
Maebaru, FUKUOKA, 06. VI. 1952, A. Habu<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-017<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Pogonus (Pogonus) itoshimaensis Habu, 1954 (Label) Holotype
Female adult
Maebaru, FUKUOKA, 06. VI. 1952, A. Habu<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-017<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Pogonistes chinensis Habu, 1986 Holotype
Male adult
Nanen, near Peking, CHINA, 09. VII. 1943, Tsushou-Nouji-Shikenjo
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-018
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Pogonistes chinensis Habu, 1986 Holotype
Male adult
Nanen, near Peking, CHINA, 09. VII. 1943, Tsushou-Nouji-Shikenjo
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-018
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Pogonistes chinensis Habu, 1986 Holotype
Male adult
Nanen, near Peking, CHINA, 09. VII. 1943, Tsushou-Nouji-Shikenjo
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-018
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Pogonistes chinensis Habu, 1986 (Label) Holotype
Male adult
Nanen, near Peking, CHINA, 09. VII. 1943, Tsushou-Nouji-Shikenjo
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-018
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake
Pogonistes rufoaeneus Dejean, 1828: 19 (Pogonus)
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Pogonus (Pogonoidius) meridionalis Dejean, 1828
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Pogonus (Pogonoidius) cumanus Lutshnik, 1916c: 76
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Pogonus (Pogonus) orientalis virens Menetries, 1849
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Diodercarus arrowi Lutshnik, 1931: 528
Male adult
Iran, Hormozgan province, Bandar Abbas env., 12.III.2015, S. Azadbakhsh leg.
Copyright © Alexander Anichtchenko
Pogonus (Pogonus) minutus Dejean, 1828
Male adult
Pogonus senegalensis Dejean, 1831 det. M.Baehr, 2015. Size: 5,3 mm. Namibia, Etosha National Park, Okaukuejo, 1400 m leg. U.Schmidt, 18.-20.III.1994; Photo: U.Schmidt, 2016
Copyright © Udo Schmidt
Diplochaetus planatus (G.Horn, 1876)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison
Thalassotrechus barbarae (G.Horn, 1892)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison
Pogonus (Pogonus) fennelli Hudson, 2000
Female adult
Australia 7.5 mm; det. K. Will, 2020
Copyright © Alexander Anichtchenko
Iranopogonus freyellus Jedlicka, 1959 Holotype
Male adult
Iran, Sistan va Baluchestan Province, Chabahar env., IX.1937, leg. Band (NHMB). Image of Matthias Borer (NHMB) from Azadbakhsh, 2019
Copyright © - - -
Pogonopsis pallida Bedel, 1898 Holotype
Male adult
Tunisia (MNHN) Image from Azadbakhsh, 2019
Copyright © - - -
Sirdenus (Syrdenopsis) grayii (Wollaston, 1862)
Female adult
Uzbekistan, Kyzyl Kumy, 25km SW Bukhara, 9.6.1994 Klimenko A. leg. (cAA)
Copyright © Alexander Anichtchenko
Sirdenus (Sirdenus) filiformis (Dejean, 1828)
Male adult
Italie, Sardaigne, Sestu 11-05-1982
Copyright © Pascal Deschamps
Pogonistes gracilis Dejean, 1828
Male adult
Italie Sardaigne Stagna Salé Porcus Sinis 25-04-1982
Copyright © Pascal Deschamps
Pogonus (Pogonus) pallidipennis Dejean, 1828
Female adult
Italie Sardaigne Capo Malfatano 18-04-1982
Copyright © Pascal Deschamps
Pogonus (Pogonus) luridipennis Germar, 1823
Male adult
France 11560 Fleury Etang dePissevaches 23-05-2021
Copyright © Pascal Deschamps
Pogonus (Pogonoidius) meridionalis verdugoi Toribio, 2025 Holotype
Male adult
Habitus: la Laguna de las Yeguas en Alcazar de San Juán (Ciudad Real); left aedeagus: la Laguna de las Yeguas en Alcázar de San Juan (Ciudad Real). Right: la Laguna de Peña Hueca en Villacañas (Toledo). Scale 0,5 mm
Copyright © Marcos Toribio
Pogonus (Pogonus) chalceus viridanus Dejean, 1828
Male adult
Spain, Alicante, Urbanova, Antiguas salinas, 12.1.2024 (DUBC)
Copyright © Alexander Anichtchenko
Pogonus (Pogonus) submarginatus Reitter, 1908
Male adult
Kalmykia, Sarpa lake, 10.7.2007 A. Abramov leg.
Copyright © Alexander Anichtchenko
Pogonus (Pogonoidius) punctulatus Dejean, 1828
Male adult
Nonhyeon-dong, Namdong-gu, Incheon, Korea, 7.IV. 2015, I. J. Choi. leg. Image from Choi et al. 2015
Copyright © - - -
Pogonus (Pogonus) itoshimaensis Habu, 1954
Male adult
Mandol-ri, Simwon-myeon, Gochang-gun, Jeollabuk-do, Korea, 30.VI. 2003, J. C. Paik leg. Image from Choi et al. 2015
Copyright © - - -
Pogonus (Pogonus) japonicus Putzeys, 1875
Male adult
Daebu-do (Is.), Daebunam-dong, Ansan-si, Gyeonggi-do, Korea, 27.VII. 2004, J. C. Paik leg. Image from Choi et al. 2015
Copyright © - - - -
Sinozolini taxon page (1 photos)
Phrypeus rickseckeri (Hayward, 1897)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet
Chaltenia patagonica Roig-Junent et Cicchino, 2001
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Chaltenia patagonica Roig-Junent et Cicchino, 2001
Male adult
San Blas, Provincia de Buenos Aires
Copyright © Armando Cicchino -
Trechini taxon page (792 photos)
Aepus marinus (Strom, 1783)
Male adult
length: 2,2.2,4 mm
England, Aymer Cove
leg.det. M.G.Telfer, 2004
Foto: U.Schmidt, 2009
Copyright © U.Schmidt
Anophthalmus hitleri Scheibel, 1937
Male adult
Slovenia, Jama Pekel Cave, 12.VII.1995 leg. R.Udrzhal
long.
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko
Blemus alexandrovi (Lutshnik, 1915)
Male adult
Jewish Aut. Reg., Obluchie Distr., env. Radde Vill., bank of Amur Riv. between mouth of Lagar and Belaja Riv. 26.07.2004 I.Melnik leg.<br /> L=4.0 mm
Copyright © Kirill Makarov
Blemus discus (Fabricius, 1792)
Male adult
length: 4,5-5,5 mm
Italia, L. die Cavazzo
leg. Budberg, 1961, det. U.Schmidt, 1987
Foto: U.Schmidt, 2008
Copyright © U.Schmidt
Duvaliopsis pilosella L. Miller, 1868 (as rybinskii Knirsch, 1924)
Female adult
W Ukraine, Transcarpatian. Rakhov Distr., N slope of Goverla Mt., Grapa polonina, 48.192°N 24.465°E 4-13.VI.2007 h~1600m I.Melnik leg. L=3.7 mm
Copyright © Kirill Makarov
Duvalius (Hungarotrechus) subterraneus subterraneus L. Miller, 1868
Male adult
W Ukraine, NE Carpathians Mts.
Transcarpathian, Rakhov Distr.,Chornogora Mts., N slope of Sheshul Mt., h=1450m, 9.VI.2009, R.Panin leg./det.V.B. Rizun
length: 6mm
Copyright © Ruslan Panin
Duvalius (Duvalius) gusevi Belousov, 1989
Female adult
N Osetia
photo of S.G. Udalov
taked from www.zin.ru
Copyright © Alexander Anichtchenko
Duvalius (Duvalius) miroshnikovi Belousov et Zamotajlov, 1995: 53
Male adult
Baribin cave, Sochi, Rusia
photo of S.G. Udalov
taked from www.zin.ru
Copyright © Alexander Anichtchenko
Epaphiopsis (Epaphiama) intermedia Belousov & Kabak, 2003 Paratype
Male adult
China, Sichuan, NW of Mt. Yuanbaoshan, right bank of Lianhegou River,
3550-3600m, 20.6.2000 Belousov & Kabak leg.
long.
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko
Epaphiopsis (Epaphiama) nigra Belousov & Kabak, 2003 Paratype
Male adult
China, Sichuan, SSW of Shimian, SE slope of Mt. 4977 W of Lijipin V, H 2700 m,
3.07.2000 Belousov & Kabak. leg.
long.
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko
Epaphiopsis (Epaphiama) proxima Belousov, 2003 Paratype
Male adult
China, S Sichuan, NE of Mianning, 8 km NW Lajiajia. upper forest h 3000m.
4.08.2002 Belousov leg.
long.
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko
Epaphiopsis (Epaphiama) sinuata Belousov & Kabak, 2003 Paratype
Male adult
China, Sichuan, right bank of Lanhegou Riv., NW of Mt. Ubaoshan,
E of Ji..i, 2800-3500m 30.6.2000 Belousov & Kabak leg.
long.
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko
Epaphiopsis (Pseudepaphius) davidiani Belousov, 2003 Paratype
Male adult
China, Sichuan, right bank of Niuzhihe Riv., E of Pusiun Vill., 2500-2700m,
17.6.2000 Belousov & Kabak leg.
long.
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko
Epaphiopsis (Pseudepaphius) korolevi Belousov, 2003 Paratype
Female adult
China, S Sichuan, S of Xichang, E slope of Mt. 4282 (NE of Dechang) 2800m
7.05.2001 Belousov & Korolev leg.
long.
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko
Jeannelius birsteini Ljovuschkin, 1963: 451
Male adult
NW Caucas, caves of Sochi distr.<br /> photo of S.G. Udalov<br /> taked from www.zin.ru
Copyright © - - -
Jeannelius gloriosus Ljovuschkin, 1965: 1263
Male adult
Abkhasia, Novyi Afon cave<br /> photo of S.G. Udalov<br /> taked from www.zin.ru
Copyright © - - -
Jeannelius magnificus Kurnakov, 1959: 233
Male adult
Abkhasia, Gulripshskii distr., caves of Tsebelda depress<br /> photo of S.G. Udalov<br /> taked from www.zin.ru
Copyright © - - -
Meganophthalmus kravetzi Komarov, 1993: 366
Male adult
caves near Nalchik, Central Caucas, Kabardino-Balkaria<br /> photo of S. G. Udalov<br /> taked from www.zin.ru
Copyright © - - -
Meganophthalmus mirabilis Kurnakov, 1959: 234
Male adult
Abkhasia, Gulripshskii distr., Tsebeldinskaja depres. <br /> photo of S.G. Udalov<br /> taked from www.zin.ru
Copyright © - - -
Nannotrechus (Archinannotrechus) ancestralis Belousov, 1998: 113
Male adult
Karachaevo-Cherkessia, low source of Daut riv.<br /> 17.V.2009 Anichtchenko A. leg.<br /> long. 2.3 mm<br /> photo A. Anichtchenko
Copyright © Alexander Anichtchenko
Queinnectrechus angusticollis Belousov & Kabak, 2003
Male adult
China, NW Sichuan, NW of Lixian, 10 km WSW of Shangmeng, alp., 3850-4000m
25.07.2002 Belousov & Kabak leg.
long.
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko
Queinnectrechus humeralis Belousov & Kabak, 2003
Male adult
China, Sichuan, WSW of Lixian, Qunlaishan Mt.R., W of Mt. "5892" h~3500-3600 12.VII.2000 leg. I.Belousov, I.Kabak
L= 4,0 mm
Copyright © Kirill Makarov
Queinnectrechus incisus Belousov & Kabak, 2003 Paratype Paratype
Male adult
China, NW Sichuan, Tonghua, basin of river near Pingshitou, 3300-3400m 19.8.2002
Belousov & Kabak leg.
long.
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko
Queinnectrechus micrangulus Belousov & Kabak, 2003 Paratype Paratype
Male adult
China, Sichuan, SSW of Shimian, E slope of Mt "4977" W of Lijipin 3000-3900m
5.07.2000 Belousov & Kabak leg.
long.
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko
Sinotrechiama imitator Belousov & Kabak, 2003 Paratype Paratype
Male adult
China, NW Sichuan, NE of Lixian, N Tonghua, basin of river nr Pingshitou <br /> 3400-3500m, 19.08.2002<br /> Belousov & Kabak leg.<br /> long.<br /> photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko
Sinotrechiama pilifer Belousov & Kabak, 2003 Paratype Paratype
Male adult
China, Sichuan, right bank of Niuzhihe Riv., E of Pusiun Vill., 2500-2700m<br /> 17.06.2000 Belousov & Kabak leg.<br /> long.<br /> photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko
Sinotrechiama tronqueti (Deuve, 1995)
Female adult
China, N Sichuan, WSW of Beichuan, Mt. Qianfoshan, source of river h~ 2000 m 1.VI.2001 leg. I.Belousov & I.Kabak
L= 6,4 mm
Copyright © Kirill Makarov
Thalassophilus longicornis Sturm, 1825
Male adult
length: 3,4-4,5 mm
Austria, Umgebung Wien
det. H.Tuchert
Photo: U.Schmidt, 2008
Copyright © U.Schmidt
Trechoblemus micros (Herbst, 1784)
Male adult
England, R. Lym Brockenhurst, leg.det. J.M.Walters, 2005 Photo: U.Schmidt, 2009. length: 4-5 mm,
Copyright © U.Schmidt
Trechus (Epaphius) densicornis (Fischhuber, 1977)
Female adult
SE Primorie reg., Lazo res., seashore, near bay Ta-Chingous
27.06.04 leg. M.& L. Smirnov
photo from www.zin.ru
Copyright © Maksim Smirnov
Trechus (Epaphius) ephippiatus Bates, 1873
Female adult
S Primorie, Lazo-raicentr Vill, 43°22'43"N 133°54'01"E on light 6.VIII.2007 leg. K.Makarov, A.Zaitzev
L=4.0
Copyright © Kirill Makarov
Trechus (Epaphius) secalis Paykull, 1790
Male adult
length: 4,2 mm
West Siberia, KHMAO-Yugra, the vicinities of Khanty-Mansiysk town
1.07.2004.
Zinoviev E.leg.
Copyright © Evgeniy Zinoviev
Trechus (Microtrechus) tuckaleechee Barr, 1962
Male adult
Great Smoky Mountains National Park
Copyright © Igor Sokolov
Trechus (Trechus) tchibiloevi Anichtchenko, 2009 Holotype
Male adult
Russia, East Siberia, Irkutsk Territory, 18 km
N Ust Kut, valley of Lena River, 28.07.2008 Shavrin A., Enustschenko I. leg.
photo Anichtchenko A.
Copyright © Alexander Anichtchenko
Ushijimaella lucida Belousov et Kabak, 2002
Male adult
China, Sichuan, WSW of Lixian, Qunlaishan Mt.R., W of Mt. "5892", h~3000 11.VII.2000 leg. I.Belousov, I.Kabak <br /> L= 3,6 mm
Copyright © Kirill Makarov
Trechus (Trechus) quadristriatus (Schrank, 1781)
Female adult
length: 3,5-4,5 mm
Germania, Bayern, Oberfranken, Kulmbach
leg.det. U.Schmidt, 1976
Photo: U.Schmidt, 2008
Copyright © U.Schmidt
Duvalius (Duvalius) mohammadzadehi Muilwijk & Felix, 2008 Holotype
Male adult
Iran, Fars, Estahban, 25.IV.2006, leg. J. Muilwijk.
Copyright © Ron Felix
Duvalius (Duvalius) koeni Muilwijk & Felix, 2008 Holotype
Male adult
SW Iran, Pole-e Tang, 60 km NW Andimeshk, 10-11.IV.1977, Loc. No. 284, Exped. Nat. Mus. Praha.
Copyright © Ron Felix
Taniatrechus setosus Belousov et Dolshanski, 1994: 62
Male adult
W Cauc., Abkhazia, Bzybsky Mt. R., S of Khipsta Mt., 1700 m, Souvenir Cave, –40 m, 19.VIII. 2009. S. Mazina, J. Denisov leg.<br /> Photo of Belousov et Koval.
Copyright © - - -
Meganophthalmus medvedevi Belousov & Koval, 2009 Paratype
Male adult
W Cauc., Abkhazia, Otshamtshira Distr., Otap Vill., Golova Otapa (= Uatapakhy) Cave, middle part, 22.VIII. 2009. A.G. Koval leg.<br /> Photo of Belousov et Koval.
Copyright © - - -
Trechus (Trechus) amplicollis Fairmaire, 1859
Female adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletskyi forest, near Pyatnychany vill., 28.V-1.VII.2011, Panin R. leg./det.Rizun V.B. L=4.5 mm
Copyright © Ruslan Panin
Trechus (Epaphius) nigricornis Motschulsky, 1844
Male adult
det. Makarov K.
Copyright © Kirill Makarov
Trechus (Trechus) pilisensis pilisensis Csiki, 1918
Male adult
W.Ukraine, NW Podolian,
near Lviv city, forest Vynnitsky, on shore lake, 22.IV-17.V.2011, Panin R. leg
L= 4 mm
Copyright © Ruslan Panin
Trechus (Trechus) schyberosiae Szallies & Schüle, 2011 Holotype
Male adult
Kanton Luzern, Schrattenfluh, unterhalb Hengst, 2100 m, 14.6.2009, leg. A. Szallies (NMBE).
Drawing Peter Schuele
Copyright © Peter Schüle
Trechus (Trechus) carrilloi Toribio et Rodriguez, 1997: 283
Male adult
Aedeagus
Copyright © Marcos Toribio
Trechus (Trechus) astrophilus Schmidt, 2009 Paratype
Male adult
TIBET (South Central) 19.VI.07, Budha Vall. N of Yangpachem, ca. 30°11’07N 90°28’42E, 5300–5600 m
photograph of Johannes Reibnitz (SMNS)
Copyright © ---
Trechus (Trechus) glabratus Schmidt, 2009 Paratype
Female adult
TIBET South Centr. 6.VII.07, Doru Tshu Vall. SW Dongu La, 4500–4600 m, ca. 29°43’16N 89°47’12E
photograph of Johannes Reibnitz (SMNS)
Copyright © ---
Trechus (Trechus) mieheorum Schmidt, 2009 Holotype
Male adult
TIBET South Centr. 18.VII.07, Reting Tsangpo Vall. E Reting, upp. Kiykiy side vall. 4900–5200 m, 30°24’38N 91°41’10E” (SMNS).
photograph of Johannes Reibnitz (SMNS)
Copyright © ---
Trechus (Trechus) muguensis Schmidt, 2009 Paratype
Female adult
photograph of Johannes Reibnitz (SMNS)
Copyright © ---
Trechus (Trechus) religiosus Schmidt, 2009 Paratype
Male adult
TIBET (South Central) 11.VII.07, SW slope Nyanchentangla 5100–5500 m, 30°18’24,0N 90°35’48,0E
photograph of Johannes Reibnitz (SMNS)
Copyright © ---
Trechus (Trechus) rolwalingensis rolwalingensis s.str. Holotype
Male adult
NEPAL Rolwaling Vall., Yarlung Ri base camp 4600–4800 m, 16.–18.9.1999 lg. Schmidt” (SMNS)
photograph of Johannes Reibnitz (SMNS)
Copyright © ---
Trechus (Trechus) sculptipennis Schmidt, 2009 Paratype
Female adult
photograph of Johannes Reibnitz (SMNS)
Copyright © ---
Trechus (Trechus) solhoeyi Schmidt, 2009 Paratype
Male adult
TIBET South Centr. 17–20.VI.07, Budha Vall. N of Yangpachem, ca. 30°10’56N 90°29’21E, 5000–5200 m. Image of Johannes Reibnitz (SMNS)
Copyright © - - -
Trechus (Trechus) stratiotes stratiotes s.str. Paratype
Male adult
“NEPAL Prov. Karnali Distr. Humla, 16 km W Simikot, 3 km NW Sankha La, 4300–4800 m, 29°57’18’’N 81°39’30’’E, HF 29.–30.06.2001, leg. A. Kopetz, stone-debris & alpine mats”
photograph of Johannes Reibnitz (SMNS)
Copyright © ---
Trechus (Trechus) tsampa Schmidt, 2009 Holotype
Male adult
“TIBET (South Central) 29.VI.07, Dulong, Kurum vall. NW Lhasa, 4900–5150 m, ca. 29°40’31N 90°46’16E”, “Namba side valley, ascent south west of Namba” (SMNS).
photograph of Johannes Reibnitz (SMNS)
Copyright © ---
Trechus (Trechus) chaklaensis Schmidt, 2009 Paratype
Female adult
TIBET South Centr. 17.VII.07, Lhundup area, above Chak La 5000–5200 m, ca. 30°07’05N 91°16’31E
photograph of Johannes Reibnitz (SMNS)
Copyright © ---
Trechus (Trechus) dongulaensis Schmidt, 2009 Paratype
Female adult
TIBET South Centr., 6.VII.07, 120 km W Lhasa, 2 km NE Dongu La pass, 4800–5000 m, ca. 29°45’01N 89°51’11E”
photograph of Johannes Reibnitz (SMNS)
Copyright © ---
Trechus (Trechus) lama Schmidt, 2009 Paratype
Female adult
“TIBET (South Central) 29.VI.07, Dulong, Kurum vall. NW Lhasa, 4900–5200 m, ca. 29°42’18N 90°35’16E”, “south ascent of Tsubu side valley, above Tsurphu Monastery”
photograph of Johannes Reibnitz (SMNS)
Copyright © ---
Trechus (Trechus) martinae Schmidt, 2009 Paratype
Female adult
“TIBET (South Central) 21.VII.07, Gyama vall. ca. 60 km E Lhasa, 4900–5230 m, ca. 29°40’36N 91°35’33E”, “north west ascent of Chungenpo Shong side vall.”
photograph of Johannes Reibnitz (SMNS)
Copyright © ---
Trechus (Trechus) yeti Schmidt, 2009 Holotype
Male adult
TIBET South Centr. 3–4.VII.07, NE of Shogu La pass 5000–5350 m 29°54’48–29°57’20N 90°08’28–90°07’49E (SMNS). image of Johannes Reibnitz (SMNS)
Copyright © Joachim Schmidt
Tasmanorites grossus Moore, 1972
Male adult
Grove, S.J. (2010). Tasmanian Forest Insect Collection web-site
Copyright © Simon Grove
Trechistus terricola Moore, 1972
Female adult
Grove, S.J. (2010). Tasmanian Forest Insect Collection web-site
Copyright © Simon Grove
Trechobembix baldiensis (Blackburn, 1894)
Male adult
Grove, S.J. (2010). Tasmanian Forest Insect Collection web-site
Copyright © Simon Grove
Dongodytes (Dongodytodes) deharvengi Tian, 2011 Paratype
Male adult
cave, about 4 km south of Xia’ao Xiang, Du’an Yao Minority<br /> Autonomic Xian, coordinate around 107.55 E /24.16 N,<br /> 26 April, 2010, Louis Deharveng leg., in SCAU<br /> photographed by Gao Qi
Copyright © - - -
Trechus (Trechus) bouilloni Faille, Bourdeau & Fresneda, 2012 Paratype
Male adult
Spain, Navarra, Sierra de Urbasa–Andía, Lizarraga, Puerto de Lizarraga, MSS, trap: 1–5–1980/15–8–1980, Bourdeau and Fresneda leg.
