Carabus (Carabus) donjuan Imura, 2011 Holotype
Endophallus
Image from Imura, 2011
Copyright © - - -

Carabus (Carabus) koidei yanyuan Cavazzuti & Rapuzzi, 2010
Endophallus
Image from Imura, 2011
Copyright © - - -

Carabus (Acoptopterus, ?Carabus) yunnanus yanyuanicus Cavazzuti, 1996
Endophallus
China, S-Sichuan, S Yanyuan, Chuandongzi Mt h=3300-3600 m
Copyright © Ruslan Panin

Carabus (Apotomopterus) protenes baoxingensis Deuve & Mourzine, 2003
Endophallus
China: С-Sichuan:Qionglai Shan Mts., Baoxing env. (TOPOTYPUS)
Copyright © Igor Solodovnikov

Carabus (Apotomopterus) delavayi parvipes Cavazzuti, 1996
Endophallus
S-Sichuan: S of Yanyuan, Chuandongzi Massif h=3200 m, 27.351206, 101.507036
Copyright © Ruslan Panin

Carabus (Apotomopterus) ludivinae wuliangensis Cavazzuti, 1997
Endophallus

Copyright © Ruslan Panin

Carabus (Apotomopterus) protenes ascendens Semenov, 1898
Endophallus
China, Chongqing, N Jinfo Shan, Shanwangping h=1400m, 29.101944, 107.166389
Copyright © Ruslan Panin

Carabus (Apotomopterus) korellianus Kleinfeld, 2002
Endophallus
China, SW-Yunnan, Dehong Distr., Ruili, Pijiazhai vill., h=1800 m, 24.048111, 97.738222
Copyright © Ruslan Panin

Carabus (Apotomopterus) madefactus Cavazzuti, 1997
Endophallus
China, CW-Yunnan, Nánjiàn Yízú Zìzhìxiàn, Mt Lingbao h=2400 m, 24.768611, 100.500000
Copyright © Ruslan Panin

Carabus (Apotomopterus) mecynodes pinchuanensis Imura, 1999
Endophallus
China, S-Sichuan, 40-50 km E Xichang (to Zhaojue) 27°51'N 102°30'E, h=2700-3000m, Pine forest
Copyright © Ruslan Panin

Carabus (Apotomopterus) patroclus apricalis Cavazzuti, 1997
Endophallus
China, S Sichuan, W of Pingchuanzhen, near Galuo vill., h=2400 m
Copyright © Ruslan Panin

Carabus (Apotomopterus) microtatos lingbaonis Imura, 2019
Endophallus
China, CW-Yunnan, Nánjiàn Yízú Zìzhìxiàn, Mt Lingbao h=2400 m, 24.768611, 100.500000
Copyright © Ruslan Panin

Carabus (Apotomopterus) bousquetellus bousquetellus Deuve, 1998
Endophallus
S-China, CW-Yunnan, W of Lincang, Daxue Shan h=2600m, 23.935556, 100.257778
Copyright © Ruslan Panin

Carabus (Apotomopterus) patroclus patroclus Breuning, 1933
Endophallus
China, S Sichuan, Pass 56 km SW Miannin, near Wawo h=2440 m
Copyright © Ruslan Panin

Carabus (Archicarabus) monticola fontanae Born, 1906
Endophallus

Copyright © Ruslan Panin

Carabus (Archicarabus) montivagus montivagus Palliardi, 1825
Endophallus

Copyright © Ruslan Panin

Carabus (Archicarabus) nemoralis nemoralis Muller, 1764
Endophallus

Copyright © Ruslan Panin

Carabus (Archicarabus) phoenix Lapouge, 1924
Endophallus
C-Lebanon, Byblos /Jbeil Distr., Kfardebian env., h=1200m,
Copyright © Ruslan Panin

Carabus (Aulonocarabus) kurilensis ? daisetsuzanus Kono, 1936
Endophallus
N-Japan, Hokkaido, Katō District, Kamishihoro, Horoka near Maruyama Bridge,
Copyright © Ruslan Panin

Carabus (Aulonocarabus) truncaticollis fleischeri Reitter, 1898
Endophallus
Russia, N Burajatia, Barguzinskiy Mts., the upper reaches of M.Chivyrkuy riv., h=2000 m
Copyright © Ruslan Panin

Carabus (Carabus) granulatus granulatus Linne, 1758
Endophallus

Copyright © Ruslan Panin

Carabus (Carabus) arvensis arvensis Herbst, 1784
Endophallus
W Ukraine, NW Podolian, Lviv reg., Javoriv distr.,near Stradch vill.,
Copyright © Ruslan Panin

Carabus (Carabus) arvensis hokkaidoensis Lapouge, 1924
Endophallus
N Japan:Hokkaido, Hamamasu env.,
Copyright © Ruslan Panin

Carabus (Carabus) arvensis deyrollei Gory, 1839
Endophallus
NW-Spain, Cantabria, Udías env., Monte Corona h=100m
Copyright © Ruslan Panin

Carabus (Carabus) stscheglowi Mannerheim, 1827
Endophallus
Russian, Chuvashia, Cheboksary, Zavolzhye distr., , near lake Astrakhanka,
Copyright © Ruslan Panin

Carabus (Carabus) vinctus Weber, 1801
Endophallus
LOUISIANA, East Baton Rouge Parish, 1.5mi W Denham Springs, 30.482 ̊ N 90.981 ̊ W, bottomland hardwood forest, 4 Sep 2017
Copyright © Igor Sokolov

Carabus (Cechenochilus) heydenianus Starck, 1889
Endophallus
Carabus (Procechenochilus) heydenianus euxinus Semenov, 1896 Abkhasia: Arabika plato (TOPOTYPUS)
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) boeberi buschi Semenov & Znojko, 1931
Endophallus
Russia: Kabardino-Balkaria: u.r. Adylsu Riv.
Copyright © Igor Solodovnikov

Carabus (Chaetocarabus) intricatus bayardi (as silaensis Haury, 1881)
Endophallus
S-Italy, Calabria, Sila Grande Massif, Camigliatello h=1040 m
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens escheri Palliardi, 1825
Endophallus

Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens festivus Dejean, 1826
Endophallus

Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens farinesi Dejean, 1826
Endophallus

Copyright © Ruslan Panin

Carabus (Chrysocarabus) hispanus Fabricius, 1787
Endophallus

Copyright © Ruslan Panin

Carabus (Coptolabrus) elysii xiaoganicus Deuve et Remond, 2016
Endophallus
China, Hubei, Xiaogan, Mt Shuangfeng Shan,
Copyright © Ruslan Panin

Carabus (Coptolabrus) lafossei tungchengensis Li, 1993
Endophallus
China, Hubei, Shangzhi co., Shimenyan 8-11.6.2009 (cAA)
Copyright © Alexander Anichtchenko

Carabus (Coptolabrus) augustus ignigena Hauser, 1914
Endophallus

Copyright © Alexander Anichtchenko

Carabus (Cryptocarabus) tsharynensis Kabak 1994
Endophallus
SE-Kazakhstan, Almaty Region, W-Zailiyskiy Alatau rng, Belshabdar valley h~1300 m
Copyright © Ruslan Panin

Carabus (Cryptocarabus) sacarum Kabak, 1998
Endophallus
SE-Kazakhstan, Almaty reg., C-Zailiyskiy Alatau N-slopes, Kaskalen river Valley, Izvestkovy vill., h=1700 m,
Copyright © Ruslan Panin

Carabus (Ctenocarabus) melancholicus costatus Germar, 1824
Endophallus
NW Portugal, Viana do Castelo distr., Sierra de Agra
Copyright © Ruslan Panin

