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Carabidae of the World
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Carabinae Latreille, 1802
→
Carabini Latreille, 1802
→
Carabina sensu stricto
→
Carabus Linne, 1758
→
Pagocarabus Morawitz, 1886
→ crassesculptus Kraatz, 1881
Region
Afghanistan (AF)
Albania (AL)
Aleutian isl (ALU)
Algeria (AG)
Andorra (AN)
Angola (ANG)
Antarctica (Antarctica)
Antilles islands (ANT)
Antigua and Barbuda (Antigua and Barbuda)
Barbados (Barbados)
Cayman islands (Cayman islands)
Cuba (Cuba)
Dominica (Lesser Antilles) (Dominica (Lesser Antilles))
Dominican Republic (Hispaniola, Greater Antilles) (Dominican Republic (Greater Antilles))
Grenada (Grenada)
Guadeloupe (Guadeloupe)
Haiti (Haiti)
Jamaica (Jamaica)
Martinique (Martinique)
Puerto Rico (Puerto Rico)
Saint Kitts and Nevis (Saint Kitts and Nevis)
Saint Lucia (Saint Lucia)
St. Vincent (St. Vincent)
Trinidad and Tobago (Trinidad and Tobago)
Arab Emirates (AE)
Argentina (ARG)
Armenia (AR)
Australia (AST)
Christmas Island (Christmas Island)
Lord Howe Island (Lord Howe Island)
Macquarie Island (Macquarie Island)
Tasmania (Tasmania)
Austria (AU)
Azerbaijan (AB)
Azores (AZ)
BALTIC AMBER, Baltic See coast and Amber quarry Jantarny near Kaliningrad, Russia (BAM)
Bahamas (BHM)
Bahrain (BA)
Bangladesh (BGD)
Belgium (BE)
Belize (BLZ)
Bhutan (BT)
Bolivia (BOL)
Bosnia and Herzegovina (BH)
Botswana (BTW)
Brazil (BRZ)
Acre (AC)
Alagoas (AL)
Amazonas (AM)
Arnapá (AP)
Bahia (BA)
Ceará (CE)
District Federal (Brasilia) (DF)
Espirito Santo (ES)
Goiás (GO)
Maranhão (MA)
Mato Grosso (MT)
Mato Grosso do Sul (MS)
Minas Gerais (MG)
Paraná (PR)
Paraîba (PB)
Pará (PA)
Pernambouc (PE)
Piaui (PI)
Rio Grande do Norte (RN)
Rio Grande do Sul (RS)
Rio de Janeiro (RJ)
Rondônia (RO)
Roraima (RR)
Santa Catarina (SC)
Sergipe (SE)
São Paulo (SP)
Tocantins (TO)
Brunei (Brunei)
Bulgaria (BU )
Burundi (BUR)
Byelorussia (BY)
Cambodia (CBD)
Cameroon (CAM)
Canada (CAN)
Canary isls (CI)
Cape Verde (Cape Verde)
Central African Republic (CAR)
Chad (CHD)
Chile (CHL)
Juan Fernández Islands (Juan Fernández Islands)
China (CH)
Anhui (Anhui)
Beijing (Beijing)
Chongqing (Chongqing)
Fujian (Fujian)
Gansu (Gansu)
Guangdong (Guangdong)
Guangxi (Guangxi)
Guizhou (Guizhou)
Hainan (Hainan)
Hebei (Hebei)
Heilongjiang (Heilongjiang)
Henan (Henan)
Honkong (Hongkong)
Hubei (Hubei)
Hunan (Hunan)
Inner Mongolia (Inner Mongolia)
Jiangsu (Jiangsu)
Jiangxi (Jiangxi)
Jilin (Jilin)
Liaoning (Liaoning)
Macau (Macau)
Ningxia (Ningxia)
Qinghai (Qinghai)
Shaanxi (Shaanxi)
Shandong (Shandong)
Shanghai (Shanghai)
Shanxi (Shanxi)
Sichuan (Sichuan)
Tianjin (Tsiensin) (Tianjin)
Xinjiang (Xinjiang)
Xizang (Tibet) (Xizang (Tibet))
Yunnan (Yunnan)
Zhejiang (Zhejiang)
Colombia (COL)
Comoros isl. (COM)
Costa Rica (CRC)
Croatia (CR)
Crozet Islands (CRZ)
Cyprus (CY)
Czech (CZ)
DOMINICAN AMBER, Dominican Republic, Lower Miocene. (DAM)
Democratic Republic of the Congo (CNG)
Denmark (DE)
Diego Garcia isls (DGC)
Djibouti (DJB)
Ecuador (ECD)
Egypt (EG)
Sinai (Sinai)
El Salvador (SVD)
Equatorial Guinea (EQG)
Bioco (Fernando Pó) island (Bioco (Fernando Pó) island)
Eritrea (ERT)
Estonia (EN)
Ethiopia (ETP)
Falkland il. (FLK)
Faroe Islands (FA)
Finland (FI)
France (FR)
Corse (Corse)
French Guiana (GNF)
Gabon (GAB)
Galapagos Islands (GAL)
Georgia (GG)
Abkhazia (Abkhazia)
Ajaria (Ajaria)
South Ossetia (South Ossetia)
Germany (GE)
Great Britain (GB)
Greece (GR)
Aegean Islands (Aegean Islands)
Crete (Crete)
Dodecanese Islands (Dodecanese Islands)
Euboea (Evia) Island (Euboea (Evia) Island)
Ionian Islands (Ionian Islands)
Peloponnese (Peloponnese)
Saronic Islands (Saronic Islands)
Sporades Islands (Sporades Islands)
The Cyclades (The Cyclades)
Guatemala (GTM)
Guyana (GYN)
Honduras (HDR)
Hungary (HU)
Iceland (IC)
India (IND)
Andaman and Nicobar isl. (Andaman and Nicobar isl.)
