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Carabidae of the World
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Harpalinae Bonelli, 1810
→ Harpalini Bonelli, 1810
Region
Afghanistan (AF)
Albania (AL)
Aleutian isl (ALU)
Algeria (AG)
Andorra (AN)
Angola (ANG)
Antarctica (Antarctica)
Antilles islands (ANT)
Antigua and Barbuda (Antigua and Barbuda)
Barbados (Barbados)
Cayman islands (Cayman islands)
Cuba (Cuba)
Dominica (Lesser Antilles) (Dominica (Lesser Antilles))
Dominican Republic (Hispaniola, Greater Antilles) (Dominican Republic (Greater Antilles))
Grenada (Grenada)
Guadeloupe (Guadeloupe)
Haiti (Haiti)
Jamaica (Jamaica)
Martinique (Martinique)
Puerto Rico (Puerto Rico)
Saint Kitts and Nevis (Saint Kitts and Nevis)
Saint Lucia (Saint Lucia)
St. Vincent (St. Vincent)
Trinidad and Tobago (Trinidad and Tobago)
Arab Emirates (AE)
Argentina (ARG)
Armenia (AR)
Australia (AST)
Christmas Island (Christmas Island)
Lord Howe Island (Lord Howe Island)
Macquarie Island (Macquarie Island)
Tasmania (Tasmania)
Austria (AU)
Azerbaijan (AB)
Azores (AZ)
BALTIC AMBER, Baltic See coast and Amber quarry Jantarny near Kaliningrad, Russia (BAM)
Bahamas (BHM)
Bahrain (BA)
Bangladesh (BGD)
Belgium (BE)
Belize (BLZ)
Bhutan (BT)
Bolivia (BOL)
Bosnia and Herzegovina (BH)
Botswana (BTW)
Brazil (BRZ)
Acre (AC)
Alagoas (AL)
Amazonas (AM)
Arnapá (AP)
Bahia (BA)
Ceará (CE)
District Federal (Brasilia) (DF)
Espirito Santo (ES)
Goiás (GO)
Maranhão (MA)
Mato Grosso (MT)
Mato Grosso do Sul (MS)
Minas Gerais (MG)
Paraná (PR)
Paraîba (PB)
Pará (PA)
Pernambouc (PE)
Piaui (PI)
Rio Grande do Norte (RN)
Rio Grande do Sul (RS)
Rio de Janeiro (RJ)
Rondônia (RO)
Roraima (RR)
Santa Catarina (SC)
Sergipe (SE)
São Paulo (SP)
Tocantins (TO)
Brunei (Brunei)
Bulgaria (BU )
Burundi (BUR)
Byelorussia (BY)
Cambodia (CBD)
Cameroon (CAM)
Canada (CAN)
Canary isls (CI)
Cape Verde (Cape Verde)
Central African Republic (CAR)
Chad (CHD)
Chile (CHL)
Juan Fernández Islands (Juan Fernández Islands)
China (CH)
Anhui (Anhui)
Beijing (Beijing)
Chongqing (Chongqing)
Fujian (Fujian)
Gansu (Gansu)
Guangdong (Guangdong)
Guangxi (Guangxi)
Guizhou (Guizhou)
Hainan (Hainan)
Hebei (Hebei)
Heilongjiang (Heilongjiang)
Henan (Henan)
Honkong (Hongkong)
Hubei (Hubei)
Hunan (Hunan)
Inner Mongolia (Inner Mongolia)
