Carabus (Mesocarabus) baguenai Breuning, 1926
Male adult
Spain, Almeria, Sierra de Baza 10.III.1995<br /> male<br /> photo Anichtchenko A.<br />
Copyright © Alexander Anichtchenko

Carabus (Mesocarabus) dufouri baguenai Breuning, 1926
Endophallus
SE Spain, Almería Province, Carboneras env.
det. R.Panin
Copyright © Ruslan Panin

Carabus (Mesocarabus) baguenai Breuning, 1926
Male adult
SE Spain, Almería Province, Carboneras env., 25.XI.2006, T.Gazurek leg./det. R.Panin <br /> L=26 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) breuningi Csiki, 1927
Male adult
Spain, San Martin de Castaneda, Zamora<br /> 11 oct 1984,h~1500 m
Copyright © Alexander Anichtchenko

Carabus (Mesocarabus) breuningi Csiki, 1927
Male adult
Villanueva, Orense, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Mesocarabus) breuningi Csiki, 1927
Female adult
NW-Spain, Prov. Zamora, San Martín de Castañeda, 5.VII.1984, M.Toribio leg. L=28 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) breuningi Csiki, 1927 (loc.typ.)
Male adult
NW-Spain, Orense, Orensano Central Massif, Monte de Cabeza de Manzaneda, 25.VII.1988, F. Rodrigues leg. L=25 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) cantabricus moroderi (as rodriguezianus Meyer & Mollard in Deuve, 2003)
Female adult
Poza de la Sal, Burgos, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Mesocarabus) cantabricus cantabricus (as asturicus Born, 1925)
Female adult
Pravia, Asturies, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Mesocarabus) cantabricus cantabricus (as liebei Eidam, 1925)
Male adult
Fonsagrada, Galice, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Mesocarabus) cantabricus cantabricus Chevrolat, 1840 (loc. typ.)
Female adult
La Caniza, Galice, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Mesocarabus) cantabricus moroderi (as joaquini Breuning & Toulgoet, 1974)
Male adult
Pena Ubina, Leon, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Mesocarabus) cantabricus amayensis Lassalle, 1984
Male adult
Pto.de Amaya, Burgos, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Mesocarabus) cantabricus brabeus (as barcelecoanus)
Female adult
Col de Velate, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
Male adult
Col de Velate, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Mesocarabus) cantabricus moroderi (as magdalenai Meyer & Mollard, 1996)
Endophallus
NW-Spain, Leon prov., Sierra de Gistredo, Pto. de Magdalena,
Copyright © Ruslan Panin

Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
Male adult
NC-Spain, Basque Co, Álava, Sierra de Gorbea, VI.2014, L=29 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) dufouri dufouri Dejean, 1829 (as papillatus Lapouge, 1902)
Male adult
Spain, Algeciras, Molino de las Cuevas
5-6.4.2003 Anichtchenko A. leg.
Copyright © Alexander Anichtchenko

Carabus (Mesocarabus) dufouri dufouri Dejean, 1829
Endophallus
Jaen reg., Monte Javalcuz h=1436 m
Copyright © Ruslan Panin

Carabus (Mesocarabus) dufouri dufouri Dejean, 1829
Male adult
SC Spain, Jaen reg., Monte Javalcuz h=1436 m, 6.X.2010, M.A. Lopez leg.
L=27 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) dufouri papillatus Lapouge, 1902 Lectotype
Male adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC576 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec576
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Carabus (Mesocarabus) dufouri papillatus Lapouge, 1902
Label
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC576 http://coldb.mnhn.fr/catalognumber/mnhn
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Carabus (Mesocarabus) dufouri Dejean, 1829
Female adult
S Spain, Granada, Sierra Arana, Píñar env., 14.X.2018, F. Garcia leg. L=27 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus lusitanicus Fabricius, 1801(as schaumi Gaubil, 1849)
Male+Female adult
Portugal,
Viseu distr., Gosende, h~1010m, 15.X.2009, D.Prunier leg.
female,26 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus gougeleti Reiche, 1863 (as alcaracinus Kraatz, 1886)
Male adult
Spain, Riopa (Albacete)
Copyright © Alexander Anichtchenko

Carabus (Mesocarabus) (? lusitanicus) gougeleti Reiche, 186
Endophallus
CW Spain, Province Huelva, Arroyomolinos de Leon.
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826
Endophallus
CW-Spain:Salamanca, Brincones env., h=750m
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826 Holotype
Male adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC34 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec34
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Carabus (Mesocarabus) lusitanicus trypalsi Breuning, 1975
Label
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC133 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec133
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Carabus (Mesocarabus) lusitanicus antiquus Dejean, 1826
Male adult
Aliseda, Extramadure, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Mesocarabus) lusitanicus trabuccarius (as troncedoensis) Paratype
Female adult
Troncedo, Huesca, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Mesocarabus) lusitanicus helluo (as albarracinus)
Female adult
Altos de Cabrera, Pineda de Ciguela, Cuenca, SPAIN
Copyright © Alain Mollard

