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Carabidae Carabidae of the World
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  • Aplothorax burchelli Waterhouse, 1841
    Aplothorax burchelli Waterhouse, 1841
    Male adult
    St. Helena island
    Copyright © Boleslav Brezina
    Aplothorax burchelli Waterhouse, 1841
    Aplothorax burchelli Waterhouse, 1841
    Female adult
    Sainte-Hélène NE, Horse Point Plain, 1300-1400 ft., V.1967. P. Basilewsky det.
    Copyright © Gianluigi Alzona
    Aplothorax taxon page (2 photos)
  • Calosoma (Castrida) argentinensis Csiki, 1927
    Calosoma (Castrida) argentinensis Csiki, 1927
    Larva
    Dep. Boqueron,
    Estancia La Paz
    S 21º01´ W59º47´
    04. 01. 2010
    Copyright © Ulf Drechsel
    Calosoma (Calosoma) inquisitor cupreum Dejean, 1826: 196
    Calosoma (Calosoma) inquisitor cupreum Dejean, 1826: 196
    Female adult
    Armenia, Zangezur range, Meghri suburbs., Vahravar vill., Rivulet vall., h=1400 m Yuriy Shevnin leg.; 23.05.2007; 21 mm.
    Copyright © Anton Menstschikov
    Calosoma (Calosoma) inquisitor cupreum Dejean, 1826: 196
    Calosoma (Calosoma) inquisitor cupreum Dejean, 1826: 196
    Male adult
    Iran, Ardabil, 30 km NE Ardabil,
    5 km SE Namin, 18-19.5.2007 Anichtchenko A. leg.
    male, 20 mm
    Copyright © Alexander Anichtchenko
    Calosoma (Calosoma) schayeri schayeri Erichson, 1842
    Calosoma (Calosoma) schayeri schayeri Erichson, 1842
    Male adult
    Australia N.T. Alice Springs, Airport
    19.5.2000 M. Langer leg.
    male, 21 mm
    Copyright © Alexander Anichtchenko
    Calosoma (Callisphaena) reticulatum reticulatum s.str.
    Calosoma (Callisphaena) reticulatum reticulatum s.str.
    Female adult
    Germania, or. DDR, Bezirk Frankfurt, 24.V.1989
    female, 25 mm
    photo & det. Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Calosoma (Callistenia) moniliatus Le Conte, 1851: 200
    Calosoma (Callistenia) moniliatus Le Conte, 1851: 200
    Male adult
    SE Canada, Nueva Escocia
    19.5.1970
    male, 17 mm
    Copyright © Alexander Anichtchenko
    Calosoma (Callisthenes) anthrax Semenov, 1900
    Calosoma (Callisthenes) anthrax Semenov, 1900 Holotype
    Male adult
    collection of ZIN
    photo Anichtchenko A. 2010
    Copyright © Alexander Anichtchenko
    Calosoma (Callisthenes) breviusculus Mannerheim, 1830
    Calosoma (Callisthenes) breviusculus Mannerheim, 1830
    Male adult
    Armenia, near Ashtarik<br /> 15.6.1997<br /> male, 22 mm<br /> photo, det. & coll. Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Calosoma (Callisthenes) elegans s.str. (as declivis Dohrn, 1884b)
    Calosoma (Callisthenes) elegans s.str. (as declivis Dohrn, 1884b)
    Male adult
    SE Kazakhstan Alma-Ata reg., 30 km<br /> N Sary-Ozek 28.4.1998 Klimenko leg.<br /> male, 22 mm
    Copyright © Alexander Anichtchenko
    Calosoma (Callisthenes) elegans s.str. (as declivis Dohrn, 1884b)
    Calosoma (Callisthenes) elegans s.str. (as declivis Dohrn, 1884b)
    Female adult
    SE. Kazakhstan, Sary-Olek vill., 26.IV.1997
    Copyright © Anton Menstschikov
    Calosoma (Callisthenes) elegans amethystinus Semenov & Redikorzev, 1928
    Calosoma (Callisthenes) elegans amethystinus Semenov & Redikorzev, 1928
    Male adult
    SE Kazakhstan, E Zailiyski mt. rng. Chilik riv.
    Malybai vill. 28.4.1999 Klimenko A. leg.
    male, 25 mm
    Copyright © Alexander Anichtchenko
    Calosoma (Callisthenes) elegans amethystinus Semenov & Redikorzev, 1928
    Calosoma (Callisthenes) elegans amethystinus Semenov & Redikorzev, 1928
    Male adult
    26 mm.
    Zailiyski Alatay.
    06.1997 Chibilov E. leg.
    Copyright © Sunvas
    Calosoma (Callisthenes) elegans amethystinus Semenov & Redikorzev, 1928
    Calosoma (Callisthenes) elegans amethystinus Semenov & Redikorzev, 1928
    Female adult
    SE Kazakhstan, E Zailiyski mt. rng. Chilik riv.
    Malybai vill. 28.4.1999 Klimenko A. leg.
    female, 26 mm
    Copyright © Alexander Anichtchenko
    Calosoma (Callisthenes) elegans elegans s.str.
    Calosoma (Callisthenes) elegans elegans s.str.
    Male adult
    SE Kazakhstan, Tchu-Iliyskiy Mts., Kordai pass, h=1500; A. Klimenko leg.; 20.04.1997;
    24 mm.
    photo A. Menstschicov
    Copyright © Anton Menstschikov
    Calosoma (Callisthenes) elegans karagaicus Lapouge, 1924
    Calosoma (Callisthenes) elegans karagaicus Lapouge, 1924
    Male adult
    Inner Tian-Shan, 20 km N jf Naryn
    27.5.1997 Klimenko A. leg.
    male, 27 mm
    photo, det. & coll. Anichtchenko A. 2009
    Copyright © Alexander Anichtchenko
    Calosoma (Callisthenes) elegans karagaicus Lapouge, 1924
    Calosoma (Callisthenes) elegans karagaicus Lapouge, 1924
    Male adult
    KIRGHIZSTAN, Inner Tian-Shan Mts., near Naryn tn., Naryn riv. Valley, 18-20.V.1999, Leg. Irtlatch A.
    Copyright © Anton Menstschikov
    Calosoma (Callisthenes) elegans manderstjernae Ballion, 1870
    Calosoma (Callisthenes) elegans manderstjernae Ballion, 1870
    Female adult
    Kirgizia, S Kirgizskii mt. rng., 2200m
    Kichi-Kimin riv., Ak-Tuz vill. 05.1998 Klimenko leg.
    female, 32 mm
    Copyright © Alexander Anichtchenko
    Calosoma (Callisthenes) elegans semenovi Motschulsky, 1859
    Calosoma (Callisthenes) elegans semenovi Motschulsky, 1859
    Female adult
    SE Kazakhstan, Zailijsky Mt. rng.,
    Talgar env., 1500 m, 21.5.2000
    female, 27 mm
    Copyright © Alexander Anichtchenko
    Calosoma (Callisthenes) eversmanni Chaudoir, 1850
    Calosoma (Callisthenes) eversmanni Chaudoir, 1850
    Male adult
    Syria, Antilibanon
    1.3.1991
    male, 22 mm
    Copyright © Alexander Anichtchenko
    Calosoma (Callisthenes) fischeri Fischer, 1842
    Calosoma (Callisthenes) fischeri Fischer, 1842
    Male adult
    Mongolia, Mandalgob, 20 km
    N Amardalai vill. 11-12.6.2003 Anichtchenko A. leg
    male, 19 mm
    photo, det. & coll. Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Calosoma (Callisthenes) grumi Semenov, 1900
    Calosoma (Callisthenes) grumi Semenov, 1900 Holotype
    Female adult
    collection ZIN
    photo Anichtchenko A. 2010
    Copyright © Alexander Anichtchenko
    Calosoma (Callisthenes) breuningi Mandl, 1954
    Calosoma (Callisthenes) breuningi Mandl, 1954
    Female adult
    E Kazakhstan, S Alakol lake, Koktuma vill., 1-10.5.1999 Nekroshevich S. leg. female, 22 mm; det. & coll. Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Calosoma (Callisthenes) breuningi Mandl, 1954
    Calosoma (Callisthenes) breuningi Mandl, 1954
    Male adult
    E.Kazakhstan, Alakol lake, Koktuma vill., 07.V.1999
    Copyright © Anton Menstschikov
    Calosoma (Callisthenes) breuningi iliense Kabak, 2022
    Calosoma (Callisthenes) breuningi iliense Kabak, 2022
    Male adult
    SE Kazakhstan, near Sary Ozek vill., A. Klimenko leg; 22.04.1997; 22 mm.; det. Anichtchenko A. 2022
    Copyright © Anton Menstschikov
    Calosoma (Callisthenes) kuschakewitschi kuschakewitschi s.str.
    Calosoma (Callisthenes) kuschakewitschi kuschakewitschi s.str.
    Male adult
    Karzhantau mt. rng., Susungen riv.,
    2300 m, 7.6.1998 Tretjakov leg.
    male, 23 mm
    Copyright © Alexander Anichtchenko
    Calosoma (Callisthenes) kuschakewitschi plasoni Born, 1917
    Calosoma (Callisthenes) kuschakewitschi plasoni Born, 1917
    Male adult
    Kirgizia, 40 km S Bishkek
    10.5.1990
    male, 26 mm
    Copyright © Alexander Anichtchenko
    Calosoma (Callisthenes) kuschakewitschi plasoni Born, 1917
    Calosoma (Callisthenes) kuschakewitschi plasoni Born, 1917
    Female adult
    Kirgizia, Chon-Aryk. 23.V.1998<br /> V. Dolin leg.
    Copyright © Denis Potanin
    Calosoma (Callisthenes) marginatus Gebler, 1829
    Calosoma (Callisthenes) marginatus Gebler, 1829
    Female adult
    NE Kazakhstan, N Zaisan lake, 20.6.2000
    Narymskii mt.rng. Klimenko A. leg.
    female, 18 mm
    photo, det. & coll. Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Calosoma (Callisthenes) mirificus Casale, 1979
    Calosoma (Callisthenes) mirificus Casale, 1979
    Female adult
    Afganistan, S slope of Paropamiz Mt. rng., near Sabzan pass, 2400m, 22.VI.1967 M.Stoljarov leg.
    Collection of ZIN
    det. D. Obydov 1999
    photo Anichtchenko A. 2010
    Copyright © Alexander Anichtchenko
    Calosoma (Callisthenes) panderi Fischer, 1822
    Calosoma (Callisthenes) panderi Fischer, 1822
    Male adult
    NW Kazakhstan, Aktiubinsk reg., Baiganinsk distr.,
    near Zharly vill. 10-12.05.1997 leg. M.& L. Smirnov
    male, 20 mm
    Photo from www.zin.ru
    Copyright © Maksim Smirnov
    Calosoma (Callisthenes) pavlovskii Kryzhanovskij, 1955
    Calosoma (Callisthenes) pavlovskii Kryzhanovskij, 1955
    Male adult
    Kirgizstan, Talas Mt. rng., Besh-Tash
    riv., 1000m, 15.5.2003
    male, 20 mm
    photo, det. & coll. Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Calosoma (Callisthenes) pseudocarabus Semenov et Redikorzev, 1928
    Calosoma (Callisthenes) pseudocarabus Semenov et Redikorzev, 1928
    Male adult
    NE Kazakhstan, Saur mt.rng. 2000m
    Sargar mt., 14.6.2001 Klimenko leg.
    male, 16 mm
    Copyright © Alexander Anichtchenko
    Calosoma (Callisthenes) pseudocarabus Semenov et Redikorzev, 1928
    Calosoma (Callisthenes) pseudocarabus Semenov et Redikorzev, 1928
    Female adult
    E Kazakstan, NE Saur, 2 km. W m. Tas, h=2500; 10.06.2001, Ivanov A.V. leg.,
    15 mm., female
    photo, det. & coll. Menstschikov
    Copyright © Anton Menstschikov
    Calosoma (Callisthenes) pseudocarabus Semenov et Redikorzev, 1928
    Calosoma (Callisthenes) pseudocarabus Semenov et Redikorzev, 1928
    Male adult
    E Kazakstan, NE Saur, 2 km. W m. Tas, h=2500; 10.06.2001, Ivanov A.V. leg., 14 mm., male
    photo, det. & coll. Menstschikov
    Copyright © Anton Menstschikov
    Calosoma (Callisthenes) regelianus A. Morawitz, 1886
    Calosoma (Callisthenes) regelianus A. Morawitz, 1886
    Female adult
    Tadzikistan, Darvaz Ezgan 2.5.1971
    female, 25 mm
    photo, det. & coll. Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Calosoma (Callisthenes) usgentensis usgentensis Solskyi, 1874
    Calosoma (Callisthenes) usgentensis usgentensis Solskyi, 1874
    Male adult
    Kirghizstan, near Jalal-Abad town, A. Klimenko leg., 05.04.1997; 30 mm.
    photo, det. & coll. A. Menstschicov
    Copyright © Anton Menstschikov
    Calosoma (Callisthenes) usgentensis usgentensis Solskyi, 1874
    Calosoma (Callisthenes) usgentensis usgentensis Solskyi, 1874
    Female adult
    S Kirgizia, Dzhalal-Abad reg.,
    Suzak vill., 1000m, 25.5.1999
    male, 27 mm
    photo, det. & coll. A. Anichtchenko
    Copyright © Alexander Anichtchenko
    Calosoma (Callitropa) externum Say, 1823: 149
    Calosoma (Callitropa) externum Say, 1823: 149
    Female adult
    Canada
    Copyright Henri Goulet, Agriculture & Agri-Food Canada
    Copyright © Henri Goulet
    Calosoma (Calosoma) scrutator Fabricius, 1775
    Calosoma (Calosoma) scrutator Fabricius, 1775
    Female adult
    USA, Illinois, Peoria Co.,<br /> Orange Prairie, 31.5.1990<br /> 32 mm<br /> coll. A.Montes<br /> photo Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Calosoma (Calosoma) scrutator Fabricius, 1775
    Calosoma (Calosoma) scrutator Fabricius, 1775
    Male adult
    3 m SE Tishomingo , Tishomingo Co., Mississippi, USA; 24.VI.2005<br /> Eric B. Lago leg.
    Copyright © Denis Potanin
    Calosoma (Calosoma) wilcoxi Le Conte, 1848
    Calosoma (Calosoma) wilcoxi Le Conte, 1848
    Male adult
    Canada
    Copyright Henri Goulet, Agriculture & Agri-Food Canada
    Copyright © Henri Goulet
    Calosoma (Calosoma) wilcoxi Le Conte, 1848
    Calosoma (Calosoma) wilcoxi Le Conte, 1848
    Male adult
    2 m. W Rawlings, Allegany Co., Maryland, USA; 26 May 1983<br /> J.D. Glaser leg.
    Copyright © Denis Potanin
    Calosoma (Calosoma) cyanescens Motschulsky, 1859
    Calosoma (Calosoma) cyanescens Motschulsky, 1859
    Male adult
    Primorskii reg., Kedrovaja Pad Nat.res., 11.VII.1994 Chichvarhin A. leg
    male, 19 mm
    photo, det. & coll. Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Calosoma (Calosoma) frigidum Kirby, 1837: 19
    Calosoma (Calosoma) frigidum Kirby, 1837: 19
    Male adult
    Canada
    Copyright Henri Goulet, Agriculture & Agri-Food Canada
    Copyright © Henri Goulet
    Calosoma (Calosoma) frigidum Kirby, 1837: 19
    Calosoma (Calosoma) frigidum Kirby, 1837: 19
    Male adult
    2 m W Rawlings, Allegany Co., Maryland, USA; 26.V.1983<br> J.D. Glaser leg.
    Copyright © Denis Potanin
    Calosoma (Calosoma) maximowiczi A. Morawitz, 1863: 20
    Calosoma (Calosoma) maximowiczi A. Morawitz, 1863: 20
    Female adult
    China, Mt. Wangfoshan (1940 m.), Baokang Countri, Hubei; 20-28.06.2008, 30 mm.
    Copyright © Anton Menstschikov
    Calosoma (Calosoma) oceanicum klynstrai Breuning, 1927
    Calosoma (Calosoma) oceanicum klynstrai Breuning, 1927
    Female adult
    INDONESIA, Tanimbar Island, III.2009
    photo & coll. N. Belonogov
    Copyright © Nil Belonogov
    Calosoma (Calosoma) oceanicum klynstrai Breuning, 1927
    Calosoma (Calosoma) oceanicum klynstrai Breuning, 1927
    Male adult
    INDONESIA, Tanimbar Island, III.2009
    photo & coll. N. Belonogov
    Copyright © Nil Belonogov
    Calosoma (Calosoma) sycophanta Linne, 1758
    Calosoma (Calosoma) sycophanta Linne, 1758
    Male adult
    Spain, Sierra Nevada, Pico de Veleta
    2800m
    Copyright © Alexander Anichtchenko
    Calosoma (Carabosoma) prominens LeConte, 1853
    Calosoma (Carabosoma) prominens LeConte, 1853
    Male adult
    USA Arizona, Cochise Co., Interstate 8 at Texas, Canyon Rest Area, 1500m, 32º00.3N 110º08.4W, 24.7.1996 Kavanaugh D. leg.; 30 mm
    Copyright © Alexander Anichtchenko
    Calosoma (Carabosoma) peregrinator Guerin-Meneville, 1844
    Calosoma (Carabosoma) peregrinator Guerin-Meneville, 1844
    Male adult
    Fort Davis, Jeff Davis Co., Texas, USA; July 1998
    Copyright © Denis Potanin
    Calosoma (Caminara) denticolle Gebler, 1833: 274
    Calosoma (Caminara) denticolle Gebler, 1833: 274
    Female adult
    Burjatia, Selenga riv., Deben vill. 12.7.2003 Anichtchenko A. leg. female, 25 mm
    Copyright © Alexander Anichtchenko
    Calosoma (Caminara) imbricatum deserticola Semenov, 1897d: 242
    Calosoma (Caminara) imbricatum deserticola Semenov, 1897d: 242
    Male adult
    W Kazakstan, Ustjurtskij res., Kordon Kenderli; 01.05.2009, Gorbunov P.J. leg.,
    25 mm., male
    photo, det. & coll. Menstschikov
    Copyright © Anton Menstschikov
    Calosoma (Caminara) imbricatum deserticola Semenov, 1897d: 242
    Calosoma (Caminara) imbricatum deserticola Semenov, 1897d: 242
    Male adult
    W Kazakstan, Ustjurtskij res., Kordon Kenderli; 01.05.2009, Gorbunov P.J. leg., 24 mm., female<br /> photo, det. & coll. Menstschikov
    Copyright © Anton Menstschikov
    Calosoma (Campalita) chlorostictum Dejean, 1831a: 558
    Calosoma (Campalita) chlorostictum Dejean, 1831a: 558
    Male adult
    RSA, Kwazulu/Natal, Badplaas, 1000m, 1.XII.2000<br /> male, 24 mm<br /> det. P. Schüle<br /> photo &. coll. Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Calosoma (Carabophanus) arrowi Breuning, 1928
    Calosoma (Carabophanus) arrowi Breuning, 1928
    Male adult
    Ethiopia, Oromia reg., Arsi highlands, Wahoro env., Tiliku Lensho river valley10.-12.II.2010, J.Štastny lgt
    photo Jaroslav Stastny
    Copyright © Jaroslav Štastny
    Calosoma (Castrida) alternans granulatum Perty, 1830
    Calosoma (Castrida) alternans granulatum Perty, 1830
    Male adult
    Paraguay, Dep.Caaguazu, Tayao S25.28 W56.07 12-16.III.2003 leg. U.Drechsel
    male, 28 mm
    photo, det. & coll. Anichtchenko A. 2009
    Copyright © Alexander Anichtchenko
    Calosoma (Castrida) alternans granulatum Perty, 1830
    Calosoma (Castrida) alternans granulatum Perty, 1830
    Female adult
    Argentina, Entre Rios,
    Liebig, nov.1997
    long. 32 mm
    photo, det. & coll. Anichtchenko A. 2009
    Copyright © Alexander Anichtchenko
    Calosoma (Castrida) argentinensis Csiki, 1927
    Calosoma (Castrida) argentinensis Csiki, 1927
    Male adult
    Argentina, Prov. Santa Fe Coronda, 22.12.1965 leg. Mahunka, 23 mm
    Copyright © Alexander Anichtchenko
    Calosoma (Castrida) argentinensis Csiki, 1927
    Calosoma (Castrida) argentinensis Csiki, 1927
    Male adult
    ARGENTINA: Salta; November 1995
    Copyright © Denis Potanin
    Calosoma (Castrida) vagans Dejean,
    Calosoma (Castrida) vagans Dejean,
    Male adult
    Chile, Santiago de Chile, Quilicura
    19.9.1991 G. Aceituno leg.
    male, 21 mm
    Copyright © Alexander Anichtchenko
    Calosoma (Charmosta) investigator Illiger, 1798
    Calosoma (Charmosta) investigator Illiger, 1798
    Male adult
    Lake Baikal, Buriatia, Iliynka vill., 1.8.1997 Anichtchenko A. leg.; 20 mm
    Copyright © Alexander Anichtchenko
    Calosoma (Charmosta) investigator Illiger, 1798
    Calosoma (Charmosta) investigator Illiger, 1798
    Male adult
    S. Ural, Bashkiria, near Mesjagutovo vill., 09.07.2001 A. Menstschicov leg.
    Copyright © Anton Menstschikov
    Calosoma (Charmosta) lugens Chaudoir, 1869f 372
    Calosoma (Charmosta) lugens Chaudoir, 1869f 372
    Male adult
    Wen-dong, North Fengcheng City, Liaoning, China; 1-5.VII.2006
    Copyright © Denis Potanin
    Calosoma (Chrysostigma) calidum Fabricius, 1775
    Calosoma (Chrysostigma) calidum Fabricius, 1775
    Male adult
    Canada<br /> Copyright Henri Goulet, Agriculture & Agri-Food Canada
    Copyright © Henri Goulet
    Calosoma (Chrysostigma) semilaeve LeConte, 1851: 199
    Calosoma (Chrysostigma) semilaeve LeConte, 1851: 199
    Male adult
    Orange Co., California, USA; May 1973
    Copyright ©
    Calosoma (Ctenosta) bastardi Alluaud, 1925
    Calosoma (Ctenosta) bastardi Alluaud, 1925
    Female adult

    Copyright © Alexander Anichtchenko
    Calosoma (Ctenosta) orientale Hope, 1838
    Calosoma (Ctenosta) orientale Hope, 1838
    Male adult
    China, Yunnan pro.
    Copyright © Ye Liu
    Calosoma (Ctenosta) planicolle Chaudoir, 1869
    Calosoma (Ctenosta) planicolle Chaudoir, 1869
    Male adult
    length: 30 mm. Kenia, Tsavo National Parc, leg. U.Schmidt, 1991; det. W.Lorenz, 2000. Foto: U.Schmidt, 2006
    Copyright © U.Schmidt
    Calosoma (Ctenosta) planicolle Chaudoir, 1869
    Calosoma (Ctenosta) planicolle Chaudoir, 1869
    Male adult
    SOUTH AFRICA: Northern Transvaal
    Copyright © Denis Potanin
    Calosoma (Ctenosta) senegalense Dejean, 1831
    Calosoma (Ctenosta) senegalense Dejean, 1831
    Female adult
    length: 22-27 mm
    Kenia Tsavo National Parc
    leg. U.Schmidt, 1991; det. W.Lorenz, 2000
    Foto: U.Schmidt, 2006
    Copyright © U.Schmidt
    Calosoma (Microcallisthenes) relictum Apfelbeck, 1918c
    Calosoma (Microcallisthenes) relictum Apfelbeck, 1918c
    Male adult
    Macedonia, Shar mt., Peak Tserapashina
    1850-2530m 9.7.1995
    male, 13 mm
    Copyright © Alexander Anichtchenko
    Calosoma (Microcallisthenes) relictum Apfelbeck, 1918c
    Calosoma (Microcallisthenes) relictum Apfelbeck, 1918c
    Male adult
    Macedonia, Shar mt., Peak Titov Vrah
    2747 m, 14.7.1995 V.Sakalian leg.
    male, 13 mm
    Copyright © Alexander Anichtchenko
    Calosoma (Calosoma) inquisitor inquisitor Linne, 1758: 414
    Calosoma (Calosoma) inquisitor inquisitor Linne, 1758: 414
    Male adult
    length: 13-24 mm
    England, Dendless Wood
    leg.det. M.G.Telfer, 2003
    Foto: U.Schmidt, 2009
    Copyright © U.Schmidt
    Calosoma (Calosoma) himalayanum Gestro, 1875b: 851
    Calosoma (Calosoma) himalayanum Gestro, 1875b: 851
    Female adult
    China, Tibet
    Copyright © Ye Liu
    Calosoma (Carabops) abyssinicum Gestro, 1880
    Calosoma (Carabops) abyssinicum Gestro, 1880
    Male adult
    Ethiopie, Shemene 08.2004
    19 mm
    coll. A.Montes
    Copyright © Alexander Anichtchenko
    Calosoma (Carabops) abyssinicum Gestro, 1880
    Calosoma (Carabops) abyssinicum Gestro, 1880
    Male adult
    Ethiopia, Bale prov., Str. ZW. Sasheme u. Dodola, Mt. Mascall 3100-3300 m
    21.VII.2005 Puchner leg.
    male, 18 mm
    det. Puchner F. 2005
    photo & coll. Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Calosoma (Teratexis) fabulosum Semenov & Znojko, 1933
    Calosoma (Teratexis) fabulosum Semenov & Znojko, 1933 Holotype
    Male adult
    collection of ZIN
    photo Anichtchenko A. 2010
    Copyright © Alexander Anichtchenko
    Calosoma (Teratexis) fabulosum Semenov & Znojko, 1933
    Calosoma (Teratexis) fabulosum Semenov & Znojko, 1933 Paratype
    Male adult
    collection of ZIN
    photo Anichtchenko A. 2010
    Copyright © Alexander Anichtchenko
    Calosoma (Campalita) chlorostictum Dejean, 1831a: 558
    Calosoma (Campalita) chlorostictum Dejean, 1831a: 558
    Male adult
    La Mandraka, Centre Est, Madagascar; December 1995
    Copyright © Denis Potanin
    Calosoma (Calosoma) cyanescens Motschulsky, 1859
    Calosoma (Calosoma) cyanescens Motschulsky, 1859
    Male adult
    China,
    Jilin prov, P'an-shih
    42° 56' 32" N
    126° 03' 22" E
    20.06.2009
    Local people leg.

    Copyright © Anton Menstschikov
    Calosoma (Caminara) reitteri Roeschke, 1897
    Calosoma (Caminara) reitteri Roeschke, 1897
    Male adult
    W Kazakstan, Ustjurtskij res., Kordon Kenderli; 01.05.2009, Gorbunov P.J. leg.,
    26 mm.; photo A. Menstschicov
    Copyright © Anton Menstschikov
    Calosoma (Caminara) reitteri Roeschke, 1897
    Calosoma (Caminara) reitteri Roeschke, 1897
    Male adult
    Turkmenia, Badkhyz,
    Akar-Cheshme 9.4.1985
    23 mm
    Copyright © Alexander Anichtchenko
    Calosoma (Caminara) olivieri Dejean, 1831
    Calosoma (Caminara) olivieri Dejean, 1831
    Male adult
    З Казахстан, среднее течение р. Эмба, окр. п. Миялы, П.Ю.Горбунов leg. 02.07.2009; photo A. Menstschicov
    Copyright © Anton Menstschikov
    Calosoma (Calosoma) sycophanta Linne, 1758
    Calosoma (Calosoma) sycophanta Linne, 1758
    Male adult

    Copyright © Kirill Makarov
    Calosoma (Charmosta) lugens Chaudoir, 1869
    Calosoma (Charmosta) lugens Chaudoir, 1869
    Male adult

    Copyright © Kirill Makarov
    Calosoma (Callisthenes) kuschakewitschi kuschakewitschi s.str.
    Calosoma (Callisthenes) kuschakewitschi kuschakewitschi s.str.
    Male adult

    Copyright © Kirill Makarov
    Calosoma (Campalita) auropunctatum auropunctatum Herbst, 1784: 131
    Calosoma (Campalita) auropunctatum auropunctatum Herbst, 1784: 131
    Female adult

    Copyright © Kirill Makarov
    Calosoma (Campalita) auropunctatum auropunctatum Herbst, 1784
    Calosoma (Campalita) auropunctatum auropunctatum Herbst, 1784
    Female adult
    Krym, Alupka, 12.5.1989 Kuznetsov D. leg.<br /> female, 26 mm
    Copyright © Alexander Anichtchenko
    Calosoma (Campalita) maderae maderae Fabricius, 1775
    Calosoma (Campalita) maderae maderae Fabricius, 1775
    Male adult
    Spain, Granada prov., Armilla, 15.3.2000
    29 mm
    Copyright © Alexander Anichtchenko
    Calosoma (Campalita) auropunctatum dzungaricum Gebler, 1833: 274
    Calosoma (Campalita) auropunctatum dzungaricum Gebler, 1833: 274
    Male adult
    W Kazakhstan, Emba riv., near Mijaly vill., 02.07.2009, P.Gorbunov leg.; 27 mm.
    Copyright © Anton Menstschikov
    Calosoma (Callisthenes) peksi Heinz & Pavesi, 1995
    Calosoma (Callisthenes) peksi Heinz & Pavesi, 1995
    Female adult
    Turquie (Izmir), Boz-dag( ob. Bozdag(ko'y 1500-1700 m, 2-3.V.1996, Heinz leg.
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) peksi Heinz & Pavesi, 1995
    Calosoma (Callisthenes) peksi Heinz & Pavesi, 1995
    Male adult
    Turquie (Izmir), Boz-dag( ob. Bozdag(ko'y 1500-1700 m, 2-3.V.1996, Heinz leg.
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) substriatus Motschulsky, 1859
    Calosoma (Callisthenes) substriatus Motschulsky, 1859
    Female adult
    Azerbaijan, Talysh, Gasmalian, 20.V.1992, V. Siniaev leg.
    photo from zin.ru
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) substriatus Motschulsky, 1859
    Calosoma (Callisthenes) substriatus Motschulsky, 1859 Lectotype
    Male adult
    "Callisthenes substriatus Motsch. Mt. Saganlug Persia"
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) substriatus Motschulsky, 1859
    Calosoma (Callisthenes) substriatus Motschulsky, 1859
    Male adult
    Azerbaijan, Talysh, Gasmalian, 20.V.1992, V. Siniaev leg.
    photo from zin.ru
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) anthrax Semenov, 1900
    Calosoma (Callisthenes) anthrax Semenov, 1900
    Female adult
    Inner Mongolia (China), Dalai-Nor Lake, VI.1957, Svetlovidova leg.
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) karelini breuningi Mandl, 1954
    Calosoma (Callisthenes) karelini breuningi Mandl, 1954 Holotype
    Male adult
    "Sergiopol", "Cal. Breuningi, Typus, m., det. Ing. K. Mandl"
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) karelini vladimiri Obydov, 1998
    Calosoma (Callisthenes) karelini vladimiri Obydov, 1998
    Male adult
    Kazakhstan, Taldy-Kurgan env., Kyzyl-Zhar Mts., 400 m, 15.IV.2000, M. Danilevsky leg.
    photo from zin.ru
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) pavlovskii Kryzhanovskij, 1955
    Calosoma (Callisthenes) pavlovskii Kryzhanovskij, 1955 Holotype
    Male adult
    "Talas, IV.08, E. Fischer", "Callisthenes pavlovskii sp. nov., Holotype, O. Kryzhanovskij det.".
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) elegans elegans s.str.
    Calosoma (Callisthenes) elegans elegans s.str.
    Male adult
    SE Kazakhstan, Chu-Ili mts., Kurdai pass, h =1800m, 20.V.1999
    24 mm
    Copyright © Maksim Smirnov
    Calosoma (Callisthenes) elegans danilevskii Obydov, 1997d
    Calosoma (Callisthenes) elegans danilevskii Obydov, 1997d Paratype
    Male adult
    Kazakhstan, Kaskelen Riv. Val., 1500 m, 9.VI.1989, M. Danilevsky leg.<br /> from zin.ru
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) elegans danilevskii Obydov, 1997d
    Calosoma (Callisthenes) elegans danilevskii Obydov, 1997d Paratype
    Female adult
    Kazakhstan, Kaskelen Riv. Val., 1500 m, 9.VI.1989, M. Danilevsky leg.
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) elegans semenovi Motschulsky, 1859
    Calosoma (Callisthenes) elegans semenovi Motschulsky, 1859 Lectotype
    Male adult
    "elegans Kirsch. Stett. Zeit., 1859, 107. Semenof", "Neotypus, C. semenovi Motsch., Kryzhanovskij, 1966".
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) elegans ballioni Solsky, 1874
    Calosoma (Callisthenes) elegans ballioni Solsky, 1874
    Male adult
    Kazakhstan, Karaoi, 550 m, 7.5.1991, 24.IV.1994, M. Danilevsky leg.
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) elegans ballioni Solsky, 1874
    Calosoma (Callisthenes) elegans ballioni Solsky, 1874
    Female adult
    Kazakhstan, Karaoi, 550 m, 24.IV.1994, M. Danilevsky leg.
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) elegans valentinae Obydov, 1997
    Calosoma (Callisthenes) elegans valentinae Obydov, 1997 Paratype
    Male adult
    Kazakhstan, Kapchagai, left bank of Ili Riv., 25.IV.1994, M. Danilevsky leg.
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) elegans valentinae Obydov, 1997
    Calosoma (Callisthenes) elegans valentinae Obydov, 1997 Paratype
    Female adult
    Kazakhstan, Kapchagai, left bank of Ili Riv., 25. IV.1994, M. Danilevsky leg.
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) elegans saryakensis Kabak, 1992
    Calosoma (Callisthenes) elegans saryakensis Kabak, 1992 Holotype
    Male adult
    Koksengir mt. environs, Karaganda dist., 21.V.1958, Tobias leg.
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) elegans amethystinus Semenov & Redikorzev, 1928
    Calosoma (Callisthenes) elegans amethystinus Semenov & Redikorzev, 1928 Holotype
    Male adult
    On the road to the Charyn River, steppe, 8.VII.09, A. Malchevsky leg.
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) elegans purpureus Obydov, 2001
    Calosoma (Callisthenes) elegans purpureus Obydov, 2001 Holotype
    Male adult
    Kazakhstan, Mezhdurechensk, 650 m, 12.IV.2000, M. Danilevsky leg.
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) elegans keminensis Obydov, 2001
    Calosoma (Callisthenes) elegans keminensis Obydov, 2001 Holotype
    Male adult
    Kirghizia, Kemin (Bystrovka) env., 1300 m, Kyzyl-Oktyabr, 16.V.2000, M. Danilevsky leg.
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) elegans kolshengelicus Obydov, 1997
    Calosoma (Callisthenes) elegans kolshengelicus Obydov, 1997 Paratype
    Female adult
    Kazakhstan, Kolshengel, 450m, 22.IV.1994, M. Danilevsky leg.
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) elegans kolshengelicus Obydov, 1997
    Calosoma (Callisthenes) elegans kolshengelicus Obydov, 1997 Paratype
    Male adult
    Kazakhstan, Kolshengel, 450m, 22.IV.1994, M. Danilevsky leg.
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) subtilestriatus Mandl, 1954
    Calosoma (Callisthenes) subtilestriatus Mandl, 1954 Lectotype
    Male adult
    "Kuldscha"
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) akkolicus Obydov et Gottwald, 2002
    Calosoma (Callisthenes) akkolicus Obydov et Gottwald, 2002 Holotype
    Male adult
    Kazakhstan, Karatau, Akkol env., 9-10.V.1978, male, J. Gottwald leg.
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) akkolicus Obydov et Gottwald, 2002
    Calosoma (Callisthenes) akkolicus Obydov et Gottwald, 2002 Paratype
    Female adult
    Kazakhstan, Karatau, Akkol env., 9-10.V.1978, female, J. Gottwald leg.
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) akkolicus korelli Obydov, 2004
    Calosoma (Callisthenes) akkolicus korelli Obydov, 2004 Paratype
    Female adult
    Kirghizia, Kyzyl-Sai Village env., V.1992, M. Pejcha leg.
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) glasunovi Semenov, 1900
    Calosoma (Callisthenes) glasunovi Semenov, 1900 Lectotype
    Male adult
    "Trkst. Mnt. Nurata, Sintab, Glasunov 1892", "Calosoma glasunovi m. Typ., XII.99, A. Semenov det."
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) glasunovi Semenov, 1900
    Calosoma (Callisthenes) glasunovi Semenov, 1900
    Female adult
    Uzbekistan, Nuratau Mt. Ridge, 1300 m, 6 km from Farish Village to the south, 6-7.V.2002, male, D. Obydov leg.
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) glasunovi Semenov, 1900
    Calosoma (Callisthenes) glasunovi Semenov, 1900
    Female adult
    Uzbekistan, Nuratau Mt. Ridge, 1300 m, 6 km from Farish Village to the south, 6-7.V.2002, female, D. Obydov leg.
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) klapperichi Mandl, 1955
    Calosoma (Callisthenes) klapperichi Mandl, 1955
    Male adult
    Afganistan, Bad. Shiva, 14.VII.1954. Topotypus
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) pumicatus pumicatus s.str.
    Calosoma (Callisthenes) pumicatus pumicatus s.str. Holotype
    Female adult
    "Plateau Persan Occid. de Zendjan a Ardebil, alt. moy 300 m (J. de Morgan)".
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) pumicatus pumicatus (as zarudnianum)
    Calosoma (Callisthenes) pumicatus pumicatus (as zarudnianum) Holotype
    Male adult
    Holotypus of Calosoma zarudnianum:
    "Calosoma zarudnianum m, sp. n., A. Semenov-Tian-Shansky det. 21", "W. Pers., 13.V.04, Zarudnyi leg.".
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) pumicatus schelkovnikovi Zaitzev, 1918
    Calosoma (Callisthenes) pumicatus schelkovnikovi Zaitzev, 1918 Lectotype
    Male adult
    "ms. Sehend 11. 600' 18.V.16", "Mus Caucas. 100-16. Exp. Urmiana", "Calosoma schelkovnikovi Zaitz. (type)", "Received personally from F.A. Zaitzev in 1920".
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) kuschakewitschi batesoni Semenov & Redikorzev, 1928
    Calosoma (Callisthenes) kuschakewitschi batesoni Semenov & Redikorzev, 1928 Paratype
    Male adult
    "Bateson 1887 Inter Kazalinsk et Karkaralinsk (Desert. Kirgis.)"
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) kuschakewitschi solskyanus Obydov, 1999
    Calosoma (Callisthenes) kuschakewitschi solskyanus Obydov, 1999 Holotype
    Male adult
    South Kazakhstan, Ulanbel env., 19.4.1977, A. Shatalkin leg.
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) kuschakewitschi solskyanus Obydov, 1999
    Calosoma (Callisthenes) kuschakewitschi solskyanus Obydov, 1999
    Female adult
    Kazakhstan, 80 km from Merke to the north, 6.V.1994, J. Kadlec leg.
    Copyright © Dmitri Obydov
    Calosoma (Callisthenes) kuschakewitschi sidzhakensis Obydov, 2008
    Calosoma (Callisthenes) kuschakewitschi sidzhakensis Obydov, 2008 Holotype
    Male adult
    Uzbekistan-Kazakhstan border, Ugamsky Mt. Ridge., 2100 m, 15 km N-W Sidzhak Village, 1.VI.1978.
    Copyright © Dmitri Obydov
    Calosoma (Callistenia) moniliatus Le Conte, 1851: 200
    Calosoma (Callistenia) moniliatus Le Conte, 1851: 200
    Male adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Chrysostigma) affinis Chaudoir, 1843
    Calosoma (Chrysostigma) affinis Chaudoir, 1843
    Male adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Chrysostigma) calidus Fabricius, 1775
    Calosoma (Chrysostigma) calidus Fabricius, 1775
    Female adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Chrysostigma) lepidus Le Conte, 1844: 201
    Calosoma (Chrysostigma) lepidus Le Conte, 1844: 201
    Female adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Blaptosoma) viridisulcatum viridisulcatum Chaudoir, 1863
    Calosoma (Blaptosoma) viridisulcatum viridisulcatum Chaudoir, 1863
    Male adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Blaptosoma) anthracinum microgonum Bates, 1891
    Calosoma (Blaptosoma) anthracinum microgonum Bates, 1891
    Male adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Blaptosoma) laeve Dejean, 1826
    Calosoma (Blaptosoma) laeve Dejean, 1826
    Female adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH;
    Copyright © - - -
    Calosoma (Blaptosoma) haydeni punctulicolle Bates, 1891
    Calosoma (Blaptosoma) haydeni punctulicolle Bates, 1891
    Female adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Blaptosoma) haydeni haydeni s.str.
    Calosoma (Blaptosoma) haydeni haydeni s.str.
    Male adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Blaptosoma) cf atrovirens obscurum Gehin, 1885
    Calosoma (Blaptosoma) cf atrovirens obscurum Gehin, 1885
    Male adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Callitropa) externum Say, 1823: 149
    Calosoma (Callitropa) externum Say, 1823: 149
    Male adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Callitropa) macrum LeConte, 1853
    Calosoma (Callitropa) macrum LeConte, 1853
    Female adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Chrysostigma) tepidum LeConte, 1851: 199
    Calosoma (Chrysostigma) tepidum LeConte, 1851: 199
    Male adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Chrysostigma) simplex LeConte, 1878
    Calosoma (Chrysostigma) simplex LeConte, 1878
    Female adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Chrysostigma) semilaeve LeConte, 1851: 199
    Calosoma (Chrysostigma) semilaeve LeConte, 1851: 199
    Male adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Chrysostigma) obsoletum Say, 1823: 149
    Calosoma (Chrysostigma) obsoletum Say, 1823: 149
    Male adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Chrysostigma) morrisoni Horn, 1885
    Calosoma (Chrysostigma) morrisoni Horn, 1885
    Female adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Callitropa) palmeri Horn, 1876: 199
    Calosoma (Callitropa) palmeri Horn, 1876: 199
    Male adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Callitropa) protractum LeConte, 1862: 52
    Calosoma (Callitropa) protractum LeConte, 1862: 52
    Male adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Calosoma) splendidum Dejean, 1831
    Calosoma (Calosoma) splendidum Dejean, 1831
    Female adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Calosoma) aurocinctum Chaudoir, 1850
    Calosoma (Calosoma) aurocinctum Chaudoir, 1850
    Male adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Carabomimus) asperum (Jeannel, 1940)
    Calosoma (Carabomimus) asperum (Jeannel, 1940)
    Male adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Carabomimus) costipenne Chaudoir, 1869a: 375
    Calosoma (Carabomimus) costipenne Chaudoir, 1869a: 375
    Female adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Carabomimus) diminutum Bates, 1891: 227
    Calosoma (Carabomimus) diminutum Bates, 1891: 227
    Female adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Carabomimus) morelianum Bates, 1891
    Calosoma (Carabomimus) morelianum Bates, 1891
    Female adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH. det. Anichtchenko, 2020
    Copyright © - - -
    Calosoma (Carabomimus) politum Chaudoir, 1869a: 373
    Calosoma (Carabomimus) politum Chaudoir, 1869a: 373
    Male adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Carabomimus) striatulum Chevrolat, 1835: 165
    Calosoma (Carabomimus) striatulum Chevrolat, 1835: 165
    Male adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Carabosoma) angulatum Chevrolat, 1834
    Calosoma (Carabosoma) angulatum Chevrolat, 1834
    Female adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Carabosoma) eremicola Fall, 1910: 91
    Calosoma (Carabosoma) eremicola Fall, 1910: 91
    Female adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Carabosoma) glabratum Dejean, 1831
    Calosoma (Carabosoma) glabratum Dejean, 1831
    Female adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Carabosoma) marginalis Casey, 1897
    Calosoma (Carabosoma) marginalis Casey, 1897
    Female adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Carabosoma) parvicollis Fall, 1910: 90
    Calosoma (Carabosoma) parvicollis Fall, 1910: 90
    Female adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Carabosoma) peregrinator Guerin-Meneville, 1844
    Calosoma (Carabosoma) peregrinator Guerin-Meneville, 1844
    Male adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Carabosoma) prominens LeConte, 1853: 400
    Calosoma (Carabosoma) prominens LeConte, 1853: 400
    Male adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Carabosoma) sponsum Casey, 1897: 340
    Calosoma (Carabosoma) sponsum Casey, 1897: 340
    Male adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Castrida) abbreviatum Chaudoir, 1869
    Calosoma (Castrida) abbreviatum Chaudoir, 1869
    Female adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Castrida) argentinensis Csiki, 1927
    Calosoma (Castrida) argentinensis Csiki, 1927
    Female adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Castrida) fulgens Chaudoir, 1869
    Calosoma (Castrida) fulgens Chaudoir, 1869
    Male adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Castrida) granatense granatense Gehin, 1885
    Calosoma (Castrida) granatense granatense Gehin, 1885
    Male adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Castrida) retusum Fabricius, 1775: 237
    Calosoma (Castrida) retusum Fabricius, 1775: 237
    Male adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Castrida) sayi Dejean, 1826
    Calosoma (Castrida) sayi Dejean, 1826
    Male adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Charmosta) investigator Illiger, 1798
    Calosoma (Charmosta) investigator Illiger, 1798
    Habitat
    The Chelyabinsk area, Kizilskyi township, Cheka mountain (steppe). 14-15/07/2008
    Copyright © Evgeniy Chibilov
    Calosoma (Charmosta) investigator Illiger, 1798
    Calosoma (Charmosta) investigator Illiger, 1798
    Habitat
    The Chelyabinsk area, Kizilskyi township, Cheka mountain (steppe). 14-15/07/2008
    Copyright © Evgeniy Chibilov
    Calosoma (Castrida) argentinensis Csiki, 1927
    Calosoma (Castrida) argentinensis Csiki, 1927
    Habitat
    Paraguay, Dep. Boqueron, Mariscal Estigarribia
    S22º01,5´ W60º37.1´ 02.02.2008
    photo: U. Drechsel
    Copyright © Ulf Drechsel
    Calosoma (Castrida) alternans Fabricius, 1792
    Calosoma (Castrida) alternans Fabricius, 1792
    Habitat
    Paraguay, Dep. Alto Parana, Estancia Dimas
    S25º32,7´ W55º13,3´ 23.01.2007
    photo: U. Drechsel
    Copyright © Ulf Drechsel
    Calosoma (Campalita) auropunctatum dzungaricum Gebler, 1833: 274
    Calosoma (Campalita) auropunctatum dzungaricum Gebler, 1833: 274
    Habitat

    Copyright © Sunvas
    Calosoma (Calosoma) sycophanta Linne, 1758
    Calosoma (Calosoma) sycophanta Linne, 1758
    Habitat
    South Ural, Chelyabinsk area, Chebarkul a pine forest.
    Photo Chibilov E.A.
    Copyright © Sunvas
    Calosoma (Blaptosoma) viridisulcatum Chaudoir, 1863: 114
    Calosoma (Blaptosoma) viridisulcatum Chaudoir, 1863: 114
    Habitat
    Mexico. Guadelajaravic en la costa Norte del Lago Chapala 28 Julio 2005
    photo Eric Vandenberghe
    Copyright © ---
    Calosoma (Calosoma) inquisitor Linne, 1758: 414
    Calosoma (Calosoma) inquisitor Linne, 1758: 414
    Habitat
    Chelyabinsk area, Kizilskyi township, Cheka mountain (steppe). 14-15/07/2008
    photo E. Chibilov
    Copyright © Sunvas
    Calosoma (Charmosta) investigator Illiger, 1798
    Calosoma (Charmosta) investigator Illiger, 1798
    Female adult

    Copyright © Kirill Makarov
    Calosoma (Calosoma) inquisitor inquisitor Linne, 1758: 414
    Calosoma (Calosoma) inquisitor inquisitor Linne, 1758: 414
    Male adult

    Copyright © Kirill Makarov
    Calosoma (Callisthenes) panderi Fischer, 1822
    Calosoma (Callisthenes) panderi Fischer, 1822
    Male adult
    NW Kazakhstan, Aktiubinsk reg., Baiganinsk distr., near Zharly vill. 10-12.05.1997 leg. M.& L. Smirnov (MPU)
    Copyright © Kirill Makarov
    Calosoma (Carabomorphus) brachycerum (Gersteacker, 1884)
    Calosoma (Carabomorphus) brachycerum (Gersteacker, 1884)
    Male adult
    N Tanzania, 105 km W of Arusha and 5 km S of Kwa Kuchinga vill. 3°41'S 25°59' E 31.VIII-2.IX.2006 leg. A.Sokolov
    L=23 mm
    Copyright © Kirill Makarov
    Calosoma (Callisthenes) breviusculus Mannerheim, 1830
    Calosoma (Callisthenes) breviusculus Mannerheim, 1830
    Male adult
    Turkey, Buglan; 21mm
    Copyright © Aleš Sedláček
    Calosoma (Microcallisthenes) relictum Apfelbeck, 1918
    Calosoma (Microcallisthenes) relictum Apfelbeck, 1918
    Male adult
    Macedonia, Bistra planina Mts.; 13mm
    Copyright © Aleš Sedláček
    Calosoma (Microcallisthenes) relictum Apfelbeck, 1918
    Calosoma (Microcallisthenes) relictum Apfelbeck, 1918
    Female adult
    Macedonia, Bistra planina Mts.; 13mm
    Copyright © Aleš Sedláček
    Calosoma (Microcallisthenes) relictum Apfelbeck, 1918
    Calosoma (Microcallisthenes) relictum Apfelbeck, 1918
    Male adult
    Macedonia, Bistra planina Mts.; 14mm
    Copyright © Aleš Sedláček
    Calosoma (Calosoma) sycophanta Linne, 1758
    Calosoma (Calosoma) sycophanta Linne, 1758
    Male adult
    CZ; S Moravia, Břeclav; 26mm
    Copyright © Aleš Sedláček
    Calosoma (Ctenosta) guineense Imhoff, 1843
    Calosoma (Ctenosta) guineense Imhoff, 1843
    Male adult
    Rep. Centre Africaine: Bangui env. IV.1974 28 mm
    Copyright © Ruslan Panin
    Calosoma (Calosoma) frigidum Kirby, 1837: 19
    Calosoma (Calosoma) frigidum Kirby, 1837: 19
    Female adult
    SC-Canada, Manitoba prov., Winnipeg, King’s Park, 23.VI.2008, Todd Lawton leg. L=24 mm
    Copyright © Ruslan Panin
    Calosoma (Campalita) chinense Kirby, 1819: 379
    Calosoma (Campalita) chinense Kirby, 1819: 379
    Female adult
    North Korea, Ryanggang Province, Mt P'urun-bong, 24.VIII.1994 L=33 mm
    Copyright © Ruslan Panin
    Calosoma (Castrida) alternans granulatum Perty, 1830
    Calosoma (Castrida) alternans granulatum Perty, 1830
    Male adult
    Brasil
    Copyright © Kirill Makarov
    Calosoma (Callisthenes) breviusculus Mannerheim, 1830
    Calosoma (Callisthenes) breviusculus Mannerheim, 1830
    Female adult
    ARMENIE, Aragatsotn route du lac Kari, 28-06-2014, Pascal Deschamps coll.
    Copyright © Pascal Deschamps
    Calosoma (Carabops) janssensi Basilewsky, 1953
    Calosoma (Carabops) janssensi Basilewsky, 1953
    Female adult
    01/12/1948, Congo, The Democratic Republic of the Upemba National Park, Lusinga. Image from: Mal, N., Drumont, A., Kerkhof, S. & P. Grootaert, 2012. Pictorial beetle collection of the Royal Belgian Institute of Natural Sciences.
    Copyright © - - -
    Calosoma (Orinodromus) deckeni (Gersteacker, 1867)
    Calosoma (Orinodromus) deckeni (Gersteacker, 1867)
    Male adult
    04/1912, Tanzania, Mount Kilimanjaro, Bismarck Hügel, 2700m. Image from: Mal, N., Drumont, A., Kerkhof, S. & P. Grootaert, 2012. Pictorial beetle collection of the Royal Belgian Institute of Natural Sciences.
    Copyright © - - -
    Calosoma (Calosoma) sycophanta Linne, 1758
    Calosoma (Calosoma) sycophanta Linne, 1758
    Male adult
    Turkey : Antalya, Yukarikaraman 3.iv.2000 - M. Grotz Leg. (Size : 33 mm) Coll. N. Mal
    Copyright © Noël Mal
    Calosoma (Calosoma) sycophanta Linne, 1758
    Calosoma (Calosoma) sycophanta Linne, 1758
    Female adult
    Turkey : Antalya, Yukarikaraman 3.iv.2000 - M. Grotz Leg. (Size : 33 mm) Coll. N. Mal
    Copyright © Noël Mal
    Calosoma (Ctenosta) grandidieri Maindron, 1900
    Calosoma (Ctenosta) grandidieri Maindron, 1900
    Female adult
    MADAGASCAR, Tulear Prov., nr. Tsimanampetsotsa NP, P. Schüle det.
    Copyright © Peter Schüle
    Calosoma (Ctenosta) senegalense Dejean, 1831
    Calosoma (Ctenosta) senegalense Dejean, 1831
    Female adult
    MADAGASCAR, Tulear Prov., nr. Tsimanampetsotsa NP, P. Schüle det.
    Copyright © Peter Schüle
    Calosoma (Castrida) galapageium Hope, 1838
    Calosoma (Castrida) galapageium Hope, 1838
    Male adult
    Contributors: Picture: Camille Locatelli, Copyright: RBINS / DIGIT-3 Belspo Licence: CC BY NC ND http://virtualcollections.naturalsciences.be
    Copyright © - - -
    Calosoma (Castrida) granatense floreanum (Basilewsky, 1968)
    Calosoma (Castrida) granatense floreanum (Basilewsky, 1968) Holotype
    Male adult
    Picture: Camille Locatelli, Copyright: RBINS / DIGIT-3 Belspo Licence: CC BY NC ND http://virtualcollections.naturalsciences.be
    Copyright © - - -
    Calosoma (Castrida) granatense floreana (Basilewsky, 1968)
    Calosoma (Castrida) granatense floreana (Basilewsky, 1968)
    Label
    Picture: Camille Locatelli, Copyright: RBINS / DIGIT-3 Belspo Licence: CC BY NC ND http://virtualcollections.naturalsciences.be
    Copyright © - - -
    Calosoma (Castrida) leleuporum (Basilewsky, 1968)
    Calosoma (Castrida) leleuporum (Basilewsky, 1968) Holotype
    Male adult
    Picture: Camille Locatelli, Copyright: RBINS / DIGIT-3 Belspo Licence: CC BY NC ND http://virtualcollections.naturalsciences.be
    Copyright © - - -
    Calosoma (Castrida) leleuporum (Basilewsky, 1968)
    Calosoma (Castrida) leleuporum (Basilewsky, 1968)
    Label
    Picture: Camille Locatelli, Copyright: RBINS / DIGIT-3 Belspo Licence: CC BY NC ND http://virtualcollections.naturalsciences.be
    Copyright © - - -
    Calosoma (Castrida) linelli Mutchler, 1925
    Calosoma (Castrida) linelli Mutchler, 1925
    Male adult
    Picture: Camille Locatelli, Copyright: RBINS / DIGIT-3 Belspo Licence: CC BY NC ND http://virtualcollections.naturalsciences.be
    Copyright © - - -
    Calosoma (Ctenosta) scabrosum scabrosum
    Calosoma (Ctenosta) scabrosum scabrosum Holotype
    Male adult
    (MNHN)
    Copyright © Boleslav Brezina
    Calosoma (Ctenosta) scabrosum roeschkei Breuning, 1927
    Calosoma (Ctenosta) scabrosum roeschkei Breuning, 1927 Holotype
    Male adult
    “Usambara”(NBCL)
    Copyright © Boleslav Brezina
    Ctenosta aethiops Jeannel, 1940
    Ctenosta aethiops Jeannel, 1940 Holotype
    Male adult
    MNHN. Scale bar 10 mm.
    Copyright © Boleslav Brezina
    Calosoma (Caminara) imbricatum hottentottum Chaudoir, 1852
    Calosoma (Caminara) imbricatum hottentottum Chaudoir, 1852 Holotype
    Male adult
    MNHN. Scale bar 10 mm.
    Copyright © Boleslav Brezina
    Calosoma (Ctenosta) scabrosum scabrosum
    Calosoma (Ctenosta) scabrosum scabrosum
    Male adult
    Djibouti: Obock (NMP)
    Copyright © Boleslav Brezina
    Calosoma (Ctenosta) jakli Häckel, Farkač & Sehnal, 2005
    Calosoma (Ctenosta) jakli Häckel, Farkač & Sehnal, 2005 Holotype
    Male adult
    SW Asia, S Oman, Dzhopar Prov., Al Mughsayi vill. env., 0–50 m a.s.l., VIII.1999, lgt. S. Jákl / HOLOTYPE Calosoma jakli det. Häckel, Farkač & Sehnal, 2005 / Calsoma scabrosum det. Häckel, Farkač & Sehnal, 2010 (HAC).
    Copyright © Boleslav Brezina
    Calosoma (Ctenosta) scabrosum roeschkei Breuning, 1927
    Calosoma (Ctenosta) scabrosum roeschkei Breuning, 1927
    Male adult
    Kenya, Voi
    Copyright © Boleslav Brezina
    Calosoma (Caminara) imbricatum hottentottum Chaudoir, 1852
    Calosoma (Caminara) imbricatum hottentottum Chaudoir, 1852
    Male adult
    Namibia: Okahandja. Scale bar 10 mm.
    Copyright © Boleslav Brezina
    Calosoma (Caminara) denticolle Gebler, 1833: 274
    Calosoma (Caminara) denticolle Gebler, 1833: 274
    Female adult

    Copyright © Kirill Makarov
    Calosoma (Caminara) denticolle Gebler, 1833: 274
    Calosoma (Caminara) denticolle Gebler, 1833: 274
    Male adult

    Copyright © Kirill Makarov
    Calosoma (Elgonorites) elgonense Burgeon, 1928
    Calosoma (Elgonorites) elgonense Burgeon, 1928
    Male adult
    Kenya, M. Elgon, vers. S, 3600 m., 6.IX.1986. (Coll. R. Sciaky)
    Copyright © Riccardo Sciaky
    Calosoma (Teratexis) fabulosum Semenov & Znojko, 1933
    Calosoma (Teratexis) fabulosum Semenov & Znojko, 1933
    Male+Female adult
    Central Afghanistan, Bamian, Bande-Amir, 3000 m (1 m, 1 f)
    Copyright © Igor Solodovnikov
    Calosoma (Teratexis) fabulosum Semenov & Znojko, 1933
    Calosoma (Teratexis) fabulosum Semenov & Znojko, 1933
    Male+Female adult
    Central Afghanistan, Bamian, Bande-Amir, 3000 m (1 m, 1 f)
    Copyright © Igor Solodovnikov
    Calosoma (Castrida) granatense granatense Gehin, 1885
    Calosoma (Castrida) granatense granatense Gehin, 1885
    Male adult
    Ecuador, 28.9.85, Galapagos, Isabela Sud, Sierra Negra Sud, 490m Pierre Moret leg. & det.
    Copyright © Alexander Anichtchenko
    Calosoma (Carabomimus) politum Chaudoir, 1869
    Calosoma (Carabomimus) politum Chaudoir, 1869
    Male adult
    Mexico, MIC ME line, N Angangueo, 19*39'686 N 100*15'877 W, 3100 m, 28 jun 2010, J.A. Shetterly et al. leg. 15 mm; det. Anichtchenko A., 2020
    Copyright © Alexander Anichtchenko
    Calosoma (Blaptosoma) atrovirens Chaudoir, 1869
    Calosoma (Blaptosoma) atrovirens Chaudoir, 1869
    Male adult
    Mexico, SLP, Hwy 80, 3k. E Ciudad del Maiz, 20.VII.1991, leg. Shetterly et al; 22 mm; det. Anichtchenko A. 2020
    Copyright © Alexander Anichtchenko
    Calosoma (Blaptosoma) porosifrons Bates, 1891
    Calosoma (Blaptosoma) porosifrons Bates, 1891
    Female adult
    Mexico, Durango, Otinapa road 23*59.2N 104*58.5W, 6.7.2010, leg. J.A. Shetterly et al; det. Anichtchenko A. 2020
    Copyright © Alexander Anichtchenko
    Calosoma (Blaptosoma) viridisulcatum Chaudoir, 1863
    Calosoma (Blaptosoma) viridisulcatum Chaudoir, 1863
    Male adult
    Mexico, Jal., N Guadalajara, Hwy 80, 20.18.744N 103.54.015W, 1664m, km 49, 30.VI.2010, leg. J.A. Shetterly et al; det. Anichtchenko A. 2020
    Copyright © Alexander Anichtchenko
    Calosoma (Blaptosoma) laeve Dejean, 1826
    Calosoma (Blaptosoma) laeve Dejean, 1826
    Male adult
    Mexico, Mich, 11 k E Zacapu, Hwy 15, 19*45.840 N 101*43.733 W, 2021 m, 28.6.2010, leg. J.A. Shetterly et al; 26 mm; det. Anichtchenko A. 2020
    Copyright © Alexander Anichtchenko
    Calosoma (Calosoma) sycophanta Linne, 1758
    Calosoma (Calosoma) sycophanta Linne, 1758
    Male adult
    Italy: Abruzzo, Pallano, h = 1000m, 18.06.1993, leg. Facovone
    Copyright © Igor Solodovnikov
    Calosoma (Calosoma) sycophanta Linne, 1758
    Calosoma (Calosoma) sycophanta Linne, 1758
    Female adult
    Calosoma (s.str.) sycophanta severum f.i. cupreum Letzner, 1850 Turkey, Pozanti, 30.IV.2009 , leg. O.Boilly
    Copyright © Igor Solodovnikov
    Calosoma (Calosoma) scrutator Fabricius, 1775
    Calosoma (Calosoma) scrutator Fabricius, 1775
    Male adult
    USA: Texas, Live Oak County 5 miles South of George West, 24.09.2010, local collector
    Copyright © Igor Solodovnikov
    Calosoma (Calosoma) sycophanta Linne, 1758
    Calosoma (Calosoma) sycophanta Linne, 1758
    Male adult
    Italy: Abruzzo, Pallano, h = 1000m, 18.06.1993, leg. Facovone
    Copyright © Igor Solodovnikov
    Calosoma (Calosoma) aurocinctum Chaudoir, 1850
    Calosoma (Calosoma) aurocinctum Chaudoir, 1850
    Female adult
    MEXICO, Sian Ka’an, Quintana Roo, 19.VI.2015, leg. Julio Medin
    Copyright © Igor Solodovnikov
    Calosoma (Carabosoma) angulatum Chevrolat, 1834
    Calosoma (Carabosoma) angulatum Chevrolat, 1834
    Male adult
    Reserva silvestra de Domitila, Pl., Granade prov., Nicaragua, 1-5.6.2005. det. Anichtchenko A. 2020
    Copyright © Alexander Anichtchenko
    Calosoma (Ctenosta) planicolle Chaudoir, 1869
    Calosoma (Ctenosta) planicolle Chaudoir, 1869
    Male adult
    C Tanzania, Chikuyu, Dodoma, 1.2015 (DUBC)
    Copyright © Alexander Anichtchenko
    Calosoma (Carabomimus) morelianum Bates, 1891
    Calosoma (Carabomimus) morelianum Bates, 1891
    Female adult
    Mexico, Santa Martha de Ocuilan, 2760m, 19*04'09"N 99*23'17"W. 19.6.1995, J. Shetterly et al leg. ; det. Anichtchenko A., 2020; 14mm
    Copyright © Alexander Anichtchenko
    Calosoma (Carabomimus) politum diminutum Bates, 1891
    Calosoma (Carabomimus) politum diminutum Bates, 1891
    Male adult
    Mexico, MX, Salazar, Alt. 3000 m, 19*18.194N, 99*23.566W, 24.6.2000, leg. J.A. Shetterly et al.; 15 mm; det. Anichtchenko A. 2020
    Copyright © Alexander Anichtchenko
    Calosoma (Carabomimus) striatulum s.str.
    Calosoma (Carabomimus) striatulum s.str.
    Male adult
    Mexico, VC, Hwy 140, NE Perote, km 140, 19*37'50"N 97*10'17"W, 2450 m, 21.6.2000, J. Shetterly et al leg.; 15 mm; det. Anichtchenko A. 2020
    Copyright © Alexander Anichtchenko
    Calosoma (Carabomimus) depressicolle Chaudoir, 1869
    Calosoma (Carabomimus) depressicolle Chaudoir, 1869
    Male adult
    det. Anichtchenko A. 2020
    Copyright © Alexander Anichtchenko
    Calosoma (Blaptosoma) haydeni punctulicolle Bates, 1891
    Calosoma (Blaptosoma) haydeni punctulicolle Bates, 1891
    Female adult
    Mexico, Durango, Hwy #40, km #35, 23*56.3N 104*51.9W, 15-18.7.2001, J.A. Shetterly et al leg.; 25 mm. det. Anichtchenko A. 2020 (atypical form, lateral margin of elytra not serrate near humera)
    Copyright © Alexander Anichtchenko
    Calosoma (Blaptosoma) haydeni punctulicolle Bates, 1891
    Calosoma (Blaptosoma) haydeni punctulicolle Bates, 1891
    Male adult
    Mexico, Chihuahua, S Hy 24, San Francisco Mesa, nr. mine 2300 m, 24*49.671N 105*50.956W, 25.7.2005, leg. J.A. Shetterly (cAA). det. Anichtchenko A. 2020
    Copyright © Alexander Anichtchenko
    Calosoma (Campalita) algiricum Gehin, 1885
    Calosoma (Campalita) algiricum Gehin, 1885
    Female adult
    Maroc, col du Kerdous, 28-04-1996
    Copyright © Pascal Deschamps
    Calosoma (Callisthenes) elegans danilevskii Obydov, 1997d
    Calosoma (Callisthenes) elegans danilevskii Obydov, 1997d
    Female adult
    SE-Kazakhstan, Almaty reg., C Trans-Ili Alatau N-slopes, Kaskalen river Valley, Izvestkovy vill., h=1700 m, 6.V.2015, D.Kovalchuk leg./det R.Panin, L=32 mm
    Copyright © Ruslan Panin
    Calosoma (Calosoma) scrutator Fabricius, 1775
    Calosoma (Calosoma) scrutator Fabricius, 1775
    Male adult

    Copyright © Christensen_Tim
    Calosoma (Ctenosta) bastardi Alluaud, 1925
    Calosoma (Ctenosta) bastardi Alluaud, 1925
    Female adult
    S-Madagaskar, Tulear prov., near Betioky town, h = 150 m, 23°38'11" S, 44°20'9" E, 27.XII.2019, leg. A. Kozlov & Yu. Kovaleva
    Copyright © Igor Solodovnikov
    Calosoma (Ctenosta) grandidieri Maindron, 1900
    Calosoma (Ctenosta) grandidieri Maindron, 1900
    Male adult
    S-Madagaskar, Tulear prov., near Betioky town, h = 150 m, 23°38'11" S, 44°20'9" E, 27.XII.2019, leg. A. Kozlov & Yu. Kovaleva
    Copyright © Igor Solodovnikov
    Calosoma (Callisthenes) breuningi breuningi
    Calosoma (Callisthenes) breuningi breuningi Holotype
    Male adult
    «Sergiopol», «Panderi v. n. aenea (St. blau)», «Karel. subsp. det. Ganglb.», «Callisthenes Karelini, Fisch.», «Typus», «Cal. Breuningi m. Typus, det. Ing. K. Mandl», «Breuningi m.» (NHMW).
    Copyright © Ilya Kabak
    Calosoma (Callisthenes) cyaneosterum Mandl, 1954
    Calosoma (Callisthenes) cyaneosterum Mandl, 1954 Holotype
    Male adult
    «Songarei», «Panderi det. Ganglb.», «Collectio Kaufmann», «Typus», «Calosoma cyaneosterum m. Typus, det. Ing. K. Mandl» (NHMW)
    Copyright © Ilya Kabak
    Calosoma (Callisthenes) breuningi iliense Kabak, 2022
    Calosoma (Callisthenes) breuningi iliense Kabak, 2022 Paratype
    Female adult
    «Kerbulak, 1.IV.1983, I. I. Kabak leg.»
    Copyright © Ilya Kabak
    Calosoma (Callisthenes) breuningi iliense Kabak, 2022
    Calosoma (Callisthenes) breuningi iliense Kabak, 2022 Paratype
    Male adult
    W of Kerbulak, 44°03ʹ03ʹʹ N, 77°01ʹ12ʹʹ E, 640 m, 25.IV.2017, I. I. Kabak leg.
    Copyright © Ilya Kabak
    Calosoma (Callisthenes) breuningi iliense Kabak, 2022
    Calosoma (Callisthenes) breuningi iliense Kabak, 2022 Paratype
    Female adult
    «SE Kazakhstan, right bank of Ili River, near Malaysary, 30.IV–1.V.1985, G. V. Nikolaev, A. S. Badenko leg.» (ca. 44°17ʹ30ʹʹ N, 76°57ʹ16ʹʹ E, 510 m),
    Copyright © Ilya Kabak
    Calosoma (Callisthenes) breuningi iliense Kabak, 2022
    Calosoma (Callisthenes) breuningi iliense Kabak, 2022 Paratype
    Male adult
    «SE Kazakhstan, right bank of Ili River, near Malaysary, 30.IV–1.V.1985, G. V. Nikolaev, A. S. Badenko leg.» (ca. 44°17ʹ30ʹʹ N, 76°57ʹ16ʹʹ E, 510 m),
    Copyright © Ilya Kabak
    Calosoma (Callisthenes) breuningi breuningi
    Calosoma (Callisthenes) breuningi breuningi
    Female adult
    SE Kazakhstan, Almaty region, Ak-Eshki lake, 42 km NNE Taldy-Kurgan, H = 456 m, sands, 45°21ʹ44ʺ N, 78°37ʹ04ʺ E, 21.IV.2017, S. V. Kolov leg.
    Copyright © Ilya Kabak
    Calosoma (Callisthenes) elegans kashense Kabak, 2022
    Calosoma (Callisthenes) elegans kashense Kabak, 2022 Paratype
    Female adult
    China, Xinjiang, left bank of Kash Riv., E of Nilki Town, foothills, 43°50ʹ17ʺ N, 82°48ʹ07ʺ E, H = 1270 m, 09.V.2017, I. I. Kabak leg. (ZIN)
    Copyright © Ilya Kabak
    Calosoma (Callisthenes) elegans kashense Kabak, 2022
    Calosoma (Callisthenes) elegans kashense Kabak, 2022 Paratype
    Male adult
    China, Xinjiang, left bank of Kash Riv., E of Nilki Town, foothills, 43°50ʹ17ʺ N, 82°48ʹ07ʺ E, H = 1270 m, 09.V.2017, I. I. Kabak leg. (ZIN)
    Copyright © Ilya Kabak
    Calosoma (Callisthenes) breuningi breuningi
    Calosoma (Callisthenes) breuningi breuningi
    Female adult
    SE Kazakhstan, Alakol lake, Koktuma env.
    Copyright © Ilya Kabak
    Calosoma (Callisthenes) breuningi breuningi
    Calosoma (Callisthenes) breuningi breuningi
    Female adult
    «Sergiopol» (patria falsa!), (ZIN)
    Copyright © Ilya Kabak
    Calosoma (Callisthenes) breuningi breuningi
    Calosoma (Callisthenes) breuningi breuningi
    Male adult
    Сев. предг. Джунгарского Алатау у с. Уч-Арал Ала-Кульск р-на, 7.V.1952, Проценко» (ca. 46°09ʹ25ʺ N, 80°58ʹ55ʺ E, 410 m), (ZIN)
    Copyright © Ilya Kabak
    Calosoma (Callisthenes) breuningi breuningi (as karelini vladimiri)
    Calosoma (Callisthenes) breuningi breuningi (as karelini vladimiri)
    Male adult
    Taldy-Kurgan, Arkarly Pass, 24.IV.1985, V. Cherkasov. (44°14ʹ43ʺ N, 77°42ʹ47ʺ E, 1085 m) (patria falsa!) Specimen collected together with Holotype of C. karelini vladimiri Obydov
    Copyright © Ilya Kabak
    Calosoma (Callisthenes) panderi Fischer, 1822
    Calosoma (Callisthenes) panderi Fischer, 1822
    Male adult
    «Окр. Уральска, IV.1907, Б. Уваров» (ca. 51°14ʹ N, 51°23ʹ E, H = 30 m)
    Copyright © Ilya Kabak
    Calosoma (Callisthenes) elegans rostislavi Semenov, 1906
    Calosoma (Callisthenes) elegans rostislavi Semenov, 1906
    Female adult
    Ketmen Mt. R., Kyrghyzsai gorge, 800 m, 24–25.V.1996, R. Kh. Kadyrbekov (ca. 43°19ʹ20ʹʹ N, 79°29ʹ27ʹʹ E, 1370 m)
    Copyright © Ilya Kabak
    Calosoma (Callisthenes) elegans rostislavi Semenov, 1906
    Calosoma (Callisthenes) elegans rostislavi Semenov, 1906
    Male adult
    "Кульджа 18.IV.1913" [China, Xinjiang, Kuldzha] (ZIN)
    Copyright © Ilya Kabak
    Calosoma (Callisthenes) elegans rostislavi Semenov, 1906
    Calosoma (Callisthenes) elegans rostislavi Semenov, 1906 Holotype
    Male adult
    «Семиреченская обл., р. Или, Джаркентский уезд, IV.06, В. Рюкбейль», «Calos. (Callisthen.) Rostislavi m. Typ. un. ♂. IV.07. A. Semenov det.» (ZIN)
    Copyright © Ilya Kabak
    Calosoma (Callisthenes) breuningi iliense Kabak, 2022
    Calosoma (Callisthenes) breuningi iliense Kabak, 2022
    Endophallus
    W of Kerbulak, 44°03ʹ03ʹʹ N, 77°01ʹ12ʹʹ E, 640 m, 25.IV.2017, I. I. Kabak leg.
    Copyright © Ilya Kabak
    Calosoma (Callisthenes) subtilestriatum Mandl, 1954
    Calosoma (Callisthenes) subtilestriatum Mandl, 1954 Holotype
    Male adult
    «Kuldsha», «marginatum det. Ganglb.», «Typus», «Calosoma subtilestriatum, Typus, det. Ing. K. Mandl», (NHMW).
    Copyright © Ilya Kabak
    Calosoma (Callisthenes) elegans rostislavi Semenov, 1906
    Calosoma (Callisthenes) elegans rostislavi Semenov, 1906
    Male adult
    CH, Xinjiang, S foothills of Boro-Horo Mt. R., left bank of Piliktshi Riv., E of Karagatsh Vill., placor, 44°04ʹ38ʹʹ N, 81°30ʹ49ʹʹ E – 44°04ʹ48ʹʹ N, 81°30ʹ55ʹʹ E, 990–1040 m, 07.V.2017, I. I. Kabak leg.
    Copyright © Ilya Kabak
    Calosoma (Campalita) maderae indicum Hope, 1831
    Calosoma (Campalita) maderae indicum Hope, 1831
    Male adult
    Pakistan, Kagan Valley Shagan, Mai 1981
    Copyright © Pascal Deschamps
    Calosoma (Carabomimus) digueti colimaensis Lassalle, 2009
    Calosoma (Carabomimus) digueti colimaensis Lassalle, 2009 Holotype
    Male adult
    Mexique, Jalisco, Nevado de Colima, 2000-2300m, VII.2009, in coll. Lassalle
    Copyright © Bernard Lassalle
    Calosoma (Calopachys) blaptoides oaxacaensis Lassalle, 2009
    Calosoma (Calopachys) blaptoides oaxacaensis Lassalle, 2009 Holotype
    Male adult
    Mexique, Oaxaca, 30-50 km nord Oaxaca, 2200m., VIII.2009, in coll. Lassalle.
    Copyright © Bernard Lassalle
    Calosoma (Calopachys) blaptoides tlaxiacoensis Lassalle, 2009
    Calosoma (Calopachys) blaptoides tlaxiacoensis Lassalle, 2009 Holotype
    Male adult
    Mexique, Oaxaca, Mts au sud de Sta Maria Tlaxiaco, Sra de Piedra de Olla, 2600m., VII.2008, in coll. Lassalle.
    Copyright © Bernard Lassalle
    Calosoma (Chrysostigma) cancellatus Eschscholtz, 1833
    Calosoma (Chrysostigma) cancellatus Eschscholtz, 1833
    Male adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Calosoma (Callistenia) dietzi Schaeffer, 1904: 197
    Calosoma (Callistenia) dietzi Schaeffer, 1904: 197
    Male adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH. Photo processed by Anichtchenko A. 2022
    Copyright © - - -
    Calosoma (Callistenia) discors Le Conte, 1857
    Calosoma (Callistenia) discors Le Conte, 1857
    Male adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH. Photo processed by Anichtchenko A. 2022
    Copyright © - - -
    Calosoma (Callistenia) lariversi Van Dyke, 1943: 17
    Calosoma (Callistenia) lariversi Van Dyke, 1943: 17
    Male adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH. Photo processed by Anichtchenko A. 2022
    Copyright © - - -
    Calosoma (Callistenia) subaeneus Chaudoir, 1869
    Calosoma (Callistenia) subaeneus Chaudoir, 1869
    Male adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH. Photo processed by Anichtchenko A. 2022
    Copyright © - - -
    Calosoma (Callistenia) latipennis Horn, 1870
    Calosoma (Callistenia) latipennis Horn, 1870
    Male adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH. Photo processed by Anichtchenko A. 2022
    Copyright © - - -
    Calosoma (Callistenia) luxatus Say, 1823
    Calosoma (Callistenia) luxatus Say, 1823
    Male adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH. Photo processed by Anichtchenko A. 2022
    Copyright © - - -
    Calosoma (Callistenia) wilkesi Le Conte, 1851
    Calosoma (Callistenia) wilkesi Le Conte, 1851
    Female adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH. Photo processed by Anichtchenko A. 2022
    Copyright © - - -
    Calosoma (Callistenia) monticola Casey, 1913
    Calosoma (Callistenia) monticola Casey, 1913
    Female adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH. Photo processed by Anichtchenko A. 2022
    Copyright © - - -
    Calosoma (Callistenia) oregonus Gidaspow, 1959
    Calosoma (Callistenia) oregonus Gidaspow, 1959
    Female adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH. Photo processed by Anichtchenko A. 2022
    Copyright © - - -
    Calosoma (Callistenia) schaefferi Breuning, 1928
    Calosoma (Callistenia) schaefferi Breuning, 1928
    Male adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH. Photo processed by Anichtchenko A. 2022
    Copyright © - - -
    Calosoma (Callistenia) placerus Gidaspow, 1959
    Calosoma (Callistenia) placerus Gidaspow, 1959
    Female adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH. Photo processed by Anichtchenko A. 2022
    Copyright © - - -
    Calosoma (Callistenia) subasperatus Schaeffer, 1915
    Calosoma (Callistenia) subasperatus Schaeffer, 1915
    Female adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH. Photo processed by Anichtchenko A. 2022
    Copyright © - - -
    Calosoma (Calopachys) omiltemium Bates, 1891
    Calosoma (Calopachys) omiltemium Bates, 1891
    Female adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH. Photo processed by Anichtchenko A. 2022
    Copyright © - - -
    Calosoma (Carabomimus) morelianum Bates, 1891
    Calosoma (Carabomimus) morelianum Bates, 1891
    Female adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH. Photo processed by Anichtchenko A. 2022
    Copyright © - - -
    Calosoma (Charmosta) investigator Illiger, 1798
    Calosoma (Charmosta) investigator Illiger, 1798
    Male adult

    Copyright © Kirill Makarov
    Calosoma (Campalita) chlorostictum ivinskisi Obydov & Saldaitis, 2010
    Calosoma (Campalita) chlorostictum ivinskisi Obydov & Saldaitis, 2010 Holotype Paratype
    Male+Female adult
    Yemen, Sokotra Isl., Ayhft Riv. valley, 600 m, 25.XI.2008, R. Saldaitiene & A. Saldaitis leg.
    Copyright © - - -
    Calosoma (Callisphaena) reticulatum reticulatum
    Calosoma (Callisphaena) reticulatum reticulatum
    Male adult
    Russia, Dagestan, 22 mm
    Copyright © Igor Solodovnikov
    Calosoma (Callisphaena) reticulatum reticulatum
    Calosoma (Callisphaena) reticulatum reticulatum
    Female adult
    Russia, Dagestan, 26 mm
    Copyright © Igor Solodovnikov
    Calosoma (Callitropa) macrum LeConte, 1853
    Calosoma (Callitropa) macrum LeConte, 1853
    Female adult
    Texas Live Oak Co. 20.9.2016 (cAA)
    Copyright © Alexander Anichtchenko
    Calosoma (Carabosoma) marginalis Casey, 1897
    Calosoma (Carabosoma) marginalis Casey, 1897
    Female adult
    Texas Live Oak Co. 20.9.2016 (cAA)
    Copyright © Alexander Anichtchenko
    Calosoma (Carabosoma) peregrinator Guerin-Meneville, 1844
    Calosoma (Carabosoma) peregrinator Guerin-Meneville, 1844
    Male adult
    Texas Canyon, Cochise County, Arizona 3.7.2013 (DUBC) 23 mm
    Copyright © Alexander Anichtchenko
    Calosoma (Callistenia) luxatus Say, 1823 (zimmermanni LeConte, 1848)
    Calosoma (Callistenia) luxatus Say, 1823 (zimmermanni LeConte, 1848)
    Female adult
    USA OR, Jackson Co., FR300, 0.18mi W of rd. 20, 42.02.342N 122.54.937, 20.6.2021 M. Kippenhan leg. (cAA) 15.5 mm; det. A. Anichtchenko 2023
    Copyright © Alexander Anichtchenko
    Calosoma (Callistenia) lariversi Van Dyke, 1943
    Calosoma (Callistenia) lariversi Van Dyke, 1943 Holotype
    Male adult
    coll and photo California Academy of Sciences, San Francisco
    Copyright © - - -
    Calosoma (Callistenia) latipennis Horn, 1870: 70
    Calosoma (Callistenia) latipennis Horn, 1870: 70 Holotype
    Female adult
    MCZ
    Copyright © Harvard College
    Carabus zimmermani LeConte, 1847
    Carabus zimmermani LeConte, 1847 Holotype
    Female adult
    Museum of Comparative Zoology, Harvard University, Cambridge, Ma. (https://mczbase.mcz.harvard.edu/)
    Copyright © Harvard College
    Calosoma striatulum LeConte, 1859
    Calosoma striatulum LeConte, 1859 Holotype
    Male adult
    Museum of Comparative Zoology, Harvard University, Cambridge, Ma. (https://mczbase.mcz.harvard.edu/)
    Copyright © Harvard College
    Calosoma (Callistenia) luxatus Say, 1823
    Calosoma (Callistenia) luxatus Say, 1823
    Female adult
    USA Utah, Iron Co., 18 mi East Cedar City, Jet Hwy 14-143, 9.6.2001 M. Kippenhan leg (cAA) 15 mm
    Copyright © Alexander Anichtchenko
    Calosoma (Callistenia) luxatus Say, 1823
    Calosoma (Callistenia) luxatus Say, 1823
    Female adult
    USA OR, Jackson Co., Tubs Springs State Wayside Hwy 66, 42.06.93N 122.26.5W, 9.6.2013 M. Kippenhan leg. (cAA) 18.5 mm
    Copyright © Alexander Anichtchenko
    Calosoma (Ctenosta) guineense Imhoff, 1843
    Calosoma (Ctenosta) guineense Imhoff, 1843
    Female adult
    DR Congo, N Kivu, Kasvo 2014 (DUBC) 25mm
    Copyright © Alexander Anichtchenko
    Calosoma (Castrida) peruviense Mandl, 1971
    Calosoma (Castrida) peruviense Mandl, 1971
    Male adult
    Peru, Junin, Satipo, near rio Venado, 11.11.47S 074.46.10W, 22.IX.2017 (DUBC) 25 mm
    Copyright © Alexander Anichtchenko
    Calosoma (Chrysostigma) calidus Fabricius, 1775
    Calosoma (Chrysostigma) calidus Fabricius, 1775
    Female adult
    Canada, Manitoba, 3km S Aweme, 12.6.2010 (DUBC)
    Copyright © Alexander Anichtchenko
    Calosoma (Castrida) argentinensis Csiki, 1927
    Calosoma (Castrida) argentinensis Csiki, 1927
    Female adult
    (DUBC)
    Copyright © Alexander Anichtchenko
    Calosoma (Callisthenes) elegans gongliuense Kabak & Liang, 2023
    Calosoma (Callisthenes) elegans gongliuense Kabak & Liang, 2023 Holotype Paratype
    Male+Female adult
    China, Xinjiang, Yili, Gongliu County, Wutu Vill., Institute of Zoology, Chinese Academy of Sciences” (in Chinese); “1982.IV.5;” “0262;” “102(E) 770020” (ca. 43°23ʹ52ʺ N / 82°14ʹ12ʺ E, H = 860 m) (IOZ).
    Copyright © Ilya Kabak
    Calosoma (Castrida) linelli Mutchler, 1925
    Calosoma (Castrida) linelli Mutchler, 1925 Holotype
    Male adult
    Chatham I. Galapagos Is.
    Copyright © Smithsonian NMNH
    Callisthenes declivis Dohrn, 1884
    Callisthenes declivis Dohrn, 1884 Lectotype
    Male adult
    Kazakhstan: “Turkestan,” “Type,” “elegans var. declivisDohrn,” “Mus. Zool. Polonicus Warszawa 12/45,” “Inst. Zool. P.A.N. Warszawa Cotypus Nr. 1253,” “Lectotypus Callisthenes declivis Dohrn, 1884 design. I. Kabak, 2004” (= Callisthenes elegans elegans Kirsch, 1859). (MZPW)
    Copyright © Ilya Kabak
    Calosoma (Callisthenes) elegans elegans (as valentinae Obydov, 1997)
    Calosoma (Callisthenes) elegans elegans (as valentinae Obydov, 1997)
    Male adult
    SE-Kazakhstan, Almaty reg., W of Zarechnoe vill., h=600 m, 43.715278, 76.972778, 5.IV.2018, L=23 mm
    Copyright © Ruslan Panin
    Calosoma (Callisthenes) elegans semenovi Motschulsky, 1859
    Calosoma (Callisthenes) elegans semenovi Motschulsky, 1859
    Male adult
    SE Kazakhstan, near Almaty, Komsomol vill., h=650 m, 43.458889, 76.748333, 3. V.2015, L=26 mm
    Copyright © Ruslan Panin
    Calosoma (Callisthenes) elegans semenovi Motschulsky, 1859
    Calosoma (Callisthenes) elegans semenovi Motschulsky, 1859
    Male adult
    SE Kazakhstan, near Almaty, Komsomol vill., h=650 m, 43.458889, 76.748333, 3. V.2015
    Copyright © Ruslan Panin
    Calosoma taxon page (304 photos)
  • Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
    Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
    Female adult

    Copyright © Evgeniy Chibilov
    Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
    Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
    Larva

    Copyright © Evgeniy Chibilov
    Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
    Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
    Larva

    Copyright © Evgeniy Chibilov
    Endofallus Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
    Endofallus Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
    Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
    Male adult
    Russia,
    S Ural Mts, Mt.Iremel h=800m, 4.VI.2000 Filimonov V. leg.
    male,21mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
    Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
    Male adult
    Carabus (Morphocarabus) karpinskii Rusia, S Ural mts., Jremel Mt., 1300 m 4.6.2000male, 20 mm
    19.08.2008
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
    Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
    Habitat

    Copyright ©
    Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
    Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
    Habitat

    Copyright © Evgeniy Chibilov
    Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
    Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
    Habitat
    Habitat (Iremel moun., South Ural)
    Copyright © Evgeniy Chibilov
    Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
    Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993
    Habitat
    Habitat (Iremel moun., South Ural)
    Copyright © Evgeniy Chibilov
    Carabus (Morphocarabus) karpinskii Kryzhanovskij & Matveev, 1993
    Carabus (Morphocarabus) karpinskii Kryzhanovskij & Matveev, 1993
    Female adult

    Copyright © Evgeniy Chibilov
    Carabus (Morphocarabus) karpinskii Kryzhanovskij & Matveev, 1993
    Carabus (Morphocarabus) karpinskii Kryzhanovskij & Matveev, 1993
    Female adult

    Copyright © Evgeniy Chibilov
    Carabus (Carabus) granulatus Linne, 1758
    Carabus (Carabus) granulatus Linne, 1758
    Larva
    Carabus (Carabus) granulatus granulatus Linne, 1758
    Larva of 2nd instar
    L = 16,2 mm
    Moskov Reg., Istra Distr., env. Pavlovskaya Sloboda 12.VI.2008 leg. K.Makarov
    Copyright © Kirill Makarov
    Carabus (Procrustes) coriaceus Linne, 1758
    Carabus (Procrustes) coriaceus Linne, 1758
    Larva
    Latvija photo Andris Bukejs
    Copyright © Alexander Anichtchenko
    Carabus (Diocarabus) loschnikovi loschnikovi Fischer, 1823
    Carabus (Diocarabus) loschnikovi loschnikovi Fischer, 1823
    Male adult
    Komi Resp., Pecero-Ilych Nat. Res., Zajachij Kamen h~786 62°10'06" N 59°12'53" E 8.VIII.2004 M. Chernjakhovski leg.
    L=15.0 mm
    Copyright © Evgeniy Zinoviev
    Carabus (Acathaicus) alexandrae alexandrae Semenov, 1887 Holotype
    Carabus (Acathaicus) alexandrae alexandrae Semenov, 1887 Holotype Holotype
    Male adult
    Kwantin and Li-tschara-pu, 22.VI.1885, leg. Potanin<br /> Deposited in Zoolog. Museum St. Petersburg.<br /> male, long. 22 mm<br /> photo B. Brezina
    Copyright © Boleslav Brezina
    Carabus (Acathaicus) alexandrae alexandrae Semenov, 1887
    Carabus (Acathaicus) alexandrae alexandrae Semenov, 1887
    Male adult
    male, 25mm<br /> photo B. Brezina
    Copyright © Boleslav Brezina
    Carabus (Acathaicus) alexandrae alexandrae Semenov, 1887
    Carabus (Acathaicus) alexandrae alexandrae Semenov, 1887
    Male adult
    female, 28mm<br /> photo B. Brezina
    Copyright © Boleslav Brezina
    Carabus (Acathaicus) alexandrae alexandrae Semenov, 1887
    Carabus (Acathaicus) alexandrae alexandrae Semenov, 1887
    Male adult
    female, 24mm<br /> photo B. Brezina
    Copyright © Boleslav Brezina
    Carabus (Acathaicus) alexandrae cratocephaloides Semenov, 1887 Holotype
    Carabus (Acathaicus) alexandrae cratocephaloides Semenov, 1887 Holotype Holotype
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Acathaicus) alexandrae cratocephaloides Semenov, 1887
    Carabus (Acathaicus) alexandrae cratocephaloides Semenov, 1887
    Male adult
    male, 26mm
    Copyright © Boleslav Brezina
    Carabus (Acathaicus) alexandrae cratocephaloides Semenov, 1887
    Carabus (Acathaicus) alexandrae cratocephaloides Semenov, 1887
    Male adult
    male, 24mm
    Copyright © Boleslav Brezina
    Carabus (Acathaicus) alexandrae cratocephaloides Semenov, 1887
    Carabus (Acathaicus) alexandrae cratocephaloides Semenov, 1887
    Female adult
    female, 28mm
    Copyright © Boleslav Brezina
    Carabus (Acathaicus) idolon Semenov, 1898 Holotype
    Carabus (Acathaicus) idolon Semenov, 1898 Holotype Holotype
    Male adult
    male
    Copyright © Boleslav Brezina
    Carabus (Acathaicus) interruptocostatus interruptocostatus s.str.
    Carabus (Acathaicus) interruptocostatus interruptocostatus s.str.
    Female adult
    female, 27mm
    Copyright © Boleslav Brezina
    Carabus (Acathaicus) interruptocostatus interruptocostatus s.str.
    Carabus (Acathaicus) interruptocostatus interruptocostatus s.str.
    Male adult
    male, 29mm
    Copyright © Boleslav Brezina
    Carabus (Acathaicus) interruptocostatus Semenov, 1887 Holotype
    Carabus (Acathaicus) interruptocostatus Semenov, 1887 Holotype Holotype
    Female adult

    Copyright © Boleslav Brezina
    Carabus (Acoptolabrus) constricticollis constricticollis Kraatz, 1886
    Carabus (Acoptolabrus) constricticollis constricticollis Kraatz, 1886
    Female adult

    Copyright © Alexander Anichtchenko
    Carabus (Acoptolabrus) gehinii gehinii (as aereicollis Hauser, 1921)
    Carabus (Acoptolabrus) gehinii gehinii (as aereicollis Hauser, 1921)
    Male adult

    Copyright © Alexander Anichtchenko
    Carabus (Acoptolabrus) gehinii gehinii Fairmaire, 1886
    Carabus (Acoptolabrus) gehinii gehinii Fairmaire, 1886
    Male+Female adult
    Komari, Chitose, Hokkaido, Japan.
    20-27.05.2007
    Copyright © Safronov
    Carabus (Acoptolabrus) gehinii gehinii Fairmaire, 1886
    Carabus (Acoptolabrus) gehinii gehinii Fairmaire, 1886
    Male+Female adult
    Bibi, Chitose, Hokkaido, Japan.
    03.05.2000
    Copyright © Safronov
    Carabus (Acoptolabrus) gehinii gehinii (as manoianus (Imura, 1989)
    Carabus (Acoptolabrus) gehinii gehinii (as manoianus (Imura, 1989)
    Male adult

    Copyright © Alexander Anichtchenko
    Carabus (Acoptolabrus) gehinii munakatai (Ishikawa, 1968)
    Carabus (Acoptolabrus) gehinii munakatai (Ishikawa, 1968)
    Male+Female adult
    Mt.Shiribetsudake, Kishibetsu, Hokkaido, Japan.
    05.06.2005
    Copyright © Safronov
    Carabus (Acoptolabrus) leechi leechi Bates, 1888
    Carabus (Acoptolabrus) leechi leechi Bates, 1888
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Acoptolabrus) leechi nangminicola Deuve & Li, 2001
    Carabus (Acoptolabrus) leechi nangminicola Deuve & Li, 2001
    Female adult
    N Coree, Cha Gangdo, Nangnimsan Mt., 6.2006. 31 mm. (coll. A. Montes, Madrid)
    Copyright © Alexander Anichtchenko
    Carabus (Acoptolabrus) lopatini Morawitz, 1886
    Carabus (Acoptolabrus) lopatini Morawitz, 1886
    Male+Female adult
    Russia,
    Sakhalin isl., Tchekhov Mt, VI.1997
    female, 40mm
    photo & coll. Panin R.
    Copyright © Ruslan Panin
    Carabus (Acoptolabrus) mirabilissimus rigoutianus Deuve & Li, 1998
    Carabus (Acoptolabrus) mirabilissimus rigoutianus Deuve & Li, 1998
    Female adult
    N Koree, Pyonganbukto, mt. Miaoxiang 6.2006
    28 mm
    coll. A.Montes
    Copyright © Alexander Anichtchenko
    Carabus (Acoptolabrus) schrencki schrencki (as hauryi Gehin, 1885 )
    Carabus (Acoptolabrus) schrencki schrencki (as hauryi Gehin, 1885 )
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Acoptolabrus) schrencki schrencki (as hauryi Gehin, 1885 )
    Carabus (Acoptolabrus) schrencki schrencki (as hauryi Gehin, 1885 )
    Male adult
    det. Brezina B.
    Copyright © Boleslav Brezina
    Carabus (Acoptolabrus) schrencki schrencki (as hauryi Gehin, 1885 )
    Carabus (Acoptolabrus) schrencki schrencki (as hauryi Gehin, 1885 )
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Acoptolabrus) schrencki schrencki Motschulsky, 1860
    Carabus (Acoptolabrus) schrencki schrencki Motschulsky, 1860
    Female adult
    Primorie, Lazovskii distr., Chandalaz mt., L.Voita leg 19.07.2008
    photo A.Menstschicov
    Copyright © Anton Menstschikov
    Carabus (Acoptolabrus) schrencki kimieae Kezuka & Imura, 1989
    Carabus (Acoptolabrus) schrencki kimieae Kezuka & Imura, 1989
    Male+Female adult
    Mt. Wulongshan, Dandong City,
    Liaoning Province, CHINE
    25-30.VI.2008
    Copyright © Safronov
    Carabus (Acrocarabus) callisthenoides Semenov, 1888
    Carabus (Acrocarabus) callisthenoides Semenov, 1888
    Male adult
    male, 22mm
    Copyright © Boleslav Brezina
    Carabus (Acrocarabus) callisthenoides Semenov, 1888
    Carabus (Acrocarabus) callisthenoides Semenov, 1888
    Male adult
    female, 23mm
    Copyright © Boleslav Brezina
    Carabus (Acrocarabus) guerini aldarkose Kabak, 1992
    Carabus (Acrocarabus) guerini aldarkose Kabak, 1992
    Male adult
    male, 22mm
    Copyright © Boleslav Brezina
    Carabus (Acrocarabus) guerini guerini Fischer, 1842
    Carabus (Acrocarabus) guerini guerini Fischer, 1842
    Male adult
    male, 21mm
    Copyright © Boleslav Brezina
    Carabus (Acrocarabus) guerini guerini Fischer, 1842
    Carabus (Acrocarabus) guerini guerini Fischer, 1842
    Female adult
    female, 25mm
    Copyright © Boleslav Brezina
    Carabus (Acrocarabus) guerini guerini Fischer, 1842
    Carabus (Acrocarabus) guerini guerini Fischer, 1842
    Male adult
    SE Kazakhstan, W Dzhungar Mts, Lepsinsk env., 1600 m., 18.06.2000, A.Klimenko
    Copyright © Anton Menstschikov
    Carabus (Alipaster) barovskii bayanbulak Deuve, 1993 Paratype
    Carabus (Alipaster) barovskii bayanbulak Deuve, 1993 Paratype Paratype
    Male adult
    male, 15mm
    Copyright © Boleslav Brezina
    Carabus (Alipaster) barovskii bayanbulak Deuve, 1993 Paratype
    Carabus (Alipaster) barovskii bayanbulak Deuve, 1993 Paratype Paratype
    Female adult
    female, 17mm
    Copyright © Boleslav Brezina
    Carabus (Alipaster) infantulus infantulus Morawitz, 1886
    Carabus (Alipaster) infantulus infantulus Morawitz, 1886
    Male adult
    male, 15mm
    Copyright © Boleslav Brezina
    Carabus (Alipaster) pupulus pupulus Morawitz, 1889
    Carabus (Alipaster) pupulus pupulus Morawitz, 1889
    Male adult
    Inner Tien-Shan, W part of Terskei mnts., Ukek rng., 20-25 km E Kotshkorka, h=2800 m, 8.07.98 leg. Tretyakov<br /> male 13,6mm
    Copyright © Maksim Smirnov
    Carabus (Alipaster) pupulus pupulus Morawitz, 1889
    Carabus (Alipaster) pupulus pupulus Morawitz, 1889
    Male adult
    male, 13mm
    Copyright © Boleslav Brezina
    Carabus (Alipaster) pupulus pupulus Morawitz, 1889
    Carabus (Alipaster) pupulus pupulus Morawitz, 1889
    Male adult
    female, 16mm
    Copyright © Boleslav Brezina
    Carabus (Alipaster) pupulus pupulus Morawitz, 1889
    Carabus (Alipaster) pupulus pupulus Morawitz, 1889
    Male adult
    Kirgizstan, Terskei AlaToo Mt. rng, Kadzhi-Sai v., 2800 m.; A.Subankulov leg.; 30.05.2000
    Copyright © Anton Menstschikov
    Carabus (Apoplesius) deuvei hajdaji Hackel, 2007 Holotype
    Carabus (Apoplesius) deuvei hajdaji Hackel, 2007 Holotype Holotype
    Male adult
    Turkey- prov. Mus 60 km NW Mus 1630m
    3856N 4108E Bulgan Gec. 23-24.06.03 E. & P. Hajdaj leg.
    Copyright © Boleslav Brezina
    Carabus (Apotomopterus) anxiensis Kleinfeld, 1998 Holotype & Paratype
    Carabus (Apotomopterus) anxiensis Kleinfeld, 1998 Holotype & Paratype Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) arrowi hector Hauser, 1924
    Carabus (Apotomopterus) arrowi hector Hauser, 1924
    Female adult
    China, Guangxi pro., Maoershan Mountain, Liang H.B. coll. Photo by Liuye det. Anichtchenko A.
    Copyright © Ye Liu
    Carabus (Apotomopterus) bousquetellus gengmaensis Kleinfeld, 2003 Holotype & Paratype
    Carabus (Apotomopterus) bousquetellus gengmaensis Kleinfeld, 2003 Holotype & Paratype Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) cantonensis pervarius Kleinfeld, 1997
    Carabus (Apotomopterus) cantonensis pervarius Kleinfeld, 1997 Holotype
    Male adult
    Frank Kleinfeld
    Copyright © - - -
    Carabus (Apotomopterus) cantonensis pervarius Kleinfeld, 1997
    Carabus (Apotomopterus) cantonensis pervarius Kleinfeld, 1997
    Male adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) casaleianus jigongshanicus Kleinfeld, 1998 Holotype & Paratype
    Carabus (Apotomopterus) casaleianus jigongshanicus Kleinfeld, 1998 Holotype & Paratype Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) clermontianus Breuning, 1933
    Carabus (Apotomopterus) clermontianus Breuning, 1933
    Female adult
    N Viet-Nam, 100 km NW Hanoi 5.1995 Rapaltovskii E. leg.
    Copyright © Alexander Anichtchenko
    Carabus (Apotomopterus) datianshanicus Kleinfeld, 1997 Holotype & Paratype
    Carabus (Apotomopterus) datianshanicus Kleinfeld, 1997 Holotype & Paratype Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) davidis davidis Deyrolle & Fairmaire, 1878
    Carabus (Apotomopterus) davidis davidis Deyrolle & Fairmaire, 1878
    Female adult

    Copyright © Alexander Anichtchenko
    Carabus (Apotomopterus) cantonensis metallaicus Kleinfeld, 1999 Holotype
    Carabus (Apotomopterus) cantonensis metallaicus Kleinfeld, 1999 Holotype Holotype
    Female adult
    as pervarius metallaicus
    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) davidis neodavidis Kleinfeld & Schutze, 2001 Holotype & Paratype
    Carabus (Apotomopterus) davidis neodavidis Kleinfeld & Schutze, 2001 Holotype & Paratype Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) davidis viridithoracis Kleinfeld & Schutze, 2001 Holotype & Paratype
    Carabus (Apotomopterus) davidis viridithoracis Kleinfeld & Schutze, 2001 Holotype & Paratype Holotype Paratype
    Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) davidis yizhangensis Kleinfeld, 2002
    Carabus (Apotomopterus) davidis yizhangensis Kleinfeld, 2002 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) delavayi s.str. (as medea Kleinfeld, 1999)
    Carabus (Apotomopterus) delavayi s.str. (as medea Kleinfeld, 1999) Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) delavayi tenuimanus Deuve & Imura, 1990
    Carabus (Apotomopterus) delavayi tenuimanus Deuve & Imura, 1990
    Male adult
    Lijiang Yunnan Chine
    photo Daniel Prunier
    Copyright © Daniel Prunier
    Carabus (Apotomopterus) delavayi tenuimanus Deuve & Imura, 1990
    Carabus (Apotomopterus) delavayi tenuimanus Deuve & Imura, 1990
    Female adult
    Lijiang, Yunnan, Chine
    female
    photo Daniel Prunier
    Copyright © Daniel Prunier
    Carabus (Morphocarabus) illigeri illigeri (as bjelasnicensis Apfelbeck, 1902)
    Carabus (Morphocarabus) illigeri illigeri (as bjelasnicensis Apfelbeck, 1902)
    Male+Female adult
    Serbia, Debelo Brdo, 5.VI.2007, S. Sapaly leg. L=32 mm photo Panin R.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) illigeri illigeri Dejean, 1826 (as bjelasnicensis Apfelbeck, 1902)
    Carabus (Morphocarabus) illigeri illigeri Dejean, 1826 (as bjelasnicensis Apfelbeck, 1902)
    Endophallus
    endophallus as bjelasnicensis Apfelbeck, 1902
    Copyright © Ruslan Panin
    Carabus (Archicarabus) monticola fontanae Born, 1906
    Carabus (Archicarabus) monticola fontanae Born, 1906
    Male+Female adult
    N-Italy,
    Piemonte, Bielmonte, h1350 m, 14-15.VI.2010, Pascal Stefani leg.
    female, 23 mm
    photo Panin R.
    Copyright © Ruslan Panin
    Carabus (Archicarabus) monticola fontanae Born, 1906
    Carabus (Archicarabus) monticola fontanae Born, 1906
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi hampei Kuster, 1846
    Carabus (Morphocarabus) rothi hampei Kuster, 1846
    Endophallus
    W Ukraine, Vinogradov distr.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi hampei Kuster, 1846
    Carabus (Morphocarabus) rothi hampei Kuster, 1846
    Endophallus
    endophallus hampei
    Vinogradov distr.
    Copyright © Igor Solodovnikov
    Carabus (Morphocarabus) kollari Palliardi, 1825
    Carabus (Morphocarabus) kollari Palliardi, 1825
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Morphocarabus) kollari Palliardi, 1825
    Carabus (Morphocarabus) kollari Palliardi, 1825
    Female adult
    female, 24 mm
    Copyright © Boleslav Brezina
    Carabus (Morphocarabus) kollari Palliardi, 1825
    Carabus (Morphocarabus) kollari Palliardi, 1825
    Male adult
    male, 27 mm
    Copyright © Boleslav Brezina
    Carabus (Trachycarabus) estreicheri Fischer, 1822
    Carabus (Trachycarabus) estreicheri Fischer, 1822
    Endophallus
    Lviv Region, Brody distr., near Yasenov vill.,
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) estreicheri Fischer, 1822
    Carabus (Trachycarabus) estreicheri Fischer, 1822
    Male+Female adult
    W UKRAINE, NW Podolian,
    Lviv Region, Brody distr., near Yasenov vill., steppe sites, 2-28.V.2010, Panin R. leg.
    female, 23mm
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) estreicheri Fischer, 1822
    Carabus (Trachycarabus) estreicheri Fischer, 1822
    Male adult

    Copyright © Safronov
    Carabus (Trachycarabus) estreicheri Fischer, 1822
    Carabus (Trachycarabus) estreicheri Fischer, 1822
    Female adult
    W-UKRAINE, NW Podolian,
    Lviv Region, Brody distr., near Yasenov vill., steppe sites, 12.V-2.VI.2007 Panin R. leg.
    female, 22mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi hampei Kuster, 1846
    Carabus (Morphocarabus) rothi hampei Kuster, 1846
    Male+Female adult
    W.Ukraine.
    Transcarpathian region,Vinogradiv distr., near Malaya Kopanya vill., Chorna Gora Mt., h=180m. 7-18.VII.2008. Panin R. leg.
    female 33 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi hampei Kuster, 1846
    Carabus (Morphocarabus) rothi hampei Kuster, 1846
    Male adult
    W.Ukraine.
    Transcarpathian region,Vinogradov distr., near Malaya Kopanya vill., Chorna Gora Mt., h=180m. 7-18.VII.08. Panin R. leg.
    male, 28 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi hampei Kuster, 1846 (as ormayi Reitter, 1896) loc.typ.
    Carabus (Morphocarabus) rothi hampei Kuster, 1846 (as ormayi Reitter, 1896) loc.typ.
    Female adult
    W.UKRAINE,
    Transcarpathian reg., near Beregovo,15.V-16.VI.2009 Kanarsky Yu. & Geryak Yu. leg.
    female, 30 mm
    photo/det. Panin R.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi hampei Kuster, 1846
    Carabus (Morphocarabus) rothi hampei Kuster, 1846
    Female adult
    Carabus (Morphocarabus) rothi hampei
    W Ukraine, Zakarpatian reg., Vinogradov
    1.7.1989 Avdeev I. leg.
    female, 31 mm
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) rothi hampei Kuster, 1846
    Carabus (Morphocarabus) rothi hampei Kuster, 1846
    Female adult
    arabus (Morphocarabus) rothi hampei
    W Ukraine, Zakarpatian reg., Vinogradov
    1.7.1989 Avdeev I. leg.
    female, 31 mm
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) spasskianus schestopalovi Plutenko, 1995
    Carabus (Morphocarabus) spasskianus schestopalovi Plutenko, 1995
    Male adult
    NE-Kazakhstan,
    S-Altay, Sarym-Sakty Mt. h~2500m., Sarym-Sakty riv. 24.VI.1999
    male, 23mm.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) spasskianus schestopalovi Plutenko, 1995
    Carabus (Morphocarabus) spasskianus schestopalovi Plutenko, 1995
    Male adult
    SE Kazakhstan,
    S Altay Mts., Sarymsakty mt. rng., Sarymsakty riv.
    22.VI.2000 2400m, Nekroshevich S. leg.
    male, 20 mm
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) kollari Palliardi, 1825
    Carabus (Morphocarabus) kollari Palliardi, 1825
    Male+Female adult
    Romania, SW Carpathians,
    Banat, Anina Mts. (Muntii Aninei) 8 km NE Oravita, 7.VI.2007, S.I. Budka leg.
    female, 34 mm
    photo Panin R
    Copyright © Ruslan Panin
    Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860
    Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860
    Male+Female adult
    SW -France,
    Pyrenees-Atlantiques, Foret de Sare, XII.2009, Pascal Stefani leg.
    female, 31 mm
    Copyright © Ruslan Panin
    Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860
    Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860
    Female adult
    Spain,
    Navarra, Sierra de Aralar, Altxueta Mt., 5.V.2006
    Anichtchenko A. leg.
    L= 17mm
    Copyright © Alexander Anichtchenko
    Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860
    Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860
    Male adult
    Spain,
    Pais Vasco, Bera de Bidasoa, Larrun 450m,
    9.9.2001 Anichtchenko A. leg.
    male, 29mm
    Copyright © Alexander Anichtchenko
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as delacrei Tarrier, 1975)
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as delacrei Tarrier, 1975)
    Male+Female adult
    NE Spain, Catalonia, Sant Jaume de Frontanyа, h=1000 m, V-VI.2010, Pascal Stefani leg.
    female, 38 mm
    Copyright © Ruslan Panin
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as delacrei Tarrier, 1975)
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as delacrei Tarrier, 1975)
    Male adult
    Spain,
    Catalunya, San Jaume de Frontana,
    May 2001
    Copyright © Ruslan Panin
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826
    Female adult
    Auriac alt.
    500m. (France : Aude) 25.V.200
    Copyright © Ruslan Panin
    Carabus (Carabus) arvensis sylvaticus Dejean, 1826
    Carabus (Carabus) arvensis sylvaticus Dejean, 1826
    Female adult
    France, Centre reg., Eure-et-Loir, Forкt de Montecot, VI.2009, Pascal Stefani leg.
    female, 20 mm
    Copyright © Ruslan Panin
    Carabus (Eucarabus) arcensis sylvaticus Dejean, 1826 (as sugintensis Brunier, 1922)
    Carabus (Eucarabus) arcensis sylvaticus Dejean, 1826 (as sugintensis Brunier, 1922)
    Male adult
    NW -France, Haut-Rhin Depart., Fulleren, XI.2004, Pascal Stefani leg.
    Copyright © Ruslan Panin
    Carabus (Carabus) arvensis sylvaticus Dejean, 1826 (as liebmanni Lengerken, 1911 )
    Carabus (Carabus) arvensis sylvaticus Dejean, 1826 (as liebmanni Lengerken, 1911 )
    Female adult
    France 67, Weifersweller 350m, Vosges massif
    25.8.91
    long. 21mm
    Copyright © Alexander Anichtchenko
    Carabus (Carabus) arvensis sylvaticus Dejean, 1826
    Carabus (Carabus) arvensis sylvaticus Dejean, 1826
    Male adult
    France 61, Longny-au-Perche. 30.V.1991
    male, 19 mm
    Copyright © Alexander Anichtchenko
    Carabus (Chrysocarabus) olympiae Sella, 1855
    Carabus (Chrysocarabus) olympiae Sella, 1855
    Male+Female adult
    N-Italy,
    Piemonte, Bielmonte, 14-15.VI.1981, Pascal Stefani leg.
    female, 34 mm
    photo Panin R.
    Copyright © Ruslan Panin
    Carabus (Chrysocarabus) olympiae Sella, 1855
    Carabus (Chrysocarabus) olympiae Sella, 1855
    Male adult
    Italy, Piemonte, Sessera, 2500m 23.III.1987
    Copyright © Alexander Anichtchenko
    Carabus (Coptolabrodes) haeckeli Brezina & Imura, 1997
    Carabus (Coptolabrodes) haeckeli Brezina & Imura, 1997
    Male adult
    C China, Shaanxi,
    Tsinling Mts., Taibaishan Mt., h=3000m, 33:55N/107:44 E, VI.2004, V.Siniaev & his team
    female, 31 mm
    photo Panin R.
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) sibiricus tscherkessicus Korge, 1964
    Carabus (Trachycarabus) sibiricus tscherkessicus Korge, 1964
    Male adult
    S Russia, NW-Caucasus,
    Karachay-Cherkessia,Teberda, Mukhinsky (Mukhu) Mt. pass, h=2750-2800 m , 8.VIII.2010, I.A. Solodovnikov leg.
    male, 23 mm

    Copyright © Ruslan Panin
    Carabus (Trachycarabus) sibiricus tscherkessicus Korge, 1964
    Carabus (Trachycarabus) sibiricus tscherkessicus Korge, 1964
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Megodontus) imperialis Fischer, 1823
    Carabus (Megodontus) imperialis Fischer, 1823
    Endophallus
    endophallus
    Copyright © Safronov
    Carabus (Megodontus) imperialis Fischer, 1823
    Carabus (Megodontus) imperialis Fischer, 1823
    Female adult
    Carabus (Megodontus) imperialis
    photo M. Smirnov
    from www.zin.ru
    Copyright © Maksim Smirnov
    Carabus (Eupachys) glyptopterus Fischer, 1827
    Carabus (Eupachys) glyptopterus Fischer, 1827
    Male+Female adult
    Russia, SE-Siberia,
    Buryat Republic, Ivolginsky Distr., 04.VII.2010, Filippov A.E. leg.
    female, 32 mm
    photo Panin R.
    Copyright © Ruslan Panin
    Carabus (Eupachys) glyptopterus Fischer, 1827
    Carabus (Eupachys) glyptopterus Fischer, 1827
    Female adult
    Mongolia, 15 km N Bayanteeg, 2000 m, 17.7.2006 Anichtchenko A. leg.
    Copyright © Alexander Anichtchenko
    Carabus (Megodontus) schoenherri sajanus Breuning, 1927
    Carabus (Megodontus) schoenherri sajanus Breuning, 1927
    Male+Female adult
    <b>Carabus (Megodontus) schoenherri sajanus Breuning, 1927</b><br /> Russia, SW Siberia, Krasnoyarsk reg., S of Abakan,<br /> Sayanogorsk distr., Cheryomushki vill. env.<br /> 07-VII-1994<br /> M. Danilevsky leg.<br /> male 27mm, female 31 mm.
    Copyright © Safronov
    Carabus (Megodontus) schoenherri sajanus Breuning, 1927
    Carabus (Megodontus) schoenherri sajanus Breuning, 1927
    Endophallus
    endophallus
    Copyright © Safronov
    Carabus (Megodontus) schoenherri sajanus Breuning, 1927
    Carabus (Megodontus) schoenherri sajanus Breuning, 1927
    Male adult
    Carabus (Megodontus) schoenherri sajanus Breuning, 1927
    W Sayan Mts, Borus mt. rng., near Cheremushki vill.,
    29-31.7.2000 Brinev A. leg.
    male, 30mm
    photo Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Carabus (Coptolabrus) formosus brockmanni (as nili Deuve, 2005 PT)
    Carabus (Coptolabrus) formosus brockmanni (as nili Deuve, 2005 PT) Paratype
    Male+Female adult
    C China, Shaanxi,
    Liupan Mts., Guan Shan Mt. h=800 m, , 34:50N/106:48 E, Team V.Siniaev leg.
    female, 38 mm
    photo Panin R.
    Copyright © Ruslan Panin
    Carabus (Megodontus) violaceus fulgens Charpentier, 1825
    Carabus (Megodontus) violaceus fulgens Charpentier, 1825
    Male adult
    NC-Spain,
    Navarra Reg., Sierra de Andнa, IX.2004, Pascal Stefani leg.
    male, 30 mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) violaceus fulgens Charpentier, 1825 (as gerundensis Born, 1916)
    Carabus (Megodontus) violaceus fulgens Charpentier, 1825 (as gerundensis Born, 1916)
    Male adult
    Spain, Gerona, Garriguella, XI.1990 J. Soler leg.
    male, 31mm
    Copyright © Alexander Anichtchenko
    Carabus (Procerus) gigas gigas Creutzer, 1799
    Carabus (Procerus) gigas gigas Creutzer, 1799
    Male adult
    Croatia
    Copyright © Stanislav Krejcik
    Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792
    Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792
    Female adult
    NE-FRANCE,
    Lorraine Reg., Moselle, 15.II.2010, Pascal Stefani leg.
    female, 26 mm
    Copyright © Ruslan Panin
    Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792
    Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792
    Male adult
    Austria,
    Weissenbach stt., III.1979, Kretschmer leg.
    male, 24mm
    photo Panin R.
    Copyright © Ruslan Panin
    Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792 (as aureus)
    Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792 (as aureus)
    Male adult
    France 28, Saint-Eliph, V.1991
    female, 25mm
    Copyright © Alexander Anichtchenko
    Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792
    Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792
    Female adult
    France 28, Saint-Eliph, V.1991
    female, 24 mm

    Copyright © Alexander Anichtchenko
    Carabus (Archicarabus) nemoralis lamadridae Born, 1895 (as layrei Tarrier, 1975)
    Carabus (Archicarabus) nemoralis lamadridae Born, 1895 (as layrei Tarrier, 1975)
    Male adult
    NW-Spain, León Province, Acebo, V.2007, Pascal Stefani leg.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) zawadzkii zawadzkii Kraatz, 1854
    Carabus (Morphocarabus) zawadzkii zawadzkii Kraatz, 1854
    Male adult
    Ukrainian Carpathians
    Lviv Reg., Skole Distr., near Grebeniv vill., h=1000m 9 - 20.VII.07, Panin R. leg.
    male, 31 mm
    Copyright © Safronov
    Carabus (Acathaicus) alexandrae Semenov, 1887
    Carabus (Acathaicus) alexandrae Semenov, 1887
    Male adult
    Carabus (Acathaicus) alexandrae alexandrae Semenov, 1897 Holotype
    Kwantin and Li-tschara-pu, 22.VI.1885, leg. Potanin
    Deposited in Zoolog. Museum St. Petersburg.
    male, long. 22 mm
    photo B. Brezina
    Copyright © Boleslav Brezina
    Carabus (Carpatophilus) linnaei linnaei Panzer, 1812
    Carabus (Carpatophilus) linnaei linnaei Panzer, 1812
    Male adult
    Carabus (Carpatophilus) linnaei linnaei Panzer, 1812
    Copyright © Safronov
    Carabus (Isiocarabus) strandiellus dayongensis Kleinfeld, 1996 HT
    Carabus (Isiocarabus) strandiellus dayongensis Kleinfeld, 1996 HT Holotype
    Male adult
    Carabus (Isiocarabus) strandiellus dayongensis HT
    Copyright © Frank Kleinfeld
    Carabus (Isiocarabus) dargei dargei Deuve, 1987 (as kulingensis Holotype)
    Carabus (Isiocarabus) dargei dargei Deuve, 1987 (as kulingensis Holotype) Holotype
    Male adult
    Carabus (Isiocarabus) dargei dargei (as kulingensis Holotype)
    Copyright © Frank Kleinfeld
    Carabus (Isiocarabus) dargei yulingensis (as dreuxi yulingensis) Kleinfeld, 2002 HT PT
    Carabus (Isiocarabus) dargei yulingensis (as dreuxi yulingensis) Kleinfeld, 2002 HT PT Holotype Paratype
    Male+Female adult
    Carabus (Isiocarabus) dargei yulingensis (as dreuxi yulingensis) HT PT
    Copyright © Frank Kleinfeld
    Carabus (Isiocarabus) fiduciarius fiduciarius (as fiduciarius tim)
    Carabus (Isiocarabus) fiduciarius fiduciarius (as fiduciarius tim) Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Isiocarabus) hunanicola jinxiuensis Imura, 1995 (as dreuxi xinyingensis) HT PT
    Carabus (Isiocarabus) hunanicola jinxiuensis Imura, 1995 (as dreuxi xinyingensis) HT PT Holotype Paratype
    Male+Female adult
    Carabus (Isiocarabus) hunanicola jinxiuensis (as dreuxi xinyingensis) HT PT
    Copyright © Frank Kleinfeld
    Carabus (Isiocarabus) hunanicola liangshanicus (as dreuxi liangshanicus) HT PT Kleinfeld, 1998
    Carabus (Isiocarabus) hunanicola liangshanicus (as dreuxi liangshanicus) HT PT Kleinfeld, 1998 Holotype Paratype
    Male+Female adult
    Carabus (Isiocarabus) hunanicola liangshanicus (as dreuxi liangshanicus) HT PT
    Copyright © Frank Kleinfeld
    Carabus (Isiocarabus) kiukiangensis orphniopterus Hauser, 1920
    Carabus (Isiocarabus) kiukiangensis orphniopterus Hauser, 1920
    Male adult
    Carabus (Isiocarabus) kuikiangensis orphniopterus Hauser, 1920
    China, Fujian, Yun Ling Shan,
    Fokien, 4.6.1998
    Copyright © Alexander Anichtchenko
    Carabus (Ulocarabus) theanus Reitter, 1895
    Carabus (Ulocarabus) theanus Reitter, 1895
    Male adult
    Vakhskiy Mt Range,Saryhosor, Mullonomi,
    29.4.1990, S.Zonshteyn leg.
    Copyright © Alexander Anichtchenko
    Carabus (Tribax) titan titan Zolotarev, 1913
    Carabus (Tribax) titan titan Zolotarev, 1913
    Male+Female adult
    Russia,NW Caucasus,
    Krasnodar reg.,Chernogorie Mnt. Range, Agulova Balka h=950m V-VII.1999, A.S.Zamotajlov leg.
    female,32mm
    photo Panin R.
    Copyright © Ruslan Panin
    Carabus (Tribax) titan titan Zolotarev, 1913
    Carabus (Tribax) titan titan Zolotarev, 1913
    Endophallus
    endophallus
    Copyright © Ruslan Panin
    Carabus (Tribax) titan titan Zolotarev, 1913
    Carabus (Tribax) titan titan Zolotarev, 1913
    Male adult
    Carabus (Tribax) titan titan Zolotarev, 1913
    Tuapse distr., Anastasievka vill., Semiglavaya mt.,
    Pshekaho riv., 20.7.1995
    male, 30 mm
    Copyright © Alexander Anichtchenko
    Carabus (Tribax) titan abadzechus Zamotajlov, 1988
    Carabus (Tribax) titan abadzechus Zamotajlov, 1988
    Male adult
    Carabus (Tribax) titan abadzechus Zamotajlov, 1988
    W Caucasus, Adygea, Asish-Tau Mt.R., vall riv. Bzykha, Fagus forest, h=1450m, 14-22.08.2008, I.A.Solodovnikov, O.Beljavskyi
    male, 25 mm

    Copyright © Igor Solodovnikov
    Carabus (Tribax) steveni steveni Menetries, 1832
    Carabus (Tribax) steveni steveni Menetries, 1832
    Male adult
    Carabus (Tribax) steveni steveni Menetries, 1832
    Karachaevo-Cherkessia,
    Hudes riv., near Tsarskie vorota,
    V.2009 Anichtchenko A. leg.
    male, 25mm
    photo Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Carabus (Tribax) steveni steveni Menetries, 1832
    Carabus (Tribax) steveni steveni Menetries, 1832
    Male adult
    Carabus (Tribax) steveni steveni Menetries, 1832
    C Caucasus, Kabardino-Balkaria,
    Elbrusskyi distr., riv. Malka, upper reaches riv. Sultan-Bashi, h=2400-2600 m, 26, VII, 2008. Leg. I.A.Solodovnikov.
    male, 20,5 mm
    Copyright © Igor Solodovnikov
    Carabus (Tribax) steveni steveni Menetries, 1832
    Carabus (Tribax) steveni steveni Menetries, 1832
    Female adult
    Carabus (Tribax) steveni steveni Menetries, 1832
    C Caucasus, Kabardino-Balkaria,
    Elbrusskyi distr., riv. Malka, upper reaches riv. Sultan-Bashi, h=2400-2600 m, 26, VII, 2008. Leg. I.A.Solodovnikov.
    female 24,5 mm
    Copyright © Igor Solodovnikov
    Carabus (Tribax) steveni schamyli Hampe, 1852
    Carabus (Tribax) steveni schamyli Hampe, 1852
    Female adult
    S Russia, N Ossetia, Alagir distr., near Nizhniy Tsey vill.,VII.1981
    female, 25mm
    photo Panin R.
    Copyright © Ruslan Panin
    Carabus (Tribax) steveni schamyli Hampe, 1852
    Carabus (Tribax) steveni schamyli Hampe, 1852
    Male adult
    C Caucasus, N Ossetia,
    N slope of Pastbistshnyi Mt.R., Tagardon env., h=1100m, 7-9.08.1997, leg. A.G.Koval
    male 25,5 mm
    Copyright © Igor Solodovnikov
    Carabus (Tribax) steveni schamyli Hampe, 1852
    Carabus (Tribax) steveni schamyli Hampe, 1852
    Male adult
    C Caucasus, N Ossetia,
    N slope of Pastbistshnyi Mt.R., Tagardon env., h=1100m, 7-9.08.1997, leg. A.G.Koval
    female 27 mm
    Copyright © Igor Solodovnikov
    Carabus (Tribax) steveni schamyli Hampe, 1852
    Carabus (Tribax) steveni schamyli Hampe, 1852
    Male adult
    N Osetia, B. Balkary, 1700 m, 2.9.85
    Abdurahmanov leg.
    male, 24 mm
    Copyright © Alexander Anichtchenko
    Carabus (Tribax) steveni satanas Semenov, 1896
    Carabus (Tribax) steveni satanas Semenov, 1896
    Male adult
    Carabus (Tribax) steveni satanas Semenov, 1896
    S Russia, C-Caucasus:
    Balkaria, Valley of riv. Cherek Balkarskyi, h=1750-1800m 10.VI-20.VII.1992 Leg. Zamotajlov A.
    male, 23 mm
    photo Panin R., det. Solodovnikov I.A
    Copyright © Ruslan Panin
    Carabus (Tribax) steveni satanas Semenov, 1896
    Carabus (Tribax) steveni satanas Semenov, 1896
    Male+Female adult
    Carabus (Tribax) steveni satanas Semenov, 1896
    Russia, C.Caucasus, Kabardino-Balkaria,
    valley of River Cherek Balkarsky, h=1800 m
    10.VI-20.VII.1992
    A. Zamotajlov leg.
    Copyright © Safronov
    Carabus (Tribax) steveni satanas Semenov, 1896
    Carabus (Tribax) steveni satanas Semenov, 1896
    Endophallus
    endophallus
    Copyright © Ruslan Panin
    Carabus (Tribax) puschkini puschkini Adams, 1817
    Carabus (Tribax) puschkini puschkini Adams, 1817
    Male adult
    Armenia,<br /> Shirak Province, Ashotsk, 18.VI.2001, P.V.Kazaryan leg.<br /> male 25mm
    Copyright © Ruslan Panin
    Carabus (Tribax) puschkini ishikawaianus Breuning & Ruspoli, 1970
    Carabus (Tribax) puschkini ishikawaianus Breuning & Ruspoli, 1970
    Male adult

    Copyright © Alexander Anichtchenko
    Carabus (Tribax) kasbekianus veselyi Gottwald, 1980
    Carabus (Tribax) kasbekianus veselyi Gottwald, 1980
    Female adult
    Carabus (Tribax) kasbekianus veselyi Gottwald, 1980
    Caucasus, Georgia, Egrissky Mt.R., riverheads Tekhuri river, h=2400, A.G.Koval leg.; 04.07.1989; 25 mm.; photo A. Menstschicov
    Copyright © Anton Menstschikov
    Carabus (Tribax) kasbekianus onerosus Belousov & Zamotajlov, 1993
    Carabus (Tribax) kasbekianus onerosus Belousov & Zamotajlov, 1993 Paratype
    Male+Female adult
    Carabus (Tribax) kasbekianus onerosus Belousov & Zamotajlov, 1993 Paratypes
    Caucasus, W Georgia, Svanetia, right bank o Magana riv.,
    800m, 12.V.-4.VII.1991 Zamotajlov & Miroshnikov leg.
    male 27mm, female 33mm
    photo & collection Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Carabus (Tribax) edmundi Semenov, 1896
    Carabus (Tribax) edmundi Semenov, 1896
    Male adult
    Carabus (Tribax) edmundi Semenov, 1896
    Copyright © Boleslav Brezina
    Carabus (Tribax) circassicus teberdensis Zolotarev, 1913
    Carabus (Tribax) circassicus teberdensis Zolotarev, 1913
    Male+Female adult
    W-Caucasus: Karacaj-Cherkessija,
    Abishira–Akhuba Mnt. Range h=2000-2650m, Kyafar-Agur Riv., 14–17.VII.2009, Fominykh D. leg.
    female, 25mm
    Copyright © Ruslan Panin
    Carabus (Tribax) circassicus kubaniensis Semenov, 1896
    Carabus (Tribax) circassicus kubaniensis Semenov, 1896
    Male adult
    N Caucasus, Krasnodar reg., 30 km S Maykop
    august 1993
    male, 26 mm
    Copyright © Alexander Anichtchenko
    Carabus (Tribax) circassicus kubaniensis Semenov, 1896
    Carabus (Tribax) circassicus kubaniensis Semenov, 1896
    Male+Female adult
    Russia, NW Caucasus,
    Krasnodar reg., Chernogorie mts., 24.V-29.VII.1999. A.S. Zamotajlov leg.
    female 29mm/male 25mm
    Copyright © Ruslan Panin
    Carabus (Tribax) circassicus kubaniensis Semenov, 1896
    Carabus (Tribax) circassicus kubaniensis Semenov, 1896
    Endophallus
    endophallus
    Copyright © Ruslan Panin
    Carabus (Tribax) circassicus circassicus Ganglbauer, 1886
    Carabus (Tribax) circassicus circassicus Ganglbauer, 1886
    Male+Female adult
    Abkhasia,
    W.slope of Gagrinsky Mt., Krepostnaya River source, h=930-980m., A.S.Zamotajlov leg. 27.VII.2000
    female, 29 mm/male, 26mm
    photo Panin R.
    Copyright © Ruslan Panin
    Carabus (Tribax) circassicus abagonensis Starck, 1894
    Carabus (Tribax) circassicus abagonensis Starck, 1894
    Male+Female adult
    S Russia, NW Caucasus,
    Adygea Rep., Mt.Abago, 26.VI.1999
    L=24mm
    photo Panin R.
    Copyright © Ruslan Panin
    Carabus (Tribax) biebersteini ulgeni Gottwald, 1982
    Carabus (Tribax) biebersteini ulgeni Gottwald, 1982
    Male adult
    S Russia, Karacaj-Cherkessija,
    near Kart-Djurt vill., upper Kylan riv. h=1300m, 26-27.V.2009, Fominykh D. leg.
    male, 28mm
    Copyright © Ruslan Panin
    Carabus (Tribax) biebersteini otcharensis Kurnakov, 1970
    Carabus (Tribax) biebersteini otcharensis Kurnakov, 1970
    Male adult
    Caucasus, W Georgia, Svanetia, upper reaches of riv. Shoudidi, 1700-2100m, 20-21.V.1986 leg. A. Zamotajlov
    Copyright © Evgeniy Chibilov
    Carabus (Tribax) biebersteini ingusch Zolotarev, 1913
    Carabus (Tribax) biebersteini ingusch Zolotarev, 1913
    Male+Female adult
    Carabus (Tribax) biebersteini ingusch Zolotarev, 1913
    Russia, C.Caucasus, N.Ossetia,
    valley of River Khazniaush
    10.VI-31.VII.1993
    A. Zamotajlov leg.
    Copyright © Safronov
    Carabus (Tribax) biebersteini constantinowi Starck, 1894
    Carabus (Tribax) biebersteini constantinowi Starck, 1894
    Male adult
    Carabus (Tribax) biebersteini constantinowi Starck, 1894
    NW-Caucasus, S Russia,
    Karacaj-Cherkessija, Nizhniy Arkhyz, 21-30.V.2001, Tikhonov V. leg.
    male,30mm
    Copyright © Ruslan Panin
    Carabus (Tribax) apschuanus apschuanus Rost, 1893
    Carabus (Tribax) apschuanus apschuanus Rost, 1893
    Male+Female adult
    Carabus (Tribax) apschuanus apschuanus Rost, 1893    
    Russia, Caucasus, Krasnodar reg.,
    h=1200 m, 29.VII.2000
    A. Zamotajlov leg.
    Copyright © Safronov
    Carabus (Tribax) agnatus agnatus Ganglbauer, 1889
    Carabus (Tribax) agnatus agnatus Ganglbauer, 1889
    Male adult
    NW-Caucasus,
    Krasnodar reg., Mostovskoj distr., Skirda Mts. N. slope Yatyrgvart Mt., h=2300-2500m. 09-10.VII.2008, D.Fominyh leg.
    male, 23mm
    photo Panin R.
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) sibiricus sibiricus Fischer, 1822
    Carabus (Trachycarabus) sibiricus sibiricus Fischer, 1822
    Male adult

    Copyright © Kirill Makarov
    Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886
    Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886
    Male+Female adult
    W Ukraine,<br /> Ternopol reg., Pidvolochisk distr., Supranivka vill.<br /> 3.VII.1908. Leg. Stobetskii. S.<br /> Photo R. Panin
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886
    Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886
    Male adult
    W Ukraine,
    Ternopil reg., Pidvolohcisk distr., Supranivka vill.
    male, 26mm
    The end of XIX the beginning of XX century
    photo Panin R.
    coll. SMNH Ukraine, Lviv
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886
    Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886
    Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886
    Endophallus
    aedeagus
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) sibiricus obliteratus Fischer, 1827
    Carabus (Trachycarabus) sibiricus obliteratus Fischer, 1827
    Male adult
    Altai mts., Kalbinskii mt.rng.
    Ognevka vill. VII.1994, Klimenko A. leg.
    male, 28 mm
    Copyright © Alexander Anichtchenko
    Carabus (Trachycarabus) sibiricus obliteratus Fischer, 1827
    Carabus (Trachycarabus) sibiricus obliteratus Fischer, 1827
    Male adult
    NE Kazakhstan,
    S Altai Mts., Ulbinsky range h~500m.,Turgusun vill., 20.VI.2001
    male 25mm
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) sibiricus obliteratus Fischer, 1827
    Carabus (Trachycarabus) sibiricus obliteratus Fischer, 1827
    Endophallus
    endophallus sibiricus obliteratus Fischer, 1827
    NE Kazakhstan,
    S Altai mts.,S Ulbinsky mt.h=500m.,Turgusun vill., 20.VI.2001
    male 25mm.
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) sibiricus kolosovi Zinoviev, 1997 loc.typ.
    Carabus (Trachycarabus) sibiricus kolosovi Zinoviev, 1997 loc.typ.
    Female adult
    S. Ural, Bashkiria, near Mesjagutovo vill., 09.07.2001 A. Menstschicov leg.
    photo A. Menstschicov
    Copyright © Anton Menstschikov
    Carabus (Trachycarabus) sibiricus kolosovi Zinoviev, 1997
    Carabus (Trachycarabus) sibiricus kolosovi Zinoviev, 1997
    Female adult
    Russia, North-East Bashkortostan. Ai river, near Sikiyaz-Tamak vill.
    12.07.1997, Zinoviev E.leg
    length: 21,8 mm
    Copyright © Evgeniy Zinoviev
    Carabus (Trachycarabus) sibiricus haeres (as fossulatus Dejean, 1826)
    Carabus (Trachycarabus) sibiricus haeres (as fossulatus Dejean, 1826)
    Female adult
    <b>Carabus (Trachycarabus) sibiricus fossulatus Dejean, 1826</b><br /> NE Ukraine,<br /> Kharkov reg.,1974<br /> female, 27mm.
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) sibiricus haeres (as fossulatus Dejean, 1826)
    Carabus (Trachycarabus) sibiricus haeres (as fossulatus Dejean, 1826)
    Male+Female adult
    E.UKRAINE,<br /> Kharkov reg., Zmiev distr.,in steppe,10-20.V I.2007, D.Kovalchuk leg.<br /> female, 26mm/male, 24mm.<br /> photo/det. Panin R.
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) sibiricus haeres (as fossulatus Dejean, 1826)
    Carabus (Trachycarabus) sibiricus haeres (as fossulatus Dejean, 1826)
    Endophallus
    endophallus
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) sibiricus errans Fischer, 1823
    Carabus (Trachycarabus) sibiricus errans Fischer, 1823
    Female adult
    Carabus (Trachycarabus) sibiricus errans Fischer, 1823
    S.Russia, Rostov-Don reg., Avilo-Uspenskaya env.
    V.2000
    Copyright © Safronov
    Carabus (Trachycarabus) sibiricus errans Fischer, 1823
    Carabus (Trachycarabus) sibiricus errans Fischer, 1823
    Male adult
    SW-Russia,
    Rostov reg. env. stanitsa Razdorskaya, A. Ponomarev leg. 9-15.V.2001
    male 25mm
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) sibiricus errans Fischer, 1823
    Carabus (Trachycarabus) sibiricus errans Fischer, 1823
    Male+Female adult
    female:SE Ukraine,
    Donetsk, 25.VII.2009, A.I.Gubin leg. L=30mm.
    male:E Ukraine, Lugansk reg., near Slavyanoserbk, 5-15.VIII.2009, S.Konovalov leg. L=26mm.
    photo/det.Panin R.
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) sibiricus errans Fischer, 1823
    Carabus (Trachycarabus) sibiricus errans Fischer, 1823
    Endophallus
    endophallus sibiricus errans
    SW-Russia,
    Rostov reg. env. stanitsa Razdorskaya, A.Ponomarev leg
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) sibiricus errans Fischer, 1823
    Carabus (Trachycarabus) sibiricus errans Fischer, 1823
    Endophallus
    endophallus sibiricus errans- Lugansk reg., near Slavyanoserbk
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) sibiricus bosphoranus Fischer, 1823
    Carabus (Trachycarabus) sibiricus bosphoranus Fischer, 1823
    Female adult
    Carabus (Trachycarabus) sibiricus bosphoranus Fischer, 1823
    S Ukraine,
    Krym Peninsula, Nikitskaya Jaila, S.Mosyanin leg. 10.VIII.2000
    female 23mm.
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) sibiricus bosphoranus Fischer, 1823
    Carabus (Trachycarabus) sibiricus bosphoranus Fischer, 1823
    Male adult
    Carabus (Trachycarabus) sibiricus bosphoranus Fisch. 1823<br /> S Ukraine, Krym Peninsula,<br /> Dolgorukovskaya jaila, 25.VIII.1991 I.A.Solodovnikov leg.<br /> male, 23mm<br />
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) scabriusculus scabriusculus (as erythropus Fischer, 1822)
    Carabus (Trachycarabus) scabriusculus scabriusculus (as erythropus Fischer, 1822)
    Male+Female adult
    W.Ukraine,<br /> NW Podolian Upland , Lviv Reg., Zolochiv distr., near Luky vill., steppe sites, 7.V-8.VI. 2010<br /> leg. Panin R.<br /> female, 22mm
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) scabriusculus scabriusculus (as erythropus Fischer, 1822)
    Carabus (Trachycarabus) scabriusculus scabriusculus (as erythropus Fischer, 1822)
    Male adult
    W-UKRAINE<br /> NW Podolian Upland, Lviv Reg., Zolochiv Distr., near Luka vill., in steppe. Panin R. leg. 1.V - 16.VI.07
    Copyright © Safronov
    Carabus (Trachycarabus) scabriusculus scabriusculus Olivier, 1795
    Carabus (Trachycarabus) scabriusculus scabriusculus Olivier, 1795
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Trachycarabus) scabriusculus inapertus Motschulsky, 1850 (loc.typ)
    Carabus (Trachycarabus) scabriusculus inapertus Motschulsky, 1850 (loc.typ)
    Male+Female adult
    NE UKAINE,
    Kharkov reg., Zmiev distr., in steppe, 15.VI-14.VII.2009, Kovalchuk D. leg.
    female, 20mm
    photo/det. Panin R.
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) scabriusculus inapertus Motschulsky, 1850 (loc.typ)
    Carabus (Trachycarabus) scabriusculus inapertus Motschulsky, 1850 (loc.typ)
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Trachycarabus) planus planus Gehin, 1885
    Carabus (Trachycarabus) planus planus Gehin, 1885
    Male+Female adult
    S Russia, NW Caucasus,<br /> Karacaj-Cherkessija, near Kart-Djurt vill., upper Kylan riv. h=1300m, 26-27.V.2009 , Fominykh D. leg.<br /> L=24mm<br /> photo/det. Panin R.
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) planus planus Gehin, 1885
    Carabus (Trachycarabus) planus planus Gehin, 1885
    Male adult
    NW Caucasus, Rostov reg.,<br /> Avilo-Uspenskaya vill. 22.5.2000 Arzanov Yu. leg.<br /> male, 25 mm
    Copyright © Alexander Anichtchenko
    Carabus (Trachycarabus) planus planus Gehin, 1885
    Carabus (Trachycarabus) planus planus Gehin, 1885
    Male adult
    S Russia,<br /> Krasnodar reg., Anapa distr., B. Utrish vill., env., 11-20.VIII.2008, Khatchikov E. leg.<br /> male, 26mm<br /> photo Panin R.
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) planus planus Gehin, 1885
    Carabus (Trachycarabus) planus planus Gehin, 1885
    Endophallus
    <b>endophalus </b> S Russia,Krasnodar reg., Anapa distr., B. Utrish vill., env.,
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) planus planus Gehin, 1885
    Carabus (Trachycarabus) planus planus Gehin, 1885
    Endophallus
    <b>endophalus</b> Karacaj-Cherkessija, near Kart-Djurt vill., upper Kylan riv. h=1300m
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) planus campicola Reitter, 1896
    Carabus (Trachycarabus) planus campicola Reitter, 1896
    Male adult
    Krym Peninsula, near Sudak. Leg.Vashchenko S. 1-5.V.2004<br /> 27 mm
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) planus campicola Reitter, 1896
    Carabus (Trachycarabus) planus campicola Reitter, 1896
    Male+Female adult
    S Ukraine, Krym Peninsula,<br /> near Perevalnoe vill., in steppe, 15.IV.2009, Shaporinskiy V. leg.<br /> female, 28 mm<br /> photo/det.Panin R.
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) planus campicola Reitter, 1896
    Carabus (Trachycarabus) planus campicola Reitter, 1896
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Trachycarabus) mandibularis mandibularis Fischer, 1827
    Carabus (Trachycarabus) mandibularis mandibularis Fischer, 1827
    Male adult
    NE Kazakhstan, S Altai Mt., Narymskii mt. rng.
    Bolshenarymskoe vill. 26.7.2001, 1800m Klimenko leg.
    male, 21 mm
    Copyright © Alexander Anichtchenko
    Carabus (Trachycarabus) mandibularis buchtarmensis Kryzhanovskij, 1953
    Carabus (Trachycarabus) mandibularis buchtarmensis Kryzhanovskij, 1953
    Male adult
    Kazakhstan, Kalbinsky mt. rng., 600m ,
    Ognevka vill., 23.6.98 Klimenko A. leg.
    male, 23 mm
    Copyright © Alexander Anichtchenko
    Carabus (Trachycarabus) mandibularis buchtarmensis Kryzhanovskij, 1953
    Carabus (Trachycarabus) mandibularis buchtarmensis Kryzhanovskij, 1953
    Male adult
    NE-Kazakhstan,
    S-Altays, Kalbinky Mt. h=500m., near Ognevka vill., 20.VI.1998.A. Klimenko leg.
    male, 23mm
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) mandibularis buchtarmensis Kryzhanovskij, 1953
    Carabus (Trachycarabus) mandibularis buchtarmensis Kryzhanovskij, 1953
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
    Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
    Male adult
    Burjatia, Dzhida distr., near Beloozersk vill. 4-9.VII.1997 Anichtchenko A. leg.
    male, 23 mm
    Copyright © Alexander Anichtchenko
    Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
    Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
    Male adult
    Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
    China, Beijing, Mentougou, 2005;
    Photo & det. by liuye

    Copyright © Ye Liu
    Carabus (Trachycarabus) besseri Fischer, 1822
    Carabus (Trachycarabus) besseri Fischer, 1822
    Male adult

    Copyright © Safronov
    Carabus (Trachycarabus) besseri Fischer, 1822
    Carabus (Trachycarabus) besseri Fischer, 1822
    Male+Female adult
    W Ukraine.NW Podolian,
    Lviv reg., Zolochiv distr., Lysa Gora near Stinka vill., steppe sites, 25.IV-9.V.2009 Panin R. leg
    female, 30mm
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) besseri Fischer, 1822
    Carabus (Trachycarabus) besseri Fischer, 1822
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Tomocarabus) scabripennis scabripennis Chaudoir, 1850
    Carabus (Tomocarabus) scabripennis scabripennis Chaudoir, 1850
    Male adult
    Carabus (Tomocarabus) scabripennis scabripennis Chaudoir, 1850
    C. Armenia,
    Kotayk reg., Yeghvard, Arailer Mt., 16.VI.1999, P.V.Kazaryan leg.
    male, 20mm
    photo Panin R.
    Copyright © Ruslan Panin
    Carabus (Tomocarabus) scabripennis marthaiellus Rapuzzi & Kleinfeld, 2000
    Carabus (Tomocarabus) scabripennis marthaiellus Rapuzzi & Kleinfeld, 2000 Holotype Paratype
    Male+Female adult
    Carabus (Tomocarabus) scabripennis marthaiellus Rapuzzi & Kleinfeld, 2000
    Copyright © Frank Kleinfeld
    Carabus (Tomocarabus) marginalis Fabricius, 1794
    Carabus (Tomocarabus) marginalis Fabricius, 1794
    Male adult
    male, 26 mm
    Copyright © Ruslan Panin
    Carabus (Tomocarabus) marginalis Fabricius, 1794
    Carabus (Tomocarabus) marginalis Fabricius, 1794
    Male+Female adult
    E Ukraine,
    Kharkov reg.,near Vasishchevo vill.10-20.VI.2006 D.Kovalchuk leg.
    female 24mm
    Copyright © Ruslan Panin
    Carabus (Tomocarabus) decolor Fischer, 1823
    Carabus (Tomocarabus) decolor Fischer, 1823
    Female adult
    Carabus (Tomocarabus) decolor Fischer, 1823
    Copyright © Alexander Anichtchenko
    Carabus (Tomocarabus) decolor Fischer, 1823
    Carabus (Tomocarabus) decolor Fischer, 1823
    Male adult
    S Russia, NW-Caucasus,
    Karacaj-Cherkessija,Skalisty Mnt. Range, Site Banditskoe h=1400m, 2-3.V.2009,
    Fominykh D., Bondarenko A.S. leg.
    male,17mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) wulffiusi wulffiusi Morawitz, 1862
    Carabus (Morphocarabus) wulffiusi wulffiusi Morawitz, 1862
    Male+Female adult
    Russia, Far East,
    Primorskiy Kray, near Kamenushka vill., 2-3.VIII.1992, K.E.Dovgaylo leg.
    female, 24mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) wulffiusi wulffiusi Morawitz, 1862
    Carabus (Morphocarabus) wulffiusi wulffiusi Morawitz, 1862
    Female adult

    Copyright © Safronov
    Carabus (Morphocarabus) wulffiusi wulffiusi Morawitz, 1862
    Carabus (Morphocarabus) wulffiusi wulffiusi Morawitz, 1862
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
    Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
    Male adult
    Serbia, Majdanpek, Ceremosja 15.6.2001 Z.Zlatic leg. male, 32 mm
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
    Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
    Male+Female adult
    NE-Serbia, Krs Mts., Majdanpek env.,h=300m.20-31.V.2006, Pr.Sima leg. female, 32mm/male, 30mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
    Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
    Endophallus
    NE-Serbia, Krs Mts., Majdanpek env.,h=300m.20-31.V.2006, Pr.Sima leg
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) venustus venustus Morawitz, 1862
    Carabus (Morphocarabus) venustus venustus Morawitz, 1862
    Male adult
    SE Primorskii reg., Lazovskii Nat.Res., 12 km S Staraya Kamenka vill., near Amerika kor. 43°15N, 134°1E, 19.VII.04 leg. M.& L. Smirnovy photo from www.zin.ru
    Copyright © Maksim Smirnov
    Carabus (Morphocarabus) tarbagataicus pietrorattii Deuve, 1991(as pseudogebleri Deuve, 1994)
    Carabus (Morphocarabus) tarbagataicus pietrorattii Deuve, 1991(as pseudogebleri Deuve, 1994)
    Male adult
    Carabus (Morphocarabus) tarbagataicus pietrorattii Deuve, 1991(as pseudogebleri Deuve, 1994)
    S Tarbagatai mt.rng., Alexeevka vill.,
    600m, 11.5.2001 Klimenko A. leg.
    male, 30 mm
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) tarbagataicus pietrorattii Deuve, 1991
    Carabus (Morphocarabus) tarbagataicus pietrorattii Deuve, 1991
    Female adult

    Copyright © Ruslan Panin
    Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997 HT
    Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997 HT Holotype
    Male adult
    Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997 HT
    Rusia, SE Tuva, Sangilen mt.rng., Kargy riv., 50°31'N 97°01'E,
    1800 m, 29.07.1996, D. Obydov leg.