Carabus (Cupreocarabus) huangi huangi Deuve, 1992
Endophallus

Copyright © Ruslan Panin

Carabus (Cupreocarabus) balangicus balangicus Cavazzuti, 2002
Endophallus
Chine, C-Sichuan, Xiaojin, Wolong valley: Balang Shan Pass h=3448 m, 30.884617, 102.973117
Copyright © Ruslan Panin

Carabus (Deroplectes) dokhtouroffi dokhtouroffi Ganglbauer, 1886
Endophallus
NC Tadzhikistan, Pamir, N-slope Peter I.Mt Rng, h=2100m, Ganishou vill.
Copyright © Ruslan Panin

Carabus (Deroplectes) klapperichianus coiffaitianus Deuve, 1990
Endophallus
Tadzhikistan: Badakhshan: Ishkashim Mt.R.: Kuh-i-Lal, VI.1979 leg. I.K. Lopatin
Copyright © Igor Solodovnikov

Carabus (Deroplectes) arcanus arcanus Semenov, 1898
Endophallus

Copyright © Igor Solodovnikov

Carabus (Deroplectes) arcanus hissarianus Semenov, 1900
Endophallus
Tajikistan, Gissar ridge, Kondara, Kwak, h = 1900 m, 7,VI.1970, I.K. Lopatin leg.
Copyright © Igor Solodovnikov

Carabus (Deroplectes) sphinx dardazicus Deuve, 1990
Endophallus

Copyright © Igor Solodovnikov

Carabus (Diocarabus) fraterculus fraterculus Reitter, 1895
Endophallus
N-Korea, N Pyongan Prov.,Sakju, Mt Ponsan,
Copyright © Ruslan Panin

Carabus (Eucarabus) ulrichi ulrichi Germar, 1824
Endophallus
endophallus ulrichi (as podolicus Semenov, 1898)
Copyright © Ruslan Panin

Carabus (Eucarabus) manifestus pyonganicola Deuve & Li, 1998
Endophallus

Copyright © Ruslan Panin

Carabus (Eucarabus) obsoletus obsoletus Sturm, 1815
Endophallus

Copyright © Ruslan Panin

Carabus (Eucarabus) manfredeggeri Müller 2012 Paratype
Endophallus

Copyright © Ruslan Panin

Carabus (Eucarabus) angustus dopyeong Rapuzzi, 2015
Endophallus
scale bar = 5mm
Copyright © Jong-Bong Choi

Carabus (Eucarabus) sternbergi sternbergi Roeschke, 1898
Endophallus
Scale bar = 5mm
Copyright © Jong-Bong Choi

Carabus (Eucarabus) sternbergi gureopensis J.K. Park & J. Park, 2013 Paratype
Endophallus
Scale bar = 5mm
Copyright © Jong-Bong Choi

Carabus (Eucarabus) cartereti cartereti Deuve, 1982
Endophallus
Scale bar = 5mm
Copyright © Jong-Bong Choi

Carabus (Eucarabus) nitididorsus nitididorsus Ishikawa & Kim, 1983
Endophallus
Scale bar = 5mm
Copyright © Jong-Bong Choi

Carabus (Eucarabus) nitididorsus paegunsanus Imura, 1997
Endophallus
Scale bar = 5mm
Copyright © Jong-Bong Choi

Carabus (Eucarabus) nitididorsus baegyangsanus (Choi & Park, 2019) Holotype
Endophallus
Scale bar = 5mm
Copyright © Jong-Bong Choi

Carabus (Eucarabus) namhaedoensis Kwon & Lee, 1984
Endophallus
Scale bar = 5mm
Copyright © Jong-Bong Choi

Carabus (Eucarabus) xiuyanensis Deuve & Li, 1998
Endophallus
NE-China: SE-Liaoning, Xiuyan County, Dayingzi, Mt. Laopingding,
Copyright © Ruslan Panin

Carabus (Eucarabus) billbergi kuandianicus Rapuzzi, 2007
Endophallus
NE China, Liaoning, Kuandian County, Dachuantou env.,
Copyright © Ruslan Panin

Carabus (Eucarabus) parreyssi parreyssi Palliardi, 1825
Endophallus
SW-Croatia, Mts Velebit, Mali Halan, h=1390 m,
Copyright © Ruslan Panin

Carabus (Euleptocarabus) porrecticollis porrecticollis Bates, 1883
Endophallus
Japan, C Honshu, Nagano Pre., Toyooka town,
Copyright © Ruslan Panin

Carabus (Eurycarabus) famini fezzanus Lapouge, 1915
Endophallus
Morocco, Middle Atlas Mt Rng, Azrou city h=1500m,
Copyright © Ruslan Panin

Carabus (Hemicarabus) nitens Linne, 1758
Endophallus
Belarus, Vitebsk reg., Miory distr. Swamp «Mokh», 09.05.2011, leg. G.G. Sushko
Copyright © Igor Solodovnikov

Carabus (Hemicarabus) tuberculosus Dejean, 1829
Endophallus
Carabus (Hemicarabus) tuberculosus n. strophium F.-W., 1842 SE Kazakhstan: NW Tarbogatai Mt.R.
Copyright © Igor Solodovnikov

Carabus (Hemicarabus) serratus Say, 1825
Endophallus
E-USA,Virginia, Leesburg env., Ball's Bluff Battlefield Regional Park
Copyright © Igor Solodovnikov

Carabus (Hygrocarabus) nodulosus Creutzer, 1799
Endophallus
Endophallus
Copyright © Ruslan Panin

Carabus (Hygrocarabus) variolosus variolosus Fabricius, 1787
Endophallus
Endophallus
Copyright © Ruslan Panin

Carabus (Imaibius) nouristani nouristani (Ledoux, 1977)
Endophallus

Copyright © Frank Kleinfeld

Carabus (Imaibius) dardiellus perobscurus Deuve, 1980
Endophallus

Copyright © Ruslan Panin

Carabus (Imaibius) boysi boysi Tatum, 1851
Endophallus
NW-India, Himachal Pradesh, Manikaran env., Pulga vill.
Copyright © Ruslan Panin

Carabus (Isiocarabus) miaorum Lassalle & Prunier, 1993
Endophallus
China, NW-Guangxi, Mao'er Shan, h~1600 m 25°52'N 110°29'E
Copyright © Ruslan Panin

Carabus (Lamprostus) robustus ducalis Schaufuss, 1882
Endophallus
NE-Turkey:Trabzon, Zigana Pass h=2000m
Copyright © Ruslan Panin

Carabus (Lamprostus) torosus torosus Frivaldsky, 1835
Endophallus
NE-Bulgaria, Dobrich Region, Kavarna env.,
Copyright © Ruslan Panin

Carabus (Lamprostus) saulcyi saulcyi Piochard, 1875
Endophallus
C Lebanon, Jbeil Distr., Ehmej vill., h=1100 m
Copyright © Ruslan Panin

Carabus (Lamprostus) torosus sinopensis Breuning & Ruspoli, 1970
Endophallus
NC-Turkey, Sinop province, Erfelek env., h=200m
Copyright © Ruslan Panin

Carabus (Lamprostus) hemprichi elonensis Schweiger, 1970
Endophallus
N-Israel, Midlle Galilea, Nazareth env., Hoshaya viil.
Copyright © Ruslan Panin

Carabus (Lamprostus) syrus cheikensis Deuve, 1992
Endophallus
N-Israel, Midlle Galilea, Nazareth env., Hoshaya viil.,
Copyright © Ruslan Panin

Carabus (Lamprostus) erenleriensis erenleriensis Schweiger, 1966
Endophallus
NW-Turkey:Bolu Prov., Abant Lake
Copyright © Ruslan Panin

Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931
Endophallus
NE-China:Liaoning prov.,Fengcheng City, Miaoyang,
Copyright © Ruslan Panin

Carabus (Leptocarabus) procerulus Chaudoir, 1862
Endophallus
Japan:S Honshu , Matsubara-ko,
Copyright © Ruslan Panin