Andhra Pradesh (Andhra Pradesh)
Arunachal Pradesh (Arunachal Pradesh)
Assam (Assam)
Bihar (Bihar)
Chhattisgarh (Chhattisgarh)
Delhi (Delhi)
Goa (Goa)
Gujarat (Gujarat)
Haryana (Haryana)
Himachal Pradesh (Himachal Pradesh)
Jammu and Kashmir (Jammu and Kashmir)
Jharkhand (Jharkhand)
Karnataka (Karnataka)
Kerala (Kerala)
Ladakh (Ladakh)
Madhya Pradesh (Madhya Pradesh)
Maharashtra (Maharashtra)
Manipur (Manipur)
Meghalaya (Meghalaya)
Mizoram (Mizoram)
Nagaland (Nagaland)
Odisha (Odisha)
Puducherry (Puducherry)
Punjab (Punjab)
Rajasthan (Rajasthan)
Sikkim (Sikkim)
Tamil Nadu (Tamil Nadu)
Telangana (Telangana)
Tripura (Tripura)
Uttar Pradesh (Uttar Pradesh)
Uttarakhand (Uttarakhand)
West Bengal (West Bengal)
Indonesia (IDS)
Borneo (Kalimantan) (Borneo)
Java (Java)
Lesser Sunda Islands (Lesser Sunda Ils.)
Maluku Islands (Maluku Isl.)
Papua (Papua)
Riau Islands (Riau Islands)
Sulawesi (Celebes) (Sulawesi)
Sumatra (Sumatra)
Timor (Timor)
Iran (IN)
Iraq (IQ)
Ireland (IR)
Israel (IS)
Italy (IT)
Sardegna (Sardegna)
Sicily (Sicily)
Japan (JA)
Hokkaido (Hokkaido)
Honshu (Honshu)
Kyushu (Kyushu)
Ryukyu Islands (Ryukyu Islands)
Shikoku (Shikoku)
Jordan (JO)
Kashmir (KA)
Kazakhstan (KZ)
Kenya (KEN)
Kerguelen Islands (Kerguelen Islands)
Kuwait (KU)
Kyrgyzstan (KI)
Laos (LAO)
Latvia (LA)
Lebanon (LE)
Lesotho (LSO)
Libya (LB)
Liechtenstein (LS)
Lithuania (LT)
Luxenbourg (LU)
Macedonia (MC)
Madagascar (MAD)
Madeira Arch. (MR)
Malawi (MLW)
Malaysia (MLS)
Sabah (Borneo) (Sabah (Borneo))
Sarawak (Borneo) (Sarawak (Borneo))
Maldives (MLD)
Malta (MA)
Mauritania (MAU)
Mauritius (MRC)
Melanesia (Melanesia)
Bismarck Archipelago (Bismarck Archipelago)
Fiji (Fiji)
New Caledonia (New Caledonia)
Papua New Guinea (Papua New Guinea)
Solomon Islands (Solomon Islands)
Vanuatu ils (Vanuatu ils)
Mexico (MEX)
Aguascalientes (Aguascalientes)
Baja California (Baja California)
Baja California Sur (Baja California Sur)
Campeche (Campeche)
Chiapas (Chiapas)
Chihuahua (Chihuahua)
Coahuila (Coahuila)
Colima (Colima)
Durango (Durango)
Federal District (Federal District)
Guanajuato (Guanajuato)
Guerrero (Guerrero)
Hidalgo (Hidalgo)
Jalisco (Jalisco)
Mexico State (Mexico State)
Michoacan (Michoacan)
Morelos (Morelos)
Nayarit (Nayarit)
Nuevo Leon (Nuevo Leon)
Oaxaca (Oaxaca)
Puebla (Puebla)
Queretaro (Queretaro)
Quintana Roo (Quintana Roo)
San Luis Potosi (San Luis Potosi)
Sinaloa (Sinaloa)
Sonora (Sonora)
Tabasco (Tabasco)
Tamaulipas (Tamaulipas)
Tlaxcala (Tlaxcala)
Veracruz (Veracruz)
Yucatan (Yucatan)
Zacatecas (Zacatecas)
Micronesia (MIC)
Caroline islands (Caroline islands)
Guam (Guam)
Kiribati (Kiribati)
Marshall ils (Marshall ils)
North Marians isl. (North Marians isl.)