    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997
    Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997
    Female adult
    Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997
    Tuva, Akademika Obrucheva mt.rng., 15 km N Cherbi vill.
    ur. Chailag 7.VII.1999 A. & N. Menschikovy
    female, 22 mm
    det. Menschikov A.
    photo Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997
    Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997
    Male adult
    Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997
    Tuva, Akademika Obrucheva mt.rng., 15 km N Cherbi vill.
    ur. Chailag 7.VII.1999 A. & N. Menschikovy
    male, 21 mm
    det. Menschikov A.
    photo Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) spasskianus tungus Obydov, 1999 HT
    Carabus (Morphocarabus) spasskianus tungus Obydov, 1999 HT Holotype
    Male adult
    Carabus (Morphocarabus) spasskianus tungus Obydov, 1999 HT
    Rusia, N Krasnoyarsk reg., Ilimpeya riv and Dovogpokan riv.
    VII.1974, Lunin leg.
    photo from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) spasskianus spasskianus Breuning, 1932
    Carabus (Morphocarabus) spasskianus spasskianus Breuning, 1932
    Male adult
    Siberia, Lake Baikal, Listvianka vill. 25.8.90
    Anichtchenko A. leg.
    male, 23 mm
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) spasskianus spasskianus Breuning, 1932
    Carabus (Morphocarabus) spasskianus spasskianus Breuning, 1932
    Male+Female adult
    SE Siberia, Russia
    Irkutsk reg., near Pivovarikha vill., 7.V.07, D. Fominyh leg.
    male, 24mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) spasskianus spasskianus Breuning, 1932
    Carabus (Morphocarabus) spasskianus spasskianus Breuning, 1932
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Morphocarabus) spasskianus shoriensis Obydov, 1999 HT
    Carabus (Morphocarabus) spasskianus shoriensis Obydov, 1999 HT Holotype
    Male adult
    Gornaya Shoria, 8.VIII.1978, S. Nikireev leg.

    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) spasskianus schestopalovi Plutenko, 1995
    Carabus (Morphocarabus) spasskianus schestopalovi Plutenko, 1995
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Morphocarabus) spasskianus saragaschensis Obydov, 1999
    Carabus (Morphocarabus) spasskianus saragaschensis Obydov, 1999
    Male adult
    W Sayan Mts., Batenevskii Mt.rng., 12 km E Pervomaiskoe vill.,
    Enisey riv., mountain steppe, 400m, 5.VII.2000 A. & N. Menschikovy leg.
    female, 26mm

    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) spasskianus saragaschensis Obydov, 1999
    Carabus (Morphocarabus) spasskianus saragaschensis Obydov, 1999 Holotype
    Male adult
    Rusia, Khakassia, Saragash, 400 m, 6.VII.1994, D. Obydov leg.

    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) spasskianus putus Motschulsky, 1846
    Carabus (Morphocarabus) spasskianus putus Motschulsky, 1846
    Male adult
    Carabus (Morphocarabus) spasskianus putus Motschulsky, 1846
    SW Siberia, Altay Mts., Shebalino distr., Kumalyr env.
    02.07.2003, A.Safronov leg.
    Copyright © Safronov
    Carabus (Morphocarabus) spasskianus microputus O.Berlov & Shilenkov, 1996
    Carabus (Morphocarabus) spasskianus microputus O.Berlov & Shilenkov, 1996
    Female adult
    Tuva, 25 km E Ak-Dovurak 23.VI.2006
    A. & N. Menschikovy leg.
    female, 24 mm
    det. Menschikov A.


    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) spasskianus microputus O.Berlov & Shilenkov, 1996
    Carabus (Morphocarabus) spasskianus microputus O.Berlov & Shilenkov, 1996
    Male adult
    Tuva, 25 km E Ak-Dovurak 23.VI.2006
    A. & N. Menschikovy leg
    female, 24 mm
    det. Menschikov A.

    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) spasskianus microputus O.Berlov & Shilenkov, 1996
    Carabus (Morphocarabus) spasskianus microputus O.Berlov & Shilenkov, 1996
    Male adult
    Rusia, SW Tuva, W Tannu-Ola mt.rng., bass. of Barlyk riv.,
    50°29'N 90°44'E, 30.VII.2000, D. Obydov leg.
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) spasskianus korostelevi Obydov, 2004loc.typ.
    Carabus (Morphocarabus) spasskianus korostelevi Obydov, 2004loc.typ.
    Male adult
    E Kazahstan, Ivanovskii Mts., Prohodnoy Belok, 03.06.2000, A. Ivanov leg.

    Copyright © Anton Menstschikov
    Carabus (Morphocarabus) spasskianus geblerianus Obydov, 1999
    Carabus (Morphocarabus) spasskianus geblerianus Obydov, 1999 Holotype
    Male adult
    Rusia, Barnaul, 27.5.1965
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) spasskianus cracens Obydov, 1999
    Carabus (Morphocarabus) spasskianus cracens Obydov, 1999
    Male adult
    Carabus (Morphocarabus) spasskianus cracens Obydov, 1999
    NE Kazakhstan, S Altay Mts., Kalbinskii range,
    Ognevka vill., 600m, 18.6.1999 A. Klimenko leg.
    male, 22 mm
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) spasskianus cracens Obydov, 1999
    Carabus (Morphocarabus) spasskianus cracens Obydov, 1999 Paratype
    Male adult
    Museum of Institute of Systematic Biology (Daugavpils, Latvia)
    photo Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) spasskianus cracens Obydov, 1999
    Carabus (Morphocarabus) spasskianus cracens Obydov, 1999
    Female adult
    Carabus (Morphocarabus) spasskianus cracens Obydov, 1999
    NE-Kazakhstan:
    S-Altay, Kalbinsky Mt., Ognevka vill., h~600m 18.VI.1999 A. Klimenko leg.
    female, 24mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) spasskianus cracens Obydov, 1999
    Carabus (Morphocarabus) spasskianus cracens Obydov, 1999 Holotype
    Male adult
    Carabus (Morphocarabus) spasskianus cracens HT
    E Kazakhstan, Kolbinskii Mt.rng., 600m, near Ognevka vill.,
    13-18.07.1999, D. Obydov leg
    photo from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Cyclocarabus) belousovi Kabak, 1992
    Carabus (Cyclocarabus) belousovi Kabak, 1992
    Female adult
    Uzbekistan, Karzhantau Mt., Aktash vill.
    1500 m, 12.4.1999
    female, 23 mm
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
    Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
    Female adult
    Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
    Moravia mer.,Vranov, 26-29.06.1979, Dr. Resl leg.
    photo A. Menstschicov
    Copyright © Anton Menstschikov
    Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
    Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
    Male adult
    Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
    SE Moravia mer.,Vranov, 26-29.06.1979, Dr. Resl leg.
    photo A. Menstschicov
    Copyright © Anton Menstschikov
    Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
    Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
    Male adult
    Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
    S.Bohemia,
    Hluboka nad Vltavou, 24.VI-18.VII.1990, S. Budka leg.
    male, 27mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
    Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799 (as scheidleri floriani)
    Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799 (as scheidleri floriani)
    Male adult
    Carabus (Morphocarabus) scheidleri scheidleri (as scheidleri floriani)
    A.: Steiermark, Rein Umg. VIII.1983 Kenyery
    female, 32 mm
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) excellens excellens Fabricius, 1798
    Carabus (Morphocarabus) excellens excellens Fabricius, 1798
    Male adult
    NW Podolian
    Copyright © Safronov
    Carabus (Morphocarabus) excellens excellens Fabricius, 1798
    Carabus (Morphocarabus) excellens excellens Fabricius, 1798
    Male adult
    Moldova, 2003
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) excellens (as lomnitzkii Reitter, 1896 )
    Carabus (Morphocarabus) excellens (as lomnitzkii Reitter, 1896 )
    Male adult
    W-UKRAINE, NW Podolian, Lviv Reg., Zolochiv Distr., W Ukraine, Lysa Gora steppe massif, 19.V.-7.VI.2007, Panin R. leg. 25 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) excellens excellens Fabricius, 1798 (ab.erythromerus Dejean, 1826)
    Carabus (Morphocarabus) excellens excellens Fabricius, 1798 (ab.erythromerus Dejean, 1826)
    Male adult
    Moldova, Anenii Noi distr., near Garbovets vill. 20.VI.08 Bodnar' I.I.leg. male, 26mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) excellens excellens Fabricius, 1798
    Carabus (Morphocarabus) excellens excellens Fabricius, 1798
    Endophallus
    Moldova,<br /> Anenii Noi distr., near Garbovets vill.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) excellens (as lomnitzkii Reitter, 1896)
    Carabus (Morphocarabus) excellens (as lomnitzkii Reitter, 1896)
    Male adult
    W Ukraine, NW Podolian, Ternopil reg., Pidvilochisk distr., near Rosokhuvatets & Kamyanki vill., steppe sites, 24.V-15.VI.2008, Panin R. leg.
    male, 24 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) helleri Ganglbauer, 1893
    Carabus (Morphocarabus) helleri Ganglbauer, 1893
    Male adult
    Moravia Mer., Bile Karpaty 28.5.1993 Budka leg. male, 28 mm
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) helleri Ganglbauer, 1893
    Carabus (Morphocarabus) helleri Ganglbauer, 1893
    Male+Female adult
    SE Moravia, Strani, V.2002, M.Snizek leg. male bronze colour-26mm male blue colour-28mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) helleri Ganglbauer, 1893
    Carabus (Morphocarabus) helleri Ganglbauer, 1893
    Endophallus
    SE Moravia, Strani, V.2002, M.Snizek leg. male bronze colour.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) helleri Ganglbauer, 1893
    Carabus (Morphocarabus) helleri Ganglbauer, 1893
    Endophallus
    SE Moravia, Strani, V.2002, M.Snizek leg. male blue colour
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi incompsus (as mendax Csiki,1906)
    Carabus (Morphocarabus) rothi incompsus (as mendax Csiki,1906)
    Male+Female adult
    W.Ukraine,
    Transcarpathian Reg., Rakhov Distr., Chornogora Mt Range, h=1900m. alpine zone, Panin R. leg/det. 28.VI-9.VIII.2008
    female, 32 mm
    photo endophallus Solodovnikov I.A
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi comptus Dejean, 1831
    Carabus (Morphocarabus) rothi comptus Dejean, 1831
    Male adult
    Carabus (Morphocarabus) rothi comptus Dejean, 1831
    Carpathians
    male, 23mm
    coll.SMNH Ukraine ,Lvi
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi alutensis Savulescu, 1972
    Carabus (Morphocarabus) rothi alutensis Savulescu, 1972
    Male adult
    C Romania,
    Olt River: Rimnica Vilcea VII.1971, Savulescu N. leg.

    Copyright © Igor Solodovnikov
    Carabus (Eucarabus) obsoletus carpathicus Palliardi, 1825
    Carabus (Eucarabus) obsoletus carpathicus Palliardi, 1825
    Male+Female adult
    female, 25 mm
    SW-Romania, Banat, Resita - Valiug, 3-8.VI.2007, Krause R. leg.
    Copyright © Ruslan Panin
    Carabus (Tachypus) cancellatus tibiscinus Csiki, 1905
    Carabus (Tachypus) cancellatus tibiscinus Csiki, 1905
    Male+Female adult
    female,28 mm
    SE Hungary, Csongrбd, Makу env., 1-8.VIII.2004, Krause R., Slovak, Pittner leg
    Copyright © Ruslan Panin
    Carabus (Tribax) apschuanus pseudoplatessa Gottwald, 1982
    Carabus (Tribax) apschuanus pseudoplatessa Gottwald, 1982
    Male+Female adult
    NW-Caucasus, Karachay-Cherkessia, Teberda: Dzhamagat gorge, the left bank of the riv. Dzhamagat, 10.VII-28.VIII.2010 S.Pavluchuk leg.
    female, 27 mm
    Copyright © Ruslan Panin
    Carabus (Tribax) apschuanus pseudoplatessa Gottwald, 1982
    Carabus (Tribax) apschuanus pseudoplatessa Gottwald, 1982
    Endophallus
    endophallus
    Copyright © Ruslan Panin
    Carabus (Pachystus) hungaricus mingens Quensel, 1806
    Carabus (Pachystus) hungaricus mingens Quensel, 1806
    Male adult
    male, 31 mm
    SW-Russia: Stavropol Krai, near Stavropol city , Russkiy Les, 11.VI-28.IX.2010, Pavluchuk S. leg.
    Copyright © Ruslan Panin
    Carabus (Archicarabus) montivagus montivagus Palliardi, 1825
    Carabus (Archicarabus) montivagus montivagus Palliardi, 1825
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Archicarabus) montivagus montivagus Palliardi, 1825
    Carabus (Archicarabus) montivagus montivagus Palliardi, 1825
    Male+Female adult
    SW-Romania: Banat, Caras-Severin Reg., Sasca Montana – Stinapari, 21.V.-5.VI.2010, Krause Rene leg.
    female, 26 mm
    Copyright © Ruslan Panin
    Carabus (Archicarabus) montivagus blandus Frivaldsky, 1865
    Carabus (Archicarabus) montivagus blandus Frivaldsky, 1865
    Female adult
    Carabus (Archicarabus) montivagus blandus
    Slovacchia, Plastovce,
    16.7.1995 Budka leg.
    female, 21 mm
    Copyright © Alexander Anichtchenko
    Carabus (Archicarabus) montivagus blandus Frivaldsky, 1865
    Carabus (Archicarabus) montivagus blandus Frivaldsky, 1865
    Female adult
    female, 23 mm
    S-Slovakia, Region de Nitra, Plastovce viil., 1.V.2009, Rudolf Kmeco leg.
    Copyright © Ruslan Panin
    Carabus (Procechenochilus) kokujewi Semenov, 1898
    Carabus (Procechenochilus) kokujewi Semenov, 1898
    Female adult
    RUSSIA, Karachay-Cherkessia, NW Caucasus Mts., Daut Mt. Range, Iskrovtsev - W.Ullu-Kel Mt. pass circus, h=2800-2600m
    Copyright © Alexander Anichtchenko
    Carabus (Procechenochilus) ? kokujewi Semenov, 1898
    Carabus (Procechenochilus) ? kokujewi Semenov, 1898
    Female adult
    Russia, NW Caucasus, Karachay-Cherkessia, Karachaevsk distr., near Uchkula vill., valley Uchkulanichi riv. h=2800-2900m, 25-28.VII.2010, Fominykh D. leg.
    female, 23 mm
    Copyright © Ruslan Panin
    Carabus (Neoplectes) starckianus ganglbaueri Reitter, 1888
    Carabus (Neoplectes) starckianus ganglbaueri Reitter, 1888
    Endophallus

    Copyright © Alexander Anichtchenko
    Carabus (Neoplectes) starckianus ganglbaueri Reitter, 1888
    Carabus (Neoplectes) starckianus ganglbaueri Reitter, 1888
    Male adult
    Russia, NW Caucasus, Krasnodar reg., Adygeia distr., Guzeripl env. 20.5.2003
    male, 26 mm
    Copyright © Alexander Anichtchenko
    Carabus (Neoplectes) starckianus ganglbaueri Reitter, 1888
    Carabus (Neoplectes) starckianus ganglbaueri Reitter, 1888
    Female adult
    NW Caucasus, S Russia, Adygea Rep., Confluence of the river Belaya & Kisha h~700m, 24.V-7.VIII.2010, D.Fominyh leg.
    female,32 mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) violaceus andrzejuscii (as wolffii Dejean, 1826)
    Carabus (Megodontus) violaceus andrzejuscii (as wolffii Dejean, 1826)
    Male+Female adult
    SW-Romania: Banat, Moldova Noua, Ravenska viil., 21.V.-5.VI.2010, Krause Rene leg. female, 32 mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) violaceus andrzejuscii (as mehelyi Ganglbauer, 1896 loc. typ.)
    Carabus (Megodontus) violaceus andrzejuscii (as mehelyi Ganglbauer, 1896 loc. typ.)
    Male+Female adult
    N-Romania, Rodna Mts., Pasul Setref - Sacel 10km E, h=810 m, 8-19.VI.2009, Krause Rene leg. female, 31mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) violaceus andrzejuscii Dejean, 1826 (as pygmaeus Petri, 1912)
    Carabus (Megodontus) violaceus andrzejuscii Dejean, 1826 (as pygmaeus Petri, 1912)
    Male adult
    W Ukraine, NE Carpathians,<br /> Transcarpathian Reg., Rakhov Distr., Chornogora Mt Range h=2030m, alpine zone, 13-24.VI.2010 Panin R. leg./det.<br /> male, 25 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) regalis regalis Fischer, 1822
    Carabus (Morphocarabus) regalis regalis Fischer, 1822
    Male adult
    Carabus (Morphocarabus) regalis regalis Fischer, 1822
    S. Ural, Bashkiria, near Mesjagutovo vill., 09.07.2001 A. Menstschicov leg.
    photo A. Menstschicov
    Copyright © Anton Menstschikov
    Carabus (Morphocarabus) regalis jenissoni Dejean, 1831
    Carabus (Morphocarabus) regalis jenissoni Dejean, 1831
    Male adult
    Carabus (Morphocarabus) regalis jenissoni Dejean, 1831
    Rusia, Tuva, Todzh, Azas lake, 990 m,
    52°24'N 96°33'E, 8.VII.1998, D. Obydov leg.

    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) regalis jenissoni Dejean, 1831
    Carabus (Morphocarabus) regalis jenissoni Dejean, 1831
    Female adult
    Carabus (Morphocarabus) regalis jenissoni Dejean, 1831
    SW Siberia, Altays Mts, Choya distr., Sukhoy Karasuk env.
    23.07.2004, A.Safronov leg.
    Copyright © Safronov
    Carabus (Morphocarabus) regalis eltoni Obydov, 2006
    Carabus (Morphocarabus) regalis eltoni Obydov, 2006 Holotype
    Male adult
    Carabus (Morphocarabus) regalis eltoni Obydov, 2006 HT
    Rusia, Tver distr., near Ilyinskoe vill.,
    29-30.VII.2006, D. Obydov leg.
    photo from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) odoratus septentrionalis Breuning 1932
    Carabus (Morphocarabus) odoratus septentrionalis Breuning 1932
    Female adult
    N. Russia, arctic Urals, Paypudynya Riv., Pompeliz Mt.h=500-700 m.,10-15.07.2002
    S. Andreev, A. Romanenko leg.
    Copyright © Safronov
    Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844: 100
    Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844: 100
    Male adult
    <b>Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844</b><br /> Buriatia, W Hamar-Daban Mts.,<br /> up riv. M.Zangisan, Baishint Ula Mt. 2300m<br /> 23.5.1999 Anichtchenko A. leg.<br /> male, 18 mm
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844: 100
    Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844: 100
    Endophallus
    endophallus
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844: 100
    Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844: 100 Lectotype
    Male adult
    Carabus (Morphocarabus) odoratus odoratus LT<br /> Lectotype<br /> Hamar-Gaban Mts.<br /> photo from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844 (as melleus Lapouge, 1909)
    Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844 (as melleus Lapouge, 1909)
    Male+Female adult
    Russia, S Siberia, East Sayans, Mondy circ., h~2300m., 5-8.VI.2001, R. Mishustin leg. 20mm.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844 (as melleus Lapouge, 1909)
    Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844 (as melleus Lapouge, 1909)
    Endophallus
    <p> endophallus odoratus odoratus (as melleus Lapouge, 1909)<br /> <br />     <br />  </p>
    Copyright © Ruslan Panin
    Carabus (Semnocarabus) rustemi Kabak, 2009
    Carabus (Semnocarabus) rustemi Kabak, 2009 Paratype
    Male adult
    Paratype
    Copyright © Ilya Kabak
    Carabus (Eotribax) valikhanovi bagua Kabak, 2009
    Carabus (Eotribax) valikhanovi bagua Kabak, 2009 Holotype
    Female adult
    China, Xinjiang, N slope of Narat Mt. R., SSW of Tekes, Kokterek basin, Buryl Riv., 3145 m, 28.07.2009,
    42°48'06»N / 81°41'20»E, I. Kabak leg. (ZIN)
    Copyright © Ilya Kabak
    Carabus (Alipaster) thianshanskii shatyensis Deuve et Tian, 2004
    Carabus (Alipaster) thianshanskii shatyensis Deuve et Tian, 2004 Paratype
    Male adult

    Copyright © Ilya Kabak
    Carabus (Alipaster) thianshanskii sarkhobensis Kabak, 2009
    Carabus (Alipaster) thianshanskii sarkhobensis Kabak, 2009 Holotype
    Female adult
    CH, Xinjiang, N sl. of Khalyktau MR., N Kongurbulak R., H=3600 m, 42°35'50»N / 81°02'30»E, 31.07.2003, I. Kabak leg
    (ZIN)
    Copyright © Ilya Kabak
    Carabus (Cechenotribax) petri alipastroides Kabak, 2009
    Carabus (Cechenotribax) petri alipastroides Kabak, 2009 Holotype
    Male adult
    E spur of Terskei Alatau Mt. R., 10 km S of Saryzhas Village, 2800-3000 m, 5.VII.1987, I. Kabak: 42°51’00”N / 79°37’00”E – 42°50’30”N / 79°39’00”E)
    (ZIN)
    Copyright © Ilya Kabak
    Carabus (Tomocarabus) convexus s.str. (as pyrenaicus Born, 1907)
    Carabus (Tomocarabus) convexus s.str. (as pyrenaicus Born, 1907)
    Male adult
    Pirineos, Canfranc, 1300m, Barranco de Izas
    15.6.2005 Anichtchenko A. leg.
    male, 16 mm
    Copyright © Alexander Anichtchenko
    Carabus (Tomocarabus) convexus s.str. (as paganetti Born, 1905)
    Carabus (Tomocarabus) convexus s.str. (as paganetti Born, 1905)
    Male adult
    Lazio, Roma, Bosco di Nettuno
    22.5.1986 C. Iacovone leg.
    male, 20 mm
    Copyright © Alexander Anichtchenko
    Carabus (Tomocarabus) convexus convexus Fabricius, 1775
    Carabus (Tomocarabus) convexus convexus Fabricius, 1775
    Male adult
    W.Ukraine,
    Transcarpathian region,Vinogradov distr., near Malaya Kopanya vill., Chorna Gora Mt., h=180m. 7-18.VII.08. Panin R. leg.
    male 18 mm
    Copyright © Ruslan Panin
    Carabus (Tomocarabus) convexus s.str. (ab. niger)
    Carabus (Tomocarabus) convexus s.str. (ab. niger)
    Male adult
    W Ukraine, NW Podolia, Lviv Region, Zolochiv distr., near Gologirky vill., in steppe 1-19.V.2007 Panin R. leg.
    16mm
    Copyright © Ruslan Panin
    Carabus (Tomocarabus) convexus erzeliki Schweiger, 1962
    Carabus (Tomocarabus) convexus erzeliki Schweiger, 1962
    Male adult
    Turkey, Sivas 1560 m. 19.7.1991
    male, 19 mm
    Copyright © Alexander Anichtchenko
    Carabus (Tomocarabus) bessarabicus bessarabicus Fischer, 1823
    Carabus (Tomocarabus) bessarabicus bessarabicus Fischer, 1823
    Male+Female adult
    UKRAINE,
    Kherson Reg., Golaya Pristan distr. near Rybalche vill. 6.IV.2000
    long (female) 20mm
    Copyright © Ruslan Panin
    Carabus (Tachypus) cristoforii Spence, 1823 (as nicolasi Reitter, 1888)
    Carabus (Tachypus) cristoforii Spence, 1823 (as nicolasi Reitter, 1888)
    Male adult
    S-France, Hautes-Pyrenees, Pic de Nere h=2200m,15.VII.1999,
    male, 14mm
    Copyright © Ruslan Panin
    Carabus (Piocarabus) sui Imura & Zhou, 1998
    Carabus (Piocarabus) sui Imura & Zhou, 1998
    Female adult
    China, Beijing, Yunmengshan,2005, Liu ye coll.;<br /> 35mm
    Copyright © Ye Liu
    Carabus (Teratocarabus) azrael azrael Semenov & Znojko, 1932
    Carabus (Teratocarabus) azrael azrael Semenov & Znojko, 1932
    Female adult

    Copyright © Alexander Anichtchenko
    Carabus (Teratocarabus) azrael mizunumaianus Imura, 1991
    Carabus (Teratocarabus) azrael mizunumaianus Imura, 1991
    Male adult
    China, Liaoning, Benxi, Gongchangung
    20.VI.2004 Hong
    Copyright © Boleslav Brezina
    Carabus (Tanaocarabus) sylvosus Say, 1825
    Carabus (Tanaocarabus) sylvosus Say, 1825
    Male adult
    Canada
    Copyright Henri Goulet, Agriculture & Agri-Food Canada
    Copyright © Henri Goulet
    Carabus (Tanaocarabus) sylvosus Say, 1825
    Carabus (Tanaocarabus) sylvosus Say, 1825
    Female adult
    female, 26mm
    Copyright © Boleslav Brezina
    Carabus (Tanaocarabus) sylvosus Say, 1825
    Carabus (Tanaocarabus) sylvosus Say, 1825
    Male adult
    male, 25mm
    Copyright © Boleslav Brezina
    Carabus (Tanaocarabus) finitimus Haldeman, 1852
    Carabus (Tanaocarabus) finitimus Haldeman, 1852
    Female adult

    Copyright © Boleslav Brezina
    Carabus (Tanaocarabus) hendrichsi Bolivar y Pieltain, Rotger & Coronado, 1967
    Carabus (Tanaocarabus) hendrichsi Bolivar y Pieltain, Rotger & Coronado, 1967
    Male adult
    male, 23mm
    Copyright © Boleslav Brezina
    Carabus (Tanaocarabus) forreri Bates, 1882
    Carabus (Tanaocarabus) forreri Bates, 1882
    Male adult
    Mexico, Durango, Sierra Madre Occ.,
    Buenos Aires w., 2700 m, La Ciudad, 14.6.90 Heinz leg.
    Copyright © Alexander Anichtchenko
    Carabus (Tanaocarabus) forreri forreri Bates, 1882
    Carabus (Tanaocarabus) forreri forreri Bates, 1882
    Female adult
    female, 23mm
    Copyright © Boleslav Brezina
    Carabus (Tanaocarabus) forreri forreri Bates, 1882
    Carabus (Tanaocarabus) forreri forreri Bates, 1882
    Male adult
    male, 22mm
    Copyright © Boleslav Brezina
    Carabus (Tachypus) cancellatus fusus Palliardi, 1825
    Carabus (Tachypus) cancellatus fusus Palliardi, 1825
    Male+Female adult
    France, Riveres Charente, leg. Laforgue VI.1996
    female, 24mm
    Copyright © Ruslan Panin
    Carabus (Tachypus) cancellatus s.str. (as sulinensis Born, 1902)
    Carabus (Tachypus) cancellatus s.str. (as sulinensis Born, 1902)
    Male+Female adult
    S UKRAINE,
    Krym Peninsula, Staryi Krym, 17.V.2002
    L= 26 mm.
    Copyright © Ruslan Panin
    Carabus (Tachypus) cancellatus s.str. (as conspersus Lapouge, 1902)
    Carabus (Tachypus) cancellatus s.str. (as conspersus Lapouge, 1902)
    Male+Female adult
    E Byelorussia,
    Mogilev Reg. Bykhov distr., near Voronino vill., 20.V-3.VI.2007, Komyagunova N. leg.
    female, 26mm
    Copyright © Ruslan Panin
    Carabus (Tachypus) cancellatus s.str. (as tuberculatus Dejean, 1826)
    Carabus (Tachypus) cancellatus s.str. (as tuberculatus Dejean, 1826)
    Male adult
    W Ukraine,NW Podolian,
    Ternopil reg., Pidvolochisk distr., forest Bogdanivskiy, near Romanove Selo, 7-24.V.08, R.Yu.Panin & I.A.Solodovnikov leg.
    male, 23mm
    Copyright © Ruslan Panin
    Carabus (Tachypus) cancellatus s.str. (as scythicus Schaum, 1857)
    Carabus (Tachypus) cancellatus s.str. (as scythicus Schaum, 1857)
    Male adult
    Ukrainian Carpathians,
    Transcarpathian reg., Rakhov Distr., Chornogora Mnt.Range h=1900m. Panin R. leg. 28.VI-9.VII.08
    male, 24mm.
    Copyright © Ruslan Panin
    Carabus (Tachypus) cancellatus s.str. (as scythicus Schaum, 1857)
    Carabus (Tachypus) cancellatus s.str. (as scythicus Schaum, 1857)
    Female adult
    Ukrainian Carpathians,
    Transcarpathian reg., Rakhov Distr., Chornohora range h=1850m, 29.VI-28.VII.2007 Panin R. leg.
    female, 27 mm.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) odoratus martjanovianus Obydov 1999 HT
    Carabus (Morphocarabus) odoratus martjanovianus Obydov 1999 HT Holotype
    Male adult
    Carabus (Morphocarabus) odoratus martjanovianus Obydov 1999
    Rusia, Sayan Mts., near Sayanogorsk, 7.VII.1993
    photo from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) odoratus magadanicus Obydov 1999 PT
    Carabus (Morphocarabus) odoratus magadanicus Obydov 1999 PT Paratype
    Male adult
    Carabus (Morphocarabus) odoratus magadanicus Obydov 1999 PT
    Rusia, Far East, E of Magadan, W Nyuklia vill.
    VII.1997, Yu. Marusik leg.
    photo from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) odoratus krugeri Obydov 1999 PT
    Carabus (Morphocarabus) odoratus krugeri Obydov 1999 PT Paratype
    Female adult
    Carabus (Morphocarabus) odoratus krugeri Obydov 1999 PT
    Rusia, Tuva, Akademika Obrucheva Mt.rng., 2100, 51°56'N 095°32'E,
    Koptu pass, tundra 2.VII.1998, D. Obydov leg.
    photo from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) odoratus kamtchatensis Breuning 1942
    Carabus (Morphocarabus) odoratus kamtchatensis Breuning 1942
    Male adult
    Carabus (Morphocarabus) odoratus kamtchatensis Breuning 1942
    Rusia, Kamchatka peninsula, VII.21
    photo from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) odoratus dohrni Gebler, 1847
    Carabus (Morphocarabus) odoratus dohrni Gebler, 1847
    Male adult
    Carabus (Morphocarabus) odoratus dohrni Gebler, 1847
    Rusia, Kuznetskii Alatau, Taizhe-Su riv., 53°55'N 89°10'E,
    1250-1650 m, 25.VI-30.VII.2002, V. Murzin and V. Sinyaev leg.
    photo from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) odoratus dabanensis Shilenkov 1996 loc.typ.
    Carabus (Morphocarabus) odoratus dabanensis Shilenkov 1996 loc.typ.
    Male+Female adult
    Carabus (Morphocarabus) odoratus dabanensis Shilenkov 1996 loc.typ.
    Buryatia, Hamar-Daban mt. rng., Snezhnaya riv.,
    4.6.2007 Anichtchenko A. leg.
    male 17 mm, female 19 mm
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) odoratus czernyscheviellus Obydov, 2002 PT
    Carabus (Morphocarabus) odoratus czernyscheviellus Obydov, 2002 PT Paratype
    Female adult
    Carabus (Morphocarabus) odoratus czernyscheviellus PT
    Rusia, SE Transbaikalia, Chita reg., Nerchinsko-Kugenskii Mt.rng.,
    near Vershino-Darasunskii vill., Bogatui Mt. 1221 m.,
    15.VII.2002, D. Obydov leg.
    photo from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) odoratus bargusinicus Shilenkov, 1996
    Carabus (Morphocarabus) odoratus bargusinicus Shilenkov, 1996
    Male+Female adult


        Carabus (Morphocarabus) odoratus bargusinicus Shilenkov, 1996
        Russia, N Burajatia,
        Barguzinskiy Mts., the upper reaches of M.Chiverkuy riv., h=2000m, Vashchenko S. leg.
        female, 21mm/male,19mm
        photo Panin R.



    Copyright © Ruslan Panin
    Carabus (Morphocarabus) odoratus bargusinicus Shilenkov, 1996
    Carabus (Morphocarabus) odoratus bargusinicus Shilenkov, 1996
    Endophallus


        endophallus



    Copyright © Ruslan Panin
    Carabus (Morphocarabus) odoratus antropovi Shilenkov 1996
    Carabus (Morphocarabus) odoratus antropovi Shilenkov 1996
    Male+Female adult
    Russia, S Tuva, Khorumnug -Tayga mts. Range, h=1600-2300m, Buren riv. valley, 4-26.VI.2000 Vashchenko S & A. leg<br /> female, 20mm/male,18mm<br /> photo Panin R.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) odoratus ?antropovi Shilenkov 1996
    Carabus (Morphocarabus) odoratus ?antropovi Shilenkov 1996
    Male adult
    S Tuva, Horumnut-Taiga Mt.rng., Bureya riv., 2000 m, tundra, Vaschenko S. leg. 18 mm
    Copyright © Alexander Anichtchenko
    Carabus (Tachypus) auratus auratus Linne, 1761
    Carabus (Tachypus) auratus auratus Linne, 1761
    Male adult
    Canada
    Copyright Henri Goulet, Agriculture & Agri-Food Canada
    Copyright © Henri Goulet
    Carabus (Tachypus) auratus s.str. (as brunieri Barthe, 1921)
    Carabus (Tachypus) auratus s.str. (as brunieri Barthe, 1921)
    Male adult
    SC-FRANCE, Aude, Rennes les Bains, 6.VII.1989, Sunard A. leg.
    male, 26mm
    Copyright © Ruslan Panin
    Carabus (Tachypus) auratus auratus Linne, 1761
    Carabus (Tachypus) auratus auratus Linne, 1761
    Female adult
    30 mm
    Germania, Oberfranken, Pegnitz
    leg.det. U.Schmidt, 1972
    Foto: U.Schmidt, 2008
    Copyright © U.Schmidt
    Carabus (Tachypus) auratus lasserrei Doue, 1855
    Carabus (Tachypus) auratus lasserrei Doue, 1855
    Male adult
    SC-FRANCE, Midi-Pyrуnуes reg. Tarn, Saint Paul cap de Joux, 24.V.2003, A. Laforgue leg.
    male, 24mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) monilis Fabricius, 1792
    Carabus (Morphocarabus) monilis Fabricius, 1792
    Male adult
    France 39, Foncina le Haut 300m, 23.6.1979; 29 mm
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) monilis Fabricius, 1792
    Carabus (Morphocarabus) monilis Fabricius, 1792
    Endophallus
    France 39, Foncina le Haut 300m
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) monilis Fabricius, 1792 (as taunicus)
    Carabus (Morphocarabus) monilis Fabricius, 1792 (as taunicus)
    Male adult
    W Germany, Vogelsberg Mt., 700m, 13.5.1985
    male, 24 mm
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) monilis Fabricius, 1792 (as interpositus Géhin, 1880)
    Carabus (Morphocarabus) monilis Fabricius, 1792 (as interpositus Géhin, 1880)
    Male adult
    NW France,
    Basse-Normandie, Orne, Forкt d'Йcouves, V.2009, Pascal Stefani leg.
    male, 29mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) monilis Fabricius, 1792 (as interpositus Géhin, 1880)
    Carabus (Morphocarabus) monilis Fabricius, 1792 (as interpositus Géhin, 1880)
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Morphocarabus) monilis (as interpositus Gehin, 1880)
    Carabus (Morphocarabus) monilis (as interpositus Gehin, 1880)
    Male+Female adult
    NW France,
    Haute-Normandie reg., Bois Les Loges, Etretat, Seine-Maritime,10.V.1988
    female, 28mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) monilis (as amoenus Baudet-Lafarge, 1836)
    Carabus (Morphocarabus) monilis (as amoenus Baudet-Lafarge, 1836)
    Female adult
    C France,
    Puy-de-Dome, Col de la Croix Saint Robert h=1300m, VII.2008, Pascal Stefani leg.
    female, 24 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) monilis (as helveticus Heer, 1837 Holotype)
    Carabus (Morphocarabus) monilis (as helveticus Heer, 1837 Holotype) Holotype
    Male adult
    Carabus (Morphocarabus) monilis (as helveticus Heer, 1837 Holotype)
    ?hybrid of monilis/cancellatus
    Museum of Zurich
    photo Anichtchenko
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) monilis (as dupeuxi Deuve, 1974)
    Carabus (Morphocarabus) monilis (as dupeuxi Deuve, 1974)
    Male+Female adult

    France,
    Charente, Foret de Braconne VII.1998, Laforgue leg.
    famale, 27mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) michailovi Kabak, 1992
    Carabus (Morphocarabus) michailovi Kabak, 1992
    Male adult
    NE Kazakhstan, S Altay Mts., Sarymsakty mt. rng.,
    Sarymsakty riv., 2400 m, 22.6.2000 Nekroshevich S. leg.
    male, 18 mm
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) michailovi Kabak, 1992
    Carabus (Morphocarabus) michailovi Kabak, 1992
    Male adult
    NE Kazakhstan, S Altay Mts., Sarymsakty mt. rng.,
    Sarymsakty riv., 2400 m, 22.6.2000 Nekroshevich S. leg.
    male, 17 mm
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) michailovi Kabak, 1992
    Carabus (Morphocarabus) michailovi Kabak, 1992
    Male adult
    NE-Kazakhstan,
    S Altai Mts.,Sarym-Sakty Range, Sarym-Sakty riv., h=2400m. 21.VI.1999., A.Klimenko leg.
    male,18mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) mestscherjakovi telezkianus Shilenkov, 1996
    Carabus (Morphocarabus) mestscherjakovi telezkianus Shilenkov, 1996
    Male adult
    Carabus (Morphocarabus) mestscherjakovi telezkianus Shilenkov, 1996
    Rusia, Altai, S shore of Teletskoe lake, 5.VII.1968
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) mestscherjakovi telezkianus Shilenkov, 1996
    Carabus (Morphocarabus) mestscherjakovi telezkianus Shilenkov, 1996
    Male adult
    Carabus (Morphocarabus) mestscherjakovi telezkianus Shilenkov, 1996
    SW Khakasia, Abakanskii Mt.rng., Choochek Mt., 20 km SSE Mrassu vill.,
    1800m, 7-19.7.1999 Lomakin D. leg.
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
    Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
    Male adult
    W Sayan mts., Kopen mt., Dzhabash riv., 75 km
    S from Arbaty vill., 2235m, 26-30.6.2003 Brinev A. leg.
    long. 17 mm
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
    Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
    Male adult
    Russia, S Siberia,
    S Khakassia, N Tuva reg., Sailyg-Khem-Taiga mts. range h=2400m, 10.VII.2004
    male, 18mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
    Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
    Male+Female adult
    Russia, S Siberia,
    S Khakassia, N Tuva reg., Sailyg-Khem-Taiga mts. range h=2000-2600m,10-30.VI.2005, S.Vashchenko & S.Alexeenko leg.
    male,17.5mm/female,19mm
    photo Panin R.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
    Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
    Endophallus
    endophallus
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) mestscherjakovi pseudoodoratus Brinev, 2002 Paratype
    Carabus (Morphocarabus) mestscherjakovi pseudoodoratus Brinev, 2002 Paratype Paratype
    Male adult
    Krasnojarsk reg., Aradan distr., 5 km SW Aradan vill.,<br /> Us riv., forest, 52є32`N 93є27`E 9-12.VII.1998 A.Brinev leg<br /> male, 18 mm
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) mestscherjakovi pseudoodoratus Brinev, 2002 Paratype
    Carabus (Morphocarabus) mestscherjakovi pseudoodoratus Brinev, 2002 Paratype Paratype
    Female adult
    Krasnojarsk reg., Aradan distr., 5 km SW Aradan vill., vall. of Us riv., A.Brinev leg.<br /> female, 20 mm<br /> photo & coll. Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
    Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
    Male adult
    Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
    W Sayan, Olenya rechka
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
    Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
    Male adult
    Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
    W Sayan, Kulumys Mt.R., env. Oiskoje Lake, mountain tundra, 12-14. 07.1999 A.Semin leg.
    male
    L=17,0 mm
    Copyright © Maksim Smirnov
    Carabus (Morphocarabus) kozhantschikovi martjanovi Lutshnik, 1924
    Carabus (Morphocarabus) kozhantschikovi martjanovi Lutshnik, 1924
    Female adult
    Khakassia, 1800 m, near Cheremushki vill., 12.6.-20.7.2000
    female, 21 mm
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
    Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
    Male adult
    W Sayan Mts., Kazhyr-Suk riv., 80 km W Oleniya
    Rechka, W part of Oiskii mt. rng., 30.6.-7.7.1999
    2100 m, Brinev A. leg.
    male, 19 mm
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
    Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
    Male adult
    W Sayan, low fl. Kazyr-Suk riv., 80km W meteostation Olenia Rechka, W part of Oiskii mt.rng., 30.VI-7.VII.1999 2100m, Brinev A. leg.
    male, 20mm
    det. Brinev A.
    photo Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
    Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
    Endophallus

    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 18789 (as viridipennis Kraatz, 1886, nec Lewis,
    Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 18789 (as viridipennis Kraatz, 1886, nec Lewis,
    Male adult
    E Rusia, Khabarovsk reg.,
    near Birobidzhan VI.1991
    male, 26 mm
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
    Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
    Male adult
    Rusia, S Primorie, Vladivostok distr., Razdolnaya riv., near Kluchevoi vill., 10-14.VII.1988, S. Muhanov leg.; Obydov det. as tristiculus Kraatz, 1879
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) hummeli cf pusongensis Imura, 1993
    Carabus (Morphocarabus) hummeli cf pusongensis Imura, 1993
    Male adult
    China, Beijing, Wulingshan mountain, 1985
    Copyright © Ye Liu
    Carabus (Morphocarabus) hummeli ssp.
    Carabus (Morphocarabus) hummeli ssp.
    Male adult
    Carabus (Morphocarabus) hummeli ssp.
    China, East Inner Mongolia, Chifen, 2006, Shi H.L. leg.
    Photo & det. by liuye
    Copyright © Ye Liu
    Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
    Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
    Male adult
    China, Jinling pro.;
    Photo by liuye
    det. Anichtchenko A.
    Copyright © Ye Liu
    Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
    Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
    Male adult
    Russia, Far East,
    Primorskiy Kray, near Kamenushka vill., 2-3.VIII.1992, K.E.Dovgaylo leg.
    male, 27mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) hummeli pusongensis Imura, 1993
    Carabus (Morphocarabus) hummeli pusongensis Imura, 1993
    Male adult
    S Primorie, Chernye gory, Poima riv.,
    30.7-5.8.1999 Anichtchenko A. leg.
    male, 26 mm
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) hummeli nevelskii Shilenkov, 1996
    Carabus (Morphocarabus) hummeli nevelskii Shilenkov, 1996
    Female adult
    Rusia, S Sakhalin isl., VII.1993, G. Vasiliev leg.

    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) hummeli nevelskii Shilenkov, 1996
    Carabus (Morphocarabus) hummeli nevelskii Shilenkov, 1996
    Male+Female adult
    Russia, Sakhalin isl.,
    Makarov distr., Lesnaja riv., VII.1992, M. Nesterov leg.
    female,23mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) hummeli hummeli Fischer, 1823
    Carabus (Morphocarabus) hummeli hummeli Fischer, 1823
    Male adult
    Carabus (Morphocarabus) hummeli hummeli Fischer, 1823
    Rusia, Chita reg., Darasun, VII.1991
    I. Avdeev leg.
    male, 23 mm
    Copyright © Alexander Anichtchenko
    Carabus (Sphodristocarabus) varians varians Fischer, 1827
    Carabus (Sphodristocarabus) varians varians Fischer, 1827
    Male+Female adult
    female: Transcaucasia, Armenia, Amasia distr., Karakhachskiy mountain pass h=2270m, 3.VIII.1985 P.V.Kazaryan leg., 22mm<br /> male: Armenia, Amasia distr., Arpalach vill. VI.1999, P.V. Kazaryan leg.
    Copyright © Ruslan Panin
    Carabus (Sphodristocarabus) varians rugatus Breuning, 1934
    Carabus (Sphodristocarabus) varians rugatus Breuning, 1934
    Female adult

    Copyright © Ruslan Panin
    Carabus (Sphodristocarabus) blumenthaliensis basilewskianus Breuning & Ruspoli, 1970
    Carabus (Sphodristocarabus) blumenthaliensis basilewskianus Breuning & Ruspoli, 1970
    Male adult
    SW Georgia, Adzharia 10 km Baku 2000m
    18.6.98
    25 mm
    Copyright © Alexander Anichtchenko
    Carabus (Sphodristocarabus) blumenthaliensis basilewskianus Breuning & Ruspoli, 1970
    Carabus (Sphodristocarabus) blumenthaliensis basilewskianus Breuning & Ruspoli, 1970
    Male+Female adult
    NE-Turkey,
    Artvin Prov., Hopa vers Borcka, Cankutaran Gecidi h=800m, 17.VI.2007 leg. D. Prunier
    female, 24mm
    Copyright © Ruslan Panin
    Carabus (Sphodristocarabus) varians janthinus s.str.
    Carabus (Sphodristocarabus) varians janthinus s.str.
    Male+Female adult
    NW-Caucasus, Russia, Krasnodar reg.,Tuapse distr., Mt.Semashkho 04.V.2008, D.Fominyh leg.<br /> female, 24mm
    Copyright © Ruslan Panin
    Carabus (Sphodristocarabus) varians armeniacus Mannerheim, 1830
    Carabus (Sphodristocarabus) varians armeniacus Mannerheim, 1830
    Female adult
    NE Turkey,<br /> Artvin Prov., Borçka, Macahel Gecidi h=1450m, 18.VI.2007, D.Prunier leg.<br /> female, 26mm
    Copyright © Ruslan Panin
    Carabus (Sphodristocarabus) tokatensis almushensis Mandl, 1975
    Carabus (Sphodristocarabus) tokatensis almushensis Mandl, 1975
    Male adult
    Turkey, Sivas, 20 km N Zara, N Arapca, 1750m; 11.VII.1993
    Copyright © Alexander Anichtchenko
    Carabus (Sphodristocarabus) scovitzi czipkai Heinz, 1980
    Carabus (Sphodristocarabus) scovitzi czipkai Heinz, 1980
    Male adult
    Turkey, Artvin, Kafkasor, 1600 m
    4.7.1996 Staven and Skoupy leg.
    male, 22 mm
    Copyright © Alexander Anichtchenko
    Carabus (Sphodristocarabus) macrogonus simonianus Breuning & Ruspoli, 1970
    Carabus (Sphodristocarabus) macrogonus simonianus Breuning & Ruspoli, 1970
    Female adult
    Turkey, Ordu, Golkoy, 1220 m, 19.7.91
    Copyright © Alexander Anichtchenko
    Carabus (Sphodristocarabus) macrogonus resplendens Breuning, 1934
    Carabus (Sphodristocarabus) macrogonus resplendens Breuning, 1934
    Female adult
    Turkey; Cayirkent Samsun 12.VII.1996
    Copyright © Alexander Anichtchenko
    Carabus (Sphodristocarabus) georgia Cavazzuti, 1984
    Carabus (Sphodristocarabus) georgia Cavazzuti, 1984
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Sphodristocarabus) bohemani bohemani Menetries, 1832
    Carabus (Sphodristocarabus) bohemani bohemani Menetries, 1832
    Female adult
    S Azerbaydzan, Lenkoran distr., Vazha riv.,
    2 km NW Dashdatuk vill. 38º40*09N 48º42*06E
    17.4.2004 Makarov K. leg.
    female, 22 mm
    Copyright © Alexander Anichtchenko
    Carabus (Sphodristocarabus) bohemani bohemani Menetries, 1832
    Carabus (Sphodristocarabus) bohemani bohemani Menetries, 1832
    Male adult
    Azerbaijan,
    Talysh Mts., Polikesh, h~1800m.,2-6.VI.2001 Dolin V.G. leg.
    male, 20mm.
    Copyright © Ruslan Panin
    Carabus (Sphodristocarabus) hollbergi Mannerheim, 1827
    Carabus (Sphodristocarabus) hollbergi Mannerheim, 1827
    Male adult
    N Azerbaijan,
    4km W Ismayilli, h=700m, 9.VII.2008, Pljushch I.G. leg.
    male,22mm
    Copyright © Ruslan Panin
    Carabus (Semnocarabus) transiliensis Semenov, 1896
    Carabus (Semnocarabus) transiliensis Semenov, 1896
    Male adult
    Kirgizstan, S slope Kungei Ala Too
    Oi-Tal riv. 3000m 14.7.1999
    13 mm
    Copyright © Alexander Anichtchenko
    Carabus (Semnocarabus) transiliensis Semenov, 1896
    Carabus (Semnocarabus) transiliensis Semenov, 1896
    Male+Female adult
    E KAZAKHSTAN,
    Bolshoe Almaatinskoe Lake,30.VI.1992
    15mm
    Copyright © Ruslan Panin
    Carabus (Semnocarabus) transiliensis Semenov, 1896
    Carabus (Semnocarabus) transiliensis Semenov, 1896
    Male adult
    SE-KAZAKHSTAN,
    Ketmen Mts.,N-slopes h=2250-2500m.1-5.VI.1990
    16mm
    Copyright © Ruslan Panin
    Carabus (Semnocarabus) regulus regulus Dohrn, 1882
    Carabus (Semnocarabus) regulus regulus Dohrn, 1882
    Male adult
    photo M. Smirnov from www.zin.ru
    Copyright © Maksim Smirnov
    Carabus (Semnocarabus) regulus regulus Dohrn, 1882
    Carabus (Semnocarabus) regulus regulus Dohrn, 1882
    Male adult
    E Kirgizia, Kaindy-Ketta mts. rng. Sary-Dzhas riv.
    Maidadyr near Enelchek vill. 6-8.7.1999 Klimenko leg.
    14 mm
    Copyright © Alexander Anichtchenko
    Carabus (Semnocarabus) erosus erosus Motschulsky, 1865
    Carabus (Semnocarabus) erosus erosus Motschulsky, 1865
    Male adult
    Kirgizia, S Kungei Ala-Too, Kichi-Yuryukty
    2800m 1.7.1999 Subankulov leg.
    male, 21 mm
    Copyright © Alexander Anichtchenko
    Carabus (Semnocarabus) erosus corax Semenov, 1896
    Carabus (Semnocarabus) erosus corax Semenov, 1896
    Male adult
    SE Kazakhstan, Ili riv., Bakanas vill.,
    1-5.5.1999 Klimenko A. leg.
    Copyright © Alexander Anichtchenko
    Carabus (Semnocarabus) carbonicolor carbonicolor Morawitz, 1886
    Carabus (Semnocarabus) carbonicolor carbonicolor Morawitz, 1886
    Male adult
    E Kirgizia, Kaindy-Ketta mts. rng., Sary-Dzas riv.,
    Maidadyr near Enelchek vill., 6-8.7.99 Klimenko leg.
    Copyright © Alexander Anichtchenko
    Carabus (Semnocarabus) carbonicolor callosus Semenov, 1896
    Carabus (Semnocarabus) carbonicolor callosus Semenov, 1896
    Male adult
    Kazakhstan, Bol. Alma-Atinka riv. 1600m 1.6.1999 Klimenko A. leg.
    male, 19 mm
    Copyright © Alexander Anichtchenko
    Carabus (Scambocarabus) sculptipennis Chaudoir, 1887
    Carabus (Scambocarabus) sculptipennis Chaudoir, 1887
    Male adult
    China, Beijing, Xiaolongmeng, 2005, Liu ye coll.;
    22mm; photo by Liuye
    Copyright © Ye Liu
    Carabus (Scambocarabus) kruberi kruberi Fischer, 1822
    Carabus (Scambocarabus) kruberi kruberi Fischer, 1822
    Male adult
    Rusia, Chita reg., 23 km SW Borzia, Borzinskoe lake,
    1.VI.2002 Anichtchenko A. leg.
    Copyright © Alexander Anichtchenko
    Carabus (Scambocarabus) kruberi Fischer, 1822
    Carabus (Scambocarabus) kruberi Fischer, 1822
    Male adult
    China, Beijing, labagoumeng, 2004, Li fei coll.;
    21mm; photo by Liuye
    Copyright © Ye Liu
    Carabus (Morphocarabus) hummeli biamensis Obydov, 2007 Paratypus
    Carabus (Morphocarabus) hummeli biamensis Obydov, 2007 Paratypus Paratype
    Male adult
    Russia, North Primorie, B. Svetlovodnaya riv. (Biame riv.), 40 km from source, 30.07.1976, A. Klitin leg.
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
    Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
    Female adult
    E.RUSSIA, Primorsky ter., Chuguevsky distr., 30 km ESE from Chuguevka, 9-14.VIII.2007
    Copyright © Nil Belonogov
    Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
    Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
    Male adult
    E.RUSSIA, Primorsky ter., Chuguevsky distr., 30 km ESE from Chuguevka, 9-14.VIII.2007
    Copyright © Nil Belonogov
    Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 18789 (as viridipennis Kraatz, 1886, nec Lewis,
    Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 18789 (as viridipennis Kraatz, 1886, nec Lewis,
    Female adult
    Primorskii reg., near Kovalerovo
    VII.1990 Evstigneev A. leg.
    female, 25 mm
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) hummeli ? middendorfi Menetries, 1851
    Carabus (Morphocarabus) hummeli ? middendorfi Menetries, 1851
    Male adult
    Carabus (Morphocarabus) hummeli ? middendorfi Menetries, 1851
    Rusia, Khabarovsk reg., Ohotsk sea, Port Ayan, 08.24
    photo from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) henningi pelevini Obydov 2006
    Carabus (Morphocarabus) henningi pelevini Obydov 2006
    Female adult
    NE Kazakhstan, S Altai mt.rng., Sarymskiy mt. rng., Sarymsakty riv.,<br /> 2400m, 22.VI.2000 Nekroshevich S. leg. female 17mm det. Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) henningi pelevini Obydov 2006 HT
    Carabus (Morphocarabus) henningi pelevini Obydov 2006 HT
    Male adult
    Carabus (Morphocarabus) henningi pelevini Obydov 2006 HT
    Kazakhstan, S Altai, Sarym-Sakty Mt.rng., 3000-3400 m,
    18-20.VI.1998, A. Klimenko leg.
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) henningi peczoricus Obydov 2000 HT
    Carabus (Morphocarabus) henningi peczoricus Obydov 2000 HT Holotype
    Male adult
    Carabus (Morphocarabus) henningi peczoricus Obydov 2000 HT
    Rusia, Komi republic, Kosiyu River mouth, (Pechora basin) 21.VII.1980
    photo from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) henningi oviformis Beheim & Breuning, 1943
    Carabus (Morphocarabus) henningi oviformis Beheim & Breuning, 1943
    Male adult
    Carabus (Morphocarabus) henningi oviformis Beheim & Breuning, 1943
    Rusia, Krasnoyarsk reg., Taimyr AO, near Norilsk,
    07.2004, M. Shestopalov leg.
    photo from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) henningi melyachi Shilenkov & Menstshikov, 1996
    Carabus (Morphocarabus) henningi melyachi Shilenkov & Menstshikov, 1996
    Male adult
    Carabus (Morphocarabus) henningi melyachi Shilenkov & Menstshikov, 1996
    C Altai mts, Seminskii mt.rng., Katun riv., 3km S Ust-Sema vill.
    4.VII.2002 A. & N. Menschikovy
    male, 17mm
    det. Menschikov A.
    photo Anichtchenko
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) henningi henningi Fischer, 1817
    Carabus (Morphocarabus) henningi henningi Fischer, 1817
    Male adult
    S. Ural, Bashkiria, near Mesjagutovo vill., 09.07.2001 A. Menstschicov leg.
    Copyright © Anton Menstschikov
    Carabus (Morphocarabus) henningi henningi Fischer, 1817
    Carabus (Morphocarabus) henningi henningi Fischer, 1817
    Male adult
    SE Siberia, Russia, Irkutsk reg., near Pivovarikha vill., 7.V.07, D. Fominykh leg. female, 21mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) henningi hamarensis Shilenkov, 1996
    Carabus (Morphocarabus) henningi hamarensis Shilenkov, 1996
    Female adult
    S Siberia, Hamar-Daban mts., Snezhnaya riv., 1-6.VI.2007, Anichtchenko A. leg. & det., 20 mm
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) henningi hamarensis Shilenkov, 1996
    Carabus (Morphocarabus) henningi hamarensis Shilenkov, 1996
    Female adult
    Carabus (Morphocarabus) henningi hamarensis Shilenkov, 1996 loc.typ.
    Copyright © Alexander Anichtchenko
    Carabus (Rhigocarabus) xiei xiei Deuve, 1992
    Carabus (Rhigocarabus) xiei xiei Deuve, 1992
    Male adult
    male, 18mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) xiei xiei Deuve, 1992
    Carabus (Rhigocarabus) xiei xiei Deuve, 1992
    Female adult
    female, 19mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) xiei xiei Deuve, 1992
    Carabus (Rhigocarabus) xiei xiei Deuve, 1992
    Male adult
    male, 20mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) xiei xiei Deuve, 1992
    Carabus (Rhigocarabus) xiei xiei Deuve, 1992
    Male adult
    male, 17mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) xiei sanggarparicus Deuve, 1996
    Carabus (Rhigocarabus) xiei sanggarparicus Deuve, 1996 Holotype Paratype
    Male adult
    Holotype
    male, 19mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) xiei sanggarparicus Deuve, 1996
    Carabus (Rhigocarabus) xiei sanggarparicus Deuve, 1996
    Female adult
    Paratype
    female, 20mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) xiei barkamensis Deuve, 1994
    Carabus (Rhigocarabus) xiei barkamensis Deuve, 1994 Holotype Paratype
    Male adult
    Holotype, male, 19mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) xiei barkamensis Deuve, 1994
    Carabus (Rhigocarabus) xiei barkamensis Deuve, 1994
    Male adult
    Paratype, female, 19mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) xiangchengicus qingmaicus Deuve, 2007
    Carabus (Rhigocarabus) xiangchengicus qingmaicus Deuve, 2007
    Male adult
    male, 15mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) xiangchengicus qingmaicus Deuve, 2007
    Carabus (Rhigocarabus) xiangchengicus qingmaicus Deuve, 2007
    Female adult
    female, 18mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) xiangchengicus qingmaicus Deuve, 2007
    Carabus (Rhigocarabus) xiangchengicus qingmaicus Deuve, 2007
    Female adult
    female, 16mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) xiangchengicus sangpiensis Deuve, 2007
    Carabus (Rhigocarabus) xiangchengicus sangpiensis Deuve, 2007 Paratype
    Male adult
    male, 15 mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) thais thais Heinz, 1997
    Carabus (Rhigocarabus) thais thais Heinz, 1997
    Male adult
    male, 16mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) thais thais Heinz, 1997
    Carabus (Rhigocarabus) thais thais Heinz, 1997
    Male adult
    male, 19mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) thais thais Heinz, 1997
    Carabus (Rhigocarabus) thais thais Heinz, 1997
    Male adult
    male, 20mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) subindigestus subindigestus Deuve, 1994
    Carabus (Rhigocarabus) subindigestus subindigestus Deuve, 1994
    Female adult
    female, 15mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) subindigestus subindigestus Deuve, 1994
    Carabus (Rhigocarabus) subindigestus subindigestus Deuve, 1994
    Male adult
    male, 16mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) subindigestus subindigestus Deuve, 1994
    Carabus (Rhigocarabus) subindigestus subindigestus Deuve, 1994
    Male adult
    male, 15mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) subindigestus subindigestus Deuve, 1994
    Carabus (Rhigocarabus) subindigestus subindigestus Deuve, 1994
    Male adult
    male, 16mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) sehnali Brezina & Hackel, 2006
    Carabus (Rhigocarabus) sehnali Brezina & Hackel, 2006 Holotype Paratype
    Male adult
    male, 20mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) sehnali Brezina & Hackel, 2006
    Carabus (Rhigocarabus) sehnali Brezina & Hackel, 2006
    Female adult
    female, 23mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) roborowskii roborowskii Semenov, 1887
    Carabus (Rhigocarabus) roborowskii roborowskii Semenov, 1887
    Male adult
    China, E Tibet, Chegarla 4800m, 20 km E
    of Rioche, 26.6.97 A. Wrzecionko leg.
    Copyright © Alexander Anichtchenko
    Carabus (Rhigocarabus) rhododendron Deuve & Imura, 1991
    Carabus (Rhigocarabus) rhododendron Deuve & Imura, 1991
    Male adult
    male, 18mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) rhododendron Deuve & Imura, 1991
    Carabus (Rhigocarabus) rhododendron Deuve & Imura, 1991
    Female adult
    female, 19mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) rhododendron Deuve & Imura, 1991
    Carabus (Rhigocarabus) rhododendron Deuve & Imura, 1991
    Male adult
    male, 17mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) quindecim Cavazzuti, 2002
    Carabus (Rhigocarabus) quindecim Cavazzuti, 2002 Holotype
    Male adult
    Holotype, male, 15mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) pseudopusio Deuve, 1996
    Carabus (Rhigocarabus) pseudopusio Deuve, 1996 Paratype
    Male adult
    male, 15mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) pseudopusio Deuve, 1996
    Carabus (Rhigocarabus) pseudopusio Deuve, 1996
    Female adult
    female, 16mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) poeta Semenov, 1898
    Carabus (Rhigocarabus) poeta Semenov, 1898 Holotype Paratype Lectotype
    Male adult
    male, 17mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) poeta Semenov, 1898
    Carabus (Rhigocarabus) poeta Semenov, 1898
    Male adult
    male, 18mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) poeta (as korsakowi Semenov, 1898)
    Carabus (Rhigocarabus) poeta (as korsakowi Semenov, 1898) Holotype
    Female adult
    Holotype, female 18 mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) poeta (as korsakowi Semenov, 1898)
    Carabus (Rhigocarabus) poeta (as korsakowi Semenov, 1898) Syntypes
    Female adult
    female, 16mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) poeta (as korsakowi Semenov, 1898)
    Carabus (Rhigocarabus) poeta (as korsakowi Semenov, 1898) Lectotype
    Male adult
    male, 17mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) pepek Imura & Kalab, 2006
    Carabus (Rhigocarabus) pepek Imura & Kalab, 2006 Paratype
    Male adult
    male, 17mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) pepek Imura & Kalab, 2006
    Carabus (Rhigocarabus) pepek Imura & Kalab, 2006 Paratype
    Female adult
    female, 18mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) morawitzianus breviculus Cavazzuti & Ratti, 1998
    Carabus (Rhigocarabus) morawitzianus breviculus Cavazzuti & Ratti, 1998 Paratype
    Male adult
    male, 15 mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) maleki Deuve, 1991
    Carabus (Rhigocarabus) maleki Deuve, 1991 Paratype
    Male adult
    male, 14mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) maleki Deuve, 1991
    Carabus (Rhigocarabus) maleki Deuve, 1991
    Female adult
    female, 15mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) maleki Deuve, 1991
    Carabus (Rhigocarabus) maleki Deuve, 1991 Paratype
    Male adult
    male, 15mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) itzingeri rugulosior Deuve, 1996 Paratype
    Carabus (Rhigocarabus) itzingeri rugulosior Deuve, 1996 Paratype Paratype
    Male adult
    male, 18mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) itzingeri itzingeri Breuning, 1934
    Carabus (Rhigocarabus) itzingeri itzingeri Breuning, 1934
    Male adult
    male, 19mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) itzingeri itzingeri Breuning, 1934
    Carabus (Rhigocarabus) itzingeri itzingeri Breuning, 1934
    Female adult
    female, 20mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) itzingeri kozaburoi Imura, 1999
    Carabus (Rhigocarabus) itzingeri kozaburoi Imura, 1999 Paratype
    Male adult
    male, 16mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) itzingeri kozaburoi Imura, 1999
    Carabus (Rhigocarabus) itzingeri kozaburoi Imura, 1999 Paratype
    Female adult
    female, 18mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) handelmazzettii handelmazzettii Mandl, 1955
    Carabus (Rhigocarabus) handelmazzettii handelmazzettii Mandl, 1955
    Male adult
    male, 18mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) handelmazzettii handelmazzettii Mandl, 1955
    Carabus (Rhigocarabus) handelmazzettii handelmazzettii Mandl, 1955
    Female adult
    female, 20mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) handelmazzettii handelmazzettii Mandl, 1955
    Carabus (Rhigocarabus) handelmazzettii handelmazzettii Mandl, 1955
    Female adult
    female, 21mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) handelmazzettii habaicus Deuve, 2005
    Carabus (Rhigocarabus) handelmazzettii habaicus Deuve, 2005 Paratype
    Male adult
    male, 16mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) pusio pusio s.str.
    Carabus (Rhigocarabus) pusio pusio s.str.
    Male adult
    male, 16mm
    (as hylonomus)
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) pusio pusio s.str.
    Carabus (Rhigocarabus) pusio pusio s.str.
    Female adult
    female, 17mm
    (as hylonomus)
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) gracilicollis gracilicollis Semenov, 1887
    Carabus (Rhigocarabus) gracilicollis gracilicollis Semenov, 1887
    Male adult
    male, 15mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) gracilicollis gracilicollis Semenov, 1887
    Carabus (Rhigocarabus) gracilicollis gracilicollis Semenov, 1887
    Female adult
    female, 16mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) gigoloides priapeius Cavazzuti, 2006
    Carabus (Rhigocarabus) gigoloides priapeius Cavazzuti, 2006
    Male adult
    male, 20mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) gigoloides jone Brezina & Hackel, 2004
    Carabus (Rhigocarabus) gigoloides jone Brezina & Hackel, 2004 Holotype Paratype
    Male adult
    male, 19mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) gigoloides jone Brezina & Hackel, 2004
    Carabus (Rhigocarabus) gigoloides jone Brezina & Hackel, 2004
    Female adult
    female, 20mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) gigolo (as qilingianus Cavazzuti, 2006)
    Carabus (Rhigocarabus) gigolo (as qilingianus Cavazzuti, 2006)
    Male adult
    18mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) gigolo Heinz & Brezina, 1996
    Carabus (Rhigocarabus) gigolo Heinz & Brezina, 1996 Paratype
    Male adult
    20mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) gigolo Heinz & Brezina, 1996
    Carabus (Rhigocarabus) gigolo Heinz & Brezina, 1996 Paratype
    Male adult
    20 mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) gigolo Heinz & Brezina, 1996
    Carabus (Rhigocarabus) gigolo Heinz & Brezina, 1996 Paratype
    Female adult
    21 mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) gentleman Brezina & Hackel, 2004
    Carabus (Rhigocarabus) gentleman Brezina & Hackel, 2004 Holotype
    Male adult
    male, 19mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) gentleman Brezina & Hackel, 2004
    Carabus (Rhigocarabus) gentleman Brezina & Hackel, 2004 Paratype
    Female adult
    female, 20mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) dacatraianus Deuve, 1995
    Carabus (Rhigocarabus) dacatraianus Deuve, 1995
    Male adult
    China, E Tibet, Angenla 5000-5100m
    45 km W of Bachen 29.6-2.7.1997 Wrzecionko leg.
    16 mm
    Copyright © Alexander Anichtchenko
    Carabus (Rhigocarabus) choui Deuve, 1989
    Carabus (Rhigocarabus) choui Deuve, 1989
    Male adult
    male, 16mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) choui Deuve, 1989
    Carabus (Rhigocarabus) choui Deuve, 1989
    Female adult
    female, 17mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) cateniger cateniger s.str. (as sommitalis)
    Carabus (Rhigocarabus) cateniger cateniger s.str. (as sommitalis)
    Male adult
    male, 16 mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) cateniger cateniger Morawitz, 1886
    Carabus (Rhigocarabus) cateniger cateniger Morawitz, 1886
    Female adult
    female, 19mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) cateniger cateniger Morawitz, 1886
    Carabus (Rhigocarabus) cateniger cateniger Morawitz, 1886
    Male adult
    male, 18mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) cateniger saltuensis Cavazzuti, 2006
    Carabus (Rhigocarabus) cateniger saltuensis Cavazzuti, 2006 Paratype
    Female adult
    female, 19mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) cateniger dabanicola Deuve, 1998
    Carabus (Rhigocarabus) cateniger dabanicola Deuve, 1998
    Male adult
    male, 16mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) cateniger dabanicola Deuve, 1998
    Carabus (Rhigocarabus) cateniger dabanicola Deuve, 1998
    Female adult
    female, 18mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) buddaicus bocsiki Deuve, 2007
    Carabus (Rhigocarabus) buddaicus bocsiki Deuve, 2007 Holotype
    Male adult
    male, 20 mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) buddaicus angenlaicus Deuve, 2007
    Carabus (Rhigocarabus) buddaicus angenlaicus Deuve, 2007 Paratype
    Male adult
    male, 18 mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) broukpytlik Brezina & Hackel, 2004
    Carabus (Rhigocarabus) broukpytlik Brezina & Hackel, 2004 Holotype Paratype
    Male adult
    male, 19mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) broukpytlik Brezina & Hackel, 2004
    Carabus (Rhigocarabus) broukpytlik Brezina & Hackel, 2004
    Female adult
    female, 20mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) broukpytlik Brezina & Hackel, 2004
    Carabus (Rhigocarabus) broukpytlik Brezina & Hackel, 2004
    Male adult
    male, 16mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) morawitzianus breviculus Cavazzuti & Ratti, 1998
    Carabus (Rhigocarabus) morawitzianus breviculus Cavazzuti & Ratti, 1998 Paratype
    Male adult
    China, N Tibet, Tangula Shan Mts., Tangula
    pass 5100-5300m, 8-11.7.1997 A. Wrzecionko leg.
    Copyright © Alexander Anichtchenko
    Carabus (Rhigocarabus) batangicus fickleri Korell & Richter, 1996
    Carabus (Rhigocarabus) batangicus fickleri Korell & Richter, 1996 Paratype
    Male adult
    male, 16 mm
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) poeta Semenov, 1898
    Carabus (Rhigocarabus) poeta Semenov, 1898
    Male adult
    male, 17mm
    Copyright © Boleslav Brezina
    Carabus (Piocarabus) reitterianus Breuning, 1934
    Carabus (Piocarabus) reitterianus Breuning, 1934
    Male adult
    China, Gansu, ca 25 km E Xiahe 2700-2900m, Wagatan Timber<br /> 30.7.-2.8.1993 Heinz leg.<br /> male, 23 mm
    Copyright © Alexander Anichtchenko
    Carabus (Piocarabus) nanwutai Kleinfeld, Korell & Wrase, 1996
    Carabus (Piocarabus) nanwutai Kleinfeld, Korell & Wrase, 1996 Holotype Paratype
    Male adult
    China, Shaanxi, Qing Ling Shan, track Hou Zen Zi vill.<br /> to Taiban Shan, 2500m, 27-29.6.1998 mixed forest<br /> male, long. 19 mm
    Copyright © Alexander Anichtchenko
    Carabus (Piocarabus) nanwutai Kleinfeld, Korell & Wrase, 1996
    Carabus (Piocarabus) nanwutai Kleinfeld, Korell & Wrase, 1996
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus kitawakianus kitawakianus Imura, 1993
    Carabus kitawakianus kitawakianus Imura, 1993
    Male adult
    China, Shaanxi, Tschinling mts.,
    Taibeichan Nat. res. 2000m 5-15.8.2005
    male, 27 mm
    Copyright © Alexander Anichtchenko
    Carabus (Pseudotribax) ferghanicus Breuning, 1933
    Carabus (Pseudotribax) ferghanicus Breuning, 1933
    Male adult
    male, 29mm
    Copyright © Boleslav Brezina
    Carabus (Pseudotribax) validus Kraatz, 1884
    Carabus (Pseudotribax) validus Kraatz, 1884
    Male adult
    male, 31mm
    Copyright © Boleslav Brezina
    Carabus (Pseudotribax) validus Kraatz, 1884
    Carabus (Pseudotribax) validus Kraatz, 1884
    Female adult
    female, 32mm
    Copyright © Boleslav Brezina
    Carabus (Pseudocranion) sackeni sackeni Semenov, 1898
    Carabus (Pseudocranion) sackeni sackeni Semenov, 1898
    Male adult
    male, 23mm
    Copyright © Boleslav Brezina
    Carabus (Pseudocranion) sackeni remondianus Deuve, 1996
    Carabus (Pseudocranion) sackeni remondianus Deuve, 1996 Paratype
    Female adult
    Museum of Institute of Systematic Biology (Daugavpils, Latvia)
    Copyright © Alexander Anichtchenko
    Carabus (Pseudocranion) labrangicus macilentior Deuve, 1996
    Carabus (Pseudocranion) labrangicus macilentior Deuve, 1996
    Female adult
    female, 21mm
    Copyright © Boleslav Brezina
    Carabus (Pseudocranion) gansuensis jedompansis Deuve, 1989
    Carabus (Pseudocranion) gansuensis jedompansis Deuve, 1989
    Male adult
    China, Sichuan, Langmusi 3200m 7.1999
    Copyright © Alexander Anichtchenko
    Carabus (Pseudocranion) fumigatus fumigatus Semenov, 1898
    Carabus (Pseudocranion) fumigatus fumigatus Semenov, 1898 Lectotype
    Male adult
    male, 22mm
    Copyright © Boleslav Brezina
    Carabus (Pseudocranion) fumigatus fumigatus Semenov, 1898
    Carabus (Pseudocranion) fumigatus fumigatus Semenov, 1898 Syntypes
    Female adult
    female, 22mm
    Copyright © Boleslav Brezina
    Carabus (Morphocarabus) gebleri ultimus Obydov, 1999
    Carabus (Morphocarabus) gebleri ultimus Obydov, 1999
    Female adult
    Carabus (Morphocarabus) gebleri ultimus Obydov, 1999 PT
    E Kazakhstan, Altai, near Zyrjanovska, 6.VII.1995, D. Obydov leg.
    female
    photo from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) gebleri ultimus Obydov, 1999
    Carabus (Morphocarabus) gebleri ultimus Obydov, 1999 Holotype
    Male adult
    Carabus (Morphocarabus) gebleri ultimus Obydov, 1999 HT
    E Kazakhstan, Altai, near Zyrjanovska, 6.VII.1995, D. Obydov leg.
    male
    photo from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) gebleri gebleri Fischer, 1817
    Carabus (Morphocarabus) gebleri gebleri Fischer, 1817
    Female adult
    Carabus (Morphocarabus) gebleri gebleri Fischer, 1817
    Russia, Altai, Zmeinogorsk
    female
    photo from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) gebleri gebleri Fischer, 1817
    Carabus (Morphocarabus) gebleri gebleri Fischer, 1817
    Male adult
    Carabus (Morphocarabus) gebleri gebleri Fischer, 1817
    Rusia, Altai, Zmeinogorsk
    male
    photo from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) chaudoiri zaikai Obydov, 1999 PT
    Carabus (Morphocarabus) chaudoiri zaikai Obydov, 1999 PT Paratype
    Female adult
    Carabus (Morphocarabus) chaudoiri zaikai Obydov, 1999 PT
    Rusia, Tuva, Akademik Obruchev Mt. rng., 2100 m;, 52°00'N 95°34'E,
    Sainak pass, tundra, 2.VII.1998, D. Obydov leg.
    female,
    photo from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) chaudoiri juliae Obydov, 2002
    Carabus (Morphocarabus) chaudoiri juliae Obydov, 2002
    Male adult
    male, 18mm
    Copyright © Boleslav Brezina
    Carabus (Morphocarabus) chaudoiri czadanicus Obydov, 1997 HT
    Carabus (Morphocarabus) chaudoiri czadanicus Obydov, 1997 HT Holotype
    Male adult
    Carabus (Morphocarabus) chaudoiri czadanicus Obydov, 1997 HT
    Rusia, Tuva, near Hadyn vill., 28-30.VI.1994, R. Shmarzh leg.
    male,
    photo from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) chaudoiri beladici Obydov, 2001
    Carabus (Morphocarabus) chaudoiri beladici Obydov, 2001
    Male adult
    S-Tuva
    East Tannu-Ola,Biche-Sailig r.h=2100m, 15.VII.1996, D.E.Lomakin leg.
    Copyright © Anton Menstschikov
    Carabus (Morphocarabus) chaudoiri beladici Obydov, 2001 PT
    Carabus (Morphocarabus) chaudoiri beladici Obydov, 2001 PT Paratype
    Female adult
    Carabus (Morphocarabus) chaudoiri beladici Obydov, 2001 PT
    Rusia, Tuva, E Tannu-Ola Mt.rng., near Kara-Hol lake, 29-31.V.2000, M. Beladzhik leg
    female,
    photo from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) aeruginosus zyrjanovskianus Shilenkov & O.Berlov, 1996
    Carabus (Morphocarabus) aeruginosus zyrjanovskianus Shilenkov & O.Berlov, 1996
    Male+Female adult
    NE KAZAKHSTAN,
    S-Altays, Ulbinsky Mt. h=900m., Bobrovka vill., 24.VI.01, A. Klimenko leg.
    female,25 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) aeruginosus zyrjanovskianus Shilenkov & O.Berlov, 1996
    Carabus (Morphocarabus) aeruginosus zyrjanovskianus Shilenkov & O.Berlov, 1996
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Pseudocoptolabrus) taliensis taliensis (Fairmaire, 1886)
    Carabus (Pseudocoptolabrus) taliensis taliensis (Fairmaire, 1886)
    Male adult
    China, Yunnan, Lijiang env., 2700 m, 10.6.1998
    Copyright © Alexander Anichtchenko
    Carabus (Pseudocoptolabrus) taliensis s.str. (as yulongxuensis)
    Carabus (Pseudocoptolabrus) taliensis s.str. (as yulongxuensis)
    Male adult
    male, 26mm
    Copyright © Boleslav Brezina
    Carabus (Pseudocoptolabrus) taliensis s.str. (as yulongxuensis)
    Carabus (Pseudocoptolabrus) taliensis s.str. (as yulongxuensis)
    Male adult
    male, 24mm
    Copyright © Boleslav Brezina
    Carabus (Pseudocoptolabrus) taliensis s.str. (as yulongxuensis)
    Carabus (Pseudocoptolabrus) taliensis s.str. (as yulongxuensis)
    Female adult
    female, 29mm
    Copyright © Boleslav Brezina
    Carabus (Pseudocoptolabrus) taliensis yanmenensis Deuve, 1996
    Carabus (Pseudocoptolabrus) taliensis yanmenensis Deuve, 1996
    Male adult
    male, 27mm
    Copyright © Boleslav Brezina
    Carabus (Pseudocoptolabrus) taliensis yanmenensis Deuve, 1996
    Carabus (Pseudocoptolabrus) taliensis yanmenensis Deuve, 1996
    Female adult
    female, 28mm
    Copyright © Boleslav Brezina
    Carabus (Pseudocoptolabrus) taliensis xueshanicola Deuve, 1992
    Carabus (Pseudocoptolabrus) taliensis xueshanicola Deuve, 1992
    Female adult
    female, 30mm
    Copyright © Boleslav Brezina
    Carabus (Pseudocoptolabrus) taliensis xueshanicola Deuve, 1992
    Carabus (Pseudocoptolabrus) taliensis xueshanicola Deuve, 1992
    Male adult
    male, 26mm
    Copyright © Boleslav Brezina
    Carabus (Pseudocoptolabrus) taliensis xueshanicola Deuve, 1992
    Carabus (Pseudocoptolabrus) taliensis xueshanicola Deuve, 1992
    Male adult
    male, 27mm
    Copyright © Boleslav Brezina
    Carabus (Pseudocoptolabrus) taliensis xueshanicola Deuve, 1992
    Carabus (Pseudocoptolabrus) taliensis xueshanicola Deuve, 1992
    Male adult
    male, 28mm
    Copyright © Boleslav Brezina
    Carabus (Pseudocoptolabrus) taliensis peterheisei
    Carabus (Pseudocoptolabrus) taliensis peterheisei Holotype
    Male adult
    Holotype
    Copyright © Frank Kleinfeld
    Carabus (Pseudocoptolabrus) taliensis myanmarensis Imura, 1999
    Carabus (Pseudocoptolabrus) taliensis myanmarensis Imura, 1999
    Female adult
    female, 36 mm
    Copyright © Boleslav Brezina
    Carabus (Pseudocoptolabrus) taliensis mulianus Deuve, 1995
    Carabus (Pseudocoptolabrus) taliensis mulianus Deuve, 1995
    Male adult
    male, 27mm
    Copyright © Boleslav Brezina
    Carabus (Pseudocoptolabrus) taliensis liqiuensis Imura, 1999
    Carabus (Pseudocoptolabrus) taliensis liqiuensis Imura, 1999
    Male adult
    male, 32mm
    Copyright © Boleslav Brezina
    Carabus (Pseudocoptolabrus) taliensis lijiangensis
    Carabus (Pseudocoptolabrus) taliensis lijiangensis Paratype
    Female adult
    female, 33mm
    Copyright © Boleslav Brezina
    Carabus (Pseudocoptolabrus) taliensis lijiangensis
    Carabus (Pseudocoptolabrus) taliensis lijiangensis
    Male adult
    male, 31mm
    Copyright © Boleslav Brezina
    Carabus (Pseudocoptolabrus) taliensis jingpingensis Deuve, 2005
    Carabus (Pseudocoptolabrus) taliensis jingpingensis Deuve, 2005 Paratype
    Male adult
    male, 26mm
    Copyright © Boleslav Brezina
    Carabus (Pseudocoptolabrus) taliensis jingpingensis Deuve, 2005
    Carabus (Pseudocoptolabrus) taliensis jingpingensis Deuve, 2005 Paratype
    Female adult
    female, 30mm
    Copyright © Boleslav Brezina
    Carabus (Pseudocoptolabrus) nosei taronensis Imura, 2001
    Carabus (Pseudocoptolabrus) nosei taronensis Imura, 2001 Paratype
    Female adult
    female, 28mm
    Copyright © Boleslav Brezina
    Carabus (Pseudocoptolabrus) burmanensis Breuning, 1932
    Carabus (Pseudocoptolabrus) burmanensis Breuning, 1932
    Male adult
    male, 23mm
    Copyright © Boleslav Brezina
    Carabus (Morphocarabus) aeruginosus verzhutzkii O.Berlov & Shilenkov, 1996
    Carabus (Morphocarabus) aeruginosus verzhutzkii O.Berlov & Shilenkov, 1996
    Male adult
    Carabus (Morphocarabus) aeruginosus verzhutzkii O.Berlov & Shilenkov, 1996
    C Altays, Katun r., 5 km. N Katanda vill., 21.06.2002, A. Menstschicov leg.
    Copyright © Anton Menstschikov
    Carabus (Pseudocoptolabrus) armiger armiger Imura, 1997
    Carabus (Pseudocoptolabrus) armiger armiger Imura, 1997
    Female adult
    female, 30mm
    Copyright © Boleslav Brezina
    Carabus (Morphocarabus) aeruginosus verzhutzkii O.Berlov & Shilenkov, 1996
    Carabus (Morphocarabus) aeruginosus verzhutzkii O.Berlov & Shilenkov, 1996
    Male adult
    C Altai Mts., Seminskii Mt.rng., Katun riv., 3 km S Ust'-Sema riv.,
    4.VII.2002 A. & N. Menschikovy leg.
    det. Mentschikov A.
    photo Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) aeruginosus salechardensis Obydov, 1999
    Carabus (Morphocarabus) aeruginosus salechardensis Obydov, 1999 Paratype
    Male adult
    Rusia, Selehard, tundra, 30.VIII.1953, I. Telyshev leg

    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) aeruginosus hermani Mannerheim, 1827 loc.typ.
    Carabus (Morphocarabus) aeruginosus hermani Mannerheim, 1827 loc.typ.
    Male adult
    C Rusia, S Ural Mts., Zlatoust, 22-28.VI.1952, S. Nikulina leg.
    male,

    Copyright © Dmitri Obydov
    Carabus (Procrustes) paiafa acuticollis Motschulsky, 1858
    Carabus (Procrustes) paiafa acuticollis Motschulsky, 1858
    Male adult

    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) aeruginosus eschscholtzi Mannerheim, 1827
    Carabus (Morphocarabus) aeruginosus eschscholtzi Mannerheim, 1827
    Male adult
    Russia, W Altai, Zmeinogorsk distr., near Cherepanovskii vill., 2-16.6.1990, D. Obydov leg.
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
    Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
    Male adult
    SE Siberia, E Sayan, Nizhneudinsk, h=500m, 21-26.VI.2006,
    male, 26mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
    Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
    Endophallus
    endophallus
    Copyright © Ruslan Panin
    Carabus (Mimocarabus) maurus osculatii Osculati, 1844
    Carabus (Mimocarabus) maurus osculatii Osculati, 1844
    Male adult
    Carabus (Mimocarabus) maurus osculatii Osculati, 1844
    Copyright © Alexander Anichtchenko
    Carabus (Mimocarabus) maurus maurus Adams, 1817
    Carabus (Mimocarabus) maurus maurus Adams, 1817
    Male+Female adult
    Armenia,
    Ashotsk distr., Tsoghamarg vill., 24-27.V.1999 P.V.Kazaryan leg.
    female, 20mm/male,18mm
    Copyright © Ruslan Panin
    Carabus (Procrustes) impressus Klug, 1832 (as carmelita)
    Carabus (Procrustes) impressus Klug, 1832 (as carmelita)
    Male adult

    Copyright © Safronov
    Carabus (Procrustes) iimpressus Klug, 1832 (as drusum)
    Carabus (Procrustes) iimpressus Klug, 1832 (as drusum) Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Procrustes) impressus Klug, 1832 (as desertum)
    Carabus (Procrustes) impressus Klug, 1832 (as desertum) Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Procrustes) coriaceus rugifer Kraatz, 1877
    Carabus (Procrustes) coriaceus rugifer Kraatz, 1877
    Male adult

    Copyright © Safronov
    Carabus (Procrustes) coriaceus kindermanni Waltl, 1838
    Carabus (Procrustes) coriaceus kindermanni Waltl, 1838
    Male+Female adult
    NW TURKEY,
    Stambul Avdzhilar, (Eropean part), h~20 m, 10-29.X.2007 M.V.Cheriavskaya leg./det. I.A.Solodovnikov
    female,31mm
    Copyright © Ruslan Panin
    Carabus (Procrustes) coriaceus hopffgarteni Kraatz, 1878
    Carabus (Procrustes) coriaceus hopffgarteni Kraatz, 1878
    Male adult

    Copyright © Safronov
    Carabus (Procrustes) coriaceus cerisyi Dejean, 1826 (as foudrasi)
    Carabus (Procrustes) coriaceus cerisyi Dejean, 1826 (as foudrasi)
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Procrustes) coriaceus coriaceus Linne, 1758
    Carabus (Procrustes) coriaceus coriaceus Linne, 1758
    Female adult
    The neighborhood Nicolo-Berezovskoe republic of
    Bashkortostan 5.07
    Copyright © Sunvas
    Carabus (Procrustes) coriaceus coriaceus Linne, 1758
    Carabus (Procrustes) coriaceus coriaceus Linne, 1758
    Male adult
    length: 32.42 mm
    Germania, Oberfranken, Weismain
    leg.det. U.Schmidt, 1983
    Foto: U.Schmidt, 2005
    Copyright © U.Schmidt
    Carabus (Procrustes) coriaceus cerisyi Dejean, 1826
    Carabus (Procrustes) coriaceus cerisyi Dejean, 1826
    Male adult
    male, 30mm
    Copyright © Boleslav Brezina
    Carabus (Procrustes) coriaceus cerisyi Dejean, 1826
    Carabus (Procrustes) coriaceus cerisyi Dejean, 1826
    Female adult
    female, 31mm
    Copyright © Boleslav Brezina
    Carabus (Procrustes) clypeatus clypeatus Adams, 1817
    Carabus (Procrustes) clypeatus clypeatus Adams, 1817
    Female adult
    C-Caucasus, N Ossetia, vall. Mamyhdon riv., h=2100
    Koval A.G. leg; 08.08.2001
    Copyright © Anton Menstschikov
    Carabus (Procrustes) chevrolati korbi Breuning, 1928
    Carabus (Procrustes) chevrolati korbi Breuning, 1928
    Male adult
    length: 28 mm
    Türkei, Tülü/Sathan, Erzincan
    leg. Staven & Willmer; det. K.Staven, 2003
    Foto: U.Schmidt, 2009
    Copyright © U.Schmidt
    Carabus (Procrustes) anatolicus lycicus Breuning, 1934
    Carabus (Procrustes) anatolicus lycicus Breuning, 1934
    Male adult

    Copyright © Safronov
    Carabus (Procrustes) chevrolati chevrolati Cristoforis & Jan, 1837
    Carabus (Procrustes) chevrolati chevrolati Cristoforis & Jan, 1837
    Male+Female adult
    female, 31mm: Ashotsk,Vardaghbyur vill., 30.V.1997
    male, 30 mm: Amasia, Chaldash vill.,8-11.VIII.1997
    P.V. Kazaryan leg.
    Copyright © Ruslan Panin
    Carabus (Procerus) scabrosus tauricus Bonelli, 1811
    Carabus (Procerus) scabrosus tauricus Bonelli, 1811
    Female adult

    Copyright © Safronov
    Carabus (Cechenochilus) heydenianus Starck, 1889
    Carabus (Cechenochilus) heydenianus Starck, 1889
    Male adult
    Caucasian nat. park., S slope Pseashkho mt., 2500 m
    21.7.1975 Ovtsharenko V. leg.
    Copyright © Alexander Anichtchenko
    Carabus (Cechenochilus) heydenianus Starck, 1889
    Carabus (Cechenochilus) heydenianus Starck, 1889
    Male+Female adult
    NW-Caucasus,
    Krasnodar reg., Mostovskoy distr., Skirda Mt Range, N slopes Yatyrgvart Mt. h=2300-2500m., 09-10.VII.2008, D. Fominykh leg.
    female,19mm/male,18mm
    Copyright © Ruslan Panin
    Carabus (Cechenochilus) adangensis Gottwald, 1983
    Carabus (Cechenochilus) adangensis Gottwald, 1983
    Male adult
    male, 19mm
    Copyright © Boleslav Brezina
    Carabus (Cechenochilus) adangensis Gottwald, 1983
    Carabus (Cechenochilus) adangensis Gottwald, 1983
    Female adult
    female, 18mm
    Copyright © Boleslav Brezina
    Carabus (Cechenochilus) adangensis Gottwald, 1983
    Carabus (Cechenochilus) adangensis Gottwald, 1983
    Female adult
    Russia, NW-Caucasus:
    Karacaj-Cherkessija, Abishira–Akhuba Mt. Range h=2000-2650m, Kyafar-Agur Riv. 14–17.VII.2009, Fominykh D. leg.
    female,21mm
    Copyright © Ruslan Panin
    Carabus (Cechenochilus) adangensis (as gusevi)
    Carabus (Cechenochilus) adangensis (as gusevi)
    Male adult
    male, 19mm
    Copyright © Boleslav Brezina
    Carabus (Cechenochilus) adangensis (as gusevi)
    Carabus (Cechenochilus) adangensis (as gusevi)
    Male adult
    male, 19mm
    Copyright © Boleslav Brezina
    Carabus (Cechenochilus) lazorum Belousov & Zamotajlov, 1999
    Carabus (Cechenochilus) lazorum Belousov & Zamotajlov, 1999
    Male adult
    male, 20mm
    Copyright © Boleslav Brezina
    Carabus (Cechenochilus) boeberi longiceps Chaudoir, 1846
    Carabus (Cechenochilus) boeberi longiceps Chaudoir, 1846
    Male adult
    C-Caucasus, N Ossetia, near Kozskii, h=2800,
    A.G. Koval leg. 10.08.2001; 12 mm
    Copyright © Anton Menstschikov
    Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009
    Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Holotype
    Male adult
    Russia, Karachai-Circassian Republic, Abishira-Akhuba Mount Range between passes Agur and Mylgvar, 2785–2830 m.
    Copyright © Igor Solodovnikov
    Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009
    Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009
    Male adult
    male, 17mm
    Copyright © Boleslav Brezina
    Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009
    Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Paratype
    Male adult
    Russia, W Caucasus,
    Karacaj-Cherkessija, Abishira-Akhuba Mnt Range, Area of pass Turii h~2650-2850m, 17.VII.2009, Fominykh D. leg.
    male,17 mm
    photo & coll. Panin R.
    Copyright © Ruslan Panin
    Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009
    Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Paratype
    Female adult
    Russia, Karachai-Circassian Republic, Abishira-Akhuba Mount Range between passes Agur and Mylgvar, 2785–2830 m. PARATYPUS
    Copyright © Igor Solodovnikov
    Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009
    Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009
    Female adult
    Russia, Karachai-Circassian Republic, Abishira-Akhuba Mount Range between passes Agur and Mylgvar, 2785–2830 m. PARATYPUS
    Copyright © Igor Solodovnikov
    Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009
    Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Paratype
    Female adult
    Russia, Karachai-Circassian Republic, Abishira-Akhuba Mount Range between passes Agur and Mylgvar, 2785–2830 m.
    Copyright © Igor Solodovnikov
    Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009
    Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Paratype
    Male adult
    Russia, Karachai-Circassian Republic, Abishira-Akhuba Mount Range between passes Agur and Mylgvar, 2785–2830 m.
    Copyright © Igor Solodovnikov
    Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009
    Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Paratype
    Habitat
    Russia, Karachai-Circassian Republic, Abishira-Akhuba Mount Range between passes Agur and Mylgvar, 2785–2830 m.
    photo: K.E.Dowgailo
    Copyright © Igor Solodovnikov
    Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009
    Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Paratype
    Female adult
    female, 21mm
    Copyright © Boleslav Brezina
    Carabus (Cechenochilus) boeberi buschi Semenov & Znojko, 1931
    Carabus (Cechenochilus) boeberi buschi Semenov & Znojko, 1931
    Female adult
    female, 19mm
    Copyright © Boleslav Brezina
    Carabus (Cechenochilus) boeberi buschi Semenov & Znojko, 1931
    Carabus (Cechenochilus) boeberi buschi Semenov & Znojko, 1931
    Female adult
    female, 23mm
    Copyright © Boleslav Brezina
    Carabus (Cechenochilus) boeberi boeberi Adams, 1817
    Carabus (Cechenochilus) boeberi boeberi Adams, 1817
    Male adult
    male, 18mm
    Copyright © Boleslav Brezina
    Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
    Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
    Male adult
    W.Ukraine, Transcarpathian Reg., Rakhov Distr., Chornogora range., h=2020m. Panin R. leg. 28.VI-9.VIII.2008, male, 19 mm
    Copyright © Ruslan Panin
    Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
    Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
    Female adult
    W Ukraine, NE Carpathians, Transcarpathian Reg., Rakhov Distr., Chornogora Mts Range h=2030m, alpin zone, 13-24.VIII.2010, Panin R. leg., female, 22 mm
    Copyright © Ruslan Panin
    Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
    Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
    Female adult
    Ukrainian Carpathians, Transcarpathian reg., Rakhov Distr., Chornogora range, h=2010m, 29.VI-28.VII.2007, Panin R. leg., female, 23 mm., photo/det.Panin R
    Copyright © Ruslan Panin
    Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
    Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
    Female adult
    Ukrainian Carpathians, Transcarpathian reg., Rakhov Distr., Chornogora range, h=2010 m., alpine zone 29.VI-28.VII.2007 Panin R. leg., female,23 mm.
    Copyright © Ruslan Panin
    Carabus (Platycarabus) irregularis montandoni Buysson, 1882
    Carabus (Platycarabus) irregularis montandoni Buysson, 1882
    Male adult
    W-Ukraine,
    Lviv Reg., Zhydachiv distr., near Vybranivka, under the bark of oak tree, in the forest,12.X.2007 Panin R. leg.
    male,22 mm.
    Copyright © Ruslan Panin
    Carabus (Platycarabus) irregularis montandoni Buysson, 1882
    Carabus (Platycarabus) irregularis montandoni Buysson, 1882
    Male adult
    W-Ukraine, NW Podolian,
    Lviv Reg., Zhydachiv distr., near Vybranivka, under the bark of oak tree, in the forest,12.X.2007 Panin R. leg
    L= 22 mm
    Copyright © Ruslan Panin
    Carabus (Platycarabus) depressus lucens Schaum, 1856
    Carabus (Platycarabus) depressus lucens Schaum, 1856
    Female adult
    Italia, Alpi Cozie, Col. delle Finestre,
    2150m, VII.1986 Ratti leg.
    female, 24 mm
    Copyright © Alexander Anichtchenko
    Carabus (Platycarabus) depressus depressus Bonelli, 1811
    Carabus (Platycarabus) depressus depressus Bonelli, 1811
    Female adult
    Italia, Bellamonte, 2200m
    VI.1992 Rubizzani leg.
    female, 20 mm
    Copyright © Alexander Anichtchenko
    Carabus (Platycarabus) cychroides Baudi, 1860
    Carabus (Platycarabus) cychroides Baudi, 1860
    Male+Female adult
    female: NW-Italy, Provincia Torino, Mt.Albergian h~2600 m, 9.VI.2007, V.Monzini leg.
    L=20 mm
    male:, N slope Mt.Albergian h~2000 m, 9.VI.2003, P.Zorgati leg.
    photo Panin R.
    Copyright © Ruslan Panin
    Carabus (Piocarabus) vladimirskyi Dejean & Boisduval, 1830
    Carabus (Piocarabus) vladimirskyi Dejean & Boisduval, 1830
    Male adult
    China, Beijing, Jiufeng mountain, 2007, Liu ye leg.;<br /> Photo & det. by liuye
    Copyright © Ye Liu
    Carabus (Eucarabus) sternbergi Roeschke, 1898
    Carabus (Eucarabus) sternbergi Roeschke, 1898
    Female adult
    Korea, Gyeongi Do,
    Bokwang Sa 30.2.1976
    female, 28 mm
    Copyright © Alexander Anichtchenko
    Carabus (Eucarabus) pawlowskianus Deuve, 1989
    Carabus (Eucarabus) pawlowskianus Deuve, 1989
    Female adult
    N Korea, Pong-San Mt.
    12.7.1997
    female, 25 mm
    Copyright © Alexander Anichtchenko
    Carabus (Eucarabus) nitididorsus Ishikawa & Kim, 1983
    Carabus (Eucarabus) nitididorsus Ishikawa & Kim, 1983
    Male adult
    S Korea, Deagu, 25.8.2003
    Copyright © Alexander Anichtchenko
    Carabus (Eucarabus) nitididorsus Ishikawa & Kim, 1983
    Carabus (Eucarabus) nitididorsus Ishikawa & Kim, 1983
    Female adult
    Korea, Peak Chonwongbong, 1600-1900m
    Mt. Chiri-San, Kyongsangnam-Do, 18.7.87 Y. Imura leg.
    Copyright © Alexander Anichtchenko
    Carabus (Eucarabus) manifestus guanmenshanus Imura, 1991
    Carabus (Eucarabus) manifestus guanmenshanus Imura, 1991
    Female adult
    China, Liaoning, Mt. Lao Gu Go, Dandong, 29.5.97
    Copyright © Alexander Anichtchenko
    Carabus (Eucarabus) billbergi billbergi Mannerheim, 1827
    Carabus (Eucarabus) billbergi billbergi Mannerheim, 1827
    Male adult
    Russia, E Siberia, Buryatia, lake Baikal, near Il'inka vill. Anichtchenko A. leg.
    Copyright © Alexander Anichtchenko
    Carabus (Eucarabus) billbergi billbergi Mannerheim, 1827
    Carabus (Eucarabus) billbergi billbergi Mannerheim, 1827
    Male adult
    Tian tou, Yueshi Counti, Anhui, China
    1-10.08.2004
    Copyright © Safronov
    Carabus (Pagocarabus) crassesculptus eleganticolor Deuve, 1997
    Carabus (Pagocarabus) crassesculptus eleganticolor Deuve, 1997 Paratype
    Male adult
    China, Gansu, Min Shan mts., 70 km NW Wudu
    Copyright © Alexander Anichtchenko
    Carabus (Eucarabus) ulrichi s.str. (as volhynicus Lutshnik, 1927)
    Carabus (Eucarabus) ulrichi s.str. (as volhynicus Lutshnik, 1927)
    Male+Female adult
    W Ukraine, NW Podolian,
    Ternopil reg., Pidvolochisk distr., forest Bogdanivskiy, near Romanove Selo, 7-24.V.2008, Panin R. & Solodovnikov I. leg.
    Copyright © Ruslan Panin
    Carabus (Pantophyrtus) turcomanorum karaalmicus Gottwald, 1994
    Carabus (Pantophyrtus) turcomanorum karaalmicus Gottwald, 1994
    Male adult
    male, 27mm
    Copyright © Boleslav Brezina
    Carabus (Pantophyrtus) turcomanorum turcomanorum Thieme, 1881
    Carabus (Pantophyrtus) turcomanorum turcomanorum Thieme, 1881
    Female adult
    female, 32mm
    Copyright © Boleslav Brezina
    Carabus (Pantophyrtus) turcomanorum turcomanorum Thieme, 1881
    Carabus (Pantophyrtus) turcomanorum turcomanorum Thieme, 1881
    Male adult
    male, 28mm
    Copyright © Boleslav Brezina
    Carabus (Pantophyrtus) turcomanorum s.str. (as orbitosus)
    Carabus (Pantophyrtus) turcomanorum s.str. (as orbitosus)
    Male adult
    male, 27mm
    Copyright © Boleslav Brezina
    Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993
    Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993
    Male adult
    Kirgizstan, S Chatkal mt. rng., Sary-Chelek lake 2000 m, 25.6.2000
    Copyright © Alexander Anichtchenko
    Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993
    Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993
    Male adult
    male, 29mm
    Copyright © Boleslav Brezina
    Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993
    Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993 Paratype
    Female adult
    female, 28mm
    Copyright © Boleslav Brezina
    Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993
    Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993 Paratype
    Female adult
    female, 34mm
    Copyright © Boleslav Brezina
    Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993
    Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993 Paratype
    Male adult
    male, 27mm
    Copyright © Boleslav Brezina
    Carabus (Pantophyrtus) brachypedilus brachypedilus s.str.
    Carabus (Pantophyrtus) brachypedilus brachypedilus s.str.
    Male adult
    N Kirgizskii mt rng., Ak-Suu riv., 1400 m,
    Belovodskoe, 16.4.2000 Klimenko A. leg.
    Copyright © Alexander Anichtchenko
    Carabus (Pantophyrtus) brachypedilus brachypedilus s.str.
    Carabus (Pantophyrtus) brachypedilus brachypedilus s.str.
    Female adult
    female, 27mm
    Copyright © Boleslav Brezina
    Carabus (Pantophyrtus) brachypedilus brachypedilus s.str.
    Carabus (Pantophyrtus) brachypedilus brachypedilus s.str.
    Male adult
    male, 26mm
    Copyright © Boleslav Brezina
    Carabus (Pantophyrtus) brachypedilus brachypedilus s.str.
    Carabus (Pantophyrtus) brachypedilus brachypedilus s.str.
    Female adult
    female, 28mm
    Copyright © Boleslav Brezina
    Carabus (Pantophyrtus) brachypedilus absentivus Belousov & Kabak 1993
    Carabus (Pantophyrtus) brachypedilus absentivus Belousov & Kabak 1993 Paratype
    Male adult
    male, 24mm
    Copyright © Boleslav Brezina
    Carabus (Pantophyrtus) brachypedilus absentivus Belousov & Kabak 1993
    Carabus (Pantophyrtus) brachypedilus absentivus Belousov & Kabak 1993 Paratype
    Male adult
    male, 24mm
    Copyright © Boleslav Brezina
    Carabus (Pantophyrtus) brachypedilus absentivus Belousov & Kabak 1993
    Carabus (Pantophyrtus) brachypedilus absentivus Belousov & Kabak 1993 Paratype
    Female adult
    female, 26mm
    Copyright © Boleslav Brezina
    Carabus (Pantophyrtus) brachypedilus absentivus Belousov & Kabak 1993
    Carabus (Pantophyrtus) brachypedilus absentivus Belousov & Kabak 1993 Paratype
    Female adult
    female, 30mm
    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) crassesculptus diruptus Morawitz, 1886
    Carabus (Pagocarabus) crassesculptus diruptus Morawitz, 1886
    Male adult
    China, Gansu, Umg. Xiahe 3000-3200m
    24-26.6.1993 Heinz leg.
    20 mm
    Copyright © Alexander Anichtchenko
    Carabus (Pagocarabus) crassethoracis Kleinfeld, 1999
    Carabus (Pagocarabus) crassethoracis Kleinfeld, 1999 Holotype
    Male adult

    Copyright © Frank Kleinfeld
    Carabus (Pagocarabus) crassesculptus crassesculptus Kraatz, 1881
    Carabus (Pagocarabus) crassesculptus crassesculptus Kraatz, 1881
    Male adult
    China, Beijing, Wulingshan, 2007, Liu ye leg.;
    Photo & det. by liuye
    Copyright © Ye Liu
    Carabus (Pagocarabus) crassesculptus crassesculptus Kraatz, 1881
    Carabus (Pagocarabus) crassesculptus crassesculptus Kraatz, 1881
    Male adult
    China, Hebei, Xianhua (=Kalgan)
    10-15.5.1995
    21 mm
    Copyright © Alexander Anichtchenko
    Carabus (Pagocarabus) crassesculptus tonghua Deuve & Mourzine, 2001
    Carabus (Pagocarabus) crassesculptus tonghua Deuve & Mourzine, 2001
    Male adult
    China, NW Sishuan, 3500m, Hongyuan
    Somang Qu riv. grassland, 9-29.6.2004
    male, 20 mm
    Copyright © Alexander Anichtchenko
    Carabus (Eucarabus) ulrichi ulrichi Germar, 1824
    Carabus (Eucarabus) ulrichi ulrichi Germar, 1824
    Endophallus
    endophallus ulrichi (as podolicus Semenov, 1898)
    Copyright © Ruslan Panin
    Carabus (Eucarabus) ulrichi ulrichi Germar, 1824 (as podolicus Semenov, 1898 loc.typ)
    Carabus (Eucarabus) ulrichi ulrichi Germar, 1824 (as podolicus Semenov, 1898 loc.typ)
    Female adult
    C-UKRAINE,<br /> Vinnitsa Reg., near Khmilnyk, forest, 10-20.VIII.2002, Demidov G.N. leg., det. Panin R<br /> female, 30mm
    Copyright © Ruslan Panin
    Carabus (Eucarabus) ulrichi ulrichi Germar, 1824
    Carabus (Eucarabus) ulrichi ulrichi Germar, 1824
    Male adult
    NW Ukrainian Carpathians
    Lviv Reg., Staryi Sambir Distr., near Dobromyl , h=500m 9.VI-25.VII.2007, Panin R. Leg.
    Copyright © Safronov
    Carabus (Platycarabus) irregularis montandoni Buysson, 1882
    Carabus (Platycarabus) irregularis montandoni Buysson, 1882
    Endophallus
    endophallus
    Copyright © Ruslan Panin
    Carabus (Tribax) nothus planipennis Chaudoir, 1846
    Carabus (Tribax) nothus planipennis Chaudoir, 1846
    Female adult
    Dagestan, Huizah., near Motchoh, h=1800, 27.07.1993, Ilina leg.
    photo A. Menstschicov
    Copyright © Anton Menstschikov
    Carabus (Tribax) nothus planipennis Chaudoir, 1846
    Carabus (Tribax) nothus planipennis Chaudoir, 1846
    Male adult
    male, 22mm
    Copyright © Boleslav Brezina
    Carabus (Tribax) nothus planipennis Chaudoir, 1846
    Carabus (Tribax) nothus planipennis Chaudoir, 1846
    Male adult
    male, 21mm
    Copyright © Boleslav Brezina
    Carabus (Tribax) nothus planipennis Chaudoir, 1846
    Carabus (Tribax) nothus planipennis Chaudoir, 1846
    Male adult
    Dagestan, Bogosskii mt. rng, Inhokvari-Honoh
    3500 m, 15-25.6.85 Abdurakhmanov leg.
    Copyright © Alexander Anichtchenko
    Carabus (Tribax) nothus nothus Adams, 1817
    Carabus (Tribax) nothus nothus Adams, 1817
    Male adult
    male, 24mm
    Copyright © Boleslav Brezina
    Carabus (Tribax) nothus nothus Adams, 1817
    Carabus (Tribax) nothus nothus Adams, 1817
    Female adult
    female, 24mm
    Copyright © Boleslav Brezina
    Carabus (Tribax) nothus nothus Adams, 1817
    Carabus (Tribax) nothus nothus Adams, 1817
    Female adult
    female, 25mm
    Copyright © Boleslav Brezina
    Carabus (Tribax) nothus fausti Dohrn, 1873
    Carabus (Tribax) nothus fausti Dohrn, 1873
    Male adult
    male, 24mm
    Copyright © Boleslav Brezina
    Carabus (Eucarabus) billbergi billbergi Mannerheim, 1827
    Carabus (Eucarabus) billbergi billbergi Mannerheim, 1827
    Male adult
    Rusia, Primorskii reg., Kiparisovo vill., 5.8.1993
    Chichvarhin A. leg.
    male, 20 mm
    Copyright © Alexander Anichtchenko
    Carabus (Eucarabus) billbergi billbergi Mannerheim, 1827
    Carabus (Eucarabus) billbergi billbergi Mannerheim, 1827
    Male adult
    Rusia, Primorskii reg., Anuchinskii distr.,
    near LZU-3, 7.8.1979 Bakurov leg.
    male, 21 mm
    Copyright © Alexander Anichtchenko
    Carabus (Tribax) nothus (as abdurachmanovi)
    Carabus (Tribax) nothus (as abdurachmanovi)
    Male adult
    Carabus (Tribax) nothus (as abdurachmanovi)
    male, 22mm
    Copyright © Boleslav Brezina
    Carabus (Tribax) nothus (as abdurachmanovi)
    Carabus (Tribax) nothus (as abdurachmanovi)
    Female adult
    Carabus (Tribax) nothus (as abdurachmanovi)
    female, 23mm
    Copyright © Boleslav Brezina
    Carabus (Tribax) kasakorum kasakorum Semenov, 1896
    Carabus (Tribax) kasakorum kasakorum Semenov, 1896
    Male adult
    male, 26mm
    Copyright © Boleslav Brezina
    Carabus (Tribax) kasakorum kasakorum Semenov, 1896
    Carabus (Tribax) kasakorum kasakorum Semenov, 1896
    Male adult
    male, 27mm
    Copyright © Boleslav Brezina
    Carabus (Tribax) kasakorum kasakorum Semenov, 1896
    Carabus (Tribax) kasakorum kasakorum Semenov, 1896
    Female adult
    female, 29mm
    Copyright © Boleslav Brezina
    Carabus (Tribax) kasakorum kasakorum Semenov, 1896
    Carabus (Tribax) kasakorum kasakorum Semenov, 1896
    Female adult
    female, 29mm
    Copyright © Boleslav Brezina
    Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
    Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
    Male adult
    male, 23mm
    Copyright © Boleslav Brezina
    Carabus (Microplectes) riedeli Menetries, 1832
    Carabus (Microplectes) riedeli Menetries, 1832
    Female adult
    female, 19mm
    Copyright © Boleslav Brezina
    Carabus (Microplectes) riedeli Menetries, 1832
    Carabus (Microplectes) riedeli Menetries, 1832
    Male adult
    male, 17mm
    Copyright © Boleslav Brezina
    Carabus (Microplectes) riedeli Menetries, 1832
    Carabus (Microplectes) riedeli Menetries, 1832
    Male adult
    male, 17mm
    Copyright © Boleslav Brezina
    Carabus (Microplectes) riedeli Menetries, 1832
    Carabus (Microplectes) riedeli Menetries, 1832
    Male adult
    N-Ossetia,
    Khazniaush Pass, h=2500-2700m, 9.VI-31.VII.1994, Zamotajlov A.leg.
    male, 17mm
    Copyright © Ruslan Panin
    Carabus (Microplectes) riedeli Menetries, 1832
    Carabus (Microplectes) riedeli Menetries, 1832
    Endophallus
    endophallus
    Copyright © Ruslan Panin
    Carabus (Microplectes) convallium convallium Starck, 1889
    Carabus (Microplectes) convallium convallium Starck, 1889
    Male adult
    male, 19mm
    Copyright © Boleslav Brezina
    Carabus (Microplectes) convallium convallium Starck, 1889
    Carabus (Microplectes) convallium convallium Starck, 1889
    Male adult
    male, 19mm
    Copyright © Boleslav Brezina
    Carabus (Microplectes) convallium convallium Starck, 1889
    Carabus (Microplectes) convallium convallium Starck, 1889
    Male+Female adult
    female:NW-Caucasus, Russia,
    Krasnodar reg., Apsheron Distr., Lagonaki Mt., near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominyh leg.
    male:Karachaevo-Cherkessia,the right source of the river Beskess, h=2100m, 10.VIII.1997, I.A. Solodovnikov leg.
    female, 21mm/male, 19mm
    Copyright © Ruslan Panin
    Carabus (Microplectes) convallium convallium Starck, 1889
    Carabus (Microplectes) convallium convallium Starck, 1889
    Endophallus
    Endophallus
    Copyright © Ruslan Panin
    Carabus (Microplectes) convallium argonautarum Semenov, 1898
    Carabus (Microplectes) convallium argonautarum Semenov, 1898
    Male adult
    male, 18mm
    Copyright © Boleslav Brezina
    Carabus (Microplectes) convallium argonautarum Semenov, 1898
    Carabus (Microplectes) convallium argonautarum Semenov, 1898
    Male adult
    male, 19mm
    Copyright © Boleslav Brezina
    Carabus (Microplectes) convallium argonautarum Semenov, 1898
    Carabus (Microplectes) convallium argonautarum Semenov, 1898
    Female adult
    female, 20mm
    Copyright © Boleslav Brezina
    Carabus (Microplectes) convallium argonautarum Semenov, 1898
    Carabus (Microplectes) convallium argonautarum Semenov, 1898
    Male+Female adult
    Abkhasia,
    W.slope of Gagrinsky Mt., Krepostnaya River source, h=930-980m., A.S.Zamotajlov leg. 27.VII.2000
    female, 19mm
    photo Panin R.
    Copyright © Ruslan Panin
    Carabus (Microplectes) convallium argonautarum Semenov, 1898
    Carabus (Microplectes) convallium argonautarum Semenov, 1898
    Endophallus
    Endophallus
    Copyright © Ruslan Panin
    Carabus (Pachystus) tamsi tamsi Menetries, 1832
    Carabus (Pachystus) tamsi tamsi Menetries, 1832
    Male+Female adult
    Transcaucasia, Armenia,
    Ashotsk, 18.VI.2001, P.V.Kazaryan leg.
    female, 26mm//male, 24mm
    Copyright © Ruslan Panin
    Carabus (Pachystus) tamsi tamsi Menetries, 1832
    Carabus (Pachystus) tamsi tamsi Menetries, 1832
    Female adult
    Azerbayjan, Talysh mt, Zuvant
    Pirasora, 1900-2000 m 9.7.1983 I. Belousov leg.
    female, 25 mm
    Copyright © Alexander Anichtchenko
    Carabus (Mesocarabus) macrocephalus Dejean, 1826
    Carabus (Mesocarabus) macrocephalus Dejean, 1826
    Male adult
    Spain, Picos de Europa, Macizo Central,
    Llagu Bajero 1890m 6.8.2003
    Copyright © Alexander Anichtchenko
    Carabus (Pachystus) tamsi tamsi Menetries, 1832
    Carabus (Pachystus) tamsi tamsi Menetries, 1832
    Male adult
    N Iran, Azarbaygan-e-Sharqi, 21 km SE
    Halhal, 3734N 4841E, 1633m
    3.6.2003 Kabatek P. leg
    male, 23 mm
    Copyright © Alexander Anichtchenko
    Carabus (Pachystus) tamsi tamsi Menetries, 1832
    Carabus (Pachystus) tamsi tamsi Menetries, 1832
    Endophallus
    endophallus
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) lusitanicus lusitanicus Fabricius, 1801
    Carabus (Mesocarabus) lusitanicus lusitanicus Fabricius, 1801
    Male adult
    Portugal, Vale de Estrela, Guarda, VI.1976
    Copyright © Alexander Anichtchenko
    Carabus (Mesocarabus) lusitanicus lusitanicus Fabricius, 1801(as schaumi Gaubil, 1849)
    Carabus (Mesocarabus) lusitanicus lusitanicus Fabricius, 1801(as schaumi Gaubil, 1849)
    Male+Female adult
    Portugal,
    Viseu distr., Gosende, h~1010m, 15.X.2009, D.Prunier leg.
    female,26 mm
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) lusitanicus brevis Dejean, 1826
    Carabus (Mesocarabus) lusitanicus brevis Dejean, 1826
    Female adult
    Spain, Sierra Guadarrama, Montes Carpetinos,
    Pto. de Somosierra 1600-1700m, 3.VI.2006
    Copyright © Alexander Anichtchenko
    Carabus (Mesocarabus) breuningi Csiki, 1927
    Carabus (Mesocarabus) breuningi Csiki, 1927
    Male adult
    Spain, San Martin de Castaneda, Zamora<br /> 11 oct 1984,h~1500 m
    Copyright © Alexander Anichtchenko
    Carabus (Mesocarabus) lusitanicus gougeleti Reiche, 1863 (as alcaracinus Kraatz, 1886)
    Carabus (Mesocarabus) lusitanicus gougeleti Reiche, 1863 (as alcaracinus Kraatz, 1886)
    Male adult
    Spain, Riopa (Albacete)
    Copyright © Alexander Anichtchenko
    Carabus (Pachystus) hungaricus scythus Motschulsky, 1847
    Carabus (Pachystus) hungaricus scythus Motschulsky, 1847
    Male adult
    S Ukraine,
    Zaporizhia reg., Khortytsa island, 19.VIII.2008, M. Mulenko leg., det.Panin R.
    male, 27mm
    Copyright © Ruslan Panin
    Carabus (Pachystus) hungaricus cribellatus Adams, 1812
    Carabus (Pachystus) hungaricus cribellatus Adams, 1812
    Female adult
    Kazakhstan,
    Astana reg. Enbekshil-der distr., Burevestnik vill., 9.VIII.2002
    female,26 mm
    Copyright © Ruslan Panin
    Carabus (Pachystus) graecus morio Mannerheim, 1830
    Carabus (Pachystus) graecus morio Mannerheim, 1830
    Male adult
    Turchia, Sehtavul Geg 1700m
    5.1996 Rabizzani leg.
    male, 26 mm
    Copyright © Alexander Anichtchenko
    Carabus (Pachystus) graecus kykladensis Breuning, 1961
    Carabus (Pachystus) graecus kykladensis Breuning, 1961
    Male adult
    male, 24mm
    Copyright © Boleslav Brezina
    Carabus (Pachystus) graecus kykladensis Breuning, 1961
    Carabus (Pachystus) graecus kykladensis Breuning, 1961
    Male adult
    female, 24mm
    Copyright © Boleslav Brezina
    Carabus (Mesocarabus) dufouri dufouri Dejean, 1829 (as papillatus Lapouge, 1902)
    Carabus (Mesocarabus) dufouri dufouri Dejean, 1829 (as papillatus Lapouge, 1902)
    Male adult
    Spain, Algeciras, Molino de las Cuevas
    5-6.4.2003 Anichtchenko A. leg.
    Copyright © Alexander Anichtchenko
    Carabus (Pachystus) glabratus glabratus Paykull, 1790
    Carabus (Pachystus) glabratus glabratus Paykull, 1790
    Female adult
    W UKRAINE<br /> Lviv region, Stryi distr., near Bilche vill., 9-20.VI.2005, Panin R. leg.<br /> female 30 mm
    Copyright © Ruslan Panin
    Carabus (Pachystus) glabratus glabratus Paykull, 1790
    Carabus (Pachystus) glabratus glabratus Paykull, 1790
    Male adult
    W Ukraine, NW Podolian, Lviv reg., Zolochiv distr., near Stinka vill., Lysa Gora, 26.IV.2009 R.Panin leg., 29mm
    Copyright © Ruslan Panin
    Carabus (Pachystus) glabratus glabratus Paykull, 1790
    Carabus (Pachystus) glabratus glabratus Paykull, 1790
    Female adult
    Germania, Oberfranken, Kulmbach leg.det. U.Schmidt, 1974 Foto: U.Schmidt, 2008
    Copyright © U.Schmidt
    Carabus (Pachystus) glabratus glabratus Paykull, 1790
    Carabus (Pachystus) glabratus glabratus Paykull, 1790
    Endophallus
    Endophallus
    Copyright © Ruslan Panin
    Carabus (Pachystus) cavernosus cavernosus Frivaldsky, 1837
    Carabus (Pachystus) cavernosus cavernosus Frivaldsky, 1837
    Male adult
    Bulgaria, Middle Stara Planina Mts.,
    feeder of Byala Reka riv., below Botev Peak
    1900-2000m 25.7.1996 B. Gueorguiev leg.
    male, 25 mm
    Copyright © Alexander Anichtchenko
    Carabus (Pachycarabus) staehlini Adams, 1817
    Carabus (Pachycarabus) staehlini Adams, 1817
    Male adult
    N Osetia, Skalistyi mt. rng.
    Hamarat-Hoh mt. 14.7.1996 Arzanov leg.
    male, 21 mm
    Copyright © Alexander Anichtchenko
    Carabus (Pachycarabus) roseri ciftekopruleriensis Machard, 1975
    Carabus (Pachycarabus) roseri ciftekopruleriensis Machard, 1975
    Female adult
    NE Turkey,
    Artvin Prov., Borcka, Macahel Gecidi h=1450m, 18.VI.2007, D.Prunier leg.
    female, 22mm
    Copyright © Ruslan Panin
    Carabus (Pachycarabus) imitator imitator Reitter, 1883
    Carabus (Pachycarabus) imitator imitator Reitter, 1883
    Male adult
    Abkhasia,
    Anchkho mnt.Range, near Lake Chkho, h~2450-2550m, 13-14.VII.2009, Bondarenko A.S. leg.
    male, 20mm
    Copyright © Ruslan Panin
    Carabus (Oxycarabus) saphyrinus pseudosaphyrinus Schweiger, 1964 (as boluensis)
    Carabus (Oxycarabus) saphyrinus pseudosaphyrinus Schweiger, 1964 (as boluensis)
    Male adult
    Turkei, Bolu, Seben, Aladag,
    1500m, 17.7.1988 Staven K. leg.
    male, 23 mm
    Copyright © Alexander Anichtchenko
    Carabus (Oxycarabus) saphyrinus pseudosaphyrinus Schweiger, 1964 (as rabaronianus)
    Carabus (Oxycarabus) saphyrinus pseudosaphyrinus Schweiger, 1964 (as rabaronianus)
    Male adult
    Anatolia, Kastamonu, Yaraligoz verso Catalzeitin,
    VI.2001 Sola Claudio leg.
    male, 28 mm
    Copyright © Alexander Anichtchenko
    Carabus (Oxycarabus) saphyrinus pseudosaphyrinus Schweiger, 1964 (as rabaronianus)
    Carabus (Oxycarabus) saphyrinus pseudosaphyrinus Schweiger, 1964 (as rabaronianus)
    Female adult
    Anatolia, Kastamonu, Yaraligoz
    verso Catalzeytin, VI.2001 Sola C. leg.
    female, 30 mm
    Copyright © Alexander Anichtchenko
    Carabus (Orinocarabus) sylvestris transsylvanicus Dejean, 1826
    Carabus (Orinocarabus) sylvestris transsylvanicus Dejean, 1826
    Male adult
    Ukrainian Carpathians,<br /> Transcarpathian reg., Rakhov Distr., Chornogora Mts. Petros Mt. h=1850-2000m. alpine zone, 29.VI-28.VII.2007<br /> Panin R. Leg.
    Copyright © Safronov
    Carabus (Orinocarabus) sylvestris transsylvanicus Dejean, 1826
    Carabus (Orinocarabus) sylvestris transsylvanicus Dejean, 1826
    Male+Female adult
    Ukrainian Carpathians
    Transcarpathian reg., Rakhov Distr., Chornogora Mts. Pietros Mt. h=1850-2000 m. alpine zone, 29.VI-28.VII.2007 Panin R. Leg.
    long 22 mm.
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) pedemontanus omensis Born, 1901
    Carabus (Orinocarabus) pedemontanus omensis Born, 1901
    Female adult
    Italy, Piemonte (CN), Col del Mulo
    17.7.2001 leg- Grottolo
    female, 22 mm
    Copyright © Alexander Anichtchenko
    Carabus (Orinocarabus) pedemontanus omensis Born, 1901
    Carabus (Orinocarabus) pedemontanus omensis Born, 1901
    Male adult
    Italy, Piemonte (CN), Col del Mulo
    17.7.2001 leg. Grottolo
    male, 22 mm
    Copyright © Alexander Anichtchenko
    Carabus (Orinocarabus) fairmairei Thomson, 1875
    Carabus (Orinocarabus) fairmairei Thomson, 1875
    Female adult
    SE France,
    Grand Belvédère du refuge du Mont Viso, 5-20.VII.1991
    female, 20mm.
    photo Panin R.
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) concolor concolor Fabricius, 1792
    Carabus (Orinocarabus) concolor concolor Fabricius, 1792
    Male adult
    Piemonte, Oropa (BI) , M. Camino,
    2300 m, 20.7.2003 leg. F. Angelini
    male, 18 mm
    Copyright © Alexander Anichtchenko
    Carabus (Orinocarabus) concolor concolor Fabricius, 1792
    Carabus (Orinocarabus) concolor concolor Fabricius, 1792
    Female adult
    Piemonte, Oropa (BI) , M. Camino,
    2300 m, 20.7.2003 leg. F. Angelini
    female, 19 mm
    Copyright © Alexander Anichtchenko
    Carabus (Orinocarabus) castanopterus A. & J.B. Villa, 1833
    Carabus (Orinocarabus) castanopterus A. & J.B. Villa, 1833
    Male adult
    Italy, Lombardia (BG), passo S. Simone 2000 m,
    Mezzoldo 26.8.2006 leg. Grottolo
    male, 17 mm
    Copyright © Alexander Anichtchenko
    Carabus (Orinocarabus) bertolinii Kraatz, 1878
    Carabus (Orinocarabus) bertolinii Kraatz, 1878
    Male adult
    Italia, Bellamonte, 2200 m, VI.1982 Rubizzani leg.
    male, 18 mm
    Copyright © Alexander Anichtchenko
    Carabus (Pachystus) hortensis hortensis Linne, 1758
    Carabus (Pachystus) hortensis hortensis Linne, 1758
    Male adult
    Germania, Oberfranken, Langenau
    leg.det. U.Schmidt, 2002
    Foto: U.Schmidt, 2006
    Copyright © U.Schmidt
    Carabus (Pachystus) hortensis preslii Dejean, 1830
    Carabus (Pachystus) hortensis preslii Dejean, 1830
    Female adult
    S Greece, Peloponnesos, 14.7.1987
    female, 28 mm
    Copyright © Alexander Anichtchenko
    Carabus (Pachystus) hortensis preslii Dejean, 1830 (as jonicus)
    Carabus (Pachystus) hortensis preslii Dejean, 1830 (as jonicus)
    Male adult
    Greece, Korfu 16.8.1987
    male, 25 mm
    Copyright © Alexander Anichtchenko
    Carabus (Pachystus) hortensis neumeyeri Schaum, 1856
    Carabus (Pachystus) hortensis neumeyeri Schaum, 1856
    Female adult
    Italy, Basilicata, Serra Calvello (PZ)
    1400 m, 7.2003 F. Angelini leg.
    female, 25 mm
    Copyright © Alexander Anichtchenko
    Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847 (as guadalupensis)
    Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847 (as guadalupensis) Paratype
    Male adult
    ESP - Caceres, S. Guadalupe,
    800 m XI.2002
    Copyright © Alexander Anichtchenko
    Carabus (Oreocarabus) ghiliani La Ferte-Senectere, 1847
    Carabus (Oreocarabus) ghiliani La Ferte-Senectere, 1847
    Male adult
    Spain, Segovia, Sierra de Somosierra,
    La Pinilla, 1800m 4.4.2005
    male, 19 mm
    Copyright © Alexander Anichtchenko
    Carabus (Oreocarabus) luetgensi luetgensi s.str.
    Carabus (Oreocarabus) luetgensi luetgensi s.str.
    Male adult
    Spain, Zamora, San Martin de Castaneda
    10.5.1997
    male, 20 mm
    Copyright © Alexander Anichtchenko
    Carabus (Ophiocarabus) regeli Morawitz, 1886
    Carabus (Ophiocarabus) regeli Morawitz, 1886
    Male adult
    Kirgizia, E Terskey Ala-Too, Chon-Akshuu pass,
    2800 m, 13.6.1999
    Copyright © Alexander Anichtchenko
    Carabus (Ophiocarabus) aeneolus aeneolus Morawitz, 1886
    Carabus (Ophiocarabus) aeneolus aeneolus Morawitz, 1886
    Male adult
    photo M. Smirnov from www.zin.ru
    Copyright © Maksim Smirnov
    Carabus (Ophiocarabus) aeneolus aeneolus Morawitz, 1886
    Carabus (Ophiocarabus) aeneolus aeneolus Morawitz, 1886
    Female adult
    SE KAZAKHSTAN,
    Alma-Ata reg., Zailiysky Mts., h=3000m, B. Almaatinka riv. valley,01.VI.2001
    Copyright © Nil Belonogov
    Carabus (Ophiocarabus) aeneolus aeneolus Morawitz, 1886
    Carabus (Ophiocarabus) aeneolus aeneolus Morawitz, 1886
    Male+Female adult
    KAZAKHSTAN,
    Bolshoe Almaatinskoe Lake,30.VI.1992
    female, 17mm
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) baguenai Breuning, 1926
    Carabus (Mesocarabus) baguenai Breuning, 1926
    Male adult
    Spain, Almeria, Sierra de Baza 10.III.1995<br /> male<br /> photo Anichtchenko A.<br />
    Copyright © Alexander Anichtchenko
    Carabus (Ohomopterus) yamato Nakane, 1953
    Carabus (Ohomopterus) yamato Nakane, 1953
    Female adult
    Japan, Ohkubo, 740-770m, Shirakawa-mura
    Ohno-gun (Gifu) 20-25.6.80 Ishikawa leg.
    Copyright © Alexander Anichtchenko
    Carabus (Ohomopterus) yaconinus yamaokai Ishikawa & Kubota, 1994
    Carabus (Ohomopterus) yaconinus yamaokai Ishikawa & Kubota, 1994
    Female adult
    Japan, Ehime pref., Taisanji, 14.5.2002
    Matsuyama city, T. Kurihara leg.
    Copyright © Alexander Anichtchenko
    Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
    Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
    Male adult
    SW-France,
    Foret d'Iraty Pyrenees-Atlantiques, d'Occabe, 15.VIII.1988
    male,30 mm
    photo Panin R.
    Copyright © Ruslan Panin
    Carabus (Ohomopterus) yaconinus yaconinus Bates, 1873
    Carabus (Ohomopterus) yaconinus yaconinus Bates, 1873
    Male adult
    Japan, Kagawa pref, Shin-ike
    Awai town, Kannonji city, 4.3.1993
    male, 28 mm
    Copyright © Alexander Anichtchenko
    Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876 Loc.typ.
    Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876 Loc.typ.
    Female adult
    Spain, Navarra, Sierra de Aralar,
    Altxueta 1200-1300m 6.1.2001
    Copyright © Alexander Anichtchenko
    Carabus (Ohomopterus) maiyasanus maiyasanus Bates, 1873
    Carabus (Ohomopterus) maiyasanus maiyasanus Bates, 1873
    Female adult
    Japan, Shiga pref.,
    Lake Yogo 18.8.1995 Shimizu leg.
    24 mm
    Copyright © Alexander Anichtchenko
    Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
    Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
    Endophallus
    endophalus
    Copyright © Ruslan Panin
    Carabus (Ohomopterus) japonicus tsushimae Breuning, 1932
    Carabus (Ohomopterus) japonicus tsushimae Breuning, 1932
    Male adult
    Japan, Tsushima isl., Nagasaki pref., Hiratake, Kamiagata town, 19.6.2002 T. Kurihara leg.
    male, 23 mm
    Copyright © Alexander Anichtchenko
    Carabus (Ohomopterus) insulicola insulicola Chaudoir, 1869
    Carabus (Ohomopterus) insulicola insulicola Chaudoir, 1869
    Male adult
    Japan, Yamanashi pref., Kodachi, Kawaguchiko-cho, Minamitusru distr.
    2.IX.2000 T. Kurihara leg.
    male, 29 mm
    Copyright © Alexander Anichtchenko
    Carabus (Ohomopterus) insulicola insulicola Chaudoir, 1869
    Carabus (Ohomopterus) insulicola insulicola Chaudoir, 1869
    Female adult
    15.VII.1998 JAPAN, Ichihara, env. Chiba
    Copyright © Nil Belonogov
    Carabus (Megodontus) violaceus rilvensis Kolbe, 1887
    Carabus (Megodontus) violaceus rilvensis Kolbe, 1887
    Female adult
    Bulgaria, Stara Zagora distr. Stara Planina Mts., Shipka Mt., 16.08.2005
    A. Safronov leg.