Photo Faille, Bourdeau & Fresneda, 2012
Copyright © ---
Trechus (Trechus) grenieri Pandelle, 1867
Male adult
Nistos, Bas–Nistos, Grotte de l’Eglise (Bonadona 1971, Corbaz & Jauzion 1988)
Photo Faille, Bourdeau & Fresneda, 2012
Copyright © ---
Trechus (Trechus) uhagonii uhagonii s.str.
Male adult
Cueva de Orobe
Photo Faille, Bourdeau & Fresneda, 2012
Copyright © ---
Trechus (Trechus) beusti (Schaufuss, 1863)
Male adult
Zegama, Cueva de San Adrián
Photo Faille, Bourdeau & Fresneda, 2012
Copyright © ---
Trechus (Trechus) pieltaini Jeannel, 1920e: 155
Male adult
Cueva de Mairuelegorreta
Photo Faille, Bourdeau & Fresneda, 2012
Copyright © ---
Trechus (Trechus) bruckoides Faille, Bourdeau & Fresneda, 2012 Paratype
Male adult
Pic Montagnon, 15–VII–1979, Bourdeau leg.
Photo Faille, Bourdeau & Fresneda, 2012
Copyright © ---
Trechus (Trechus) bruckii Fairmaire, 1862a: 548
Male adult
Pic de Jaout, dans la zone subalpine, vers 1500 m
Photo Faille, Bourdeau & Fresneda, 2012
Copyright © ---
Trechus (Trechus) anichtchenkoi Toribio, 2012 Holotype
Male adult
Photo Vicente Ortuño
Copyright © ---
Trechus (Trechus) apicalis Motschulsky, 1845b: 347
Male adult
Canada
Photo of Henri Goulet
Copyright © Henri Goulet
Beronaphaenops paphlagonicus Gueorguiev, 2012 Holotype
Male adult
Turkey, Pinarbasi Distr., Milli Park Kűre Dağlari, cave Eşek Çukuru Mağarası 2, 14.VII.2008, P. Beron leg.<br /> Scale bar: 1 mm.<br /> Image of Borislav Gueorguiev, 2012
Copyright © - - -
Epaphiopsis (Epaphiama) jacobsoni Sokolov et Shilenkov, 1987: 10
Male adult
Altai Republic: Holzun mt.rng., Bannaya riv., 1300-1600 m,12-14.06.1999, R.Yu & A.Yu. Dudko leg. & det.
Copyright © Roman Dudko
Perileptus (Perileptus) areolatus areolatus Creutzer, 1799
Male adult
Slovakia, Pastovce, 8.V.2003
Copyright © Alexander Anichtchenko
Duvalius (Neoduvalius) gejzadunayi Lohaj, Čeplík & Lakota, 2013 Holotype
Male adult
MONTENEGRO, Rožaje, Donje Biševo, Pećina u Dubokom potoku cave, 17.4.–11.8. 2009 traps, R.Lohaj, D.Čeplík & G.Dunay lgt. (white label, printed) / HOLOTYPUS Duvalius (Neoduvalius) gejzadunayi sp. n. R. Lohaj, D. Čeplík & J.Lakota det. 2011 (red label, printed) (NMP).<br /> Image from original describtion.
Copyright © - - -
Duvalius (Neoduvalius) starivlahi B.Gueorguiev; S.Curcic & B.Curcic, 2000
Male adult
Hadži Prodanova pećina cave<br /> ZooKeys 278: 91–104
Copyright © - - -
Trechus (Trechus) alpicola alpicola Sturm, 1825: 93
Male adult
CZ, W Bohemia, Šumava Mts.; 3,5mm
Copyright © Aleš Sedláček
Trechus (Trechus) amplicollis Fairmaire, 1859h: CXIIX
Female adult
CZ, N Moravia, Jeseníky Mts.; 4,5mm
Copyright © Aleš Sedláček
Trechus (Trechus) latus latus Putzeys, 1847
Male adult
SK, C Slovakia, Nízké Tatry Mts.; 4mm
Copyright © Aleš Sedláček
Trechus (Trechus) matejkai Vsetecka, 1938: 81
Female adult
SK, C Slovakia, Nízké Tatry Mts.; 3mm
Copyright © Aleš Sedláček
Trechus (Trechus) pulchellus pulchellus Putzeys, 1846
Female adult
SK, C Slovakia, Donovaly; 3mm
Copyright © Aleš Sedláček
Trechus (Trechus) striatulus Putzeys, 1847
Female adult
SK, C Slovakia, Nízké Tatry Mts.; 3,5mm
Copyright © Aleš Sedláček
Trechus (Trechus) nigrinus Putzeys, 1847: 306
Male adult
SK, SW Slovakia, Malé Karpaty; 3,5mm
Copyright © Aleš Sedláček
Duvalius (Hungarotrechus) bokori gelidus Csiki, 19 12c: 510
Male adult
SK, SE Slovakia, Slovenský kras; 3mm
Copyright © Aleš Sedláček
Duvalius (Hungarotrechus) bokori valyianus Bokor, 1922: 132
Male adult
SK, E Slovakia, Muráňská planina; 3,5mm
Copyright © Aleš Sedláček
Trechus (Epaphius) rivularis Gyllanhal, 1810: 33
Female adult
CZ, N Bohemia, Doksy; 4mm
Copyright © Aleš Sedláček
Orotrechus torretassoi J. Muller, 1928
Female adult
ITALY, Friuli Venezia Giulia, Pordenone province, Mt. Cereis "Grotta sfessa" cave, 11.vi. 1988. L. Dreon leg
Copyright © Enrico Ruzzier
Orotrechus muellerianus muellerianus Schatzmayr, 1907
Male adult
ITALY, Friuli Venezia Giulia, Trieste province, Samatorza, "Grotta azzurra" cave, 26. xi. 2011 E. Ruzzier leg.
Copyright © Enrico Ruzzier
Orotrechus vicentinus juccii Pomini, 1940
Male adult
ITALY, Friuli Venezia Giulia, Verona Province, Sant'Ambrogio di Valpolicella, Cave Brentani 12. vii. 2011. E. Ruzzier leg.
Copyright © Enrico Ruzzier
Orotrechus targionii gestroi Tamanini, 1953
Male adult
ITALY, Veneto , Vicenza province,Mt. Grappa, loc. Campo Solagna, 1050 m , 26. v. 2012,
E. Ruzzier leg.
Copyright © Enrico Ruzzier
Masuzoa baicalensis Shilenkov & Anichtchenko, 2008 Paratype
Female adult
S Siberia, Hamar-Daban mts., up of Poperechnaya riv., (left Snezhnaya riv) 1600-1800m, 1-6.V.07, Anichtchenko A. leg.
5.5 mm
Copyright © Alexander Anichtchenko
Trechus (Trechus) quadrimaculatus Motschulsky, 1850
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Trechus (Trechus) isfanensis isfanensis Paratype
Male adult
N slps Turkestan Mt. rng., Aktash Mts., Maran pass, 20 km SSW Isphana, 3000 m, 5.06.1996, A. Putchkov leg.
Copyright © Alexander Anichtchenko
Orotrechus holdhausi bucciarellii Tamanini, 1954b: 104
Male adult
Italy, Veneto, Treviso, San Pedro di Feletto, 2.V.2012, Grotta La Bora, 1252 V/TV, 110m, D. Bianco, E. Piva, E. Ruzzier (Enrico Ruzzier Collection)
Copyright © Alexander Anichtchenko
Trechus (Trechus) putchkovi Belousov et Kabak, 1996: 382 Paratype
Male adult
S Kyrgyzstan, Alai Mts. rng., Gultcha riv., Kysyl-Alai env., 3100 m, 7.07.1995, A. Putchkov leg. (Collection Enrico Ruzzier)
Copyright © Alexander Anichtchenko
Boldoriella (Boldoriella) humeralis (Dodero, 1924)
Male adult
Italy, Lombardia, Brescia, Paitone, 28.V.2011, gr. "Buco del Frate", E. Rizzier leg. & det.; (Collection Enrico Ruzzier) 5 mm
Copyright © Alexander Anichtchenko
Allegrettia pavani castionii Monzini, 2013 Holotype
Male adult
Italia, Lombardia, Valle Imagna Bergamo, Grotta Lacca del Palio 3722 Lo, 14-IV-1990, V. Monzini leg.
Copyright © Vittorino Monzini
Duvalius (Hungarotrechus) corpulentus Weise, 1875: 356
Female adult
W Ukraine, NE Carpathians, Transcarpathian Reg., Rakhiv Distr., Chornogora Mt. Range, S slope h=1600m, bank of stream, 27.VI.2014, Panin R. leg. L=5.5 mm
Copyright © Ruslan Panin
Trechus (Trechus) ericalis Magrini, Queinnec & Vigna Taglianti, 2013 Holotype
Male adult
Etiopia, Oromia Prov., Bale, Goba, Monte Sgona. loc. 38, about 3800 m a.s.l., 8.XI.1973, leg. A. Vigna Taglianti (CVT). Image from Magrini et al, 2013
Copyright © - - -
Thalassophilus azoricus Oromi et Borges, 1991: 2
Male adult
Azorean Biodiversity Portal
Copyright © Enésima Pereira Mendonça
Trechus (Trechus) isabelae Borges & Serrano, 2007
Male adult
Azorean Biodiversity Portal
Copyright © Enésima Pereira Mendonça
Trechus (Trechus) jorgensis Oromi et Borges, 1991: 5
Male adult
Azorean Biodiversity Portal
Copyright © Enésima Pereira Mendonça
Trechus (Trechus) montanheirorum Oromi et Borges, 1991: 7
Male adult
Azorean Biodiversity Portal
Copyright © Enésima Pereira Mendonça
Trechus (Trechus) picoensis Machado, 1988: 2
Male adult
Azorean Biodiversity Portal
Copyright © Enésima Pereira Mendonça
Trechus (Trechus) terceiranus Machado, 1988
Female adult
Azorean Biodiversity Portal
Copyright © Enésima Pereira Mendonça
Trechus (Trechus) terrabravensis Borges; Serrano & Amorim, 2004
Male adult
Azorean Biodiversity Portal
Copyright © Enésima Pereira Mendonça
Dongodytes (Dongodytodes) yaophilus Tian, Yin & Huang, 2014 Paratype
Female adult
Guangxi: Dahua: Qibainong: cave Qiaoxu Dong, 24°04.370N, 107°40.140E, 535 m, 2013-VI-22, leg. Mingyi Tian, Wei Lin, Haomin Yin & Sunbin Huang (SCAU). Scale bar: 1.0 mm. Image from Tian, Yin & Huang, 2014
Copyright ©
Dongodytes (Dongodytes) lani Tian, Yin & Huang, 2014 Paratype
Female adult
Guangxi: Du’an: Longwan: cave Longhuan Dong, 23°49.5213N, 108°14.4792E, 248 m, 2013-XII-24, leg. Mingyi Tian, Weixin Liu, Haomin Yin & Xiaozhu Luo (SCAU). Scale bar: 1.0 mm. Image from Tian, Yin & Huang, 2014
Copyright ©
Dongodytes (Dongodytes) troglodytes Tian, Yin & Huang, 2014 Holotype
Male adult
Guangxi: Du’an: Longfu: Shangme: cave Shuiyuan Dong, 24°11.335N, 107°49.865E, 509 m, 2013-VI-28, leg. Mingyi Tian, Wei Lin, Weixin Liu, Haomin Yin & Sunbin Huang (SCAU). Scale bar: 1.0 mm. Image from Tian, Yin & Huang, 2014
Copyright ©
Dongodytes (Dongodytes) elongatus Tian, Yin & Huang, 2014 Paratype
Female adult
Guangxi: Du’an: Lalie: Fuyan: Jianong: cave Lapo Dong I, 24°11.987N, 108°20.378E, 140 m, 2013-VI-23, leg. Mingyi Tian, Weixin Liu, Haomin Yin & Sunbin Huang (SCAU). Scale bar: 1.0 mm. Image from Tian, Yin & Huang, 2014
Copyright © - - -
Dongodytes (Dongodytodes) jinzhuensis Tian, Yin & Huang, 2014 Paratype
Female adult
cave Jinzhu Dong I, 24°06.547 N, 108°04.785 E, 190 m (CSAU). Scale bar: 1.0 mm. Image from Tian, Yin & Huang, 2014
Copyright © - - -
Dongodytes (Dongodytodes) brevipenis Tian, Yin & Huang, 2014 Paratype
Male adult
Guangxi: Du’an: Chengjing: Wanmao: cave Diaomao Dong, 24°01.723N, 108°07.236E, 140 m, 2013-VI-23, leg. Mingyi Tian, Wei Lin, Weixin Liu, Haomin Yin & Sunbin Huang, (SCAU). Scale bar: 1.0 mm. Image from Tian, Yin & Huang, 2014
Copyright © - - -
Dongodytes (Dongodytodes) brevipenis Tian, Yin & Huang, 2014 Paratype
Male adult
Guangxi: Du’an: Longwan: cave Nongqu Dong I, 23°56.021N, 108°10.962E, 459 m, 2013-VI-27, leg. Mingyi Tian, Wei Lin, Weixin Liu, Haomin Yin & Sunbin Huang, (SCAU). Scale bar: 1.0 mm. Image from Tian, Yin & Huang, 2014
Copyright © - - -
Dongodytes (Dongodytodes) inexpectatus Tian, Yin & Huang, 2014 Holotype
Male adult
Guangxi: Du’an: Gaoling: Nongchi: cave Nongguangshang Dong II, 24°05.4263N, 108°04.5726E, 175 m, 2013-XII-27, leg. Mingyi Tian, Weixin Liu, Haomin Yin & Xiaozhu Luo (SCAU). Scale bar: 1.0 mm. Image from Tian, Yin & Huang, 2014
Copyright © - - -
Lessinodytes caoduroi Vigna Taglianti, 1982: 369
Male adult
Mt. Lessini caves, Pre-Alps Verona, Italy - V. Monzini leg.
Copyright © Vittorino Monzini
Italaphaenops dimaioi Ghidini, 1964: 35
Male adult
Mt Lessini caves, Verona Pre-Alps, V. Monzini leg.
Copyright © Vittorino Monzini
Trechus (Trechus) championi Jeannel, 1920d: 109
Male adult
India, Uttarakhand, 13-15.04.2012, Uttarkashi distr., N30°57'41.57", E78°41'54.75" left tributary of Bhagirathi riv., 2300 m, Anichtchenko A. leg. det. Anichtchenko A.
Copyright © Alexander Anichtchenko
Perileptus (Perileptus) imaicus Jeannel, 1923
Male adult
N India, Uttarakhand, 10-12.04.2012, Uttarkashi distr., N30*44,999' E078*34,412', 1446m, 14 km E Uttarkashi, Anichtchenko A. leg & det
Copyright © Alexander Anichtchenko
Perileptus cf. robustus - indicus
Male adult
N India, Uttarakhand, 5 km N Ramnagar vill., 23-26.4.2012, A. Anichtchenko leg.
Copyright © Alexander Anichtchenko
Trechus (Trechus) sachalinensis Lafer, 1989: 141
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Trechus (Trechus) tarbagataicus Belousov et Kabak, 1992b: 829
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Trechus (Trechus) planipennis Rosenhauer, 1856: 41
Male adult
Spain, Sierra Nevada, 2300m det. A. Anichtchenko
Copyright © Kirill Makarov
Trechus (Epaphius) nigricornis Motschulsky, 1844
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Trechus (Epaphius) rivularis (Gyllenhal, 1810)
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Trechus (Trechus) concinnus Tschitscherine, 1904b: 148
Female adult
det. K. Makarov
Copyright © Kirill Makarov
Trechus (Trechus) hoppi hoppi Jeannel, 1927: 334
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Trechus (Trechus) kantegiricus Belousov et Kabak, 1994a: 15
Male adult
det. K. Makarov
Copyright © Kirill Makarov
Trechus (Minitrechus) gypaeti Vigna Taglianti & Magrini, 2009 Holotype
Male adult
Etiopia - Arussi - loc. 33 bis / M. Encuolò, vetta / m 4000 circa - 5.XI.1973 / A. Vigna leg. (CVT). Image from Vigna Taglianti & Magrini, 2009.
Copyright © - - -
Paraphaenops breuilianus Jeannel, 1916
Male adult
Cueva de la Cambra, Mola de Cati, Tarragona, Hispania - leg. V. Monzini, 1988
Copyright © Vittorino Monzini
Trechisibus (Trechisibus) eleonorae Allegro; Giachino & Sciaky, 2008 Holotype
Male adult
Perú, Ancash, Cordillera Blanca, prov. de Asunción, Punta Olimpica, laguna m 4200, 22.XI.2005, G. Allegro leg. (CAl).
Copyright © Gianni Allegro
Typhlotrechus bilimekii hacquenii Sturm, 1853
Male adult
Slovenia, Ljubljana, Gorni lg.Jama, Velika Basica, IV-IX.2003, M. Egger leg. L=7.5 mm
Copyright © Ruslan Panin
Anophthalmus schmidtii insignis J. Muller, 1912: 299
Male adult
W-Slovenia, Dinaric Alps, Trnovo Forest Plateau, Lokve env., 2.V-16.X.2010, M. Egger leg. L=8 mm
Copyright © Ruslan Panin
Anophthalmus fabbrii J. Muller, 1931c: 190
Male adult
Villanova caves, Udine, Italy - 14-V-1983, V. Monzini leg.
Copyright © Vittorino Monzini
Duvalius (Duvalius) winklerianus aequalis Jeannel, 1931
Male adult
Italy, Brescia, Lumezzane, 26.VI.2013, leg. & det. M. Agosti, photo by M.Agosti
Copyright © - - -
Duvalius (Duvalius) gestroi gestroi Dodero, 1900
Male adult
photo by Mauro Agosti
Copyright © - - -
Oodinotrechus (Pingleotrechus) yinae Sun & Tian, 2015 Holotype
Male adult
Guangxi: Guilin: Pingle: Ertang: Letang: cave Chaotianyan, 110°45’501"E / 24°37’075"N, 5-XII-2011, Mingyi Tian, Weixin Liu & Haomin Yin leg, (SCAU). Image from Sun & Tian, 2015
Copyright © - - -
Perileptus (Perileptodes) darlingtoni Casale & Laneyrie, 1989
Male adult
Papua New Guinea, Papua prov., Genyem env., 500m, 30.XII.2008. det. A. Anichtchenko
Copyright © Alexander Anichtchenko
Anophthalmus gobanzi gobanzi Ganglbauer, 1911: 237 (Trechus)
Female adult
Austria, Carinthia, Vellachtal, Paulitschsattel env., 3.VI. 1989, M. Egger leg. L=4 mm
Copyright © Ruslan Panin
Duvalius (Biharotrechus) transcarpathicus Shilenkov et Rizun, 1989
Female adult
W Ukraine, NE Carpathians, Tiachiv distr., cave near Uglya vill., 15.V-26.VI.2015, Panin R. & Kanarsky Yu. leg. L=5 mm
Copyright © Ruslan Panin
Trechus (Trechus) plicatulus L. Miller, 1868
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, alpin zone, 11.VI-1.IX.2015, Panin R. & Kanarsky Yu. leg. L=4.8 mm
Copyright © Ruslan Panin
Duvalius (Hungarotrechus) ruthenus ruthenus Reitter, 1878
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, alpin zone, 11.VI-1.IX.2015, Panin R. & Kanarsky Yu. leg. L=5.2 mm
Copyright © Ruslan Panin
Trechus (Trechus) pseudomontanellus Rizun, 1994
Male adult
W Ukraine, NE Carpathians,Transcarpathian Reg., Marmarosh Mts., near Lug vill., bank of Kuziy (Kizi) stream, 400m 18.V.2015, Panin R. leg. L= 4.5 mm
Copyright © Ruslan Panin
Dactylotrechus yalongensis Belousov & Kabak, 2014 Holotype
Male adult
China, Sichuan Province, SW Mianning Town, 28º13´19´´ N / 101º43´21´´ E, H=3700 m, 07.07.2011, Belousov & Kabak leg. (ZISP). Image from IGOR A. BELOUSOV & ILYA I. KABAK, 2014
Copyright © - - -
Puertrechus mengsaensis Belousov & Kabak, 2014 Paratype
Female adult
CH, Yunnan Province, Mt. E Mengsa Town, 23º42´26´´ N / 99º47´26´´ E, H=3045 m, 30.05.2010, Belousov & Kabak leg. Image from IGOR A. BELOUSOV & ILYA I. KABAK, 2014
Copyright © - - -
Trechus (Trechus) fontinalis Rybinski, 1901
Male adult
W Ukraine, NE Carpathians Mts., Chornogora Mt Range, h=2015m, 7.VI.2015, R. Panin leg./det. L= 5mm.