Palau archipelago (Palau archipelago)
Moldavia (MD)
Mongolia (MG)
Montenegro (ME)
Mordor (RU)
Altai Republic and Altaisky Province (Altai Republic and Altaisky Province)
Amurskaya Area (Amurskaya Area)
Bashkiria, E of Chelyabinsk regs. (S Ural mts. Bashkiria, E of Chelyabinsk regs.)
Buryat Republic (Buryat Republic)
Chukot autonomous region (Chukotka)
East Sayan Mountains (East Sayan Mountains)
Ingushetia, Chechnya, Daghestan (E Caucas) (Ingushetia, Chechnya, Daghestan (E Caucas))
Irkutsk Area (Irkutsk Area)
Kabardino-Balkaria, Ossetia (C Caucas) (Kabardino-Balkaria, Ossetia (C Caucas))
Kaliningrad, Briansk, Smolensk, Tula, Orlov, Kaluga, Moskow, Riaz (C of Russian plains)
Kamchatskaya Area (Kamchatka)
Karachaevo-Cherkessia (W Caucas) (Karachaevo-Cherkessia)
Karelia, Leningrad reg., Pskov reg., Kostroma reg., Perm reg., Komi, Pechora plains. (N of Russian plains)
Khabarovsky Province (Khabarovsky Province)
Khamar-Daban Mountains (Khamar-Daban Mountains (E Siberia))
Komi, Yamalo-Nenetsk, Hanty-Mansi (N Ural mts.)
Krasnodar reg. (W Caucas)
Krasnoyarsky Province (Krasnoyarsk)
Kuriles Islands (Kuriles Islands)
Magadanskaya Area (Magadan)
Omsk, S of Tyumen, W Novosibirsk regs. (S of W Siberia)
Perm, Sverdlovsk regs. (C Ural mts)
Primorsky District (Primorie) (Maritime Prov)
Sakhalin Island (Sakhalin Island)
Tuva Republic (Tuva)
Voronezh, Rostov, Saratov, Kalmykia, Volgograd, Astrahan regs. (S of Russian plains)
West Sayan Mountains (West Sayan)
Yakut-Saha Republic (Yakut-Saha Republic)
Zabaykalsky Krai (ex Chita reg.) (Zabaykalsky Krai)
plains of Krasnodar reg., S Rostov reg., Stavropol reg. (Ciscaucas reg. (plains of Krasnodar reg., S Rostov)
taiga from Ural mts to Yenisei riv. (Middle of W Siberia)
tundra of Tyumen, Yamalo-Nenetsk regs. (N of W Siberia)
Morocco (MO)
Mozambique (MZB)
Myanmar (Burma) (BRM)
Namibia (NAM)
Nepal (NP)
Netherlands (NL)
New Zealand (NZL)
Antipodes Islands (Antipodes Islands)
Auckland Islands (Auckland Islands)
Campbell Island (Campbell Island)
Chatham Islands (Chatham Islands)
Nicaragua (NCG)
North Korea (NC)
Norway (NR)
Svalbard (Spitzbergen) (SB)
Oman (OM)
Pakistan (PA)
Panama (PNM)
Paraguay (PRG)
Peru (PER)
Philippines (PLP)
Basilan (Basilan)
Cebu (Cebu)
Guimaras (Guimaras)
Leyte (Leyte)
Luzon (Luzon)
Marinduque (Marinduque)
Masbate (Masbate)
Mindanao (Mindanao)
Mindoro (Mindoro)
Negros (Negros)
Palawan (Palawan)
Panay (Panay)
Polillo Islands (Polillo Islands)
Romblon (Romblon)
Samar (Samar)
Sulu Archipelago (Sulu Archipelago)
Poland (PL)
Polynesia (POL)
Cook Islands (Cook Islands)
Hawaiian Islands (HAW)
Marquesas Islands (Marquesas Islands)
Niue (Niue)
Norfolk ils (Norfolk ils)
Pitcairn ils (Pitcairn ils)
Rapa Nui (Easter Island) (Rapa Nui)
Samoa (Samoa)
Society Islands (Society Islands)
Tahiti (Tahiti)
Tokelau (TKL)
Tonga (Tonga)
Tuvalu ils (Tuvalu ils)
Portugal (PT)
Qatar (QA)
Republic of South Africa (RSA)
Republic of the Congo (CG)
Reunion isl. (Reunion isl.)
