Carabus (Neoplectes) apollo apollo Zolotarev, 1913
Male adult
Caucasus, Abchasia

Copyright © Dmitri Obydov

Carabus (Neoplectes) apollo phoebus Kurnakov, 1962
Male adult
Abchasia, Kodori, A. Zolotarew leg.
Photo by D.V. Obydov
from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Neoplectes) apollo tenebricosus Kurnakov, 1962
Male adult

Copyright © Safronov

Carabus (Neoplectes) apollo tenebricosus Kurnakov, 1962
Endophallus

Copyright © Alexandr Safronov

Carabus (Neoplectes) compressus rebellis Reitter, 1895
Male adult
Georgia, Kvemo-Svaneti, S slopes of Svanetskiy Mt.R., S slope of Latpari pass, h=2225m, 17.VI -13.VII.2017 leg. D.Fominykh L=24.0
Copyright © Kirill Makarov

Carabus (Neoplectes) compressus rebellis Reitter, 1895
Female adult
Georgia, Kvemo-Svaneti, S slopes of Svanetskiy Mt.R., S slope of Latpari pass, h=2225m, 17.VI -13.VII.2017 leg. D.Fominykh L=26.0 mm
Copyright © Kirill Makarov

Carabus (Neoplectes) compressus rebellis Reitter, 1895
Male genitalia
Georgia, Kvemo-Svaneti, S slopes of Svanetskiy Mt.R., S slope of Latpari pass, h=2225m, 17.VI -13.VII.2017 leg. D.Fominykh
Copyright © Kirill Makarov

Carabus (Neoplectes) compressus rebellis Reitter, 1895
Endophallus
Georgia, Kvemo-Svaneti, S slopes of Svanetskiy Mt.R., S slope of Latpari pass, h=2225m, 17.VI -13.VII.2017 leg. D.Fominykh
Copyright © Kirill Makarov

Carabus (Neoplectes) zubovi Fominykh, Zamotajlov et Titarenko, 2018 Holotype
Male adult
S slopes of Svaneti Mt. Range near Lentekhi Town, S slopes of Goldash Mt., 42°49'N, 42°45'E, 2300–2800 m, 21.VI.2016, D. Fominykh, A. Zherebilo (ZISP).
Copyright © Kirill Makarov

Carabus (Neoplectes) compressus lailensis Belousov, 1992
Male adult
Laila Mt.
Copyright © Kirill Makarov

Carabus (Tribax) lailensis stefaniianus Deuve, 2018 Holotype
Male adult
Géorgie, Basse-Svanétie, au nord de Lentekhi, versant sud de la chaîne de Svanétie, 1000 m., Th. Deuve et A. Matocq, juin 2016. (MNHN)
Copyright © Thierry Deuve

Carabus (Tribax) zubovi mananauriensis Deuve, 2022 Holotype
Male adult
Géorgie, Svanétie, 20 km à l’ouest de Lentekhi, 6 km à l’ouest de Mananaouri, 1350 m (A. Wřzecionko leg.), in coll. A. Wřzecionko, à Horni Sucha (République Tchèque).
Copyright © Thierry Deuve

Carabus (Neoplectes) compressus rebellis Reitter, 1895 Lectotype
Male adult
Image from Deuve, 2024
Copyright © Thierry Deuve

Carabus (Tribax) compressus pseudorebellis Deuve, 2024 Holotype
Male adult
Géorgie, Basse-Svanétie, 12 km au nord-est de Lentekhi, Mont Goldash, vallée de la Cholshoura, Durashi (« Dvashi »), env. 1150 mètres (W. Heinz, 1-4 juillet 2019), in coll. Muséum Stuttgart (Germany).
Copyright © Thierry Deuve

Carabus (Neoplectes) compressus zubovi Fominykh, Zamotajlov et Titarenko, 2018 Paratype
Female adult
S slopes of Svaneti Mt. Range near Lentekhi Town, S slopes of Goldash Mt., 42°50'N, 42°45'E, 2200–2600 m, 18.VI–13.VII.2017 (D. Fominykh). Image from Fominykh, Zamotajlov et Titarenko, 2018
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Carabus (Neoplectes) compressus zubovi Fominykh, Zamotajlov et Titarenko, 2018 Paratype
Male+Female adult
5-6: Paki Vill. environs; 7-8: Mannanauri Vill. environs. Scale bar 10.0 mm. Image from Fominykh, Zamotajlov et Titarenko, 2018
Copyright © - - -

Carabus (Neoplectes) daphnis askhicus Belousov, 1992 Paratype
Male adult
SC-Caucasus, W Georgia, valley of riv. Chkhorodukhu, h=1000-1200m, 14.V.-6.VII.1991, Zamotajlov A.S & Miroshnikov leg.
male, 29 mm
photo & collection Panin R.
Copyright © Ruslan Panin

Carabus (Neoplectes) daphnis askhicus Belousov, 1992 Paratype
Male+Female adult
W Georgia, valley of Chkhorodukhu riv., 1000-1200m
14.V-6.VII.1991
male 30mm, female 33mm
photo & collection Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) daphnis askhicus Belousov, 1992 Paratype
Male adult
Georgia, env. Martvilli, Askhi plateau
10-15.7.2001 A.Rubenian leg.
27 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) daphnis daphnis Kurnakov, 1962
Male adult
Georgia, Mingrelia, Lecheha vallis, Tehuri superior
Caucasus centr., 1200m, 27.6-7.7.1989 Retezar I. leg.
male, 28 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) daphnis daphnis Kurnakov, 1962
Endophallus
Georgia, Mingrelia, Lecheha vallis, Tehuri superior
Caucasus centr., 1200m, 27.6-7.7.1989 Retezar I. leg.
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) daphnis askhicus Belousov, 1992
Endophallus
Georgia, env. Martvilli, Askhi plateau
10-15.7.2001 A.Rubenian leg.
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) daphnis askhicus Belousov, 1992
Endophallus
W Georgia, valley of Chkhorodukhu riv., 1000-1200m
14.V-6.VII.1991
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) daphnis askhicus Belousov, 1992 Paratype
Endophallus