Carabus (Leptocarabus) procerulus Chaudoir, 1862
Endophallus
Japan, S Honshu, Mie Pref., Hakusan,
Copyright © Ruslan Panin

Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
Endophallus
W Ukraine, Transcarpathian Reg., 4 km NE Chop, forest Peresh.
Copyright © Ruslan Panin

Carabus (Macrothorax) rugosus celtibericus Germar, 1824
Endophallus
SW-Portugal, Serra do Cercal h=217 m, 37°47'15.6"N 8°42'02.0"W, road N390,
Copyright © Ruslan Panin

Carabus (Meganebrius) quinlani smetanai Häckel, 2011 Paratype
Endophallus
C-Nepal:Dhaulagiri Massif:Baglung Distr.,(1.5km W of Okhale vill., 28.32’152’’N/ 83.48’673’E, h=2600m)
Copyright © Ruslan Panin

Carabus (Meganebrius) santostamangi santostamangi
Endophallus

Copyright © Joachim Schmidt

Carabus (Meganebrius) thudamensis Deuve & Schmidt, 2017 Holotype
Endophallus

Copyright © Joachim Schmidt

Carabus (Meganebrius) everesti everesti Andrewes, 1929
Endophallus
Kemba Kharka
Copyright © Joachim Schmidt

Carabus (Meganebrius) salpansis Deuve, 1984
Endophallus
Kemba Kharka
Copyright © Joachim Schmidt

Carabus (Meganebrius) indicus indicus Fairmaire, 1889
Endophallus
Tonglu
Copyright © Joachim Schmidt

Carabus (Megodontus) imperialis Fischer, 1823
Endophallus
endophallus
Copyright © Safronov

Carabus (Megodontus) schoenherri sajanus Breuning, 1927
Endophallus
endophallus
Copyright © Safronov

Carabus (Megodontus) violaceus fiorii Born, 1901
Endophallus
endophallus
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus baeterrensis Lapouge, 1901
Endophallus
ehdophallus
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823
Endophallus
endophallus
Lviv reg., Nikolaev distr., near Krasiv vill.
Copyright © Ruslan Panin

Carabus (Megodontus) exaratus Quensel, 1806
Endophallus

Copyright © Ruslan Panin

Carabus (Megodontus) violaceus aurolimbatus Dejean, 1829
Endophallus

Copyright © Ruslan Panin

Carabus (Megodontus) bonvouloiri bonvouloiri (as kaschkarensis Blumenthal & Breuning, 1967)
Endophallus
NE-Turke,Rize vill., S Ayder h=1740 m,
Copyright © Ruslan Panin

Carabus (Megodontus) stroganowi transfuga Semenov, 1898
Endophallus
SE-Azerbaijan: Talysh, Kalvaz,
Copyright © Ruslan Panin

Carabus (Megodontus) kolbei kolbei Roeschke, 1887
Endophallus
N-Japan, C-Hokkaido, Hidaka distr., Niikappu env.,
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus dryas Gistl, 1857
Endophallus
NE-Greece, Mt. Olympos, Litochoro env.
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus azurescens Dejean, 1826
Endophallus
W-Croatia, Velebit Mts., betw.Karlobag & Brusane vill.,
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus germarii Sturm, 1815
Endophallus
N Italy, Trent, Ampola, Mt. Tremalzo h=1700m
Copyright © Ruslan Panin

Carabus (Megodontus) obovatus Fischer, 1827
Endophallus
C.obovatus taizhensis Obydov, 2006 Khakassia: Kuznetskyi Alatau, env. vill. Priiskovyi, Bobrovaya Mt., h = 1200-1300 m
Copyright © Igor Solodovnikov

Carabus (Megodontus) kolbei hidakamontanus Ishikawa, 1966
Endophallus
E-Hokkaido: Samani distr., Horomankyo env.
Copyright © Igor Solodovnikov

Carabus (Megodontus) croaticus croaticus Dejean, 1826
Endophallus
W-Croatia, Velebit Mts., h=950 m, Oltari - Zavizan,
Copyright © Ruslan Panin

Carabus (Megodontus) tuvensis Shilenkov, 1996
Endophallus

Copyright © Igor Solodovnikov

Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
Endophallus
endophalus
Copyright © Ruslan Panin

Carabus (Mesocarabus) dufouri dufouri Dejean, 1829
Endophallus
Jaen reg., Monte Javalcuz h=1436 m
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus brevis (as sorianus Jeanne, 1972)
Endophallus
NC-Spain, Soria, Puerto de Oncala
Copyright © Ruslan Panin

Carabus (Mesocarabus) (? lusitanicus) gougeleti Reiche, 186
Endophallus
CW Spain, Province Huelva, Arroyomolinos de Leon.
Copyright © Ruslan Panin

Carabus (Mesocarabus) cantabricus cantabricus Chevrolat, 1840
Endophallus
NW-Spain, Pontevedra prov., A Cañiza viil., Alto de Fontefría h=800m
det.Panin R.
Copyright © Ruslan Panin

Carabus (Mesocarabus) dufouri baguenai Breuning, 1926
Endophallus
SE Spain, Almería Province, Carboneras env.
det. R.Panin
Copyright © Ruslan Panin

Carabus (Mesocarabus) macrocephalus Dejean, 1826
Endophallus
NW-Spain,Cantabrian Mts, Picos de Europa Mt Rng, Puerto de Panderruedas h=1460 m,
Copyright © Ruslan Panin

Carabus (Mesocarabus) latus latus Dejean, 1826
Endophallus
C-Spain, Toledo, Mts de Toledo, Sierra del Castañar Rng, Puerto del Milagro env, h=920m
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus helluo (as albarracinus Ganglbauer, 1886)
Endophallus
CE-Spain, Provincia Teruel, Sierra de Javalón Rng, El Cañigral env., h=1460m,
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826
Endophallus
CW-Spain:Salamanca, Brincones env., h=750m
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus harcyniae (as belgicus Lapouge, 1913)
Endophallus
Luxembourg, Bollendorf-Pont env., 3.III.1999
Copyright © Ruslan Panin

Carabus (Mesocarabus) (? problematicus trans lusitanicus) planiusculus (as xaxarsi Born, 1917)
Endophallus
NE Spain, Catalonia,Girona Prov., San Hilario Sacalm env.<br />
Copyright © Ruslan Panin

Carabus (Mesocarabus) riffensis Fairmaire, 1875
Endophallus
Morocco, Rif, Foresta di Ketama, 25.06.1998, 1600m, leg. D. Sechi (immature)
Copyright © Alexander Anichtchenko

Carabus (Mesocarabus) cantabricus moroderi (as magdalenai Meyer & Mollard, 1996)
Endophallus
NW-Spain, Leon prov., Sierra de Gistredo, Pto. de Magdalena,
Copyright © Ruslan Panin

Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
Endophallus
NC-Spain, Basque Co, Álava, Sierra de Gorbea.
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus helluo (as alicantinus Ganglbauer, 1886)
Endophallus
SE-Spain, Murcia reg., Jumilla, Sierra del Carche, h=1300 m,
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus trabuccarius Fairmaire, 1857
Endophallus
NE Spain, Catalonia, Tarragona, Sierra de Montsant h=800 m, Albarca env.,
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus lusitanicus (as vieirae Oliveira, 1875)
Endophallus
CW-Portugal: Leiria distr., near Marinha Grande, 39°45'57.8"N 8°58'10.3"W, h-79 m
Copyright © Ruslan Panin

Carabus (Mesocarabus) riffensis Fairmaire, 1875
Endophallus
N-Morocco, Riff Mts h=1650 m, 26 km E of Bab Berred
Copyright © Ruslan Panin

Carabus (Microplectes) riedeli Menetries, 1832
Endophallus
endophallus
Copyright © Ruslan Panin