Copyright © Ruslan Panin
Shenaphaenops humeralis Ueno, 1999j: 630 Holotype
Male adult
Cave Shendong Migong (Shendong in the original description), Muqiao, Laoyingshan, Shuicheng County, northwestern Guizhou, 26°35'15"N/ 104°59'47"E, 1910 m in altitude, VIII-22-2014, Mingyi Tian leg. Scale bar: 2.0 mm. Image from Mingyi Tian et al., 2016
Copyright © - - -
Shiqianaphaenops majusculus Ueno, 1999c: 47 Holotype
Male adult
Cave Feng Dong, Tangshan, Shiqian County, 27°29'10"N/108°15'23"E, 700 m in altitude, Mingyi Tian & Jingli Cheng leg.Scale bar: 2.0 mm. Image from Mingyi Tian et al., 2016
Copyright © - - -
Dianotrechus gueorguievi Tian, 2016 Holotype
Female adult
China, Yunnan Province, Ermu Vill., Kunming District, Dashi Dong (Big Rock Cave), 24°49'13"N/102°27'56"E, 1940 m in altitude, XI-8- 2011, B. Petrov leg.” (SCAU). Scale bar: 2.0 mm. Image from Mingyi Tian et al., 2016
Copyright © - - -
Wanhuaphaenops zhangi Tian & Wang, 2016 Holotype
Male adult
Cave Songjia Dong, Beihu Qu, Chenzhou, southern Hunan Province, 25°40'08.05"N/112°53'59"E, 493 m in altitude, VIII-25-2015, Xinhui Wang, Sunbin Huang, Mingruo Tang & Pingjing Yang leg. (SCAU). Scale bar: 2.0 mm. Image from Mingyi Tian et al., 2016
Copyright © - - -
Tianeotrechus trisetosus Tian & Tang, 2016 Holotype
Male adult
Cave Bahao Dong, Gandong Village, Bala Xiang, Tian’e County, northern Guangxi, 24°55'57.10"N/107°02'40.80"E, 686 m in altitude, VIII-7-2015, Mingruo Tang leg. (SCAU). Scale bar: 2.0 mm. Image from Mingyi Tian et al., 2016
Copyright © - - -
Huoyanodytes tujiaphilus Tian & Huang, 2016 Holotype
Female adult
Cave Tujiamei Dong, Huoyan Karst, Huoyan Xiang, Wulongshan Geopark, Longshan County, NW Hunan Province, China, 29°12'20.11"N/109°18'37.21"E, 427 m in altitude, VII-3-2014, leg. Mingyi Tian, Weixin Liu, Haomin Yin, Sunbin Huang & Xinhui Wang, deposited in SCAU. Scale bar: 2.0 mm. Image from Mingyi Tian et al., 2016
Copyright © - - -
Kozlovites caviceps Jeannel, 1935 Holotype
Female adult
Copyright © - - -
Dactylotrechus setosus Belousov & Kabak, 2003 Paratype
Female adult
Copyright © - - -
Pseudanophthalmus ciliaris orlindae Barr, 1959 Paratype
Female adult
Kent. VI.25.1957, Cook Cave, nr. Adairville, Logan Co., TC Barr. Type N 4260: 2 (MZLU)
Copyright © Lund University
Orotrechus gigas Vigna Taglianti, 1981: 70
Male adult
V. Monzini leg. & det.
Copyright © Vittorino Monzini
Trechodes alluaudi Jeannel, 1926
Male adult
Madagascar Est, Ré. Ambatondrazaka, 5km N Didy, J. Janak leg., 1995, P. Schüle det.
Copyright © Peter Schüle
Trechodes jeanneli Mateu, 1958 (f. typ.)
Male adult
Madagascar Est, 14.16.1., Rég. Ambatondrazaka, 5km N Didy, J. Janak lgt., 1995
Copyright © Peter Schüle
Trechodes jeanneli var.
Male adult
Madagascar Est, 14.16.1., Rég. Ambatondrazaka, 5km N Didy, J. Janak lgt., 1995
Copyright © Peter Schüle
Duvalius (Hungarotrechus) subterraneus subterraneus L. Miller, 1868: 10
Female adult
Copyright © Kirill Makarov
Trechus (Trechus) terskeiensis ukokensis Belousov & Kabak, 1996
Male adult
Copyright © Kirill Makarov
Cimmeritodes (Zhecimmerites) parvus Tian & Li, 2016 Paratype
Female adult
Copyright © - - -
Wanoblemus wui Tian & Fang, 2016 Holotype
Male adult
Copyright © - - -
Guiaphaenops deuvei Tian, Feng & Wei, 2017 Holotype
Male adult
cave near Yangli Cun, Jiayou Zhen, Lingyun Xian, Baise Shi, Guangxi, 24°28'39"N, 106°37'52"E, 643 m, VII-25-2012, Mingyi Tian, Weixin Liu, Feifei Sun & Haomin Yin leg., in South China Agricultural University, Guangzhou, China (SCAU). Image from Tian, Feng & Wei, 2017
Copyright © - - -
Dongoblemus kemadongicus Deuve & Tian, 2016 Holotype
Female adult
Chine, Yunnan, Zhenxiong Xian, Chishuiyuan, grotte Kema Dong, 1730 m, 27°39’13’’N - 104°51’10’’E (Tian Mingyi, Liu Weixin, Wang Xinhui, Tang Mingruo, 23.VI.2015), in coll. South China Agricultural University, Canton (Chine). Image from Deuve & Tian, 2016
Copyright © - - -
Cimmeritodes (Xiangcimmerites) zhongfangensis Deuve & Tian, 2016 Holotype
Female adult
Chine, Hunan, Huaihua Shi, Zhongfang Xian, Huangyan Xiang, Fujia Cun, grotte Jinji Dong, 890 m, 27°28’20’’N - 110°4’7’’E (Tian Mingyi, Liu Weixin, Wang Xinhui, Tang Mingruo, 1.VII.2015), in coll. South China Agricultural University, Canton (Chine). Image from Deuve & Tian, 2016
Copyright © - - -
Cimmeritodes (Dianocimmerites) crassifemoralis Deuve & Tian, 2016 Holotype
Female adult
Chine, Yunnan, Zhenxiong Xian, Yudong Xiang, Shangshiyuan Cun, grotte Xiao Dong, 1350 m, 27°37’15’’N - 105°2’9’’E (Tian Mingyi, Liu Weixin, Wang Xinhui, Tang Mingruo, 24.VI.2015), in coll. South China Agricultural University, Canton (Chine). Image from Deuve & Tian, 2016
Copyright © - - -
Graciliblemus lipingensis Deuve & Tian, 2016 Holotype
Female adult
Chine, Guizhou, Liping Xian, grotte Feilong Dong, 26°19’35’’N - 109°11’17’’E (Tian Mingyi, Cheng Jingli, 22.VIII.2015), in coll. South China Agricultural University, Canton (Chine). Image from Deuve & Tian, 2016
Copyright © - - -
Guizhaphaenops yudongensis Deuve & Tian, 2016 Holotype
Male adult
Chine, Yunnan, Zhenxiong Xian, Yudong Xiang, Shangshiyuan Cun, grotte Xiao Dong, 1350 m, 27°37’15’’N - 105°2’9’’E (Tian Mingyi, Liu Weixin, Wang Xinhui, Tang Mingruo, 24.VI.2015), in coll. South China Agricultural University, Canton (Chine). Image from Deuve & Tian, 2016
Copyright © - - -
Cathaiaphaenops cychroides Deuve & Tian, 2016 Holotype
Male adult
Chine, Hubei, Lichuan Xian, Liangwu, grotte Zakuo Dong, 1169 m, 30°18’09’’N - 108°49’39,20’’E (Tian Mingyi, Liu Weixin, Yin Haomin, Huang Sunbin, Cheng Jingli, 4.VII.2014), in coll. South China Agricultural University, Canton (Chine). Image from Deuve & Tian, 2016
Copyright © - - -
Cathaiaphaenops enshiensis Deuve & Tian, 2016 Holotype
Male adult
Chine, Hubei, Enshi, Banqiao, grotte Huangjingping Dadong, 1710 m, 30°30’42’’N - 109°18’38’’E (Tian Mingyi, Huang Sunbin, Wang Xinhui, Tang Mingruo, 8.XI.2015), in coll. South China Agricultural University, Canton (Chine). Image from Deuve & Tian, 2016
Copyright © - - -
Trechisibus (Trechisibus) lemairei Allegro & Giachino, 2016 Holotype
Male adult
Peru, Dept. Huánuco, Quebrada Queulacocha, m 3,800, VI.1981, legit M. Curti (CAl).
Copyright © - - -
Trechisibus (Trechisibus) huascarani Allegro & Giachino, 2016 Holotype
Male adult
Peru, Dip. Ancash, Punta Olímpica, m 4,550, 5.VII.2008, legit G. Allegro (CAl). Image from Allegro & Giachino, 2016
Copyright © - - -
Trechisibus (Trechisibus) recuayi Allegro & Giachino, 2016 Holotype
Male adult
Peru, Dip . Ancash, Punta Olímpica , m 4,550, 5.VII.2008, legit G. Allegro (CAl). Image from Allegro & Giachino, 2016
Copyright © - - -
Trechisibus (Trechisibus) curtii Allegro & Giachino, 2016 Holotype
Male adult
Peru, Dept. Huánuco, Quebrada Queulacocha , m 3,800, VI.1981, legit M. Curti (CAl). Image from Allegro & Giachino, 2016
Copyright © - - -
Trechisibus (Trechisibus) brevis Allegro & Giachino, 2016 Holotype
Male adult
"Argentina, Neuquen, Pen. Quetrihué, 12.II.1998, legit Riccardo Sciaky (CAl). Image from Allegro & Giachino, 2016" - wrong label. specimen from Koralpe (Austria)
Copyright © - - -
Trechus rotundipennis (as Trechisibus longipenis Allegro & Giachino, 2016) Holotype
Male adult
"Argentina, Neuquen, Pen. Quetrihué, 12.II.1998, legit Riccardo Sciaky (CSc)." - wrong label. Specimen from Koralpe (Austria). Image from Allegro & Giachino, 2016
Copyright © - - -
Trechisibus (Trechisibus) parvulus Allegro & Giachino, 2016 Holotype
Male adult
"Argentina, Neuquen, Pen. Quetrihué, 12.II.1998, legit Riccardo Sciaky (CSc)" - wrong label. Specimen from Koralpe (Austria). Image from Allegro & Giachino, 2016
Copyright © - - -
Trechisibus (Trechisibus) sciakyi Allegro & Giachino, 2016 Holotype
Male adult
"Argentina, Neuquen, Pen. Quetrihué, 12.II.1998, legit Riccardo Sciaky (CSc). Image from Allegro & Giachino, 2016" - wrong label
Copyright © - - -
Trechus splendes (as Trechisibus striatus Allegro & Giachino, 2016) Holotype
Male adult
"Argentina, Neuquen, Pen. Quetrihué, 12.II.1998, legit Riccardo Sciaky (CSc)." - wrong label. Specimen from Koralpe (Austria). Image from Allegro & Giachino, 2016
Copyright © - - -
Duvaliopsis pilosella pilosella L. Miller, 1868 (as rybinskii Knirsch, 1924)
Male adult
W Ukraine, Chornagora Range, author photo Yu. Kanarsky
Copyright © Ruslan Panin
Duvaliopsis pilosella pilosella L. Miller, 1868 (as rybinskii Knirsch, 1924)
Male adult
W Ukraine, Chornagora Range, author photo Yu. Kanarsky
Copyright © Ruslan Panin
Trechus (Trechus) tenuiscapus Lindroth, 1961
Male adult
along the Bristelcone Pine trail, 2300m , GBNP (EMEC344626), Nevada, Great Basin National Park. 4.9 mm
Copyright © Kipling W. Will
Kozlovites amplipennis Belousov et Kabak, 2016 Paratype
Female adult
China, W Yunnan, SW Weideng, 26°58´51´´N / 98°58´35´´E – 26°58´51´´N / 98°59´04´´E, H=3725–4150 m, 03.06. 2005, Belousov I. & Kabak I. leg.
Copyright © - - -
Kozlovites modestus Belousov et Kabak, 2016 Paratype
Female adult
China,Yunnan, Shangrila–>Deqen,214 Ntn. Road, NE slope of SE Baima Mt.R., SW of Benzilanzhen Vill., 28°08´05´´N/ 99°13´21´´E, 07.06.2013, H=3865 m, Belousov I. & Kabak I. leg.
Copyright © - - -
Kozlovites yuae Deuve, 1992f: 171 Paratype
Male adult
without geographical label data. “IOZ(E) 11364” (IZAS).
Copyright © - - -
Kozlovites niger Belousov et Kabak, 2016 Paratype
Female adult
CH, Yunnan, N Weixi, l. trib. Luozhua, 7.75 km E Nianjua Mt., 27°44´ 9´´N / 98°57´ 3´´E, 15.06.2015, H=4080 m, Belousov I. & Kabak I. leg.
Copyright © - - -
Agonotrechus fugongensis Deuve and Liang, 2016 Holotype
Male adult
Image from Deuve et al, 2016
Copyright © - - -
Agonotrechus xiaoheishan Deuve and Kavanaugh, 2016 Holotype
Male adult
Image from Deuve et al, 2016
Copyright © - - -
Agonotrechus wuyipeng Deuve, 1992f: 172
Male adult
Gongshan County: Qiqi Trail at No 12 Bridge, N27.71500°/ E98.50222°, 2770 m, 2 May 2002, H.B. Liang & w.D. Ba collectors. Image from Deuve et al, 2016
Copyright © - - -
Minutotrechus minutus (Ueno, 1997) Paratype
Female adult
China, Yunnan, Gaoligong Shan, Tengchong County, Dabei, 2430 m (IOZ). Image from Deuve et al, 2016
Copyright © - - -
Eocnides fragilis Ueno, 1989a: 14
Male adult
Bingzhongluo Township (Niwaluo He, just below Nu Jiang Road, 1630 m, N28.05140°/ E98.59319°, 8 October 2002, D.H. Kavanaugh, P.E. Marek and D.Z. Dong collectors. Image from Deuve et al, 2016
Copyright © - - -
Queinnectrechus (Queinnectrechus) griswoldi Deuve & Kavanaugh, 2016 Holotype
Male adult
CHINA, Yunnan, Gongshan County, Cikai Township, 0.1 km SE of Heipu Yakou in valley below tunnel, N27.76978°/ E98.44681°,”/ “3720 m, 13 August 2006, Stop #DHK-2006-073 D.H. Kavanaugh & J.A. Miller collectors (IOZ). Image from Deuve et al, 2016
Copyright © - - -
Queinnectrechus (Queinnectrechus) gongshanicus Deuve & Liang, 2016 Holotype
Male adult
CHINA, Yunnan, Gongshan County, Bingzhongluo Township, Sw slope of Kawakarpu Shan, on slope NE of Chukuai Lake, 3950 m,”/ “N27.98206/ E098.48027°, 20 August 2006, Stop #DHK-2006-086 Y. Liu, P. Hu, D.Z. Dong, & J. wang collectors (IOZ). Image from Deuve et al, 2016
Copyright © - - -
Queinnectrechus (Gaoligongtrechus) balli Deuve and Kavanaugh, 2016 Holotype
Male adult
CHINA, Yunnan Province, Gaoligong Shan, Nujiang Prefecture, Nujiang State Nature Reserve, Dulong/Gongshan Yakou [= Qiqi/Dulong divide] area, 21 airkm w of Gongshan,”/ “N27.69655°/ E98.45389°, 3300-3680m, 16-17 July 2000, Stop#00-24C, D.H. Kavanaugh, C.E. Griswold, Liang H.-B., D. Ubick, & Dong D.-Z. collectors (IOZ). Image from Deuve et al, 2016
Copyright © - - -
Trechus (Trechus) indicus indicus Putzeys, 1870b: 175
Male adult
China, Yunnan. Image from Deuve et al, 2016
Copyright © - - -
Trechus (Trechus) shiyueliang Deuve & Kavanaugh, 2016 Holotype
Male adult
CHINA, Yunnan, Fugong County, Lishadi Township, 10-11 km w of Shibali on Shibali Road, 3200-3280 m, NN27.19980°/E98.71375° to N27.20654°/E98.71772°,”/ 8 August 2005, Stop#DHK-2005-068 D.H. Kavanaugh, H.B. Liang, P. Paquin, & D.Z. Dong collectors (IOZ). Image from Deuve et al, 2016
Copyright © - - -
Epaphiotrechus fortipesoides Deuve & Kavanaugh, 2016 Holotype
Female adult
China, Yunnan, Tengchong County, Houqiao Township, 8.5 airkm NNE of Houqiao at Gaoshidong, N25.39858°/ E98.30533°, 2580 m. Image from Deuve et al, 2016
Copyright © - - -
Trechepaphiama gaoligong Deuve & Kavanaugh, 2016 Holotype
Female adult
China, Yunnan, Gaoligong Shan, Lushui County, 9.3 km ESE of Pianma, N25.99363°/E98.66651°, 2460-2470 m. Image from Deuve et al, 2016
Copyright © - - -
Trechepaphiopsis monochaeta Deuve & Kavanaugh, 2016 Holotype
Male adult
China, Yunnan, Gongshan County, Qiqi Trail at No. 12 Bridge Camp area, 16.3 airkm W of Gongshan, N27.71503°/E98.50244°, 2775 m. Image from Deuve et al, 2016
Copyright © - - -
Trechepaphiopsis unipilosa Deuve & Liang, 2016 Holotype
Male adult
China, Yunnan, Fugong County, Shiyueliang Township, 8.5 km above Shibali on Shibali Road, North Fork of Yamu He, N27.18416°/E98.72026°, 3100 m. Image from Deuve et al, 2016
Copyright © - - -
Trechepaphiopsis unisetulosa Deuve & Kavanaugh, 2016 Holotype
Male adult
China, Yunnan, Fugong County, Shiyueliang Township, 0.2 km W of Shibali, N27.16650°/E098.77936°, 2537 m. Image from Deuve et al, 2016
Copyright © - - -
Trechepaphiopsis uniporosa Deuve & Liang, 2016 Holotype
Male adult
Image from Deuve et al, 2016
Copyright © - - -
Trechus (Trechus) mingguangensis Deuve & Liang, 2016 Holotype
Male adult
Image from Deuve et al, 2016
Copyright © - - -
Trechus (Trechus) qiqiensis Deuve & Kavanaugh, 2016 Holotype
Male adult
Image from Deuve et al, 2016
Copyright © - - -
Trechus (Trechus) pseudoqiqiensis Deuve & Liang, 2016 Holotype
Female adult
Image from Deuve et al, 2016
Copyright © - - -
Trechus (Trechus) luzhangensis Deuve & Liang, 2016 Holotype
Male adult
Image from Deuve et al, 2016
Copyright © - - -
Trechus (Trechus) gongshanensis Deuve & Liang, 2016 Holotype
Male adult
Image from Deuve et al, 2016
Copyright © - - -
Trechus (Trechus) shibalicus Deuve & Kavanaugh, 2016 Holotype
Male adult
Image from Deuve et al, 2016
Copyright © - - -
Trechepaphiopsis unisetosa (Deuve, 2004)
Male adult
Image from Deuve et al, 2016
Copyright © - - -
Trechus (Trechus) gamae Reboleira & Serrano, 2009 Paratype
Male adult
Algar do Pena, UTM: 29SND16596841, 18.8.2007 ; P. Sofia Reboleira leg. (MNHN http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec7460)
Copyright © Alexander Anichtchenko
Kenodactylus audouini Guerin-Meneville, 1830
Male adult
Argentina, PN Tierra del Fuego, Bahia Lapataia, 12-15.I.2003, T. Ueno; det. T. Ueno, 2007
Copyright © Alexander Anichtchenko
Iberotrechus bolivari (Jeannel, 1913)
Female adult
Spain, Cantabria, Santiurde de Toranzo, Cueva del Pis (Penilla), 19.VIII.2004, M. Toribio leg.
Copyright © Alexander Anichtchenko
Pachydesus rufipes rufipes (Boheman, 1848)
Male adult
RSA, Mpumalanga Middelburg to Stoffberg 43,5 km after Middelburg, 16.11.2001. 6 mm; det. Geginat
Copyright © Alexander Anichtchenko
Xuedytes bellus Tian & Huang, 2017 Holotype
Male adult
Cave II, southeastern Du’an Yao Autonomous County, Hechi Shi (=Prefecture), northern Guangxi Zhuang Autonomous Region, southern China, VIII-08-2017, leg. Mingyi Tian, Sunbin Huang, Dianmei Wang and Mengzhen Chen. Deposited in the insect collection of South China Agricultural University, Guangzhou. Image from Tian, Huang & Wang, 2017.