Copyright © Ruslan Panin

Carabus (Neoplectes) daphnis tsekuricus Belousov, 1992
Male adult
LEFT: Lakhashuri River valley; RIGHT: Tzikuri Mt. Scale bar 10.0 mm.
Copyright © Kirill Makarov

Carabus (Neoplectes) daphnis boulbenellus Deuve et Stéfani, 2017 Holotype
Male adult
Géorgie, Basse-Ratcha, à l’est de Nikortsminda, Skhartali, 1200-1300 m., Th. Deuve, P. Stéfani leg. (MNHN).
Copyright © Thierry Deuve

Carabus (Neoplectes) edithae exedithae Gottwald, 1985
Male adult
Karachaevo-Cherkessia, low cource of Daut river,
16-20.V.2009 Anichtchenko A. leg

Copyright © Alexander Anichtchenko

Carabus (Neoplectes) edithae exedithae Gottwald, 1985
Female adult
Karachaevo-Cherkessia, Zelenchukskii distr.,
near Bukovo vill., 8-13.VII.2007, S.Dementiev leg.

Copyright © Dmitri Obydov

Carabus (Neoplectes) edithae kabardensis Kurnakov, 1970
Female adult
Kabardino-Balkaria, Skalisty Mt Ridge, 1800 m,
20.VII.1996. Andreev leg.
Photo by D.V. Obydov

Copyright © Dmitri Obydov

Carabus (Archiplectes) edithae Reitter, 1893
Male adult

Copyright © Kirill Makarov

Carabus (Archiplectes) edithae Reitter, 1893
Habitat
Carabus (Archiplectes) edithae exedithae Gottwald, 1985
Karachaevo-Cherkessia, valley of Daut riv. 43.62' 42.09'
17-18.V.2009
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) edithae exedithae Gottwald, 1985
Male adult
NС-Caucasus, Karachaevo-Cherkessia, Abishir-Akhuba Mt.Range h=2250-2600 m, right bank Pkhiya river, 10-13.VI.2013, S. Pavluchuk leg.
L=25 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) edithae exedithae Gottwald, 1985
Endophallus
NС-Caucasus, Karachaevo-Cherkessia, right bank Pkhiya river, Abishir-Akhuba Mt.Range h=2250-2600 m,
Copyright © Ruslan Panin

Carabus (Neoplectes) edithae Reitter, 1893
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Carabus (Neoplectes) edithae miloslavskii Obydov, 2010 Holotype Paratype
Male+Female adult
Holotype: Caucasus centr., Archyz, zona subalp., 15.VI.1967, A. Bogachev leg; Paratype: KarachaiCherkessian Region, Zagedan Mt. Ridge, 2500 m., 6-7.VII.1994, Melnik leg.
Copyright © - - -

Carabus (Neoplectes) ibericus s.str. (as djanoliensis Fominykh & Zamotajlov, 2016) Paratype
Male+Female adult
Image from Fominykh & Zamotajlov, 2016
Copyright © - - -

Carabus (Neoplectes) ibericus ibericus (as prunierianus Deuve, 2012)
Male+Female adult
Georgia, Magaroskari env. Image from Fominykh & Zamotajlov, 2016
Copyright © - - -

Carabus (Neoplectes) ibericus ibericus (as prunierianus Deuve, 2012)
Male adult
Georgia, Passanauri env. Image from Fominykh & Zamotajlov, 2016
Copyright © - - -

Carabus (Neoplectes) ibericus ibericus (as szekelyi Retezar, 2011)
Male+Female adult
Itrapula env. Image from Fominykh & Zamotajlov, 2016
Copyright © - - -

Carabus (Neoplectes) ibericus ibericus (as szekelyi Retezar, 2011)
Male+Female adult
Dzhalobet env.; Image from Fominykh & Zamotajlov, 2016
Copyright © - - -

Carabus (Neoplectes) ibericus ibericus (as szekelyi Retezar, 2011)
Male+Female adult
Nakerala env.; Image from Fominykh & Zamotajlov, 2016
Copyright © - - -

Carabus (Neoplectes) ibericus ibericus (as imereticus Retezar, 2011) Paratype
Endophallus
NC-Georgia, S-Caucasus: W slope Likhi Mt.Rng., 3 km W Rikoti Pass h~900 m,
Copyright © Ruslan Panin

Carabus (Neoplectes) ibericus ibericus (as imereticus Retezar, 2011) Paratype
Male adult
S-Caucasus: NC-Georgia, W slope Likhi Mt. Rng., 3 km W of Rikoti Pass h~900 m, 15.VI - 8.VII.2006, A.Puchkov leg., L=29.5 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) ibericus lafertei Chaudoir, 1846
Female adult
SC-Georgia, Samtskhe–Javakheti, Meskheti Mt Range, Borjomi, Likani env., h=897 m, 41°50.125'N 43°19.905'E, VI.2018, D. Prunier leg., L=37 mm
Copyright © Ruslan Panin

Carabus (Archiplectes) jason Semenov, 1898
Male adult
Caucasus, W Georgia, Svanetia, upper reaches of riv. Shoudidi, 1700-2100m, 20-21.V.1986
leg. A. Zamotajlov
Copyright © Sunvas