Carabus (Microplectes) convallium convallium Starck, 1889
Endophallus
Endophallus
Copyright © Ruslan Panin

Carabus (Microplectes) convallium argonautarum Semenov, 1898
Endophallus
Endophallus
Copyright © Ruslan Panin

Carabus (Mimocarabus) pumilio Kuster, 1846
Endophallus
S-Armenia, Vayots Dzor Region, E Ughedzor vill., h=2630 m
Copyright © Ruslan Panin

Carabus (Mimocarabus) maurus maurus (Adams, 1817)
Endophallus
E Georgia, Kakheti, near Patardzeuli vill., h=750-850 m
Copyright © Ruslan Panin

Endofallus Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) illigeri illigeri Dejean, 1826 (as bjelasnicensis Apfelbeck, 1902)
Endophallus
endophallus as bjelasnicensis Apfelbeck, 1902
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei Kuster, 1846
Endophallus
W Ukraine, Vinogradov distr.
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei Kuster, 1846
Endophallus
endophallus hampei
Vinogradov distr.
Copyright © Igor Solodovnikov

Carabus (Morphocarabus) kollari Palliardi, 1825
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) wulffiusi wulffiusi Morawitz, 1862
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Endophallus
NE-Serbia, Krs Mts., Majdanpek env.,h=300m.20-31.V.2006, Pr.Sima leg
Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus spasskianus Breuning, 1932
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus schestopalovi Plutenko, 1995
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens Fabricius, 1798
Endophallus
Moldova,<br /> Anenii Noi distr., near Garbovets vill.
Copyright © Ruslan Panin

Carabus (Morphocarabus) helleri Ganglbauer, 1893
Endophallus
SE Moravia, Strani, V.2002, M.Snizek leg. male bronze colour.
Copyright © Ruslan Panin

Carabus (Morphocarabus) helleri Ganglbauer, 1893
Endophallus
SE Moravia, Strani, V.2002, M.Snizek leg. male blue colour
Copyright © Ruslan Panin

Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844: 100
Endophallus
endophallus
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844 (as melleus Lapouge, 1909)
Endophallus
<p> endophallus odoratus odoratus (as melleus Lapouge, 1909)<br /> <br />     <br />  </p>
Copyright © Ruslan Panin

Carabus (Morphocarabus) odoratus bargusinicus Shilenkov, 1996
Endophallus

    endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) monilis Fabricius, 1792
Endophallus
France 39, Foncina le Haut 300m
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) monilis Fabricius, 1792 (as interpositus Géhin, 1880)
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
Endophallus
endophallus
Copyright © Ruslan Panin

Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
Endophallus

Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) aeruginosus zyrjanovskianus Shilenkov & O.Berlov, 1996
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
Endophallus
endophallus
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens trans illigeri Dejean, 1826
Endophallus
W-Croatia, Karlovac, near Bosiljevo vill., h~300m,
Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus aeruginosiformis Breuning, 1932
Endophallus
Russia, Khakassia,
W-Sayan, Alan Mt range, valley Enisey riv., 5 km S Mayna vill.,
Copyright © Ruslan Panin

Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus shoriensis Obydov, 1999
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei Kuster, 1846 (transition to rothi incompsus Kraatz, 1880)
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) helleri Ganglbauer, 1893 (as pseudopreyssleri Breuning, 1932)
Endophallus
Czech Rep., NE Moravia, Moravian-Silesian Beskids Mt Rng, PR Trojacka, Hodslavice vill.,
Copyright © Ruslan Panin

Carabus (Morphocarabus) kollari Palliardi, 1825 (as semetricus Kraatz, 1878)
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus geblerianus Obydov, 1999
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei (as ormayi Reitter, 1896) loc.typ.
Endophallus
near Beregovo, 27.IV-22.V..2011,, Panin R. & Chyzhevsky O. leg
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens trans scheidleri
Endophallus
NC Croatia, Ivanščica Mts , Ivanec,4-18.VI.2011, Krause Rene leg.
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei Kuster, 1846
Endophallus
C Romania, Tвrgu Mureş,
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei (as zilahiensis Csiki, 1906)
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) regalis tschusovae Obydov, 2004
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi rothi Dejean, 1829
Endophallus
C Romania, Sibiu Area, right bank Olt River, Bradu vill., h=380 m,
Copyright © Ruslan Panin

Carabus (Morphocarabus) venustus venustus Morawitz, 1862
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens praecellens Pillardi, 1825
Endophallus
Serbia, Fruska Gora Mts, 20-31.V.2006, M.Sima leg.
Copyright © Ruslan Panin

Carabus (Morphocarabus) illigeri versicolor Frivaldsky, 1835
Endophallus
C Bulgaria: Stara planina Mts, Gabrovo
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi comptus (as szoerenyensis Csiki, 1908)
Endophallus
SW Romania, Tarcu Mts, vf. Nedeia h=2000m,
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei (as gutiiensis Takács & Lie, 1992)
Endophallus
N Romania, E Carpathian, Maramureş Reg., Gutai Mts, cabana Mogosa, 4 km N Baia Sprie,
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi incompsus Kraatz, 1880
Endophallus
C Romania, Brașov County, Cristian vill., h=380 m
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi alutensis Savulescu, 1972
Endophallus
C Romania, S Carpathians, Rimnicu Vilcea env., Calimaneşti, Cozia Mts.,
Copyright © Ruslan Panin

Carabus (Morphocarabus) viridilimbatus panini I.Solodovnikov 2017 Holotype
Endophallus
Russia, SE Yakutia, Neryungri distr., Tokinski Stanovik Mt Rng., Beder riv., (right tributary rivers Algama), 26.VI.1996
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei (as diffinis Csiki, 1906)
Endophallus
NW Romania, Apuseni Mts, Trascău Mt Rng, Petresti de Jos, 12 km E Turda,.
Copyright © Ruslan Panin

Carabus (Morphocarabus) illigeri illigeri (as tarensis Toševski, 1990)
Endophallus
WC-Serbia: National park Tara Mts.:NE Mitrovac h=1500m,
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei (as hunyadensis Csiki, 1926)
Endophallus
CW Romania, Apuseni Mts, Metalliferous Mt Rng, Hunedoara reg., near Săcărâmb vill.
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
Endophallus
W Ukraine,Transcarpathian Reg., Rakhiv distr., Marmarosh Mts., near Lug vill., h=400 m.
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Endophallus
Transcarpathian region, Uzhgorod distr., 28.IV-23.V.2011
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Endophallus
SE Slovakia, Slanske vrchy Mts, Herlianske sedlo, 8-19.VI.2009, Krause Rene leg.
det.Panin R.
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii zawadzkii, Kraatz 1854
Endophallus
Gorgany Mts. h=700 m, 16.VI.2011,
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii zawadzkii, Kraatz 1854
Endophallus
W.UKRAINE, Ukrainian Carpathians
Lviv Reg., Skole distr., Skolivski Beskidy Mts.
Copyright © Igor Solodovnikov

Carabus (Morphocarabus) praecellens jucundus Csiki, 1906
Endophallus
N Hungary, Pilis Mts., Piliszentlelek,20-27.VII.2011, Krause Rene leg.
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as pseudojucundus Retezár, 1974)
Endophallus
NW-Hungary::Komarom distr., Tardos, Gerecse Mts.
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as vertesensis Retezár, 1974)
Endophallus
NW Hungary: Fejer: Vertes Mt Rng, Tatabanya
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi comptus (as borlovensis Lie, 1994)
Endophallus
SW Romania,, Banat, Borlova env.,
Copyright © Ruslan Panin

Carabus (Morphocarabus) brinevi Kabak, 2014 Paratype
Endophallus

Copyright © Ilya Kabak

Carabus (Morphocarabus) odoratus baeri Menetries, 1851: 45
Endophallus
Russia, SW Taymyr, near Talnakh, 25 km N Norilsk, hilly tundra, hillside overgrown with alder slate, dwarfish birches, blueberry, h = 150-250 м, 24-28.06.2003, leg. S.Saluk
Copyright © Igor Solodovnikov

Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Endophallus
NE-Hungary, 4 km S Sátoraljaújhely, right bank riv. Bodrog (Long-erdő),
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi distinguendus Csiki, 1906
Endophallus
NC-Hungary: E- part of Budapest,Cinkota,
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi distinguendus (as bieneri Mandl, 1965)
Endophallus
C-Slovakia, Detva Distr., Stara Huta env. h=700-800m,
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi distinguendus (as subparvulus Mandl, 1965)
Endophallus
NC-Hungary:Borzsony Mts, Nograd-Diosjeno,
Copyright © Ruslan Panin

Carabus (Morphocarabus) odoratus kurmaevi I.Solodovnikov, 2017 Holotype
Endophallus
Russia, Siberia, Krasnoyarsk Krai, the North Yenisei district, 55 km of CE of the vill. Bryanka, valley of the river Chirimba, stream Nakhodny, h = 350 m, the mixed forest (birch and pine); on a steep slope along an animal track on a hill brow, 23.VII-20.VIII.2015, leg. A.V. Kurmaev
Copyright © Igor Solodovnikov

Carabus (Morphocarabus) viridilimbatus saluki I.Solodovnikov 2017 Paratype
Endophallus
Russia, SW Taymyr, near Talnakh, 25 km N Norilsk, hilly tundra, hillside overgrown with alder slate, dwarfish birches, blueberry, h = 150-250 м, 24-28.06.2003, leg. S.Saluk
Copyright © Igor Solodovnikov

Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859
Endophallus
Russia, Chita Region, Udokansky district. Kodar Range, 56 °34'42,9 " N, 118 °29'34,95" E, 6/13/2002, leg. A. Pokhorov
Copyright © Igor Solodovnikov

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Holotype
Endophallus
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill.
Copyright © Ruslan Panin

Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
Endophallus
Russia, S Tuva, Khorumnug-Taiga Mts, Buren riv. Valley, Buren-Bay-Khaak vill. h=900 m
Copyright © Ruslan Panin

Carabus (Morphocarabus) odoratus antropovi Shilenkov 1996
Endophallus
Russia, S Siberia, Tuva, W Tannu-Ola Mts. 35 km NE Khandagayty vill., Teeli Pass 2200-2800 m
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Endophallus
SE Poland, Subcarpathia Prov., Nadleśnictwo Kołaczyce N of Jasło vill., M. Kosibowicz leg./det. R Panin
Copyright © Ruslan Panin

Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
Endophallus
Russia, NE Altay, Turochak Distr., Lake Teletskoe N part, near Yaylyu vill.
Copyright © Ruslan Panin

Carabus (Morphocarabus) aeruginosus eschscholtzi Mannerheim, 1827
Endophallus
NE Kazakhstan, W Altay Mts, near Shemonaikha city., flood plain,
Copyright © Ruslan Panin

Carabus (Neoplectes) starckianus ganglbaueri Reitter, 1888
Endophallus

Copyright © Alexander Anichtchenko

Carabus (Neoplectes) daphnis daphnis Kurnakov, 1962
Endophallus
Georgia, Mingrelia, Lecheha vallis, Tehuri superior
Caucasus centr., 1200m, 27.6-7.7.1989 Retezar I. leg.
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) daphnis askhicus Belousov, 1992
Endophallus
Georgia, env. Martvilli, Askhi plateau
10-15.7.2001 A.Rubenian leg.
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) daphnis askhicus Belousov, 1992
Endophallus
W Georgia, valley of Chkhorodukhu riv., 1000-1200m
14.V-6.VII.1991
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) daphnis askhicus Belousov, 1992 Paratype
Endophallus

Copyright © Ruslan Panin

Carabus (Neoplectes) juenthneri avadcharensis Kurnakov, 1970
Endophallus

Copyright © Ruslan Panin

Carabus (Neoplectes) reitteri laevisternis Starck, 1890
Endophallus

Copyright © Ruslan Panin

Carabus (Neoplectes) prometheus prometheus Reitter, 1887 (as nubicola Zolotarev, 1913)
Endophallus
Endophallus prometheus prometheus (as nubicola Zolotarev, 1913)
Copyright © Ruslan Panin

Carabus (Neoplectes) prometheus prometheus Reitter, 1887
Endophallus
Endophallus prometheus prometheus Reitter, 1887
Copyright © Ruslan Panin

Carabus (Neoplectes) prometheus prometheus Reitter, 1887
Endophallus
aedeagus
Copyright © Ruslan Panin

Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990
Endophallus

Copyright © Ruslan Panin

Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990
Endophallus
aedeagus miroshnikovi Psebai env., Nikitino
Copyright © Ruslan Panin

Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890
Endophallus
Endophallus
Copyright © Ruslan Panin

Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
Endophallus

Copyright © Ruslan Panin

Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988 (as shchessiensis) Paratype
Endophallus

Copyright © Ruslan Panin

Carabus (Neoplectes) satyrus duripshensis Kurnakov, 1972
Endophallus

Copyright © Safronov

Carabus (Neoplectes) apollo tenebricosus Kurnakov, 1962
Endophallus

Copyright © Alexandr Safronov

Carabus (Neoplectes) edithae exedithae Gottwald, 1985
Endophallus
NС-Caucasus, Karachaevo-Cherkessia, right bank Pkhiya river, Abishir-Akhuba Mt.Range h=2250-2600 m,
Copyright © Ruslan Panin

Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014
Endophallus

Copyright © Igor Solodovnikov

Carabus (Neoplectes) satyrus s.str.
Endophallus

Copyright © Igor Solodovnikov

Carabus (Neoplectes) pseudopshuensis Zamotajlov, 1991
Endophallus

Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus besleticus Kurnakov, 1972
Endophallus

Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972
Endophallus

Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus dsychvensis Gottwald, 1985
Endophallus

Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus duripshensis Kurnakov, 1972
Endophallus

Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus napraensis Belousov & Zamotajlov, 1993 Paratype
Endophallus

Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus anastasiae Solodovnikov, 2014 Paratype
Endophallus

Copyright © Igor Solodovnikov

Carabus (Neoplectes) ibericus ibericus (as imereticus Retezar, 2011) Paratype
Endophallus
NC-Georgia, S-Caucasus: W slope Likhi Mt.Rng., 3 km W Rikoti Pass h~900 m,
Copyright © Ruslan Panin

Carabus (Neoplectes) compressus rebellis Reitter, 1895
Endophallus
Georgia, Kvemo-Svaneti, S slopes of Svanetskiy Mt.R., S slope of Latpari pass, h=2225m, 17.VI -13.VII.2017 leg. D.Fominykh
Copyright © Kirill Makarov

Carabus (Neoplectes) polychrous svetlanae Solodovnikov, 2015
Endophallus

Copyright © Igor Solodovnikov

Carabus (Neoplectes) polychrous rousianus Gottwald, 1985
Endophallus

Copyright © Igor Solodovnikov

Carabus (Neoplectes) polychrous polychrous
Endophallus

Copyright © Igor Solodovnikov

Carabus (Neoplectes) polychrous khipstaensis Solodovnikov, 2015
Endophallus

Copyright © Igor Solodovnikov

Carabus (Neoplectes) martviliensis Retezar & Djavelidze, 1992
Endophallus
Zeda Gordi env., 980 m and Doberazeni env., 600 m.
Copyright © Kirill Makarov

Carabus (Neoplectes) C. martviliensis mzekalae
Endophallus
Melouri env., 475 m and Dzedzileti env. , 770 m
Copyright © Kirill Makarov