    Copyright © Safronov
    Carabus (Ohomopterus) dehaanii Chaudoir, 1848
    Carabus (Ohomopterus) dehaanii Chaudoir, 1848
    Male adult
    Japan, Hyogo pref., Ueda-rindo, Mihara, 4.3.93 N. Sano leg.
    Copyright © Alexander Anichtchenko
    Carabus (Ohomopterus) arrowianus murakii Ishikawa & Kubota, 1984
    Carabus (Ohomopterus) arrowianus murakii Ishikawa & Kubota, 1984
    Male adult
    Sho, alt 3m, Futami-T, Watarai-Co. Mie pref. Japan, 17.5.1987 Y. Imura leg.
    male, 27 mm
    Copyright © Alexander Anichtchenko
    Carabus (Ohomopterus) arrowianus arrowianus Breuning, 1934
    Carabus (Ohomopterus) arrowianus arrowianus Breuning, 1934
    Male adult
    Japan, Shizuoka pref., Mt. Mitake, Inasa town 18.10.1998 T. Kurihara leg.
    Copyright © Alexander Anichtchenko
    Carabus (Ohomopterus) albrechti esakianus Nakane, 1961
    Carabus (Ohomopterus) albrechti esakianus Nakane, 1961
    Male adult
    Japan, Tokyo pref, Mt. Takao, Nachioji City, 11.8.1990 K. Kurihara leg.
    Copyright © Alexander Anichtchenko
    Carabus (Megodontus) violaceus picenus A.& J.B.Villa, 1838
    Carabus (Megodontus) violaceus picenus A.& J.B.Villa, 1838
    Male adult
    Italia, Marche Pesaro
    leg.det. K.Staven, 1998
    Foto: U.Schmidt, 2008
    length: 26-30 mm
    Copyright © U.Schmidt
    Carabus (Ohomopterus) lewisianus lewisianus Breuning, 1932
    Carabus (Ohomopterus) lewisianus lewisianus Breuning, 1932
    Male adult
    Japan, Yamanashi pref., Koganesawa-rindo, Otsuki city,
    13.6.1990 K. Kurihara leg.
    male, 20 mm
    Copyright © Alexander Anichtchenko
    Carabus (Nesaeocarabus) gomerae A.Muller, 2004
    Carabus (Nesaeocarabus) gomerae A.Muller, 2004 Paratype
    Female adult

    Copyright © Boleslav Brezina
    Carabus (Megodontus) violaceus mixtus Gehin, 1876
    Carabus (Megodontus) violaceus mixtus Gehin, 1876
    Female adult
    France, Eloise 74, pont de Gresin 14.07.1991
    female, 29mm
    Copyright © Alexander Anichtchenko
    Carabus (Megodontus) violaceus fiorii Born, 1901
    Carabus (Megodontus) violaceus fiorii Born, 1901
    Male+Female adult
    NW-Italy,
    Reggio Emilia, Regnano, 5.IX.1996 leg. Cartini A.
    female, 30 mm
    photo Panin R.
    Copyright © Ruslan Panin
    Carabus (Megodontus) violaceus fiorii Born, 1901
    Carabus (Megodontus) violaceus fiorii Born, 1901
    Endophallus
    endophallus
    Copyright © Ruslan Panin
    Carabus (Megodontus) violaceus baeterrensis Lapouge, 1901
    Carabus (Megodontus) violaceus baeterrensis Lapouge, 1901
    Male+Female adult
    S France,
    Midi-Pyrénées Region, Ariège, Col du Chioula h=1500m, V-VI.2005, Pascal Stefani leg.
    photo Panin R.
    Copyright © Ruslan Panin
    Carabus (Megodontus) violaceus baeterrensis Lapouge, 1901
    Carabus (Megodontus) violaceus baeterrensis Lapouge, 1901
    Endophallus
    ehdophallus
    Copyright © Ruslan Panin
    Carabus (Megodontus) violaceus aurolimbatus Dejean, 1829
    Carabus (Megodontus) violaceus aurolimbatus Dejean, 1829
    Female adult
    Kazakhstan,
    North Kazakhstan Province (Kokchetavski region)1.VII.1986
    female, 24mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) violaceus aurolimbatus Dejean, 1829
    Carabus (Megodontus) violaceus aurolimbatus Dejean, 1829
    Male adult
    C.UKRAINE,
    Poltava reg., near Novi Sanzhary, bank of Vorskla River, 1-2.VII.07, D.Kovalchuk leg.
    male 25mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) violaceus aurichalceus Kraatz, 1879
    Carabus (Megodontus) violaceus aurichalceus Kraatz, 1879
    Male adult
    Spain, Pais Vasco, Aranzazu, 6.I.2005
    Anichtchenko A. leg.
    male, 27 mm
    photo, det. & coll. Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823
    Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823
    Male adult
    W Ukraine, NW Podolian, Ternopol reg., Kremenets distr., near Chornyi Lis vill., steppe sites, 4-22.V.2009, Panin R. leg. female, 29 mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823
    Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823
    Male adult
    W-UKRAINE
    NW Podolian, Lviv Reg., Zolochiv Distr., near Osovitsa, steppe sites, Panin R. leg. 1.V-6.VI.07
    male, 26mm
    Copyright © Safronov
    Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823
    Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823
    Male+Female adult
    W. Ukraine, W. Podolian, Lviv reg., Nikolaev distr., near Krasiv vill., under the bark of oak tree, 8.II.2009, Panin R. leg. female, 29mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823 (as carbonatus Schaufuss, 1882)
    Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823 (as carbonatus Schaufuss, 1882)
    Male adult
    W.UKRAINE, Ukrainian Carpathians, Lviv Reg., Skolivski Beskidy Mts., near Skole, h=700m, 20.IX.2009, Panin R. leg. male, 33mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823
    Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823
    Endophallus
    endophallus
    Lviv reg., Nikolaev distr., near Krasiv vill.
    Copyright © Ruslan Panin
    Carabus (Megodontus) vietinghoffi vietinghoffi Adams, 1812
    Carabus (Megodontus) vietinghoffi vietinghoffi Adams, 1812
    Male adult
    Yakutia, near Kular vill., 27.7.2000, 20 mm det. & coll. Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Carabus (Megodontus) vietinghoffi vietinghoffi Adams, 1812
    Carabus (Megodontus) vietinghoffi vietinghoffi Adams, 1812
    Female adult
    Canada
    Copyright Henri Goulet, Agriculture & Agri-Food Canada

    Copyright © Henri Goulet
    Carabus (Megodontus) vietinghoffi fulgidus Fischer, 1827
    Carabus (Megodontus) vietinghoffi fulgidus Fischer, 1827
    Male adult
    Russia, Evreiskaya AO., near Obluchie 14.6.1993
    Chichvarhin A. leg.
    male, 20 mm
    photo, det. & coll. Anichtchenko A
    Copyright © Alexander Anichtchenko
    Carabus (Megodontus) tuvensis Shilenkov, 1996
    Carabus (Megodontus) tuvensis Shilenkov, 1996
    Male adult
    Photo A. Menstschicov
    Copyright © Anton Menstschikov
    Carabus (Megodontus) tuvensis Shilenkov, 1996
    Carabus (Megodontus) tuvensis Shilenkov, 1996
    Female adult
    Photo A. Mentschikov
    Copyright © Anton Menstschikov
    Carabus (Megodontus) stroganowi tiedemanni Breuning, 1934
    Carabus (Megodontus) stroganowi tiedemanni Breuning, 1934
    Male adult
    IRAN, Gilan, Masuleh, 1550 m., 17.06.2002., O.Montrieull leg.
    photo A. Menstschicov
    Copyright © Anton Menstschikov
    Carabus (Megodontus) stroganowi stroganowi Zoubkoff, 1837
    Carabus (Megodontus) stroganowi stroganowi Zoubkoff, 1837
    Male adult
    Iran,
    Mazanderan, Naharkoran, b. Gorgan
    500m, 4-11.VII.1975 Heinz leg.
    male, 30 mm
    photo Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Carabus (Megodontus) septemcarinatus Motschulsky, 1840
    Carabus (Megodontus) septemcarinatus Motschulsky, 1840
    Male adult

    Copyright © Safronov
    Carabus (Megodontus) schoenherri schoenherri Fischer, 1822
    Carabus (Megodontus) schoenherri schoenherri Fischer, 1822
    Male adult

    Copyright © Anton Menstschikov
    Carabus (Megodontus) schoenherri schoenherri Fischer, 1822
    Carabus (Megodontus) schoenherri schoenherri Fischer, 1822
    Male adult
    enza Reg., Kuznetsk Distr., env. Tikhmenevo Vill., upper course of Sura River, pine forest VI.2005 I.Lebezhinskaja leg.
    L=28.0 mm
    Copyright © Kirill Makarov
    Carabus (Megodontus) schoenherri schoenherri Fischer, 1822 (as sangilenus Obydov, 1997)
    Carabus (Megodontus) schoenherri schoenherri Fischer, 1822 (as sangilenus Obydov, 1997)
    Male adult
    Tyva, Akademika Obrucheva mt.rng., 15km N Cherbi vill.,
    Chailag ur., 7.7.1999 A. & N. Menshikovy leg.
    male, 26mm
    photo Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Carabus (Megodontus) schoenherri schoenherri Fischer, 1822
    Carabus (Megodontus) schoenherri schoenherri Fischer, 1822
    Female adult

    Copyright © Safronov
    Carabus (Megodontus) persianus persianus Roeschke, 1896
    Carabus (Megodontus) persianus persianus Roeschke, 1896
    Male adult
    Iran, Gilan, pas. N.Dailiman, 1100m, 12-19.VII.1999 Heinz leg.
    male, 32 mm
    photo, det. & coll. Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Carabus (Megodontus) obovatus Fischer, 1827
    Carabus (Megodontus) obovatus Fischer, 1827
    Female adult
    Kemerovo reg., Kuznetskii Mt. rng., Berikul, 1.6.1982
    female, 16mm
    Copyright © Alexander Anichtchenko
    Carabus (Megodontus) obovatus Fischer, 1827
    Carabus (Megodontus) obovatus Fischer, 1827
    Male adult
    Kemerovo reg., Kuznetskii Mt. rng., Berikul, 1.6.1982
    male, 17mm
    Copyright © Alexander Anichtchenko
    Carabus (Megodontus) obovatus Fischer, 1827
    Carabus (Megodontus) obovatus Fischer, 1827
    Female adult
    Kuznetskii Mt., Gluhari vill, A. Menstschicov leg., 07.06.2006
    female
    Copyright © Anton Menstschikov
    Carabus (Megodontus) leachi surakovi Obydov, 1995
    Carabus (Megodontus) leachi surakovi Obydov, 1995
    Male adult

    Copyright © Kirill Makarov
    Carabus (Megodontus) leachi klimenkoi Obydov, 1999
    Carabus (Megodontus) leachi klimenkoi Obydov, 1999 Holotype
    Male adult
    Museum of Institute of Systematic Biology (Daugavpils, Latvia)
    Copyright © Alexander Anichtchenko
    Carabus (Megodontus) leachi panzeri Dejean, 1829 (as klimenkoi Obydov, 1999)
    Carabus (Megodontus) leachi panzeri Dejean, 1829 (as klimenkoi Obydov, 1999)
    Male adult
    Kazakhstan, S Altay mts., Ognevka vill.,<br /> Kalbinskii mt. rng., 16.6.1999 Klimenko A. leg.
    Copyright © Alexander Anichtchenko
    Carabus (Megodontus) gyllenhali Fischer, 1827
    Carabus (Megodontus) gyllenhali Fischer, 1827
    Male adult
    Krym Peninsula
    male, 29 mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) gyllenhali Fischer, 1827
    Carabus (Megodontus) gyllenhali Fischer, 1827
    Male adult
    Krym Peninsula, near Sevastopol, 05.05.2000, A.V. Konovalov leg.
    Copyright © Anton Menstschikov
    Carabus (Megodontus) exaratus Quensel, 1806
    Carabus (Megodontus) exaratus Quensel, 1806
    Male adult
    Russia, C.Caucasus, N.Ossetia,
    valley of River Khazniaush,
    h=1400 m, 10.VI-31.VII.1993
    A. Zamotajlov leg.
    Copyright © Safronov
    Carabus (Megodontus) exaratus Quensel, 1806
    Carabus (Megodontus) exaratus Quensel, 1806
    Male+Female adult
    NW Caucasus, Russia,
    Krasnodar reg., Krasnodar City, 27.IX-07.X.07, D.D.Fominykh leg.
    female,29mm/male, 27mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) exaratus Quensel, 1806
    Carabus (Megodontus) exaratus Quensel, 1806
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Megodontus) kantaikensis Gehin, 1885
    Carabus (Megodontus) kantaikensis Gehin, 1885
    Female adult
    Russia, Lake Baikal, vall. of Goloustnaya riv., Nizhnii Kochergat vill.,
    27.VII.1994 Anichtchenko A. leg.
    female, 18 mm
    photo, det. & coll. Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Carabus (Megodontus) kantaikensis Gehin, 1885
    Carabus (Megodontus) kantaikensis Gehin, 1885
    Male adult
    E Sayan Mts., Sentsa riv. (Oka riv. confluence), near Shasnur vill.
    7-16.7.1998 Anichtchenko A. leg.
    male, 18 mm
    photo, det. & coll. Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Carabus (Megodontus) kantaikensis Gehin, 1885
    Carabus (Megodontus) kantaikensis Gehin, 1885
    Male adult
    Krasnoyarsk Area, Sayanogorsk Distr, env. Cheremushki Vill., right bank of Enisej Riv., h= 500 - 800 m, 23.07.1999 A. Semin leg.<br /> male<br /> L=17,0 mm
    Copyright © Maksim Smirnov
    Carabus (Megodontus) viridicans Obydov, 1993
    Carabus (Megodontus) viridicans Obydov, 1993
    Male adult
    W Sayan, Cheremushki vill., Borus Mt. rng., h= 500m, 29-30.7.2000, Brinev A. leg.; 16 mm
    Copyright © Alexander Anichtchenko
    Carabus (Megodontus) viridicans Obydov, 1993
    Carabus (Megodontus) viridicans Obydov, 1993
    Female adult
    Photo A. Menstschicov
    Copyright © Anton Menstschikov
    Carabus (Megodontus) viridicans Obydov, 1993
    Carabus (Megodontus) viridicans Obydov, 1993
    Male adult

    Copyright © Anton Menstschikov
    Carabus (Megodontus) danilevskii Obydov, 1993
    Carabus (Megodontus) danilevskii Obydov, 1993
    Male adult
    Krasnoyarsk Area, Sayanogorsk Distr, env. Cheremushki Vill., right bank of Enisej Riv., h= 500 - 800 m, 23.07.1999 A. Semin leg.
    L=17,0 mm
    Copyright © Maksim Smirnov
    Carabus (Nesaeocarabus) faustus faustus Brulle, 1838
    Carabus (Nesaeocarabus) faustus faustus Brulle, 1838
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Nesaeocarabus) faustus faustus Brulle, 1838
    Carabus (Nesaeocarabus) faustus faustus Brulle, 1838
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Nesaeocarabus) faustus cabrerai Enderlein, 1929
    Carabus (Nesaeocarabus) faustus cabrerai Enderlein, 1929
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Nesaeocarabus) coarctatus Brulle, 1838
    Carabus (Nesaeocarabus) coarctatus Brulle, 1838
    Female adult
    Canary isl., Las Palmas, Las Mesas 20.2.90
    Copyright © Alexander Anichtchenko
    Carabus (Nesaeocarabus) abbreviatus Brulle, 1835
    Carabus (Nesaeocarabus) abbreviatus Brulle, 1835
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Nesaeocarabus) abbreviatus Brulle, 1835
    Carabus (Nesaeocarabus) abbreviatus Brulle, 1835
    Female adult
    Spain, Canary Islands,
    Tenerife, Ft de Esperanza 1360m, TF-24, 29.III.2007, D.Prunier leg.
    female, 22mm
    Copyright © Ruslan Panin
    Carabus (Pagocarabus) wrzecionkoianus Deuve, 2007
    Carabus (Pagocarabus) wrzecionkoianus Deuve, 2007 Holotype Paratype
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) wrzecionkoianus Deuve, 2007
    Carabus (Pagocarabus) wrzecionkoianus Deuve, 2007
    Female adult

    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) wagae folwarcznyi Deuve, 1997
    Carabus (Pagocarabus) wagae folwarcznyi Deuve, 1997 Paratype
    Male adult
    male, 21 mm
    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) ? wagae Fairmaire, 1882
    Carabus (Pagocarabus) ? wagae Fairmaire, 1882
    Male adult
    China, E Tibet, pas Dongdola 5000 m, 4.7.1995
    Copyright © Alexander Anichtchenko
    Carabus (Pagocarabus) wagae degeicola Deuve, 1996
    Carabus (Pagocarabus) wagae degeicola Deuve, 1996
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) tsogoensis Deuve, 1997
    Carabus (Pagocarabus) tsogoensis Deuve, 1997
    Male adult
    male, 20 mm
    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) tryznai Brezina & Hackel, 2006
    Carabus (Pagocarabus) tryznai Brezina & Hackel, 2006 Holotype
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) tryznai Brezina & Hackel, 2006
    Carabus (Pagocarabus) tryznai Brezina & Hackel, 2006 Paratype
    Female adult

    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) sinotibeticola xiaodongi (Imura, Zhou & Su, 2004)
    Carabus (Pagocarabus) sinotibeticola xiaodongi (Imura, Zhou & Su, 2004)
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) sinotibeticola sinotibeticola Mandl, 1975
    Carabus (Pagocarabus) sinotibeticola sinotibeticola Mandl, 1975
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) sinotibeticola sinotibeticola Mandl, 1975
    Carabus (Pagocarabus) sinotibeticola sinotibeticola Mandl, 1975
    Female adult

    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) sinotibeticola pantholops Brezina, Hackel & Sehnal, 2007
    Carabus (Pagocarabus) sinotibeticola pantholops Brezina, Hackel & Sehnal, 2007 Holotype Paratype
    Male adult
    male, 23mm
    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) sinotibeticola pantholops Brezina, Hackel & Sehnal, 2007
    Carabus (Pagocarabus) sinotibeticola pantholops Brezina, Hackel & Sehnal, 2007
    Female adult
    female, 24mm
    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) sichuanicola sigena Kleinfeld, 2000
    Carabus (Pagocarabus) sichuanicola sigena Kleinfeld, 2000 Holotype
    Male adult

    Copyright © Frank Kleinfeld
    Carabus (Pagocarabus) sichuanicola sigena Kleinfeld, 2000
    Carabus (Pagocarabus) sichuanicola sigena Kleinfeld, 2000
    Male adult
    male, 26mm
    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) sichuanicola morvanellus Deuve, 1998
    Carabus (Pagocarabus) sichuanicola morvanellus Deuve, 1998 Holotype
    Female adult

    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) sichuanicola sichuanicola Deuve, 1989 (as dromlae)
    Carabus (Pagocarabus) sichuanicola sichuanicola Deuve, 1989 (as dromlae) Paratype
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) puetzi Kleinfeld, 2000
    Carabus (Pagocarabus) puetzi Kleinfeld, 2000 Holotype
    Male adult

    Copyright © Frank Kleinfeld
    Carabus (Pagocarabus) piceophilus piceophilus Deuve, 1994
    Carabus (Pagocarabus) piceophilus piceophilus Deuve, 1994
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) piceophilus piceophilus Deuve, 1994
    Carabus (Pagocarabus) piceophilus piceophilus Deuve, 1994
    Male adult
    male, 22mm
    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) paulusi Kalab, 1995
    Carabus (Pagocarabus) paulusi Kalab, 1995
    Male adult
    China, E-Tibet, Chakla, 5000m, 100 km
    N of Lhasa, 4.6.1997 Wrzecionko A. leg.
    Copyright © Alexander Anichtchenko
    Carabus (Pagocarabus) nanschanicus globulipennis Deuve, 2006
    Carabus (Pagocarabus) nanschanicus globulipennis Deuve, 2006 Holotype
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Rhigoidocarabus) zhubajie laojunensis Kleinfeld, 1999
    Carabus (Rhigoidocarabus) zhubajie laojunensis Kleinfeld, 1999 Holotype
    Male adult

    Copyright © Frank Kleinfeld
    Carabus (Oreocarabus) taedatus agassi Le Conte, 1850
    Carabus (Oreocarabus) taedatus agassi Le Conte, 1850
    Female adult
    Canada
    Copyright Henri Goulet, Agriculture & Agri-Food Canada
    Copyright © Henri Goulet
    Carabus (Oreocarabus) taedatus agassi Le Conte, 1850
    Carabus (Oreocarabus) taedatus agassi Le Conte, 1850
    Male adult
    USA, Arizona, Coconino Co., Humphreys Peak,
    bei Flagstaff 14.6.1990 Heinz leg.
    Copyright © Alexander Anichtchenko
    Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
    Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
    Female adult
    female, 49mm
    Copyright © Boleslav Brezina
    Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
    Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
    Male adult
    male, 43mm
    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) lama yajiangensis Deuve, 1994
    Carabus (Pagocarabus) lama yajiangensis Deuve, 1994
    Male adult
    male, 23mm
    Copyright © Boleslav Brezina
    Carabus (Archicarabus) pseudomonticola Lapouge, 1908 ? nemoralis
    Carabus (Archicarabus) pseudomonticola Lapouge, 1908 ? nemoralis
    Male adult
    NE Spain, Catalonia, Sant Jaume de Frontanyà, h=1000 m, V-VI.2010, Pascal Stefani leg.
    L=26 мм
    Copyright © Ruslan Panin
    Carabus (Pagocarabus) lama daochengensis Deuve, 1994
    Carabus (Pagocarabus) lama daochengensis Deuve, 1994
    Male adult
    male, 21mm
    Copyright © Boleslav Brezina
    Carabus (Archicarabus) pseudomonticola Lapouge, 1908
    Carabus (Archicarabus) pseudomonticola Lapouge, 1908
    Male adult
    Spain, Montseny, Barcelona
    19.VI.1986 M. Tomas leg.
    male, 21 mm
    Copyright © Alexander Anichtchenko
    Carabus (Pagocarabus) kamensis tatsienlui Breuning, 1934
    Carabus (Pagocarabus) kamensis tatsienlui Breuning, 1934
    Female adult

    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) jiudingensis jiudingensis Deuve, 1994
    Carabus (Pagocarabus) jiudingensis jiudingensis Deuve, 1994
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) jiudingensis jiudingensis Deuve, 1994
    Carabus (Pagocarabus) jiudingensis jiudingensis Deuve, 1994
    Female adult

    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) draco Brezina, 1999
    Carabus (Pagocarabus) draco Brezina, 1999
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) chomae (Imura, 2002)
    Carabus (Pagocarabus) chomae (Imura, 2002)
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) chomae (Imura, 2002)
    Carabus (Pagocarabus) chomae (Imura, 2002)
    Female adult

    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) chomae (Imura, 2002)
    Carabus (Pagocarabus) chomae (Imura, 2002)
    Male adult
    male, 24mm
    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) caelestinus garpen Kalab, 2007
    Carabus (Pagocarabus) caelestinus garpen Kalab, 2007 Holotype Paratype
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) caelestinus garpen Kalab, 2007
    Carabus (Pagocarabus) caelestinus garpen Kalab, 2007
    Female adult
    female, 21 mm
    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) caelestinus caelestinus (Imura, 2007)
    Carabus (Pagocarabus) caelestinus caelestinus (Imura, 2007) Paratype
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) borodini Heinz, 1996
    Carabus (Pagocarabus) borodini Heinz, 1996
    Male adult
    China, C Tibet, Mila 5000m, 150 km E of Lhasa 6.6.1997 Wrzecionko A. leg.
    Copyright © Alexander Anichtchenko
    Carabus (Pagocarabus) alpherakii takin Hackel & Sehnal, 2007
    Carabus (Pagocarabus) alpherakii takin Hackel & Sehnal, 2007 Holotype Paratype
    Male adult
    male, 19mm
    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) alpherakii takin Hackel & Sehnal, 2007
    Carabus (Pagocarabus) alpherakii takin Hackel & Sehnal, 2007
    Male adult
    male, 17mm
    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) alpherakii takin Hackel & Sehnal, 2007
    Carabus (Pagocarabus) alpherakii takin Hackel & Sehnal, 2007 Paratype
    Female adult
    female, 22mm
    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) alpherakii takin Hackel & Sehnal, 2007
    Carabus (Pagocarabus) alpherakii takin Hackel & Sehnal, 2007 Paratype
    Female adult
    female, 23mm
    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) alpherakii takin Hackel & Sehnal, 2007
    Carabus (Pagocarabus) alpherakii takin Hackel & Sehnal, 2007 Paratype
    Male adult
    male, 18 mm
    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) alpherakii pignataroi Cavazzuti, 1996
    Carabus (Pagocarabus) alpherakii pignataroi Cavazzuti, 1996 Paratype
    Female adult
    female, 20mm
    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) alpherakii pignataroi Cavazzuti, 1996
    Carabus (Pagocarabus) alpherakii pignataroi Cavazzuti, 1996
    Male adult
    male, 20mm
    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) alpherakii pignataroi Cavazzuti, 1996
    Carabus (Pagocarabus) alpherakii pignataroi Cavazzuti, 1996
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) alpherakii pignataroi Cavazzuti, 1996
    Carabus (Pagocarabus) alpherakii pignataroi Cavazzuti, 1996 Paratype
    Female adult

    Copyright © Boleslav Brezina
    Carabus (Cupreocarabus) alpherakii alpherakii Semenov, 1898
    Carabus (Cupreocarabus) alpherakii alpherakii Semenov, 1898
    Male adult
    male, 23mm
    Copyright © Boleslav Brezina
    Carabus (Cupreocarabus) alpherakii alpherakii Semenov, 1898
    Carabus (Cupreocarabus) alpherakii alpherakii Semenov, 1898
    Female adult
    female, 24mm
    Copyright © Boleslav Brezina
    Carabus (Cupreocarabus) alpherakii alpherakii Semenov, 1898
    Carabus (Cupreocarabus) alpherakii alpherakii Semenov, 1898
    Male adult
    Chine, W Sichuan, Litang env., <br /> 3-6.7.99 Dr. V. Banes leg.
    Copyright © Alexander Anichtchenko
    Carabus (Pagocarabus) alpherakii budha Breuning, 1950
    Carabus (Pagocarabus) alpherakii budha Breuning, 1950
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) alpherakii budha Breuning, 1950
    Carabus (Pagocarabus) alpherakii budha Breuning, 1950
    Female adult
    female, 20mm
    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) alpherakii budha Breuning, 1950
    Carabus (Pagocarabus) alpherakii budha Breuning, 1950
    Female adult
    female, 24mm
    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) alpherakii budha Breuning, 1950
    Carabus (Pagocarabus) alpherakii budha Breuning, 1950
    Male adult
    male, 20 mm
    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) alpherakii budha Breuning, 1950
    Carabus (Pagocarabus) alpherakii budha Breuning, 1950
    Male adult
    male, 19mm
    Copyright © Boleslav Brezina
    Carabus (Apotomopterus) leda Kleinfeld, 2000
    Carabus (Apotomopterus) leda Kleinfeld, 2000 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) korellianus Kleinfeld, 2002
    Carabus (Apotomopterus) korellianus Kleinfeld, 2002 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) infirmior homodynamus Deuve, 1994 (as jiucaishan)
    Carabus (Apotomopterus) infirmior homodynamus Deuve, 1994 (as jiucaishan) Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) hera Kleinfeld, 2000
    Carabus (Apotomopterus) hera Kleinfeld, 2000 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) guzhangensis Kleinfeld, 1998
    Carabus (Apotomopterus) guzhangensis Kleinfeld, 1998 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) guizhouensis yunwuensis Kleinfeld, 1996
    Carabus (Apotomopterus) guizhouensis yunwuensis Kleinfeld, 1996 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) guangxicus Deuve, 1989
    Carabus (Apotomopterus) guangxicus Deuve, 1989
    Male adult
    Guposhan, Hezhou, Guangxi prov. China
    male,
    photo Daniel Prunier
    Copyright © Daniel Prunier
    Carabus (Apotomopterus) grossefoveatus grossefoveatus Hauser, 1913 (as zhongnanshan)
    Carabus (Apotomopterus) grossefoveatus grossefoveatus Hauser, 1913 (as zhongnanshan) Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) grossefoveatus annikae Kleinfeld, 1996
    Carabus (Apotomopterus) grossefoveatus annikae Kleinfeld, 1996 Holotype
    Male adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) delavayi yunlongensis Kleinfeld, 2000
    Carabus (Apotomopterus) delavayi yunlongensis Kleinfeld, 2000 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) longeantennatus angulihabitus Kleinfeld, 1998
    Carabus (Apotomopterus) longeantennatus angulihabitus Kleinfeld, 1998 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) longeantennatus changyangensis Kleinfeld, 1998
    Carabus (Apotomopterus) longeantennatus changyangensis Kleinfeld, 1998 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) longeantennatus yongshan Kleinfeld, 1999
    Carabus (Apotomopterus) longeantennatus yongshan Kleinfeld, 1999 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) mecynodes helena Kleinfeld, 1999
    Carabus (Apotomopterus) mecynodes helena Kleinfeld, 1999 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) penelope penelope Kleinfeld, 1997
    Carabus (Apotomopterus) penelope penelope Kleinfeld, 1997 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) rapuzzi rapuzzi Kleinfeld, 2000
    Carabus (Apotomopterus) rapuzzi rapuzzi Kleinfeld, 2000 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) roxane roxane Kleinfeld, 2002
    Carabus (Apotomopterus) roxane roxane Kleinfeld, 2002 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) yunanensis yunanensis Born, 1905
    Carabus (Apotomopterus) yunanensis yunanensis Born, 1905 Paralectotypes
    Female adult

    Copyright © Alexander Anichtchenko
    Carabus (Apotomopterus) sauteri fujianensis Kleinfeld, 1997
    Carabus (Apotomopterus) sauteri fujianensis Kleinfeld, 1997 Holotype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) sauteri sanjiangensis Kleinfeld, 2001
    Carabus (Apotomopterus) sauteri sanjiangensis Kleinfeld, 2001 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) sauteri tianpingensis Deuve & Yu, 1992 (as eleganticollis chalingensis)
    Carabus (Apotomopterus) sauteri tianpingensis Deuve & Yu, 1992 (as eleganticollis chalingensis) Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) sauteri tianpingensis Deuve & Yu, 1992 (as longhuiensis)
    Carabus (Apotomopterus) sauteri tianpingensis Deuve & Yu, 1992 (as longhuiensis) Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) sauteri tianpingensis Deuve & Yu, 1992 (as peterfrankei)
    Carabus (Apotomopterus) sauteri tianpingensis Deuve & Yu, 1992 (as peterfrankei) Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) schuetzei Kleinfeld, 1998
    Carabus (Apotomopterus) schuetzei Kleinfeld, 1998 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) shun gigantothoracis Kleinfeld, 1997
    Carabus (Apotomopterus) shun gigantothoracis Kleinfeld, 1997 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) shun paraschuetzei Kleinfeld, 2002
    Carabus (Apotomopterus) shun paraschuetzei Kleinfeld, 2002 Holotype
    Male+Female adult

    Copyright © Kirschenhofer
    Carabus (Apotomopterus) solidior perangustus Kleinfeld & Schutze, 2003
    Carabus (Apotomopterus) solidior perangustus Kleinfeld & Schutze, 2003 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) songshanicus Kleinfeld, 1994
    Carabus (Apotomopterus) songshanicus Kleinfeld, 1994 Holotype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) tonkinensis pandora Kleinfeld, 2000
    Carabus (Apotomopterus) tonkinensis pandora Kleinfeld, 2000 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) tonkinensis tonkinensis Deuve, 1990
    Carabus (Apotomopterus) tonkinensis tonkinensis Deuve, 1990
    Female adult
    N Viet Nam, Fan-si-pan mt., N seite Cha-pa (=Sapa)
    1600 m, 20-30.4.95; 22.17N 103.44E Siniaev leg.
    Copyright © Alexander Anichtchenko
    Carabus (Apotomopterus) torquatus lushanicus Kleinfeld, 1996
    Carabus (Apotomopterus) torquatus lushanicus Kleinfeld, 1996 Holotype
    Male adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) vitalisi bernardlassallei A.Muller, 2005
    Carabus (Apotomopterus) vitalisi bernardlassallei A.Muller, 2005 Paratype
    Female adult

    Copyright © Boleslav Brezina
    Carabus (Apotomopterus) vogtae artemis Kleinfeld, 2000
    Carabus (Apotomopterus) vogtae artemis Kleinfeld, 2000 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) vogtae menghaiensis Kleinfeld, 2000
    Carabus (Apotomopterus) vogtae menghaiensis Kleinfeld, 2000 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) vogtae mojiangensis Kleinfeld, 2000
    Carabus (Apotomopterus) vogtae mojiangensis Kleinfeld, 2000 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) vogtae paravogtae Kleinfeld, 2000
    Carabus (Apotomopterus) vogtae paravogtae Kleinfeld, 2000 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) vogtae thetis Kleinfeld, 2000
    Carabus (Apotomopterus) vogtae thetis Kleinfeld, 2000 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) yuanshanensis Kleinfeld, 1996
    Carabus (Apotomopterus) yuanshanensis Kleinfeld, 1996 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) yunanensis kassandra Kleinfeld, 1999
    Carabus (Apotomopterus) yunanensis kassandra Kleinfeld, 1999 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) yunanensis leiboensis Kleinfeld, 1999
    Carabus (Apotomopterus) yunanensis leiboensis Kleinfeld, 1999 Holotype Paratype
    Male+Female adult

    Copyright © Kirschenhofer
    Carabus (Carabus) bornianus bornianus Hauser, 1922
    Carabus (Carabus) bornianus bornianus Hauser, 1922 Lectotype
    Male adult

    Copyright © Frank Kleinfeld
    Carabus (Carabus) dreuxioides wushanling Kleinfeld, 1998
    Carabus (Carabus) dreuxioides wushanling Kleinfeld, 1998 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Carabus) emanuelei grabneri Kleinfeld, 2003
    Carabus (Carabus) emanuelei grabneri Kleinfeld, 2003 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Carabus) gressittianus daiyunshan Kleinfeld, 1998
    Carabus (Carabus) gressittianus daiyunshan Kleinfeld, 1998 Holotype
    Male adult

    Copyright © Frank Kleinfeld
    Carabus (Carabus) gressittianus fuscomarginatus Kleinfeld, 1997
    Carabus (Carabus) gressittianus fuscomarginatus Kleinfeld, 1997 Holotype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Carabus) kucerai wuangxianus Kleinfeld, 1998
    Carabus (Carabus) kucerai wuangxianus Kleinfeld, 1998 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Carabus) mianningensis Kleinfeld, 2000
    Carabus (Carabus) mianningensis Kleinfeld, 2000 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Carabus) nanosomus wufengensis Kleinfeld, 1998
    Carabus (Carabus) nanosomus wufengensis Kleinfeld, 1998 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Carabus) oblongior yunlingicus Deuve, 1994
    Carabus (Carabus) oblongior yunlingicus Deuve, 1994
    Male adult
    China, NW Yunnan, Zhongdian env.
    3400m, 5.7.2000
    male, 22 mm
    Copyright © Alexander Anichtchenko
    Carabus (Carabus) pseudolatipennis wenxianensis Deuve, 1992
    Carabus (Carabus) pseudolatipennis wenxianensis Deuve, 1992
    Male adult
    China, S Gansu, Min Shan 2100 m,
    SW Wudu 1.8.2000, V. Patrikeev leg.
    Copyright © Alexander Anichtchenko
    Carabus (Carabus) pseudolatipennis wenxianensis Deuve, 1992
    Carabus (Carabus) pseudolatipennis wenxianensis Deuve, 1992
    Male adult
    male, 20 mm
    Copyright © Boleslav Brezina
    Carabus (Carabus) tianbaoshan Kleinfeld, 1998
    Carabus (Carabus) tianbaoshan Kleinfeld, 1998 Holotype
    Female adult

    Copyright © Frank Kleinfeld
    Carabus (Carabus) xupuensis Kleinfeld, 1998
    Carabus (Carabus) xupuensis Kleinfeld, 1998 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Archicarabus) gotschi gotschi Chaudoir, 1846
    Carabus (Archicarabus) gotschi gotschi Chaudoir, 1846
    Male+Female adult
    Transcaucasia, Armenia,
    Ashotsk distr., Bandivan, 8-21.X.1991 P.V.Kazaryan leg.
    female, 20mm
    Copyright © Ruslan Panin
    Carabus (Archicarabus) nemoralis nemoralis Muller, 1764
    Carabus (Archicarabus) nemoralis nemoralis Muller, 1764
    Male+Female adult
    W Ukraine, NW Podolian,
    Lviv reg., Zolochiv distr., near Vilshanitsya vill., Lysa Gora, 16-30.V.2009 R.Panin leg.
    female, 26 mm
    Copyright © Ruslan Panin
    Carabus (Archicarabus) nemoralis nemoralis Muller, 1764
    Carabus (Archicarabus) nemoralis nemoralis Muller, 1764
    Female adult

    Copyright © Ruslan Panin
    Carabus (Archicarabus) nemoralis prasinotinctus Heyden, 1880
    Carabus (Archicarabus) nemoralis prasinotinctus Heyden, 1880
    Male+Female adult
    AUDE.Pic de Nore, h=1200m.IX.2005, A. Laforgue leg.
    female,25mm/male, 24mm
    Copyright © Ruslan Panin
    Carabus (Archicarabus) nemoralis prasinotinctus Heyden, 1880
    Carabus (Archicarabus) nemoralis prasinotinctus Heyden, 1880
    Male adult
    N Spain, 3 km N Bera de Bidasoa 14.9.2002
    Anichtchenko A. leg.
    long. 25mm
    Copyright © Alexander Anichtchenko
    Carabus (Archicarabus) rossii Dejean, 1826
    Carabus (Archicarabus) rossii Dejean, 1826
    Male adult
    Lazio (Roma), Bosco di
    Nettuno 22.5.1986 C. Iacovone leg.
    male, 23 mm
    Copyright © Alexander Anichtchenko
    Carabus (Archicarabus) steuartii Deyrolle, 1852
    Carabus (Archicarabus) steuartii Deyrolle, 1852
    Male adult
    Portugal, S. Martinho, 27.5.1987
    male, 18 mm
    Copyright © Alexander Anichtchenko
    Carabus (Archicarabus) victor victor Fischer, 1836
    Carabus (Archicarabus) victor victor Fischer, 1836
    Male adult
    Anatolia Or.
    Artvin Heinz leg.
    male, 18 mm
    Copyright © Alexander Anichtchenko
    Carabus (Archicarabus) victor wiedemanni Menetries, 1836
    Carabus (Archicarabus) victor wiedemanni Menetries, 1836
    Male adult
    Anatolia, Kocaeli, tra Kandira e Sile VII.2001
    Solo Claudio leg.
    long. 24mm
    Copyright © Alexander Anichtchenko
    Carabus (Archicarabus) nemoralis nemoralis Muller, 1764
    Carabus (Archicarabus) nemoralis nemoralis Muller, 1764
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
    Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
    Endophallus
    Endophallus fabricii malachiticus
    Copyright © Ruslan Panin
    Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
    Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
    Male adult
    W.Ukraine, Transcarpathian Reg., Rakhov Distr., Chornogora Mt range h=2020m alpine zone, Panin R. leg. 28.VI-9.VIII.2008, male 19 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi rothi Dejean, 1829
    Carabus (Morphocarabus) rothi rothi Dejean, 1829
    Male+Female adult
    Romania
    Banat, Nemeseti h=250m. VI.1989 Lie P. leg.

    photo Solodovnikov I.A.
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) apollo apollo Zolotarev, 1913
    Carabus (Neoplectes) apollo apollo Zolotarev, 1913
    Male adult
    Caucasus, Abchasia

    Copyright © Dmitri Obydov
    Carabus (Neoplectes) apollo phoebus Kurnakov, 1962
    Carabus (Neoplectes) apollo phoebus Kurnakov, 1962
    Male adult
    Abchasia, Kodori, A. Zolotarew leg.
    Photo by D.V. Obydov
    from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Neoplectes) apollo tenebricosus Kurnakov, 1962
    Carabus (Neoplectes) apollo tenebricosus Kurnakov, 1962
    Male adult

    Copyright © Safronov
    Carabus (Pachystus) hungaricus gastridulus Fischer, 1823
    Carabus (Pachystus) hungaricus gastridulus Fischer, 1823
    Male adult
    N. Krym Peninsula,
    near Armyansk, Demidov G. leg.5.V.2006
    male, 28mm
    Copyright © Ruslan Panin
    Carabus (Neoplectes) daphnis askhicus Belousov, 1992
    Carabus (Neoplectes) daphnis askhicus Belousov, 1992 Paratype
    Male adult
    SC-Caucasus, W Georgia, valley of riv. Chkhorodukhu, h=1000-1200m, 14.V.-6.VII.1991, Zamotajlov A.S & Miroshnikov leg.
    male, 29 mm
    photo & collection Panin R.
    Copyright © Ruslan Panin
    Carabus (Neoplectes) daphnis askhicus Belousov, 1992
    Carabus (Neoplectes) daphnis askhicus Belousov, 1992 Paratype
    Male+Female adult
    W Georgia, valley of Chkhorodukhu riv., 1000-1200m
    14.V-6.VII.1991
    male 30mm, female 33mm
    photo & collection Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Carabus (Neoplectes) daphnis askhicus Belousov, 1992
    Carabus (Neoplectes) daphnis askhicus Belousov, 1992 Paratype
    Male adult
    Georgia, env. Martvilli, Askhi plateau
    10-15.7.2001 A.Rubenian leg.
    27 mm
    Copyright © Alexander Anichtchenko
    Carabus (Neoplectes) daphnis daphnis Kurnakov, 1962
    Carabus (Neoplectes) daphnis daphnis Kurnakov, 1962
    Male adult
    Georgia, Mingrelia, Lecheha vallis, Tehuri superior
    Caucasus centr., 1200m, 27.6-7.7.1989 Retezar I. leg.
    male, 28 mm
    Copyright © Alexander Anichtchenko
    Carabus (Neoplectes) edithae exedithae Gottwald, 1985
    Carabus (Neoplectes) edithae exedithae Gottwald, 1985
    Male adult
    Karachaevo-Cherkessia, low cource of Daut river,
    16-20.V.2009 Anichtchenko A. leg

    Copyright © Alexander Anichtchenko
    Carabus (Neoplectes) edithae exedithae Gottwald, 1985
    Carabus (Neoplectes) edithae exedithae Gottwald, 1985
    Female adult
    Karachaevo-Cherkessia, Zelenchukskii distr.,
    near Bukovo vill., 8-13.VII.2007, S.Dementiev leg.

    Copyright © Dmitri Obydov
    Carabus (Neoplectes) edithae kabardensis Kurnakov, 1970
    Carabus (Neoplectes) edithae kabardensis Kurnakov, 1970
    Female adult
    Kabardino-Balkaria, Skalisty Mt Ridge, 1800 m,
    20.VII.1996. Andreev leg.
    Photo by D.V. Obydov

    Copyright © Dmitri Obydov
    Carabus (Archiplectes) jason Semenov, 1898
    Carabus (Archiplectes) jason Semenov, 1898
    Male adult
    Caucasus, W Georgia, Svanetia, upper reaches of riv. Shoudidi, 1700-2100m, 20-21.V.1986
    leg. A. Zamotajlov
    Copyright © Sunvas
    Carabus (Neoplectes) jason Semenov, 1898
    Carabus (Neoplectes) jason Semenov, 1898
    Male adult
    Abkhasia, up river Shaudidi, 21.V - 2.VII.1986, Zamotailov A. leg.
    Photo by D.V. Obydov

    Copyright © Dmitri Obydov
    Carabus (Neoplectes) juenthneri atchibachi Zamotajlov, 1991
    Carabus (Neoplectes) juenthneri atchibachi Zamotajlov, 1991 Paratype
    Female adult
    Abkhasia,
    Valley of river Bzyb near Mt. Achibakhu. h=1500m., A.S. Zamotajlov leg.
    female: 33mm, 31mm
    coll. R.Panin
    Copyright © Ruslan Panin
    Carabus (Neoplectes) juenthneri atchibachi Zamotajlov, 1991
    Carabus (Neoplectes) juenthneri atchibachi Zamotajlov, 1991 Paratype
    Female adult
    W Caucas, Abkhasia, Valley of river Bzyb near Mt. Achibakhu. h=1500m.,
    24.IV.-6.VI.1991 A.S. Zamotajlov leg.
    coll. Anichtchenko
    Copyright © Alexander Anichtchenko
    Carabus (Neoplectes) juenthneri atchibachi Zamotajlov, 1991
    Carabus (Neoplectes) juenthneri atchibachi Zamotajlov, 1991 Paratype
    Male adult
    W Caucas, Abkhasia, Valley of river Bzyb near Mt. Achibakhu. h=1500m.,
    24.IV.-6.VI.1991 A.S. Zamotajlov leg.
    coll. Anichtchenko
    Copyright © Alexander Anichtchenko
    Carabus (Neoplectes) juenthneri atchibachi Zamotajlov, 1991
    Carabus (Neoplectes) juenthneri atchibachi Zamotajlov, 1991 Paratype
    Male adult
    W Caucas, Abkhasia, Valley of river Bzyb near Mt. Achibakhu. h=1500m.,
    24.IV.-6.VI.1991 A.S. Zamotajlov leg.
    coll. Anichtchenko
    Copyright © Alexander Anichtchenko
    Carabus (Megodontus) croaticus croaticus Dejean, 1826
    Carabus (Megodontus) croaticus croaticus Dejean, 1826
    Male adult
    YU- Slovenija, Krim 1000 m, 30.10.1975 Drovenik leg.
    Copyright © Alexander Anichtchenko
    Carabus (Megodontus) caelatus caelatus Fabricius, 1801
    Carabus (Megodontus) caelatus caelatus Fabricius, 1801
    Male adult
    Bosnia, Trebevic, Vidikovac-Brus, 1200m
    21.11.1985 Mihljevic B. leg.
    Copyright © Alexander Anichtchenko
    Carabus (Megodontus) bonvouloiri bonvouloiri Chaudoir, 1863
    Carabus (Megodontus) bonvouloiri bonvouloiri Chaudoir, 1863
    Female adult
    Anatolia Orientale, Macka-Zigana VIII.1981
    Copyright © Alexander Anichtchenko
    Carabus (Megodontus) avinovi Semenov & Znojko, 1932
    Carabus (Megodontus) avinovi Semenov & Znojko, 1932
    Male adult
    Russia, Sakhalin isl., Susunaiskyi rng.,
    Chehov Mt., 400m, 19.6.1990 Berlov E. leg.
    female, 22 mm
    photo Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Carabus (Neoplectes) juenthneri avadcharensis Kurnakov, 1970 (loc.typ.)
    Carabus (Neoplectes) juenthneri avadcharensis Kurnakov, 1970 (loc.typ.)
    Male+Female adult
    Abkhasia, near Avadhara, Site Pyv h~1850m, 43.490730 40.686433, 1.VI-12.VII.2009, Zamotajlov A.S., Bondarenko A.S. leg.
    L=27mm

    Copyright © Ruslan Panin
    Carabus (Neoplectes) juenthneri avadcharensis Kurnakov, 1970
    Carabus (Neoplectes) juenthneri avadcharensis Kurnakov, 1970
    Male adult
    Abkhasia, 7,4km NE from Ritza lake, 1,5 km N Kutykuh vill.,
    1635 m, 19-25.VII.2008, A. Vlasenko leg.
    Copyright © Dmitri Obydov
    Carabus (Neoplectes) juenthneri dsychvensis Gottwald, 1985
    Carabus (Neoplectes) juenthneri dsychvensis Gottwald, 1985
    Male adult
    Abchasia, N Dsykhwa, coll. A. Zolotarew
    Photo by D.V. Obydov
    Copyright © Dmitri Obydov
    Carabus (Neoplectes) juenthneri juenthneri Reitter, 1899
    Carabus (Neoplectes) juenthneri juenthneri Reitter, 1899
    Female adult

    Copyright © Alexander Anichtchenko
    Carabus (Neoplectes) koltzei faunulus Kurnakov, 1970
    Carabus (Neoplectes) koltzei faunulus Kurnakov, 1970
    Female adult
    Abchasia, Akarmara, 6500 f, A. Zolotarew leg.
    Photo by D.V. Obydov

    Copyright © Dmitri Obydov
    Carabus (Neoplectes) koltzei faunus Kurnakov, 1970
    Carabus (Neoplectes) koltzei faunus Kurnakov, 1970
    Male adult
    Abkhazia, Kadorsky Mt Ridge, 1000 m, Chlou, 22.V.1993, A. Evstigneev leg.
    Photo by D.V. Obydov

    Copyright © Dmitri Obydov
    Carabus (Neoplectes) koltzei kobachidzei Kurnakov, 1970
    Carabus (Neoplectes) koltzei kobachidzei Kurnakov, 1970
    Female adult
    Caucasus, E Abkhasia, SW slopes of Akiba mt. rng., 1200m, 7.V.-2.VII.1991<br /> Zamotajlov & Miroshnikov leg.
    Copyright © Sunvas
    Carabus (Neoplectes) koltzei kobachidzei Kurnakov, 1970
    Carabus (Neoplectes) koltzei kobachidzei Kurnakov, 1970
    Female adult
    Caucasus, E Abkhasia, SW slopes of Akiba mt. rng., 1200m, 7.V.-2.VII.1991
    Zamotajlov & Miroshnikov leg.
    Copyright © Sunvas
    Carabus (Neoplectes) koltzei kobachidzei Kurnakov, 1970
    Carabus (Neoplectes) koltzei kobachidzei Kurnakov, 1970
    Male+Female adult
    Caucasus, E Abkhasia, SW slope of Akiba mt.rng., 1200m, 7.V.-2.VII.1991 Zamotajlov & Miroshnikov leg.
    male 37mm, female 38mm
    photo & collection Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Carabus (Archiplectes) koltzei Rost, 1889
    Carabus (Archiplectes) koltzei Rost, 1889
    Male adult
    Abkhasia, Habyu 3.7.2002
    Ivliev P. leg.
    32 mm
    Copyright © Alexander Anichtchenko
    Carabus (Neoplectes) koltzei koltzei Rost, 1889
    Carabus (Neoplectes) koltzei koltzei Rost, 1889
    Male adult
    Abkhasia, Tsebeldy, 23.V.1986, A.Zamotailov leg.
    Photo by D.V. Obydov

    Copyright © Dmitri Obydov
    Carabus (Neoplectes) koltzei lennoni Gottwald, 1985
    Carabus (Neoplectes) koltzei lennoni Gottwald, 1985
    Male adult
    Abkhasia, Amtkel lake, 22.V - 4.VII.1986, A.Zamotailov leg.
    Photo by D.V. Obydov
    from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Neoplectes) daphnis daphnis Kurnakov, 1962
    Carabus (Neoplectes) daphnis daphnis Kurnakov, 1962
    Endophallus
    Georgia, Mingrelia, Lecheha vallis, Tehuri superior
    Caucasus centr., 1200m, 27.6-7.7.1989 Retezar I. leg.
    Copyright © Alexander Anichtchenko
    Carabus (Neoplectes) daphnis askhicus Belousov, 1992
    Carabus (Neoplectes) daphnis askhicus Belousov, 1992
    Endophallus
    Georgia, env. Martvilli, Askhi plateau
    10-15.7.2001 A.Rubenian leg.
    Copyright © Alexander Anichtchenko
    Carabus (Neoplectes) daphnis askhicus Belousov, 1992
    Carabus (Neoplectes) daphnis askhicus Belousov, 1992
    Endophallus
    W Georgia, valley of Chkhorodukhu riv., 1000-1200m
    14.V-6.VII.1991
    Copyright © Alexander Anichtchenko
    Carabus (Neoplectes) daphnis askhicus Belousov, 1992
    Carabus (Neoplectes) daphnis askhicus Belousov, 1992 Paratype
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Neoplectes) juenthneri avadcharensis Kurnakov, 1970
    Carabus (Neoplectes) juenthneri avadcharensis Kurnakov, 1970
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Cratophyrtus) turcosinensis medvedevi Kryzhanovskij, 1968
    Carabus (Cratophyrtus) turcosinensis medvedevi Kryzhanovskij, 1968
    Female adult
    S Kazakhstan, NW slope of Ugamskii mt.rng., 3 km W Sairamskii peak, 2200m
    tributary of Karzhantau riv., 15.6.2004 Matalin A. leg.
    20 mm
    Copyright © Alexander Anichtchenko
    Carabus (Neoplectes) komarowi komarowi Reitter, 1882
    Carabus (Neoplectes) komarowi komarowi Reitter, 1882
    Female adult
    Centr. Caucasus, Svanetia, coll. A. Zolotarew
    Photo by D.V. Obydov
    from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Neoplectes) kratkyi felicitanus Reitter, 1893
    Carabus (Neoplectes) kratkyi felicitanus Reitter, 1893
    Male adult
    Russia, N Caucasus, Karachaevo-Cherkessia 2200m, Bolshoi Pzizer mt. 8.8.2002
    25 mm
    Copyright © Alexander Anichtchenko
    Carabus (Neoplectes) kratkyi felicitanus Reitter, 1893 (as porcellus)
    Carabus (Neoplectes) kratkyi felicitanus Reitter, 1893 (as porcellus)
    Male adult
    NW Caucasus, Resp. Adygea, Mt.Tybga, 28.VI.1997
    male,20mm
    Copyright © Ruslan Panin
    Carabus (Neoplectes) kratkyi felicitanus Reitter, 1893 (as porcellus)
    Carabus (Neoplectes) kratkyi felicitanus Reitter, 1893 (as porcellus)
    Male adult
    Prov. Kuban, mt. Tybga, 1300 m, VI.1913, Shaposhnikov leg.

    Copyright © Dmitri Obydov
    Carabus (Neoplectes) kratkyi felicitanus Reitter, 1893
    Carabus (Neoplectes) kratkyi felicitanus Reitter, 1893
    Female adult
    Rusia, Caucas, Dzhuga mt., 2200 m, 24.VI. - 22.VIII.1986, Zamotajlov A. leg.

    Copyright © Dmitri Obydov
    Carabus (Neoplectes) kratkyi Ganglbauer, 1890
    Carabus (Neoplectes) kratkyi Ganglbauer, 1890
    Male+Female adult
    W Caucas, 1900m, Khatsavitaya Mt., 4.VI-13.VII.1994
    A.Miroshnikov leg.
    male 25mm, female 26mm
    photo & collection Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Carabus (Neoplectes) kratkyi Ganglbauer, 1890
    Carabus (Neoplectes) kratkyi Ganglbauer, 1890
    Male adult
    Russia, NW-Caucasus: <br /> Krasnodar reg., Mostovskoy distr., Khatsavitaja Mt. h=1900m, 4.VI.-13.VII.1994, A. Miroshnikov leg.<br /> male, 26mm<br /> photo Panin R./det.Solodovnikov I.A.
    Copyright © Ruslan Panin
    Carabus (Neoplectes) kratkyi Ganglbauer, 1890
    Carabus (Neoplectes) kratkyi Ganglbauer, 1890
    Male+Female adult
    W Caucas Mts., 2500m, Mt. Yatyrgvarta,
    25.VI-12.VIII.1992 Shchurov V. leg.
    male 20mm, female 21mm
    Copyright © Alexander Anichtchenko
    Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890
    Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890
    Female adult
    NW-Caucasus, Russia,
    Karacaj-Cherkessija, Abishira-Akhuba range h=2650m, upper Kyafar-Agur Riv., 4-9.VIII.2009, Solodovnikov I.A. leg.
    female, 23 mm
    Copyright © Ruslan Panin
    Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890
    Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890
    Male adult
    NW-Caucasus: Karacaj-Cherkessija,
    Abishira–Akhuba Mnt. Range h=2000-2650m, Kyafar-Agur Riv., 14–17.VII.2009, Fominykh D. leg.
    male, 22mm
    Copyright © Ruslan Panin
    Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890
    Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890
    Male adult
    Caucasus, Kuban, Meyer-Darcis-Patria

    Copyright © Dmitri Obydov
    Carabus (Neoplectes) kratkyi solodovnikovi Zamotajlov, 1992
    Carabus (Neoplectes) kratkyi solodovnikovi Zamotajlov, 1992
    Male adult
    W. Caucasus, M. Bambak Mt., VI.1931, Vvedensky leg.
    from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Archiplectes) plasoni dalensis Rost, 1893
    Carabus (Archiplectes) plasoni dalensis Rost, 1893
    Male adult
    Abkhasia, near Yu. Ptysh vill.
    14.7.1986 Zamotailov A. leg.
    31 mm
    Copyright © Alexander Anichtchenko
    Carabus (Archiplectes) plasoni dalensis Rost, 1893
    Carabus (Archiplectes) plasoni dalensis Rost, 1893
    Male adult
    Caucasus merid., Dal-Suchumi, N.S. Brianskyj leg.
    from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Archiplectes) plasoni plasoni Ganglbauer, 1886
    Carabus (Archiplectes) plasoni plasoni Ganglbauer, 1886
    Female adult
    Gruzia, Centr. Caucasus, Svanetia, coll. A. Zolotarew
    det. D.V. Obydov
    from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Neoplectes) polychrous Rost, 1892
    Carabus (Neoplectes) polychrous Rost, 1892
    Male adult
    Abkhasia, Gudaut distr., near Huap vill., 600 m, 9.VI.1990, M. Danilevskyi leg. det. D.V. Obydov
    Copyright © Dmitri Obydov
    Carabus (Neoplectes) rousianus rousianus Gottwald, 1985
    Carabus (Neoplectes) rousianus rousianus Gottwald, 1985
    Male adult
    Abkhasia, 7,6 km NE Pitsunda, 2,1 km NNE Ryapsh riv mouth, Ryapsh canyon h=60 m, 43°10N 40°25E, 25-27.VII.2008, A.Vlasenko leg.<br /> det. D.V. Obydov
    Copyright © Dmitri Obydov
    Carabus (Archiplectes) prometheus miroshnikovi Zamotajlov, 1990
    Carabus (Archiplectes) prometheus miroshnikovi Zamotajlov, 1990
    Female adult
    NW-Caucasus, Krasnodar reg., Mostovskoy distr., Site Val , h=1800m. 07-08.VII.2008, D.Fominyh leg., 35mm; det. I. Solodovnikov 2023
    Copyright © Ruslan Panin
    Carabus (Archiplectes) prometheus zolotarevi Zamotajlov, 1988
    Carabus (Archiplectes) prometheus zolotarevi Zamotajlov, 1988
    Male adult
    Paratypus, male Prov. Kuban distr., Maikop, Laba Min., Ch. Shaposhnikov leg.
    from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Archiplectes) prometheus zolotarevi Zamotajlov, 1988
    Carabus (Archiplectes) prometheus zolotarevi Zamotajlov, 1988 Paratype
    Male adult
    Paratypus, male - Fl. Min. Laba, VII.1918, Shaposhnikov leg.
    from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Archiplectes) prometheus basilianus Starck, 1890
    Carabus (Archiplectes) prometheus basilianus Starck, 1890
    Male adult
    Russia, W Caucas, Port-Artur Mt. 22.VI.1986, A.Zamotajlov leg.
    det. D.V. Obydov
    from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Neoplectes) prometheus dzhugensis Zamotajlov, 1988
    Carabus (Neoplectes) prometheus dzhugensis Zamotajlov, 1988 Paratype
    Female adult
    Paratypus, female
    Prov. Kuban circ., mt. Dzhuga (Tigeni), 27.VI.1911.

    Copyright © Dmitri Obydov
    Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990
    Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990
    Male+Female adult
    W.Caucasus,
    Psebai env., Nikitino, 2.V-3.VII.1993, leg. A.S.Zamotajlov
    female 33mm
    Copyright © Ruslan Panin
    Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990 (as hephaestus)
    Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990 (as hephaestus)
    Male adult
    Paratypus Carabus (Archiplectes) hephaestus
    Maikop, Ust. Urusht, 26.IV.1910

    Copyright © Dmitri Obydov
    Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990
    Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990
    Female adult
    W Caucasus, Krasnodar reg., Maikop distr.
    Nikel, 1.7.1989
    38 mm
    Copyright © Alexander Anichtchenko
    Carabus (Archiplectes) prometheus pschadensis Zamotajlov, 1990
    Carabus (Archiplectes) prometheus pschadensis Zamotajlov, 1990
    Female adult
    NW Caucas, 5 km N from Pshada vill., 25.04.1989, Smorodinov leg.
    specimen from collection of M.Shestopalov (Moscow).
    det. D.V. Obydov
    from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Neoplectes) prometheus prometheus Reitter, 1887
    Carabus (Neoplectes) prometheus prometheus Reitter, 1887
    Male adult
    NW-Caucasus,Krasnodar reg., Apsheron Distr., Lagonaki Mt., near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominykh leg.
    L= 42mm.
    Copyright © Ruslan Panin
    Carabus (Neoplectes) prometheus prometheus Reitter, 1887 (as zugmayeriae)
    Carabus (Neoplectes) prometheus prometheus Reitter, 1887 (as zugmayeriae)
    Male adult
    W. Caucasus, Maikop, 20.VI.1967., A. Bogachev leg.
    det. D.V. Obydov

    Copyright © Dmitri Obydov
    Carabus (Neoplectes) prometheus prometheus Reitter, 1887
    Carabus (Neoplectes) prometheus prometheus Reitter, 1887
    Male+Female adult
    NW Caucasus, Russia,
    Krasnodar reg.,Krasnoarmeisk distr., Krasny Les, near Podlesnyi vill.,17-22.IV.2008, D.D.Fominykh leg.
    female, 40mm
    Copyright © Ruslan Panin
    Carabus (Archiplectes) prometheus prometheus Reitter, 1887 (as nubicola)
    Carabus (Archiplectes) prometheus prometheus Reitter, 1887 (as nubicola)
    Male+Female adult
    W Caucasus, Adygea,
    h~1700m. upper res. Riv. Kurdzhips, 11-21.VIII.2007, I.A.Solodovnikov leg./det.
    L=32mm.
    Copyright © Ruslan Panin
    Carabus (Neoplectes) prometheus prometheus Reitter, 1887
    Carabus (Neoplectes) prometheus prometheus Reitter, 1887
    Male adult
    Krasnodar distr., Mezmai
    20.6.1989
    38 mm
    Copyright © Alexander Anichtchenko
    Carabus (Neoplectes) protensus olegi Belousov, 1992
    Carabus (Neoplectes) protensus olegi Belousov, 1992 Paratype
    Male adult
    Caucasus, W Georgia, Svanetia, right bank of riv. Magana, 800m, 12.V.-4.VII.1991, Zamotajlov & Miroshnikov leg.
    male, 31mm
    photo and collection Anichtchenko A.
    Copyright © Alexander Anichtchenko
    Carabus (Neoplectes) protensus protensoides Novotny & Vorisek, 1988
    Carabus (Neoplectes) protensus protensoides Novotny & Vorisek, 1988
    Male adult
    Georgia, Martvilli env., Akshi plateau 10-15.7.2001
    Rubenian leg.
    32 mm
    Copyright © Alexander Anichtchenko
    Carabus (Neoplectes) reitteri gagrinus Starck, 1894
    Carabus (Neoplectes) reitteri gagrinus Starck, 1894
    Female adult
    Abkhasia, near Gagry, 16.V.1992, A. Evstigneev leg.
    det. D.V. Obydov
    Copyright © Dmitri Obydov
    Carabus (Neoplectes) reitteri laevisternis Starck, 1890
    Carabus (Neoplectes) reitteri laevisternis Starck, 1890
    Male adult
    Abkhasia,W.slope of Gagrinsky Mt., Krepostnaya River source, h=1325m., A.S.Zamotajlov leg. 27.VII.2000
    male, 29 mm
    Copyright © Ruslan Panin
    Carabus (Neoplectes) reitteri laevisternis Starck, 1890
    Carabus (Neoplectes) reitteri laevisternis Starck, 1890
    Male adult
    W-Caucasus:Sochi env., Aibga Mt. h=2000m., 06.VII.2000
    male, 29mm
    Copyright © Ruslan Panin
    Carabus Neoplectes) reitteri reitteri Retowski, 1885
    Carabus Neoplectes) reitteri reitteri Retowski, 1885
    Female adult
    NW Caucasus, Adygeya, Fischt-Oshten Mts. base, h=1800 m.<br /> 20.07.2006, D.Safronov leg.
    Copyright © Safronov
    Carabus (Neoplectes) reitteri reitteri Retowski, 1885 (as achunensis)
    Carabus (Neoplectes) reitteri reitteri Retowski, 1885 (as achunensis)
    Male adult
    W. Caucasus, Sochi Distr., Chernomoretz Vill. env.,
    15.VII.1992, P. Gorbachev leg.
    det. D.V. Obydov

    Copyright © Dmitri Obydov
    Carabus (Neoplectes) reitteri laevisternis Starck, 1890
    Carabus (Neoplectes) reitteri laevisternis Starck, 1890
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Neoplectes) prometheus prometheus Reitter, 1887 (as nubicola Zolotarev, 1913)
    Carabus (Neoplectes) prometheus prometheus Reitter, 1887 (as nubicola Zolotarev, 1913)
    Endophallus
    Endophallus prometheus prometheus (as nubicola Zolotarev, 1913)
    Copyright © Ruslan Panin
    Carabus (Neoplectes) prometheus prometheus Reitter, 1887
    Carabus (Neoplectes) prometheus prometheus Reitter, 1887
    Endophallus
    Endophallus prometheus prometheus Reitter, 1887
    Copyright © Ruslan Panin
    Carabus (Neoplectes) prometheus prometheus Reitter, 1887
    Carabus (Neoplectes) prometheus prometheus Reitter, 1887
    Endophallus
    aedeagus
    Copyright © Ruslan Panin
    Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990
    Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990
    Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990
    Endophallus
    aedeagus miroshnikovi Psebai env., Nikitino
    Copyright © Ruslan Panin
    Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890
    Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890
    Endophallus
    Endophallus
    Copyright © Ruslan Panin
    Carabus (Neoplectes) satyrus besleticus Kurnakov, 1972 (as gaskoi)
    Carabus (Neoplectes) satyrus besleticus Kurnakov, 1972 (as gaskoi)
    Male adult
    USSR, Caucasus occid, Abchasia, Suchumi, vallis Kelasuri, 100m 25.6-17.7.1977 Retezar I. leg. 36 mm
    Copyright © Alexander Anichtchenko
    Carabus (Neoplectes) satyrus mtsaranus Kurnakov, 1972
    Carabus (Neoplectes) satyrus mtsaranus Kurnakov, 1972
    Male adult
    Abkhasia, Bzhybskii Mt.rng., near Mtsara vill., 16.VI.1991, S.Gusev leg.
    det. D.V. Obydov

    Copyright © Dmitri Obydov
    Carabus (Neoplectes) satyrus napraensis Belousov & Zamotajlov, 1993
    Carabus (Neoplectes) satyrus napraensis Belousov & Zamotajlov, 1993 Paratype
    Male adult
    Abkhasia, Bzybian Mt. Range near Mt. Chibzharga., h=1900-2000m., A.S. Zamotajlov leg.
    male, 28mm
    photo & coll.Panin R.
    Copyright © Ruslan Panin
    Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972
    Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972
    Male adult
    Abkhasia, Habyu vill.,2.7.2002 Ibliev leg.<br /> 35 mm
    Copyright © Alexander Anichtchenko
    Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972
    Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972
    Male adult
    Abkhasia, Habyu vill.<br /> 2.7.2002 Ibliev leg.<br /> 35 mm
    Copyright © Alexander Anichtchenko
    Carabus (Neoplectes) starcki Heyden, 1885
    Carabus (Neoplectes) starcki Heyden, 1885
    Male+Female adult
    NW-Caucasus,
    Krasnodar reg., Mt. Aishkho, Malaya Laba River, h=2200m., 14.VII.2008, D.Fominyh leg.
    female-21mm
    Copyright © Ruslan Panin
    Carabus (Neoplectes) starcki Heyden, 1885
    Carabus (Neoplectes) starcki Heyden, 1885
    Male adult
    Caucasian Nat. Park, S slope
    Pseashkho mt. 2500m 21.7.1975, Ovtsharenko leg.
    20 mm
    Copyright © Alexander Anichtchenko
    Carabus (Neoplectes) starcki Heyden, 1885
    Carabus (Neoplectes) starcki Heyden, 1885
    Male adult
    NW Caucasus, Adygeya, Pshekho-Su Mt. env., h=1500 m., 21.07.2006, D.Safronov leg.
    Copyright © Safronov
    Carabus (Neoplectes) starcki Heyden, 1885 (as parvulus)
    Carabus (Neoplectes) starcki Heyden, 1885 (as parvulus)
    Male adult
    Prov. Kuban, mt. Aischa, VI.1906, Shaposhnikov leg.
    det. D.V. Obydov

    Copyright © Dmitri Obydov
    Carabus (Neoplectes) starckianus adelaidae Starck, 1889
    Carabus (Neoplectes) starckianus adelaidae Starck, 1889
    Male adult
    Russia, Caucasus, Chvizhepse River Val., 27.V.1994, P. Gorbachev leg.
    Photo from www.zin.ru
    det. D.V. Obydov
    Copyright © Dmitri Obydov
    Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988 (as shchessiensis)
    Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988 (as shchessiensis) Paratype
    Male+Female adult
    NW Caucasus,
    Krasnodar reg., Tuapsinskyi distr. Shessi Mt.
    30.V.-25.VI.1993 Mt.Shessi, h=1800m., Imura & A.S.Zamotajlov, 1993 leg.
    female, 29mm
    Copyright © Ruslan Panin
    Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988
    Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988
    Female adult
    Russia,W Caucasus,
    Krasnodar reg.,Chernogorie mts.,Chernogor mt. h=1700-1800m. 23.V.-30.VII.1999. A.S.Zamotajlov leg.
    female 29mm.
    Copyright © Ruslan Panin
    Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988
    Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988
    Male adult
    West. Caucasus, Shakhe Valley, 1.VI.1989, A. Shamaev leg.
    det. D.V. Obydov

    Copyright © Dmitri Obydov
    Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
    Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
    Male adult
    NW-Caucasus, Russia,
    Krasnodar reg., Apsheron Distr., Lagonaki Mt. ,near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominykh leg.
    male, 28mm
    Copyright © Ruslan Panin
    Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
    Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
    Male+Female adult
    NW-Caucasus, Russia,
    Krasnodar reg.,the right bank of Mzymta river, Engelmanovs glades, h=1200m., A.S.Zamotajlov leg.
    female-30mm\male-28mm
    Copyright © Ruslan Panin
    Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
    Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
    Female adult

    Copyright © Safronov
    Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
    Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
    Male adult
    Rusia, W Caucasus, Sochi distr., Aishkha mts.
    900m, 15.7.2003 Gorbachev leg.
    30 mm
    Copyright © Alexander Anichtchenko
    Carabus (Neoplectes) starckianus starckianus Ganglbauer, 1886
    Carabus (Neoplectes) starckianus starckianus Ganglbauer, 1886
    Male adult
    Russia, Krasnodar reg., 6 km NE Soloniki vill.,
    Sochi Nat.Park, 28-30.V.2009 Anichtchenko A. leg.

    Copyright © Alexander Anichtchenko
    Carabus (Neoplectes) starckianus starckianus Ganglbauer, 1886
    Carabus (Neoplectes) starckianus starckianus Ganglbauer, 1886
    Female adult
    W Caucas, Sochi distr., near Lazorevskoe vill., 9.06.1993, P.Gorbachev leg.
    det. D.V. Obydov


    Copyright © Dmitri Obydov
    Carabus (Neoplectes) starckianus starckianus Ganglbauer, 1886 (as phlagochensis)
    Carabus (Neoplectes) starckianus starckianus Ganglbauer, 1886 (as phlagochensis)
    Female adult
    Caucasus, Sotschi, Flagoch.
    det. D.V. Obydov
    from www.zin.ru

    Copyright © Dmitri Obydov
    Carabus (Neoplectes) starckianus theseus Briansky, 1910
    Carabus (Neoplectes) starckianus theseus Briansky, 1910
    Male+Female adult
    NW-Caucasus, Russia,
    Krasnodar reg.,Tuapse distr., Mt.Semashkho 04.V.2008, D.Fominyh leg.
    female, 32mm
    Copyright © Ruslan Panin
    Carabus (Neoplectes) starckianus theseus Briansky, 1910
    Carabus (Neoplectes) starckianus theseus Briansky, 1910
    Male adult
    Tuapse distr., Anastasievka vill., Semiglavaya mt.
    riv. Pshekaho 20.7.1995
    30 mm
    Copyright © Alexander Anichtchenko
    Carabus (Neoplectes) starckianus theseus Briansky, 1910
    Carabus (Neoplectes) starckianus theseus Briansky, 1910
    Female adult
    Tuapse distr., Anastasievka vill., Semiglavaya mt.
    bass. Pshekaho riv. 20.7.1995
    female, 30 mm
    Copyright © Alexander Anichtchenko
    Carabus (Aristocarabus) viridifossulatus viridifossulatus Fairmaire, 1887
    Carabus (Aristocarabus) viridifossulatus viridifossulatus Fairmaire, 1887
    Female adult
    35mm
    Copyright © Boleslav Brezina
    Carabus (Aristocarabus) viridifossulatus viridifossulatus Fairmaire, 1887
    Carabus (Aristocarabus) viridifossulatus viridifossulatus Fairmaire, 1887
    Male adult
    29 mm
    Copyright © Boleslav Brezina
    Carabus (Aristocarabus) viridifossulatus aleatus Cavazzuti, 2003
    Carabus (Aristocarabus) viridifossulatus aleatus Cavazzuti, 2003
    Male adult
    26 mm
    Copyright © Boleslav Brezina
    Carabus (Aristocarabus) viridifossulatus gonggashanensis Deuve, 1992
    Carabus (Aristocarabus) viridifossulatus gonggashanensis Deuve, 1992
    Male adult
    32 mm
    Copyright © Boleslav Brezina
    Carabus (Aristocarabus) viridifossulatus jiajiensis Deuve, 1998
    Carabus (Aristocarabus) viridifossulatus jiajiensis Deuve, 1998
    Male adult
    27 mm
    Copyright © Boleslav Brezina
    Carabus (Aristocarabus) viridifossulatus kvetoslavae Sehnal, Farkac & Hackel, 2006
    Carabus (Aristocarabus) viridifossulatus kvetoslavae Sehnal, Farkac & Hackel, 2006 Holotype Paratype
    Female adult
    36 mm
    Copyright © Boleslav Brezina
    Carabus (Aristocarabus) viridifossulatus kvetoslavae Sehnal, Farkac & Hackel, 2006
    Carabus (Aristocarabus) viridifossulatus kvetoslavae Sehnal, Farkac & Hackel, 2006
    Male adult
    30 mm
    Copyright © Boleslav Brezina
    Carabus (Aristocarabus) viridifossulatus kvetoslavae Sehnal, Farkac & Hackel, 2006
    Carabus (Aristocarabus) viridifossulatus kvetoslavae Sehnal, Farkac & Hackel, 2006 Paratype
    Female adult
    35 mm
    Copyright © Boleslav Brezina
    Carabus (Aristocarabus) viridifossulatus phamianus Deuve, 1990
    Carabus (Aristocarabus) viridifossulatus phamianus Deuve, 1990
    Male adult
    27 mm
    Copyright © Boleslav Brezina
    Carabus (Aristocarabus) viridifossulatus seticollis Deuve & Mourzine, 2000
    Carabus (Aristocarabus) viridifossulatus seticollis Deuve & Mourzine, 2000
    Male adult
    28 mm
    Copyright © Boleslav Brezina
    Carabus (Aulonocarabus) canaliculatus rufinus Beheim & Breuning, 1943
    Carabus (Aulonocarabus) canaliculatus rufinus Beheim & Breuning, 1943
    Female adult

    Copyright © Safronov
    Carabus (Aulonocarabus) canaliculatus canaliculatus Adams, 1812
    Carabus (Aulonocarabus) canaliculatus canaliculatus Adams, 1812
    Male adult

    Copyright © Safronov
    Carabus (Aulonocarabus) canaliculatus jankowskiellus Deuve, 1991
    Carabus (Aulonocarabus) canaliculatus jankowskiellus Deuve, 1991
    Female adult
    CHINA, Jilin prov., Hongshi
    forest plant
    Copyright © Anton Menstschikov
    Carabus (Aulonocarabus) careniger careniger Chaudoir, 1863
    Carabus (Aulonocarabus) careniger careniger Chaudoir, 1863
    Male adult
    S Primorie, Chernye mts.,
    Poima riv., 30.07.- 6.08.1999 Anichtchenko A. leg
    male, 25 mm
    Copyright © Alexander Anichtchenko
    Carabus (Aulonocarabus) gossarei imanensis Lafer, 1989
    Carabus (Aulonocarabus) gossarei imanensis Lafer, 1989
    Male adult
    Primorskii reg., Chug. distr., Oblachnaya mt.,
    Zabytyi, 11.7.1977 Kuznetsov V. leg
    male, 19 mm
    Copyright © Alexander Anichtchenko
    Carabus (Aulonocarabus) kolymensis kodarensis Brinev, 1997
    Carabus (Aulonocarabus) kolymensis kodarensis Brinev, 1997
    Male adult
    NE Chita reg., Kodar mt. rng., up of Chara riv.,
    15 km N Leprindo, 2200m 5-15.7.2001 Brinev A. leg.
    Copyright © Alexander Anichtchenko
    Carabus (Aulonocarabus) kolymensis viridicupreior Deuve & Mourzine, 1993
    Carabus (Aulonocarabus) kolymensis viridicupreior Deuve & Mourzine, 1993
    Male adult
    SE Yakutia, Cherskii Mt.rng., Garmychan Mts., 20 km NNE Sasyr,
    Kalgar riv., 7.VIII.1998 Nogovitsyna leg.
    Copyright © Alexander Anichtchenko
    Carabus (Aulonocarabus) kolymensis viridicupreior Deuve & Mourzine, 1993
    Carabus (Aulonocarabus) kolymensis viridicupreior Deuve & Mourzine, 1993
    Female adult
    10.VII.2001 NE RUSSIA, Magadan, near Karmaken vill
    Copyright © Nil Belonogov
    Carabus (Aulonocarabus) kurilensis kurilensis Lapouge, 1913
    Carabus (Aulonocarabus) kurilensis kurilensis Lapouge, 1913
    Male adult
    Rusia, Sakhalin isl., 44 kordon
    6.08.1991
    male, 20mm
    Copyright © Alexander Anichtchenko
    Carabus (Aulonocarabus) kurilensis pseudodiamesus Ivanovs, 1994
    Carabus (Aulonocarabus) kurilensis pseudodiamesus Ivanovs, 1994
    Male adult
    Russia, Sakhalin isl., Susunaiskii mg.
    Chehov mt., 19.6.1990 Berlov E. leg.
    male, 22 mm
    Copyright © Alexander Anichtchenko
    Carabus (Aulonocarabus) truncaticollis polaris Poppius, 1905
    Carabus (Aulonocarabus) truncaticollis polaris Poppius, 1905
    Male adult
    Arctic Russia, NW Siberia,
    Yamal, 230 N from Labytnangi, Panta-Yakha Riv., 15-29.VI.2002
    male,18mm.
    Copyright © Ruslan Panin
    Carabus (Aulonocarabus) truncaticollis polaris Poppius, 1905
    Carabus (Aulonocarabus) truncaticollis polaris Poppius, 1905
    Female adult
    N.Russia, Polar Ural, Paypudynya River, Pompeliz Mt., h=500-700 m.
    10-15.VII.2002 S. Andreev, A. Romanenko leg.
    Copyright © Safronov
    Carabus (Axinocarabus) fedtschenkoi fedtschenkoi Solsky, 1874
    Carabus (Axinocarabus) fedtschenkoi fedtschenkoi Solsky, 1874
    Male adult
    27 mm
    Copyright © Boleslav Brezina
    Carabus (Axinocarabus) fedtschenkoi fedtschenkoi Solsky, 1874
    Carabus (Axinocarabus) fedtschenkoi fedtschenkoi Solsky, 1874
    Female adult
    30 mm
    Copyright © Boleslav Brezina
    Carabus (Axinocarabus) fedtschenkoi kondarensis Kryzhanovskij, 1953
    Carabus (Axinocarabus) fedtschenkoi kondarensis Kryzhanovskij, 1953
    Male adult
    Tadzhikistan, Varzhobskoe usch., kishlak Navobob
    4.1994 Avdeev I. leg.
    Copyright © Alexander Anichtchenko
    Carabus (Axinocarabus) fedtschenkoi kondarensis Kryzhanovskij, 1953
    Carabus (Axinocarabus) fedtschenkoi kondarensis Kryzhanovskij, 1953
    Female adult
    31 mm
    Copyright © Boleslav Brezina
    Carabus (Axinocarabus) fedtschenkoi miles Semenov, 1886
    Carabus (Axinocarabus) fedtschenkoi miles Semenov, 1886
    Male adult
    27 mm
    Copyright © Boleslav Brezina
    Carabus (Axinocarabus) fedtschenkoi miles Semenov, 1886
    Carabus (Axinocarabus) fedtschenkoi miles Semenov, 1886
    Female adult
    29 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) aristochroides aristochroides Deuve, 1992
    Carabus (Calocarabus) aristochroides aristochroides Deuve, 1992
    Male adult
    China Sichuan N-Occ. 25 km Nord Maniganggo
    4050 m, VI.2001 Cavazzuti leg.
    Copyright © Pierfranco Cavazzuti
    Carabus (Calocarabus) aristochroides qagcaensis Deuve, 1995
    Carabus (Calocarabus) aristochroides qagcaensis Deuve, 1995
    Male adult
    17 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) aristochroides qagcaensis Deuve, 1995
    Carabus (Calocarabus) aristochroides qagcaensis Deuve, 1995
    Female adult
    19 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) deuveianus Cavazzuti & Casale, 2006
    Carabus (Calocarabus) deuveianus Cavazzuti & Casale, 2006 Holotype
    Male adult
    Cina, Gansu mer., Qiling Shan, Passo a Nord di Tewo 4000m
    Guan Gai Shan. Colle mer. 18.VII.2005 Cavazzuti leg.
    Copyright © Pierfranco Cavazzuti
    Carabus (Calocarabus) deuveianus Cavazzuti & Casale, 2006
    Carabus (Calocarabus) deuveianus Cavazzuti & Casale, 2006
    Male adult
    17 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) dietererberi Heinz, 2001
    Carabus (Calocarabus) dietererberi Heinz, 2001
    Female adult
    20 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) gologensis Deuve e Tian 2004
    Carabus (Calocarabus) gologensis Deuve e Tian 2004
    Male adult
    18 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) gratus gratus Semenov, 1887
    Carabus (Calocarabus) gratus gratus Semenov, 1887
    Female adult
    17 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) gratus nishidai Deuve & Imura, 1993
    Carabus (Calocarabus) gratus nishidai Deuve & Imura, 1993 Paratype
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Calocarabus) gratus nishidai Deuve & Imura, 1993
    Carabus (Calocarabus) gratus nishidai Deuve & Imura, 1993 Paratype
    Female adult
    19 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) gratus serxuensis Deuve, 1995
    Carabus (Calocarabus) gratus serxuensis Deuve, 1995
    Male adult
    17 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) guinanensis guinanensis Deuve, 1991
    Carabus (Calocarabus) guinanensis guinanensis Deuve, 1991
    Male adult
    16 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) guinanensis guinanensis Deuve, 1991
    Carabus (Calocarabus) guinanensis guinanensis Deuve, 1991
    Female adult
    17 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) guinanensis maqenicola Kalab, 2005
    Carabus (Calocarabus) guinanensis maqenicola Kalab, 2005 Paratype
    Male adult
    China, SE Qinghai mts. 15 km E Maqen, 34.27N/100.24E 4300m
    J.Kalab leg., 22-24.7.2004 alpine meadows, screes
    Copyright © Pierfranco Cavazzuti
    Carabus (Calocarabus) janatai Brezina, 1996
    Carabus (Calocarabus) janatai Brezina, 1996 Paratype
    Female adult
    21 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) janatai Brezina, 1996
    Carabus (Calocarabus) janatai Brezina, 1996
    Male adult
    18 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) janatai Brezina, 1996
    Carabus (Calocarabus) janatai Brezina, 1996
    Male adult
    19 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) juengerianus Kleinfeld, 1995
    Carabus (Calocarabus) juengerianus Kleinfeld, 1995 Paratype
    Male adult
    18 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) juengerianus Kleinfeld, 1995
    Carabus (Calocarabus) juengerianus Kleinfeld, 1995
    Female adult

    Copyright © Boleslav Brezina
    Carabus (Calocarabus) juengerianus Kleinfeld, 1995
    Carabus (Calocarabus) juengerianus Kleinfeld, 1995
    Female adult
    20 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) juengerianus Kleinfeld, 1995 (as yangguifei)
    Carabus (Calocarabus) juengerianus Kleinfeld, 1995 (as yangguifei)
    Male adult
    17 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) juengerianus Kleinfeld, 1995
    Carabus (Calocarabus) juengerianus Kleinfeld, 1995
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Calocarabus) juengerianus Kleinfeld, 1995 (as yangguifei)
    Carabus (Calocarabus) juengerianus Kleinfeld, 1995 (as yangguifei)
    Male adult
    18 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) juengerianus Kleinfeld, 1995 (as yangguifei)
    Carabus (Calocarabus) juengerianus Kleinfeld, 1995 (as yangguifei) Paratype
    Female adult
    20 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) linxiaensis darcangicus Deuve, 1995
    Carabus (Calocarabus) linxiaensis darcangicus Deuve, 1995 Paratype
    Female adult
    20 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) linxiaensis darcangicus Deuve, 1995
    Carabus (Calocarabus) linxiaensis darcangicus Deuve, 1995 Paratype
    Male adult
    18 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) linxiaensis linxiaensis Deuve, 1992
    Carabus (Calocarabus) linxiaensis linxiaensis Deuve, 1992
    Male adult
    17 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) linxiaensis linxiaensis Deuve, 1992
    Carabus (Calocarabus) linxiaensis linxiaensis Deuve, 1992
    Female adult
    18 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) linxiaensis linxiaensis Deuve, 1992
    Carabus (Calocarabus) linxiaensis linxiaensis Deuve, 1992
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Calocarabus) linxiaensis ponggartangensis Deuve, 1995
    Carabus (Calocarabus) linxiaensis ponggartangensis Deuve, 1995 Holotype
    Male adult
    17 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) linxiaensis pseudogratus Deuve, 1994
    Carabus (Calocarabus) linxiaensis pseudogratus Deuve, 1994 Paratype
    Male adult
    19 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) linxiaensis pseudogratus Deuve, 1994
    Carabus (Calocarabus) linxiaensis pseudogratus Deuve, 1994
    Male adult
    17 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) linxiaensis pseudogratus Deuve, 1994
    Carabus (Calocarabus) linxiaensis pseudogratus Deuve, 1994
    Male adult
    17 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) linxiaensis pseudogratus Deuve, 1994
    Carabus (Calocarabus) linxiaensis pseudogratus Deuve, 1994
    Male adult
    17 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) linxiaensis pseudogratus Deuve, 1994
    Carabus (Calocarabus) linxiaensis pseudogratus Deuve, 1994 Paratype
    Male adult
    China, NW Sichuan, 45 km S Zamtang 31.52N 100.59E,
    4600 m, 15-19.7.2001 Rejsek leg.
    Copyright © Alexander Anichtchenko
    Carabus (Calocarabus) linxiaensis subacidus Cavazzuti, 2002
    Carabus (Calocarabus) linxiaensis subacidus Cavazzuti, 2002 Holotype
    Male adult
    China, Sichuan N-Occ., Sertar-Garze, 50 km a Nord del bivio Luhuo-Garze,
    4400.VI.2001 Cavazzuti leg.
    Copyright © Pierfranco Cavazzuti
    Carabus (Calocarabus) mandarin Kalab, 2002
    Carabus (Calocarabus) mandarin Kalab, 2002 Paratype
    Male adult
    17 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) turnaianus Deuve, 1995
    Carabus (Calocarabus) turnaianus Deuve, 1995
    Male adult
    China, Sichuan N-Occ., 20 km Sud di Qagca, Lala Shan 42-4500m
    VI.2001 Cavazzuti leg.
    Copyright © Pierfranco Cavazzuti
    Carabus (Calocarabus) trichothorax trichothorax Brezina & Imura, 1997
    Carabus (Calocarabus) trichothorax trichothorax Brezina & Imura, 1997
    Male adult
    22 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) trichothorax trichothorax Brezina & Imura, 1997
    Carabus (Calocarabus) trichothorax trichothorax Brezina & Imura, 1997
    Female adult
    22 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) pingpong Brezina & Hackel, 2004
    Carabus (Calocarabus) pingpong Brezina & Hackel, 2004 Holotype Paratype
    Male adult
    CH, S Gansu, SSW Minxian (12) W Qilian Shan
    11 km W of Kunda 34:08N/103:45E/ 4000-4300m
    alpine zone 9-10.VII.2002 Belousov & Kabak
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) pingpong Brezina & Hackel, 2004
    Carabus (Calocarabus) pingpong Brezina & Hackel, 2004
    Male adult
    CH, S Gansu, SSW Minxian (12) W Qilian Shan
    11 km W of Kunda 34:08N/103:45E/ 4000-4300m
    alpine zone 9-10.VII.2002 Belousov & Kabak
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) pingpong Brezina & Hackel, 2004
    Carabus (Calocarabus) pingpong Brezina & Hackel, 2004 Paratype
    Female adult
    20 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) pingpong Brezina & Hackel, 2004
    Carabus (Calocarabus) pingpong Brezina & Hackel, 2004 Paratype
    Male adult
    18 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) przewalskii somoli Deuve, 2007
    Carabus (Calocarabus) przewalskii somoli Deuve, 2007 Holotype
    Male adult
    16 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) przewalskii somoli Deuve, 2007
    Carabus (Calocarabus) przewalskii somoli Deuve, 2007
    Female adult
    19 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) przewalskii przewalskii Morawitz, 1886 (as elnarae)
    Carabus (Calocarabus) przewalskii przewalskii Morawitz, 1886 (as elnarae)
    Male adult
    CINA, Qinghai Nord-Or. Passo fre Daliang e Obo
    3600 m, 11.VII.2005 Cavazzuti leg.
    Copyright © Pierfranco Cavazzuti
    Carabus (Calocarabus) przewalskii przewalskii Morawitz, 1886
    Carabus (Calocarabus) przewalskii przewalskii Morawitz, 1886
    Male adult
    17 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) przewalskii przewalskii Morawitz, 1886
    Carabus (Calocarabus) przewalskii przewalskii Morawitz, 1886
    Female adult

    Copyright © Boleslav Brezina
    Carabus (Calocarabus) przewalskii przewalskii Morawitz, 1886
    Carabus (Calocarabus) przewalskii przewalskii Morawitz, 1886
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Calocarabus) przewalskii przewalskii Morawitz, 1886
    Carabus (Calocarabus) przewalskii przewalskii Morawitz, 1886
    Female adult
    19 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) przewalskii sewertzowi Semenov, 1898
    Carabus (Calocarabus) przewalskii sewertzowi Semenov, 1898 Holotype
    Male adult
    17 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) przewalskii sewertzowi Semenov, 1898
    Carabus (Calocarabus) przewalskii sewertzowi Semenov, 1898
    Male adult
    19 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) przewalskii sewertzowi Semenov, 1898
    Carabus (Calocarabus) przewalskii sewertzowi Semenov, 1898
    Female adult
    20 mm
    Copyright © Boleslav Brezina
    Carabus (Calocarabus) przewalskii paulusianus Deuve, 2007
    Carabus (Calocarabus) przewalskii paulusianus Deuve, 2007
    Male adult
    19 mm
    Copyright © Boleslav Brezina
    Carabus (Carabus) arvensis arvensis Herbst, 1784
    Carabus (Carabus) arvensis arvensis Herbst, 1784
    Male+Female adult
    W Ukraine, NW Podolian, Lviv reg., Javoriv distr., near Stradch vill., 20-30.IV.2009, Panin R. leg.
    Copyright © Ruslan Panin
    Carabus (Carabus) arvensis arvensis Herbst, 1784
    Carabus (Carabus) arvensis arvensis Herbst, 1784
    Male adult
    W Ukraine, Male Polissya, Lviv reg., near Kamenka-Bugzkaya, under the bark of oak tree, in the forest, 29.III.08, Panin R. leg. male,17mm
    Copyright © Ruslan Panin
    Carabus (Carabus) arvensis arvensis Herbst, 1784
    Carabus (Carabus) arvensis arvensis Herbst, 1784
    Male adult
    W. Ukraine , NW. Podolian, Lviv reg., Nikolaev distr., near Krasiv vill., 9.II.08, Panin R. leg. L=17mm
    Copyright © Ruslan Panin
    Carabus (Carabus) arvensis conciliator Fischer, 1822 (as jurgitae)
    Carabus (Carabus) arvensis conciliator Fischer, 1822 (as jurgitae)
    Male adult
    Russland, Altai, Aktasch
    leg. Nikolaevsky, 1991; det. U.Schmidt, 1995
    Foto: U.Schmidt, 2008
    Copyright © U.Schmidt
    Carabus (Carabus) arvensis baschkiricus Breuning, 1936
    Carabus (Carabus) arvensis baschkiricus Breuning, 1936
    Female adult
    Chebarkul, Chelyabinsk area<br /> 2.08.2008
    Copyright © Evgeniy Chibilov
    Carabus (Carabus) arvensis carpathus Born, 1902
    Carabus (Carabus) arvensis carpathus Born, 1902
    Male+Female adult
    Ukrainian Carpathians,
    Transcarpathian reg., Rakhov Distr., Chornohora range, h=1850 -2000m., 29.VI-28.VII.2007 Panin R. leg.
    18mm.
    Copyright © Ruslan Panin
    Carabus (Carabus) arvensis deyrollei Gory, 1839
    Carabus (Carabus) arvensis deyrollei Gory, 1839
    Male adult
    Spain, Asturias, 1-8.7.1999
    long. 17mm
    Copyright © Alexander Anichtchenko
    Carabus (Carabus) arvensis faldermanni Dejean, 1829
    Carabus (Carabus) arvensis faldermanni Dejean, 1829
    Female adult
    Rusia, S Primorskii reg., Poima riv., 30.VII-6.VIII.1999
    Anichtchenko A. leg.
    long. 22mm
    Copyright © Alexander Anichtchenko
    Carabus (Carabus) arvensis faldermanni Dejean, 1829
    Carabus (Carabus) arvensis faldermanni Dejean, 1829
    Female adult
    Rusia, Evreiskaja AO., near Obluchie 20.6.93
    Chichvarhin A. leg.
    long. 22mm
    Copyright © Alexander Anichtchenko
    Carabus (Carabus) granulatus leander Kraatz, 1878
    Carabus (Carabus) granulatus leander Kraatz, 1878
    Male+Female adult
    Russia, C-Caucasus: Balkaria, Valley of riv. Cherek Balkarskyi, h=1750-1800m 10.VI-20.VII.1992, Zamotajlov A. leg.
    female,22mm
    Copyright © Ruslan Panin
    Carabus (Carabus) granulatus calabricus Spettoli & Vigna Taglianti, 2001
    Carabus (Carabus) granulatus calabricus Spettoli & Vigna Taglianti, 2001
    Male adult
    Italia, Pollino, San Sever. Luc. Bosco Magnano,
    (PZ) 13.X.2002 800m, F.Angelini leg.
    long.18mm
    Copyright © Alexander Anichtchenko
    Carabus (Carabus) granulatus duarius Fischer, 1844
    Carabus (Carabus) granulatus duarius Fischer, 1844
    Male+Female adult
    SE Siberia, Russia
    Irkutsk reg., near Pivovarikha vill., 7.V.07, D.Fominyh leg.
    female, 22mm
    Copyright © Ruslan Panin
    Carabus (Carabus) granulatus granulatus Linne, 1758
    Carabus (Carabus) granulatus granulatus Linne, 1758
    Male adult
    W Ukraine,
    Lviv Reg., Javoriv distr., near Ivano-Frankovo vill., 30.IV-18.V.2009, Panin R. leg.
    male, 21mm
    Copyright © Ruslan Panin
    Carabus (Carabus) granulatus granulatus Linne, 1758
    Carabus (Carabus) granulatus granulatus Linne, 1758
    Male adult
    23 mm
    Germania, Oberfranken, Bad Steben
    leg.det. U.Schmidt, 2006
    Foto: U.Schmidt, 2008
    Copyright © U.Schmidt
    Carabus (Carabus) granulatus granulatus Linne, 1758
    Carabus (Carabus) granulatus granulatus Linne, 1758
    Female adult
    Polonia, Verh. Zalesie, V.1994 Strelnikov D. leg.
    long. 21mm
    Copyright © Alexander Anichtchenko
    Carabus (Carabus) granulatus granulatus Linne, 1758
    Carabus (Carabus) granulatus granulatus Linne, 1758
    Female adult
    long. 22mm
    Copyright © Ruslan Panin
    Carabus (Carabus) granulatus granulatus Linne, 1758
    Carabus (Carabus) granulatus granulatus Linne, 1758
    Male adult
    Canada
    Copyright Henri Goulet, Agriculture & Agri-Food Canada
    Copyright © Henri Goulet
    Carabus (Carabus) granulatus interstitialis Duftschmid, 1812
    Carabus (Carabus) granulatus interstitialis Duftschmid, 1812
    Male adult
    Austria, Murek, X.1971 leg. Orselli
    long. 22mm
    Copyright © Alexander Anichtchenko
    Carabus (Carabus) granulatus yezoensis Bates, 1883
    Carabus (Carabus) granulatus yezoensis Bates, 1883
    Male adult
    S Sakhalin isl., Novoaleksandrovka, Susun riv., 29.V.90<br /> long. 19mm
    Copyright © Alexander Anichtchenko
    Carabus (Carabus) menetriesi menetriesi Hummel, 1827
    Carabus (Carabus) menetriesi menetriesi Hummel, 1827
    Male adult
    W-Ukraine, Roztochia, Lviv Reg., Yavorov distr., near Ivano-Frankovo vill., Site Zalyvki, Panin R. leg., 1-20.V.2005, male, 21 mm.
    Copyright © Ruslan Panin
    Carabus (Carabus) stscheglowi Mannerheim, 1827
    Carabus (Carabus) stscheglowi Mannerheim, 1827
    Female adult
    E Ukraine,
    Dniepropetrovsk reg.,Novomoskovskiy distr. near Andriivka vill.16.VII.06 Kanarsky Yu.leg.
    female 23mm
    Copyright © Ruslan Panin
    Carabus (Carabus) stscheglowi Mannerheim, 1827
    Carabus (Carabus) stscheglowi Mannerheim, 1827
    Male+Female adult
    Central Russia , Voronezh region, near Grafskoe, 12.VIII.1984 long=20mm
    Copyright © Ruslan Panin
    Carabus (Carabus) vanvolxemi Putzeys, 1875
    Carabus (Carabus) vanvolxemi Putzeys, 1875
    Male adult

    Copyright © Alexander Anichtchenko
    Carabus (Cathaicus) brandti aishanensis Kleinfeld, 1999
    Carabus (Cathaicus) brandti aishanensis Kleinfeld, 1999 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Cathaicus) brandti brandti Faldermann, 1835
    Carabus (Cathaicus) brandti brandti Faldermann, 1835
    Male adult
    China, Beijing, Zhongguanchun, in the flower bed, 2007, Liu ye leg.;<br /> Photo & det. by liuye
    Copyright © Ye Liu
    Carabus (Cathaicus) brandti brandti Faldermann, 1835
    Carabus (Cathaicus) brandti brandti Faldermann, 1835
    Male adult
    28 mm
    Copyright © Boleslav Brezina
    Carabus (Cathaicus) brandti brandti Faldermann, 1835
    Carabus (Cathaicus) brandti brandti Faldermann, 1835
    Female adult
    27 mm
    Copyright © Boleslav Brezina
    Carabus (Acathaicus) interruptocostatus fantingi Imura & Su, 1998
    Carabus (Acathaicus) interruptocostatus fantingi Imura & Su, 1998 Holotype
    Male adult
    China, N.-Sichuan, above Jiuzhaigou, 2100-2200m, N bank of Riv. Bai He, Jiuzhaigou Xian" - 24 mm lang - in coll. Imura.
    Copyright © Frank Kleinfeld
    Carabus (Rhigocarabus) wengdaensis Kleinfeld & Puchner, 2009
    Carabus (Rhigocarabus) wengdaensis Kleinfeld & Puchner, 2009 Holotype
    Male adult
    HOLOTYPUS Sichuan, Str. Wengda-Luhuo, Pass 20 km südl. Wengda, 4100m (ca. 31°43'N/100°44'E), VIII.2007, leg. Ding - coll. Puchner 20mm lang / 7 mm breit
    Copyright © Frank Kleinfeld
    Carabus (Acathaicus) interruptocostatus nanpingicus Deuve, 1995
    Carabus (Acathaicus) interruptocostatus nanpingicus Deuve, 1995 Holotype
    Male adult
    Chine, Sichuan, Nanping, VII.1995, S. J. L. Remond" - Länge 25 mm - in MNHN Paris.
    Copyright © Frank Kleinfeld
    Carabus (Acathaicus) interruptocostatus amplior Deuve, 1994
    Carabus (Acathaicus) interruptocostatus amplior Deuve, 1994 Holotype
    Male adult
    Chine, Gansu, Min Shan, 2000m, J. M. Bousquet, 1994" - Länge 29 mm - in MNHN Paris
    Copyright © Frank Kleinfeld
    Carabus (Acathaicus) interruptocostatus tewoicus Deuve, 1996
    Carabus (Acathaicus) interruptocostatus tewoicus Deuve, 1996 Holotype
    Male adult
    Chine, Gansu, Tewo, 2000m, J. L. Remond, SR. 24,5 mm, in MNHN Paris
    Copyright © Frank Kleinfeld
    Carabus (Imaibius) reuteri Kleinfeld, 2008
    Carabus (Imaibius) reuteri Kleinfeld, 2008 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Platycarabus) fabricii fabricii Panzer, 1812
    Carabus (Platycarabus) fabricii fabricii Panzer, 1812
    Male adult
    NE-Italy,
    Trentino-Alto Adige Reg,. Bolzano-Bozen Prov.,Val Aurina – Rottal 2050-2150m, 28.VI.1997, I.Rapuzzi leg.
    male, 19mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) pedemontanus maurinensis Deuve, 2002
    Carabus (Orinocarabus) pedemontanus maurinensis Deuve, 2002 Holotype
    Male adult

    Copyright © Frank Kleinfeld
    Carabus (Imaibius) pachtoun kabakovianus Gottwald, 1992
    Carabus (Imaibius) pachtoun kabakovianus Gottwald, 1992 Holotype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Imaibius) pachtoun kabakovianus Gottwald, 1992
    Carabus (Imaibius) pachtoun kabakovianus Gottwald, 1992
    Male adult

    Copyright © Frank Kleinfeld
    Carabus (Imaibius) nouristani nouristani (Ledoux, 1977)
    Carabus (Imaibius) nouristani nouristani (Ledoux, 1977)
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Imaibius) nouristani nouristani (Ledoux, 1977)
    Carabus (Imaibius) nouristani nouristani (Ledoux, 1977)
    Endophallus

    Copyright © Frank Kleinfeld
    Carabus (Imaibius) anami Ledoux, 1977
    Carabus (Imaibius) anami Ledoux, 1977
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Imaibius) pachtoun pachtoun (Ledoux, 1975)
    Carabus (Imaibius) pachtoun pachtoun (Ledoux, 1975) Paratype
    Male adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) angermaieri Kleinfeld, 2008
    Carabus (Apotomopterus) angermaieri Kleinfeld, 2008 Holotype Paratype
    Male+Female adult
    China, Guangxi, Rong'an (=Chang'an: 25°13'109°23'), Durongshan, 1500m, V.2007, leg. Li et al. in coll. Kleinfeld
    male 30mm, female 32 mm
    Copyright © Frank Kleinfeld
    Carabus (Pagocarabus) qiangding Kleinfeld & Puchner, 2007
    Carabus (Pagocarabus) qiangding Kleinfeld & Puchner, 2007 Holotype Paratype
    Male+Female adult
    China. Sichuan, Str. zwischen Yajiang und Kangding, Tagong Grasslands (30°02'59,3''N/101°34'14,3''E)
    28 mm
    Copyright © Frank Kleinfeld
    Carabus (Pagocarabus) thibetanus Breuning, 1950
    Carabus (Pagocarabus) thibetanus Breuning, 1950 Holotype
    Male adult
    Tong-ho River, Szetschuan, VII.1947, 4.000m / C. Plesius thibetanus mihi, Typ / HOLOTYPE Coll. Frey, NHM Basel ? 27 mm
    Copyright © Frank Kleinfeld
    Carabus (Coptolabrus) formosus elytraviridis Kleinfeld, 2007
    Carabus (Coptolabrus) formosus elytraviridis Kleinfeld, 2007 Holotype Paratype
    Male+Female adult
    China, Sichuan, Aba pref., Nanping Co., Jiuzhaigou (ca. 33.17N/103.54E), 2000 m, 30.5.-6.6.1990, coll. Kleinfeld.
    HT 32 mm, PT 36 mm
    Copyright © Frank Kleinfeld
    Carabus (Carabus) kucerai puchneri Kleinfeld, 2007
    Carabus (Carabus) kucerai puchneri Kleinfeld, 2007
    Male adult
    China, Sichuan, road Pingwu-Nanping, Umg. Mupi Tibetan (Mupi 32.33N/104.33, 1250 m, 28.5.-4.6.2006),
    27 mm, coll. Puchner
    Copyright © Frank Kleinfeld
    Carabus (Cathaicus) brandti brandti Faldermann, 1835 (as grossegranulatus)
    Carabus (Cathaicus) brandti brandti Faldermann, 1835 (as grossegranulatus) Syntypes Lectotype
    Male adult
    C. (Cathaicus) brandti grossegranulatus Breuning 1943<br /> Lectotypus/SyntypusLectotypus (25 mm) von "South Mandschou". Syntypus (25 mm) "South Mandschou, 30. V. 1937"
    Copyright © Frank Kleinfeld
    Carabus (Cathaicus) brandti brandti Faldermann, 1835
    Carabus (Cathaicus) brandti brandti Faldermann, 1835 Syntypes Lectotype
    Male adult
    C. (Cathaicus) brandti grossegranulatus Breuning 1943
    Lectotypus/SyntypusLectotypus (25 mm) von "South Mandschou". Syntypus (25 mm) "South Mandschou, 30. V. 1937"
    Copyright © Frank Kleinfeld
    Carabus (Cathaicus) brandti reginae Deuve, 1997
    Carabus (Cathaicus) brandti reginae Deuve, 1997 Holotype
    Male adult
    Ex Ningxia, Suyuko, Helan Shan 2800 m, 2.VII.1996.
    MNHN Paris. 30 mm
    Copyright © Frank Kleinfeld
    Carabus (Carabus) sculpturatus Menetries, 1832
    Carabus (Carabus) sculpturatus Menetries, 1832
    Male+Female adult
    Azerbaijan,
    Talysh Mts. Hirkan National Park, Moscow Forest, 8-9.VII.1988, P.V.Kazaryan leg.
    female, 22 mm
    photo Panin R.
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) estreicheri Fischer, 1822
    Carabus (Trachycarabus) estreicheri Fischer, 1822
    Female adult
    Volgograd Reg., Serafimovka Distr., Ust' -Khoperskaya Station 5.V.2009 leg.O.Sveshnikov
    18.5 mm
    Copyright © Kirill Makarov
    Carabus (Hygrocarabus) nodulosus Creutzer, 1799
    Carabus (Hygrocarabus) nodulosus Creutzer, 1799
    Endophallus
    Endophallus
    Copyright © Ruslan Panin
    Carabus (Hygrocarabus) nodulosus Creutzer, 1799
    Carabus (Hygrocarabus) nodulosus Creutzer, 1799
    Male+Female adult
    Austria, Styria Region (Steiermark), Leibnitz, 31.X.1999, I.Rapuzzi leg.<br /> female, 29 mm
    Copyright © Ruslan Panin
    Carabus (Hygrocarabus) variolosus Fabricius, 1787
    Carabus (Hygrocarabus) variolosus Fabricius, 1787
    Male adult
    W-Ukraine, NW Podolian,<br /> Lviv reg., Zhydachiv distr.,near Vybranivka vill., 15.V-8.VI.2008, Panin R. leg.<br /> male, 27mm
    Copyright © Ruslan Panin
    Carabus (Hygrocarabus) variolosus Fabricius, 1787
    Carabus (Hygrocarabus) variolosus Fabricius, 1787
    Male+Female adult
    W-Ukraine,<br /> Lviv Reg., Zhydachiv distr., near Vybranivka vill., Panin R. leg.<br /> female, 30mm
    Copyright © Ruslan Panin
    Carabus (Hygrocarabus) variolosus variolosus Fabricius, 1787
    Carabus (Hygrocarabus) variolosus variolosus Fabricius, 1787
    Endophallus
    Endophallus
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) lambrechti Kleinfeld, 2006
    Carabus (Apotomopterus) lambrechti Kleinfeld, 2006 Holotype
    Male adult
    26 mm
    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) pseudoguizhouensis Kleinfeld, 2006
    Carabus (Apotomopterus) pseudoguizhouensis Kleinfeld, 2006 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Platycarabus) irregularis irregularis Fabricius, 1792
    Carabus (Platycarabus) irregularis irregularis Fabricius, 1792
    Male adult
    NE-Italy,
    Friuli Reg., Udine, Carnia, Pierabech h~1100m, I.Rapuzzi leg.
    male, 23 mm
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) shun emeifeng Kleinfeld, 2006
    Carabus (Apotomopterus) shun emeifeng Kleinfeld, 2006 Holotype
    Male adult
    China, Fujian, Taining Co., SW Xinqiao: Emeifeng Shan (ca. 26.59N/117.05E), 1500 m, 34 mm, coll. Kleinfeld
    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) delavayi planithoracis Kleinfeld, 2006
    Carabus (Apotomopterus) delavayi planithoracis Kleinfeld, 2006 Holotype
    Male adult
    30 mm,
    China, Süd-Yunnan, Lüchün (23.00N/102.20E): Shanbashan, 2200 m, 9/2005, coll. Kleinfeld
    Copyright © Frank Kleinfeld
    Carabus (Imaibius) nouristani bandaensis Kleinfeld & Reuter, 2007
    Carabus (Imaibius) nouristani bandaensis Kleinfeld & Reuter, 2007 Holotype
    Male adult
    Afghanistan, prov. Konar, NE Jalalabad, Schamschehr - Banda-Wald (ca. 34.44N/70.50E), 2800 m
    23 mm
    Copyright © Frank Kleinfeld
    Carabus (Eucarabus) manifestus pyonganicola Deuve & Li, 1998
    Carabus (Eucarabus) manifestus pyonganicola Deuve & Li, 1998
    Male+Female adult
    N-Korea: Pyonganam-Do, YangTok County,(39;127), 12-19,VI.2006
    Copyright © Ruslan Panin
    Carabus (Eucarabus) manifestus pyonganicola Deuve & Li, 1998
    Carabus (Eucarabus) manifestus pyonganicola Deuve & Li, 1998
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Platycarabus) irregularis ramanus Sokolar, 1909 (as hayeki Kobmann, 1925)
    Carabus (Platycarabus) irregularis ramanus Sokolar, 1909 (as hayeki Kobmann, 1925)
    Male adult
    N Slovenia, Maribor – Dravograd, Pomorje Mts.,13.XI.2005, I.Rapuzzi leg.
    Copyright © Ruslan Panin
    Carabus (Platycarabus) irregularis ramanus Sokolar, 1909 (as bucephalus Kraatz, 1879)
    Carabus (Platycarabus) irregularis ramanus Sokolar, 1909 (as bucephalus Kraatz, 1879)
    Male+Female adult
    W Slovenia, Goriška region Trnovo, h~1000 m, IX.2002, I.Rapuzzi leg.
    female, 29 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) praecellens trans illigeri Dejean, 1826
    Carabus (Morphocarabus) praecellens trans illigeri Dejean, 1826
    Male+Female adult
    W-Croatia, Karlovac, near Bosiljevo vill., h~300m, V.2000, I.Rapuzzi leg. female, 33 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) praecellens trans illigeri Dejean, 1826
    Carabus (Morphocarabus) praecellens trans illigeri Dejean, 1826
    Endophallus
    W-Croatia, Karlovac, near Bosiljevo vill., h~300m,
    Copyright © Ruslan Panin
    Carabus (Platycarabus) creutzeri creutzeri Fabricius, 1801 (as trnovensis Bernau, 1914)
    Carabus (Platycarabus) creutzeri creutzeri Fabricius, 1801 (as trnovensis Bernau, 1914)
    Male adult
    W Slovenia,
    Lokve, Trnovski Gozd h=1100m, 17.IV.2006, leg. I.Rapuzzi
    male, 26mm
    Copyright © Ruslan Panin
    Carabus (Platycarabus) creutzeri creutzeri Fabricius, 1801
    Carabus (Platycarabus) creutzeri creutzeri Fabricius, 1801
    Male adult
    Italy, Fruili - UD - Carnia, N. Paulano 29.III.2002 leg. I.Rapuzzi
    L=22.0 mm
    Copyright © Kirill Makarov
    Carabus (Platycarabus) creutzeri creutzeri Fabricius, 1801(as posarinii Rapuzzi, 1999 PT)
    Carabus (Platycarabus) creutzeri creutzeri Fabricius, 1801(as posarinii Rapuzzi, 1999 PT) Paratype
    Male+Female adult
    Slovenia:Koper:Kras Mt.R.:Brkini Mts.:Suhorije env., VIII.1998, I.Rapuzzi leg.
    photo & koll. Panin R.
    female, 32 mm
    Copyright © Ruslan Panin
    Carabus (Diocarabus) opaculus opaculus Putzeys, 1875
    Carabus (Diocarabus) opaculus opaculus Putzeys, 1875
    Male adult
    Rusia, Kunashir isl.,<br /> 8.1991 Shestopalov M. leg.<br /> male, 15 mm
    Copyright © Alexander Anichtchenko
    Carabus (Diocarabus) opaculus opaculus Putzeys, 1875
    Carabus (Diocarabus) opaculus opaculus Putzeys, 1875
    Male adult
    Japan, Hokkaido, <br /> Hokuryu-cho 260 m., Museushi (West), 43:48’37’N;141:40’13’E, 6-26.VII.2010, I.Rapuzzi & L.Caldon leg.
    Copyright © Ruslan Panin
    Carabus (Platycarabus) creutzeri kircheri Germar, 1824
    Carabus (Platycarabus) creutzeri kircheri Germar, 1824
    Male+Female adult
    NE-Italy, Veneto Reg., Belluno, Forcella Lavardet h=2000m, VII.2002, I.Rapuzzi leg.
    female, 24 mm
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) casaleianus casaleianus Deuve, 1994
    Carabus (Apotomopterus) casaleianus casaleianus Deuve, 1994
    Female adult
    China, SW Hunan Prov., Chengbu Co, Nan-Shan-Ding (26.34;110,08)1-10.VIII,2006, I.Rapuzzi leg.
    L=33 mm
    Copyright © Ruslan Panin
    Carabus (Diocarabus) lucepunctus ignipunctatus Rapuzzi, 2009 PT
    Carabus (Diocarabus) lucepunctus ignipunctatus Rapuzzi, 2009 PT Paratype
    Male+Female adult
    China, SW Hunan Prov., Chengbu Co, Nan-Shan-Ding (26.34;110,08)1-10.VIII,2006, I.Rapuzzi leg.
    femele, 18 mm
    Copyright © Ruslan Panin
    Carabus (Macrothorax) morbillosus alternans Palliardi, 1825
    Carabus (Macrothorax) morbillosus alternans Palliardi, 1825
    Male adult
    S Italy, Sicily, Province Palermo, Piana degli Albanesi h~700m, XI.2010, I.Rapuzzi leg.
    L=32 mm
    Copyright © Ruslan Panin
    Carabus (Aulonocarabus) canaliculatus pseudocareniger Deuve, 1991
    Carabus (Aulonocarabus) canaliculatus pseudocareniger Deuve, 1991
    Male adult
    N Korea, Ryanggang Prov., Mt. Tongjom Ridge h~1861m, (128.60E;41.20N) I. Rapuzzi leg.
    L=27 mm
    Copyright © Ruslan Panin
    Carabus (Eucarabus) manifestus guanmenshanus Imura, 1991
    Carabus (Eucarabus) manifestus guanmenshanus Imura, 1991
    Female adult
    NE China, Liaoning, Anshan City, Mt.Qian Shan, 4-16.VII.2005, I.Rapuzzi leg.
    L=26 mm
    Copyright © Ruslan Panin
    Carabus (Ohomopterus) albrechti albrechti Morawitz, 1862
    Carabus (Ohomopterus) albrechti albrechti Morawitz, 1862
    Female adult
    Japan, S-Hokkaido:Oshima Peninsula,Pass btw Otoshibe - Assabu, h~250-300 m, 10-20.VII.2010, 42:01’53’Т; 140:22’09’E, I.Rapuzzi & L.Caldon leg.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) spasskianus aeruginosiformis Breuning, 1932
    Carabus (Morphocarabus) spasskianus aeruginosiformis Breuning, 1932
    Male adult
    Russia, Khakassia,
    W-Sayan, Alan Mt range, valley Enisey riv., 5 km S Mayna vill., 15.VII.2000, A.& N. Menstschicovy leg.
    L=24 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) spasskianus aeruginosiformis Breuning, 1932
    Carabus (Morphocarabus) spasskianus aeruginosiformis Breuning, 1932
    Endophallus
    Russia, Khakassia,
    W-Sayan, Alan Mt range, valley Enisey riv., 5 km S Mayna vill.,
    Copyright © Ruslan Panin
    Carabus (Pagocarabus) promachus promachus s.str.
    Carabus (Pagocarabus) promachus promachus s.str.
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) promachus danbaensis Kleinfeld, 1999
    Carabus (Pagocarabus) promachus danbaensis Kleinfeld, 1999
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) promachus danbaensis Kleinfeld, 1999
    Carabus (Pagocarabus) promachus danbaensis Kleinfeld, 1999
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Pagocarabus) promachus promachus s.str. (as konkerianus)
    Carabus (Pagocarabus) promachus promachus s.str. (as konkerianus)
    Female adult

    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) poschingerianus Korell & Kleinfeld, 1989
    Carabus (Pagocarabus) poschingerianus Korell & Kleinfeld, 1989 Holotype
    Male adult

    Copyright © Frank Kleinfeld
    Carabus (Pagocarabus) erwini plesiolabrus Deuve, 1991
    Carabus (Pagocarabus) erwini plesiolabrus Deuve, 1991
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Pagocarabus) erwini erwini s.str.
    Carabus (Pagocarabus) erwini erwini s.str.
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Meganebrius) wallichi wittmeri Mandl, 1974
    Carabus (Meganebrius) wallichi wittmeri Mandl, 1974
    Female adult
    Nepal, Lantang, near Dunche, Songta vill.
    3500 m 15.5.2000
    Copyright © Alexander Anichtchenko
    Carabus (Meganebrius) arunensis Heinertz, 1980
    Carabus (Meganebrius) arunensis Heinertz, 1980
    Female adult

    Copyright © Boleslav Brezina
    Carabus (Meganebrius) franzi Mandl, 1974
    Carabus (Meganebrius) franzi Mandl, 1974
    Female adult
    Nepal, Lantang, near Dunche, Songta vill,
    3500 m, 15.5.2000
    Copyright © Alexander Anichtchenko
    Carabus (Meganebrius) tamang gurkhaorum Lassalle, 1985
    Carabus (Meganebrius) tamang gurkhaorum Lassalle, 1985
    Female adult
    CW Nepal, Manaslu massif 2300 m,
    Bara Pokhari Lekh, 5.2001 M.Hackel leg.
    Copyright © Alexander Anichtchenko
    Carabus (Macrothorax) aumonti Lucas, 1849
    Carabus (Macrothorax) aumonti Lucas, 1849
    Male adult

    Copyright © Alexander Anichtchenko
    Carabus (Macrothorax) planatus Chaudoir, 1843
    Carabus (Macrothorax) planatus Chaudoir, 1843
    Male adult
    Sicilia, Madonie, Piano Battaglia (PA) 1500m
    4.6.1992 Aliquo leg.
    male, 31 mm
    Copyright © Alexander Anichtchenko
    Carabus (Macrothorax) morbillosus macilentus Lapouge, 1899
    Carabus (Macrothorax) morbillosus macilentus Lapouge, 1899
    Male adult
    Spain, Alicante, Santa Pola, 20.2.1988 M. Godoy leg.
    male, 29 mm
    Copyright © Alexander Anichtchenko
    Carabus (Macrothorax) morbillosus constantinus Kraatz, 1899
    Carabus (Macrothorax) morbillosus constantinus Kraatz, 1899
    Female adult

    Copyright © Safronov
    Carabus (Macrothorax) rugosus boeticus Deyrolle, 1852
    Carabus (Macrothorax) rugosus boeticus Deyrolle, 1852
    Female adult
    Spain, Cadiz, Novo Sancti Petri, 10.6.2003
    female, 32 mm
    Copyright © Alexander Anichtchenko
    Carabus (Macrothorax) rugosus rugosus Fabricius, 1792
    Carabus (Macrothorax) rugosus rugosus Fabricius, 1792
    Male adult
    N Africa, 2 km W Ceuta, Allabacal,
    17-18.2.2004 Anichtchenko A. leg.
    male, 26 mm
    Copyright © Alexander Anichtchenko
    Carabus (Macrothorax) rugosus laufferi Breuning, 1927
    Carabus (Macrothorax) rugosus laufferi Breuning, 1927
    Female adult
    Spain, Gredos Mts., 2 km E from La Mira Mt.
    4-7.7.2007 Anichtchenko A. leg.
    female, 29 mm
    Copyright © Alexander Anichtchenko
    Carabus (Macrothorax) rugosus celtibericus Germar, 1824
    Carabus (Macrothorax) rugosus celtibericus Germar, 1824
    Male adult
    Portugal, Vila Fresca de Azeitas,
    Santana, 13.12.2001
    male, 30 mm
    Copyright © Alexander Anichtchenko
    Carabus (Lipaster) stjernvalli stjernvalli Mannerheim, 1830
    Carabus (Lipaster) stjernvalli stjernvalli Mannerheim, 1830
    Male+Female adult
    Armenia,
    Ashotsk, 22.VI.1990, P.V.Kazaryan leg.
    female 34mm
    Copyright © Ruslan Panin
    Carabus (Lipaster) stjernvalli (as tabackuriensis Novotny & Vorisek, 1988)
    Carabus (Lipaster) stjernvalli (as tabackuriensis Novotny & Vorisek, 1988)
    Male adult

    Copyright © Alexander Anichtchenko
    Carabus (Limnocarabus) clathratus clathratus Linne, 1761
    Carabus (Limnocarabus) clathratus clathratus Linne, 1761
    Female adult
    W-Ukraine, Maloe Polesie, Lviv Region, near Kamenka-Bugzkaya, 28.III.2008, Panin R. leg. L=31 mm
    Copyright © Ruslan Panin
    Carabus (Limnocarabus) clathratus clathratus Linne, 1761
    Carabus (Limnocarabus) clathratus clathratus Linne, 1761
    Male adult
    length: 20-36 mm, Germania, Mecklenburg, Wismar, leg. F.Wolf, 1998; det. A.Skale, Photo: U.Schmidt, 2009
    Copyright © U.Schmidt
    Carabus (Limnocarabus) clathratus clathratus (as clathratus foveolatoseriatus)
    Carabus (Limnocarabus) clathratus clathratus (as clathratus foveolatoseriatus)
    Female adult
    The Chelyabinsk area, Kizilskyi township, Cheka mountain (steppe). 14-15/07/2008
    Copyright © Sunvas
    Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
    Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
    Male adult
    Transcaucasia, Armenia,
    Amasia distr., Arpalach vill., VI-VII.1999, P.V.Kazaryan leg.
    male, 25mm
    Copyright © Ruslan Panin
    Carabus (Carabus) limbatus Say, 1825 (as goryi)
    Carabus (Carabus) limbatus Say, 1825 (as goryi)
    Female adult

    Copyright © Henri Goulet
    Carabus (Carabus) limbatus Say, 1825
    Carabus (Carabus) limbatus Say, 1825
    Male adult

    Copyright © Denis Potanin
    Carabus (Leptoplesius) merzbacheri kirgisiensis Mandl, 1967
    Carabus (Leptoplesius) merzbacheri kirgisiensis Mandl, 1967
    Female adult
    22mm
    Copyright © Boleslav Brezina
    Carabus (Leptoplesius) merzbacheri kirgisiensis Mandl, 1967
    Carabus (Leptoplesius) merzbacheri kirgisiensis Mandl, 1967
    Female adult
    female, 27mm
    Copyright © Boleslav Brezina
    Carabus (Leptoplesius) merzbacheri kirgisiensis Mandl, 1967
    Carabus (Leptoplesius) merzbacheri kirgisiensis Mandl, 1967
    Female adult
    female, 24mm
    Copyright © Boleslav Brezina
    Carabus (Leptoplesius) shokalskii kuqaensis Deuve, 1994
    Carabus (Leptoplesius) shokalskii kuqaensis Deuve, 1994 Paratype
    Female adult
    female, 24mm
    Copyright © Boleslav Brezina
    Carabus (Leptocarabus) yokoae marcilhaci Deuve, 1992
    Carabus (Leptocarabus) yokoae marcilhaci Deuve, 1992
    Male adult
    CHINA, Shanxi, Tschingling Mts., Taibeichan Nat. reserv.;
    Copyright © Anton Menstschikov
    Carabus (Leptocarabus) yokoae marcilhaci Deuve, 1992
    Carabus (Leptocarabus) yokoae marcilhaci Deuve, 1992
    Male adult
    China, Shanxi, Taschingling Mts., Taibeishan Nat. res.
    2000 m., 5-15.08.2005
    Copyright © Alexander Anichtchenko
    Carabus (Leptocarabus) semiopacus Reitter, 1895
    Carabus (Leptocarabus) semiopacus Reitter, 1895
    Female adult
    N Korea, Nangnim, 14.VII.1981
    female, long. 24 mm
    Copyright © Alexander Anichtchenko
    Carabus (Leptocarabus) procerulus Chaudoir, 1862
    Carabus (Leptocarabus) procerulus Chaudoir, 1862
    Male adult
    Japan, Yamanashi pref., Otawa-rindo,
    Narusawa vill., 9.IX.2002 T. Kurihara leg
    male, 31 mm
    Copyright © Alexander Anichtchenko
    Carabus (Leptocarabus) harmandi fujisan Ishikawa, 1972
    Carabus (Leptocarabus) harmandi fujisan Ishikawa, 1972
    Female adult
    Japan, Yamanashi pref., Fuji-Rinds 1600m, Narusawa vill. 3.V.2005 Kurihara K. leg. 25 mm
    Copyright © Alexander Anichtchenko
    Carabus (Leptocarabus) arboreus exarboreus (Ishikawa, 1992)
    Carabus (Leptocarabus) arboreus exarboreus (Ishikawa, 1992)
    Male adult
    E.Russia, Far East, Sakhalin Isl.,<br /> forest of M.Tym river<br /> 11-IX-2000 . Egorov leg.<br /> male 27mm, female 30mm.
    Copyright © Safronov
    Carabus (Lasiocoptolabrus) sunwukong sunwukong Imura, 1993
    Carabus (Lasiocoptolabrus) sunwukong sunwukong Imura, 1993 Paratype
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Lasiocoptolabrus) sunwukong baotianmanus (Imura, 2005)
    Carabus (Lasiocoptolabrus) sunwukong baotianmanus (Imura, 2005) Paratype
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Lamprostus) calleyi pseudoprasinus Lapouge, 1912
    Carabus (Lamprostus) calleyi pseudoprasinus Lapouge, 1912
    Male+Female adult
    Transcaucasia, Armenia,
    Ashotsk distr. Tsoghamarg vill, .21-22.V.1997, P.V.Kazaryan leg
    female, 24 mm
    Copyright © Ruslan Panin
    Carabus (Lamprostus) calleyi nigrinus Motschulsky, 1865
    Carabus (Lamprostus) calleyi nigrinus Motschulsky, 1865
    Male adult
    Azerbaijan,
    Talysh Mts., Palikesh, Zuvand, 08.VI.2001, Dolin V.G. leg.
    male, 24mm
    Copyright © Ruslan Panin
    Carabus (Lamprostus) torosus spinolae Cristoforis & Jan, 1837
    Carabus (Lamprostus) torosus spinolae Cristoforis & Jan, 1837
    Male adult
    length: 29 mm
    Türkei, Cankiri, Cerkes, Kurtcimeni
    leg.det. K.Staven, 1989
    Foto: U.Schmidt, 2009
    Copyright © U.Schmidt
    Carabus (Ischnocarabus) tenuitarsis tenuitarsis Kraatz, 1877
    Carabus (Ischnocarabus) tenuitarsis tenuitarsis Kraatz, 1877
    Male adult
    male, 24mm
    Copyright © Boleslav Brezina
    Carabus (Ischnocarabus) tenuitarsis tenuitarsis Kraatz, 1877
    Carabus (Ischnocarabus) tenuitarsis tenuitarsis Kraatz, 1877
    Female adult
    female, 26mm
    Copyright © Boleslav Brezina
    Carabus (Ischnocarabus) cychropalpus Peyron, 1858
    Carabus (Ischnocarabus) cychropalpus Peyron, 1858
    Male adult
    male, 27mm
    Copyright © Boleslav Brezina
    Carabus (Ischnocarabus) cychropalpus Peyron, 1858
    Carabus (Ischnocarabus) cychropalpus Peyron, 1858
    Female adult
    female, 28mm
    Copyright © Boleslav Brezina
    Carabus (Iniopachus) pyrenaeus jeanneli Deuve, 1994
    Carabus (Iniopachus) pyrenaeus jeanneli Deuve, 1994
    Male adult
    1-2: Pirineos, Canfranc, Ibon de Iserias 2200 m 16-17.7.05<br /> 3: Pirineos, Monte Perdido, Collata Anisolo 2400m 25.8.05
    Copyright © Alexander Anichtchenko
    Carabus (Iniopachus) auriculatus ubinensis (Puissegur, 1955)
    Carabus (Iniopachus) auriculatus ubinensis (Puissegur, 1955)
    Female adult
    Spain, Montes Cantabricos, Peña Ubiña, 2400m
    5-6.VII.2009 Anichtchenko A. leg.
    female, 19 mm
    Copyright © Alexander Anichtchenko
    Carabus (Iniopachus) auriculatus auriculatus Putzeys, 1872
    Carabus (Iniopachus) auriculatus auriculatus Putzeys, 1872
    Female adult
    Spain, Picos de Europa, Macizo Central, Las Colladinas,
    ref. Hermoso 2100m, 8.8.2003
    Copyright © Alexander Anichtchenko
    Carabus (Imaibius) wittmerorum Heinertz, 1978
    Carabus (Imaibius) wittmerorum Heinertz, 1978
    Male adult

    Copyright © Alexander Anichtchenko
    Carabus (Imaibius) stoliczkanus Bates, 1878
    Carabus (Imaibius) stoliczkanus Bates, 1878
    Male adult

    Copyright © Alexander Anichtchenko
    Carabus (Imaibius) pachtoun marysae Letellier, 1979
    Carabus (Imaibius) pachtoun marysae Letellier, 1979
    Male adult

    Copyright © Alexander Anichtchenko
    Carabus (Imaibius) isabellae Lassalle, 1985
    Carabus (Imaibius) isabellae Lassalle, 1985
    Male adult
    male, 27mm
    Copyright © Boleslav Brezina
    Carabus (Imaibius) isabellae Lassalle, 1985
    Carabus (Imaibius) isabellae Lassalle, 1985
    Female adult
    female, 30mm
    Copyright © Boleslav Brezina
    Carabus (Imaibius) gandharae Heinertz, 1978
    Carabus (Imaibius) gandharae Heinertz, 1978
    Male adult

    Copyright © Alexander Anichtchenko
    Carabus (Imaibius) dardiellus polymorphus Heinz, 1993
    Carabus (Imaibius) dardiellus polymorphus Heinz, 1993 Paratype
    Female adult
    Museum of Institute of Systematic Biology (Daugavpils, Latvia)

    Copyright © Alexander Anichtchenko
    Carabus (Imaibius) caschmirensis caschmirensis Redtenbacher, 1844
    Carabus (Imaibius) caschmirensis caschmirensis Redtenbacher, 1844
    Female adult
    length: 32 mm<br /> Pakistan, Mansehra<br /> leg. Heinz & Staven, 2001; det. K.Staven, 2001<br /> Foto: U.Schmidt, 2008
    Copyright © U.Schmidt
    Carabus (Imaibius) boysi Tatum, 1851
    Carabus (Imaibius) boysi Tatum, 1851
    Male adult

    Copyright © Alexander Anichtchenko
    Carabus (Imaibius) barysomus hazarensis Deuve, 1982
    Carabus (Imaibius) barysomus hazarensis Deuve, 1982
    Male adult
    length: 35 mm<br /> Pakistan, Mansehra<br /> leg.det. K.Staven, 2001<br /> Topotypus<br /> Foto: U.Schmidt, 2008
    Copyright © U.Schmidt
    Carabus (Imaibius) baronii sharanicus Deuve & Letellier, 1980
    Carabus (Imaibius) baronii sharanicus Deuve & Letellier, 1980 Paratype
    Male adult

    Copyright © Alexander Anichtchenko
    Carabus (Imaibius) baronii sharanicus Deuve & Letellier, 1980
    Carabus (Imaibius) baronii sharanicus Deuve & Letellier, 1980
    Male adult

    Copyright © Alexander Anichtchenko
    Carabus (Imaibius) baronii baronii Heinertz, 1977
    Carabus (Imaibius) baronii baronii Heinertz, 1977
    Male adult
    length: 23 mm
    Pakistan, Mansehra
    leg.det. K.Staven, 1997
    Topotypus
    Foto: U.Schmidt, 2008
    Copyright © U.Schmidt
    Carabus (Imaibius) baronii battalensis Heinz & Staven, 1998
    Carabus (Imaibius) baronii battalensis Heinz & Staven, 1998 Paratype
    Male adult

    Copyright © Alexander Anichtchenko
    Carabus (Pseudocoptolabrus) businskyi Deuve, 1990
    Carabus (Pseudocoptolabrus) businskyi Deuve, 1990
    Male adult
    CH, NW-Yunnan (Lijiang) Yulongxue Shan, E-slope
    27:10:50N/100:12:26E/ 4400m, alpine meadows 12.VII.2000
    K. & B. Brezina, M.Brychcin, M. Hackel
    Copyright © Boleslav Brezina
    Carabus (Oreocarabus) taibaiensis Kleinfeld, 2001
    Carabus (Oreocarabus) taibaiensis Kleinfeld, 2001 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Oreocarabus) latro minshanensis Deuve, 1987
    Carabus (Oreocarabus) latro minshanensis Deuve, 1987
    Male adult
    China. N Sishuan, Juizhalgou
    12-16.6.1995 Benes leg.
    male, 22 mm
    Copyright © Alexander Anichtchenko
    Carabus (Oreocarabus) laotse paralaotse Kleinfeld, 2000
    Carabus (Oreocarabus) laotse paralaotse Kleinfeld, 2000 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Oreocarabus) mikhaili Deuve & Mourzine, 1997
    Carabus (Oreocarabus) mikhaili Deuve & Mourzine, 1997
    Male adult
    China, S Gansu, Min Shan, 2100m
    SW Wudu, 1.8.2000 V.Patrikeev leg.
    male, 20 mm
    Copyright © Alexander Anichtchenko
    Carabus (Trachycarabus) sibiricus gvozdevae Obydov, 1997
    Carabus (Trachycarabus) sibiricus gvozdevae Obydov, 1997
    Male adult
    Russia, Khakassia,
    W Sayan, Enisey riv., Batenevskiy Mt Range, 12 km E Pervomayskiy vill., 11.VII.2000, A.& N. Menstschicovy leg.
    L=21 mm
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) sibiricus gvozdevae Obydov, 1997
    Carabus (Trachycarabus) sibiricus gvozdevae Obydov, 1997
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Homoeocarabus) maeander paludis Gehin, 1885
    Carabus (Homoeocarabus) maeander paludis Gehin, 1885
    Female adult
    female, 21mm
    Copyright © Boleslav Brezina
    Carabus (Homoeocarabus) maeander maeander s.str.
    Carabus (Homoeocarabus) maeander maeander s.str.
    Male adult
    Canada
    Copyright Henri Goulet, Agriculture & Agri-Food Canada
    Copyright © Henri Goulet
    Carabus (Homoeocarabus) maeander maeander s.str.
    Carabus (Homoeocarabus) maeander maeander s.str.
    Male adult
    male, 18mm
    Copyright © Boleslav Brezina
    Carabus (Homoeocarabus) maeander maeander s.str.
    Carabus (Homoeocarabus) maeander maeander s.str.
    Male adult
    male, 17mm
    Copyright © Boleslav Brezina
    Carabus (Cathoplius) aliai Escalera, 1944
    Carabus (Cathoplius) aliai Escalera, 1944
    Male adult
    male, 27mm
    Copyright © Boleslav Brezina
    Carabus (Cathoplius) asperatus asperatus (Dejean, 1826)
    Carabus (Cathoplius) asperatus asperatus (Dejean, 1826)
    Male adult
    male, 24mm
    Copyright © Boleslav Brezina
    Carabus (Cathoplius) asperatus asperatus (Dejean, 1826)
    Carabus (Cathoplius) asperatus asperatus (Dejean, 1826)
    Female adult
    female, 26mm
    Copyright © Boleslav Brezina
    Carabus (Cathoplius) asperatus ifniensis (Zarco, 1941)
    Carabus (Cathoplius) asperatus ifniensis (Zarco, 1941)
    Male adult
    male, 30mm
    Copyright © Boleslav Brezina
    Carabus (Cathoplius) asperatus ifniensis (Zarco, 1941)
    Carabus (Cathoplius) asperatus ifniensis (Zarco, 1941)
    Male adult
    male, 34mm
    Copyright © Boleslav Brezina
    Carabus (Cathoplius) stenocephalus Lucas, 1866 (as escalerae)
    Carabus (Cathoplius) stenocephalus Lucas, 1866 (as escalerae)
    Male adult
    male, 29mm
    Copyright © Boleslav Brezina
    Carabus (Cathoplius) stenocephalus Lucas, 1866 (as escalerae)
    Carabus (Cathoplius) stenocephalus Lucas, 1866 (as escalerae)
    Female adult
    female, 33mm
    Copyright © Boleslav Brezina
    Carabus (Cathoplius) stenocephalus Lucas, 1866
    Carabus (Cathoplius) stenocephalus Lucas, 1866
    Female adult
    female, 28mm
    Copyright © Boleslav Brezina
    Carabus (Cathoplius) stenocephalus Lucas, 1866
    Carabus (Cathoplius) stenocephalus Lucas, 1866
    Male adult
    male, 25 mm
    Copyright © Boleslav Brezina
    Carabus (Cathoplius) asperatus susicus Antoine, 1941
    Carabus (Cathoplius) asperatus susicus Antoine, 1941
    Male adult
    male, 27mm
    Copyright © Boleslav Brezina
    Carabus (Cathoplius) asperatus susicus Antoine, 1941
    Carabus (Cathoplius) asperatus susicus Antoine, 1941
    Female adult
    female, 29mm
    Copyright © Boleslav Brezina
    Carabus (Aulonocarabus) mouthiezianus Deuve, 1991
    Carabus (Aulonocarabus) mouthiezianus Deuve, 1991
    Male adult
    Yakutia, Verhoyanskii Mt.rng., âåðõîâüÿ Êåëå, ð. Íÿìíè, ëåâ. èñòîê. ð. Êîê÷èí 29.7.1994
    Copyright © Alexander Anichtchenko
    Carabus (Orinocarabus) bremii Stierlin, 1881
    Carabus (Orinocarabus) bremii Stierlin, 1881
    Male adult
    NW Italy, Piemonte, Mucrone Mt., 14-21.VI.1999
    Copyright © Alexander Anichtchenko
    Carabus (Cechenotribax) petri Semenov & Znojko, 1932
    Carabus (Cechenotribax) petri Semenov & Znojko, 1932
    Female adult
    female, 18mm
    Copyright © Boleslav Brezina
    Carabus (Cechenotribax) petri Semenov & Znojko, 1932
    Carabus (Cechenotribax) petri Semenov & Znojko, 1932
    Male adult
    male, 14mm
    Copyright © Boleslav Brezina
    Carabus (Cephalornis) potanini (Semenov, 1887)
    Carabus (Cephalornis) potanini (Semenov, 1887)
    Male adult
    China, S Gansu, 3400m, Minshan mts.,
    Wenxian 8.92, J. M. Bousquet
    Copyright © Boleslav Brezina
    Carabus (Chaetocarabus) arcadicus arcadicus Gistl, 1850
    Carabus (Chaetocarabus) arcadicus arcadicus Gistl, 1850
    Female adult
    female, 35mm
    Copyright © Boleslav Brezina
    Carabus (Chaetocarabus) arcadicus arcadicus Gistl, 1850
    Carabus (Chaetocarabus) arcadicus arcadicus Gistl, 1850
    Male adult
    male, 29mm
    Copyright © Boleslav Brezina
    Carabus (Chaetocarabus) arcadicus merlini Schaum, 1861
    Carabus (Chaetocarabus) arcadicus merlini Schaum, 1861
    Male adult
    male, 30mm
    Copyright © Boleslav Brezina
    Carabus (Chaetocarabus) arcadicus merlini Schaum, 1861
    Carabus (Chaetocarabus) arcadicus merlini Schaum, 1861
    Female adult
    female, 36mm
    Copyright © Boleslav Brezina
    Carabus (Chaetocarabus) intricatus bayardi Solier, 1835
    Carabus (Chaetocarabus) intricatus bayardi Solier, 1835
    Male+Female adult
    S-Italy,
    Basilicata Reg., Potenza Prov., Latronico, F. Angelini leg. 14.VI.2002
    female, 26mm
    Copyright © Ruslan Panin
    Carabus (Chaetocarabus) intricatus bayardi Solier, 1835
    Carabus (Chaetocarabus) intricatus bayardi Solier, 1835
    Male adult
    male, 25mm
    Copyright © Boleslav Brezina
    Carabus (Chaetocarabus) intricatus bayardi Solier, 1835
    Carabus (Chaetocarabus) intricatus bayardi Solier, 1835
    Female adult
    female, 27mm
    Copyright © Boleslav Brezina
    Carabus (Chaetocarabus) intricatus intricatus Linne, 1761 (as bohemicus)
    Carabus (Chaetocarabus) intricatus intricatus Linne, 1761 (as bohemicus)
    Female adult
    female, 33mm
    Copyright © Boleslav Brezina
    Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
    Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
    Male adult
    male, 26mm
    Copyright © Boleslav Brezina
    Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
    Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
    Male adult
    W-Ukraine,
    Lviv Reg., Zhydachiv distr., near Vybranivka Vill. Panin R.leg. 12.X.2007
    31mm.
    Copyright © Ruslan Panin
    Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
    Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
    Female adult
    36 mm
    Fundort: Germania, Oberfranken, Kulmbach
    leg.det. U.Schmidt, 1977
    Foto: U.Schmidt, 2008
    Copyright © U.Schmidt
    Carabus (Chaetocarabus) intricatus krueperi Reitter, 1896
    Carabus (Chaetocarabus) intricatus krueperi Reitter, 1896
    Male adult
    male, 35mm
    Copyright © Boleslav Brezina
    Carabus (Chaetocarabus) intricatus krueperi Reitter, 1896
    Carabus (Chaetocarabus) intricatus krueperi Reitter, 1896
    Female adult
    female, 38mm
    Copyright © Boleslav Brezina
    Carabus (Chaetocarabus) intricatus lefebvrei Dejean, 1826
    Carabus (Chaetocarabus) intricatus lefebvrei Dejean, 1826
    Male adult
    male, 28mm
    Copyright © Boleslav Brezina
    Carabus (Chaetocarabus) intricatus lefebvrei Dejean, 1826
    Carabus (Chaetocarabus) intricatus lefebvrei Dejean, 1826
    Female adult
    female, 30mm
    Copyright © Boleslav Brezina
    Carabus (Chrysocarabus) auronitens costellatus Gehin, 1882 (as subfestivus Oberthur, 1884)
    Carabus (Chrysocarabus) auronitens costellatus Gehin, 1882 (as subfestivus Oberthur, 1884)
    Female adult
    W- FRANCE,
    Bretagne Reg., Finistrere, Foret de Coatloc'h, XI.2006
    female, 26 mm
    Copyright © Ruslan Panin
    Carabus (Chrysocarabus) auronitens costellatus Gehin, 1882
    Carabus (Chrysocarabus) auronitens costellatus Gehin, 1882
    Male+Female adult
    C-FRANCE,
    Limousin Reg., Corrиze Dep., Aubazines, X.2005, Pascal Stefani leg.
    female,26mm
    Copyright © Ruslan Panin
    Carabus (Chrysocarabus) auronitens costellatus Gehin, 1882
    Carabus (Chrysocarabus) auronitens costellatus Gehin, 1882
    Male adult
    France 42, Mount de Madeleine, 1300m, les Biefs
    18.VIII.1991
    male, 19mm
    Copyright © Alexander Anichtchenko
    Carabus (Chrysocarabus) auronitens cupreonitens Chevrolat, 1861
    Carabus (Chrysocarabus) auronitens cupreonitens Chevrolat, 1861
    Female adult
    NW-FRANCE,
    Basse-Normandie Reg., Orne, Forêt de Bellême, 14.XII.2008, Pascal Stefani leg.
    female, 26mm
    Copyright © Ruslan Panin
    Carabus (Chrysocarabus) auronitens cupreonitens Chevrolat, 1861
    Carabus (Chrysocarabus) auronitens cupreonitens Chevrolat, 1861
    Male+Female adult
    NW-FRANCE,
    Basse-Normandie Reg,, Orne, Foret d' Ecouves, V.2009, Pascal Stefani leg.
    male, 25 mm
    Copyright © Ruslan Panin
    Carabus (Chrysocarabus) auronitens escheri (as laevipennis Seidlitz, 1891)
    Carabus (Chrysocarabus) auronitens escheri (as laevipennis Seidlitz, 1891)
    Male adult
    W Ukraine, NE Carpathians Mts.
    Transcarpathian, Rakhov Distr., Chornogora range, Petros Mt.h=2010m, 9.VI.2009, R.Panin leg
    male, 25 mm
    Copyright © Ruslan Panin
    Carabus (Chrysocarabus) auronitens escheri Palliardi, 1825 (ab. opacus Haury, 1878)
    Carabus (Chrysocarabus) auronitens escheri Palliardi, 1825 (ab. opacus Haury, 1878)
    Male+Female adult
    W Ukraine, NE Carpathians Mts.,<br /> Transcarpathian reg., Rakhov Distr., Chornogora range, h=2020m, alpin zone, 28.VI-22.VIII.2009, Panin R. leg.<br /> female, 27mm
    Copyright © Ruslan Panin
    Carabus (Chrysocarabus) auronitens escheri Palliardi, 1825
    Carabus (Chrysocarabus) auronitens escheri Palliardi, 1825
    Male adult
    W.Ukraine,
    Carpatians, Grebeniv env., Zelemyanka Mts.range h=1000 m., 9-20.VII.2007, R.Panin leg.
    Copyright © Safronov
    Carabus (Chrysocarabus) auronitens farinesi Dejean, 1826 (as barthei Lapouge, 1912)
    Carabus (Chrysocarabus) auronitens farinesi Dejean, 1826 (as barthei Lapouge, 1912)
    Male+Female adult
    SC -France,
    Midi-Pyrйnйes reg., Ariиge, Foret de Belesta, XI.2008, J.Nicolas leg.
    L=31 mm
    Copyright © Ruslan Panin
    Carabus (Chrysocarabus) auronitens festivus Dejean, 1826
    Carabus (Chrysocarabus) auronitens festivus Dejean, 1826
    Male adult
    male, 22mm
    Copyright © Ruslan Panin
    Carabus (Chrysocarabus) auronitens montanus Gehin, 1882
    Carabus (Chrysocarabus) auronitens montanus Gehin, 1882
    Male+Female adult
    SC-FRANCE,
    Haute-Garonne, Col de Mentй h~1300m., II.1980. Kretschmer leg.
    female, 23mm.
    photo/det. Panin R.
    Copyright © Ruslan Panin
    Carabus (Chrysocarabus) lineatus lateralis Chevrolat, 1840
    Carabus (Chrysocarabus) lineatus lateralis Chevrolat, 1840
    Male adult
    Spain, Villoslada de Cameros, Logroño, 19.6.87
    F. Rodriguez leg.
    male, 24mm
    Copyright © Alexander Anichtchenko
    Carabus (Chrysocarabus) lineatus lineatus Dejean, 1826
    Carabus (Chrysocarabus) lineatus lineatus Dejean, 1826
    Female adult
    N-Spain,
    Navarra, Sierra de Urbasa, Puerto de Opacua, III.2003
    female, 27mm
    Copyright © Ruslan Panin
    Carabus (Chrysocarabus) lineatus salmantinus Bolivar, 1922
    Carabus (Chrysocarabus) lineatus salmantinus Bolivar, 1922
    Male adult
    Spain, Candelario, Salamanca, 19.XI.1988
    leg. E. Carabajal
    long. 24mm
    Copyright © Alexander Anichtchenko
    Carabus (Chrysocarabus) solieri clairi Gehin, 1885
    Carabus (Chrysocarabus) solieri clairi Gehin, 1885
    Male adult
    France, Caussols, VII.1984
    male, 26 mm
    Copyright © Alexander Anichtchenko
    Carabus (Chrysocarabus) solieri liguranus Breuning, 1933
    Carabus (Chrysocarabus) solieri liguranus Breuning, 1933
    Female adult
    NW Italy,
    Altare, Savona,15.VI.1995, Cartini A.leg.
    female, 33mm
    Copyright © Ruslan Panin
    Carabus (Chrysocarabus) solieri solieri Dejean, 1826
    Carabus (Chrysocarabus) solieri solieri Dejean, 1826
    Male adult
    SE-France: Alpes-Maritimes
    Copyright © Safronov
    Carabus (Chrysocarabus) splendens Olivier, 1790
    Carabus (Chrysocarabus) splendens Olivier, 1790
    Male+Female adult
    SE France,
    Midi-Pyrйnйes reg., Ariиge, Vaychis, 22.X.2002, A.Laforgue leg.
    female, 30mm.
    Copyright © Ruslan Panin
    Carabus (Chrysocarabus) splendens Olivier, 1790
    Carabus (Chrysocarabus) splendens Olivier, 1790
    Male adult
    Spain, Pais Vasco, Bera de Bidasoa, 15.9.2001
    Anichtchenko A. leg.
    male, 30mm
    Copyright © Alexander Anichtchenko
    Carabus (Chrysocarabus) hispanus Fabricius, 1787
    Carabus (Chrysocarabus) hispanus Fabricius, 1787
    Male adult
    FRANCE,
    Tarn, Foret de la Gresigne, h=400m.VII.2005 Laforgue leg.
    male-30mm
    Copyright © Ruslan Panin
    Carabus (Chrysocarabus) hispanus Fabricius, 1787
    Carabus (Chrysocarabus) hispanus Fabricius, 1787
    Male adult
    Auriac alt. 500m.
    (France : Aude) 25.V.2000
    Copyright © Pascal Stefani
    Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896 (as juberriensis Mollard, 1999)
    Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896 (as juberriensis Mollard, 1999)
    Male+Female adult
    ANDORRA,
    E-Pyrenees, Juberri, h=1350-1400m, 22.VI-18.VII.2007
    female, 33mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
    Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
    Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
    Male+Female adult
    Russia, S Siberia, W Sayan, Ergaki Mt range, Nizhnyaya Buyba river h=1700 m, 19.VII.1999, A.& N. Menstschikovy leg.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) spasskianus shoriensis Obydov, 1999
    Carabus (Morphocarabus) spasskianus shoriensis Obydov, 1999
    Male adult
    Russia, Khakassia,
    Abakanskii Mt Range, Askiz river vall., 7 km E Kazanovka vill., 15.VI.2006, A.& N. Menstschikovy leg.
    L=24 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) spasskianus shoriensis Obydov, 1999
    Carabus (Morphocarabus) spasskianus shoriensis Obydov, 1999
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Diocarabus) loschnikovi tsaganensis Obydov, 2001
    Carabus (Diocarabus) loschnikovi tsaganensis Obydov, 2001
    Male adult
    Russia, W Tuva,
    Kyzyl-Tayga Mt Range h=1500 m, Ak-Suk riv vall., 21.VI.2006, A.& N. Menstschikovy leg.
    L=16 mm
    Copyright © Ruslan Panin
    Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
    Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988 (as shchessiensis)
    Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988 (as shchessiensis) Paratype
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Carabus) granulatus granulatus Linne, 1758
    Carabus (Carabus) granulatus granulatus Linne, 1758
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Chrysocarabus) auronitens escheri Palliardi, 1825
    Carabus (Chrysocarabus) auronitens escheri Palliardi, 1825
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Chrysocarabus) auronitens festivus Dejean, 1826
    Carabus (Chrysocarabus) auronitens festivus Dejean, 1826
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Chrysocarabus) auronitens farinesi Dejean, 1826
    Carabus (Chrysocarabus) auronitens farinesi Dejean, 1826
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Chrysocarabus) hispanus Fabricius, 1787
    Carabus (Chrysocarabus) hispanus Fabricius, 1787
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Eucarabus) obsoletus obsoletus Sturm, 1815
    Carabus (Eucarabus) obsoletus obsoletus Sturm, 1815
    Male adult
    NW Ukrainian Carpathians, Lviv Reg., Staryi Sambor Distr., near Dobromyl , h=500 m., 15-24.VI.2005, Panin R. leg., male, 28mm
    Copyright © Ruslan Panin
    Carabus (Eucarabus) obsoletus obsoletus Sturm, 1815
    Carabus (Eucarabus) obsoletus obsoletus Sturm, 1815
    Male adult
    NW Ukrainian Carpathians,
    Lviv reg., Staryi Sambor Distr., near Dobromyl , h=500m 6-28.VI.2009, Panin R. leg.
    male, 27 mm
    photo Panin R.
    Copyright © Ruslan Panin
    Carabus (Eucarabus) obsoletus obsoletus Sturm, 1815
    Carabus (Eucarabus) obsoletus obsoletus Sturm, 1815
    Female adult

    Copyright © Safronov
    Carabus (Eucarabus) obsoletus obsoletus Sturm, 1815
    Carabus (Eucarabus) obsoletus obsoletus Sturm, 1815
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Megodontus) violaceus aurolimbatus Dejean, 1829
    Carabus (Megodontus) violaceus aurolimbatus Dejean, 1829
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997
    Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997
    Male+Female adult
    Russia,Tuva,
    Akademika Obrucheva Mt range, 15 km N Cherbi vill., Chailang ur. 7.VII.1999, A & N. Menstschikovy leg.
    L=23 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997
    Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Hemicarabus) tuberculosus Dejean, 1829
    Carabus (Hemicarabus) tuberculosus Dejean, 1829
    Male adult
    male, 19mm
    Copyright © Boleslav Brezina
    Carabus (Hemicarabus) tuberculosus Dejean, 1829
    Carabus (Hemicarabus) tuberculosus Dejean, 1829
    Male adult
    China, Shanxi pro., Jiaochen
    Copyright © Ye Liu
    Carabus (Hemicarabus) tuberculosus Dejean, 1829
    Carabus (Hemicarabus) tuberculosus Dejean, 1829
    Male adult
    Far East, Lazovsky Dist., shore of Japan Sea, 3 km NE Preobrazhenie Vill., Ezhovaya (Tasovaya) Bay 13.07.06 M. & L. Smirnov's leg.
    male
    L=20,0 mm
    Copyright © Maksim Smirnov
    Carabus (Hemicarabus) serratus Say, 1825
    Carabus (Hemicarabus) serratus Say, 1825
    Male adult
    male, 18mm
    Copyright © Boleslav Brezina
    Carabus (Hemicarabus) serratus Say, 1825 (as tatumi Motschulsky, 1865 )
    Carabus (Hemicarabus) serratus Say, 1825 (as tatumi Motschulsky, 1865 )
    Female adult
    female, 18mm
    Copyright © Boleslav Brezina
    Carabus (Hemicarabus) serratus Say, 1825 (as tatumi Motschulsky, 1865 )
    Carabus (Hemicarabus) serratus Say, 1825 (as tatumi Motschulsky, 1865 )
    Female adult
    female, 19mm
    Copyright © Boleslav Brezina
    Carabus (Hemicarabus) serratus Say, 1825 (as tatumi Motschulsky, 1865)
    Carabus (Hemicarabus) serratus Say, 1825 (as tatumi Motschulsky, 1865)
    Male adult
    male, 17mm
    Copyright © Boleslav Brezina
    Carabus (Hemicarabus) serratus Say, 1825
    Carabus (Hemicarabus) serratus Say, 1825
    Female adult
    female, 21mm
    Copyright © Boleslav Brezina
    Carabus (Hemicarabus) nitens Linne, 1758
    Carabus (Hemicarabus) nitens Linne, 1758
    Male adult
    male, 15mm
    Copyright © Boleslav Brezina
    Carabus (Hemicarabus) nitens Linne, 1758
    Carabus (Hemicarabus) nitens Linne, 1758
    Male+Female adult
    Belarus,
    Vitebsk reg., Ushachskiy distr., near Bolshie Dolcy vill. 9.V.1990, I.A.Solodovnikov leg.
    female, 18mm.
    Copyright © Ruslan Panin
    Carabus (Hemicarabus) nitens Linne, 1758
    Carabus (Hemicarabus) nitens Linne, 1758
    Male adult
    Russia, Ural, riv. Rezh, vill. Kamenka; 10.VII.2009, N. Menstschicova leg.
    male, 14 mm.,
    Copyright © Anton Menstschikov
    Carabus (Hemicarabus) macleayi macleayi Dejean, 1826
    Carabus (Hemicarabus) macleayi macleayi Dejean, 1826
    Male adult
    male, 15mm
    Copyright © Boleslav Brezina
    Carabus (Hemicarabus) macleayi macleayi Dejean, 1826
    Carabus (Hemicarabus) macleayi macleayi Dejean, 1826
    Male adult
    Russia, Far East,<br /> Lazovskij Nat. Res., Proselochnaja Riv. near nouth, Tumannaja Mt. 42°59.888' N 134°06.935' E 18.VIII.2007, K.Makarov & A.Zaitsev leg.<br /> L=16 mm
    Copyright © Kirill Makarov
    Carabus (Morphocarabus) rothi hampei Kuster, 1846 (transition to rothi incompsus Kraatz, 1880)
    Carabus (Morphocarabus) rothi hampei Kuster, 1846 (transition to rothi incompsus Kraatz, 1880)
    Male adult
    NW Romania , Bistrita Nasaud, Mts. Tibles, 6 km SE Bichigiu, 17.VI.2009, S.Budka leg.
    L=35 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi hampei Kuster, 1846 (transition to rothi incompsus Kraatz, 1880)
    Carabus (Morphocarabus) rothi hampei Kuster, 1846 (transition to rothi incompsus Kraatz, 1880)
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi hampei Kuster, 1846 (as dacicus Csiki, 1906)
    Carabus (Morphocarabus) rothi hampei Kuster, 1846 (as dacicus Csiki, 1906)
    Female adult
    NW Romania , Bistrita Nasaud, Beclean env., 17.VI.2009, S.Budka leg.
    female, 31 mm
    Copyright © Ruslan Panin
    Carabus (Heterocarabus) marietti sapancaensis Blumenthal & Breuning, 1967
    Carabus (Heterocarabus) marietti sapancaensis Blumenthal & Breuning, 1967
    Female adult
    female, 22mm
    Copyright © Boleslav Brezina
    Carabus (Heterocarabus) marietti sapancaensis Blumenthal & Breuning, 1967
    Carabus (Heterocarabus) marietti sapancaensis Blumenthal & Breuning, 1967
    Male adult
    male, 22mm
    Copyright © Boleslav Brezina
    Carabus (Heterocarabus) marietti ornatus Heinz, 1976
    Carabus (Heterocarabus) marietti ornatus Heinz, 1976 Paratype
    Female adult
    female, 20mm
    Copyright © Boleslav Brezina
    Carabus (Heterocarabus) marietti ormanensis Blumenthal & Breuning, 1967
    Carabus (Heterocarabus) marietti ormanensis Blumenthal & Breuning, 1967
    Male adult
    male, 21mm
    Copyright © Boleslav Brezina
    Carabus (Heterocarabus) marietti muchei Breuning, 1961
    Carabus (Heterocarabus) marietti muchei Breuning, 1961
    Male adult
    male, 19mm
    Copyright © Boleslav Brezina
    Carabus (Heterocarabus) marietti inexspectatus Basquin & Darge, 1986
    Carabus (Heterocarabus) marietti inexspectatus Basquin & Darge, 1986
    Male adult
    male, 20mm
    Copyright © Boleslav Brezina
    Carabus (Heterocarabus) marietti inexspectatus Basquin & Darge, 1986
    Carabus (Heterocarabus) marietti inexspectatus Basquin & Darge, 1986
    Female adult
    female, 21mm
    Copyright © Boleslav Brezina
    Carabus (Heterocarabus) marietti marietti Cristoforis & Jan, 1837 (as charetianus Auvray, 1992)
    Carabus (Heterocarabus) marietti marietti Cristoforis & Jan, 1837 (as charetianus Auvray, 1992)
    Male adult
    male, 21mm
    Copyright © Boleslav Brezina
    Carabus (Heterocarabus) marietti marietti Cristoforis & Jan, 1837 (as charetianus Auvray, 1992)
    Carabus (Heterocarabus) marietti marietti Cristoforis & Jan, 1837 (as charetianus Auvray, 1992)
    Female adult
    female, 23mm
    Copyright © Boleslav Brezina
    Carabus (Heterocarabus) marietti bischoffi Chaudoir, 1848
    Carabus (Heterocarabus) marietti bischoffi Chaudoir, 1848
    Female adult
    female, 20mm
    Copyright © Boleslav Brezina
    Carabus (Heterocarabus) marietti bischoffi Chaudoir, 1848
    Carabus (Heterocarabus) marietti bischoffi Chaudoir, 1848
    Male adult
    male, 18mm
    Copyright © Boleslav Brezina
    Carabus (Heterocarabus) marietti bischoffi Chaudoir, 1848
    Carabus (Heterocarabus) marietti bischoffi Chaudoir, 1848
    Male adult
    male, 19mm
    Copyright © Boleslav Brezina
    Carabus (Heterocarabus) marietti marietti (as ativsengueni Schweiger, 1962)
    Carabus (Heterocarabus) marietti marietti (as ativsengueni Schweiger, 1962)
    Male adult
    male, 17mm
    Copyright © Boleslav Brezina
    Carabus (Heterocarabus) marietti akensis Haury, 1889
    Carabus (Heterocarabus) marietti akensis Haury, 1889
    Male adult
    22mm
    Copyright © Boleslav Brezina
    Carabus (Goniocarabus) tadzhikistanus Kryzhanovskij, 1968
    Carabus (Goniocarabus) tadzhikistanus Kryzhanovskij, 1968
    Female adult
    female, 28mm
    Copyright © Boleslav Brezina
    Carabus (Goniocarabus) tadzhikistanus Kryzhanovskij, 1968
    Carabus (Goniocarabus) tadzhikistanus Kryzhanovskij, 1968
    Male adult
    male, 25mm
    Copyright © Boleslav Brezina
    Carabus (Goniocarabus) sogdianus Semenov, 1898
    Carabus (Goniocarabus) sogdianus Semenov, 1898
    Male adult
    male, 21mm
    Copyright © Boleslav Brezina
    Carabus (Goniocarabus) sogdianus Semenov, 1898
    Carabus (Goniocarabus) sogdianus Semenov, 1898
    Female adult
    female, 23mm
    Copyright © Boleslav Brezina
    Carabus (Goniocarabus) gussakovskii Kryzhanovskij, 1971
    Carabus (Goniocarabus) gussakovskii Kryzhanovskij, 1971
    Female adult
    female, 23mm
    Copyright © Boleslav Brezina
    Carabus (Goniocarabus) gussakovskii Kryzhanovskij, 1971
    Carabus (Goniocarabus) gussakovskii Kryzhanovskij, 1971
    Male adult
    male, 22mm
    Copyright © Boleslav Brezina
    Carabus (Goniocarabus) gussakovskii Kryzhanovskij, 1971
    Carabus (Goniocarabus) gussakovskii Kryzhanovskij, 1971
    Male adult
    male, 21mm
    Copyright © Boleslav Brezina
    Carabus (Goniocarabus) grombczewskii Semenov, 1891
    Carabus (Goniocarabus) grombczewskii Semenov, 1891
    Female adult
    female, 22mm
    Copyright © Boleslav Brezina
    Carabus (Goniocarabus) grombczewskii Semenov, 1891
    Carabus (Goniocarabus) grombczewskii Semenov, 1891
    Male adult
    male, 20mm
    Copyright © Boleslav Brezina
    Carabus (Goniocarabus) caerulans Moravitz, 1886
    Carabus (Goniocarabus) caerulans Moravitz, 1886
    Male adult
    male, 22mm
    Copyright © Boleslav Brezina
    Carabus (Goniocarabus) caerulans Moravitz, 1886
    Carabus (Goniocarabus) caerulans Moravitz, 1886
    Female adult
    female, 24mm
    Copyright © Boleslav Brezina
    Carabus (Gnathocarabus) kusnetzovi Semenov, 1903
    Carabus (Gnathocarabus) kusnetzovi Semenov, 1903
    Female adult
    female, 21mm
    Copyright © Boleslav Brezina
    Carabus (Gnathocarabus) kusnetzovi Semenov, 1903
    Carabus (Gnathocarabus) kusnetzovi Semenov, 1903
    Male adult
    male, 17mm
    Copyright © Boleslav Brezina
    Carabus (Fulgenticarabus) flutschi koreus Deuve, 2006
    Carabus (Fulgenticarabus) flutschi koreus Deuve, 2006
    Male adult
    N Korea, Samjiyon, Ryang Gang Province
    6-12.6.2006 leg. Wong
    Copyright © Boleslav Brezina
    Carabus (Fulgenticarabus) flutschi koreus Deuve, 2006
    Carabus (Fulgenticarabus) flutschi koreus Deuve, 2006
    Female adult
    female, 22mm
    Copyright © Boleslav Brezina
    Carabus (Megodontus) avinovi Semenov & Znojko, 1932
    Carabus (Megodontus) avinovi Semenov & Znojko, 1932
    Male adult