Carabus (Neoplectes) jason Semenov, 1898
Male adult
Abkhasia, up river Shaudidi, 21.V - 2.VII.1986, Zamotailov A. leg.
Photo by D.V. Obydov

Copyright © Dmitri Obydov

Carabus (Neoplectes) jason Semenov, 1898
Male adult
Abhazia, Tchkhaltinskiy (Abkhazskiy) Mt.R., 20-26.VII.2022 leg. D. Fominykh
Copyright © Igor Solodovnikov

Carabus (Neoplectes) jason Semenov, 1898
Endophallus

Copyright © Igor Solodovnikov

Carabus (Neoplectes) khipstaensis Solodovnikov, 2015 Paratype
Male adult
Абхазия, Гудаутский р-н, Ю скл. Бзыбского хр., водораздельный холм в истоках р. Хипста, 43°17'52.03" N / 40°40'8.72" E, h = 1435 m, Линия № 64-4, буковый спелый лес с папоротниками и ежевикой, 09-13.08.2014, leg. И.А. Солодовников, Е.В. Татун
Copyright © Igor Solodovnikov

Carabus (Neoplectes) khipstaensis Solodovnikov, 2015 Paratype
Male adult
Абхазия, Гудаутский р-н, Ю скл. Бзыбского хр., водораздельный холм в истоках р. Хипста, 43°17'52.03" N / 40°40'8.72" E, h = 1435 m, Линия № 64-4, буковый спелый лес с папоротниками и ежевикой, 09-13.08.2014, leg. И.А. Солодовников, Е.В. Татун
Copyright © Igor Solodovnikov

Carabus (Neoplectes) khipstaensis Solodovnikov, 2015 Paratype
Female adult
Абхазия, Гудаутский р-н, Ю скл. Бзыбского хр., водораздельный холм в истоках р. Хипста, 43°17'52.03" N / 40°40'8.72" E, h = 1435 m, Линия № 64-4, буковый спелый лес с папоротниками и ежевикой, 09-13.08.2014, leg. И.А. Солодовников, Е.В. Татун
Copyright © Igor Solodovnikov

Carabus (Neoplectes) khipstaensis Solodovnikov, 2015 Paratype
Female adult
Абхазия, Гудаутский р-н, Ю скл. Бзыбского хр., водораздельный холм в истоках р. Хипста, 43°17'52.03" N / 40°40'8.72" E, h = 1435 m, Линия № 64-4, буковый спелый лес с папоротниками и ежевикой, 09-13.08.2014, leg. И.А. Солодовников, Е.В. Татун
Copyright © Igor Solodovnikov

Carabus (Neoplectes) polychrous khipstaensis Solodovnikov, 2015
Endophallus

Copyright © Igor Solodovnikov

Carabus (Neoplectes) koltzei faunulus Kurnakov, 1970
Female adult
Abchasia, Akarmara, 6500 f, A. Zolotarew leg.
Photo by D.V. Obydov

Copyright © Dmitri Obydov

Carabus (Neoplectes) koltzei faunus Kurnakov, 1970
Male adult
Abkhazia, Kadorsky Mt Ridge, 1000 m, Chlou, 22.V.1993, A. Evstigneev leg.
Photo by D.V. Obydov

Copyright © Dmitri Obydov

Carabus (Neoplectes) koltzei kobachidzei Kurnakov, 1970
Female adult
Caucasus, E Abkhasia, SW slopes of Akiba mt. rng., 1200m, 7.V.-2.VII.1991<br /> Zamotajlov & Miroshnikov leg.
Copyright © Sunvas

Carabus (Neoplectes) koltzei kobachidzei Kurnakov, 1970
Female adult
Caucasus, E Abkhasia, SW slopes of Akiba mt. rng., 1200m, 7.V.-2.VII.1991
Zamotajlov & Miroshnikov leg.
Copyright © Sunvas

Carabus (Neoplectes) koltzei kobachidzei Kurnakov, 1970
Male+Female adult
Caucasus, E Abkhasia, SW slope of Akiba mt.rng., 1200m, 7.V.-2.VII.1991 Zamotajlov & Miroshnikov leg.
male 37mm, female 38mm
photo & collection Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Archiplectes) koltzei Rost, 1889
Male adult
Abkhasia, Habyu 3.7.2002
Ivliev P. leg.
32 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) koltzei koltzei Rost, 1889
Male adult
Abkhasia, Tsebeldy, 23.V.1986, A.Zamotailov leg.
Photo by D.V. Obydov

Copyright © Dmitri Obydov

Carabus (Neoplectes) koltzei lennoni Gottwald, 1985
Male adult
Abkhasia, Amtkel lake, 22.V - 4.VII.1986, A.Zamotailov leg.
Photo by D.V. Obydov
from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Neoplectes) komarowi komarowi Reitter, 1882
Female adult
Centr. Caucasus, Svanetia, coll. A. Zolotarew
Photo by D.V. Obydov
from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Neoplectes) komarowi komarowi Reitter, 1882
Female adult
NW Georgia, Upper Svaneti, Mestia distr., Lakhamula env. h~1250 m, 19.VI.2014, V.Sima leg. L=33 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) komarowi mestiaicus Deuve, 2016 Holotype
Female adult
Géorgie, Haute Svanétie, 10 km au nord de Mestia, vers les lacs de Koruldi, env. 2500 mètres, in coll. A. Wřzecionko, à Horní Sucha (République Tchèque).
Copyright © Thierry Deuve