Carabus (Neoplectes) jason Semenov, 1898
Endophallus

Copyright © Igor Solodovnikov

Carabus (Neoplectes) zolotarevi Zamotajlov, 1988
Endophallus
Carabus (Archiplectes) zolotarevi zolotarevi Zamotajlov, 1988 Russia: Krasnodar reg., SW slopes of Malye Balkany Mt.R., env. Allous pass, h = 1960 m, 08-18.VII.2023 leg. D. Fominykh
Copyright © Igor Solodovnikov

Carabus (Neoplectes) ibericus lafertei Chaudoir, 1846
Endophallus
Georgia, Lesser Caucasus, Imereti prov., Meskheti Mt Rng, N slope, Zekari Pass h~2150 -2250 m alpine meadows
Copyright © Ruslan Panin

Carabus (Neoplectes) protensus protensoides Novotny & Vorisek, 1988
Endophallus
W Georgia, Martvili distr., W slope of Askhi Plateau h=1200-1800 m,
Copyright © Ruslan Panin

Carabus (Ohomopterus) japonicus japonicus Motschulsky, 1857
Endophallus
S Japan, Kyushu, Kumamoto Pref., Izumi viil., h=600 m
Copyright © Ruslan Panin

Carabus (Ohomopterus) yamato kitai (Imura, 2004)
Endophallus
Japan, S Honshu, Mie Pref., Hakusan,
Copyright © Ruslan Panin

Carabus (Ohomopterus) uenoi Ishikawa, 1960
Endophallus

Copyright © Jong-Bong Choi

Carabus (Ohomopterus) dehaanii dehaanii Chaudoir, 1848
Endophallus
Japan, S Honshu, Mie Pref., Inabe city
Copyright © Ruslan Panin

Carabus (Ophiocarabus) nilkiensis Kabak, 2014 Paratype
Endophallus

Copyright © Ilya Kabak

Carabus (Ophiocarabus) wusunshanicus Kabak, 2015
Endophallus

Copyright © Ilya Kabak

Carabus (Ophiocarabus) regeli regeli Morawitz, 1886
Endophallus
Karatau Mt. R.
Copyright © Ilya Kabak

Carabus (Ophiocarabus) successor successor Reitter, 1896
Endophallus
SE-Kazakhstan, Narynkol distr., 4 km SW Bayankol, h~2400-2500 m
Copyright © Ruslan Panin

Carabus (Ophiocarabus) ernsti ulastaiensis Kabak, 2019 Paratype
Endophallus
China, Xinjiang, S slopes of Boro-Horo Mt. R., ridge between Kaptshik and Zekku rivers, NE of Ulastai (= Wulasitai) Vill., alpine meadows, 43°56'07"N / 83°18'33"E – 43°56'22"N / 83°18'30"E, 3135–3280 m, 17.07.2017 leg I.I. Kabak
Copyright © Kirill Makarov

Carabus (Ophiocarabus) successor eoketmenicus Kabak, 2016 Holotype Paratype
Endophallus
endophallus - paratypes; aedeagus - holotype
Copyright © Ilya Kabak

Carabus (Ophiocarabus) kurdaiensis Kabak, 2016 Paratype
Endophallus

Copyright © Ilya Kabak

Carabus (Ophiocarabus) ernsti Kabak, 2001
Endophallus
7–8, 12–13 – C. ernsti ulastaiensis, paratypes; 9–10, 14–15 – C. ernsti ernsti Kabak, 2002: 9, 14 – from the locality in mountains to the West of the Saryk River, 10, 15 – from the locality in right tributary of the Saryk River; 11,16 – C. ernsti nilkiensis Kabak, 2014: 11 – holotype, 12 – paratype. ml – median lobe; pp – praeputial pad; rbll – right basolateral lobe.
Copyright © Ilya Kabak

Carabus (Pachystus) hortensis hortensis Linne, 1758
Endophallus
W-Ukraine, Male Polissya, near Kamenka-Buzka vill.
Copyright © Ruslan Panin

Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847
Endophallus
WC Spain, Cáceres, Hervás, h=1000m
Copyright © Ruslan Panin

Carabus (?Oreocarabus) qinlingensis Imura, 1993
Endophallus
China, W-Shaanxi, Taibai Shan Mts, btw.Houzhenzi/Ba Xian Tai, nr.Nan Tian Men h=3200m, 33°55'17"N/107°46'53"E,
Copyright © Ruslan Panin

Carabus (Oreocarabus) latro minshanensis Deuve, 1987
Endophallus
China, N-Sichuan, Jiuzhaigou env.,
Copyright © Ruslan Panin

Carabus (Oreocarabus) luetgensi pseudosteuarti Lapouge, 1924
Endophallus
Portugal, Guarda, Sierra de Estrela h=800 m,
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei linnaei Panzer, 1812
Endophallus
W.UKRAINE, Lviv Reg., NE Carpathians, Skolivski Beskidy Mts. Zelemyanka Mt. Rng
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018
Endophallus
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone,
Copyright © Ruslan Panin

Carabus (Orinocarabus) sylvestris transsylvanicus Dejean, 1826
Endophallus
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone,
Copyright © Ruslan Panin

Carabus (Orinocarabus) bertolinii Kraatz, 1878
Endophallus
NE-Italy, Trentino prov. Dolomites Alps, Plateau Pale di San Martino h=2500 m,
Copyright © Ruslan Panin

Carabus (Pachycarabus) staehlini Adams, 1817
Endophallus
S Russia, E Caucasus, Dagestan, Nukatl Mt Rng h=2600m,
Copyright © Ruslan Panin

Carabus (Pachycarabus) imitator katherinae Reitter, 1898
Endophallus
Abkhasia, Sukhumi distr., Anchkho Mt. Pass E- part h=1870-1920 m. det. I. Solodovnikov, 2025
Copyright © Ruslan Panin

Carabus (Pachystus) tamsi tamsi Menetries, 1832
Endophallus
endophallus
Copyright © Ruslan Panin

Carabus (Pachystus) glabratus glabratus Paykull, 1790
Endophallus
Endophallus
Copyright © Ruslan Panin

Carabus (Pachystus) hungaricus cribellatus Adams, 1812
Endophallus

Copyright © Ruslan Panin

Carabus (Pachystus) hungaricus scythus Motschulsky, 1847
Endophallus

Copyright © Ruslan Panin

Carabus (Pachystus) hungaricus hungaricus Fabricius, 1792
Endophallus
Czech, S Moravia, Pouzdrany
Copyright © Ruslan Panin

Carabus (Pachystus) cribratus cribratus Quensel, 1806
Endophallus
NW Armenia, Amasia, (Shirak prov.)
Copyright © Ruslan Panin

Carabus (?Oreocarabus) titanus Breuning, 1933
Endophallus
C-China Sichuan prov., Nanping 1500m,
Copyright © Ruslan Panin

Carabus (Platycarabus) irregularis montandoni Buysson, 1882
Endophallus
endophallus
Copyright © Ruslan Panin

Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Endophallus
Endophallus fabricii malachiticus
Copyright © Ruslan Panin

Carabus (Platycarabus) fabricii ucrainicus Lazorko, 1951
Endophallus

Copyright © Ruslan Panin

Carabus (Procrustes) piochardi praestigiator Morawitz, 1886
Endophallus
C-Lebanon, Byblos /Jbeil Distr., Kfardebian env., h=1200m,
Copyright © Ruslan Panin

Carabus (Procrustes) paiafa paiafa White, 1845
Endophallus

Copyright © Kirill Makarov

Carabus (Procrustes) coriaceus ressli Mandl, 1961
Endophallus
SC-Turkey:Hatay Prov, Dörtyol, Nur Mts, Topaktaş h=900-1200 m,
Copyright © Ruslan Panin

Carabus (Pseudocoptolabrus) watanabei valdemar Deuve, 2016 Paratype
Endophallus
Chine, Yunnan, Dêqên, monts Hengduan Shan, près du col de Doker La, 3600 m
Copyright © Thierry Deuve