Carabus (Mesocarabus) lusitanicus lusitanicus (as vieirae Oliveira, 1875)
Female adult
CW-Portugal: Leiria distr., near Marinha Grande, 39°45'57.8"N 8°58'10.3"W, h-79 m, 19.IX.2016, D. Prunir leg., L=26 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) macrocephaloides (Jeanne, 1972)
Female adult
Sierra de Andia, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Mesocarabus) macrocephaloides (Jeanne, 1972)
Female adult
Sierra de Andia, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Mesocarabus) macrocephaloides (Jeanne, 1972)
Female adult
Sierra de Andia, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Mesocarabus) macrocephaloides (Jeanne, 1972)
Female adult
Sierra de Andia, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Mesocarabus) macrocephalus Dejean, 1826
Male adult
Spain, Picos de Europa, Macizo Central,
Llagu Bajero 1890m 6.8.2003
Copyright © Alexander Anichtchenko

Carabus (Mesocarabus) macrocephalus Dejean, 1826
Male adult
NW-Spain,Cantabrian Mts, Picos de Europa Mt Rng, Puerto de Panderruedas h=1460 m, 25.IX.2004,T.Gazurek leg.
L=28 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) macrocephalus Dejean, 1826
Endophallus
NW-Spain,Cantabrian Mts, Picos de Europa Mt Rng, Puerto de Panderruedas h=1460 m,
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus harcyniae Sturm, 1815 (as obenbergerianus Breuning, 1927)
Male adult
Czech, Bohemia, Plzeň Reg., Domažlice Distr., Babylon, h=500 m, 30.VIII.2008, S.I. Budka leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus inflatus Kraatz, 1878
Male adult
SE–France, Var departament, Foret royale -Saint-Paul-en-Foret vill.,18.VI.2002, Thelot J.P. leg.
L=26 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus islandicus Lindroth, 1968
Female adult
SE-Iceland:Höfn env.:Hornafjördur, 24-28.VIII.2008, C. Reuter leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus harcyniae (as procedens Csiki, 1927)
Female adult
N-Ireland, Fermanagh Distr., Lough Navar Forest, 17.VII.2013, D.Novak & J.Stypula leg.<br /> L=27mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) (? problematicus trans lusitanicus) planiusculus (as xaxarsi Born, 1917)
Endophallus
NE Spain, Catalonia,Girona Prov., San Hilario Sacalm env.<br />
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus planiusculus (as xaxarsi)
Male adult
Sant Sadurni, Gerone, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Mesocarabus) problematicus planiusculus (as bepmalei Lapouge, 1904)
Female adult
Pic de Bassibes, Huesca, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Mesocarabus) problematicus planiusculus (as lagardei Mollard, 1996) Paratype
Female adult
Sierra de Andia, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Mesocarabus) problematicus planiusculus (as andorranus Lapouge, 1902)
Female adult
Col d'envalira, Pas de la Case, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Mesocarabus) problematicus problematicus Herbst, 1786
Female adult
W-Hungary, NE-Alps, Mts. Kőszeg, Velem env., Borha-forrás, 26.VII.1981, Gy. Somorijai leg. L=26 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) riffensis Fairmaire, 1875 Lectotype
Female adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC142 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec142
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Carabus (Mesocarabus) riffensis Fairmaire, 1875
Label
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC142 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec142
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Carabus (Mesocarabus) riffensis Fairmaire, 1875
Male adult
Morocco, Rif, Foresta di Ketama, 25.06.1998, 1600m, leg. D. Sechi (immature)
Copyright © Alexander Anichtchenko

Carabus (Mesocarabus) riffensis Fairmaire, 1875
Endophallus
Morocco, Rif, Foresta di Ketama, 25.06.1998, 1600m, leg. D. Sechi (immature)
Copyright © Alexander Anichtchenko

Carabus (Mesocarabus) riffensis Fairmaire, 1875 (as dominicae)
Male adult
Ft.de Ketama , Rif, MAROC
Copyright © Alain Mollard

Carabus (Mesocarabus) riffensis Fairmaire, 1875
Female adult
N-Morocco, Riff Mts h=1650 m, 26 km E of Bab Berred, 34.961333, -4.669417, Cedar forest, C. Reuter leg. L=33 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) riffensis Fairmaire, 1875
Endophallus
N-Morocco, Riff Mts h=1650 m, 26 km E of Bab Berred
Copyright © Ruslan Panin

Carabus (Mesocarabus) riffensis Fairmaire, 1875
Male adult
N-Morocco, Riff Mts h=1650 m, 26 km E of Bab Berred, 34.961333, -4.669417, Cedar forest, C. Reuter leg. L=27 mm
Copyright © Ruslan Panin