Rodriguez isls. (ROD)
Romania (RO)
Rwanda (RWA)
Santa Helena isl (SH)
Sao Tome e Principe (STM)
Saudi Arabia (SA)
Serbia (SRB)
Seychelles (SEY)
Singapore (SGP)
Slovakia (SK)
Slovenia (SL)
Somalia (SOM)
South Georgia and the South Sandwich Islands (SGSSI)
South Korea (SC)
Spain (SP)
Balearic Islands (Balearic Islands)
Sri Lanka (SRL)
St. Paul isls (STP)
Sudan (SUD)
Surinam (SUR)
Swaziland (SWZ)
Sweden (SV)
Switzerland (SZ)
Syria (SY)
Tadzhikistan (TD)
Taiwan (Formosa) (Taiwan)
Tanzania (TAN)
Zanzibar isl. (Zanzibar isl.)
Thailand (TAI)
Tunisia (TU)
Turkey (TR)
Turkmenistan (TM)
Uganda (UGD)
Ukraine (UK)
Crimea (Crimea)
United States (USA)
Alabama (Alabama)
Alaska (Alaska)
Arizona (Arizona)
Arkansas (Arkansas)
California (California)
Colorado (Colorado)
Connecticut (Connecticut)
Delaware (Delaware)
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Virginia (Virginia)
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West Virginia (West Virginia)
Wisconsin (Wisconsin)
Wyoming (Wyoming)
Uruguay (URG)
Uzbekistan (UZ)
Venezuela (VNZ)
Vietnam (VTN)
West Africa (WAF)
Benin (Benin)
Burkina Faso (Burkina Faso)
Côte d'Ivoire (Côte d'Ivoire)
Gambia (Gambia)
Ghana (Ghana)
Guinea (Guinea)
Guinea Bissau (Guinea Bissau)
Liberia (Liberia)
Mali (Mali)
Niger (Niger)
Nigeria (Nigeria)
Senegal (Senegal)
Sierra Leone (Sierra Leone)
Togo (Togo)
Yemen (YE)
Socotra (Socotra)
Zambia (ZMB)
Zimbabwe (ZIM)
Carabus (Pagocarabus) crassesculptus crassesculptus Kraatz, 1881
Male adult
China, Beijing, Wulingshan, 2007, Liu ye leg.;
Photo & det. by liuye
Copyright ©
Ye Liu
Carabus (Pagocarabus) crassesculptus crassesculptus Kraatz, 1881
Male adult
China, Hebei, Xianhua (=Kalgan)
10-15.5.1995
21 mm
Copyright ©
Alexander Anichtchenko
crassesculptus
taxon page
(2 photos)
Carabus (Pagocarabus) crassethoracis Kleinfeld, 1999
Holotype
Male adult
Copyright ©
Frank Kleinfeld
crassethoracis
taxon page
(1 photos)
Carabus (Pagocarabus) crassesculptus diruptus Morawitz, 1886
Male adult
China, Gansu, Umg. Xiahe 3000-3200m
24-26.6.1993 Heinz leg.
20 mm
Copyright ©
Alexander Anichtchenko
diruptus
taxon page
(1 photos)
Carabus (Pagocarabus) crassesculptus eleganticolor Deuve, 1997
Paratype
Male adult
China, Gansu, Min Shan mts., 70 km NW Wudu
Copyright ©
Alexander Anichtchenko
Carabus (Pagocarabus) crassesculptus eleganticolor Deuve, 1997
Female adult
S Gansu, Min Shan, 5-20 km S Cagan, 2300-3200m 7-20.7.2019 V. Patrikeev leg. (cAA). det. Anichtchenko A. 2020
Copyright ©
Alexander Anichtchenko
rugosissimus
taxon page
(2 photos)
Carabus (Pagocarabus) crassesculptus tonghua Deuve & Mourzine, 2001
Male adult
China, NW Sishuan, 3500m, Hongyuan
Somang Qu riv. grassland, 9-29.6.2004
male, 20 mm
Copyright ©
Alexander Anichtchenko
tonghua
taxon page
(1 photos)
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