    Copyright © Kirill Makarov
    Carabus (Megodontus) kolbei aino Rost, 1908
    Carabus (Megodontus) kolbei aino Rost, 1908
    Female adult

    Copyright © Kirill Makarov
    Carabus (Megodontus) kolbei aino Rost, 1908
    Carabus (Megodontus) kolbei aino Rost, 1908
    Male adult

    Copyright © Kirill Makarov
    Carabus (Pachycarabus) koenigi Ganglbauer, 1886
    Carabus (Pachycarabus) koenigi Ganglbauer, 1886
    Male adult
    NW Caucasus, S Russia, Adygea Rep., Lagonaki plateau, Site Murzikal h=1800-2000m, 24.V-18.IX.2010, D.Fominykh leg.
    L= 22 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822 (as pitschugini)
    Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822 (as pitschugini) Holotype
    Male adult
    East Siberia, Tunkinskie Goltzy (East Sajans), Mondy env., 24.VI.1979, E. Ya. Berlov leg.
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822 (as pitschugini)
    Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822 (as pitschugini) Paratype
    Female adult
    East Siberia, Tunkinskie Goltzy (East Sajans), Mondy env., 24.VI.1979, E. Ya. Berlov leg.
    Copyright © Dmitri Obydov
    Carabus (Pachystus) hortensis neumeyeri Schaum, 1856 (as chalkidikensis)
    Carabus (Pachystus) hortensis neumeyeri Schaum, 1856 (as chalkidikensis) Holotype Paratype
    Male+Female adult
    Griechenland, Makedonien, Chalkidike, Holomondas Berge, zwischen den Orten Amea und Taxiarhis, 932 m, N 40°27'861", E0 23O32'661" leg. et coll. A. Muller (Wien).
    Copyright © A.Muller
    Carabus (Diocarabus) caustomarginatus Imura & Mizusawa, 1994 (as paegamensis)
    Carabus (Diocarabus) caustomarginatus Imura & Mizusawa, 1994 (as paegamensis) Holotype Paratype
    Male+Female adult
    1.-20.6.2006, Nordkorea, Chonchon County, Mt.Paegam, 1825m, (40.80/126.20), leg. einheimischer Sammler; coll. A. Mьller (Wien)
    Copyright © A.Muller
    Carabus (Carabus) granulatus interstitialis Duftschmid, 1812
    Carabus (Carabus) granulatus interstitialis Duftschmid, 1812
    Male adult
    Italy, Emilia reg., Modena, Castelnuovo Rangone, 31.XII.1991, Sola C. leg.
    L=19 mm
    Copyright © Ruslan Panin
    Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
    Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
    Male adult
    Italy-Switzerland, Riviera – Preonzo, 18.II.1998, C.Sola leg.
    L=26 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) castanopterus A. & J.B. Villa, 1833
    Carabus (Orinocarabus) castanopterus A. & J.B. Villa, 1833
    Female adult
    NC-Italy - S-Switzerland, Val Colla – Gazzirola, 22.VI.1999, C. Sola leg.
    L=21mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) helleri Ganglbauer, 1893 (as pseudopreyssleri Breuning, 1932)
    Carabus (Morphocarabus) helleri Ganglbauer, 1893 (as pseudopreyssleri Breuning, 1932)
    Male adult
    Czech Rep., NE Moravia, Moravian-Silesian Beskids Mt Rng, PR Trojacka, Hodslavice vill., 29.V.12.VI.2010, Krause Rene leg./det.Panin R. L=30 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) helleri Ganglbauer, 1893 (as pseudopreyssleri Breuning, 1932)
    Carabus (Morphocarabus) helleri Ganglbauer, 1893 (as pseudopreyssleri Breuning, 1932)
    Endophallus
    Czech Rep., NE Moravia, Moravian-Silesian Beskids Mt Rng, PR Trojacka, Hodslavice vill.,
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) kollari Palliardi, 1825 (as semetricus Kraatz, 1878)
    Carabus (Morphocarabus) kollari Palliardi, 1825 (as semetricus Kraatz, 1878)
    Male adult
    SW-Romania:Banat, Bocsa/Coltan, 21.V-5.VI.2010, Krause Rene leg.
    L=29 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) kollari Palliardi, 1825 (as semetricus Kraatz, 1878)
    Carabus (Morphocarabus) kollari Palliardi, 1825 (as semetricus Kraatz, 1878)
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Eucarabus) catenulatus catenulatus Scopoli, 1763
    Carabus (Eucarabus) catenulatus catenulatus Scopoli, 1763
    Male adult
    Lombardy - North Italy
    Copyright © Monzini
    Carabus (Tomocarabus) convexus gracilior Gehin, 1885
    Carabus (Tomocarabus) convexus gracilior Gehin, 1885
    Male adult
    SE-Bulgaria, Burgas Prov., Primorsko env., 18-30.VI.2000, Krause Rene leg.
    L=18 mm
    Copyright © Ruslan Panin
    Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
    Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
    Female adult
    SW Slovakia, Bratislava env., IV.2004, Krause Rene leg.
    L=32 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) sylvestris sylvestris Panzer, 1793 (as slabai Niedl, 1968)
    Carabus (Orinocarabus) sylvestris sylvestris Panzer, 1793 (as slabai Niedl, 1968)
    Female adult
    Czech, WS Bohemia,
    Plzen Reg., Domazlice Distr., Babylon, h=500m, 30.VIII.2008, S.I. Budka leg.
    L=24mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) violaceus violaceus Linne, 1758
    Carabus (Megodontus) violaceus violaceus Linne, 1758
    Male adult
    S.Bohemia, Hluboka nad Vltavou, h=380 m, 6.VIII.2004, S. Budka leg.
    L=28 mm
    Copyright © Ruslan Panin
    Carabus (Carabus) arvensis deyrollei Gory, 1839
    Carabus (Carabus) arvensis deyrollei Gory, 1839
    Female adult
    NW Spain, Galicia Reg., Pontevedra Prov., A Lama, Xesta, h~650 m, 1.X.1994
    L=17 mm
    Copyright © Ruslan Panin
    Carabus (Archicarabus) wiedemanni wiedemanni Menetries, 1836
    Carabus (Archicarabus) wiedemanni wiedemanni Menetries, 1836
    Male adult
    SE-Bulgaria:Burgas env. Tsarevo (Michurin), 10-15.VII.1981, J.Slaba leg.<br /> L=22 mm
    Copyright © Ruslan Panin
    Carabus (Cechenochilus) boeberi buschi Semenov & Znojko, 1931
    Carabus (Cechenochilus) boeberi buschi Semenov & Znojko, 1931
    Male adult
    Male, 19 mm.
    C Caucasus, Kabardino-Balkaria, upper reach. riv. Adylsu, h=2550, 20-24.07.2008, Leg. E.Derzhinsky
    Copyright © Igor Solodovnikov
    Carabus (Megodontus) kantaikensis Gehin, 1885
    Carabus (Megodontus) kantaikensis Gehin, 1885
    Male adult

    Copyright © Kirill Makarov
    Carabus (Acoptolabrus) schrencki schrencki Motschulsky, 1860
    Carabus (Acoptolabrus) schrencki schrencki Motschulsky, 1860
    Female adult

    Copyright © Kirill Makarov
    Carabus (Aulonocarabus) kabakovi Lafer, 1989
    Carabus (Aulonocarabus) kabakovi Lafer, 1989
    Male adult

    Copyright © Kirill Makarov
    Carabus (Aulonocarabus) gossarei gossarei Haury, 1879
    Carabus (Aulonocarabus) gossarei gossarei Haury, 1879
    Male adult

    Copyright © Kirill Makarov
    Carabus (Diocarabus) dorogostaiskii Shilenkov, 1983
    Carabus (Diocarabus) dorogostaiskii Shilenkov, 1983
    Male adult

    Copyright © Kirill Makarov
    Carabus (Damaster) blaptoides rugipennis Motschulsky, 1861
    Carabus (Damaster) blaptoides rugipennis Motschulsky, 1861
    Male adult

    Copyright © Kirill Makarov
    Carabus (Diocarabus) fraterculus odaesaniculus Deuve, 1991
    Carabus (Diocarabus) fraterculus odaesaniculus Deuve, 1991
    Female adult
    S Korea, South Odea Mt., 21.06.2003
    Copyright © Anton Menstschikov
    Carabus (Ulocarabus) stschurovskii Solsky, 1874
    Carabus (Ulocarabus) stschurovskii Solsky, 1874
    Male adult

    Copyright © Kirill Makarov
    Carabus (Morphocarabus) karpinskii Kryzhanovskij & Matveev, 1993
    Carabus (Morphocarabus) karpinskii Kryzhanovskij & Matveev, 1993
    Male adult

    Copyright © Kirill Makarov
    Carabus (Cechenochilus) adangensis Gottwald, 1983
    Carabus (Cechenochilus) adangensis Gottwald, 1983
    Male adult

    Copyright © Kirill Makarov
    Carabus (Cechenochilus) boeberi felix Semenov & Znojko, 1931
    Carabus (Cechenochilus) boeberi felix Semenov & Znojko, 1931
    Female adult

    Copyright © Kirill Makarov
    Carabus (Aulonocarabus) gossarei gossarei Haury, 1879
    Carabus (Aulonocarabus) gossarei gossarei Haury, 1879
    Male adult

    Copyright © Kirill Makarov
    Carabus (Apoplesius) omphreodes Reitter, 1898
    Carabus (Apoplesius) omphreodes Reitter, 1898
    Male adult

    Copyright © Kirill Makarov
    Carabus (Mesocarabus) problematicus harcyniae Sturm, 1815 (as obenbergerianus Breuning, 1927)
    Carabus (Mesocarabus) problematicus harcyniae Sturm, 1815 (as obenbergerianus Breuning, 1927)
    Male adult
    Czech, Bohemia, Plzeň Reg., Domažlice Distr., Babylon, h=500 m, 30.VIII.2008, S.I. Budka leg.
    L=23 mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) violaceus germarii Sturm, 1815
    Carabus (Megodontus) violaceus germarii Sturm, 1815
    Female adult
    Croatia, Ivanščica Mts , Ivanec, 6-13.VI.2003, Krause Rene leg.
    L=36 mm
    Copyright © Ruslan Panin
    Carabus (Chaetocarabus) intricatus krueperi Reitter, 1896
    Carabus (Chaetocarabus) intricatus krueperi Reitter, 1896
    Male adult
    CE-Greece: Pieria, Stomio (Larissa) Ossa Mt. 7-14.VII.2009, Krause Rene leg.
    L=35 mm
    Copyright © Ruslan Panin
    Carabus (Procrustes) coriaceus banaticus Redtenbacher, 1849
    Carabus (Procrustes) coriaceus banaticus Redtenbacher, 1849
    Male adult
    SW Romania, Banat, Băile Herculane env., Domogled valley, 24-30.V.2008, Krause Rene leg.
    L=35 mm
    Copyright © Ruslan Panin
    Carabus (Procrustes) chevrolati thirki Chaudoir, 1857
    Carabus (Procrustes) chevrolati thirki Chaudoir, 1857
    Male adult
    NE Turkey, Trabzon, Zigana Pass, h~2000m, V. Slovak leg.
    L=27 mm
    Copyright © Ruslan Panin
    Carabus (Tachypus) cancellatus graniger Palliardi, 1825
    Carabus (Tachypus) cancellatus graniger Palliardi, 1825
    Male adult
    SW-Romania, Banatului Montan, Resita - Valiug, 29.V-9.VI.2005. Krause R. leg.
    L=25 mm
    Copyright © Ruslan Panin
    Carabus (Eucarabus) catenulatus catenulatus Scopoli, 1763
    Carabus (Eucarabus) catenulatus catenulatus Scopoli, 1763
    Female adult
    NW Croatia, Bosanci, 1-12,VI.2004, Baros Slovak leg.
    L=33 mm
    Copyright © Ruslan Panin
    Carabus (Eucarabus) catenulatus catenulatus Scopoli, 1763
    Carabus (Eucarabus) catenulatus catenulatus Scopoli, 1763
    Male adult
    Italia, Riva San Vitale (Chleti)
    VI.1996 Rubizzani leg.
    male, 29 mm
    Copyright © Alexander Anichtchenko
    Carabus (Eucarabus) cumanus Fischer, 1823
    Carabus (Eucarabus) cumanus Fischer, 1823
    Male+Female adult
    NW Caucasus, Russia,
    Krasnodar reg.,Tuapse distr., Mt.Semashkho 04.V.2008, D.Fominyh leg.
    female, 24mm
    Copyright © Ruslan Panin
    Carabus (Eucarabus) italicus italicus Dejean, 1826
    Carabus (Eucarabus) italicus italicus Dejean, 1826
    Male adult
    Italia, Emilia Romagna, Serramazzoni (Modena)
    700m, 9.VI.1990 M. Po leg.
    male, 22 mm
    Copyright © Alexander Anichtchenko
    Carabus (Eucarabus) italicus italicus Dejean, 1826
    Carabus (Eucarabus) italicus italicus Dejean, 1826
    Male adult
    NE-ITALY,
    Veneto, Hiane, 14.V.1986 Sandec F. leg.
    L= 23mm
    Copyright © Ruslan Panin
    Carabus (Eucarabus) manifestus manifestus Kraatz, 1881
    Carabus (Eucarabus) manifestus manifestus Kraatz, 1881
    Male adult
    China, Beijing, Xiaolongmeng, 2005, Liu Ye coll.;
    L=20mm
    Copyright © Ye Liu
    Carabus (Eucarabus) ulrichi arrogans Schaum, 1858 (as gornjakensis Pavicevic & Toševski, 19
    Carabus (Eucarabus) ulrichi arrogans Schaum, 1858 (as gornjakensis Pavicevic & Toševski, 19
    Male+Female adult
    NE-Serbia, Serbian Carpathians,
    Branicevskom reg., Šetonje, Gornjak, 27.V-10.VI.2006, R.Krause leg.
    Copyright © Ruslan Panin
    Carabus (Eucarabus) ulrichi arrogans Schaum, 1858 (as ceremosnjensis Pavicevic & Toševski,
    Carabus (Eucarabus) ulrichi arrogans Schaum, 1858 (as ceremosnjensis Pavicevic & Toševski,
    Male+Female adult
    NE-Serbia,
    Serbian Carpathians, Homolje Mts., Krepoljin 20-31.V.2006, Sima leg.
    female, 29mm
    Copyright © Ruslan Panin
    Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825 (as superboides Lie, 1989)
    Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825 (as superboides Lie, 1989)
    Female adult
    SW-Romania, Banat,
    Oravita, 5-7.VI.2007, Krause Rene leg.
    female,30mm
    Copyright © Ruslan Panin
    Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825
    Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825
    Male adult
    NE Serbien, Negotim .
    V.1981
    male, 25 mm
    Copyright © Alexander Anichtchenko
    Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825 (as superbus Kraatz, 1878)
    Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825 (as superbus Kraatz, 1878)
    Male+Female adult
    SW-Romania,
    Banat, Moldova Noua, Ravenska viil., 29.V-12.VI.2006, Krause Rene leg.
    female,31mm
    Copyright © Ruslan Panin
    Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825
    Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825
    Female adult
    female, 32mm
    Copyright © Boleslav Brezina
    Carabus (Eucarabus) ulrichi rhilensis Kraatz, 1876 (as sokolari Born, 1904)
    Carabus (Eucarabus) ulrichi rhilensis Kraatz, 1876 (as sokolari Born, 1904)
    Male+Female adult
    Czech Rep., W Bohemia,
    Plzen reg., Tachov Distr., Plana u Marianskych Lazni, 17-29.V.2008, S.I.Budka leg.
    female, 26 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) linnaei macairei Dejean, 1826
    Carabus (Orinocarabus) linnaei macairei Dejean, 1826
    Female adult
    SW-Romania, Banat, Resita - Valiug, 29.V- 9.VI.2005, Krause R. , Baros, Pittner leg.
    L=19 mm
    Copyright © Ruslan Panin
    Carabus (Archiplectes) edithae Reitter, 1893
    Carabus (Archiplectes) edithae Reitter, 1893
    Male adult

    Copyright © Kirill Makarov
    Carabus (Chrysocarabus) lineatus lateralis Chevrolat, 1840 (as quasilineatus Tarrier, 1975)
    Carabus (Chrysocarabus) lineatus lateralis Chevrolat, 1840 (as quasilineatus Tarrier, 1975)
    Male+Female adult
    NW Spain,
    E Asturias, Monte de Moal h~1000m., 5.VI.2008, Pascal Stefani leg.
    female, 27mm.
    Copyright © Ruslan Panin
    Carabus (Aulonocarabus) truncaticollis polaris Poppius, 1905 (as tungusensis Obydov, 2010)
    Carabus (Aulonocarabus) truncaticollis polaris Poppius, 1905 (as tungusensis Obydov, 2010) Holotype
    Female adult
    Evenk Autonomous Region, N. Tunguska Riv., Tura Vil. env., 12.VII.1979
    Copyright © Dmitri Obydov
    Carabus (Carabus) arvensis conciliator Fischer, 1822 (as jurgitae Paratype)
    Carabus (Carabus) arvensis conciliator Fischer, 1822 (as jurgitae Paratype) Paratype
    Male adult
    Russia, Altai, Aktash, VI.1992, V.Shott leg.
    Copyright © Dmitri Obydov
    Carabus (Carabus) arvensis conciliator Fischer, 1822
    Carabus (Carabus) arvensis conciliator Fischer, 1822
    Male adult
    Russia, Barguzin Nat. res., Davshe vill env., hot springs, 14.VII.1990, T. Ananina leg.
    Copyright © Dmitri Obydov
    Carabus (Carabus) arvensis hokkaidoensis Lapouge, 1924
    Carabus (Carabus) arvensis hokkaidoensis Lapouge, 1924
    Male adult
    Russia, South Kuriles ils., Kunashir isl., S slope of Golovinka volcan, 28.VII.1990, S. Pushkarev leg.
    Copyright © Dmitri Obydov
    Carabus (Carabus) arvensis klitini Obydov, 2007
    Carabus (Carabus) arvensis klitini Obydov, 2007 Holotype
    Male adult
    Russia, South Kuriles, Urup isl., Schukina bay, spring 700 м SW Vasina, 24-31.VII.2006, A. Klitin leg.
    Copyright © Dmitri Obydov
    Carabus (Carabus) arvensis klitini Obydov, 2007
    Carabus (Carabus) arvensis klitini Obydov, 2007 Paratype
    Female adult
    Russia, South Kuriles, Urup isl., Schukina bay, spring near Vasina, 22-24.VII.2006, A. Klitin leg.
    Copyright © Dmitri Obydov
    Carabus (Tomocarabus) bessarabicus concretus Fischer, 1823
    Carabus (Tomocarabus) bessarabicus concretus Fischer, 1823
    Habitat
    Orenburg region
    Copyright © Evgeniy Chibilov
    Carabus (Trachycarabus) estreicheri Fischer, 1822 (as jaegeri Ménétriés, 1827)
    Carabus (Trachycarabus) estreicheri Fischer, 1822 (as jaegeri Ménétriés, 1827)
    Female adult
    E UKAINE, Lugansk city, Site Luganskoe, 20-30.VII.2010, V.A.Landik leg.
    L=21 mm
    det.Panin.R.
    Copyright © Ruslan Panin
    Carabus (Goniocarabus) sogdianus Semenov, 1898
    Carabus (Goniocarabus) sogdianus Semenov, 1898
    Male adult
    N Tajikistan, SW Pamir-Alay, Zarafshan Range, Fann Mts, Marguzor Lake circ., Padrut vill., 16.V.1997
    L=21 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) spasskianus geblerianus Obydov, 1999
    Carabus (Morphocarabus) spasskianus geblerianus Obydov, 1999
    Male adult
    Russia, NW Altay, Barnaul env.,23.07.1998, D.Lomakin leg.

    Copyright © Ruslan Panin
    Carabus (Morphocarabus) spasskianus geblerianus Obydov, 1999
    Carabus (Morphocarabus) spasskianus geblerianus Obydov, 1999
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859
    Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859
    Male adult
    Russia, SE Siberia, Transbaikalia, N Chita reg., Kodar Mt. Range, Big Leprindo Lake env.h~1800m., 1.VII.2006, S. Rybalkin leg. L=20 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859
    Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Trachycarabus) sibiricus tscherkessicus Korge, 1964
    Carabus (Trachycarabus) sibiricus tscherkessicus Korge, 1964
    Male adult

    Copyright © Kirill Makarov
    Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
    Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
    Female adult

    Copyright © Kirill Makarov
    Carabus (Tribax) lederi Reitter, 1882
    Carabus (Tribax) lederi Reitter, 1882
    Female adult

    Copyright © Kirill Makarov
    Carabus (Eotribax) hiekei Kabak & Kryzhanovskij, 1990
    Carabus (Eotribax) hiekei Kabak & Kryzhanovskij, 1990
    Female adult

    Copyright © Kirill Makarov
    Carabus (Cratophyrtus) kaufmanni kaufmanni Solsky, 1874
    Carabus (Cratophyrtus) kaufmanni kaufmanni Solsky, 1874
    Female adult

    Copyright © Kirill Makarov
    Carabus (Morphocarabus) rothi hampei (as ormayi Reitter, 1896) loc.typ.
    Carabus (Morphocarabus) rothi hampei (as ormayi Reitter, 1896) loc.typ.
    Male adult
    W. UKRAINE,
    Transcarpathian reg., near Beregovo, 27.IV-22.V..2011,, Panin R. & Chyzhevsky O. leg.
    L=28 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi hampei (as ormayi Reitter, 1896) loc.typ.
    Carabus (Morphocarabus) rothi hampei (as ormayi Reitter, 1896) loc.typ.
    Endophallus
    near Beregovo, 27.IV-22.V..2011,, Panin R. & Chyzhevsky O. leg
    Copyright © Ruslan Panin
    Carabus (Deroplectes) staudingeri Ganglbauer, 1886
    Carabus (Deroplectes) staudingeri Ganglbauer, 1886
    Female adult
    N Tajikistan, SW Pamir-Alay, Fann Mts h=1800m, Padrut vill., 2-3.V.1997
    L=28 mm
    Copyright © Ruslan Panin
    Carabus (Deroplectes) staudingeri Ganglbauer, 1886
    Carabus (Deroplectes) staudingeri Ganglbauer, 1886
    Male adult
    Uzbekhistan, W Gissar mt.rng., 60 km Sakhrisaby 17.5.1994, Klimenko leg.
    male, 25 mm
    Copyright © Alexander Anichtchenko
    Carabus (Deroplectes) staudingeri Ganglbauer, 1886
    Carabus (Deroplectes) staudingeri Ganglbauer, 1886
    Male adult
    male, 29mm
    Copyright © Boleslav Brezina
    Carabus (Deroplectes) staudingeri Ganglbauer, 1886
    Carabus (Deroplectes) staudingeri Ganglbauer, 1886
    Female adult
    female, 30mm
    Copyright © Boleslav Brezina
    Carabus (Morphocarabus) henningi melyachi Shilenkov & Menstshikov, 1996 loc.typ.
    Carabus (Morphocarabus) henningi melyachi Shilenkov & Menstshikov, 1996 loc.typ.
    Male adult
    Russia, Altay, Katun Mt Range, Karakol lake,22.VI.1993, Melyach F.V. leg.<br /> L=18 mm
    Copyright © Ruslan Panin
    Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
    Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
    Male adult
    relict lowland populations of blue-purple color, of the the river Ticino park - Italy - Lombardy
    Copyright © Monzini
    Carabus (Tribax) biebersteini otcharensis Kurnakov, 1970 (as retezari)
    Carabus (Tribax) biebersteini otcharensis Kurnakov, 1970 (as retezari)
    Female adult
    Abkhazia, Lake Ritsa, 43-31'N / 40-37'E, 19-25.VII.2008, А. Vlasenko leg.
    photo from zin.ru
    Copyright © Dmitri Obydov
    Carabus (Tribax) kraatzi Chaudoir, 1877
    Carabus (Tribax) kraatzi Chaudoir, 1877
    Male adult
    Georgia: Achalzich (Chambobel), 1910, Korb leg.
    photo from zin.ru
    Copyright © Dmitri Obydov
    Carabus (Tribax) macropus Chaudoir, 1877
    Carabus (Tribax) macropus Chaudoir, 1877
    Female adult
    Georgia, Khoc(aldagh, prope Lagodechi, Kachetia, VI.1907.
    photo from zin.ru
    Copyright © Dmitri Obydov
    Carabus (Neoplectes) starckianus starckianus (as phlagochensis)
    Carabus (Neoplectes) starckianus starckianus (as phlagochensis)
    Female adult
    Russia, Кrasnodar Reg., Dagomys, Flagokh, N43-44' / E39-40', 11.VII.2010, N. Pichuginleg.
    Specimen from the collection N. U. Pichugin (Vladimir).
    Copyright © Dmitri Obydov
    Carabus (Cechenochilus) boeberi aequaliceps Reitter, 1896
    Carabus (Cechenochilus) boeberi aequaliceps Reitter, 1896
    Male adult
    Checheno-Ingusheti, 20 kilometres eastward Tebulosmta, 2600-3000 m, 9.VI.1986, Komarov leg.
    Specimen from the collection of М. Shestopalova (Moscow).

    Copyright © Dmitri Obydov
    Carabus (Cechenochilus) adangensis Gottwald, 1983 (as gusevi)
    Carabus (Cechenochilus) adangensis Gottwald, 1983 (as gusevi) Paratype
    Male adult
    Paratypus: Carabus (Cechenochilus) gusevi Zamotajlov et Koval, 1989; Abkhazia, Dzyshra, 26.VII.1987, I. Belousov leg.
    Specimen from the collection of М. Shestopalov (Moscow).

    Copyright © Dmitri Obydov
    Carabus (Eurycarabus) genei Gene, 1839
    Carabus (Eurycarabus) genei Gene, 1839
    Male adult
    Corse de Sud, env. de
    Rondinara, VI.1998 Tingaud M. leg.
    male, 23 mm

    Copyright © Alexander Anichtchenko
    Carabus (Eurycarabus) famini fezzanus Lapouge, 1915
    Carabus (Eurycarabus) famini fezzanus Lapouge, 1915
    Female adult
    Marokko, Moy Atlas, Azrou, Ibel Hebri, 1950 m
    leg. Staven/Gremmels, 2000; det. K.Staven, 2000
    length: 22 mm
    Foto: U.Schmidt, 2009
    Copyright © U.Schmidt
    Carabus (Eurycarabus) famini piraticus Fairmaire, 1880
    Carabus (Eurycarabus) famini piraticus Fairmaire, 1880
    Male adult
    Morocco, Safi 10 km
    Khattazakane env. 29.XII.2001, h 100m
    male, 22 mm
    Copyright © Alexander Anichtchenko
    Carabus (Eurycarabus) famini numidus Castelnau, 1835
    Carabus (Eurycarabus) famini numidus Castelnau, 1835
    Male adult

    Copyright © Alexander Anichtchenko
    Carabus (Eurycarabus) famini fezzanus Lapouge, 1915
    Carabus (Eurycarabus) famini fezzanus Lapouge, 1915
    Male adult
    Marokko, Moyen Atlas, s. Azrou
    Jbel Hebri, zedernwald, 1950m, 31.III.2000 Staven/Gremmels leg.
    male, 22 mm
    Copyright © Alexander Anichtchenko
    Carabus (Eurycarabus) famini favieri Fairmaire, 1859
    Carabus (Eurycarabus) famini favieri Fairmaire, 1859
    Female adult
    N Africa, 2 km W Ceuta Allabacal, 17-18.II.2004, Anichtchenko A. leg.
    female, 22 mm
    Copyright © Alexander Anichtchenko
    Carabus (Euleptocarabus) porrecticollis porrecticollis Bates, 1883
    Carabus (Euleptocarabus) porrecticollis porrecticollis Bates, 1883
    Male adult
    Japan: Niigata pref., Chayagahara Jyouetsu City,
    18.XI.2006 A.Toba leg.
    Copyright © Alexander Anichtchenko
    Carabus (Eotribax) valikhanovi Kabak, 1990
    Carabus (Eotribax) valikhanovi Kabak, 1990
    Male adult
    male, 18mm
    Copyright © Boleslav Brezina
    Carabus (Eotribax) valikhanovi Kabak, 1990
    Carabus (Eotribax) valikhanovi Kabak, 1990
    Male adult
    male, 16mm
    Copyright © Boleslav Brezina
    Carabus (Eotribax) malkovskyi malkovskyi Kabak, 1990
    Carabus (Eotribax) malkovskyi malkovskyi Kabak, 1990
    Male adult
    male, 19mm
    Copyright © Boleslav Brezina
    Carabus (Eotribax) malkovskyi malkovskyi (as erenhabergaensis Deuve, 1993)
    Carabus (Eotribax) malkovskyi malkovskyi (as erenhabergaensis Deuve, 1993) Paratype
    Male adult
    male, 16mm
    Copyright © Boleslav Brezina
    Carabus (Eotribax) malkovskyi malkovskyi (as erenhabergaensis Deuve, 1993)
    Carabus (Eotribax) malkovskyi malkovskyi (as erenhabergaensis Deuve, 1993)
    Male adult
    male, 17mm
    Copyright © Boleslav Brezina
    Carabus (Eotribax) malkovskyi malkovskyi (as erenhabergaensis Deuve, 1993)
    Carabus (Eotribax) malkovskyi malkovskyi (as erenhabergaensis Deuve, 1993)
    Male adult
    male, 17mm
    Copyright © Boleslav Brezina
    Carabus (Eotribax) malkovskyi malkovskyi (as erenhabergaensis Deuve, 1993)
    Carabus (Eotribax) malkovskyi malkovskyi (as erenhabergaensis Deuve, 1993)
    Male adult
    male, 18mm
    Copyright © Boleslav Brezina
    Carabus (Eotribax) hiekei Kabak & Kryzhanovskij, 1990
    Carabus (Eotribax) hiekei Kabak & Kryzhanovskij, 1990
    Female adult
    female, 18mm
    Copyright © Boleslav Brezina
    Carabus (Eotribax) hiekei Kabak & Kryzhanovskij, 1990
    Carabus (Eotribax) hiekei Kabak & Kryzhanovskij, 1990
    Male adult
    male, 17mm
    Copyright © Boleslav Brezina
    Carabus (Eotribax) eous ukokensis Belousov & Kabak, 1994
    Carabus (Eotribax) eous ukokensis Belousov & Kabak, 1994 Paratype
    Female adult
    female, 18mm
    Copyright © Boleslav Brezina
    Carabus (Eotribax) eous ukokensis Belousov & Kabak, 1994
    Carabus (Eotribax) eous ukokensis Belousov & Kabak, 1994 Paratype
    Male adult
    male, 17mm
    Copyright © Boleslav Brezina
    Carabus (Eotribax) eous eous Morawitz, 1889
    Carabus (Eotribax) eous eous Morawitz, 1889
    Male adult
    male, 16mm
    Copyright © Boleslav Brezina
    Carabus (Eotribax) eous eous Morawitz, 1889
    Carabus (Eotribax) eous eous Morawitz, 1889
    Female adult
    female, 16mm
    Copyright © Boleslav Brezina
    Carabus (Eotribax) eous eous Morawitz, 1889
    Carabus (Eotribax) eous eous Morawitz, 1889
    Female adult
    female, 16mm
    Copyright © Boleslav Brezina
    Carabus (Eotribax) eous eous Morawitz, 1889
    Carabus (Eotribax) eous eous Morawitz, 1889
    Male adult
    male, 15mm
    Copyright © Boleslav Brezina
    Carabus (Eotribax) eous eous Morawitz, 1889
    Carabus (Eotribax) eous eous Morawitz, 1889
    Male adult
    Kirgizstan, Terskey Ala-Too, Borskanu riv,
    2600 m, 29.5.1999
    Copyright © Alexander Anichtchenko
    Carabus (Eotribax) eous eous Morawitz, 1889
    Carabus (Eotribax) eous eous Morawitz, 1889
    Male adult
    Kirghyzstan, N Terskei Ala Too, Borskaun riv., h=2500 m., 7-14.07.2000
    male
    L=17 mm
    Copyright © Maksim Smirnov
    Carabus (Eotribax) dzhungaricola Deuve, 1989
    Carabus (Eotribax) dzhungaricola Deuve, 1989
    Male adult
    male, 15mm
    Copyright © Boleslav Brezina
    Carabus (Eotribax) dzhungaricola Deuve, 1989
    Carabus (Eotribax) dzhungaricola Deuve, 1989
    Male adult
    male, 15mm
    Copyright © Boleslav Brezina
    Carabus (Eocechenus) yushuensis Deuve, 1991
    Carabus (Eocechenus) yushuensis Deuve, 1991
    Female adult
    female, 15mm
    Copyright © Boleslav Brezina
    Carabus (Eocechenus) takashimai Deuve & Imura, 1993
    Carabus (Eocechenus) takashimai Deuve & Imura, 1993
    Female adult
    female, 20mm
    Copyright © Boleslav Brezina
    Carabus (Eocechenus) takashimai Deuve & Imura, 1993
    Carabus (Eocechenus) takashimai Deuve & Imura, 1993
    Male adult
    male, 20mm
    Copyright © Boleslav Brezina
    Carabus (Eocechenus) noctivagus Deuve, 1992
    Carabus (Eocechenus) noctivagus Deuve, 1992
    Male adult
    male, 21mm
    Copyright © Boleslav Brezina
    Carabus (Eocechenus) mouthiezianoides Deuve & Kalab, 1992
    Carabus (Eocechenus) mouthiezianoides Deuve & Kalab, 1992 Paratype
    Male adult
    male, 18mm
    Copyright © Boleslav Brezina
    Carabus (Eocechenus) leptoplesioides yanjinganus Imura & Mizusawa, 1998
    Carabus (Eocechenus) leptoplesioides yanjinganus Imura & Mizusawa, 1998 Paratype
    Female adult
    female, 20mm
    Copyright © Boleslav Brezina
    Carabus (Eocechenus) leptoplesioides yanjinganus Imura & Mizusawa, 1998
    Carabus (Eocechenus) leptoplesioides yanjinganus Imura & Mizusawa, 1998 Paratype
    Male adult
    male, 20mm
    Copyright © Boleslav Brezina
    Carabus (Eocechenus) kaznakovi kaznakovi (as zadoiensis Lassalle, 2003)
    Carabus (Eocechenus) kaznakovi kaznakovi (as zadoiensis Lassalle, 2003)
    Female adult
    female, 19mm
    Copyright © Boleslav Brezina
    Carabus (Eocechenus) kaznakovi nangqenicus Deuve, 1995
    Carabus (Eocechenus) kaznakovi nangqenicus Deuve, 1995 Paratype
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Eocechenus) kaznakovi kaznakovi Semenov & Znojko, 1932
    Carabus (Eocechenus) kaznakovi kaznakovi Semenov & Znojko, 1932
    Male adult
    L=17 mm
    Copyright © Boleslav Brezina
    Carabus (Eocechenus) kaznakovi angenensis Deuve, 1996
    Carabus (Eocechenus) kaznakovi angenensis Deuve, 1996
    Male adult
    male, 17mm
    Copyright © Boleslav Brezina
    Carabus (Eocechenus) kaznakovi graziosii Deuve, 1995
    Carabus (Eocechenus) kaznakovi graziosii Deuve, 1995
    Female adult
    female, 16 mm
    Copyright © Boleslav Brezina
    Carabus (Eocechenus) ditomoides Deuve, 1991
    Carabus (Eocechenus) ditomoides Deuve, 1991
    Male adult
    male, 19mm
    Copyright © Boleslav Brezina
    Carabus (Eccoptolabrus) exiguus wudumontanus Imura, 1998
    Carabus (Eccoptolabrus) exiguus wudumontanus Imura, 1998
    Male adult
    male, 19mm
    Copyright © Boleslav Brezina
    Carabus (Eccoptolabrus) exiguus tagong Kleinfeld & Puchner, 2007
    Carabus (Eccoptolabrus) exiguus tagong Kleinfeld & Puchner, 2007 Holotype Paratype
    Female adult
    female, 20 mm
    Copyright © Frank Kleinfeld
    Carabus (Eccoptolabrus) exiguus tagong Kleinfeld & Puchner, 2007
    Carabus (Eccoptolabrus) exiguus tagong Kleinfeld & Puchner, 2007 Paratype
    Female adult
    female, 20 mm
    Copyright © Frank Kleinfeld
    Carabus (Eccoptolabrus) exiguus tagong Kleinfeld & Puchner, 2007
    Carabus (Eccoptolabrus) exiguus tagong Kleinfeld & Puchner, 2007 Holotype
    Male adult
    male, 19 mm
    Copyright © Frank Kleinfeld
    Carabus (Eccoptolabrus) exiguus ochotonarum Brezina, 1996
    Carabus (Eccoptolabrus) exiguus ochotonarum Brezina, 1996 Holotype Paratype
    Female adult

    Copyright © Frank Kleinfeld
    Carabus (Eccoptolabrus) exiguus ochotonarum Brezina, 1996
    Carabus (Eccoptolabrus) exiguus ochotonarum Brezina, 1996
    Female adult
    CH. S-Gansu (Labrang) valley E of Ponggartang
    35:14N/102:55E. ca. 2700m conoferous forest
    9.VII.1995 K. & B. Brezina
    Copyright © Frank Kleinfeld
    Carabus (Eccoptolabrus) exiguus ochotonarum Brezina, 1996
    Carabus (Eccoptolabrus) exiguus ochotonarum Brezina, 1996 Paratype
    Male adult

    Copyright © Frank Kleinfeld
    Carabus (Eccoptolabrus) exiguus nivium Breuning, 1933
    Carabus (Eccoptolabrus) exiguus nivium Breuning, 1933
    Male adult
    male, 18mm
    Copyright © Boleslav Brezina
    Carabus (Eccoptolabrus) exiguus nivium Breuning, 1933
    Carabus (Eccoptolabrus) exiguus nivium Breuning, 1933
    Female adult
    female, 20mm
    Copyright © Boleslav Brezina
    Carabus (Eccoptolabrus) exiguus nivium Breuning, 1933
    Carabus (Eccoptolabrus) exiguus nivium Breuning, 1933
    Male adult
    female, 18mm
    Copyright © Boleslav Brezina
    Carabus (Eccoptolabrus) exiguus nivium Breuning, 1933
    Carabus (Eccoptolabrus) exiguus nivium Breuning, 1933 Holotype Paratype
    Female adult
    female, 19 mm
    Copyright © Frank Kleinfeld
    Carabus (Eccoptolabrus) exiguus nivium Breuning, 1933
    Carabus (Eccoptolabrus) exiguus nivium Breuning, 1933
    Male adult
    male, 18 mm
    Copyright © Frank Kleinfeld
    Carabus (Eccoptolabrus) exiguus lanzhouicus Deuve, 1989
    Carabus (Eccoptolabrus) exiguus lanzhouicus Deuve, 1989
    Male adult
    male, 17mm
    Copyright © Boleslav Brezina
    Carabus (Eccoptolabrus) exiguus lanzhouicus Deuve, 1989
    Carabus (Eccoptolabrus) exiguus lanzhouicus Deuve, 1989
    Male adult
    male, 19mm
    Copyright © Boleslav Brezina
    Carabus (Eccoptolabrus) exiguus fanianus Imura, 1993
    Carabus (Eccoptolabrus) exiguus fanianus Imura, 1993 Paratype
    Male adult
    male, 18mm
    Copyright © Boleslav Brezina
    Carabus (Eccoptolabrus) exiguus fanianus Imura, 1993
    Carabus (Eccoptolabrus) exiguus fanianus Imura, 1993
    Female adult
    female, 19mm
    Copyright © Boleslav Brezina
    Carabus (Eccoptolabrus) exiguus exiguus Semenov, 1898
    Carabus (Eccoptolabrus) exiguus exiguus Semenov, 1898
    Female adult
    female, 19mm
    Copyright © Boleslav Brezina
    Carabus (Eccoptolabrus) exiguus exiguus Semenov, 1898
    Carabus (Eccoptolabrus) exiguus exiguus Semenov, 1898 Holotype
    Male adult
    male, 18 mm
    Copyright © Frank Kleinfeld
    Carabus (Eccoptolabrus) exiguus evae Brezina et Hackel, 2007
    Carabus (Eccoptolabrus) exiguus evae Brezina et Hackel, 2007 Holotype
    Male adult
    China, S-Qinghai, 5km S Baima, 30.VI.-1.VII.2006 V. Banes
    long of imago 19 mm
    Copyright © Boleslav Brezina
    Carabus (Eccoptolabrus) exiguus cordimeatus Cavazzuti, 1997
    Carabus (Eccoptolabrus) exiguus cordimeatus Cavazzuti, 1997 Holotype
    Male adult
    male, 19mm
    Copyright © Boleslav Brezina
    Carabus (Eccoptolabrus) exiguus cithara Cavazzuti, 2002
    Carabus (Eccoptolabrus) exiguus cithara Cavazzuti, 2002 Holotype
    Female adult
    female, 22mm
    Copyright © Boleslav Brezina
    Carabus (Eccoptolabrus) exiguus alucinatus Cavazzuti, 2006
    Carabus (Eccoptolabrus) exiguus alucinatus Cavazzuti, 2006 Holotype
    Female adult
    female, 21mm
    Copyright © Boleslav Brezina
    Carabus (Megodontus) vietinghoffi bowringi Chaudoir, 1863
    Carabus (Megodontus) vietinghoffi bowringi Chaudoir, 1863
    Male adult
    Russia, Far East, Primorskiy Kray, Ussuriysk Distr., Kaymanovka vill , 26.VII-3.VIII.2011, A. Korshunov leg.
    L=29
    Copyright © Ruslan Panin
    Carabus (Leptoplesius) schotti Obydov, 1996
    Carabus (Leptoplesius) schotti Obydov, 1996 Holotype
    Female adult
    Киргизия, хр. Терскей, дол. р. Тургей
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) chaudoiri marusiki Obydov, 1997
    Carabus (Morphocarabus) chaudoiri marusiki Obydov, 1997 Holotype
    Male adult
    image from zin.ru
    Copyright © Dmitri Obydov
    Carabus (Megodontus) obovatus Fischer, 1827
    Carabus (Megodontus) obovatus Fischer, 1827
    Male adult
    SW Siberia, Russia, Kemerevo reg., Krapivino distr., Azhendarovo, forest, 26-31.VII.2011, D. Efimov leg.
    L=18 mm
    Copyright © Ruslan Panin
    Carabus (Carabus) arvensis faldermanni Dejean, 1829
    Carabus (Carabus) arvensis faldermanni Dejean, 1829
    Male+Female adult
    Russia, Far East, Primorskiy Kray, Ussuriysk Distr., Kaymanovka vill., 11.VII-1.VIII..2011, A. Korshunov leg.
    female, 23 mm
    Copyright © Ruslan Panin
    Carabus (Diocarabus) slovtzovi rasae Obydov, 2001
    Carabus (Diocarabus) slovtzovi rasae Obydov, 2001 Holotype
    Male adult
    Russia, SE Tuva, pass Balyktyk-Hem, 2300 м, 50°17'N 96°23'E, 27.VI-5.VII.1996, D. Obydov leg.
    Copyright © Dmitri Obydov
    Carabus (Tribax) kasakorum kasakorum Semenov, 1896
    Carabus (Tribax) kasakorum kasakorum Semenov, 1896
    Male adult
    Pr. Wladikavkas, montes, coll. A. Zolotarew
    photo from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Tribax) kasakorum kasakorum Semenov, 1896
    Carabus (Tribax) kasakorum kasakorum Semenov, 1896
    Male adult
    Montes prope Wladikavkas, 6000 f., coll. A. Zolotarew
    photo from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
    Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
    Female adult
    Caucasus, N. Osetia, 60 km S Alagir, 1850 m, 12.IX.1990, A. Klimenko leg.<br /> photo from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
    Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
    Female adult
    Cuaucas, N Osetia, Ardon riv., Alagir, Duravtuan, 28.IV.1982, A.Sokolov leg.<br /> photo from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Tribax) nothus planipennis Chaudoir, 1846
    Carabus (Tribax) nothus planipennis Chaudoir, 1846
    Male adult
    Georgia, Lagodehi, Ninigora, 2750 м, near snow 24.V.1979, I.Belousov leg.
    photo from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Tribax) nothus formaneki Reitter, 1896
    Carabus (Tribax) nothus formaneki Reitter, 1896
    Male adult
    Topotypus: Azerbaijan, Caucas. or., distr. Nukha, 3000 m, 20.VII.1934, A. Bogachev leg.
    Copyright © Dmitri Obydov
    Carabus (Sphodristocarabus) adamsi Adams, 1817 (as chiragricus)
    Carabus (Sphodristocarabus) adamsi Adams, 1817 (as chiragricus)
    Male adult
    Russia, Stavropol reg., Kislovodsk vic., 1,5 км ESE Podkumok 905 м, N43*57E42*47, 13-18.05.2006, A.Vlasenko leg.
    det. Obydov
    Copyright © Dmitri Obydov
    Carabus (Sphodristocarabus) varians janthinus (as dvorshaki)
    Carabus (Sphodristocarabus) varians janthinus (as dvorshaki)
    Male adult
    Abkhasia, Gagra, 12.VI.1986<br /> photo from www.zin.ru
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) excellens excellens (f.i. frivaldszkyi Kraatz, 1887)
    Carabus (Morphocarabus) excellens excellens (f.i. frivaldszkyi Kraatz, 1887)
    Female adult
    W-UKRAINE, NW Podolian Upland,<br /> Lviv Reg., Zolochiv Distr., Lysa Gora steppe massif, 14-30.V.2008, Panin. R. leg./det.<br /> L=26 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) excellens excellens Fabricius, 1798
    Carabus (Morphocarabus) excellens excellens Fabricius, 1798
    Male adult
    NC Ukraine, Kiev, Lysa Gora, on the fringes forest, 5-24.VI.2007, Gerasimov R.P.. leg.<br /> L=25 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) excellens excellens Fabricius, 1798
    Carabus (Morphocarabus) excellens excellens Fabricius, 1798
    Male adult
    C- UKRAINE, NW Odessa reg., Kodyma distr., near Aleksandrivka vill.,18.V.2003, Demidov G.N. leg.<br /> L=26 mm
    Copyright © Ruslan Panin
    Carabus (Platycarabus) fabricii fabricii (as fassatii Smetana, 1955)
    Carabus (Platycarabus) fabricii fabricii (as fassatii Smetana, 1955)
    Male adult
    C-Slovakia:Nízke Tatry Mts., 4.VI.2011, Krause Rene leg.
    L=19 mm
    Copyright © Ruslan Panin
    Carabus (Diocarabus) slovtzovi slovtzovi Mannerheim, 1849
    Carabus (Diocarabus) slovtzovi slovtzovi Mannerheim, 1849
    Female adult
    Russia, East Sayans,Tunkunskye Goltsy Mts., near Hoito-Gol vill., D.E.Lomakin leg
    L=12,5 mm.
    Copyright © Anton Menstschikov
    Carabus (Diocarabus) slovtzovi slovtzovi Mannerheim, 1849
    Carabus (Diocarabus) slovtzovi slovtzovi Mannerheim, 1849
    Female adult
    SW Burjatia, 20 km. S. Kyren vill., Zungol r., 21.06.2007 D.E. Lomakin leg.
    Copyright © Anton Menstschikov
    Carabus (Diocarabus) slovtzovi slovtzovi Mannerheim, 1849
    Carabus (Diocarabus) slovtzovi slovtzovi Mannerheim, 1849
    Male adult
    S Siberia, Burjatia reg., E Sayan Mts., Khugaituma mt., h=1700 m, 1-5. VII.2001
    L=14,3 mm

    Copyright © Maksim Smirnov
    Carabus (Diocarabus) massagetus massagetus Motschulsky, 1846
    Carabus (Diocarabus) massagetus massagetus Motschulsky, 1846
    Male adult
    Khakassia, Bograd Distr., env. Borozda Vill., forest VII.2006, M.Shestopalov leg.
    L=18.0 mm
    Copyright © Kirill Makarov
    Carabus (Diocarabus) loschnikovi loschnikovi Fischer, 1823
    Carabus (Diocarabus) loschnikovi loschnikovi Fischer, 1823
    Male adult
    RUSSIA, Tuva, Tanu-Ola Mts., near Shurmak village, h=1500m , 23-30.VI.1997
    Copyright © - - -
    Carabus (Diocarabus) fraterculus fraterculus Reitter, 1895
    Carabus (Diocarabus) fraterculus fraterculus Reitter, 1895
    Male adult
    Kaolingzu st., Kirin pr., 18.VII.1990
    L=16 mm
    Copyright © Alexander Anichtchenko
    Carabus (Diocarabus) dorogostaiskii Shilenkov, 1983
    Carabus (Diocarabus) dorogostaiskii Shilenkov, 1983
    Female adult
    Chita reg., Sokhondo, Kyra vill., 18.VII.1990 Shilenkov V. leg.
    L= 15 mm
    Copyright © Alexander Anichtchenko
    Carabus (Diocarabus) chamissonis Fischer, 1822
    Carabus (Diocarabus) chamissonis Fischer, 1822
    Male adult
    Canada
    Copyright Henri Goulet, Agriculture & Agri-Food Canada
    Copyright © Henri Goulet
    Carabus (Diocarabus) caustomarginatus Imura & Mizusawa, 1994
    Carabus (Diocarabus) caustomarginatus Imura & Mizusawa, 1994
    Male adult

    Copyright © Alexander Anichtchenko
    Carabus (Diocarabus) beybienkoi Kryzhanovskij, 1973
    Carabus (Diocarabus) beybienkoi Kryzhanovskij, 1973
    Male adult
    S Sakhalin, Susunaj Mt.R. 10.08 - 13.09.2003 D. Kochetkov leg.
    L=14,5 mm
    Copyright © Maksim Smirnov
    Carabus (Deroplectes) sphinx sphinx (as pseudocoiffaitianus Deuve, 1990)
    Carabus (Deroplectes) sphinx sphinx (as pseudocoiffaitianus Deuve, 1990)
    Female adult
    L=30 mm
    Copyright © Boleslav Brezina
    Carabus (Deroplectes) sphinx sphinx (as pseudocoiffaitianus Deuve, 1990)
    Carabus (Deroplectes) sphinx sphinx (as pseudocoiffaitianus Deuve, 1990)
    Male adult
    L=25 mm
    Copyright © Boleslav Brezina
    Carabus (Deroplectes) sphinx dardazicus Deuve, 1990
    Carabus (Deroplectes) sphinx dardazicus Deuve, 1990
    Female adult
    female, 28mm
    Copyright © Boleslav Brezina
    Carabus (Deroplectes) sphinx dardazicus Deuve, 1990
    Carabus (Deroplectes) sphinx dardazicus Deuve, 1990
    Male adult
    male, 27mm
    Copyright © Boleslav Brezina
    Carabus (Deroplectes) dokhtouroffi pseudomidas Mandl, 1967
    Carabus (Deroplectes) dokhtouroffi pseudomidas Mandl, 1967
    Male adult
    male, 28mm
    Copyright © Boleslav Brezina
    Carabus (Deroplectes) dokhtouroffi pseudomidas Mandl, 1967
    Carabus (Deroplectes) dokhtouroffi pseudomidas Mandl, 1967
    Male adult
    male, 28mm
    Copyright © Boleslav Brezina
    Carabus (Deroplectes) dokhtouroffi dokhtouroffi Ganglbauer, 1886
    Carabus (Deroplectes) dokhtouroffi dokhtouroffi Ganglbauer, 1886
    Female adult
    female, 32mm
    Copyright © Boleslav Brezina
    Carabus (Deroplectes) dokhtouroffi dokhtouroffi Ganglbauer, 1886
    Carabus (Deroplectes) dokhtouroffi dokhtouroffi Ganglbauer, 1886
    Male adult
    male, 28mm
    Copyright © Boleslav Brezina
    Carabus (Deroplectes) dokhtouroffi ? pseudomidas Mandl, 1967
    Carabus (Deroplectes) dokhtouroffi ? pseudomidas Mandl, 1967
    Male adult
    Vahshskii Mt. rng., Saryhosor, Mullohomi 28.4.1990 S. L. Zonshtein leg.

    Copyright © Alexander Anichtchenko
    Carabus (Deroplectes) arcanus arcanus Semenov, 1898
    Carabus (Deroplectes) arcanus arcanus Semenov, 1898
    Female adult
    female, 25mm
    Copyright © Boleslav Brezina
    Carabus (Deroplectes) arcanus arcanus Semenov, 1898
    Carabus (Deroplectes) arcanus arcanus Semenov, 1898
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Morphocarabus) praecellens trans scheidleri
    Carabus (Morphocarabus) praecellens trans scheidleri
    Endophallus
    NC Croatia, Ivanščica Mts , Ivanec,4-18.VI.2011, Krause Rene leg.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) praecellens trans scheidleri
    Carabus (Morphocarabus) praecellens trans scheidleri
    Male adult
    NC Croatia, Ivanščica Mts , Ivanec,4-18.VI.2011, Krause Rene leg.<br /> L=31 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) praecellens praecellens Pillardi, 1825
    Carabus (Morphocarabus) praecellens praecellens Pillardi, 1825
    Male adult
    Serbia ,
    Frushka Gora, Ruma VI.1985
    Copyright © Igor Solodovnikov
    Carabus (Procrustes) coriaceus hopffgarteni (as emgei Ganglbauer, 1888)
    Carabus (Procrustes) coriaceus hopffgarteni (as emgei Ganglbauer, 1888)
    Male adult
    CE-Greece: Larissa reg. Stomio, Ossa Mt. 6-17.VII.2011, Krause Rene leg.
    L=32 mm
    Copyright © Ruslan Panin
    Carabus (Chaetocarabus) arcadicus arcadicus Gistl, 1850
    Carabus (Chaetocarabus) arcadicus arcadicus Gistl, 1850
    Female adult
    CE-Greece, Kato Olympos, Karia env., 6-17.VII.2011, Krause Rene leg. L=37 mm
    Copyright © Ruslan Panin
    Carabus (Procerus) gigas gigas Creutzer, 1799
    Carabus (Procerus) gigas gigas Creutzer, 1799
    Male+Female adult
    N Croatia, Ivanščica Mts , Ivanec, 4-18.VI.2011, Krause Rene leg.
    female, 54 mm
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) problematicus inflatus Kraatz, 1878
    Carabus (Mesocarabus) problematicus inflatus Kraatz, 1878
    Male adult
    SE–France, Var departament, Foret royale -Saint-Paul-en-Foret vill.,18.VI.2002, Thelot J.P. leg.
    L=26 mm
    Copyright © Ruslan Panin
    Carabus (Chrysocarabus) solieri solieri Dejean, 1826
    Carabus (Chrysocarabus) solieri solieri Dejean, 1826
    Female adult
    SE France, Alpes-Maritimes, Ascros, VI.2011
    L=30 mm
    Copyright © Ruslan Panin
    Carabus (Pachystus) cribratus cribratus Quensel, 1806
    Carabus (Pachystus) cribratus cribratus Quensel, 1806
    Male adult
    Turkey, Gьmьshane<br /> leg. det. K.Staven, 1992<br /> length: 26 mm
    Copyright © U.Schmidt
    Carabus (Pachystus) cribratus cribratus Quensel, 1806
    Carabus (Pachystus) cribratus cribratus Quensel, 1806
    Male adult

    Copyright © Safronov
    Carabus (Pachystus) cribratus cribratus Quensel, 1806
    Carabus (Pachystus) cribratus cribratus Quensel, 1806
    Male adult
    near Tbilisi, Bakuriani<br /> 7.VI.1973, Shilenkov V. leg.<br /> male, 25 mm
    Copyright © Alexander Anichtchenko
    Carabus (Cyclocarabus) pseudolamprostus Kalab, 1996
    Carabus (Cyclocarabus) pseudolamprostus Kalab, 1996
    Male adult
    Kirgizstan, S slope of Pskem mt. rng.,
    Chandalash riv., 2400m, 1.6.2000
    male, 24 mm
    Copyright © Alexander Anichtchenko
    Carabus (Cyclocarabus) namanganensis namanganensis Heyden, 1886
    Carabus (Cyclocarabus) namanganensis namanganensis Heyden, 1886
    Female adult
    Uzbekistan, Karzhantau Mt., Aktash vill.
    1500 m, 12.4.1999
    L= 23 mm
    Copyright © Alexander Anichtchenko
    Carabus (Cyclocarabus) namanganensis namanganensis Heyden, 1886
    Carabus (Cyclocarabus) namanganensis namanganensis Heyden, 1886
    Male adult
    Uzbekistan, Karzhantau Mt., Aktash vill.
    1500 m, 12.4.1999
    L= 22 mm
    Copyright © Alexander Anichtchenko
    Carabus (Cyclocarabus) karaterekensis hemicallisthenes Kabak & Kalab, 1997
    Carabus (Cyclocarabus) karaterekensis hemicallisthenes Kabak & Kalab, 1997
    Male adult
    Kirgizstan, Chandalash, Chakmak-Suu riv.,
    2800 m, 1-2.6.2000
    male, 17 mm
    Copyright © Alexander Anichtchenko
    Carabus (Cyclocarabus) brosciformis Semenov, 1896
    Carabus (Cyclocarabus) brosciformis Semenov, 1896
    Male adult

    Copyright © Alexander Anichtchenko
    Carabus (Cyclocarabus) aulacocnemus uzbek Deuve & Kalab, 1992
    Carabus (Cyclocarabus) aulacocnemus uzbek Deuve & Kalab, 1992
    Female adult
    Uzbekistan, 2800m, Chatkalskii mt. rng.,
    Jangi-Abad vill., bass. Angren riv., 6.6.2000 Tihonov D. leg.
    L= 24 mm
    Copyright © Alexander Anichtchenko
    Carabus (Cyclocarabus) aulacocnemus nurbeki Kalab, 1997
    Carabus (Cyclocarabus) aulacocnemus nurbeki Kalab, 1997
    Male adult
    Kirgizstan, S slope Pskem, Chandalash riv.,
    2200 m, 31.5.2000
    male, 21 mm
    Copyright © Alexander Anichtchenko
    Carabus (Cyclocarabus) aulacocnemus aulacocnemus ( as minusculus Semenov, 1903)
    Carabus (Cyclocarabus) aulacocnemus aulacocnemus ( as minusculus Semenov, 1903)
    Male adult
    W Kirgizstan, Talas mt. rng, Manas mt., 1800 m
    22.4.2000
    male, 18 mm
    Copyright © Alexander Anichtchenko
    Carabus (Cychrostomus) pseudoprosodes Semenov & Znojko, 1932
    Carabus (Cychrostomus) pseudoprosodes Semenov & Znojko, 1932
    Male adult
    male, 19mm
    Copyright © Boleslav Brezina
    Carabus (Cychrostomus) anchocephalus anchocephalus (as facetus Semenov, 1903)
    Carabus (Cychrostomus) anchocephalus anchocephalus (as facetus Semenov, 1903)
    Female adult
    L=24mm
    Copyright © Boleslav Brezina
    Carabus (Cychrostomus) anchocephalus lanzhouensis Deuve, 1991
    Carabus (Cychrostomus) anchocephalus lanzhouensis Deuve, 1991
    Male adult
    male, 24mm
    Copyright © Boleslav Brezina
    Carabus (Cychrostomus) anchocephalus lanzhouensis Deuve, 1991
    Carabus (Cychrostomus) anchocephalus lanzhouensis Deuve, 1991 Paratype
    Male adult
    male, 21mm
    Copyright © Boleslav Brezina
    Carabus (Cychrostomus) anchocephalus anchocephalus Reitter, 1896
    Carabus (Cychrostomus) anchocephalus anchocephalus Reitter, 1896
    Male adult
    male, 22mm
    Copyright © Boleslav Brezina
    Carabus (Cychrostomus) anchocephalus anchocephalus Reitter, 1896
    Carabus (Cychrostomus) anchocephalus anchocephalus Reitter, 1896
    Male adult
    China, Shanxi pro., Jiaochen
    Copyright © Ye Liu
    Carabus (Platycarabus) depressus lucens Schaum, 1856
    Carabus (Platycarabus) depressus lucens Schaum, 1856
    Female adult
    SE-France: Alpi Marittime, Monte Viso, 14.VII.2010
    L=23 mm
    Copyright © Ruslan Panin
    Carabus (Oreocarabus) luetgensi getschmanni Lapouge, 1924
    Carabus (Oreocarabus) luetgensi getschmanni Lapouge, 1924
    Male adult
    NW-Spain, León Province, Acebo, h=1000m, V.2007, Pascal Stefani leg.
    L=22 mm
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) problematicus planiusculus Haury in Gehin, 1885
    Carabus (Mesocarabus) problematicus planiusculus Haury in Gehin, 1885
    Male adult
    NE Spain, Catalonia, Sant Jaume de Frontanyà, h=1000 m, V-VI.2010, Pascal Stefani leg.
    L=24 mm
    Copyright © Ruslan Panin
    Carabus (Tachypus) auratus lotharingus Dejean, 1826
    Carabus (Tachypus) auratus lotharingus Dejean, 1826
    Male adult
    SC France: Herault departament, Foret de Saint Gervais, V. 2009 leg. J. Nicolas
    L=27mm
    Copyright © Ruslan Panin
    Carabus (Eucarabus) obsoletus nagyagensis Seidlitz, 1888
    Carabus (Eucarabus) obsoletus nagyagensis Seidlitz, 1888
    Female adult
    NW-Romania, Mts Ses (Plopis),Cluj Reg., Bucea env, h=580 m, 8.VI.2011, Sl.Budka leg.
    L=26 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi hampei Kuster, 1846
    Carabus (Morphocarabus) rothi hampei Kuster, 1846
    Male adult
    C Romania, Târgu Mureş, Ernei, 46°35′471″N 24°37’351″E, h=300 m,20.VIII.2004, Sl.Budka leg.
    L=30 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi hampei Kuster, 1846
    Carabus (Morphocarabus) rothi hampei Kuster, 1846
    Endophallus
    C Romania, Tвrgu Mureş,
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi hampei (as zilahiensis Csiki, 1906)
    Carabus (Morphocarabus) rothi hampei (as zilahiensis Csiki, 1906)
    Male+Female adult
    NW Romania, Zalău env., 31.VII.2003, Sl.Budka leg.
    L=24 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi hampei (as zilahiensis Csiki, 1906)
    Carabus (Morphocarabus) rothi hampei (as zilahiensis Csiki, 1906)
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Platycarabus) fabricii fabricii (as fassatii Smetana, 1955)
    Carabus (Platycarabus) fabricii fabricii (as fassatii Smetana, 1955)
    Male adult
    C-Slovakia:Nizke Tatry Mts., h=1750 m,.9.VI.2007, V.Slovak leg.
    L=20 mm
    Copyright © Ruslan Panin
    Carabus (Macrothorax) aumonti maroccanus Bedel, 1895
    Carabus (Macrothorax) aumonti maroccanus Bedel, 1895
    Male adult
    N Morocco (Spain:Melilla), hipodromo, X.1969, leg. Pardo Alcaide
    L=38 mm
    Copyright © Ruslan Panin
    Carabus (Tomocarabus) bessarabicus ? concretus Fischer, 1823
    Carabus (Tomocarabus) bessarabicus ? concretus Fischer, 1823
    Male adult
    E Ukraine, Lugannsk Reg., Sverdlovsk distr., Steppe Reserve “Provalsky", 3.V-3.VI.2011, leg.Glotov S./det.Panin R.<br /> female, 22 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi comptus (as szoerenyensis Csiki, 1908)
    Carabus (Morphocarabus) rothi comptus (as szoerenyensis Csiki, 1908)
    Female adult
    SW Romania, Tarcu Mts, vf. Matania Mt. h=2000 m, 12.VI.2008, S.Budka leg.
    L=23 mm
    Copyright © Ruslan Panin
    Carabus (Pachystus) hungaricus hungaricus Fabricius, 1792
    Carabus (Pachystus) hungaricus hungaricus Fabricius, 1792
    Female adult
    C Hungary, Pest reg., Örkény, 9-10.V.1997, S.Budka leg.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi hampei (as hunyadensis Csiki, 1926)
    Carabus (Morphocarabus) rothi hampei (as hunyadensis Csiki, 1926)
    Male adult
    CW Romania, Apuseni Mts, Muntii Metaliferi, Hunedoara reg., near Sacaramb vill., 6.VI.2011, S.Budka leg.
    L=37 mm
    Copyright © Ruslan Panin
    Carabus (Tomocarabus) bessarabicus ? concretus Fischer, 1823
    Carabus (Tomocarabus) bessarabicus ? concretus Fischer, 1823
    Endophallus
    Steppe Reserve “Provalsky",
    Copyright © Ruslan Panin
    Carabus (Megodontus) vietinghoffi bowringi (as lazoensis Obydov, 1999)
    Carabus (Megodontus) vietinghoffi bowringi (as lazoensis Obydov, 1999)
    Male+Female adult
    Russia, Far East, Primorsky reg., S. Sichote-Alin Mts.R., Kavalerovo vill. env.,
    5-25.VII.1992, S. Kunitzyn leg.
    male - 28 mm, female - 30 mm.
    Copyright © Alexandr Safronov
    Carabus (Megodontus) vietinghoffi bowringi Chaudoir, 1863
    Carabus (Megodontus) vietinghoffi bowringi Chaudoir, 1863
    Male+Female adult
    Russia, Far East, Primorsky reg., Khasansky distr.,
    Zanadvorovka vill., Gusevsky pit.
    M - 19-21.VI.2007, body lenght 29 mm.
    F - 15.VII.2008, body lenght 30 mm.
    S. Veriga leg.
    Copyright © Alexandr Safronov
    Carabus (Megodontus) vietinghoffi bowringi Chaudoir, 1863
    Carabus (Megodontus) vietinghoffi bowringi Chaudoir, 1863
    Female adult
    Tian Tou, Yueshi County, Anhui Prov., CHINA
    1-10.VIII.2004 local coll.
    female - 10 mm.
    Copyright © Alexandr Safronov
    Carabus (Neoplectes) satyrus duripshensis Kurnakov, 1972
    Carabus (Neoplectes) satyrus duripshensis Kurnakov, 1972
    Male+Female adult
    W.Caucasus, Abkhasia, Gudauta distr.,
    Duripsh village env., Khipsta valley, h=500-700 m.
    07-12.VI.2009 A.Safronov leg.
    Body lenght 30-33 mm.
    Copyright © Alexandr Safronov
    Carabus (Cyclocarabus) aulacocnemus galinae Obydov, 2003
    Carabus (Cyclocarabus) aulacocnemus galinae Obydov, 2003 Paratype
    Male+Female adult
    W-Tianshan:Karatau Mts.,N-part:Boialdyr River valley,
    N of Ken-su, 600-750m, 22-V-2000 М. Danilevsky leg.
    male - 20 мм, female - 19 мм.
    Copyright © Alexandr Safronov
    Carabus (Cyclocarabus) namanganensis namanganensis Heyden, 1886
    Carabus (Cyclocarabus) namanganensis namanganensis Heyden, 1886
    Male+Female adult
    Kazakhstan, S. Karatau Mts. R., Mashat vill. env.
    23-IV-1993 М. Danilevsky leg.
    male - 21 мм, female - 22 мм.
    Copyright © Alexandr Safronov
    Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878 (as viridipennis Kraatz, 1886, nec Lewis,18
    Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878 (as viridipennis Kraatz, 1886, nec Lewis,18
    Male+Female adult
    Russia, Far East,Primorsky reg., S. Sikhote-Alin Mts. R., Kavalerovo vill. env.,
    5-25.VII.1992, С. Kunitzyn leg.
    male - 25 mm, female - 25 mm.
    Copyright © Alexandr Safronov
    Carabus (Morphocarabus) hummeli pusongensis Imura, 1993
    Carabus (Morphocarabus) hummeli pusongensis Imura, 1993
    Male+Female adult
    Russia, Far East, Primorsky ter., Khasansky distr., Zanadvorovka vill.,
    Gusevsky pit, S.Veriga leg.
    male - 13.VII.2008, female 19-21.VI.2007
    male - 28 мм, female - 28 мм.
    Copyright © Alexandr Safronov
    Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
    Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
    Male+Female adult
    Russia, Far East, Primorsky ter., Chuguevsky distr., 30 km ESE from Chuguevka, 9-14.VIII.2007 S. Shabalin leg. male - 25 mm, female - 26 mm.
    Copyright ©
    Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878 (as purpuripennis Kraatz, 1886)
    Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878 (as purpuripennis Kraatz, 1886)
    Male adult
    Russia, Khabarovsk reg., near Komsomolsk-on-Amur.
    7-21.VI.2010, A. Kapkaev leg.
    Male left - 24 mm, male right - 23 mm.
    Copyright © Alexandr Safronov
    Carabus (Morphocarabus) hummeli hummeli Fischer, 1823
    Carabus (Morphocarabus) hummeli hummeli Fischer, 1823
    Male+Female adult
    Russia, Far East, Khabarovsk reg., Obluchye env.
    M. Khingan Mts. R.
    10-20.VII.1992 S. Andreev leg.
    male - 26 mm, female - 27 mm.
    Copyright © Alexandr Safronov
    Carabus (Pantophyrtus) turcomanorum turcomanorum Thieme, 1881
    Carabus (Pantophyrtus) turcomanorum turcomanorum Thieme, 1881
    Male+Female adult
    Kyrgyzstan, Kyrgyzskyi Mts. R., Uzun Bulak Gorge, h=1800 ms.
    01-09.VI.2002 loc. coll.
    male - 28 mm, female - 33 mm.
    Copyright © Alexandr Safronov
    Carabus (Neoplectes) satyrus duripshensis Kurnakov, 1972
    Carabus (Neoplectes) satyrus duripshensis Kurnakov, 1972
    Endophallus

    Copyright © Safronov
    Carabus (Cupreocarabus) lixianensis jinchuanicus Deuve & Tian, 2007
    Carabus (Cupreocarabus) lixianensis jinchuanicus Deuve & Tian, 2007
    Male adult
    male, 22 mm
    Copyright © Boleslav Brezina
    Carabus (Cupreocarabus) lixianensis jinchuanicus Deuve & Tian, 2007
    Carabus (Cupreocarabus) lixianensis jinchuanicus Deuve & Tian, 2007
    Female adult
    female, 24 mm
    Copyright © Boleslav Brezina
    Carabus (Cupreocarabus) lixianensis gamdaensis Deuve & Tian, 2002
    Carabus (Cupreocarabus) lixianensis gamdaensis Deuve & Tian, 2002 Paratype
    Male adult
    male, 22mm
    Copyright © Boleslav Brezina
    Carabus (Cupreocarabus) laevithorax taiyanghensis Deuve & Tian, 2006
    Carabus (Cupreocarabus) laevithorax taiyanghensis Deuve & Tian, 2006
    Male adult
    male, 21 mm
    Copyright © Boleslav Brezina
    Carabus (Cupreocarabus) laevithorax taiyanghensis Deuve & Tian, 2006
    Carabus (Cupreocarabus) laevithorax taiyanghensis Deuve & Tian, 2006
    Female adult
    female, 23 mm
    Copyright © Boleslav Brezina
    Carabus (Cupreocarabus) kangdingi sculptior Deuve, 1992
    Carabus (Cupreocarabus) kangdingi sculptior Deuve, 1992
    Female adult
    female, 25mm
    Copyright © Boleslav Brezina
    Carabus (Cupreocarabus) kangdingi sculptior Deuve, 1992
    Carabus (Cupreocarabus) kangdingi sculptior Deuve, 1992
    Male adult
    male, 22mm
    Copyright © Boleslav Brezina
    Carabus (Cupreocarabus) kangdingi sculptior Deuve, 1992
    Carabus (Cupreocarabus) kangdingi sculptior Deuve, 1992
    Male adult
    male, 23mm
    Copyright © Boleslav Brezina
    Carabus (Cupreocarabus) kangdingi kangdingi Korell, Kleinfeld & Gorgner, 1992
    Carabus (Cupreocarabus) kangdingi kangdingi Korell, Kleinfeld & Gorgner, 1992 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Cupreocarabus) drahoslavae Brezina B. & Hackel M., 2006
    Carabus (Cupreocarabus) drahoslavae Brezina B. & Hackel M., 2006 Paratype
    Female adult

    Copyright © Boleslav Brezina
    Carabus (Cupreocarabus) drahoslavae Brezina B. & Hackel M., 2006
    Carabus (Cupreocarabus) drahoslavae Brezina B. & Hackel M., 2006 Holotype
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Cupreocarabus) brezinai Deuve, 1994
    Carabus (Cupreocarabus) brezinai Deuve, 1994
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Cupreocarabus) brezinai Deuve, 1994
    Carabus (Cupreocarabus) brezinai Deuve, 1994
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Ctenocarabus) galicianus Gory, 1839
    Carabus (Ctenocarabus) galicianus Gory, 1839
    Male adult
    Spain,
    Sierra de la Cabrera Baja, 8 km SW Truchillas, 9.VII.2008, Anichtchenko A. leg.
    L= 20 mm
    Copyright © Alexander Anichtchenko
    Carabus (Cryptocechenus) sifanicus Semenov, 1898 (as oxydocupreus)
    Carabus (Cryptocechenus) sifanicus Semenov, 1898 (as oxydocupreus)
    Male adult
    male, 17mm
    Copyright © Boleslav Brezina
    Carabus (Cryptocechenus) kalabi Deuve, 1990
    Carabus (Cryptocechenus) kalabi Deuve, 1990
    Male adult
    male, 18mm
    Copyright © Boleslav Brezina
    Carabus (Cryptocechenus) kalabi Deuve, 1990 (as cuprifuscatus)
    Carabus (Cryptocechenus) kalabi Deuve, 1990 (as cuprifuscatus) Paratype
    Female adult

    Copyright © Boleslav Brezina
    Carabus (Cryptocarabus) subparallelus Ballion, 1878
    Carabus (Cryptocarabus) subparallelus Ballion, 1878
    Female adult
    Kazakhstan,
    near Alma-Ata, V.2009, Pak O.leg.
    female, 17mm
    Copyright © Ruslan Panin
    Carabus (Cryptocarabus) sacarum Kabak, 1998
    Carabus (Cryptocarabus) sacarum Kabak, 1998
    Male adult
    SE Kazakhstan, Zailiiskii Ala too, Kaskelen riv.,
    1400m 24.V.2000
    Copyright © Alexander Anichtchenko
    Carabus (Sphodristocarabus) adamsi Adams, 1817 (as chiragricus Fischer, 1828)
    Carabus (Sphodristocarabus) adamsi Adams, 1817 (as chiragricus Fischer, 1828)
    Male+Female adult
    Russia, N. Caucasus, Stavropol reg., Zheleznovodsk, Zheleznaya Mt., h=600 m. 5-11.VI.2011, Е. Shankhiza leg. male - 24 mm, female - 25 mm.
    Copyright ©
    Carabus (Sphodristocarabus) varians varians Fischer, 1827
    Carabus (Sphodristocarabus) varians varians Fischer, 1827
    Male+Female adult
    N. Armenia, Idzhevan distr., Acharkut vill. env., h=800 m.<br /> 26-28.VII.2008 S. Andreev leg.<br /> male - 22 mm, female - 23 mm.
    Copyright ©
    Carabus (Cryptocarabus) mullerellus kasantsevi Kabak, 1993
    Carabus (Cryptocarabus) mullerellus kasantsevi Kabak, 1993
    Male adult
    SE Kazakhstan, Alma-Ata reg., Narynkol vill.
    12.V.2000 Klimenko A. leg.
    Copyright © Alexander Anichtchenko
    Carabus (Cryptocarabus) lindemanni Ballion, 1878
    Carabus (Cryptocarabus) lindemanni Ballion, 1878
    Male adult
    SE Kazakhstan, Alma-Ata env., Kok-Tube 16.IV.2000
    L=15mm
    Copyright © Alexander Anichtchenko
    Carabus (Cryptocarabus) lindemanni Ballion, 1878
    Carabus (Cryptocarabus) lindemanni Ballion, 1878
    Female adult
    SE Kazakhstan, Alma-Ata env., 16.04. 2000
    L=16,0 mm
    Copyright © Maksim Smirnov
    Carabus (Cryptocarabus) kadyrbekovi Kabak 1994
    Carabus (Cryptocarabus) kadyrbekovi Kabak 1994
    Male adult
    SE Kazakhstan, Zailijskii Ala Too, Karakastek riv., 27.5.2000
    Copyright © Alexander Anichtchenko
    Carabus (Cechenotribax) petri Semenov & Znojko, 1932
    Carabus (Cechenotribax) petri Semenov & Znojko, 1932
    Male adult
    SE Kazakhstan, N-Tianshan:N-slopes Ketmen Mt.Rng. h=2250-2500m., near Ketmen vill.,1-5.VI.1990, Dolin V.G. leg.
    L=13.5 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) regalis tschusovae Obydov, 2004
    Carabus (Morphocarabus) regalis tschusovae Obydov, 2004
    Male+Female adult
    NE-Kazakhstan, W Altay, Ivanovsky Mt.Rng, near Ridder (Leninogorsk), h-1300-1500m, 4-6.VII.2003
    female, 25 mm
    det.Panin R.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) regalis tschusovae Obydov, 2004
    Carabus (Morphocarabus) regalis tschusovae Obydov, 2004
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Neoplectes) satyrus duripshensis Kurnakov, 1972
    Carabus (Neoplectes) satyrus duripshensis Kurnakov, 1972
    Female adult

    Copyright © Alexander Anichtchenko
    Carabus (Neoplectes) satyrus duripshensis Kurnakov, 1972
    Carabus (Neoplectes) satyrus duripshensis Kurnakov, 1972
    Male adult
    W.Caucasus, Abkhasia, Gudauta distr.,
    Duripsh village env., Khipsta valley, h=500-700 m.
    07-12.VI.2009 A.Safronov leg.
    male - 30 mm.
    Copyright © Alexandr Safronov
    Carabus (Morphocarabus) regalis jenissoni Dejean, 1831 (as cyanescens Kraatz, 1882, nec Sturm, 1815)
    Carabus (Morphocarabus) regalis jenissoni Dejean, 1831 (as cyanescens Kraatz, 1882, nec Sturm, 1815)
    Male+Female adult
    Kazakhstan, E. Kazakhstan area,
    Zyryanovsk env., near Putintzevo vill.
    10-VI-2005 M. Danilevsky leg.
    male - 24 mm, female - 25 mm.
    Copyright © Alexandr Safronov
    Carabus (Ophiocarabus) regeli obscurior Semenov, 1888
    Carabus (Ophiocarabus) regeli obscurior Semenov, 1888
    Male+Female adult
    Kyrgyzstan, NE Tian-Shan, E. Terskey Alatau Mts. R.,
    Kok-kia River, h=2800 м.
    7-8.VI.1997 М. Danilevsky leg.
    Body lenght 15-18 mm.
    Copyright © Alexandr Safronov
    Carabus (Semnocarabus) carbonicolor carbonicolor Morawitz, 1886
    Carabus (Semnocarabus) carbonicolor carbonicolor Morawitz, 1886
    Male+Female adult
    Kyrgyzstan, NC Tianshan, Kyrgyz Mt. R., Tuyuk riv., h=2000 м.
    25-VI-1997 М. Danilevsky leg.
    male - 20 mm, female - 21 mm.
    Copyright © Alexandr Safronov
    Carabus (Semnocarabus) carbonicolor carbonicolor Morawitz, 1886
    Carabus (Semnocarabus) carbonicolor carbonicolor Morawitz, 1886
    Endophallus

    Copyright © Alexandr Safronov
    Carabus (Pachycarabus) roseri soganliensis Heinz & Korge, 1964
    Carabus (Pachycarabus) roseri soganliensis Heinz & Korge, 1964
    Male+Female adult
    Turkey, Rize prov., Ikizdere distr., Rize Daglari,
    47 km to SE from lyidere, Sivrikaya Koyu, h=2025 m.
    30.VI-5.VII.2011, A. Vlasenko leg.
    male - 23 mm, female - 24 mm.
    Copyright © Alexandr Safronov
    Carabus (Neoplectes) juenthneri avadcharensis Kurnakov, 1970
    Carabus (Neoplectes) juenthneri avadcharensis Kurnakov, 1970
    Male+Female adult
    W. Caucasus, Abkhasia, 8 km NE of Ritza Lake,<br /> Avadkhara valley, h=1700 m<br /> 10-VI-2009 A.Safronov leg.
    Copyright ©
    Carabus (Procrustes) chevrolati thirki Chaudoir, 1857
    Carabus (Procrustes) chevrolati thirki Chaudoir, 1857
    Male+Female adult
    Turkey, Rize prov., Ikizdere distr., Rize Daglari,
    47 km to SE from lyidere, Sivrikaya Koyu, h=2025 m.
    30.VI-5.VII.2011, A. Vlasenko leg.
    male - 28 mm, female - 31 mm.
    Copyright © Alexandr Safronov
    Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
    Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
    Male+Female adult
    SW Siberia, Kemerovo reg., Krapivino distr.,<br /> 8 km SSW Saltymakovo vill., h=150 m.<br /> 1-20.VII.2010 D. Efimov leg.<br /> male - 25 mm, female - 26 mm.
    Copyright ©
    Carabus (Morphocarabus) spasskianus aeruginosiformis Breuning, 1932
    Carabus (Morphocarabus) spasskianus aeruginosiformis Breuning, 1932
    Male+Female adult
    Russia, S. Siberia, Krasnoyarsk reg., W. Sayan,<br /> Sayanogorsk distr., Cheremushki vill. env.,<br /> 07-VII-1994 М. Danilevsky leg.<br /> male - 22 mm, female - 25 mm.
    Copyright ©
    Carabus (Megodontus) danilevskii Obydov, 1993
    Carabus (Megodontus) danilevskii Obydov, 1993
    Male+Female adult
    Russia, SW Siberia, Krasnoyarsk reg., W. Sayan, near Cheryomushki.
    26-29.VII.1996 loc. coll.
    male - 18mm, female - 20 mm.
    Copyright © Alexandr Safronov
    Carabus (Megodontus) kantaikensis Gehin, 1885
    Carabus (Megodontus) kantaikensis Gehin, 1885
    Female adult
    Russia, SW Siberia, W. Sayan, Krasnoyarsk reg.,
    S of Abakan, near Cheryomushki, Dzhoysky Pass.
    07-VII-1994 M. Danilevsky leg.
    body lenght 18 mm.
    Copyright © Alexandr Safronov
    Carabus (Megodontus) obovatus Fischer, 1827
    Carabus (Megodontus) obovatus Fischer, 1827
    Male adult
    SW Siberia, Kemerovo reg., Krapivino distr.,
    8 km SSW Saltymakovo vill., h=150 m.
    20-30.VI.2010 D. Efimov, A. Korshunov leg.
    body lenght 18 mm.
    Copyright © Alexandr Safronov
    Carabus (Megodontus) imperialis Fischer, 1823
    Carabus (Megodontus) imperialis Fischer, 1823
    Male adult
    NE Kazakhstan, Zyrjanovsk distr., Putincevo vill. env.
    25-VI-2005 M. Danilevsky leg.
    male 24 mm, female 26 mm.
    Copyright © Safronov
    Carabus (Morphocarabus) helleri Ganglbauer, 1893
    Carabus (Morphocarabus) helleri Ganglbauer, 1893
    Male+Female adult
    CZECH, SE Moravia, Velka nad Velickou. 20-V-2005, V. Slovak leg. male - 26 mm, female - 30 mm.
    Copyright © Alexandr Safronov
    Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825
    Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825
    Male+Female adult
    Male - SW Romania, Banat - Bocsa/Coltan,
    21.V-05.VI.2010, R. Krause leg., L=29 mm.
    Female - SW Romania, Banat - Saska Montana, Stinapari,
    25-VI-2010, R. Krause leg., L=31 mm.
    Copyright © Alexandr Safronov
    Carabus (Megodontus) violaceus violaceus Linne, 1758 (as intermarinus Blumenthal & Staven, 1985)
    Carabus (Megodontus) violaceus violaceus Linne, 1758 (as intermarinus Blumenthal & Staven, 1985)
    Male+Female adult
    Poland, Gdansk distr., Gdynya env., Reservat «Cisowa»,
    5-9.08.1998 I. Solodovnikov leg.
    female - 26 mm.
    Copyright © Alexandr Safronov
    Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860 (as ursuius Lapouge, 1911)
    Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860 (as ursuius Lapouge, 1911)
    Male+Female adult
    FRANCE (64), Ursuya Mt.
    XII-2009, P. Stefani leg.
    male - 30 mm, female - 32 mm.
    Copyright © Alexandr Safronov
    Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896
    Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896
    Male adult
    Anyos (ANDORRE), h=1450ms,
    5-VI-2010, P. Stefani leg.
    body lenght - 32 mm.
    Copyright © Alexandr Safronov
    Carabus (Tribax) biebersteini otcharensis Kurnakov, 1970
    Carabus (Tribax) biebersteini otcharensis Kurnakov, 1970
    Male adult
    W. Caucasus, Abkhazia, Gudauta distr.,
    Duripsh vill. env., Khipsta River, h=700 m,
    11-06-2009 A. Safronov leg.
    male - 40 mm
    Copyright © Alexandr Safronov
    Carabus (Neoplectes) polychrous polychrous Rost, 1892
    Carabus (Neoplectes) polychrous polychrous Rost, 1892
    Female adult
    W.Caucasus, Abkhasia, Gudauta distr., Duripsh village env., Khipsta valley, h=500-700 m. 8-11.06.2009 A. Safronov leg. lenght 40-43 mm.
    Copyright ©
    Carabus (Sphodristocarabus) varians janthinus s.str. (as dvorshaki Mandl, 1975)
    Carabus (Sphodristocarabus) varians janthinus s.str. (as dvorshaki Mandl, 1975)
    Male+Female adult
    W. Caucasus, Abkhazia, Gudauta distr..,<br /> Duripsh vill. env., Khipsta riv., h=400-500 м.<br /> 03-13.VI.2009 A. Safronov leg.<br /> body lenght 20-25 мм.
    Copyright ©
    Carabus (Sphodristocarabus) varians (as dvorshaki Mandl, 1975)
    Carabus (Sphodristocarabus) varians (as dvorshaki Mandl, 1975)
    Endophallus

    Copyright ©
    Carabus (Cratophyrtus) turcosinensis turcosinensis (as korelli Kleinfeld, 1988)
    Carabus (Cratophyrtus) turcosinensis turcosinensis (as korelli Kleinfeld, 1988) Holotype
    Male adult

    Copyright © Frank Kleinfeld
    Carabus (Cratophyrtus) kaufmanni segregatus Morawitz, 1886
    Carabus (Cratophyrtus) kaufmanni segregatus Morawitz, 1886
    Female adult
    female, 19mm
    Copyright © Boleslav Brezina
    Carabus (Cratophyrtus) kaufmanni segregatus Morawitz, 1886
    Carabus (Cratophyrtus) kaufmanni segregatus Morawitz, 1886
    Male adult
    male, 18mm
    Copyright © Boleslav Brezina
    Carabus (Cratophyrtus) kaufmanni lebedevianus Breuning, 1933
    Carabus (Cratophyrtus) kaufmanni lebedevianus Breuning, 1933
    Male adult
    male, 19mm
    Copyright © Boleslav Brezina
    Carabus (Cratophyrtus) kaufmanni lebedevianus Breuning, 1933
    Carabus (Cratophyrtus) kaufmanni lebedevianus Breuning, 1933
    Male adult
    male, 20mm
    Copyright © Boleslav Brezina
    Carabus (Cratophyrtus) kaufmanni kaufmanni Solsky, 1874
    Carabus (Cratophyrtus) kaufmanni kaufmanni Solsky, 1874
    Female adult
    female, 19mm
    Copyright © Boleslav Brezina
    Carabus (Cratophyrtus) kaufmanni kaufmanni Solsky, 1874
    Carabus (Cratophyrtus) kaufmanni kaufmanni Solsky, 1874
    Male adult
    male, 19mm
    Copyright © Boleslav Brezina
    Carabus (Cratophyrtus) katajevi katajevi Gottwald, 1989
    Carabus (Cratophyrtus) katajevi katajevi Gottwald, 1989
    Male adult
    male, 25mm
    Copyright © Boleslav Brezina
    Carabus (Cratophyrtus) hauseri Reitter, 1894
    Carabus (Cratophyrtus) hauseri Reitter, 1894
    Female adult
    female, 19mm
    Copyright © Boleslav Brezina
    Carabus (Cratophyrtus) hauseri Reitter, 1894
    Carabus (Cratophyrtus) hauseri Reitter, 1894
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Tanaocarabus) sylvosus Say, 1825
    Carabus (Tanaocarabus) sylvosus Say, 1825
    Female adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Carabus (Carabus) vinctus Weber, 1801
    Carabus (Carabus) vinctus Weber, 1801
    Male adult
    Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
    Copyright © - - -
    Carabus (Apotomopterus) dechambreianus dechambreianus Deuve & Li, 2000
    Carabus (Apotomopterus) dechambreianus dechambreianus Deuve & Li, 2000
    Male+Female adult
    Gailin, Guanxi prov., CHINA,
    17-V-2011 loc. coll.
    male - 36 mm, female - 38 mm.
    Copyright © Alexandr Safronov
    Carabus (Apotomopterus) shun floridus Cavazzuti & Ratti, 1998
    Carabus (Apotomopterus) shun floridus Cavazzuti & Ratti, 1998
    Male+Female adult
    CHINA, Guangdong prov., Mts. Nanling Reserve, 1-7.VII.2011 loc. coll. male - 38 mm, female - 40 mm.
    Copyright © Alexandr Safronov
    Carabus (Neoplectes) reitteri gagrinus Starck, 1894 (as gegicus Gottwald, 1985)
    Carabus (Neoplectes) reitteri gagrinus Starck, 1894 (as gegicus Gottwald, 1985)
    Male+Female adult
    W.Caucasus, Abkhasia, Gagra distr., Gega riv. valley,
    13-15.07.2005 D. Kasatkin. leg.
    male - 30 mm, female - 34 mm.
    Copyright © Alexandr Safronov
    Carabus (Neoplectes) apollo tenebricosus Kurnakov, 1962
    Carabus (Neoplectes) apollo tenebricosus Kurnakov, 1962
    Endophallus

    Copyright © Alexandr Safronov
    Carabus (Morphocarabus) rothi hampei as telekii Csiki, 1937 (? hampei )
    Carabus (Morphocarabus) rothi hampei as telekii Csiki, 1937 (? hampei )
    Female adult
    CW Romania, Arad distr., Zarandulu Mt Range, near Ilteu vill., h=276 m, 46°66′487″N 22°20’455″E, 25.VI.2005, Sl.Budka leg.
    L=30 mm

    Copyright © Ruslan Panin
    Carabus (Cratocephalus) solskyi Ballion, 1878
    Carabus (Cratocephalus) solskyi Ballion, 1878
    Female adult
    female, 35mm
    Copyright © Boleslav Brezina
    Carabus (Cratocephalus) solskyi Ballion, 1878
    Carabus (Cratocephalus) solskyi Ballion, 1878
    Male adult
    SE Kazakhstan, Ketmen Mt. rng., Tuyuk vill., 16.06.97 leg. S.Toropov
    Copyright © Maksim Smirnov
    Carabus (Cratocephalus) narynensis Csiki, 1927
    Carabus (Cratocephalus) narynensis Csiki, 1927
    Female adult

    Copyright © Alexander Anichtchenko
    Carabus (Cratocephalus) narynensis Csiki, 1927
    Carabus (Cratocephalus) narynensis Csiki, 1927
    Female adult
    female, 25mm
    Copyright © Boleslav Brezina
    Carabus (Cratocephalus) corrugis Dohrn, 1882
    Carabus (Cratocephalus) corrugis Dohrn, 1882
    Male adult

    Copyright © Alexandr Safronov
    Carabus (Cratocephalus) corrugis Dohrn, 1882
    Carabus (Cratocephalus) corrugis Dohrn, 1882
    Male adult
    male, 24mm
    Copyright © Boleslav Brezina
    Carabus (Cratocephalus) cicatricosus cicatricosus Fischer, 1842
    Carabus (Cratocephalus) cicatricosus cicatricosus Fischer, 1842
    Male adult
    SE Kazakhstan, Ketmen mt.rng., Tuyuk vill., 1800m, 15.5.2000
    Copyright © Alexander Anichtchenko
    Carabus (Cratocephalus) cicatricosus cicatricosus Fischer, 1842
    Carabus (Cratocephalus) cicatricosus cicatricosus Fischer, 1842
    Male adult
    female, 33mm
    Copyright © Boleslav Brezina
    Carabus (Cratocephalus) cicatricosus cicatricosus Fischer, 1842
    Carabus (Cratocephalus) cicatricosus cicatricosus Fischer, 1842
    Male adult
    male, 28mm
    Copyright © Boleslav Brezina
    Carabus (Cratocephalus) cicatricosus cicatricosus Fischer, 1842
    Carabus (Cratocephalus) cicatricosus cicatricosus Fischer, 1842
    Male adult
    KAZAKHSTAN, Bolshoe Almatinskoe Lake, 30.VI.1992
    L=26mm
    Copyright © Ruslan Panin
    Carabus (Cratocephalus) chan Breuning, 1934
    Carabus (Cratocephalus) chan Breuning, 1934
    Male adult
    male, 23mm
    Copyright © Boleslav Brezina
    Carabus (Cratocephalus) balassogloi Dohrn, 1882
    Carabus (Cratocephalus) balassogloi Dohrn, 1882
    Male adult

    Copyright © Alexander Anichtchenko
    Carabus (Cratocephalus) balassogloi Dohrn, 1882
    Carabus (Cratocephalus) balassogloi Dohrn, 1882
    Male adult
    KIRGIZSTAN, N Issyk-Kul Lake, Ananievo env., 1800 m; 20.04.2000
    Copyright © Anton Menstschikov
    Carabus (Cratocephalus) balassogloi Dohrn, 1882
    Carabus (Cratocephalus) balassogloi Dohrn, 1882
    Male adult
    male, 23mm
    Copyright © Boleslav Brezina
    Carabus (Cratocechenus) wrzecionkoi HT (HYBRID ?Eotribax valikhanovi/Cratocechenus akinini puellus )
    Carabus (Cratocechenus) wrzecionkoi HT (HYBRID ?Eotribax valikhanovi/Cratocechenus akinini puellus ) Holotype
    Male adult

    Copyright © Boleslav Brezina
    Carabus (Cratocechenus) tshistjakovae tshistjakovae Kabak, 2001
    Carabus (Cratocechenus) tshistjakovae tshistjakovae Kabak, 2001 Paratype
    Male adult
    male, 24mm
    Copyright © Boleslav Brezina
    Carabus (Cratocechenus) redikortzewi Semenov & Breuning, 1931
    Carabus (Cratocechenus) redikortzewi Semenov & Breuning, 1931
    Female adult
    female, 17mm
    Copyright © Boleslav Brezina
    Carabus (Cratocechenus) redikortzewi Semenov & Breuning, 1931
    Carabus (Cratocechenus) redikortzewi Semenov & Breuning, 1931
    Male adult
    male, 15mm
    Copyright © Boleslav Brezina
    Carabus (Cratocechenus) ovtchinnikovi ovtchinnikovi Gottwald, 1987
    Carabus (Cratocechenus) ovtchinnikovi ovtchinnikovi Gottwald, 1987
    Male adult

    Copyright © Alexander Anichtchenko
    Carabus (Cratocechenus) ovtchinnikovi ovtchinnikovi Gottwald, 1987
    Carabus (Cratocechenus) ovtchinnikovi ovtchinnikovi Gottwald, 1987
    Female adult
    female, 18mm
    Copyright © Boleslav Brezina
    Carabus (Cratocechenus) ovtchinnikovi letellierianus Deuve, 1990
    Carabus (Cratocechenus) ovtchinnikovi letellierianus Deuve, 1990
    Female adult
    female, 21mm
    Copyright © Boleslav Brezina
    Carabus (Cratocechenus) ovtchinnikovi letellierianus Deuve, 1990
    Carabus (Cratocechenus) ovtchinnikovi letellierianus Deuve, 1990
    Male adult
    male, 18mm
    Copyright © Boleslav Brezina
    Carabus (Cratocechenus) ovtchinnikovi kafkai Deuve, 1994
    Carabus (Cratocechenus) ovtchinnikovi kafkai Deuve, 1994
    Female adult
    female, 21mm
    Copyright © Boleslav Brezina
    Carabus (Cratocechenus) jacobsoni Semenov, 1898
    Carabus (Cratocechenus) jacobsoni Semenov, 1898
    Female adult
    female, 16mm
    Copyright © Boleslav Brezina
    Carabus (Cratocechenus) jacobsoni Semenov, 1898
    Carabus (Cratocechenus) jacobsoni Semenov, 1898
    Male adult
    male, 15mm
    Copyright © Boleslav Brezina
    Carabus (Cratocechenus) jacobsoni x akinini elisabethae HYBRID
    Carabus (Cratocechenus) jacobsoni x akinini elisabethae HYBRID
    Female adult
    female, 18mm
    Copyright © Boleslav Brezina
    Carabus (Cratocechenus) jacobsoni Semenov, 1898
    Carabus (Cratocechenus) jacobsoni Semenov, 1898
    Male adult
    S Kazakhstan, Almaty reg., Big Lake, h=2800 m, 25.VI.96
    male
    L=14 mm
    Copyright © Maksim Smirnov
    Carabus (Cratocechenus) akinini puellus Lapouge, 1924
    Carabus (Cratocechenus) akinini puellus Lapouge, 1924
    Male adult
    male, 19mm
    Copyright © Boleslav Brezina
    Carabus (Cratocechenus) akinini puellus Lapouge, 1924
    Carabus (Cratocechenus) akinini puellus Lapouge, 1924
    Female adult
    female, 21mm
    Copyright © Boleslav Brezina
    Carabus (Cratocechenus) akinini akinini Morawitz, 1886 (as oitalensis )
    Carabus (Cratocechenus) akinini akinini Morawitz, 1886 (as oitalensis ) Paratype
    Male adult
    male, 20mm
    Copyright © Boleslav Brezina
    Carabus (Cratocechenus) akinini musartianus Gottwald, 1987
    Carabus (Cratocechenus) akinini musartianus Gottwald, 1987
    Female adult
    female, 22mm
    Copyright © Boleslav Brezina
    Carabus (Cratocechenus) akinini musartianus Gottwald, 1987
    Carabus (Cratocechenus) akinini musartianus Gottwald, 1987
    Male adult
    male, 21mm
    Copyright © Boleslav Brezina
    Carabus (Cratocechenus) akinini loudai Gottwald, 1987
    Carabus (Cratocechenus) akinini loudai Gottwald, 1987
    Female adult
    female, 23mm
    Copyright © Boleslav Brezina
    Carabus (Cratocechenus) akinini loudai Gottwald, 1987
    Carabus (Cratocechenus) akinini loudai Gottwald, 1987
    Male adult
    male, 20mm
    Copyright © Boleslav Brezina
    Carabus (Cratocechenus) akinini loudai Gottwald, 1987
    Carabus (Cratocechenus) akinini loudai Gottwald, 1987
    Male adult
    male, 21mm
    Copyright © Boleslav Brezina
    Carabus (Cratocechenus) akinini ketmenensis Dolin, 1991
    Carabus (Cratocechenus) akinini ketmenensis Dolin, 1991
    Male adult
    male, 22mm
    Copyright © Boleslav Brezina
    Carabus (Cratocechenus) akinini ketmenensis Dolin, 1991
    Carabus (Cratocechenus) akinini ketmenensis Dolin, 1991
    Male adult
    male, 21mm
    Copyright © Boleslav Brezina
    Carabus (Cratocechenus) akinini loudai (as issykatae Chromý, 1993)
    Carabus (Cratocechenus) akinini loudai (as issykatae Chromý, 1993) Paratype
    Male adult
    male, 21mm
    Copyright © Boleslav Brezina
    Carabus (Cratocechenus) akinini loudai (as issykatae Chromý, 1993)
    Carabus (Cratocechenus) akinini loudai (as issykatae Chromý, 1993) Paratype
    Male adult
    female, 23mm
    Copyright © Boleslav Brezina
    Carabus (Cratocechenus) akinini elisabethae Semenov, 1908
    Carabus (Cratocechenus) akinini elisabethae Semenov, 1908
    Female adult
    female, 23mm
    Copyright © Boleslav Brezina
    Carabus (Cratocechenus) akinini elisabethae Semenov, 1908
    Carabus (Cratocechenus) akinini elisabethae Semenov, 1908
    Female adult
    female, 22mm
    Copyright © Boleslav Brezina
    Carabus (Cratocechenus) akinini elisabethae Semenov, 1908
    Carabus (Cratocechenus) akinini elisabethae Semenov, 1908
    Male adult
    male, 20mm
    Copyright © Boleslav Brezina
    Carabus (Cratocechenus) akinini elisabethae Semenov, 1908
    Carabus (Cratocechenus) akinini elisabethae Semenov, 1908
    Male adult
    male, 21mm
    Copyright © Boleslav Brezina
    Carabus (Cratocechenus) akinini buffi Deuve & Kalab, 1992
    Carabus (Cratocechenus) akinini buffi Deuve & Kalab, 1992
    Male adult
    male, 18mm
    Copyright © Boleslav Brezina
    Carabus (Cratocechenus) akinini akinini Morawitz, 1886
    Carabus (Cratocechenus) akinini akinini Morawitz, 1886
    Female adult
    female, 21mm
    Copyright © Boleslav Brezina
    Carabus (Cratocechenus) akinini akinini Morawitz, 1886
    Carabus (Cratocechenus) akinini akinini Morawitz, 1886
    Male adult
    male, 18mm
    Copyright © Boleslav Brezina
    Carabus (Cratocechenus) akinini akinini Morawitz, 1886
    Carabus (Cratocechenus) akinini akinini Morawitz, 1886
    Male adult
    male, 20mm
    Copyright © Boleslav Brezina
    Carabus (Cratocechenus) akinini akinini Morawitz, 1886
    Carabus (Cratocechenus) akinini akinini Morawitz, 1886
    Male adult
    male, 22mm
    Copyright © Boleslav Brezina
    Carabus (Cratocarabus) puer Morawitz, 1886
    Carabus (Cratocarabus) puer Morawitz, 1886
    Female adult
    female, 16mm
    Copyright © Boleslav Brezina
    Carabus (Cratocarabus) kryzhanovskii molossulus Deuve, 1993
    Carabus (Cratocarabus) kryzhanovskii molossulus Deuve, 1993 Paratype
    Male adult
    mele, 16mm
    Copyright © Boleslav Brezina
    Carabus (Cratocarabus) kryzhanovskii molossulusas (as laevimolossulus Deuve, 1993)
    Carabus (Cratocarabus) kryzhanovskii molossulusas (as laevimolossulus Deuve, 1993) Paratype
    Male adult
    female, 17mm
    Copyright © Boleslav Brezina
    Carabus (Cratocarabus) kryzhanovskii kryzhanovskii Bogachev, 1965
    Carabus (Cratocarabus) kryzhanovskii kryzhanovskii Bogachev, 1965
    Male adult
    male, 15mm
    Copyright © Boleslav Brezina
    Carabus (Coptolabrus) smaragdinus yanggangensis Deuve & Li, 2000
    Carabus (Coptolabrus) smaragdinus yanggangensis Deuve & Li, 2000
    Male+Female adult
    Mt. Tongnimpeak, h=1520 m, Ryang Gong Province, N.Korea, 1-18.VI.2008
    Copyright ©
    Carabus (Coptolabrus) smaragdinus mandschuricus Semenov, 1898
    Carabus (Coptolabrus) smaragdinus mandschuricus Semenov, 1898
    Male adult
    Russia, Far East, South Primoriye, Kaymanovka village, 10-20.07.2000; P.Gorbunov leg.
    Copyright © Anton Menstschikov
    Carabus (Coptolabrus) smaragdinus mandschuricus Semenov, 1898
    Carabus (Coptolabrus) smaragdinus mandschuricus Semenov, 1898
    Male adult
    Primorje, Lasovskii distr., Chandalaz mt., L.Voita leg. 19.VII.2008
    Copyright © Anton Menstschikov
    Carabus (Coptolabrus) smaragdinus longipennis Chaudoir, 1863
    Carabus (Coptolabrus) smaragdinus longipennis Chaudoir, 1863
    Male+Female adult

    Copyright © Alexandr Safronov
    Carabus (Coptolabrus) smaragdinus longipennis Chaudoir, 1863
    Carabus (Coptolabrus) smaragdinus longipennis Chaudoir, 1863
    Female adult
    Huolongling, Beitaizi, Huadian City, Jilin Prov., CHINE
    1-8.VI.2007
    Copyright © Alexandr Safronov
    Carabus (Coptolabrus) smaragdinus longipennis (as mucdenensis Hauser, 1921)
    Carabus (Coptolabrus) smaragdinus longipennis (as mucdenensis Hauser, 1921)
    Male adult
    Mt. Wulongshan, Dandong City,
    Liaoning Province, CHINE
    25-30.VI.2008
    Copyright © Alexandr Safronov
    Carabus (Coptolabrus) smaragdinus honanensis Hauser, 1921
    Carabus (Coptolabrus) smaragdinus honanensis Hauser, 1921
    Male+Female adult
    Mt. Wangwu, Henan Prov., CHINA
    03.06.2006
    Copyright © Alexandr Safronov
    Carabus (Coptolabrus) smaragdinus hoenggandoensis (Kwon & Lee, 1984)
    Carabus (Coptolabrus) smaragdinus hoenggandoensis (Kwon & Lee, 1984)
    Female adult
    Chujado Isl., S. Korea, 21.07.2006; 40 mm.
    Copyright ©
    Carabus (Coptolabrus) smaragdinus furumiellus Deuve, 1994
    Carabus (Coptolabrus) smaragdinus furumiellus Deuve, 1994
    Male+Female adult
    Hua-Bo-zi-Shan, Huanren County,Liaoning Province, CHINE, 20-30.VI.2008
    Copyright ©
    Carabus (Coptolabrus) pustulifer shangzhouicus Deuve & Li, 2003
    Carabus (Coptolabrus) pustulifer shangzhouicus Deuve & Li, 2003
    Male+Female adult
    Shahezi, Shangzhou City, Shaanxi Province, CHINE 11-22.VI.2008
    Copyright ©
    Carabus (Coptolabrus) pustulifer pustulifer Lucas, 1869
    Carabus (Coptolabrus) pustulifer pustulifer Lucas, 1869
    Female adult
    China, Sichuan, Leshan, June,
    800m,
    female, 40 mm

    Copyright © Alexander Anichtchenko
    Carabus (Coptolabrus) pustulifer pustulifer Lucas, 1869
    Carabus (Coptolabrus) pustulifer pustulifer Lucas, 1869
    Female adult
    Chine, C.Sichuan, Ya'an, 07.2005 Local people
    Copyright © Igor Solodovnikov
    Carabus (Coptolabrus) pustulifer pustulifer Lucas, 1869
    Carabus (Coptolabrus) pustulifer pustulifer Lucas, 1869
    Male adult
    Chine, C.Sichuan, Ya'an, 07.2005 Local people

    Copyright © Igor Solodovnikov
    Carabus (Coptolabrus) pustulifer mirificus Kraatz, 1894
    Carabus (Coptolabrus) pustulifer mirificus Kraatz, 1894
    Male+Female adult
    Mt. Julong, Baokang, Hubei Prov., CHINE
    25.VII.2007
    Copyright © Alexandr Safronov
    Carabus (Coptolabrus) pustulifer mirificus Kraatz, 1894
    Carabus (Coptolabrus) pustulifer mirificus Kraatz, 1894
    Male adult
    Chine, SW-Hubei, Enshi, VI.2005 Local people
    Copyright © Igor Solodovnikov
    Carabus (Coptolabrus) pustulifer mirificus Kraatz, 1894
    Carabus (Coptolabrus) pustulifer mirificus Kraatz, 1894
    Male adult
    Chine, SW-Hubei, Enshi, 06.2005, Local people
    Copyright © Igor Solodovnikov
    Carabus (Coptolabrus) pustulifer guerryi Born, 1903
    Carabus (Coptolabrus) pustulifer guerryi Born, 1903
    Female adult
    N Yunnan: Gaoligongshan, h =1300 m, 06.2003 Local people
    Copyright © Igor Solodovnikov
    Carabus (Coptolabrus) pustulifer (as cyanelongatus Deuve, 2006)
    Carabus (Coptolabrus) pustulifer (as cyanelongatus Deuve, 2006)
    Female adult
    Zhagyab, Quamdu City, Tibet<br /> 8-15.VI.2008
    Copyright ©
    Carabus (Chrysocarabus) lineatus salmantinus Bolivar, 1922
    Carabus (Chrysocarabus) lineatus salmantinus Bolivar, 1922
    Male adult
    CW Spain, Cáceres, San Martín de Trevejo,24.VI.2011, A. Diez leg.
    L=24 mm
    Copyright © Ruslan Panin
    Carabus (Macrothorax) rugosus boeticus Deyrolle, 1852
    Carabus (Macrothorax) rugosus boeticus Deyrolle, 1852
    Male adult
    SC Spain, Jaen reg., Puente de la Sierra, 10.VII.2011, M.Lopez leg.
    L=28 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi rothi Dejean, 1829
    Carabus (Morphocarabus) rothi rothi Dejean, 1829
    Male+Female adult
    C Romania, Sibiu Area, right bank Olt River, Bradu vill., h=380 m, 19.VIII.2001, S.Budka leg.
    L=28 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi rothi Dejean, 1829
    Carabus (Morphocarabus) rothi rothi Dejean, 1829
    Endophallus
    C Romania, Sibiu Area, right bank Olt River, Bradu vill., h=380 m,
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) venustus venustus Morawitz, 1862
    Carabus (Morphocarabus) venustus venustus Morawitz, 1862
    Male adult
    Russia, Far East, Primorskiy reg., Ussuriysk Distr., Kaymanovka vill., 6-15.VIII.2011,
    A. Korshunov leg.
    L= 21.5 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) venustus venustus Morawitz, 1862
    Carabus (Morphocarabus) venustus venustus Morawitz, 1862
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Pachystus) hungaricus cribellatus Adams, 1812
    Carabus (Pachystus) hungaricus cribellatus Adams, 1812
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Pachystus) hungaricus cribellatus Adams, 1812
    Carabus (Pachystus) hungaricus cribellatus Adams, 1812
    Male adult
    E Siberia, Russia, Krasnoyarsk Reg., near Krasnoyarsk, forest-steppe,1.VII-20.VIII,2011,M.Ivanov leg.
    L=23 mm
    Copyright © Ruslan Panin
    Carabus (Pachystus) hungaricus scythus Motschulsky, 1847
    Carabus (Pachystus) hungaricus scythus Motschulsky, 1847
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Pachycarabus) roseri soganliensis Heinz & Korge, 1964
    Carabus (Pachycarabus) roseri soganliensis Heinz & Korge, 1964
    Female adult
    NE-Turkey: Rize Prov.Ovit Pass (40.37;40.46) h-2700m,15.VII.2008, I.Rapuzzi & Caldon leg.
    L=23 mm
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) sibiricus sibiricus Fischer, 1822
    Carabus (Trachycarabus) sibiricus sibiricus Fischer, 1822
    Male adult
    E Siberia, Russia, Krasnoyarsk Reg., near Krasnoyarsk, on dry steppe slopes, 7.VIII,2011, M.Ivanov leg.
    L=21 mm
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) sibiricus sibiricus Fischer, 1822
    Carabus (Trachycarabus) sibiricus sibiricus Fischer, 1822
    Endophallus
    near Krasnoyarsk
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) latus latus Dejean, 1826
    Carabus (Mesocarabus) latus latus Dejean, 1826
    Female adult
    CW Spain, Cáceres, San Martín de Trevejo, 24.VI.2011, A. Diez leg./det.Panin R.
    L=27 mm
    Copyright © Ruslan Panin
    Carabus (Coptolabrus) pustulifer Lucas, 1869
    Carabus (Coptolabrus) pustulifer Lucas, 1869
    Habitat
    China, North Guizhou pro., Fanjinshan mountain, 2008.7.19, 2000m, Liu ye leg.;
    photo & det. by liuye
    Copyright © Ye Liu
    Carabus (Coptolabrus) smaragdinus Fischer, 1823
    Carabus (Coptolabrus) smaragdinus Fischer, 1823
    Habitat
    China, Liaoning pro., 2009.8.1, Liu ye leg.
    Copyright © Ye Liu
    Carabus (Chaetocarabus) arcadicus Gistl, 1850
    Carabus (Chaetocarabus) arcadicus Gistl, 1850
    Habitat
    Mt. Olymbos, Grecia, 1.I.2007 photo Anichtchenko
    Copyright © Alexander Anichtchenko
    Carabus (Imaibius) olafi Deuve & J.Schmidt, 2007
    Carabus (Imaibius) olafi Deuve & J.Schmidt, 2007
    Habitat
    Nepal

    Copyright © Boleslav Brezina
    Carabus (Limnocarabus) clathratus clathratus Linne, 1761
    Carabus (Limnocarabus) clathratus clathratus Linne, 1761
    Habitat
    Carabus (Limnocarabus) clathratus foveolatoseriatus
    The Chelyabinsk area, Kizilskyi township, Cheka mountain (steppe). 14-15/07/2008
    photo E. Chibilov
    Copyright © Sunvas
    Carabus (Meganebrius) arunensis Heinertz, 1980
    Carabus (Meganebrius) arunensis Heinertz, 1980
    Habitat
    Nepal
    Copyright © Boleslav Brezina
    Carabus (Archiplectes) edithae Reitter, 1893
    Carabus (Archiplectes) edithae Reitter, 1893
    Habitat
    Carabus (Archiplectes) edithae exedithae Gottwald, 1985
    Karachaevo-Cherkessia, valley of Daut riv. 43.62' 42.09'
    17-18.V.2009
    Copyright © Alexander Anichtchenko
    Carabus (Aulonocarabus) canaliculatus Adams, 1812
    Carabus (Aulonocarabus) canaliculatus Adams, 1812
    Habitat
    China, Liaoning pro., Dandong, Wulong mountain, 2009.7.31, Liu ye leg.;
    good at climbing up the trees for food
    Copyright © Ye Liu
    Carabus (Aulonocarabus) canaliculatus Adams, 1812
    Carabus (Aulonocarabus) canaliculatus Adams, 1812
    Habitat
    C Ural
    Copyright © Anton Menstschikov
    Carabus (Aulonocarabus) truncaticollis Eschscholtz, 1833
    Carabus (Aulonocarabus) truncaticollis Eschscholtz, 1833
    Habitat
    N Ural
    Copyright © Anton Menstschikov
    Carabus (Apotomopterus) yuae Deuve, 1989
    Carabus (Apotomopterus) yuae Deuve, 1989
    Habitat
    China, Guizhou pro, Leigongshan mountain, 2008.7.3, 1600m, Liu ye leg.;
    photo & det. by liuye
    Copyright © Ye Liu
    Carabus (Acoptolabrus) schrencki Motschulsky, 1860
    Carabus (Acoptolabrus) schrencki Motschulsky, 1860
    Habitat
    China, Heilongjiang pro., Hutou town, Ussuri region, 2009.5.23, 50m, Liu ye leg.;
    Copyright © Ye Liu
    Carabus (Megodontus) croaticus durmitorensis Apfelbeck, 1904
    Carabus (Megodontus) croaticus durmitorensis Apfelbeck, 1904
    Habitat
    Shar Mts., Macedonia
    Copyright © Alexander Anichtchenko
    Carabus (Mesocarabus) dufouri dufouri Dejean, 1829
    Carabus (Mesocarabus) dufouri dufouri Dejean, 1829
    Endophallus
    Jaen reg., Monte Javalcuz h=1436 m
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) dufouri dufouri Dejean, 1829
    Carabus (Mesocarabus) dufouri dufouri Dejean, 1829
    Male adult
    SC Spain, Jaen reg., Monte Javalcuz h=1436 m, 6.X.2010, M.A. Lopez leg.
    L=27 mm
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) problematicus harcyniae (as procedens Csiki, 1927)
    Carabus (Mesocarabus) problematicus harcyniae (as procedens Csiki, 1927)
    Male adult
    England, North Yorkshire, Fylingdales Moor, Grouse Hill, VII.2006 L=24 mm
    Copyright © Ruslan Panin
    Carabus (Pachystus) hungaricus scythus Motschulsky, 1847
    Carabus (Pachystus) hungaricus scythus Motschulsky, 1847
    Female adult
    S Ukraine, Zaporizhzhya reg., Khortytsa island, 16-20.VII.2011, M. Mulenko leg., det. Panin R.
    L=28 mm
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) lusitanicus brevis (as sorianus Jeanne, 1972)
    Carabus (Mesocarabus) lusitanicus brevis (as sorianus Jeanne, 1972)
    Endophallus
    NC-Spain, Soria, Puerto de Oncala
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) lusitanicus brevis (as sorianus Jeanne, 1972)
    Carabus (Mesocarabus) lusitanicus brevis (as sorianus Jeanne, 1972)
    Male adult
    NC-Spain, Soria, Puerto de Oncala, 2.VI.2006
    L=24 mm
    Copyright © Ruslan Panin
    Carabus (Coptolabrus) principalis Bates, 1889
    Carabus (Coptolabrus) principalis Bates, 1889
    Male+Female adult
    Zi-Ding-Shan, h=1406 m, Guiyang County, Binzhou City, Hunan, CHINE
    19-29.IV.2008
    Copyright © Alexandr Safronov
    Carabus (Coptolabrus) principalis Bates, 1889
    Carabus (Coptolabrus) principalis Bates, 1889
    Male adult
    C.Chine, W Hubei, Changyang, VIII.2005,
    Det. I.A.Solodovnikov 2006
    Copyright © Igor Solodovnikov
    Carabus (Coptolabrus) principalis Bates, 1889
    Carabus (Coptolabrus) principalis Bates, 1889
    Male adult
    C.Chine, W Hubei, Changyang, 08.2005,
    Det. I.A.Solodovnikov 2006
    Copyright © Igor Solodovnikov
    Carabus (Coptolabrus) principalis Bates, 1889
    Carabus (Coptolabrus) principalis Bates, 1889
    Female adult
    C.Chine, W Hubei, Changyang, 08.2005,
    Det. I.A.Solodovnikov 2006
    Copyright © Igor Solodovnikov
    Carabus (Coptolabrus) principalis Bates, 1889
    Carabus (Coptolabrus) principalis Bates, 1889
    Female adult
    C Chine, W Hubei, Changyang, 08.2005,
    Det. I.A.Solodovnikov 2006
    Copyright © Igor Solodovnikov
    Carabus (Carabus) arvensis arvensis Herbst, 1784
    Carabus (Carabus) arvensis arvensis Herbst, 1784
    Endophallus
    W Ukraine, NW Podolian, Lviv reg., Javoriv distr.,near Stradch vill.,
    Copyright © Ruslan Panin
    Carabus (Coptolabrus) nankotaizanus Kano, 1932
    Carabus (Coptolabrus) nankotaizanus Kano, 1932
    Male adult
    Taiwan,
    Tatumi, h=200 m, 20.02.1971, D.Muting leg.
    Det. I.A.Solodovniov 2006
    Copyright © Igor Solodovnikov
    Carabus (Coptolabrus) nankotaizanus Kano, 1932
    Carabus (Coptolabrus) nankotaizanus Kano, 1932
    Male adult
    Taiwan,
    Tatumi, h=200 m, 20.II.1971, D.Muting leg.
    Det. I.A.Solodovniov 2006
    Copyright © Igor Solodovnikov
    Carabus (Carabus) stscheglowi Mannerheim, 1827
    Carabus (Carabus) stscheglowi Mannerheim, 1827
    Female adult

    Copyright © Kirill Makarov
    Carabus (Carabus) stscheglowi Mannerheim, 1827
    Carabus (Carabus) stscheglowi Mannerheim, 1827
    Male adult

    Copyright © Kirill Makarov
    Carabus (Coptolabrus) lafossei tungchengensis Li, 1993
    Carabus (Coptolabrus) lafossei tungchengensis Li, 1993
    Male+Female adult
    Tiantou, Yuexi County,
    Anhui Province, CHINE
    2-8.VI.2008
    male 45mm, female 50 mm
    Copyright © Alexandr Safronov
    Carabus (Coptolabrus) lafossei tungchengensis Li, 1993 (as dabieshanus Imura, 1996)
    Carabus (Coptolabrus) lafossei tungchengensis Li, 1993 (as dabieshanus Imura, 1996)
    Male adult
    Chine, Hubei, Dabieshan: Yuexi,h=700m., VI.2004, Local people
    Det. I.A.Solodovniov 2006
    Copyright © Igor Solodovnikov
    Carabus (Coptolabrus) lafossei tungchengensis Li, 1993 (as dabieshanus Imura, 1996)
    Carabus (Coptolabrus) lafossei tungchengensis Li, 1993 (as dabieshanus Imura, 1996)
    Male adult
    Chine, Hubei, Dabieshan: Yuexi,h=700m., VI.2004, Local people
    Det. I.A.Solodovniov 2006
    Copyright © Igor Solodovnikov
    Carabus (Coptolabrus) lafossei tiantai Kleinfeld, 1997
    Carabus (Coptolabrus) lafossei tiantai Kleinfeld, 1997 Holotype
    Female adult

    Copyright © Frank Kleinfeld
    Carabus (Coptolabrus) lafossei saturatus Hauser, 1913
    Carabus (Coptolabrus) lafossei saturatus Hauser, 1913
    Female adult
    N Fujian, Jung Ling Shan, 18-24.7.96
    female 53 mm
    Copyright © Alexander Anichtchenko
    Carabus (Coptolabrus) lafossei pseudocoelestis Kleinfeld, 1999
    Carabus (Coptolabrus) lafossei pseudocoelestis Kleinfeld, 1999 Holotype
    Male adult

    Copyright © Frank Kleinfeld
    Carabus (Coptolabrus) lafossei montigradus Hauser, 1920
    Carabus (Coptolabrus) lafossei montigradus Hauser, 1920
    Female adult
    Chine, Anhui, Huangshan, 07.2005 h=700m Local people
    Det. I.A.Solodovnikov 2006
    Copyright © Igor Solodovnikov
    Carabus (Coptolabrus) lafossei montigradus Hauser, 1920
    Carabus (Coptolabrus) lafossei montigradus Hauser, 1920
    Female adult
    Chine, Anhui, Huangshan, 07.2005 h=700m Local people
    Det. I.A.Solodovnikov 2006
    Copyright © Igor Solodovnikov
    Carabus (Coptolabrus) lafossei montigradus Hauser, 1920
    Carabus (Coptolabrus) lafossei montigradus Hauser, 1920
    Male adult
    Chine, Anhui, Huangshan, 07.2005 h=700m Local people
    Det. I.A.Solodovnikov 2006
    Copyright © Igor Solodovnikov
    Carabus (Coptolabrus) lafossei montigradus Hauser, 1920
    Carabus (Coptolabrus) lafossei montigradus Hauser, 1920
    Male adult
    Chine, Anhui, Huangshan, 07.2005 h=700m Local people
    Det. I.A.Solodovnikov 2006
    Copyright © Igor Solodovnikov
    Carabus (Coptolabrus) lafossei lafossei Feisthamel, 1845 (as donckieri Born,1898)
    Carabus (Coptolabrus) lafossei lafossei Feisthamel, 1845 (as donckieri Born,1898)
    Male adult
    Chine, Zhejiang: Zhousan Archipelago
    Copyright © Igor Solodovnikov
    Carabus (Coptolabrus) lafossei lafossei Feisthamel, 1845 (as donckieri Born,1898)
    Carabus (Coptolabrus) lafossei lafossei Feisthamel, 1845 (as donckieri Born,1898)
    Male adult
    Chine, Zhejiang: Zhousan Archipelago
    Copyright © Igor Solodovnikov
    Carabus (Coptolabrus) lafossei buchi Hauser, 1913
    Carabus (Coptolabrus) lafossei buchi Hauser, 1913
    Male+Female adult
    Yushanwu, Anji County,
    Zhejiang Province, CHINE
    16-29.IV.2008
    Copyright © Alexandr Safronov
    Carabus (Coptolabrus) lafossei lafossei Feisthamel, 1845
    Carabus (Coptolabrus) lafossei lafossei Feisthamel, 1845
    Male adult
    China, Zhejiang pro., Liang H.B. coll;
    Copyright © Ye Liu
    Carabus (Coptolabrus) jankowskii jankowskii Oberthur, 1883
    Carabus (Coptolabrus) jankowskii jankowskii Oberthur, 1883
    Male adult
    N.Korea, Chagang Do, Mt. Langlin, 30.07.2005
    Copyright © Anton Menstschikov
    Carabus (Coptolabrus) jankowskii benxiensis Imura, 1991
    Carabus (Coptolabrus) jankowskii benxiensis Imura, 1991
    Female adult
    CHINA, Jilin prov., Mt. Longgang., 15.07.2005
    Copyright © Anton Menstschikov
    Carabus (Coptolabrus) jankowskii benxiensis Imura, 1991
    Carabus (Coptolabrus) jankowskii benxiensis Imura, 1991
    Male+Female adult
    Miaoyang, Shicheng, Fengcheng City,
    Liaoning Province, CHINE
    25-30.VI.2008
    Copyright © Alexandr Safronov
    Carabus (Coptolabrus) jankowskii benxiensis Imura, 1991
    Carabus (Coptolabrus) jankowskii benxiensis Imura, 1991
    Male adult
    NE-China,
    Liaoning Prov., Donggou, Mt. Guanmen, 15.VI.2009
    male, 36mm
    Copyright © Ruslan Panin
    Carabus (Coptolabrus) ishizukai Deuve & Ohshima, 1989
    Carabus (Coptolabrus) ishizukai Deuve & Ohshima, 1989
    Male adult
    Chine, E.Sichuan, Wushan 07,2003 h=800m, Local people
    Det.I.A.Solodovnikov 2006
    Copyright © Igor Solodovnikov
    Carabus (Coptolabrus) ishizukai Deuve & Ohshima, 1989
    Carabus (Coptolabrus) ishizukai Deuve & Ohshima, 1989
    Male adult
    Chine, E.Sichuan, Wushan 07,2003 h=800m, Local people
    Det.I.A.Solodovnikov 2006
    Copyright © Igor Solodovnikov
    Carabus (Coptolabrus) augustus ignigena Hauser, 1914
    Carabus (Coptolabrus) augustus ignigena Hauser, 1914
    Female adult
    Chine, CE Hunan, Changde 07.2004 h=700m Local people
    Det.I.A.Solodovnikov 2006
    Copyright © Igor Solodovnikov
    Carabus (Coptolabrus) augustus ignigena Hauser, 1914
    Carabus (Coptolabrus) augustus ignigena Hauser, 1914
    Male adult
    Chine, CE Hunan, Changde 07.2004 h=700m Local people<br /> Det.I.A.Solodovnikov 2006
    Copyright © Igor Solodovnikov
    Carabus (Coptolabrus) formosus sunpanensis Semenov, 1898
    Carabus (Coptolabrus) formosus sunpanensis Semenov, 1898
    Male adult

    Copyright © Alexander Anichtchenko
    Carabus (Coptolabrus) formosus grumorum Semenov, 1898
    Carabus (Coptolabrus) formosus grumorum Semenov, 1898
    Female adult
    female, 30mm
    Copyright © Boleslav Brezina
    Carabus (Coptolabrus) formosus grumorum Semenov, 1898
    Carabus (Coptolabrus) formosus grumorum Semenov, 1898
    Male adult
    male, 24mm
    Copyright © Boleslav Brezina
    Carabus (Coptolabrus) formosus brockmanni Kleinfeld & Schutze, 1997
    Carabus (Coptolabrus) formosus brockmanni Kleinfeld & Schutze, 1997 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Sphodristocarabus) armeniacus (as laevilineatus)
    Carabus (Sphodristocarabus) armeniacus (as laevilineatus)
    Female adult

    Copyright © Kirill Makarov
    Carabus (Coptolabrus) formosus bousqueti Deuve, 1993
    Carabus (Coptolabrus) formosus bousqueti Deuve, 1993
    Male adult
    Wenxian, N.Sichuan, CHINA
    21.07.2006
    Copyright © Alexandr Safronov
    Carabus (Coptolabrus) formosus sunpanensis Semenov, 1898 (as rostandi Deuve, 1991)
    Carabus (Coptolabrus) formosus sunpanensis Semenov, 1898 (as rostandi Deuve, 1991)
    Male adult
    China, N Sichuan province, Venchuan, h=1400 m., 30.VI.2007, A. Gorodinski leg.;
    L=33 mm
    Copyright © Anton Menstschikov
    Carabus (Coptolabrus) elysii pulcher Kleinfeld, 1997
    Carabus (Coptolabrus) elysii pulcher Kleinfeld, 1997 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Coptolabrus) elysii magnificens Kleinfeld, 1999
    Carabus (Coptolabrus) elysii magnificens Kleinfeld, 1999 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Coptolabrus) elysii elysii Thomson, 1856 (as idai Kleinfeld, 2000)
    Carabus (Coptolabrus) elysii elysii Thomson, 1856 (as idai Kleinfeld, 2000) Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Coptolabrus) elysii elysii Thomson, 1856
    Carabus (Coptolabrus) elysii elysii Thomson, 1856
    Male+Female adult
    S. Shennongjia, Fangxian county, Hubei, CHINE
    1-11.06.2007
    Copyright © Alexandr Safronov
    Carabus (Coptolabrus) elysii elysii Thomson, 1856
    Carabus (Coptolabrus) elysii elysii Thomson, 1856
    Female adult
    China, Sichuan province, Wanyuan; 15.06.2009; Local people leg.;
    L=32 mm.
    Copyright © Anton Menstschikov
    Carabus (Coptolabrus) augustus pustuliferus Kleinfeld, 1998
    Carabus (Coptolabrus) augustus pustuliferus Kleinfeld, 1998 Holotype
    Male adult

    Copyright © Frank Kleinfeld
    Carabus (Coptolabrus) augustus cristianofonti Deuve & Font, 1998
    Carabus (Coptolabrus) augustus cristianofonti Deuve & Font, 1998
    Female adult
    China, Guizhou Prov., Sinan, 14.V.2009. L= 41mm
    Copyright © Ruslan Panin
    Carabus (Coptolabrus) augustus chalingxian Kleinfeld, 1998
    Carabus (Coptolabrus) augustus chalingxian Kleinfeld, 1998 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Coptolabrus) augustus augustus Bates, 1888
    Carabus (Coptolabrus) augustus augustus Bates, 1888
    Male+Female adult
    Male: Mt. Wugong, Pingxiang, Jiangxi, CHINA. 06.06.2007
    Female: Mt. Jinngang, Jiangxi, CHINA. 26.06.2007
    Copyright © Alexandr Safronov
    Carabus (Coptolabrus) augustus adriaenssensi (as pseudoignigena Kleinfeld, 1996)
    Carabus (Coptolabrus) augustus adriaenssensi (as pseudoignigena Kleinfeld, 1996) Holotype
    Female adult

    Copyright © Frank Kleinfeld
    Carabus (Coptolabrus) augustus yangminshanicus Font, 1997
    Carabus (Coptolabrus) augustus yangminshanicus Font, 1997
    Male adult
    Guangxi prov., Gongcheng county, N. Lianhua, CHINA<br /> 9-17.06.2007, L (mail) = 40 mm, L (female) = 45 mm.
    Copyright ©
    Carabus (Coptolabrus) augustus yangminshanicus Font, 1997
    Carabus (Coptolabrus) augustus yangminshanicus Font, 1997
    Female adult
    Mt. Yangmingshan, Yongzhou City, Hunan, CHINA<br /> 20-28.06.2007
    Copyright ©
    Carabus (Tribax) puschkini ishikawaianus Breuning & Ruspoli, 1970
    Carabus (Tribax) puschkini ishikawaianus Breuning & Ruspoli, 1970
    Endophallus
    NE-Turkey: Artvin:Hopa, Borзka env., h=450 m,
    Copyright © Ruslan Panin
    Carabus (Lipaster) stjernvalli osellai Breuning & Ruspoli, 1970
    Carabus (Lipaster) stjernvalli osellai Breuning & Ruspoli, 1970
    Male+Female adult
    NE-Turkey: Artvin:Hopa, Borçka env., h=750 m, 9.VI-7.VII.2011, M.Janalik leg.
    female, 42 mm
    Copyright © Ruslan Panin
    Carabus (Tribax) puschkini ishikawaianus Breuning & Ruspoli, 1970
    Carabus (Tribax) puschkini ishikawaianus Breuning & Ruspoli, 1970
    Male adult
    NE-Turkey: Artvin:Hopa, Borçka env., h=450 m, 29.VI-7.VII.2011, M.Janalik leg.
    L=33 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) praecellens praecellens Pillardi, 1825
    Carabus (Morphocarabus) praecellens praecellens Pillardi, 1825
    Endophallus
    Serbia, Fruska Gora Mts, 20-31.V.2006, M.Sima leg.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) praecellens praecellens Pillardi, 1825
    Carabus (Morphocarabus) praecellens praecellens Pillardi, 1825
    Male adult
    Serbia, Fruska Gora Mts, 20-31.V.2006, M.Sima leg.
    L=29 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) illigeri versicolor Frivaldsky, 1835
    Carabus (Morphocarabus) illigeri versicolor Frivaldsky, 1835
    Male+Female adult
    C Bulgaria:Stara planina Mts, Gabrovo,12-15.VI.2007, M. Janalik leg. female, 31 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) illigeri versicolor Frivaldsky, 1835
    Carabus (Morphocarabus) illigeri versicolor Frivaldsky, 1835
    Endophallus
    C Bulgaria: Stara planina Mts, Gabrovo
    Copyright © Ruslan Panin
    Carabus (Tribax) puschkini ponticus Deyrolle, 1869
    Carabus (Tribax) puschkini ponticus Deyrolle, 1869
    Endophallus
    NE-Turkey:Trabzon, Zigana Pass h=1700m,
    Copyright © Ruslan Panin
    Carabus (Tribax) puschkini ponticus Deyrolle, 1869
    Carabus (Tribax) puschkini ponticus Deyrolle, 1869
    Male adult
    NE-Turkey:Trabzon, Zigana Pass h=1700m,14-24.VI.2010, M.Janalik leg.
    L=28 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi comptus (as szoerenyensis Csiki, 1908)
    Carabus (Morphocarabus) rothi comptus (as szoerenyensis Csiki, 1908)
    Endophallus
    SW Romania, Tarcu Mts, vf. Nedeia h=2000m,
    Copyright © Ruslan Panin
    Carabus (Megodontus) bonvouloiri bonvouloiri (as kaschkarensis Blumenthal & Breuning, 1967)
    Carabus (Megodontus) bonvouloiri bonvouloiri (as kaschkarensis Blumenthal & Breuning, 1967)
    Endophallus
    NE-Turke,Rize vill., S Ayder h=1740 m,
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi comptus (as szoerenyensis Csiki, 1908)
    Carabus (Morphocarabus) rothi comptus (as szoerenyensis Csiki, 1908)
    Male adult
    SW Romania, Tarcu Mts, vf. Nedeia h=2000m, 12.VI.2008, J.Kynkor leg.
    L=22 mm
    Copyright © Ruslan Panin
    Carabus (Carabus) arvensis carpathus (as pseudonoricus Kenyery, 1983)
    Carabus (Carabus) arvensis carpathus (as pseudonoricus Kenyery, 1983)
    Male adult
    Slovenia: Julske Alpy, Podvelka, h=450m, 24.X.2010, M.Janalik leg.
    L=19 mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) bonvouloiri bonvouloiri (as kaschkarensis Blumenthal & Breuning, 1967)
    Carabus (Megodontus) bonvouloiri bonvouloiri (as kaschkarensis Blumenthal & Breuning, 1967)
    Male adult
    NE-Turke,Rize vill., S Ayder h=1740 m,28.VI-6.VII.2011, M.Janalik leg.
    L=32 mm
    Copyright © Ruslan Panin
    Carabus (Acoptolabrus) schrencki schrenck (as dongningensis Deuve & Li, 2005)
    Carabus (Acoptolabrus) schrencki schrenck (as dongningensis Deuve & Li, 2005)
    Male adult
    NE China, Heilongjiang prov., Dongning, V.2009<br /> L=25 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi hampei (as gutiiensis Takács & Lie, 1992)
    Carabus (Morphocarabus) rothi hampei (as gutiiensis Takács & Lie, 1992)
    Male adult
    N Romania, E Carpathian, Maramureş Reg., Gutâi Mts, cabana Mogosa,4 km N Baia Sprie, 9-17.VI.2010, J.Kynkor leg.
    L=33 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi hampei (as gutiiensis Takács & Lie, 1992)
    Carabus (Morphocarabus) rothi hampei (as gutiiensis Takács & Lie, 1992)
    Endophallus
    N Romania, E Carpathian, Maramureş Reg., Gutai Mts, cabana Mogosa, 4 km N Baia Sprie,
    Copyright © Ruslan Panin
    Carabus (Eucarabus) obsoletus nagyagensis (as paranagyagensis Lie, 1990)
    Carabus (Eucarabus) obsoletus nagyagensis (as paranagyagensis Lie, 1990)
    Male adult
    W Romania, Arad reg., Zarand Mts (Muntii Zarandului), Nadas-Barzavah h=142 m, J. Kynkor leg.
    L=24 mm
    Copyright © Ruslan Panin
    Carabus (Sphodristocarabus) macrogonus macrogonus Chaudoir, 1847
    Carabus (Sphodristocarabus) macrogonus macrogonus Chaudoir, 1847
    Male adult
    NE-Turkey:Trabzon env., Sumela, 17-21.VI.2010, M.Janalik leg.
    L=20 mm
    Copyright © Ruslan Panin
    Carabus (Sphodristocarabus) macrogonus macrogonus Chaudoir, 1847
    Carabus (Sphodristocarabus) macrogonus macrogonus Chaudoir, 1847
    Endophallus
    NE-Turkey:Trabzon env., Sumela,
    Copyright © Ruslan Panin
    Carabus (Sphodristocarabus) macrogonus pseudokolatensis Heinz & Korge, 1967
    Carabus (Sphodristocarabus) macrogonus pseudokolatensis Heinz & Korge, 1967
    Male adult
    NE-Turkey::Kackar Dagi Mt, Ayder S of Ardesen h=1600m, 17-21.VI.2010, M.Janalik leg.
    L=20 mm
    Copyright © Ruslan Panin
    Carabus (Sphodristocarabus) macrogonus pseudokolatensis Heinz & Korge, 1967
    Carabus (Sphodristocarabus) macrogonus pseudokolatensis Heinz & Korge, 1967
    Endophallus
    NE-Turkey::Kackar Dagi Mt, Ayder S of Ardesen h=1600m
    Copyright © Ruslan Panin
    Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931
    Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931
    Male adult
    NE-China:Liaoning prov., Fengcheng City, Miaoyang, 12-17.VI.2011
    L=26 mm
    Copyright © Ruslan Panin
    Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931
    Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931
    Endophallus
    NE-China:Liaoning prov.,Fengcheng City, Miaoyang,
    Copyright © Ruslan Panin
    Carabus (Scambocarabus) kruberi laobeiensis Deuve & Li, 2000
    Carabus (Scambocarabus) kruberi laobeiensis Deuve & Li, 2000
    Male adult
    NE-China: Liaoning prov., Fengcheng City, Miaoyang, 12-17.VI.2011
    L=27 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) wulffiusi dekraatzi Kraatz, 1881
    Carabus (Morphocarabus) wulffiusi dekraatzi Kraatz, 1881
    Male adult
    NE-China, Heilongjiang, Wuchang city, Mt Zheng-nan-gou, 1-11.VI.2011
    L=22 mm
    Copyright © Ruslan Panin
    Carabus (Platycarabus) fabricii fabricii (as feichtauensis Müller & Sach, 1999)
    Carabus (Platycarabus) fabricii fabricii (as feichtauensis Müller & Sach, 1999)
    Male adult
    NC Austria, E-Alps, Windischgarsten, Mt Hoher Nock, VII.2004, Manfred Egger leg.
    L=18 mm
    Copyright © Ruslan Panin
    Carabus (Platycarabus) fabricii fabricii (as koralpicus Sokolar, 1910)
    Carabus (Platycarabus) fabricii fabricii (as koralpicus Sokolar, 1910)
    Female adult
    S Austria, Carinthia reg.(Kärnten), Wolfsberg, Koralpe Mt Range,18.VI.2005, Manfred Egger leg.
    L=22 mm
    Copyright © Ruslan Panin
    Carabus (Procrustes) anatolicus anatolicus Chaudoir, 1857
    Carabus (Procrustes) anatolicus anatolicus Chaudoir, 1857
    Male adult
    Cyprus, Troodos, 17.2.1V.2010,. Manfred Egger leg.
    L=27 mm
    Copyright © Ruslan Panin
    Carabus (Tomocarabus) convexus moreanus Reitter, 1896
    Carabus (Tomocarabus) convexus moreanus Reitter, 1896
    Female adult
    Greece:, N Peloponnes, Xylokastro, Zemeno,h=1000m, 6.VI.1997, Manfred Egger leg.
    L=21 mm
    Copyright © Ruslan Panin
    Carabus (Procrustes) banoni Dejean, 1829
    Carabus (Procrustes) banoni Dejean, 1829
    Male adult
    Greece,W Crete Island, Mili gorge ,12.V.2009, Manfred Egger leg.
    L=26 mm
    Copyright © Ruslan Panin
    Carabus (Macrothorax) rugosus rugosus Fabricius, 1792
    Carabus (Macrothorax) rugosus rugosus Fabricius, 1792
    Male adult
    N Morocco, El Biutz –Taghramt,20.I.2005, Manfred Egger leg.
    L=29 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) alpestris hoppei Germar, 1824
    Carabus (Orinocarabus) alpestris hoppei Germar, 1824
    Male adult
    W Austria, Tirol, Innsbruck, Hafelekar Mts h=2200m, Manfred Egger leg.
    L=19 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) alpestris alpestris Sturm, 1815
    Carabus (Orinocarabus) alpestris alpestris Sturm, 1815
    Female adult
    C Austria, E-Alps, Windischgarsten, Mt Hoher Nock, VII.2004, Manfred Egger leg.
    L=20 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) linnaei linnaei Panzer, 1812
    Carabus (Orinocarabus) linnaei linnaei Panzer, 1812
    Male adult
    W-Ukraine, NW podolian,
    Lviv Reg., Zhydachiv distr., near Vybranivka, 17.II.2007,Panin R. leg.
    female, 21 mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) croaticus croaticus Dejean, 1826
    Carabus (Megodontus) croaticus croaticus Dejean, 1826
    Male adult
    SW Slovenia, Ilirska Bistrica, Sviščaki, Snežnik,17-28.VI.2009, Manfred Egger leg.
    L=32 mm
    Copyright © Ruslan Panin
    Carabus (Macrothorax) morbillosus constantinus Kraatz, 1899
    Carabus (Macrothorax) morbillosus constantinus Kraatz, 1899
    Male adult
    NW Tunisia; Tebarka, E Sejenane,26.I.2004, Manfred Egger leg.
    L=33 mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) caelatus schreiberi Kraatz, 1887
    Carabus (Megodontus) caelatus schreiberi Kraatz, 1887
    Male adult
    SW Slovenia, Obrov, Poljane, 14-27.VIII.2010, Manfed Egger leg.
    L=33 mm
    Copyright © Ruslan Panin
    Carabus (Oreocarabus) carinthiacus Sturm, 1815
    Carabus (Oreocarabus) carinthiacus Sturm, 1815
    Male adult
    NE-Italy, Bolzano, Brenner, Zirog h=2000 m, 26.VII.1992, Manfred Egger leg.
    L=20 mm
    Copyright © Ruslan Panin
    Carabus (Damaster) blaptoides oxuroides Schaum, 1862
    Carabus (Damaster) blaptoides oxuroides Schaum, 1862
    Female adult
    Japan, C-Honshu, Ofuse-cho, Kamitakai, Nagano Pref., 12.VI.2006, Ohshima leg.
    L=34 mm
    Copyright © Ruslan Panin
    Carabus (Platycarabus) fabricii fabricii (as poloniensis Géhin, 1876)
    Carabus (Platycarabus) fabricii fabricii (as poloniensis Géhin, 1876)
    Female adult
    N-Slovakia, Orava Beskids Mts, Mt Babia Góra h=1720 m, 5.VII.20.VIII.2010, Rudolf Kmeco leg.
    L=22 mm
    Copyright © Ruslan Panin
    Carabus (Chaetocarabus) arcadicus merlini Schaum, 1861
    Carabus (Chaetocarabus) arcadicus merlini Schaum, 1861
    Male adult
    S-Greece:Peloponnesos: Taygetos Mt.Rng., 17-23.V.2005, V. Sima leg.
    L=30 mm
    Copyright © Ruslan Panin
    Carabus (Tomocarabus) convexus dilatatus (as perplexus Schaum, 1857)
    Carabus (Tomocarabus) convexus dilatatus (as perplexus Schaum, 1857)
    Male adult
    CE-Greece: Larissa reg., Stomio, Ossa Mt., 6-17.VII.2011, Krause Rene leg.
    L=19 mm
    Copyright © Ruslan Panin
    Carabus (Procrustes) coriaceus mopsucrenae Peyron, 1858
    Carabus (Procrustes) coriaceus mopsucrenae Peyron, 1858
    Male adult
    SC-Turkey, Adana Prov., Doganbeyli, 8.V.2001, Snizhek M. leg.
    L=28 mm
    Copyright © Ruslan Panin
    Carabus (Eucarabus) ulrichi rhilensis (as stussineri Géhin, 1885)
    Carabus (Eucarabus) ulrichi rhilensis (as stussineri Géhin, 1885)
    Male adult
    W Croatia, Ivanščica Mts , Ivanec, 24.VI-7.VII.2006, Krause Rene leg.
    L=29 mm
    Copyright © Ruslan Panin
    Carabus (Eucarabus) ulrichi ulrichi (as intercessor Sokolar, 1912)
    Carabus (Eucarabus) ulrichi ulrichi (as intercessor Sokolar, 1912)
    Male adult
    N Hungary, Pilis Mts., Piliszentlelek, 20-27.VII.2011, Krause Rene leg.
    L=28 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi incompsus Kraatz, 1880
    Carabus (Morphocarabus) rothi incompsus Kraatz, 1880
    Endophallus
    C Romania, Brașov County, Cristian vill., h=380 m
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi incompsus Kraatz, 1880
    Carabus (Morphocarabus) rothi incompsus Kraatz, 1880
    Male+Female adult
    C Romania, Brașov County, Cristian vill., h=380 m,V-VI.2008, M.Pasek leg.
    female, 27 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi (? incompsus as spectabilis Csiki, 1906)
    Carabus (Morphocarabus) rothi (? incompsus as spectabilis Csiki, 1906)
    Female adult
    NC Romania, Bistrita Nasaud reg., Călimani Mts, Colibita vill., mixed forest, meadows h=800m, 15-21.V.2011, M.Pasek leg.<br /> L=34 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi incompsus (as mendax Csiki, 1906)
    Carabus (Morphocarabus) rothi incompsus (as mendax Csiki, 1906)
    Female adult
    W Ukraine, NE Carpathians,
    Transcarpathian reg., Chornogora Mt Range, h=1100m,13.VI.2011, Panin R. leg./det.
    L=33 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi alutensis Savulescu, 1972
    Carabus (Morphocarabus) rothi alutensis Savulescu, 1972
    Endophallus
    C Romania, S Carpathians, Rimnicu Vilcea env., Calimaneşti, Cozia Mts.,
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi alutensis Savulescu, 1972
    Carabus (Morphocarabus) rothi alutensis Savulescu, 1972
    Male+Female adult
    C Romania, S Carpathians, Rimnicu Vilcea env., Calimaneşti, Cozia Mts., VII.2007, I. Marvan leg.
    L=29 mm
    Copyright © Ruslan Panin
    Carabus (Carabus) menetriesi menetriesi s.str.
    Carabus (Carabus) menetriesi menetriesi s.str.
    Habitat
    Site Zalyvki, 20.IV.2012
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi comptus (as borlovensis Lie, 1994)
    Carabus (Morphocarabus) rothi comptus (as borlovensis Lie, 1994)
    Male adult
    SW Romania, S Carpathians,
    Banat , Tarcu Mts, Borlova env., 19-31.V.2005, P. Sima leg.
    L=24 мм
    Copyright © Ruslan Panin
    Carabus (Lamprostus) robustus ducalis Schaufuss, 1882
    Carabus (Lamprostus) robustus ducalis Schaufuss, 1882
    Male adult
    NE-Turkey:Trabzon, Zigana Pass h=2000m,14-24.VI.2010, V. Slovák leg.
    L=32 mm
    Copyright © Ruslan Panin
    Carabus (Lamprostus) robustus ducalis Schaufuss, 1882
    Carabus (Lamprostus) robustus ducalis Schaufuss, 1882
    Endophallus
    NE-Turkey:Trabzon, Zigana Pass h=2000m
    Copyright © Ruslan Panin
    Carabus (Eucarabus) ulrichi arrogans Schaum, 1858
    Carabus (Eucarabus) ulrichi arrogans Schaum, 1858
    Male adult
    NE-Serbia:Majdanpek, 9-11,V.2010, L.Brhel leg.
    L=28 mm
    Copyright © Ruslan Panin
    Carabus (Diocarabus) fraterculus fraterculus Reitter, 1895
    Carabus (Diocarabus) fraterculus fraterculus Reitter, 1895
    Male adult
    N-Korea, N Pyongan Prov.,Sakju, Mt Ponsan, 20-29.VI.2009, L. Bieber leg.
    L=19 mm
    Copyright © Ruslan Panin
    Carabus (Diocarabus) fraterculus fraterculus Reitter, 1895
    Carabus (Diocarabus) fraterculus fraterculus Reitter, 1895
    Endophallus
    N-Korea, N Pyongan Prov.,Sakju, Mt Ponsan,
    Copyright © Ruslan Panin
    Carabus (Sphodristocarabus) varians armeniacus Mannerheim, 1830
    Carabus (Sphodristocarabus) varians armeniacus Mannerheim, 1830
    Male adult
    NE Turkey, Artvin Prov., Şavşat, Çamlıca,18.VI.1993, M. Janata leg.<br /> L=22 mm
    Copyright © Ruslan Panin
    Carabus (Sphodristocarabus) varians armeniacus Mannerheim, 1830
    Carabus (Sphodristocarabus) varians armeniacus Mannerheim, 1830
    Endophallus
    NE Turkey, Artvin Prov., Şavşat, Зamlıca.
    Copyright © Ruslan Panin
    Carabus (Leptocarabus) hiurai Kamiyoshi & Mizoguchi, 1960
    Carabus (Leptocarabus) hiurai Kamiyoshi & Mizoguchi, 1960
    Male adult
    Collection ground Hojyo city, Ehime Pref. 34358
    (C) Working Group for Biological Indicator Ground Beetles Database, Japan
    Prohibited secondary use of all data
    Copyright © Katsuro Yahiro
    Carabus (Leptocarabus) kumagai kumagai Kimura & Komiya, 1974
    Carabus (Leptocarabus) kumagai kumagai Kimura & Komiya, 1974
    Male adult
    Collection ground Otsu city, Shiga Pref. 35330
    (C) Working Group for Biological Indicator Ground Beetles Database, Japan
    Prohibited secondary use of all data
    Copyright © Katsuro Yahiro
    Carabus (Coptolabrus) fruhstorferi Roeschke, 1900
    Carabus (Coptolabrus) fruhstorferi Roeschke, 1900
    Male adult
    Collection ground Izuhara cho, Nagasaki Pref. 35764
    (C) Working Group for Biological Indicator Ground Beetles Database, Japan
    Prohibited secondary use of all data
    Copyright © Katsuro Yahiro
    Carabus (Limnocarabus) clathratus maacki (as maacki aquatilis)
    Carabus (Limnocarabus) clathratus maacki (as maacki aquatilis)
    Male adult
    Collection ground Jusanko, Aomori Pref. 35305
    (C) Working Group for Biological Indicator Ground Beetles Database, Japan
    Prohibited secondary use of all data
    Copyright © Katsuro Yahiro
    Carabus (Pachystus) hungaricus hungaricus Fabricius, 1792
    Carabus (Pachystus) hungaricus hungaricus Fabricius, 1792
    Male adult
    Czech, S Moravia, Pouzdrany, 8,VIII.2005, V. Slovak lgt.
    L=27 mm
    Copyright © Ruslan Panin
    Carabus (Pachystus) hungaricus hungaricus Fabricius, 1792
    Carabus (Pachystus) hungaricus hungaricus Fabricius, 1792
    Endophallus
    Czech, S Moravia, Pouzdrany
    Copyright © Ruslan Panin
    Carabus (Eurycarabus) faminii romanoi Rapuzzi & Sparacio, 2012
    Carabus (Eurycarabus) faminii romanoi Rapuzzi & Sparacio, 2012 Holotype Paratype
    Male+Female adult
    Holotypus male, Godrano (PA), 25.XI.1978, legit I.Sparacio.<br /> photo from Ivan Rapuzzi & Ignazio Sparacio, 2012
    Copyright © - - -
    Carabus (Eurycarabus) famini famini Dejean, 1826
    Carabus (Eurycarabus) famini famini Dejean, 1826 Holotype
    Male adult
    (MNHN)
    photo from Ivan Rapuzzi & Ignazio Sparacio, 2012
    Copyright © ---
    Carabus (Isiocarabus) kiukiangensis alzonai Deuve, 1990
    Carabus (Isiocarabus) kiukiangensis alzonai Deuve, 1990 Holotype
    Male adult