Jiangsu (Jiangsu)
Jiangxi (Jiangxi)
Jilin (Jilin)
Liaoning (Liaoning)
Macau (Macau)
Ningxia (Ningxia)
Qinghai (Qinghai)
Shaanxi (Shaanxi)
Shandong (Shandong)
Shanghai (Shanghai)
Shanxi (Shanxi)
Sichuan (Sichuan)
Tianjin (Tsiensin) (Tianjin)
Xinjiang (Xinjiang)
Xizang (Tibet) (Xizang (Tibet))
Yunnan (Yunnan)
Zhejiang (Zhejiang)
Colombia (COL)
Comoros isl. (COM)
Costa Rica (CRC)
Croatia (CR)
Crozet Islands (CRZ)
Cyprus (CY)
Czech (CZ)
DOMINICAN AMBER, Dominican Republic, Lower Miocene. (DAM)
Democratic Republic of the Congo (CNG)
Denmark (DE)
Diego Garcia isls (DGC)
Djibouti (DJB)
Ecuador (ECD)
Egypt (EG)
Sinai (Sinai)
El Salvador (SVD)
Equatorial Guinea (EQG)
Bioco (Fernando Pó) island (Bioco (Fernando Pó) island)
Eritrea (ERT)
Estonia (EN)
Ethiopia (ETP)
Falkland il. (FLK)
Faroe Islands (FA)
Finland (FI)
France (FR)
Corse (Corse)
French Guiana (GNF)
Gabon (GAB)
Galapagos Islands (GAL)
Georgia (GG)
Abkhazia (Abkhazia)
Ajaria (Ajaria)
South Ossetia (South Ossetia)
Germany (GE)
Great Britain (GB)
Greece (GR)
Aegean Islands (Aegean Islands)
Crete (Crete)
Dodecanese Islands (Dodecanese Islands)
Euboea (Evia) Island (Euboea (Evia) Island)
Ionian Islands (Ionian Islands)
Peloponnese (Peloponnese)
Saronic Islands (Saronic Islands)
Sporades Islands (Sporades Islands)
The Cyclades (The Cyclades)
Guatemala (GTM)
Guyana (GYN)
Honduras (HDR)
Hungary (HU)
Iceland (IC)
India (IND)
Andaman and Nicobar isl. (Andaman and Nicobar isl.)
Andhra Pradesh (Andhra Pradesh)
Arunachal Pradesh (Arunachal Pradesh)
Assam (Assam)
Bihar (Bihar)
Chhattisgarh (Chhattisgarh)
Delhi (Delhi)
Goa (Goa)
Gujarat (Gujarat)
Haryana (Haryana)
Himachal Pradesh (Himachal Pradesh)
Jammu and Kashmir (Jammu and Kashmir)
Jharkhand (Jharkhand)
Karnataka (Karnataka)
Kerala (Kerala)
Ladakh (Ladakh)
Madhya Pradesh (Madhya Pradesh)
Maharashtra (Maharashtra)
Manipur (Manipur)
Meghalaya (Meghalaya)
Mizoram (Mizoram)
Nagaland (Nagaland)
Odisha (Odisha)
Puducherry (Puducherry)
Punjab (Punjab)
Rajasthan (Rajasthan)
Sikkim (Sikkim)
Tamil Nadu (Tamil Nadu)
Telangana (Telangana)
Tripura (Tripura)
Uttar Pradesh (Uttar Pradesh)
Uttarakhand (Uttarakhand)
West Bengal (West Bengal)
Indonesia (IDS)
Borneo (Kalimantan) (Borneo)
Java (Java)
Lesser Sunda Islands (Lesser Sunda Ils.)
Maluku Islands (Maluku Isl.)