Copyright © - - -
Giraffaphaenops clarkei Deuve, 2002
Male adult
Image from Tian, Huang & Wang, 2017
Copyright © - - -
Dongodytes (Dongodytes) grandis Ueno, 1998a: 12
Male adult
Image from Tian, Huang & Wang, 2017
Copyright © - - -
Amblystogenium pacificum (Putzeys, 1869)
Male adult
Ile de la Possesion, Crozet, T.A.A.F, II.1962 Lerri leg. (Collection R. Sciaky)
Copyright © Riccardo Sciaky
Spelaeovulcania canariensis Machado, 1987c: 332 Paratype
Male adult
C. Labrada, 9.III.83, J. L. Martin L. L. Hernandez (Collection R. Sciaky)
Copyright © Riccardo Sciaky
Geotrechus orpheus Dieck, 1869
Male adult
France, Ariege, Aven de Santhe Catherine, VIII.1981. Leg. Sciaky (Collection R. Sciaky)
Copyright © Riccardo Sciaky
Neaphaenops tellkampfii (Erichson, 1844)
Male adult
Kentucky, Barren Co., Downer's (=Johnson's) Cave, SW Cave City, Sept 27 1949m Jones, Valentine and Royer (Collection R. Sciaky)
Copyright © Riccardo Sciaky
Darlingtonea kentuckensis Valentine, 1952
Male adult
Indian Cave (Ky.), 26.VIII.46, H.H. Leg. (Collection R. Sciaky)
Copyright © Riccardo Sciaky
Doderotrechus ghilianii ghilianii Fairmaire, 1859a: 26
Male adult
Piemonte, Valle Po, Crissolo (TO), Gr. Buco di Valenza, 5.V.1995 (Collection R. Sciaky)
Copyright © Riccardo Sciaky
Kosswigia insularis Jeannel, 1947
Male adult
Turkey - Konya, Beyseyr Gölü, Cave on the island Aci Akif, 13.VII.1988, leg. Zoia (Collection R. Sciaky)
Copyright © Riccardo Sciaky
Agostinia launi (Gestro, 1892)
Male adult
Piemonte - Cuneo, Val Pesio, Grotta Superiore del Camoscere, 27.VII.1983, leg. Sciaky (Collection R. Sciaky)
Copyright © Riccardo Sciaky
Anchotrechus punctipennis Jeannel, 1927
Male adult
Tenerife, Anaga, Barranco de Inguana, III.1984, leg. Sciaky (Collection R. Sciaky)
Copyright © Riccardo Sciaky
Aphaenopsis apfelbecki (Ganglbauer, 1891)
Male adult
Bjelasnica, K. Kysely (Collection R. Sciaky)
Copyright © Riccardo Sciaky
Trichaphaenops (Trichaphaenops) gounellei annae Jeannel, 1949a: 30
Male adult
France, Gouffre Berger, Isere- Engins, Vercors 4.VIII.1981, leg. Regalin (Collection R. Sciaky)
Copyright © Riccardo Sciaky
Duvalius (Duvalius) auruncus Vigna Taglianti & Magrini, 2008 Paratype
Male adult
Grava dei Serini cave, Esperia, Lazio FR, leg. Magrini & Bastianini.
Copyright © Vittorino Monzini
Epaphiopsis (Allepaphiama) arunachala J.Schmidt, 2016 Holotype
Male adult
“INDIA Arunachal Pradesh / Chander N of Bomdila / ca. 3200 m, 17-26.VI.2008 / 27°25N 92°22E lg. Reutter” (CSCHM).
Copyright © Joachim Schmidt
Epaphiopsis (Allepaphiama) aruna J.Schmidt, 2016 Paratype
Male adult
“NEPAL Arun Vall. to Makalu / Bawala Pass 41-4300m / 26.VI.2013 lg. S.Tamang / 27°42’(47-44’’)N 87°14’(45-29’’)E”
Copyright © Joachim Schmidt
Epaphiopsis (Allepaphiama) chomolungma (Deuve et Queinnec, 1985)
Male adult
E Nepal: Dhupi Kharka, 3100 m, 2.VI.2008
Copyright © Joachim Schmidt
Epaphiopsis (Allepaphiama) pathibara J.Schmidt, 2016 Holotype
Male adult
“NEPAL Taplejung Distr. / W-slope Pathibara 2900-3000m / 11.V.2016 leg. Schmidt / 27°24’40N 87°45’29’’E” (CSCHM).
Copyright © Joachim Schmidt
Epaphiopsis (Allepaphiama) schawalleri (Deuve et Hodebert, 1991) Holotype
Male adult
“319 Ilam Distr., Mai Pokhari, / 2100-2200 m, Castanopsis forest / remants [sic], 9.-10 April 1988, J. / MARTENS & W. SCHAWALLER leg.”, “NEPAL-Expeditionen / Jochen Martens”, “Trechus / schawalleri n. sp. / Th. DEUVE det. 1990”, “HOLOTYPE” (SMNS).
Copyright © Joachim Schmidt
Epaphiopsis (Allepaphiama) simbuaensis simbuaensis Holotype
Male adult
“275 Taplejung Dist., Grat / Lasse Dhara und Alm Lassetham 3000-3300mAbies/Rhodod / 6/7Sep83 MARTENS & DAAMS l.”, “NEPAL-Expeditionen / Jochen Martens”, “Trechus / simbuaensis n. sp. / Th. DEUVE det. 1990”, “HOLOTYPE” (SMNS).
Copyright © Joachim Schmidt
Epaphiopsis (Allepaphiama) simbuaensis gracililamella J.Schmidt, 2016 Paratype
Male adult
“E-NEPAL Taplejung Distr. / E-slope Pathibara 3-3400m / 14-16.V.2016 leg. Schmidt / 27°26’20”N 87°46’44”E”
Copyright © Joachim Schmidt
Epaphiopsis (Allepaphiama) simbuaensis gracililamella J.Schmidt, 2016 Paratype
Male adult
“E-NEPAL Taplejung Distr. / E-slope Pathibara 3-3400m / 14-16.V.2016 leg. Schmidt / 27°26’20”N 87°46’44”E”
Copyright © Joachim Schmidt
Epaphiopsis (Allepaphiama) tamurensis (Deuve et Hodebert, 1991)
Male adult
E-Nepal, Taplejung District: E-slope Pathibara, 3500-3750 m, 27°26’N 87°46’E, 13.V.2016, leg. J. Schmidt (CSCHM)
Copyright © Joachim Schmidt
Epaphiopsis (Allepaphiama) tamurensis (Deuve et Hodebert, 1991) Holotype
Male adult
“359 Taplejung Distr., pasture / Lassetham NW Yamputhin, 3300- / 3500 m, mature Abies-Rhododen- / dron forest, 6-9 May 1988 / J. MARTENS & W. SCHAWALLER leg.”, “NEPAL-Expeditionen / Jochen Martens”, “Trechus / tamurensis n. sp. / Th. DEUVE det. 1990”, “HOLOTYPE” (SMNS).
Copyright © Joachim Schmidt
Epaphiopsis (Allepaphiama) thorectes (Ueno, 1983)
Male adult
Gufa Pokhari to Chauki [#21b], 2600-2900 m, 27°17’N 87°30’E to 27°12’N 87°28’E, 21.VI.2001, 1 male leg. NHMB Basel expedition to Nepal 2001 (NHMB).
Copyright © Joachim Schmidt
Trechus (Trechus) additus J.Schmidt, 2016 Paratype
Male adult
S TIBET 27.VI.2011 / Kurum Vall., S Nampa side / vall., 51-5350m, lg. Schmidt / 29°40’16N 90°45’48E (CSCHM).
Copyright © Joachim Schmidt
Trechus (Trechus) budhaensis linchungensis J.Schmidt, 2016 Holotype
Male adult
S TIBET, ca. 30km W of / Yangpachem, 9.VII.2010 / Linchung Vall. 4950-5270m / 30°06’56’’N 90°20’24’’E (CSCHM).
Copyright © Joachim Schmidt
Trechus (Trechus) astrophilus Schmidt, 2009 Paratype
Male adult
Nyainqentanglha Shan: Bhuda Valley
Copyright © Joachim Schmidt
Trechus (Trechus) christinae J.Schmidt, 2016 Paratype
Male adult
S TIBET, E of Yangpachem / above Shemalung Monastery / 4900-5400m 19.VII.2010 / 30°07’42’’N 90°36’16’’E
Copyright © Joachim Schmidt
Trechus (Trechus) dongola J.Schmidt, 2016 Paratype
Male adult
S TIBET 18.VI.2011 / vall. SE of Dongo La pass / 4900-5300m, leg. Schmidt / 28°46’N 87°57’E
Copyright © Joachim Schmidt
Trechus (Trechus) dromai J.Schmidt, 2016 Paratype
Male adult
S TIBET, N of Yangpachem / Nyainqentanglha Shan / 4900-5100m 6+7.VII.2010 / 30°06’57’’N 90°27’24’’E
Copyright © Joachim Schmidt
Trechus (Trechus) frigophilus J.Schmidt, 2016 Holotype
Male adult
S TIBET 25.VI.2011 / Lobha La E of Zuru Tso / 5570m, leg. J. Schmidt / 29°54’34’’N 86°42’26’’E (CSCHM).
Copyright © Joachim Schmidt
Trechus (Trechus) frater J.Schmidt, 2016 Holotype
Male adult
S TIBET 10.VI.2011 / upper Neyam Chu Vall. / 4900-5200m leg. Schmidt / 30°14’N 91°02’E (CSCHM).
Copyright © Joachim Schmidt
Trechus (Trechus) hiekei hiekei Paratype
Male adult
TIBET Chüsül Vall. / W slope Rana side vall. / 5-5300m, 3.VII.2010 / 29°30’47N 90°40’35E
Copyright © Joachim Schmidt
Trechus (Trechus) hiekei subtiliformis J.Schmidt, 2016 Holotype
Male adult
TIBET 27.VI.2011 / Kurum Vall., S Nampa side / vall., 51-5350m, lg. Schmidt / 29°40’16N 90°45’48E (CSCHM).
Copyright © Joachim Schmidt
Trechus (Trechus) menpa J.Schmidt, 2016 Paratype
Male adult
S TIBET 7.VI.2011 / N Menpa Monastery / 5100-5350m lg. Schmidt / 30°07’56N 92°12’11E
Copyright © Joachim Schmidt
Trechus (Trechus) nami J.Schmidt, 2016 Paratype
Male adult
S TIBET 22.VII.2010 Namtso / Nyainqentanglha Shan N-slope / valley of eastern tributary of / Shashe Tsu riv., 5100-5450m, 30°34’25N 90°44’39E to / 30°34’28N 90°45’28E
Copyright © Joachim Schmidt
Trechus (Trechus) neyamensis J.Schmidt, 2016 Paratype
Male adult
S TIBET 10.VI.2011 / upper Neyam Chu Vall. / 4900-5200m leg. Schmidt / 30°14’N 91°02’E
Copyright © Joachim Schmidt
Trechus (Trechus) parvifrater J.Schmidt, 2016 Paratype
Male adult
S TIBET 9.VI.2011 / N-slope Neyam Chu Vall. / ca. 5100m, leg. Schmidt / 30°15’N 91°02’E
Copyright © Joachim Schmidt
Trechus (Trechus) religiosus birze J. Schmidt, 2016 Paratype
Male adult
S TIBET 12.VI.2011 / N-slope Mt. Birze / 4880-5120m lg. Schmidt / 30°03’N 90°32’E
Copyright © Joachim Schmidt
Trechus (Trechus) religiosus lhai J. Schmidt, 2016 Paratype
Male adult
S TIBET, E Nyainqentanglha / Feng, E-slope Lha Tsu Vall. / 5000-5350m 20.VII.2010 / 30°22’47N 90°43’22E
Copyright © Joachim Schmidt
Trechus (Trechus) religiosus nigropiceus J. Schmidt, 2016 Paratype
Male adult
S TIBET 11.VI.2011 / N-slope Lombuk Chu Vall. / 5100-5400m leg. Schmidt / 30°13’46N 90°31’52E
Copyright © Joachim Schmidt
Trechus (Trechus) tsampa striatopunctatus J.Schmidt, 2016 Paratype
Male adult
S TIBET 11.VII.2010 / Mt. Bir Tse 5-5350m / glacier vall. N of Ice Fall / N of 29°57’27’’N 90°31’06’’E
Copyright © Joachim Schmidt
Trechus (Trechus) tseringi Schmidt, 2009 Paratype
Male adult
Dongu La Pass, N-slope
Copyright © Joachim Schmidt
Trechus (Trechus) utsi J.Schmidt, 2016 Paratype
Male adult
S TIBET Utzi N of Lhasa / 5-5100m, 1.VII.2010 / 29°42’52N 91°01’56E
Copyright © Joachim Schmidt
Trechiama (Trechiama) longicollis Ueno, 1986a: 5
Male adult
Japan, Tokushima Ken
Copyright © Vittorino Monzini
Rakantrechus (Paratrechiama) tofaceus Ueno, 1970b: 106
Male adult
Japan, Kumamoto Ken
Copyright © Vittorino Monzini
Thalassoduvalius masidai Ueno, 1956a: 65
Male adult
Japan, Kioto, leg. S. Kawai
Copyright © Vittorino Monzini
Boldoriella (Insubrites) grignensis Monzini, 1987: 13 Paratype
Male adult
Italy, Grigna Mt., leg V. Monzini
Copyright © Vittorino Monzini
Paratrechus pallescens (Bolivar & Pieltain, 1967)
Female adult
Messico, Queretaro, leg. G. Trezzi
Copyright © Vittorino Monzini
Aphaenopidius kamnikensis kamnikensis Drovenik, 1987: 53 Paratype
Male adult
Copyright © Vittorino Monzini
Pseudaphaenops jacobsoni Pliginskiy, 1912: 505
Male adult
Crimea, Boriu Teshik cave, 19-6-1996, leg. A. G. Koval
Copyright © Vittorino Monzini
Geotrechus orcinus Linder, 1859: 72
Male adult
France, gr.te de Gargas, H.P., 15-4-1962, leg. M. Cabidoche
Copyright © Vittorino Monzini
Orotrechus stephani prenottoi Daffner, 1986: 31
Male adult
Italia, TV, Possagno, gr. buco del Dinosauro, 1-5-2004, leg. L. Bodei
Copyright © Vittorino Monzini
Trichaphaenops (Trichaphaenops) gounellei annae Jeannel, 1949 (as attenuatus)
Male adult
gr. de Moras cave, Rencurel, III-1991, leg. C. Genest
Copyright © Vittorino Monzini
Aphaenops (Hydraphaenops) giraudi delicatulus Coiffait, 1962
Male adult
Pierre San Martin Cave, 23-XII-1987, France, leg. E. Queinnec
Copyright © Vittorino Monzini
Boldoriella (Boldoriella) carminatii (Dodero, 1917)
Male adult
Tomba del Polacco cave, 17-V-1980 BG Italia, V. Monzini leg.
Copyright © Vittorino Monzini
Pheggomisetes globiceps Buresch, 1925
Male adult
Bulgaria, Ponor plannina, Dusnica Pestera cave, 30/7/2004, Vasiljev leg.
Copyright © Vittorino Monzini
Pseudaphaenops tauricus Winkler, 1912a: 134
Male adult
Crimea, Dolgorukovskaya Yalla, yama Tshokrak gully, 13/5/2007, Turbanov leg.
Copyright © Vittorino Monzini
Boldoriella (Boldoriella) monguzzii inflata Monzini, 2005 Paratype
Female adult
Italia, Magasa BS, Prefon del Gas cave, 3/5/2003, V. Monzini leg.
Copyright © Vittorino Monzini
Orotrechus haraldi Daffner, 1990
Male adult
Eisenkappel, Lobnig Lobni gschacht, Austria - IX/99 - leg. M. Egger
Copyright © Vittorino Monzini
Orotrechus gracilis Meggiolaro, 1961a: 52
Male adult
Buco del Dinosauro cave, Possagno (PD) Italia - 15/VI/90 - leg. G. Trezzi
Copyright © Vittorino Monzini
Cnides cf jeanneli Ueno, 1985
Female adult
Creative Commons CC-BY 3.0. det. Anichtchenko A. 2018
Copyright © David Maddison
Cyphotrechodes gibbipennis (Blackburn, 1901)
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Oxytrechus cf. lallemandi Jeannel, 1927
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Paratrechodes macleayi (Sloane, 1920)
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Perileptus (Perileptus) dentifer Darlington, 1935
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Aphaenops (Cerbaphaenops) cerberus (Dieck, 1869)
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Pseudanophthalmus engelhardti (Barber, 1928)
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Pseudocnides (Pseudocnides) rugosifrons (Jeannel, 1920)
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Trechiella subornatella (Blackburn, 1901)
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Trechinotus flavocinctus Jeannel, 1962
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Tropidotrechus victoriae (Blackburn, 1894)
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Trechus (Trechus) humboldti Vandyke, 1945
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Xenotrechus denticollis Barr & Krekeler, 1967
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Duvalius (Duvalius) doderoi Gestro, 1885
Male adult
Tann-a de Raixe cave, Carsi GE 12/4/2014, leg. V.Monzini. Photo Riccardo Sciaky
Copyright © Vittorino Monzini
Dongodytes (Dongodytes) tonywhitteni Yang, Huang & Tian, 2018
Male adult
Image from Yang, Huang & Tian, 2018
Copyright © - - -
Aphaenops (Hydraphaenops) pandellei Linder, 1859: 72
Male adult
France, Pyrenei
Copyright © Vittorino Monzini
Duvalius (Duvalius) hetschkoi matilei Deuve, 2000
Male adult
Iran Kerman Jiroft, Narab 900m 16.xi.1999 Badii, Bar. Mof leg. Macroptere
Copyright © Jan Muilwijk
Duvalius (Biharotrechus) paroecus dryops Bokor, 1913a: 211
Male adult
Romania M.ts Bihar
Copyright © Vittorino Monzini
Duvalius (Paraduvalius) joannidisi Casale, Giachino et M. Etonti, 1990
Female adult
Macedonia
Copyright © Vittorino Monzini
Duvalius (Duvalius) ramorinii (Gestro, 1887)
Male adult
Genova, Italia. Foto R. Sciaky
Copyright © Vittorino Monzini
Aphaenops (Hydraphaenops) chaudoirii Brisout da Banneville, 1867
Male adult
France
Copyright © Erminio Piva
Orotrechus mandriolae Ganglbauer, 1911
Male adult
Cima Mandriolo, Trento, Italia. Photo of Erminio Piva
Copyright © Vittorino Monzini
Aphaenops (Hydraphaenops) ehlersi (as longiceps Jeannel, 1926)
Male adult
France
Copyright © Vittorino Monzini
Aphaenops (Hydraphaenops) giraudi delicatulus Coiffait, 1962
Male adult
France
Copyright © Vittorino Monzini
Aphaenops (Cerbaphaenops) laurenti Genest, 1983
Male adult
France. Photo of Erminio Piva
Copyright © Vittorino Monzini
Paraphaenops breuilianus Jeannel, 1916: 282
Male adult
Mola de Catì, España. Photo of Erminio Piva
Copyright © Vittorino Monzini
Orotrechus frassanellicus Piva, 2018 Holotype
Male adult
Colli Euganei, Vicenza, Italia
Copyright © Vittorino Monzini
Orotrechus vicentinus juccii Pomini, 1940
Male adult
Monti Lessini, Vicenza Italia
Copyright © Alexander Anichtchenko
Derossiella lukici Lohaj & Delić, 2019
Male adult
Image from Lohaj & Delić, 2019 (photo courtesy of Dušan Beňo).
Copyright © - - -
Derossiella nonveilleri Queinnec, 2008
Male adult
Image from Lohaj & Delić, 2019 (photo courtesy of Dušan Beňo).
Copyright © - - -
Adriaphaenops petrimaris Lohaj & Delić, 2019
Male adult
Image from Lohaj & Delić, 2019 (photo courtesy of Dušan Beňo).
Copyright © - - -
Adriaphaenops kevser Queinnec, Pavicevic & Ollivier, 2008
Male adult
Image from Lohaj & Delić, 2019 (photo courtesy of Dušan Beňo).
Copyright © - - -
Adriaphaenops rumijaensis Lohaj, Lakota, Queinnec, Pavićević & Ceplic, 2016
Male adult
Image from Lohaj & Delić, 2019 (photo courtesy of Dušan Beňo).
Copyright © - - -
Luraphaenops gaudini gineti (Jeannel, 1955)
Male adult
Aven Borel, Saint-Christol, Vaucluse, France
Copyright © Jean Michel Lemaire
Aphaenops (Hydraphaenops) bourgoini bergesi Coiffait, 1958
Male adult
Juzet d’Izaut, Haute Garonne, France
Copyright © Vittorino Monzini
Aphaenops (Hydraphaenops) bourgoini bourgoini Jeannel, 1946
Male adult
France
Copyright © Vittorino Monzini
Aphaenops (Simaphaenops) penacollaradensis Dupre, 1991
Male adult
Photo of Erminio Piva
Copyright © Vittorino Monzini
Aphaenops (Hydraphaenops) pandellei pandellei (Linder, 1859)
Male adult
Pyrénées Atlantiques - France
Copyright © Vittorino Monzini
Geotrechus gallicus gaudini Jeannel, 1928
Male adult
Pyrénées Atlantiques - France
Copyright © Vittorino Monzini
Allegrettia boldorii Jeannel, 1928 (as zavattarii Ghidini, 1934)
Male adult
Prealpi Bresciane, Italia
Copyright © Vittorino Monzini
Pachydesus rufipes (Boheman, 1848)
Male adult
Namibia, Khomas Reg., 99 km ESE from Windhoek Eningu Clayhouse Lodge 22°46'50"S 18°00'58"E 20.II.2018 leg. A. Rubenyan
Copyright © Kirill Makarov
Orotrechus schwienbacheri Grottolo et Martinelli, 1991: 153
Male adult
Val Celina, PN, Italia (Photo E. Piva)
Copyright © Vittorino Monzini
Orotrechus sebenelloi Daffner, 1983: 219
Female adult
Valdobbiadene, VI, Italia
Copyright © Vittorino Monzini
Duvalius (Duvalius) vaccae Gestro, 1885
Male adult
grotta della Besta cave - Tenda, France. Photo of Riccardo Sciaky.