    Copyright © Gianluigi Alzona
    Carabus (Isiocarabus) kiukiangensis alzonai Deuve, 1990 (aedeagus)
    Carabus (Isiocarabus) kiukiangensis alzonai Deuve, 1990 (aedeagus) Holotype
    Male adult

    Copyright © Gianluigi Alzona
    Carabus (Isiocarabus) kiukiangensis alzonai Deuve, 1990 (Label)
    Carabus (Isiocarabus) kiukiangensis alzonai Deuve, 1990 (Label) Holotype
    Male adult

    Copyright © Gianluigi Alzona
    Carabus (Megodontus) planicollis verae Csiki, 1905
    Carabus (Megodontus) planicollis verae Csiki, 1905
    Male+Female adult
    ♂ C-Romania:S Carpathians, Fagaraş Mts, near Avrig Lake, h=2100 m, 5.VI.2005
    ♀ C-Romania:S Carpathians, Fagaraş Mts, Saratii-Valley h=1700 m, 6.VI.2005
    female, 28 мм
    Copyright © Ruslan Panin
    Carabus (Lamprostus) torosus torosus Frivaldsky, 1835
    Carabus (Lamprostus) torosus torosus Frivaldsky, 1835
    Male adult
    NE-Bulgaria, Dobrich Region, Kavarna env.
    L=31 mm
    Copyright © Ruslan Panin
    Carabus (Lamprostus) torosus torosus Frivaldsky, 1835
    Carabus (Lamprostus) torosus torosus Frivaldsky, 1835
    Endophallus
    NE-Bulgaria, Dobrich Region, Kavarna env.,
    Copyright © Ruslan Panin
    Carabus (Eucarabus) obsoletus obsoletus Sturm, 1815 (as prunneri Mallasz, 1901)
    Carabus (Eucarabus) obsoletus obsoletus Sturm, 1815 (as prunneri Mallasz, 1901)
    Male adult
    NW Romania, Apuseni Mts, Trascău Mt Rng, Lungesti vill., 25 km W Turda,.VI.2012
    L=26 mm
    Copyright © Ruslan Panin
    Carabus (Semnocarabus) erosus kuldshaensis Ballion, 1878
    Carabus (Semnocarabus) erosus kuldshaensis Ballion, 1878
    Endophallus
    S-Kazakhstan:Ili River valley,:near Bakanas,
    Copyright © Ruslan Panin
    Carabus (Megodontus) stroganowi transfuga Semenov, 1898
    Carabus (Megodontus) stroganowi transfuga Semenov, 1898
    Endophallus
    SE-Azerbaijan: Talysh, Kalvaz,
    Copyright © Ruslan Panin
    Carabus (Deroplectes) dokhtouroffi dokhtouroffi Ganglbauer, 1886
    Carabus (Deroplectes) dokhtouroffi dokhtouroffi Ganglbauer, 1886
    Endophallus
    NC Tadzhikistan, Pamir, N-slope Peter I.Mt Rng, h=2100m, Ganishou vill.
    Copyright © Ruslan Panin
    Carabus (Deroplectes) dokhtouroffi dokhtouroffi Ganglbauer, 1886
    Carabus (Deroplectes) dokhtouroffi dokhtouroffi Ganglbauer, 1886
    Male adult
    NC Tadzhikistan, Pamir, N-slope Peter I.Mt Rng, h=2100m,Ganishou vill., 10-12.VI.2006, Ivanova E. leg.
    L=28 mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) stroganowi transfuga Semenov, 1898
    Carabus (Megodontus) stroganowi transfuga Semenov, 1898
    Male adult
    SE-Azerbaijan: Talysh, Kalvaz, 21-30.VI., Dolin V.G. leg.
    L=32 mm
    Copyright © Ruslan Panin
    Carabus (Semnocarabus) erosus kuldshaensis Ballion, 1878
    Carabus (Semnocarabus) erosus kuldshaensis Ballion, 1878
    Male adult
    S-Kazakhstan:Ili River valley,:near Bakanas,3-5.V.2002
    L=21 mm
    Copyright © Ruslan Panin
    Carabus (Procrustes) talyschensis talyschensis Menetries, 1832
    Carabus (Procrustes) talyschensis talyschensis Menetries, 1832
    Female adult
    Azerbaijan, Talysh Mts., Zuvand, Kalvaz vill., h=1650 m, 15.VII.2003, Dolin V.G. leg.
    L=32 mm
    Copyright © Ruslan Panin
    Carabus (Ulocarabus) ? theanus Reitter, 1895
    Carabus (Ulocarabus) ? theanus Reitter, 1895
    Male adult
    NC Tadzhikistan, Pamir, N-slope Peter I.Mt Rng, h=2100m, Ganishou vill., 10-12.VI.2006, Ivanova E. leg.
    L=23 mm
    Copyright © Ruslan Panin
    Carabus (Goniocarabus) grombczewskii grombczewskii Semenov, 1891
    Carabus (Goniocarabus) grombczewskii grombczewskii Semenov, 1891
    Male adult
    NC Tadzhikistan, Pamir, N-slope Peter I.Mt Rng, h=2100m, Ganishou vill.,12.VII.2006, Pak O. leg.
    L=19 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) viridilimbatus panini I.Solodovnikov 2017
    Carabus (Morphocarabus) viridilimbatus panini I.Solodovnikov 2017 Paratype
    Male adult
    Russia, SE Yakutia, Neryungri distr., Tokinski Stanovik Mt Rng., Beder riv., (right tributary rivers Algama), 26.VI.1996, Krasnov M. leg. L=15 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) viridilimbatus panini I.Solodovnikov 2017
    Carabus (Morphocarabus) viridilimbatus panini I.Solodovnikov 2017 Holotype
    Endophallus
    Russia, SE Yakutia, Neryungri distr., Tokinski Stanovik Mt Rng., Beder riv., (right tributary rivers Algama), 26.VI.1996
    Copyright © Ruslan Panin
    Carabus (Ulocarabus) stschurovskii Solsky, 1874
    Carabus (Ulocarabus) stschurovskii Solsky, 1874
    Endophallus
    Tadzhikistan, Pamir-Alay, S slope Gissar Mt.Range, Takob, h-1700m,
    Copyright © Ruslan Panin
    Carabus (Ulocarabus) stschurovskii Solsky, 1874
    Carabus (Ulocarabus) stschurovskii Solsky, 1874
    Male adult
    Tadzhikistan, Pamir-Alay, S slope Gissar Mt.Range, Takob, h-1700m, 10.V.1996, Dolin V.G. leg.
    det. Panin R.Yu.
    L=22 mm
    Copyright © Ruslan Panin
    Carabus (Hemicarabus) nitens Linne, 1758
    Carabus (Hemicarabus) nitens Linne, 1758
    Male adult
    16,6 mm, England, Dorset, Studland
    leg.det. J.W.Yerbury, 18.-29.VIII.2006
    Coll. Oxford University Museum of Natural History
    Photo: U.Schmidt, 2011
    Copyright © Udo Schmidt
    Carabus (Morphocarabus) monilis Fabricius, 1792
    Carabus (Morphocarabus) monilis Fabricius, 1792
    Male adult
    24 mm, England, Warwickshire
    leg.det. D.J.Mann, 15.IV.1993
    Photo: U.Schmidt, 2011
    Copyright © Udo Schmidt
    Carabus (Morphocarabus) rothi hampei (as diffinis Csiki, 1906)
    Carabus (Morphocarabus) rothi hampei (as diffinis Csiki, 1906)
    Male adult
    NW Romania, Apuseni Mts, Trascău Mt Rng, Petresti de Jos, 12 km E Turda,. 6.VIII.2008
    L=27 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi hampei (as diffinis Csiki, 1906)
    Carabus (Morphocarabus) rothi hampei (as diffinis Csiki, 1906)
    Endophallus
    NW Romania, Apuseni Mts, Trascău Mt Rng, Petresti de Jos, 12 km E Turda,.
    Copyright © Ruslan Panin
    Carabus (Ulocarabus) theanus Reitter, 1895
    Carabus (Ulocarabus) theanus Reitter, 1895
    Male adult
    Tadzhikistan, S slope Gissar Mt.Rng, left bank of the river Sorbo,Haidar gorge,1.VI.1984,Dolin V.G. leg.
    det.Panin R.Yu.
    L=21 mm
    Copyright © Ruslan Panin
    Carabus (Ulocarabus) theanus Reitter, 1895
    Carabus (Ulocarabus) theanus Reitter, 1895
    Endophallus
    Tadzhikistan, S slope Gissar Mt.Range, left bank of the river Sorbo, Haidar gorge,
    Copyright © Ruslan Panin
    Carabus (Carabus) granulatus yezoensis Bates, 1883
    Carabus (Carabus) granulatus yezoensis Bates, 1883
    Male adult
    Russia, S-Sakhalin isl., Makarov distr., Mt. Chekhova h=400m, 12.VI.1998, M. Nesterov leg.<br /> L=23 mm
    Copyright © Ruslan Panin
    Carabus (Ulocarabus) stschurovskii Solsky, 1874
    Carabus (Ulocarabus) stschurovskii Solsky, 1874
    Female adult
    Tadzhikistan, Pamir, N-slope Peter I.Mt Rng,h=2100m,Ganishou vill., Dolin V.G. leg.
    det.Panin R.Yu.
    L=24 mm
    Copyright © Ruslan Panin
    Carabus (Deroplectes) klapperichianus klapperichianus Mandl, 1955
    Carabus (Deroplectes) klapperichianus klapperichianus Mandl, 1955 Holotype
    Male adult
    image from carabus-online.de
    Copyright © Frank Kleinfeld
    Carabus (Deroplectes) klapperichianus salangicus Kleinfeld & Reuter, 2010
    Carabus (Deroplectes) klapperichianus salangicus Kleinfeld & Reuter, 2010 Holotype
    Male adult
    Afghanistan, Salang Pass, Nordseite, 25°27'N/68°58'E, ca. 2300m, 8.-18.IV.2010 (reg. subalp.) (in coll. Reuter)
    23 mm
    Copyright © Frank Kleinfeld
    Carabus (Deroplectes) klapperichianus salangicus Kleinfeld & Reuter, 2010
    Carabus (Deroplectes) klapperichianus salangicus Kleinfeld & Reuter, 2010 Paratype
    Female adult
    Afghanistan, Salang Pass, Nordseite, 25°27'N/68°58'E, ca. 2300m, 8.-18.IV.2010 (reg. subalp.) (in coll. Reuter)
    27 mm
    Copyright © Frank Kleinfeld
    Carabus (Pagocarabus) nanschanicus edacis Cavazzuti, 2006
    Carabus (Pagocarabus) nanschanicus edacis Cavazzuti, 2006 Holotype
    Male adult
    China, Gansu, Str. Lintan - Diebu, 2. Pass & Bergmassiv, 3900m, 30.V.-19.VI.2009 (in coll. Puchner)
    22,5 mm
    photo from carabus-online.de
    Copyright © ---
    Carabus (Pagocarabus) nanschanicus edacis Cavazzuti, 2006
    Carabus (Pagocarabus) nanschanicus edacis Cavazzuti, 2006 Paratype
    Female adult
    China, Gansu, Str. Lintan - Diebu, 2. Pass & Bergmassiv, 3900m, 30.V.-19.VI.2009 (in coll. Puchner)
    26 mm
    photo from carabus-online.de
    Copyright © ---
    Carabus (Apotomopterus) kafka Kleinfeld & Reuter, 2010
    Carabus (Apotomopterus) kafka Kleinfeld & Reuter, 2010 Holotype
    Male adult
    China, Guangxi, Mao'er Shan (ca. 25°50N/110°22E), 1600-1900m, 29.V.-10.VI.2009, leg. Chr. Reuter (in coll. Reuter); 25,5 mm
    Copyright © Frank Kleinfeld
    Carabus (Procerus) scabrosus caucasicus Adams, 1817
    Carabus (Procerus) scabrosus caucasicus Adams, 1817
    Male adult
    N Caucasus,
    Stavropol Krai, Pyatigorsk, Mt.Mashuk, VI.2007 Tikhonov Yu. leg.
    44mm
    Copyright © Ruslan Panin
    Carabus (Procerus) gigas duponcheli Dejean, 1831 (as kusdasi Mandl, 1966)
    Carabus (Procerus) gigas duponcheli Dejean, 1831 (as kusdasi Mandl, 1966)
    Male adult
    S-Greece: Peloponnes, Mt.Chelmos, 20.V.2007.
    L=46 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) excellens (as lomnitzkii Reitter, 1896)
    Carabus (Morphocarabus) excellens (as lomnitzkii Reitter, 1896)
    Female adult
    W Ukraine, NW Podolian Upland, Lviv reg., Zolochiv distr., Steppe Reserve Lysa Gora, 19.V-7.VI.2007, Panin R. leg.<br /> L=26 mm
    Copyright © Ruslan Panin
    Carabus (Carabus) stscheglowi Mannerheim, 1827
    Carabus (Carabus) stscheglowi Mannerheim, 1827
    Male adult
    S Ukraine, Crimea,<br /> Chatyr-Dag jayla, h=1000m, 11-23.VIII.2000, Solodovnikov I. leg.<br /> L=19 mm
    Copyright © Ruslan Panin
    Carabus (Aulonocarabus) gaschkewitschi czarensis Obydov, 1997
    Carabus (Aulonocarabus) gaschkewitschi czarensis Obydov, 1997
    Female adult
    Russia, CE-Siberia, N Chita reg., Kodar Mt. Range, Big Leprindo Lake env. h~2000m., 2-5.VII.2002 A. Prokhorov leg./det. A. Anichtchenko
    L=19 mm
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) eccoptopteroides Rapuzzi, 2012
    Carabus (Apotomopterus) eccoptopteroides Rapuzzi, 2012 Holotype Paratype
    Male+Female adult
    China, South Sichuan province, Pu-Ge County, Lianxiang, Kakaliangzi, 1-11.VI.2012 (deposited in private collection of Rapuzzi)<br /> Image Ivan Rapuzzi, 2012
    Copyright © - - -
    Carabus (Apotomopterus) francottei Rapuzzi, 2012
    Carabus (Apotomopterus) francottei Rapuzzi, 2012 Holotype
    Male adult
    China, North Guangdong province, Mts. Nanling Shan, VI.2009 (deposited in collection of Rapuzzi). Image Ivan Rapuzzi, 2012
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus gougeleti Reiche, 186
    Carabus (Mesocarabus) lusitanicus gougeleti Reiche, 186
    Male adult
    CW Spain, Province Huelva, Arroyomolinos de León,30.X.2011, T. Alcantara leg.; L=23 mm
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) (? lusitanicus) gougeleti Reiche, 186
    Carabus (Mesocarabus) (? lusitanicus) gougeleti Reiche, 186
    Endophallus
    CW Spain, Province Huelva, Arroyomolinos de Leon.
    Copyright © Ruslan Panin
    Carabus (Platycarabus) fabricii ucrainicus Lazorko, 1951
    Carabus (Platycarabus) fabricii ucrainicus Lazorko, 1951
    Male adult
    Ukrainian Carpathians, Ivano-Frankivsk reg., Yaremcha distr., Gorgany Mts., h=1590 m, 22.VII-3.VIII.08, Panin R. leg., male, 23mm
    Copyright © Ruslan Panin
    Carabus (Platycarabus) fabricii ucrainicus Lazorko, 1951
    Carabus (Platycarabus) fabricii ucrainicus Lazorko, 1951
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Platycarabus) fabricii ucrainicus Lazorko, 1951
    Carabus (Platycarabus) fabricii ucrainicus Lazorko, 1951
    Male adult
    Ukrainian Carpathians, Ivano-Frankivsk reg., Gorgany Mts., h=1590 m, 17.VI-26.VII.2011, Panin R. leg.<br /> male, 23mm
    Copyright © Ruslan Panin
    Carabus (?Oreocarabus) titanus Breuning, 1933
    Carabus (?Oreocarabus) titanus Breuning, 1933
    Endophallus
    C-China Sichuan prov., Nanping 1500m,
    Copyright © Ruslan Panin
    Carabus (Megodontus) kolbei kolbei Roeschke, 1887
    Carabus (Megodontus) kolbei kolbei Roeschke, 1887
    Endophallus
    N-Japan, C-Hokkaido, Hidaka distr., Niikappu env.,
    Copyright © Ruslan Panin
    Carabus (Leptocarabus) procerulus Chaudoir, 1862
    Carabus (Leptocarabus) procerulus Chaudoir, 1862
    Endophallus
    Japan:S Honshu , Matsubara-ko,
    Copyright © Ruslan Panin
    Carabus (Megodontus) kolbei kolbei Roeschke, 1887
    Carabus (Megodontus) kolbei kolbei Roeschke, 1887
    Male adult
    N-Japan, C-Hokkaido, Hidaka distr., Niikappu env., 27.VIII.1989, Sasaki leg,
    L=27 mm
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) latreillei ?semicoriaceus Kraatz, 1881
    Carabus (Trachycarabus) latreillei ?semicoriaceus Kraatz, 1881
    Endophallus
    NE-China: Liaodong Peninsula, Liaoning prov., Gaizhou distr., Mt. Buyun Shan.
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) latreillei ? semicoriaceus Kraatz, 1881
    Carabus (Trachycarabus) latreillei ? semicoriaceus Kraatz, 1881
    Male adult
    NE-China: Liaodong Peninsula, Liaoning prov., Gaizhou distr., Mt. Buyun Shan,VI.2007, Hao leg.
    L=23 mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) kolbei hidakamontanus Ishikawa, 1966
    Carabus (Megodontus) kolbei hidakamontanus Ishikawa, 1966
    Male adult
    N-Japan, C-Hokkaido, Hidaka Mt.Rng, Mt.Apoidak near Samani town, 20.VIII.1982,T. Akaishi leg.
    L=24 mm
    Copyright © Ruslan Panin
    Carabus (Acoptolabrus) constricticollis constricticollis (as grallatorius Roeschke, 1921)
    Carabus (Acoptolabrus) constricticollis constricticollis (as grallatorius Roeschke, 1921)
    Male adult
    NE-Korea:North Hamgyong Prov., Sechon-dong, VII.2004, Li leg.
    L=28 mm
    Copyright © Ruslan Panin
    Carabus (Eurycarabus) famini lucasi Gaubil, 1849
    Carabus (Eurycarabus) famini lucasi Gaubil, 1849
    Male adult
    N-Tunisia, Bizerte reg., Menzel Bourguiba city,h=5 m, XII.1981, S. Remond leg.
    L=25 mm
    Copyright © Ruslan Panin
    Carabus (Carabus) arvensis hokkaidoensis Lapouge, 1924
    Carabus (Carabus) arvensis hokkaidoensis Lapouge, 1924
    Endophallus
    N Japan:Hokkaido, Hamamasu env.,
    Copyright © Ruslan Panin
    Carabus (Leptocarabus) procerulus Chaudoir, 1862
    Carabus (Leptocarabus) procerulus Chaudoir, 1862
    Male adult
    Japan:S Honshu, Matsubara-ko, VIII.1997, S. Remond leg.
    L=30 mm
    Copyright © Ruslan Panin
    Carabus (Carabus) arvensis hokkaidoensis Lapouge, 1924
    Carabus (Carabus) arvensis hokkaidoensis Lapouge, 1924
    Male adult
    N Japan: Hokkaido, Hamamasu env.,1-5.VI.2002, Eiji Yamauchi leg.
    L=20 mm
    Copyright © Ruslan Panin
    Carabus (Ohomopterus) japonicus japonicus Motschulsky, 1857
    Carabus (Ohomopterus) japonicus japonicus Motschulsky, 1857
    Endophallus
    S Japan, Kyushu, Kumamoto Pref., Izumi viil., h=600 m
    Copyright © Ruslan Panin
    Carabus (Ohomopterus) japonicus japonicus Motschulsky, 1857
    Carabus (Ohomopterus) japonicus japonicus Motschulsky, 1857
    Male adult
    S Japan, Kyushu, Kumamoto Pref., Izumi viil., h=600 m,VII.2001, S. Remond leg.
    L=23 mm
    Copyright © Ruslan Panin
    Carabus (Piocarabus) titanus Breuning, 1933
    Carabus (Piocarabus) titanus Breuning, 1933
    Male adult
    C-China Sichuan prov., Nanping, h=1500m, VI.2000, S. Remond leg.<br /> L=34 mm
    Copyright © Ruslan Panin
    Carabus (Eurycarabus) famini fezzanus Lapouge, 1915
    Carabus (Eurycarabus) famini fezzanus Lapouge, 1915
    Endophallus
    Morocco, Middle Atlas Mt Rng, Azrou city h=1500m,
    Copyright © Ruslan Panin
    Carabus (Eurycarabus) famini fezzanus Lapouge, 1915
    Carabus (Eurycarabus) famini fezzanus Lapouge, 1915
    Male adult
    Morocco, Middle Atlas Mt Rng, Azrou city h=1500m, IV.1999, S. Remond leg.
    L=23 mm
    Copyright © Ruslan Panin
    Carabus (Leptocarabus) procerulus Chaudoir, 1862
    Carabus (Leptocarabus) procerulus Chaudoir, 1862
    Endophallus
    Japan, S Honshu, Mie Pref., Hakusan,
    Copyright © Ruslan Panin
    Carabus (Leptocarabus) procerulus Chaudoir, 1862
    Carabus (Leptocarabus) procerulus Chaudoir, 1862
    Male adult
    Japan, S Honshu, Mie Pref., Hakusan, VII.2001, S. Remond leg.<br /> L=33 mm
    Copyright © Ruslan Panin
    Carabus (Oreocarabus) taedatus agassi Le Conte, 1850
    Carabus (Oreocarabus) taedatus agassi Le Conte, 1850
    Male adult
    SC-USA, New Mexico, Santa Fe,VIII.1999, Sehnal R. leg.
    L=23 mm
    Copyright © Ruslan Panin
    Carabus (Oreocarabus) taedatus agassi Le Conte, 1850
    Carabus (Oreocarabus) taedatus agassi Le Conte, 1850
    Endophallus
    SC-USA, New Mexico, Santa Fe,
    Copyright © Ruslan Panin
    Carabus (Ohomopterus) yamato kitai (Imura, 2004)
    Carabus (Ohomopterus) yamato kitai (Imura, 2004)
    Endophallus
    Japan, S Honshu, Mie Pref., Hakusan,
    Copyright © Ruslan Panin
    Carabus (Ohomopterus) yamato kitai (Imura, 2004)
    Carabus (Ohomopterus) yamato kitai (Imura, 2004)
    Male adult
    Japan, S Honshu, Mie Pref., Hakusan, VII.2001, S. Remond leg.
    L=21 mm
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
    Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
    Endophallus
    N Mongolia, Khövsgöl reg., Lake Khovsgol, h=1640m, 51*:12’N; 100*:45’E, steppificated sites,
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
    Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
    Male adult
    N Mongolia, Khövsgöl reg., Lake Khovsgol, h=1640m, 51*:12’N; 100*:45’E, steppificated sites,
    20-25.VII.2010. A.Dergalev leg.
    L=22 mm
    Copyright © Ruslan Panin
    Carabus (Platycarabus) fabricii ucrainicus Lazorko, 1951
    Carabus (Platycarabus) fabricii ucrainicus Lazorko, 1951
    Female adult
    W-UKRAINE, Ivano-Frankivsk reg., Gorgany Mts, Syvula Mt., h =1820 м., 25.VII.2011,Panin R. leg., L=27 mm<br />
    Copyright © Ruslan Panin
    Carabus (Carabus) granulatus granulatus Linne, 1758
    Carabus (Carabus) granulatus granulatus Linne, 1758
    Male adult
    W-Ukraine, Roztochia,
    Lviv Reg., Javorov distr., near Ivano-Frankovo vill., Site Zalyvki, 17.IV-3.V.2010, Panin R. leg.
    L=20 mm

    Copyright © Ruslan Panin
    Carabus (Procrustes) piochardi piochardi Gehin, 1883
    Carabus (Procrustes) piochardi piochardi Gehin, 1883
    Male adult
    C-Lebanon, Faraya vill., Mt. Djebel Sannin h=1800m, X.2012, C. Reuter leg. <br /> L=31 mm
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) problematicus islandicus Lindroth, 1968
    Carabus (Mesocarabus) problematicus islandicus Lindroth, 1968
    Female adult
    SE-Iceland:Höfn env.:Hornafjördur, 24-28.VIII.2008, C. Reuter leg.
    L=24 mm
    Copyright © Ruslan Panin
    Carabus (Coptolabrus) pustulifer shangzhouicus Deuve & Li, 2003
    Carabus (Coptolabrus) pustulifer shangzhouicus Deuve & Li, 2003
    Male adult
    C-Chine, SE-Shaanxi Province, Shahezi, Shangzhou City, 6-18.VII.2010<br /> L=43 mm
    Copyright © Ruslan Panin
    Carabus (Pseudocranion) lazikouensis lazikouensis (as kundaensis Deuve, 2003)
    Carabus (Pseudocranion) lazikouensis lazikouensis (as kundaensis Deuve, 2003) Paratype
    Female adult
    C-China, S-Gansu: SSW of Minxian: Qilian Shan: W Kunda, 34:10N/103:47E, h=2800-3000m, forest, 7.VII.2002, Belousov & Kabak leg.
    L=24 mm
    Copyright © Ruslan Panin
    Carabus (Rhigocarabus) buddaicus gansuicus Deuve, 1991
    Carabus (Rhigocarabus) buddaicus gansuicus Deuve, 1991
    Male adult
    China, NC-Sichuan (ZOIGE), LANGMUSI (=DOGCANGLHAMO), 34:03:48N/102:38:08E, h=4100m, alpine meadows, screes, 16-18.VII.1999, B.Brezina leg.
    L=20 mm
    Copyright © Ruslan Panin
    Carabus (Ulocarabus) theanus delerei Mandl, 1955
    Carabus (Ulocarabus) theanus delerei Mandl, 1955
    Male adult
    N-Afghanistan, Hindukush Mt.Rng, NE Salang Pass h=2300 m, 35°27'494"N, 68°58'399"E, IV.2011, C. Reuter leg.
    L=23 mm
    Copyright © Ruslan Panin
    Carabus (Ulocarabus) theanus theanus Reitter, 1895
    Carabus (Ulocarabus) theanus theanus Reitter, 1895
    Male adult
    Vakhskiy Mt Range,Saryhosor, Mullonomi,
    29.4.1990, S.Zonshteyn leg.
    Copyright © Alexander Anichtchenko
    Carabus (Ulocarabus) theanus theanus Reitter, 1895
    Carabus (Ulocarabus) theanus theanus Reitter, 1895
    Male adult
    Tadzhikistan, S slope Gissar Mt.Rng, left bank of the river Sorbo, Haidar gorge,1.VI.1984, Dolin V.G. leg./det.Panin R.Yu.
    L=21 mm
    Copyright © Ruslan Panin
    Carabus (Ulocarabus) theanus theanus Reitter, 1895
    Carabus (Ulocarabus) theanus theanus Reitter, 1895
    Endophallus
    Tadzhikistan, S slope Gissar Mt.Range, left bank of the river Sorbo, Haidar gorge,
    Copyright © Ruslan Panin
    Carabus (Ulocarabus)? theanus theanus Reitter, 1895
    Carabus (Ulocarabus)? theanus theanus Reitter, 1895
    Male adult
    NC Tadzhikistan, Pamir, N-slope Peter I.Mt Rng, h=2100m, Ganishou vill., 10-12.VI.2006, Ivanova E. leg.
    L=23 mm
    Copyright © Ruslan Panin
    Carabus (Aulonocarabus) canaliculatus hailinensis Deuve & Li, 2000
    Carabus (Aulonocarabus) canaliculatus hailinensis Deuve & Li, 2000
    Male adult
    NE China, Heilongjiang Prov., Mudanjiang City, Mt.Tai-Gu-Ding-zi h=800m, 15-25.VI.2010
    L=29 mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) croaticus zepcensis (as bosiljevici Drovenik & Pavićević, 1985)
    Carabus (Megodontus) croaticus zepcensis (as bosiljevici Drovenik & Pavićević, 1985)
    Male adult
    CW-Serbia, Zlatar Mts, Nova Varoš, Zlatar Lake env., 1-12.VII.2007, P.Sima leg. L=30 mm
    Copyright © Ruslan Panin
    Carabus (Lamprostus) saulcyi saulcyi Piochard, 1875 (as chambouqensis)
    Carabus (Lamprostus) saulcyi saulcyi Piochard, 1875 (as chambouqensis)
    Female adult
    N-Lebanon, Fnaпdek, Djebel Qammoua, h=1100-1200m, 29.X.2012, C. Reuter leg.
    L=27 mm
    Copyright © Ruslan Panin
    Carabus (Meganebrius) kadoudali (Morvan, 1981)
    Carabus (Meganebrius) kadoudali (Morvan, 1981)
    Male+Female adult
    C-Nepal, SW Dhaulagiri, W Jaljala h=3150 m, 28°30'05"N, 83°11'48"E, 23.IX.2012, J. Schmidt leg.
    female, 20 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) illigeri versicolor Frivaldsky, 1835 (ab.)
    Carabus (Morphocarabus) illigeri versicolor Frivaldsky, 1835 (ab.)
    Female adult
    C Bulgaria: Stara Planina Mts, Schipka pass h=1450m, 12-15.VI.2007, M.Janalik leg. L=31 mm
    Copyright © Ruslan Panin
    Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
    Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
    Female adult
    N- Romania, E-Carpathians, Rodna Mts, h=2000m, 7-8.VI.2012, Cartuz Z.leg.
    L=23 mm
    Copyright © Ruslan Panin
    Carabus (Eucarabus) manfredeggeri Müller 2012
    Carabus (Eucarabus) manfredeggeri Müller 2012 Paratype
    Male+Female adult
    N-Korea, Chagang Prov., Manpo City, Songhak-ri, 2-16.VII.2011
    female, 21 mm
    male, 19 mm
    Copyright © Ruslan Panin
    Carabus (Eucarabus) manfredeggeri Müller 2012
    Carabus (Eucarabus) manfredeggeri Müller 2012 Paratype
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Megodontus) caelatus dalmatinus (as procerus Reitter, 1885)
    Carabus (Megodontus) caelatus dalmatinus (as procerus Reitter, 1885)
    Male adult
    S Bosnia, Potoci env. h=500 m, VII.2006, Bartoloni leg.
    L=38 mm
    Copyright © Ruslan Panin
    Carabus (Imaibius) dardiellus perobscurus Deuve, 1980
    Carabus (Imaibius) dardiellus perobscurus Deuve, 1980
    Male adult
    N India.Kashmir, Loran, W part Pir Panchal range,h=2400-2500m, 33°49'"N, 74°20'E
    L=25 mm
    Copyright © Ruslan Panin
    Carabus (Imaibius) dardiellus perobscurus Deuve, 1980
    Carabus (Imaibius) dardiellus perobscurus Deuve, 1980
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Coptolabrus) smaragdinus dachangensis Lu & Müller, 2009
    Carabus (Coptolabrus) smaragdinus dachangensis Lu & Müller, 2009
    Female adult
    NE-China, Liaoning Prov., Changhai County, Dachangshan Dao Isl.,12.VI.2011
    L=40 mm
    Copyright © Ruslan Panin
    Carabus (Tomocarabus) rumelicus syriensis Breuning, 1943
    Carabus (Tomocarabus) rumelicus syriensis Breuning, 1943
    Male adult
    N Lebanon, Fnaпdek, Djebel Qammoua, h=1100-1200m, C. Reuter leg.
    L=17.5 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) illigeri illigeri (as tarensis Toševski, 1990)
    Carabus (Morphocarabus) illigeri illigeri (as tarensis Toševski, 1990)
    Male adult
    WC-Serbia: National park Tara Mts.:NE Mitrovac h=1500m, 1-12.VII.2007, Pr.Sima leg. L=30 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) illigeri illigeri (as tarensis Toševski, 1990)
    Carabus (Morphocarabus) illigeri illigeri (as tarensis Toševski, 1990)
    Endophallus
    WC-Serbia: National park Tara Mts.:NE Mitrovac h=1500m,
    Copyright © Ruslan Panin
    Carabus (Cupreocarabus) huangi huangi Deuve, 1992
    Carabus (Cupreocarabus) huangi huangi Deuve, 1992
    Male adult
    C-China, E Tibet, N-Sichuan, Hongyuan,valley 10 km SE Sanggarpar, h=4200m, 32.300053, 102.584031, alpine meadow / scree, 19.VII.1995, B.Brezina leg.<br /> L=21 mm
    Copyright © Ruslan Panin
    Carabus (Cupreocarabus) huangi huangi Deuve, 1992
    Carabus (Cupreocarabus) huangi huangi Deuve, 1992
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Megodontus) violaceus dryas Gistl, 1857
    Carabus (Megodontus) violaceus dryas Gistl, 1857
    Endophallus
    NE-Greece, Mt. Olympos, Litochoro env.
    Copyright © Ruslan Panin
    Carabus (Acoptolabrus) mirabilissimus rigoutianus Deuve & Li, 1998
    Carabus (Acoptolabrus) mirabilissimus rigoutianus Deuve & Li, 1998
    Male adult
    N-Korea, South Pyongan prov., Mt. Maeng-san, 6.VI.2010
    L=25 mm
    Copyright © Ruslan Panin
    Carabus (Pseudocranion) taibaishanicus taibaishanicus Deuve, 1989
    Carabus (Pseudocranion) taibaishanicus taibaishanicus Deuve, 1989
    Female adult
    China, W-Shaanxi, Taibai Shan Mts, btw.Houzhenzi/Ba Xian Tai, nr. Nan Tian Men h=3200m,
    33°55'17"N/ 107°46'53"E, 25-30.VII.2011, Li Wuyang leg.
    L=29 mm
    Copyright © Ruslan Panin
    Carabus (Lamprostus) saulcyi saulcyi Piochard, 1875
    Carabus (Lamprostus) saulcyi saulcyi Piochard, 1875
    Endophallus
    C Lebanon, Jbeil Distr., Ehmej vill., h=1100 m
    Copyright © Ruslan Panin
    Carabus (Megodontus) violaceus dryas Gistl, 1857
    Carabus (Megodontus) violaceus dryas Gistl, 1857
    Male adult
    NE-Greece, Mt. Olympus, Litochoro env., 20-30.V.2005, Sima leg.
    L=27 mm
    Copyright © Ruslan Panin
    Carabus (Lamprostus) saulcyi saulcyi Piochard, 1875
    Carabus (Lamprostus) saulcyi saulcyi Piochard, 1875
    Male adult
    C Lebanon, Jbeil Distr., Ehmej vill., h=1100 m.V.2012, C. Reuter leg.
    L=24 mm
    Copyright © Ruslan Panin
    Carabus (Imaibius) dardiellus ? subpunctulus Roeschke, 1907
    Carabus (Imaibius) dardiellus ? subpunctulus Roeschke, 1907
    Male adult
    N India, Kashmir, Pahalgam, Aru Valley h=2400m, VII.2007
    L=25 mm
    Copyright © Ruslan Panin
    Carabus (Pachystus) hortensis hortensis Linne, 1758
    Carabus (Pachystus) hortensis hortensis Linne, 1758
    Endophallus
    W-Ukraine, Male Polissya, near Kamenka-Buzka vill.
    Copyright © Ruslan Panin
    Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847
    Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847
    Male adult
    WC Spain, Cáceres, Hervás, h=1000m, V.2007, Pascal Stefani leg.
    L=23 mm
    Copyright © Ruslan Panin
    Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847
    Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847
    Endophallus
    WC Spain, Cáceres, Hervás, h=1000m
    Copyright © Ruslan Panin
    Carabus (?Oreocarabus) qinlingensis Imura, 1993
    Carabus (?Oreocarabus) qinlingensis Imura, 1993
    Male adult
    China, W-Shaanxi, Taibai Shan Mts, btw. Houzhenzi/Ba Xian Tai, nr.Nan Tian Men h=3200m, 33°55'17"N/107°46'53"E, 25-30.VII.2011, Li Wuyang leg.
    L=19 mm
    Copyright © Ruslan Panin
    Carabus (?Oreocarabus) qinlingensis Imura, 1993
    Carabus (?Oreocarabus) qinlingensis Imura, 1993
    Endophallus
    China, W-Shaanxi, Taibai Shan Mts, btw.Houzhenzi/Ba Xian Tai, nr.Nan Tian Men h=3200m, 33°55'17"N/107°46'53"E,
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi hampei (as hunyadensis Csiki, 1926)
    Carabus (Morphocarabus) rothi hampei (as hunyadensis Csiki, 1926)
    Male adult
    CW Romania, Apuseni Mts, Metalliferous Mt Rng, Hunedoara reg., near Săcărâmb vill., 45°97'472"N/23°02'630"E, h=680 m, mixed forest, meadows, 15-20.V.2011, B.Brezina leg.
    L=36 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi hampei (as hunyadensis Csiki, 1926)
    Carabus (Morphocarabus) rothi hampei (as hunyadensis Csiki, 1926)
    Endophallus
    CW Romania, Apuseni Mts, Metalliferous Mt Rng, Hunedoara reg., near Săcărâmb vill.
    Copyright © Ruslan Panin
    Carabus (Meganebrius) pseudoharmandi Mandl, 1965
    Carabus (Meganebrius) pseudoharmandi Mandl, 1965
    Female adult
    E-Nepal, Kosi zone, Khandbari,(1.5km SW of Chichila vill., 27*45’843’’N/ 87*22’473’E, h=1940m), damaged primary forest, 13 .V.2010, B.Brezina & M.Häckel leg.
    L=23 mm

    Copyright © Ruslan Panin
    Carabus (Morphocarabus) excellens excellens ( as jasilkowskii Born,1905)
    Carabus (Morphocarabus) excellens excellens ( as jasilkowskii Born,1905)
    Male adult
    NE Romania, 5 km W Suceava, 47°63′940″N 26°18′795″E,h=340m, agricultural field, 15-20.V.2011, B. Brezina leg. L=24 mm
    Copyright © Ruslan Panin
    Carabus (Meganebrius) arunensis Heinertz, 1980
    Carabus (Meganebrius) arunensis Heinertz, 1980
    Male adult
    E-Nepal, Kosi zone, Khandbari, 1.5km SW of Chichila vill., 27*47’175’’N/ 87*23’622’E, h=1880m, damaged primary forest, 11.V.2010, B.Brezina & M.Häckel leg.
    L=27 mm

    Copyright © Ruslan Panin
    Carabus (Ophiocarabus) ballionis Kraatz, 1879
    Carabus (Ophiocarabus) ballionis Kraatz, 1879
    Male adult
    NW-China, Xinjiang prov., 40 km N Narat, h=1500, 12-12.VI.2004, Benes & Kaiser leg.
    L=22 mm
    Copyright © Ruslan Panin
    Carabus (Coptolabrus) smaragdinus liaodongensis (Li, 1992)
    Carabus (Coptolabrus) smaragdinus liaodongensis (Li, 1992)
    Male adult
    NE-China, E-Liaoning: Anshan: Mt.Qian Shan, 9-14.VI.2011. L=39 mm
    Copyright © Ruslan Panin
    Carabus (Oreocarabus) latro minshanensis Deuve, 1987
    Carabus (Oreocarabus) latro minshanensis Deuve, 1987
    Male adult
    China, N-Sichuan, Jiuzhaigou env., 13.VI.2001,E. Kučera leg.
    L=23 mm
    Copyright © Ruslan Panin
    Carabus (Oreocarabus) latro minshanensis Deuve, 1987
    Carabus (Oreocarabus) latro minshanensis Deuve, 1987
    Endophallus
    China, N-Sichuan, Jiuzhaigou env.,
    Copyright © Ruslan Panin
    Carabus (Lamprostus) syrus pseudopinguis Heinz & Staven, 2000
    Carabus (Lamprostus) syrus pseudopinguis Heinz & Staven, 2000
    Male adult
    NW-Jordan, Ajlun env., oak forests on dry limestone, h=897m, 32°21′33.1″N/ 035°43’47.4″E, 4.V.2010, Schnitter/Schellhorn leg.
    L=26 mm
    Copyright © Ruslan Panin
    Carabus (Meganebrius) quinlani smetanai Häckel, 2011
    Carabus (Meganebrius) quinlani smetanai Häckel, 2011 Paratype
    Male+Female adult
    C-Nepal:Dhaulagiri Massif:Baglung Distr., (1.5km W of Okhale vill., 28.32’152’’N/ 83.48’673’E, h=2600m) damaged primary forest, 19.V.2010, B.Brezina & M.Häckel leg.
    male, 21mm
    female, 23 mm
    Copyright © Ruslan Panin
    Carabus (Meganebrius) quinlani smetanai Häckel, 2011
    Carabus (Meganebrius) quinlani smetanai Häckel, 2011 Paratype
    Endophallus
    C-Nepal:Dhaulagiri Massif:Baglung Distr.,(1.5km W of Okhale vill., 28.32’152’’N/ 83.48’673’E, h=2600m)
    Copyright © Ruslan Panin
    Carabus (Megodontus) violaceus vlasuljensis Apfelbeck, 1894
    Carabus (Megodontus) violaceus vlasuljensis Apfelbeck, 1894
    Male adult
    WC-Serbia, Borak - Gornje Babine, 1-10.VII.2007, M. Sima leg.
    L=28 mm
    Copyright © Ruslan Panin
    Carabus (Rhigocarabus) indigestus confucius Breuning, 1933
    Carabus (Rhigocarabus) indigestus confucius Breuning, 1933
    Female adult
    C-China, E Tibet,W-Sichuan prov., Kangding, Zheduo Pass h=4300m 30.VI.2005 , V.Benes leg,
    L=17 mm
    Copyright © Ruslan Panin
    Carabus (Rhigocarabus) ladygini degensis Deuve, 1989
    Carabus (Rhigocarabus) ladygini degensis Deuve, 1989
    Endophallus
    C-China, NW- Sichuan prov., Garzê Tibetan AP, Pass Garzê – Sêrtar
    Copyright © Ruslan Panin
    Carabus (Megodontus) violaceus azurescens Dejean, 1826
    Carabus (Megodontus) violaceus azurescens Dejean, 1826
    Endophallus
    W-Croatia, Velebit Mts., betw.Karlobag & Brusane vill.,
    Copyright © Ruslan Panin
    Carabus (Rhigocarabus) ladygini degensis Deuve, 1989
    Carabus (Rhigocarabus) ladygini degensis Deuve, 1989
    Male adult
    C-China, NW- Sichuan prov., Garzê Tibetan AP, Pass Garzê – Sêrtar, 19.VII.2007, V.Benes leg.
    L=17 mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) violaceus azurescens Dejean, 1826 (loc.typ.)
    Carabus (Megodontus) violaceus azurescens Dejean, 1826 (loc.typ.)
    Male adult
    W-Croatia, Velebit Mts., betw.Karlobag & Brusane vill., VI.2002, Manfred Egger leg.
    L=30 mm
    Copyright © Ruslan Panin
    Carabus (Apoplesius) pseudodepressus Machard, 1988
    Carabus (Apoplesius) pseudodepressus Machard, 1988
    Female adult
    Turkey, Prov. Bitlis, S near Tatvan, 23.IV.2009 leg. M. & S. Nabozhenko
    L=19.0 mm
    Copyright © Kirill Makarov
    Carabus (Megodontus) caelatus caelatus (as malissorum Apfelbeck, 1919)
    Carabus (Megodontus) caelatus caelatus (as malissorum Apfelbeck, 1919)
    Male adult
    Montenegro, National park Durmitor; near Zabljak vill., Mt. Sedlo h=2200 m, 6-22.VIII.2010, Franko Sandel leg.
    29 мм
    Copyright © Ruslan Panin
    Carabus (Rhigocarabus) buddaicus linxiaicus Deuve, 1995
    Carabus (Rhigocarabus) buddaicus linxiaicus Deuve, 1995
    Male adult
    China, S-Gansu (LABRANG):, valley.E of Ponggartang, 35:12-13N/102:54-55E, h=3000-3700m, alpine meadows, coniferous forests,10.-14.VII.1999, B.Brezina leg.
    L=20 mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) violaceus germarii Sturm, 1815
    Carabus (Megodontus) violaceus germarii Sturm, 1815
    Male adult
    N Italy, Trent, Ampola, Mt. Tremalzo h=1700m, 3-13.VI.2010, Manfred Egger leg.
    29 mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) violaceus germarii Sturm, 1815
    Carabus (Megodontus) violaceus germarii Sturm, 1815
    Endophallus
    N Italy, Trent, Ampola, Mt. Tremalzo h=1700m
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
    Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
    Male adult
    W-UKRAINE, Transcarpathian region, Rakhiv distr., Chornogora Mt Range, h=800 m, near Bogdan vill., 10.VI.2012, Panin R. leg.
    L= 33 мм
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
    Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
    Male adult
    W Ukraine,Transcarpathian Reg., Rakhiv distr., Marmarosh Mts., near Lug (Lunca la Tisa) vill., h=400 m,12-15.VII.2011
    L=32 mm
    det.Panin R.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
    Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
    Endophallus
    W Ukraine,Transcarpathian Reg., Rakhiv distr., Marmarosh Mts., near Lug vill., h=400 m.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
    Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
    Female adult
    W Ukraine, NE Carpathians,
    Transcarpathian Reg., Marmarosh Mts., Rakhiv distr., 5 km W Dilowe vill., Site Kuziy, 5-15.VII.2010, Demidov G.N. leg./det.Panin R.
    L= 35 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
    Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
    Male adult
    Ukrainian Carpathians,
    Transcarpathian reg., Rakhov distr., Svidovec Mt Rng., Mt. Podpula h=1500-1600m. Lazutkin G. leg, 13-14.VII.1978, det. Panin R.
    L=28mm.
    Copyright © Ruslan Panin
    Carabus (Cupreocarabus) lixianensis lixianensis Deuve, 1990
    Carabus (Cupreocarabus) lixianensis lixianensis Deuve, 1990
    Male adult
    China, E Tibet, N-Sichuan, (Barkam),valley SW of Barkam, h=3200m, 31*53’ N; 102*12’E, mixed/coniferous forest, 22.VII.1995, K.& B.Brezina
    L=23 mm
    Copyright © Ruslan Panin
    Carabus (Carabus) menetriesi pacholei Sokolar, 1911
    Carabus (Carabus) menetriesi pacholei Sokolar, 1911
    Male+Female adult
    Czech Republic, SW Bohemia, Šumava-Volary, 01.V.2007, Slavoj Budka leg.
    Copyright © Ruslan Panin
    Carabus (Tachypus) cristoforii Spence, 1823
    Carabus (Tachypus) cristoforii Spence, 1823
    Female adult
    S France, Pyrénés Mts., Tarascon; 15mm
    Copyright © Aleš Sedláček
    Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878 (as vladobydovi Obydov, 2007)
    Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878 (as vladobydovi Obydov, 2007) Paratype
    Female adult
    Russia, South Primorie, Popova isl., 12-14.07.1990, A. Aljakrinskii leg.
    Copyright © Dmitri Obydov
    Carabus (Coptolabrus) smaragdinus sungkii J.K. Park & J. Park, 2013
    Carabus (Coptolabrus) smaragdinus sungkii J.K. Park & J. Park, 2013 Holotype Paratype
    Male+Female adult
    Holotype male: Is.Gureopdo (37°19′71.42″N, 125°98′72.62″E), Deokjeok archipelago, Gyeonggi-do, 19.viii.2010, J.K.Park; Paratype female: Is. Gureopdo, Deokjeok archipelago, Gyeonggi-do, J.K.Park. Image from original describtion J.K. Park & J. Park, 2013
    Copyright ©
    Carabus (Eucarabus) sternbergi gureopensis J.K. Park & J. Park, 2013
    Carabus (Eucarabus) sternbergi gureopensis J.K. Park & J. Park, 2013 Holotype Paratype
    Male+Female adult
    Is. Gureopdo (37°19′71.42″N, 125°98′72.62″E), Deokjeok archipelago, Gyeonggi-do, Korea, 19.viii.2010, J.K.Park
    Image from original describtion J.K. Park & J. Park, 2013
    Copyright © - - -
    Carabus (Megodontus) bonvouloiri graciliformis Breuning, 1964
    Carabus (Megodontus) bonvouloiri graciliformis Breuning, 1964
    Male adult
    NC Turkey, Gölköy env., Harcbeli pass, 35mm
    Copyright © Aleš Sedláček
    Carabus (Megodontus) bonvouloiri graciliformis Breuning, 1964
    Carabus (Megodontus) bonvouloiri graciliformis Breuning, 1964
    Female adult
    NC Turkey, Gölköy env., Harcbeli pass, 37mm
    Copyright © Aleš Sedláček
    Carabus (Megodontus) kolbei chishimanus Nakane, 1961
    Carabus (Megodontus) kolbei chishimanus Nakane, 1961
    Female adult
    SC Kurils islands, 28mm
    Copyright © Aleš Sedláček
    Carabus (Pachystus) cribratus staneki Sterba, 1931
    Carabus (Pachystus) cribratus staneki Sterba, 1931
    Male adult
    NC Turkey, Küre Mts., 60km N Kastamonu; 25mm
    Copyright © Aleš Sedláček
    Carabus (Pachystus) cribratus staneki Sterba, 1931
    Carabus (Pachystus) cribratus staneki Sterba, 1931
    Female adult
    NC Turkey, Küre Mts., 60km N Kastamonu; 27mm
    Copyright © Aleš Sedláček
    Carabus (Tachypus) auratus honnoratii Dejean, 1826
    Carabus (Tachypus) auratus honnoratii Dejean, 1826
    Male adult
    SE France, Carpentras; 23mm
    Copyright © Aleš Sedláček
    Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906 loc.typ.
    Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906 loc.typ.
    Male+Female adult
    W-UKRAINE,<br /> Transcarpathian region, Uzhgorod distr., h=250 m, 8.VII-25.VIII.2011<br /> Panin R. leg./det.<br /> female, 33 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906 loc.typ.
    Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906 loc.typ.
    Male+Female adult
    W-UKRAINE,<br /> Transcarpathian region, Uzhgorod distr., 28.IV-23.V.2011<br /> Panin R. leg./det.<br /> female, 33 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
    Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
    Endophallus
    Transcarpathian region, Uzhgorod distr., 28.IV-23.V.2011
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
    Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
    Male adult
    SE Slovakia, Slanske vrchy Mts, Herlianske sedlo, 8-19.VI.2009, Krause Rene leg.
    L=30 mm
    det.Panin R.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
    Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
    Female adult
    NW Ukrainian Carpathians,
    Lviv Region, Staryi Sambor Distr., near Dobromyl , h=450m, 9.VI-25.VII.07, Panin R. leg./det.Solodovnikov I. & Panin R.
    female, 35mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
    Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
    Female adult
    NW Ukrainian Carpathians
    Lviv Region, Staryi Sambor Distr., near Dobromyl , h=500m 9.VI-25.VII.2007, Panin R. leg./det.Solodovnikov I. & Panin R.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
    Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
    Endophallus
    SE Slovakia, Slanske vrchy Mts, Herlianske sedlo, 8-19.VI.2009, Krause Rene leg.
    det.Panin R.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) zawadzkii zawadzkii, Kraatz 1854
    Carabus (Morphocarabus) zawadzkii zawadzkii, Kraatz 1854
    Male adult
    W.Ukraine, Ivano-Frankivsk reg.,
    Yaremche distr., near Tatariv vill., Gorgany Mts. h=700 m, 16.VI.2011, Panin R. leg.
    L=31 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) zawadzkii zawadzkii, Kraatz 1854
    Carabus (Morphocarabus) zawadzkii zawadzkii, Kraatz 1854
    Endophallus
    Gorgany Mts. h=700 m, 16.VI.2011,
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) zawadzkii zawadzkii, Kraatz 1854
    Carabus (Morphocarabus) zawadzkii zawadzkii, Kraatz 1854
    Endophallus
    W.UKRAINE, Ukrainian Carpathians
    Lviv Reg., Skole distr., Skolivski Beskidy Mts.
    Copyright © Igor Solodovnikov
    Carabus (Morphocarabus) zawadzkii zawadzkii, Kraatz 1854
    Carabus (Morphocarabus) zawadzkii zawadzkii, Kraatz 1854
    Male+Female adult
    W.UKRAINE, Ukrainian Carpathians
    Lviv Reg., Skole distr., Skolivski Beskidy Mts. Zelemyanka Mt. h=1000m. 9-20.VII.07 Panin R. leg.
    female, 35mm.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) tarbagataicus birlikensis Kabak, 2013
    Carabus (Morphocarabus) tarbagataicus birlikensis Kabak, 2013 Holotype
    Male adult
    China, Xinjiang, cresp of Birliktau Mt.R., WSW Toli, 45 49 23 N / 83 06 13 E, H=2565 m, 25.07.2012, Kabak I.I. leg. (ZIN)
    Copyright © Ilya Kabak
    Carabus (Ophiocarabus) successor koeksuensis Kabak, 2013
    Carabus (Ophiocarabus) successor koeksuensis Kabak, 2013 Paratype
    Male adult
    China, Xinjiang, Koeksu Bas., Tshon-Aidak, SSE Tekes, 42* 32' 47'' N / 82* 05' 56'' E -- 42* 32' 51'' N / 82* 05' 36'' E, 3330-3550 m, 11.07.2012, Kabak I.I. leg.
    Copyright © Ilya Kabak
    Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
    Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
    Male adult
    NE Serbia, Majdanpek; male, 33mm
    Copyright © Aleš Sedláček
    Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
    Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
    Female adult
    NE Serbia, Kučevo; female, 33mm
    Copyright © Aleš Sedláček
    Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
    Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
    Female adult
    NE Serbia, Majdanpek; 35mm
    Copyright © Aleš Sedláček
    Carabus (Morphocarabus) helleri Ganglbauer, 1893
    Carabus (Morphocarabus) helleri Ganglbauer, 1893
    Male adult
    CZ; CE Moravia, Bílé Karpaty Mts.; 24mm
    Copyright © Aleš Sedláček
    Carabus (Morphocarabus) helleri Ganglbauer, 1893
    Carabus (Morphocarabus) helleri Ganglbauer, 1893
    Male adult
    CZ; CE Moravia, Bílé Karpaty Mts.; 25mm
    Copyright © Aleš Sedláček
    Carabus (Heterocarabus) marietti marietti Cristoforis & Jan, 1837
    Carabus (Heterocarabus) marietti marietti Cristoforis & Jan, 1837
    Female adult
    as ativsengueni Schweiger, 1962
    NW Turkey, Bolu, Abant Mts.; 16mm

    Copyright © Aleš Sedláček
    Carabus (Heterocarabus) marietti marietti Cristoforis & Jan, 1837
    Carabus (Heterocarabus) marietti marietti Cristoforis & Jan, 1837
    Male adult
    as ativsengueni Schweiger, 1962
    NW Turkey, Bolu, Abant Mts.; 14mm
    Copyright © Aleš Sedláček
    Carabus (Nesaeocarabus) abbreviatus Brulle, 1835
    Carabus (Nesaeocarabus) abbreviatus Brulle, 1835
    Male adult
    Canarias Isle, NC Tenerife; 22mm
    Copyright © Aleš Sedláček
    Carabus (Carabus) arvensis venetianus Bernau, 1914
    Carabus (Carabus) arvensis venetianus Bernau, 1914
    Male adult
    N Italy, Veneto, Dolomiten Mts.; 13mm
    Copyright © Aleš Sedláček
    Carabus (Carabus) arvensis venetianus Bernau, 1914
    Carabus (Carabus) arvensis venetianus Bernau, 1914
    Female adult
    N Italy, Veneto, Dolomiten Mts.; 15mm
    Copyright © Aleš Sedláček
    Carabus (Carabus) arvensis arvensis Herbst, 1784 (as noricus Sokolar, 1910)
    Carabus (Carabus) arvensis arvensis Herbst, 1784 (as noricus Sokolar, 1910)
    Female adult
    CZ, C Moravia, Dřevohostice - Líšná, 13mm
    Copyright © Aleš Sedláček
    Carabus (Carabus) arvensis carpathus Born, 1902
    Carabus (Carabus) arvensis carpathus Born, 1902
    Male adult
    as <i>eremita </i>Fischer, 1823<br /> NW Slovakia, Malá Fatra Mts., 12mm
    Copyright © Aleš Sedláček
    Carabus (Carabus) arvensis deyrollei Gory, 1839
    Carabus (Carabus) arvensis deyrollei Gory, 1839
    Female adult
    NW Spain, Lugo env./Sarria/; 15mm
    Copyright © Aleš Sedláček
    Carabus (Carabus) granulatus interstitialis Duftschmid, 1812
    Carabus (Carabus) granulatus interstitialis Duftschmid, 1812
    Female adult
    S Austria, Kärnten, Vellach; 17mm
    Copyright © Aleš Sedláček
    Carabus (Procrustes) banoni Dejean, 1829
    Carabus (Procrustes) banoni Dejean, 1829
    Male adult
    Crete, Agia Irini (Sélinos); 31mm
    Copyright © Aleš Sedláček
    Carabus (Procrustes) banoni Dejean, 1829
    Carabus (Procrustes) banoni Dejean, 1829
    Female adult
    Crete, Agia Irini (Sélinos); 34mm
    Copyright © Aleš Sedláček
    Carabus (Macrothorax) morbillosus macilentus Lapouge, 1899
    Carabus (Macrothorax) morbillosus macilentus Lapouge, 1899
    Female adult
    SE Spain, Murcia reg., P.N. de Calblanque, 1.V.2003, T.Gazurek leg.
    L=34 mm
    Copyright © Ruslan Panin
    Carabus (Coptolabrus) ? lafossei coelestis Stewart, 1845
    Carabus (Coptolabrus) ? lafossei coelestis Stewart, 1845
    Female adult
    CE-China, Shangai, 3.V.2007 L=34 mm
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) cantabricus cantabricus Chevrolat, 1840
    Carabus (Mesocarabus) cantabricus cantabricus Chevrolat, 1840
    Endophallus
    NW-Spain, Pontevedra prov., A Cañiza viil., Alto de Fontefría h=800m
    det.Panin R.
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) cantabricus cantabricus Chevrolat, 1840 (loc. typ.)
    Carabus (Mesocarabus) cantabricus cantabricus Chevrolat, 1840 (loc. typ.)
    Male adult
    NW-Spain, Pontevedra prov., A Cañiza viil., Alto de Fontefría h=800m, 31.XII.2007, T.Gazurek leg./det.Panin R.<br /> L=27 mm
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) dufouri baguenai Breuning, 1926
    Carabus (Mesocarabus) dufouri baguenai Breuning, 1926
    Endophallus
    SE Spain, Almería Province, Carboneras env.
    det. R.Panin
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) baguenai Breuning, 1926
    Carabus (Mesocarabus) baguenai Breuning, 1926
    Male adult
    SE Spain, Almería Province, Carboneras env., 25.XI.2006, T.Gazurek leg./det. R.Panin <br /> L=26 mm
    Copyright © Ruslan Panin
    Carabus (Macrothorax) rugosus celtibericus Germar, 1824
    Carabus (Macrothorax) rugosus celtibericus Germar, 1824
    Female adult
    SW-Portugal, Setúbal Distr., Cercal do Alentejo/ Tanganheira vill. 8.XII.2006, T.Gazurek leg.
    L=31 m
    Copyright © Ruslan Panin
    Carabus (Archicarabus) steuartii Deyrolle, 1852
    Carabus (Archicarabus) steuartii Deyrolle, 1852
    Female adult
    NW-Spain, Pontevedra prov., Serra do Candán Mt range, 1.III.1992, S. Devesa leg./det. S. Sobrino
    Copyright © Ruslan Panin
    Carabus (Ctenocarabus) galicianus Gory, 1839
    Carabus (Ctenocarabus) galicianus Gory, 1839
    Female adult
    NW-Spain, Pontevedra prov., Campo Lameiro env., 15.VIII.1994, S.Sobrino leg.
    L=28 mm
    Copyright © Ruslan Panin
    Carabus (Ctenocarabus) melancholicus costatus Germar, 1824
    Carabus (Ctenocarabus) melancholicus costatus Germar, 1824
    Female adult
    C-Spain, Madrid region, Rozas de Puerto Real env.,21.XII.2008, T.Gazurek leg.
    L=27 mm
    Copyright © Ruslan Panin
    Carabus (Platycarabus) creutzeri kircheri Germar, 1824
    Carabus (Platycarabus) creutzeri kircheri Germar, 1824
    Male adult
    S Austria, Karnische Alpen; male, 24mm
    Copyright © Aleš Sedláček
    Carabus (Platycarabus) creutzeri kircheri Germar, 1824
    Carabus (Platycarabus) creutzeri kircheri Germar, 1824
    Female adult
    S Austria, Karnische Alpen; 25mm
    Copyright © Aleš Sedláček
    Carabus (Mesocarabus) macrocephalus Dejean, 1826
    Carabus (Mesocarabus) macrocephalus Dejean, 1826
    Male adult
    NW-Spain,Cantabrian Mts, Picos de Europa Mt Rng, Puerto de Panderruedas h=1460 m, 25.IX.2004,T.Gazurek leg.
    L=28 mm
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) macrocephalus Dejean, 1826
    Carabus (Mesocarabus) macrocephalus Dejean, 1826
    Endophallus
    NW-Spain,Cantabrian Mts, Picos de Europa Mt Rng, Puerto de Panderruedas h=1460 m,
    Copyright © Ruslan Panin
    Carabus (Carabus) arvensis deyrollei Gory, 1839
    Carabus (Carabus) arvensis deyrollei Gory, 1839
    Endophallus
    NW-Spain, Cantabria, Udías env., Monte Corona h=100m
    Copyright © Ruslan Panin
    Carabus (Carabus) arvensis deyrollei Gory, 1839
    Carabus (Carabus) arvensis deyrollei Gory, 1839
    Male adult
    NW-Spain,
    Cantabria, Udías env., Monte Corona h=100m,14.VI.2002, Javier de Diego leg.
    L=16 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) alpestris adamellicola Ganglbauer, 1904
    Carabus (Orinocarabus) alpestris adamellicola Ganglbauer, 1904
    Male adult
    N-Italy, Brescia prov., Bagolino, Adamello Mts, Passo del Maniva env., h=2000m, VIII.2011, M.Monzini leg.
    L=19 mm
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) latus cf. molossoides Lassalle, 1984
    Carabus (Mesocarabus) latus cf. molossoides Lassalle, 1984
    Male adult
    C-Spain, Toledo, Mts de Toledo, Sierra del Castañar Rng, Puerto del Milagro env, h=920m, 7.II.2009, T.Gazurek leg./det.R. Panin.
    L=24 mm
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) latus latus Dejean, 1826
    Carabus (Mesocarabus) latus latus Dejean, 1826
    Endophallus
    C-Spain, Toledo, Mts de Toledo, Sierra del Castañar Rng, Puerto del Milagro env, h=920m
    Copyright © Ruslan Panin
    Carabus (Pachystus) cavernosus cavernosus Frivaldsky, 1837
    Carabus (Pachystus) cavernosus cavernosus Frivaldsky, 1837
    Female adult
    W Bulgaria, Cerovo, 31 mm
    Copyright © Aleš Sedláček
    Carabus (Oreocarabus) carinthiacus Sturm, 1815
    Carabus (Oreocarabus) carinthiacus Sturm, 1815
    Female adult
    NW Slovenia, Trnovski Gozd; 24 mm
    Copyright © Aleš Sedláček
    Carabus (Orinocarabus) fairmairei fairmairei Thomson, 1875
    Carabus (Orinocarabus) fairmairei fairmairei Thomson, 1875
    Male adult
    N Italy, Alpi Graie; 20 mm
    Copyright © Aleš Sedláček
    Carabus (Carabus) granulatus granulatus (as hibernicus Lindroth, 1956)
    Carabus (Carabus) granulatus granulatus (as hibernicus Lindroth, 1956)
    Female adult
    W-Ireland, County Mayo, Letterneevoge (R312),16-30.VI.2013, J.Stypula leg.
    L=23 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) fairmairei heteromorphus Daniel, 1896
    Carabus (Orinocarabus) fairmairei heteromorphus Daniel, 1896
    Female adult
    NW-Italy, Aosta region, Champorcher env., Mt.Dondenaz, VIII.2008, V.Monzini leg.
    L=19 mm
    Copyright © Ruslan Panin
    Carabus (Macrothorax) morbillosus constantinus (as galloprovincialis Lapouge, 1910)
    Carabus (Macrothorax) morbillosus constantinus (as galloprovincialis Lapouge, 1910)
    Female adult
    SE-France, Var, near Le Muy, VIII.1980, Michon leg.
    L=31 mm
    Copyright © Ruslan Panin
    Carabus (Pachystus) hungaricus scythus Motschulsky, 1847 (loc.typ.)
    Carabus (Pachystus) hungaricus scythus Motschulsky, 1847 (loc.typ.)
    Male adult
    S-Ukraine, near Dnipropetrovsk, 6-15.IX.2013, A. Sukhenko leg.
    L=28 mm
    Copyright © Ruslan Panin
    Carabus (Acoptolabrus) gehinii gehinii Fairmaire, 1886
    Carabus (Acoptolabrus) gehinii gehinii Fairmaire, 1886
    Female adult
    N-Japan, SW-Hokkaido, Chitose env.,VI.2011.
    L=34 mm
    Copyright © Ruslan Panin
    Carabus (Macrothorax) planatus Chaudoir, 1843
    Carabus (Macrothorax) planatus Chaudoir, 1843
    Male adult
    S Italy, N Sicily, Nebrodi; 33 mm
    Copyright © Aleš Sedláček
    Carabus (Orinocarabus) fairmairei fenestrellanus Beuthin, 1899
    Carabus (Orinocarabus) fairmairei fenestrellanus Beuthin, 1899
    Male adult
    NW-Italy, Piemonte prov., Cottian Alps, Mt Albergian h=2450 m.
    L=20 mm
    Copyright © Ruslan Panin
    Carabus (Macrothorax) morbillosus morbillosus Fabricius, 1792 (as cheminorum Deuve, 1988)
    Carabus (Macrothorax) morbillosus morbillosus Fabricius, 1792 (as cheminorum Deuve, 1988)
    Female adult
    SE-France, Var, Fréjus env., 26.II.1997,
    L=31 mm
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) problematicus harcyniae (as procedens Csiki, 1927)
    Carabus (Mesocarabus) problematicus harcyniae (as procedens Csiki, 1927)
    Male adult
    W-Ireland, County Mayo, Clare Isl., 17-28.VI.2002, S. McCormack leg.<br /> L=25 mm
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) problematicus harcyniae (as procedens Csiki, 1927)
    Carabus (Mesocarabus) problematicus harcyniae (as procedens Csiki, 1927)
    Female adult
    N-Ireland, Fermanagh Distr., Lough Navar Forest, 17.VII.2013, D.Novak & J.Stypula leg.<br /> L=27mm
    Copyright © Ruslan Panin
    Carabus (Limnocarabus) clathratus clathratus (as jansoni Kraatz, 1890)
    Carabus (Limnocarabus) clathratus clathratus (as jansoni Kraatz, 1890)
    Male adult
    W-Ireland, County Mayo, Bellacorick bog, 9.VII.2013, J.Stypula leg. L=27 mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) violaceus purpurascens Fabricius, 1787 (as osismius Lapouge, 1922)
    Carabus (Megodontus) violaceus purpurascens Fabricius, 1787 (as osismius Lapouge, 1922)
    Male adult
    W-FRANCE, Brittany , Finistère, Botvarec Forest, 19.XI.2006, Pascal Stefani leg.
    L=29 mm
    Copyright © Ruslan Panin
    Carabus (Tomocarabus) convexus moreanus Reitter, 1896
    Carabus (Tomocarabus) convexus moreanus Reitter, 1896
    Male adult
    S Greece, Taygetos Mts.; 13mm
    Copyright © Aleš Sedláček
    Carabus (Tomocarabus) convexus dilatatus Dejean, 1826
    Carabus (Tomocarabus) convexus dilatatus Dejean, 1826
    Male adult
    NE Serbia, Majdanpek; 13mm
    Copyright © Aleš Sedláček
    Carabus (Tomocarabus) convexus gracilior Gehin, 1885
    Carabus (Tomocarabus) convexus gracilior Gehin, 1885
    Male adult
    SW Romania, Banat; 14mm
    Copyright © Aleš Sedláček
    Carabus (Tomocarabus) convexus gracilior Gehin, 1885
    Carabus (Tomocarabus) convexus gracilior Gehin, 1885
    Female adult
    SW Romania, Banat; 15mm
    Copyright © Aleš Sedláček
    Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825
    Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825
    Male adult
    SW Romania, Banat - Eibenthal; 29 mm
    Copyright © Aleš Sedláček
    Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825
    Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825
    Female adult
    SW Romania, Banat - Eibenthal; 33 mm
    Copyright © Aleš Sedláček
    Carabus (Eucarabus) ulrichi arrogans Schaum, 1858
    Carabus (Eucarabus) ulrichi arrogans Schaum, 1858
    Male adult
    NE Serbia, Majdanpek; 23 mm
    Copyright © Aleš Sedláček
    Carabus (Eucarabus) ulrichi arrogans Schaum, 1858
    Carabus (Eucarabus) ulrichi arrogans Schaum, 1858
    Female adult
    NE Serbia, Majdanpek; 29 mm
    Copyright © Aleš Sedláček
    Carabus (Cratocechenus) akinini akinini Morawitz, 1886 (as yiningensis )
    Carabus (Cratocechenus) akinini akinini Morawitz, 1886 (as yiningensis ) Holotype
    Female adult
    China, Kuldzha (Yining) VI.1936
    Copyright © Dmitri Obydov
    Carabus (Semnocarabus) carbonicolor callosus Semenov, 1896
    Carabus (Semnocarabus) carbonicolor callosus Semenov, 1896
    Male+Female adult
    KAZAKHSTAN,
    Bolshoe Almaatinskoe Lake,30.VI.1992/ det.Panin R.
    female 19mm
    Copyright © Ruslan Panin
    Carabus (Tachypus) cancellatus durus Reitter, 1896 (as carpathicus Kolbe, 1913)
    Carabus (Tachypus) cancellatus durus Reitter, 1896 (as carpathicus Kolbe, 1913)
    Male+Female adult
    Ukrainian Carpathians,
    Ivano-Frankivsk reg., Gorgany Mts., 20km SW Nadvorna, 22.VI-3.VIII.20Panin R. leg.
    female, 26mm
    Copyright © Ruslan Panin
    Carabus (Tachypus) cancellatus durus Reitter, 1896 (as ungensis Csiki, 1905)
    Carabus (Tachypus) cancellatus durus Reitter, 1896 (as ungensis Csiki, 1905)
    Male+Female adult
    W Ukraine,
    Transcarpathian reg., Vinogradov distr., near Malaya Kopanya vill.,7-18.VII.2008, Panin R. leg.
    female, 32mm.
    Copyright © Ruslan Panin
    Carabus (Tachypus) cancellatus durus Reitter, 1896 (? hybrid)
    Carabus (Tachypus) cancellatus durus Reitter, 1896 (? hybrid)
    Female adult
    W. UKRAINE, NE Carpathians,
    Marmarosh Mts h=450m, Rakhiv distr., near Dilove vill.,19-23.VII.2010, Panin R. leg.
    L=29 mm
    Copyright © Ruslan Panin
    Carabus (Aulonocarabus) truncaticollis kodaricola Deuve, 1998
    Carabus (Aulonocarabus) truncaticollis kodaricola Deuve, 1998
    Female adult
    SE-Siberia:N-Chita reg., Kodar Mt.Rng.,h=1800m, Leprindo vill., 1.VII.2006,
    L=17 mm
    Copyright © Ruslan Panin
    Carabus (Cratocephalus) solskyi Ballion, 1878
    Carabus (Cratocephalus) solskyi Ballion, 1878
    Female adult
    SE-KAZAKHSTAN, Ketmen Mt Rng, h=1800m, Tujuk vill.,15.V.2004
    L=36 mm
    Copyright © Ruslan Panin
    Carabus (Cratocechenus) ovtchinnikovi ovtchinnikovi Gottwald, 1987
    Carabus (Cratocechenus) ovtchinnikovi ovtchinnikovi Gottwald, 1987
    Male adult
    N-Kyrgyzstan, Kyungey Ala-Too Mt Rng, Cholpon-Ata env., h=2500m, 19.VI.2000,
    L=19 mm
    Copyright © Ruslan Panin
    Carabus (Semnocarabus) carbonicolor korolkowi Semenov, 1896
    Carabus (Semnocarabus) carbonicolor korolkowi Semenov, 1896
    Male adult
    NC-Kyrgyzstan, Kyrgyz Mt.Rng , 10 km SW of Bishkek, Chon-Aryk env, h=1100m, 24.V.1998, /det. Panin R.
    22 mm

    Copyright © Ruslan Panin
    Carabus (Semnocarabus) carbonicolor korolkowi Semenov, 1896
    Carabus (Semnocarabus) carbonicolor korolkowi Semenov, 1896
    Endophallus
    NC-Kyrgyzstan, Kyrgyz Mt.Rng , 10 km SW of Bishkek, Chon-Aryk env., h=1100m,
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) praecellens jucundus Csiki, 1906
    Carabus (Morphocarabus) praecellens jucundus Csiki, 1906
    Male adult
    N Hungary, Pilis Mts., Piliszentlelek, 20-27.VII.2011, Krause Rene leg./det. R.Panin
    L=30 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) praecellens jucundus Csiki, 1906
    Carabus (Morphocarabus) praecellens jucundus Csiki, 1906
    Endophallus
    N Hungary, Pilis Mts., Piliszentlelek,20-27.VII.2011, Krause Rene leg.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as pseudojucundus Retezár, 1974)
    Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as pseudojucundus Retezár, 1974)
    Male adult
    NW-Hungary::Komarom distr., Tardos, Gerecse Mts., 20.-27.V.2011, Krause Rene leg.
    L=26 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as pseudojucundus Retezár, 1974)
    Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as pseudojucundus Retezár, 1974)
    Endophallus
    NW-Hungary::Komarom distr., Tardos, Gerecse Mts.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as vertesensis Retezár, 1974)
    Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as vertesensis Retezár, 1974)
    Male adult
    NW Hungary: Fejer: Vértes Mt Rng, Tatabanya, 28.VII-2.VIII.1996
    L=29 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as vertesensis Retezár, 1974)
    Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as vertesensis Retezár, 1974)
    Endophallus
    NW Hungary: Fejer: Vertes Mt Rng, Tatabanya
    Copyright © Ruslan Panin
    Carabus (Coptolabrus) augustus tenuiorithorax Deuve & Li, 2006
    Carabus (Coptolabrus) augustus tenuiorithorax Deuve & Li, 2006
    Male+Female adult
    SE-China, SW-Jiangxi, Mt Guan, V.2009, local coll. female, 47 mm
    Copyright © Ruslan Panin
    Carabus (Coptolabrus) formosus bousqueti Deuve, 1993
    Carabus (Coptolabrus) formosus bousqueti Deuve, 1993
    Male adult
    C-China, S-Gansu prov., Wenxian County, Mt.Guanzigou,1-14.V.2010, local coll. leg.
    L=32 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) excellens excellens (as moldaviensis Born, 1903)
    Carabus (Morphocarabus) excellens excellens (as moldaviensis Born, 1903)
    Female adult
    E-Romania, Vaslui reg.,15 km SE Vaslui, Crasna env., V-VI,2013, M,Pasek leg. L=26 mm
    Copyright © Ruslan Panin
    Carabus (Acoptolabrus) constricticollis pihyonensis Deuve & Li, 1998
    Carabus (Acoptolabrus) constricticollis pihyonensis Deuve & Li, 1998
    Male adult
    N-Korea, North Pyongan Prov., Tongnim County, 5-25.V.2010 <br /> 29 mm
    Copyright © Ruslan Panin
    Carabus (Tribax) certus Reitter, 1896
    Carabus (Tribax) certus Reitter, 1896
    Male adult
    det. K. Makarov
    Copyright © Kirill Makarov
    Carabus (Acoptolabrus) ? leechi huisaensis Deuve & Li, 1999
    Carabus (Acoptolabrus) ? leechi huisaensis Deuve & Li, 1999
    Male adult
    NC-Korea, Ryanggang Prov., Puksubaek-san, 4-10.V.2010
    28 mm

    Copyright © Ruslan Panin
    Carabus (Carabus) arvensis hokkaidoensis Lapouge, 1924
    Carabus (Carabus) arvensis hokkaidoensis Lapouge, 1924
    Female adult

    Copyright © Kirill Makarov
    Carabus (Tachypus) vagans Olivier, 1795
    Carabus (Tachypus) vagans Olivier, 1795
    Female adult
    SE-France, Var: Le Muy env., 30.VII.2008., A. Paulian leg.
    L=24 mm
    Copyright © Ruslan Panin
    Carabus (Eurycarabus) famini piraticus (as mamorensis (Kocher, 1964)
    Carabus (Eurycarabus) famini piraticus (as mamorensis (Kocher, 1964)
    Female adult
    NW-Morocco, betw.Meknes/Kenitra, Foret de Mamora, II.2008
    L= 27 mm
    Copyright © Ruslan Panin
    Carabus (Coptolabrus) lafossei saturatus Hauser, 1913
    Carabus (Coptolabrus) lafossei saturatus Hauser, 1913
    Male adult
    SE-China, Jiangxi prov.,Yushan County, Mt Wangjing, VI.2012
    40 мм
    Copyright © Ruslan Panin
    Carabus (Chrysocarabus) lineatus lineatus (as pseudonumancianus Mollard, 2010)
    Carabus (Chrysocarabus) lineatus lineatus (as pseudonumancianus Mollard, 2010)
    Male adult
    NC-Spain, Sierra de Cantabria , Sierra de Toloño Rng, Puerto de Herrera , h=1100m, 19.I,2007 25 mm
    Copyright © Ruslan Panin
    Carabus (Coptolabrus) pustulifer (as cyanelongatus Deuve, 2006)
    Carabus (Coptolabrus) pustulifer (as cyanelongatus Deuve, 2006)
    Male adult
    C-Chine, Tibet, Qamdo, Gonjo County, Ke Ri Xiang env., VI.2012, Alain Mollard leg.<br /> L=43 mm
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) lusitanicus helluo (as albarracinus Ganglbauer, 1886)
    Carabus (Mesocarabus) lusitanicus helluo (as albarracinus Ganglbauer, 1886)
    Male adult
    CE-Spain, Provincia Teruel, Sierra de Javalón Rng, El Cañigral env., h=1460m, X.2010, A. Mollard leg.<br /> L=23 mm
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) lusitanicus helluo (as albarracinus Ganglbauer, 1886)
    Carabus (Mesocarabus) lusitanicus helluo (as albarracinus Ganglbauer, 1886)
    Endophallus
    CE-Spain, Provincia Teruel, Sierra de Javalón Rng, El Cañigral env., h=1460m,
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826
    Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826
    Male adult
    CW-Spain: Salamanca, Brincones env., h=750m, X.2003<br /> L=25 mm
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826
    Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826
    Endophallus
    CW-Spain:Salamanca, Brincones env., h=750m
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) davidis davidis Deyrolle & Fairmaire, 1878
    Carabus (Apotomopterus) davidis davidis Deyrolle & Fairmaire, 1878
    Male adult
    SE-China, Jiangxi prov., Yushan County, Mt Wangjing, VI,2012
    L=38 mm
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) problematicus harcyniae (as belgicus Lapouge, 1913)
    Carabus (Mesocarabus) problematicus harcyniae (as belgicus Lapouge, 1913)
    Male adult
    Luxembourg, Bollendorf-Pont env., 3.III.1999 /det.R Panin
    L=24 mm
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) problematicus harcyniae (as belgicus Lapouge, 1913)
    Carabus (Mesocarabus) problematicus harcyniae (as belgicus Lapouge, 1913)
    Endophallus
    Luxembourg, Bollendorf-Pont env., 3.III.1999
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) (? problematicus trans lusitanicus) planiusculus (as xaxarsi Born, 1917)
    Carabus (Mesocarabus) (? problematicus trans lusitanicus) planiusculus (as xaxarsi Born, 1917)
    Male adult
    NE Spain, Catalonia, Girona Prov., San Hilario Sacalm env., 20.II.1988, A. Mollard leg.<br /> L=25 m
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) (? problematicus trans lusitanicus) planiusculus (as xaxarsi Born, 1917)
    Carabus (Mesocarabus) (? problematicus trans lusitanicus) planiusculus (as xaxarsi Born, 1917)
    Endophallus
    NE Spain, Catalonia,Girona Prov., San Hilario Sacalm env.<br />
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) bertolinii Kraatz, 1878
    Carabus (Orinocarabus) bertolinii Kraatz, 1878
    Male adult
    NE-Italy, Dolomites-Alps, Trentino prov., Rolle Pass env., Malga Fosse h=2000m, 8-28.VI.2013, Manfred Egger leg.
    L=19 mm
    Copyright © Ruslan Panin
    Carabus (Coptolabrus) formosus brockmanni Kleinfeld & Schutze, 1997
    Carabus (Coptolabrus) formosus brockmanni Kleinfeld & Schutze, 1997
    Female adult
    C-China, S-Shaanxi: Shangzhou City, Shahezi,1-14.VI.,2013, local coll. leg.
    L=39 mm
    Copyright © Ruslan Panin
    Carabus (Coptolabrus) smaragdinus dolichognathus Deuve, 1995
    Carabus (Coptolabrus) smaragdinus dolichognathus Deuve, 1995
    Male adult
    NW-Korea, North Pyongan Province, Mt. Donggorim, 1-20.VI.2013, local coll. leg.<br /> L=37 mm
    Copyright © Ruslan Panin
    Carabus (Coptolabrus) augustus augustus Bates, 1888
    Carabus (Coptolabrus) augustus augustus Bates, 1888
    Male adult
    SE-China, Jiangxi prov., Yudu County, Ganzhou city, Yun Fengzang h=1000m, 5-20.V.2011
    L=43 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) alpestris dolomitanus Mandl, 1956
    Carabus (Orinocarabus) alpestris dolomitanus Mandl, 1956
    Male adult
    NE-Italy, South Tyrol, Dolomites-Alps, 500 m left of Würzjoch, Gömaalm - at the foot Mt.Peitlerkofel h=2100m, 28.VII-17.VIII.2013, Manfred Egger leg.
    l=18 mm
    Copyright © Ruslan Panin
    Carabus (Archicarabus) monticola roccae Born, 1923
    Carabus (Archicarabus) monticola roccae Born, 1923
    Male adult
    NW-Italy, Liguria:Savona env., Monte San Giorgio, 30.X.1977, Monauzzi leg.
    L=23 mm

    Copyright © Ruslan Panin
    Carabus (Orinocarabus) linnaei linnaei (as scopolii Dejean, 1826)
    Carabus (Orinocarabus) linnaei linnaei (as scopolii Dejean, 1826)
    Female adult
    W Ukraine, NW-Podolian, near Lviv city, Vynnykivskyi forest, 20.IV.-13.V.2011, R. Panin leg.
    L=22 mm
    Copyright © Ruslan Panin
    Carabus (Tribax) circassicus circassicus (as tshchaltensis Novotný & Voříšek
    Carabus (Tribax) circassicus circassicus (as tshchaltensis Novotný & Voříšek
    Male adult
    W-Caucasus: Abkhasia, Gudauta distr., E slope Mt Anchkho h=2200-2350 m, 14-20.VII.2011, S.Pavluchuk, I. Solodovnikov leg./det. I. A. Solodovnikov.
    25 mm

    Copyright © Ruslan Panin
    Carabus (Rhigocarabus) shaluishan Kleinfeld & Puchner, 2011
    Carabus (Rhigocarabus) shaluishan Kleinfeld & Puchner, 2011 Holotype
    Male adult
    CHINA, Prov. Sichuan, zwischen Garzê und Baiyü, Berg 25 km südlich von Garzê, Nordseite, 3820m, - 31°31.985 N / /99°58.670´ E, 30.V-14.VI.2010, leg. et in coll. Puchner.
    19 mm
    Foto Kleinfeld XII.2010
    Copyright © Frank Kleinfeld
    Carabus (Rhigocarabus) shaluishan Kleinfeld & Puchner, 2011
    Carabus (Rhigocarabus) shaluishan Kleinfeld & Puchner, 2011 Paratype
    Female adult
    CHINA, Prov. Sichuan, zwischen Garzê und Baiyü, Berg 25 km südlich von Garzê, Nordseite, 3820m, - 31°31.985 N / /99°58.670´ E, 30.V-14.VI.2010, leg. et in coll. Puchner.
    18 mm Foto Kleinfeld XII.2010
    Copyright © Frank Kleinfeld
    Carabus (Calocarabus) worahniki Kleinfeld & Puchner, 2011
    Carabus (Calocarabus) worahniki Kleinfeld & Puchner, 2011 Holotype
    Male adult
    CHINA, Prov. Sichuan, zwischen Garzê und Baiyü, Berg 25km südlich von Garzê, Nordseite, 3780m, /31°32.611N/099°58.596´E, 30.V-14.VI.2010 - leg. et in coll. Puchner
    21 mm
    Foto Kleinfeld XI.2010
    Copyright © Frank Kleinfeld
    Carabus (Calocarabus) miraculix Kleinfeld & Puchner, 2011
    Carabus (Calocarabus) miraculix Kleinfeld & Puchner, 2011 Holotype
    Male adult
    CHINA, Prov. Sichuan, zwischen Garzê und Baiyü, Berg 25 km südlich von Garzê, Nordseite, 4100m, leg. Puchner 30.V-14.VI.2010 - N 31°28.298´ E 099°59.930´
    24 mm
    Foto Kleinfeld XI.2010
    Copyright © Frank Kleinfeld
    Carabus (Pagocarabus) alexpuchneri Kleinfeld & Puchner, 2011
    Carabus (Pagocarabus) alexpuchneri Kleinfeld & Puchner, 2011 Holotype
    Male adult
    China, Sichuan, Straße Xinlong - Litang, Umgebung Heping (30°33'/100°20'), 2870m, 1.-16.VI.2010, leg. et in coll. Puchner
    22,5 mm
    Copyright © Frank Kleinfeld
    Carabus (Pagocarabus) alexpuchneri Kleinfeld & Puchner, 2011
    Carabus (Pagocarabus) alexpuchneri Kleinfeld & Puchner, 2011 Paratype
    Female adult
    China, Sichuan, Straße Xinlong - Litang, Umgebung Heping (30°33'/100°20'), 2870m, 1.-16.VI.2010, leg. et in coll. Puchner
    24 mm
    Copyright © Frank Kleinfeld
    Carabus (Leptocarabus) arboreus arboreus Lewis, 1882
    Carabus (Leptocarabus) arboreus arboreus Lewis, 1882
    Male adult
    N-Japan, SW-Hokkaido, Chitose sity, 25.V-8.VI.2013, T. Hirabayashi leg.<br /> 32 mm
    Copyright © Ruslan Panin
    Carabus (Nesaeocarabus) faustus cabrerai Enderlein, 1929
    Carabus (Nesaeocarabus) faustus cabrerai Enderlein, 1929
    Male adult
    Spain, Canarias Isl.,Teneriffe: NW-part:Teno Massif, Erjos env., h=1000m, 18-21.III.2013, V.Slovak leg.
    L=23 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi rothi (as equistriatus Kraatz, 1878)
    Carabus (Morphocarabus) rothi rothi (as equistriatus Kraatz, 1878)
    Female adult
    C Romania, Sibiu reg., left bank Olt River, Avrig env., 12.VI.2013, V. Slovak leg.
    L=30 mm
    Copyright © Ruslan Panin
    Carabus (Aulonocarabus) kurilensis ? daisetsuzanus Kono, 1936
    Carabus (Aulonocarabus) kurilensis ? daisetsuzanus Kono, 1936
    Male adult
    N-Japan, Hokkaido, Katō District,Kamishihoro, Horoka near Maruyama Bridge,19.VI.2013, T. Hirabayashi leg.
    L=24 mm
    Copyright © Ruslan Panin
    Carabus (Aulonocarabus) kurilensis ? daisetsuzanus Kono, 1936
    Carabus (Aulonocarabus) kurilensis ? daisetsuzanus Kono, 1936
    Endophallus
    N-Japan, Hokkaido, Katō District, Kamishihoro, Horoka near Maruyama Bridge,
    Copyright © Ruslan Panin
    Carabus (Tribax) agnatus pseudagnatus Novotný & Voříšek, 1988
    Carabus (Tribax) agnatus pseudagnatus Novotný & Voříšek, 1988
    Female adult
    NW-Caucasus, Karachay-Cherkessia, Bol'shaya Marka River h=2500-2600m, 12.VIII.2010, I.A. Solodovnikov leg./det.
    Copyright © Ruslan Panin
    Carabus (Tribax) agnatus pseudagnatus Novotný & Voříšek, 1988
    Carabus (Tribax) agnatus pseudagnatus Novotný & Voříšek, 1988
    Male adult
    W-Caucasus: Abkhasia, Gudauta distr., Mt Anchkho h=2200-2350 m, 14-20.VII.2011, S.Pavluchuk, I. Solodovnikov leg./det. I. A. Solodovnikov.
    L=25 mm
    Copyright © Ruslan Panin
    Carabus (Tribax) agnatus pseudagnatus Novotný & Voříšek, 1988
    Carabus (Tribax) agnatus pseudagnatus Novotný & Voříšek, 1988
    Endophallus
    W-Caucasus: Abkhasia, Gudauta distr., Mt Anchkho h=2200-2350 m,
    Copyright © Ruslan Panin
    Carabus (Tribax) agnatus agnatus Ganglbauer, 1889
    Carabus (Tribax) agnatus agnatus Ganglbauer, 1889
    Male adult

    Copyright © Kirill Makarov
    Carabus (Neoplectes) edithae exedithae Gottwald, 1985
    Carabus (Neoplectes) edithae exedithae Gottwald, 1985
    Male adult
    NС-Caucasus, Karachaevo-Cherkessia, Abishir-Akhuba Mt.Range h=2250-2600 m, right bank Pkhiya river, 10-13.VI.2013, S. Pavluchuk leg.
    L=25 mm
    Copyright © Ruslan Panin
    Carabus (Neoplectes) edithae exedithae Gottwald, 1985
    Carabus (Neoplectes) edithae exedithae Gottwald, 1985
    Endophallus
    NС-Caucasus, Karachaevo-Cherkessia, right bank Pkhiya river, Abishir-Akhuba Mt.Range h=2250-2600 m,
    Copyright © Ruslan Panin
    Carabus (Oreocarabus) taedatus bicanaliceps Casey, 1920
    Carabus (Oreocarabus) taedatus bicanaliceps Casey, 1920
    Male adult
    NW-USA, Washington, Thurston County, Mima Mound Prairie, 15.IX.2005, A. Laforgue leg. L=23 мм
    Copyright © Ruslan Panin
    Carabus (Eucarabus) catenulatus fontanellae Reitter, 1896
    Carabus (Eucarabus) catenulatus fontanellae Reitter, 1896
    Male adult
    W-Croatia, Cres island,VIII.1994, G. Zannna leg. L=33 mm
    Copyright © Ruslan Panin
    Carabus (Eucarabus) cartereti omisanensis Lassalle, 1999
    Carabus (Eucarabus) cartereti omisanensis Lassalle, 1999
    Male adult
    South Korea, Gangwon Province, Kyebang-san, h=1100m, VI.1998 L=24 mm
    Copyright © Ruslan Panin
    Carabus (Sphodristocarabus) separatus persa Lapouge, 1924
    Carabus (Sphodristocarabus) separatus persa Lapouge, 1924
    Endophallus
    N-Iran: Mazandaran Province, Elborz Mt.Rng, Vandarbon env.
    Copyright © Ruslan Panin
    Carabus (Sphodristocarabus) separatus persa Lapouge, 1924
    Carabus (Sphodristocarabus) separatus persa Lapouge, 1924
    Male adult
    N-Iran: Mazandaran Province, Elborz Mt.Rng, Vandarbon env., 10.VII.2002 L=23 mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) caelatus dalmatinus Duftschmid, 1812
    Carabus (Megodontus) caelatus dalmatinus Duftschmid, 1812
    Female adult
    Bosnia, Mostar; 40 mm
    Copyright © Aleš Sedláček
    Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823
    Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823
    Male adult
    W-Ukraine, Podolian Tovtry, Ternopol reg., Pidvolochysk distr., between Chagari-Zbarazhki & Shevchenkove vill., steppe sites, 6.V-7.VI.2012, Panin R. leg. L=25 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi comptus (as borlovensis Lie, 1994)
    Carabus (Morphocarabus) rothi comptus (as borlovensis Lie, 1994)
    Endophallus
    SW Romania,, Banat, Borlova env.,
    Copyright © Ruslan Panin
    Carabus (Aulonocarabus) gaschkewitschi gaschkewitschi Motschulsky, 1859
    Carabus (Aulonocarabus) gaschkewitschi gaschkewitschi Motschulsky, 1859
    Male adult

    Copyright © Kirill Makarov
    Carabus (Carabus) granulatus yezoensis Bates, 1883
    Carabus (Carabus) granulatus yezoensis Bates, 1883
    Male adult
    det. K. Makarov
    Copyright © Kirill Makarov
    Carabus (Carabus) menetriesi menetriesi Hummel, 1827
    Carabus (Carabus) menetriesi menetriesi Hummel, 1827
    Male adult
    det. K. Makarov
    Copyright © Kirill Makarov
    Carabus (Diocarabus) aurocinctus Motschulsky, 1846
    Carabus (Diocarabus) aurocinctus Motschulsky, 1846
    Female adult
    det. K. Makarov
    Copyright © Alexander Anichtchenko
    Carabus (Diocarabus) opaculus opaculus Putzeys, 1875
    Carabus (Diocarabus) opaculus opaculus Putzeys, 1875
    Female adult

    Copyright © Kirill Makarov
    Carabus (Diocarabus) opaculus opaculus Putzeys, 1875
    Carabus (Diocarabus) opaculus opaculus Putzeys, 1875
    Male adult

    Copyright © Kirill Makarov
    Carabus (Leptocarabus) arboreus pararboreus (Ishikawa, 1992)
    Carabus (Leptocarabus) arboreus pararboreus (Ishikawa, 1992)
    Male adult
    det. K. Makarov
    Copyright © Kirill Makarov
    Carabus (Leptocarabus) arboreus pararboreus (Ishikawa, 1992)
    Carabus (Leptocarabus) arboreus pararboreus (Ishikawa, 1992)
    Female adult
    det. K. Makarov
    Copyright © Kirill Makarov
    Carabus (Morphocarabus) henningi henningi Fischer, 1817
    Carabus (Morphocarabus) henningi henningi Fischer, 1817
    Male adult

    Copyright © Kirill Makarov
    Carabus (Morphocarabus) odoratus magadanicus Obydov 1999: 614
    Carabus (Morphocarabus) odoratus magadanicus Obydov 1999: 614
    Female adult
    det. K. Makarov
    Copyright © Kirill Makarov
    Carabus (Morphocarabus) odoratus magadanicus Obydov 1999: 614
    Carabus (Morphocarabus) odoratus magadanicus Obydov 1999: 614
    Male adult
    det. K. Makarov
    Copyright © Kirill Makarov
    Carabus (Morphocarabus) venustus venustus Morawitz, 1862
    Carabus (Morphocarabus) venustus venustus Morawitz, 1862
    Male adult

    Copyright © Kirill Makarov
    Carabus (Procrustes) talyschensis Menetries, 1832
    Carabus (Procrustes) talyschensis Menetries, 1832
    Female adult
    det. K. Makarov
    Copyright © Alexander Anichtchenko
    Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
    Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
    Endophallus
    W Ukraine, Transcarpathian Reg., 4 km NE Chop, forest Peresh.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) brinevi Kabak, 2014
    Carabus (Morphocarabus) brinevi Kabak, 2014 Paratype
    Male adult
    China, Xinjiang, S Altai Mts, Keshtau Mt., ENE Qinghe, 46°47'21''N / 90°44'42''E, 2730 m, 12.08.2013, I.I. Kabak leg.
    Copyright © Ilya Kabak
    Carabus (Morphocarabus) brinevi Kabak, 2014
    Carabus (Morphocarabus) brinevi Kabak, 2014 Holotype
    Male adult
    China, Xinjiang, S Altai Mts, Keshtau Mt., ENE Qinghe, 46°47'21''N / 90°44'42''E, 2730 m, 12.08.2013, I.I. Kabak leg.
    Copyright © Ilya Kabak
    Carabus (Morphocarabus) brinevi Kabak, 2014
    Carabus (Morphocarabus) brinevi Kabak, 2014 Paratype
    Endophallus

    Copyright © Ilya Kabak
    Carabus (Ophiocarabus) nilkiensis Kabak, 2014
    Carabus (Ophiocarabus) nilkiensis Kabak, 2014 Paratype
    Male adult
    China, Xinjiang, Boro-Horo Mt. R., S slope, Nilki River, 44°01'25''N / 82°46'00''E, 2030 m, 5.08.2013, I.I. Kabak leg.
    Copyright © Ilya Kabak
    Carabus (Ophiocarabus) nilkiensis Kabak, 2014
    Carabus (Ophiocarabus) nilkiensis Kabak, 2014 Paratype
    Female adult
    China, Xinjiang, Boro-Horo Mt. R., S slope, Nilki River, 44°01'25''N / 82°46'00''E, 2030 m, 5.08.2013, I.I. Kabak leg.
    Copyright © Ilya Kabak
    Carabus (Ophiocarabus) nilkiensis Kabak, 2014
    Carabus (Ophiocarabus) nilkiensis Kabak, 2014 Paratype
    Endophallus

    Copyright © Ilya Kabak
    Carabus (Chrysocarabus) auronitens escheri Palliardi, 1825 (as laevipennis Seidlitz, 1891)
    Carabus (Chrysocarabus) auronitens escheri Palliardi, 1825 (as laevipennis Seidlitz, 1891)
    Male adult
    W Ukraine, NE Carpathians,Transcarpathian Region, Rakhiv Distr., Chornogora Mts Range, h=2000 m, alpin zone, V-28.VI..2014, Panin R. leg.
    Copyright © Ruslan Panin
    Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
    Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
    Female adult
    W Ukraine, NE Carpathians, Transcarpathian Region, Chornogora Mt Range, h=2000 m, alpin zone, 28.VII-11.IX.2014, Panin R. leg. L=22 mm
    Copyright © Ruslan Panin
    Carabus (Neoplectes) reitteri reitteri Retowski, 1885
    Carabus (Neoplectes) reitteri reitteri Retowski, 1885
    Female adult
    det. K. Makarov
    Copyright © Kirill Makarov
    Carabus (Procrustes) paiafa acuticollis Motschulsky, 1858
    Carabus (Procrustes) paiafa acuticollis Motschulsky, 1858
    Male adult
    SC-Turkey: Alania (Irmasan), rte 695 au km 25 Akseki, h= 1225 m, 02.X.2003
    Copyright © Igor Solodovnikov
    Carabus (Imaibius) wittmerorum Heinertz, 1978
    Carabus (Imaibius) wittmerorum Heinertz, 1978
    Male adult
    N. Pakistan: Frontier prov., Miandam, Swat vall., h=1800-2600 m, leg.S.Remond (TOPOTYPUS)
    Copyright © Igor Solodovnikov
    Carabus (Imaibius) barysomus hazarensis Deuve, 1982 (TOPOTYP)
    Carabus (Imaibius) barysomus hazarensis Deuve, 1982 (TOPOTYP)
    Male adult
    N-Pakistan: Mansehra prov., Hazara, Kaghan valley: Shocgran env., h = 2750 m L= 42 mm
    Copyright © Igor Solodovnikov
    Carabus (Damaster) blaptoides babaianus Ishikawa, 1983
    Carabus (Damaster) blaptoides babaianus Ishikawa, 1983
    Male adult
    Japan: C Honshu: Niigata pref., Mysato: Sakae Abiko-City; Iwafune distr., Awashima Isl. (LABEL ERROR)
    Copyright © Igor Solodovnikov
    Carabus (Damaster) blaptoides babaianus Ishikawa, 1983
    Carabus (Damaster) blaptoides babaianus Ishikawa, 1983
    Female adult
    Japan: C Honshu: Niigata pref., Mysato: Sakae Abiko-City; Iwafune distr., Awashima Isl. (LABEL ERROR)
    Copyright © Igor Solodovnikov
    Carabus (Damaster) blaptoides capito Lewis, 1881
    Carabus (Damaster) blaptoides capito Lewis, 1881
    Male adult
    Japan: Niigato pref., Sado Isl., Sado, Yamato
    Copyright © Igor Solodovnikov
    Carabus (Damaster) blaptoides capito Lewis, 1881
    Carabus (Damaster) blaptoides capito Lewis, 1881
    Male adult
    Japan: Niigato pref., Sado Isl., Sado, Yamato
    Copyright © Igor Solodovnikov
    Carabus (Imaibius) wittmerorum Heinertz, 1978
    Carabus (Imaibius) wittmerorum Heinertz, 1978
    Male adult
    N. Pakistan: Frontier prov., Miandam, Swat vall., h=1800-2600 m, leg. S.Remond (TOPOTYPUS)
    Copyright © Igor Solodovnikov
    Carabus (Megodontus) bonvouloiri casalei Cavazzuti, 1986
    Carabus (Megodontus) bonvouloiri casalei Cavazzuti, 1986
    Male adult
    NE-Turkey: Artvin: Kabaca, env. Murgul, h = 1000 m, VII.1991, leg. St. Il. Remond
    Copyright © Igor Solodovnikov
    Carabus (Megodontus) bonvouloiri casalei Cavazzuti, 1986
    Carabus (Megodontus) bonvouloiri casalei Cavazzuti, 1986
    Male adult
    NE-Turkey: Artvin: Kafkäsor, h = 1700 m, VII.2000, leg. St. Il. Remond 32,5 mm
    Copyright © Igor Solodovnikov
    Carabus (Megodontus) bonvouloiri graciliformis Breuning, 1964
    Carabus (Megodontus) bonvouloiri graciliformis Breuning, 1964
    Female adult
    NE-Turkey: Ordu: Sud Gölköy env., Harçbeli pass, h= 1500 m, VI.1999, leg. St. Il. Remond 40,5 mm
    Copyright © Igor Solodovnikov
    Carabus (Megodontus) bonvouloiri graciliformis Breuning, 1964
    Carabus (Megodontus) bonvouloiri graciliformis Breuning, 1964
    Male adult
    NE-Turkey: Ordu: Sud Gölköy env., Harçbeli pass, h= 1400 m, VI.1999, leg. St. Il. Remond
    Copyright © Igor Solodovnikov
    Carabus (Megodontus) bonvouloiri bonvouloiri Chaudoir, 1863
    Carabus (Megodontus) bonvouloiri bonvouloiri Chaudoir, 1863
    Male adult
    NE-Turkey: Rize: Ovitdagi Geç, h = 1700 m, VII.2000, leg. St. Il. Remond (as bonvouloiri kaschkarensis Blumenthal & Breuning, 1967; un)
    Copyright © Igor Solodovnikov
    Carabus (Procerus) scabrosus bureschianus Breuning, 1928
    Carabus (Procerus) scabrosus bureschianus Breuning, 1928
    Male adult
    Bulgaria, W Rodopi, Dobrostan masive, backovo env. M.Janalik leg.
    Copyright © Igor Solodovnikov
    Carabus (Procerus) scabrosus colchicus Motschulsky, 1844
    Carabus (Procerus) scabrosus colchicus Motschulsky, 1844
    Male adult
    Abkhasia: S part of Bzybian Mt.R. vall. riv. Khipsta, h= 600 m, VIII.2014, leg. I.A. Solodovnikov
    Copyright © Igor Solodovnikov
    Carabus (Procerus) scabrosus colchicus Motschulsky, 1844
    Carabus (Procerus) scabrosus colchicus Motschulsky, 1844
    Male adult
    Abkhasia: S part of Bzybian Mt.R. vall. riv. Khipsta, h= 600 m, VIII.2014, leg. I.A. Solodovnikov, 58 mm
    Copyright © Igor Solodovnikov
    Carabus (Procerus) scabrosus schuberti Breuning, 1968
    Carabus (Procerus) scabrosus schuberti Breuning, 1968
    Male adult
    NC-Turkey: 10 km E of Sinop, h = 0 m, VI.1999, leg. St. Remond, 42 mm
    Copyright © Igor Solodovnikov
    Carabus (Procerus) scabrosus tauricus Bonelli, 1811
    Carabus (Procerus) scabrosus tauricus Bonelli, 1811
    Male adult
    f. virens Crimea, Kazu-Kau Mt. (South part Chatyr-Dagh Mt.) 14.06.2009. Leg V.V.Savchuk, male. 41 mm
    Copyright © Igor Solodovnikov
    Carabus (Procerus) scabrosus tauricus Bonelli, 1811
    Carabus (Procerus) scabrosus tauricus Bonelli, 1811
    Male adult
    f. purpureus Kraatz Crimea, Inkerman, steppe, 26.03.1987. Leg I.M. Lutzky. 42 mm
    Copyright © Igor Solodovnikov
    Carabus (Ophiocarabus) successor koeksuensis Kabak, 2013
    Carabus (Ophiocarabus) successor koeksuensis Kabak, 2013 Paratype
    Female adult
    China, Xinjiang, Koeksu Bas. SSE Tekes h~3330-3550 11.VII.2012 leg. I.Kabak L=16.5 mm
    Copyright © Kirill Makarov
    Carabus (Ophiocarabus) successor koeksuensis Kabak, 2013
    Carabus (Ophiocarabus) successor koeksuensis Kabak, 2013 Paratype
    Male adult
    China, Xinjiang, Koeksu Bas. SSE Tekes h~3330-3550 11.VII.2012 leg. I.Kabak L=16.0 mm
    Copyright © Kirill Makarov
    Carabus (Cathoplius) asperatus asperatus (Dejean, 1826)
    Carabus (Cathoplius) asperatus asperatus (Dejean, 1826)
    Male adult
    Morocco, 4 km S Saft 32°10'28"N 009°15'27"E 20-21.III.2011 leg. S.Gusakov L=24.0 mm
    Copyright © Kirill Makarov
    Carabus (Oxycarabus) saphyrinus Cristoforis & Jan, 1837
    Carabus (Oxycarabus) saphyrinus Cristoforis & Jan, 1837
    Female adult
    Turkey, NE Giresun, Dereli env., h~400 40°43'01"N 38°28'20"E 12.IX.2013 leg. B.Saratovskikh L=28.0 mm
    Copyright © Kirill Makarov
    Carabus (Oxycarabus) saphyrinus Cristoforis & Jan, 1837
    Carabus (Oxycarabus) saphyrinus Cristoforis & Jan, 1837
    Male adult
    Turkey, NE Giresun, Dereli env., h~400 40°43'01"N 38°28'20"E 12.IX.2013 leg. B.Saratovskikh L=27.0 mm
    Copyright © Kirill Makarov
    Carabus (Orinocarabus) fairmairei stecki Born, 1902
    Carabus (Orinocarabus) fairmairei stecki Born, 1902
    Female adult
    France, Hautes-Alpes, env. Lake Foreant, near Col de D'Agnel, ca 2700-2800 m 17.VIII.2013 leg. B.Kataev L=18.0 mm
    Copyright © Kirill Makarov
    Carabus (Orinocarabus) fairmairei stecki Born, 1902
    Carabus (Orinocarabus) fairmairei stecki Born, 1902
    Male adult
    France, Hautes-Alpes, env. Lake Foreant, near Col de D'Agnel, ca 2700-2800 m 17.VIII.2013 leg. B.Kataev L=16.5 mm
    Copyright © Kirill Makarov
    Carabus (Morphocarabus) brinevi Kabak, 2014
    Carabus (Morphocarabus) brinevi Kabak, 2014 Paratype
    Female adult
    China, Xinjiang, S Altai Mts., Keshtau Mt., ENE of Qinghe, h~2730 49°47'21"N 90°44'42"E 12.VIII.2013 leg. I.Kabak L=19.5 mm
    Copyright © Kirill Makarov
    Carabus (Apotomopterus) ngi ngi Deuve, 1994
    Carabus (Apotomopterus) ngi ngi Deuve, 1994 Holotype
    Female adult
    in coll Lassalle
    Copyright © Bernard Lassalle
    Carabus (Pachystus) cribratus echarouxi Lassalle, 1996
    Carabus (Pachystus) cribratus echarouxi Lassalle, 1996 Holotype
    Male adult
    in coll Lassalle
    Copyright © Bernard Lassalle
    Carabus (Coptolabrus) augustus ignigena (as subpustulatus Breuuning 1961)
    Carabus (Coptolabrus) augustus ignigena (as subpustulatus Breuuning 1961) Holotype
    Male adult
    Hunan, China. ignigena subpustulatus (In coll Lassalle).
    Copyright © Bernard Lassalle
    Carabus (Coptolabrus) augustus adriaenssensi Lassalle & Prunier, 1993
    Carabus (Coptolabrus) augustus adriaenssensi Lassalle & Prunier, 1993 Holotype
    Male adult
    Chine, Guangxi, Mts Miao Er Shan, 1800m, 5.92 (in coll B. Lassalle) ? pustulifer adriaenssensi Lassalle & Prunier, 1993
    Copyright © Bernard Lassalle
    Carabus (Megodontus) stroganowi galugahensis Lassalle 2001
    Carabus (Megodontus) stroganowi galugahensis Lassalle 2001 Holotype
    Male adult
    Iran, Mazanderan, Mts 50km sud de Galugah, 5.2000 (in coll Lassalle)
    Copyright © Bernard Lassalle
    Carabus (Hemicarabus) tuberculosus Dejean, 1829
    Carabus (Hemicarabus) tuberculosus Dejean, 1829
    Male adult
    Иркутск, Новоленино, 20 июня 1985, leg. Э.Берлов
    Copyright © Oleg Berlov
    Carabus (Cryptocarabus) ? turkestanus turkestanus Breuning, 1929
    Carabus (Cryptocarabus) ? turkestanus turkestanus Breuning, 1929
    Male adult
    Kyrgyzstan, W Kungei Ala-Too Mt Rng, Chon-Kemin riv., Kyzyl-Bairak vill., 25.IV.2010. L= 18 mm
    Copyright © Ruslan Panin
    Carabus (Aulonocarabus) kolymensis kodarensis Brinev, 1997
    Carabus (Aulonocarabus) kolymensis kodarensis Brinev, 1997
    Male adult
    Russia, SE Siberia, N Chita reg., Kodar Mt. Rng. h=1800m, Novaya Chara vill., 23.VI.2006. L=17 mm
    Copyright © Ruslan Panin
    Carabus (Mimocarabus) pumilio Kuster, 1846
    Carabus (Mimocarabus) pumilio Kuster, 1846
    Endophallus
    S-Armenia, Vayots Dzor Region, E Ughedzor vill., h=2630 m
    Copyright © Ruslan Panin
    Carabus (Mimocarabus) pumilio Kuster, 1846
    Carabus (Mimocarabus) pumilio Kuster, 1846
    Male adult
    S-Armenia, Vayots Dzor Region, E Ughedzor vill., h=2630 m, 9.VI.2014, L=16 mm
    Copyright © Ruslan Panin
    Carabus (Pachycarabus) imitator imitator Reitter, 1883
    Carabus (Pachycarabus) imitator imitator Reitter, 1883
    Female adult
    det. K. Makarov
    Copyright © Kirill Makarov
    Carabus (Pachycarabus) staehlini Adams, 1817
    Carabus (Pachycarabus) staehlini Adams, 1817
    Endophallus
    S Russia, E Caucasus, Dagestan, Nukatl Mt Rng h=2600m,
    Copyright © Ruslan Panin
    Carabus (Neoplectes) satyrus satyrus Kurnakov, 1962
    Carabus (Neoplectes) satyrus satyrus Kurnakov, 1962
    Male+Female adult
    Abkhazia, Gulripsh Distr., NW slopes of Gurzul Mt. Range near Merkheul Village, 170-230 m, left tributary of Machara River, hornbeam, alder, rhododendron (Rhododendron ponticum) forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) satyrus satyrus Kurnakov, 1962
    Carabus (Neoplectes) satyrus satyrus Kurnakov, 1962
    Female adult
    Abkhazia, Gulripsh Distr., NW slopes of Gurzul Mt. Range near Merkheul Village, 170-230 m, left tributary of Machara River, hornbeam, alder, rhododendron (Rhododendron ponticum) forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) satyrus satyrus Kurnakov, 1962
    Carabus (Neoplectes) satyrus satyrus Kurnakov, 1962
    Male adult
    Abkhazia, Gulripsh Distr., NW slopes of Gurzul Mt. Range near Merkheul Village, 170-230 m, left tributary of Machara River, hornbeam, alder, rhododendron (Rhododendron ponticum) forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) satyrus satyrus Kurnakov, 1962
    Carabus (Neoplectes) satyrus satyrus Kurnakov, 1962
    Male adult
    Abkhazia, Gulripsh Distr., NW slopes of Gurzul Mt. Range near Merkheul Village, 170-230 m, left tributary of Machara River, hornbeam, alder, rhododendron (Rhododendron ponticum) forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) edithae Reitter, 1893
    Carabus (Neoplectes) edithae Reitter, 1893
    Female adult
    det. K. Makarov
    Copyright © Kirill Makarov
    Carabus (Pachycarabus) staehlini Adams, 1817
    Carabus (Pachycarabus) staehlini Adams, 1817
    Male adult
    S Russia, E-Caucasus, Dagestan, Nukatl Mt Rng h=2600m, 25.VII.1997, V. Savizky leg. L=21 mm
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) lusitanicus helluo Dejean, 1826
    Carabus (Mesocarabus) lusitanicus helluo Dejean, 1826
    Female adult
    SE-Spain, Prov.Alicante, Port de la Carrasqueta, 26.XII.1987, M.Toribio leg. L=25 mm
    Copyright © Ruslan Panin
    Carabus (Imaibius) boysi boysi Tatum, 1851
    Carabus (Imaibius) boysi boysi Tatum, 1851
    Endophallus
    NW-India, Himachal Pradesh, Manikaran env., Pulga vill.
    Copyright © Ruslan Panin
    Carabus (Imaibius) boysi boysi Tatum, 1851
    Carabus (Imaibius) boysi boysi Tatum, 1851
    Male adult
    NW-India, Himachal Pradesh, Manikaran env., Pulga vill., 4.VII.2008. L=26 mm
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) arrowi hector Hauser, 1924
    Carabus (Apotomopterus) arrowi hector Hauser, 1924 Holotype
    Female adult
    Hunan, Liang to Tang; Ap. hector G.H. Typ; Zoolog. Museum Berlin. Museum für Naturkunde Berlin, Humboldt-Universität; 37 mm
    Copyright © Frank Kleinfeld
    Carabus (Isiocarabus) dargei dargei Deuve, 1987 (as anhuinus xingziensis Kleinfeld 1996)
    Carabus (Isiocarabus) dargei dargei Deuve, 1987 (as anhuinus xingziensis Kleinfeld 1996) Holotype
    Male adult
    China, Jiangxi, Lu Shan mts (ca. 29°26'N/115°57'E). 24 mm
    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) yaophilus Deuve, 1990 (as dayaoshan)
    Carabus (Apotomopterus) yaophilus Deuve, 1990 (as dayaoshan) Holotype
    Female adult
    C-Guangxi: Dayao Shan: NE-part: Mengshan env.: 24:12N/110:31E 44 mm (in coll. Klienfeld)
    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) elektra Kleinfeld, 2004
    Carabus (Apotomopterus) elektra Kleinfeld, 2004 Holotype Paratype
    Male+Female adult
    29,5 mm
    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) roxane miyi Kleinfeld, 2005
    Carabus (Apotomopterus) roxane miyi Kleinfeld, 2005 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) rapuzzi bobaiensis Kleinfeld, 2005
    Carabus (Apotomopterus) rapuzzi bobaiensis Kleinfeld, 2005 Holotype Paratype
    Male+Female adult

    Copyright © Frank Kleinfeld
    Carabus (Aulonocarabus) gossarei huangshuni Kleinfeld & Reuter, 2009
    Carabus (Aulonocarabus) gossarei huangshuni Kleinfeld & Reuter, 2009 Holotype
    Male adult
    China, Heilongjiang, Dongning (44°03'N/131°07'E), 22 mm
    Copyright © Frank Kleinfeld
    Carabus (Aulonocarabus) gossarei huangshuni Kleinfeld & Reuter, 2009
    Carabus (Aulonocarabus) gossarei huangshuni Kleinfeld & Reuter, 2009 Paratype
    Female adult

    Copyright © Frank Kleinfeld
    Carabus (Aulonocarabus) gossarei huangshuni Kleinfeld & Reuter, 2009
    Carabus (Aulonocarabus) gossarei huangshuni Kleinfeld & Reuter, 2009 Paratype
    Male adult

    Copyright © Frank Kleinfeld
    Carabus (Procrustes) piochardi pierpaolo Kleinfeld & Rapuzzi, 2004
    Carabus (Procrustes) piochardi pierpaolo Kleinfeld & Rapuzzi, 2004
    Female adult
    29 mm
    Copyright © Frank Kleinfeld
    Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014
    Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014 Holotype Paratype
    Male adult
    Abkhazia, N Slopes of Bzybian Mt. Range, right bank of River Reshevie (= Reshava) (left tributary of River Bzyb), 700-1000 m, 13.V-02.VII.2010, leg. D. Fominykh, I. Solodovnikov
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014
    Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014 Paratype
    Male adult
    Abkhazia, N Slopes of Bzybian Mt. Range, right bank of River Reshevie (= Reshava) (left tributary of River Bzyb), 700-1000 m, 13.V-02.VII.2010, leg. D. Fominykh, I. Solodovnikov
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014
    Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014 Paratype
    Male+Female adult
    Abkhazia, N Slopes of Bzybian Mt. Range, right bank of River Reshevie (= Reshava) (left tributary of River Bzyb), 700-1000 m, 13.V-02.VII.2010, leg. D. Fominykh, I. Solodovnikov
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014
    Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014 Paratype
    Male adult
    Abkhazia, N Slopes of Bzybian Mt. Range, right bank of River Reshevie (= Reshava) (left tributary of River Bzyb), 700-1000 m, 13.V-02.VII.2010, leg. D. Fominykh, I. Solodovnikov
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014
    Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014
    Endophallus

    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) satyrus s.str.
    Carabus (Neoplectes) satyrus s.str.
    Endophallus

    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) pseudopshuensis Zamotajlov, 1991
    Carabus (Neoplectes) pseudopshuensis Zamotajlov, 1991
    Male+Female adult
    Abkhazia, Sukhum Distr., SE of Village Pskhu, hamlet Bitaga, Mt. Bzybskaya, 550-650 m, beech forest site, 12.V-30.VI.2010, leg. I. Solodovnikov, D. Fominykh
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) pseudopshuensis Zamotajlov, 1991
    Carabus (Neoplectes) pseudopshuensis Zamotajlov, 1991
    Male+Female adult
    Abkhazia, Sukhum Distr., SE of Village Pskhu, hamlet Bitaga, Mt. Bzybskaya, 550-650 m, beech forest site, 12.V-30.VI.2010, leg. I. Solodovnikov, D. Fominykh
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) pseudopshuensis Zamotajlov, 1991
    Carabus (Neoplectes) pseudopshuensis Zamotajlov, 1991
    Male adult
    1 male - Abkhazia, Sukhum Distr., vicinities of Village Sanchara, River Baul, beech forest site, 20.VII.1991, leg. M.N. Maksimenkov 2 male - Abkhazia, Sukhum Distr., SE of Village Pskhu, hamlet Bitaga, Mt. Bzybskaya, 550-650 m, beech forest site, 12.V-30.VI.2010, leg. I. Solodovnikov, D. Fominykh
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) pseudopshuensis Zamotajlov, 1991
    Carabus (Neoplectes) pseudopshuensis Zamotajlov, 1991
    Male+Female adult
    Abkhazia, Sukhum Distr., N Slopes of Bzybian Mt. Range near Village Serebryanyi, 500-600 m, 25.IV-05.VII.2013, leg. D. Fominykh
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) pseudopshuensis Zamotajlov, 1991
    Carabus (Neoplectes) pseudopshuensis Zamotajlov, 1991
    Male+Female adult
    Abkhazia, Sukhum Distr., N Slopes of Bzybian Mt. Range near Village Serebryanyi, 500-600 m, 25.IV-05.VII.2013, leg. D. Fominykh
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) pseudopshuensis Zamotajlov, 1991
    Carabus (Neoplectes) pseudopshuensis Zamotajlov, 1991
    Male+Female adult
    Abkhazia, Sukhum Distr., N Slopes of Bzybian Mt. Range near Village Serebryanyi, 500-600 m, 25.IV-05.VII.2013, leg. D. Fominykh
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) pseudopshuensis Zamotajlov, 1991
    Carabus (Neoplectes) pseudopshuensis Zamotajlov, 1991
    Endophallus

    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) besleticus besleticus Kurnakov, 1972
    Carabus (Neoplectes) besleticus besleticus Kurnakov, 1972
    Male+Female adult
    Abkhazia, Sukhum Distr., vicinities of Village Kaman, N slope of Mt. Birtskha, 160 m, beech, linden, box, elder forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) besleticus besleticus Kurnakov, 1972
    Carabus (Neoplectes) besleticus besleticus Kurnakov, 1972
    Male+Female adult
    Abkhazia, Sukhum Distr., vicinities of Village Kaman, N slope of Mt. Birtskha, 160 m, beech, linden, box, elder forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) besleticus besleticus Kurnakov, 1972
    Carabus (Neoplectes) besleticus besleticus Kurnakov, 1972
    Male+Female adult
    Abkhazia, Sukhum Distr., vicinities of Village Kaman, N slope of Mt. Birtskha, 160 m, beech, linden, box, elder forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) besleticus besleticus Kurnakov, 1972
    Carabus (Neoplectes) besleticus besleticus Kurnakov, 1972
    Male+Female adult
    Abkhazia, Sukhum Distr., vicinities of Village Kaman, N slope of Mt. Birtskha, 160 m, beech, linden, box, elder forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) besleticus besleticus Kurnakov, 1972
    Carabus (Neoplectes) besleticus besleticus Kurnakov, 1972
    Endophallus

    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972
    Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972
    Male+Female adult
    Abkhazia, Gudauta Distr, vicinities of Mtsara (Chiryuta), Zashirbara Mt. Range, source of River Mtsara, beech, blackberry, fern site, 575-690 m, 11.V-05.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972
    Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972
    Male+Female adult
    Abkhazia, Gudauta Distr, vicinities of Mtsara (Chiryuta), Zashirbara Mt. Range, source of River Mtsara, beech, blackberry, fern site, 575-690 m, 11.V-05.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972
    Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972
    Male+Female adult
    Abkhazia, Gudauta Distr, vicinities of Mtsara (Chiryuta), Zashirbara Mt. Range, source of River Mtsara, beech, blackberry, fern site, 575-690 m, 11.V-05.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972
    Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972
    Male+Female adult
    Abkhazia, Gudauta Distr, vicinities of Mtsara (Chiryuta), Zashirbara Mt. Range, source of River Mtsara, beech, blackberry, fern site, 575-690 m, 11.V-05.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972
    Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972
    Endophallus

    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) besleticus dsychvensis Gottwald, 1985
    Carabus (Neoplectes) besleticus dsychvensis Gottwald, 1985
    Male+Female adult
    Abkhazia, Bzybian Mt. Range, SW slopes of Mt. Dzykhva, 2000-2300 m, alpine zone, N 43º13’, E 041º08’, 29.VI-03.VII.2013, leg. D. Fominykh
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) besleticus dsychvensis Gottwald, 1985
    Carabus (Neoplectes) besleticus dsychvensis Gottwald, 1985
    Male+Female adult
    Abkhazia, Bzybian Mt. Range, SW slopes of Mt. Dzykhva, 2000-2300 m, alpine zone, N 43º13’, E 041º08’, 29.VI-03.VII.2013, leg. D. Fominykh
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) besleticus dsychvensis Gottwald, 1985
    Carabus (Neoplectes) besleticus dsychvensis Gottwald, 1985
    Endophallus

    Copyright © Igor Solodovnikov
    Carabus (Eucarabus) obsoletus nagyagensis (as fossulifer Fleischer, 1893)
    Carabus (Eucarabus) obsoletus nagyagensis (as fossulifer Fleischer, 1893)
    Male adult
    NW-Romania, Bihor, Mts Seș (Plopis), 20 km N Aleșd, Pădurea Neagră env., VI.2014, J.Kynkor leg. L=24 mm
    Copyright © Ruslan Panin
    Carabus (Eucarabus) obsoletus obsoletus (as csikii Mallasz, 1901)
    Carabus (Eucarabus) obsoletus obsoletus (as csikii Mallasz, 1901)
    Female adult
    NE Carpathians, Marmarosh Mts., Transcarphatian Reg., near Lug vill., h=350 m, 26.IV-8.VI.2014, R.Panin & Yu. Kanarsky leg. L=30 mm
    Copyright © Ruslan Panin
    Carabus (Eucarabus) obsoletus obsoletus (as csikii Mallasz, 1901)
    Carabus (Eucarabus) obsoletus obsoletus (as csikii Mallasz, 1901)
    Female adult
    NE Carpathians, Marmarosh Mts., Transcarphatian Reg., near Lug vill., h=350 m, 26.IV-8.VI.2014, R.Panin & Yu. Kanarsky leg. L=32 mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) vietinghoffi vietinghoffi Adams, 1812
    Carabus (Megodontus) vietinghoffi vietinghoffi Adams, 1812
    Male adult

    Copyright © Kirill Makarov
    Carabus (Trachycarabus) sibiricus haeres Fischer, 1823
    Carabus (Trachycarabus) sibiricus haeres Fischer, 1823
    Female adult

    Copyright © Kirill Makarov
    Carabus (Trachycarabus) sibiricus haeres Fischer, 1823
    Carabus (Trachycarabus) sibiricus haeres Fischer, 1823
    Male adult
    det. K. Makarov
    Copyright © Kirill Makarov
    Carabus (Neoplectes) besleticus duripshensis Kurnakov, 1972
    Carabus (Neoplectes) besleticus duripshensis Kurnakov, 1972
    Male+Female adult
    Abkhazia, Gudauta Distr., valley of River Khipsta, vicinities of Village Tvanaarkhu, karst crater with cave, beech, blackberry site, 425 m, N 43º14’ E 040º39’9”, 08.V-05.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) besleticus duripshensis Kurnakov, 1972
    Carabus (Neoplectes) besleticus duripshensis Kurnakov, 1972
    Endophallus

    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) besleticus napraensis Belousov & Zamotajlov, 1993
    Carabus (Neoplectes) besleticus napraensis Belousov & Zamotajlov, 1993 Paratype
    Male+Female adult
    Abkhazia, Gudauta Distr., Bzybian Mt. Range, vicinities of Mt. Chibzharga, 1900-2000 m, 20.VI-26.VII.1992, leg. I. Belousov, A. Zamotajlov, A. Miroshnikov
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) besleticus napraensis Belousov & Zamotajlov, 1993
    Carabus (Neoplectes) besleticus napraensis Belousov & Zamotajlov, 1993 Paratype
    Female adult
    Abkhazia, Gudauta Distr., Bzybian Mt. Range, vicinities of Mt. Chibzharga, 1900-2000 m, 20.VI-26.VII.1992, leg. I. Belousov, A. Zamotajlov, A. Miroshnikov
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) besleticus napraensis Belousov & Zamotajlov, 1993
    Carabus (Neoplectes) besleticus napraensis Belousov & Zamotajlov, 1993 Paratype
    Endophallus

    Copyright © Igor Solodovnikov
    Carabus (Tribax) apschuanus pseudoplatessa Gottwald, 1982
    Carabus (Tribax) apschuanus pseudoplatessa Gottwald, 1982
    Male adult
    det. K. Makarov
    Copyright © Kirill Makarov
    Carabus (Tribax) biebersteini fossiger Chaudoir, 1877
    Carabus (Tribax) biebersteini fossiger Chaudoir, 1877
    Male adult
    det. K. Makarov
    Copyright © Kirill Makarov
    Carabus (Aulonocarabus) kolymensis kolymensis Lafer, 1989
    Carabus (Aulonocarabus) kolymensis kolymensis Lafer, 1989
    Male adult

    Copyright © Kirill Makarov
    Carabus (Neoplectes) besleticus anastasiae Solodovnikov, 2014
    Carabus (Neoplectes) besleticus anastasiae Solodovnikov, 2014 Holotype Paratype
    Female adult
    Abkhasia, Gudauta distr., S slope Bzybian Mt.R., upper reach. Riv. Khipsta, 43°18' N / 40°41' E, h = 1907-1915 m, subalpine meadows with karst blocks, 07-12.08.2014, leg. I.A. Solodovnikov, E.V. Tatun
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) besleticus anastasiae Solodovnikov, 2014
    Carabus (Neoplectes) besleticus anastasiae Solodovnikov, 2014 Paratype
    Female adult
    Abkhasia, Gudaura distr., S slope Bzybian Mt.R., upper reach. Riv. Khipsta, 43°18' N / 40°41' E, h = 1907-1915 m, subalpine meadows with karst blocks, 07-12.08.2014, leg. I.A. Solodovnikov, E.V. Tatun
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) besleticus anastasiae Solodovnikov, 2014
    Carabus (Neoplectes) besleticus anastasiae Solodovnikov, 2014 Paratype
    Male adult
    Abkhasia, Gudauta distr., S slope Bzybian Mt.R., upper reach. Riv. Khipsta, 43°18' N / 40°41' E, h = 1984-1990 m, subalpine meadows with big karst blocks, 07-12.08.2014, leg. I.A. Solodovnikov, E.V. Tatun
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) besleticus anastasiae Solodovnikov, 2014
    Carabus (Neoplectes) besleticus anastasiae Solodovnikov, 2014 Paratype
    Female adult
    Abkhasia, Gudauta distr., S slope Bzybian Mt.R., upper reach. Riv. Khipsta, 43°18' N / 40°41' E, h = 1984-1990 m, subalpine meadows with big karst blocks, 07-12.08.2014, leg. I.A. Solodovnikov, E.V. Tatun
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) besleticus anastasiae Solodovnikov, 2014
    Carabus (Neoplectes) besleticus anastasiae Solodovnikov, 2014 Paratype
    Endophallus

    Copyright © Igor Solodovnikov
    Carabus (Coptolabrus) gemmifer pseudoformosus Deuve, 2006
    Carabus (Coptolabrus) gemmifer pseudoformosus Deuve, 2006
    Female adult
    China, Tibet AR, Qamdo, Gonjo County, Kè Rì Xiāng vill.(Kerri Township), h=3100m, VI.2012, A. Mollard leg. L=38 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) pedemontanus raynaudianus Deuve & Simard, 1977
    Carabus (Orinocarabus) pedemontanus raynaudianus Deuve & Simard, 1977
    Male adult
    SE France, Alpes-Maritimes, Isola env., Colle Santa-Anna h=2309 m, 44*13',127" N, 07*06',242"E, 31-7.VIII..2012. L=21 mm
    Copyright © Ruslan Panin
    Carabus (Iniopachus) pyrenaeus pyrenaeus (as cephalicus Csiki, 1927)
    Carabus (Iniopachus) pyrenaeus pyrenaeus (as cephalicus Csiki, 1927)
    Male adult
    SW France, Pyrenees-Atlantiques, Mt.Cirque d'Esturou h=1800m, 29.VIII.2014, Pascal Stefani leg. L=20 mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) violaceus aurichalceus Kraatz, 1879
    Carabus (Megodontus) violaceus aurichalceus Kraatz, 1879
    Male adult
    NC-Spain, Cantabria, Santander, Cueto env.XII.2013 L=28 mm
    Copyright © Ruslan Panin
    Carabus (Chrysocarabus) auronitens festivus Dejean, 1826 (ab. holochrysum)
    Carabus (Chrysocarabus) auronitens festivus Dejean, 1826 (ab. holochrysum)
    Female adult
    SC-FRANCE, Tarn, Le Linas h=550 m, XI.2008, Pascal Stefani leg. L=24 mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) violaceus mixtus (as neomixtus Tarrier, 1975)
    Carabus (Megodontus) violaceus mixtus (as neomixtus Tarrier, 1975)
    Male adult
    L=29mm SE-France, Drôme, Foret de Saou, VII.2014, Pascal Stefani leg.
    Copyright © Ruslan Panin
    Carabus (Megodontus) kolbei hidakamontanus Ishikawa, 1966
    Carabus (Megodontus) kolbei hidakamontanus Ishikawa, 1966
    Female adult
    JAPAN, Hokkaido, Samani distr., Samani town, Horomankyo, 16.08.2013, Tsukasa, Nakamura leg. 25 мм.
    Copyright © Sergey Kakunin
    Carabus (Coptolabrus) ignimitella Bates, 1888
    Carabus (Coptolabrus) ignimitella Bates, 1888
    Female adult
    China, Yunnan, Tali (error?) Image from: Mal, N., Drumont, A., Kerkhof, S. & P. Grootaert, 2012. Pictorial beetle collection of the Royal Belgian Institute of Natural Sciences.
    Copyright © - - -
    Carabus (Coptolabrus) smaragdinus monilifer Tatum, 1847 (as tyrannus)
    Carabus (Coptolabrus) smaragdinus monilifer Tatum, 1847 (as tyrannus)
    Female adult
    Korea, Jeju, Quelpaert Island (= Jeju-Do). Image from: Mal, N., Drumont, A., Kerkhof, S. & P. Grootaert, 2012. Pictorial beetle collection of the Royal Belgian Institute of Natural Sciences.
    Copyright © - - -
    Carabus (Limnocarabus) clathratus auraniensis Muller, 1902 (as antonellii Luigioni, 1921)
    Carabus (Limnocarabus) clathratus auraniensis Muller, 1902 (as antonellii Luigioni, 1921)
    Male adult
    Gaggiano Milan, Italy
    Copyright © Vittorino Monzini
    Carabus (Lamprostus) torosus sinopensis Breuning & Ruspoli, 1970
    Carabus (Lamprostus) torosus sinopensis Breuning & Ruspoli, 1970
    Endophallus
    NC-Turkey, Sinop province, Erfelek env., h=200m
    Copyright © Ruslan Panin
    Carabus (Tomocarabus) convexus erzeliki Schweiger, 1962
    Carabus (Tomocarabus) convexus erzeliki Schweiger, 1962
    Female adult
    NC-Turkey, Sinop province, Erfelek env., h=200m, 10-27.VII.2014, C. Reuter leg. L=20 mm
    Copyright © Ruslan Panin
    Carabus (Archicarabus) phoenix Lapouge, 1924
    Carabus (Archicarabus) phoenix Lapouge, 1924
    Endophallus
    C-Lebanon, Byblos /Jbeil Distr., Kfardebian env., h=1200m,
    Copyright © Ruslan Panin
    Carabus (Archicarabus) phoenix Lapouge, 1924
    Carabus (Archicarabus) phoenix Lapouge, 1924
    Male+Female adult
    C-Lebanon, Byblos /Jbeil Distr., Kfardebian env., h=1200m, XI.2014, C. Reuter leg. female,23mm
    Copyright © Ruslan Panin
    Carabus (Procerus) scabrosus estegeicus (Cavazzuti, 1989)
    Carabus (Procerus) scabrosus estegeicus (Cavazzuti, 1989)
    Female adult