Papua (Papua)
Riau Islands (Riau Islands)
Sulawesi (Celebes) (Sulawesi)
Sumatra (Sumatra)
Timor (Timor)
Iran (IN)
Iraq (IQ)
Ireland (IR)
Israel (IS)
Italy (IT)
Sardegna (Sardegna)
Sicily (Sicily)
Japan (JA)
Hokkaido (Hokkaido)
Honshu (Honshu)
Kyushu (Kyushu)
Ryukyu Islands (Ryukyu Islands)
Shikoku (Shikoku)
Jordan (JO)
Kashmir (KA)
Kazakhstan (KZ)
Kenya (KEN)
Kerguelen Islands (Kerguelen Islands)
Kuwait (KU)
Kyrgyzstan (KI)
Laos (LAO)
Latvia (LA)
Lebanon (LE)
Lesotho (LSO)
Libya (LB)
Liechtenstein (LS)
Lithuania (LT)
Luxenbourg (LU)
Macedonia (MC)
Madagascar (MAD)
Madeira Arch. (MR)
Malawi (MLW)
Malaysia (MLS)
Sabah (Borneo) (Sabah (Borneo))
Sarawak (Borneo) (Sarawak (Borneo))
Maldives (MLD)
Malta (MA)
Mauritania (MAU)
Mauritius (MRC)
Melanesia (Melanesia)
Bismarck Archipelago (Bismarck Archipelago)
Fiji (Fiji)
New Caledonia (New Caledonia)
Papua New Guinea (Papua New Guinea)
Solomon Islands (Solomon Islands)
Vanuatu ils (Vanuatu ils)
Mexico (MEX)
Aguascalientes (Aguascalientes)
Baja California (Baja California)
Baja California Sur (Baja California Sur)
Campeche (Campeche)
Chiapas (Chiapas)
Chihuahua (Chihuahua)
Coahuila (Coahuila)
Colima (Colima)
Durango (Durango)
Federal District (Federal District)
Guanajuato (Guanajuato)
Guerrero (Guerrero)
Hidalgo (Hidalgo)
Jalisco (Jalisco)
Mexico State (Mexico State)
Michoacan (Michoacan)
Morelos (Morelos)
Nayarit (Nayarit)
Nuevo Leon (Nuevo Leon)
Oaxaca (Oaxaca)
Puebla (Puebla)
Queretaro (Queretaro)
Quintana Roo (Quintana Roo)
San Luis Potosi (San Luis Potosi)
Sinaloa (Sinaloa)
Sonora (Sonora)
Tabasco (Tabasco)
Tamaulipas (Tamaulipas)
Tlaxcala (Tlaxcala)
Veracruz (Veracruz)
Yucatan (Yucatan)
Zacatecas (Zacatecas)
Micronesia (MIC)
Caroline islands (Caroline islands)
Guam (Guam)
Kiribati (Kiribati)
Marshall ils (Marshall ils)
North Marians isl. (North Marians isl.)
Palau archipelago (Palau archipelago)
Moldavia (MD)
Mongolia (MG)
Montenegro (ME)
Mordor (RU)
Altai Republic and Altaisky Province (Altai Republic and Altaisky Province)
Amurskaya Area (Amurskaya Area)
Bashkiria, E of Chelyabinsk regs. (S Ural mts. Bashkiria, E of Chelyabinsk regs.)
Buryat Republic (Buryat Republic)
Chukot autonomous region (Chukotka)
East Sayan Mountains (East Sayan Mountains)
Ingushetia, Chechnya, Daghestan (E Caucas) (Ingushetia, Chechnya, Daghestan (E Caucas))
Irkutsk Area (Irkutsk Area)
Kabardino-Balkaria, Ossetia (C Caucas) (Kabardino-Balkaria, Ossetia (C Caucas))
Kaliningrad, Briansk, Smolensk, Tula, Orlov, Kaluga, Moskow, Riaz (C of Russian plains)
Kamchatskaya Area (Kamchatka)
Karachaevo-Cherkessia (W Caucas) (Karachaevo-Cherkessia)
Karelia, Leningrad reg., Pskov reg., Kostroma reg., Perm reg., Komi, Pechora plains. (N of Russian plains)
Khabarovsky Province (Khabarovsky Province)
Khamar-Daban Mountains (Khamar-Daban Mountains (E Siberia))
Komi, Yamalo-Nenetsk, Hanty-Mansi (N Ural mts.)