Carabus (Pseudocoptolabrus) belousovianus Imura, 2011
Endophallus
Image from Imura, 2011
Copyright © - - -

Carabus (Rhigocarabus) ladygini degensis Deuve, 1989
Endophallus
C-China, NW- Sichuan prov., Garzê Tibetan AP, Pass Garzê – Sêrtar
Copyright © Ruslan Panin

Carabus (Rhigocarabus) gigolo Heinz & Brezina, 1996
Endophallus
China, Tewo. det. Anichtchenko, A. 2020
Copyright © Alexander Anichtchenko

Carabus (Rhigocarabus) buddaicus gansuicus Deuve, 1991
Endophallus
N Sichuan, Langmusi vill 3700 m, 1-25.7.2018
Copyright © Alexander Anichtchenko

Carabus (Rhigocarabus) roborowskii langmusiensis Deuve, 1996
Endophallus
N Sichuan, near Langmusi, 3700m 1-25.7.2018, V. Patrikeev leg. det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Carabus (Rhigocarabus) casanova (Imura & Brezina, 2008)
Endophallus

Copyright © Boleslav Brezina

Carabus (Rhigocarabus) gigolo Heinz & Brezina, 1996
Endophallus
Langmusi
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) gentleman Brezina & Hackel, 2004
Endophallus
Jiangjing
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) gigoloides priapeius Cavazzuti, 2006
Endophallus
Yeliguan
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) nianjuaensis Imura, 2016
Endophallus
Image from Imura, 2016
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Carabus (Rhigocarabus) kozaburoi Imura, 1999 Paratype
Endophallus
Gaoligong Shan, N. Myanmar. Image from Imura, 2016
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Carabus (Rhigocarabus) itzingeri rugulosior Deuve, 1996 Holotype
Endophallus
NW-Yunnan: Hengduan Shan: Yanmen env.: 28:06N/98:52E. Image from Imura, 2016
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Carabus (Rhigocarabus) itzingeri tanakai Imura, 1997
Endophallus
Image from Imura, 2016
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Carabus (Rhigocarabus) itzingeri choguy Imura, 2001 Holotype
Endophallus
Image from Imura, 2016
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Carabus (Rhigocarabus) rhododendron xuebangshanensis Imura, 2016 Holotype
Endophallus
Image from Imura 2016
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Carabus (Rhigocarabus) rhododendron xinshanchangensis Imura, 2016 Holotype
Endophallus
Image from Imura, 2016
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Carabus (Rhigocarabus) rhododendron lidimaensis Imura, 2016 Holotype
Endophallus
Image from Imura, 2016
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Carabus (Rhigocarabus) rhododendron s.str. Holotype
Endophallus
Image from Imura, 2016
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Carabus (Rhigocarabus) roborowskii roborowskii Semenov, 1887
Endophallus
China, E-Qinghai, Mts NW of Tongren city, 35.562778, 101.843667, h=3870 m
Copyright © Ruslan Panin

Carabus (Semnocarabus) carbonicolor carbonicolor Morawitz, 1886
Endophallus

Copyright © Alexandr Safronov

Carabus (Semnocarabus) erosus kuldshaensis Ballion, 1878
Endophallus
S-Kazakhstan:Ili River valley,:near Bakanas,
Copyright © Ruslan Panin

Carabus (Semnocarabus) carbonicolor korolkowi Semenov, 1896
Endophallus
NC-Kyrgyzstan, Kyrgyz Mt.Rng , 10 km SW of Bishkek, Chon-Aryk env., h=1100m,
Copyright © Ruslan Panin

Carabus (Semnocarabus) erosus karascharensis Eidam Lectotype Paralectotypes
Endophallus
aedeagus - lectotype. Endophallus - paralectotype.
Copyright © Kirill Makarov

Carabus (Semnocarabus) cicatricosulus pseudoerosus Mandl, 1955
Endophallus
left tributary of Kuqa River, E of Lake Kyrghyz-Artukul (= Dalongchi), 42°27'15''N, 83°22'00''E, H ~ 2550–2600 m, 7–8.VI.2002, I. Kabak leg
Copyright © Kirill Makarov

Carabus (Semnocarabus) erosus corax Semenov, 1896
Endophallus
SE-Kazakhstan, Almaty reg., C Trans-Ili Alatau N-slopes, Kaskalen river Valley, Izvestkovaya vill., h=1700 m, 6.V.2015, D.Kovalchuk leg./det. R.Panin,
Copyright © Ruslan Panin

Carabus (Sigenacarabus) trachynodes moxiensis Deuve, 1992
Endophallus
China, W-Sichuan, Daxue Range, Mt.Gongga Shan h=3000-3100m, Hailuogou Glacier Forest Park, 29.574889, 101.995556
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) varians (as dvorshaki Mandl, 1975)
Endophallus

Copyright ©

Carabus (Sphodristocarabus) macrogonus macrogonus Chaudoir, 1847
Endophallus
NE-Turkey:Trabzon env., Sumela,
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) macrogonus pseudokolatensis Heinz & Korge, 1967
Endophallus
NE-Turkey::Kackar Dagi Mt, Ayder S of Ardesen h=1600m
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) varians armeniacus Mannerheim, 1830
Endophallus
NE Turkey, Artvin Prov., Şavşat, Зamlıca.
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) separatus persa Lapouge, 1924
Endophallus
N-Iran: Mazandaran Province, Elborz Mt.Rng, Vandarbon env.
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) blumenthaliensis aequicostalis Staven & Heinz, 1995
Endophallus

Copyright © Ruslan Panin

Carabus (Sphodristocarabus) adamsi Adams, 1817
Endophallus
CN-Georgia, N of Tbilisi, NW of Tianeti, Tvalivi env.,
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) separatus separatus Gehin, 1885
Endophallus
N-Iran:Gilan Prov., Asalem - Khalkal road,
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) scovitzi scovitzi Faldermann, 1835
Endophallus
Georgia, Adzharia, Adzharo-Imeretinskiy Mt.R., near Tchidila pass, alpin zone, h=2350 m, 05.VI.2016 leg. D. Fominykh
Copyright © Igor Solodovnikov

Carabus (Sphodristocarabus) tokatensis kosensis Cavazzuti, 1984
Endophallus
CE-Turkey: Gümüshane: Köse Dag(pass N of Köse), h = 1700 m, VII.2000, leg. St. Il. Remond (TOPOTYPUS)
Copyright © Igor Solodovnikov

Carabus (Tachypus) cancellatus cancellatus Illiger, 1798
Endophallus
Carabus (Tachypus) cancellatus tuberculatus f.conspersus Lapouge, 1902 (Blaс form) N-Belarus, Mogilev reg., Bykhov distr., 5 km N Voronino Vill., mixed-pine forest, 20.V-03.VI.2007, Komyagunova N.P.
Copyright © Igor Solodovnikov

Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
Endophallus
Carabus (Tachypus) cancellatus emarginatus n. dahli Heer, 1841
Copyright © Igor Solodovnikov

Carabus (Tomocarabus) bessarabicus ? concretus Fischer, 1823
Endophallus
Steppe Reserve “Provalsky",
Copyright © Ruslan Panin

Carabus (Oreocarabus) taedatus agassi Le Conte, 1850
Endophallus
SC-USA, New Mexico, Santa Fe,
Copyright © Ruslan Panin

Carabus (Trachycarabus) estreicheri Fischer, 1822
Endophallus
Lviv Region, Brody distr., near Yasenov vill.,
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus tscherkessicus Korge, 1964
Endophallus

Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886
Endophallus

Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886
Endophallus
aedeagus
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus obliteratus Fischer, 1827
Endophallus
endophallus sibiricus obliteratus Fischer, 1827
NE Kazakhstan,
S Altai mts.,S Ulbinsky mt.h=500m.,Turgusun vill., 20.VI.2001
male 25mm.
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus haeres (as fossulatus Dejean, 1826)
Endophallus
endophallus
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus errans Fischer, 1823
Endophallus
endophallus sibiricus errans
SW-Russia,
Rostov reg. env. stanitsa Razdorskaya, A.Ponomarev leg
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus errans Fischer, 1823
Endophallus
endophallus sibiricus errans- Lugansk reg., near Slavyanoserbk
Copyright © Ruslan Panin

Carabus (Trachycarabus) scabriusculus scabriusculus Olivier, 1795
Endophallus

Copyright © Ruslan Panin

Carabus (Trachycarabus) scabriusculus inapertus Motschulsky, 1850 (loc.typ)
Endophallus

Copyright © Ruslan Panin

Carabus (Trachycarabus) planus planus Gehin, 1885
Endophallus
<b>endophalus </b> S Russia,Krasnodar reg., Anapa distr., B. Utrish vill., env.,
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus planus Gehin, 1885
Endophallus
<b>endophalus</b> Karacaj-Cherkessija, near Kart-Djurt vill., upper Kylan riv. h=1300m
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus campicola Reitter, 1896
Endophallus

Copyright © Ruslan Panin

Carabus (Trachycarabus) mandibularis buchtarmensis Kryzhanovskij, 1953
Endophallus

Copyright © Ruslan Panin

Carabus (Trachycarabus) besseri Fischer, 1822
Endophallus

Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus gvozdevae Obydov, 1997
Endophallus

Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus sibiricus Fischer, 1822
Endophallus
near Krasnoyarsk
Copyright © Ruslan Panin

Carabus (Trachycarabus) latreillei ?semicoriaceus Kraatz, 1881
Endophallus
NE-China: Liaodong Peninsula, Liaoning prov., Gaizhou distr., Mt. Buyun Shan.
Copyright © Ruslan Panin

Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
Endophallus
N Mongolia, Khövsgöl reg., Lake Khovsgol, h=1640m, 51*:12’N; 100*:45’E, steppificated sites,
Copyright © Ruslan Panin

Carabus (Tribax) titan titan Zolotarev, 1913
Endophallus
endophallus
Copyright © Ruslan Panin

Carabus (Tribax) steveni satanas Semenov, 1896
Endophallus
endophallus
Copyright © Ruslan Panin

Carabus (Tribax) circassicus kubaniensis Semenov, 1896
Endophallus
endophallus
Copyright © Ruslan Panin

Carabus (Tribax) apschuanus pseudoplatessa Gottwald, 1982
Endophallus
endophallus
Copyright © Ruslan Panin

Carabus (Tribax) puschkini ishikawaianus Breuning & Ruspoli, 1970
Endophallus
NE-Turkey: Artvin:Hopa, Borзka env., h=450 m,
Copyright © Ruslan Panin

Carabus (Tribax) puschkini ponticus Deyrolle, 1869
Endophallus
NE-Turkey:Trabzon, Zigana Pass h=1700m,
Copyright © Ruslan Panin

Carabus (Tribax) agnatus pseudagnatus Novotný & Voříšek, 1988
Endophallus
W-Caucasus: Abkhasia, Gudauta distr., Mt Anchkho h=2200-2350 m,
Copyright © Ruslan Panin

Carabus (Tribax) puschkini platypterus Ganglbauer, 1886
Endophallus
SW Georgia, Lesser Caucasus, Adzhar-Imereti (Meskheti) Mt Range, 10 km S Sairme h~1850 m.
Copyright © Ruslan Panin

Carabus (Tribax) puschkini kolenatii (as suramensis Kraatz, 1887)
Endophallus
S-Caucasus: NC-Georgia, W slope Likhi Mt.Range, 3 km W Rikoti Pass h~900m
Copyright © Ruslan Panin

Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
Endophallus
S Russia, C-Caucasus: N Ossetia, Chekhatsirtit Mt range, Mt Tepli h=3000 m
Copyright © Ruslan Panin

Carabus (Tribax) adelphus rex Gottwald, 1980
Endophallus
Abkhazia, Tkuarchalsky district, Kodorsky Mt.R., it is right board of the Bashishkvara River, beechen forest + rhododendron, h = 1693 m, 15-23.07.2018, leg. I.A. Solodovnikov
Copyright © Igor Solodovnikov

Carabus (Tribax) adelphus adelphus Rost, 1892 SW-Caucasus:"Samursakan"(=nr.Gali & Anara); Gali env.:Okumi
Endophallus
Georgia, Zemo Svaneti., W slopes of Ingurishi Mt., right bank of Inguri riv., 09.V-20.VI.2018 leg. D. Fominykh
Copyright © Igor Solodovnikov

Carabus (Tribax) nacharensis Rost, 1893
Endophallus
Abkhazia, vall. Kluch River, 13.08.1928
Copyright © Igor Solodovnikov

Carabus (Tribax) adelphus shtavlerensis Fominykh, Zamotajlov et Titarenko, 2018
Endophallus
Georgia, Zemo Svaneti, left bank of Nenskra River near Tita Vill., Shtavler Mt. Range, NW slopes of Utur Mt., 43°04'N, 42°11'E, 1048 m, 14.IV–13.VI.2016 (D. Fominykh) HOLOTYPE
Copyright © Igor Solodovnikov

Carabus (Tribax) macropus Chaudoir, 1877
Endophallus
Russia, Ingushetia, S slopes of Skalistyi Mt.R., Khahalgi Mt., alpin zone, h = 2500 m
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Endophallus
Russia, Caucasus, Ingushetia, South slopes of Skalistyi Mt Range, Stolovaya Mt., near Beyni, alpin zone, h = 2800 m, N 42°51’35.30",E 44°42’13.70", h = 2800m, 21.V–19.VII.2018, D. Fominykh leg. (TOPOTYPUS)
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Endophallus
Russia, Caucasus, Ingushetia, Skalistyi Mt. Range, S slopes of Tsoreylam Mt. Range, 5 km W of Mt. Khakhalgi, N 42°50’07.60", E 45°05’02.20", h=2500 m, 25.V–21.VII.2017, D. Fominykh leg.
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
Endophallus
Russia: N.Osetia, Bokovoy Mt.R., near Arkhonskiy pass., alpin zone, h = 2700 m, 23.V-21VII.2018, D. Fominykh leg.
Copyright © Igor Solodovnikov

Carabus (Tribax) apschuanus galianus Belousov & Zamotajlov, 1993
Endophallus

Copyright © Igor Solodovnikov

Carabus (Tribax) edmundi Semenov, 1896
Endophallus
Russia, S-Dagestan
Copyright © Igor Solodovnikov

Carabus (Tribax) osseticus abanosicus Deuve, 2013
Endophallus

Copyright © Igor Solodovnikov

Carabus (Tribax) osseticus osseticus Adams, 1817
Endophallus

Copyright © Igor Solodovnikov

Carabus (Tribax) puschkini kraatzi Chaudoir, 1877
Endophallus
Georgia, Samtskhe-Javakheti, env. Uraveli vill., 14.VI.2024 leg. D. Fominykh
Copyright © Igor Solodovnikov

Carabus (Ulocarabus) stschurovskii Solsky, 1874
Endophallus
Tadzhikistan, Pamir-Alay, S slope Gissar Mt.Range, Takob, h-1700m,
Copyright © Ruslan Panin

Carabus (Ulocarabus) theanus Reitter, 1895
Endophallus
Tadzhikistan, S slope Gissar Mt.Range, left bank of the river Sorbo, Haidar gorge,
Copyright © Ruslan Panin

Carabus (Ulocarabus) theanus theanus Reitter, 1895
Endophallus
Tadzhikistan, S slope Gissar Mt.Range, left bank of the river Sorbo, Haidar gorge,
Copyright © Ruslan Panin