    Copyright © Özgür KOÇAK
    Carabus (Procerus) scabrosus bulgharmaadensis Bodemeyer, 1915 (as ludademae)
    Carabus (Procerus) scabrosus bulgharmaadensis Bodemeyer, 1915 (as ludademae)
    Female adult
    Turkey, Karaman, 1700 m., 03 March 2014
    Copyright © Özgür KOÇAK
    Carabus (Lamprostus) torosus sinopensis Breuning & Ruspoli, 1970
    Carabus (Lamprostus) torosus sinopensis Breuning & Ruspoli, 1970
    Male adult
    NC-Turkey, Sinop province, Erfelek env., h=200m, 10-27.VII.2014, C. Reuter leg. L=31 mm
    Copyright © Ruslan Panin
    Carabus (Diocarabus) massagetus massagetus Motschulsky, 1846 (alpin form)
    Carabus (Diocarabus) massagetus massagetus Motschulsky, 1846 (alpin form)
    Male adult
    Russsia, Altay, Ukok upland, Kaldzhin-Kul Lake h=2400-2550 m, 49 *19*N/87*26*E, 16-27.VII.2008, R.Dudko leg. L=17.5 mm
    Copyright © Ruslan Panin
    Carabus (Procrustes) piochardi praestigiator Morawitz, 1886
    Carabus (Procrustes) piochardi praestigiator Morawitz, 1886
    Endophallus
    C-Lebanon, Byblos /Jbeil Distr., Kfardebian env., h=1200m,
    Copyright © Ruslan Panin
    Carabus (Procrustes) piochardi praestigiator Morawitz, 1886
    Carabus (Procrustes) piochardi praestigiator Morawitz, 1886
    Male adult
    C-Lebanon, Byblos /Jbeil Distr., Kfardebian env., h=1200m, XI.2014, C. Reuter leg. L=31 mm
    Copyright © Ruslan Panin
    Carabus (Procerus) syriacus galilaeus Schweiger, 1970
    Carabus (Procerus) syriacus galilaeus Schweiger, 1970
    Male adult
    Elon, North Galilea (Israel)
    Copyright © Alain Mollard
    Carabus (Procerus) syriacus syriacus Kollar, 1843
    Carabus (Procerus) syriacus syriacus Kollar, 1843
    Female adult
    East Saida, Montes de Lebanon (SC. Lebanon)
    Copyright © Alain Mollard
    Carabus (Procerus) syriacus bulgharmaadensis
    Carabus (Procerus) syriacus bulgharmaadensis
    Female adult
    Etekli (Adana, S.Turquey,)
    Copyright © Alain Mollard
    Carabus (Procerus) scabrosus akbesianus Breuning, 1975
    Carabus (Procerus) scabrosus akbesianus Breuning, 1975
    Male adult
    Akbés (Antakya, Turquey)
    Copyright © Alain Mollard
    Carabus (Procerus) scabrosus amasicus Csiki, 1927
    Carabus (Procerus) scabrosus amasicus Csiki, 1927
    Male adult
    Almus (Tokat, Turquey)
    Copyright © Alain Mollard
    Carabus (Procerus) scabrosus audouini Brulle, 1835
    Carabus (Procerus) scabrosus audouini Brulle, 1835
    Male adult
    Isiklar (Trabzon,Turquey)
    Copyright © Alain Mollard
    Carabus (Procerus) scabrosus callipygius Cavazzuti, 1986
    Carabus (Procerus) scabrosus callipygius Cavazzuti, 1986
    Male adult
    Devrekani (Kastamonu, Turquey)
    Copyright © Alain Mollard
    Carabus (Sphodristocarabus) blumenthaliensis aequicostalis Staven & Heinz, 1995
    Carabus (Sphodristocarabus) blumenthaliensis aequicostalis Staven & Heinz, 1995
    Male adult
    NE-Turkey:NE of Artvin:Dalis Dağ ,h=1600-1800m, 8-18.VII.2014, C. Reuter leg. L= 25 mm
    Copyright © Ruslan Panin
    Carabus (Sphodristocarabus) blumenthaliensis aequicostalis Staven & Heinz, 1995
    Carabus (Sphodristocarabus) blumenthaliensis aequicostalis Staven & Heinz, 1995
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Procerus) gigas parnassicus Kraatz, 1884
    Carabus (Procerus) gigas parnassicus Kraatz, 1884
    Female adult
    N. Greece, Phalakron Mt., 750 m, 09.05.2004. Image of В.И. Гуменюк from www.zin.ru
    Copyright © - - -
    Carabus (Procerus) scabrosus audouini Brulle, 1835
    Carabus (Procerus) scabrosus audouini Brulle, 1835
    Female adult
    Turkey. Rize Prov., South of Ikizdere, 800 m, 17.08.2009. Image of В.И. Гуменюк from www.zin.ru
    Copyright © - - -
    Carabus (Procerus) scabrosus armenius Zaitzev, 1930
    Carabus (Procerus) scabrosus armenius Zaitzev, 1930
    Male adult
    Balkaya (Gumushane, Turquey)
    Copyright © Alain Mollard
    Carabus (Procerus) scabrosus dardanellicus Kraatz, 1886
    Carabus (Procerus) scabrosus dardanellicus Kraatz, 1886
    Male adult
    Sancaeli (Canakkale, Turquey)
    Copyright © Alain Mollard
    Carabus (Procerus) scabrosus fallettianus (Cavazzuti, 1997)
    Carabus (Procerus) scabrosus fallettianus (Cavazzuti, 1997)
    Male adult
    Khndzorek (Goris, Arménie)
    Copyright © Alain Mollard
    Carabus (Procerus) scabrosus montisabanti Schweiger, 1962
    Carabus (Procerus) scabrosus montisabanti Schweiger, 1962
    Male adult
    Abant (Bolu, Turquey)
    Copyright © Alain Mollard
    Carabus (Procrustes) coriaceus rugifer Kraatz, 1877
    Carabus (Procrustes) coriaceus rugifer Kraatz, 1877
    Female adult
    W.UKRAINE, NE Carpathians, Lviv Reg., Skole distr., Zelemyanka Mt.Range h=700m, 26.V-1.VII.2010, Panin R. leg. L=43 mm
    Copyright © Ruslan Panin
    Carabus (Procerus) scabrosus scabrosus Olivier, 1795
    Carabus (Procerus) scabrosus scabrosus Olivier, 1795
    Male adult
    Dermirkoy (Kirklareli, Turquey). L = 54mm
    Copyright © Alain Mollard
    Carabus (Procerus) scabrosus propinquus Csiki, 1927
    Carabus (Procerus) scabrosus propinquus Csiki, 1927
    Male adult
    Alem Dag (Istambul, Turquey) L=57 mm
    Copyright © Alain Mollard
    Carabus (Procerus) scabrosus mentor Blumenthal & Breuning, 1967
    Carabus (Procerus) scabrosus mentor Blumenthal & Breuning, 1967
    Male adult
    Catalzeytin,(Katsamonu,Turquey)
    Copyright © Alain Mollard
    Carabus (Procerus) scabrosus munzurensis Cavazzuti & Lassalle, 1987
    Carabus (Procerus) scabrosus munzurensis Cavazzuti & Lassalle, 1987 Paratype
    Male adult
    leg.B.Lassalle, from Pulumur (Tunceli,Turquey)
    Copyright © Alain Mollard
    Carabus (Cathoplius) stenocephalus ifniensis (Zarco, 1941)
    Carabus (Cathoplius) stenocephalus ifniensis (Zarco, 1941)
    Female adult
    beach of Sidi Ifni city, 10 m a.s.l., 29°23'52"N; 10°10'40"W, Sidi Ifni province, central Morocco. Image from Ghittino, Busato & Casale, 2015.
    Copyright © - - -
    Carabus (Cathoplius) aliai Escalera, 1944
    Carabus (Cathoplius) aliai Escalera, 1944
    Male adult
    Tan Tan. Image from Ghittino, Busato & Casale, 2015.
    Copyright © - - -
    Carabus (Morphocarabus) chaudoiri mongolorum Csiki, 1927
    Carabus (Morphocarabus) chaudoiri mongolorum Csiki, 1927 Lectotype
    Male adult
    Muséum national d’Histoire naturelle, Paris (France); Collection : Insects: Coleoptera (EC); Specimen MNHN-EC- EC522; Credit: Antoine Mantilleri - 2010 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec522
    Copyright © - - -
    Carabus (Coptolabrus) gemmifer mesites Fairmaire, 1897
    Carabus (Coptolabrus) gemmifer mesites Fairmaire, 1897 Holotype
    Female adult
    "Siao-Lou-Lou-Chan, Chasseurs thibétains, 1898". Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC149 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec149
    Copyright © MNHN
    Carabus (Morphocarabus) odoratus chaffanjoni Lesne, 1898
    Carabus (Morphocarabus) odoratus chaffanjoni Lesne, 1898 Holotype
    Male adult
    "Bouloun, Env d'Jakoutsk, J. Chaffanjon" Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC157 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec157
    Copyright © MNHN
    Carabus (Morphocarabus) odoratus chaffanjoni Lesne, 1898
    Carabus (Morphocarabus) odoratus chaffanjoni Lesne, 1898
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC157 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec157
    Copyright © MNHN
    Carabus (Mesocarabus) latus antiquus Dejean, 1826
    Carabus (Mesocarabus) latus antiquus Dejean, 1826 Holotype
    Female adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC36 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec36
    Copyright © - - -
    Carabus (Mesocarabus) latus antiquus Dejean, 1826
    Carabus (Mesocarabus) latus antiquus Dejean, 1826
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC36 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec36
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus brevis Dejean, 1826
    Carabus (Mesocarabus) lusitanicus brevis Dejean, 1826 Lectotype
    Female adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC32 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec32
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus brevis Dejean, 1826
    Carabus (Mesocarabus) lusitanicus brevis Dejean, 1826
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC32 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec32
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus egesippei La Ferté-Sénectere, 1847
    Carabus (Mesocarabus) lusitanicus egesippei La Ferté-Sénectere, 1847 Holotype
    Female adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC154 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec154
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus egesippei La Ferté-Sénectere, 1847
    Carabus (Mesocarabus) lusitanicus egesippei La Ferté-Sénectere, 1847
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC154 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec154
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus mertolaensis Breuning, 1975 (= brevis)
    Carabus (Mesocarabus) lusitanicus mertolaensis Breuning, 1975 (= brevis) Holotype
    Female adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC130 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec130
    Copyright ©
    Carabus (Mesocarabus) lusitanicus mertolaensis (error of label?)
    Carabus (Mesocarabus) lusitanicus mertolaensis (error of label?)
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC130 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec130
    Copyright ©
    Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826
    Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826 Holotype
    Male adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC34 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec34
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826
    Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC34 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec34
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus latus Dejean, 1826
    Carabus (Mesocarabus) lusitanicus latus Dejean, 1826 Lectotype
    Male adult
    "Navalmoral" [Navalmoral de la Mata]. Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC35 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec35
    Copyright © - - -
    Carabus (Mesocarabus) latus latus Dejean, 1826
    Carabus (Mesocarabus) latus latus Dejean, 1826
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC35 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec35
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus trypalsi Breuning, 1975
    Carabus (Mesocarabus) lusitanicus trypalsi Breuning, 1975 Paratype
    Female adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC132 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec132
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus trypalsi Breuning, 1975
    Carabus (Mesocarabus) lusitanicus trypalsi Breuning, 1975
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC132 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec132
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus trypalsi Breuning, 1975
    Carabus (Mesocarabus) lusitanicus trypalsi Breuning, 1975 Holotype
    Male adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC133 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec133
    Copyright ©
    Carabus (Mesocarabus) lusitanicus trypalsi Breuning, 1975
    Carabus (Mesocarabus) lusitanicus trypalsi Breuning, 1975
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC133 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec133
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus helluo Dejean, 1826
    Carabus (Mesocarabus) lusitanicus helluo Dejean, 1826 Holotype
    Female adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC31 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec31
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus helluo Dejean, 1826
    Carabus (Mesocarabus) lusitanicus helluo Dejean, 1826
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC31 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec31
    Copyright © - - -
    Carabus (Mesocarabus) dufouri papillatus Lapouge, 1902
    Carabus (Mesocarabus) dufouri papillatus Lapouge, 1902 Lectotype
    Male adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC576 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec576
    Copyright © - - -
    Carabus (Mesocarabus) dufouri papillatus Lapouge, 1902
    Carabus (Mesocarabus) dufouri papillatus Lapouge, 1902
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC576 http://coldb.mnhn.fr/catalognumber/mnhn
    Copyright © - - -
    Carabus groenlandicus Dejean, 1831 (=chamissonis)
    Carabus groenlandicus Dejean, 1831 (=chamissonis) Lectotype
    Male adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC64 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec64
    Copyright © - - -
    Carabus groenlandicus Dejean, 1831 (=chamissonis)
    Carabus groenlandicus Dejean, 1831 (=chamissonis)
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC64 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec64
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus gougeleti (as valderepisensis Branes, 1988)
    Carabus (Mesocarabus) lusitanicus gougeleti (as valderepisensis Branes, 1988)
    Male adult
    Spain, Jaen prov., Despeñaperros, Arroyo de los Arcos, 38.381847, -3.515651, 1-2.05.2014, Anichtchenko A. leg. & det.; 18 mm
    Copyright © Alexander Anichtchenko
    Carabus (Coptolabrus) elysii rothschildi Born, 1899
    Carabus (Coptolabrus) elysii rothschildi Born, 1899 Holotype
    Female adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC123 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec123
    Copyright © - - -
    Carabus (Coptolabrus) elysii rothschildi Born, 1899
    Carabus (Coptolabrus) elysii rothschildi Born, 1899
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC123 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec123
    Copyright © - - -
    Carabus (Apotomopterus) kouanping kouanping Maindron, 1906
    Carabus (Apotomopterus) kouanping kouanping Maindron, 1906 Holotype
    Female adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC158 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec158
    Copyright © - - -
    Carabus (Apotomopterus) kouanping kouanping Maindron, 1906
    Carabus (Apotomopterus) kouanping kouanping Maindron, 1906
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC158 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec158
    Copyright © - - -
    Carabus (Megodontus) leachi panzeri Dejean, 1829
    Carabus (Megodontus) leachi panzeri Dejean, 1829 Holotype
    Male adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC48 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec48
    Copyright © - - -
    Carabus (Megodontus) leachi panzeri Dejean, 1829
    Carabus (Megodontus) leachi panzeri Dejean, 1829
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC48 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec48
    Copyright © - - -
    Carabus (Morphocarabus) tarbagataicus var.motschulskyi Lapouge, 1909
    Carabus (Morphocarabus) tarbagataicus var.motschulskyi Lapouge, 1909 Holotype
    Male adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC525 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec525
    Copyright © - - -
    Carabus (Morphocarabus) monilis var.motschulskyi Lapouge, 1909
    Carabus (Morphocarabus) monilis var.motschulskyi Lapouge, 1909
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC525 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec525
    Copyright © - - -
    Carabus (Morphocarabus) henningi sahlbergi Dejean, 1826
    Carabus (Morphocarabus) henningi sahlbergi Dejean, 1826 Lectotype
    Male adult
    "Siberie, Slataoust" = [Златоуст, Zlatoust, Cheliabinsk prov.]. Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC1 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec1
    Copyright © - - -
    Carabus (Morphocarabus) henningi sahlbergi Dejean, 1826
    Carabus (Morphocarabus) henningi sahlbergi Dejean, 1826
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC1 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec1
    Copyright © - - -
    Carabus (Eurycarabus) famini itoensis Antoine, 1939
    Carabus (Eurycarabus) famini itoensis Antoine, 1939 Paratype
    Female adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC1049 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec1049
    Copyright © - - -
    Carabus (Eurycarabus) famini itoensis Antoine, 1939
    Carabus (Eurycarabus) famini itoensis Antoine, 1939
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC1049 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec1049
    Copyright © - - -
    Carabus (Eurycarabus) famini favieri Fairmaire, 1859
    Carabus (Eurycarabus) famini favieri Fairmaire, 1859 Holotype
    Female adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC143 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec143
    Copyright © - - -
    Carabus (Eurycarabus) famini favieri Fairmaire, 1859
    Carabus (Eurycarabus) famini favieri Fairmaire, 1859
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC143 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec143
    Copyright © - - -
    Carabus (Eurycarabus) famini grasselyi Antoine, 1941
    Carabus (Eurycarabus) famini grasselyi Antoine, 1941 Holotype
    Female adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC1044 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec1044
    Copyright © - - -
    Carabus (Eurycarabus) famini grasselyi Antoine, 1941
    Carabus (Eurycarabus) famini grasselyi Antoine, 1941
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC1044 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec1044
    Copyright © - - -
    Carabus (Eurycarabus) famini mairei Antoine, 1925
    Carabus (Eurycarabus) famini mairei Antoine, 1925 Holotype
    Female adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC1047 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec1047
    Copyright © - - -
    Carabus (Eurycarabus) famini mairei Antoine, 1925
    Carabus (Eurycarabus) famini mairei Antoine, 1925
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC1047 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec1047
    Copyright © - - -
    Carabus (Eurycarabus) famini itoensis Antoine, 1939
    Carabus (Eurycarabus) famini itoensis Antoine, 1939 Holotype
    Male adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC1048 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec1048
    Copyright © - - -
    Carabus (Eurycarabus) famini itoensis Antoine, 1939
    Carabus (Eurycarabus) famini itoensis Antoine, 1939
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC1048 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec1048
    Copyright © - - -
    Carabus (Eurycarabus) famini atlantis Alluaud, 1923
    Carabus (Eurycarabus) famini atlantis Alluaud, 1923 Holotype
    Male adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC1058 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec1058
    Copyright © - - -
    Carabus (Eurycarabus) famini atlantis Alluaud, 1923
    Carabus (Eurycarabus) famini atlantis Alluaud, 1923
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC1058 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec1058
    Copyright © - - -
    Carabus (Eurycarabus) famini algiricus Géhin, 1885
    Carabus (Eurycarabus) famini algiricus Géhin, 1885 Holotype
    Male adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC433 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec433
    Copyright © - - -
    Carabus (Eurycarabus) famini algiricus Géhin, 1885
    Carabus (Eurycarabus) famini algiricus Géhin, 1885
    Label

    Copyright © - - -
    Carabus (Eurycarabus) famini piraticus Fairmaire, 1880
    Carabus (Eurycarabus) famini piraticus Fairmaire, 1880 Lectotype
    Male adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC144 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec144
    Copyright © - - -
    Carabus (Eurycarabus) famini piraticus Fairmaire, 1880
    Carabus (Eurycarabus) famini piraticus Fairmaire, 1880
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC144 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec144
    Copyright © - - -
    Carabus (Mesocarabus) riffensis Fairmaire, 1875
    Carabus (Mesocarabus) riffensis Fairmaire, 1875 Lectotype
    Female adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC142 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec142
    Copyright © - - -
    Carabus (Mesocarabus) riffensis Fairmaire, 1875
    Carabus (Mesocarabus) riffensis Fairmaire, 1875
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC142 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec142
    Copyright © - - -
    Carabus (Coptolabrus) augustus augustus Bates, 1888
    Carabus (Coptolabrus) augustus augustus Bates, 1888 Lectotype
    Male adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC105 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec105
    Copyright © - - -
    Carabus (Coptolabrus) augustus augustus Bates, 1888
    Carabus (Coptolabrus) augustus augustus Bates, 1888
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC105 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec105
    Copyright © - - -
    Carabus (Isiocarabus) hienfoungi hienfoungi Thomson, 1857
    Carabus (Isiocarabus) hienfoungi hienfoungi Thomson, 1857 Lectotype
    Male adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC164 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec164
    Copyright © - - -
    Carabus (Isiocarabus) hienfoungi hienfoungi Thomson, 1857
    Carabus (Isiocarabus) hienfoungi hienfoungi Thomson, 1857
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC164 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec164
    Copyright © - - -
    Carabus (Procerus) caucasicus var.tuberculosus
    Carabus (Procerus) caucasicus var.tuberculosus Lectotype
    Female adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC468 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec468
    Copyright © - - -
    Carabus (Procerus) scabrosus var.tuberculosus Géhin, 1885
    Carabus (Procerus) scabrosus var.tuberculosus Géhin, 1885
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC468 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec468
    Copyright © - - -
    Carabus (Procerus) gigas duponcheli Dejean, 1831
    Carabus (Procerus) gigas duponcheli Dejean, 1831 Holotype
    Male adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC54 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec54
    Copyright © - - -
    Carabus (Procerus) gigas duponcheli Dejean, 1831
    Carabus (Procerus) gigas duponcheli Dejean, 1831
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC54 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec54
    Copyright © - - -
    Carabus (Imaibius) dardiellus var.granulisparsus Bates, 1889
    Carabus (Imaibius) dardiellus var.granulisparsus Bates, 1889 Lectotype
    Female adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC100 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec100
    Copyright © - - -
    Carabus (Imaibius) dardiellus var.granulisparsus Bates, 1889
    Carabus (Imaibius) dardiellus var.granulisparsus Bates, 1889
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC100 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec100
    Copyright © - - -
    Carabus (Pagocarabus) trachynodes trachynodes Bates, 1891
    Carabus (Pagocarabus) trachynodes trachynodes Bates, 1891 Lectotype
    Male adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC89 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec89
    Copyright © - - -
    Carabus (Pagocarabus) trachynodes trachynodes Bates, 1891
    Carabus (Pagocarabus) trachynodes trachynodes Bates, 1891
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC89 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec89
    Copyright © - - -
    Carabus (Mesocarabus) riffensis Fairmaire, 1875
    Carabus (Mesocarabus) riffensis Fairmaire, 1875
    Male adult
    Morocco, Rif, Foresta di Ketama, 25.06.1998, 1600m, leg. D. Sechi (immature)
    Copyright © Alexander Anichtchenko
    Carabus (Mesocarabus) riffensis Fairmaire, 1875
    Carabus (Mesocarabus) riffensis Fairmaire, 1875
    Endophallus
    Morocco, Rif, Foresta di Ketama, 25.06.1998, 1600m, leg. D. Sechi (immature)
    Copyright © Alexander Anichtchenko
    Carabus (Apotomopterus) arrowi shimenensis Deuve, 1997
    Carabus (Apotomopterus) arrowi shimenensis Deuve, 1997
    Female adult
    Qiangqiang, Sichuan, Chine. 40 mm
    Copyright © Alain Mollard
    Carabus (Apotomopterus) arrowi arrowi Hauser, 1913
    Carabus (Apotomopterus) arrowi arrowi Hauser, 1913
    Female adult
    Simianshan, Guizhou, Chine. 31 mm
    Copyright © Alain Mollard
    Carabus (Apotomopterus) prattianus prattianus Bates, 1890
    Carabus (Apotomopterus) prattianus prattianus Bates, 1890
    Female adult
    Yichang, Hubei, Chine. 37 mm
    Copyright © Alain Mollard
    Carabus (Apotomopterus) prattianus prattianus Bates, 1890
    Carabus (Apotomopterus) prattianus prattianus Bates, 1890
    Male adult
    Wufeng, Hubei, Chine. 35 mm
    Copyright © Alain Mollard
    Carabus (Morphocarabus) odoratus melleus Lapouge, 1909
    Carabus (Morphocarabus) odoratus melleus Lapouge, 1909 Paralectotypes
    Male adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC521
    Copyright © MNHN
    Carabus (Morphocarabus) odoratus melleus Lapouge, 1909
    Carabus (Morphocarabus) odoratus melleus Lapouge, 1909 Lectotype
    Female adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC520
    Copyright © MNHN
    Carabus (Morphocarabus) hummeli stolidus Lapouge, 1924
    Carabus (Morphocarabus) hummeli stolidus Lapouge, 1924 Lectotype
    Female adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC519
    Copyright © - - -
    Carabus (Morphocarabus) hummeli stolidus Lapouge, 1924
    Carabus (Morphocarabus) hummeli stolidus Lapouge, 1924
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC519
    Copyright © - - -
    Carabus (Diocarabus) massagetus pusillus Lapouge, 1915
    Carabus (Diocarabus) massagetus pusillus Lapouge, 1915 Lectotype
    Female adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC526
    Copyright © - - -
    Carabus (Diocarabus) massagetus pusillus Lapouge, 1915
    Carabus (Diocarabus) massagetus pusillus Lapouge, 1915
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC526
    Copyright © - - -
    Carabus (Aulonocarabus) canaliculatus rufipennis Lapouge, 1910
    Carabus (Aulonocarabus) canaliculatus rufipennis Lapouge, 1910 Lectotype
    Male adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC517
    Copyright © - - -
    Carabus (Aulonocarabus) canaliculatus rufipennis
    Carabus (Aulonocarabus) canaliculatus rufipennis
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC517
    Copyright © - - -
    Carabus (Aulonocarabus) canaliculatus penialis Lapouge, 1913
    Carabus (Aulonocarabus) canaliculatus penialis Lapouge, 1913 Lectotype
    Male adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC516
    Copyright © - - -
    Carabus (Aulonocarabus) canaliculatus penialis Lapouge, 1913
    Carabus (Aulonocarabus) canaliculatus penialis Lapouge, 1913
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC516
    Copyright © - - -
    Carabus (Morphocarabus) excellens paradoxus Lapouge, 1906
    Carabus (Morphocarabus) excellens paradoxus Lapouge, 1906 Lectotype
    Female adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC539
    Copyright © - - -
    Carabus (Morphocarabus) excellens paradoxus
    Carabus (Morphocarabus) excellens paradoxus
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC539
    Copyright © - - -
    Carabus (Morphocarabus) odoratus dohrni Gebler, 1847: 300
    Carabus (Morphocarabus) odoratus dohrni Gebler, 1847: 300 Lectotype
    Female adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC413
    Copyright © MNHN
    Carabus (Morphocarabus) odoratus dohrni Gebler, 1847
    Carabus (Morphocarabus) odoratus dohrni Gebler, 1847
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC413
    Copyright © MNHN
    Carabus (Scambocarabus) kruberi bungii Gebler, 1830
    Carabus (Scambocarabus) kruberi bungii Gebler, 1830 Lectotype
    Male adult
    Not TYPE, because type was described from Altay. Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC416 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec416
    Copyright © MNHN
    Carabus (Scambocarabus) kruberi bungii Gebler, 1830
    Carabus (Scambocarabus) kruberi bungii Gebler, 1830
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC416 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec416
    Copyright © MNHN
    Carabus (Coptolabrus) pustulifer pustulifer Lucas, 1869
    Carabus (Coptolabrus) pustulifer pustulifer Lucas, 1869 Holotype
    Female adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC163 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec163
    Copyright © - - -
    Carabus (Coptolabrus) pustulifer pustulifer Lucas, 1869
    Carabus (Coptolabrus) pustulifer pustulifer Lucas, 1869
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC163 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec163
    Copyright © - - -
    Carabus (Tanaocarabus) sylvosus Say, 1825
    Carabus (Tanaocarabus) sylvosus Say, 1825 Lectotype
    Male adult
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC63 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec63
    Copyright © - - -
    Carabus (Tanaocarabus) sylvosus Say, 1825
    Carabus (Tanaocarabus) sylvosus Say, 1825
    Label
    Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC63 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec63
    Copyright © - - -
    Carabus (Neoplectes) khipstaensis Solodovnikov, 2015
    Carabus (Neoplectes) khipstaensis Solodovnikov, 2015 Paratype
    Male adult
    Абхазия, Гудаутский р-н, Ю скл. Бзыбского хр., водораздельный холм в истоках р. Хипста, 43°17'52.03" N / 40°40'8.72" E, h = 1435 m, Линия № 64-4, буковый спелый лес с папоротниками и ежевикой, 09-13.08.2014, leg. И.А. Солодовников, Е.В. Татун
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) khipstaensis Solodovnikov, 2015
    Carabus (Neoplectes) khipstaensis Solodovnikov, 2015 Paratype
    Male adult
    Абхазия, Гудаутский р-н, Ю скл. Бзыбского хр., водораздельный холм в истоках р. Хипста, 43°17'52.03" N / 40°40'8.72" E, h = 1435 m, Линия № 64-4, буковый спелый лес с папоротниками и ежевикой, 09-13.08.2014, leg. И.А. Солодовников, Е.В. Татун
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) khipstaensis Solodovnikov, 2015
    Carabus (Neoplectes) khipstaensis Solodovnikov, 2015 Paratype
    Female adult
    Абхазия, Гудаутский р-н, Ю скл. Бзыбского хр., водораздельный холм в истоках р. Хипста, 43°17'52.03" N / 40°40'8.72" E, h = 1435 m, Линия № 64-4, буковый спелый лес с папоротниками и ежевикой, 09-13.08.2014, leg. И.А. Солодовников, Е.В. Татун
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) khipstaensis Solodovnikov, 2015
    Carabus (Neoplectes) khipstaensis Solodovnikov, 2015 Paratype
    Female adult
    Абхазия, Гудаутский р-н, Ю скл. Бзыбского хр., водораздельный холм в истоках р. Хипста, 43°17'52.03" N / 40°40'8.72" E, h = 1435 m, Линия № 64-4, буковый спелый лес с папоротниками и ежевикой, 09-13.08.2014, leg. И.А. Солодовников, Е.В. Татун
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) rousianus rousianus Gottwald, 1985
    Carabus (Neoplectes) rousianus rousianus Gottwald, 1985
    Male+Female adult
    Абхазия, Гудаутский р-н, 5-8 км ЮВ от пос. Лдзаа (Ludzava) долина р. Риапши, левый борт; балки, поросшие орехом, каштаном, рододендроном понтийским и кленом, h = 58-70 m, 43º10’43” N / 40º25’37” E, 16-22.07.2011, leg. И.А. Солодовников, Н.Ю. Пичугин, С.M. Павлючук
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) rousianus rousianus Gottwald, 1985
    Carabus (Neoplectes) rousianus rousianus Gottwald, 1985
    Male+Female adult
    Абхазия, Гудаутский р-н, 5-8 км ЮВ от пос. Лдзаа (Ludzava) долина р. Риапши, левый борт; балки, поросшие орехом, каштаном, рододендроном понтийским и кленом, h = 58-70 m, 43º10’43” N / 40º25’37” E, 16-22.07.2011, leg. И.А. Солодовников, Н.Ю. Пичугин, С.M. Павлючук
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) polychrous Rost, 1892
    Carabus (Neoplectes) polychrous Rost, 1892
    Female adult
    Абхазия, Гудаутский р-н, окр. п. Тванаарху, дол. р. Хипсты, ущелье небольшого ручейка, поросшее буками, ежевикой и папоротником, Линия 25-3, 43°14’ N / 040°39’ E, h = 500 m,08.05-05.07.2013, leg. И.А. Солодовников, С.В. Солодовникова, С.M. Павлючук, Н.Ю. Пичугин, В.М. Коцур
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) polychrous Rost, 1892
    Carabus (Neoplectes) polychrous Rost, 1892
    Male+Female adult
    Абхазия, Гудаутский р-н, окр. п. Тванаарху, дол. р. Хипсты, ущелье небольшого ручейка, поросшее буками, ежевикой и папоротником, Линия 25-3, 43°14’ N / 040°39’ E, h = 500 m,08.05-05.07.2013, leg. И.А. Солодовников, С.В. Солодовникова, С.M. Павлючук, Н.Ю. Пичугин, В.М. Коцур
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) polychrous Rost, 1892
    Carabus (Neoplectes) polychrous Rost, 1892
    Male+Female adult
    Абхазия, Гудаутский р-н, окр. п. Тванаарху, дол. р. Хипсты, ущелье небольшого ручейка, поросшее буками, ежевикой и папоротником, Линия 25-3, 43°14’ N / 040°39’ E, h = 500 m,08.05-05.07.2013, leg. И.А. Солодовников, С.В. Солодовникова, С.M. Павлючук, Н.Ю. Пичугин, В.М. Коцур
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) rousianus svetlanae Solodovnikov, 2015
    Carabus (Neoplectes) rousianus svetlanae Solodovnikov, 2015 Paratype
    Female adult
    Абхазия, Гудаутский р-н, 12 км дороги на оз. Рица, лев. борт р. Бзыбь, буковый лес + каштан на щебнисто-глинистой почве, уроч. Авджуляр, Линия № 20, h = 210-230 m, 43º19’07” N / 040º25’06” E, 09.V-03.VII.2013, leg. И.А. Солодовников, С.В. Солодовникова, С.М. Павлючук, Н.Ю. Пичугин, В.М. Коцур
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) rousianus svetlanae Solodovnikov, 2015
    Carabus (Neoplectes) rousianus svetlanae Solodovnikov, 2015 Paratype
    Male+Female adult
    Абхазия, Гудаутский р-н, 12 км дороги на оз. Рица, лев. борт р. Бзыбь, буковый лес + каштан на щебнисто-глинистой почве, уроч. Авджуляр, Линия № 20, h = 210-230 m, 43º19’07” N / 040º25’06” E, 09.V-03.VII.2013, leg. И.А. Солодовников, С.В. Солодовникова, С.М. Павлючук, Н.Ю. Пичугин, В.М. Коцур
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) rousianus svetlanae Solodovnikov, 2015
    Carabus (Neoplectes) rousianus svetlanae Solodovnikov, 2015 Paratype
    Male adult
    Абхазия, Гудаутский р-н, 12 км дороги на оз. Рица, лев. борт р. Бзыбь, буковый лес + каштан на щебнисто-глинистой почве, уроч. Авджуляр, Линия № 20, h = 210-230 m, 43º19’07” N / 040º25’06” E, 09.V-03.VII.2013, leg. И.А. Солодовников, С.В. Солодовникова, С.М. Павлючук, Н.Ю. Пичугин, В.М. Коцур
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) rousianus svetlanae Solodovnikov, 2015
    Carabus (Neoplectes) rousianus svetlanae Solodovnikov, 2015 Paratype
    Male adult
    Абхазия, Гудаутский р-н, 12 км дороги на оз. Рица, лев. борт р. Бзыбь, буковый лес + каштан на щебнисто-глинистой почве, уроч. Авджуляр, Линия № 20, h = 210-230 m, 43º19’07” N / 040º25’06” E, 09.V-03.VII.2013, leg. И.А. Солодовников, С.В. Солодовникова, С.М. Павлючук, Н.Ю. Пичугин, В.М. Коцур
    Copyright © Igor Solodovnikov
    Carabus (Limnocarabus) clathratus clathratus Linne, 1761
    Carabus (Limnocarabus) clathratus clathratus Linne, 1761
    Female adult

    Copyright © Kirill Makarov
    Carabus (Megodontus) violaceus aurolimbatus Dejean, 1829
    Carabus (Megodontus) violaceus aurolimbatus Dejean, 1829
    Male adult

    Copyright © Kirill Makarov
    Carabus (Euleptocarabus) porrecticollis Bates, 1883
    Carabus (Euleptocarabus) porrecticollis Bates, 1883
    Female adult
    Atsushiokanomachikano, Kitakata City, Fukushima Pref., Japan, 29.III.2014, T. Nakamura leg.
    Copyright © Alexander Anichtchenko
    Carabus (Limnocarabus) clathratus maacki Morawitz, 1862
    Carabus (Limnocarabus) clathratus maacki Morawitz, 1862
    Female adult
    Приморский край, Романовка, VI.1986
    Copyright © Alexander Anichtchenko
    Carabus (Ophiocarabus) wusunshanicus Kabak, 2015
    Carabus (Ophiocarabus) wusunshanicus Kabak, 2015 Paratype
    Male adult

    Copyright © Ilya Kabak
    Carabus (Ophiocarabus) wusunshanicus Kabak, 2015
    Carabus (Ophiocarabus) wusunshanicus Kabak, 2015
    Endophallus

    Copyright © Ilya Kabak
    Carabus (Ophiocarabus) regeli regeli Morawitz, 1886
    Carabus (Ophiocarabus) regeli regeli Morawitz, 1886
    Endophallus
    Karatau Mt. R.
    Copyright © Ilya Kabak
    Carabus (Lamprostus) hemprichi elonensis Schweiger, 1970
    Carabus (Lamprostus) hemprichi elonensis Schweiger, 1970
    Endophallus
    N-Israel, Midlle Galilea, Nazareth env., Hoshaya viil.
    Copyright © Ruslan Panin
    Carabus (Lamprostus) hemprichi elonensis Schweiger, 1970
    Carabus (Lamprostus) hemprichi elonensis Schweiger, 1970
    Male adult
    N-Israel, Midlle Galilea, Nazareth env., Hoshaya viil., 3-12..III.2015, B.Vas’ko leg./det. Panin R.
    Copyright © Ruslan Panin
    Carabus (Lamprostus) syrus cheikensis Deuve, 1992
    Carabus (Lamprostus) syrus cheikensis Deuve, 1992
    Endophallus
    N-Israel, Midlle Galilea, Nazareth env., Hoshaya viil.,
    Copyright © Ruslan Panin
    Carabus (Lamprostus) syrus cheikensis Deuve, 1992
    Carabus (Lamprostus) syrus cheikensis Deuve, 1992
    Male adult
    N-Israel, Midlle Galilea, Nazareth env., Hoshaya viil., 13.XII.2012, Vas’ko, leg./det. Panin R. L=30мм
    Copyright © Ruslan Panin
    Carabus (Neoplectes) reitteri gagrinus Starck, 1894
    Carabus (Neoplectes) reitteri gagrinus Starck, 1894
    Male adult
    ABKHASIA, Gagra distr., Mamdsyshkha Mt., near Gagra, Fagus forest + karst, h = 1300-1330 m, biotope N 23, 43º18’186" N / 040º18’44" E, 12.05-05.07.2014, Leg. I.A. Solodovnikov, S.V. Solodovnikova
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) reitteri gagrinus Starck, 1894
    Carabus (Neoplectes) reitteri gagrinus Starck, 1894
    Female adult
    ABKHASIA, Gagra distr., Mamdsyshkha Mt., near Gagra, Fagus forest + karst, h = 1300-1330 m, biotope N 23, 43º18’186" N / 040º18’44" E, 12.05-05.07.2014, Leg. I.A. Solodovnikov, S.V. Solodovnikova
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) reitteri gagrinus Starck, 1894
    Carabus (Neoplectes) reitteri gagrinus Starck, 1894
    Female adult
    ABKHASIA, Gagra distr., Mamdsyshkha Mt., near Gagra, Fagus forest + karst, h = 1300-1330 m, biotope N 23, 43º18’186" N / 040º18’44" E, 12.05-05.07.2014, Leg. I.A. Solodovnikov, S.V. Solodovnikova
    Copyright © Igor Solodovnikov
    Carabus (Procrustes) paiafa paiafa White, 1845
    Carabus (Procrustes) paiafa paiafa White, 1845
    Endophallus

    Copyright © Kirill Makarov
    Carabus (Procrustes) paiafa paiafa White, 1845
    Carabus (Procrustes) paiafa paiafa White, 1845
    Male adult
    Turkey, Antalya Prov., Finikie env., h~200 20.V.2015 L=41 mm
    Copyright © Kirill Makarov
    Carabus (Apotomopterus) breuningianus breuningianus Le Moult, 1930
    Carabus (Apotomopterus) breuningianus breuningianus Le Moult, 1930
    Female adult
    SC-China, Guangxi Reg., Pingle County, Mt. Shizishan h=1290 m,5-20,VII.2011, local coll. leg. L=43 mm
    Copyright © Ruslan Panin
    Carabus (Leptocarabus) arboreus fujisanus Bates, 1883
    Carabus (Leptocarabus) arboreus fujisanus Bates, 1883
    Male adult
    Japan, CE-Honshu, Yamanashi pref., Mt. Fujisan h~1600m, 23.IX.1982, Y. Imura leg. L=23 mm
    Copyright © Ruslan Panin
    Carabus (Pachystus) cavernosus variolatus Costa, 1839
    Carabus (Pachystus) cavernosus variolatus Costa, 1839
    Male adult
    C-Italy:Abruzzo reg., Gran Sasso d'Italia Massif, h~1600 m, VIII.1997, Seichert leg. L=26 mm
    Copyright © Ruslan Panin
    Carabus (Carabus) stscheglowi Mannerheim, 1827
    Carabus (Carabus) stscheglowi Mannerheim, 1827
    Endophallus
    Russian, Chuvashia, Cheboksary, Zavolzhye distr., , near lake Astrakhanka,
    Copyright © Ruslan Panin
    Carabus (Carabus) stscheglowi Mannerheim, 1827
    Carabus (Carabus) stscheglowi Mannerheim, 1827
    Male adult
    Russian, Chuvashia, Cheboksary, Zavolzhye distr., , near lake Astrakhanka, L=27 mm
    Copyright © Ruslan Panin
    Carabus (Acoptolabrus) mirabilissimus mirabilissimus Ishikawa & Deuve, 1982
    Carabus (Acoptolabrus) mirabilissimus mirabilissimus Ishikawa & Deuve, 1982
    Female adult
    Mt.Taebaek-san, Kangwon-do,VI-2002, CE-Korea (L=26mm)
    Copyright © Alain Mollard
    Carabus (Acoptolabrus) mirabilissimus pseudokana Deuve & Li, 2002
    Carabus (Acoptolabrus) mirabilissimus pseudokana Deuve & Li, 2002
    Male adult
    Mt.Wujia-san, Chagang-do, VII-2004, N-Korea (L=21.5mm)
    Copyright © Alain Mollard
    Carabus (Acoptolabrus) mirabilissimus kana Imura, 1989
    Carabus (Acoptolabrus) mirabilissimus kana Imura, 1989
    Female adult
    Mt.Togyu-san, Cholla-bukto, VII-2000, SC-Korea (L=23.5mm)
    Copyright © Alain Mollard
    Carabus (Acoptolabrus) mirabilissimus igniferescens Deuve, 1992
    Carabus (Acoptolabrus) mirabilissimus igniferescens Deuve, 1992
    Male adult
    Mt. Taebaek-san, Kangwon-do, VI-2002, CE-Korea (L=21.5mm)
    Copyright © Alain Mollard
    Carabus (Acoptolabrus) mirabilissimus mirabilissimus (as furumiensis)
    Carabus (Acoptolabrus) mirabilissimus mirabilissimus (as furumiensis)
    Male adult
    Mt.Sobaek-san, Ch’uich’ong bukto, VII-2001, CE-Korea (L=22mm)
    Copyright © Alain Mollard
    Carabus (Acoptolabrus) constricticollis igniferrimus Deuve & Li, 2003
    Carabus (Acoptolabrus) constricticollis igniferrimus Deuve & Li, 2003
    Female adult
    Mt.Nangnim-san, Chagang-do, VI-2002, N-Korea L=32mm
    Copyright © Alain Mollard
    Carabus (Acoptolabrus) constricticollis chonmasanensis Deuve, 1994
    Carabus (Acoptolabrus) constricticollis chonmasanensis Deuve, 1994
    Female adult
    Mt. Chonma-san, Pyongang-bukto VII-1996, N-Korea L=35mm
    Copyright © Alain Mollard
    Carabus (Acoptolabrus) constricticollis furumielloides Deuve, 1997
    Carabus (Acoptolabrus) constricticollis furumielloides Deuve, 1997
    Male adult
    Mt. Liuhe, Jilin Prov. VI-1997, Chine L=23mm
    Copyright © Alain Mollard
    Carabus (Acoptolabrus) constricticollis furumielloides Deuve, 1997
    Carabus (Acoptolabrus) constricticollis furumielloides Deuve, 1997
    Male adult
    Lanheyu, E-Benxi, Liaoning Prov. VII-1998, Chine L=33mm
    Copyright © Alain Mollard
    Carabus (Acoptolabrus) constricticollis samjiyonensis Deuve & Li, 2000
    Carabus (Acoptolabrus) constricticollis samjiyonensis Deuve & Li, 2000
    Male adult
    Mt.Samjiyon-san, Yanggang-do, VI-2004, N-Korea L=22mm
    Copyright © Alain Mollard
    Carabus (Acoptolabrus) constricticollis myohyangensis Deuve & Li, 1998
    Carabus (Acoptolabrus) constricticollis myohyangensis Deuve & Li, 1998
    Female adult
    Mt.Myohyang-san, Pyongang-bukto VI-2002, N-Korea L=28mm
    Copyright © Alain Mollard
    Carabus (Mesocarabus) cantabricus moroderi (as rodriguezianus Meyer & Mollard in Deuve, 2003)
    Carabus (Mesocarabus) cantabricus moroderi (as rodriguezianus Meyer & Mollard in Deuve, 2003)
    Female adult
    Poza de la Sal, Burgos, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Acoptolabrus) gehinii gehinii (as konsenensis Ishikawa,1968)
    Carabus (Acoptolabrus) gehinii gehinii (as konsenensis Ishikawa,1968)
    Male adult
    Shibecha (C-Hokkaido) Japan 01/05/1993
    Copyright © Alain Mollard
    Carabus (Acoptolabrus) gehinii gehinii (as radiatocostatus Ishikawa,1968)
    Carabus (Acoptolabrus) gehinii gehinii (as radiatocostatus Ishikawa,1968)
    Female adult
    Erimo-Cho (SC-Hokkaido) Japan 15/06/1990.
    Copyright © Alain Mollard
    Carabus (Acoptolabrus) gehinii gehinii (as shimizui Imura, 1994)
    Carabus (Acoptolabrus) gehinii gehinii (as shimizui Imura, 1994)
    Male adult
    Tamaru-toge-Pass , (Shakotan),Japan 01/05/1995.
    Copyright © Alain Mollard
    Carabus (Acoptolabrus) gehinii gehinii (as sapporensis Uchida & Tamanuki,1927)
    Carabus (Acoptolabrus) gehinii gehinii (as sapporensis Uchida & Tamanuki,1927)
    Female adult
    Mt.Apodiake (SC-Kokkaido),Japan 12/06/1994
    Copyright © Alain Mollard
    Carabus (Acoptolabrus) gehinii munakatai (as nishijimai Imura, 1991)
    Carabus (Acoptolabrus) gehinii munakatai (as nishijimai Imura, 1991)
    Female adult
    Mt. Obira-yama (Shimamaki) Japan, 12/07/1989
    Copyright © Alain Mollard
    Carabus (Acoptolabrus) gehinii munakatai (as okikulumi Nakajima,1993)
    Carabus (Acoptolabrus) gehinii munakatai (as okikulumi Nakajima,1993)
    Male adult
    Mt.Otobe-dake (Yamakoshi) Japan,07/06/1999
    Copyright © Alain Mollard
    Carabus (Acoptolabrus) gehinii munakatai (as ninetopa Nakajiima, 1993)
    Carabus (Acoptolabrus) gehinii munakatai (as ninetopa Nakajiima, 1993)
    Male adult
    Shimamaki-gun (SW-Hokkaido) Japan, 12/07/1999.
    Copyright © Alain Mollard
    Carabus (Acoptolabrus) gehinii munakatai (as furumii Mandl, 1982)
    Carabus (Acoptolabrus) gehinii munakatai (as furumii Mandl, 1982)
    Male adult
    Shiribeshi (Oshima) Japan, 23/07/1999.
    Copyright © Alain Mollard
    Carabus (Megodontus) violaceus violaceus trans violaceus purpurascens Fabricius,1787
    Carabus (Megodontus) violaceus violaceus trans violaceus purpurascens Fabricius,1787
    Male+Female adult
    E-Netherlands, Gelderland, Winterswijk Woold env.,15.II.2015, H.de Haan leg.
    Copyright © Ruslan Panin
    Carabus (Procrustes) coriaceus coriaceus Linne, 1758
    Carabus (Procrustes) coriaceus coriaceus Linne, 1758
    Female adult
    Belgique : Prov. Luxembourg, Harnoncourt 31.vii.1975 - M. Grotz Leg. (Size : 36 mm)
    Copyright © Noël Mal
    Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896
    Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896
    Habitat
    Andorra, Anyos, ii.2012, J. Nicolas Leg. & Photos
    Copyright © Noël Mal
    Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896
    Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896
    Male adult
    Andorra, Leg. Gerts, vi.1966, 32 mm
    Copyright © Noël Mal
    Carabus (Procerus) syriacus kozloviorum Cavazzuti, 2014
    Carabus (Procerus) syriacus kozloviorum Cavazzuti, 2014 Holotype
    Male adult
    Turkey. Nigde. Ulukisla
    Copyright © Anton Kozlov
    Carabus (Procerus) syriacus kozloviorum Cavazzuti, 2014
    Carabus (Procerus) syriacus kozloviorum Cavazzuti, 2014 Paratype
    Female adult
    Turkey. Nigde. Ulukisla
    Copyright © Anton Kozlov
    Carabus (Procerus) syriacus ludademae Cavazzuti & Kozlov, 2014
    Carabus (Procerus) syriacus ludademae Cavazzuti & Kozlov, 2014 Holotype
    Male adult
    Prov. Karaman : circa 10 km a Ovest di Karaman IV.2014. (in coll A. Kozlov)
    Copyright © Anton Kozlov
    Carabus (Procerus) syriacus ludademae Cavazzuti & Kozlov, 2014
    Carabus (Procerus) syriacus ludademae Cavazzuti & Kozlov, 2014 Paratype
    Female adult
    Prov. Karaman : circa 10 km a Ovest di Karaman IV.2014. (in coll A. Kozlov)
    Copyright © Anton Kozlov
    Carabus (Procerus) scabrosus antonkozlovi Cavazzuti, 2014
    Carabus (Procerus) scabrosus antonkozlovi Cavazzuti, 2014 Paratype
    Female adult
    Tavush prov., Ditavan vill. 25-29.VII.2013 (in coll Kozlov)
    Copyright © Anton Kozlov
    Carabus (Procerus) scabrosus antonkozlovi Cavazzuti, 2014
    Carabus (Procerus) scabrosus antonkozlovi Cavazzuti, 2014 Holotype
    Male adult
    Tavush prov., Mehrab vill. 950-1005m (in coll Kozlov). 44 mm
    Copyright © Anton Kozlov
    Carabus (Apotomopterus) penelope penelope Kleinfeld, 1997
    Carabus (Apotomopterus) penelope penelope Kleinfeld, 1997 Paratype
    Male adult
    Chine, C-Fujian: SE of Yong’an, Mt.Tianbaoyan, 25°58’ N / 117°22’ E, h = 1100 m, VII.1997, leg. Wong
    Copyright © Igor Solodovnikov
    Carabus (Apotomopterus) penelope penelope Kleinfeld, 1997
    Carabus (Apotomopterus) penelope penelope Kleinfeld, 1997 Paratype
    Female adult
    Chine, C-Fujian: SE of Yong’an, Mt.Tianbaoyan, 25°58’ N / 117°22’ E, h = 1100 m, VII.1997, leg. Wong
    Copyright © Igor Solodovnikov
    Carabus (Procerus) scabrosus tatianagorokhovae Cavazzuti & Kozlov, 2014
    Carabus (Procerus) scabrosus tatianagorokhovae Cavazzuti & Kozlov, 2014 Holotype
    Male adult
    Armenia. Lori prov. Akhtala
    Copyright © Anton Kozlov
    Carabus (Oreocarabus) luetgensi luetgensi
    Carabus (Oreocarabus) luetgensi luetgensi
    Female adult
    Maetzu, Alava, Spain. det. A. Mollard. Copyright © Magellanes
    Copyright © - - -
    Carabus (Oreocarabus) luetgensi getschmanni Lapouge, 1924
    Carabus (Oreocarabus) luetgensi getschmanni Lapouge, 1924
    Male adult
    Pto.de Opakua, Alava, Spain. det. A. Mollard. Copyright © Magellanes
    Copyright © - - -
    Carabus (Oreocarabus) luetgensi pseudosteuarti Lapouge, 1924
    Carabus (Oreocarabus) luetgensi pseudosteuarti Lapouge, 1924
    Male adult
    Sierra de Estrela, Guarda, Portugal. det. A. Mollard. Copyright © Magellanes
    Copyright © - - -
    Carabus (Oreocarabus) ghiliani La Ferte-Senectere, 1847
    Carabus (Oreocarabus) ghiliani La Ferte-Senectere, 1847
    Male adult
    Pto.de Navafria, Madrid, Spain. det.A.Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Oreocarabus) ghiliani (as negrei Breuning, 1966)
    Carabus (Oreocarabus) ghiliani (as negrei Breuning, 1966)
    Male adult
    Sierra de Gredos, Avila, Spain. det.A.Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Oreocarabus) luetgensi luetgensi (f. typ.)
    Carabus (Oreocarabus) luetgensi luetgensi (f. typ.)
    Female adult
    Sierra de Marao, Vila Real, Portugal; det. A. Mollard. Copyright © Magellanes
    Copyright © - - -
    Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847
    Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847
    Female adult
    los Santos, Salamanque, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847
    Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847
    Male adult
    Villamadana, Palencia, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Oreocarabus) guadarramus (as duerensis Raynaud, 1974)
    Carabus (Oreocarabus) guadarramus (as duerensis Raynaud, 1974)
    Female adult
    Burgo de Osma, Soria, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Oreocarabus) guadarramus (as duerensis Raynaud, 1974)
    Carabus (Oreocarabus) guadarramus (as duerensis Raynaud, 1974)
    Male adult
    Sierra de Cucalon, Teruel, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Oreocarabus) guadarramus (as andiaensis Forel & Leplat, 1998)
    Carabus (Oreocarabus) guadarramus (as andiaensis Forel & Leplat, 1998)
    Male adult
    Sierra de Andia, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Oreocarabus) guadarramus (as guadalupensis Mollard, 2003)
    Carabus (Oreocarabus) guadarramus (as guadalupensis Mollard, 2003) Paratype
    Male adult
    S.de Guadalupe, Extramadure, Spain. det.A.Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Oreocarabus) guadarramus (as sagranus)
    Carabus (Oreocarabus) guadarramus (as sagranus)
    Female adult
    Sierra de la Sagra, Granada, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Oreocarabus) guadarramus (as hirschfelderi Branes, 1987)
    Carabus (Oreocarabus) guadarramus (as hirschfelderi Branes, 1987)
    Male adult
    Puerto Viejo, Salamanque, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) problematicus inflatus (as solidus)
    Carabus (Mesocarabus) problematicus inflatus (as solidus)
    Female adult
    Ripoll, Barcelone, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) problematicus planiusculus (as xaxarsi)
    Carabus (Mesocarabus) problematicus planiusculus (as xaxarsi)
    Male adult
    Sant Sadurni, Gerone, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) problematicus planiusculus Haury in Gehin, 1885
    Carabus (Mesocarabus) problematicus planiusculus Haury in Gehin, 1885
    Male adult
    La Seo de Urgel, Gerone, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) problematicus planiusculus (as bepmalei Lapouge, 1904)
    Carabus (Mesocarabus) problematicus planiusculus (as bepmalei Lapouge, 1904)
    Female adult
    Pic de Bassibes, Huesca, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) problematicus planiusculus (as navarrensis Breuning, 1933)
    Carabus (Mesocarabus) problematicus planiusculus (as navarrensis Breuning, 1933)
    Male adult
    Mt.Urculo, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) problematicus planiusculus (as trapeti Bleuse, 1885)
    Carabus (Mesocarabus) problematicus planiusculus (as trapeti Bleuse, 1885)
    Female adult
    Panticosa, Huesca, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) problematicus planiusculus (as lagardei Mollard, 1996)
    Carabus (Mesocarabus) problematicus planiusculus (as lagardei Mollard, 1996) Paratype
    Female adult
    Sierra de Andia, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) problematicus planiusculus (as andorranus Lapouge, 1902)
    Carabus (Mesocarabus) problematicus planiusculus (as andorranus Lapouge, 1902)
    Female adult
    Col d'envalira, Pas de la Case, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Chaetocarabus) arcadicus arcadicus Gistl, 1850
    Carabus (Chaetocarabus) arcadicus arcadicus Gistl, 1850
    Male adult
    Graecia, Pindus, Mt. Velouchi (Timfristos), 1200m, vi.1970-A. Sermet Leg., 30 mm
    Copyright © Noël Mal
    Carabus (Mesocarabus) lusitanicus lusitanicus (as vieirae Oliveira, 1875)
    Carabus (Mesocarabus) lusitanicus lusitanicus (as vieirae Oliveira, 1875)
    Male adult
    Guia, Leiria, Portugal. det. A. Anichtchenko; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus lusitanicus (as schaumi Gaubil, 1849)
    Carabus (Mesocarabus) lusitanicus lusitanicus (as schaumi Gaubil, 1849)
    Male adult
    S.de Gata, Extramadure, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus castilianus (as fuentei Breuning, 1926)
    Carabus (Mesocarabus) lusitanicus castilianus (as fuentei Breuning, 1926)
    Male adult
    Retortillo, Salamanque, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus castilianus (as bolivari Breuning, 1926)
    Carabus (Mesocarabus) lusitanicus castilianus (as bolivari Breuning, 1926)
    Male adult
    Ledrada, Salamanque, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826
    Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826
    Female adult
    Villasdardo, Salamanque, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus castilianus (as sayagoensis Mollard 2006)
    Carabus (Mesocarabus) lusitanicus castilianus (as sayagoensis Mollard 2006) Paratype
    Male adult
    Moral de Sayago, Zamora, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus brevis (as olmedoensis Mollard, 2007)
    Carabus (Mesocarabus) lusitanicus brevis (as olmedoensis Mollard, 2007) Paratype
    Female adult
    Olmedo, Valladolid, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826
    Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826
    Male adult
    S.de Candelario, Salamanca ("Avila" error). Spain. det. A. Anichtchenko, 2016; (in Mollard, 2013 as lusitanicus brevis) Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus antiquus Dejean, 1826
    Carabus (Mesocarabus) lusitanicus antiquus Dejean, 1826
    Female adult
    Portugal, Ribatejo, Coruche, 13.x.1974, A. Zuzarte leg., 25 mm, Coll. N. Mal
    Copyright © Noël Mal
    Carabus (Mesocarabus) lusitanicus latus (as molossoides Lassalle, 1984)
    Carabus (Mesocarabus) lusitanicus latus (as molossoides Lassalle, 1984)
    Male adult
    Cabezabellosa, Extramadure, det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus gougeleti (as nevadensis Brañes, 1989)
    Carabus (Mesocarabus) lusitanicus gougeleti (as nevadensis Brañes, 1989) Paratype
    Male adult
    Guadix, Granada, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus gougeleti Reiche, 1863
    Carabus (Mesocarabus) lusitanicus gougeleti Reiche, 1863
    Female adult
    S.Norte de Sevilla, Huelva, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus antiquus Dejean, 1826
    Carabus (Mesocarabus) lusitanicus antiquus Dejean, 1826
    Male adult
    Aliseda, Extramadure, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus helluo Dejean, 1826
    Carabus (Mesocarabus) lusitanicus helluo Dejean, 1826
    Female adult
    Puerto de Tortosa, Tarragone, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus trabuccarius Fairmaire, 1857
    Carabus (Mesocarabus) lusitanicus trabuccarius Fairmaire, 1857
    Female adult
    Gaia, Barcelona, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Apotomopterus) vitalisi phoopanensis Imura, 2000
    Carabus (Apotomopterus) vitalisi phoopanensis Imura, 2000
    Male adult
    NE-Laos: Xam Hua Prov.: Mt. Phoo-Pan (Phu Phan), N-slope, h = 2060 m (TOPOTYPUS)
    Copyright © Igor Solodovnikov
    Carabus (Apotomopterus) vitalisi phoopanensis Imura, 2000
    Carabus (Apotomopterus) vitalisi phoopanensis Imura, 2000
    Female adult
    NE-Laos: Xam Hua Prov.: Mt. Phoo-Pan (Phu Phan), N-slope, h = 2060 m (TOPOTYPUS)
    Copyright © Igor Solodovnikov
    Carabus (Mesocarabus) lusitanicus trabuccarius (as galardonensis Meyer & Mollard, 1996)
    Carabus (Mesocarabus) lusitanicus trabuccarius (as galardonensis Meyer & Mollard, 1996) Paratype
    Female adult
    Puerto de Monrepos, Huesca, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Morphocarabus) mestscherjakovi zherichini Shilenkov, 1990
    Carabus (Morphocarabus) mestscherjakovi zherichini Shilenkov, 1990 Paratype
    Female adult
    RUSSIA, Krasnoyarsk Terr., SW Tajmyr Pen., Nizhnyaya Agapa Riv., 40 km of source, south tundra, plakor, brooks holmme, on sandy soil, 7.VIII.1973 (V.V. Zherikhin).
    Copyright © Roman Dudko
    Carabus (Morphocarabus) mestscherjakovi zherichini Shilenkov, 1990
    Carabus (Morphocarabus) mestscherjakovi zherichini Shilenkov, 1990
    Label

    Copyright © Roman Dudko
    Carabus (Mesocarabus) cantabricus moroderi (as magdalenai Meyer & Mollard, 1996)
    Carabus (Mesocarabus) cantabricus moroderi (as magdalenai Meyer & Mollard, 1996) Paratype
    Female adult
    Pto. de Magdalena, Leon, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) cantabricus moroderi Breuning, 1926
    Carabus (Mesocarabus) cantabricus moroderi Breuning, 1926
    Male adult
    Foncebadon, Leon, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) cantabricus moroderi (as almanzaensis Meyer & Mollard in Deuve, 2003)
    Carabus (Mesocarabus) cantabricus moroderi (as almanzaensis Meyer & Mollard in Deuve, 2003) Paratype
    Female adult
    Almaza, Leon, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) breuningi Csiki, 1927
    Carabus (Mesocarabus) breuningi Csiki, 1927
    Male adult
    Villanueva, Orense, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) cantabricus cantabricus (as asturicus Born, 1925)
    Carabus (Mesocarabus) cantabricus cantabricus (as asturicus Born, 1925)
    Female adult
    Pravia, Asturies, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) cantabricus cantabricus (as liebei Eidam, 1925)
    Carabus (Mesocarabus) cantabricus cantabricus (as liebei Eidam, 1925)
    Male adult
    Fonsagrada, Galice, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) cantabricus cantabricus Chevrolat, 1840 (loc. typ.)
    Carabus (Mesocarabus) cantabricus cantabricus Chevrolat, 1840 (loc. typ.)
    Female adult
    La Caniza, Galice, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) cantabricus moroderi (as joaquini Breuning & Toulgoet, 1974)
    Carabus (Mesocarabus) cantabricus moroderi (as joaquini Breuning & Toulgoet, 1974)
    Male adult
    Pena Ubina, Leon, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) cantabricus amayensis Lassalle, 1984
    Carabus (Mesocarabus) cantabricus amayensis Lassalle, 1984
    Male adult
    Pto.de Amaya, Burgos, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) macrocephaloides (Jeanne, 1972)
    Carabus (Mesocarabus) macrocephaloides (Jeanne, 1972)
    Female adult
    Sierra de Andia, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) macrocephaloides (Jeanne, 1972)
    Carabus (Mesocarabus) macrocephaloides (Jeanne, 1972)
    Female adult
    Sierra de Andia, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) macrocephaloides (Jeanne, 1972)
    Carabus (Mesocarabus) macrocephaloides (Jeanne, 1972)
    Female adult
    Sierra de Andia, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) macrocephaloides (Jeanne, 1972)
    Carabus (Mesocarabus) macrocephaloides (Jeanne, 1972)
    Female adult
    Sierra de Andia, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) riffensis Fairmaire, 1875
    Carabus (Mesocarabus) riffensis Fairmaire, 1875
    Female adult
    Morocco : Rif, Ketama vi.1980 - V. Oliviella Leg. (Size : 33 mm)
    Copyright © Noël Mal
    Carabus (Tachypus) cancellatus intermedius Dejean, 1826
    Carabus (Tachypus) cancellatus intermedius Dejean, 1826
    Female adult
    NW -Serbia,Fruška Gora,Vrdnik env. h=380m, 9-10.VII.2013, M. Janalik leg. L=29 mm
    Copyright © Ruslan Panin
    Carabus (Carabus) arvensis carpathus Born, 1902
    Carabus (Carabus) arvensis carpathus Born, 1902
    Male adult
    W Ukraine, NE Carpathians,Transcarpathian reg., Vihorlat, near Strychava vill., Mt Knyaginytsya h= 500 m, 18.V-22.VI. 2014, Yu.Kanarsky leg. L=18 mm
    Copyright © Ruslan Panin
    Carabus (Carabus) arvensis arvensis Herbst, 1784
    Carabus (Carabus) arvensis arvensis Herbst, 1784
    Female adult
    NW Ukraine, Polesia, Rovno reg., Rokytne distr., near Bil’sk vill., 4-6.VII.2015, Panin R. leg. L=20 mm
    Copyright © Ruslan Panin
    Carabus (Procerus) scabrosus tauricus Bonelli, 1811
    Carabus (Procerus) scabrosus tauricus Bonelli, 1811
    Male adult
    Crimea, Sevastopol, leg.det. A.Chuvilin, 12.VIII.1999. Photo: U.Schmidt, 2016. 40mm
    Copyright © Udo Schmidt
    Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
    Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
    Female adult
    Sierra de Aralar, Navarra, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) cantabricus brabeus (as barcelecoanus)
    Carabus (Mesocarabus) cantabricus brabeus (as barcelecoanus)
    Male adult
    Col de Velate, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) cantabricus brabeus (as barcelecoanus)
    Carabus (Mesocarabus) cantabricus brabeus (as barcelecoanus)
    Female adult
    Col de Velate, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
    Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
    Male adult
    Col de Velate, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Coptolabrus) augustus ignigena Hauser, 1914
    Carabus (Coptolabrus) augustus ignigena Hauser, 1914
    Female adult
    China : Guizhoh, Sinan vi.2006 (Size : 35 mm) Coll. N. Mal
    Copyright © Noël Mal
    Carabus (Coptolabrus) ignimitella antaeus Hauser, 1914 (as guidongicus Cavazzuti & Ratti, 1998)
    Carabus (Coptolabrus) ignimitella antaeus Hauser, 1914 (as guidongicus Cavazzuti & Ratti, 1998)
    Female adult
    China : Jianxi, Jing Gangshan City 10-18.vi.2007 - J. Li Leg. (Size : 42 mm) Coll. N. Mal
    Copyright © Noël Mal
    Carabus (Coptolabrus) pustulifer shangzhouicus Deuve & Li, 2003
    Carabus (Coptolabrus) pustulifer shangzhouicus Deuve & Li, 2003
    Female adult
    China : Shaanxi, Shazhou City, Shahezi 10.vii.2007 - J. Li Leg. (Size : 42 mm), Coll. N. Mal
    Copyright © Noël Mal
    Carabus (Mesocarabus) lusitanicus brevis (as segovianus Mollard, 2011)
    Carabus (Mesocarabus) lusitanicus brevis (as segovianus Mollard, 2011) Paratype
    Male adult
    S.de Gorda, Soria, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus gougeleti (as alcaracinus)
    Carabus (Mesocarabus) lusitanicus gougeleti (as alcaracinus)
    Female adult
    Alcaraz, Albacete, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus helluo (as alicantinus)
    Carabus (Mesocarabus) lusitanicus helluo (as alicantinus)
    Female adult
    S.de Bernia, Alicante, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus trabuccarius (as leyrensis)
    Carabus (Mesocarabus) lusitanicus trabuccarius (as leyrensis)
    Female adult
    Sierra de Leyre, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus trabuccarius (as trypalsi)
    Carabus (Mesocarabus) lusitanicus trabuccarius (as trypalsi)
    Male adult
    Plan du Trypals, Huesca, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus trabuccarius (as solanensis)
    Carabus (Mesocarabus) lusitanicus trabuccarius (as solanensis)
    Male adult
    S.de Bolave, Huesca, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus trabuccarius (as troncedoensis)
    Carabus (Mesocarabus) lusitanicus trabuccarius (as troncedoensis) Paratype
    Female adult
    Troncedo, Huesca, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus trabuccarius (as benabarrensis)
    Carabus (Mesocarabus) lusitanicus trabuccarius (as benabarrensis) Paratype
    Male adult
    Benabarre, Huesca, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus brevis (as pironensis)
    Carabus (Mesocarabus) lusitanicus brevis (as pironensis) Paratype
    Female adult
    Pennasrubias, Segovia, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus brevis (as lermaensis)
    Carabus (Mesocarabus) lusitanicus brevis (as lermaensis) Paratype
    Female adult
    Espinosa de Cervera, Burgos, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus brevis (as logronicus)
    Carabus (Mesocarabus) lusitanicus brevis (as logronicus)
    Male adult
    Pto.de Piqueras, La Rioja, Spain. det. A. Mollard; Copyright © Magellanes
    Copyright © - - -
    Carabus (Mesocarabus) lusitanicus trabuccarius (as catalonicus)
    Carabus (Mesocarabus) lusitanicus trabuccarius (as catalonicus)
    Male adult
    Ft.de Roquerolles, Prades, Tarragone, SPAIN
    Copyright © Alain Mollard
    Carabus (Mesocarabus) lusitanicus brevis (as segovianus)
    Carabus (Mesocarabus) lusitanicus brevis (as segovianus)
    Male adult
    Sierra de Pardos, South Calatayud, Saragosse, SPAIN
    Copyright © Alain Mollard
    Carabus (Mesocarabus) lusitanicus helluo (as albarracinus)
    Carabus (Mesocarabus) lusitanicus helluo (as albarracinus)
    Female adult
    Altos de Cabrera, Pineda de Ciguela, Cuenca, SPAIN
    Copyright © Alain Mollard
    Carabus (Mesocarabus) riffensis Fairmaire, 1875 (as dominicae)
    Carabus (Mesocarabus) riffensis Fairmaire, 1875 (as dominicae)
    Male adult
    Ft.de Ketama , Rif, MAROC
    Copyright © Alain Mollard
    Carabus (Mesocarabus) lusitanicus cf. antiquus-gougeleti
    Carabus (Mesocarabus) lusitanicus cf. antiquus-gougeleti
    Male adult
    Bicada, South Beja, PORTUGAL
    Copyright © Alain Mollard
    Carabus (Apotomopterus) arrowi hector Hauser, 1924 (as arrogantior)
    Carabus (Apotomopterus) arrowi hector Hauser, 1924 (as arrogantior)
    Female adult
    Yushan, Hunan, CHINA
    Copyright © Alain Mollard
    Carabus (Apotomopterus) arrowi trilobatus Cavazzuti, 1994
    Carabus (Apotomopterus) arrowi trilobatus Cavazzuti, 1994
    Male adult
    Jinsha, Guizhou, CHINA
    Copyright © Alain Mollard
    Carabus (Apotomopterus) arrowi yubeicus Deuve & Tian, 1999
    Carabus (Apotomopterus) arrowi yubeicus Deuve & Tian, 1999
    Female adult
    Guanyang, Guangxi, CHINA
    Copyright © Alain Mollard
    Carabus (Apotomopterus) aeneocupreus Hauser, 1932
    Carabus (Apotomopterus) aeneocupreus Hauser, 1932
    Female adult
    Qiong Lai Shan, Sichuan, CHINA
    Copyright © Alain Mollard
    Carabus (Apotomopterus) prodigus prodigus Erichson, 1834
    Carabus (Apotomopterus) prodigus prodigus Erichson, 1834
    Male adult
    Luofu Shan, Guangdong, CHINA
    Copyright © Alain Mollard
    Carabus (Apotomopterus) yuae arlequinus Deuve, 1994
    Carabus (Apotomopterus) yuae arlequinus Deuve, 1994
    Female adult
    Yuanbao Shan, Guangxi, CHINA
    Copyright © Alain Mollard
    Carabus (Rhigocarabus) allegroi Cavazzuti & Rapuzzi, 2007
    Carabus (Rhigocarabus) allegroi Cavazzuti & Rapuzzi, 2007
    Male adult
    CINA S. Gansu, Qiling Shan, Pass N. Tewo (N 34°16’-E 103°03’), 4.100 m, 17.VI-1.VII.2010 (Colleccion Gianni Allegro)
    Copyright © Gianni Allegro
    Carabus (Chaetocarabus) intricatus krueperi Reitter, 1896
    Carabus (Chaetocarabus) intricatus krueperi Reitter, 1896
    Male+Female adult
    Graecia : Mt. Ossa, 1000m, 31.x.1975, Leg P. Ratti, Coll. N. Mal
    Copyright © Noël Mal
    Carabus (Megodontus) violaceus exasperatus Duftschmid, 1812
    Carabus (Megodontus) violaceus exasperatus Duftschmid, 1812
    Male adult
    Hungary, Budapest, Mt Buda, Harsbokor-hill,16-19.V.1992, I.Retezár leg., L=27 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) odoratus baeri Menetries, 1851: 45
    Carabus (Morphocarabus) odoratus baeri Menetries, 1851: 45
    Male adult
    Russia, SW Taymyr, near Talnakh, 25 km N Norilsk, hilly tundra, hillside overgrown with alder slate, dwarfish birches, blueberry, h = 150-250 м, 24-28.06.2003, leg. S.Saluk
    Copyright © Igor Solodovnikov
    Carabus (Morphocarabus) odoratus baeri Menetries, 1851: 45
    Carabus (Morphocarabus) odoratus baeri Menetries, 1851: 45
    Endophallus
    Russia, SW Taymyr, near Talnakh, 25 km N Norilsk, hilly tundra, hillside overgrown with alder slate, dwarfish birches, blueberry, h = 150-250 м, 24-28.06.2003, leg. S.Saluk
    Copyright © Igor Solodovnikov
    Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
    Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
    Endophallus
    NE-Hungary, 4 km S Sátoraljaújhely, right bank riv. Bodrog (Long-erdő),
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
    Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
    Male adult
    NE-Hungary, 4 km S Sátoraljaújhely, right bank riv. Bodrog (Long-erdő), 8.VI.1992, leg./det. I.Retezár leg. L=28 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi distinguendus Csiki, 1906
    Carabus (Morphocarabus) rothi distinguendus Csiki, 1906
    Endophallus
    NC-Hungary: E- part of Budapest,Cinkota,
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi distinguendus Csiki, 1906
    Carabus (Morphocarabus) rothi distinguendus Csiki, 1906
    Male adult
    NC-Hungary: E- part of Budapest, Cinkota,28.VIII-17.IX.1980, I.Retezár leg. L=27 mm
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) problematicus problematicus Herbst, 1786
    Carabus (Mesocarabus) problematicus problematicus Herbst, 1786
    Female adult
    W-Hungary, NE-Alps, Mts. Kőszeg, Velem env., Borha-forrás, 26.VII.1981, Gy. Somorijai leg. L=26 mm
    Copyright © Ruslan Panin
    Carabus (Pachycarabus) swaneticus kvirensis Belousov, 1992
    Carabus (Pachycarabus) swaneticus kvirensis Belousov, 1992
    Male adult
    SC-Caucasus: NW-Georgia, Mingrelia, Egrissky Mt.Rng., h~1800-2000 m, 23-26.VII.1990, I.Retezár leg. L=20 mm
    Copyright © Ruslan Panin
    Carabus (Pachystus) cavernosus rozneri Retezár & Gy. Szél, 2013
    Carabus (Pachystus) cavernosus rozneri Retezár & Gy. Szél, 2013 Paratype
    Male adult
    Romania, Apuseni : Trascău Mts. L=27 mm
    Copyright © Ruslan Panin
    Carabus (Pachystus) cavernosus rozneri Retezár & Gy. Szél, 2013
    Carabus (Pachystus) cavernosus rozneri Retezár & Gy. Szél, 2013 Paratype
    Label

    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi distinguendus (as bieneri Mandl, 1965)
    Carabus (Morphocarabus) rothi distinguendus (as bieneri Mandl, 1965)
    Endophallus
    C-Slovakia, Detva Distr., Stara Huta env. h=700-800m,
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi distinguendus (as bieneri Mandl, 1965)
    Carabus (Morphocarabus) rothi distinguendus (as bieneri Mandl, 1965)
    Male adult
    C-Slovakia, Detva Distr., Stara Huta env. h=700-800m, 3.VI.-5.VII.2012, V.Slovák lgt. L=29 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi distinguendus (as subparvulus Mandl, 1965)
    Carabus (Morphocarabus) rothi distinguendus (as subparvulus Mandl, 1965)
    Endophallus
    NC-Hungary:Borzsony Mts, Nograd-Diosjeno,
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) rothi distinguendus (as subparvulus Mandl, 1965)
    Carabus (Morphocarabus) rothi distinguendus (as subparvulus Mandl, 1965)
    Male+Female adult
    NC-Hungary:Borzsony Mts, Nograd-Diosjeno,1-14.V.2004, Krause Rene leg. female, 32 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859
    Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859
    Female adult
    Russia, SE Siberia, N Chita reg., Kodar Mt. Range, Big Leprindo Lake env., h~1800 m., L=21 mm
    Copyright © Ruslan Panin
    Carabus (Procerus) scabrosus nakagomei Cavazzuti & Kozlov, 2016
    Carabus (Procerus) scabrosus nakagomei Cavazzuti & Kozlov, 2016 Paratype
    Female adult

    Copyright © Anton Kozlov
    Carabus (Procerus) scabrosus nakagomei Cavazzuti & Kozlov, 2016
    Carabus (Procerus) scabrosus nakagomei Cavazzuti & Kozlov, 2016 Holotype
    Male adult

    Copyright © Anton Kozlov
    Carabus (Meganebrius) deliae colasianus (Morvan & Mandl, 1974)
    Carabus (Meganebrius) deliae colasianus (Morvan & Mandl, 1974) Holotype
    Male adult

    Copyright © Igor Solodovnikov
    Carabus (Meganebrius) deliae colasianus (Morvan & Mandl, 1974)
    Carabus (Meganebrius) deliae colasianus (Morvan & Mandl, 1974)
    Male adult

    Copyright © - - -
    Carabus (Apotomopterus) tonkinensis tonkinensis Deuve, 1990
    Carabus (Apotomopterus) tonkinensis tonkinensis Deuve, 1990
    Male adult
    North Vietnam, Tû Lé, (Yen Bai Prov.) 1200 m
    Copyright © Alain Mollard
    Carabus (Apotomopterus) tonkinensis jinpingensis Deuve & Tian, 2001
    Carabus (Apotomopterus) tonkinensis jinpingensis Deuve & Tian, 2001
    Female adult
    Ailao Shan : Jinping Xian (Yunnan) China
    Copyright © Alain Mollard
    Carabus (Procerus) syriacus bulgharmaadensis Bodemeyer, 1915 (as toulgoetianus)
    Carabus (Procerus) syriacus bulgharmaadensis Bodemeyer, 1915 (as toulgoetianus)
    Female adult
    North Gulek (Prov.Adana), South Turquey.
    Copyright © Alain Mollard
    Carabus (Procerus) syriacus limitaneus (Korell, 1989)
    Carabus (Procerus) syriacus limitaneus (Korell, 1989)
    Male adult
    Daret’Azze; North-West Syria
    Copyright © Alain Mollard
    Carabus (Apotomopterus) mecynodes mecynodes Bates, 1890
    Carabus (Apotomopterus) mecynodes mecynodes Bates, 1890
    Female adult
    Wa-shan, Chengdu (Sichuan) China
    Copyright © Alain Mollard
    Carabus (Apotomopterus) mecynodes combustus Cavazzuti & Ratti, 1999
    Carabus (Apotomopterus) mecynodes combustus Cavazzuti & Ratti, 1999
    Female adult
    Dabei Shan (Sichuan) China
    Copyright © Alain Mollard
    Carabus (Apotomopterus) mecynodes lilianae Cavazzuti, 1997
    Carabus (Apotomopterus) mecynodes lilianae Cavazzuti, 1997
    Male adult
    Huidong Xian (Sichuan) China
    Copyright © Alain Mollard
    Carabus (Meganebrius) deliae deliae (Morvan, 1972)
    Carabus (Meganebrius) deliae deliae (Morvan, 1972) Holotype
    Male adult

    Copyright © - - -
    Carabus (Meganebrius) deliae deliae (Morvan, 1972)
    Carabus (Meganebrius) deliae deliae (Morvan, 1972) Holotype
    Label

    Copyright © - - -
    Carabus (Meganebrius) quinlani quinlani Mandl, 1965
    Carabus (Meganebrius) quinlani quinlani Mandl, 1965 Holotype
    Male adult

    Copyright © - - -
    Carabus (Meganebrius) koganae koganae Colas, 1961
    Carabus (Meganebrius) koganae koganae Colas, 1961 Holotype
    Male adult

    Copyright © Igor Solodovnikov
    Carabus (Meganebrius) lebretae lebretae Colas, 1961
    Carabus (Meganebrius) lebretae lebretae Colas, 1961 Holotype
    Male adult

    Copyright © - - -
    Carabus (Mesocarabus) leptopus Thomson, 1875
    Carabus (Mesocarabus) leptopus Thomson, 1875 Holotype
    Female adult
    3957:2 (MZLU)
    Copyright © Lund University
    Carabus (Mesocarabus) leptopus
    Carabus (Mesocarabus) leptopus
    Label

    Copyright © Lund University
    Carabus (Eucarabus) obsoletus obsoletus var sacheri Thomson, 1875
    Carabus (Eucarabus) obsoletus obsoletus var sacheri Thomson, 1875 Syntypes
    Male adult
    3958:2 (MZLU)
    Copyright © Lund University
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as aragonensis Lapouge,1913)
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as aragonensis Lapouge,1913)
    Male adult
    Male adult ,Sierra de Montseny (Barcelone) Spain.
    Copyright © Alain Mollard
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as frontanyaensis Mollard,1999)
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as frontanyaensis Mollard,1999)
    Female adult
    N of Boreda, (Lerida) Spain.
    Copyright © Alain Mollard
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as brevicollis Lapouge,1910)
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as brevicollis Lapouge,1910)
    Female adult
    Corbières (Aude) France.
    Copyright © Alain Mollard
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as brevicollis var. viridis)
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as brevicollis var. viridis)
    Female adult
    Rennes les bains (Aude) France.
    Copyright © Alain Mollard
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as delacrei Tarrier,1975)
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as delacrei Tarrier,1975)
    Female adult
    Nostra Signora de Queralt, Berga (Barcelona) Spain.
    Copyright © Alain Mollard
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as occideneissimus Meyer,1995)
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as occideneissimus Meyer,1995)
    Male adult
    Collado de Boixols, (Lérida) Spain.
    Copyright © Alain Mollard
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as urgelensis Mollard,1999)
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as urgelensis Mollard,1999)
    Female adult
    Sierra del Cadi,Toloriu, (Lérida) Spain.
    Copyright © Alain Mollard
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as jeannei)
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as jeannei)
    Female adult
    Sierra de la Mussara (Tarragona) Spain.
    Copyright © Alain Mollard
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as merangensis Mollard,1999)
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as merangensis Mollard,1999)
    Male adult
    Méranges, near Puigcerda, (Catalunya) Spain
    Copyright © Alain Mollard
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as monserraticus Csiki,1917)
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as monserraticus Csiki,1917)
    Female adult
    Sierra de Montserrat, (Barcelone) Spain
    Copyright © Alain Mollard
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as palensis Mollard,1999)
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as palensis Mollard,1999)
    Male adult
    Col de N’hug, (N-Catalunya) Spain.
    Copyright © Alain Mollard
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as quillansis Mollard,1999)
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as quillansis Mollard,1999)
    Male adult
    Granès (Aude) France.
    Copyright © Alain Mollard
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (f. typ.)
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (f. typ.)
    Female adult
    Prats de Mollo (Pyr.Or) France
    Copyright © Alain Mollard
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as soredensis Mollard,1999)
    Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as soredensis Mollard,1999)
    Male adult
    Ft.de Sorède, Mts. Albères (Pyr.Or.) France
    Copyright © Alain Mollard
    Carabus (Chrysocarabus) rutilans opulentus Oberthür, 1898
    Carabus (Chrysocarabus) rutilans opulentus Oberthür, 1898
    Female adult
    East-Sierra de Aigues-Tortes, Tahull,Bohi (Lerida) Spain
    Copyright © Alain Mollard
    Carabus (Chrysocarabus) rutilans opulentus Oberthür, 1898 (as alinsis, Mollard 1999)
    Carabus (Chrysocarabus) rutilans opulentus Oberthür, 1898 (as alinsis, Mollard 1999)
    Male adult
    Alins
    Copyright © Alain Mollard
    Carabus (Chrysocarabus) rutilans opulentus Oberthür, 1898 (as turbonensis, Mollard 1999)
    Carabus (Chrysocarabus) rutilans opulentus Oberthür, 1898 (as turbonensis, Mollard 1999)
    Male adult
    Massif du Turbon, (Huesca) Spain
    Copyright © Alain Mollard
    Carabus (Chrysocarabus) rutilans opulentus Oberthür, 1898 (as cotiellensis, Mollard 1999)
    Carabus (Chrysocarabus) rutilans opulentus Oberthür, 1898 (as cotiellensis, Mollard 1999)
    Female adult
    East-Mt. Cotiella (Huesca) Spain
    Copyright © Alain Mollard
    Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896 (as casteillensis, Mollard 1999)
    Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896 (as casteillensis, Mollard 1999)
    Female adult
    Cap del Prat d’Orient,Castells (Lerida) Spain
    Copyright © Alain Mollard
    Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896 (as benesensis, Mollard 1999)
    Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896 (as benesensis, Mollard 1999)
    Female adult
    S-Sierra de Aigues-Tortes, Malpas-Benes
    Copyright © Alain Mollard
    Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896 (as pervesensis, Mollard 1999)
    Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896 (as pervesensis, Mollard 1999)
    Female adult
    Col de Pervers, Sierra de St. Gervas
    Copyright © Alain Mollard
    Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896 (as basetaensis, Mollard 1999)
    Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896 (as basetaensis, Mollard 1999)
    Female adult
    Guils, Sierra de la Baseta (Lerida) Spain
    Copyright © Alain Mollard
    Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896 (as capdellensis, Mollard 1999)
    Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896 (as capdellensis, Mollard 1999)
    Female adult
    La Torre de Capdella (Lerida) Spain
    Copyright © Alain Mollard
    Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896 (as juberriensis, Mollard 1999)
    Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896 (as juberriensis, Mollard 1999)
    Male adult
    Cesters, 1600m, Andorra
    Copyright © Alain Mollard
    Carabus (Lamprostus) erenleriensis erenleriensis Schweiger, 1966
    Carabus (Lamprostus) erenleriensis erenleriensis Schweiger, 1966
    Endophallus
    NW-Turkey:Bolu Prov., Abant Lake
    Copyright © Ruslan Panin
    Carabus (Lamprostus) erenleriensis erenleriensis Schweiger, 1966, loc.typ.
    Carabus (Lamprostus) erenleriensis erenleriensis Schweiger, 1966, loc.typ.
    Male adult
    NW-Turkey:Bolu Prov., Abant Lake, 25.VIII.1990 L=35 mm
    Copyright © Ruslan Panin
    Carabus (Imaibius) stoliczkanus brauni Heinz, 1993
    Carabus (Imaibius) stoliczkanus brauni Heinz, 1993
    Female adult
    NE Pakistan, Azad Kashmir, Bagh Distr., Sudhan-Gali h~2300 m,15.VIII.1994, S Dacatra leg. L=37 mm
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) arrowi yubeicus Deuve & Tian, 1999
    Carabus (Apotomopterus) arrowi yubeicus Deuve & Tian, 1999
    Female adult
    S-China, Guangxi, Jinxiu Co., Mt. Dayao,28.VII.2007, L=40 mm
    Copyright © Ruslan Panin
    Carabus (Ctenocarabus) melancholicus melancholicus Fabricius, 1798
    Carabus (Ctenocarabus) melancholicus melancholicus Fabricius, 1798
    Male adult
    NW Morocco, Kénitra, Souk El Arbaa env., II.1985, V.Olivella leg. L=25 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) concolor lepontinus Born, 1908
    Carabus (Orinocarabus) concolor lepontinus Born, 1908
    Male adult
    NW-Italy, Piemonte (VB), Lepontine Alps, Mt. Zeda , 6.VII.1998, L Diotti leg., L=20 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) concolor concolor Fabricius, 1792
    Carabus (Orinocarabus) concolor concolor Fabricius, 1792
    Female adult
    NW-Italy, Breuil-Cervinia h~2500 m, 2.IX.1971, Georgeon leg. L=21 mm
    Copyright © Ruslan Panin
    Carabus (Macrothorax) morbillosus lampedusae Born, 1925
    Carabus (Macrothorax) morbillosus lampedusae Born, 1925
    Female adult
    AG, Sicily, Island of Lampedusa, leg V. Monzini
    Copyright © Vittorino Monzini
    Carabus (Oxycarabus) saphyrinus pirithous Reitter, 1896
    Carabus (Oxycarabus) saphyrinus pirithous Reitter, 1896
    Male adult
    NW-Turkey,Istanbul (Asian part),Polonezköy env., 2.VI.2009. S. Dacatra leg. L=27 mm
    Copyright © Ruslan Panin
    Carabus (Rhigocarabus) morawitzianus morawitzianus Semenov, 1887
    Carabus (Rhigocarabus) morawitzianus morawitzianus Semenov, 1887
    Male adult
    C China, Qinghai, Zadoi County, Toramarkog h~4600 m, VI.2001, P. Cavazzuti leg. L=17 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) pedemontanus germanae Casale & Cavazzuti, 1977
    Carabus (Orinocarabus) pedemontanus germanae Casale & Cavazzuti, 1977
    Male adult
    NW-Italy, Piemonte, Imperia prov., Mt.Saccarello h~2000m, 30.V.1986. L=21 mm
    Copyright © Ruslan Panin
    Carabus (Procrustes) piochardi labruleriei Gehin, 1883
    Carabus (Procrustes) piochardi labruleriei Gehin, 1883 Lectotype
    Unsexed
    Muséum national d’Histoire naturelle, Paris (France), Specimen MNHN-EC-EC509. http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec509
    Copyright © - - -
    Carabus (Megodontus) violaceus aurolimbatus Dejean, 1829
    Carabus (Megodontus) violaceus aurolimbatus Dejean, 1829 Holotype
    Male adult
    Loc: "Moscow". http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec51 Muséum national d’Histoire naturelle, Paris (France) Collection: Insects - Coleoptera (EC) Specimen MNHN-EC-EC51
    Copyright © - - -
    Carabus (Megodontus) violaceus aurolimbatus Dejean, 1829
    Carabus (Megodontus) violaceus aurolimbatus Dejean, 1829
    Label
    Loc: "Moscow". http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec51 Muséum national d’Histoire naturelle, Paris (France) Collection: Insects - Coleoptera (EC) Specimen MNHN-EC-EC51
    Copyright © - - -
    Carabus (Neoplectes) starckianus olgae Semenov, 1898
    Carabus (Neoplectes) starckianus olgae Semenov, 1898 Lectotype
    Male adult
    [Circassia. Rost. Car. imperatot St.] [44] [Plectes Olgae m. A. Semenov det., ♂ Typ. un. X.96] red label этикетка [LECTOTYPUS Gottwald, 1983 design], yellow label [Zoological Institute Russian Academy of Sciences St Peterburg]
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) starckianus olgae Semenov, 1898
    Carabus (Neoplectes) starckianus olgae Semenov, 1898
    Label

    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) starckianus olgae Semenov, 1898
    Carabus (Neoplectes) starckianus olgae Semenov, 1898 Paralectotypes
    Male adult
    • [Circassien. Rost. [18]98 Pl. obtusus var. 5.] [5] [Plectes Olgae m. A. Semenov det., ♂ I.98] red label [PARALECTOTYPUS Gottwald, 1983 design], yellow label [Zoological Institute Russian Academy of Sciences St Peterburg]
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) starckianus olgae Semenov, 1898
    Carabus (Neoplectes) starckianus olgae Semenov, 1898
    Label

    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) starckianus starckianus Ganglbauer, 1886
    Carabus (Neoplectes) starckianus starckianus Ganglbauer, 1886
    Male adult
    Russia, Krasnodar area, Sochi, Sochi distr., right bank of Riv. Persiyanka, NE slope Phlagokh Mt., 03.05-02.07.2013, Leg. I.A. Solodovnikov, S.V. Solodovnikova (TOPOTYPUS as phlagochensis Imura & Zamotajlov, 1993)
    Copyright © Igor Solodovnikov
    Carabus (Orinocarabus) fairmairei pelvicus Cavazzuti, 1973
    Carabus (Orinocarabus) fairmairei pelvicus Cavazzuti, 1973
    Male adult
    NW-Italy, Cuneo, Maira Vall., Acceglio env., La Colletta h~2800 m, 12.VII.2007, R. Monguzzi leg. L=19 mm
    Copyright © Ruslan Panin
    Carabus (Isiocarabus) miaorum Lassalle & Prunier, 1993
    Carabus (Isiocarabus) miaorum Lassalle & Prunier, 1993
    Male adult
    S-China, N-Guangxi, Mao'er Shan, h~1600 m, V.2009, L= 26 mm
    Copyright © Ruslan Panin
    Carabus (Eucarabus) italicus rostagnoi Luigioni, 1904
    Carabus (Eucarabus) italicus rostagnoi Luigioni, 1904
    Female adult
    C-Italy, Lazio, Fondi env.,10.II.1979, L = 25 mm
    Copyright © Ruslan Panin
    Carabus (Archicarabus) victor victor Fischer, 1836
    Carabus (Archicarabus) victor victor Fischer, 1836
    Female adult
    NE-Turkey, Giresun, Sehitler Geçidi (Pass) h~2000 m, 23.VII.2001, S. Dacatra leg., L=19 mm
    Copyright © Ruslan Panin
    Carabus (Eucarabus) parreyssi parreyssi Palliardi, 1825
    Carabus (Eucarabus) parreyssi parreyssi Palliardi, 1825
    Male adult
    W-Croatia, NP Northern Velebit, Zavižan,15.V-13.VI.1996, Colombetta leg., L=21 mm
    Copyright © Ruslan Panin
    Carabus (Lamprostus) torosus giresuni Breuning, 1964 (as paululus Heinz & Korge, 1964)
    Carabus (Lamprostus) torosus giresuni Breuning, 1964 (as paululus Heinz & Korge, 1964)
    Male adult
    NE-Turkey, Giresun, Sehitler Geçidi (Pass) h~2000 m,24.VII.2001, S. Dacatra leg., L=26 mm
    Copyright © Ruslan Panin
    Carabus (Platycarabus) creutzeri baldensis Schaum, 1857
    Carabus (Platycarabus) creutzeri baldensis Schaum, 1857
    Male adult
    NC-Italy: Trento, Sega di Ala, IX.1999, A. Vicini leg. , L=24 mm
    Copyright © Ruslan Panin
    Carabus (Ophiocarabus) tekesensis Deuve & Tian, 2004
    Carabus (Ophiocarabus) tekesensis Deuve & Tian, 2004
    Female adult
    China, Xinjiang, Narat Mts. NE Tshon-Kushei Korbulak Riv., h~2630 42°56'25"N 82°15'27"E 13.VII.2014 leg. I.Kabak L=19.0 mm
    Copyright © Kirill Makarov
    Carabus (Archicarabus) rossii Dejean, 1826
    Carabus (Archicarabus) rossii Dejean, 1826
    Female adult
    NC Italy, Emilia reg., Bologna, San Martino, 29.V.2016, G..L.Agnoli leg. L=30 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) linnaei folgariacus Born, 1913
    Carabus (Orinocarabus) linnaei folgariacus Born, 1913
    Male adult
    NE-Italy, Trentino-Alto Adige (Südtirol), N of vill. San Michele vill., 13-17.VI.2016, R. Janak leg. L=20 mm
    Copyright © Ruslan Panin
    Carabus (Chrysocarabus) auronitens kraussi Lapouge, 1898
    Carabus (Chrysocarabus) auronitens kraussi Lapouge, 1898
    Female adult
    C-Austria, Styria, Ennstal Alps, Totes Gebirge Mt Rng, Mt Loser h~1100 m, 16-20.VII.2014, R.Janak leg. L=27 mm
    Copyright © Ruslan Panin
    Carabus (Sphodristocarabus) adamsi Adams, 1817
    Carabus (Sphodristocarabus) adamsi Adams, 1817
    Endophallus
    CN-Georgia, N of Tbilisi, NW of Tianeti, Tvalivi env.,
    Copyright © Ruslan Panin
    Carabus (Sphodristocarabus) adamsi Adams, 1817
    Carabus (Sphodristocarabus) adamsi Adams, 1817
    Male adult
    CN-Georgia, N of Tbilisi, NW of Tianeti, Tvalivi env., 20.VI.2015, F. Grycz leg. L=24 mm
    Copyright © Ruslan Panin
    Carabus (Coptolabrus) ignimitella guangxicola Deuve, 1998
    Carabus (Coptolabrus) ignimitella guangxicola Deuve, 1998
    Male adult
    S-China, Guangxi prov., Jingxiu, Mt. Dayao Shan, 1-17.V.2006, L=47 mm
    Copyright © Ruslan Panin
    Carabus (Lamprostus) robustus triumphans Cavazzuti, 1987
    Carabus (Lamprostus) robustus triumphans Cavazzuti, 1987
    Male adult
    NE-Turkey:Trabzon, Tonya env., VIII.2004. L=41 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) latreilleanus Csiki, 1927
    Carabus (Orinocarabus) latreilleanus Csiki, 1927
    Female adult
    NW-Italy, Piemonte, Biella, Biellese Alps Rng, Mt.Camino, 25.VI.1997, V.Monzini leg. L=18 mm
    Copyright © Ruslan Panin
    Carabus (Lamprostus) torosus nordmanni Chaudoir, 1848 (as enricoi )
    Carabus (Lamprostus) torosus nordmanni Chaudoir, 1848 (as enricoi )
    Male adult
    NE-Turkey:Ordu prov.,3 km N Gölköy h~1100 m, VII.2006, L=37 mm
    Copyright © Ruslan Panin
    Carabus (Macrothorax) morbillosus lampedusae Born, 1925
    Carabus (Macrothorax) morbillosus lampedusae Born, 1925
    Male adult
    S Italy, Isl. Lampedusa, Baia dei Conigli, VI.2003, L=29 mm
    Copyright © Ruslan Panin
    Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014
    Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014 Paratype
    Female adult
    Abkhazia, Bzybsky Mt.Rng, right bank of river Reshevie (left tributary of river Bzyb) h~670-720 м, 13-14.V.2010, D.Fominykh & I.Solodovnikov leg. L=41 mm
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) arrowi ghirettii Cavazzuti, 1995
    Carabus (Apotomopterus) arrowi ghirettii Cavazzuti, 1995
    Female adult
    Fanjing Shan, Guizhou, 1900 m, Leg: C.Reuter. L= 42 mm
    Copyright © Alain Mollard
    Carabus (Mesocarabus) breuningi Csiki, 1927
    Carabus (Mesocarabus) breuningi Csiki, 1927
    Female adult
    NW-Spain, Prov. Zamora, San Martín de Castañeda, 5.VII.1984, M.Toribio leg. L=28 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) odoratus kurmaevi I.Solodovnikov, 2017
    Carabus (Morphocarabus) odoratus kurmaevi I.Solodovnikov, 2017 Holotype
    Male adult
    Russia, Siberia, Krasnoyarsk Krai, the North Yenisei district, 55 km of CE of the vill. Bryanka, valley of the river Chirimba, stream Nakhodny, h = 350 m, the mixed forest (birch and pine); on a steep slope along an animal track on a hill brow, 23.VII-20.VIII.2015, leg. A.V. Kurmaev
    Copyright © Igor Solodovnikov
    Carabus (Morphocarabus) odoratus kurmaevi I.Solodovnikov, 2017
    Carabus (Morphocarabus) odoratus kurmaevi I.Solodovnikov, 2017 Paratype
    Male adult
    Russia, Siberia, Krasnoyarsk Krai, the North Yenisei district, 55 km of CE of the vill. Bryanka, valley of the river Chirimba, stream Nakhodny, h = 350 m, the mixed forest (birch and pine); on a steep slope along an animal track on a hill brow, 23.VII-20.VIII.2015, leg. A.V. Kurmaev
    Copyright © Igor Solodovnikov
    Carabus (Morphocarabus) odoratus kurmaevi I.Solodovnikov, 2017
    Carabus (Morphocarabus) odoratus kurmaevi I.Solodovnikov, 2017 Paratype
    Female adult
    Russia, Siberia, Krasnoyarsk Krai, the North Yenisei district, 55 km of CE of the vill. Bryanka, valley of the river Chirimba, stream Nakhodny, h = 350 m, the mixed forest (birch and pine); on a steep slope along an animal track on a hill brow, 23.VII-20.VIII.2015, leg. A.V. Kurmaev
    Copyright © Igor Solodovnikov
    Carabus (Morphocarabus) odoratus kurmaevi I.Solodovnikov, 2017
    Carabus (Morphocarabus) odoratus kurmaevi I.Solodovnikov, 2017 Paratype
    Female adult
    Russia, Siberia, Krasnoyarsk Krai, the North Yenisei district, 55 km of CE of the vill. Bryanka, valley of the river Chirimba, stream Nakhodny, h = 350 m, the mixed forest (birch and pine); on a steep slope along an animal track on a hill brow, 23.VII-20.VIII.2015, leg. A.V. Kurmaev
    Copyright © Igor Solodovnikov
    Carabus (Morphocarabus) odoratus kurmaevi I.Solodovnikov, 2017
    Carabus (Morphocarabus) odoratus kurmaevi I.Solodovnikov, 2017 Holotype
    Endophallus
    Russia, Siberia, Krasnoyarsk Krai, the North Yenisei district, 55 km of CE of the vill. Bryanka, valley of the river Chirimba, stream Nakhodny, h = 350 m, the mixed forest (birch and pine); on a steep slope along an animal track on a hill brow, 23.VII-20.VIII.2015, leg. A.V. Kurmaev
    Copyright © Igor Solodovnikov
    Carabus (Morphocarabus) viridilimbatus panini I.Solodovnikov 2017
    Carabus (Morphocarabus) viridilimbatus panini I.Solodovnikov 2017 Holotype
    Male adult
    Russia, SE Yakutia, Neryungri distr., Tokinski Stanovik Mt Rng., Beder riv., (right tributary rivers Algama), 26.VI.1996, Krasnov M. leg., 17 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) viridilimbatus panini I.Solodovnikov 2017
    Carabus (Morphocarabus) viridilimbatus panini I.Solodovnikov 2017 Paratype
    Female adult
    Russia, SE Yakutia, Neryungri distr., Tokinski Stanovik Mt Rng., Beder riv., (right tributary rivers Algama), taiga, h = 940 m, 55°55'27.78" N, 129°42'6.20" E, 26.VI.1996, leg. Krasnov M. 21,1 mm
    Copyright © Igor Solodovnikov
    Carabus (Morphocarabus) viridilimbatus saluki I.Solodovnikov 2017
    Carabus (Morphocarabus) viridilimbatus saluki I.Solodovnikov 2017 Holotype
    Male adult
    Russia, SW Taymyr, near Talnakh, 25 km N Norilsk, hilly tundra, hillside overgrown with alder slate, dwarfish birches, blueberry, h = 150-250 м, 24-28.06.2003, leg. S.Saluk 20,5 mm
    Copyright © Igor Solodovnikov
    Carabus (Morphocarabus) viridilimbatus saluki I.Solodovnikov 2017
    Carabus (Morphocarabus) viridilimbatus saluki I.Solodovnikov 2017 Paratype
    Male adult
    Russia, SW Taymyr, near Talnakh, 25 km N Norilsk, hilly tundra, hillside overgrown with alder slate, dwarfish birches, blueberry, h = 150-250 м, 24-28.06.2003, leg. S.Saluk
    Copyright © Igor Solodovnikov
    Carabus (Morphocarabus) viridilimbatus saluki I.Solodovnikov 2017
    Carabus (Morphocarabus) viridilimbatus saluki I.Solodovnikov 2017 Paratype
    Female adult
    Russia, SW Taymyr, near Talnakh, 25 km N Norilsk, hilly tundra, hillside overgrown with alder slate, dwarfish birches, blueberry, h = 150-250 м, 24-28.06.2003, leg. S.Saluk
    Copyright © Igor Solodovnikov
    Carabus (Morphocarabus) viridilimbatus saluki I.Solodovnikov 2017
    Carabus (Morphocarabus) viridilimbatus saluki I.Solodovnikov 2017 Paratype
    Female adult
    Russia, SW Taymyr, near Talnakh, 25 km N Norilsk, hilly tundra, hillside overgrown with alder slate, dwarfish birches, blueberry, h = 150-250 м, 24-28.06.2003, leg. S.Saluk
    Copyright © Igor Solodovnikov
    Carabus (Morphocarabus) viridilimbatus saluki I.Solodovnikov 2017
    Carabus (Morphocarabus) viridilimbatus saluki I.Solodovnikov 2017 Paratype
    Endophallus
    Russia, SW Taymyr, near Talnakh, 25 km N Norilsk, hilly tundra, hillside overgrown with alder slate, dwarfish birches, blueberry, h = 150-250 м, 24-28.06.2003, leg. S.Saluk
    Copyright © Igor Solodovnikov
    Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859
    Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859
    Endophallus
    Russia, Chita Region, Udokansky district. Kodar Range, 56 °34'42,9 " N, 118 °29'34,95" E, 6/13/2002, leg. A. Pokhorov
    Copyright © Igor Solodovnikov
    Carabus (Tanaocarabus) sylvosus Say, 1825
    Carabus (Tanaocarabus) sylvosus Say, 1825
    Female adult

    Copyright © Henri Goulet
    Carabus (Coptolabrus) augustus tenuitarsatus Rapuzzi, 2016
    Carabus (Coptolabrus) augustus tenuitarsatus Rapuzzi, 2016 Holotype
    Male adult
    China, Chongqing province, Pengshui county, Mt. Heimending. Image from Rapuzzi, 2016
    Copyright ©
    Carabus (Coptolabrus) smaragdinus losevi Rapuzzi, 2016
    Carabus (Coptolabrus) smaragdinus losevi Rapuzzi, 2016 Holotype
    Male adult
    Far East of Russia, South Primorye, Khasanskiy district, Furugelm Island, 11/13.VII.2013, O. Losev legit. Image from Rapuzzi, 2016
    Copyright © - - -
    Carabus (Coptolabrus) elysii wangguofeni Rapuzzi, 2016
    Carabus (Coptolabrus) elysii wangguofeni Rapuzzi, 2016 Holotype
    Male adult
    China, Anhui province, Taihu, Wangling vill., South slope of Mt. Dabieshan, 400 m, 10/30.IV.2015, (30º31'18'' N; 116º16'39'' E), Xihuangshun legit; preserved in Ivan Rapuzzi collection. Image from Rapuzzi, 2016
    Copyright ©
    Carabus (Neoplectes) ibericus s.str. (as djanoliensis Fominykh & Zamotajlov, 2016)
    Carabus (Neoplectes) ibericus s.str. (as djanoliensis Fominykh & Zamotajlov, 2016) Paratype
    Male+Female adult
    Image from Fominykh & Zamotajlov, 2016
    Copyright © - - -
    Carabus (Neoplectes) ibericus ibericus (as prunierianus Deuve, 2012)
    Carabus (Neoplectes) ibericus ibericus (as prunierianus Deuve, 2012)
    Male+Female adult
    Georgia, Magaroskari env. Image from Fominykh & Zamotajlov, 2016
    Copyright © - - -
    Carabus (Neoplectes) ibericus ibericus (as prunierianus Deuve, 2012)
    Carabus (Neoplectes) ibericus ibericus (as prunierianus Deuve, 2012)
    Male adult
    Georgia, Passanauri env. Image from Fominykh & Zamotajlov, 2016
    Copyright © - - -
    Carabus (Neoplectes) ibericus ibericus (as szekelyi Retezar, 2011)
    Carabus (Neoplectes) ibericus ibericus (as szekelyi Retezar, 2011)
    Male+Female adult
    Itrapula env. Image from Fominykh & Zamotajlov, 2016
    Copyright © - - -
    Carabus (Neoplectes) ibericus ibericus (as szekelyi Retezar, 2011)
    Carabus (Neoplectes) ibericus ibericus (as szekelyi Retezar, 2011)
    Male+Female adult
    Dzhalobet env.; Image from Fominykh & Zamotajlov, 2016
    Copyright © - - -
    Carabus (Neoplectes) ibericus ibericus (as szekelyi Retezar, 2011)
    Carabus (Neoplectes) ibericus ibericus (as szekelyi Retezar, 2011)
    Male+Female adult
    Nakerala env.; Image from Fominykh & Zamotajlov, 2016
    Copyright © - - -
    Carabus (Procerus) scabrosus akbesianus Breuning, 1975
    Carabus (Procerus) scabrosus akbesianus Breuning, 1975 Holotype
    Male adult
    Syrie, Akbes (MNHN-EC-EC131)
    Copyright © - - -
    Carabus (Coptolabrus) ignimitella ertli Born, 1910
    Carabus (Coptolabrus) ignimitella ertli Born, 1910 Holotype
    Male adult
    Foothow (MNHN-EC-EC136)
    Copyright © - - -
    Carabus (Apotomopterus) davidis davidis Deyrolle & Fairmaire, 1878
    Carabus (Apotomopterus) davidis davidis Deyrolle & Fairmaire, 1878 Holotype
    Female adult
    (MNHN-EC-EC137) coldb.mnhn.fr/catalognumber/mnhn/ec/ec137
    Copyright © - - -
    Carabus (Apotomopterus) delavayi delavayi Fairmaire, 1886
    Carabus (Apotomopterus) delavayi delavayi Fairmaire, 1886 Lectotype
    Female adult
    (MNHN-EC-EC138) http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec138
    Copyright © - - -
    Carabus (Archicarabus) gotschi caramanus Fairmaire, 1886
    Carabus (Archicarabus) gotschi caramanus Fairmaire, 1886 Holotype
    Male adult
    (MNHN-EC-EC141) http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec141
    Copyright © - - -
    Carabus (Pagocarabus) wagae wagae Fairmaire, 1882
    Carabus (Pagocarabus) wagae wagae Fairmaire, 1882 Holotype
    Female adult
    Thibet MNHN) http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec146
    Copyright © - - -
    Carabus (Pseudocoptolabrus) taliensis (Fairmaire, 1886)
    Carabus (Pseudocoptolabrus) taliensis (Fairmaire, 1886) Holotype
    Female adult
    Yunnan; http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec147
    Copyright ©
    Carabus (Acoptolabrus) gehinii gehinii Fairmaire, 1886
    Carabus (Acoptolabrus) gehinii gehinii Fairmaire, 1886 Holotype
    Male adult
    Yokohama; http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec148
    Copyright © - - -
    Carabus (Coptolabrus) gemmifer gemmifer Fairmaire, 1887
    Carabus (Coptolabrus) gemmifer gemmifer Fairmaire, 1887 Holotype
    Male adult
    Yunnan (MNHN) http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec150
    Copyright © MNHN
    Carabus (Aristocarabus) viridifossulatus viridifossulatus Fairmaire, 1887
    Carabus (Aristocarabus) viridifossulatus viridifossulatus Fairmaire, 1887 Holotype
    Male adult
    Moupin
    Copyright © - - -
    Carabus (Coptolabrus) gemmifer loczyi Csiki, 1927
    Carabus (Coptolabrus) gemmifer loczyi Csiki, 1927 Holotype
    Male adult
    Yunnan; reçu de Bang Haes le 23/IV/1912 comme yunnanus (MNHN) http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec124
    Copyright © MNHN
    Carabus (Apotomopterus) vitalisi vitalisi Lapouge, 1916
    Carabus (Apotomopterus) vitalisi vitalisi Lapouge, 1916 Lectotype
    Male adult
    Xieng Khouang, Nan-Binh. deposited in MNHN (http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec514)
    Copyright © - - -
    Carabus (Morphocarabus) monilis Fabricius, 1792
    Carabus (Morphocarabus) monilis Fabricius, 1792
    Female adult
    France, Dröme, Ft. de Saou, 17.vii.1975-M. Grotz Leg.-Coll.RBINS, N. Mal as monilis saouensis nicodi
    Copyright © Noël Mal
    Carabus (Tachypus) auratus honnoratii Dejean, 1826
    Carabus (Tachypus) auratus honnoratii Dejean, 1826
    Male+Female adult
    France : Vaucluse, Mt.Ventoux-Mt. Serein, vii.1969, N. Mal Leg., Coll.RBINS-N. Mal incl. ventouxensis and Nicolasi
    Copyright © Noël Mal
    Carabus (Tachypus) auratus honnoratii Dejean, 1826
    Carabus (Tachypus) auratus honnoratii Dejean, 1826
    Male+Female adult
    France : Vaucluse, Mt. Ventoux-Mt. Serein, 1400-1800m, vii.1969-N. Mal Leg.-Coll.RBINS, N. Mal - including f.i. ventouxensis and nicolasi
    Copyright © Noël Mal
    Carabus (Imaibius) wittmerorum Heinertz, 1978
    Carabus (Imaibius) wittmerorum Heinertz, 1978
    Female adult

    Copyright © Noël Mal
    Carabus (Ctenocarabus) melancholicus costatus Germar, 1824
    Carabus (Ctenocarabus) melancholicus costatus Germar, 1824
    Male adult

    Copyright © Noël Mal
    Carabus (Alipaster) barovskii bayanbulak Deuve, 1993
    Carabus (Alipaster) barovskii bayanbulak Deuve, 1993
    Male adult
    China, Xinjiang, S sl. of Khalyktau, road Kuga-Bayanbulak, Telemet Pass. S slope, h~3400-3500 9.VI.2001 leg.I.Kabak L=14.0 mm
    Copyright © Kirill Makarov
    Carabus (Megodontus) kolbei taikinus Ishikawa, 1993
    Carabus (Megodontus) kolbei taikinus Ishikawa, 1993
    Male adult
    N-Japan, SC-Hokkaido:Tokachi, Hiroo Distr., Taiki-cho env., 28.VIII-2.IX.2017, Teruo Hirabayashi leg. L=24 mm
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) estreicheri (ab.rufofemoratus Lomnicki, 1892)
    Carabus (Trachycarabus) estreicheri (ab.rufofemoratus Lomnicki, 1892)
    Female adult
    W Ukraine, NW Podolian, Lviv reg., Brody distr., Makitra Mt, steppe site, 15.VI - 24.IX.2017, Panin R. & Kanarsky Yu. leg.
    Copyright © Ruslan Panin
    Carabus (Coptolabrus) ignimitella antaeus Hauser, 1914 (as guidongicus)
    Carabus (Coptolabrus) ignimitella antaeus Hauser, 1914 (as guidongicus)
    Female adult
    China, E-Hunan, Zhuzhou, Yanling County, Wanyang Shan Mts, VI.2003 , L= 47 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) wulffiusi liaoningensis Deuve & Imura, 1990
    Carabus (Morphocarabus) wulffiusi liaoningensis Deuve & Imura, 1990
    Female adult
    NE-China:E-Liaoning: NP Kuandian Tianqiaogou , VI.2010, L=24
    Copyright © Ruslan Panin
    Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931
    Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931
    Male adult
    NE-China:E-Liaoning: NP Kuandian Tianqiaogou , VI.2010, L=25 mm, det. Panin R.
    Copyright © Ruslan Panin
    Carabus (Coptolabrus) jankowskii fusanus Born, 1907
    Carabus (Coptolabrus) jankowskii fusanus Born, 1907
    Female adult
    South Korea, S Gyeongsang Prov., Busan, Mt Geumwonsan, 17.VI.2017, L=37 mm
    Copyright © Ruslan Panin
    Carabus (Coptolabrus) formosus wanxianicus Imura, 1995
    Carabus (Coptolabrus) formosus wanxianicus Imura, 1995
    Male adult
    China, NE-Sichuan, Dazhou, Wanyuan env., 6.VI.2005. L = 33mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
    Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
    Female adult
    NE-Hungary. (photo by Imre Retezár)
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
    Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
    Male adult
    NE-Hungary. (photo by Imre Retezár)
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
    Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
    Male adult
    W Ukraine, Transcarpathian, Mts E of Uzgorod, Veliki Lazy h~270-300 m, 8.IV-23.V.2011, Panin R. leg. L=28 mm
    Copyright © Ruslan Panin
    Carabus (Coptolabrus) nankotaizanus Kano, 1932
    Carabus (Coptolabrus) nankotaizanus Kano, 1932
    Male adult
    NE-Taiwan, Yilan County, V.2017, L=39 mm
    Copyright © Ruslan Panin
    Carabus (Coptolabrus) jankowskii fusanus (as obtusipennis)
    Carabus (Coptolabrus) jankowskii fusanus (as obtusipennis)
    Female adult
    SC-Korea: Chiri-san Massif, Mt. Jirisan, 28.VI.2011, L=35 mm
    Copyright © Ruslan Panin
    Carabus (Homoeocarabus) maeander paludis Gehin, 1885
    Carabus (Homoeocarabus) maeander paludis Gehin, 1885
    Female adult
    Japan, Hokkaido; Tomakomai city, Benten, 17.V.2015, Takemoto leg., L=21 mm
    Copyright © Ruslan Panin
    Carabus (Axinocarabus) fedtschenkoi kondarensis Kryzhanovskij, 1953
    Carabus (Axinocarabus) fedtschenkoi kondarensis Kryzhanovskij, 1953
    Female adult
    NW-Tadzhikistan: Gissar Mt.Rng, Varzob river ravine, Gushari env., h~1400-1500m, 24.III.2010, L=29 mm
    Copyright © Ruslan Panin
    Carabus (Leptocarabus) arboreus fujisanus Bates, 1883
    Carabus (Leptocarabus) arboreus fujisanus Bates, 1883
    Female adult
    Japan, Honshu, Mt. Fuji h~1900m, Shizuoka pref., Fujinomiya city, 23.VI.2007, M. Matsunaga leg., L=25 mm
    Copyright © Ruslan Panin
    Carabus (Platycarabus) irregularis irregularis (as jurassicus Born, 1911)
    Carabus (Platycarabus) irregularis irregularis (as jurassicus Born, 1911)
    Male+Female adult
    France, Ain (01), Jura Mts, Grand Colombier, Forêt d' Arvières h~1250 m, 25.X.2014, Ya. Vasseur leg., L=23 mm
    Copyright © Ruslan Panin
    Carabus (Neoplectes) komarowi komarowi Reitter, 1882
    Carabus (Neoplectes) komarowi komarowi Reitter, 1882
    Female adult
    NW Georgia, Upper Svaneti, Mestia distr., Lakhamula env. h~1250 m, 19.VI.2014, V.Sima leg. L=33 mm
    Copyright © Ruslan Panin
    Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
    Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
    Female adult
    W Georgia, Martvili Distr., 5 km N T'aleri, 11-18.VI.2014, V. Sima leg., L=50 mm
    Copyright © Ruslan Panin
    Carabus (Lamprostus) calleyi pseudoprasinus Lapouge, 1912
    Carabus (Lamprostus) calleyi pseudoprasinus Lapouge, 1912
    Male adult
    S Georgia, Tsalka (Paravani) 41°29′18.7″N, 43°51′12.3″E,
    Copyright © Ruslan Panin
    Carabus (Lamprostus) calleyi pseudoprasinus Lapouge, 1912
    Carabus (Lamprostus) calleyi pseudoprasinus Lapouge, 1912
    Male adult
    S Georgia, Tsalka (Paravani) 41°29′18.7″N, 43°51′12.3″E, 8-9.V.2016, Pavel Turek leg. / det. I. Solodovnikov, L=23 mm
    Copyright © Ruslan Panin
    Carabus (Coptolabrus) gemmifer mesites Fairmaire, 1897
    Carabus (Coptolabrus) gemmifer mesites Fairmaire, 1897
    Male adult
    China, Shimian 1200m Shimian Yaan Sichuan VII.2017 (DUBC). det. Anichtchenko A. 2018
    Copyright © Alexander Anichtchenko
    Carabus (Trachycarabus) mandibularis abakkereiorum Kabak & Hu, 2017
    Carabus (Trachycarabus) mandibularis abakkereiorum Kabak & Hu, 2017 Holotype Paratype
    Male+Female adult
    (ZIN). China, Xinjiang, Altay Mt. R., Kran Valley, N of Altay Town, 47°55´37''N / 88°08´23´´E, h = 1015 m, 31.VII.2015 I.I. Kabak leg.
    Copyright © Ilya Kabak
    Carabus (Trachycarabus) mandibularis buchtarmensis Kryzhanovskij, 1953
    Carabus (Trachycarabus) mandibularis buchtarmensis Kryzhanovskij, 1953 Paralectotypes
    Male genitalia
    Aedeagus: from Tshernovaya; Endofallus: from Bolshenarymskoe.
    Copyright © Ilya Kabak
    Carabus (Trachycarabus) mandibularis abakkereiorum Kabak & Hu, 2017
    Carabus (Trachycarabus) mandibularis abakkereiorum Kabak & Hu, 2017
    Male genitalia

    Copyright © Ilya Kabak
    Carabus (Trachycarabus) sibiricus obliteratus Fischer, 1827
    Carabus (Trachycarabus) sibiricus obliteratus Fischer, 1827 Paratype
    Female adult
    China, Xinjiang, “Prov. Xinjiang: Ake Tei A Zhu Pai 1750 m, 5.VIII.1960, leg. Wan Su Yung, China Naturwiss. Institut”; “Paratypus Carabus (Trachycarabus) ♀ sibiricus pseudobliteratus nov. ssp. det. Korell et Kleinfeld, 1982”. (MFNB)
    Copyright © Ilya Kabak
    Carabus (Cryptocarabus) kasantsevi Kabak, 1993
    Carabus (Cryptocarabus) kasantsevi Kabak, 1993
    Male adult
    SE Kasakhstan, E part of Terskey Alatau, Bayankol riv., tugai.
    Copyright © Ilya Kabak
    Carabus (Cratophyrtus) katajevi katajevi Gottwald, 1989
    Carabus (Cratophyrtus) katajevi katajevi Gottwald, 1989
    Male adult

    Copyright © Ilya Kabak
    Carabus (Leptoplesius) itshkibashi Kabak, 2004
    Carabus (Leptoplesius) itshkibashi Kabak, 2004
    Female adult
    China, Xinjiang, S slope of Khalyktau
    Copyright © Ilya Kabak
    Carabus (Cratocarabus) kryzhanovskii kryzhanovskii Bogachev, 1965
    Carabus (Cratocarabus) kryzhanovskii kryzhanovskii Bogachev, 1965
    Female adult

    Copyright © Ilya Kabak
    Carabus (Cratocechenus) tshistjakovae gracilidens Kabak, 2004
    Carabus (Cratocechenus) tshistjakovae gracilidens Kabak, 2004
    Female adult
    China, Xinjiang, Narat mt.rng.
    Copyright © Ilya Kabak
    Carabus (Ophiocarabus) ernsti ernsti Kabak, 2001
    Carabus (Ophiocarabus) ernsti ernsti Kabak, 2001
    Male adult
    Boro-Horo
    Copyright © Ilya Kabak
    Carabus (Ophiocarabus) kurdaiensis Kabak
    Carabus (Ophiocarabus) kurdaiensis Kabak Paratype
    Female adult
    China, Xinjiang, Narat Mt. Range, Ekiasha River, near Kurdai Pass, 42°54ʹ21″N / 82°28ʹ30″E, 3640 m, 26.07.2013 I.I. Kabak leg. L=16.5 mm
    Copyright © Kirill Makarov
    Carabus (Orinocarabus) linnaei linnaei Panzer, 1812
    Carabus (Orinocarabus) linnaei linnaei Panzer, 1812
    Endophallus
    W.UKRAINE, Lviv Reg., NE Carpathians, Skolivski Beskidy Mts. Zelemyanka Mt. Rng
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) linnaei linnaei Panzer, 1812
    Carabus (Orinocarabus) linnaei linnaei Panzer, 1812
    Male adult
    W.UKRAINE, Lviv Reg., NE Carpathians, Skolivski Beskidy Mts. Zelemyanka Mt. Rng h=600m, 15.VII-12VIII.2014, Panin R. leg. L=19 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) linnaei linnaei Panzer, 1812
    Carabus (Orinocarabus) linnaei linnaei Panzer, 1812
    Female adult
    W Ukraine, Transcarpathian Reg., Rakhiv distr., Marmarosh Mts., h~400 m, 19-23.VII.2010, Reserve "Kuziy", Panin R. leg. L=23 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) linnaei linnaei Panzer, 1812
    Carabus (Orinocarabus) linnaei linnaei Panzer, 1812
    Male adult
    W-UKRAINE, Ivano-Frankivsk region, S Gorgany Mts, Synyk Mt. h =1630 м., 15.V-18.VI.2010, Panin R. leg. L=19 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) linnaei linnaei (as scopolii Dejean, 1826)
    Carabus (Orinocarabus) linnaei linnaei (as scopolii Dejean, 1826)
    Male adult
    W Ukraine, Lviv reg., NW Podolian, forest near Vybranivka vill., 12.X.2007, Panin R. leg. L=19 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
    Male adult
    W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI.-5,VIII.2017, R. Panin R. & Yu.Kanarsky leg.
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
    Male adult
    W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI.-5,VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=17 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
    Female adult
    Male adult W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI.-5,VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=18 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
    Male adult
    W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI.-5,VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=17.5 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Holotype
    Male adult
    W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=17.5 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
    Male adult
    W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L =17 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
    Male adult
    W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=16.5 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018
    Endophallus
    W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone,
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
    Male adult
    W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=17 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
    Female adult
    W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=17 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
    Female adult
    W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=18 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
    Female adult
    W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=19 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
    Male adult
    W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=19 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
    Female adult
    W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=20 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
    Male adult
    W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=16.5 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
    Male adult
    W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=19 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018
    Habitat
    Mt. Hoverla
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018
    Habitat

    Copyright © Ruslan Panin
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
    Male adult
    W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Gutyn-Tomnatyk h~2015 m alpin zone, 8.VI-28.VII.2011, R. Panin R. leg. L=14 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
    Female adult
    W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Gutyn-Tomnatyk h~2015 m, alpin zone, 8.VI-28.VII.2011, R. Panin R. leg. L=18 mm
    Copyright © Ruslan Panin
    Carabus (Ophiocarabus) latiballioni Deuve, 1993
    Carabus (Ophiocarabus) latiballioni Deuve, 1993
    Male adult
    China, Xinjiang, Narat Mt.R., S of Kurdelin forest, h~1760 43°07'50"N 82°50'51"E 30.VII.2013 leg. I.Kabak L=17.0 mm
    Copyright © Kirill Makarov
    Carabus (Ophiocarabus) latiballioni Deuve, 1993
    Carabus (Ophiocarabus) latiballioni Deuve, 1993
    Female adult
    China, Xinjiang, Narat Mt.R., S of Kurdelin forest, h~1760 43°07'50"N 82°50'51"E 30.VII.2013 leg. I.Kabak L=22.0 mm
    Copyright © Kirill Makarov
    Carabus (Cratocechenus) tshistjakovae gracilidens Kabak, 2004
    Carabus (Cratocechenus) tshistjakovae gracilidens Kabak, 2004
    Female adult
    China, Xinjiang, Narat Mt.R.,, h~2275 Tshashi vall.,Tuyukasha Riv. 43°00'29"N 82°46'02"E 18.VII.2014 leg. I.Kabak L=22.0
    Copyright © Kirill Makarov
    Carabus (Cryptocarabus) sororius Morawitz, 1886
    Carabus (Cryptocarabus) sororius Morawitz, 1886
    Male adult
    China, Xinjiang, S slope of Boro-Horo Mt.R., env. of Ulastai vill.,h~1690 43°49'50"N 83°08'45"E 8.V.2017 leg. I.Kabak L=18.5 mm
    Copyright © Kirill Makarov
    Carabus (Cryptocarabus) sororius Morawitz, 1886
    Carabus (Cryptocarabus) sororius Morawitz, 1886
    Female adult
    China, Xinjiang, S slope of Boro-Horo Mt.R., env. of Ulastai vill.,h~1690 43°49'50"N 83°08'45"E 8.V.2017 leg. I.Kabak L=20.0 mm
    Copyright © Kirill Makarov
    Carabus (Neoplectes) ibericus ibericus (as imereticus Retezar, 2011)
    Carabus (Neoplectes) ibericus ibericus (as imereticus Retezar, 2011) Paratype
    Endophallus
    NC-Georgia, S-Caucasus: W slope Likhi Mt.Rng., 3 km W Rikoti Pass h~900 m,
    Copyright © Ruslan Panin
    Carabus (Neoplectes) ibericus ibericus (as imereticus Retezar, 2011)
    Carabus (Neoplectes) ibericus ibericus (as imereticus Retezar, 2011) Paratype
    Male adult
    S-Caucasus: NC-Georgia, W slope Likhi Mt. Rng., 3 km W of Rikoti Pass h~900 m, 15.VI - 8.VII.2006, A.Puchkov leg., L=29.5 mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) persianus espeyliensis Deuve, 2001
    Carabus (Megodontus) persianus espeyliensis Deuve, 2001
    Male adult
    N-Iran, Gilan Prov., Deylaman Distr., Balarud Mts, Espili env., 18-24.VI.2017, Václav Čutka leg., L= 32 mm
    Copyright © Ruslan Panin
    Carabus (Carabus) sculpturatus (as gilanicus Mandl, 1967)
    Carabus (Carabus) sculpturatus (as gilanicus Mandl, 1967)
    Male adult
    N-Iran:Gilan Prov., Asalem Distr., Siahbil, deciduous forest (very wet) h= -10 m, 20-24.VI.2017, V. Čutka leg., L=19 mm
    Copyright © Ruslan Panin
    Carabus (Carabus) sculpturatus (as gilanicus Mandl, 1967)
    Carabus (Carabus) sculpturatus (as gilanicus Mandl, 1967)
    Female adult
    N-Iran:Gilan Prov., Asalem Distr., Siahbil, deciduous forest (very wet) h= -10 m, 20-24.VI.2017, V. Čutka leg., L=22 mm
    Copyright © Ruslan Panin
    Carabus (Tribax) puschkini platypterus Ganglbauer, 1886
    Carabus (Tribax) puschkini platypterus Ganglbauer, 1886
    Endophallus
    SW Georgia, Lesser Caucasus, Adzhar-Imereti (Meskheti) Mt Range, 10 km S Sairme h~1850 m.
    Copyright © Ruslan Panin
    Carabus (Tribax) puschkini platypterus Ganglbauer, 1886
    Carabus (Tribax) puschkini platypterus Ganglbauer, 1886
    Male adult
    SW Georgia, Lesser Caucasus, Adzhar-Imereti (Meskheti) Mt Range, 10 km S Sairme h~1850 m, 27.VI-2.VII.2013, A.Puchkov & Martynov leg. L=27 mm
    Copyright © Ruslan Panin
    Carabus (Ophiocarabus) successor successor Reitter, 1896
    Carabus (Ophiocarabus) successor successor Reitter, 1896
    Endophallus
    SE-Kazakhstan, Narynkol distr., 4 km SW Bayankol, h~2400-2500 m
    Copyright © Ruslan Panin
    Carabus (Megodontus) stroganowi morgani Lapouge, 1907
    Carabus (Megodontus) stroganowi morgani Lapouge, 1907
    Male adult
    Iran, Mazandaran prov., Nowshahr env, Khirood forest, 24.5.2013, 1200 m.
    Copyright © Alexander Anichtchenko
    Carabus (Megodontus) stroganowi tiedemanni Breuning, 1934
    Carabus (Megodontus) stroganowi tiedemanni Breuning, 1934
    Male adult

    Copyright © Alexander Anichtchenko
    Carabus (Ophiocarabus) successor successor Reitter, 1896
    Carabus (Ophiocarabus) successor successor Reitter, 1896
    Male adult
    SE-Kazakhstan, Narynkol distr., 4 km SW Bayankol, h~2400-2500 m, 2-14.VI.2007, Shumov leg. L=17.5 mm
    Copyright © Ruslan Panin
    Carabus (Mimocarabus) maurus Adams, 1817
    Carabus (Mimocarabus) maurus Adams, 1817
    Female adult

    Copyright © Kirill Makarov
    Carabus (Mimocarabus) maurus Adams, 1817
    Carabus (Mimocarabus) maurus Adams, 1817
    Male adult

    Copyright © Kirill Makarov
    Carabus (Ophiocarabus) successor successor Reitter, 1896
    Carabus (Ophiocarabus) successor successor Reitter, 1896
    Female adult
    SE-Kazakhstan, Narynkol distr., 4 km SW Bayankol, h~2400-2500 m, 2-14.VI.2007, Shumov leg. L=17 mm
    Copyright © Ruslan Panin
    Carabus (Meganebrius) quinlani quinlani Mandl, 1965
    Carabus (Meganebrius) quinlani quinlani Mandl, 1965
    Male adult
    C-Nepal, Gandaki, Manang Distr., Timang env., h~2500-2600m, old open forest, grassland, 3-5.V.2007, Fouqué R. leg. L=19 mm
    Copyright © Ruslan Panin
    Carabus (Sphodristocarabus) separatus bararii Deuve, 2000
    Carabus (Sphodristocarabus) separatus bararii Deuve, 2000
    Male adult
    N-Iran: Mazendaran: Ramsar env., deciduous forest, h~305 m, 16-24.VI.2017, Václav Čutka leg. L=22 mm
    Copyright © Ruslan Panin
    Carabus (Pagocarabus) sichuanicola sichuanicola (as drolmae Lassalle, 2003)
    Carabus (Pagocarabus) sichuanicola sichuanicola (as drolmae Lassalle, 2003)
    Female adult
    China, Sichuan, NW Chengdu, Mt. Jiajin Shan h~4000 m,VI. 2016, local collector, L= 24 mm
    Copyright © Ruslan Panin
    Carabus (Cupreocarabus) alpherakii alpherakii Semenov, 1898
    Carabus (Cupreocarabus) alpherakii alpherakii Semenov, 1898
    Female adult
    China, W Sichuan, Litang env., h~5000 m,29.VII -3.VIIII.1995, J. Schneider leg. L=23 mm
    Copyright © Ruslan Panin
    Carabus (Sphodristocarabus) separatus bararii Deuve, 2000
    Carabus (Sphodristocarabus) separatus bararii Deuve, 2000
    Female adult
    N-Iran: Mazendaran: Ramsar env., deciduous forest, h~305 m, 16-24.VI.2017, Václav Čutka leg. L=23,5 mm
    Copyright © Ruslan Panin
    Carabus (Tribax) puschkini kolenatii (as suramensis Kraatz, 1887)
    Carabus (Tribax) puschkini kolenatii (as suramensis Kraatz, 1887)
    Male adult
    S-Caucasus: NC-Georgia, W slope Likhi Mt.Range, 3 km W Rikoti Pass h~900m 11-14.VI -2006, A.Puchkov leg. L=27 mm
    Copyright © Ruslan Panin
    Carabus (Tribax) puschkini kolenatii (as suramensis Kraatz, 1887)
    Carabus (Tribax) puschkini kolenatii (as suramensis Kraatz, 1887)
    Endophallus
    S-Caucasus: NC-Georgia, W slope Likhi Mt.Range, 3 km W Rikoti Pass h~900m
    Copyright © Ruslan Panin
    Carabus (Cryptocarabus) tsharynensis Kabak 1994
    Carabus (Cryptocarabus) tsharynensis Kabak 1994
    Endophallus
    SE-Kazakhstan, Almaty Region, W-Zailiyskiy Alatau rng, Belshabdar valley h~1300 m
    Copyright © Ruslan Panin
    Carabus (Cryptocarabus) tsharynensis Kabak 1994
    Carabus (Cryptocarabus) tsharynensis Kabak 1994
    Male adult
    SE-Kazakhstan, Almaty Region, W-Zailiyskiy Alatau rng, Belshabdar valley h~1300 m, 25-27.VI.2002, A.Puchkov leg. L=17 mm
    Copyright © Ruslan Panin
    Carabus (Cryptocarabus) tsharynensis Kabak 1994
    Carabus (Cryptocarabus) tsharynensis Kabak 1994
    Female adult
    E-Kazakhstan, Almaty Region, W-Zailiyskiy Alatau, Turgen distr., Tauturgen vill., h~1200 m, 8-13.VI.2002, A.Puchkov leg. L=19 mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) persianus javahardehensis Deuve, 2000
    Carabus (Megodontus) persianus javahardehensis Deuve, 2000
    Male adult
    N-Iran, Mazandaran Prov., Ramsar Co., Javahar Deh env., 16-25.VI.2017, Václav Čutka leg. L=28 mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) persianus javahardehensis Deuve, 2000
    Carabus (Megodontus) persianus javahardehensis Deuve, 2000
    Female adult
    N-Iran, Mazandaran Prov., Ramsar Co., Javahar Deh env., 16-25.VI.2017, Václav Čutka leg. L=30 mm
    Copyright © Ruslan Panin
    Carabus (Sphodristocarabus) separatus separatus Gehin, 1885
    Carabus (Sphodristocarabus) separatus separatus Gehin, 1885
    Male adult
    N-Iran:Gilan Prov., Asalem - Khalkal road, 25-30.V.2018, Václav Čutka leg. L= 21 mm
    Copyright © Ruslan Panin
    Carabus (Sphodristocarabus) separatus separatus Gehin, 1885
    Carabus (Sphodristocarabus) separatus separatus Gehin, 1885
    Female adult
    N-Iran: Gilan Prov., Asalem - Khalkal road, 25-30.V.2018, Václav Čutka leg. L= 23 mm
    Copyright © Ruslan Panin
    Carabus (Sphodristocarabus) separatus separatus Gehin, 1885
    Carabus (Sphodristocarabus) separatus separatus Gehin, 1885
    Male adult
    N-Iran:Gilan Prov., Asalem - Khalkal road, 25-30.V.2018, Václav Čutka leg. L= 22 mm
    Copyright © Ruslan Panin
    Carabus (Sphodristocarabus) separatus separatus Gehin, 1885
    Carabus (Sphodristocarabus) separatus separatus Gehin, 1885
    Endophallus
    N-Iran:Gilan Prov., Asalem - Khalkal road,
    Copyright © Ruslan Panin
    Carabus (Hemicarabus) serratus Say, 1825
    Carabus (Hemicarabus) serratus Say, 1825
    Male adult
    E-USA,Virginia, Leesburg env., Ball's Bluff Battlefield Regional Park, 27.VIII-4.IX.2015,V. Čutka leg. L=17 mm
    Copyright © Ruslan Panin
    Carabus (Hemicarabus) serratus Say, 1825
    Carabus (Hemicarabus) serratus Say, 1825
    Female adult
    E-USA, Virginia, Leesburg env., Ball's Bluff Battlefield Regional Park, 27.VIII-4.IX.2015,V. Čutka leg. L=18 mm
    Copyright © Ruslan Panin
    Carabus (Sphodristocarabus) separatus haerii Heinz, 1973
    Carabus (Sphodristocarabus) separatus haerii Heinz, 1973
    Female adult
    N-Iran:Gilan Prov., Rudbar, Harzevil env.,23-24.V.2018, Václav Čutka leg. L=24 mm
    Copyright © Ruslan Panin
    Carabus (Sphodristocarabus) separatus haerii Heinz, 1973
    Carabus (Sphodristocarabus) separatus haerii Heinz, 1973
    Male adult
    N-Iran: Gilan Prov., E of Rudbar, Barehsar env., 19-23.VI.2017, Václav Čutka leg. L=22 mm
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) dufouri Dejean, 1829
    Carabus (Mesocarabus) dufouri Dejean, 1829
    Female adult
    S Spain, Granada, Sierra Arana, Píñar env., 14.X.2018, F. Garcia leg. L=27 mm
    Copyright © Ruslan Panin
    Carabus (Piocarabus) sui Imura & Zhou, 1998
    Carabus (Piocarabus) sui Imura & Zhou, 1998
    Female adult
    CE-China, Henan, Mt Wangwu, V.2007, local collector. L= 33 mm
    Copyright © Ruslan Panin
    Carabus (Oreocarabus) ghiliani La Ferte-Senectere, 1847
    Carabus (Oreocarabus) ghiliani La Ferte-Senectere, 1847
    Female adult
    C-Spain: Sierra de Guadarrama, Puerto de Cotos (Madrid), 20.IX.2018, F.Rodriguez leg., L=22 mm
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) breuningi Csiki, 1927 (loc.typ.)
    Carabus (Mesocarabus) breuningi Csiki, 1927 (loc.typ.)
    Male adult
    NW-Spain, Orense, Orensano Central Massif, Monte de Cabeza de Manzaneda, 25.VII.1988, F. Rodrigues leg. L=25 mm
    Copyright © Ruslan Panin
    Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
    Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
    Endophallus
    S Russia, C-Caucasus: N Ossetia, Chekhatsirtit Mt range, Mt Tepli h=3000 m
    Copyright © Ruslan Panin
    Carabus (Alipaster) pupulus pupulus Morawitz, 1889
    Carabus (Alipaster) pupulus pupulus Morawitz, 1889
    Female adult
    S Kyrgyzstan, Inner Tian Shan, Dzhany-Dzher Mt range, upper Mudyuryum riv. Ak-Baytal gorge h~3300 m, 15.VII.2011, S.Toporov leg. L=16 mm
    Copyright © Ruslan Panin
    Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
    Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
    Male adult
    Russia, C-Caucasus: N Ossetia, Chekhatsirtit Mt range, Mt Tepli h=3000 m, 21.VII.2009. /det. Solodovnikov I.A, L=20.5 mm
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) cantabricus moroderi (as magdalenai Meyer & Mollard, 1996)
    Carabus (Mesocarabus) cantabricus moroderi (as magdalenai Meyer & Mollard, 1996)
    Endophallus
    NW-Spain, Leon prov., Sierra de Gistredo, Pto. de Magdalena,
    Copyright © Ruslan Panin
    Carabus (Goniocarabus) perelloi Casale, 1979
    Carabus (Goniocarabus) perelloi Casale, 1979
    Male adult
    N-Afghanistan, Hindukush, Provinz Baghlan, N of Kabul, N Salang Pass, subalp., h = 2200 m, 35°27'.494N / 68°58.399'E, Mai 2011, leg. Christoph Reuter (TOPOTYPUS)
    Copyright © Igor Solodovnikov
    Carabus (Tribax) adelphus rex Gottwald, 1980
    Carabus (Tribax) adelphus rex Gottwald, 1980
    Endophallus
    Abkhazia, Tkuarchalsky district, Kodorsky Mt.R., it is right board of the Bashishkvara River, beechen forest + rhododendron, h = 1693 m, 15-23.07.2018, leg. I.A. Solodovnikov
    Copyright © Igor Solodovnikov
    Carabus (Tribax) adelphus adelphus Rost, 1892 SW-Caucasus:"Samursakan"(=nr.Gali & Anara); Gali env.:Okumi
    Carabus (Tribax) adelphus adelphus Rost, 1892 SW-Caucasus:"Samursakan"(=nr.Gali & Anara); Gali env.:Okumi
    Endophallus
    Georgia, Zemo Svaneti., W slopes of Ingurishi Mt., right bank of Inguri riv., 09.V-20.VI.2018 leg. D. Fominykh
    Copyright © Igor Solodovnikov
    Carabus (Tribax) nacharensis Rost, 1893
    Carabus (Tribax) nacharensis Rost, 1893
    Endophallus
    Abkhazia, vall. Kluch River, 13.08.1928
    Copyright © Igor Solodovnikov
    Carabus (Tribax) adelphus shtavlerensis Fominykh, Zamotajlov et Titarenko, 2018
    Carabus (Tribax) adelphus shtavlerensis Fominykh, Zamotajlov et Titarenko, 2018
    Endophallus
    Georgia, Zemo Svaneti, left bank of Nenskra River near Tita Vill., Shtavler Mt. Range, NW slopes of Utur Mt., 43°04'N, 42°11'E, 1048 m, 14.IV–13.VI.2016 (D. Fominykh) HOLOTYPE
    Copyright © Igor Solodovnikov
    Carabus (Tribax) adelphus shtavlerensis Fominykh, Zamotajlov et Titarenko, 2018
    Carabus (Tribax) adelphus shtavlerensis Fominykh, Zamotajlov et Titarenko, 2018
    Male adult
    Georgia, Zemo Svaneti, left bank of Nenskra River near Zemo Margi Vill., 1490 m, 43°04'23.5''N, 42°11'55.2''E, 10.V.2018 (D. Fominykh) PARATYPE
    Copyright © Igor Solodovnikov
    Carabus (Tribax) adelphus shtavlerensis Fominykh, Zamotajlov et Titarenko, 2018
    Carabus (Tribax) adelphus shtavlerensis Fominykh, Zamotajlov et Titarenko, 2018
    Male adult
    Abkhazia, Tkuarchalsky district, Kodorsky Mt.R., a source of the river Mokvi, a talus under the pass, h = 2635 m, 17-23.07.2018, leg. I.A. Solodovnikov
    Copyright © Igor Solodovnikov
    Carabus (Rhigocarabus) turnai turnai Deuve, 1994
    Carabus (Rhigocarabus) turnai turnai Deuve, 1994
    Male adult
    China, Tibet, W-Sichuan, pass btw Sêrtar / Luhuo h=4200 m, 28.VI.2011, V.Patrikeev leg.L=19 mm
    Copyright © Ruslan Panin
    Carabus (Rhigocarabus) turnai turnai Deuve, 1994
    Carabus (Rhigocarabus) turnai turnai Deuve, 1994
    Female adult
    China, Tibet, W-Sichuan, pass btw Sêrtar / Luhuo h=4200 m, 28.VI.2011, V.Patrikeev leg. L=21 mm
    Copyright © Ruslan Panin
    Carabus (Coptolabrus) elysii xiaoganicus Deuve et Remond, 2016
    Carabus (Coptolabrus) elysii xiaoganicus Deuve et Remond, 2016
    Male adult
    China, Hubei, Xiaogan, Mt Shuangfeng Shan, VII.2016 L=37 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018
    Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
    Male adult
    W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg.
    Copyright © Ruslan Panin
    Carabus (Coptolabrus) gemmifer gemmifer (as loczyi Csiki, 1927)
    Carabus (Coptolabrus) gemmifer gemmifer (as loczyi Csiki, 1927)
    Female adult
    S-China, W-Yunnan: Dali city, Mt Jizushan, VI.2018, L=34 mm
    Copyright © Ruslan Panin
    Carabus (Coptolabrus) elysii xiaoganicus Deuve et Remond, 2016
    Carabus (Coptolabrus) elysii xiaoganicus Deuve et Remond, 2016
    Endophallus
    China, Hubei, Xiaogan, Mt Shuangfeng Shan,
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) davidis helmutschuetzei Kleinfeld, 1995
    Carabus (Apotomopterus) davidis helmutschuetzei Kleinfeld, 1995
    Male adult
    SE-China, SW-Jiangxi:S of Yongxin: Mt.Jinggangshan,V.2012, L=35 mm
    Copyright © Ruslan Panin
    Carabus (Coptolabrus) lafossei buchi Hauser, 1913
    Carabus (Coptolabrus) lafossei buchi Hauser, 1913
    Male adult
    China Zhejiang Yuyao - Leg. Buch 1925
    Copyright © Vittorino Monzini
    Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
    Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
    Endophallus
    NC-Spain, Basque Co, Álava, Sierra de Gorbea.
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
    Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
    Male adult
    NC-Spain, Basque Co, Álava, Sierra de Gorbea, VI.2014, L=29 mm
    Copyright © Ruslan Panin
    Carabus (Coptolabrus) smaragdinus furumiellus Deuve, 1994
    Carabus (Coptolabrus) smaragdinus furumiellus Deuve, 1994
    Female adult
    NE-China, E-Liaoning, Benxi, Mt Liuershan, VIII.1998. L=39 mm
    Copyright © Ruslan Panin
    Carabus (Oreocarabus) luetgensi pseudosteuarti Lapouge, 1924
    Carabus (Oreocarabus) luetgensi pseudosteuarti Lapouge, 1924
    Female adult
    Portugal, Guarda, Sierra de Estrela h=800 m, 26.VI-4.VIII.2015, L=26 mm
    Copyright © Ruslan Panin
    Carabus (Oreocarabus) luetgensi pseudosteuarti Lapouge, 1924
    Carabus (Oreocarabus) luetgensi pseudosteuarti Lapouge, 1924
    Endophallus
    Portugal, Guarda, Sierra de Estrela h=800 m,
    Copyright © Ruslan Panin
    Carabus (Oreocarabus) luetgensi pseudosteuarti Lapouge, 1924
    Carabus (Oreocarabus) luetgensi pseudosteuarti Lapouge, 1924
    Male adult
    Portugal, Guarda, Sierra de Estrela h=800 m, 26.VI-4.VIII.2015, L=24 mm
    Copyright © Ruslan Panin
    Carabus (Neoplectes) compressus rebellis Reitter, 1895
    Carabus (Neoplectes) compressus rebellis Reitter, 1895
    Male adult
    Georgia, Kvemo-Svaneti, S slopes of Svanetskiy Mt.R., S slope of Latpari pass, h=2225m, 17.VI -13.VII.2017 leg. D.Fominykh L=24.0
    Copyright © Kirill Makarov
    Carabus (Neoplectes) compressus rebellis Reitter, 1895
    Carabus (Neoplectes) compressus rebellis Reitter, 1895
    Female adult
    Georgia, Kvemo-Svaneti, S slopes of Svanetskiy Mt.R., S slope of Latpari pass, h=2225m, 17.VI -13.VII.2017 leg. D.Fominykh L=26.0 mm
    Copyright © Kirill Makarov
    Carabus (Neoplectes) compressus rebellis Reitter, 1895
    Carabus (Neoplectes) compressus rebellis Reitter, 1895
    Male genitalia
    Georgia, Kvemo-Svaneti, S slopes of Svanetskiy Mt.R., S slope of Latpari pass, h=2225m, 17.VI -13.VII.2017 leg. D.Fominykh
    Copyright © Kirill Makarov
    Carabus (Neoplectes) compressus rebellis Reitter, 1895
    Carabus (Neoplectes) compressus rebellis Reitter, 1895
    Endophallus
    Georgia, Kvemo-Svaneti, S slopes of Svanetskiy Mt.R., S slope of Latpari pass, h=2225m, 17.VI -13.VII.2017 leg. D.Fominykh
    Copyright © Kirill Makarov
    Carabus (Neoplectes) zubovi Fominykh, Zamotajlov et Titarenko, 2018
    Carabus (Neoplectes) zubovi Fominykh, Zamotajlov et Titarenko, 2018 Holotype
    Male adult
    S slopes of Svaneti Mt. Range near Lentekhi Town, S slopes of Goldash Mt., 42°49'N, 42°45'E, 2300–2800 m, 21.VI.2016, D. Fominykh, A. Zherebilo (ZISP).
    Copyright © Kirill Makarov
    Carabus (Neoplectes) compressus lailensis Belousov, 1992
    Carabus (Neoplectes) compressus lailensis Belousov, 1992
    Male adult
    Laila Mt.
    Copyright © Kirill Makarov
    Carabus (Neoplectes) daphnis tsekuricus Belousov, 1992
    Carabus (Neoplectes) daphnis tsekuricus Belousov, 1992
    Male adult
    LEFT: Lakhashuri River valley; RIGHT: Tzikuri Mt. Scale bar 10.0 mm.
    Copyright © Kirill Makarov
    Carabus (Neoplectes) polychrous svetlanae Solodovnikov, 2015
    Carabus (Neoplectes) polychrous svetlanae Solodovnikov, 2015
    Endophallus

    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) polychrous rousianus Gottwald, 1985
    Carabus (Neoplectes) polychrous rousianus Gottwald, 1985
    Endophallus

    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) polychrous polychrous
    Carabus (Neoplectes) polychrous polychrous
    Endophallus

    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) polychrous khipstaensis Solodovnikov, 2015
    Carabus (Neoplectes) polychrous khipstaensis Solodovnikov, 2015
    Endophallus