Krasnodar reg. (W Caucas)
Krasnoyarsky Province (Krasnoyarsk)
Kuriles Islands (Kuriles Islands)
Magadanskaya Area (Magadan)
Omsk, S of Tyumen, W Novosibirsk regs. (S of W Siberia)
Perm, Sverdlovsk regs. (C Ural mts)
Primorsky District (Primorie) (Maritime Prov)
Sakhalin Island (Sakhalin Island)
Tuva Republic (Tuva)
Voronezh, Rostov, Saratov, Kalmykia, Volgograd, Astrahan regs. (S of Russian plains)
West Sayan Mountains (West Sayan)
Yakut-Saha Republic (Yakut-Saha Republic)
Zabaykalsky Krai (ex Chita reg.) (Zabaykalsky Krai)
plains of Krasnodar reg., S Rostov reg., Stavropol reg. (Ciscaucas reg. (plains of Krasnodar reg., S Rostov)
taiga from Ural mts to Yenisei riv. (Middle of W Siberia)
tundra of Tyumen, Yamalo-Nenetsk regs. (N of W Siberia)
Morocco (MO)
Mozambique (MZB)
Myanmar (Burma) (BRM)
Namibia (NAM)
Nepal (NP)
Netherlands (NL)
New Zealand (NZL)
Antipodes Islands (Antipodes Islands)
Auckland Islands (Auckland Islands)
Campbell Island (Campbell Island)
Chatham Islands (Chatham Islands)
Nicaragua (NCG)
North Korea (NC)
Norway (NR)
Svalbard (Spitzbergen) (SB)
Oman (OM)
Pakistan (PA)
Panama (PNM)
Paraguay (PRG)
Peru (PER)
Philippines (PLP)
Basilan (Basilan)
Cebu (Cebu)
Guimaras (Guimaras)
Leyte (Leyte)
Luzon (Luzon)
Marinduque (Marinduque)
Masbate (Masbate)
Mindanao (Mindanao)
Mindoro (Mindoro)
Negros (Negros)
Palawan (Palawan)
Panay (Panay)
Polillo Islands (Polillo Islands)
Romblon (Romblon)
Samar (Samar)
Sulu Archipelago (Sulu Archipelago)
Poland (PL)
Polynesia (POL)
Cook Islands (Cook Islands)
Hawaiian Islands (HAW)
Marquesas Islands (Marquesas Islands)
Niue (Niue)
Norfolk ils (Norfolk ils)
Pitcairn ils (Pitcairn ils)
Rapa Nui (Easter Island) (Rapa Nui)
Samoa (Samoa)
Society Islands (Society Islands)
Tahiti (Tahiti)
Tokelau (TKL)
Tonga (Tonga)
Tuvalu ils (Tuvalu ils)
Portugal (PT)
Qatar (QA)
Republic of South Africa (RSA)
Republic of the Congo (CG)
Reunion isl. (Reunion isl.)
Rodriguez isls. (ROD)
Romania (RO)
Rwanda (RWA)
Santa Helena isl (SH)
Sao Tome e Principe (STM)
Saudi Arabia (SA)
Serbia (SRB)
Seychelles (SEY)
Singapore (SGP)
Slovakia (SK)
Slovenia (SL)
Somalia (SOM)
South Georgia and the South Sandwich Islands (SGSSI)
South Korea (SC)
Spain (SP)
Balearic Islands (Balearic Islands)
Sri Lanka (SRL)
St. Paul isls (STP)
Sudan (SUD)
Surinam (SUR)
Swaziland (SWZ)
Sweden (SV)
Switzerland (SZ)
Syria (SY)
Tadzhikistan (TD)
Taiwan (Formosa) (Taiwan)
Tanzania (TAN)
Zanzibar isl. (Zanzibar isl.)
Thailand (TAI)
Tunisia (TU)
Turkey (TR)
Turkmenistan (TM)
Uganda (UGD)
Ukraine (UK)
Crimea (Crimea)
United States (USA)
Alabama (Alabama)
Alaska (Alaska)
Arizona (Arizona)
Arkansas (Arkansas)
California (California)
Colorado (Colorado)
Connecticut (Connecticut)
Delaware (Delaware)
Florida (Florida)
Georgia (Georgia)
Idaho (Idaho)
Illinois (Illinois)
Indiana (Indiana)
Iowa (Iowa)
Kansas (Kansas)
Kentucky (Kentucky)
Louisiana (Louisiana)
Maine (Maine)
Maryland (Maryland)
Massachusetts (Massachusetts)
Michigan (Michigan)
Minnesota (Minnesota)
Mississippi (Mississippi)
Missouri (Missouri)
Montana (Montana)
Nebraska (Nebraska)
Nevada (Nevada)