Copyright © Vittorino Monzini
Epaphiopsis (Formosiellus) grebennikovi Deuve, 2017 Holotype
Male adult
Taïwan, canton de Yilan, forêt de Taïping Shan, 2216 m., 24°29’47’’N-121°31’30’’E (V. Grebennikov, 27 mars 2013), in coll. National Museum of Natural Science, Taichung, Taiwan.
Copyright © Thierry Deuve
Paratrechus maddisoni Deuve & Moret, 2017 Paratype
Male adult
Equateur, Pichincha, massif de Guamani, col de Papallacta (Paso de la Virgen), 4060 m., 0°20’8,52’’S-78°12’7,199W, M. Gobbi et P. Moret, 11 mars 2017. (MNHN)
Copyright © Thierry Deuve
Paratrechus hypogeus Deuve & Moret, 2017 Holotype
Male adult
Equateur, Prov. Tungurahua, Parc national de Llanganatis, P13, Cerro Jaramillo, 4150 mètres, 1°09’40.6’’S - 78°21’23.5’’W, P. Moret, 14 février 2010. (MNHN).
Copyright © Thierry Deuve
Trechisibus (Trechisibus) pubescens Deuve & Moret, 2017 Holotype
Male adult
Equateur, Prov. Loja, Cordillera Lagunillas, 14 km au sud de Jimbura, 3510 mètres, 4°44’36’’S-79°25’28’’W (P. Moret, 11 août 2013), in coll. QCAZ de l’Université catholique de Quito.
Copyright © Thierry Deuve
Trechisibus (Trechisibus) barragani Deuve & Moret, 2017 Holotype
Male adult
Equateur, Province de Loja, Parc National de Yacuri, « Waypoint 168 », 3445 mètres, 4°44’12’’S-79°25’38’’W (P. Moret, S. Aguirre, E. Moreno, 4 août 2016, « on open ground, 19h-20h30 »), in coll. QCAZ.
Copyright © Thierry Deuve
Arctaphaenops muellneri Schmid, 1972a: 96
Male adult
Österreich, Kleinreifling, leg. M.Egger
Copyright © Vittorino Monzini
Trechus (Epaphius) rivularis (Gyllenhal, 1810)
Female adult
W-Ukraine, Roztochia, Lviv Reg., Yavoriv distr., N of Ivano-Frankovo vill., Site «Zalyvky», peat bog, 17.V-18.VI.2019, Panin R. & Kanarsky Yu. leg
Copyright © Ruslan Panin
Trechus (Trechus) suturalis Putzeys, 1870
Male adult
Spain, Cantabria, 2 km E Puerto de San Glorio, near Vejo riv., 43.072369, -4.733061, 1300 m., 23.07.2019, A. Anichtchenko & R. Sciaky leg. ; det. M. Toribio, 2019
Copyright © Alexander Anichtchenko
Trechus (Trechus) kricheldorffi Wagner, 1913
Female adult
Spain, Mt. Cantabricos, Picos de Europa, 2200m, 3km NW mirador Fuente De, 22.07.2019, A. Anichtchenko & R. Sciaky leg.; det. M. Toribio, 2019
Copyright © Alexander Anichtchenko
Trechus (Trechus) fulvus fulvus Dejean, 1831
Male adult
Madeira, Funchal,11-16.9.2004. det. Anichtchenko A., 2019
Copyright © Alexander Anichtchenko
Trechus (Trechus) custos Wollaston, 1854
Male adult
Madeira, Funchal, 11-16.9.2004. det. Anichtchenko A. 2019
Copyright © Alexander Anichtchenko
Trechus (Trechus) tetracoderus Gemminger et Harold, 1868
Female adult
Madeira, Funchal, 11-16.9.2004. 4.8 mm. det. Anichtchenko A., 2019
Copyright © Alexander Anichtchenko
Iga formicina Ueno, 1953
Male adult
Japan, Tokushima-ken (Photo of Riccardo Sciaky)
Copyright © Vittorino Monzini
Allegrettia pavani orobiensis Monzini, 2015 Holotype
Male adult
Italia, Lombardia, Valle Seriana, Parre, Bergamo, miniera (1) del M.te Trevasco, 12-IV-2002, V. Monzini leg. (CA)
Copyright © Vittorino Monzini
Speotrechus mayeti (Abeille de Perrin, 1875)
Male adult
France, mines de fer Privas
Copyright © Vittorino Monzini
Trechus (Trechus) nigrocruciatus Wollaston, 1854: 64
Female adult
Madeira
Copyright © Vittorino Monzini
Anophthalmus schmidtii flachi (Winkler, 1912)
Female adult
Ucka - Croatia
Copyright © Vittorino Monzini
Mexaphaenops elegans Barr, 1967
Female adult
Sierra Gorda, Pinal de Amoles, 2700m, Ojo de Agua - Mexico
Copyright © Vittorino Monzini
Tasmanorites magnus Moore, 1972
Male adult
Australia Tas. m1450 Ben Lemond N.P. Hamilton Crags. Meadow Vale
Copyright © Vittorino Monzini
Sporades (Perileptosporades) jaechi Liebherr, 2020 Holotype
Male adult
NEW CALEDONIA (NC 27) / N-Prov., Poum Com. / unnamed river / 9.XI.2016, leg. M.A. Jäch // ca. 7 km NE Poum / ca. 15 m a.s.l. / 20°12'18.3"S 164°5'27.7"/ almost dried out lowland river (NHMW).
Copyright © James K. Liebherr
Allegrettia pedersolii Vailati, 2017
Male adult
mines of the Presolana, Bergamo, Italia
Copyright © Vittorino Monzini
Epaphiopsis (Epaphiama) nishikawai Ueno, 1987
Male adult
Hokkaido, Yakumo-cho, Unseki Pass
Copyright © Vittorino Monzini
Aphaenops (Cerbaphaenops) tiresias tiresias (as proserpina)
Male adult
Ariege, St. Lary, Fr
Copyright © Vittorino Monzini
Tonkinaphaenops yinquanicus Huang & Tian, 2019 Holotype
Male adult
Copyright © - - -
Boldoriella (Boldoriella) brigantiae Monguzzi & Regalin, 2001
Male adult
Copyright © Vittorino Monzini
Duvalius (Neoduvalius) gejzadunayi Lohaj, Čeplik & Lakota, 2013 Paratype
Male adult
Copyright © Vittorino Monzini
Duvalius (Duvalius) miroshnikovi Belousov et Zamotajlov, 1995: 53
Female adult
Copyright © Vittorino Monzini
Duvalius (Duvalius) dellermae Magrini & Sbordoni, 2020 Holotype
Male adult
Italy: Lazio, Southern Aurunci Mountains, Pandora Cave, N° 1850 L /LT, 1225 m a.s.l., locality Fosse di Sola, plateau between Monte Redentore and Monte Altino (Maranola, Latina). UTM WGS 84: 385895 4573991. 6.VI.2013, Federica Dellerma legit (CM). Image from Magrini & Sbordoni, 2020
Copyright © - - -
Orcusiella prokletijensis Pavićević, Lohaj & Popović, 2020 Holotype Paratype
Male+Female adult
Montenegro, Prokletije Mts., Bjelič massif, Maja Kolata, “Jama Kolektor” pit, 2052 m a.s.l., 20. 07. 2014 - 28. 07. 2015, traps, leg. M. Popović. HOLOTYPUS Orcusiella gen. nov. prokletijensis sp. nov., D. Pavićević, R. Lohaj & M. Popović des. 2020 (CDP). Image from Pavićević, Lohaj & Popović, 2020
Copyright © - - -
Orotrechus messai J. Muller, 1913
Male adult
Pomera cave, Montello, Veneto Italia
Copyright © Vittorino Monzini
Shenoblemus minusculus Tian & Fang, 2020 Holotype
Male adult
cave Shenxian Dong, Qiaoshan, Xinming, Huangshan, Anhui, 30°23'9.55"N, 118°14'7.66"E, 366 m in altitude, 2016-IV-22, leg. Wenbo Li (SCAU). Image from Fang et al, 2020
Copyright © - - -
Wanoblemus huangshanicus Tian & Li, 2020 Holotype
Male adult
cave Shenxian Dong, Qiaoshan, Xinming, Huangshan, Anhui, 30°23'9.55"N, 118°14'7.66"E, 366 m in altitude, 2016-IV-22, leg. Wenbo Li (SCAU). Image from Fang et al, 2020
Copyright © - - -
Orotrechus venetianus Winkler, 1911
Male adult
Cansiglio, Friuli, Italia
Copyright © Vittorino Monzini
Boldoriella (Boldoriella) concii culminicola Monguzzi, 1982b: 200
Female adult
Valle Imagna, Prealpi Bergamasche, Italia
Copyright © Vittorino Monzini
Duvalioblemus (Shublemus) liyuani Deuve, He & Tian, 2020 Holotype
Male adult
cave Lianhua Dong, Tangba Cun, Bailu Zhen, Pengzhou Shi, Chengdu, Sichuan, 31° 10ʹ5852ʺN 103°56ʹ09.13ʺE, 1064 m in altitude, 9.VI.2019, leg. Li He & Yuan Li (SCAU). Image from Deuve, He & Tian, 2020
Copyright © - - -
Trechodes laophilus Deuve, 2002
Male adult
Thailand, Chiang Mai, Mae lai river, km 27 road 118, 18°54’59’’n -99°14’09’’E, 470 m a.s.l., unspecified local collector
Copyright © Gianni Allegro
Aphaenops (Aphaenops) ochsi orbaraensis Dupre, 1988a: 100
Female adult
Puits de Carralba - Navarra - SP
Copyright © Vittorino Monzini
Uenoaphaenops fani (Ueno, 2003)
Female adult
cave Hua’er Dong, Xiangdingshan, Xiangding, Shiping, Gulin, Luzhou, Sichuan (四川省泸州市古蔺县石屏镇向顶村象顶山华儿洞), 28.028931°N, 106.00716°E, 640 m, 2020-VI-22, leg. Yuan Li & Zhen Wang. Image from Tian & He, 2020
Copyright © - - -
Chu pheggomisetoides Tian & He, 2020 Holotype
Male adult
cave Hanwang Dong, Zhongbai, Zengjia, Chaotian, Guangyuan, Sichuan (四川省广元市朝天区曾家镇中柏村汉王洞), 32.577297°N, 106.106979°E, 1210 m, 2020-VI-07, leg. Li He & Yuan Li (SCAU). Image from Tian & He, 2020
Copyright © - - -
Boreaphaenops liyuani Tian & He, 2020 Holotype
Female adult
cave Hanwang Dong, Zhongbai, Zengjia, Chaotian, Guangyuan, Sichuan (四川省广元市朝天区曾家镇中柏村汉王洞), 32.577297°N, 106.106979°E, 1210 m, 2020-VI-6, leg. Li He, Yuan Li & Yimei Wen (SCAU). Image from Tien & He, 2020
Copyright © - - -
Agonotrechus sinotroglophilus Deuve, 2000
Male adult
Hanwang Dong, Zhongbai, Zengjia, Chaotian, Guangyuan, Sichuan (四川省广元市朝天区曾家镇中柏村汉王洞), 32.577297°N, 106.106979°E, 1210 m, 2018-IV-06, leg. Li He (SCAU). Image from Tian & He, 2020
Copyright © - - -
Miaotrechus heweii Tian, Chen & Ma, 2020 Holotype
Male adult
cave Jingua Dong , Jingua Dong Cun , Longquan Zhen, Danzhai Xian, Qiandongnan Miao & Buyi Autonomous Prefecture, Guizhou Province, China, 26.14059N ° / 107.80400°E, 732 m in altitude, 2019-X-4, leg. Jingli Cheng. Image from Tian, Mingyi, Chen, Mengzhen & Ma, Zijun, 2020
Copyright © Mingyi Tian
Miaotrechus mahua Tian, Chen & Ma, 2020 Holotype
Male adult
Tian, Mingyi, Chen, Mengzhen & Ma, Zijun, 2020
Copyright © Mingyi Tian
Aspidaphaenops dudou Tian & Huang, 2018 Holotype
Male adult
Image from Tian & Huang, 2018
Copyright © Mingyi Tian
Shilinotrechus anthonyi Huang, Tian & Faille, 2020 Holotype
Male adult
China: Yunnan: Qujing: Zhanyi: Panjiang: cave Caiyun Dong (25°46'35"N 103°57'45"E), 2,134 m, coll. Mingyi Tian and Jingli Cheng, 16 August 2015 (Deposited in SCAU). Image from Huang, Tian & Faille, 2020
Copyright © Mingyi Tian
Sinotroglodytes yanwangi Huang, Tian & Faille, 2020 Paratype
Male adult
China: Hubei: Yichang: Wufeng: Changleping: Dawan: cave Da Dong (30°11'12"N 110°42'46"E), 790 m, coll. Mingyi Tian et al., 2 July 2017 (SCAU). Image from Huang, Tian & Faille, 2020
Copyright © Mingyi Tian
Deuveaphaenops (Deuveaphaenops) qimenxicus Tian & Huang, 2017 Holotype
Male adult
Cave I near Qimenxia, Dongping, Huangying Cun, Huangying Xiang, Wulong Xian, 29°10ʹ33”N 107° 42ʹ12”E, 1310 m, 28.VI.2015, leg. Mingyi Tian, Weixin Liu, Xinhui Wang & Mingruo Tang (SCAU). Image from Tian & Huang, 2017
Copyright © Alexander Anichtchenko
Deuveaphaenops (Furongius) gelaophilus Tian & Huang, 2017 Holotype
Male adult
Cave Xianmi Dong, Shuanghe Cun, Shangba Xiang, Daozhen Xian, 28°51ʹ18ʹ’N 107°31ʹ18ʹ’E, 860 m, 22.VII.2015, leg. Mingyi Tian, Jujian Chen, Mingruo Tang & Pingjing Yang (SCAU). Image from Tian & Huang, 2017
Copyright © Mingyi Tian
Qianaphaenops (Qiandongaphaenops) variabilis Tian, Huang & Wang, 2015 Holotype Paratype
Male adult
cave Wanfuchangcheng Dong, in Jiangjun Shan, Pingzhuang Xiang, Cengong County, Guizhou Province, 27°24’50.91”N, 108°35’43.0”E, 830 m, 2014-VI-29, leg. Mingyi Tian, Weixin Liu, Haomin Yin, Sunbin Huang and Xinhui Wang (SCAU). Image from Tian, Huang & Wang, 2015
Copyright © Mingyi Tian
Jiulongotrechus pubescens Tian, Huang & Wang, 2015 Holotype
Male adult
cave Jiulong Dong, Jiulong Village, Yangtou Town, Bijiang District, Tongren City, eastern Guizhou Province, 27°42’29.59” N, 109°19’40.3” E, 450 m, 2014-VI-30, leg. Mingyi Tian, Weixin Liu, Haomin Yin, Sunbin Huang and Xinhui Wang (SCAU). Image from Tian, Huang & Wang, 2015
Copyright © Mingyi Tian
Shilinotrechus intricatus Huang & Tian, 2015 Holotype
Male adult
CHINA: Da Dong cave, Jiuxiang, Yiliang, Kunming, Yunnan Province, 25°5′47.12″ N, 103°23′18.7″ E, 1727 m, 7-VII-2013, Mingyi Tian, Wei Lin,Weixin Liu, Haomin Yin and Sunbin Huang leg. (SCAU). Image from Huang & Tian, 2015
Copyright © Mingyi Tian
Sichuanotrechus dakangensis Huang & Tian, 2015 Holotype
Male adult
CHINA: Zhaojia Dong cave, Xinghuo Cun San Zu, Dakang Zhen, Jiangyou Shi, Sichuan Province, 31°50′15.69″ N, 104°43′22.26″ E, 574 m, 17-VII-2014,Weixin Liu, HaominYin, Sunbin Huang and Xinhui Wang leg. (SCAU). Image from Huang & Tian, 2015
Copyright © Mingyi Tian
Shuangheaphaenops elegans Tian, 2017 Holotype
Male adult
Cave Mahuang Dong, Shuanghe Dong cave system, Wenquan Zhen, Suiyang County, 28°14'32"N, 107°17'24"E, 720 m, X-18-2016, leg. Wenbo Li (SCAU). Image from Tian, 2017
Copyright © Mingyi Tian
Sinaphaenops (Sinaphaenops) mochongensis Tian & Huang, 2015 Holotype
Female adult
Guizhou: Qiannan: Duyun: Mochong: Yueliang Dong, 26u02936.24″ N, 106u26939.46″ E, 901 m, 29-I-2011, leg. Mingyi Tian (SCAU). Image from Tian & Huang, 2015
Copyright © Mingyi Tian
Luoxiaotrechus yini Tian & S. Huang, 2015 Paratype
Male adult
China: Jiangxi: Pingxiang: Lianghua: Gaotan: cave Shuilian Dong, 27u25900″ N, 113u58923″ E, 520 m (SCAU). Image from Tian & S. Huang, 2015
Copyright © Mingyi Tian
Velesaphaenops tarensis Ćurčić & Pavićević, 2018 Holotype
Male adult
Image from Ćurčić et al, 2018
Copyright © - - -
Yunotrechus diannanensis Tian & S.Huang, 2014 Holotype
Male adult
cave named Shiyakou Dadong, 23°06′54″N, 104°17′58″E, 1320 m, Shiyakou Cun, Pojiao Xiang, Maguan County, 3. VII.2013, leg. Mingyi Tian, Wei Lin, Weixin Liu, Haomin Yin and Sunbin Huang (SCAU). Image from Tian & S.Huang, 2014
Copyright © Mingyi Tian
Uenoites reuteri Belousov, Kabak & Schmidt, 2019 Holotype
Male adult
MYANMAR: 1 male, with label data “N Burma, leg. Reuter, ca. 75 km NW Putao, 3700 m NN, 10.08.2006 ” (coll. J. Schmidt).
Copyright © Joachim Schmidt
Pilosaphaenops mengzhenae Huang, Tian & Faille, 2020 Holotype
Male adult
Image from Huang, Tian & Faille, 2020
Copyright © Mingyi Tian
Pilosaphaenops qianzhii Huang, Tian & Faille, 2020 Holotype
Male adult
Image from Huang, Tian & Faille, 2020
Copyright © Mingyi Tian
Pilosaphaenops weiguofui Huang, Tian & Faille, 2020 Holotype
Male adult
Image from Huang, Tian & Faille, 2020
Copyright © Mingyi Tian
Trechus (Archeotrechus) relictus Magrini, Quéinnec & Vigna Taglianti, 2012 Holotype
Male adult
Ethiopia, Prov. Oromia Prov., Bale massif, Goba, Mt. Sgona (Mt. Batu), about 3800 m a.s.l., 8.XI.1973, A. Vigna Taglianti leg. (CVT). Image from Magrini, Quéinnec & Vigna Taglianti, 2012
Copyright © - - -
Pachytrechodes brevis Belousov & Nyundo, 2013 Paratype
Female adult
Tanzania, Udzungwa Mts Nat. Park; forest floor, 1500 m asl; E36º52´00´´S7º45´30´´; 03.viii.1999 Coll. B.A. Nyundo; searching on ground, daytime; B.A. Nyundo Expedition (1998–2001).
Copyright © - - -
Archaeotrechiama embersoni Deuve, 1995 Holotype
Male adult
Image from Lincoln University Entomology Research Collection.
Copyright © - - -
Oarotrechus gracilentus Townsend, 2010 Holotype
Male adult
Image from Lincoln University Entomology Research Collection
Copyright © - - -
Cimmeritodes (Zhecimmerites) zhejiangensis Deuve & Tian, 2015 Holotype
Male adult
Chine, Zhejiang occidental, Quzhou City, Changshan Xian, Xinjiao Xiang, Dongling Cun, grotte Niujiao Dong, 546 mètres, 29°09’44,86’’N - 118°40’26,18’’E, Sunbin Huang & Xinhui Wang leg., 4.X.2014, in coll. South China Agricultural University, à Canton (Chine). Image from Deuve & Tian, 2015
Copyright © Mingyi Tian
Zhijinaphaenops jingliae Deuve & Tian, 2015 Holotype
Male adult
Chine, Guizhou, Guiyang, Xifeng Xian, Shidong Zhen, Musan, grotte Zhangkou Dong, 1203 mètres, 27°04’09,61’’N - 106°33’55,05’’E, Jingli Cheng leg., 4.XII.2014, in coll. South China Agricultural University, à Canton (Chine). Image from Deuve & Tian, 2015
Copyright © Mingyi Tian
Zhijinaphaenops liuae Deuve & Tian, 2015 Holotype
Male adult
Chine, Guizhou, Guiyang, Xifeng Xian, Shidong Zhen, grotte Hejia Dong, 1198 mètres, 27°2’30,89’’N - 106°31’40,38’’E, Weixin Liu, 13.VII.2013, in coll. South China Agricultural University, Canton (Chine). Image from Deuve & Tian, 2015
Copyright © Mingyi Tian
Typhlotrechus bilimekii istrus J. Muller, 1926b: 144
Male adult
Slovenia, Obrov, Golac, Jama Polina peć, V-X.2004, Manfref Egger leg. L=9 mm
Copyright © Ruslan Panin
Tonkinaphaenops impunctatus Faille & Huang, 2019 Holotype
Male adult
Vietnam: Cao Bằng province: Trà Lĩnh district, Quốc Toản commune, Bản Danh: Hang Rắn (snake cave) (=TH06); 22°46’42.3’’N 106°18’49.8’’E, 642m; coll. A. Faille, P.V. Phu, V.T. Tu; 17 October 2018 (MNHN).