Carabus (Neoplectes) komarowi komarowi Reitter, 1882 Lectotype Paralectotypes
Male+Female adult
LT - female «Kaukasus Swanetien Leder Reitter » , «Komarowi m.», «Roeschke C.», « MA.INS.COLE.194.1», «Lectotype», «Carabus komarowi Reitter, Lectotype, Th. Deuve des. 2024», ex coll. Vogt, in coll. Naturalis Biodiversity Center, Leyde (Pays-Bas). Image from Deuve, 2024
Copyright © Thierry Deuve

Carabus (Neoplectes) komarowi komarowi Reitter, 1882 Lectotype Paralectotypes
Female adult
LT, left, female: «Kaukas Leder», «Komarowi v. mutabilis m.», «Roeschke C. », « Collectie C. & O. Vogt, Acq. 1960», «Plectes Komarowi mutabilis Reitter, 1896, ZMAN type COLE.0264.1», «Lectotype», «Carabus komarowi mutabilis Reitter, Lectotype, Th. Deuve des. 2024 » , ex coll. Vogt, in coll. Naturalis Biodiversity Center, Leyde (Pays-Bas). Image from Deuve, 2024
Copyright © Thierry Deuve

Carabus (Tribax) perditus Deuve, 2024 Holotype
Male adult
MNHN
Copyright © Thierry Deuve

Carabus (Tibax) komarowi antoni Deuve, 2024 Holotype
Male adult
«Kaukasus Swanetien Leder Reitter», ex coll. M. Maindron, ex coll. G. Babault, in coll. Muséum National d’Histoire Naturelle, Paris (France).
Copyright © Thierry Deuve

Carabus (Neoplectes) kratkyi felicitanus Reitter, 1893
Male adult
Russia, N Caucasus, Karachaevo-Cherkessia 2200m, Bolshoi Pzizer mt. 8.8.2002
25 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) kratkyi felicitanus Reitter, 1893 (as porcellus)
Male adult
NW Caucasus, Resp. Adygea, Mt.Tybga, 28.VI.1997
male,20mm
Copyright © Ruslan Panin

Carabus (Neoplectes) kratkyi felicitanus Reitter, 1893 (as porcellus)
Male adult
Prov. Kuban, mt. Tybga, 1300 m, VI.1913, Shaposhnikov leg.

Copyright © Dmitri Obydov

Carabus (Neoplectes) kratkyi felicitanus Reitter, 1893
Female adult
Rusia, Caucas, Dzhuga mt., 2200 m, 24.VI. - 22.VIII.1986, Zamotajlov A. leg.

Copyright © Dmitri Obydov

Carabus (Neoplectes) kratkyi Ganglbauer, 1890
Male+Female adult
W Caucas, 1900m, Khatsavitaya Mt., 4.VI-13.VII.1994
A.Miroshnikov leg.
male 25mm, female 26mm
photo & collection Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) kratkyi Ganglbauer, 1890
Male adult
Russia, NW-Caucasus: <br /> Krasnodar reg., Mostovskoy distr., Khatsavitaja Mt. h=1900m, 4.VI.-13.VII.1994, A. Miroshnikov leg.<br /> male, 26mm<br /> photo Panin R./det.Solodovnikov I.A.
Copyright © Ruslan Panin

Carabus (Neoplectes) kratkyi Ganglbauer, 1890
Male+Female adult
W Caucas Mts., 2500m, Mt. Yatyrgvarta,
25.VI-12.VIII.1992 Shchurov V. leg.
male 20mm, female 21mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890
Female adult
NW-Caucasus, Russia,
Karacaj-Cherkessija, Abishira-Akhuba range h=2650m, upper Kyafar-Agur Riv., 4-9.VIII.2009, Solodovnikov I.A. leg.
female, 23 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890
Male adult
NW-Caucasus: Karacaj-Cherkessija,
Abishira–Akhuba Mnt. Range h=2000-2650m, Kyafar-Agur Riv., 14–17.VII.2009, Fominykh D. leg.
male, 22mm
Copyright © Ruslan Panin

Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890
Male adult
Caucasus, Kuban, Meyer-Darcis-Patria

Copyright © Dmitri Obydov

Carabus (Neoplectes) kratkyi solodovnikovi Zamotajlov, 1992
Male adult
W. Caucasus, M. Bambak Mt., VI.1931, Vvedensky leg.
from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890
Endophallus
Endophallus
Copyright © Ruslan Panin

Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
Female adult
female, 49mm
Copyright © Boleslav Brezina

Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
Male adult
male, 43mm
Copyright © Boleslav Brezina

Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
Female adult

Copyright © Kirill Makarov

Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
Female adult
W Georgia, Martvili Distr., 5 km N T'aleri, 11-18.VI.2014, V. Sima leg., L=50 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) martviliensis mzekalae Fominykh, Zamotajlov et Titarenko, 2019 Paratype
Male+Female adult
Imereti, left bank of river Semi near Melouri, 475 m, 19.V–12.VII.2019, D. Fominykh, A. Zubov, B. Dubinin & D. Safronov leg.
Copyright © Kirill Makarov

Carabus (Neoplectes) martviliensis mzekalae Fominykh, Zamotajlov et Titarenko, 2019 Paratype
Male+Female adult
Imereti, left bank of river Tskhenistskali near Dzedzileti, 620 m, 16.V–13.VII.2019, D. Fominykh, A. Zubov, B. Dubinin & D. Safronov leg.
Copyright © Kirill Makarov

Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
Male+Female adult
Imereti, S slopes of Askhi plateau near Zeda-Gordi, NE slope of Mt. Gulkuna, 980 m, 20.V–14.VII.2019, D. Fominykh & A. Zubov leg.
Copyright © Kirill Makarov

Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
Male+Female adult
Samegrelo, Doberazeni vic., left bank of river Tekhuri, 500 m, 16.IV–31.V.2017, D. Fominykh leg.
Copyright © Kirill Makarov

Carabus (Neoplectes) martviliensis Retezar & Djavelidze, 1992
Endophallus
Zeda Gordi env., 980 m and Doberazeni env., 600 m.
Copyright © Kirill Makarov

Carabus (Neoplectes) C. martviliensis mzekalae
Endophallus
Melouri env., 475 m and Dzedzileti env. , 770 m
Copyright © Kirill Makarov

Carabus (Archiplectes) plasoni dalensis Rost, 1893
Male adult
Abkhasia, near Yu. Ptysh vill.
14.7.1986 Zamotailov A. leg.
31 mm
Copyright © Alexander Anichtchenko

Carabus (Archiplectes) plasoni dalensis Rost, 1893
Male adult
Caucasus merid., Dal-Suchumi, N.S. Brianskyj leg.
from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Archiplectes) plasoni plasoni Ganglbauer, 1886
Female adult
Gruzia, Centr. Caucasus, Svanetia, coll. A. Zolotarew
det. D.V. Obydov
from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Neoplectes) polychrous Rost, 1892
Male adult
Abkhasia, Gudaut distr., near Huap vill., 600 m, 9.VI.1990, M. Danilevskyi leg. det. D.V. Obydov
Copyright © Dmitri Obydov

Carabus (Neoplectes) rousianus rousianus Gottwald, 1985
Male adult
Abkhasia, 7,6 km NE Pitsunda, 2,1 km NNE Ryapsh riv mouth, Ryapsh canyon h=60 m, 43°10N 40°25E, 25-27.VII.2008, A.Vlasenko leg.<br /> det. D.V. Obydov
Copyright © Dmitri Obydov

Carabus (Neoplectes) polychrous polychrous Rost, 1892
Female adult
W.Caucasus, Abkhasia, Gudauta distr., Duripsh village env., Khipsta valley, h=500-700 m. 8-11.06.2009 A. Safronov leg. lenght 40-43 mm.
Copyright ©

Carabus (Neoplectes) rousianus rousianus Gottwald, 1985
Male+Female adult
Абхазия, Гудаутский р-н, 5-8 км ЮВ от пос. Лдзаа (Ludzava) долина р. Риапши, левый борт; балки, поросшие орехом, каштаном, рододендроном понтийским и кленом, h = 58-70 m, 43º10’43” N / 40º25’37” E, 16-22.07.2011, leg. И.А. Солодовников, Н.Ю. Пичугин, С.M. Павлючук
Copyright © Igor Solodovnikov

Carabus (Neoplectes) rousianus rousianus Gottwald, 1985
Male+Female adult
Абхазия, Гудаутский р-н, 5-8 км ЮВ от пос. Лдзаа (Ludzava) долина р. Риапши, левый борт; балки, поросшие орехом, каштаном, рододендроном понтийским и кленом, h = 58-70 m, 43º10’43” N / 40º25’37” E, 16-22.07.2011, leg. И.А. Солодовников, Н.Ю. Пичугин, С.M. Павлючук
Copyright © Igor Solodovnikov

Carabus (Neoplectes) polychrous Rost, 1892
Female adult
Абхазия, Гудаутский р-н, окр. п. Тванаарху, дол. р. Хипсты, ущелье небольшого ручейка, поросшее буками, ежевикой и папоротником, Линия 25-3, 43°14’ N / 040°39’ E, h = 500 m,08.05-05.07.2013, leg. И.А. Солодовников, С.В. Солодовникова, С.M. Павлючук, Н.Ю. Пичугин, В.М. Коцур
Copyright © Igor Solodovnikov

Carabus (Neoplectes) polychrous Rost, 1892
Male+Female adult
Абхазия, Гудаутский р-н, окр. п. Тванаарху, дол. р. Хипсты, ущелье небольшого ручейка, поросшее буками, ежевикой и папоротником, Линия 25-3, 43°14’ N / 040°39’ E, h = 500 m,08.05-05.07.2013, leg. И.А. Солодовников, С.В. Солодовникова, С.M. Павлючук, Н.Ю. Пичугин, В.М. Коцур
Copyright © Igor Solodovnikov

Carabus (Neoplectes) polychrous Rost, 1892
Male+Female adult
Абхазия, Гудаутский р-н, окр. п. Тванаарху, дол. р. Хипсты, ущелье небольшого ручейка, поросшее буками, ежевикой и папоротником, Линия 25-3, 43°14’ N / 040°39’ E, h = 500 m,08.05-05.07.2013, leg. И.А. Солодовников, С.В. Солодовникова, С.M. Павлючук, Н.Ю. Пичугин, В.М. Коцур
Copyright © Igor Solodovnikov

Carabus (Neoplectes) rousianus svetlanae Solodovnikov, 2015 Paratype
Female adult
Абхазия, Гудаутский р-н, 12 км дороги на оз. Рица, лев. борт р. Бзыбь, буковый лес + каштан на щебнисто-глинистой почве, уроч. Авджуляр, Линия № 20, h = 210-230 m, 43º19’07” N / 040º25’06” E, 09.V-03.VII.2013, leg. И.А. Солодовников, С.В. Солодовникова, С.М. Павлючук, Н.Ю. Пичугин, В.М. Коцур
Copyright © Igor Solodovnikov