New Hampshire (New Hampshire)
New Jersey (New Jersey)
New Mexico (New Mexico)
New York (New York)
North Carolina (North Carolina)
North Dakota (North Dakota)
Ohio (Ohio)
Oklahoma (Oklahoma)
Oregon (Oregon)
Pennsylvania (Pennsylvania)
Rhode Island (Rhode Island)
South Carolina (South Carolina)
South Dakota (South Dakota )
Tennessee (Tennessee)
Texas (Texas)
Utah (Utah)
Vermont (Vermont)
Virginia (Virginia)
Washington (Washington)
West Virginia (West Virginia)
Wisconsin (Wisconsin)
Wyoming (Wyoming)
Uruguay (URG)
Uzbekistan (UZ)
Venezuela (VNZ)
Vietnam (VTN)
West Africa (WAF)
Benin (Benin)
Burkina Faso (Burkina Faso)
Côte d'Ivoire (Côte d'Ivoire)
Gambia (Gambia)
Ghana (Ghana)
Guinea (Guinea)
Guinea Bissau (Guinea Bissau)
Liberia (Liberia)
Mali (Mali)
Niger (Niger)
Nigeria (Nigeria)
Senegal (Senegal)
Sierra Leone (Sierra Leone)
Togo (Togo)
Yemen (YE)
Socotra (Socotra)
Zambia (ZMB)
Zimbabwe (ZIM)
Main (random 10) |
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Adult
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Immature stages
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Endophallus
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Biotope
Amblystomus jeanneli Basilewsky, 1948
Female adult
Sénégal, Nianing, piège lumineux 26/09/2009, Pascal Deschamps coll.
Copyright ©
Pascal Deschamps
Anomostomus torridus LaFerte-Senectere, 1853
Female adult
Sénégal, Nianing, piège lumineux, 30/01/2009, Pascal Deschamps coll.
Copyright ©
Pascal Deschamps
Amblystomus orpheus (LaFerte-Senectere, 1853)
Male adult
Namibia, Etosha National Park, Okaukuejo, 1400 m, leg. U.Schmidt, 18.-20.III.1994
det. M.Baehr, 2012
4,8 mm
Copyright ©
Udo Schmidt
Amblystomus amabilis (Boheman, 1848)
Female adult
Size: 4,8 mm
Namibia, Etosha National Park, Okaukuejo, 1400 m, leg. U.Schmidt, 18.-20.III.1994;
det. M.Baehr, 2012 Photo: U.Schmidt, 2014
Copyright ©
Udo Schmidt
Amblystomus natalicus Peringuey, 1896
Male adult
3,3 mm; Namibia, Kamanjab, Outjo, Otjitambi Guest Farm, 1000 m, leg. U.Schmidt, 23.III.1994; det. M.Baehr, 2015; Photo: U.Schmidt, 2016
Copyright ©
Udo Schmidt
Anomostomus laevigatus (Kuntzen, 1919)
Female adult
Mozambique, 16.XII.2005, Manica Province, 60km W Chitobe. 6.5 mm; det. P. Schule 2009
Copyright ©
Alexander Anichtchenko
Barysomus metallicus Reiche, 1843
Female adult
Venezuela, Barinas, Santa Barbara, IV.1985, L. Ratel leg.
Copyright ©
Alexander Anichtchenko
Oosoma gyllenhalii (Dejean, 1829)
Unsexed
Copyright ©
- - -
Anomostomus (Tnaei) devagiriensis Nijisha, Sabu & Aswathi, 2023
Holotype
Female adult
India: Kerala: Chalode, paddy field (11°55’47.6”N 75°30’06.3”E), ‘Light trap’, 09.x.2020, coll. K. Nijisha. Deposited in ZSIK
Copyright ©
- - -
Amblystomina
taxon page
(41 photos)
Dixus eremita Dejean, 1825
Male adult
Volgogrd Area, Pallasovka Distr., 5 km NNW Elton Vill., Sorochya Balka 8.IV.2007 leg. A.Matalin
L=9.0
Copyright ©
Kirill Makarov
Eucarterus sparsutus Reitter, 1898f: 338
Male adult
Copyright ©
Kirill Makarov
Odontocarus robustus (Dejean, 1831)
Male adult
Greece, Peloponnes, Oros Menalon
occ. Dimitsama 900-1000m 12-23.5.1992 A.Korell leg.