Copyright © Mingyi Tian
Tonkinaphaenops jingxicus Huang & Tian, 2019 Holotype
Female adult
China: Guangxi: Jingxi: Ludong: Biaoliang: cave Nianluo Dong; 23°10’5.35”N 106°16’13.98”E, 927 m; coll. Sunbin Huang, Dianmei Wang, Mengzhen Chen, Ruiqi Han; 1 February 2018; (SCAU).
Copyright © Mingyi Tian
Tonkinaphaenops anthonyi Faille & Tian, 2019 Holotype
Male adult
Vietnam: Cao Bằng province: Hạ Lang district, Đồng Loan commune, Lũng Rúm: Hang Dõi (bat cave); 22°44’47.7”N 106°44’35.9”E, 386 m; 20 October 2018; coll. A. Faille, P.V. Phu, V.T. Tu; (MNHN).
Copyright © Mingyi Tian
Agonotrechus spinangulus Belousov, Kabak & Liang, 2019 Paratype
Female adult
China, Sichuan, Muli County, Liziping Xiang , Kangwu Pass , mixed forest, N 28.12630, E 101.15987 2012. VI.5, Huang Hao leg. ( IOZ)
Copyright © - - -
Trechus (Trechus) cratocephalus Belousov & Kabak, 2019 Paratype
Male adult
CH, Xinjiang , W of Narat Mt. R, 7–9 km SE of Sarkhob village, 42°47´15´´ N / 81°14´45´´ E— 42°46´00´´ N / 81°15´20´´ E, 2200–2550 m, 23.07.2003, Kabak I.I. leg.” ( ZISP)
Copyright © Ilya Kabak
Trechus (Trechus) aghiazicus Belousov & Kabak, 2019 Paratype
Male adult
China, Xinjiang, W of Narat Mt. R, sources of right tributary of Tshong-Muyuny River near Biyke Pass , 42º40´20´´ N / 81º20´50´´ E— 42º38´40´´ N / 81º21´30´´ E, 3100–3800 m, 26.07.2003, Kabak I. leg.” ( ZISP)
Copyright © Ilya Kabak
Trechus (Trechus) tsanmensis Belousov & Kabak, 2019 Paratype
Male adult
China, Xinjiang, Narat Mt. Range, sources of Zhasylkol River (left trib. of Za- nma River ) 3000–3200 m, 24.06.2001, Kabak I.I. leg.” [43º12´00´´ N / 83º39´00´´ E— 43º11´40´´ N / 83º39´00´´ E] ( ZISP)
Copyright © Ilya Kabak
Trechus (Trechus) torgaut Belousov & Kabak, 2019 Paratype
Male adult
China, Xinjiang, sources of Kunges River, left bank near Adunkur 1 Pass , ESE of Kunges-Linchan, 2500–3100 m, 13.07.2001, Kabak I.I. leg.” [43º12´00´´ N / 84º47´40´´ E— 43º11´15´´ N / 84º47´20´´ E] ( ZISP)
Copyright © Ilya Kabak
Adriaphaenops antroherponomimus (Noesske, 1928) Holotype
Male adult
Image from Lohaj et al, 2016
Copyright © - - -
Adriaphaenops pretneri Scheibel, 1935b: 35 Holotype
Female adult
Image from Lohaj et al, 2016
Copyright © - - -
Adriaphaenops stirni (Pretner, 1959) Lectotype
Male adult
Image from Lohaj et al, 2016
Copyright © - - -
Adriaphaenops zupcensis (Pavićević, 1990)
Male adult
Topotype. Image from Lohaj et al, 2016
Copyright © - - -
Ishikawatrechus bidilatatus Sugaya & Yamasako, 2014 Holotype
Male adult
[ EHIME: JAPAN] / Ôyashiki, Kawano-uchi / Tôon-shi, alt. ca. 530 m / 14. V. 2012 / K. Sugaya leg. Image from Sugaya & Yamasako, 2014
Copyright © - - -
Trechus (Epaphius) dorsistriatus A. Morawitz, 1862
Male adult
Russia, Primorskii reg., right tributary of Poima riv., Chernye Gory, 30.7.1999, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko
Xiangxius jiangi Tian & X. Huang, 2021 Holotype
Male adult
cave Tangle Dong, suburb of Jishou, Xiangxi Tujia & Miao Autonomous Prefecture, western Hunan Province, 28°18’21”N / 109°39’15”E, 270 m in altitude, 2018-I-15, leg. Xinglong Huang (SCAU)
Copyright © Mingyi Tian
Guizhaphaenopsodes solidior Tian & X. Huang, 2021 Holotype
Male adult
cave Tangle Dong, suburb of Jishou, Xiangxi Tujia & Miao Autonomous Prefecture, west Hunan Province, 28°18’21”N /109°39’15”E, 270 m in altitude, 2018-I-15, leg. Xinglong Huang (SCAU).
Copyright © Mingyi Tian
Sinotroglodytes hefengensis Tian & Li, 2021 Holotype
Male adult
the limestone cave Qizimei Dong, Qizimeishan, Chunmuyun, Hefeng County, Enshi Tujia & Miao Autonomous Prefecture, southwestern Hubei Province, Central China, 30°02’30”N /109°47’09”E, 1820 m in altitude, 2020-VI-18, leg. Chenliang Li (SCAU).
Copyright © Mingyi Tian
Typhlotrechus bilimekii droveniki M. Etonti, 1995: 83 Paratype
Female adult
Copyright © Vittorino Monzini
Panaphaenops guixicus Tian, Huang & Ma, 2021 Holotype
Male adult
the cave Zimu Dong, Danxiashan, Danxia Town, Panzhou, Liupanshui City, Guizhou Province, 25°40'0.0"N 104°37'3.6";E, 1791 m in altitude, 24.VIII.2020, leg. Yunchun Li & Yucheng Lin.
Copyright © Mingyi Tian
Panaphaenops guixicus Tian, Huang & Ma, 2021 Holotype
Male adult
the cave Zimu Dong, Danxiashan, Danxia Town, Panzhou, Liupanshui City, Guizhou Province, 25°40'0.0"N 104°37'3.6";E, 1791 m in altitude, 24.VIII.2020, leg. Yunchun Li & Yucheng Lin.
Copyright © Mingyi Tian
Casaleaphaenops liyunchuni Tian, Huang & Ma, 2021 Holotype
Male adult
cave Hei Dong, Longjiang Cun, Shenxi Town, Honghuagang District, Zunyi City, northern Guizhou Province, 2736°3.2“N 10658°4.8“E, altitude 822 m, 22.VIII.2020, leg. Yunchun Li & Yucheng Lin.
Copyright © Mingyi Tian
Casaleaphaenops liyunchuni Tian, Huang & Ma, 2021 Holotype
Male adult
cave Hei Dong, Longjiang Cun, Shenxi Town, Honghuagang District, Zunyi City, northern Guizhou Province, 2736°3.2“N 10658°4.8“E, altitude 822 m, 22.VIII.2020, leg. Yunchun Li & Yucheng Lin.
Copyright © Mingyi Tian
Orotrechus fabianii Gestro, 1900
Male adult
Colli Berici, Lonigo Vicenza, Italia
Copyright © Vittorino Monzini
Trichaphaenops (Trichaphaenops) sollaudi Jeannel, 1916
Male adult
Topotype
Copyright © Vittorino Monzini
Trichaphaenops (Trichaphaenops) gounellei gounellei (Bedel, 1880)
Male adult
Photo by Erminio Piva
Copyright © Vittorino Monzini
Anophthalmus (Duvaliaphaenops) raffaldianus (Lemaire, 1981)
Male adult
Photos Erminio Piva
Copyright © Vittorino Monzini
Anophthalmus (Trichaphaenops) obesus (Abeille de Perrin, 1886)
Male adult
Photo Erminio Piva
Copyright © Vittorino Monzini
Sinotrechiama yunnanus Belousov, Kabak & Liang, 2019 Holotype
Female adult
China, Yunnan, Dayao, Santai Town , pass to Xiaobaicaol- ing, 3105 m, N 26.05615, E 101.05448 ”; “ 2012.V.29, day, under stone, mixed forest, Shi Hongliang leg., Institute of Zoology , CAS.
Copyright © - - -
Uenotrechus gejianbangi Tian et Wei, 2017 Holotype
Male adult
( SCAU), Cave Ji Dong (also called Shuiku Dong ), Neiwen Tun , Sancai Cun , Shuiyuan Zhen , Huanjiang Maonan Autonomous County, Guangxi Zhuang Autonomous Region, 24°49.430'N / 108°05.504'E, 290 m, XII-02-2016, leg. Arnaud Faille, Jujian Chen and Sunbin Huang.
Copyright © Mingyi Tian
Uenotrechus nandanensis Deuve & Tian, 2010 Holotype
Male adult
Cave Dadongqiao Dong, Encun , Nandan County, Guangxi, XII-27-2008, leg. Zhihong Xue, Wei Lin and Lei Gao
Copyright © Mingyi Tian
Uenotrechus deuvei Tian et Chen, 2017 Holotype
Male adult
( SCAU), Cave Longmu Dong I, Longmu Cun, Sanziyang Xiang, Du’an Yao Autonomous County, northern Guangxi Zhaung Autonomous Region, southern China, 24°24' 52.59" N / 108°00' 09.69" E, 324 m, VII-28-2015, leg. Xinhui Wang, Jujian Chen & Minruo Tang.
Copyright © Mingyi Tian
Puertrechus daxueshanicus Belousov & Kabak, 2014 Holotype
Male adult
CH, Yunnan Province, E sl. of Mt Daxueshan, W of Niutoushan Village, 24º07´15´´ N / 99º39´12´´ E, H=3375 m, 3.06.2010, Belousov & Kabak leg.(ZISP).
Copyright © - - -
Puertrechus daxueshanicus Belousov & Kabak, 2014 Paratype
Female adult
CH, Yunnan Province, E sl. of Mt Daxueshan, W of Niutoushan Village, 24º07´15´´ N / 99º39´12´´ E, H=3375 m, 3.06.2010, Belousov & Kabak leg.(ZISP).
Copyright © - - -
Junnanotrechus wrasei Belousov & Kabak, 2014 Holotype Paratype
Male+Female adult
China (N Yunnan), Dali Bai Nat. Aut. Pref., Diancang Shan, pass 43 km NW Dali, 3104 m, 25 ° 59 ´ 33.5 ´´ N / 99 ° 52 ´ 12.5 ´´ E, (pasture & shrubs, litter, moss and mushrooms sifted), 23.08. 2009, (M. Schülke) ( ZISP)
Copyright © - - -
Junnanotrechus oblongus Belousov & Kabak, 2014 Holotype Paratype
Male adult
CH, Yunnan Province, E sl. of Daxueshan, W of Niutoushan, 24°07´15´´N / 99°39´12´´E, H= 3375 m, 3.06. 2010, Belousov & Kabak leg.( ZISP)
Copyright © - - -
Junnanotrechus schuelkei Belousov & Kabak, 2014 Holotype
Female adult
China (Yunnan), Pu’er Pref., Ailao Shan, 37 km NW Jingdong, 24 ° 45 ´ 12 ´´ N / 100 ° 41 ´ 24.5 ´´ E, 2300 m, (devastated forest remnant, litter/dead wood sifted), 13.09. 2009, (M. Schülke) ( ZISP)
Copyright © - - -
Junnanotrechus microps Ueno & Yin, 1993
Male adult
Diancang Shan
Copyright © - - -
Junnanotrechus elegantulus Belousov & Kabak, 2014 Holotype Paratype
Male+Female adult
CH, Yunnan Province, Mt. Daxueshan E Mengsa Town, 23°42´26´´N / 99°47´26´´E, H= 3045 m, 30.05. 2010, Belousov & Kabak leg.( ZISP)
Copyright © - - -
Junnanotrechus koroleviellus Belousov & Kabak, 2014 Holotype Paratype
Male+Female adult
CH, Yunnan Province, W of Yunxian Town, 24°28´29 N / 99°58´23´´E, H= 2965 m, 17.06. 2011, Belousov, Kabak, Korolev leg.( ZISP)
Copyright © - - -
Cimbrodytes heterognathus Piva, 2021 Holotype
Male adult
Photo Erminio Piva
Copyright © Vittorino Monzini
Ckacus youcha Tian & Huang, 2021 Holotype
Male adult
Luoshui Dong, a vertical cave in Anlong Xian, southwestern most of Guizhou Province, Southwest China
Copyright © Mingyi Tian
Yalongaphaenops erwini Belousov & Kabak, 2021 Holotype
Male adult
China, Sichuan Province, SW Mianning Town; 28°15'27"N, 101°43'51"E; H = 3805 m a.s.l.; 11.07.2011; Belousov & Kabak leg. (ZIN).
Copyright © Ilya Kabak
Agonotrechus reflexicollis (Ueno, 1977)
Male adult
Bhutan, 3 October 2017, Yendu Pang ‘Base camp' on North bank Yendu Chu, 3940 m, 27°21 32.0''N, 90°20' 04.7''E
Copyright © Joachim Schmidt
Trechus (Trechus) tatai Reboleira & Ortuño, 2010
Female adult
Algar do Javali Cave, Montejunto karst massif, Portugal. Image from Reboleira & Eusébio, 2021
Copyright © - - -
Duvalius (Hungarotrechus) roubali Jeannel, 1926b: 14
Female adult
W Ukraine, NE Carpathians,Chornogora Mt Range, alpine zone, under a stone-boulder, 22.VI.2018, R.Panin leg.
Copyright © Ruslan Panin
Duvalius (Hungarotrechus) roubali Jeannel, 1926b: 14
Male adult
W Ukraine, NE Carpathians, Chornogora Mt Range, alpine zone, 22.VI-14.IX.2018, traps, R.Panin leg., L=5.6 mm, photo by D. Khrapov
Copyright © Ruslan Panin
Duvaliopsis pilosella pilosella L. Miller, 1868 (as rybinskii Knirsch, 1924)
Male adult
W Ukraine, NE Carpathians, Chornogora Mt Range, Pozhyzhevska polonyna, h=1430m, 30.VI -3.IX.2016, Panin R. & Kanarsky Yu.leg. L= 4.1 мм
Copyright © Denis Khrapov
Trechus (Trechus) pseudomontanellus Rizun, 1994: 30
Female adult
W Ukraine, NE Carpathians, Transcarpathian Reg., Marmarosh Mts., near Lug vill., bank of Kuziy (Kizi) stream, h=400 m, 2.VII.2021, Panin R. leg. L= 4.5 mm
Copyright © Denis Khrapov
Trechus (Trechus) amplicollis Fairmaire, 1859h: CXIIX
Male adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletskyi forest, near Pyatnychany vill., 28.V-9.VI.2013, Panin R. leg., L=5.5mm
Copyright © Denis Khrapov
Duvalius (Biharotrechus) transcarpathicus Shilenkov et Rizun, 1989: 83
Male adult
W Ukraine, NE Carpathians, Tiachiv distr., cave near Uglya vill., 30.IV-29.IX.2017, Panin R. & Kanarsky Yu. leg. L=5.1mm
Copyright © Denis Khrapov
Orotrechus jamae C. Etonti et M. Etonti, 1979: 27
Male adult
NE Italy, Belluno, Prealps, Cansiglio plateau, Bus dell'Ors, 19.V.2013, L=6.2mm
Copyright © Denis Khrapov
Qianlongius zhoui Tian & Jia, 2021 Holotype
Male adult
cave Qianlong Dong, Qianlong, Wuluo, Songtao, Tongren, Guizhou Province, 28.16° N, 108.84°E, 908 m, 2021-IV-2, leg. Yi Zhao, Xinyang Jia and Mingyi Tian (SCAU). Image from Tian et al, 2021
Copyright © Mingyi Tian
Wulongius qilinger Tian & Huang, 2021 Holotype
Female adult
cave Qiankou Dong, Tongluo, Jiangkou, Wulong, Chongqing, 29.32°N, 107.91°E, 1103 m, 2021-IV-14, leg. “Wu Ya” (a nickmane of Mr Hongying Wu) & Mingyi Tian, in SCAU.
Copyright © Mingyi Tian
Qianotrechus (Qianotrechus) congcongae Tian & Zhao, 2021 Holotype
Male adult
cave Shigao Dong, Hexi, Nanchuan, Chongqing, 28.82° N, 107.32°E, 729 m, 2021-IV-13, leg. Yi Zhao, Xinyang Jia and Mingyi Tian, in SCAU.
Copyright © Mingyi Tian
Blemus alexandrovi (Lutshnik, 1915) Holotype
Female adult
Mandtschuria, Chandaoehedzy
Copyright © Kirill Makarov
Trechus (Epaphius) nigricornis nigricornis Holotype
Female adult
Kiachta / 100 v[erst] v[on] Kiachta
Copyright © Kirill Makarov
Trechus (Epaphius) nigricornis nigricornis Holotype
Female adult
Kiachta / 100 v[erst] v[on] Kiachta
Copyright © Kirill Makarov
Trechus (Epaphius) nigricornis arsenjevi Jeannel, 1962c: 178 Paratype
Female adult
Paratypus of Epaphius arsenjevi Jeannle, 1962: Lower Amur riv. lake Kisi 12.VIII.1928 leg.Formosov
Copyright © Kirill Makarov
Trechus (Epaphius) nigricornis arsenjevi Jeannel, 1962c: 178 Paratype
Female adult
Paratypus of Epaphius arsenjevi Jeannle, 1962: Tumen-ula River, Russia-Korean frontier 14.VII.1913 leg. Tsherski
Copyright © Kirill Makarov
Trechus (Epaphius) nigricornis arsenjevi Jeannel, 1962c: 178 Holotype
Male adult
Holotype of Epaphius plutenkoi Lafer, 1989: Primor.[sky Krai], Khasan, env. Golubinogo Utesa [Pigeon Cliff], 23.7.72, G.Lafer
Copyright © Kirill Makarov
Anophthalmus (Anophthalmus) meggiolaroi P. Moravec & Lompe, 2003 (as charon)
Male adult
Copyright © Vittorino Monzini
Trechus (Trechus) kouroshi Muilwijk, Seiedy & Wrase, 2021 Holotype
Male adult
Iran, Kohgiluyeh and Boyer-Ahmad / Suq cave, 812 m/ 30°49ˊ25.9´´N, 50° 25ˊ37.9´´ E, 5.vi.2019 / S. Rahideh, Y. Fatemii & J. Muilwijk leg. deposited in HMIM
Copyright © Jan Muilwijk
Duvalius (Duvalius) nezelensis Muilwijk & Malek Hosseini, 2021 Holotype
Male adult
Iran Kohgiluyeh Va Boyer Ahmad, Zagros/ Nezel cave, 12.vi.2018 / M. Porebrahim, S. Azizi, Z. Rozbehi, S. Rahideh, & J. Muilwijk leg.”
Copyright © Jan Muilwijk
Duvalius (Duvalius) achaemenius Muilwijk & Malek Hosseini, 2021 Holotype
Male adult
Iran, Kohgiluyeh Va Boyer Ahmad / Suq cave near Suq / 16.vi.2018 / S. Rahideh & J. Muilwijk leg. (HMIM)
Copyright © Jan Muilwijk
Trechus (Trechus) reebi Queinnec & Olliver, 2013 Paratype
Male adult
Ethiopia, Oromia, Bale Mts, NP Sanatee Plateau, 06°46′259″N / 039°45′282″E, h=3460 m.15.V.2012, leg. M.Egger
Copyright © Denis Khrapov
Duvalius (Hungarotrechus) corpulentus Weise, 1875: 356
Male adult
Transcarpathian Region, Rakhiv Distr., Chornogora Mt. Range, S slope h=1600m, bank of stream, 6-8.VI.2016, Panin R. leg., L=5.55 mm
Copyright © Denis Khrapov
Dongodytes (Dongodytodes) brevipenis Tian, Yin & Huang, 2014
Male adult
Copyright © Vittorino Monzini
Zhijinaphaenops jingliae Deuve & Tian, 2015
Male adult
Guizhou: Guiyang, Xifeng, Xishan, Shenli, Mafen Dong cave (息烽县西山镇马粪洞), 27°04'N, 106°37'E, 1157 m, 2020–II–29, leg. Guangyuan Cheng
Copyright © Mingyi Tian
Haixiaphaenops jinxiaohongae Tian, Cheng & Huang, 2021 Holotype
Male adult
Guizhou, Qingzhen, Anliu, Yangtianwo, Dawan Dong cave (贵州省清镇市暗流镇大湾洞), 26°52'N, 106°24'E, 941 m, 2020–IV–19, leg. Xiaohong Jin & Guangyuan Cheng, in SCAU
Copyright © Mingyi Tian
Haixiaphaenops jinxiaohongae Tian, Cheng & Huang, 2021 Paratype
Male adult
Guizhou, Qingzhen, Anliu, Ximi, Changtu Dong cave (贵州省清镇市暗流镇长土洞), 26°51‘N, 106°21‘E, 1249 m, 2020–VI–27, leg. Chenggang Wang, Xiaohong Jing, Guangyuan Cheng, Yi Zhao & Mingyi Tian, SCAU.
Copyright © Mingyi Tian
Zhijinaphaenops zhaofeii Tian, Cheng & Huang, 2021
Male adult
Guizhou, Guiyang, Xifeng, Jiuzhuang, Changtu Dong cave (贵州省贵阳市息烽县九庄镇张口洞), 27°11'N, 106°29'E, 1008 m, 2019–VI–08, leg. Jingli Cheng & Mingyi Tian.