Carabus (Neoplectes) rousianus svetlanae Solodovnikov, 2015 Paratype
Male+Female adult
Абхазия, Гудаутский р-н, 12 км дороги на оз. Рица, лев. борт р. Бзыбь, буковый лес + каштан на щебнисто-глинистой почве, уроч. Авджуляр, Линия № 20, h = 210-230 m, 43º19’07” N / 040º25’06” E, 09.V-03.VII.2013, leg. И.А. Солодовников, С.В. Солодовникова, С.М. Павлючук, Н.Ю. Пичугин, В.М. Коцур
Copyright © Igor Solodovnikov

Carabus (Neoplectes) rousianus svetlanae Solodovnikov, 2015 Paratype
Male adult
Абхазия, Гудаутский р-н, 12 км дороги на оз. Рица, лев. борт р. Бзыбь, буковый лес + каштан на щебнисто-глинистой почве, уроч. Авджуляр, Линия № 20, h = 210-230 m, 43º19’07” N / 040º25’06” E, 09.V-03.VII.2013, leg. И.А. Солодовников, С.В. Солодовникова, С.М. Павлючук, Н.Ю. Пичугин, В.М. Коцур
Copyright © Igor Solodovnikov

Carabus (Neoplectes) rousianus svetlanae Solodovnikov, 2015 Paratype
Male adult
Абхазия, Гудаутский р-н, 12 км дороги на оз. Рица, лев. борт р. Бзыбь, буковый лес + каштан на щебнисто-глинистой почве, уроч. Авджуляр, Линия № 20, h = 210-230 m, 43º19’07” N / 040º25’06” E, 09.V-03.VII.2013, leg. И.А. Солодовников, С.В. Солодовникова, С.М. Павлючук, Н.Ю. Пичугин, В.М. Коцур
Copyright © Igor Solodovnikov

Carabus (Neoplectes) polychrous svetlanae Solodovnikov, 2015
Endophallus

Copyright © Igor Solodovnikov

Carabus (Neoplectes) polychrous rousianus Gottwald, 1985
Endophallus

Copyright © Igor Solodovnikov

Carabus (Neoplectes) polychrous polychrous
Endophallus

Copyright © Igor Solodovnikov

Carabus (Archiplectes) prometheus miroshnikovi Zamotajlov, 1990
Female adult
NW-Caucasus, Krasnodar reg., Mostovskoy distr., Site Val , h=1800m. 07-08.VII.2008, D.Fominyh leg., 35mm; det. I. Solodovnikov 2023
Copyright © Ruslan Panin

Carabus (Archiplectes) prometheus basilianus Starck, 1890
Male adult
Russia, W Caucas, Port-Artur Mt. 22.VI.1986, A.Zamotajlov leg.
det. D.V. Obydov
from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Neoplectes) prometheus dzhugensis Zamotajlov, 1988 Paratype
Female adult
Paratypus, female
Prov. Kuban circ., mt. Dzhuga (Tigeni), 27.VI.1911.

Copyright © Dmitri Obydov

Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990
Male+Female adult
W.Caucasus,
Psebai env., Nikitino, 2.V-3.VII.1993, leg. A.S.Zamotajlov
female 33mm
Copyright © Ruslan Panin

Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990 (as hephaestus)
Male adult
Paratypus Carabus (Archiplectes) hephaestus
Maikop, Ust. Urusht, 26.IV.1910

Copyright © Dmitri Obydov

Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990
Female adult
W Caucasus, Krasnodar reg., Maikop distr.
Nikel, 1.7.1989
38 mm
Copyright © Alexander Anichtchenko

Carabus (Archiplectes) prometheus pschadensis Zamotajlov, 1990
Female adult
NW Caucas, 5 km N from Pshada vill., 25.04.1989, Smorodinov leg.
specimen from collection of M.Shestopalov (Moscow).
det. D.V. Obydov
from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Neoplectes) prometheus prometheus Reitter, 1887
Male adult
NW-Caucasus,Krasnodar reg., Apsheron Distr., Lagonaki Mt., near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominykh leg.
L= 42mm.
Copyright © Ruslan Panin

Carabus (Neoplectes) prometheus prometheus Reitter, 1887 (as zugmayeriae)
Male adult
W. Caucasus, Maikop, 20.VI.1967., A. Bogachev leg.
det. D.V. Obydov

Copyright © Dmitri Obydov

Carabus (Neoplectes) prometheus prometheus Reitter, 1887
Male+Female adult
NW Caucasus, Russia,
Krasnodar reg.,Krasnoarmeisk distr., Krasny Les, near Podlesnyi vill.,17-22.IV.2008, D.D.Fominykh leg.
female, 40mm
Copyright © Ruslan Panin

Carabus (Archiplectes) prometheus prometheus Reitter, 1887 (as nubicola)
Male+Female adult
W Caucasus, Adygea,
h~1700m. upper res. Riv. Kurdzhips, 11-21.VIII.2007, I.A.Solodovnikov leg./det.
L=32mm.
Copyright © Ruslan Panin

Carabus (Neoplectes) prometheus prometheus Reitter, 1887
Male adult
Krasnodar distr., Mezmai
20.6.1989
38 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) prometheus prometheus Reitter, 1887 (as nubicola Zolotarev, 1913)
Endophallus
Endophallus prometheus prometheus (as nubicola Zolotarev, 1913)
Copyright © Ruslan Panin

Carabus (Neoplectes) prometheus prometheus Reitter, 1887
Endophallus
Endophallus prometheus prometheus Reitter, 1887
Copyright © Ruslan Panin

Carabus (Neoplectes) prometheus prometheus Reitter, 1887
Endophallus
aedeagus
Copyright © Ruslan Panin

Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990
Endophallus

Copyright © Ruslan Panin

Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990
Endophallus
aedeagus miroshnikovi Psebai env., Nikitino
Copyright © Ruslan Panin

Carabus (Neoplectes) protensus olegi Belousov, 1992 Paratype
Male adult
Caucasus, W Georgia, Svanetia, right bank of riv. Magana, 800m, 12.V.-4.VII.1991, Zamotajlov & Miroshnikov leg.
male, 31mm
photo and collection Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) protensus protensoides Novotny & Vorisek, 1988
Male adult
Georgia, Martvilli env., Akshi plateau 10-15.7.2001
Rubenian leg.
32 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) juenthneri atchibachi Zamotajlov, 1991 Paratype
Female adult
Abkhasia,
Valley of river Bzyb near Mt. Achibakhu. h=1500m., A.S. Zamotajlov leg.
female: 33mm, 31mm
coll. R.Panin
Copyright © Ruslan Panin

Carabus (Neoplectes) juenthneri atchibachi Zamotajlov, 1991 Paratype
Female adult
W Caucas, Abkhasia, Valley of river Bzyb near Mt. Achibakhu. h=1500m.,
24.IV.-6.VI.1991 A.S. Zamotajlov leg.
coll. Anichtchenko
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) juenthneri atchibachi Zamotajlov, 1991 Paratype
Male adult
W Caucas, Abkhasia, Valley of river Bzyb near Mt. Achibakhu. h=1500m.,
24.IV.-6.VI.1991 A.S. Zamotajlov leg.
coll. Anichtchenko
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) juenthneri atchibachi Zamotajlov, 1991 Paratype
Male adult
W Caucas, Abkhasia, Valley of river Bzyb near Mt. Achibakhu. h=1500m.,
24.IV.-6.VI.1991 A.S. Zamotajlov leg.
coll. Anichtchenko
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) juenthneri avadcharensis Kurnakov, 1970 (loc.typ.)
Male+Female adult
Abkhasia, near Avadhara, Site Pyv h~1850m, 43.490730 40.686433, 1.VI-12.VII.2009, Zamotajlov A.S., Bondarenko A.S. leg.
L=27mm

Copyright © Ruslan Panin

Carabus (Neoplectes) juenthneri avadcharensis Kurnakov, 1970
Male adult
Abkhasia, 7,4km NE from Ritza lake, 1,5 km N Kutykuh vill.,
1635 m, 19-25.VII.2008, A. Vlasenko leg.
Copyright © Dmitri Obydov

Carabus (Neoplectes) juenthneri dsychvensis Gottwald, 1985
Male adult
Abchasia, N Dsykhwa, coll. A. Zolotarew
Photo by D.V. Obydov
Copyright © Dmitri Obydov

Carabus (Neoplectes) juenthneri juenthneri Reitter, 1899
Female adult

Copyright © Alexander Anichtchenko

Carabus (Neoplectes) juenthneri avadcharensis Kurnakov, 1970
Endophallus

Copyright © Ruslan Panin

Carabus (Neoplectes) reitteri gagrinus Starck, 1894
Female adult
Abkhasia, near Gagry, 16.V.1992, A. Evstigneev leg.
det. D.V. Obydov
Copyright © Dmitri Obydov

Carabus (Neoplectes) reitteri laevisternis Starck, 1890
Male adult
Abkhasia,W.slope of Gagrinsky Mt., Krepostnaya River source, h=1325m., A.S.Zamotajlov leg. 27.VII.2000
male, 29 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) reitteri laevisternis Starck, 1890
Male adult
W-Caucasus:Sochi env., Aibga Mt. h=2000m., 06.VII.2000
male, 29mm
Copyright © Ruslan Panin

Carabus Neoplectes) reitteri reitteri Retowski, 1885
Female adult
NW Caucasus, Adygeya, Fischt-Oshten Mts. base, h=1800 m.<br /> 20.07.2006, D.Safronov leg.
Copyright © Safronov

Carabus (Neoplectes) reitteri reitteri Retowski, 1885 (as achunensis)
Male adult
W. Caucasus, Sochi Distr., Chernomoretz Vill. env.,
15.VII.1992, P. Gorbachev leg.
det. D.V. Obydov

Copyright © Dmitri Obydov

Carabus (Neoplectes) reitteri laevisternis Starck, 1890
Endophallus

Copyright © Ruslan Panin

Carabus (Neoplectes) juenthneri avadcharensis Kurnakov, 1970
Male+Female adult
W. Caucasus, Abkhasia, 8 km NE of Ritza Lake,<br /> Avadkhara valley, h=1700 m<br /> 10-VI-2009 A.Safronov leg.
Copyright ©

Carabus (Neoplectes) reitteri gagrinus Starck, 1894 (as gegicus Gottwald, 1985)
Male+Female adult
W.Caucasus, Abkhasia, Gagra distr., Gega riv. valley,
13-15.07.2005 D. Kasatkin. leg.
male - 30 mm, female - 34 mm.
Copyright © Alexandr Safronov

Carabus (Neoplectes) reitteri reitteri Retowski, 1885
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Carabus (Neoplectes) besleticus dsychvensis Gottwald, 1985
Male+Female adult
Abkhazia, Bzybian Mt. Range, SW slopes of Mt. Dzykhva, 2000-2300 m, alpine zone, N 43º13’, E 041º08’, 29.VI-03.VII.2013, leg. D. Fominykh
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus dsychvensis Gottwald, 1985
Male+Female adult
Abkhazia, Bzybian Mt. Range, SW slopes of Mt. Dzykhva, 2000-2300 m, alpine zone, N 43º13’, E 041º08’, 29.VI-03.VII.2013, leg. D. Fominykh
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus dsychvensis Gottwald, 1985
Endophallus