    Copyright © Igor Solodovnikov
    Carabus (Tribax) macropus Chaudoir, 1877
    Carabus (Tribax) macropus Chaudoir, 1877
    Male adult
    Daghestan: Bogosskyi Mts., Inkhokvari
    Copyright © Igor Solodovnikov
    Carabus (Tribax) macropus Chaudoir, 1877
    Carabus (Tribax) macropus Chaudoir, 1877
    Male adult
    Daghestan: Bogosskyi Mts., Inkhokvari
    Copyright © Igor Solodovnikov
    Carabus (Cupreocarabus) tianmingyii Deuve, 2015
    Carabus (Cupreocarabus) tianmingyii Deuve, 2015 Holotype
    Male adult
    Chine, Sichuan, Barkam Xian, Kangshan Xiang, Ya’erzhu, 3500-4000 mètres (Th. Deuve, Tian Mingyi et Song Zhao, 14-18 juillet 2007), in coll. South China Agricultural University, Canton.
    Copyright © Thierry Deuve
    Carabus (Pseudocranion) zhanglaensis potrikeevi Deuve, 2015
    Carabus (Pseudocranion) zhanglaensis potrikeevi Deuve, 2015 Holotype
    Male adult
    Chine, Gansu, env. 60 km à l’est de Têwo, Palanama, 4000 mètres, in coll. Andreï Gorodinski, Moscou (Russie).
    Copyright © Thierry Deuve
    Carabus (Apotomopterus) jingliae Deuve & Tian, 2019
    Carabus (Apotomopterus) jingliae Deuve & Tian, 2019 Holotype
    Male adult
    Chine, Guangxi, Luzhai Xian, Xiangqiaoyan National Geopark, grotte She Dong, 194 mètres, 24°45’32’’N-109°39’21’’E, courant dans une salle de la partie totalement obscure de la grotte (Cheng Jingli leg., 29 janvier 2019), in coll. South China Agricultural University, Canton (Chine).
    Copyright © Thierry Deuve
    Carabus (Apotomopterus) bousquetellus luchunicus Deuve, 2019
    Carabus (Apotomopterus) bousquetellus luchunicus Deuve, 2019 Holotype
    Male adult
    Chine, Yunnan, Monts Ailao Shan, Lüchun (MNHN)
    Copyright © Thierry Deuve
    Carabus (Apotomopterus) caoyutangicus jiangsuensis Deuve, 2019
    Carabus (Apotomopterus) caoyutangicus jiangsuensis Deuve, 2019 Holotype
    Female adult
    Chine, Jiangsu, Changzhou, Longhutang (Jatua), in coll. Jaroslav Turna, Kostelec na Hane (République Tchèque).
    Copyright © Thierry Deuve
    Carabus (Apotomopterus) shun pseudocantonensis Deuve & Tian, 2019
    Carabus (Apotomopterus) shun pseudocantonensis Deuve & Tian, 2019 Holotype
    Male adult
    Chine, Guangdong, Guangning Xian, Shijian Zhen, Shanxin Cun, en forêt de bambous (Qin Changsheng, 25 mai 2016), in coll. South China Agricultural University (SCAU),
    Copyright © Thierry Deuve
    Carabus (Morphocarabus) tarbagataicus altaicola Deuve & Tian, 2019
    Carabus (Morphocarabus) tarbagataicus altaicola Deuve & Tian, 2019 Holotype
    Female adult
    Chine, Xinjiang, Préfecture d’Altaï, Monts Altaï, au nord de la ville d’Altaï, Parc forestier de Xiaodonggou (Chen Liusheng, 8.VIII. 2014), in coll. South China Agricultural University, Canton.
    Copyright © Thierry Deuve
    Carabus (Pseudocoptolabrus) taliensis kawagarbo Deuve, 2018
    Carabus (Pseudocoptolabrus) taliensis kawagarbo Deuve, 2018 Holotype
    Male adult
    Chine, Nord-Ouest du Yunnan, Monts Hengduan Shan, au nord du Mont Meili Xueshan (Kawagarbo), à l’ouest du village de Shuaqu, 4268 mètres, 28°35’12’’N-98°40’36’’E, in coll. R. Sehnal, à Unhošt (République Tchèque).
    Copyright © Thierry Deuve
    Carabus (Aulonocarabus) canaliculatus biraensis Deuve & Reuter, 2019
    Carabus (Aulonocarabus) canaliculatus biraensis Deuve & Reuter, 2019 Holotype
    Male adult
    Russie, Extrême-Orient Russe, Oblast Autonome Juif (EAO), environs de Khabarovsk, au nord de Bira, Mont Studentcheskaïa, haut bassin de la rivière Rousskaïa, 1200 mètres, A. Brinev, 12-19.VII. 2014 (MNHN)
    Copyright © Thierry Deuve
    Carabus (Aulonocarabus) gaschkewitschi brinevellus Deuve & Reuter, 2019
    Carabus (Aulonocarabus) gaschkewitschi brinevellus Deuve & Reuter, 2019 Holotype
    Male adult
    Russie, Sibérie, région de Khabarovsk, district de Verkhneboureinskij, Monts Ezop, 1900 mètres, 52°34’N-133°46’E (A. Brinev, 15-21 juillet 2005), in coll. C. Reuter, à Berlin (Allemagne).
    Copyright © Thierry Deuve
    Carabus (Aulonocarabus) gaschkewitschi ippolitovi Berlov, 2004
    Carabus (Aulonocarabus) gaschkewitschi ippolitovi Berlov, 2004 Paratype
    Female adult
    Иркутская область, Качугский район, заповедник "Байкало-Ленский", Байкальский хребет близ мыса Елохин, каменистая горная тундра на склоне западной экспозиции, 1500 м, 25-VI-2003, leg. О.Берлов. 20 мм. Image courtesy of O. Berlov, at www.zin.ru
    Copyright © - - -
    Carabus (Aulonocarabus) gaschkewitschi ippolitovi Berlov, 2004
    Carabus (Aulonocarabus) gaschkewitschi ippolitovi Berlov, 2004 Holotype
    Male adult
    Иркутская область, Качугский район, заповедник "Байкало-Ленский", Байкальский хребет близ мыса Елохин, каменистая горная тундра на склоне западной экспозиции, 1500 м, 25-VI-2003, leg. О.Берлов. 19 мм. Image courtesy of O. Berlov, at www.zin.ru
    Copyright © - - -
    Carabus (Aulonocarabus) gaschkewitschi raddei Morawitz, 1862
    Carabus (Aulonocarabus) gaschkewitschi raddei Morawitz, 1862
    Unsexed
    Topotypus - Россия, Читинская обл., Сохондинский зап-к., 13.08.1962. Image courtesy of D.V. Obydov, at www.zin.ru
    Copyright © - - -
    Carabus (Aulonocarabus) gaschkewitschi s.str.
    Carabus (Aulonocarabus) gaschkewitschi s.str. Lectotype
    Male adult
    fl. Amur. Image courtesy of D.V. Obydov, at www.zin.ru
    Copyright © - - -
    Carabus (Aulonocarabus) gaschkewitschi gaschkewitschi Motschulsky, 1859
    Carabus (Aulonocarabus) gaschkewitschi gaschkewitschi Motschulsky, 1859
    Male adult
    Russia, Khabarovsk reg., Bureinskii Mt. rng., VII.64 det. Obydov. Image courtesy of D.V. Obydov, at www.zin.ru
    Copyright © - - -
    Carabus (Rhigocarabus) indigestus dilbu Deuve & Kaláb, 2018
    Carabus (Rhigocarabus) indigestus dilbu Deuve & Kaláb, 2018 Holotype
    Male adult
    Chine, Ouest-Sichuan, col à 9 km à l’ouest de Babang, (route de Dêgê), zone sud du col, 4600-4800 mètres, 31º39’N/98º43’E (pelouses alpines, éboulis), in coll. J. Kaláb, à Jinačovice (République Tchèque).
    Copyright © Thierry Deuve
    Carabus (Rhigocarabus) indigestus ronggaicus Deuve & Kaláb, 2018
    Carabus (Rhigocarabus) indigestus ronggaicus Deuve & Kaláb, 2018 Holotype
    Male adult
    Chine, Ouest-Sichuan, col à 17 km au SE-SSE de Ronggai, 4800 mètres, 30º59’N/99º00’E (pelouses alpines, éboulis), in coll. J. Kaláb, à Jinačovice (République Tchèque).
    Copyright © Thierry Deuve
    Carabus (Rhigocarabus) ladygini babangensis Deuve & Kaláb, 2018
    Carabus (Rhigocarabus) ladygini babangensis Deuve & Kaláb, 2018 Holotype
    Male adult
    Chine, Ouest-Sichuan, col à 9 km à l’ouest de Babang, (route de Dêgê), zone sud du col, 4600-4800 mètres, 31º39’N/98º43’E (pelouses alpines, éboulis), in coll. J. Kaláb, à Jinačovice (République Tchèque).
    Copyright © Thierry Deuve
    Carabus (Pagocarabus) lama achaicus Deuve & Kaláb, 2018
    Carabus (Pagocarabus) lama achaicus Deuve & Kaláb, 2018 Holotype
    Male adult
    Chine, Ouest-Sichuan, col à 9 km au nord-ouest d’Acha, zone sud du col, 4700 mètres, 31º06’N/99º29’E (pelouses alpines, éboulis), in coll. J. Kaláb, à Jinačovice (République Tchèque).
    Copyright © Thierry Deuve
    Carabus (Calocarabus) aurocostulus Deuve & Kaláb, 2018
    Carabus (Calocarabus) aurocostulus Deuve & Kaláb, 2018 Holotype
    Male adult
    Chine, Ouest-Sichuan, montagnes à 15 km à l’ouest de Keluodong, 4600-4700 mètres, 31º58’N/98º32’E (pelouses alpines, éboulis), in coll. J. Kaláb, à Jinačovice (République Tchèque).
    Copyright © Thierry Deuve
    Carabus (Calocarabus) punctatocostulus Deuve & Kaláb, 2018
    Carabus (Calocarabus) punctatocostulus Deuve & Kaláb, 2018 Holotype
    Female adult
    Chine, Ouest-Sichuan, montagnes à 15 km à l’ouest de Keluodong, 4600-4700 mètres, 31º58’N/98º32’E (pelouses alpines, éboulis), in coll. J. Kaláb, à Jinačovice (République Tchèque).
    Copyright © Thierry Deuve
    Carabus (Apotomopterus) cavazzutiellus neptunus Deuve, 2018
    Carabus (Apotomopterus) cavazzutiellus neptunus Deuve, 2018 Holotype
    Female adult
    Chine, Sud-Est Sichuan, Huangjing Forest Park, 1480 mètres, 28°11’N-105°44’E (Jatua), in coll. J. Turna, à Kostelec na Hane (République Tchèque).
    Copyright © Thierry Deuve
    Carabus (Apotomopterus) longeantennatus acorepoides Deuve, 2018
    Carabus (Apotomopterus) longeantennatus acorepoides Deuve, 2018 Holotype
    Female adult
    Chine, Chongqing, Wulingshan Forest Park, 1690 m, 29°33’N-107°36’E (Jatua), in coll. J. Turna, à Kostelec na Hane (République Tchèque).
    Copyright © Thierry Deuve
    Carabus (Tribax) lailensis stefaniianus Deuve, 2018
    Carabus (Tribax) lailensis stefaniianus Deuve, 2018 Holotype
    Male adult
    Géorgie, Basse-Svanétie, au nord de Lentekhi, versant sud de la chaîne de Svanétie, 1000 m., Th. Deuve et A. Matocq, juin 2016. (MNHN)
    Copyright © Thierry Deuve
    C. (Cupreocarabus) kuceraianus subemili Deuve & Kaláb, 2017
    C. (Cupreocarabus) kuceraianus subemili Deuve & Kaláb, 2017 Holotype
    Male adult
    Chine, Sichuan centro-occidental, montagnes à 15 km à l’est-nord-est de Jintang, 4100 mètres, 30º25’N/102º25’E (pelouses alpines, éboulis), in coll. J. Kaláb, à Jinačovice (République Tchèque).
    Copyright © Thierry Deuve
    Carabus (Imaibius) leepai sabatinellii Deuve & Kaláb, 2017
    Carabus (Imaibius) leepai sabatinellii Deuve & Kaláb, 2017 Holotype
    Male adult
    Pakistan, Azad Kashmir, District de Muzaffarabad, Pir Chanasi (G. Sabatinelli, 15 août 2012), in coll. J. Kaláb, à Jinačovice (République Tchèque).
    Copyright © Thierry Deuve
    Carabus (Apotomopterus) yundongbeicus mopanensis Deuve, 2017
    Carabus (Apotomopterus) yundongbeicus mopanensis Deuve, 2017 Holotype
    Male adult
    Chine, Yunnan, Mopan Shan, 2300 mètres, 23°57’N-101°58’E (Jatua leg.), in coll. J. Turna, à Kostelec na Hane (République Tchèque).
    Copyright © Thierry Deuve
    Carabus (Neoplectes) daphnis boulbenellus Deuve et Stéfani, 2017
    Carabus (Neoplectes) daphnis boulbenellus Deuve et Stéfani, 2017 Holotype
    Male adult
    Géorgie, Basse-Ratcha, à l’est de Nikortsminda, Skhartali, 1200-1300 m., Th. Deuve, P. Stéfani leg. (MNHN).
    Copyright © Thierry Deuve
    Carabus (Cupreocarabus) mollardianus Deuve, 2016
    Carabus (Cupreocarabus) mollardianus Deuve, 2016 Holotype
    Female adult
    Chine, Sichuan, Baoxing Xian, Monts Jiajin Shan, Lushan
    Copyright © Thierry Deuve
    Carabus (Apotomopterus) protenes erlangbaensis Deuve, 2016
    Carabus (Apotomopterus) protenes erlangbaensis Deuve, 2016 Holotype
    Male adult
    Chine, Guizhou nord occidental, Erlangba, 1200- 1350 m, 28°33’N-106°00’E (Jatua), in coll. Jaroslav Turna, à Kostelec na Hane (République Tchèque).
    Copyright © Thierry Deuve
    Carabus (Apotomopterus) rugulothorax Deuve, 2016
    Carabus (Apotomopterus) rugulothorax Deuve, 2016 Holotype
    Female adult
    Chine, Guizhou nord occidental, Erlangba, 1200-1350 m, 28°33’N-106°00’E (Jatua), in coll. Jaroslav Turna, à Kostelec na Hane (République Tchèque).
    Copyright © Thierry Deuve
    C. (Rhigocarabus) itzingeri gongshanensis Deuve, 2016
    C. (Rhigocarabus) itzingeri gongshanensis Deuve, 2016 Holotype
    Male adult
    Chine, Yunnan, Monts Gaoligong Shan, Préfecture de Nujiang, Gongshan Xian, 21 km à vol d’oiseau à l’ouest de Gongshan, col entre Gongshan et Dulong, 3300-3680 mètres, 27,69655°N-98,45349°E (D. H. Kavanaugh, C. E. Griswold, H. B. Liang, D. Ubick, D. Z. Dobg leg., 16-17 juillet 2000, in coll. Institut Zoologique, Pékin.
    Copyright © Thierry Deuve
    Carabus (Neoplectes) komarowi mestiaicus Deuve, 2016
    Carabus (Neoplectes) komarowi mestiaicus Deuve, 2016 Holotype
    Female adult
    Géorgie, Haute Svanétie, 10 km au nord de Mestia, vers les lacs de Koruldi, env. 2500 mètres, in coll. A. Wřzecionko, à Horní Sucha (République Tchèque).
    Copyright © Thierry Deuve
    Carabus (Coptolabrus) pustulifer lengchongensis Deuve, 2016
    Carabus (Coptolabrus) pustulifer lengchongensis Deuve, 2016 Holotype
    Male adult
    Chine, Guizhou nord-occidental, Lengchong, 1900 mètres, 27°06’N-105°15’E (Jatua), in coll. Jaroslav Turna, à Kostelec na Hane (République Tchèque).
    Copyright © Thierry Deuve
    Carabus (Pseudocoptolabrus) watanabei valdemar Deuve, 2016
    Carabus (Pseudocoptolabrus) watanabei valdemar Deuve, 2016 Holotype
    Male adult
    Chine, Yunnan, Dêqên, monts Hengduan Shan, près du col de Doker La, 3600 m. (in coll. I. Brunk)
    Copyright © Thierry Deuve
    Carabus (Neoplesius) nanschanicus heishuicus Deuve, 2016
    Carabus (Neoplesius) nanschanicus heishuicus Deuve, 2016 Holotype
    Male adult
    Chine, Sichuan central, Heishui Xian, montagnes à 14 km au S-SSE de Heishui, 4200-4400m, 31º56´N/102º59´E (prairies alpines, éboulis), in coll. J. Kaláb, à Jinačovice (République Tchèque).
    Copyright © Thierry Deuve
    Carabus (Pseudocoptolabrus) watanabei valdemar Deuve, 2016
    Carabus (Pseudocoptolabrus) watanabei valdemar Deuve, 2016 Paratype
    Endophallus
    Chine, Yunnan, Dêqên, monts Hengduan Shan, près du col de Doker La, 3600 m
    Copyright © Thierry Deuve
    Carabus (Apotomopterus) nigrovagans Deuve, 2016
    Carabus (Apotomopterus) nigrovagans Deuve, 2016 Holotype
    Male adult
    Chine, Guangxi, Jinxiu Xian, Mont Dayao Shan, Laoshan. in coll. J.-L. & S. Remond, à Sainte-Adresse.
    Copyright © Thierry Deuve
    Carabus (Apotomopterus) nigrovagans Deuve, 2016
    Carabus (Apotomopterus) nigrovagans Deuve, 2016 Paratype
    Female adult
    Chine, Guangxi, Jinxiu Xian, Mont Dayao Shan, Laoshan. in coll. J.-L. & S. Remond, à Sainte-Adresse.
    Copyright © Thierry Deuve
    Carabus (Coptolabrus) elysii xiaoganicus Deuve et Remond, 2016
    Carabus (Coptolabrus) elysii xiaoganicus Deuve et Remond, 2016 Holotype
    Male adult
    Chine, Hubei, Xiaogan, Mt Shuangfeng Shan, in coll. J.-L. & S. Remond, à Sainte-Adresse.
    Copyright ©
    Carabus (Cupreocarabus) balangicus qiaoqiensis Deuve & Kaláb, 2015
    Carabus (Cupreocarabus) balangicus qiaoqiensis Deuve & Kaláb, 2015 Holotype
    Male adult
    Chine, Sichuan central, montagnes à 13 km au nord-est de Qiaoqi, 3750-4200 mètres, 30°47’N/102°50-51’E, en pelouse alpine et éboulis, in coll. Jaroslav Kaláb, à Jinačovice (République Tchèque).
    Copyright © Thierry Deuve
    Carabus (Cupreocarabus) kuceraianus emili Deuve et Kaláb, 2015
    Carabus (Cupreocarabus) kuceraianus emili Deuve et Kaláb, 2015 Holotype
    Male adult
    Chine, Sichuan central, env. 11 km au nord-ouest / nord-nord-ouest de Jintang, abords orientaux du col de Muzhuliangzi, 4000 mètres, 30°27’N/102°13’E, en pelouse alpine et éboulis, in coll. Jaroslav Kaláb, à Jinačovice (République Tchèque).
    Copyright © Thierry Deuve
    Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
    Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
    Female adult
    W Ukraine, Transcarpathian lowland, near Chop, 11.VI-19.VII.2019, Panin R leg. L=29 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
    Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
    Male adult
    W Ukraine, Transcarpathian lowland, near Chop, 11.VI-19.VII.2019, Panin R leg. L=30 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
    Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
    Female adult
    W Ukraine, Transcarpathian lowland, near Chop, 11.VI-19.VII.2019, Panin R leg. L=31 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
    Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
    Female adult
    W Ukraine, Transcarpathian lowland, near Chop, 11.VI-19.VII.2019, Panin R leg. L=29 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
    Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
    Male adult
    W Ukraine, Transcarpathian lowland, near Chop, 11.VI-19.VII.2019, Panin R leg. L=28 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
    Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
    Female adult
    W Ukraine, Transcarpathian lowland, near Chop, 11.VI-19.VII.2019, Panin R leg. L=30 mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) avinovi Semenov & Znojko, 1932
    Carabus (Megodontus) avinovi Semenov & Znojko, 1932
    Female adult
    S Sakhalin, Dolinsky distr., Firsovo vill. env., 3.VII.2017 . Фотография с сайта www.zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
    Copyright © Maksim Smirnov
    Carabus (Acoptolabrus) lopatini Morawitz, 1886
    Carabus (Acoptolabrus) lopatini Morawitz, 1886
    Male adult
    S Sakhalin, Yuzhno Sakhalinsk, Tschekhova Mt., 800 m.,, 28.VI.2017 ; Фотография с сайта www.zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
    Copyright © Maksim Smirnov
    Carabus (Ohomopterus) kaempferi Thomson, 1875
    Carabus (Ohomopterus) kaempferi Thomson, 1875 Holotype
    Male adult
    Image courtesy of Biological Museum, Lund University
    Copyright © - - -
    Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019
    Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
    Male adult
    W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 4-24.V.2010, Panin R. leg. L= 29 mm.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) excellens ciscarpathicusn (as rareulensis Born, 1907)
    Carabus (Morphocarabus) excellens ciscarpathicusn (as rareulensis Born, 1907)
    Male adult
    NE-Romania, Rarau Mts. h-1300m, VI.2014, J. Kynkor leg. L=23 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) excellens ciscarpathicus (as polonicus Lomnicki, 1886)
    Carabus (Morphocarabus) excellens ciscarpathicus (as polonicus Lomnicki, 1886)
    Male adult
    SW Poland: Przemyśl env. photo by Mieczyslaw Stachowiak
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) excellens ciscarpathicus (as polonicus Lomnicki, 1886)
    Carabus (Morphocarabus) excellens ciscarpathicus (as polonicus Lomnicki, 1886)
    Female adult
    W Ukraine, Lviv Region, Zhydahciv distr., Borodchytsi vill.,the right bank of Dniester river, 11-16.VII.2012, Panin. R leg. L=27 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019
    Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Holotype
    Endophallus
    W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019
    Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Holotype
    Male adult
    W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 28.V-9.VI.2013, Panin R. leg. L=29 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019
    Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
    Female adult
    W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 28.V-9.VI.2013, Panin R. leg. L= 28 mm.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019
    Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
    Male adult
    W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 18.V-12.VI.2014, L=28 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019
    Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019
    Habitat
    Biletsky flood-plain forest,
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019
    Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
    Male adult
    W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 27.IV-13.VI.2018, Panin R. leg. L= 28 mm.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019
    Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
    Male adult
    W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 27.IV-13.VI.2018, Panin R. leg. L= 29 mm.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019
    Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
    Female adult
    W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 20.V-25.VI.2008, Panin R. leg. L= 31 mm.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019
    Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
    Female adult
    W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 20.V-25.VI.2008, Panin R. leg. L= 30 mm.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019
    Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
    Male adult
    W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 12.V-2.VI.2009, Panin R. leg. L= 29 mm.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019
    Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
    Female adult
    W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 27.IV-13.VI.2018, Panin R. leg. L= 31 mm.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019
    Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
    Male adult
    W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., Panin R. leg. L= 28 mm.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) excellens excellens (ab.goldeggi Duftschmid, 1812)
    Carabus (Morphocarabus) excellens excellens (ab.goldeggi Duftschmid, 1812)
    Male adult
    W-Ukraine, Podolian Tovtry, Ternopol reg., Pidvolochysk distr., Shevchenkove vill., steppe sites, 6.V-7.VI.2012, Panin R. leg. L=24 mm
    Copyright © Ruslan Panin
    Carabus (Neoplectes) martviliensis mzekalae Fominykh, Zamotajlov et Titarenko, 2019
    Carabus (Neoplectes) martviliensis mzekalae Fominykh, Zamotajlov et Titarenko, 2019 Paratype
    Male+Female adult
    Imereti, left bank of river Semi near Melouri, 475 m, 19.V–12.VII.2019, D. Fominykh, A. Zubov, B. Dubinin & D. Safronov leg.
    Copyright © Kirill Makarov
    Carabus (Neoplectes) martviliensis mzekalae Fominykh, Zamotajlov et Titarenko, 2019
    Carabus (Neoplectes) martviliensis mzekalae Fominykh, Zamotajlov et Titarenko, 2019 Paratype
    Male+Female adult
    Imereti, left bank of river Tskhenistskali near Dzedzileti, 620 m, 16.V–13.VII.2019, D. Fominykh, A. Zubov, B. Dubinin & D. Safronov leg.
    Copyright © Kirill Makarov
    Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
    Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
    Male+Female adult
    Imereti, S slopes of Askhi plateau near Zeda-Gordi, NE slope of Mt. Gulkuna, 980 m, 20.V–14.VII.2019, D. Fominykh & A. Zubov leg.
    Copyright © Kirill Makarov
    Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
    Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
    Male+Female adult
    Samegrelo, Doberazeni vic., left bank of river Tekhuri, 500 m, 16.IV–31.V.2017, D. Fominykh leg.
    Copyright © Kirill Makarov
    Carabus (Neoplectes) martviliensis Retezar & Djavelidze, 1992
    Carabus (Neoplectes) martviliensis Retezar & Djavelidze, 1992
    Endophallus
    Zeda Gordi env., 980 m and Doberazeni env., 600 m.
    Copyright © Kirill Makarov
    Carabus (Neoplectes) C. martviliensis mzekalae
    Carabus (Neoplectes) C. martviliensis mzekalae
    Endophallus
    Melouri env., 475 m and Dzedzileti env. , 770 m
    Copyright © Kirill Makarov
    Carabus (Diocarabus) pusillus Lapouge, 1915
    Carabus (Diocarabus) pusillus Lapouge, 1915 Paralectotypes
    Female adult
    Image courtesy of MNHN
    Copyright © MNHN
    Carabus (Orinocarabus) sylvestris transsylvanicus Dejean, 1826
    Carabus (Orinocarabus) sylvestris transsylvanicus Dejean, 1826
    Endophallus
    W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone,
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) sylvestris transsylvanicus Dejean, 1826
    Carabus (Orinocarabus) sylvestris transsylvanicus Dejean, 1826
    Male adult
    W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=21 mm
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) sylvestris transsylvanicus Dejean, 1826 (aberrant)
    Carabus (Orinocarabus) sylvestris transsylvanicus Dejean, 1826 (aberrant)
    Male adult
    W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg.
    Copyright © Ruslan Panin
    Carabus (Procerus) scabrosus vardziaensis Deuve, 2017
    Carabus (Procerus) scabrosus vardziaensis Deuve, 2017 Holotype
    Female adult

    Copyright © Ruslan Panin
    Carabus (Rhigocarabus) gigolo Heinz & Brezina, 1996
    Carabus (Rhigocarabus) gigolo Heinz & Brezina, 1996
    Endophallus
    China, Tewo. det. Anichtchenko, A. 2020
    Copyright © Alexander Anichtchenko
    Carabus (Sigenacarabus) draco lubasha Deuve, 2012
    Carabus (Sigenacarabus) draco lubasha Deuve, 2012
    Male adult
    China, C Sichuan, Jiyuding shan, 30 km N of Wenchuan, 3250 m 2.7.2017, leg. V. Patrikeev (cAA). det. Anichtchenko A. 2020
    Copyright © Alexander Anichtchenko
    Carabus (Pseudocoptolabrus) taliensis lijiangensis Deuve, 1990
    Carabus (Pseudocoptolabrus) taliensis lijiangensis Deuve, 1990
    Male adult
    N Yunnan, S slope Yulong mts., near Lijiang, 3900m, 15-25.6.2016, leg. V. Patrikeev. det. A. Anichtchenko, 2020
    Copyright © Alexander Anichtchenko
    Carabus (Deroplectes) arcanus hissarianus Semenov, 1900
    Carabus (Deroplectes) arcanus hissarianus Semenov, 1900
    Male adult
    Tadzhikistan, near Dushanbe, env. Vill. Varzob, h = 1120 m, 38º46’27” N / 68º49’03” Е, 19.IV.1992, leg. V.Kukuruza
    Copyright © Igor Solodovnikov
    Carabus (Deroplectes) klapperichianus coiffaitianus Deuve, 1990
    Carabus (Deroplectes) klapperichianus coiffaitianus Deuve, 1990
    Male adult
    Tadzhikistan: Badakhshan: Ishkashim Mt.R.: Kuh-i-Lal, VI.1979 leg. I.K. Lopatin (TOPOTYPUS)
    Copyright © Igor Solodovnikov
    Carabus (Deroplectes) klapperichianus coiffaitianus Deuve, 1990
    Carabus (Deroplectes) klapperichianus coiffaitianus Deuve, 1990
    Endophallus
    Tadzhikistan: Badakhshan: Ishkashim Mt.R.: Kuh-i-Lal, VI.1979 leg. I.K. Lopatin
    Copyright © Igor Solodovnikov
    Carabus (Deroplectes) klapperichianus coiffaitianus Deuve, 1990
    Carabus (Deroplectes) klapperichianus coiffaitianus Deuve, 1990
    Female adult
    Tadzhikistan: Badakhshan: Ishkashim Mt.R.: Kuh-i-Lal, VI.1979 leg. I.K. Lopatin (TOPOTYPUS)
    Copyright © Igor Solodovnikov
    Carabus (Deroplectes) klapperichianus salangicus Kleinfeld & Reuter, 2010
    Carabus (Deroplectes) klapperichianus salangicus Kleinfeld & Reuter, 2010 Paratype
    Female adult
    N-Afghanistan, Hindukush Mt.Rng, Salang Pass, Nordseite, 25°27'.494N, 68°58.399'E, ca. reg. subalp., h = 2200-2400 m, V.2010, leg. Christoph Reuter
    Copyright © Igor Solodovnikov
    Carabus (Cechenochilus) kokujewi Semenov, 1898
    Carabus (Cechenochilus) kokujewi Semenov, 1898
    Male adult
    Russia, Karachai-Cherkessia, vall. Daut Riv., near Glacier Ryndzhi-Murudzhu, h = 3050 m, 22.VIII.2011, leg. I.A. Solodovnikov, S.V. Solodovnikova
    Copyright © Igor Solodovnikov
    Carabus (Cechenochilus) kokujewi Semenov, 1898
    Carabus (Cechenochilus) kokujewi Semenov, 1898
    Female adult
    Russia, Karachai-Cherkessia, vall. Daut Riv., near Glacier Ryndzhi-Murudzhu, h = 3100 m, 22.VIII.2011, leg. I.A. Solodovnikov, S.V. Solodovnikova
    Copyright © Igor Solodovnikov
    Carabus (Calocarabus) aristochroides luoxuensis Deauve & Kalab, 2019
    Carabus (Calocarabus) aristochroides luoxuensis Deauve & Kalab, 2019 Holotype
    Male adult
    Chine, Sichuan occidental, 4800-4900 mètres, col à 23 km au nord-nord-est de Luoxu, 32º40’N/98º03-04’E (prairies alpines et éboulis). in coll. J. Kaláb, à Jinačovice (République Tchèque)
    Copyright © Thierry Deuve
    Carabus (Rhigocarabus) dungchen Deuve & Kalab, 2019
    Carabus (Rhigocarabus) dungchen Deuve & Kalab, 2019 Paratype
    Male adult
    Chine, Sichuan occidental, montagnes à 13 km au S-SSW de Cuo’a, 4700 mètres, 31º42’N/99º24’E (prairies alpines et éboulis).
    Copyright © Thierry Deuve
    Carabus (Rhigocarabus) ladygini ganzicola Deuve & Kalab, 2019
    Carabus (Rhigocarabus) ladygini ganzicola Deuve & Kalab, 2019 Holotype
    Male adult
    Chine, Sichuan occidental, montagnes à 17 km au nord-nord-ouest de Ganzi (= Garzê), 31º46’N/99º56’E, 4700-4800 mètres (prairies alpines et éboulis)
    Copyright © Thierry Deuve
    Carabus (Neoplesius) lama latitatus Deuve & Kalab, 2019
    Carabus (Neoplesius) lama latitatus Deuve & Kalab, 2019 Holotype
    Male adult
    Chine, Sichuan occidental, montagnes à 17 km au nord-nord-ouest de Ganzi (= Garzê), 31º46’N/99º56’E, 4700-4800 mètres (prairies alpines et éboulis), in coll. J. Kaláb, à Jinačovice (République Tchèque).
    Copyright © Thierry Deuve
    Carabus (Neoplesius) lama lapidiphilus Deuve & Kalab, 2019
    Carabus (Neoplesius) lama lapidiphilus Deuve & Kalab, 2019 Holotype
    Male adult
    Chine, Sichuan occidental, montagnes à 15 km au nord-est de Baiyü, 31º19’N/98º55’E, 4750 mètres (prairies alpines et éboulis), in coll. J. Kaláb, à Jinačovice (République Tchèque).
    Copyright © Thierry Deuve
    Carabus (Neoplesius) lama paucipunctatus Deuve & Kalab, 2019
    Carabus (Neoplesius) lama paucipunctatus Deuve & Kalab, 2019 Holotype
    Male adult
    Chine, Sichuan occidental, montagnes à 20 km au sud-sud-est de Batang, 29º50’N/99º11’E, 4700-4800 mètres (prairies alpines et éboulis), in coll. J. Kaláb, à Jinačovice (République Tchèque).
    Copyright © Thierry Deuve
    Carabus (Cechenochilus) kokujewi Semenov, 1898
    Carabus (Cechenochilus) kokujewi Semenov, 1898
    Male adult
    Russia, Karachai-Cherkessia, vall. Daut Riv., near Glacier Ryndzhi-Murudzhu, h = 3050 m, 22.VIII.2011, leg. I.A. Solodovnikov, S.V. Solodovnikova
    Copyright © Igor Solodovnikov
    Carabus (Cechenochilus) kokujewi Semenov, 1898
    Carabus (Cechenochilus) kokujewi Semenov, 1898
    Female adult
    Russia, Karachai-Cherkessia, vall. Daut Riv., near Glacier Ryndzhi-Murudzhu, h = 2550 m, 23.VIII.2011, leg. I.A. Solodovnikov, S.V. Solodovnikova
    Copyright © Igor Solodovnikov
    Carabus (Cechenochilus) adangensis Gottwald, 1983
    Carabus (Cechenochilus) adangensis Gottwald, 1983
    Female adult
    Russia, Karachai-Cherkessia, Abischira-Akhuba Mt.R., env. vill. Romantic, alpin zone, h = 2470 m, 10.07-22.08.2019, leg. S.M. Pavlyuchuk, A.P. Tatulyan
    Copyright © Igor Solodovnikov
    Carabus (Morphocarabus) henningi hamarensis Shilenkov, 1996
    Carabus (Morphocarabus) henningi hamarensis Shilenkov, 1996
    Female adult
    Lake Baikal, Maritui vill. 25.7.1994, Anichtchenko A. leg. & det.
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) henningi hamarensis Shilenkov, 1996
    Carabus (Morphocarabus) henningi hamarensis Shilenkov, 1996
    Female adult
    Lake Baikal, Maritui vill. 25.7.1994, Anichtchenko A. leg. & det.
    Copyright © Alexander Anichtchenko
    Carabus (Rhigocarabus) morawitzianus morawitzianus Semenov, 1887
    Carabus (Rhigocarabus) morawitzianus morawitzianus Semenov, 1887
    Male adult
    Qinghai, Anemaqen mts pass, 20 km N of Maduo town, 4500 m, 6.7.2012, V. Patrikeev leg.; det. Anichtchenko A. 2020
    Copyright © Alexander Anichtchenko
    Carabus (Rhigocarabus) buddaicus gansuicus Deuve, 1991
    Carabus (Rhigocarabus) buddaicus gansuicus Deuve, 1991
    Male adult
    N Sichuan, Langmusi vill. 3700 m, 1-25.7.2018, Leg. V. Patrikeev. det. Anichtchenko A. 2020
    Copyright © Alexander Anichtchenko
    Carabus (Rhigocarabus) buddaicus gansuicus Deuve, 1991
    Carabus (Rhigocarabus) buddaicus gansuicus Deuve, 1991
    Endophallus
    N Sichuan, Langmusi vill 3700 m, 1-25.7.2018
    Copyright © Alexander Anichtchenko
    Carabus (Rhigocarabus) gigolo Heinz & Brezina, 1996
    Carabus (Rhigocarabus) gigolo Heinz & Brezina, 1996
    Male adult
    S Gansu, near Tewo, 8.8.2014, V. Patrikeev leg.
    Copyright © Alexander Anichtchenko
    Carabus (Rhigocarabus) roborowskii langmusiensis Deuve, 1996
    Carabus (Rhigocarabus) roborowskii langmusiensis Deuve, 1996
    Male adult
    N Sichuan, near Langmusi, 3700m 1-25.7.2018, V. Patrikeev leg. det. Anichtchenko A. 2020
    Copyright © Alexander Anichtchenko
    Carabus (Rhigocarabus) roborowskii langmusiensis Deuve, 1996
    Carabus (Rhigocarabus) roborowskii langmusiensis Deuve, 1996
    Endophallus
    N Sichuan, near Langmusi, 3700m 1-25.7.2018, V. Patrikeev leg. det. Anichtchenko A. 2020
    Copyright © Alexander Anichtchenko
    Carabus (Neoplesius) nanschanicus nocticolor Deuve & Kalab, 1992
    Carabus (Neoplesius) nanschanicus nocticolor Deuve & Kalab, 1992
    Female adult
    Quinghai-Gansu border, Dalijia pass, 50km W Linxia town, 3800m 20.7.2019, V. Patrikeev leg.
    Copyright © Alexander Anichtchenko
    Carabus (Ohomopterus) uenoi Ishikawa, 1960
    Carabus (Ohomopterus) uenoi Ishikawa, 1960
    Endophallus

    Copyright © Jong-Bong Choi
    Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
    Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
    Female adult
    W Ukraine, Transcarpathian, Marmarosh Mts., near Lug vill., h=400 m, 28.IV -26.VII.2017, Panin R. leg., L= 38 mm
    Copyright © Ruslan Panin
    Carabus (Ophiocarabus) ernsti ulastaiensis Kabak, 2019
    Carabus (Ophiocarabus) ernsti ulastaiensis Kabak, 2019 Paratype
    Male adult
    China, Xinjiang, S slopes of Boro-Horo Mt. R., ridge between Kaptshik and Zekku rivers, NE of Ulastai (= Wulasitai) Vill., alpine meadows, 43°56'07"N / 83°18'33"E – 43°56'22"N / 83°18'30"E, 3135–3280 m, 17.07.2017 leg I.I. Kabak L=16.5 mm
    Copyright © Kirill Makarov
    Carabus (Ophiocarabus) ernsti ulastaiensis Kabak, 2019
    Carabus (Ophiocarabus) ernsti ulastaiensis Kabak, 2019 Paratype
    Male genitalia
    China, Xinjiang, S slopes of Boro-Horo Mt. R., ridge between Kaptshik and Zekku rivers, NE of Ulastai (= Wulasitai) Vill., alpine meadows, 43°56'07"N / 83°18'33"E – 43°56'22"N / 83°18'30"E, 3135–3280 m, 17.07.2017 leg I.I. Kabak
    Copyright © Kirill Makarov
    Carabus (Ophiocarabus) ernsti ulastaiensis Kabak, 2019
    Carabus (Ophiocarabus) ernsti ulastaiensis Kabak, 2019 Paratype
    Endophallus
    China, Xinjiang, S slopes of Boro-Horo Mt. R., ridge between Kaptshik and Zekku rivers, NE of Ulastai (= Wulasitai) Vill., alpine meadows, 43°56'07"N / 83°18'33"E – 43°56'22"N / 83°18'30"E, 3135–3280 m, 17.07.2017 leg I.I. Kabak
    Copyright © Kirill Makarov
    Carabus (Ophiocarabus) ernsti ulastaiensis Kabak, 2019
    Carabus (Ophiocarabus) ernsti ulastaiensis Kabak, 2019 Paratype
    Female adult
    China, Xinjiang, S slopes of Boro-Horo Mt. R., ridge between Kaptshik and Zekku rivers, NE of Ulastai (= Wulasitai) Vill., alpine meadows, 43°56'07"N / 83°18'33"E – 43°56'22"N / 83°18'30"E, 3135–3280 m, 17.07.2017 leg I.I. Kabak L=17.0 mm
    Copyright © Kirill Makarov
    Carabus (Cryptocarabus) sacarum Kabak, 1998
    Carabus (Cryptocarabus) sacarum Kabak, 1998
    Endophallus
    SE-Kazakhstan, Almaty reg., C-Zailiyskiy Alatau N-slopes, Kaskalen river Valley, Izvestkovy vill., h=1700 m,
    Copyright © Ruslan Panin
    Carabus (Cryptocarabus) sacarum Kabak, 1998
    Carabus (Cryptocarabus) sacarum Kabak, 1998
    Male adult
    SE-Kazakhstan, Almaty reg., C-Zailiyskiy Alatau N-slopes, Kaskalen river Valley, Izvestkovy vill., h=1700 m, 6.V.2015, D.Kovalchuk leg./det. R.Panin L=18 mm
    Copyright © Ruslan Panin
    Carabus (Cryptocarabus) sacarum Kabak, 1998
    Carabus (Cryptocarabus) sacarum Kabak, 1998
    Female adult
    SE-Kazakhstan, Almaty reg., C-Zailiyskiy Alatau N-slopes, Kaskalen river Valley, Izvestkovy vill., h=1700 m, 6.V.2015, D.Kovalchuk leg./det. R.Panin L=20 mm
    Copyright © Ruslan Panin
    Carabus (Tomocarabus) convexus rhinopterus Hampe, 1852 (as salax Motschulsky, 1865)
    Carabus (Tomocarabus) convexus rhinopterus Hampe, 1852 (as salax Motschulsky, 1865)
    Male adult
    Turkey, Sivas, 20 km N Zara, 1750 m 10 km S Serefiye 17.8.1992 K. Staven leg
    Copyright © Alexander Anichtchenko
    Carabus (Eucarabus) angustus dopyeong Rapuzzi, 2015
    Carabus (Eucarabus) angustus dopyeong Rapuzzi, 2015
    Male adult
    scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Eucarabus) angustus dopyeong Rapuzzi, 2015
    Carabus (Eucarabus) angustus dopyeong Rapuzzi, 2015
    Endophallus
    scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Eucarabus) sternbergi sternbergi Roeschke, 1898
    Carabus (Eucarabus) sternbergi sternbergi Roeschke, 1898
    Male adult
    Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Eucarabus) sternbergi sternbergi Roeschke, 1898
    Carabus (Eucarabus) sternbergi sternbergi Roeschke, 1898
    Endophallus
    Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Eucarabus) sternbergi gureopensis J.K. Park & J. Park, 2013
    Carabus (Eucarabus) sternbergi gureopensis J.K. Park & J. Park, 2013 Paratype
    Endophallus
    Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Eucarabus) cartereti cartereti Deuve, 1982
    Carabus (Eucarabus) cartereti cartereti Deuve, 1982
    Male adult
    Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Eucarabus) cartereti cartereti Deuve, 1982
    Carabus (Eucarabus) cartereti cartereti Deuve, 1982
    Endophallus
    Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Eucarabus) angustus dopyeong Rapuzzi, 2015
    Carabus (Eucarabus) angustus dopyeong Rapuzzi, 2015
    Male genitalia
    Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Eucarabus) sternbergi sternbergi Roeschke, 1898
    Carabus (Eucarabus) sternbergi sternbergi Roeschke, 1898
    Male genitalia
    Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Eucarabus) sternbergi sternbergi Roeschke, 1898
    Carabus (Eucarabus) sternbergi sternbergi Roeschke, 1898
    Male genitalia
    Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Eucarabus) cartereti cartereti Deuve, 1982
    Carabus (Eucarabus) cartereti cartereti Deuve, 1982
    Male genitalia
    Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Eucarabus) nitididorsus nitididorsus Ishikawa & Kim, 1983
    Carabus (Eucarabus) nitididorsus nitididorsus Ishikawa & Kim, 1983
    Male genitalia
    Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Eucarabus) nitididorsus nitididorsus Ishikawa & Kim, 1983
    Carabus (Eucarabus) nitididorsus nitididorsus Ishikawa & Kim, 1983
    Endophallus
    Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Eucarabus) nitididorsus nitididorsus Ishikawa & Kim, 1983
    Carabus (Eucarabus) nitididorsus nitididorsus Ishikawa & Kim, 1983
    Male adult
    Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Eucarabus) nitididorsus nitididorsus Ishikawa & Kim, 1983
    Carabus (Eucarabus) nitididorsus nitididorsus Ishikawa & Kim, 1983
    Male adult
    Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Eucarabus) nitididorsus nitididorsus Ishikawa & Kim, 1983
    Carabus (Eucarabus) nitididorsus nitididorsus Ishikawa & Kim, 1983
    Male adult
    Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Eucarabus) nitididorsus paegunsanus Imura, 1997
    Carabus (Eucarabus) nitididorsus paegunsanus Imura, 1997
    Male adult
    Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Eucarabus) nitididorsus paegunsanus Imura, 1997
    Carabus (Eucarabus) nitididorsus paegunsanus Imura, 1997
    Endophallus
    Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Eucarabus) nitididorsus paegunsanus Imura, 1997
    Carabus (Eucarabus) nitididorsus paegunsanus Imura, 1997
    Male genitalia
    Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Eucarabus) nitididorsus baegyangsanus (Choi & Park, 2019)
    Carabus (Eucarabus) nitididorsus baegyangsanus (Choi & Park, 2019) Holotype
    Male adult
    Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Eucarabus) nitididorsus baegyangsanus (Choi & Park, 2019)
    Carabus (Eucarabus) nitididorsus baegyangsanus (Choi & Park, 2019) Paratype
    Male adult
    Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Eucarabus) nitididorsus baegyangsanus (Choi & Park, 2019)
    Carabus (Eucarabus) nitididorsus baegyangsanus (Choi & Park, 2019) Paratype
    Male genitalia
    Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Eucarabus) nitididorsus baegyangsanus (Choi & Park, 2019)
    Carabus (Eucarabus) nitididorsus baegyangsanus (Choi & Park, 2019) Paratype
    Male genitalia
    Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Eucarabus) nitididorsus baegyangsanus (Choi & Park, 2019)
    Carabus (Eucarabus) nitididorsus baegyangsanus (Choi & Park, 2019) Paratype
    Male genitalia
    Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Eucarabus) nitididorsus baegyangsanus (Choi & Park, 2019)
    Carabus (Eucarabus) nitididorsus baegyangsanus (Choi & Park, 2019) Holotype
    Endophallus
    Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Eucarabus) namhaedoensis Kwon & Lee, 1984
    Carabus (Eucarabus) namhaedoensis Kwon & Lee, 1984
    Male adult
    Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Eucarabus) namhaedoensis Kwon & Lee, 1984
    Carabus (Eucarabus) namhaedoensis Kwon & Lee, 1984
    Male adult
    Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Eucarabus) namhaedoensis Kwon & Lee, 1984
    Carabus (Eucarabus) namhaedoensis Kwon & Lee, 1984
    Male genitalia
    Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Eucarabus) namhaedoensis Kwon & Lee, 1984
    Carabus (Eucarabus) namhaedoensis Kwon & Lee, 1984
    Endophallus
    Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Neoplesius) lama lama Semenov, 1898
    Carabus (Neoplesius) lama lama Semenov, 1898
    Male adult
    NW Sichuan, Chola mts, Maniganggo vill., 4600 m 15.6.2015, V. Patrikeev leg (cAA). det. Shi Hongliang, 2020
    Copyright © Alexander Anichtchenko
    Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
    Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
    Endophallus
    Russia, S Tuva, Khorumnug-Taiga Mts, Buren riv. Valley, Buren-Bay-Khaak vill. h=900 m
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
    Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
    Male adult
    Russia, S Tuva, Khorumnug-Taiga Mts, Buren riv. Valley, Buren-Bay-Khaak vill. h=900 m, 17-24.VI.2002, Vashchenko A.& B.leg.,/ det. Panin R. L=25 mm
    Copyright © Ruslan Panin
    Carabus (Aulonocarabus) truncaticollis fleischeri Reitter, 1898
    Carabus (Aulonocarabus) truncaticollis fleischeri Reitter, 1898
    Female adult
    Russia, N Burajatia, Barguzinskiy Mts., the upper reaches of M.Chivyrkuy riv., h=2000 m, 26.VI-12.VII.2007, Vashchenko S leg. L=18.5 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) henningi cf. saldaitisi Obydov 1997
    Carabus (Morphocarabus) henningi cf. saldaitisi Obydov 1997
    Female adult
    Russia, S Tuva, E Tanu-Ola Mts., Cha-Ova-Art Pass, h=1700m, 28.VI-3.VII.2000, Vashchenko S. leg. L=20 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) henningi cf saldaitisi Obydov 1997
    Carabus (Morphocarabus) henningi cf saldaitisi Obydov 1997
    Male adult
    Russia, S Tuva, E Tanu-Ola Mts., Cha-Ova-Art Pass, h=1700m, 28.VI-3.VII.2000, Vashchenko S. leg. L=19.5 mm
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) sibiricus haeres (trans s. errans)
    Carabus (Trachycarabus) sibiricus haeres (trans s. errans)
    Female adult
    C Ukraine, S of Poltava city, edge forest-lane, 2-26.IX.2010, O. Pozharov leg. /det. R.Panin. L=30 mm
    Copyright © Ruslan Panin
    Carabus (Aulonocarabus) truncaticollis fleischeri Reitter, 1898
    Carabus (Aulonocarabus) truncaticollis fleischeri Reitter, 1898
    Endophallus
    Russia, N Burajatia, Barguzinskiy Mts., the upper reaches of M.Chivyrkuy riv., h=2000 m
    Copyright © Ruslan Panin
    Carabus (Aulonocarabus) truncaticollis fleischeri Reitter, 1898
    Carabus (Aulonocarabus) truncaticollis fleischeri Reitter, 1898
    Male adult
    Russia, N Burajatia, Barguzinskiy Mts., the upper reaches of M.Chivyrkuy riv., h=2000 m, 26.VI-12.VII.2007, Vashchenko S leg. L=17 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) henningi pelevini Obydov 2006
    Carabus (Morphocarabus) henningi pelevini Obydov 2006
    Female adult
    E Kazakhstan, S Altay, Sarym-Sakty Mt Range, 12 km SE Ornek vill. h=1200m, 10.VII.1998, L=21 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) henningi pelevini Obydov 2006a: 99
    Carabus (Morphocarabus) henningi pelevini Obydov 2006a: 99
    Male adult
    E Kazakhstan, S Altay, Sarym-Sakty Mt Range, 12 km SE Ornek vill. h=1200m, 10.VII.1998, L=20 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) henningi henningi Fischer, 1817
    Carabus (Morphocarabus) henningi henningi Fischer, 1817
    Male adult
    Russia, S Siberia, NW Buryatia, E Sayans Mts., Tunka Range, near Mondy vill., h=2400 m, alpine tundra, 1-15.VI.2001, Mishustin R. leg./det. R.Panin
    Copyright © Ruslan Panin
    Carabus (Eucarabus) ulrichii (as podolicus Semenov, 1898 loc.typ. )
    Carabus (Eucarabus) ulrichii (as podolicus Semenov, 1898 loc.typ. )
    Male adult
    CW-UKRAINE, Vinnytsia Distr., Zarvantsi vill., 1-25.V.2020, Kokhan S. leg./det. R.Panin,
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) argenticornis Rapuzzi, 2020
    Carabus (Apotomopterus) argenticornis Rapuzzi, 2020 Holotype
    Male adult
    China, South Guangxi province, Qinzhou city, Mt. Shiwandashan, 1200 m, V.2020. Image from Rapuzzi, 2020
    Copyright © - - -
    Carabus (Tribax) kasakorum daghestanicola Rapuzzi, 2018
    Carabus (Tribax) kasakorum daghestanicola Rapuzzi, 2018 Holotype
    Female adult
    Russia, East Caucasus, Daghestan, Andiysky Mountain Range, Kharami pass, 2,500 m, 5.VII.2010, S. Rybalkin legit, Ivan Rapuzzi collection. Image from Rapuzzi, 2018
    Copyright © - - -
    Carabus (Carabus) vigilax rautenstrauchi Rapuzzi, 2018
    Carabus (Carabus) vigilax rautenstrauchi Rapuzzi, 2018 Holotype
    Male adult
    China, Sichuan, Leibo, Shanlinggang environs, VII.2018, Wang legit in collection Ivan Rapuzzi. Image from Rapuzzi, 2018
    Copyright © - - -
    Carabus (Apotomopterus) longeantennatus transjinshaensis Rapuzzi, 2018
    Carabus (Apotomopterus) longeantennatus transjinshaensis Rapuzzi, 2018 Holotype
    Male adult
    China, Sichuan, Leibo, Shanlinggang env., VII.2018, Wang legit, Ivan Rapuzzi collection. Image from Rapuzzi, 2018
    Copyright © - - -
    Carabus (Apotomopterus) benardi briseis Rapuzzi, 2018
    Carabus (Apotomopterus) benardi briseis Rapuzzi, 2018 Holotype
    Female adult
    China, Sichuan, Leibo, Shanlinggang environs, VII.2018, Wang legit, Ivan Rapuzzi collection. Image from Rapuzzi, 2018
    Copyright © - - -
    Carabus (Tribax) kasakorum scintillans Rapuzzi, 2018
    Carabus (Tribax) kasakorum scintillans Rapuzzi, 2018 Holotype
    Male adult
    Russia, Caucasus, Ingushetia, South slopes of Skalistyi Mountain, Range, Beyni village environs, 2,800 m, 21.V/19.VII.2018, Ivan Rapuzzi collection. Image from Rapuzzi, 2018
    Copyright © - - -
    Carabus (Apotomopterus) benardi ebian Cavazutti & Rapuzzi, 2010
    Carabus (Apotomopterus) benardi ebian Cavazutti & Rapuzzi, 2010
    Male adult
    specimen from type locality. Image from Rapuzzi, 2018
    Copyright © - - -
    Carabus (Apotomopterus) benardi alterego Rapuzzi, 2020
    Carabus (Apotomopterus) benardi alterego Rapuzzi, 2020 Holotype
    Female adult
    China, Southeast Sichuan province, leibo, 50 km South from Shanlinggangxiang, Vi.2019, Wang legit. Image from Rapuzzi, 2020
    Copyright © - - -
    Carabus (Apotomopterus) yunanensis planielongatus Rapuzzi, 2020
    Carabus (Apotomopterus) yunanensis planielongatus Rapuzzi, 2020 Holotype
    Female adult
    Southeast Sichuan province, leibo, 50 km South from Shanlinggangxiang, Vi.2019, Wang legit . Image from Rapuzi, 2020
    Copyright © - - -
    Carabus (Apotomopterus) cyanipennis atripennis Rapuzzi, 2020
    Carabus (Apotomopterus) cyanipennis atripennis Rapuzzi, 2020 Holotype
    Male adult
    China, Guangxi province, Yuecheng ling, 1800 m, 1/16.V.2008, V. Siniaev legit. Image from Rapuzzi, 2020
    Copyright © - - -
    Carabus (Apotomopterus) tonkinensis hagiangensis Rapuzzi, 2020
    Carabus (Apotomopterus) tonkinensis hagiangensis Rapuzzi, 2020 Holotype
    Male adult
    Vietnam, Ha Giang province, lung Cu, 1600m, iV.2019. Image from Rapuzzi, 2020
    Copyright © - - -
    Carabus (Pseudocoptolabrus) chortenensis Cavazzuti, 2005
    Carabus (Pseudocoptolabrus) chortenensis Cavazzuti, 2005
    Male adult
    specimen from type locality. Image from Rapuzzi, 2020
    Copyright © - - -
    Carabus (Pseudocoptolabrus) taliensis xerophilous Rapuzzi, 2020
    Carabus (Pseudocoptolabrus) taliensis xerophilous Rapuzzi, 2020 Holotype
    Female adult
    nE Yunnan, dahai xiang, Vii.2019, Wang legit. Image from Rapuzzi, 2020
    Copyright ©
    Carabus (Apotomopterus) patroclus vicarius Cavazzuti, 1996
    Carabus (Apotomopterus) patroclus vicarius Cavazzuti, 1996
    Male adult
    China, S Sichuan, / Yunnan border, W of Lugu lake, S of Yongning vill., h=2700m, 11-18.VI.2017, L=31 mm
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) arrowi ghirettii Cavazzuti, 1995
    Carabus (Apotomopterus) arrowi ghirettii Cavazzuti, 1995
    Female adult
    S-China, NE-Guizhou prov., Fanjing Shan, 1800-2000 m, 5-11.VI.14, 27.54°N 108.42°E , L=40 mm
    Copyright © Ruslan Panin
    Carabus (Isiocarabus) fiduciarius fiduciarius Thomson, 1856
    Carabus (Isiocarabus) fiduciarius fiduciarius Thomson, 1856
    Female adult
    CE-China, Hubei, Xiaogan City, Shuangfeng Mountain, 26.VI.2016, L=30 mm
    Copyright © Ruslan Panin
    Carabus (Carabus) andreiianus Deuve, 2007
    Carabus (Carabus) andreiianus Deuve, 2007
    Female adult
    C-China, N-Sichuan, Pingwu Co. h=1500 m, 27.VI.2011, L=24 mm
    Copyright © Ruslan Panin
    Carabus (Carabus) koidei koidei Imura, 1996
    Carabus (Carabus) koidei koidei Imura, 1996
    Male adult
    C-China, S-Sichuan, Pass btw. Yanyaun / Muli, h=3244 m, 27.68638N; 101.22335E,,11-18.VI.2017, Ch. Reuter leg. L=22 mm
    Copyright © Ruslan Panin
    Carabus (Neoplesius) zhangjieae Deuve & Li, 2020
    Carabus (Neoplesius) zhangjieae Deuve & Li, 2020 Holotype
    Male adult
    Chine, Sichuan occidental, Danba Xian, env. 7 km à l’est de Danba, rive gauche de la rivière Xiaojin, Jikaiyi, env. 2300 mètres, 30°53’-101°55’E, Zhang Jie leg., 23 septembre 2019; 18 mm; Image from Deuve & Li, 2020
    Copyright © - - -
    Carabus (Cupreocarabus) mollardianus Deuve, 2016
    Carabus (Cupreocarabus) mollardianus Deuve, 2016
    Female adult
    Chine, Sichuan, Lushan, Mt. Jiajin Shan, 10.VII.2016 local collector leg./ det. R.Panin, L=22 mm
    Copyright © Ruslan Panin
    Carabus (Cupreocarabus) ? siguniangensis jiachingensis Lassalle, 2003
    Carabus (Cupreocarabus) ? siguniangensis jiachingensis Lassalle, 2003
    Male adult
    Chine, Sichuan, Lushan, Mt. Jiajin Shan, 10.VII.2016 local collector leg., L=21 mm
    Copyright © Ruslan Panin
    Carabus (Sigenacarabus) trachynodes moxiensis Deuve, 1992
    Carabus (Sigenacarabus) trachynodes moxiensis Deuve, 1992
    Male adult
    China, W-Sichuan, W of Moxi, Daxue Range, Mt.Gongga Shan h=3000-3100m, 29°34′29.6″N /101°59′44.0″E, 10-12.VI.2018, Ch. Reuter leg. L=21 mm
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) ludivinae ludivinae Deuve, 1996
    Carabus (Apotomopterus) ludivinae ludivinae Deuve, 1996
    Male adult
    S-China, SW-Yunnan: Lincang city, Mt. Daxue, 18.IV.2017, L=29 mm
    Copyright © Ruslan Panin
    Carabus (Meganebrius) tuberculipennis ganjalaensis Deuve, 2000
    Carabus (Meganebrius) tuberculipennis ganjalaensis Deuve, 2000
    Male adult
    Nepal, Helambu, GanjaLa Pass S-slope, Dhukpu Kharka h=4100m, 28°03'50"N, 85°34'21"E, 12.VIII.2010, S.Tamang leg., L=21 mm
    Copyright © Ruslan Panin
    Carabus (s.str.) morphocaraboides zhaojueanus Imura, 1999
    Carabus (s.str.) morphocaraboides zhaojueanus Imura, 1999
    Female adult
    China, S-Sichuan, SE Mianning, Pass btw Mianshanzhen & Yuexi 28°30’N/102°26’E, 22-23.VI.2017, Ch.Reuter leg., L=22 mm; det. B. Brezina, 2020
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) benardi benardi Breuning, 1931
    Carabus (Apotomopterus) benardi benardi Breuning, 1931 Holotype
    Male adult
    http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec128
    Copyright © MNHN
    Carabus (Coptolabrus) smaragdinus shantungensis Born, 1910
    Carabus (Coptolabrus) smaragdinus shantungensis Born, 1910 Holotype
    Male adult
    Shandong, Qingdao reçu de Bang Haes le 23/IV/1912 . http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec122
    Copyright © MNHN
    Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
    Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
    Male adult
    W Ukraine, Transcarpathian Reg., 4 km NE Chop, forest Peresh, Panin R. leg. L=30 mm
    Copyright © Ruslan Panin
    Carabus (Rhigocarabus) pseudopusio Deuve, 1996
    Carabus (Rhigocarabus) pseudopusio Deuve, 1996
    Male adult
    NW Sichuan, Zonggrannkow pass, 60km W Heishue city, 4000m, 22.7.2017 V.Patrikeev leg. (cAA). det. Anichtchenko A. 2020
    Copyright © Alexander Anichtchenko
    Carabus (Rhigocarabus) gracilicollis pusio Semenov, 1898
    Carabus (Rhigocarabus) gracilicollis pusio Semenov, 1898
    Male adult
    N Sichuan, pass between Songpan and Juzaigow, W side, 25-30.6.2015, 4200 m V. Patrikeev leg. (cAA). det. Anichtchenko A. 2020
    Copyright © Alexander Anichtchenko
    Carabus (Rhigocarabus) ladygini degensis Deuve, 1989
    Carabus (Rhigocarabus) ladygini degensis Deuve, 1989
    Male adult
    NW Sichuan, Chola Mts, Maniganggo vill., 4600m, 15.6.2015 V. Patrikeev leg. (cAA). det. Anichtchenko A. 2020
    Copyright © Alexander Anichtchenko
    Carabus (Rhigocarabus) thais thais Heinz, 1997
    Carabus (Rhigocarabus) thais thais Heinz, 1997
    Male adult
    E Qinghai-Gansu border, Dalia pass 50km W Lixia town 3800m 20.7.2019 (cAA) det. Anichtchenko 2020
    Copyright © Alexander Anichtchenko
    Carabus (Apotomopterus) prattianus prattianus Bates, 1890
    Carabus (Apotomopterus) prattianus prattianus Bates, 1890 Holotype
    Female adult
    Ichang (MNHN) http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec80
    Copyright © MNHN
    Carabus (Apotomopterus) protenes protenes Bates, 1889
    Carabus (Apotomopterus) protenes protenes Bates, 1889 Lectotype
    Female adult
    Ichang (MNHN) http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec81
    Copyright © MNHN
    Carabus (Apotomopterus) mecynodes mecynodes Bates, 1890
    Carabus (Apotomopterus) mecynodes mecynodes Bates, 1890
    Female adult
    NEOTYPE. Sichuan (MNHN) http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec82
    Copyright © MNHN
    Carabus (Apotomopterus) ichangensis Bates, 1889
    Carabus (Apotomopterus) ichangensis Bates, 1889 Lectotype
    Male adult
    Ichang (MNHN) http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec83
    Copyright © MNHN
    Carabus (Carabus) vigilax vigilax Bates, 1890
    Carabus (Carabus) vigilax vigilax Bates, 1890 Lectotype
    Male adult
    Yang-tsé sup. Wa-Shan 6000ft, 1889-06-01/1889-06-30 (MNHN)
    Copyright © MNHN
    Carabus (Isiocarabus) kiukiangensis kiukiangensis Bates, 1888
    Carabus (Isiocarabus) kiukiangensis kiukiangensis Bates, 1888 Lectotype
    Male adult
    Jiujiang (MNHN) http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec88
    Copyright © MNHN
    Carabus (Megodontoides) promachus promachus
    Carabus (Megodontoides) promachus promachus Holotype
    Male adult
    Kangding (MNHN) http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec96
    Copyright © MNHN
    Carabus (Meganebrius) kadoudali (Morvan, 1981)
    Carabus (Meganebrius) kadoudali (Morvan, 1981) Paratype
    Female adult
    Vallée de Uttar Ganga, Dhorpatan, col de Jhala-Jélé , 3300m, Morvan P. 1977-04-01/1977-04-30 (MNHN) http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec184
    Copyright © MNHN
    Carabus (Meganebrius) davatchii (Morvan, 1981)
    Carabus (Meganebrius) davatchii (Morvan, 1981) Paratype
    Male adult
    Vallée de Uttar Ganga, Dhorpatan, col de Jhala-Jélé (MNHN) http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec186
    Copyright © MNHN
    Carabus (Pagocarabus) crassesculptus eleganticolor Deuve, 1997
    Carabus (Pagocarabus) crassesculptus eleganticolor Deuve, 1997
    Female adult
    S Gansu, Min Shan, 5-20 km S Cagan, 2300-3200m 7-20.7.2019 V. Patrikeev leg. (cAA). det. Anichtchenko A. 2020
    Copyright © Alexander Anichtchenko
    Carabus (Rhigocarabus) cateniger copulans Semenov, 1898
    Carabus (Rhigocarabus) cateniger copulans Semenov, 1898
    Male adult
    E Qinghai, Laji Shan, near Garang, 3900m, 19.6.2011, Patrikeev V. leg (cAA) det. Anichtchenko A. 2020
    Copyright © Alexander Anichtchenko
    Carabus (Rhigocarabus) cateniger papaoensis Cavazzuti, 2006
    Carabus (Rhigocarabus) cateniger papaoensis Cavazzuti, 2006
    Male adult
    E Qinghai, near Babao town, 3800m 5.8.12 Patrikeev V. leg. (cAA) det. Anichtchenko A. 2020
    Copyright © Alexander Anichtchenko
    Carabus (Rhigocarabus) cateniger cateniger Morawitz, 1886
    Carabus (Rhigocarabus) cateniger cateniger Morawitz, 1886
    Male adult
    NW Gansu, 80 km S Jayuguan, Tulai shan, Qionglai mts, 10.7.2014, 4200 m, Patrikeev leg (cAA)
    Copyright © Alexander Anichtchenko
    Carabus (Pseudocoptolabrus) taliensis ridilioides Deuve & Tian, 2020
    Carabus (Pseudocoptolabrus) taliensis ridilioides Deuve & Tian, 2020 Holotype
    Male adult
    Chine, Yunnan, col à env. 17 km au nord-ouest de Shangri-La (Zhongdian), 3684 mètres, 27°55’33’’N-99°36’42’’E, in coll. R. Sehnal (Unhošt, République Tchèque). Image from Deuve & Tian, 2020
    Copyright ©
    Carabus (Pseudocoptolabrus) pseudochortenensis Deuve & Tian, 2020
    Carabus (Pseudocoptolabrus) pseudochortenensis Deuve & Tian, 2020 Paratype
    Male adult
    Chine, Sichuan, entre Yanyuan et Muli, col de Mianbu, 3244 mètres, 27,68638°N-101,22335°E (Shi Hongliang et Liu Ye), in coll. Institut Zoologique, Academia Sinica, Pékin (Chine). Image from Deuve & Tian, 2020
    Copyright © - - -
    Carabus (Chrysocarabus) olympiae Sella, 1855
    Carabus (Chrysocarabus) olympiae Sella, 1855
    Male adult

    Copyright © Vittorino Monzini
    Carabus (Lipaster) stjernvalli paravanicus Fominykh et al 2020
    Carabus (Lipaster) stjernvalli paravanicus Fominykh et al 2020 Paratype
    Male+Female adult
    Georgia, Samtskhe-Dzhavakheti, W bank of Paravani Lake near Aspara vill., E slopes of Kyerogly Mt., 2100-2200 m, N 41°28’24.99” / E 43°46’19.65”, 04.-05.06.2015 (D. Fominykh, A. Zubov & A. Safronov). Image from Fominykh et al 2020
    Copyright © - - -
    Carabus (Lipaster) stjernvalli bogatshevi Zamotajlov, 1990
    Carabus (Lipaster) stjernvalli bogatshevi Zamotajlov, 1990 Holotype
    Male adult
    Transcaucasus, mt. Tshareli (=Chareli) near Borzhomi, 2100 m, 30 VII 1929, A. Bogačev leg.” (ZISP). Image from Fominykh et al 2020
    Copyright © - - -
    Carabus (Lipaster) stjernvalli tabackuriensis Novotný & Vořišek, 1988
    Carabus (Lipaster) stjernvalli tabackuriensis Novotný & Vořišek, 1988
    Male+Female adult
    Georgia, Kvemo-Kartli, Tsalka distr., 4.5 km SW of Kavta (=Chapaevka) vill., NE slopes of Buladag Mt., 1700 m, N 41°32’ / E44°01’, pitfall traps, 20.05.-04.06.2015 (A. Zubov & R. Khryapin). Image from Fominykh et al 2020
    Copyright © - - -
    Carabus (Lipaster) stjernvalli tabackuriensis Novotný & Vořišek, 1988
    Carabus (Lipaster) stjernvalli tabackuriensis Novotný & Vořišek, 1988
    Male+Female adult
    Georgia, Kvemo-Kartli, Trialeti Mt. Range near Tukhmatash Pass, E slopes of Dalidag Mt., 2070 m, N 41°32’ / E43°52’, pitfall traps, 06.06.–07.2017 (D. Fominykh). Image from Fominykh et al 2020
    Copyright © - - -
    Carabus (Lipaster) stjernvalli kartalinicus Fominykh et al 2020
    Carabus (Lipaster) stjernvalli kartalinicus Fominykh et al 2020 Paratype
    Male+Female adult
    Georgia, Kvemo-Kartli, Trialeti Mt. Range near Manglisi vill., NW part of Kartli Mt. Range, N slopes of Shikhan Mt., 2070 m, N 41°45’ / E 44°22’, pitfall traps, 07.06.-17.07.2017 (D. Fominykh). Image from Fominykh et al 2020
    Copyright © - - -
    Carabus (Lipaster) stjernvalli kartalinicus Fominykh et al 2020
    Carabus (Lipaster) stjernvalli kartalinicus Fominykh et al 2020 Paratype
    Male adult
    Georgia, Shida-Kartli, Trialeti Mt. Range near Dre vill., NW part of Razmeti Mt. Range, 1920 m, N 41°50’ / E 44°05’, 08.06.2017 (D. Fominykh). Image from Fominykh et al 2020
    Copyright © - - -
    Carabus (Lipaster) stjernvalli stjernvalli Mannerheim, 1830
    Carabus (Lipaster) stjernvalli stjernvalli Mannerheim, 1830
    Male+Female adult
    1-2: Georgia, Adjaria, Arsiani (=Arsiyan) Mt. Range near Goderdzi Pass, Chanchakhi Mt., 2300 m, N 41°36’35.00” / E42°31’34.15”, 10.06.2015 (D. Fominykh & A. Zubov); 3-4: Georgia, Adjaria (=Adjara, Adzharia), Baki (Bako) vill. env., 08.-26.06.1998 (V. Filippov). Image from Fominykh et al 2020
    Copyright © - - -
    Carabus (Lipaster) stjernvalli stjernvalli Mannerheim, 1830
    Carabus (Lipaster) stjernvalli stjernvalli Mannerheim, 1830
    Male+Female adult
    Georgia, Guria, N slopes of Adzhar-Imereti Mt. Range near Gomi vill. between Bzhuzhi and Kalova rivers, 1440 m, N 41°50’16,60” / E 42°08’29,20”, pitfall traps, 10.04.-07.06.2016 (D. Fominykh). Image from Fominykh et al 2020
    Copyright © - - -
    Carabus (Lipaster) stjernvalli stjernvalli Mannerheim, 1830
    Carabus (Lipaster) stjernvalli stjernvalli Mannerheim, 1830
    Male+Female adult
    Georgia, Samtskhe-Dzhavakheti, Adigeni Distr., Khormani Mt. Range, E slopes of Tail’ Mt., 2290 m, N 41°36’57.00” / E 42°40’32.40”, 22.-24.05.2015 (A. Zubov & R. Khryapin) Image from Fominykh et al 2020
    Copyright © - - -
    Carabus (Lipaster) stjernvalli gvalijai Retezár, 1992
    Carabus (Lipaster) stjernvalli gvalijai Retezár, 1992
    Male+Female adult
    Georgia, Racha, Racha Mt. Range, N slopes of Khikhata Mt. Range near Mravaldzali vill., 2100 m, N 42°29’13,64” / E 43°21’12,19”, 08.-09.06.2016 (A. Zubov). Image from Fominykh et al 2020
    Copyright © - - -
    Carabus (Lipaster) stjernvalli gvalijai Retezár, 1992
    Carabus (Lipaster) stjernvalli gvalijai Retezár, 1992
    Male+Female adult
    Georgia, Kvemo-Svaneti, S slopes of Svaneti Mt. Range near Lentekhi vill., S slopes of Goldash Mt., 2615 m, N 42°50’00,10” / E 42°45’16,90”, pitfall traps, 18.06.-13.07.2017 (D. Fominykh). Image from Fominykh et al 2020
    Copyright © - - -
    Carabus (Macrothorax) rugosus laufferi Breuning, 1927
    Carabus (Macrothorax) rugosus laufferi Breuning, 1927
    Female adult
    C-Spain: Avila, El Hornillo env., 27.V.2017, J. Ferrero leg. 30 mm
    Copyright © Ruslan Panin
    Carabus (Macrothorax) rugosus levantinus Lauffer, 1905
    Carabus (Macrothorax) rugosus levantinus Lauffer, 1905
    Male adult
    SE-Spain, Murcia, Jumillla, Sierra del Carche, 6.VII.2019, F. Garcia leg., L=29 mm
    Copyright © Ruslan Panin
    Carabus (s.str.) morphocaraboides zhaojueanus Imura, 1999
    Carabus (s.str.) morphocaraboides zhaojueanus Imura, 1999
    Male adult
    China, S-Sichuan, road 56 km E of Xichang to Zhaojue, Mt.WenLin, Pass h=3200 m, 27°52’N/102°30E, 5-20.VI.2017, Ch.Reuter leg.; det B. Brezina, 2020
    Copyright © Ruslan Panin
    Carabus (Ctenocarabus) melancholicus submeridionalis Breuning, 1975
    Carabus (Ctenocarabus) melancholicus submeridionalis Breuning, 1975
    Male adult
    S- Spain, Cádiz, Tarifa, El Cabrito, 28.XII.2019, F. Garcia leg. L=26 mm
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) lusitanicus helluo (as alicantinus Ganglbauer, 1886)
    Carabus (Mesocarabus) lusitanicus helluo (as alicantinus Ganglbauer, 1886)
    Endophallus
    SE-Spain, Murcia reg., Jumilla, Sierra del Carche, h=1300 m,
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) lusitanicus helluo (as alicantinus Ganglbauer, 1886)
    Carabus (Mesocarabus) lusitanicus helluo (as alicantinus Ganglbauer, 1886)
    Male adult
    SE-Spain, Murcia reg., Jumilla, Sierra del Carche, h =1300 m, 30.XI.2019, F. Garcia leg.
    Copyright © Ruslan Panin
    Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
    Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
    Male adult
    W Ukraine, Lviv city, Sykhiv, Zubra forest, (Osvytskiy Grove), under bark, 49°47'28.10" N, 24° 2'20.09" E, 10.XI.2017, Panin R leg.
    Copyright © Ruslan Panin
    Carabus (Imaibius) rostianus Semenov, 1906
    Carabus (Imaibius) rostianus Semenov, 1906
    Female adult
    Natural History Museum (2014). Dataset: Collection specimens. Resource: Specimens. Natural History Museum Data Portal (data.nhm.ac.uk). © The Trustees of the Natural History Museum, London
    Copyright © - - -
    Carabus (Imaibius) gridellii naranensis Heinertz, 1978
    Carabus (Imaibius) gridellii naranensis Heinertz, 1978
    Female adult
    Natural History Museum (2014). Dataset: Collection specimens. Resource: Specimens. Natural History Museum Data Portal (data.nhm.ac.uk). © The Trustees of the Natural History Museum, London
    Copyright © - - -
    Carabus (Imaibius) epipleuralis Semenov, 1906
    Carabus (Imaibius) epipleuralis Semenov, 1906
    Male adult
    Natural History Museum (2014). Dataset: Collection specimens. Resource: Specimens. Natural History Museum Data Portal (data.nhm.ac.uk). © The Trustees of the Natural History Museum, London
    Copyright © - - -
    Carabus (Imaibius) gridellii babussarensis Deuve & Kalab, 2000
    Carabus (Imaibius) gridellii babussarensis Deuve & Kalab, 2000
    Female adult
    Pakistan, Hazara, Babursar pass., 4100m, 10.Viii.1982, W.G. Tremewan. Natural History Museum (2014). Dataset: Collection specimens. Resource: Specimens. Natural History Museum Data Portal (data.nhm.ac.uk). © The Trustees of the Natural History Museum, London
    Copyright © - - -
    Carabus (Macrothorax) planatus Chaudoir, 1843
    Carabus (Macrothorax) planatus Chaudoir, 1843
    Female adult
    S Italy, N Sicily, Nebrodi Mts, Portella di Fémmina Morta h=1500 m, 2.IX.1989, V.Aliquo leg. L=33 mm
    Copyright © Ruslan Panin
    Carabus (Coptolabrus) smaragdinus branickii Taczanowski, 1887
    Carabus (Coptolabrus) smaragdinus branickii Taczanowski, 1887
    Male adult
    N Korea, S. Hamgyong Prov., Mt.Chail Peak, 1-16.VII.2010, l=38 mm
    Copyright © Ruslan Panin
    Carabus (Pachystus) hortensis preslii Dejean, 1830
    Carabus (Pachystus) hortensis preslii Dejean, 1830
    Male adult
    S-Greece:, N Peloponnes, Erymanthos Reg., Kalentzi, 26.VI.2007, Manfred Egger leg. L=26 mm
    Copyright © Ruslan Panin
    Carabus (Acoptolabrus) schrencki minpongsanensis Deuve & Li, 2003
    Carabus (Acoptolabrus) schrencki minpongsanensis Deuve & Li, 2003
    Male adult
    North Korea, Chagang Prov., Manpo, Songhak-ri (villl.,) 1-16.VII.2011, L=26 mm
    Copyright © Ruslan Panin
    Carabus (Eucarabus) manifestus guanmenshanus Imura, 1991
    Carabus (Eucarabus) manifestus guanmenshanus Imura, 1991
    Male adult
    China, Liaoning, Tieling County, Sandagou env., h=850 m, 13-18.VI.2011
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) bertolinii Kraatz, 1878
    Carabus (Orinocarabus) bertolinii Kraatz, 1878
    Endophallus
    NE-Italy, Trentino prov. Dolomites Alps, Plateau Pale di San Martino h=2500 m,
    Copyright © Ruslan Panin
    Carabus (Orinocarabus) bertolinii Kraatz, 1878
    Carabus (Orinocarabus) bertolinii Kraatz, 1878
    Male adult
    NE-Italy, Trentino prov. Dolomites Alps, Plateau Pale di San Martino h=2500 m, 17-19.VIII.2019, M. Egger leg.
    Copyright © Ruslan Panin
    Carabus (Ohomopterus) dehaanii dehaanii Chaudoir, 1848
    Carabus (Ohomopterus) dehaanii dehaanii Chaudoir, 1848
    Female adult
    Japan, S Honshu, Hyogo Pref., Nishiwaki city, 20.VI.2010, Shinichi S. leg., L=37 mm
    Copyright © Ruslan Panin
    Carabus (Archicarabus) alysidotus Illiger, 1798
    Carabus (Archicarabus) alysidotus Illiger, 1798
    Female adult
    Italy, Reggio Emilia, Albinea env. h=150 m, VI.2018, A. Cartini leg., L=25 mm
    Copyright © Ruslan Panin
    Carabus (Eucarabus) billbergi kuandianicus Rapuzzi, 2007
    Carabus (Eucarabus) billbergi kuandianicus Rapuzzi, 2007
    Female adult
    NE China, Liaoning, Kuandian County, Dachuantou env., 20-29.VI.2015,
    Copyright © Ruslan Panin
    Carabus (Eucarabus) xiuyanensis Deuve & Li, 1998
    Carabus (Eucarabus) xiuyanensis Deuve & Li, 1998
    Endophallus
    NE-China: SE-Liaoning, Xiuyan County, Dayingzi, Mt. Laopingding,
    Copyright © Ruslan Panin
    Carabus (Eucarabus) xiuyanensis Deuve & Li, 1998
    Carabus (Eucarabus) xiuyanensis Deuve & Li, 1998
    Male adult
    NE-China: SE-Liaoning, Xiuyan County, Dayingzi, Mt. Laopingding, 16.VI.2015, L=24.5 mm
    Copyright © Ruslan Panin
    Carabus (Coptolabrus) fruhstorferi Roeschke, 1900
    Carabus (Coptolabrus) fruhstorferi Roeschke, 1900
    Male adult
    Japan, Tsushima Isl., 20.IX.2018, S. Shinichi leg., L=35 mm
    Copyright © Ruslan Panin
    Carabus (Damaster) blaptoides oxuroides Schaum, 1862
    Carabus (Damaster) blaptoides oxuroides Schaum, 1862
    Female adult
    Japan, C Honshu, Nagano Prefecture, Nagano city, 3.IV.2012, S. Shinichi leg. L=42 mm. det. Anichtchenko A. 2021
    Copyright © Ruslan Panin
    Carabus (Leptocarabus) harmandi harmandi Lapouge, 1909
    Carabus (Leptocarabus) harmandi harmandi Lapouge, 1909 Lectotype
    Male adult
    Honshu, Tochigi, Mts near Nikko. Muséum national d’Histoire naturelle, Paris (France) Collection: Insects - Coleoptera (EC) Specimen MNHN-EC-EC549
    Copyright © MNHN
    Carabus (Eucarabus) xiuyanensis Deuve & Li, 1998
    Carabus (Eucarabus) xiuyanensis Deuve & Li, 1998
    Female adult
    NE-China: SE-Liaoning, Xiuyan County, Dayingzi, Mt. Laopingding, 16.VI.2015,
    Copyright © Ruslan Panin
    Carabus (Euleptocarabus) porrecticollis porrecticollis Bates, 1883
    Carabus (Euleptocarabus) porrecticollis porrecticollis Bates, 1883
    Male adult
    Japan, C Honshu, Nagano Pre., Toyooka town, 28.XII.2019, S. Shinichi leg. L=30 mm
    Copyright © Ruslan Panin
    Carabus (Euleptocarabus) porrecticollis porrecticollis Bates, 1883
    Carabus (Euleptocarabus) porrecticollis porrecticollis Bates, 1883
    Endophallus
    Japan, C Honshu, Nagano Pre., Toyooka town,
    Copyright © Ruslan Panin
    Carabus (Eucarabus) billbergi kuandianicus Rapuzzi, 2007
    Carabus (Eucarabus) billbergi kuandianicus Rapuzzi, 2007
    Endophallus
    NE China, Liaoning, Kuandian County, Dachuantou env.,
    Copyright © Ruslan Panin
    Carabus (Eucarabus) billbergi kuandianicus Rapuzzi, 2007
    Carabus (Eucarabus) billbergi kuandianicus Rapuzzi, 2007
    Male adult
    NE China, Liaoning, Kuandian County, Dachuantou env., 20-29.VI.2015, L=23 mm
    Copyright © Ruslan Panin
    Carabus (Damaster) blaptoides rugipennis Motschulsky, 1861
    Carabus (Damaster) blaptoides rugipennis Motschulsky, 1861
    Larva
    Kunashir isl. Stage 1 and stage 2.
    Copyright © Kirill Makarov
    Carabus (Damaster) blaptoides rugipennis Motschulsky, 1861
    Carabus (Damaster) blaptoides rugipennis Motschulsky, 1861
    Male adult
    Kunashir isl.
    Copyright © Kirill Makarov
    Carabus (Damaster) blaptoides babaianus Ishikawa, 1983
    Carabus (Damaster) blaptoides babaianus Ishikawa, 1983
    Male adult
    Hagyu., Iide Town, Nishiokitama Co., Yamagata pref., Japan, 17 3.2016.T. Nakamura leg. (DUBC). 35mm
    Copyright © Alexander Anichtchenko
    Carabus (Damaster) blaptoides oxuroides Schaum, 1862
    Carabus (Damaster) blaptoides oxuroides Schaum, 1862
    Female adult
    Japan, Kiyose, Tokyo, 20.1.1961 (DUBC). 40 mm
    Copyright © Alexander Anichtchenko
    Carabus (Cratocephalus) solskyi chengyii Rapuzzi & Yang, 2021
    Carabus (Cratocephalus) solskyi chengyii Rapuzzi & Yang, 2021 Holotype Paratype
    Male+Female adult
    China, Xinjiang province, Yining County, Kashi Township Reservoir, VII.2020 (deposited in collection Ivan Rapuzzi, Prepotto, Italy). Image from Rapuzzi & Yang, 2021
    Copyright © - - -
    Carabus (Tribax) macropus Chaudoir, 1877
    Carabus (Tribax) macropus Chaudoir, 1877
    Male adult
    Russia, Ingushetia, S slopes of Skalistyi Mt.R., Khahalgi Mt., alpin zone, h = 2500 m, 25.V-19.VII.2018 leg. D.Fominykh
    Copyright © Igor Solodovnikov
    Carabus (Tribax) macropus Chaudoir, 1877
    Carabus (Tribax) macropus Chaudoir, 1877
    Female adult
    Russia, Ingushetia, S slopes of Skalistyi Mt.R., Khahalgi Mt., alpin zone, h = 2500 m, 25.V-19.VII.2018 leg. D.Fominykh
    Copyright © Igor Solodovnikov
    Carabus (Tribax) macropus Chaudoir, 1877
    Carabus (Tribax) macropus Chaudoir, 1877
    Endophallus
    Russia, Ingushetia, S slopes of Skalistyi Mt.R., Khahalgi Mt., alpin zone, h = 2500 m
    Copyright © Igor Solodovnikov
    Carabus (Lipaster) stjernvalli gordius Reitter, 1898
    Carabus (Lipaster) stjernvalli gordius Reitter, 1898
    Male adult

    Copyright © Vittorino Monzini
    Carabus (Tribax) steveni schamyli Hampe, 1852
    Carabus (Tribax) steveni schamyli Hampe, 1852
    Female adult
    C Caucasus, North Ossetia, the road on the village of Coban (Koban), the left river bank Gizeldon, the deciduous wood, h = 895-905 m, N 42°55'32.91", E 44°31'53.53", 20.05-03.07.2017, leg. A.P. Tatulyan
    Copyright © Igor Solodovnikov
    Carabus (Tribax) steveni schamyli Hampe, 1852
    Carabus (Tribax) steveni schamyli Hampe, 1852
    Female adult
    C Caucasus, North Ossetia, the road on the village of Coban (Koban), the left river bank Gizeldon, the deciduous wood, h = 895-905 m, N 42°55'32.91", E 44°31'53.53", 20.05-03.07.2017, leg. A.P. Tatulyan
    Copyright © Igor Solodovnikov
    Carabus (Tribax) steveni schamyli Hampe, 1852
    Carabus (Tribax) steveni schamyli Hampe, 1852
    Male adult
    C Caucasus, North Ossetia, the road on the village of Coban (Koban), the left river bank Gizeldon, the deciduous wood, h = 895-905 m, N 42°55'32.91", E 44°31'53.53", 20.05-03.07.2017, leg. A.P. Tatulyan
    Copyright © Igor Solodovnikov
    Carabus (Tribax) steveni schamyli Hampe, 1852
    Carabus (Tribax) steveni schamyli Hampe, 1852
    Male adult
    C Caucasus, North Ossetia, the road on the village of Coban (Koban), the left river bank Gizeldon, the deciduous wood, h = 895-905 m, N 42°55'32.91", E 44°31'53.53", 20.05-03.07.2017, leg. A.P. Tatulyan
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) starckianus theseus Briansky, 1910
    Carabus (Neoplectes) starckianus theseus Briansky, 1910
    Male adult
    Russia, Krasnodar Territory, Tuapse district, env. vill. Psebe, left bank of the Psebe River, hornbeam + oak + walnut, h = 146 m, 03.05-01.07.2014, leg. I.A. Solodovnikov, S.V. Solodovnikova, S.M. Pavlyuchuk
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) starckianus theseus Briansky, 1910
    Carabus (Neoplectes) starckianus theseus Briansky, 1910
    Female adult
    Russia, Krasnodar Territory, Tuapse district, env. vill. Psebe, left bank of the Psebe River, hornbeam + oak + walnut, h = 146 m, 03.05-01.07.2014, leg. I.A. Solodovnikov, S.V. Solodovnikova, S.M. Pavlyuchuk
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
    Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
    Female adult
    Russia, Krasnodar Territory, West-Caucasus, Apsheronsky district, Guam ridge, env. Mezmay village, by the road, beech, oak, blackberry, h = 1200-1250 m, 03.IV-14.VI.2020, leg. S.M. Pavlyuchuk, A.P. Tatulyan
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
    Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
    Female adult
    Russia, Krasnodar Territory, West-Caucasus, Apsheronsky district, Guam ridge, env. Mezmay village, by the road, beech, oak, blackberry, h = 1200-1250 m, 03.IV-14.VI.2020, leg. S.M. Pavlyuchuk, A.P. Tatulyan
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
    Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
    Male adult
    Russia, Krasnodar Territory, West-Caucasus, Apsheronsky district, Guam ridge, env. Mezmay village, by the road, beech, oak, blackberry, h = 1200-1250 m, 03.IV-14.VI.2020, leg. S.M. Pavlyuchuk, A.P. Tatulyan
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
    Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
    Male adult
    Russia, Krasnodar Territory, West-Caucasus, Apsheronsky district, Guam ridge, env. Mezmay village, by the road, beech, oak, blackberry, h = 1200-1250 m, 03.IV-14.VI.2020, leg. S.M. Pavlyuchuk, A.P. Tatulyan
    Copyright © Igor Solodovnikov
    Carabus (Isiocarabus) fiduciarius fiduciarius Thomson, 1856
    Carabus (Isiocarabus) fiduciarius fiduciarius Thomson, 1856
    Male adult
    27.5 mm. China, Sichuan, Wanyuan, Mt. Hua-e-shan. Image from Rapuzzi, 2015
    Copyright © - - -
    Carabus (Isiocarabus) fiduciarius kirinicus Csiki, 1927
    Carabus (Isiocarabus) fiduciarius kirinicus Csiki, 1927
    Male+Female adult
    Male: 34 mm, R. of Korea, Kyonggi-Do, Nam-yang-Ju-shi. Female: 35 mm, R. of Korea, Kyonggi-Do, East from Seoul, Yangpyeong (CIR). Image from Rapuzzi, 2015
    Copyright © - - -
    Carabus (Isiocarabus) fiduciarius saishutoicus Csiki, 1927
    Carabus (Isiocarabus) fiduciarius saishutoicus Csiki, 1927
    Male+Female adult
    Male: 26.5 mm, R. of Korea, Cheju Island, Mt. Hallasan. Female: 31 mm, R. of Korea, Cheju Island, Mt. Hangla (CIR). Image from Rapuzzi, 2015
    Copyright © - - -
    Carabus (Carabus) szeli obong Rapuzzi, 2015
    Carabus (Carabus) szeli obong Rapuzzi, 2015 Holotype Paratype
    Male+Female adult
    Democratic Republic of Korea, North Hamgyong Province, Mt. Obong, (ca 42.40 NL; 129.80 EL), 1/15.VII.2012. Image from Rapuzzi, 2015
    Copyright © - - -
    Carabus (Carabus) angustus dopyeong Rapuzzi, 2015
    Carabus (Carabus) angustus dopyeong Rapuzzi, 2015 Holotype Paratype
    Male+Female adult
    R. of Korea, Geonggi-do/Gangwondo provincial border, Pass North from Dopyeong, 600 m, 3/13.VII.2012, I. Rapuzzi and L. Caldon leg. Image from Rapuzzi, 2015
    Copyright © - - -
    Carabus (Carabus) sternbergi gimhwa Rapuzzi, 2015
    Carabus (Carabus) sternbergi gimhwa Rapuzzi, 2015 Holotype Paratype
    Male+Female adult
    R. of Korea, Gyeonggi-do/Gangwon-do provincial border, Pass North from Dopyeong, 600 m, 3/13.VII.2012, I. Rapuzzi and L. Caldon leg. Image from Rapuzzi, 2015
    Copyright © - - -
    Carabus (Carabus) sternbergi goheungicus Rapuzzi, 2015
    Carabus (Carabus) sternbergi goheungicus Rapuzzi, 2015 Holotype
    Female adult
    Republic of Korea, Goheung peninsula, Paryeongsan, 200m, 10/17.VII.2012, I. Rapuzzi and L. Caldon legit. Image from Rapuzzi, 2015
    Copyright © - - -
    Carabus (Carabus) sternbergi jindoensis Rapuzzi, 2015
    Carabus (Carabus) sternbergi jindoensis Rapuzzi, 2015 Holotype Paratype
    Male+Female adult
    R. of Korea, Jindo Island, 8.VIII.2010. Image from Rapuzzi, 2015
    Copyright © - - -
    Carabus (Carabus) sternbergi deogyusan Rapuzzi, 2015
    Carabus (Carabus) sternbergi deogyusan Rapuzzi, 2015 Holotype Paratype
    Male+Female adult
    Republic of Korea, Cholla Bukdo, Deogyusan, 10 km North from Gucheondong, 550 m, 7/15.VII.2007, I. Rapuzzi and L. Caldon leg. Image from Rapuzzi, 2015
    Copyright © - - -
    Carabus (Eucarabus) cartereti peacedam Rapuzzi, 2015
    Carabus (Eucarabus) cartereti peacedam Rapuzzi, 2015 Holotype Paratype
    Male+Female adult
    R. of Korea, Gangwondo, “Peace Dam”, NE Hwacheon 400 m, 3/13.VII.2012, I. Rapuzzi and L. Caldon leg. Image from Rapuzzi, 2015
    Copyright © - - -
    Carabus (Coptolabrus) smaragdinus euviridis (Ishikawa & Kim, 1983)
    Carabus (Coptolabrus) smaragdinus euviridis (Ishikawa & Kim, 1983)
    Male adult
    38 mm, R. of Korea, Jirisan Mt., I. Rapuzzi and L. Caldon leg. (CIR). Image from Rapuzzi, 2015
    Copyright © - - -
    Carabus (Coptolabrus) smaragdinus euaureus Rapuzzi, 2015
    Carabus (Coptolabrus) smaragdinus euaureus Rapuzzi, 2015 Holotype Paratype
    Male+Female adult
    R. of Korea, Jirisan Nat. Park, Yeongrieongchi, 1200 m, 7/16.VII.2012, I. Rapuzzi and L. Caldon leg. (CIR). Image from Rapuzzi, 2015
    Copyright © - - -
    Carabus (Coptolabrus) smaragdinus buangun Rapuzzi, 2015
    Carabus (Coptolabrus) smaragdinus buangun Rapuzzi, 2015 Holotype Paratype
    Male+Female adult
    R. of Korea, Jeollabuk-do Province, Buan-gun, Byeonsanbando, 100/200m, 13/20.VII.2012, I. Rapuzzi and L. Caldon leg. (CIR). Image from Rapuzzi, 2015
    Copyright © - - -
    Carabus (Coptolabrus) jankowskii byeoksanensis Rapuzzi, 2015
    Carabus (Coptolabrus) jankowskii byeoksanensis Rapuzzi, 2015 Holotype Paratype
    Male+Female adult
    R. of Korea, Jeollabuk-do Province, Buan-gun, Byeonsanbando, 100/200 m, 13/20.VII.2012, I. Rapuzzi and L. Caldon leg. (CIR). Image from Rapuzzi, 2015
    Copyright © - - -
    Carabus (Acoptolabrus) constricticollis constricticollis (as grallatorius)
    Carabus (Acoptolabrus) constricticollis constricticollis (as grallatorius)
    Male adult
    33 mm, Seishin Korea (= Chongjin, North Hamgyong province, D.R. of Korea) (CIR). Image from Rapuzzi, 2015
    Copyright © - - -
    Carabus (Acoptolabrus) constricticollis microcolasellus Rapuzzi, 2015
    Carabus (Acoptolabrus) constricticollis microcolasellus Rapuzzi, 2015 Holotype Paratype
    Male+Female adult
    D.R. of Korea, North Hamgyong Province, Paeksa, 1139 m, VII.2004, local collector leg. (CIR). Image from Rapuzzi, 2015
    Copyright © - - -
    Carabus (Acoptolabrus) costricticollis aff. limes Rapuzzi, 2009
    Carabus (Acoptolabrus) costricticollis aff. limes Rapuzzi, 2009
    Male adult
    D.R. of Korea, South Hwanghae province, Unryul County, Mt. Kuwol, 954 m, local collector leg. (CIR). Image from Rapuzzi, 2015
    Copyright © - - -
    Carabus (Acoptolabrus) leechi drouini Rapuzzi, 2015
    Carabus (Acoptolabrus) leechi drouini Rapuzzi, 2015 Holotype Paratype
    Male+Female adult
    D.R. of Korea, North Hamgyong Prov., Kichu, Mt. Muhak, 8/19.VII.2006, local collector leg. (CIR). Image from Rapuzzi, 2015
    Copyright © - - -
    Carabus (Acoptolabrus) leechi onjinsanicola Rapuzzi, 2009
    Carabus (Acoptolabrus) leechi onjinsanicola Rapuzzi, 2009
    Male adult
    D. R. of Korea, South Hwanghae province, Unryul County, Mt. Kuwol, 954 m, local collector leg. (CIR). Image from Rapuzzi, 2015
    Copyright © - - -
    Carabus (Acoptolabrus) leechi viniciosalamii Rapuzzi, 2015
    Carabus (Acoptolabrus) leechi viniciosalamii Rapuzzi, 2015 Holotype Paratype
    Male+Female adult
    D.R. of Korea, North Hamgyong Prov., Mt. Wanta, 24.VII.2004, local collector leg. (CIR). Image from Rapuzzi, 2015
    Copyright © - - -
    Carabus (Acoptolabrus) planicranion Rapuzzi, 2015
    Carabus (Acoptolabrus) planicranion Rapuzzi, 2015 Holotype Paratype
    Male+Female adult
    D.R. of Korea, North Hamgyong Prov., Mt. Obong, (ca 42,40 NL; 129,80 EL), 1/15.VII.2012, local collector leg. (CIR). Image from Rapuzzi, 2015
    Copyright © - - -
    Carabus (Acoptolabrus) changeonleei Ishikawa & Kim, 1983
    Carabus (Acoptolabrus) changeonleei Ishikawa & Kim, 1983
    Female adult
    R. of Korea, Jirisan Mt., I. Rapuzzi and L. Caldon leg. (CIR). Image from Rapuzzi, 2015
    Copyright © - - -
    Carabus (Acoptolabrus) mirabilissimus igniferescens Deuve, 1992
    Carabus (Acoptolabrus) mirabilissimus igniferescens Deuve, 1992
    Male adult
    R. of Korea, Gangwondo province, South from Gimhwa, 450 m (CIR). Image from Rapuzzi, 2015
    Copyright © - - -
    Carabus (Acoptolabrus) mirabilissimus mirabilissimus Ishikawa & Deuve, 1982
    Carabus (Acoptolabrus) mirabilissimus mirabilissimus Ishikawa & Deuve, 1982
    Male adult
    R. of Korea, Gangwondo province, “Peace Dam”, NE Hwacheon 400 m (CIR). Image from Rapuzzi, 2015
    Copyright © - - -
    Carabus (Acoptolabrus) mirabilissimus furumiensis Deuve, 2002
    Carabus (Acoptolabrus) mirabilissimus furumiensis Deuve, 2002
    Male adult
    R. of Korea, Gangwondo province, Odaesan Mt., Jingogae, 800 m (CIR). Image from Rapuzzi, 2015
    Copyright © - - -
    Carabus (Leptocarabus) vogtianus cf horvatovichi Deuve, 1992
    Carabus (Leptocarabus) vogtianus cf horvatovichi Deuve, 1992
    Male adult
    R. of Korea, Geonggido/Gangwondo provincial border, Pass North from Dopyeong, 600 m, 3/13.VII.2012, I. Rapuzzi and L. Caldon leg. Image from Rapuzzi, 2015
    Copyright © - - -
    Carabus (Aulonocarabus) canaliculatus vojnitsi Mandl, 1980
    Carabus (Aulonocarabus) canaliculatus vojnitsi Mandl, 1980 Holotype
    Male adult
    Image from Xueping et al, 2013
    Copyright © - - -
    Carabus (Aulonocarabus) canaliculatus rufinus Beheim & Breuning, 1943
    Carabus (Aulonocarabus) canaliculatus rufinus Beheim & Breuning, 1943
    Male adult
    Mt. Changbai Shan, Antu county, Jilin province, China. 13/15.VI.2012. Er-dao-bai-he, elevation 800 m. Image from Xueping et al, 2013
    Copyright © - - -
    Carabus (Aulonocarabus) canaliculatus vojnitsi Mandl, 1980
    Carabus (Aulonocarabus) canaliculatus vojnitsi Mandl, 1980
    Male adult
    Mt. Changbai Shan, Antu county, Jilin province, China. 13/15.VI.2012, Bai-shan-shang-fang, elevation 1850m. Ermans Birch forest. Image from Xueping et al, 2013
    Copyright © - - -
    Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988
    Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988 Paratype
    Male adult
    Krasnodar area, Babuk-Aul, 19.06.1983 , leg. A. Miroshnikov
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
    Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
    Female adult
    W Caucasus, NW slope Aibga Mt.R. vall. riv. Galion, h = 650 m, Fagus, Alnus viridis forest, 18-24.VI.1999, I.A. Solodovnikov leg.
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
    Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
    Male adult
    W Caucasus, Aibga Mt.R., 02.VIII.1994, A.S. Zamotailov leg.
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
    Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
    Male adult
    Russia, W Caucasus, Krasnaya Polyana, Atchischkho Mt.R., h = 1200 m, 10.06.1995, N.Yu. Pichugin leg.
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) starckianus adelaidae Starck, 1889
    Carabus (Neoplectes) starckianus adelaidae Starck, 1889
    Female adult
    Russia, Krasnodar area, Adler, Ader distr., right bank vall. Riv. Msymta, near vill. Chvesipse, h = 800 m, 06.VII.1999, B. Saratovskikh leg.
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) starckianus adelaidae Starck, 1889
    Carabus (Neoplectes) starckianus adelaidae Starck, 1889
    Female adult
    Russia, Krasnodar area, Adler, Ader distr., right bank vall. Riv. Msymta, near vill. Chvesipse (Fagus+Quercus+Castanea forest) [Sochi national park], h = 720 m, 03.V-08.VII.2016, I.A. Solodovnikov, S.V.Solodovnikova, V.M.Kotsur leg.
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) starckianus adelaidae Starck, 1889
    Carabus (Neoplectes) starckianus adelaidae Starck, 1889
    Female adult
    Russia, Krasnodar area, Adler, Ader distr., right bank vall. Riv. Msymta, near vill. Chvesipse (Fagus+Quercus+Castanea forest) [Sochi national park], h = 720 m, 03.V-08.VII.2016, I.A. Solodovnikov, S.V.Solodovnikova, V.M.Kotsur leg.
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) starckianus adelaidae Starck, 1889
    Carabus (Neoplectes) starckianus adelaidae Starck, 1889
    Male adult
    Russia, Krasnodar area, Adler, Ader distr., right bank vall. Riv. Msymta, near vill. Chvesipse (Fagus+Quercus+Castanea forest) [Sochi national park], h = 720 m, 03.V-08.VII.2016, I.A. Solodovnikov, S.V.Solodovnikova, V.M.Kotsur leg.
    Copyright © Igor Solodovnikov
    Carabus (Cechenochilus) boeberi boeberi Adams, 1817
    Carabus (Cechenochilus) boeberi boeberi Adams, 1817
    Female adult
    Грузия, Мцхета-Мтианети, к северу от пос. Гудаури, ЮВ отроги г. Байдарская (нижний пик), по ущелью ручья, альпика, под камнями, точка 2, h = 2700 м, 42.495413° N, 44.487844° E, 10.08.2019, leg. И.А., А.И., С.В. Солодовниковы, О.И. Данилова 26 mm
    Copyright © Igor Solodovnikov
    Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931 Neotype
    Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931 Neotype
    Male adult
    Chongjin Muséum national d’Histoire naturelle, Paris (France). Collection: Insects - Coleoptera (EC). Specimen MNHN-EC-EC518
    Copyright © MNHN
    Carabus (Procrustes) piochardi piochardi Gehin, 1883
    Carabus (Procrustes) piochardi piochardi Gehin, 1883 Lectotype
    Male adult
    Liban, Jebel Sannin. http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec511
    Copyright © MNHN
    Carabus (Morphocarabus) hummeli putyatini Rapuzzi, 2012
    Carabus (Morphocarabus) hummeli putyatini Rapuzzi, 2012 Holotype
    Male adult
    Russia, South Pimorskiy region, Putyatin Island, 50-200 m, 14/22.VIII.2008, A. Plutenko leg. (in coll Rapuzzi). Image from Rapuzzi, 2012
    Copyright © - - -
    Carabus (Aulonocarabus) gossarei mareschii Rapuzzi, 2010
    Carabus (Aulonocarabus) gossarei mareschii Rapuzzi, 2010 Holotype
    Male adult
    Askol’d Island (CR). Image from Rapuzzi, 2012
    Copyright © - - -
    Carabus (Megodontus) vietinghoffi rugicolor Rapuzzi, 2010
    Carabus (Megodontus) vietinghoffi rugicolor Rapuzzi, 2010 Holotype
    Male adult
    Reyneke island (CR). Image from Rapuzzi, 2012
    Copyright © - - -
    Carabus (Coptolabrus) smaragdinus robinzoni Rapuzzi, 2010
    Carabus (Coptolabrus) smaragdinus robinzoni Rapuzzi, 2010 Holotype
    Male adult
    Reyneke island (CR). Image from Rapuzzi, 2012
    Copyright © - - -
    Carabus (Pseudocoptolabrus) zuzkae Imura & Březina, 2012
    Carabus (Pseudocoptolabrus) zuzkae Imura & Březina, 2012 Holotype Paratype
    Male+Female adult
    Holotype: m, Northeast of Fugong, 3,440 m in altitude, 26.57.32N/ 98.56.44E, on the Nu Shan Mountain Range, in Fugong Xian, of Nujianglilizu- zizhizhou, northwestern Yunnan, Southwest China, 1.VI.2006, preserved in the Zoological Institute of Academy of Sciences, St. Petersburg, Russia. Paratype: female, same locality, 31.V.2006, preserved in the collection of the first author, Y. IBJG6 (Yokohama).Image from Imura & Březina, 2012
    Copyright © - - -
    Carabus (Pseudocoptolabrus) watanabei watanabei (Imura, 2003)
    Carabus (Pseudocoptolabrus) watanabei watanabei (Imura, 2003) Holotype
    Male adult
    Imau Bum of northern Myanmar. Image from Imura & Březina, 2012
    Copyright © - - -
    Carabus (Apotomopterus) patroclus shiziensis Deuve, 2017
    Carabus (Apotomopterus) patroclus shiziensis Deuve, 2017 Holotype
    Male adult
    Chine, Yunnan, Wuding Xian, Shizi Shan, 2270 m, 25°32’N-102°23’E (Jatua), in coll. Jaroslav Turna, à Kostelec na Hane (République Tchèque)
    Copyright © Thierry Deuve
    Carabus (Apotomopterus) patroclus guzhouensis Deueve, 2017
    Carabus (Apotomopterus) patroclus guzhouensis Deueve, 2017 Holotype
    Male adult
    Chine, Yunnan, Xinhua, parc forestier de Guzhou, 2010 m, 24°07’N-101°51’E (Jatua), in coll. Jaroslav Turna, à Kostelec na Hane (République Tchèque).
    Copyright © Thierry Deuve
    Carabus (Hypsocarabus) qinlingensis micangicola Deuve, 2017
    Carabus (Hypsocarabus) qinlingensis micangicola Deuve, 2017 Holotype
    Female adult
    Chine, Sichuan, Mts Micang Shan, env. de Guangwushan, 1700 m (in coll. Andreï Gorodinski, Moscow).
    Copyright © Thierry Deuve
    Carabus (Cupreocarabus) alpherakii exilior Deuve, 2017
    Carabus (Cupreocarabus) alpherakii exilior Deuve, 2017 Holotype
    Male adult
    Chine, Sichuan, 40 km S Hekou (Yajiang), mts à l’ouest de Tanggang, 3800 m, 29°41’31’’N-101°01’08’’E, in coll. Ross Sehnal, à Unhošt (République Tchèque).
    Copyright © Thierry Deuve
    Carabus (Cupreocarabus) alpherakii younglimae Deuve, 2017
    Carabus (Cupreocarabus) alpherakii younglimae Deuve, 2017 Holotype
    Male adult
    Chine, Sichuan, vall. 8 km NE Shade, 4575 m, 29°42’33’’N-101°19’10’’E, in coll. M. Janata, à Prague (République Tchèque).
    Copyright © Thierry Deuve
    Carabus (Archicarabus) phoenix Lapouge, 1924
    Carabus (Archicarabus) phoenix Lapouge, 1924
    Male adult
    Lebanon
    Copyright © Mingyi Tian
    Carabus (Procrustes) piochardi gezei Deuve, 2002
    Carabus (Procrustes) piochardi gezei Deuve, 2002
    Female adult
    Lebanon
    Copyright © Mingyi Tian
    Carabus (Procrustes) piochardi labruleriei Gehin, 1883
    Carabus (Procrustes) piochardi labruleriei Gehin, 1883
    Male adult
    Lebanon. det. Mingyi Tian
    Copyright © Mingyi Tian
    Carabus (Lamprostus) hemprichi hemprichi Dejean, 1826
    Carabus (Lamprostus) hemprichi hemprichi Dejean, 1826
    Male adult
    Lebanon. det. Mingyi Tian
    Copyright © Mingyi Tian
    Carabus (Lamprostus) saulcyi saulcyi Piochard, 1875 (as chambouqensis)
    Carabus (Lamprostus) saulcyi saulcyi Piochard, 1875 (as chambouqensis)
    Male adult
    Lebanon
    Copyright © Mingyi Tian
    Carabus (Lamprostus) syrus moukarzelae Deuve, 2002
    Carabus (Lamprostus) syrus moukarzelae Deuve, 2002
    Female adult
    Lebanon.
    Copyright © Mingyi Tian
    Carabus (Procerus) syriacus Kollar, 1843
    Carabus (Procerus) syriacus Kollar, 1843
    Female adult
    Lebanon
    Copyright © Mingyi Tian
    Carabus (Tomocarabus) rumelicus syriensis Breuning, 1943 (as libanoticus)
    Carabus (Tomocarabus) rumelicus syriensis Breuning, 1943 (as libanoticus)
    Male adult
    Lebanon
    Copyright © Mingyi Tian
    Carabus (Isiocarabus) kiukiangensis Bates, 1888
    Carabus (Isiocarabus) kiukiangensis Bates, 1888
    Male adult
    Gutianshan National Park (Gutianshan NP), formerly Gutianshan National Nature Reserve, is located in the western part of Zhejiang Province in southeast China (29°14'N, 118°07'E). Image from Zumstein et al, 2021
    Copyright © - - -
    Carabus (Morphocarabus) brinevi altaicola Deuve & Tian, 2019
    Carabus (Morphocarabus) brinevi altaicola Deuve & Tian, 2019
    Female adult
    «qing tong se», «28#» (ICXU)
    Copyright © Ilya Kabak
    Carabus (Morphocarabus) brinevi kanasensis Kabak, 2021
    Carabus (Morphocarabus) brinevi kanasensis Kabak, 2021 Holotype Paratype
    Male adult
    China, Xinjiang, Altai Mts, left bank of Kanas lake, 48°44ʹ35ʺN / 87°06ʹ03ʺE, H = 2710 m, alpine meadows, 15.07.2019 I.I. Kabak leg. (ZIN)
    Copyright © Ilya Kabak
    Carabus (Carabus) vinctus Weber, 1801
    Carabus (Carabus) vinctus Weber, 1801
    Male adult
    LOUISIANA, East Baton Rouge Parish, 1.5mi W Denham Springs, 30.482 ̊ N 90.981 ̊ W, bottomland hardwood forest, hand collected, 4 Sep 2017
    Copyright © Igor Sokolov
    Carabus (Carabus) vinctus Weber, 1801
    Carabus (Carabus) vinctus Weber, 1801
    Endophallus
    LOUISIANA, East Baton Rouge Parish, 1.5mi W Denham Springs, 30.482 ̊ N 90.981 ̊ W, bottomland hardwood forest, 4 Sep 2017
    Copyright © Igor Sokolov
    Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993
    Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993
    Male adult
    Uzbekistan, Chatkal Mt. rng., Irtash, 41°10'25.7"N 70°16'20.4"E, 5-7.8.2021, 2400m, A. Anichtchenko leg. (cAA). 27mm
    Copyright © Alexander Anichtchenko
    Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993
    Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993
    Female adult
    Uzbekistan, Chatkal Mt. rng., Irtash, 41°10'25.7"N 70°16'20.4"E, 5-7.8.2021, 2400m, A. Anichtchenko leg. (cAA). 30 mm
    Copyright © Alexander Anichtchenko
    Carabus (Cechenochilus) boeberi tschitscherini Semenov, 1896
    Carabus (Cechenochilus) boeberi tschitscherini Semenov, 1896
    Male adult
    Russia, C Caucasus, Ingushetia, S slopes of Skalistyi Mt.R.
    Copyright © Igor Solodovnikov
    Carabus (Cechenochilus) boeberi tschitscherini Semenov, 1896
    Carabus (Cechenochilus) boeberi tschitscherini Semenov, 1896
    Female adult
    Russia, C Caucasus, Ingushetia, S slopes of Skalistyi Mt.R.
    Copyright © Igor Solodovnikov
    Carabus (Cechenochilus) heydenianus Starck, 1889
    Carabus (Cechenochilus) heydenianus Starck, 1889
    Male adult
    Carabus (Procechenochilus) heydenianus euxinus Semenov, 1896 Abkhasia: Arabika plato (TOPOTYPUS)
    Copyright © Igor Solodovnikov
    Carabus (Cechenochilus) heydenianus Starck, 1889
    Carabus (Cechenochilus) heydenianus Starck, 1889
    Female adult
    Carabus (Procechenochilus) heydenianus euxinus Semenov, 1896 Abkhasia: Arabika plato (TOPOTYPUS)
    Copyright © Igor Solodovnikov
    Carabus (Cechenochilus) heydenianus Starck, 1889
    Carabus (Cechenochilus) heydenianus Starck, 1889
    Endophallus
    Carabus (Procechenochilus) heydenianus euxinus Semenov, 1896 Abkhasia: Arabika plato (TOPOTYPUS)
    Copyright © Igor Solodovnikov
    Carabus (Cechenochilus) boeberi aequaliceps Reitter, 1896
    Carabus (Cechenochilus) boeberi aequaliceps Reitter, 1896
    Male adult
    Russia, Dagestan, Andiyskiy Mt.R., near Andiyskiye vorota pass, 06.VI–18.VII.2021, leg. D. Fominykh
    Copyright © Igor Solodovnikov
    Carabus (Cechenochilus) boeberi aequaliceps Reitter, 1896
    Carabus (Cechenochilus) boeberi aequaliceps Reitter, 1896
    Female adult
    Russia, Dagestan, Andiyskiy Mt.R., near Andiyskiye vorota pass, 06.VI–18.VII.2021, leg. D. Fominykh
    Copyright © Igor Solodovnikov
    Carabus (Pachystus) hortensis hortensis Linne, 1758
    Carabus (Pachystus) hortensis hortensis Linne, 1758
    Female adult
    Natural hybrid Carabus hortensis X Carabus granulatus. Belarus, Minsk reg., 8,7 km of Borisov, near vill. Lyubatovstchina, pine forest (Pinetum pleuroziosum), h = 213 m, 54.31934° N, 28.511486° E, 20-27.09.2018, leg. Klychenok Snezhana
    Copyright © Igor Solodovnikov
    Carabus (Megodontus) obovatus Fischer, 1827
    Carabus (Megodontus) obovatus Fischer, 1827
    Male adult
    C.obovatus taizhensis Obydov, 2006 Khakassia: Kuznetskyi Alatau, env. vill. Priiskovyi, Bobrovaya Mt., h = 1200-1300 m
    Copyright © Igor Solodovnikov
    Carabus (Megodontus) obovatus Fischer, 1827
    Carabus (Megodontus) obovatus Fischer, 1827
    Endophallus
    C.obovatus taizhensis Obydov, 2006 Khakassia: Kuznetskyi Alatau, env. vill. Priiskovyi, Bobrovaya Mt., h = 1200-1300 m
    Copyright © Igor Solodovnikov
    Carabus (Acoptolabrus) leechi chiricola (Kwon & Lee, 1984)
    Carabus (Acoptolabrus) leechi chiricola (Kwon & Lee, 1984) Holotype
    Male adult
    deposited in Kyungpook National University, Daegu, Republic of Korea, Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Acoptolabrus) leechi chiricola (Kwon & Lee, 1984)
    Carabus (Acoptolabrus) leechi chiricola (Kwon & Lee, 1984) Paratype
    Female adult
    deposited in Kyungpook National University, Daegu, Republic of Korea, Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Acoptolabrus) leechi chiricola (Kwon & Lee, 1984)
    Carabus (Acoptolabrus) leechi chiricola (Kwon & Lee, 1984) Paratype
    Female adult
    deposited in Kyungpook National University, Daegu, Republic of Korea, Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Coptolabrus) smaragdinus hoenggandoensis (Kwon & Lee, 1984)
    Carabus (Coptolabrus) smaragdinus hoenggandoensis (Kwon & Lee, 1984) Holotype
    Male adult
    deposited in Kyungpook National University, Daegu, Republic of Korea, Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Coptolabrus) smaragdinus hoenggandoensis (Kwon & Lee, 1984)
    Carabus (Coptolabrus) smaragdinus hoenggandoensis (Kwon & Lee, 1984) Paratype
    Female adult
    deposited in Kyungpook National University, Daegu, Republic of Korea, Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Coptolabrus) smaragdinus hongdoensis (Kwon & Lee, 1984)
    Carabus (Coptolabrus) smaragdinus hongdoensis (Kwon & Lee, 1984) Holotype
    Male adult
    deposited in Kyungpook National University, Daegu, Republic of Korea, Scale bar = 5mm
    Copyright © Jong-Bong Choi
    Carabus (Acoptolabrus) gehinii munakatai (Ishikawa, 1968)
    Carabus (Acoptolabrus) gehinii munakatai (Ishikawa, 1968)
    Male adult
    Japan, SW-Hokkaido, Yakumo town, 4.VI.2017, S.Shinichi leg., L=30 mm
    Copyright © Ruslan Panin
    Carabus (Damaster) blaptoides blaptoides (Kollar, 1836)
    Carabus (Damaster) blaptoides blaptoides (Kollar, 1836)
    Female adult
    Japan, SW Honshu, Shimane Pref., Unnan city, 16.I2016, S.Shinichi leg., L=48 mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) kolbei taikinus Ishikawa, 1993
    Carabus (Megodontus) kolbei taikinus Ishikawa, 1993
    Female adult
    Japan, SC-Hokkaido:Tokachi, Hiroo Distr., Taiki-cho env., 3-7.IX.2019, S.Shinichi leg.
    Copyright © Ruslan Panin
    Carabus (Megodontus) kolbei kosugei Nakane, 1955
    Carabus (Megodontus) kolbei kosugei Nakane, 1955
    Male adult
    Japan, SW-Hokkaido, Otaru city, 2-16.VIII.2020, S.Shinichi leg.
    Copyright © Ruslan Panin
    Carabus (Megodontus) kolbei hidakamontanus Ishikawa, 1966
    Carabus (Megodontus) kolbei hidakamontanus Ishikawa, 1966
    Male adult
    SE-Hokkaido: Samani distr.
    Copyright © Igor Solodovnikov
    Carabus (Megodontus) kolbei hidakamontanus Ishikawa, 1966
    Carabus (Megodontus) kolbei hidakamontanus Ishikawa, 1966
    Male adult
    E-Hokkaido: Samani distr., Horomankyo env.
    Copyright © Igor Solodovnikov
    Carabus (Megodontus) kolbei hidakamontanus Ishikawa, 1966
    Carabus (Megodontus) kolbei hidakamontanus Ishikawa, 1966
    Female adult
    E-Hokkaido: Samani distr., Horomankyo env.
    Copyright © Igor Solodovnikov
    Carabus (Megodontus) kolbei hidakamontanus Ishikawa, 1966
    Carabus (Megodontus) kolbei hidakamontanus Ishikawa, 1966
    Endophallus
    E-Hokkaido: Samani distr., Horomankyo env.
    Copyright © Igor Solodovnikov
    Carabus (Imaibius) wittmerorum vojtechi Schneider & Rapuzzi, 2021
    Carabus (Imaibius) wittmerorum vojtechi Schneider & Rapuzzi, 2021 Holotype Paratype
    Male+Female adult
    Pakistan, Province Khyber, District Swabi, Bir Gali env., 1900 m, 03.VI.2020, Ikram Ullah legit.
    Copyright © - - -
    Carabus (Ohomopterus) dehaanii dehaanii Chaudoir, 1848
    Carabus (Ohomopterus) dehaanii dehaanii Chaudoir, 1848
    Male adult
    Japan, S Honshu, Mie Pref., Inabe city, 21.II.2021, Shinichi S. leg., L=33 mm
    Copyright © Ruslan Panin
    Carabus (Ohomopterus) dehaanii dehaanii Chaudoir, 1848
    Carabus (Ohomopterus) dehaanii dehaanii Chaudoir, 1848
    Endophallus
    Japan, S Honshu, Mie Pref., Inabe city
    Copyright © Ruslan Panin
    Carabus (Tribax) nothus sharoiensis Fominykh, Zamotajlov, Avtaeva et Titarenko, 2019
    Carabus (Tribax) nothus sharoiensis Fominykh, Zamotajlov, Avtaeva et Titarenko, 2019 Paratype
    Male adult

    Copyright © Igor Solodovnikov
    Damaster (Coptolabrus) jankowskii irwolsanensis Kim, 2009
    Damaster (Coptolabrus) jankowskii irwolsanensis Kim, 2009 Holotype
    Male adult
    Mt. Irwol, Yeongyang-gun, Gyeongsangbuk-do, 26, vii, 2007
    Copyright © - - -
    Damaster (Coptolabrus) jankowskii unmunsanensis Kim, 2009
    Damaster (Coptolabrus) jankowskii unmunsanensis Kim, 2009 Paratype
    Female adult
    Mt. Unmun, Cheongdo-gun, Gyeongsangbuk-do, 12, viii, 2004
    Copyright © - - -
    Damaster (Coptolabrus) jankowskii ganghwadoensis Kim, 2009
    Damaster (Coptolabrus) jankowskii ganghwadoensis Kim, 2009 Paratype
    Female adult
    Mt. Goryeo, Ganghwa-gun, Gyeonggi-do, 16, viii, 2007
    Copyright © - - -
    Carabus (Neoplectes) ibericus lafertei Chaudoir, 1846
    Carabus (Neoplectes) ibericus lafertei Chaudoir, 1846
    Female adult
    SC-Georgia, Samtskhe–Javakheti, Meskheti Mt Range, Borjomi, Likani env., h=897 m, 41°50.125'N 43°19.905'E, VI.2018, D. Prunier leg., L=37 mm
    Copyright © Ruslan Panin
    Carabus (Procrustes) coriaceus ressli Mandl, 1961
    Carabus (Procrustes) coriaceus ressli Mandl, 1961
    Endophallus
    SC-Turkey:Hatay Prov, Dörtyol, Nur Mts, Topaktaş h=900-1200 m,
    Copyright © Ruslan Panin
    Carabus (Procrustes) coriaceus ressli Mandl, 1961
    Carabus (Procrustes) coriaceus ressli Mandl, 1961
    Male adult
    SC-Turkey: Hatay Prov, Dörtyol, Nur Mts, Topaktaş h=900-1200 m, 12.V.2005, D. Prunier leg.
    Copyright © Ruslan Panin
    Carabus (Procrustes) coriaceus ressli Mandl, 1961
    Carabus (Procrustes) coriaceus ressli Mandl, 1961
    Female adult
    SC-Turkey: Hatay Prov, Dörtyol, Nur Mts, Topaktaş h=900-1200 m, 12.V.2005, D. Prunier leg. L=35 mm
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) lusitanicus antiquus Dejean, 1826
    Carabus (Mesocarabus) lusitanicus antiquus Dejean, 1826
    Female adult
    SW Portugal, Beja Distr., 3,7 km NW Brunheiras, road CM1072, 37°46'31.9"N 8°47'02.2"W, 15.III.2011, D. Prunier leg., L=26 mm
    Copyright © Ruslan Panin
    Carabus (Ctenocarabus) melancholicus costatus Germar, 1824
    Carabus (Ctenocarabus) melancholicus costatus Germar, 1824
    Endophallus
    NW Portugal, Viana do Castelo distr., Sierra de Agra
    Copyright © Ruslan Panin
    Carabus (Tribax) nothus fausti Dohrn, 1873
    Carabus (Tribax) nothus fausti Dohrn, 1873
    Male adult
    Carabus (Microtribax) fausti fausti Dohrn, 1873 Russia: Daghestan, Kyabyak Mt.R.
    Copyright © Igor Solodovnikov
    Carabus (Tribax) nothus fausti Dohrn, 1873
    Carabus (Tribax) nothus fausti Dohrn, 1873
    Female adult
    Carabus (Microtribax) fausti fausti Dohrn, 1873 Russia: Daghestan, Kyabyak Mt.R.
    Copyright © Igor Solodovnikov
    Carabus (Diocarabus) fraterculus neochinensis Deuve & Li, 1998
    Carabus (Diocarabus) fraterculus neochinensis Deuve & Li, 1998
    Female adult
    NE-China: Liaoning prov., Fengcheng City, Shicheng, 20-28.VI.2011, Li Jingke leg., L=22 mm
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) protenes baoxingensis Deuve & Mourzine, 2003
    Carabus (Apotomopterus) protenes baoxingensis Deuve & Mourzine, 2003
    Male adult

    Copyright © Igor Solodovnikov
    Carabus (Apotomopterus) protenes baoxingensis Deuve & Mourzine, 2003
    Carabus (Apotomopterus) protenes baoxingensis Deuve & Mourzine, 2003
    Female adult

    Copyright © Igor Solodovnikov
    Carabus (Apotomopterus) protenes baoxingensis Deuve & Mourzine, 2003
    Carabus (Apotomopterus) protenes baoxingensis Deuve & Mourzine, 2003
    Endophallus
    China: С-Sichuan:Qionglai Shan Mts., Baoxing env. (TOPOTYPUS)
    Copyright © Igor Solodovnikov
    Carabus (Tribax) lederi lederi Reitter, 1882
    Carabus (Tribax) lederi lederi Reitter, 1882
    Male adult
    Georgia, Kvemo-Svaneti, Svanetian Mt.R., near Latpari pass., alpin zone, 2700-3000m (TOPOTYPUS)
    Copyright © Igor Solodovnikov
    Carabus (Tribax) lederi lederi Reitter, 1882
    Carabus (Tribax) lederi lederi Reitter, 1882
    Female adult
    Georgia, Kvemo-Svaneti, Svanetian Mt.R., near Latpari pass., alpin zone, 2700-3000m (TOPOTYPUS)
    Copyright © Igor Solodovnikov
    Carabus (Tribax) kasakorum kasakorum Semenov, 1896
    Carabus (Tribax) kasakorum kasakorum Semenov, 1896
    Male adult
    Russia: N.Osetia: Pastbyshchniy Mt.R., S slopes of Khoshkharanrag Mt.R., right bank of Amusadon riv., h=1500 m
    Copyright © Igor Solodovnikov
    Carabus (Tribax) kasakorum kasakorum Semenov, 1896
    Carabus (Tribax) kasakorum kasakorum Semenov, 1896
    Female adult
    Russia: N.Osetia: Pastbyshchniy Mt.R., S slopes of Khoshkharanrag Mt.R., right bank of Amusadon riv., h=1500 m
    Copyright © Igor Solodovnikov
    Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
    Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
    Male adult
    Russia, Caucasus, Ingushetia, South slopes of Skalistyi Mt Range, Stolovaya Mt., near Beyni, alpin zone, h = 2800 m, N 42°51’35.30",E 44°42’13.70", h = 2800m, 21.V–19.VII.2018, D. Fominykh leg.
    Copyright © Igor Solodovnikov
    Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
    Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
    Female adult
    Russia, Caucasus, Ingushetia, South slopes of Skalistyi Mt Range, Stolovaya Mt., near Beyni, alpin zone, h = 2800 m, N 42°51’35.30",E 44°42’13.70", h = 2800m, 21.V–19.VII.2018, D. Fominykh leg.
    Copyright © Igor Solodovnikov
    Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
    Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
    Endophallus
    Russia, Caucasus, Ingushetia, South slopes of Skalistyi Mt Range, Stolovaya Mt., near Beyni, alpin zone, h = 2800 m, N 42°51’35.30",E 44°42’13.70", h = 2800m, 21.V–19.VII.2018, D. Fominykh leg. (TOPOTYPUS)
    Copyright © Igor Solodovnikov
    Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
    Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
    Male adult
    Russia, Caucasus, Ingushetia, Skalistyi Mt. Range, S slopes of Tsoreylam Mt. Range, 5 km W of Mt. Khakhalgi, N 42°50’07.60", E 45°05’02.20", h=2500 m, 25.V–21.VII.2017, D. Fominykh leg.
    Copyright © Igor Solodovnikov
    Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
    Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
    Endophallus
    Russia, Caucasus, Ingushetia, Skalistyi Mt. Range, S slopes of Tsoreylam Mt. Range, 5 km W of Mt. Khakhalgi, N 42°50’07.60", E 45°05’02.20", h=2500 m, 25.V–21.VII.2017, D. Fominykh leg.
    Copyright © Igor Solodovnikov
    Carabus (Macrothorax) rugosus celtibericus Germar, 1824
    Carabus (Macrothorax) rugosus celtibericus Germar, 1824
    Male adult
    SW-Portugal, Serra do Cercal h=217 m, 37°47'15.6"N 8°42'02.0"W, road N390, 19.III.2011, D. Prunier leg., L=27 mm
    Copyright © Ruslan Panin
    Carabus (Macrothorax) rugosus celtibericus Germar, 1824
    Carabus (Macrothorax) rugosus celtibericus Germar, 1824
    Endophallus
    SW-Portugal, Serra do Cercal h=217 m, 37°47'15.6"N 8°42'02.0"W, road N390,
    Copyright © Ruslan Panin
    Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
    Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
    Female adult
    Russia, Chechnya, Caucasus, Skalystyi Mt. Range upper river Gekhi, Mt. Ekkyrkort, N 42°52', E 45°26', h=2230 m, 26.V–21.VII.2018, D.Fominykh leg.
    Copyright © Igor Solodovnikov
    Carabus (Diocarabus) fraterculus neochinensis Deuve & Li, 1998
    Carabus (Diocarabus) fraterculus neochinensis Deuve & Li, 1998
    Male adult
    NE-China: Liaoning prov., Fengcheng City, Shicheng, 20-28.VI.2011, Li Jingke leg., L=20 mm
    Copyright © Ruslan Panin
    Carabus (Ctenocarabus) melancholicus costatus Germar, 1824
    Carabus (Ctenocarabus) melancholicus costatus Germar, 1824
    Male adult
    NW Portugal, Viana do Castelo distr., Sierra de Agra, 41°47'52.3"N 8°41'27.6"W, h=786 m, 3.VII.2016, D. Prunier leg.
    Copyright © Ruslan Panin
    Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
    Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
    Endophallus
    Russia: N.Osetia, Bokovoy Mt.R., near Arkhonskiy pass., alpin zone, h = 2700 m, 23.V-21VII.2018, D. Fominykh leg.
    Copyright © Igor Solodovnikov
    Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
    Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
    Female adult
    Russia: N.Osetia, Bokovoy Mt.R., near Arkhonskiy pass., alpin zone, h = 2700 m, 23.V-21VII.2018, D. Fominykh leg.
    Copyright © Igor Solodovnikov
    Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
    Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
    Male adult
    Russia: N.Osetia, Bokovoy Mt.R., near Arkhonskiy pass., alpin zone, h = 2700 m, 23.V-21VII.2018, D. Fominykh leg.
    Copyright © Igor Solodovnikov
    Carabus (Tribax) puschkini kolenatii Chaudoir, 1846
    Carabus (Tribax) puschkini kolenatii Chaudoir, 1846
    Female adult
    C-Georgia, Trialeti Mt Range, 2 km SE of Borjomi h=916 m, 41°48'35.3"N 43°26'11.8"E, 17.VI.2013, D. Prunier leg./det. Panin R., L=29 mm
    Copyright © Ruslan Panin
    Carabus (Tribax) puschkini kolenatii Chaudoir, 1846
    Carabus (Tribax) puschkini kolenatii Chaudoir, 1846
    Male adult
    C-Georgia, Trialeti Mt Range, 2 km SE of Borjomi h=916 m, 41°48'35.3"N 43°26'11.8"E, 17.VI.2013, D. Prunier leg./det. Panin R., L=28 mm
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) lusitanicus trabuccarius Fairmaire, 1857
    Carabus (Mesocarabus) lusitanicus trabuccarius Fairmaire, 1857
    Endophallus
    NE Spain, Catalonia, Tarragona, Sierra de Montsant h=800 m, Albarca env.,
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) lusitanicus trabuccarius Fairmaire, 1857
    Carabus (Mesocarabus) lusitanicus trabuccarius Fairmaire, 1857
    Male adult
    NE Spain, Catalonia, Tarragona, Sierra de Montsant h=800 m, Albarca env.,7.XI.2018, T. Rafales leg., L=25 mm
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) lusitanicus trabuccarius Fairmaire, 1857
    Carabus (Mesocarabus) lusitanicus trabuccarius Fairmaire, 1857
    Female adult
    NE Spain, Catalonia, Tarragona, Sierra de Montsant h=800 m, Albarca env.,7.XI.2018, T. Rafales leg., L=27 mm
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) lusitanicus lusitanicus (as vieirae Oliveira, 1875)
    Carabus (Mesocarabus) lusitanicus lusitanicus (as vieirae Oliveira, 1875)
    Female adult
    CW-Portugal: Leiria distr., near Marinha Grande, 39°45'57.8"N 8°58'10.3"W, h-79 m, 19.IX.2016, D. Prunir leg., L=26 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) excellens excellens (as lomnitzkii Reitter, 1896)
    Carabus (Morphocarabus) excellens excellens (as lomnitzkii Reitter, 1896)
    Male adult
    W Ukraine, NW Podolian Upland, Lviv reg., Zolochiv distr., Lysa Gora steppe massif, 11.VII-14.VIII. 2021, Panin R. & D. Khrapov leg., L=25 mm
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) lusitanicus lusitanicus (as vieirae Oliveira, 1875)
    Carabus (Mesocarabus) lusitanicus lusitanicus (as vieirae Oliveira, 1875)
    Endophallus
    CW-Portugal: Leiria distr., near Marinha Grande, 39°45'57.8"N 8°58'10.3"W, h-79 m
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) lusitanicus lusitanicus (as vieirae Oliveira, 1875)
    Carabus (Mesocarabus) lusitanicus lusitanicus (as vieirae Oliveira, 1875)
    Male adult
    CW-Portugal: Leiria distr., near Marinha Grande, 39°45'57.8"N 8°58'10.3"W, h-79 m, 19.IX.2016, D. Prunir leg., L=24 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) excellens excellens (as lomnitzkii Reitter, 1896)
    Carabus (Morphocarabus) excellens excellens (as lomnitzkii Reitter, 1896)
    Male adult
    W Ukraine, NW Podolian Upland, Lviv reg., Zolochiv distr., Lysa Gora steppe massif, 11.VII-14.VIII. 2021, Panin R. & D. Khrapov leg., L=25 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) excellens excellens (as lomnitzkii Reitter, 1896)
    Carabus (Morphocarabus) excellens excellens (as lomnitzkii Reitter, 1896)
    Female adult
    W Ukraina, Podolian Upland, steppe reserve Kasova Gora near Burshtyn vill., 15.VI -28.VII.2020, R. Panin & D. Khrapov leg., L=27 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) excellens excellens (as lomnitzkii Reitter, 1896)
    Carabus (Morphocarabus) excellens excellens (as lomnitzkii Reitter, 1896)
    Male adult
    W Ukraina, Podolian Upland, steppe reserve Kasova Gora near Burshtyn vill., 15.VI -28.VII.2020, R. Panin & D. Khrapov leg., L=24 mm
    Copyright © Ruslan Panin
    Carabus (Relictocarabus) meurguesianus Ledoux, 1990
    Carabus (Relictocarabus) meurguesianus Ledoux, 1990
    Male adult
    Image from molbiol.ru
    Copyright © - - -
    Carabus (Shunichiocarabus) uenoianus uenoianus Imura, 1995
    Carabus (Shunichiocarabus) uenoianus uenoianus Imura, 1995
    Male adult

    Copyright © - - -
    Carabus (Tribax) boulbenicus Deuve, 2022
    Carabus (Tribax) boulbenicus Deuve, 2022 Holotype
    Male adult
    Géorgie, Svanétie, à l’est de Khaishi, vallée de la Kasleti, 950 m. R. Boulben et P. Stéfani leg.
    Copyright © Thierry Deuve
    Carabus (Tribax) adelphus shtavlerensis Fominykh, Zamotajlov et Titarenko, 2018
    Carabus (Tribax) adelphus shtavlerensis Fominykh, Zamotajlov et Titarenko, 2018 Holotype Paratype
    Male+Female adult
    Georgia, left bank of Nenskra River near Tita Vill., Shtavler Mt. Range, NW slopes of Utur Mt., 43°04'N, 42°11'E, 1048 m, 14.IV–13.VI.2016 D. Fominykh.
    Copyright © - - -
    Carabus (Tribax) adelphus adelphus Rost, 1892
    Carabus (Tribax) adelphus adelphus Rost, 1892
    Male+Female adult
    Muzhava Vill. environs
    Copyright © - - -
    Carabus (Carabus) arvensis sachalinensis Lapouge, 1906
    Carabus (Carabus) arvensis sachalinensis Lapouge, 1906
    Male adult
    Kuriles isl., Kunashir
    Copyright © Kirill Makarov
    Carabus (Euleptocarabus) porrecticollis kansaiensis Nakane, 1961
    Carabus (Euleptocarabus) porrecticollis kansaiensis Nakane, 1961
    Female adult
    Takatsuki (Osaka) JAPON, XI-1985
    Copyright © Alain Mollard
    Carabus (Pachystus) pisidicus pisidicus Peyron, 1854
    Carabus (Pachystus) pisidicus pisidicus Peyron, 1854
    Male adult
    Col Yellibel (Konya), TURQUIE VII-1984
    Copyright © Alain Mollard
    Carabus (Oreocarabus) gemellatus Menetries, 1832
    Carabus (Oreocarabus) gemellatus Menetries, 1832
    Female adult
    Col de Gascar (Mazaderan), IRAN, VI-2000
    Copyright © Alain Mollard
    Carabus (Pachycarabus) imitator katherinae Reitter, 1898
    Carabus (Pachycarabus) imitator katherinae Reitter, 1898
    Female adult
    Shoudidi Valley (Abkhasie) GEORGIE, VI-1990
    Copyright © Alain Mollard
    Carabus (Limnocarabus) clathratus arelatensis Lapouge, 1903
    Carabus (Limnocarabus) clathratus arelatensis Lapouge, 1903
    Female adult
    Aigues-Tortes, Camargue, France VI-1984
    Copyright © Alain Mollard
    Carabus (Limnocarabus) clathratus stygius Ganglbauer, 1890
    Carabus (Limnocarabus) clathratus stygius Ganglbauer, 1890
    Male adult
    Lenkoran (Narimanabad) AZERBAIDJAN III-1970
    Copyright © Alain Mollard
    Carabus (Trachycarabus) scabriusculus lippii Dejean, 1826
    Carabus (Trachycarabus) scabriusculus lippii Dejean, 1826
    Female adult
    SW Ukraine, Odessa reg., Ivanovka distr., steppe ravine near Gudevichevo vill., 22.IV-5.V.2022, A.Gontarenko leg., L=22 mm
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) scabriusculus lippii Dejean, 1826
    Carabus (Trachycarabus) scabriusculus lippii Dejean, 1826
    Male adult
    SW Ukraine, Odessa reg., Ivanovka distr., steppe ravine near Gudevichevo vill., 22.IV-5.V.2022, A.Gontarenko leg., L=21.5 mm
    Copyright © Ruslan Panin
    Carabus (Oreocarabus) tewoensis Deuve, 1992
    Carabus (Oreocarabus) tewoensis Deuve, 1992
    Female adult
    Tewo, (Gansu) ,CHINE, VI-2012
    Copyright © Alain Mollard
    Carabus (Oreocarabus) mikhaili loustei Deuve & Mourzine, 1998
    Carabus (Oreocarabus) mikhaili loustei Deuve & Mourzine, 1998
    Female adult
    Mts.Nanling (Sichuan) , CHINE , VI-2001
    Copyright © Alain Mollard
    Carabus (Oreocarabus) luetgensi pseudoguadarramus Mollard, 2006
    Carabus (Oreocarabus) luetgensi pseudoguadarramus Mollard, 2006 Paratype
    Female adult
    Montes de Herrera (Valladolid) Espagne, XI-2005
    Copyright © Alain Mollard
    Carabus (Pachystus) cribratus pulumuricus Cavazzuti, 1994
    Carabus (Pachystus) cribratus pulumuricus Cavazzuti, 1994
    Female adult
    Pulumur (Tunceli) TURQUIE, VI-1991
    Copyright © Alain Mollard
    Carabus (Pachystus) cribratus porrectangulus Gehin, 1885
    Carabus (Pachystus) cribratus porrectangulus Gehin, 1885
    Female adult
    Col d’Egribel (Bolu)), TURQUIE, VII-1999
    Copyright © Alain Mollard
    Carabus (Pachystus) cribratus yaraligozi Battoni, 1982
    Carabus (Pachystus) cribratus yaraligozi Battoni, 1982
    Male adult
    Yaraligoz (Kastamonu), TURQUIE VI-2006
    Copyright © Alain Mollard
    Carabus (Leptocarabus) kyushuensis cerberus Ishikawa, 1962
    Carabus (Leptocarabus) kyushuensis cerberus Ishikawa, 1962
    Female adult
    Fuchu (Mitsugi,Hiroshima) JAPON, VII-2001. 34 mm
    Copyright © Alain Mollard
    Carabus (Leptocarabus) kyushuensis nakatomii Ishikawa, 1966
    Carabus (Leptocarabus) kyushuensis nakatomii Ishikawa, 1966
    Male adult
    Tombara (Honshu,Shimane) JAPON, VII-2001. 36 mm
    Copyright © Alain Mollard
    Carabus (Leptocarabus) kyushuensis kyushuensis Nakane, 1960
    Carabus (Leptocarabus) kyushuensis kyushuensis Nakane, 1960
    Female adult
    Izumi, (Honshu, Kumamoto) JAPON, VII-2001. 31 mm
    Copyright © Alain Mollard
    Carabus (Leptocarabus) arboreus akitanus (Ishikawa, 1992)
    Carabus (Leptocarabus) arboreus akitanus (Ishikawa, 1992)
    Male adult
    Muriyoshi (Akita), JAPON, VIII-2001; 26 mm
    Copyright © Alain Mollard
    Carabus (Leptocarabus) arboreus babai Ishikawa, 1968
    Carabus (Leptocarabus) arboreus babai Ishikawa, 1968
    Male adult
    Hiychiyama (Niigata, Honshu), JAPON, XI-1984; 20 mm
    Copyright © Alain Mollard
    Carabus (Leptocarabus) arboreus exilis Bates, 1883
    Carabus (Leptocarabus) arboreus exilis Bates, 1883
    Male adult
    Sado Isle ( Honshu), JAPON, X-1994; 27 mm
    Copyright © Alain Mollard
    Carabus (Leptocarabus) arboreus gracillimus Bates, 1883
    Carabus (Leptocarabus) arboreus gracillimus Bates, 1883
    Female adult
    Mt.Tate-yama, 2000m (Toyama, Honshu), JAPON, X-1994; 20 mm
    Copyright © Alain Mollard
    Carabus (Leptocarabus) arboreus karibanus (Ishikawa, 1992)
    Carabus (Leptocarabus) arboreus karibanus (Ishikawa, 1992)
    Female adult
    Kuromatsunai (Hokkaido), JAPON, X-1994; 30 mm
    Copyright © Alain Mollard
    Carabus (Leptocarabus) arboreus kitakamisanus (Ishikawa, 1992)
    Carabus (Leptocarabus) arboreus kitakamisanus (Ishikawa, 1992)
    Male adult
    Niisato (Iwate, Honshu), JAPON, VIII-2001; 30 mm
    Copyright © Alain Mollard
    Carabus (Leptocarabus) arboreus nepta (Ishikawa, 1984)
    Carabus (Leptocarabus) arboreus nepta (Ishikawa, 1984)
    Male adult
    North, Hakkoda-san(Aomori, Honshu), JAPON, VI-1986 ; 25 mm
    Copyright © Alain Mollard
    Carabus (Leptocarabus) arboreus parexilis Nakane, 1961
    Carabus (Leptocarabus) arboreus parexilis Nakane, 1961
    Male adult
    Towada-Cho (Honshu), JAPON, X-1990; 30 mm
    Copyright © Alain Mollard
    Carabus (Leptocarabus) arboreus ogurai Ishikawa, 1969
    Carabus (Leptocarabus) arboreus ogurai Ishikawa, 1969
    Female adult
    Tabayama (Yamanashi, Honshu), JAPON, X-1990; 24 mm
    Copyright © Alain Mollard
    Carabus (Leptocarabus) arboreus pronepta (Ishikawa, 1992)
    Carabus (Leptocarabus) arboreus pronepta (Ishikawa, 1992)
    Male adult
    Hakkode-san (Aomori, Honshu), JAPON, XI-1990; 24 mm
    Copyright © Alain Mollard
    Carabus (Leptocarabus) arboreus shimoheiensis (Ishikawa, 1992)
    Carabus (Leptocarabus) arboreus shimoheiensis (Ishikawa, 1992)
    Female adult
    Kitakami (Iwate, Honshu), JAPON, XI-1986; 23 mm
    Copyright © Alain Mollard
    Carabus (Leptocarabus) arboreus shinanensis Born, 1922
    Carabus (Leptocarabus) arboreus shinanensis Born, 1922
    Male adult
    Mt. Yatsugatake (Nagano, Honshu), JAPON, VII-1985; 23 mm
    Copyright © Alain Mollard
    Carabus (Leptocarabus) arboreus tenuiformis Bates, 1883
    Carabus (Leptocarabus) arboreus tenuiformis Bates, 1883
    Female adult
    Mt. Osaki-Yama,1900m (Tochigi, Honshu), JAPON, VI-1988; 26 mm
    Copyright © Alain Mollard
    Carabus (Aulonocarabus) kurilensis pseudodiamesus Ivanovs, 1994
    Carabus (Aulonocarabus) kurilensis pseudodiamesus Ivanovs, 1994
    Male adult
    Mt.Chehova (Sachalin Isl.) VIII-1992; 22 mm
    Copyright © Alain Mollard
    Carabus (Aulonocarabus) kurilensis rausuanus Ishikawa, 1966
    Carabus (Aulonocarabus) kurilensis rausuanus Ishikawa, 1966
    Male adult
    Horoka-Kamishihoro-cho,(Kato-Gun,Hokkaido) JAPON, VII-2016; 26 mm
    Copyright © Alain Mollard
    Carabus (Aulonocarabus) kurilensis rishirensis Nakane, 1957
    Carabus (Aulonocarabus) kurilensis rishirensis Nakane, 1957
    Female adult
    Mt. Rishiri, (Rishiri Isl. Hokkaido) JAPON, VIII-1998; 22 mm
    Copyright © Alain Mollard
    Carabus (Leptocarabus) yokoae xunyangbanus Imura, 1998
    Carabus (Leptocarabus) yokoae xunyangbanus Imura, 1998
    Male adult
    Qinling Shan, (Xi’an, Shanxi) CHINE VI-1992; 27 mm
    Copyright © Alain Mollard
    Carabus (Leptocarabus) yokoae nanjiangensis Imura & Su, 2000
    Carabus (Leptocarabus) yokoae nanjiangensis Imura & Su, 2000
    Female adult
    Gangwu Shan (Taoyuan, Sichuan) CHINE VI-1992; 27 mm
    Copyright © Alain Mollard
    Carabus (Leptocarabus) seishinensis seunglaki (Kwon & Lee, 1984)
    Carabus (Leptocarabus) seishinensis seunglaki (Kwon & Lee, 1984)
    Male adult
    Taebaeck-san (Kangwon-sud) KOREE, VII-1998; 21 mm
    Copyright © Alain Mollard
    Carabus (Macrothorax) aumonti Lucas, 1849 (as sculptus Lapouge, 1925)
    Carabus (Macrothorax) aumonti Lucas, 1849 (as sculptus Lapouge, 1925)
    Male adult
    Lalla Maghrnia , ALGERIE, V-1927; 42 mm
    Copyright © Alain Mollard
    Carabus (Morphocarabus) zawadzkii zawadzkii, Kraatz 1854
    Carabus (Morphocarabus) zawadzkii zawadzkii, Kraatz 1854
    Male adult
    W Ukraine, NE Carpathians, Lviv region, Skole Beskids Mts, Zelemyanka Mt. Range h=915 m, 22.VI-17.VII.2022, Panin R. & Kanarsky Yu. leg., L= 32 mm
    Copyright © Ruslan Panin
    Carabus (Iniopachus) pyrenaeus cerdanus Lapouge, 1924
    Carabus (Iniopachus) pyrenaeus cerdanus Lapouge, 1924
    Male+Female adult
    Cirque des Peyssons, ANDORRE, VI-1985
    Copyright © Alain Mollard
    Carabus (Iniopachus) pyrenaeus costulus Gehin, 1885
    Carabus (Iniopachus) pyrenaeus costulus Gehin, 1885
    Male+Female adult
    Mt. Canigou, 2400m (Pyrénées Orientales, Pyrénées) FRANCE, VI-1985
    Copyright © Alain Mollard
    Carabus (Iniopachus) pyrenaeus vallierensis Colas, 1965
    Carabus (Iniopachus) pyrenaeus vallierensis Colas, 1965
    Male+Female adult
    Mt.Vallier (Ariege,Pyrénées) FRANCE, VI-1998
    Copyright © Alain Mollard
    Carabus (Sphodristocarabus) biroi biroi Menetries, 1832
    Carabus (Sphodristocarabus) biroi biroi Menetries, 1832
    Male+Female adult
    Amasya (Samsun), TURQUIE VI-1987
    Copyright © Alain Mollard
    Carabus (Sphodristocarabus) bohemani esfandiarii Heinz, 1973
    Carabus (Sphodristocarabus) bohemani esfandiarii Heinz, 1973
    Female adult
    Hasan-Kif (Mazandaran) IRAN VII-2000
    Copyright © Alain Mollard
    Carabus (Sphodristocarabus) bohemani esperanzae Heinz, 1970
    Carabus (Sphodristocarabus) bohemani esperanzae Heinz, 1970
    Male+Female adult
    Rasht, (Masuleh) IRAN VI-2000
    Copyright © Alain Mollard
    Carabus (Sphodristocarabus) elegantulus Motschulsky, 1850 (as astrabadensis Semenov, 1903)
    Carabus (Sphodristocarabus) elegantulus Motschulsky, 1850 (as astrabadensis Semenov, 1903)
    Male+Female adult
    Ft.Pashmaky, (Golestan) IRAN VI-2000
    Copyright © Alain Mollard
    Carabus (Sphodristocarabus) giachinoi Cavazzuti, 1991
    Carabus (Sphodristocarabus) giachinoi Cavazzuti, 1991
    Female adult
    Ulukoy, (Gumushane) TURQUIE VI-1991
    Copyright © Alain Mollard
    Carabus (Sphodristocarabus) gilnickii brunneicupreus Deuve, 1994
    Carabus (Sphodristocarabus) gilnickii brunneicupreus Deuve, 1994
    Male+Female adult
    Camlik, (Rize), TURQUIE VI-1997
    Copyright © Alain Mollard
    Carabus (Sphodristocarabus) gilnickii hasandaghensis Darge & Basquin, 1990
    Carabus (Sphodristocarabus) gilnickii hasandaghensis Darge & Basquin, 1990
    Male+Female adult
    Ispir (Erzurum), TURQUIE VI-1997
    Copyright © Alain Mollard
    Carabus (Sphodristocarabus) heinzi heinzi Breuning, 1964
    Carabus (Sphodristocarabus) heinzi heinzi Breuning, 1964
    Male+Female adult
    Egribel (Gerisum), TURQUIE VI-1992
    Copyright © Alain Mollard
    Carabus (Sphodristocarabus) heinzi mateuellus Deuve, 2003
    Carabus (Sphodristocarabus) heinzi mateuellus Deuve, 2003
    Male adult
    Col Harçebeli,(Ordu) TURQUIE V-2001
    Copyright © Alain Mollard
    Carabus (Sphodristocarabus) kurdicus kurdicus Heinz, 1975
    Carabus (Sphodristocarabus) kurdicus kurdicus Heinz, 1975
    Female adult
    Resadiye,(Bitlis) TURQUIE V-1991
    Copyright © Alain Mollard
    Carabus (Sphodristocarabus) macrogonus akkusensis Breuning & Ruspoli, 1970
    Carabus (Sphodristocarabus) macrogonus akkusensis Breuning & Ruspoli, 1970 Paratype
    Male+Female adult
    Akkus,(Ordu) TURQUIE X-1962
    Copyright © Alain Mollard
    Carabus (Sphodristocarabus) macrogonus diogenei Cavazzuti, 1991
    Carabus (Sphodristocarabus) macrogonus diogenei Cavazzuti, 1991
    Female adult
    Gokçebel,(Sinop) TURQUIE VI-1988
    Copyright © Alain Mollard
    Carabus (Sphodristocarabus) macrogonus keltepensis Vigna-Taglianti & Bruschi, 1976
    Carabus (Sphodristocarabus) macrogonus keltepensis Vigna-Taglianti & Bruschi, 1976
    Female adult
    Mt. Keltepe (Izmit) TURQUIE VI-2005
    Copyright © Alain Mollard
    Carabus (Sphodristocarabus) macrogonus drannadaghensis Breuning, 1964
    Carabus (Sphodristocarabus) macrogonus drannadaghensis Breuning, 1964
    Male+Female adult
    Drannaz Dag,(Sinop) TURQUIE VII-1993
    Copyright © Alain Mollard
    Carabus (Sphodristocarabus) macrogonus schweigerinae Schweiger, 1962
    Carabus (Sphodristocarabus) macrogonus schweigerinae Schweiger, 1962
    Male+Female adult
    Lac d’Abant, (Bolu) TURQUIE VIII-1993
    Copyright © Alain Mollard
    Carabus (Sphodristocarabus) tokatensis tokatensis Roeschke, 1898
    Carabus (Sphodristocarabus) tokatensis tokatensis Roeschke, 1898
    Male+Female adult
    Yilidiz Dag, (Tokat) TURQUIE VI-1985
    Copyright © Alain Mollard
    Carabus (Sphodristocarabus) tokatensis simonae Cavazzuti, 1984
    Carabus (Sphodristocarabus) tokatensis simonae Cavazzuti, 1984
    Male+Female adult
    Pulumur, (Tunceli) TURQUIE VI-1988
    Copyright © Alain Mollard
    Carabus (Sphodristocarabus) tokatensis kosensis Cavazzuti, 1984
    Carabus (Sphodristocarabus) tokatensis kosensis Cavazzuti, 1984
    Female adult
    Kosedagi-Geç (Gumushane) TURQUIE VII-2000
    Copyright © Alain Mollard
    Carabus (Sphodristocarabus) scovitzi hedini Heinz, 1973
    Carabus (Sphodristocarabus) scovitzi hedini Heinz, 1973
    Male+Female adult
    Borçka (Artvin) TURQUIE V-2005
    Copyright © Alain Mollard
    Carabus (Sphodristocarabus) scovitzi sarigoelensis Deuve & Charet, 1992
    Carabus (Sphodristocarabus) scovitzi sarigoelensis Deuve & Charet, 1992
    Male+Female adult
    Kaçkar (Artvin) TURQUIE VII-1992
    Copyright © Alain Mollard
    Carabus (Sphodristocarabus) adamsi adamsi (as adamsianus Mandl, 1975)
    Carabus (Sphodristocarabus) adamsi adamsi (as adamsianus Mandl, 1975)
    Male adult
    Dagestan. det & Photo I. Belousov
    Copyright © Creative Commons
    Carabus (Sphodristocarabus) adamsi adamsi (as adamsianus Mandl, 1975)
    Carabus (Sphodristocarabus) adamsi adamsi (as adamsianus Mandl, 1975)
    Female adult
    Dagestan. det & Photo I. Belousov
    Copyright © Creative Commons
    Carabus (Sphodristocarabus) adamsi hollbergi Mannerheim, 1827
    Carabus (Sphodristocarabus) adamsi hollbergi Mannerheim, 1827
    Male adult
    det & Photo I. Belousov
    Copyright © Creative Commons
    Carabus (Oxycarabus) saphyrinus notabilis Roeschke, 1898
    Carabus (Oxycarabus) saphyrinus notabilis Roeschke, 1898
    Male adult
    Ulu Dag (Bursa) TURQUIE VI-1990
    Copyright © Alain Mollard
    Carabus (Oxycarabus) saphyrinus saphyrinus Cristoforis & Jan, 1837
    Carabus (Oxycarabus) saphyrinus saphyrinus Cristoforis & Jan, 1837
    Male adult
    Gulug (Kumbet, Giresun) TURQUIE VII-1981
    Copyright © Alain Mollard
    Carabus (Imaibius) barysomus heroicus Semenov, 1906
    Carabus (Imaibius) barysomus heroicus Semenov, 1906
    Female adult
    Sudhangali (Muzaffarba) CASHMIRE, VIII-2008
    Copyright © Alain Mollard
    Carabus (Imaibius) barysomus huegeli Roeschke, 1907
    Carabus (Imaibius) barysomus huegeli Roeschke, 1907
    Female adult
    Daksum (Hindoustan) CASHMIRE, VIII-1980
    Copyright © Alain Mollard
    Carabus (Imaibius) cavifrons Mandl, 1974
    Carabus (Imaibius) cavifrons Mandl, 1974
    Female adult
    Bumra Uhel (Kuz-Heol) NEPAL, VI-1980
    Copyright © Alain Mollard
    Carabus (Sphodristocarabus) enigmaticus enigmaticus Heinz, 1980
    Carabus (Sphodristocarabus) enigmaticus enigmaticus Heinz, 1980
    Male+Female adult
    Ispir (Erzurum) TURQUIE VI-1991
    Copyright © Alain Mollard
    Carabus (Sphodristocarabus) enigmaticus citamgineus Lassalle & Charet, 1992
    Carabus (Sphodristocarabus) enigmaticus citamgineus Lassalle & Charet, 1992
    Male+Female adult
    Gole (Erzurum) TURQUIE VII-1991
    Copyright © Alain Mollard
    Carabus (Archicarabus) wiedemanni pseudorugosus Machard, 1994
    Carabus (Archicarabus) wiedemanni pseudorugosus Machard, 1994
    Male+Female adult
    Col Buglan (Mus) TURQUIE VI-1981
    Copyright © Alain Mollard
    Carabus (Archicarabus) wiedemanni congruus Motschulsky, 1865
    Carabus (Archicarabus) wiedemanni congruus Motschulsky, 1865
    Female adult
    Turkey, Cankiri, Cerkes - Kursunlu, Atkaracalar 1250m 10.7.1996 Leg Staven and Skoupy (DUBC)
    Copyright © Alexander Anichtchenko
    Carabus (Procrustes) chevrolati internatus Heinz & Korge, 1964
    Carabus (Procrustes) chevrolati internatus Heinz & Korge, 1964
    Female adult
    Turkey, Bolu, Gerede, Esentepe 1350m, 1.7.1994, leg. K. Staven
    Copyright © Alexander Anichtchenko
    Carabus (Archicarabus) wiedemanni wiedemanni (as mussardianus Breuning, 1965)
    Carabus (Archicarabus) wiedemanni wiedemanni (as mussardianus Breuning, 1965)
    Female adult
    NC-Turkey, Sinop province, Erfelek env., h=200m, 10-27.VII.2014, C. Reuter leg. L=24 mm
    Copyright © Ruslan Panin
    Carabus (Archicarabus) wiedemanni (as mussardianus Breuning, 1965)
    Carabus (Archicarabus) wiedemanni (as mussardianus Breuning, 1965)
    Male adult
    NC-Turkey, Sinop province, Erfelek env., h=200m, 10-27.VII.2014, C. Reuter leg. L=23 mm
    Copyright © Ruslan Panin
    Carabus (Cechenochilus) boeberi felix Semenov & Znojko, 1931
    Carabus (Cechenochilus) boeberi felix Semenov & Znojko, 1931
    Male adult