Copyright © Mingyi Tian
Trechus (Trechus) pilonensis Toribio, 2014
Male adult
Spain, Raicedo-Colunga (M. del Sueve), Asturias, 3.12.2016, M. Toribio leg. (cAA)
Copyright © Alexander Anichtchenko
Trechus (Trechus) xiongmao Belousov & Kabak, 2021 Paratype
Male adult
China, NW Sichuan, NW of Lixian, 9 km W of Shangmeng, scree above timber-line 3600–3800 m, 23.07.2002, Belousov I. & Kabak I. leg.” [31°40´30´´N / 103° 02´25´´E].
Copyright © - - -
Trechus (Trechus) comma Belousov & Kabak, 2021 Paratype
Male adult
China, NW Sichuan, NE of Lixian, N Tonghua, basin of river near Pingshitou, S of Shibanpengzi, H~3900 m, 22.08.2002, Belousov I. & Kabak I. leg.” [31°44´41´´N / 103°22´50´´E].
Copyright © - - -
Trechus (Trechus) sharovae Belousov & Kabak, 2021 Paratype
Male adult
China, NW Sichuan, NE of Lixian, N of Tonghua, basin of river near Pingshitou, H~3300–3500 m, 19.08.2002, Belousov I. & Kabak I. leg.” [31°43´49´´N / 103°23´11´´E — 31°44´15´´N / 103°22´54´´E].
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Orotrechus vicentinus juccii Pomini, 1940: 82
Male adult
N-Italy, Veneto(VR), Mts Lessini, Cave of Mt. Solane, 5.IX.1984, R. Monguzzi leg., L=4.4 mm
Copyright © Denis Khrapov
Duvalius (Hungarotrechus) ruthenus ruthenus Reitter, 1878: 37
Female adult
W Ukraine, NE Carpathians, Chornagora Mt Range, alpin zone, V-2.VII.2016, Panin R. & Kanarsky Yu. leg., L= 4.7 mm
Copyright © Denis Khrapov
Geotrechus sarpedon Faille, Fresneda & Bourdeau, 2013 Holotype
Male adult
(collection C. Bourdeau, Rebigue), [label data: voucher number ZSM-L445] Camou - grotte inférieure d´Oxibar (=Idobeltse Ko Karbia), (France, Pyrénées-Atlantiques F-64), 29.VII.2011, leg. C. Bourdeau.
Copyright © - - -
Trechus (Trechus) mattisi Schmidt & Faille, 2018 Paratype
Male adult
ETHIOPIA, Bale Mts, Harenna Forest, E Rira, alt. 3150 m, 06°44′51″ N, 39°46′40″ E, Feb. 2007
Copyright © Joachim Schmidt
Trechus (Trechus) minitrechus Schmidt & Faille, 2018 Paratype
Male adult
ETHIOPIA, Oromia, Mt Enkuolo NE slope, alt. 3200–3300 m, 07°23′45″ N, 39°22′27″ E, 8 Dec. 2016
Copyright © Joachim Schmidt
Trechus (Trechus) transversicollis Schmidt & Faille, 2018 Holotype
Male adult
ETHIOPIA, Bale Mts, Harenna Forest, E Rira, alt. 2690 m, 06°43′25″ N, 39°45′25″ E, Feb. 2007
Copyright © Joachim Schmidt
Trechus (Trechus) nanulus Schmidt & Faille, 2018 Paratype
Male adult
ETHIOPIA, Oromia, S slope Bale Mts below Rira, alt. 2380 m, 06°43′17″ N, 39°43′14″ E, Dec. 2006
Copyright © Joachim Schmidt
Aphaenops (Cerbaphaenops) cerberus bruneti Jeannel, 1926a: 237
Male adult
S France, Ariège, Moulis, Grotte de Lique, 6.IX.2014, Branes leg.
Copyright © Denis Khrapov
Duvalius (Duvalius) longhii larianus B. Bari, 1940
Female adult
Buco della Nicolina cave, CO, Lombardia, Italia
Copyright © Vittorino Monzini
Aphaenops (Aphaenops) jeanneli (Abeille de Perrin, 1905)
Male adult
Pyr. Atl., Mines - F64
Copyright © Vittorino Monzini
Duvalius (Biharotrechus) redtenbacheri bihariensis (Csiki, 1911)
Male adult
Copyright © Vittorino Monzini
Duvalius (Neoduvalius) lohaji Queinnec & Ollivier, 2021 Holotype
Male adult
Mracna Pecina, Livno, Prolog Planina, Bosnia and Herzegovina
Copyright © - - -
Duvaliomimus (Duvaliomimus) watti Britton, 1958 Holotype
Male adult
Copyright © - - -
Himalotrechus humeratus Belousov, Kabak & Schmidt, 2019 Holotype
Male adult
India, Arunachal Pradesh, N of Bomdila, NW Chander, 10–15 km NW of, 27º25´ N 92º22´ E, ca 3200 m, 18 VI.2008 / Reuter (coll. J. Schmidt).
Copyright © - - -
Velebitaphaenops giganteus Casale & Jalzic, 2012
Female adult
Image from cavebiology.net
Copyright © Creative Commons
Scotoplanetes aquacultor Lakota, Lohaj & Dunay, 2010
Male adult
Image from cavebiology.net
Copyright © Creative Commons
Aphaenops (Cerbaphaenops) hustachei hustachei Jeannel, 1917
Male adult
grotte de l'Haiovat de Pelor, houtes Pyrenees, France
Copyright © Vittorino Monzini
Boldoriella (Insubrites) pesarinii Sciaky, 1982: 25
Male adult
Mt. Pora, 1800 m., Bratto BG Italia
Copyright © Vittorino Monzini
Duvalius (Biharotrechus) reufi Pavicevic & Popovic, 2003
Male adult
Image from Ćurčić et al, 2022
Copyright © - - -
Duvalius (Duvalius) suvodolensis (S. Ćurčić, Brajković & B. Ćurčić, 2003)
Male adult
Image from Ćurčić et al, 2022
Copyright © - - -
Duvalius (Duvalius) suvoborensis Pavecevic & Popovic, 2003
Male adult
Image from Ćurčić et al, 2022
Copyright © - - -
Duvalius (Duvalius) sturanyi (Apfelbeck, 1904)
Female adult
Image from Ćurčić et al, 2022
Copyright © - - -
Duvalius (Duvalius) javorensis (S. Ćurčić, Brajković, B. Ćurčić & Mitić, 2003)
Male adult
Image from Ćurčić et al, 2022
Copyright © - - -
Duvalius (Duvalius) bolei Pralmer, 1963a: 144
Female adult
Image from Ćurčić et al, 2022
Copyright © - - -
Boldoriella (Insubrites) chiarae Monguzzi, 1982b: 211
Male adult
Mt. Guglielmo Brescia Italia
Copyright © Vittorino Monzini
Duvalius (Neoduvalius) djokovici Ćurčić, Pavićević & Vesović, 2022 Holotype Paratype
Male+Female adult
Copyright © - - -
Duvalius (Duvalius) casellii carrarae Jeannel, 1928a: 617
Female adult
Tana della Bastiola cave, Massa, Italia, leg. Lanza 20-X-1942
Copyright © Vittorino Monzini
Doderotrechus casalei Vigna Taglianti, 1969
Female adult
Valmala, Piemonte, Italia, leg. A. Casale
Copyright © Vittorino Monzini
Pheggomisetes globiceps ninae S. Ćurčić, H. Schonmann, Brajković, B. Ćurčić & Tomić, 2004
Female adult
Serbia, Hodzina Dupka pit, Dimitrovgrad
Copyright © Vittorino Monzini
Duvalioblemus (Duvalioblemus) arnaudi Deuve, 2022 Holotype
Female adult
Chine, Sichuan, Mt Gongga Shan, versant nordoriental, 3554 m, 29°55’31’’N/101°58’46’’E (V. Grebennikov leg.), in coll. Muséum d’Histoire Naturelle de Stuttgart (Allemagne).
Copyright © Thierry Deuve
Birmaphaenops brehieri Deuve, 2017 Holotype
Male adult
Birmanie, État Shan, environs d’Ywangan, grotte Kyauk Khaung (“Stone Cave”), 21°11’28,2’’N - 96°33’9,4’’E (Franck Bréhier, 30.XI.2015, lot n°MY15-17/04), in coll. Muséum national d’Histoire naturelle, Paris.
Copyright © Thierry Deuve
Lanxangaphaenops augustini augustini Holotype
Female adult
, Laos, Province de Khammouane, Ban Thongxam, Tham Nam Ock, 247 m, 17°35’24,108’’N - 105°50’7,26’’E (Bernard Lips et Alexis Augustin leg., 3.III.2016, “flacon 11925”), in coll. Muséum national d’Histoire naturelle, Paris.
Copyright © Thierry Deuve
Lanxangaphaenops augustini steineri Deuve, 2017 Holotype
Female adult
Laos, Province de Khammouane, District de Boualapha, Tham Nguen Maï, 17°22’13,512’’N - 105°53’36,707’’E (Helmut Steiner leg., 24.II.2016, “spécimen 128/16”), incoll. Muséum national d’Histoire naturelle, Paris
Copyright © Thierry Deuve
Deuveopsis (Deuveopsis) lobeliae Quéinnec & Ollivier, 2021 Paratype
Male adult
Ethiopia, Amhara, Mount Choke, station I (afro-alpine belt, rocks below Lobelia), 10˚41’54’’N, 037˚50’21’’E, 3950 m a.s.l., 21.VIII.2011, E. Quéinnec & C. Reeb leg.
Copyright © Eric Quéinnec
Aethiopsis chioriae Quéinnec & Ollivier, 2021 Paratype
Male adult
Ethiopia, Amhara, Abuna Yusef Mountains (afro-alpine belt, rocks below a small cliff near the summit), 12˚08’32’’N, 39˚10’59’’E, 4190 m a.s.l., 5.I.2013, R. Chiori, E. Ollivier, E. Quéinnec, C. Reeb leg.
Copyright © Eric Quéinnec
Nilotrechus niloticus Quéinnec & Ollivier, 2021 Paratype
Male adult
Ethiopia, Amhara, Mount Choke, station III (afro-alpine belt, lawn in open area), 10˚38’17’’N, 037˚50’05’’E, 3760 m a.s.l., 22.VIII.2011, E. Quéinnec & C. Reeb leg.
Copyright © Eric Quéinnec
Afrotrechus (Abyssiniopsis) bunae Quéinnec & Ollivier, 2021 Paratype
Male adult
Ethiopia, Oromia province, Belleta forest (degraded forest, 25 km WSW of Jimma), 7˚31’50’’N, 36˚31’46’’E, 1800 m a.s.l., VIII.2011, E. Quéinnec & C. Reeb leg.
Copyright © Eric Quéinnec
Tianzhuaphaenops jinshanensis D. Zhao & Tian, 2016 Holotype
Male adult
Cave Jinshan Rongdong, Luihe Vallige, Bangdong Town, Tianzhu County, Qiandongnan Miao and Dong Autonomous Prefecture, Guizhou Province, 26°57'42" N/109°12'24" E, 460 m, VIII-21-2015, Mingyi Tian & Jingli Cheng leg., deposited in the insect collections of South China Agricultural University, Guangzhou (SCAU)
Copyright © Mingyi Tian
Libotrechus duanensis Lin & Tian, 2014 Holotype
Male adult
CHINA: Guangxi Zhuang Autonomous Region: Du’an Yao Autonomous Xian: Longfu Xiang: Longfu Cun: Shangmo Tun, a cave named Shuiyuan Dong, 24°11.335′N 107°49.865′E, 509 m, 28 June 2013, Wei Lin leg., deposited in SCAU.
Copyright © Mingyi Tian
Youtrechus zijunae Tian, Huang & Jia, 2023 Holotype
Male adult
29.73 °N / 114.24 °E, 160 m, a limestone cave called Feixian Dong, Xian’an District, Xianning, Hubei, 2019. VII. 13, Mingyi Tian, Mengzhen Chen, Zijun Ma & Jingli Cheng leg., in SCAU.
Copyright © Mingyi Tian
Wumengius fengi Tian, Huang & Jia, 2023 Holotype
Male adult
26.31 °N / 104.70 °E, 1435 m, the cave Da Dong near village Tuoluo, Hongguang, Duge, Liupanshui, Guizhou, 2019. VII. 6, Zegang Feng & Chen Zhang leg., in SCAU.
Copyright © Mingyi Tian
Chenotrechus parvulus Tian, Huang & Jia, 2023 Holotype
Male adult
the cave Xiniuyan, Guhang, Luxin, Wuxuan, Laibin, Central Guangxi Zhuang Autonomous Region, Southern China, 109.55°E, 23.56°N, 115 m, 2021-I-15, leg. Mingyi Tian, Weixin Liu & Yi Zhao, in SCAU
Copyright © Mingyi Tian
Sinaphaenopoides zhaoyiae Tian, Huang & Jia, 2023 Holotype
Female adult
Zhongguo Dong, Tianzidi Cun, Suxian Qu, Chenzhou Shi, 2019-XII-24, leg. Yi Zhao, in SCAU
Copyright © Mingyi Tian
Ecotrechus ruruiae Tian, Huang & Jia, 2023 Holotype
Female adult
the cave Laba Dong, Shaba, Ziyun Buyi & Miao Autonomous County, Guizhou Province, 25.84 N, 106.08 E, 1450 m in altitude, 2022-V-16, Rurui Ye leg., in SCAU
Copyright © Mingyi Tian
Hurites yunxianicus Tian, Huang & Jia, 2023 Holotype
Male adult
Yunxian Dong, Longhui County, Shaoyang, Central Hunan, 27.28 N, 110.89 E, 425 m, 2019-IV-30, leg. Mingyi Tian, Mengzhen Chen & Zijun Ma, in SCAU
Copyright © Mingyi Tian
Pangaphaenops seraphicus Tian, Huang & Jia, 2023 Holotype
Male adult
Cave I, Suolue Town, Bama Yao Autonomous County, Hechi Shi, Guangxi Zhuang Autonomous Region, Southern China, 113.36°E, 23.16°N, 532 m, 2022-VI-21, leg. Jingli Cheng, in SCAU.
Copyright © Mingyi Tian
Zunotrechus zhaojieyui Tian, Huang & Jia, 2023 Holotype
Male adult
Da Dong, Bali, Bozhou, Zunyi Shi, Guizhou Province, 23.16 N, 113.36 E, 922 m, 2019-IV-2, Mingyi Tian leg., in SCAU.
Copyright © Mingyi Tian
Dongodytes (Dongodytodes) pingjingae Tian, Huang & Jia, 2023 Holotype
Male adult
Xiannü Dong, Guhe, Dahua Yao Autonomous County, Hechi Shi, Guangxi Zhuang Autonomous Region, 23.90 N, 107.74° E, 240 m, VII. 9. 2016, Mingyi Tian, Pingjing Yang, Mingruo Tang & Jingli Cheng leg., in SCAU
Copyright © Mingyi Tian
Dongodytes (Dongodytes) mingruoae Tian, Huang & Jia, 2023 Holotype
Male adult
Cave I, Jiudu Xiang, Du’an Yao Autonomous County, Hechi Shi, Guangxi Zhuang Autonomous Region, southern China, 24.27°N / 108.23°E, 333 m in altitude, 2018-IX-25, leg. Mingyi Tian, Sunbin Huang & Mengzhen Chen, in SCAU
Copyright © Mingyi Tian
Deuveaphaenops (Furongius) huangheae Tian, Huang & Jia, 2023 Holotype
Female adult
Shisanya Dong, Jinming, Sanqiao, Daozhen, northern Guizhou, 2022-IV-12, leg. Huanghe Zhou, in SCAU.
Copyright © Mingyi Tian
Shilinotrechus zhangfani Tian, Huang & Jia, 2023 Holotype
Male adult
cave Xianren Dong, Xi’er Township, Mile Shi, Honghe Hani & Yi Autonomous Prefecture, Yunnan Province, 103.20 E,24.48 N, 1901 m above sea level, 2020-VIII-17, Mingyi Tian & Jingli Cheng leg., in SCAU.
Copyright © Mingyi Tian
Guizhaphaenops wanghaoi Tian, Huang & Jia, 2023 Holotype
Male adult
Weixiao Tiankeng, Gaoling, Qinglong County, Qianxinan Buyi & Miao Autonomous Prefecture, southwestern Guizhou Province, southwestern China, 23.16 N, 113.36 E, 1345 m, 2022-III-21, leg. Hao Wang, in SCAU
Copyright © Mingyi Tian
Graciliblemus gui Tian, Huang & Jia, 2023 Holotype
Male adult
Gui Dong (Ghost Cave), Pilin, Zhaoxing, Liping, Guizhou, 2020-VI-22, 305 m, Mingyi Tian leg., in SCAU
Copyright © Mingyi Tian
Panotrechus pilosus Tian & Zhou, 2023 Holotype
Male adult
Wojiao Dong Cave, Baotian Zhen, Panzhou Shi, Luipanshui Shi, western Guizhou, 25.40° N / 104.74° E, 1593 m, 7.VII.2021, Jingli Cheng & Mingyi Tian leg. (SCAU).
Copyright © Mingyi Tian
Yunaphaenops graciliphallus Tian & He, 2023 Holotype
Male adult
Fengwei Dong Cave, Wenliu Xiang, Qiubei Xian, Wenshan Zhuang & Miao Autonomous Prefecture, eastern Yunnan, 24.33° N / 104.29° E, 1302 m, 3.IX.2020, leg. Chenghai Fu (SCAU).
Copyright © Mingyi Tian
Leshanius yunzhii Tian & He, 2023 Holotype
Female adult
Xiaoxuankong Tiankeng Cave, Nongke Cun, Fulu Zhen, Shawan Qu, Leshan Shi, Sichuan, 29.29° N / 103.65° E, 530 m, 13.III.2021, leg. Yunzhi Zhang & Li He (SCAU).
Copyright © Mingyi Tian
Aphaenops (Aphaenops) leschenaulti Bonvouloir, 1862: 568
Male adult
Image from Faille, Fresneda & Bourdeau, 2023
Copyright © - - -
Aphaenops (Riberaphaenops) holcartensis Genest, 1977: 281
Male adult
Image from Faille, Fresneda & Bourdeau, 2023
Copyright © - - -
Aphaenops (Pyreneotrechus) aldensis Jeannel, 1955
Male adult
Image from Faille, Fresneda & Bourdeau, 2023
Copyright © - - -
Aphaenops (Nafarroaphaenops) giraudi J. Ochs, 1938
Male adult
Image from Faille, Fresneda & Bourdeau, 2023
Copyright © - - -
Aphaenops (Hydraphaenops) ehlersi (Abeille de Perrin, 1872)
Male adult
Image from Faille, Fresneda & Bourdeau, 2023
Copyright © - - -
Aphaenops (Geotrechus) discontignyi Fairmaire, 1867
Male adult
Image from Faille, Fresneda & Bourdeau, 2023
Copyright © - - -
Aphaenops (Geotrechidius) gallicus (Delarouzee, 1857)
Male adult
Image from Faille, Fresneda & Bourdeau, 2023
Copyright © - - -
Aphaenops (Dupreaphaenops) vasconicus Jeannel, 1913
Male adult
Image from Faille, Fresneda & Bourdeau, 2023
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Aphaenops (Ceretotrechus) ubachi (Español, 1965)
Male adult
Image from Faille, Fresneda & Bourdeau, 2023
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Aphaenops (Aurigerotrechus) vulcanus Abeille de Perrin, 1904
Male adult
Image from Faille, Fresneda & Bourdeau, 2023
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Aphaenops (Argonotrechus) orpheus Dieck, 1869
Male adult
Image from Faille, Fresneda & Bourdeau, 2023
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Nipponaphaenops (Tanakaphaenops) kojii Naitô, 2022 Holotype
Female adult
Kuroyama, 480 m alt., Furuhashi, Ishidatami,Uchiko-chô [typed on white label] / Ehime Pref.northwestern Shikoku, western Japan, 9.V.2009,K. Tanaka leg. [typed on white label] / found in abaited trap set by K. Tanaka on 10.IV.2009 (NSMT). Image from Naitô, 2022
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Trechisibus (Trechisibiodes) valenciai M.Etonti & Mateu, 1992 Paratype
Male adult
Image from: Córdoba, Silvia Patricia, Sánchez, Francisco Rolando & Bezdjian, Laura Patricia, 2023, Types of Carabidae (Insecta: Coleoptera) deposited at the entomological collection of Instituto- Fundación Miguel Lillo, Tucumán, Argentina
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Trechisibus (Trechisibus) bordoni Mateu, 1978 Paratype
Female adult
Image from: Córdoba, Silvia Patricia, Sánchez, Francisco Rolando & Bezdjian, Laura Patricia, 2023, Types of Carabidae (Insecta: Coleoptera) deposited at the entomological collection of Instituto- Fundación Miguel Lillo, Tucumán, Argentina
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Aphaenops (Aphaenops) pallisei Vives & Comas, 2023 Holotype
Female adult
Cueva de los Arnales, 24-VIII-2021, J. Pallisé & A. Meseguer leg. Holotipo temporalmente conservado en la colección J. Vives & E. Vives (Terrassa) antes de pasar definitivamente al MCNB. Image from Vives & Comas, 2023
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Aepopsis robinii (Laboulbene, 1849)
Male adult
Sanxenxo, Pontevedra, Spain, 12.8.1992 M. Toribio leg. (cAA)
Copyright © Alexander Anichtchenko
Omalodera dentimaculata Solier, 1849
Male adult
Chile, Region X, Osorno prov., 3-6km E Maicolpue, 24.10.1993, R.D. Ward leg. (cAA)
Copyright © Alexander Anichtchenko
Ushijimaella uenoi P. Moravac et Wrase, 1998: 213 Paratype
Female adult
China, Shaanxi, Qin Ling Shan, 107.56E, 35.45N, autoroute km 93 S of Zhouzhi, 108km SW Xian Mountain Forest, sifted, 1650m, 1-2.9.1995, leg. A. Putz (cAA)
Copyright © Alexander Anichtchenko
Protrechiama crassipes (Ueno, 1997)
Female adult
China, Sichuan prov., Yizi Pass btw. Meigu County and Ebian County , mixed forest, 2923 m, N 28.67477, E 103.05248 ”; “ 2012.VI.12, in dead log, Shi Hongliang, Liu Ye leg. Image from Belousov, I., Kabak, I. & Liang, H., 2019
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Junnanotrechus baehri Deuve, 2013
Male+Female adult
China (Yunnan), Dali Bai Aut. Pref., Jeu Shan, summit plateau, 37 km NE Dali, 3150 m, 25 ° 58´30 ´´ N / 100 ° 21´36 ´´ E, (mixed forest, sifted from litter, moss, mushrooms), 5.09. 2009
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Anophthalmus (Anophthalmus) driolii Bognolo et M. Etonti, 1996: 179 Paratype
Male adult
Copyright © Vittorino Monzini
Baehria separata Schmidt & Faille, 2023 Paratype
Male+Female adult
Ethiopia, Amhara, Mt. Choke, crater valley, alt. 3780–3900 m, 10°42'12"N, 37°50'58"E, 27.II.2019, leg. D. Hauth, J. Schmidt, Yeshitla M., Yitbarek W.