17 mm
Copyright ©
Alexander Anichtchenko
Carterus (Carterus) dama (P.Rossi, 1792)
Male adult
det. K. Makarov
Copyright ©
Kirill Makarov
Tschitscherinellus oxygonus oxygonus
Male adult
Copyright ©
Kirill Makarov
Dixus genohensis Azadbakhsh & Kirschenhofer, 2018
Paratype
Female adult
Iran, Hormozgan Province, Geno Mountain, 20.III.2017, 1800 m, S. Azadbakhsh leg. (cAA)
Copyright ©
Alexander Anichtchenko
Parapenthus solitarius (Peyron, 1858)
Male adult
Image from Kataev & Wrase, 2018.
Copyright ©
- - -
Eocarterus (Baeticocarus) amicorum Wrase, 1993
Male adult
Spain, Andalucia, Sierra de las Nieves, 1750-1800m, 6-8.4.2010, Anichtchenko A. leg. & det.
Copyright ©
Alexander Anichtchenko
Carterus (Pristocarterus) validiusculus Brulerie, 1873
Male adult
Lebanon, Jbeil prov., Laqlouq, 8.5.2000, P. Rapuzzi leg.; det. Anichtchenko A. 2018
Copyright ©
Alexander Anichtchenko
Oedesis tomentosus (Dejean, 1831)
Male adult
Spain, San Roque, Cortijo de la Ventilla, 4.4.2003 Anichtchenko A. leg & det. (cAA). 11 mm
Copyright ©
Alexander Anichtchenko
Ditomina
taxon page
(66 photos)
Nipponoharpalus discrepans (A.Morawitz, 1862)
Male adult
Collection ground Chitose city, Hokkaido Pref. pref. 34168
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Harpalus (Harpalus) caspius (as roubali Schauberger, 1928)
Male adult
Moravia, Kobylí; 11 mm
Copyright ©
Aleš Sedláček
Siopelus (Neosiopelus) punctatellus (Reiche, 1847)
Male adult
7,2 mm. Namibia, Outjo, 25 km south Okaukuejo, Ongave Lodge, 1000 m, leg. U.Schmidt, 21.III.1994; det. M.Baehr, 2012 Photo: U.Schmidt, 2014
Copyright ©
Udo Schmidt
Neodiachipteryx carinigera (Putzeys, 1878)
Male adult
Antilles, Hispaniola. Image from Shpeley, Hunting & Ball, 2017
Copyright ©
- - -
Omostropus mandibularis (Roth, 1851)
Female adult
RSA, Nothern prov., Louis Trichardt Ben Lavin Nat. Res., 29.I.2000. det. S. Facchini 2003. 8 mm
Copyright ©
Alexander Anichtchenko
Parophonus (Parophonus) mendax (P.Rossi, 1790)
Male adult
Bulgaria. Chernomerets, 24.XI.2006, A. Anichtchenko leg. & det.
Copyright ©
Alexander Anichtchenko
Axinotoma hulstaerti Basilewsky, 1968
Male adult
Gabon, Makokou 7.7 km SW, Ivindo NP, Ipassa forest research s., 0*30.52N 12*48'21E, 21-22.6.2016, secondary lowland rainforest, MV light, D.Telnov leg.10 mm. det. S. Facchini, 2019
Copyright ©
Alexander Anichtchenko
Siopelus (Siopelus) usambarensis Facchini, 2021
Holotype
Male adult
Coll. I.R.Sc.N.B., Tanzania, Tanga Region, Amani Nature Reserve, 06.X.2013, Agroforestry area, 05°06.239'S, 38°37.509'E, 994 m, Leg. L. Geeraert (RBINS).
Copyright ©
Sergio Facchini
Trichotichnus (Neotichnus) riedeli Kataev, 2022
Holotype
Female adult
Papua N. G. / Morobe Prov. / leg. A. Riedel, ca. 10 km S Garaina, / Saureri, 1800–2150 m, / 26.III.1998 (ZSM). Image from Kataev, 2022
Copyright ©
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Harpalus (Epiharpalus) punctipennis Mulsant, 1852
Male adult
Image from Kataev, 2023
Copyright ©
- - -
Harpalina
taxon page
(722 photos)
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