    Copyright © Kirill Makarov
    Carabus (Archicarabus) wiedemanni congruus Motschulsky, 1865 (as ilgazicus Breining, 1970)
    Carabus (Archicarabus) wiedemanni congruus Motschulsky, 1865 (as ilgazicus Breining, 1970)
    Female adult
    Ilgaz (Cankiri) TURQUIE VI-1990
    Copyright © Alain Mollard
    Carabus (Archicarabus) wiedemanni congruus Motschulsky, 1865 (as ilgazicus Breining, 1970)
    Carabus (Archicarabus) wiedemanni congruus Motschulsky, 1865 (as ilgazicus Breining, 1970)
    Male adult
    Ilgaz (Cankiri) TURQUIE VI-1990
    Copyright © Alain Mollard
    Carabus (Archicarabus) wiedemanni congruus Motschulsky, 1865 (as turcicus Motschulsky, 1865)
    Carabus (Archicarabus) wiedemanni congruus Motschulsky, 1865 (as turcicus Motschulsky, 1865)
    Female adult
    Camlibel(Tokat) TURQUIE VII-1991
    Copyright © Alain Mollard
    Carabus (Neoplectes) reitteri reitteri Retowski, 1885
    Carabus (Neoplectes) reitteri reitteri Retowski, 1885
    Male adult

    Copyright © Kirill Makarov
    Carabus (Carabus) granulatus yezoensis Bates, 1883
    Carabus (Carabus) granulatus yezoensis Bates, 1883
    Female adult

    Copyright © Kirill Makarov
    Carabus (Carabus) menetriesi menetriesi Hummel, 1827
    Carabus (Carabus) menetriesi menetriesi Hummel, 1827
    Female adult

    Copyright © Kirill Makarov
    Carabus (Megodontus) violaceus aurolimbatus Dejean, 1829
    Carabus (Megodontus) violaceus aurolimbatus Dejean, 1829
    Larva
    N Ukraine, Chernigov reg., Dmitrovka viil., photo by Yakubovsky E.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) odoratus Motschulsky, 1844
    Carabus (Morphocarabus) odoratus Motschulsky, 1844
    Male adult

    Copyright © Kirill Makarov
    Carabus (Ohomopterus) albrechti okumurai (Ishikawa, 1966)
    Carabus (Ohomopterus) albrechti okumurai (Ishikawa, 1966)
    Male+Female adult
    Mt. Daibosatzu (Yamanashi, Honshu) JAPON
    Copyright © Alain Mollard
    Carabus (Ohomopterus) albrechti tohokuensis Ishikawa, 1984
    Carabus (Ohomopterus) albrechti tohokuensis Ishikawa, 1984
    Male+Female adult
    Hashimori Pass (Iwate) JAPON
    Copyright © Alain Mollard
    Carabus (Ohomopterus) albrechti freyi van Emden, 1932
    Carabus (Ohomopterus) albrechti freyi van Emden, 1932
    Male+Female adult
    Sado Island (Niigata) JAPON
    Copyright © Alain Mollard
    Carabus (Ohomopterus) iwawakianus narukawai Ishikawa & Kubota, 1996
    Carabus (Ohomopterus) iwawakianus narukawai Ishikawa & Kubota, 1996
    Male+Female adult
    Tsuchiyama (Mie,Honshu) JAPON
    Copyright © Alain Mollard
    Carabus (Ohomopterus) arrowianus nakamurai (Ishikawa, 1966)
    Carabus (Ohomopterus) arrowianus nakamurai (Ishikawa, 1966)
    Female adult
    Mt.Takamoriyama (Tida-City, Nagano) JAPON VI-1987
    Copyright © Alain Mollard
    Carabus (Ohomopterus) chugokuensis chugokuensis (Nakane, 1961)
    Carabus (Ohomopterus) chugokuensis chugokuensis (Nakane, 1961)
    Male+Female adult
    Mt.Takasho (Shiman) JAPON VI-1987
    Copyright © Alain Mollard
    Carabus (Ohomopterus) japonicus awajiensis Imura, Dejima & Mizusawa, 1993
    Carabus (Ohomopterus) japonicus awajiensis Imura, Dejima & Mizusawa, 1993
    Male adult
    Mt.Sen-Zan,(Awaji Isle, Hyogo ) JAPON VI-1996
    Copyright © Alain Mollard
    Carabus (Pachystus) glabratus glabratus Paykull, 1790
    Carabus (Pachystus) glabratus glabratus Paykull, 1790
    Male adult

    Copyright © Kirill Makarov
    Carabus (Oreocarabus) hortensis hortensis Linne, 1758
    Carabus (Oreocarabus) hortensis hortensis Linne, 1758
    Male adult

    Copyright © Kirill Makarov
    Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
    Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
    Male adult

    Copyright © Kirill Makarov
    Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
    Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
    Female adult

    Copyright © Kirill Makarov
    Carabus (Megodontus) vietinghoffi vietinghoffi Adams, 1812
    Carabus (Megodontus) vietinghoffi vietinghoffi Adams, 1812
    Female adult

    Copyright © Kirill Makarov
    Carabus (Hemicarabus) nitens Linne, 1758
    Carabus (Hemicarabus) nitens Linne, 1758
    Female adult

    Copyright © Kirill Makarov
    Carabus (Pachycarabus) koenigi Ganglbauer, 1886
    Carabus (Pachycarabus) koenigi Ganglbauer, 1886
    Male adult

    Copyright © Kirill Makarov
    Carabus (Pachycarabus) staehlini staehlini Adams, 1817
    Carabus (Pachycarabus) staehlini staehlini Adams, 1817
    Male adult

    Copyright © Kirill Makarov
    Carabus (Tribax) apschuanus galianus Belousov & Zamotajlov, 1993
    Carabus (Tribax) apschuanus galianus Belousov & Zamotajlov, 1993 Paratype
    Male+Female adult
    Mt.Okhachkue (L’Achake) GEORGIE, VII-1991
    Copyright © Alain Mollard
    Carabus (Tribax) biebersteini hurkai Gottwald, 1980
    Carabus (Tribax) biebersteini hurkai Gottwald, 1980
    Male+Female adult
    Chkumi (Racha-Lechkumi) GEORGIE, VII-2018
    Copyright © Alain Mollard
    Carabus (Tribax) kasbekianus mingrelicus Reitter, 1889
    Carabus (Tribax) kasbekianus mingrelicus Reitter, 1889
    Male+Female adult
    Chkhorodukhu GEORGIE, VII-2018
    Copyright © Alain Mollard
    Carabus (Tribax) osseticus Adams, 1817
    Carabus (Tribax) osseticus Adams, 1817
    Male+Female adult
    Kzabegi , GEORGIE, VIII-1991
    Copyright © Alain Mollard
    Carabus (Tribax) agnatus agnatus Ganglbauer, 1889
    Carabus (Tribax) agnatus agnatus Ganglbauer, 1889
    Female adult

    Copyright © Kirill Makarov
    Carabus (Coptolabrus) nankotaizanus Kano, 1932 (as shimonoyai Kurosawa & Kudo, 1980)
    Carabus (Coptolabrus) nankotaizanus Kano, 1932 (as shimonoyai Kurosawa & Kudo, 1980)
    Male adult
    Mt.Lala-Shan, (Fuxing,Taoyuan) TAIWAN (FORMOSE) II-1997
    Copyright © Alain Mollard
    Carabus (Coptolabrus) augustus ignigena Hauser, 1914 (as lautus)
    Carabus (Coptolabrus) augustus ignigena Hauser, 1914 (as lautus)
    Male+Female adult
    Mt.Yingpan Shan (Qiubei Xian, Yunnan) CHINE, VII-2008
    Copyright © Alain Mollard
    Carabus (Coptolabrus) ignimitella angulicollis Hauser, 1911
    Carabus (Coptolabrus) ignimitella angulicollis Hauser, 1911
    Male+Female adult
    Mt.Datian Ding (Baishi,Guangdong) CHINE, VII-2008 (F 52mm, M 46mm)
    Copyright © Alain Mollard
    Carabus (Coptolabrus) ignimitella angulicollis Hauser, 1911 (as mellianus Kolbe, 1914)
    Carabus (Coptolabrus) ignimitella angulicollis Hauser, 1911 (as mellianus Kolbe, 1914)
    Male+Female adult
    Tianjing Shan (Luyuan, Lecheng), Guangdong) CHINE, VII-2008 (m 48mm, fem 50mm)
    Copyright © Alain Mollard
    Carabus (Damaster) blaptoides blaptoides (Kollar, 1836)
    Carabus (Damaster) blaptoides blaptoides (Kollar, 1836)
    Female adult
    Shiobara (Tochigi), JAPON, IV-1990
    Copyright © Alain Mollard
    Carabus (Damaster) blaptoides blaptoides (Kollar, 1836) (as lewisi Rye, 1872)
    Carabus (Damaster) blaptoides blaptoides (Kollar, 1836) (as lewisi Rye, 1872)
    Female adult
    Kitakata (Myakasi, Kyushu) JAPON, VI-1989
    Copyright © Alain Mollard
    Carabus (Damaster) blaptoides fortunei Adams, 1861
    Carabus (Damaster) blaptoides fortunei Adams, 1861
    Male+Female adult
    Awa-Shima Isle (Niigata) JAPON III-1990
    Copyright © Alain Mollard
    Carabus (Acoptolabrus) leechi yooni Deuve, 1990
    Carabus (Acoptolabrus) leechi yooni Deuve, 1990
    Male+Female adult
    Sorak-San (Kyongi) KOREE, VI-2002
    Copyright © Alain Mollard
    Carabus (Megodontus) kolbei urupiensis Obydov, 2007
    Carabus (Megodontus) kolbei urupiensis Obydov, 2007 Holotype
    Female adult
    South Kuril Islands, Urup Island, north-western coast of Tokotan Lake, 21-29.VII. 2006, A. K. Klitin leg.
    Copyright © - - -
    Carabus (Megodontus) kolbei chishimanus Nakane, 1961
    Carabus (Megodontus) kolbei chishimanus Nakane, 1961
    Male adult
    Iturup isl.
    Copyright © - - -
    Carabus (Megodontus) kolbei ushishirensis Obydov & Saldaitis, 1996
    Carabus (Megodontus) kolbei ushishirensis Obydov & Saldaitis, 1996 Holotype Paratype
    Male+Female adult
    Holotype: Ushishir Islands, Yankicha, N and SE part of Kraternaya, 47°30'N, 152°48'E, 20.VIII.1995, Yu. Marusik leg.". Paratype: Ketoy Island
    Copyright © - - -
    Carabus (Tribax) biebersteini vlasenkoi Obydov, 2011
    Carabus (Tribax) biebersteini vlasenkoi Obydov, 2011 Holotype Paratype
    Male+Female adult
    Russia, North Caucasus,Stavropol Region, 33 km from Kislovodsk City to the north-west, Predgornyi Dist.,8 km from st. Bekeshevskaya to the south-west, Forestry “Ugolnaya Dacha”,Pastbischnyi Mt. Ridge, left coast of Kuma River, 800 m, N44º02’ E42°21’, 11-17.05.2006, A. Vlasenko leg.
    Copyright © - - -
    Carabus (Neoplectes) edithae miloslavskii Obydov, 2010
    Carabus (Neoplectes) edithae miloslavskii Obydov, 2010 Holotype Paratype
    Male+Female adult
    Holotype: Caucasus centr., Archyz, zona subalp., 15.VI.1967, A. Bogachev leg; Paratype: KarachaiCherkessian Region, Zagedan Mt. Ridge, 2500 m., 6-7.VII.1994, Melnik leg.
    Copyright © - - -
    Carabus (Morphocarabus) chaudoiri juliae Obydov, 2002
    Carabus (Morphocarabus) chaudoiri juliae Obydov, 2002 Holotype Paratype
    Male+Female adult
    Russia, Tuva Republic, Mungun-Taiga, Khindiktig-Khol Lake, 2400 m, 22.-29.VI.2000, M. Beladič leg.
    Copyright © - - -
    Carabus (Megodontus) leachi semiensis Obydov, 2010
    Carabus (Megodontus) leachi semiensis Obydov, 2010 Holotype
    Male adult
    E. Kazakhstan, Semei (Semipalatinsk) env., 12-26.VII.1992, V. Sinyaev leg.
    Copyright © - - -
    Carabus (Diocarabus) slovtzovi rasae Obydov, 2001
    Carabus (Diocarabus) slovtzovi rasae Obydov, 2001 Holotype
    Male adult
    Tuva, Akademika Obrucheva Mt Ridge, 2100 m, N52°00’ E95º34’, Sainak Pass, 2.VII.1998, D. Obydov leg.
    Copyright © - - -
    Carabus (Acoptolabrus) constricticollis fei Rapuzzi, Li & Zhang, 2012
    Carabus (Acoptolabrus) constricticollis fei Rapuzzi, Li & Zhang, 2012 Holotype Paratype
    Male+Female adult
    North East China, South West Liaoning Province, Liadong Peninsula, Haicheng City, Gusahanzhen, ErDaoHe, 700 m. 17/19-VI-2011 (Holotype female, in the collection of Key Laboratory of remote sensing monitoring of geographic environment, College of Heilongjiang Province, Harbin Normal University, Harbin, China)
    Copyright © - - -
    Carabus (Homoeocarabus) maeander evenkiensis Obydov, 2010
    Carabus (Homoeocarabus) maeander evenkiensis Obydov, 2010 Holotype Paratype
    Male+Female adult
    North Siberia, Evenk Autonomous Region, N Tunguska River, 60 km up to Tura Village, Tabachnyi Stream, 19.VI.1972
    Copyright © - - -
    Carabus (Tribax) apschuanus galianus Belousov & Zamotajlov, 1993
    Carabus (Tribax) apschuanus galianus Belousov & Zamotajlov, 1993 Paratype
    Male adult
    Eastern Abkhazia, south slope of Mt. Okhachku, 4.V.1989, I. Belousov leg.
    Copyright © - - -
    Carabus (Megodontus) croaticus antoniocaldoni Rapuzzi, 2014
    Carabus (Megodontus) croaticus antoniocaldoni Rapuzzi, 2014 Holotype Paratype
    Male+Female adult
    Serbia, Kopaonik mountain, 1500m, IX.2013, I. Rapuzzi and L. Caldon leg; (Ivan Rapuzzi collection).
    Copyright © - - -
    Carabus (Morphocarabus) mestscherjakovi tuvinicus Obydov, 2002
    Carabus (Morphocarabus) mestscherjakovi tuvinicus Obydov, 2002 Holotype
    Female adult
    Россия, Тува, хр. Таскыл, долина р. Уюк, 4.VI-8VI.1999, И. Белло leg.
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) odoratus chlebnikovae Obydov 2005
    Carabus (Morphocarabus) odoratus chlebnikovae Obydov 2005 Paratype
    Female adult
    Россия, Северный Байкал, 80 км Ю Северобайкальска, 5 км Ю истока р. Куркула, 19.VII.2003, М. Хлебникова leg.
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) odoratus putoranicus Obydov, 2002
    Carabus (Morphocarabus) odoratus putoranicus Obydov, 2002 Paratype
    Male adult
    Россия, Север Вост. Сибири, Плато Путорана, окр. оз. Аян, 26.VI-10.VIII.1978, M. Сладков leg.
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) odoratus taskylensis Obydov, 2001
    Carabus (Morphocarabus) odoratus taskylensis Obydov, 2001 Holotype
    Male adult
    Россия, Тува, хр. Таскыл, р. Уюк, 4.VI-8.VI.1999, И. Белло и М. Беладжик leg.
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) odoratus jacuticus Obydov, 2002
    Carabus (Morphocarabus) odoratus jacuticus Obydov, 2002 Paratype
    Male adult
    Россия, Якутия, В. Хандыга, VII.1993, А. Наглис leg.
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) hummeli jurgae Obydov, 1995
    Carabus (Morphocarabus) hummeli jurgae Obydov, 1995 Holotype
    Male adult
    Россия, Якутия, 300 км севернее хр. Сунта-Хаята, VII.1993, Н. Шивис leg.
    Copyright © Dmitri Obydov
    Carabus (Mesocarabus) problematicus kolskianus Obydov, 2008
    Carabus (Mesocarabus) problematicus kolskianus Obydov, 2008 Holotype Paratype
    Male+Female adult
    Holotypus, male - Россия, Кольский полуостров, Мурман (Мурманск), Александровск (г. Полярный), Л. Зенкевич leg. Paratypus, female - Россия, Кольский полуостров, Мурман, окр. Александровска, 1914, Л. Зенкевич leg.
    Copyright © Dmitri Obydov
    Carabus (Diocarabus) loschnikovi tsaganensis Obydov, 2001
    Carabus (Diocarabus) loschnikovi tsaganensis Obydov, 2001 Paratype
    Male adult
    Россия, Ю-З Тува, хр. Цаган-Шибэту, 2230 м, окр. Мугур-Аксу, 50°20'N 90°40'E, 22.VII.2000, Д. Обыдов leg.
    Copyright © Dmitri Obydov
    Carabus (Diocarabus) slovtzovi mugurensis Gottwald, 1993
    Carabus (Diocarabus) slovtzovi mugurensis Gottwald, 1993
    Male adult
    Ю-З Тува, окр. Мугур-Аксу, около оз. Хиндиктиг-Холь, 2500 м, 50°21'N 89°56'E, 24-25.VII.2000, Д. Обыдов leg.
    Copyright © Dmitri Obydov
    Carabus (Megodontus) tuvensis Shilenkov, 1996
    Carabus (Megodontus) tuvensis Shilenkov, 1996 Holotype Paratype
    Male+Female adult
    Holotypus (m) - Россия, Тува, 80 км С пос. Тээли, 25.06.1972, Б.А. Коротяев leg. Paratypus (f) - Россия, Тува, Саяны, р. Ак-Суг, 26.06.1972, Б.А. Коротяев leg.
    Copyright © Dmitri Obydov
    Carabus (Megodontus) obovatus pseudoleachi Obydov, 1999
    Carabus (Megodontus) obovatus pseudoleachi Obydov, 1999 Holotype
    Male adult
    Россия, Алтай, окр. пос. Яйлю, 06.1988, Водянов leg.
    Copyright © Dmitri Obydov
    Carabus (Megodontus) obovatus taizhensis Obydov, 2006
    Carabus (Megodontus) obovatus taizhensis Obydov, 2006 Paratype
    Male adult
    Россия, Кузне-кий Алатау, долина р. Тайже-Су, около Хакасской границы, 1250-1650 м, 53°55'N, 89°10'E, 25.VI-30.VII.2002, В. Мурзин и В. Синяев leg.
    Copyright © Dmitri Obydov
    Carabus (Megodontus) leachi sarymsaktensis Obydov, 1999
    Carabus (Megodontus) leachi sarymsaktensis Obydov, 1999 Paratype
    Male adult
    Казахстан, Южный Алтай, 2500 - 3400 м, хр. Сарым-Сакты, 21.VI.1998, A. Клименко leg.
    Copyright © Dmitri Obydov
    Carabus (Megodontus) danilevskii Obydov, 1993
    Carabus (Megodontus) danilevskii Obydov, 1993 Holotype Paratype
    Male+Female adult
    Россия, Зап. Саян, левый берег р. Енисей, хр. Джойский, 700-800 м, около пос. Черемушки, 20-28.VII.1992, Д. Обыдов leg.
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) chaudoiri chaudoiri Gebler, 1847
    Carabus (Morphocarabus) chaudoiri chaudoiri Gebler, 1847 Lectotype
    Female adult
    Altay
    Copyright © Dmitri Obydov
    Carabus (Morphocarabus) chaudoiri shilenkovi O.Berlov, 1989
    Carabus (Morphocarabus) chaudoiri shilenkovi O.Berlov, 1989
    Male adult
    Россия, Якутия, Оймяконское нагорье, 12-18.VII.1998
    Copyright © Dmitri Obydov
    Carabus (Trachycarabus) planarius Obydov, 1994
    Carabus (Trachycarabus) planarius Obydov, 1994 Paratype
    Female adult
    Казахстан, Семипалатинский р-н, окр. пос. Шолпан, 500 м, 20 км. южнее пос. Таскескен, 2.06.1993, М. Данилевский leg.
    Copyright © Dmitri Obydov
    Carabus (Trachycarabus) sibiricus seriatoporus Fischer von Waldheim, 1828
    Carabus (Trachycarabus) sibiricus seriatoporus Fischer von Waldheim, 1828
    Male adult
    окр. Якутска, пос. Бестях на р. Лена, 30.VI.1962, Сладков leg.
    Copyright © Dmitri Obydov
    Carabus (Coptolabrus) kubani Deuve, 1990
    Carabus (Coptolabrus) kubani Deuve, 1990 Paratype
    Male adult
    Daju, (Yulongxue Shan, Yunnan) CHINE, VII-1990
    Copyright © Alain Mollard
    Carabus (Coptolabrus) ignimitella stillatus Cavazzuti & Ratti, 1998
    Carabus (Coptolabrus) ignimitella stillatus Cavazzuti & Ratti, 1998
    Male+Female adult
    Mt.Nanshan (Nanping, Fujian) CHINE VI-2001
    Copyright © Alain Mollard
    Carabus (Coptolabrus) ignimitella ignimitella Bates, 1888 (as nyukmingi Deuve, 1991)
    Carabus (Coptolabrus) ignimitella ignimitella Bates, 1888 (as nyukmingi Deuve, 1991)
    Male+Female adult
    Wuyi Shan (Fujian) CHINE VI-2001
    Copyright © Alain Mollard
    Carabus (Coptolabrus) ignimitella ignimitella Bates, 1888
    Carabus (Coptolabrus) ignimitella ignimitella Bates, 1888
    Male+Female adult
    Luozhou (Fuzhou, Fujian) CHINE VI-2001
    Copyright © Alain Mollard
    Carabus (Megodontus) vietinghoffi fulgidus Fischer, 1827 (as bureianus Shilenkov, 1996)
    Carabus (Megodontus) vietinghoffi fulgidus Fischer, 1827 (as bureianus Shilenkov, 1996)
    Male adult
    Far East, Khabarovsk reg., Verhnebureinskiy distr., Bureinskiy N.R., Lan riv. 10.6.2007 (DUBC) 20 mm
    Copyright © Alexander Anichtchenko
    Carabus (Aulonocarabus) canaliculatus sichotensis Born, 1914
    Carabus (Aulonocarabus) canaliculatus sichotensis Born, 1914
    Male adult
    Хабаровский край, Нанайский район, гол. Тардоки-Яни, 25-VI-1980, leg. В.Кузнецов. 20mm
    Copyright © - - -
    Carabus (Chrysocarabus) solieri liguranus Breuning, 1933
    Carabus (Chrysocarabus) solieri liguranus Breuning, 1933
    Male adult

    Copyright © Igor Solodovnikov
    Carabus (Chrysocarabus) olympiae Sella, 1855
    Carabus (Chrysocarabus) olympiae Sella, 1855
    Male adult
    Carabus (Chrysocarabus) olympiae n. fucatus Lapouge, 1902; un Italy: Piemont (TO), Bielmonte, h = 800 m.
    Copyright © Igor Solodovnikov
    Carabus (Megodontus) planicollis verae Csiki, 1905
    Carabus (Megodontus) planicollis verae Csiki, 1905
    Female adult
    C-Romania: Carpathians, E of Olt River: Fagaraş Mts, Balea Cascada, h = 1150 m
    Copyright © Igor Solodovnikov
    Carabus (Platycarabus) irregularis ramanus Sokolar, 1909
    Carabus (Platycarabus) irregularis ramanus Sokolar, 1909
    Female adult
    NW Croatia, Ivanščica Mts , Ivanec, 4-18.VI.2011, Krause Rene leg., L=33 mm
    Copyright © Ruslan Panin
    Carabus (Coptolabrus) pustulifer aeriviolaceus Cavazzuti & Rapuzzi, 2010
    Carabus (Coptolabrus) pustulifer aeriviolaceus Cavazzuti & Rapuzzi, 2010
    Male+Female adult
    Meigu (Xichang, Sichuan) CHINE, VII-2008
    Copyright © Alain Mollard
    Carabus (Coptolabrus) pustulifer elisabettianus Font, 1997
    Carabus (Coptolabrus) pustulifer elisabettianus Font, 1997
    Male+Female adult
    Huanjan Shan (Wuchuan, Guizhou) CHINE, VII-2008
    Copyright © Alain Mollard
    Carabus (Coptolabrus) pustulifer inusitatus Cavazzuti, 1997
    Carabus (Coptolabrus) pustulifer inusitatus Cavazzuti, 1997
    Male+Female adult
    Fanjing Shan (Jiangkou, Guizhou) CHINE, VI-2007
    Copyright © Alain Mollard
    Carabus (Coptolabrus) pustulifer katsumii Deuve & Ohshima, 1992
    Carabus (Coptolabrus) pustulifer katsumii Deuve & Ohshima, 1992 Paratype
    Female adult
    Wenxian (Gansu) CHINE, VII-2008
    Copyright © Alain Mollard
    Carabus (Coptolabrus) pustulifer mirandior Deuve, 1993
    Carabus (Coptolabrus) pustulifer mirandior Deuve, 1993
    Male+Female adult
    Dayao (Yunnan) CHINE, VII-2008
    Copyright © Alain Mollard
    Carabus (Coptolabrus) pustulifer mirificoides Deuve & Li, 2002
    Carabus (Coptolabrus) pustulifer mirificoides Deuve & Li, 2002
    Male+Female adult
    Yangtian (Gansu) CHINE, VII-2008
    Copyright © Alain Mollard
    Carabus (Coptolabrus) pustulifer qiannaicus Deuve & Tian, 2000
    Carabus (Coptolabrus) pustulifer qiannaicus Deuve & Tian, 2000
    Male+Female adult
    Longzjia (Maoping, Guizhou) CHINE, V-2008
    Copyright © Alain Mollard
    Carabus (Coptolabrus) pustulifer szechwanensis Hauser, 1931
    Carabus (Coptolabrus) pustulifer szechwanensis Hauser, 1931
    Male+Female adult
    Jinfo Shan (Chongqing) CHINE, V-2008
    Copyright © Alain Mollard
    Carabus (Coptolabrus) pustulifer wakoi Imura, 1995
    Carabus (Coptolabrus) pustulifer wakoi Imura, 1995
    Female adult
    Mts. Dabashan (Chengkou, Sichuan) CHINE, V-2008
    Copyright © Alain Mollard
    Carabus (Coptolabrus) pustulifer xianfengensis Deuve & Li, 2009
    Carabus (Coptolabrus) pustulifer xianfengensis Deuve & Li, 2009
    Male+Female adult
    Mt.Maoping (Zhaoshi, Hunan/Hubei) CHINE, V-2008
    Copyright © Alain Mollard
    Carabus (Coptolabrus) augustus chenzhouensis Deuve & Sehnal, 2010
    Carabus (Coptolabrus) augustus chenzhouensis Deuve & Sehnal, 2010
    Male+Female adult
    Yizhang Xian (Hunan) CHINE
    Copyright © Alain Mollard
    Carabus (Coptolabrus) augustus tongrenensis Deuve & Li, 2006
    Carabus (Coptolabrus) augustus tongrenensis Deuve & Li, 2006
    Male adult
    Tongren Xian (Jiangkou, Guizhou) CHINE
    Copyright © Alain Mollard
    Carabus (Coptolabrus) augustus haiyangicus Deuve, 2012
    Carabus (Coptolabrus) augustus haiyangicus Deuve, 2012
    Male+Female adult
    Xing’an (Guilin, Guangxi) CHINE
    Copyright © Alain Mollard
    Carabus (Coptolabrus) pustulifer adriaenssensi (as guizhouicus Deuve, 1994)
    Carabus (Coptolabrus) pustulifer adriaenssensi (as guizhouicus Deuve, 1994)
    Male+Female adult
    Leigong Shan (Kaili, Guizhou) CHINE
    Copyright © Alain Mollard
    Carabus (Coptolabrus) augustus mugepoensis Deuve & Li in Deuve, 2009
    Carabus (Coptolabrus) augustus mugepoensis Deuve & Li in Deuve, 2009
    Male+Female adult
    Mt.Gupo Shan (Zhongshan, Guangxi) CHINE
    Copyright © Alain Mollard
    Carabus (Coptolabrus) augustus divus Roeschke, 1908
    Carabus (Coptolabrus) augustus divus Roeschke, 1908
    Male+Female adult
    Pingxiang (Zhongshan, Jiangxi) CHINE
    Copyright © Alain Mollard
    Carabus (Coptolabrus) formosus formosus Semenov, 1887
    Carabus (Coptolabrus) formosus formosus Semenov, 1887
    Male+Female adult
    Wu-Chang-xi, (Gansu) CHINE VI-1998
    Copyright © Alain Mollard
    Carabus (Coptolabrus) formosus subformosus Semenov, 1887
    Carabus (Coptolabrus) formosus subformosus Semenov, 1887
    Male+Female adult
    Wenxian, (Gansu) CHINE VIII-1992
    Copyright © Alain Mollard
    Carabus (Coptolabrus) formosus brockmannioides Deuve & Murzin, 1997
    Carabus (Coptolabrus) formosus brockmannioides Deuve & Murzin, 1997 Paratype
    Male+Female adult
    Shihoshian, (Lixian, Gansu) CHINE VI-2003
    Copyright © Alain Mollard
    Carabus (Coptolabrus) formosus akane Imura & Yamaya, 1994
    Carabus (Coptolabrus) formosus akane Imura & Yamaya, 1994 Paratype
    Female adult
    Tongchuan (Xi’an, Shaanxi) CHINE VI-1993
    Copyright © Alain Mollard
    Carabus (Coptolabrus) formosus purpureodiscoides Kleinfeld, 1999
    Carabus (Coptolabrus) formosus purpureodiscoides Kleinfeld, 1999
    Male+Female adult
    Dalong Shan, Yunxian (Shiyan, Hubei) CHINE X-2010
    Copyright © Alain Mollard
    Carabus (Coptolabrus) elysii xiaoganicus Deuve et Remond, 2016
    Carabus (Coptolabrus) elysii xiaoganicus Deuve et Remond, 2016
    Male+Female adult
    Shuangfeng Shan (Xiaogan, Hubei) CHINE VI-2016
    Copyright © Alain Mollard
    Carabus (Coptolabrus) elysii xiaoganicus Deuve et Remond, 2016
    Carabus (Coptolabrus) elysii xiaoganicus Deuve et Remond, 2016
    Male+Female adult
    Shuangfeng Shan (Xiaogan, Hubei) CHINE VI-2016
    Copyright © Alain Mollard
    Carabus (Coptolabrus) elysii chikongschanensis Hauser, 1914
    Carabus (Coptolabrus) elysii chikongschanensis Hauser, 1914
    Male+Female adult
    Zhuguang, (Hunan) CHINE V-1991
    Copyright © Alain Mollard
    Carabus (Coptolabrus) elysii amucronatus Deuve, 2010
    Carabus (Coptolabrus) elysii amucronatus Deuve, 2010
    Male adult
    Zhongtiao Shan (Shanxi) CHINE VII-2002
    Copyright © Alain Mollard
    Carabus (Coptolabrus) elysii viridicyaneus Lassalle, 2016
    Carabus (Coptolabrus) elysii viridicyaneus Lassalle, 2016 Paratype
    Male adult

    Copyright © Bernard Lassalle
    Carabus (Tribax) apschuanus galianus Belousov & Zamotajlov, 1993
    Carabus (Tribax) apschuanus galianus Belousov & Zamotajlov, 1993
    Male adult

    Copyright © Igor Solodovnikov
    Carabus (Tribax) apschuanus galianus Belousov & Zamotajlov, 1993
    Carabus (Tribax) apschuanus galianus Belousov & Zamotajlov, 1993
    Endophallus

    Copyright © Igor Solodovnikov
    Carabus (Tribax) apschuanus galianus Belousov & Zamotajlov, 1993
    Carabus (Tribax) apschuanus galianus Belousov & Zamotajlov, 1993
    Female adult

    Copyright © Igor Solodovnikov
    Carabus (Coptolabrus) smaragdinus smaragdinus Fischer, 1823
    Carabus (Coptolabrus) smaragdinus smaragdinus Fischer, 1823
    Male+Female adult
    Mt. Nerchinski (Tchita, Sibérie) RUSSIE VII-1995
    Copyright © Alain Mollard
    Carabus (Coptolabrus) smaragdinus lengxuemeiae Deuve & Li, 2000
    Carabus (Coptolabrus) smaragdinus lengxuemeiae Deuve & Li, 2000
    Male+Female adult
    Lujiaogou Shan (Kuandian, Liaoning) CHINE VI-1997
    Copyright © Alain Mollard
    Carabus (Coptolabrus) jankowskii chindoensis Kwon & Park, 1989
    Carabus (Coptolabrus) jankowskii chindoensis Kwon & Park, 1989
    Male+Female adult
    Chongue san (Chindo Isle, ) KOREE VII-1992
    Copyright © Alain Mollard
    Carabus (Coptolabrus) jankowskii quelpartianus Breuning, 1932
    Carabus (Coptolabrus) jankowskii quelpartianus Breuning, 1932
    Male+Female adult
    Hallasam, (Cheju-do Isle, ) KOREE VII-1990
    Copyright © Alain Mollard
    Carabus (Coptolabrus) lafossei tungchengensis Li, 1993
    Carabus (Coptolabrus) lafossei tungchengensis Li, 1993
    Male+Female adult
    Mt.Huang-Bai-Shan (Shangcheng, Henan) VI-2012
    Copyright © Alain Mollard
    Carabus (Tribax) edmundi Semenov, 1896
    Carabus (Tribax) edmundi Semenov, 1896
    Female adult
    Russia, S-Dagestan, 18.IV-19.VI.2022 leg. D. Fominykh
    Copyright © Igor Solodovnikov
    Carabus (Tribax) edmundi Semenov, 1896
    Carabus (Tribax) edmundi Semenov, 1896
    Endophallus
    Russia, S-Dagestan
    Copyright © Igor Solodovnikov
    Carabus (Tribax) edmundi Semenov, 1896
    Carabus (Tribax) edmundi Semenov, 1896
    Male adult
    Russia, S-Dagestan, 18.IV-19.VI.2022 leg. D. Fominykh
    Copyright © Igor Solodovnikov
    Carabus (Tribax) osseticus abanosicus Deuve, 2013
    Carabus (Tribax) osseticus abanosicus Deuve, 2013
    Endophallus

    Copyright © Igor Solodovnikov
    Carabus (Tribax) osseticus abanosicus Deuve, 2013
    Carabus (Tribax) osseticus abanosicus Deuve, 2013
    Male adult

    Copyright © Igor Solodovnikov
    Carabus (Tribax) osseticus osseticus Adams, 1817
    Carabus (Tribax) osseticus osseticus Adams, 1817
    Male adult

    Copyright © Igor Solodovnikov
    Carabus (Tribax) osseticus osseticus Adams, 1817
    Carabus (Tribax) osseticus osseticus Adams, 1817
    Female adult

    Copyright © Igor Solodovnikov
    Carabus (Tribax) osseticus osseticus Adams, 1817
    Carabus (Tribax) osseticus osseticus Adams, 1817
    Endophallus

    Copyright © Igor Solodovnikov
    Carabus (Tribax) osseticus abanosicus Deuve, 2013
    Carabus (Tribax) osseticus abanosicus Deuve, 2013
    Female adult

    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) jason Semenov, 1898
    Carabus (Neoplectes) jason Semenov, 1898
    Male adult
    Abhazia, Tchkhaltinskiy (Abkhazskiy) Mt.R., 20-26.VII.2022 leg. D. Fominykh
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) jason Semenov, 1898
    Carabus (Neoplectes) jason Semenov, 1898
    Endophallus

    Copyright © Igor Solodovnikov
    Carabus (Leptocarabus) vogtianus horvatovichi Deuve, 1992
    Carabus (Leptocarabus) vogtianus horvatovichi Deuve, 1992
    Male adult

    Copyright © Jong-Bong Choi
    Carabus (Carabus) granulatus granulatus Linne, 1758
    Carabus (Carabus) granulatus granulatus Linne, 1758
    Male adult
    ugly specimens
    Copyright © Igor Solodovnikov
    Carabus (Tachypus) cancellatus cancellatus Illiger, 1798
    Carabus (Tachypus) cancellatus cancellatus Illiger, 1798
    Male adult
    Carabus (Tachypus) cancellatus tuberculatus f.conspersus Lapouge, 1902 (Blaс form) ugly specimens N-Belarus, Vitebsk reg., Senno distr., 37 km SSE Vitebsk, near Lake Streshno, 01-10.VII.2006, I.A. Solodovnikov
    Copyright © Igor Solodovnikov
    Carabus (Tachypus) cancellatus cancellatus Illiger, 1798
    Carabus (Tachypus) cancellatus cancellatus Illiger, 1798
    Endophallus
    Carabus (Tachypus) cancellatus tuberculatus f.conspersus Lapouge, 1902 (Blaс form) N-Belarus, Mogilev reg., Bykhov distr., 5 km N Voronino Vill., mixed-pine forest, 20.V-03.VI.2007, Komyagunova N.P.
    Copyright © Igor Solodovnikov
    Carabus (Tachypus) cancellatus cancellatus Illiger, 1798
    Carabus (Tachypus) cancellatus cancellatus Illiger, 1798
    Male adult
    Carabus (Tachypus) cancellatus tuberculatus f.conspersus Lapouge, 1902 (Blaс form) E-Belarus, Mogilev reg., Bykhov distr., 5 km N Voronino Vill., mixed-pine forest, 20.V-03.VI.2007, Komyagunova N.P.
    Copyright © Igor Solodovnikov
    Carabus (Tachypus) cancellatus cancellatus Illiger, 1798
    Carabus (Tachypus) cancellatus cancellatus Illiger, 1798
    Male adult
    Carabus (Tachypus) cancellatus tuberculatus f. tenebrosus Ball. (Blue form) S-Belarus, Brest reg., env. Tomashovka, pine forest, 26.VI-13.VII.2022, Kuznetzov V.A.
    Copyright © Igor Solodovnikov
    Carabus (Tachypus) cancellatus cancellatus Illiger, 1798
    Carabus (Tachypus) cancellatus cancellatus Illiger, 1798
    Female adult
    Carabus (Tachypus) cancellatus tuberculatus f.conspersus Lapouge, 1902 (Blaс form) E-Belarus, Mogilev reg., Bykhov distr., 5 km N Voronino Vill., mixed-pine forest, 20.V-03.VI.2007, Komyagunova N.P.
    Copyright © Igor Solodovnikov
    Carabus (Tachypus) cancellatus cancellatus Illiger, 1798
    Carabus (Tachypus) cancellatus cancellatus Illiger, 1798
    Female adult
    Carabus (Tachypus) cancellatus tuberculatus f. tenebrosus Ball. (Blue form) S-Belarus, Brest reg., env. Tomashovka, pine forest, 26.VI-13.VII.2022, Kuznetzov V.A.
    Copyright © Igor Solodovnikov
    Carabus (Tachypus) cancellatus cancellatus Illiger, 1798
    Carabus (Tachypus) cancellatus cancellatus Illiger, 1798
    Female adult
    Carabus (Tachypus) cancellatus tuberculatus f. tenebrosus Ball. (Blue form) S-Belarus, Brest reg., env. Tomashovka, pine forest, 26.VI-13.VII.2022, Kuznetzov V.A.
    Copyright © Igor Solodovnikov
    Carabus (Tomocarabus) decolor Fischer, 1823
    Carabus (Tomocarabus) decolor Fischer, 1823
    Male adult

    Copyright © Kirill Makarov
    Carabus (Carabus) granulatus granulatus Linne, 1758
    Carabus (Carabus) granulatus granulatus Linne, 1758
    Male adult
    Carabus (s.str.) granulatus granulatus ab. wimmeli Schulz, 1900 S-Belarus, Brest reg., env. Tomashovka, pine forest, 26.VI-13.VII.2022, Kuznetzov V.A.
    Copyright © Igor Solodovnikov
    Carabus (Tachypus) cancellatus cancellatus Illiger, 1798
    Carabus (Tachypus) cancellatus cancellatus Illiger, 1798
    Male adult
    Carabus (s.str.) cancellatus cancellatus f.balticus Bernau, 1915 N-Belarus, Vitebsk reg. 21 mm
    Copyright © Igor Solodovnikov
    Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
    Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
    Male adult
    Carabus (s.str.) cancellatus emarginatus n.monzinii Monzini, 1983 25,5 mm
    Copyright © Igor Solodovnikov
    Carabus (Sphodristocarabus) kurdicus kurdicus Heinz, 1975
    Carabus (Sphodristocarabus) kurdicus kurdicus Heinz, 1975
    Female adult

    Copyright © Igor Solodovnikov
    Carabus (Sphodristocarabus) varians janthinus Ganglbauer, 1887
    Carabus (Sphodristocarabus) varians janthinus Ganglbauer, 1887
    Female adult
    Carabus (Sphrodistocarabus) janthinus dvorschaki Mandl, 1975 Abkhazia, Gudautsky district, S- Bzybsky Ridge, a watershed hill at the head of the river Khipsta, 43°17'56.63" N, 40°40'16.74" E, h= 1490 m, mature beech-maple forest with butterbur, moist soil, 05.07-17.08.2017, leg. I.A. Solodovnikov, S.V. Solodovnikova, E.V. Tatun
    Copyright © Igor Solodovnikov
    Carabus (Sphodristocarabus) scovitzi scovitzi Faldermann, 1835
    Carabus (Sphodristocarabus) scovitzi scovitzi Faldermann, 1835
    Endophallus
    Georgia, Adzharia, Adzharo-Imeretinskiy Mt.R., near Tchidila pass, alpin zone, h=2350 m, 05.VI.2016 leg. D. Fominykh
    Copyright © Igor Solodovnikov
    Carabus (Sphodristocarabus) scovitzi scovitzi Faldermann, 1835
    Carabus (Sphodristocarabus) scovitzi scovitzi Faldermann, 1835
    Male adult
    Georgia, Adzharia, Adzharo-Imeretinskiy Mt.R., near Tchidila pass, alpin zone, h=2350 m, 05.VI.2016 leg. D. Fominykh
    Copyright © Igor Solodovnikov
    Carabus (Sphodristocarabus) tokatensis kosensis Cavazzuti, 1984
    Carabus (Sphodristocarabus) tokatensis kosensis Cavazzuti, 1984
    Endophallus
    CE-Turkey: Gümüshane: Köse Dag(pass N of Köse), h = 1700 m, VII.2000, leg. St. Il. Remond (TOPOTYPUS)
    Copyright © Igor Solodovnikov
    Carabus (Sphodristocarabus) tokatensis kosensis Cavazzuti, 1984
    Carabus (Sphodristocarabus) tokatensis kosensis Cavazzuti, 1984
    Male adult
    CE-Turkey: Gümüshane: Köse Dag(pass N of Köse), h = 1700 m, VII.2000, leg. St. Il. Remond (TOPOTYPUS)
    Copyright © Igor Solodovnikov
    Carabus (Trachycarabus) planus retowskianus Mandl, 1955
    Carabus (Trachycarabus) planus retowskianus Mandl, 1955
    Male adult
    S Ukraine, Zaporizhia reg., Melitopol distr., near Troitskoe vill., 1.VI.2010, I.V. Kovalev leg., L=27 mm
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) estreicheri Fischer, 1822 (ab.rufofemoratus Lomnicki, 1892)
    Carabus (Trachycarabus) estreicheri Fischer, 1822 (ab.rufofemoratus Lomnicki, 1892)
    Female adult
    W Ukraina, Podolian Upland, Lviv reg., Brody distr., Mt. Makitra, steppe site, 19.V-5.VI.2020, Panin R. leg. L=23 mm
    Copyright © Ruslan Panin
    Carabus (Carabus) pseudolatipennis nanpingensis Deuve, 1992
    Carabus (Carabus) pseudolatipennis nanpingensis Deuve, 1992 Holotype
    Female adult
    N-Sichuan: Nanping Xian, 2600m
    Copyright © Thierry Deuve
    Carabus (Carabus) latipennis latipennis Breuning, 1932
    Carabus (Carabus) latipennis latipennis Breuning, 1932 Holotype
    Female adult

    Copyright © Thierry Deuve
    Carabus (Pseudocranion) gansuensis tiro Semenov, 1898
    Carabus (Pseudocranion) gansuensis tiro Semenov, 1898 Lectotype
    Male adult
    Sun-pan
    Copyright © Thierry Deuve
    Carabus (Pseudocranion) gansuensis altiphilus Deuve, 2010
    Carabus (Pseudocranion) gansuensis altiphilus Deuve, 2010 Holotype
    Male adult
    Chine, Nord Sichuan, alentours meridionaux du col situe a 12 km a l'est-sud-est de Zhangla (pelouses alpines, eboulis) 32°43N 103°45E (in coll. J. Kalab)
    Copyright © Thierry Deuve
    Carabus (Trachycarabus) planus campicola Dejean, 1831
    Carabus (Trachycarabus) planus campicola Dejean, 1831
    Female adult
    S Ukraine, Crimea, S slope Mt Demerdzhi h=650 m, sibljak, VII.2004, R. Panin leg., L=29 mm
    Copyright © Ruslan Panin
    Carabus (Pachystus) glabratus gibbosus Heyden, 1866
    Carabus (Pachystus) glabratus gibbosus Heyden, 1866
    Male adult
    N Croatia, Ivanščica Mts, Ivanec, 4-18.VI.2011, Krause Rene leg., L=32 mm
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) yunanensis huizeanus Cavazzuti, 2009
    Carabus (Apotomopterus) yunanensis huizeanus Cavazzuti, 2009 Holotype
    Male adult
    Mt. SE di Huize, 2400-2800 m. N. Yunnan, Cina meridionale.
    Copyright © - - -
    Carabus (Neoplesius) piceophilus zheduo Cavazzuti, 2009
    Carabus (Neoplesius) piceophilus zheduo Cavazzuti, 2009 Holotype
    Male adult
    Passo a Ovest di Kangding, Mass. Zheduo Shan, 30°04'28"N, 101°48'16"E, 4400-4550 m, Sichuan occ., Cina.
    Copyright © - - -
    Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
    Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
    Endophallus
    Carabus (Tachypus) cancellatus emarginatus n. dahli Heer, 1841
    Copyright © Igor Solodovnikov
    Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
    Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
    Male adult
    Carabus (Tachypus) cancellatus emarginatus n. dahli Heer, 1841
    Copyright © Igor Solodovnikov
    Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
    Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
    Female adult
    Carabus (Tachypus) cancellatus emarginatus n. dahli Heer, 1841
    Copyright © Igor Solodovnikov
    Carabus (Morphocarabus) spasskianus cf putus Motschulsky, 1846
    Carabus (Morphocarabus) spasskianus cf putus Motschulsky, 1846
    Female adult
    Russia, SW Siberia, C-Altays Mts, Kuray Mt Rng NE Aktash vill., h=2100-3000 m, 8-13.VII.2022, E.Filippov leg./det.R Panin, L= 22 mm (alpine petrophilic population close to the form schestopalovi Plutenko, 1995)
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) spasskianus cf putus Motschulsky, 1846
    Carabus (Morphocarabus) spasskianus cf putus Motschulsky, 1846
    Male adult
    Russia, SW Siberia, C-Altays Mts, Kuray Mt Rng NE Aktash vill., h=2100-3000 m, 8-13.VII.2022, E.Filippov leg./det.R Panin, L= 23 mm (alpine petrophilic population close to the form schestopalovi Plutenko, 1995)
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) spasskianus putus Motschulsky, 1846 (loc.typ.)
    Carabus (Morphocarabus) spasskianus putus Motschulsky, 1846 (loc.typ.)
    Female adult
    Russia, W Altay, 18 km N of Zmeinogorsk, 23.VI.1998, A.& N. Menstschikovy leg. L=24 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) spasskianus putus Motschulsky, 1846 (loc.typ.)
    Carabus (Morphocarabus) spasskianus putus Motschulsky, 1846 (loc.typ.)
    Male adult
    Russia, W Altay, 18 km N of Zmeinogorsk, 23.VI.1998, A.& N. Menstschikovy leg. L=23 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) odoratus antropovi Shilenkov 1996
    Carabus (Morphocarabus) odoratus antropovi Shilenkov 1996
    Female adult
    Russia, S Siberia, Tuva, W Tannu-Ola Mts. 35 km NE Khandagayty vill., Teeli Pass 2200-2800 m, 23-25.VI.2021, A.& E.Filippovs leg. L=19 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) odoratus antropovi Shilenkov 1996
    Carabus (Morphocarabus) odoratus antropovi Shilenkov 1996
    Endophallus
    Russia, S Siberia, Tuva, W Tannu-Ola Mts. 35 km NE Khandagayty vill., Teeli Pass 2200-2800 m
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) odoratus antropovi Shilenkov 1996
    Carabus (Morphocarabus) odoratus antropovi Shilenkov 1996
    Male adult
    Russia, S Siberia, Tuva, W Tannu-Ola Mts. 35 km NE Khandagayty vill., Teeli Pass 2200-2800 m, 23-25.VI.2021, A.& E.Filippovs leg. L=19 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
    Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
    Endophallus
    SE Poland, Subcarpathia Prov., Nadleśnictwo Kołaczyce N of Jasło vill., M. Kosibowicz leg./det. R Panin
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
    Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
    Male adult
    SE Poland, Subcarpathia Prov., Nadleśnictwo Kołaczyce N of Jasło vill. M. Kosibowicz leg./det. R Panin, VII.2016, L=29 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
    Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
    Male adult
    SE Poland, N Carpathians, Subcarpathia Prov., Nadleśnictwo Baligród, (209a) 22.VII.2021, M.Kosibowicz leg./det. R.Panin, L=31 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
    Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
    Male adult
    SE Poland, N Carpathians, Subcarpathia Prov., National Park Magura, VII.2016, M.Ranocha leg./ det. R. Panin, L=30 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
    Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
    Male adult
    SE Poland, N Carpathians, Subcarpathia Prov., Nadleśnictwo Lesko, VII.2017 M.Ranocha leg./ det. R. Panin, L=31 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
    Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
    Endophallus
    Russia, NE Altay, Turochak Distr., Lake Teletskoe N part, near Yaylyu vill.
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
    Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
    Male adult
    Russia, NE Altay, Turochak Distr., Lake Teletskoe N part, near Yaylyu vill. 23-25.VII.2019, Ivliev P. leg./ det. R. Panin, L=24 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) aeruginosus eschscholtzi Mannerheim, 1827
    Carabus (Morphocarabus) aeruginosus eschscholtzi Mannerheim, 1827
    Female adult
    NE Kazakhstan, W Altay Mts, near Shemonaikha city., flood plain, 25-28.VII.2018, S.Vashchenko leg. L=25 mm
    Copyright © Ruslan Panin
    Carabus (Tribax) zubovi mananauriensis Deuve, 2022
    Carabus (Tribax) zubovi mananauriensis Deuve, 2022 Holotype
    Male adult
    Géorgie, Svanétie, 20 km à l’ouest de Lentekhi, 6 km à l’ouest de Mananaouri, 1350 m (A. Wřzecionko leg.), in coll. A. Wřzecionko, à Horni Sucha (République Tchèque).
    Copyright © Thierry Deuve
    Carabus (Rhigocarabus) indigestus dongziensis Deuve & Kaláb, 2022
    Carabus (Rhigocarabus) indigestus dongziensis Deuve & Kaláb, 2022 Holotype
    Male adult
    Chine, Sud-Sichuan, montagnes à 10 km au nordouest de Dongzi, 4500 m, 28°34’N/100°46’E, pelouses alpines et éboulis, in coll. J. Kaláb, à Jinačovice (République Tchèque).
    Copyright © Thierry Deuve
    Carabus (Rhigocarabus) ladygini shiliensis Deuve et Müller, 2022
    Carabus (Rhigocarabus) ladygini shiliensis Deuve et Müller, 2022 Holotype
    Male adult
    Chine, Sichuan nord-occidental, environs de Shili, Zango (31,85°N-101,05°E), 4000 m (V. Benes leg.), in coll. A. Müller, Vienne (Autriche).
    Copyright © Thierry Deuve
    Carabus (Morphocarabus) aeruginosus eschscholtzi Mannerheim, 1827
    Carabus (Morphocarabus) aeruginosus eschscholtzi Mannerheim, 1827
    Endophallus
    NE Kazakhstan, W Altay Mts, near Shemonaikha city., flood plain,
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) aeruginosus eschscholtzi Mannerheim, 1827
    Carabus (Morphocarabus) aeruginosus eschscholtzi Mannerheim, 1827
    Male adult
    NE Kazakhstan, W Altay Mts, near Shemonaikha city., flood plain, 25-28.VII.2018, S.Vashchenko leg., L=22.5 mm
    Copyright © Ruslan Panin
    Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792
    Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792
    Male adult
    C-Poland, Łódź sity, Łagiewniki Forest, in dead logs of Birch, 18-30.XII.2013, D. Nawak leg., L=25 mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) leachi leachi Fischer, 1823
    Carabus (Megodontus) leachi leachi Fischer, 1823
    Female adult
    NE Kazakhstan, W Altay, near Shemonaikha city., flood plain, 18-28.VII.2018, S.Vashchenko leg., L=22.5 mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) leachi leachi Fischer, 1823
    Carabus (Megodontus) leachi leachi Fischer, 1823
    Male adult
    NE Kazakhstan, W Altay, near Shemonaikha city., flood plain, 18-28.VII.2018, S.Vashchenko leg., L=21.5 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
    Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
    Male adult
    Russia, W Sayan Mts., Kazyr-Suk river, W slope of Oyskiy Mt. rng., ?h=2200 m 7-15.VII.1999, Brinev A. leg., L=19 mm
    Copyright © Ruslan Panin
    Carabus (Morphocarabus) monilis tanarguensis Maguerre & Stefani, 2021 (syn.)
    Carabus (Morphocarabus) monilis tanarguensis Maguerre & Stefani, 2021 (syn.) Holotype Paratype
    Male+Female adult
    Holotype, m (21 mm), environs du Col de la Croix de Bauzon, 1300 m, communes de La Souche, Borne et Mayres, Ardèche, France (07/2003), in coll. D. Maguerre, à Mons-en-Barœul, France. Paratype f (23 mm), au dessus du Col de Meyrand, 1400 mètres, communes de Loubaresse et de Borne, Ardèche, France, (07/2004), in coll. P. Stéfani.
    Copyright © - - -
    Carabus (Meganebrius) thudamensis Deuve & Schmidt, 2017
    Carabus (Meganebrius) thudamensis Deuve & Schmidt, 2017 Holotype
    Male adult
    Est-Népal, District de Sankhuwasabha, Thudam, 3500 – 3600 m, 27°45’23’’N – 87°32’56’’E (S. Tamang leg., 1-vii-2011), in coll. J. Schmidt (Admannshagen, Allemagne).
    Copyright © Joachim Schmidt
    Carabus (Meganebrius) santostamangi thanglaensis Deuve & Schmidt, 2017
    Carabus (Meganebrius) santostamangi thanglaensis Deuve & Schmidt, 2017 Holotype
    Male adult
    Est-Népal, limite des districts de Sankhuwasabha et de Taplejung, Jaljale Himal, Thangla Phedi, 4 150 m, 27°39’09’’ N – 87°35’56’’ E (S. Tamang leg., 27-vi-2011), in coll. J. Schmidt (Admannshagen, Allemagne).
    Copyright © Joachim Schmidt
    Carabus (Meganebrius) santostamangi santostamangi
    Carabus (Meganebrius) santostamangi santostamangi Holotype
    Male adult
    Est-Népal, district de Taplejung, versant oriental de la Palung Khola, 3 720 m, 27° 35’ 43’’ N – 87° 41’ 50’’ E (S. Tamang leg., 7-vii-2011), in coll. J. Schmidt (Admannshagen, Allemagne).
    Copyright © Joachim Schmidt
    Carabus (Meganebrius) santostamangi santostamangi
    Carabus (Meganebrius) santostamangi santostamangi
    Endophallus

    Copyright © Joachim Schmidt
    Carabus (Meganebrius) thudamensis Deuve & Schmidt, 2017
    Carabus (Meganebrius) thudamensis Deuve & Schmidt, 2017 Holotype
    Endophallus

    Copyright © Joachim Schmidt
    Carabus (Meganebrius) everesti everesti Andrewes, 1929
    Carabus (Meganebrius) everesti everesti Andrewes, 1929
    Endophallus
    Kemba Kharka
    Copyright © Joachim Schmidt
    Carabus (Meganebrius) salpansis Deuve, 1984
    Carabus (Meganebrius) salpansis Deuve, 1984
    Endophallus
    Kemba Kharka
    Copyright © Joachim Schmidt
    Carabus (Meganebrius) indicus indicus Fairmaire, 1889
    Carabus (Meganebrius) indicus indicus Fairmaire, 1889
    Endophallus
    Tonglu
    Copyright © Joachim Schmidt
    Carabus (Carabus) baogai subbaogai Deuve, 2013
    Carabus (Carabus) baogai subbaogai Deuve, 2013 Holotype
    Female adult
    Chine, Hunan, Nanshan, 1750-1850, 26°10'N/ 10°09'E, in coll. Jaroslav Tuma, Kostelec na Hane (République Tchèque).
    Copyright © Thierry Deuve
    Carabus (Eupachys) acutithorax Deuve, 1989
    Carabus (Eupachys) acutithorax Deuve, 1989
    Male adult
    China, Shanxi, Yongzhi [Yongji]
    Copyright © Thierry Deuve
    Carabus (Rhigocarabus) casanova Imura & Brezina, 2008
    Carabus (Rhigocarabus) casanova Imura & Brezina, 2008 Holotype Paratype
    Male+Female adult
    north-northwest of Kaba, 34°11'09-14"N I 103°23' 10-21 "E, 3,800-4,100 m in altitude, on the Die Shan Mountain Range in Tewo Xian, of Gannan Zangzu Zizhizhou, in southern Gansu, China, 14-VI-2006. HT to be deposited in the Department of Zoology, National Museum of Nature and Science, Tokyo
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) casanova (Imura & Brezina, 2008)
    Carabus (Rhigocarabus) casanova (Imura & Brezina, 2008)
    Endophallus

    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) gigolo Heinz & Brezina, 1996
    Carabus (Rhigocarabus) gigolo Heinz & Brezina, 1996
    Endophallus
    Langmusi
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) gentleman Brezina & Hackel, 2004
    Carabus (Rhigocarabus) gentleman Brezina & Hackel, 2004
    Endophallus
    Jiangjing
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) gigoloides priapeius Cavazzuti, 2006
    Carabus (Rhigocarabus) gigoloides priapeius Cavazzuti, 2006
    Endophallus
    Yeliguan
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) gigoloides kabaki Imura, 2007
    Carabus (Rhigocarabus) gigoloides kabaki Imura, 2007
    Male adult
    S of Wushan
    Copyright © Boleslav Brezina
    Carabus (Rhigocarabus) nianjuaensis Imura, 2016
    Carabus (Rhigocarabus) nianjuaensis Imura, 2016 Holotype Paratype
    Male adult
    China, NW. Yunnan, NE. of Weixi-lisuzu Zizhixian ［维西傈僳族自治县］, 8.65 km west of Laoantong ［老安 统］, 7.35 km east of Mt. Nianjua ［念菊阿］, 27°44ʼ41” N / 98°56ʼ58” E, 3950 m in altitude, 16-VI-2015, to be preserved in the Zoological Institute of Academy of Sciences, St. Petersburg, Russia. Paratype: 1♂, same data as for the holotype, preserved in the collections of Y. Imura (Yokohama). Image from Imura, 2016
    Copyright © - - -
    Carabus (Rhigocarabus) nianjuaensis Imura, 2016
    Carabus (Rhigocarabus) nianjuaensis Imura, 2016
    Endophallus
    Image from Imura, 2016
    Copyright © - - -
    Carabus (Rhigocarabus) kozaburoi Imura, 1999
    Carabus (Rhigocarabus) kozaburoi Imura, 1999 Paratype
    Endophallus
    Gaoligong Shan, N. Myanmar. Image from Imura, 2016
    Copyright © - - -
    Carabus (Rhigocarabus) kozaburoi Imura, 1999
    Carabus (Rhigocarabus) kozaburoi Imura, 1999 Paratype
    Male adult
    Gaoligong Shan, N. Myanmar. Image from Imura, 2016
    Copyright © - - -
    Carabus (Rhigocarabus) itzingeri rugulosior Deuve, 1996
    Carabus (Rhigocarabus) itzingeri rugulosior Deuve, 1996 Holotype
    Endophallus
    NW-Yunnan: Hengduan Shan: Yanmen env.: 28:06N/98:52E. Image from Imura, 2016
    Copyright © - - -
    Carabus (Rhigocarabus) itzingeri rugulosior Deuve, 1996
    Carabus (Rhigocarabus) itzingeri rugulosior Deuve, 1996 Holotype
    Male adult
    NW-Yunnan: Hengduan Shan: Yanmen env.: 28:06N/98:52E. Image from Imura, 2016
    Copyright © - - -
    Carabus (Rhigocarabus) itzingeri Breuning, 1934
    Carabus (Rhigocarabus) itzingeri Breuning, 1934 Holotype
    Female adult
    Batang, W. Schuan. Image from Imura, 2016
    Copyright © - - -
    Carabus (Rhigocarabus) itzingeri tanakai Imura, 1997
    Carabus (Rhigocarabus) itzingeri tanakai Imura, 1997
    Male adult
    NE. Putao, N. Myanmar. Image from Imura, 2016
    Copyright © - - -
    Carabus (Rhigocarabus) itzingeri choguy Imura, 2001
    Carabus (Rhigocarabus) itzingeri choguy Imura, 2001 Holotype
    Male adult
    Chayu, E. Xizang. Image from Imura, 2016
    Copyright © - - -
    Carabus (Rhigocarabus) itzingeri tanakai Imura, 1997
    Carabus (Rhigocarabus) itzingeri tanakai Imura, 1997
    Endophallus
    Image from Imura, 2016
    Copyright © - - -
    Carabus (Rhigocarabus) itzingeri choguy Imura, 2001
    Carabus (Rhigocarabus) itzingeri choguy Imura, 2001 Holotype
    Endophallus
    Image from Imura, 2016
    Copyright © - - -
    Carabus (Rhigocarabus) rhododendron Deuve & Imura, 1991
    Carabus (Rhigocarabus) rhododendron Deuve & Imura, 1991 Holotype
    Male adult
    Mt. Bamaxue Shan, NW. Yunnan. Image from Imura, 2016
    Copyright © - - -
    Carabus (Rhigocarabus) rhododendron lidimaensis Imura, 2016
    Carabus (Rhigocarabus) rhododendron lidimaensis Imura, 2016 Holotype
    Male adult
    China, NW. Yunnan, near the borders between Weixi-lisuzu Zizhixian ［维西傈僳族自治县］ and Deqin Xian ［德钦县］, 4.3 km east of Lidima [里底 玛], 27°40ʼ43” N / 99 °19ʼ20” E, 4110 m in altitude, 23-VI-2015, to be preserved in the Zoological Institute of Academy of Sciences, St. Petersburg, Russia. Image from Imura, 2016
    Copyright © - - -
    Carabus (Rhigocarabus) rhododendron xuebangshanensis Imura, 2016
    Carabus (Rhigocarabus) rhododendron xuebangshanensis Imura, 2016 Holotype
    Male adult
    China, NW. Yunnan, near the borders between Lanping-baizu-pumizu Zizhixian [兰坪白族普米族自治县] and Jianchuan Xian [剑川县], NE. of Lanping [兰 坪], 0.95 km NNE. of Mt. Xuebang Shan [雪 邦 山], 26 °29ʼ14” N / 99 °30ʼ08” E, 4035 m in altitude, 29-V-2015, to be preserved in the Zoological Institute of Academy of Sciences, St. Petersburg, Russia. Image from Imura, 2016
    Copyright © - - -
    Carabus (Rhigocarabus) rhododendron xinshanchangensis Imura, 2016
    Carabus (Rhigocarabus) rhododendron xinshanchangensis Imura, 2016 Holotype
    Male adult
    China, NW. Yunnan, Lanping-baizu-pumizu Zizhixian [兰坪白族普米族自治县], ca. 10km NNE. of Lanping [兰坪], SE. of Xinshanchang [新山 场], 26 °30ʼ37” N / 99 °29ʼ26” E, 3780 m in altitude, 19-VI-2014, to be preserved in the Zoological Institute of Academy of Sciences, St. Petersburg, Russia. Image from Imura, 2016
    Copyright © - - -
    Carabus (Rhigocarabus) rhododendron xuebangshanensis Imura, 2016
    Carabus (Rhigocarabus) rhododendron xuebangshanensis Imura, 2016 Holotype
    Endophallus
    Image from Imura 2016
    Copyright © - - -
    Carabus (Rhigocarabus) rhododendron xinshanchangensis Imura, 2016
    Carabus (Rhigocarabus) rhododendron xinshanchangensis Imura, 2016 Holotype
    Endophallus
    Image from Imura, 2016
    Copyright © - - -
    Carabus (Rhigocarabus) rhododendron lidimaensis Imura, 2016
    Carabus (Rhigocarabus) rhododendron lidimaensis Imura, 2016 Holotype
    Endophallus
    Image from Imura, 2016
    Copyright © - - -
    Carabus (Rhigocarabus) rhododendron s.str.
    Carabus (Rhigocarabus) rhododendron s.str. Holotype
    Endophallus
    Image from Imura, 2016
    Copyright © - - -
    Carabus (Aristocarabus) romanowi sandaguensis Imura, Zhou & Su, 2004
    Carabus (Aristocarabus) romanowi sandaguensis Imura, Zhou & Su, 2004 Holotype
    Male adult
    mixed forest, 2,875 m in altitude, Sandagu in northwestern Heishui Xian of northern Siclman, Southwest China, 24-26-VI I-200 1, XiaoDong Yu & Hong-Zhang ZHOU leg., in col l. Institute of Zoology, Chinese Academy of Sciences, Beijing. Image from Imura, Zhou & Su, 2004
    Copyright © - - -
    Carabus (Megodontoides) promachus wujiapeng (Imura, Zhou & Su, 2004)
    Carabus (Megodontoides) promachus wujiapeng (Imura, Zhou & Su, 2004) Holotype
    Male adult
    mixed forest, 2,320 m in altitude, Wujiapeng in southwestrn Danba Xian of west-central Siclman, Southwest China, 1 9-2 1 -VII.2001, Xiao-Dong Yu & Hong-Zhang ZHOU leg. Image from (Imura, Zhou & Su, 2004)
    Copyright © - - -
    Carabus (Cupreocarabus) sinotibeticola feicuipennis (Imura, Zhou & Su, 2004)
    Carabus (Cupreocarabus) sinotibeticola feicuipennis (Imura, Zhou & Su, 2004) Holotype
    Male adult
    shrubs of Rhododendron spp., 4, 1 35 m in altitude, in southwestern Jiulong Xian of western Sichuan, Southwest China, 1 0-13-VI I-200 1, Xiao-Dong Yu & Hong-Zhang ZHOU leg., in coll. Institute of Zoology, Chinese Academy of Sciences, Beijing. Image from (Imura, Zhou & Su, 2004)
    Copyright © - - -
    Carabus (Cupreocarabus) sculptior geshizhanus (Imura, Zhou & Su, 2004)
    Carabus (Cupreocarabus) sculptior geshizhanus (Imura, Zhou & Su, 2004) Holotype
    Male adult
    mixed forest, 2,610 m in altitude, Geshizha in central Danba Xian of west-central Sichuan, Southwest China, 20-22-V II-200 1, Xiao-Dong Yu & Hong-Zhang ZHOU leg. Image from (Imura, Zhou & Su, 2004)
    Copyright © - - -
    Carabus (Carabus) donjuan Imura, 2011
    Carabus (Carabus) donjuan Imura, 2011 Holotype Paratype
    Male+Female adult
    alpine zone of the Chuandongzi Massif, 27.21.04-34N / 101.30.25-33E, 3035-3290 m in alt., south of Yanyuan, Yanyuan Xian, Liangshan-yizu-zizhizhou, S Sichuan, SW China, 29.VI.2010 (ZIN). Image from Imura, 2011
    Copyright © - - -
    Carabus (Carabus) donjuan Imura, 2011
    Carabus (Carabus) donjuan Imura, 2011 Holotype
    Endophallus
    Image from Imura, 2011
    Copyright © - - -
    Carabus (Pseudocoptolabrus) belousovianus Imura, 2011
    Carabus (Pseudocoptolabrus) belousovianus Imura, 2011 Holotype Paratype
    Male+Female adult
    uppermost zone of the Chuandongzi Massif. '27・20' 21-34" NI IOI・ 32' 42-52" E. 4070・4135 m in alt . south of Yanyuan, Yanyuan Xian. Liangshan-yizu-Zizhizhou. southern Sichuan, Southwest China, 4-VII-2010. preserved in the Zoological Institute of Academy of Sciences. St. Petersburg. Russia. Image from Imura, 2011
    Copyright © - - -
    Carabus (Pseudocoptolabrus) belousovianus Imura, 2011
    Carabus (Pseudocoptolabrus) belousovianus Imura, 2011
    Endophallus
    Image from Imura, 2011
    Copyright © - - -
    Carabus (Carabus) koidei yanyuan Cavazzuti & Rapuzzi, 2010
    Carabus (Carabus) koidei yanyuan Cavazzuti & Rapuzzi, 2010
    Male adult
    Image from Imura, 2011
    Copyright © - - -
    Carabus (Carabus) koidei yanyuan Cavazzuti & Rapuzzi, 2010
    Carabus (Carabus) koidei yanyuan Cavazzuti & Rapuzzi, 2010
    Endophallus
    Image from Imura, 2011
    Copyright © - - -
    Carabus (Pseudocoptolabrus) masahiroi (Imura, 2006)
    Carabus (Pseudocoptolabrus) masahiroi (Imura, 2006) Holotype
    Male adult
    Chudu Razi Massif, E. Kachin, N. Myanmar, VIII-2005, to be preserved in the National Science Museum (Nat. Hist), Tokyo. Image from Imura, 2011
    Copyright © - - -
    Carabus (Pseudocoptolabrus) branaungi liukuensis (Imura, 2007)
    Carabus (Pseudocoptolabrus) branaungi liukuensis (Imura, 2007) Holotype
    Male adult
    south-southwest of Liuku , (25°41'31"N/98°45 '47"E), 3,545 m in altitude, in Lushui Xian, of western Yunnan, China, 1 9-V-2006, preserved in the collection of the Department of Zoology, National Science Museum (Nat. Hist.), Tokyo. Image from Imura, 2011
    Copyright © - - -
    Carabus (Hemicarabus) macleayi amanoi (Imura, 2004)
    Carabus (Hemicarabus) macleayi amanoi (Imura, 2004)
    Male adult
    alpine zone of Mt. Rishiri-zan, on Is. Rishiri-to, off the western coast of the northern tip of Hokkaido, Northeast Japan. Image from Imura, 2005
    Copyright © - - -
    Carabus (Hemicarabus) macleayi amanoi (Imura, 2004)
    Carabus (Hemicarabus) macleayi amanoi (Imura, 2004)
    Male adult
    alpine zone of Mt. Rishiri-zan, on Is. Rishiri-to, off the western coast of the northern tip of Hokkaido, Northeast Japan. Image from Imura, 2005
    Copyright © - - -
    Carabus (Coptolabrus) elysii amucronatus Deuve, 2010
    Carabus (Coptolabrus) elysii amucronatus Deuve, 2010 Holotype
    Male adult
    Chine, Shanxi, Yongji. 29mm
    Copyright © Thierry Deuve
    Carabus (Rhigocarabus) pseudopusio metallicior Deuve, 1996
    Carabus (Rhigocarabus) pseudopusio metallicior Deuve, 1996 Holotype
    Male adult
    NW-Sichuan: 23km SSW of Shangbazhai, 32.25N 102.57E 4300m
    Copyright © Thierry Deuve
    Carabus (Pseudocranion) shuamaluko luhuanus Cavazzuti & Rapuzzi, 2005
    Carabus (Pseudocranion) shuamaluko luhuanus Cavazzuti & Rapuzzi, 2005 Holotype
    Male adult
    Cina, Sichuan, Heishui Xian (Luhua), Dakog Bing Chuan 2700 m. (Holotypus in coll. Cavazzuti)
    Copyright © - - -
    Carabus (Pseudocranion) absonus Cavazzuti & Rapuzzi, 2005
    Carabus (Pseudocranion) absonus Cavazzuti & Rapuzzi, 2005 Holotype
    Male adult
    Cina, Shaanxi meridionale, Zhiyang, Dabashan 1600 m i meridionale, Zhiyang, Dabashan 1600 m. VII-2003 (in coll. Rapuzzi). Image from Cavazzuti & Rapuzzi, 2005
    Copyright © - - -
    Carabus (Pseudocoptolabrus) armiger legionarius Cavazzuti & Rapuzzi, 2005
    Carabus (Pseudocoptolabrus) armiger legionarius Cavazzuti & Rapuzzi, 2005 Holotype
    Female adult
    Cina, Sichuan meridionale, Passo a sud-ovest di Mianning, 2650 m. (in coll. Rapuzzi). Image from Cavazzuti & Rapuzzi, 2005
    Copyright © - - -
    Carabus (Hemicarabus) nitens Linne, 1758
    Carabus (Hemicarabus) nitens Linne, 1758
    Male adult
    Belarus, Vitebsk reg., Miory distr. Swamp «Mokh», 09.05.2011, leg. G.G. Sushko
    Copyright © Igor Solodovnikov
    Carabus (Hemicarabus) nitens Linne, 1758
    Carabus (Hemicarabus) nitens Linne, 1758
    Endophallus
    Belarus, Vitebsk reg., Miory distr. Swamp «Mokh», 09.05.2011, leg. G.G. Sushko
    Copyright © Igor Solodovnikov
    Carabus (Hemicarabus) nitens Linne, 1758
    Carabus (Hemicarabus) nitens Linne, 1758
    Male adult

    Copyright © Igor Solodovnikov
    Carabus (Hemicarabus) tuberculosus Dejean, 1829
    Carabus (Hemicarabus) tuberculosus Dejean, 1829
    Male adult
    Carabus (Hemicarabus) tuberculosus n. strophium F.-W., 1842 SE Kazakhstan: NW Tarbogatai Mt.R.
    Copyright © Igor Solodovnikov
    Carabus (Hemicarabus) tuberculosus Dejean, 1829
    Carabus (Hemicarabus) tuberculosus Dejean, 1829
    Endophallus
    Carabus (Hemicarabus) tuberculosus n. strophium F.-W., 1842 SE Kazakhstan: NW Tarbogatai Mt.R.
    Copyright © Igor Solodovnikov
    Carabus (Hemicarabus) tuberculosus Dejean, 1829
    Carabus (Hemicarabus) tuberculosus Dejean, 1829
    Female adult

    Copyright © Igor Solodovnikov
    Carabus (Hemicarabus) tuberculosus Dejean, 1829
    Carabus (Hemicarabus) tuberculosus Dejean, 1829
    Female adult
    21mm
    Copyright © Boleslav Brezina
    Carabus (Hemicarabus) tuberculosus Dejean, 1829
    Carabus (Hemicarabus) tuberculosus Dejean, 1829
    Male adult
    21mm
    Copyright © Boleslav Brezina
    Carabus (Hemicarabus) serratus Say, 1825
    Carabus (Hemicarabus) serratus Say, 1825
    Male adult

    Copyright © Igor Solodovnikov
    Carabus (Hemicarabus) serratus Say, 1825
    Carabus (Hemicarabus) serratus Say, 1825
    Endophallus
    E-USA,Virginia, Leesburg env., Ball's Bluff Battlefield Regional Park
    Copyright © Igor Solodovnikov
    Carabus (Hemicarabus) serratus Say, 1825
    Carabus (Hemicarabus) serratus Say, 1825
    Female adult

    Copyright © Igor Solodovnikov
    Carabus (Isiocarabus) fiduciarius saishutoicus Csiki, 1927
    Carabus (Isiocarabus) fiduciarius saishutoicus Csiki, 1927 Lectotype
    Unsexed

    Copyright © - - -
    Carabus (Cupreocarabus) minimocupreus zhegushanus Cavazzuti & Rapuzzi, 2005
    Carabus (Cupreocarabus) minimocupreus zhegushanus Cavazzuti & Rapuzzi, 2005 Holotype
    Female adult
    Cina, Sichuan centrale, Lixian, Colle di Zhegushan 4100 m.
    Copyright © - - -
    Carabus (Cupreocarabus) melii Cavazzuti & Ratti, 2005
    Carabus (Cupreocarabus) melii Cavazzuti & Ratti, 2005 Holotype
    Male adult
    Cina, Sichuan centrale, Valle a nord-ovest di Heishui, 2700 m
    Copyright © - - -
    Carabus (Zheduocarabus) jiulongensis paolorapuzzii Cavazzuti & Rapuzzi, 2005
    Carabus (Zheduocarabus) jiulongensis paolorapuzzii Cavazzuti & Rapuzzi, 2005 Holotype
    Male adult
    Cina, Sichuan meridionale, Sabde. 24-Vl/28-VI-2001, Kucera leg, in coll. Rapuzzi.
    Copyright © - - -
    Carabus (Eucarabus) parreyssi parreyssi Palliardi, 1825
    Carabus (Eucarabus) parreyssi parreyssi Palliardi, 1825
    Female adult
    SW-Croatia, Mts Velebit, Mali Halan, h=1390 m, 23.V-20.VI.2022, L=24 mm
    Copyright © Ruslan Panin
    Carabus (Eucarabus) parreyssi parreyssi Palliardi, 1825
    Carabus (Eucarabus) parreyssi parreyssi Palliardi, 1825
    Endophallus
    SW-Croatia, Mts Velebit, Mali Halan, h=1390 m,
    Copyright © Ruslan Panin
    Carabus (Eucarabus) parreyssi parreyssi Palliardi, 1825
    Carabus (Eucarabus) parreyssi parreyssi Palliardi, 1825
    Male adult
    SW-Croatia, Mts Velebit, Mali Halan, h=1390 m, 23.V-20.VI.2022, L=22.5 mm
    Copyright © Ruslan Panin
    Carabus (Archicarabus) monticola monticola (as maritimensis Born, 1923)
    Carabus (Archicarabus) monticola monticola (as maritimensis Born, 1923)
    Female adult
    SE France, Alpes-Maritimes, Col de Braus h=1000 m, 23.V-18.VI.2014, P. Cavazzuti & G. Proscia leg., L= 24 mm
    Copyright © Ruslan Panin
    Carabus (Pachystus) cavernosus variolatus Costa, 1839
    Carabus (Pachystus) cavernosus variolatus Costa, 1839
    Female adult
    C-Italy: Abruzzo reg., Gran Sasso massif, Campo Imperatore, h=1900 m, VIII.2021, G.Proscia leg., L=27 mm
    Copyright © Ruslan Panin
    Carabus (Archicarabus) montivagus vellepiticus Hampe, 1850
    Carabus (Archicarabus) montivagus vellepiticus Hampe, 1850
    Female adult
    W Croatia, NP Northern Velebit, Stolak, 44.969492, 14.986169, 23.V-27.VI.2022, L=24 mm
    Copyright © Ruslan Panin
    Carabus (Archicarabus) montivagus vellepiticus Hampe, 1850
    Carabus (Archicarabus) montivagus vellepiticus Hampe, 1850
    Male adult
    W Croatia, NP Northern Velebit, Stolak, 44.969492, 14.986169, 23.V-27.VI.2022, L=23.5 mm
    Copyright © Ruslan Panin
    Carabus (Chrysocarabus) solieri bonnetianus Colas, 1936 (loc.typ.)
    Carabus (Chrysocarabus) solieri bonnetianus Colas, 1936 (loc.typ.)
    Female adult
    SE-France, Var, Tanneron, Saint-Cassien des Bois, 18.IV.2019, L=32 mm
    Copyright © Ruslan Panin
    Carabus (Chrysocarabus) solieri bonnetianus Colas, 1936 (loc.typ.)
    Carabus (Chrysocarabus) solieri bonnetianus Colas, 1936 (loc.typ.)
    Male adult
    SE-France, Var, Tanneron, Saint-Cassien des Bois, 18.IV.2019, L= 31 mm
    Copyright © Ruslan Panin
    Carabus (Chrysocarabus) solieri clairi Gehin, 1885 (as devillei Tarrier, 1972)
    Carabus (Chrysocarabus) solieri clairi Gehin, 1885 (as devillei Tarrier, 1972)
    Male adult
    SE-France, Alpes-Maritimes, Beuil env., h=1550m, 14.VII.1986
    Copyright © Ruslan Panin
    Carabus (Procrustes) coriaceus excavatus Charpentier, 1825
    Carabus (Procrustes) coriaceus excavatus Charpentier, 1825
    Male adult
    W Croatia, Velebit Mts, Jablanac Stirovaca h=998 m, 44.71124, 14.95673,3.VI-7.VII.2017, G. Proscia leg., L= 37 mm
    Copyright © Ruslan Panin
    Carabus (Procrustes) coriaceus excavatus Charpentier, 1825
    Carabus (Procrustes) coriaceus excavatus Charpentier, 1825
    Female adult
    W Croatia, Velebit Mts, Jablanac Stirovaca h=998 m, 44.71124, 14.95673,3.VI-7.VII.2017, G. Proscia leg., L= 38 mm
    Copyright © Ruslan Panin
    Carabus (Imaibius) dardiellus polymorphus Heinz, 1993
    Carabus (Imaibius) dardiellus polymorphus Heinz, 1993 Holotype
    Male adult
    Reshian, 2100-2500 m, W. Heinz leg. Image from Deuve & Schmidt, 2023
    Copyright © Thierry Deuve
    Carabus (Imaibius) dardiellus walterheinzi Deuve & Schmidt, 2023
    Carabus (Imaibius) dardiellus walterheinzi Deuve & Schmidt, 2023 Holotype
    Male adult
    Pakistan, Azad Jammu & Kashmir, District de Muzaffarabad, environs de Reshian, 2100-2500 mètres (W. Heinz leg.), in coll. Muséum de Stuttgart, Allemagne
    Copyright © Thierry Deuve
    Carabus (Aristocarabus) viridifossulatus nobilissimus Cavazzuti & Rapuzzi, 2005
    Carabus (Aristocarabus) viridifossulatus nobilissimus Cavazzuti & Rapuzzi, 2005 Paratype
    Female adult
    Cina, Sichuan occidentale, Jintang, Jia Ching Shan 3650 m
    Copyright © - - -
    Carabus (Apotomopterus) patroclus catagraphus Cavazzuti & Rapuzzi, 2005
    Carabus (Apotomopterus) patroclus catagraphus Cavazzuti & Rapuzzi, 2005 Holotype
    Male adult
    China, Sichuan occidentale, Jiulong pref., Mai-Pu, 3000-3500 m. VII-2004, in coli. Rapuzzi
    Copyright © - - -
    Carabus (Apotomopterus) tibeticus Cavazzuti & Rapuzzi, 2005
    Carabus (Apotomopterus) tibeticus Cavazzuti & Rapuzzi, 2005 Holotype
    Female adult
    Tibet orientale, Chamdo pref. (Yi Gong), Po-mi, 2300-2700 m. 20/27-VIII- 2004, in coli. Rapuzzi.
    Copyright © - - -
    Carabus (Archaeocarabus) mirandus Cavazzuti & Rapuzzi, 2005
    Carabus (Archaeocarabus) mirandus Cavazzuti & Rapuzzi, 2005 Holotype
    Male adult
    Cina, Sichuan meridionale, Jinyang, 3000 m.
    Copyright © - - -
    Carabus (Leptocarabus) vogtianus horvatovichi Deuve, 1992
    Carabus (Leptocarabus) vogtianus horvatovichi Deuve, 1992 Holotype
    Male adult
    Image from: Korean type specimens of insects deposited in Hungarian Natural History Museum.
    Copyright © - - -
    Carabus (Eucarabus) szeli szeli
    Carabus (Eucarabus) szeli szeli Holotype
    Male adult
    Image from: Korean type specimens of insects deposited in Hungarian Natural History Museum.
    Copyright © - - -
    Carabus (Megodontus) croaticus croaticus Dejean, 1826
    Carabus (Megodontus) croaticus croaticus Dejean, 1826
    Endophallus
    W-Croatia, Velebit Mts., h=950 m, Oltari - Zavizan,
    Copyright © Ruslan Panin
    Carabus (Megodontus) croaticus croaticus Dejean, 1826
    Carabus (Megodontus) croaticus croaticus Dejean, 1826
    Female adult
    W-Croatia, Velebit Mts., h=950 m, Oltari - Zavizan, 30.VI. 2018, L=32 mm
    Copyright © Ruslan Panin
    Carabus (Megodontus) croaticus croaticus Dejean, 1826
    Carabus (Megodontus) croaticus croaticus Dejean, 1826
    Male adult
    W-Croatia, Velebit Mts., h=950 m, Oltari - Zavizan, 30.VI.2018
    Copyright © Ruslan Panin
    Carabus (Alipaster) foreli baiensis Deuve & Kabak, 2013
    Carabus (Alipaster) foreli baiensis Deuve & Kabak, 2013 Holotype
    Male adult
    Chine Xinjiang, affluent droit de la rivière Terek, 3500 mètres (env. 58 km à l'ouest-nord-ouest de Baicheng), 42°0l'N/81°15'E, pelouses alpines et éboulis, in coll. Jaroslav Kalâb, à Jinacovice (République Tchèque)
    Copyright © Thierry Deuve
    Carabus (Alipaster) foreli baiensis Deuve & Kabak, 2013
    Carabus (Alipaster) foreli baiensis Deuve & Kabak, 2013
    Male genitalia

    Copyright © Thierry Deuve
    Carabus (Alipaster) foreli baiensis Deuve & Kabak, 2013
    Carabus (Alipaster) foreli baiensis Deuve & Kabak, 2013 Paratype
    Male adult
    Chine, Xinjiang, versant sud des Mts Khalyk-taou (Halik Shan), Bay Xian, au nord-ouest de Terekmeikon, 3200-3690 mètres, 42°02'22"N/81°26'17"E à 42°01'29"N/81°23'43"E (!. Kabak, 12 juillet 2006)
    Copyright © Thierry Deuve
    Carabus (Chaetocarabus) intricatus bayardi (as silaensis Haury, 1881)
    Carabus (Chaetocarabus) intricatus bayardi (as silaensis Haury, 1881)
    Endophallus
    S-Italy, Calabria, Sila Grande Massif, Camigliatello h=1040 m
    Copyright © Ruslan Panin
    Carabus (Chaetocarabus) intricatus bayardi (as silaensis Haury, 1881)
    Carabus (Chaetocarabus) intricatus bayardi (as silaensis Haury, 1881)
    Endophallus
    S-Italy, Calabria, Sila Grande Massif, Camigliatello h=1040 m, 19.XI.2009, G. Proscia leg. L=31 mm
    Copyright © Ruslan Panin
    Carabus (Chaetocarabus) intricatus bayardi (as silaensis Haury, 1881)
    Carabus (Chaetocarabus) intricatus bayardi (as silaensis Haury, 1881)
    Male adult
    S-Italy, Calabria, Sila Grande Massif, Camigliatello h=1040 m, 19.XI.2009, G. Proscia leg. L=28 mm
    Copyright © Ruslan Panin
    Carabus (Cupreocarabus) balangicus balangicus Cavazzuti, 2002
    Carabus (Cupreocarabus) balangicus balangicus Cavazzuti, 2002
    Endophallus
    Chine, C-Sichuan, Xiaojin, Wolong valley: Balang Shan Pass h=3448 m, 30.884617, 102.973117
    Copyright © Ruslan Panin
    Carabus (Cupreocarabus) balangicus balangicus Cavazzuti, 2002
    Carabus (Cupreocarabus) balangicus balangicus Cavazzuti, 2002
    Male adult
    Chine, C-Sichuan, Xiaojin Co, Wolong valley, Balang Shan Pass h=3448 m, 30.884617, 102.973117, 9.VI.-1.VII.2011, G. Proscia leg. L=19 mm
    Copyright © Ruslan Panin
    Carabus (Cupreocarabus) balangicus balangicus Cavazzuti, 2002
    Carabus (Cupreocarabus) balangicus balangicus Cavazzuti, 2002
    Male adult
    Chine, C-Sichuan, Xiaojin Co, Wolong valley, Balang Shan Pass h=3448 m, 30.884617, 102.973117, 9.VI.-1.VII.2011, G. Proscia leg. L=20 mm
    Copyright © Ruslan Panin
    Carabus (Rhigocarabus) cateniger cateniger (as sommitalis Cavazzuti, 2006)
    Carabus (Rhigocarabus) cateniger cateniger (as sommitalis Cavazzuti, 2006)
    Female adult
    China, C-Gansu, Mts Qilian h=3378 m, road G227, Yangxiongzi env. 38.050917, 100.887267, 18-23.VI.2011, G.Proscia leg., L=19 mm
    Copyright © Ruslan Panin
    Carabus (Rhigocarabus) cateniger cateniger (as sommitalis Cavazzuti, 2006)
    Carabus (Rhigocarabus) cateniger cateniger (as sommitalis Cavazzuti, 2006)
    Male adult
    China, C-Gansu, Mts Qilian h=3378 m, road G227, Yangxiongzi env. 38.050917, 100.887267, 18-23.VI.2011, G.Proscia leg., L=17 mm
    Copyright © Ruslan Panin
    Carabus (Rhigocarabus) cateniger dabanicola Deuve, 1998
    Carabus (Rhigocarabus) cateniger dabanicola Deuve, 1998
    Male adult
    China, Qinghai, Xining city, 55km NE of Datong, Daban Shan h=3936 m, 37.348450, 101.397600, 17-23.VI.2011, G.Proscia leg., L= 16 mm
    Copyright © Ruslan Panin
    Carabus (Rhigocarabus) cateniger dabanicola Deuve, 1998
    Carabus (Rhigocarabus) cateniger dabanicola Deuve, 1998
    Female adult
    China, Qinghai, Xining city, 55km NE of Datong, Daban Shan h=3936 m, 37.348450, 101.397600, 17-23.VI.2011, G.Proscia leg., L= 18.5 mm
    Copyright © Ruslan Panin
    Carabus (Rhigocarabus) roborowskii roborowskii Semenov, 1887
    Carabus (Rhigocarabus) roborowskii roborowskii Semenov, 1887
    Male adult
    China, E-Qinghai, Mts NW of Tongren city, 35.562778, 101.843667, h=3870 m, 15-26.VI.2011, G. Proscia leg./det.R.Panin, L=20 mm
    Copyright © Ruslan Panin
    Carabus (Rhigocarabus) roborowskii roborowskii Semenov, 1887
    Carabus (Rhigocarabus) roborowskii roborowskii Semenov, 1887
    Endophallus
    China, E-Qinghai, Mts NW of Tongren city, 35.562778, 101.843667, h=3870 m
    Copyright © Ruslan Panin
    Carabus (Rhigocarabus) roborowskii roborowskii Semenov, 1887
    Carabus (Rhigocarabus) roborowskii roborowskii Semenov, 1887
    Female adult
    China, E-Qinghai, Mts NW of Tongren city, 35.562778, 101.843667, h=3870 m, 15-26.VI.2011, G. Pros cia leg./det.R.Panin, L=20 mm
    Copyright © Ruslan Panin
    Carabus (Cupreocarabus) huangi huangi Deuve, 1992
    Carabus (Cupreocarabus) huangi huangi Deuve, 1992
    Male adult
    China, E Tibet, N-Sichuan, 53 km NE of Barkam, Pass Sandaoban env., h=4293 m, 32.232950, 102.598533, 11-29.VI.2011, G. Proscia leg., L=21 mm
    Copyright © Ruslan Panin
    Carabus (Cupreocarabus) huangi huangi Deuve, 1992
    Carabus (Cupreocarabus) huangi huangi Deuve, 1992
    Female adult
    China, E Tibet, N-Sichuan, 53 km NE of Barkam, Pass Sandaoban env., h=4293 m, 32.232950, 102.598533, 11-29.VI.2011, G. Proscia leg., L=21.5 mm
    Copyright © Ruslan Panin
    Carabus (Tomocarabus) filippovi Rapuzzi, 2023
    Carabus (Tomocarabus) filippovi Rapuzzi, 2023 Holotype
    Male adult
    Mongolia,Hentei highlands, NE Ulan Bator, VI.2009, Evgenii Filippov legit. (in coll. Rapuzzi). Image from Rapuzzi, 2023. New combination by Anichtchenko 9.9.2023
    Copyright © - - -
    Carabus (Morphocarabus) latreillei mengguicus Rapuzzi, 2023
    Carabus (Morphocarabus) latreillei mengguicus Rapuzzi, 2023 Holotype
    Male adult
    China, Inner Mongolia Province, Tongliao, Ka’erqin, Jinbaotun, VIII.20. Image from Rapuzzi, 2023
    Copyright © - - -
    Carabus (Cychrostomus) anchocephalus farinosus Rapuzzi, 2023
    Carabus (Cychrostomus) anchocephalus farinosus Rapuzzi, 2023 Holotype
    Male adult
    China, Inner Mongolia Province, 30 Km NW Hohhot, 3/7.VI.2010, M. Janata legit. Image from Rapuzzi, 2023
    Copyright © - - -
    Carabus (Apotomopterus) artemisioides Rapuzzi, 2023
    Carabus (Apotomopterus) artemisioides Rapuzzi, 2023 Holotype Paratype
    Male+Female adult
    Vietnam, Lai Chau, Tam Duong, VII.2022. Image from Rapuzzi, 2023
    Copyright © - - -
    Carabus (Apotomopterus) tenuicostatus Rapuzzi, 2023
    Carabus (Apotomopterus) tenuicostatus Rapuzzi, 2023 Holotype Paratype
    Male+Female adult
    Vietnam, Lai Chau, Tam Duong, IX.2022; Image from Rapuzzi, 2023
    Copyright © - - -
    Carabus (Apotomopterus) tonkinensis tonkinensis Deuve, 1990
    Carabus (Apotomopterus) tonkinensis tonkinensis Deuve, 1990
    Male adult
    Vietnam, Lai Chau, Tam Duong, IX.2022. Image from Rapuzzi, 2023
    Copyright © - - -
    Carabus (Neoplectes) zolotarevi Zamotajlov, 1988
    Carabus (Neoplectes) zolotarevi Zamotajlov, 1988
    Female adult
    Carabus (Archiplectes) zolotarevi zolotarevi Zamotajlov, 1988 Russia: Krasnodar reg., SW slopes of Malye Balkany Mt.R., env. Allous pass, h = 1960 m, 08-18.VII.2023 leg. D. Fominykh
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) zolotarevi Zamotajlov, 1988
    Carabus (Neoplectes) zolotarevi Zamotajlov, 1988
    Endophallus
    Carabus (Archiplectes) zolotarevi zolotarevi Zamotajlov, 1988 Russia: Krasnodar reg., SW slopes of Malye Balkany Mt.R., env. Allous pass, h = 1960 m, 08-18.VII.2023 leg. D. Fominykh
    Copyright © Igor Solodovnikov
    Carabus (Neoplectes) zolotarevi Zamotajlov, 1988
    Carabus (Neoplectes) zolotarevi Zamotajlov, 1988
    Male adult
    Carabus (Archiplectes) zolotarevi zolotarevi Zamotajlov, 1988 Russia: Krasnodar reg., SW slopes of Malye Balkany Mt.R., env. Allous pass, h = 1960 m, 08-18.VII.2023 leg. D. Fominykh
    Copyright © Igor Solodovnikov
    Carabus (Leptocarabus) semiopacus sstr (left); semiopacus gwangyangensis (right)
    Carabus (Leptocarabus) semiopacus sstr (left); semiopacus gwangyangensis (right)
    Male adult
    Image from T. Kwon et al (2023)
    Copyright © - - -
    Carabus (Leptocarabus) semiopacus Reitter, 1895
    Carabus (Leptocarabus) semiopacus Reitter, 1895
    Male adult
    1A and 3B: A. (A.) semiopacus semiopacus; 2A and 4B: A. (A.) semiopacus gwangyangensis Kwon & Park, 2023 (A: right lateral view; B: left and right basal lobes), (Scale bars: 1 mm). Image from T. Kwon et al, 2023
    Copyright © - - -
    Carabus (Leptocarabus) koreanus Reitter, 1895
    Carabus (Leptocarabus) koreanus Reitter, 1895
    Male adult
    Habitus of A. koreanus ssp. and male aedeagus tips: A. koreanus koreanus; B: A. koreanus minor; C: A. koreanus mineti; D: A. koreanus rectokoreanus (Scale bars: 1 cm). Image from T. Kwon et al 2023
    Copyright © - - -
    Carabus (Deroplectes) arcanus arcanus Semenov, 1898
    Carabus (Deroplectes) arcanus arcanus Semenov, 1898
    Endophallus

    Copyright © Igor Solodovnikov
    Carabus (Deroplectes) arcanus arcanus Semenov, 1898
    Carabus (Deroplectes) arcanus arcanus Semenov, 1898
    Female adult

    Copyright © Igor Solodovnikov
    Carabus (Deroplectes) arcanus arcanus Semenov, 1898
    Carabus (Deroplectes) arcanus arcanus Semenov, 1898
    Male adult

    Copyright © Igor Solodovnikov
    Carabus (Pseudocoptolabrus) taliensis jiaozicola Deuve, 2019
    Carabus (Pseudocoptolabrus) taliensis jiaozicola Deuve, 2019 Holotype
    Male adult
    Chine, Yunnan septentrional, Mont Jiaozi Xueshan, 3640 m, 26°05’N-102°51’E, in coll. Jaroslav Turna, Kostelec na Hane (République Tchèque)
    Copyright © Thierry Deuve
    Carabus (Ulocarabus) theanus xerophiloides Deuve, 2019
    Carabus (Ulocarabus) theanus xerophiloides Deuve, 2019 Holotype
    Female adult
    NO LABEL DATA! ex coll. M. Maindron, ex coll. G. Babault (MNHN).
    Copyright © Thierry Deuve
    Carabus (Carabus) gressittianus tiangongicus Deuve, 2019
    Carabus (Carabus) gressittianus tiangongicus Deuve, 2019 Holotype
    Female adult
    Chine, Fujian méridional, Tiangong Shan, 1508 m, 25°15’N-117°03’E, in coll. Jaroslav Turna, Kostelec na Hane (République Tchèque)
    Copyright © Thierry Deuve
    Carabus (Apotomopterus) guangxicus hongjianicus Deuve, 2019
    Carabus (Apotomopterus) guangxicus hongjianicus Deuve, 2019 Holotype
    Male adult
    Chine, Fujian méridional, Hongjian Shan, 1380 m, 25°04’N-117°11’E, in coll. Jaroslav Turna, Kostelec na Hane (République Tchèque).
    Copyright © Thierry Deuve
    Carabus (Deroplectes) arcanus hissarianus Semenov, 1900
    Carabus (Deroplectes) arcanus hissarianus Semenov, 1900
    Endophallus
    Tajikistan, Gissar ridge, Kondara, Kwak, h = 1900 m, 7,VI.1970, I.K. Lopatin leg.
    Copyright © Igor Solodovnikov
    Carabus (Deroplectes) sphinx dardazicus Deuve, 1990
    Carabus (Deroplectes) sphinx dardazicus Deuve, 1990
    Endophallus

    Copyright © Igor Solodovnikov
    Carabus (Deroplectes) sphinx dardazicus Deuve, 1990
    Carabus (Deroplectes) sphinx dardazicus Deuve, 1990
    Male adult

    Copyright © Igor Solodovnikov
    Carabus (Deroplectes) sphinx dardazicus Deuve, 1990
    Carabus (Deroplectes) sphinx dardazicus Deuve, 1990
    Female adult

    Copyright © Igor Solodovnikov
    Carabus (Apotomopterus) bousquetellus fengqingensis Deuve, Sota & Liang, 2023 (? bousquetellus)
    Carabus (Apotomopterus) bousquetellus fengqingensis Deuve, Sota & Liang, 2023 (? bousquetellus) Holotype
    Male adult
    Chine, Yunnan Prov., Fengqing Distr., col de Guihuashu, 24,67361°N 99,84449°E, 2460 mètres, 11.IX.2004, H. Liang et T. Sota, in coll. Institut Zoologique, Pékin (Chine). Image from Deuve, Sota & Liang, 2023
    Copyright © - - -
    Carabus (Cechenochilus) boeberi buschi Semenov & Znojko, 1931
    Carabus (Cechenochilus) boeberi buschi Semenov & Znojko, 1931
    Endophallus
    Russia: Kabardino-Balkaria: u.r. Adylsu Riv.
    Copyright © Igor Solodovnikov
    Carabus (Cechenochilus) boeberi buschi Semenov & Znojko, 1931
    Carabus (Cechenochilus) boeberi buschi Semenov & Znojko, 1931
    Male adult
    Russia: Kabardino-Balkaria: u.r. Adylsu Riv.
    Copyright © Igor Solodovnikov
    Carabus (Megodontus) tuvensis Shilenkov, 1996
    Carabus (Megodontus) tuvensis Shilenkov, 1996
    Endophallus

    Copyright © Igor Solodovnikov
    Carabus (Megodontus) tuvensis Shilenkov, 1996
    Carabus (Megodontus) tuvensis Shilenkov, 1996
    Male adult

    Copyright © Igor Solodovnikov
    Carabus (Eucarabus) catenulatus fontanellae Reitter, 1896
    Carabus (Eucarabus) catenulatus fontanellae Reitter, 1896
    Male adult
    W-Croatia, Dalmatia, Isl. Rab, 16-30.V.2019, G. Proscia leg.
    Copyright © Ruslan Panin
    Carabus (Trachycarabus) scabriusculus inapertus Motschulsky, 1850
    Carabus (Trachycarabus) scabriusculus inapertus Motschulsky, 1850
    Male adult
    NC Ukraine, Kiev city, Lysa Gora, 1-15.VI.2012, Gerasimov R.leg./det. R.Panin, L=19 mm
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) anacoreta Rapuzzi, 2024
    Carabus (Apotomopterus) anacoreta Rapuzzi, 2024 Holotype
    Male adult
    Ha Giang Province, Xin Man; Aug. 2023. Image from Rapuzzi, 2024; 31mm
    Copyright © - - -
    Carabus (Apotomopterus) vietnamicus Rapuzzi, 2023
    Carabus (Apotomopterus) vietnamicus Rapuzzi, 2023 Holotype Paratype
    Male+Female adult
    Vietnam, Xin Man, 2000/2100 m, VI.2023. Image from Rapuzzi, 2023; 32 mm
    Copyright © - - -
    Carabus (Apotomopterus) narcissus Rapuzzi, 2023
    Carabus (Apotomopterus) narcissus Rapuzzi, 2023 Holotype Paratype
    Male+Female adult
    Vietnam, Xin Man, 2000/2100 m, VI.2023. Image from Rapuzzi, 2023; 33 mm
    Copyright © - - -
    Carabus (Apotomopterus) tarasovinus toussainti Rapuzzi, 2024
    Carabus (Apotomopterus) tarasovinus toussainti Rapuzzi, 2024 Holotype
    Female adult
    Vietnam, Nghe An Province, Tuong Duong; Jun. 2023. Image from Rapuzzi, 2024
    Copyright © - - -
    Carabus (Apotomopterus) tonkinensis tropicus Rapuzzi, 2024
    Carabus (Apotomopterus) tonkinensis tropicus Rapuzzi, 2024 Holotype
    Female adult
    Vietnam, Da Nang Province, Ba Na hills, top of Mt. Chua; 1480 m asl; Aug. 2023. Image from Rapuzzi, 2024
    Copyright © - - -
    Carabus (Rhigocarabus) degensis ladyginioides Deuve, Liu & Liang, 2024
    Carabus (Rhigocarabus) degensis ladyginioides Deuve, Liu & Liang, 2024 Holotype
    Male adult
    Chine, province de Qinghai, canton de Madoi, col de Bayan Har, 4827 m, 34,1267°N 97,6575°E, pelouses alpines et éboulis (Hongliang Shi, Zhongbao Lu, Weifeng Yan & Pingzhou Zhu leg., 30.VII.2017, dans des pièges), in IOZ. Image from Deuve et al, 2024
    Copyright © - - -
    Carabus (Meganebrius) hongliangi Deuve, Liu & Liang, 2024
    Carabus (Meganebrius) hongliangi Deuve, Liu & Liang, 2024 Holotype
    Male adult
    Chine, Tibet, Préfecture de Xigazê, canton de Gyirong, ville de Gyirong, village de Cha, col de Latuo La, 3995 m, 28,3908°N 85,4074°E, pelouses alpines (Hongliang Shi leg., 10.VII.2019, sous des pierres), in IOZ. Image from Deuve et al 2024
    Copyright © - - -
    Carabus (Acoptopterus) dongchuanicus fazhensis Deuve, Liu & Liang, 2024
    Carabus (Acoptopterus) dongchuanicus fazhensis Deuve, Liu & Liang, 2024 Holotype
    Male adult
    Chine, province du Yunnan, district de Dongchuan, ville de Fazhe, Mont Jiaozi Shan, Yanzidong, 2910 m, 26,09758°N 102,89406°E, en bord de route (Hongliang Shi leg., 26.V.2010), in IOZ. Image from Deuve et al 2024
    Copyright © - - -
    Carabus (Acoptopterus) yunnanus dayaoicus Deuve, Liu & Liang, 2024
    Carabus (Acoptopterus) yunnanus dayaoicus Deuve, Liu & Liang, 2024 Holotype
    Male adult
    Chine, province du Yunnan, canton de Dayao, ville de Santai, col de Xiaobaicaoling, 3105 m, 26,0561°N 101,0544°E, en forêt mixte (Hongliang Shi & Ye Liu leg., 30.V.2012, dans des pièges), in IOZ. Image from Deuve et al, 2024
    Copyright © - - -
    Carabus (Neoplesius) takashimai qamdoensis Deuve, Liu & Liang, 2024
    Carabus (Neoplesius) takashimai qamdoensis Deuve, Liu & Liang, 2024 Holotype
    Female adult
    Chine, Tibet, “Qamdo” (Chaodong Zhu leg., 8.VIII.1997), in IOZ. Image from Deuve et al 2024
    Copyright © - - -
    Carabus (Calocarabus) przewalskii menyuanensis Deuve, Liu & Liang, 2024
    Carabus (Calocarabus) przewalskii menyuanensis Deuve, Liu & Liang, 2024 Holotype
    Male adult
    Chine, province du Qinghai, canton de Menyuan, ville de Zhugu, Sigou, 2602 m, 37,1353°N 102,3887°E (Yizhou Liu & Wenqi Yin leg., 26.VII.2020, dans des pièges), in IOZ. Image from Deuve 2024
    Copyright © - - -
    Carabus (Pseudocoptolabrus) taliensis huiliensis Deuve, Liu & Liang, 2024
    Carabus (Pseudocoptolabrus) taliensis huiliensis Deuve, Liu & Liang, 2024 Holotype
    Male adult
    Chine, province du Yunnan, canton de Huili, Mont Longzhou Shan, chemin vers le sommet, 3024 m, 26,8117°N 102,2238°E, forêt de Rhododendron (Hongliang Shi & Bo Liu leg., 19.VII.2015), in IOZ. Image from Deuve 2024
    Copyright © - - -
    Carabus (Morvanicarabus) scheibei scheibei Eidam, 1937
    Carabus (Morvanicarabus) scheibei scheibei Eidam, 1937
    Female adult
    NE-Afghanistan, Hindu Kush, Nuristan Prov., near Parun, 35°19'N 70°52'E h=2900 m, Pine forest, VI.2022, local collector/det. Kleinfeld, L= 24 mm
    Copyright © Ruslan Panin
    Carabus (Oreocarabus) gemellatus Menetries, 1832
    Carabus (Oreocarabus) gemellatus Menetries, 1832
    Male adult
    Iran, Mazandaran, pass b. Gach-i-Sar (N seite), 2700m, 17-21.vi.1998, Heinz leg. (cAA)
    Copyright © Alexander Anichtchenko
    Carabus (Lamprostus) punctatus Castelnau, 1835
    Carabus (Lamprostus) punctatus Castelnau, 1835
    Female adult
    Anatolia mer., Umg. Yayladagi (Antakya), 600-900m, 13.4.1981, Heinz leg. (cAA)
    Copyright © Alexander Anichtchenko
    Carabus (Procrustes) mulsantianus mulsantianus Morawitz, 1886 (as akbesianus)
    Carabus (Procrustes) mulsantianus mulsantianus Morawitz, 1886 (as akbesianus)
    Male adult
    Anatolia Mer., Martavan, ca. 600m, 32 km NW Kilis, 15.4.1989, Heinz leg.(cAA)
    Copyright © Alexander Anichtchenko
    Carabus (Mesocarabus) riffensis Fairmaire, 1875
    Carabus (Mesocarabus) riffensis Fairmaire, 1875
    Female adult
    N-Morocco, Riff Mts h=1650 m, 26 km E of Bab Berred, 34.961333, -4.669417, Cedar forest, C. Reuter leg. L=33 mm
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) riffensis Fairmaire, 1875
    Carabus (Mesocarabus) riffensis Fairmaire, 1875
    Endophallus
    N-Morocco, Riff Mts h=1650 m, 26 km E of Bab Berred
    Copyright © Ruslan Panin
    Carabus (Mesocarabus) riffensis Fairmaire, 1875
    Carabus (Mesocarabus) riffensis Fairmaire, 1875
    Male adult
    N-Morocco, Riff Mts h=1650 m, 26 km E of Bab Berred, 34.961333, -4.669417, Cedar forest, C. Reuter leg. L=27 mm
    Copyright © Ruslan Panin
    Carabus (Isiocarabus) miaorum Lassalle & Prunier, 1993
    Carabus (Isiocarabus) miaorum Lassalle & Prunier, 1993
    Endophallus
    China, NW-Guangxi, Mao'er Shan, h~1600 m 25°52'N 110°29'E
    Copyright © Ruslan Panin
    Carabus (Isiocarabus) miaorum Lassalle & Prunier, 1993
    Carabus (Isiocarabus) miaorum Lassalle & Prunier, 1993
    Male adult
    China, NW-Guangxi, Mao'er Shan, h~1600 m 25°52'N 110°29'E, 28.V-9.VI.2009, C.Reuter leg.
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) delavayi parvipes Cavazzuti, 1996
    Carabus (Apotomopterus) delavayi parvipes Cavazzuti, 1996
    Endophallus
    S-Sichuan: S of Yanyuan, Chuandongzi Massif h=3200 m, 27.351206, 101.507036
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) delavayi parvipes Cavazzuti, 1996
    Carabus (Apotomopterus) delavayi parvipes Cavazzuti, 1996
    Male adult
    S-Sichuan: S of Yanyuan, Chuandongzi Massif h=3200 m, 27.351206, 101.507036, 8-19.VI.2017, C.Reuter leg., L=32 mm
    Copyright © Ruslan Panin
    Carabus (Imaibius) caschmirensis fortetuberculatus Heinz, 1983
    Carabus (Imaibius) caschmirensis fortetuberculatus Heinz, 1983 Paratype
    Male adult

    Copyright © Alexander Anichtchenko
    Carabus (Imaibius) boysi boysi Tatum, 1851
    Carabus (Imaibius) boysi boysi Tatum, 1851
    Female adult
    N India, Uttarakhand, Ghangaria, N30.42.245, E079.35.634, 3000m, 11-13.6.2011, Anichtchenko A. leg. & det. (cAA)
    Copyright © Alexander Anichtchenko
    Carabus (Imaibius) caschmirensis fortetuberculatus Heinz, 1983
    Carabus (Imaibius) caschmirensis fortetuberculatus Heinz, 1983 Paratype
    Female adult

    Copyright © Alexander Anichtchenko
    Carabus (Lamprostus) syrus moukarzelae Deuve, 2002
    Carabus (Lamprostus) syrus moukarzelae Deuve, 2002
    Female adult
    S-Lebanon: Marjaayoun, 18.XI.-11.XII.2017, C. Reuter leg., L=35 mm
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) ludivinae wuliangensis Cavazzuti, 1997
    Carabus (Apotomopterus) ludivinae wuliangensis Cavazzuti, 1997
    Endophallus

    Copyright © Ruslan Panin
    Carabus (Apotomopterus) protenes ascendens Semenov, 1898
    Carabus (Apotomopterus) protenes ascendens Semenov, 1898
    Male adult
    China, Chongqing, N Jinfo Shan, Shanwangping h=1400m, 29.101944, 107.166389, 2-8.VI.2018, C.Reuter leg./det. F.Kleinfeld, L=26 mm
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) protenes ascendens Semenov, 1898
    Carabus (Apotomopterus) protenes ascendens Semenov, 1898
    Endophallus
    China, Chongqing, N Jinfo Shan, Shanwangping h=1400m, 29.101944, 107.166389
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) laoshanicus Imura, 1995
    Carabus (Apotomopterus) laoshanicus Imura, 1995
    Male adult
    China, E Guangxi, Jinxiu Yao County, Tongmu vill., local collector, L=36 mm
    Copyright © Ruslan Panin
    Carabus (Imaibius) nouristani nouristani (Ledoux, 1977)
    Carabus (Imaibius) nouristani nouristani (Ledoux, 1977)
    Female adult
    NE-Afghanistan: Nuristan: Darah-I-Nur valley, 2900m 34°45'N 70°40'E, VIII.2012, C.Reuter leg., L=26 mm
    Copyright © Ruslan Panin
    Carabus (Pachystus) cribratus cribratus Quensel, 1806
    Carabus (Pachystus) cribratus cribratus Quensel, 1806
    Endophallus
    NW Armenia, Amasia, (Shirak prov.)
    Copyright © Ruslan Panin
    Carabus (Pachystus) cribratus cribratus Quensel, 1806
    Carabus (Pachystus) cribratus cribratus Quensel, 1806
    Male adult
    NW Armenia, Amasia, (Shirak prov.), 20-24.VIII.1998, Kazaryan P.V. leg.
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) korellianus Kleinfeld, 2002
    Carabus (Apotomopterus) korellianus Kleinfeld, 2002
    Male adult
    China, SW-Yunnan, Dehong Distr., Ruili, Pijiazhai vill., h=1800 m, 24.048111, 97.738222, 9-12.VI.2023, C.Reuter leg., L=31 mm
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) korellianus Kleinfeld, 2002
    Carabus (Apotomopterus) korellianus Kleinfeld, 2002
    Endophallus
    China, SW-Yunnan, Dehong Distr., Ruili, Pijiazhai vill., h=1800 m, 24.048111, 97.738222
    Copyright © Ruslan Panin
    Carabus (Imaibius) nouristani nouristani (Ledoux, 1977)
    Carabus (Imaibius) nouristani nouristani (Ledoux, 1977)
    Male adult
    NE-Afghanistan: Nuristan: Darah-I-Nur valley, 2900m 34°45'N 70°40'E, VIII.2012, C.Reuter leg., L=26 mm
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) madefactus Cavazzuti, 1997
    Carabus (Apotomopterus) madefactus Cavazzuti, 1997
    Endophallus
    China, CW-Yunnan, Nánjiàn Yízú Zìzhìxiàn, Mt Lingbao h=2400 m, 24.768611, 100.500000
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) madefactus Cavazzuti, 1997
    Carabus (Apotomopterus) madefactus Cavazzuti, 1997
    Male adult
    China, CW-Yunnan, Nánjiàn Yízú Zìzhìxiàn, Mt Lingbao h=2400 m, 24.768611, 100.500000, 6-15.VI.2023, C.Reuter leg./det. F.Kleinfeld, L=27 mm
    Copyright © Ruslan Panin
    Carabus (Sigenacarabus) trachynodes moxiensis Deuve, 1992
    Carabus (Sigenacarabus) trachynodes moxiensis Deuve, 1992
    Male adult
    China, W-Sichuan, Daxue Range, Mt.Gongga Shan h=3000-3100m, Hailuogou Glacier Forest Park, 29.574889, 101.995556, 10-12.VI.2018, Ch. Reuter leg., L=21 mm
    Copyright © Ruslan Panin
    Carabus (Sigenacarabus) trachynodes moxiensis Deuve, 1992
    Carabus (Sigenacarabus) trachynodes moxiensis Deuve, 1992
    Endophallus
    China, W-Sichuan, Daxue Range, Mt.Gongga Shan h=3000-3100m, Hailuogou Glacier Forest Park, 29.574889, 101.995556
    Copyright © Ruslan Panin
    Carabus (Acoptopterus, ?Carabus) yunnanus yanyuanicus Cavazzuti, 1996
    Carabus (Acoptopterus, ?Carabus) yunnanus yanyuanicus Cavazzuti, 1996
    Endophallus
    China, S-Sichuan, S Yanyuan, Chuandongzi Mt h=3300-3600 m
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) mecynodes pinchuanensis Imura, 1999
    Carabus (Apotomopterus) mecynodes pinchuanensis Imura, 1999
    Endophallus
    China, S-Sichuan, 40-50 km E Xichang (to Zhaojue) 27°51'N 102°30'E, h=2700-3000m, Pine forest
    Copyright © Ruslan Panin
    Carabus (Acoptopterus) yunnanus yanyuanicus Cavazzuti, 1996 (? or oblongior ssp.)
    Carabus (Acoptopterus) yunnanus yanyuanicus Cavazzuti, 1996 (? or oblongior ssp.)
    Male adult
    China, S-Sichuan, S Yanyuan, Chuandongzi Mt h=3300-3600 m, 16-20.VI.2023, C.Reuter leg./det.Klienfeld, 2023, L=21.5 mm
    Copyright © Ruslan Panin
    Carabus (Rhigocarabus) rhododendron Deuve & Imura, 1991
    Carabus (Rhigocarabus) rhododendron Deuve & Imura, 1991
    Male adult
    China, N Yunnan: Baimaxue Shan, S of Deqen, h=4300m, 28°20'N 99°03'E, 16-20.VI.2016, C.Reuter leg., L=18 mm
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) mecynodes pinchuanensis Imura, 1999
    Carabus (Apotomopterus) mecynodes pinchuanensis Imura, 1999
    Female adult
    China, S-Sichuan, 40-50 km E Xichang (to Zhaojue) 27°51'N 102°30'E, h=2700-3000m, Pine forest, 5-20.VI.2017, C.Reuter leg., L=33 mm
    Copyright © Ruslan Panin
    Carabus (Tomocarabus) rumelicus rumelicus Chaudoir, 1867
    Carabus (Tomocarabus) rumelicus rumelicus Chaudoir, 1867
    Male adult
    Turkey, Yavsan, K. Maras, 7.7.2004 (cAA)
    Copyright © Alexander Anichtchenko
    Carabus (Piocarabus) vladimirskyi Dejean & Boisduval, 1830
    Carabus (Piocarabus) vladimirskyi Dejean & Boisduval, 1830
    Male adult
    Buryatia, Kyahta distr. [Khyagh], 5 km SE Chikoi vill., 16.8.1998, Anichtchenko A. leg.
    Copyright © Alexander Anichtchenko
    Carabus (Apoplesius) omphreodes omphreodes
    Carabus (Apoplesius) omphreodes omphreodes Holotype
    Male adult
    Paraplesius omphreodes 1898 Mardin / coll. Rttr 5-IX-1900 / coll. Roeschke type / Holotypus / omphreodes Rttr. / coll. C. & O. Vogt Acq. 1969 - in coll. Zoolog. Inst. Univ. Amsterdam; 23,5 mm.
    Copyright © Frank Kleinfeld
    Carabus (Apoplesius) omphreodes rawandozi Kleinfeld & Reuter, 2018
    Carabus (Apoplesius) omphreodes rawandozi Kleinfeld & Reuter, 2018 Holotype Paratype
    Male adult
    Holotype: N. Irak, Umg. Rawandoz, 36°38’N/44°33’E, Akoian valley, 30/IV/12-V-2018, 1900-2000 m, (23,5mm). Paratypus ♀: id. but IV-2017 (25 mm)
    Copyright © Frank Kleinfeld
    Carabus (Apotomopterus) patroclus apricalis Cavazzuti, 1997
    Carabus (Apotomopterus) patroclus apricalis Cavazzuti, 1997
    Endophallus
    China, S Sichuan, W of Pingchuanzhen, near Galuo vill., h=2400 m
    Copyright © Ruslan Panin
    Carabus (Lamprostus) hemprichi sidonius Lapouge, 1907
    Carabus (Lamprostus) hemprichi sidonius Lapouge, 1907
    Male adult
    Israel, Menashe Forest, b. Megiddo, 200-300mm 7.04.1987 Heinz leg (cAA). 27 mm
    Copyright © Alexander Anichtchenko
    Carabus (Apotomopterus) patroclus apricalis Cavazzuti, 1997
    Carabus (Apotomopterus) patroclus apricalis Cavazzuti, 1997
    Male adult
    China, S Sichuan, W of Pingchuanzhen, near Galuo vill., h=2400 m, 7-18.VI.2017, C.Reuter leg./det. F.Kleinfeld, L=30 mm
    Copyright © Ruslan Panin
    Carabus (Neoplectes) komarowi komarowi Reitter, 1882
    Carabus (Neoplectes) komarowi komarowi Reitter, 1882 Lectotype Paralectotypes
    Male+Female adult
    LT - female «Kaukasus Swanetien Leder Reitter » , «Komarowi m.», «Roeschke C.», « MA.INS.COLE.194.1», «Lectotype», «Carabus komarowi Reitter, Lectotype, Th. Deuve des. 2024», ex coll. Vogt, in coll. Naturalis Biodiversity Center, Leyde (Pays-Bas). Image from Deuve, 2024
    Copyright © Thierry Deuve
    Carabus (Neoplectes) komarowi komarowi Reitter, 1882
    Carabus (Neoplectes) komarowi komarowi Reitter, 1882 Lectotype Paralectotypes
    Female adult
    LT, left, female: «Kaukas Leder», «Komarowi v. mutabilis m.», «Roeschke C. », « Collectie C. & O. Vogt, Acq. 1960», «Plectes Komarowi mutabilis Reitter, 1896, ZMAN type COLE.0264.1», «Lectotype», «Carabus komarowi mutabilis Reitter, Lectotype, Th. Deuve des. 2024 » , ex coll. Vogt, in coll. Naturalis Biodiversity Center, Leyde (Pays-Bas). Image from Deuve, 2024
    Copyright © Thierry Deuve
    Carabus (Neoplectes) compressus rebellis Reitter, 1895
    Carabus (Neoplectes) compressus rebellis Reitter, 1895 Lectotype
    Male adult
    Image from Deuve, 2024
    Copyright © Thierry Deuve
    Carabus (Tribax) perditus Deuve, 2024
    Carabus (Tribax) perditus Deuve, 2024 Holotype
    Male adult
    MNHN
    Copyright © Thierry Deuve
    Carabus (Tibax) komarowi antoni Deuve, 2024
    Carabus (Tibax) komarowi antoni Deuve, 2024 Holotype
    Male adult
    «Kaukasus Swanetien Leder Reitter», ex coll. M. Maindron, ex coll. G. Babault, in coll. Muséum National d’Histoire Naturelle, Paris (France).
    Copyright © Thierry Deuve
    Carabus (Tribax) compressus pseudorebellis Deuve, 2024
    Carabus (Tribax) compressus pseudorebellis Deuve, 2024 Holotype
    Male adult
    Géorgie, Basse-Svanétie, 12 km au nord-est de Lentekhi, Mont Goldash, vallée de la Cholshoura, Durashi (« Dvashi »), env. 1150 mètres (W. Heinz, 1-4 juillet 2019), in coll. Muséum Stuttgart (Germany).
    Copyright © Thierry Deuve
    Carabus (Neoplectes) compressus zubovi Fominykh, Zamotajlov et Titarenko, 2018
    Carabus (Neoplectes) compressus zubovi Fominykh, Zamotajlov et Titarenko, 2018 Paratype
    Female adult
    S slopes of Svaneti Mt. Range near Lentekhi Town, S slopes of Goldash Mt., 42°50'N, 42°45'E, 2200–2600 m, 18.VI–13.VII.2017 (D. Fominykh). Image from Fominykh, Zamotajlov et Titarenko, 2018
    Copyright © - - -
    Carabus (Neoplectes) compressus zubovi Fominykh, Zamotajlov et Titarenko, 2018
    Carabus (Neoplectes) compressus zubovi Fominykh, Zamotajlov et Titarenko, 2018 Paratype
    Male+Female adult
    5-6: Paki Vill. environs; 7-8: Mannanauri Vill. environs. Scale bar 10.0 mm. Image from Fominykh, Zamotajlov et Titarenko, 2018
    Copyright © - - -
    Carabus (Apotomopterus) microtatos lingbaonis Imura, 2019
    Carabus (Apotomopterus) microtatos lingbaonis Imura, 2019
    Endophallus
    China, CW-Yunnan, Nánjiàn Yízú Zìzhìxiàn, Mt Lingbao h=2400 m, 24.768611, 100.500000
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) microtatos lingbaonis Imura, 2019
    Carabus (Apotomopterus) microtatos lingbaonis Imura, 2019
    Male adult
    China, CW-Yunnan, Nánjiàn Yízú Zìzhìxiàn, Mt Lingbao h=2400 m, 24.768611, 100.500000, 6-15.VI.2023, C.Reuter leg., L=24 mm
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) bousquetellus bousquetellus Deuve, 1998
    Carabus (Apotomopterus) bousquetellus bousquetellus Deuve, 1998
    Endophallus
    S-China, CW-Yunnan, W of Lincang, Daxue Shan h=2600m, 23.935556, 100.257778
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) patroclus patroclus Breuning, 1933
    Carabus (Apotomopterus) patroclus patroclus Breuning, 1933
    Endophallus
    China, S Sichuan, Pass 56 km SW Miannin, near Wawo h=2440 m
    Copyright © Ruslan Panin
    Carabus (Eurycarabus) famini atlantis Alluaud, 1923
    Carabus (Eurycarabus) famini atlantis Alluaud, 1923
    Female adult
    C Morocco, Haut Atlas, SW of Oukaïmeden h=2900m, 31.173644, -7.844442, 29.IV-11.V.2024, C.Reuter leg., L=21 mm
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) mecynodes pinchuanensis Imura, 1999
    Carabus (Apotomopterus) mecynodes pinchuanensis Imura, 1999
    Male adult
    China, S-Sichuan, 40-50 km E Xichang (to Zhaojue) 27°51'N 102°30'E, h=2700-3000m, Pine forest, 5-20.VI.2017, C.Reuter leg., L= 29 mm
    Copyright © Ruslan Panin
    Carabus (Lamprostus) syrus cheikensis Deuve, 1992
    Carabus (Lamprostus) syrus cheikensis Deuve, 1992
    Female adult
    SE Lebanon, Rashaya Distr., Tannoura, oak forest, h=700 m, 11-19.IV.2016, C. Reuter leg., L= 32 mm
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) microtatos lingbaonis Imura, 2019
    Carabus (Apotomopterus) microtatos lingbaonis Imura, 2019
    Female adult
    China, CW-Yunnan, Nánjiàn Yízú Zìzhìxiàn, Mt Lingbao h=2400 m, 24.768611, 100.500000, 6-15.VI.2023, C.Reuter leg., L=25 mm
    Copyright © Ruslan Panin
    Carabus (Mimocarabus) maurus smyrnensis Deuve, 2009
    Carabus (Mimocarabus) maurus smyrnensis Deuve, 2009 Holotype
    Male adult
    Turquie, Anatolie occidentale, Izmir (« Smyrne, Asie Mineure »), in Muséum national d’Histoire naturelle, Paris.
    Copyright © Thierry Deuve
    Carabus (Mimocarabus) maurus paphius Redtenbacher, 1843
    Carabus (Mimocarabus) maurus paphius Redtenbacher, 1843 Lectotype
    Male+Female adult
    Lectotype female (left) from collection Kotschy (Muséum de Vienne); right - « Chypre » Muséum de Paris
    Copyright © Thierry Deuve
    Carabus (Apotomopterus) patroclus patroclus Breuning, 1933
    Carabus (Apotomopterus) patroclus patroclus Breuning, 1933
    Male adult
    China, S Sichuan, Pass 56 km SW Miannin, near Wawo h=2440 m, 21-24. VI.2017, C.Reuter leg./det. F.Kleinfeld, L=29 mm
    Copyright © Ruslan Panin
    Carabus (Apotomopterus) bousquetellus bousquetellus Deuve, 1998
    Carabus (Apotomopterus) bousquetellus bousquetellus Deuve, 1998
    Male adult
    S-China, CW-Yunnan, W of Lincang, Daxue Shan h=2600m, 23.935556, 100.257778, 7-13.VI.2023, C.Reuter leg./det. F.Kleinfeld, L=24 mm
    Copyright © Ruslan Panin
    Carabus (Semnocarabus) erosus karascharensis Eidam, 1931
    Carabus (Semnocarabus) erosus karascharensis Eidam, 1931 Lectotype Paralectotypes
    Male adult
    Lectotype, left: “Ost Turkestan, Bagratsch-kul, Kurla 5.1902. Coll. Hsr,” “Syntypus,” “O. Leonhard,” “erosus subsp. karascharensis type Eidam, det. P. Eidam;” “coll DEI, Eberswalde;” “coll DEI, Müncheberg Col.–07059”
    Copyright © Kirill Makarov
    Carabus (Semnocarabus) erosus karascharensis Eidam
    Carabus (Semnocarabus) erosus karascharensis Eidam Lectotype Paralectotypes
    Endophallus
    aedeagus - lectotype. Endophallus - paralectotype.
    Copyright © Kirill Makarov
    Carabus (Semnocarabus) cicatricosulus pseudoerosus Mandl, 1955
    Carabus (Semnocarabus) cicatricosulus pseudoerosus Mandl, 1955
    Endophallus
    left tributary of Kuqa River, E of Lake Kyrghyz-Artukul (= Dalongchi), 42°27'15''N, 83°22'00''E, H ~ 2550–2600 m, 7–8.VI.2002, I. Kabak leg
    Copyright © Kirill Makarov
    Carabus (Alipaster) semenoviellus Breuning, 1934
    Carabus (Alipaster) semenoviellus Breuning, 1934 Holotype
    Male adult
    (NBC), «Tekkes-tal.», «semenoviellus t. Breuning c.», «collectie C. & O. Vogt Acq. 1960», «Typus», «Holotypus», «C. (Alipaster) semenoviellus Br. Gottwald det. 1984», «Alipaster semenoviellus Breuning, 1934 ZMAN type COLE. 0375.1».
    Copyright © Ilya Kabak
    Carabus (Alipaster) semenoviellus Breuning, 1934
    Carabus (Alipaster) semenoviellus Breuning, 1934
    Male adult
    ESE of Tekes, Tagymbel, S of Karatogai Vill., 43°09ʹ19ʺN / 82°14ʹ45ʺE, 2200 m, 10.07.2014 (I.I. Kabak leg.)
    Copyright © Ilya Kabak
    Carabus (Alipaster) semenoviellus Breuning, 1934
    Carabus (Alipaster) semenoviellus Breuning, 1934
    Male genitalia

    Copyright © Ilya Kabak
    Carabus (Ophiocarabus) successor eoketmenicus Kabak, 2016
    Carabus (Ophiocarabus) successor eoketmenicus Kabak, 2016 Holotype Paratype
    Endophallus
    endophallus - paratypes; aedeagus - holotype
    Copyright © Ilya Kabak
    Carabus (Ophiocarabus) successor eoketmenicus Kabak, 2016
    Carabus (Ophiocarabus) successor eoketmenicus Kabak, 2016 Holotype
    Male adult
    China, Xinjiang, Wusunshan Mt. Range, E part, NE of Tekes Town, 43°20ʹ26″N / 81°59ʹ48″E, 2285 m, 27.07.2016 (I.I. Kabak leg). (ZIN)
    Copyright © Ilya Kabak
    Carabus (Ophiocarabus) kurdaiensis Kabak, 2016
    Carabus (Ophiocarabus) kurdaiensis Kabak, 2016 Paratype
    Female adult
    China, Xinjiang, Narat Mt. Range, Ekiasha River, near Kurdai Pass, 42°54ʹ21″N / 82°28ʹ30″E, 3640 m, 26.07.2013 (I.I. Kabak leg.).
    Copyright © Ilya Kabak
    Carabus (Ophiocarabus) kurdaiensis Kabak, 2016
    Carabus (Ophiocarabus) kurdaiensis Kabak, 2016 Paratype
    Endophallus

    Copyright © Ilya Kabak
    Carabus (Ophiocarabus) ernsti ernsti Kabak, 2001
    Carabus (Ophiocarabus) ernsti ernsti Kabak, 2001
    Male adult
    Left from the locality in right tributary of the Saryk River; Right – from the locality in mountains to the West of the Saryk River.
    Copyright © Ilya Kabak
    Carabus (Ophiocarabus) ernsti ulastaiensis Kabak, 2019
    Carabus (Ophiocarabus) ernsti ulastaiensis Kabak, 2019 Paratype
    Male+Female adult
    China, Xinjiang, S slopes of Boro-Horo Mt. R., ridge between Kaptshik and Zekku rivers, NE of Ulastai (= Wulasitai) Vill., alpine meadows, 43°56ʹ07ʺN / 83°18ʹ33ʺE – 43°56ʹ22ʺN / 83°18ʹ30ʺE, 3135–3280 m, 17.07.2017 (I.I. Kabak leg.)
    Copyright © Ilya Kabak
    Carabus (Ophiocarabus) ernsti Kabak, 2001
    Carabus (Ophiocarabus) ernsti Kabak, 2001
    Endophallus
    7–8, 12–13 – C. ernsti ulastaiensis, paratypes; 9–10, 14–15 – C. ernsti ernsti Kabak, 2002: 9, 14 – from the locality in mountains to the West of the Saryk River, 10, 15 – from the locality in right tributary of the Saryk River; 11,16 – C. ernsti nilkiensis Kabak, 2014: 11 – holotype, 12 – paratype. ml – median lobe; pp – praeputial pad; rbll – right basolateral lobe.
    Copyright © Ilya Kabak
    Carabus (Alipaster) semenoviellus tokkuztaraensis Kabak, 2019
    Carabus (Alipaster) semenoviellus tokkuztaraensis Kabak, 2019 Holotype Paratype
    Male adult
    Holotype, left: China, Xinjiang, Narat Mt. R., Chashi Valley, Tuyukasha, 43°00ʹ29ʺN / 82°46ʹ02ʺE, 2275 m, 18.07.2014 (I.I. Kabak leg.); Paratypes: middle: China, Xinjiang, Narat Mt. R., Erbotu Valley, 43°06ʹ14ʺN / 83°00ʹ05ʺE, 1915 m, 21.07.2014 (I.I. Kabak leg.). Right: China, Xinjiang, Narat Mt. R., Aktuyuk River, S of Kurdelin, 43°04ʹ12ʺN / 82°51ʹ19ʺE, 2255 m, 31.07.2013 (I.I. Kabak leg.)
    Copyright © Ilya Kabak
    Carabus (Acoptolabrus) constricticollis Kraatz, 1886
    Carabus (Acoptolabrus) constricticollis Kraatz, 1886
    Male adult
    China, Liaoning prov., Mt. Guanmen 20.6.2009 (cAA)
    Copyright © Alexander Anichtchenko
    Carabus (Coptolabrus) elysii elysii Thomson, 1856 (as idai)
    Carabus (Coptolabrus) elysii elysii Thomson, 1856 (as idai)
    Female adult
    China, Qinling, Zhouzhi co., Shaanxi pr., 8-14.6.2009 (cAA).
    Copyright © Alexander Anichtchenko
    Carabus (Coptolabrus) lafossei tungchengensis Li, 1993
    Carabus (Coptolabrus) lafossei tungchengensis Li, 1993
    Male adult
    China, Hubei, Shangzhi co., Shimenyan 8-11.6.2009 (cAA) 42 mm
    Copyright © Alexander Anichtchenko
    Carabus (Coptolabrus) lafossei tungchengensis Li, 1993
    Carabus (Coptolabrus) lafossei tungchengensis Li, 1993
    Endophallus
    China, Hubei, Shangzhi co., Shimenyan 8-11.6.2009 (cAA)
    Copyright © Alexander Anichtchenko
    Carabus (Coptolabrus) gemmifer mesites Fairmaire, 1897
    Carabus (Coptolabrus) gemmifer mesites Fairmaire, 1897
    Female adult
    China, Sichuan, Xichang City, Muli Co., Ke-Er-xiang 2-12.7.2009 (cAA) 32 mm
    Copyright © Alexander Anichtchenko
    Carabus (Coptolabrus) ignimitella angulicollis Hauser, 1911
    Carabus (Coptolabrus) ignimitella angulicollis Hauser, 1911
    Male adult
    China Hunan, Guidong pr., 20.5.2009 (cAA) 44 mm
    Copyright © Alexander Anichtchenko
    Carabus (Coptolabrus) augustus ignigena Hauser, 1914
    Carabus (Coptolabrus) augustus ignigena Hauser, 1914
    Endophallus

    Copyright © Alexander Anichtchenko
    Carabus (Acoptopterus) rautenstrauchi alberti Rapuzzi, 2024
    Carabus (Acoptopterus) rautenstrauchi alberti Rapuzzi, 2024 Holotype
    Male adult
    Sichuan, Gu Dui Xiang, Meigu to Shanlenggang; 2250 m asl; Jul. 2022
    Copyright © - - -
    Carabus (Pseudocoptolabrus) taliensis shanlenggang Rapuzzi, 2024
    Carabus (Pseudocoptolabrus) taliensis shanlenggang Rapuzzi, 2024 Holotype
    Male adult
    Sichuan, 30 km behind Meigu to Shanglenggang; 2700 m; Jul. 2022
    Copyright © Alexander Anichtchenko
    Carabus (Coptolabrus) gemmifer rectipustulatus Rapuzzi, 2024
    Carabus (Coptolabrus) gemmifer rectipustulatus Rapuzzi, 2024 Holotype
    Male adult
    Sichuan, Yanyuan to Wodixiang; 2700 m asl; 16 Aug. 2022
    Copyright © - - -
    Carabus (Tribax) puschkini kraatzi Chaudoir, 1877
    Carabus (Tribax) puschkini kraatzi Chaudoir, 1877
    Endophallus
    Georgia, Samtskhe-Javakheti, env. Uraveli vill., 14.VI.2024 leg. D. Fominykh
    Copyright © Igor Solodovnikov
    Carabus (Tribax) puschkini kraatzi Chaudoir, 1877
    Carabus (Tribax) puschkini kraatzi Chaudoir, 1877
    Female adult
    Georgia, Samtskhe-Javakheti, env. Uraveli vill., 14.VI.2024 leg. D. Fominykh
    Copyright © Igor Solodovnikov
    Carabus (Tribax) puschkini kraatzi Chaudoir, 1877
    Carabus (Tribax) puschkini kraatzi Chaudoir, 1877
    Male adult
    Georgia, Samtskhe-Javakheti, env. Uraveli vill., 14.VI.2024 leg. D. Fominykh
    Copyright © Igor Solodovnikov
    Carabus (Sphodristocarabus) varians armeniacus Mannerheim, 1830
    Carabus (Sphodristocarabus) varians armeniacus Mannerheim, 1830
    Male adult
    C Georgia, Imereti, Caucasus, Likhi Mt Range, Rikoti Pass h=900-1000 m, 14-16.V.2017, E. Rutjan E. leg., L=22 mm
    Copyright © Ruslan Panin
    Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
    Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
    Female adult
    W Ukraine, Lviv city, Holosko, Bryukhovytsʹkyi forest 49.886833, 24.012083 under bark, 10.XI.2024, R. Panin leg., L=35 mm
    Copyright © Ruslan Panin
    Carabus (Piocarabus) titanus Breuning, 1933
    Carabus (Piocarabus) titanus Breuning, 1933
    Female adult
    China Sichuan prov., Nanping, h=1500m, VI.2000, S. Remond leg.
    L=38 mm
    Copyright © Ruslan Panin
    Carabus taxon page (3801 photos)
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