Copyright © Joachim Schmidt
Trechus (Abunetrechus) bipartitus Raffray, 1880
Male adult
Ethiopia, Amhara, Mt. Abuna Yosef, N-slope, 3800–3950 m, 12°07'52"N, 39°11'39"E, 4.III.2019, leg. D. Hauth, J. Schmidt, Yeshitla M., Yitbarek W
Copyright © Joachim Schmidt
Trechus (Abunetrechus) lalibelae Quéinnec & Ollivier, 2021
Male adult
Ethiopia, Amhara, Mt. Abuna Yosef, N-slope, 3800–3950 m, 12°07'52"N, 39°11'39"E, 4.III.2019, leg. D. Hauth, J. Schmidt, Yeshitla M., Yitbarek W.
Copyright © Joachim Schmidt
Orotrechus ruffoi Tamanini, 1954
Female adult
Lessini M.ts, Verona, Italy
Copyright © Vittorino Monzini
Duvalius (Duvalius) vallombrosus (C. & F. Rasetti, 1920)
Female adult
Buca dell'Orso cave, Toscana, Italy
Copyright © Vittorino Monzini
Aphaenops (Hydraphaenops) carrerei Coiffait, 1953
Male adult
gr. du Trapech, Montvallier FR
Copyright © Vittorino Monzini
Aphaenops (Hydraphaenops) aeacus (Saulcy, 1864)
Male adult
Bagnérres de Bigorre - Gr.tte de Castelmouli - FR
Copyright © Vittorino Monzini
Pheggomisetes globiceps breiti Mandl, 1942
Male adult
Bulgaria, Gintsi, cave Dineva, 24.3.1995 B. Gueorguiev leg. (cAA)
Copyright © Alexander Anichtchenko
Pachydesus bohemani (Jeannel, 1926)
Female adult
RSA Western Cape, Cape Town, foodhills of Table Mt., 13-15.1.2009, leg & det P. Schule
Copyright © Alexander Anichtchenko
Pheggomisetes buresi (Knirsch, 1923)
Male adult
Bulgaria, Vrachanska pl., Mt, cave Ledenika, 17.3.1995 B Gueorguiev leg. (cAA)
Copyright © Alexander Anichtchenko
Ryukyuaphaenops pulcherrimus Sugaya, Kakizoe, Ooka, Tamura & Sone, 2023
Male adult
“JAPAN: Okinawa-ken, Okinawa-jima Is., / Motobu-chō, Furujima, Ufugushikumui- / abu-dai-ichi Cave [上城森アブ第 / 一], 26.6744°N, 127.9056°E, (el. 220m), / 28 II 2022, H. Tamura” (NSMT). Image from Sugaya, Kakizoe, Ooka, Tamura & Sone, 2023
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Anophthalmus (Anophthalmus) hauckei P. Maravec et Lompe, 2003
Female adult
Najdena jama, Laze planina, Slo
Copyright © Vittorino Monzini
Anophthalmus (Arctaphaenops) angulipennis styriacus Winkler, 1933
Male adult
Copyright © Vittorino Monzini
Boldoriella (Insubrites) brembana (Binaghi, 1937)
Male adult
M.te Cavallo, 2000m, Val Seriana Bergamo, Italia
Copyright © Vittorino Monzini
Trechus (Trechus) santaluciaensis Donabauer, 2014 Paratype
Male adult
Monterey County, Big Sur, 28.IV.2013, leg. M. Donabauer; Image from Donabauer, 2014
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Trechus (Trechus) conformis Jeannel, 1927
Male adult
Gualale, near campground at Gualale river, 2.V.2013, leg. M. Donabauer. Image from Donabauer, 2014
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Trechus (Trechus) ovipennis Motschulsky, 1845
Male adult
E of Eureka, Elk River, 4.V.2013, leg. M. Donabauer; Image from Donabauer, 2014
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Duvalius (Duvalius) lepinensis Cenuti, 1950
Male adult
Ouso di pozzo comune cave, Roma, Bastianini leg
Copyright © Vittorino Monzini
Orotrechus robustus Jeannel, 1928a: 342
Female adult
Altipiano Cansiglio, BL - Italy
Copyright © Vittorino Monzini
Boldoriella (Insubrites) serianensis serianensis (Breit, 1913)
Male adult
Val di Scalve, Bg. - Italia
Copyright © Vittorino Monzini
Troglocimmerites zapparolii Casale & Magrini, 2023 Holotype
Female adult
Turchia, Anatolia nord-orientale, vilayet Giresun, strada per Çanakci, 70-80 m s.l.m., 5,8 km dal bivio per Çanakci, 6.V.1987, leg. M. Zapparoli (AVT). Image from Casale & Magrini, 2023
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Taniatrechus xeniae Belousov & Koval, 2023 Holotype
Female adult
“W Caucasus, Abkhazia, Bzyb Mt. Range, 3.4 km NEE of Mt. Tshiptshira, 2,220 m, Prostornaya Cave (= L-07 Cave), at a depth of 50 m, traps, 31.VIII.2010 – 11.VIII.2011. X.N. Gorbunova, T.V. Petrova leg.” (ZIN).
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Chiapadytes pitteti Casale & Magrini, 2021 Holotype
Male adult
“Mexico, Chiapas, San Cristobal de Las Casas, m 2250 s.l.m., Cueva N° 7 sul Rio Quinta, 24.I.1982, leg. A. Gobetti e J.F. Pittet” (MSNG). Image from Casale & Magrini, 2021
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Chiapadytes bolivari Vigna Taglianti, 1977
Female adult
Mexico, Chiapas, San Cristobal de Las Casas, Las Piedrecitas, Cueva de la Planta I, 25.III.1982, leg. A. Gobetti e J.F. Pittet (TT ♀, MSNG). Image from Casale & Magrini, 2021
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Epaphiopsis (Epaphiopsis) narukawai Morita, 2021 Paratype
Female adult
Mt. Gozaisho-dake, alt. 1,000 m, Komonochô, Mie-gun, Mie Prefecture, Honshu, Japan. Image from Morita, 2021
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Anophthalmus (Anophthalmus) spectabilis istrianus (Ganglbauer, 1913)
Male adult
Grotta Roditti
Copyright © Vittorino Monzini
Thalassophilus longicornis Sturm, 1825: 83
Male adult
CZECH REPUBLIC - Olomouc reg., Přerov dist., Skalička, 11.VI.2020, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar
Yinaphaenops guilinensis Li, Liu & Huang, 2024 Holotype
Male adult
cave Guizai Dong, Lingchuan, Guilin, Guangxi, China, 26-IV-2024, leg. Sun-Bin Huang, Ming-Zhi Zhao, Ran Li & Rong Chen (SCAU). Image from Li, Liu & Huang, 2024
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Ushijimaella ollivieri Deuve & Queinnec, 2014 Holotype
Male adult
Chine, Sichuan, environs de Maoxian, 3000-3200 m, in coll. É. Ollivier, à Bolbec (France).
Copyright © Thierry Deuve
Guizhaphaenops pouillyi Deuve & Queinnec, 2014 Holotype
Male adult
Chine, Guizhou, canton de Panxian, rivière Gesuo He, Yin Dong / Luoshui Dong II, in coll. Muséum national d’Histoire naturelle, à Paris.
Copyright © Thierry Deuve
Sinaphaenops (Dongaphaenops) xuxiakei Deuve & Tian, 2014 Holotype
Male adult
Chine, Guizhou, Panxian, au sud-ouest de Gucheng, Biyun Dong (= Shui Dong) (Tian Mingyi & Huang Sunbin leg., 9.VII.2013), in coll. South China Agricultural University, à Canton (Chine).
Copyright © Thierry Deuve
Trechus (Trechus) bourdeaui Fresneda, Valenzuela & Faille, 2015 Holotype
Male adult
España,Asturias,Quirós-Salcedo,CuevaVallemuñón,21.III.2014,E.Valenzuela leg. (CFL). Image from Fresneda, Valenzuela & Faille, 2015
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Trechus (Trechus) saxicola Putzeys, 1870
Male adult
Imago: Puerto de Ventana 1586 m; Aedeagus: Puerto de Pajares 1378 m. Image from Fresneda, Valenzuela & Faille, 2015
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Aspidaphaenops qujingensis Deuve & Tian, 2020 Holotype
Male adult
Chine, Yunnan, Préfecture de Qujing, Luoping Xian, Machang, grotte Shuiyuan Dong, 1530 mètres, 24°49’33’’N 104°21’48’’E (Tian M. Y., Lui W. X., Wang X. H., Tang M. R. leg., 18.VI.2015), in coll. South China Agricultural University, Canton.
Copyright © Thierry Deuve
Aspidaphaenops qujingensis achaetus Deuve & Tian, 2020 Holotype
Male adult
Chine, Yunnan, Préfecture de Qujing, Shizong, grotte Xinzhuangke Dong, 1110 mètres, 24°39’4’’N 104°15’57’’E (Tian M. Y., Lui W. X., Wang X. H., Tang M. R. leg., 17.VI.2015), in coll. South China Agricultural University, Canton.
Copyright © Thierry Deuve
Cimmeritodes (Shimenrites) shimenensis Deuve & Tian, 2017 Holotype
Male adult
Chine, Hunan, Shimen, Xiannu Dong, 280 mètres, 29°38’55’’N -111°04’01’’E (Tian M. & Chen J., 6.VIII.2016), in coll. South China Agricultural University, Canton, Chine.
Copyright © Thierry Deuve
Sinotroglodytes ariagnoi Deuve, 2016 Holotype
Male adult
Chine, Hubei sud-occidental, préfecture d’Enshi, Enshi Xian, Hongtu, grotte Chenjiayan Dong (J. Lips, 18.viii.2014), in coll. Muséum national d’Histoire naturelle, Paris.
Copyright © Thierry Deuve
Cathaiaphaenops (Amygdalotrechus) lagredeae Deuve, 2016 Holotype
Female adult
Chine, Hubei sud-occidental, préfecture d’Enshi, Enshi Xian, Shadi, grotte Shadi Xiao Dong (J. Lips, 26.VIII.2014), in coll. Muséum national d’Histoire naturelle, Paris.
Copyright © Thierry Deuve
Minimaphaenops (Enshiaphaenops) senecali Deuve, 2016 Holotype
Female adult
Chine, Hubei sud-occidental, préfecture d’Enshi, Enshi Xian, Shadi, grotte Shadi Xiao Dong (J. Lips, 26.VIII.2014), in coll. Muséum national d’Histoire naturelle, Paris.
Copyright © Thierry Deuve
Gotoblemus (Kamigotoblemus) exilis Naito, 2018 Holotype
Male adult
Nanamé-gô, 130 m in altitude, northern foot of Yagura-daké Mountain on Na kadôri-jima Island of the Gotô Islands, Shin-kamigotô-chô, Nagasaki Prefecture, western Japan. Image from Naito, 2018
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Paratrechus obrieni Mateu, 1981
Male adult
Costa Rica, Cerro Buena Vista, 3400 m a.s.l., N 9.5548°, W 83.7552°, 15.IX.2023. Image from Donabauer, 2024
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Paratrechus alexandri Trezzi, 1995
Male adult
Costa Rica, Volcan Barva, ca. 2700 m a.s.l., 9.IX.2023. Image from Donabauer, 2024
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Duvalius (Duvalius) bonzanoi Casale et Vigna Taglianti, 1990
Male adult
GR. Joannina, Anemotrypa-Pramanta
Copyright © Vittorino Monzini
Trechus (Epaphius) dorsistriatus laferi Sundukov & Makarov, 2021 Holotype
Male adult
Ю Kурилы, СВ Kунашир, р. Саратовская 2 км выше устья, 26.07.2014, Ю. Сундуков, Л. Сундукова (FEB).
Copyright © Kirill Makarov
Aphaenops (Ceretotrechus) delioti Faille, Bourdeau, Bellés & Fresneda, 2015
Male adult
Spain
Copyright © Vittorino Monzini
Inotrechus usholtaensis Maghradze, Lohaj & Faille, 2025 Holotype
Male adult
GEORGIA, Racha-Lechkhumi and Kvemo Svaneti region, Racha karst massif, Usholta Cave, dark zone, 21.viii.2022, E. Maghradze leg. (IZISU).
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Inotrechus injaevae Daizhanski et Ljovuschkin, 1989: 147
Male adult
Image from Maghradze et al, 2025
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Inotrechus injaevae Daizhanski et Ljovuschkin, 1989: 147
Male adult
Image from Maghradze et al, 2025
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Inotrechus kurnakovi Dolzhanski et Ljovuschkin, 1989
Male adult
Image from Maghradze et al, 2025
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Duvalius (Duvalius) gregoi Maghradze, Lohaj & Faille, 2025 Holotype
Male adult
GEORGIA, Racha-Lechkhumi and Kvemo Svaneti region, Racha karst massif, Kvedi II Cave, dark zone, 24.v.2023, E. Maghradze leg. (IZISU). Image from Maghradze et al, 2025
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Darlingtonea kentuckensis Valentine, 1952
Male adult
Kentucky, Pulaski Co., 10mi NE Somerset, stab, Taylor Cave, west bank Buck Creek 10.11.1949, Valentine, Jones & Diamond (cAA).
Copyright © Alexander Anichtchenko
Neoblemus championi Jeannel, 1923
Male adult
India, Uttarakhand, 18-20.4.2012, 15km SW New Tehri, 869m, Anichtchenko A. leg. & det. (cAA).
Copyright © Alexander Anichtchenko
Perileptus (Perileptus) melas Jeannel, 1926
Male adult
Philippines, Mindanao, Ugwad Falls (Kapuka), Caraga, Davao Or., 260m, 7.440359, 126.434027, 29–30.IV.2023, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko
Perileptus (Parablemus) cf cameroni Jeannel, 1923
Male adult
India, Uttarakhand, 8km NW Jaspur, lake shore, 22-23.4.2012, Anichtchenko A. leg. & det. (cAA). 2.3 mm
Copyright © Alexander Anichtchenko
Nanotrechiama loebli Belousov & Kabak, 2018 Holotype
Male adult
CH, Yunnan Province, Mt. E Mengsa Town, 23º42´26´´ N / 99º47´26´´ E, H=3045 m, 30.05.2010, Belousov I. & Kabak I. leg. (ZIN). 4.5 mm; Image from Belousov & Kabak, 2018
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Petraphaenops unguiculatus Delić, Kapla & Colla, 2020 Holotype
Male adult
SLOVENIA, Luče, Raduha, Devettisoča jama (Cave Registry N o. 9000), 46.41231°N, 14.75095°E, found at –450 m (pitfalltrap), collected by T. Delić, 14 June 2017. Image from Delić et al, 2019 (2020)
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Kusumia kukai Ashida, 2023 Holotype
Male adult
(OMNH TI 545), 19.XI.2022 (baited traps set on 23.X.2022), H. Ashida leg. ( Nishigô (about 800 m in altitude), west of Kôyasan, Kôya-chô, Wakayama Prefec ture, Central Japan.). Image from Ashida, 2023
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Anophthalmus (Anophthalmus) hitleri Scheibel, 1937
Female adult
(SLO) Velenje, Steska jama
Copyright © Vittorino Monzini
Trechus (Trechus) pochoni Jeannel, 1939
Male adult
Efra lake, Switzerland
Copyright © Vittorino Monzini
Kusumia hybrida Ashida, 2025 Holotype
Male adult
holotype ♂, from Tonono in Ôtô-chô
Copyright © Qin Yuyao
Kusumia hayashii Ashida, 2025 Holotype
Male adult
holotype ♂, from Sakamoto in Ôtô-chô
Copyright © Qin Yuyao
Kusumia militis Ueno & Naito, 2005
Male adult
Shio-dani (610 m in altitude), Tenkawa-mura, Nara Prefecture, 8.XII.2024 (baited traps set on 17.XI.2024), H. ASHIDA leg.
Copyright © Qin Yuyao
Duvalius (Duvalius) krueperi (Schaum, 1862)
Male adult
GR - Oros Parnassos - M. Egger leg.
Copyright © Vittorino Monzini
Aphaenops (Nafarroaphaenops) cuberesi Comas & Vives, 2025 Holotype
Male adult
cova Cuberes / Serradell / Conca de Dalt /(Pallars Jussà) / 22-VIII-2024 / Filbà & Pallisè leg.; Aphaenops (Nafarroaphaenops) cuberesi n. sp. J.Comas & E. Vives det. Museu de Ciències Naturals de Barcelona. Image from Comas & Vives, 2025
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Zolini taxon page (32 photos)
Sloaneana tasmaniae (Sloane, 1915)
Male adult
Grove, S.J. (2010). Tasmanian Forest Insect Collection web-site
Copyright © Simon Grove
Sloaneana cf. tasmaniae (Sloane, 1915)
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison
Oopterus punctatus Larochelle & Larivière, 2017 Holotype
Male adult
Lincoln University Entomology Research Collection
Copyright © - - -
Zolus helmsi (Sharp, 1886)
Female adult
Copyright © - - -
Zolus carinatus (Broun, 1882)
Male adult
Copyright © - - -
Zolus kauriensis Larochelle & Larivière, 2017
Male adult
Copyright © - - -
Zolus subopacus Broun, 1915
Male adult
Copyright © - - -
Zolus wongi Larochelle & Larivière, 2017 Paratype
Female adult
[NEW ZEALAND BR] Mt Dewar 1000m J.C. Watt (typed) / under bark [of] logs (typed) / Paparoa Range Dec. 1969 (NZAC). Scale bar 1mm
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Oopterus patulus (Broun, 1881)
Male adult
scale bar 1mm
Copyright © - - -
Oopterus discoideus Larochelle & Larivière, 2017 Paratype
Female adult
“[NEW ZEALAND FD] Wolfe Flat 600m-680m (typed) / Turret Ra [= Range] (typed) / Manapouri Exp. Jan. 70 J. McBurney (typed) / moss (typed) 70/69. Scale bar 1mm
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Oopterus marrisi Larochelle & Larivière, 2017 Paratype
Male adult
Murchison Mountains (FD)
Copyright © - - -
Oopterus lewisi (Broun, 1912)
Male adult
Copyright © - - -
Pseudoopterus otiraensis Larochelle & Larivière, 2017 Paratype
Male adult
NEW ZEALAND WD Arthur’s Pass NP, Kellys Creek x Otira River [= Kelly Creek near Otira River Junction] 4248S 17134E 350m 5.III. 2009 Larochelle and Larivière (typed) / Floodplain Podoc.-brleaf for. [= Podocarp-broadleaf forest] (Fuchsia-Horopito dom. [= dominant]): wet clayish soil. Under large, deeply embedded stones. Scale bar 1mm
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Maungazolus parvulus (Broun, 1903)
Male adult
Copyright © - - -
Maungazolus acutus Larochelle & Larivière, 2017 Paratype
Male adult
NEW ZEALAND FD Takahe Valley Murchison Mts [= Mountains] 11 Dec 1972 A.C. Eyles (typed) / Litter (typed) 72/271
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Maungazolus septempunctatus Larochelle & Larivière, 2017 Paratype
Male adult
“[NEW ZEALAND SI] Table Hill Stewart I. (typed) / 2000’ 16.II.68 J. McBurney (typed) / Mat plants (typed) 68/55
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Pseudoopterus latipennis (Broun, 1903)
Male adult
Scale bar 1mm
Copyright © - - -
Pseudoopterus plicaticollis (Blanchard, 1843)
Male adult
Scale bar 1mm
Copyright © - - -
Pseudoopterus fiordlandensis Larochelle & Larivière, 2017 Paratype
Female adult
NEW ZEALAND FD Takahe Valley Murchison Mts [= Mountains]11 Dec 1972 A.C. Eyles (typed) / Litter (typed) 72/271 (hand-written) / JOHN NUNN (typed)
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Maungazolus pygmaetus Broun, 1907
Female adult
Copyright © - - -
Oopterus nunni Larochelle & Larivière, 2017 Holotype
Male adult
NEW ZEALAND SL Catlins SFP [= Catlins State Forest Park], Tawanui Campground 100m 12.I.1999 Larochelle and Larivière (typed) / Wet beech forest – along stream banks – in leaf litter
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Oopterus garnerae Larochelle & Larivière, 2017
Female adult
Copyright © - - -
Merizodus soledadinus (Guerin-Meneville, 1830)
Male adult
Copyright © - - -