Copyright © Igor Solodovnikov

Carabus (Neoplectes) reitteri gagrinus Starck, 1894
Male adult
ABKHASIA, Gagra distr., Mamdsyshkha Mt., near Gagra, Fagus forest + karst, h = 1300-1330 m, biotope N 23, 43º18’186" N / 040º18’44" E, 12.05-05.07.2014, Leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Neoplectes) reitteri gagrinus Starck, 1894
Female adult
ABKHASIA, Gagra distr., Mamdsyshkha Mt., near Gagra, Fagus forest + karst, h = 1300-1330 m, biotope N 23, 43º18’186" N / 040º18’44" E, 12.05-05.07.2014, Leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Neoplectes) reitteri gagrinus Starck, 1894
Female adult
ABKHASIA, Gagra distr., Mamdsyshkha Mt., near Gagra, Fagus forest + karst, h = 1300-1330 m, biotope N 23, 43º18’186" N / 040º18’44" E, 12.05-05.07.2014, Leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Neoplectes) reitteri reitteri Retowski, 1885
Male adult

Copyright © Kirill Makarov

Carabus (Neoplectes) satyrus besleticus Kurnakov, 1972 (as gaskoi)
Male adult
USSR, Caucasus occid, Abchasia, Suchumi, vallis Kelasuri, 100m 25.6-17.7.1977 Retezar I. leg. 36 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) satyrus mtsaranus Kurnakov, 1972
Male adult
Abkhasia, Bzhybskii Mt.rng., near Mtsara vill., 16.VI.1991, S.Gusev leg.
det. D.V. Obydov

Copyright © Dmitri Obydov

Carabus (Neoplectes) satyrus napraensis Belousov & Zamotajlov, 1993 Paratype
Male adult
Abkhasia, Bzybian Mt. Range near Mt. Chibzharga., h=1900-2000m., A.S. Zamotajlov leg.
male, 28mm
photo & coll.Panin R.
Copyright © Ruslan Panin

Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972
Male adult
Abkhasia, Habyu vill.,2.7.2002 Ibliev leg.<br /> 35 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972
Male adult
Abkhasia, Habyu vill.<br /> 2.7.2002 Ibliev leg.<br /> 35 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) satyrus duripshensis Kurnakov, 1972
Male+Female adult
W.Caucasus, Abkhasia, Gudauta distr.,
Duripsh village env., Khipsta valley, h=500-700 m.
07-12.VI.2009 A.Safronov leg.
Body lenght 30-33 mm.
Copyright © Alexandr Safronov

Carabus (Neoplectes) satyrus duripshensis Kurnakov, 1972
Endophallus

Copyright © Safronov

Carabus (Neoplectes) satyrus duripshensis Kurnakov, 1972
Female adult

Copyright © Alexander Anichtchenko

Carabus (Neoplectes) satyrus duripshensis Kurnakov, 1972
Male adult
W.Caucasus, Abkhasia, Gudauta distr.,
Duripsh village env., Khipsta valley, h=500-700 m.
07-12.VI.2009 A.Safronov leg.
male - 30 mm.
Copyright © Alexandr Safronov

Carabus (Neoplectes) satyrus satyrus Kurnakov, 1962
Male+Female adult
Abkhazia, Gulripsh Distr., NW slopes of Gurzul Mt. Range near Merkheul Village, 170-230 m, left tributary of Machara River, hornbeam, alder, rhododendron (Rhododendron ponticum) forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Neoplectes) satyrus satyrus Kurnakov, 1962
Female adult
Abkhazia, Gulripsh Distr., NW slopes of Gurzul Mt. Range near Merkheul Village, 170-230 m, left tributary of Machara River, hornbeam, alder, rhododendron (Rhododendron ponticum) forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Neoplectes) satyrus satyrus Kurnakov, 1962
Male adult
Abkhazia, Gulripsh Distr., NW slopes of Gurzul Mt. Range near Merkheul Village, 170-230 m, left tributary of Machara River, hornbeam, alder, rhododendron (Rhododendron ponticum) forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Neoplectes) satyrus satyrus Kurnakov, 1962
Male adult
Abkhazia, Gulripsh Distr., NW slopes of Gurzul Mt. Range near Merkheul Village, 170-230 m, left tributary of Machara River, hornbeam, alder, rhododendron (Rhododendron ponticum) forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014 Holotype Paratype
Male adult
Abkhazia, N Slopes of Bzybian Mt. Range, right bank of River Reshevie (= Reshava) (left tributary of River Bzyb), 700-1000 m, 13.V-02.VII.2010, leg. D. Fominykh, I. Solodovnikov
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014 Paratype
Male adult
Abkhazia, N Slopes of Bzybian Mt. Range, right bank of River Reshevie (= Reshava) (left tributary of River Bzyb), 700-1000 m, 13.V-02.VII.2010, leg. D. Fominykh, I. Solodovnikov
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014 Paratype
Male+Female adult
Abkhazia, N Slopes of Bzybian Mt. Range, right bank of River Reshevie (= Reshava) (left tributary of River Bzyb), 700-1000 m, 13.V-02.VII.2010, leg. D. Fominykh, I. Solodovnikov
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014 Paratype
Male adult
Abkhazia, N Slopes of Bzybian Mt. Range, right bank of River Reshevie (= Reshava) (left tributary of River Bzyb), 700-1000 m, 13.V-02.VII.2010, leg. D. Fominykh, I. Solodovnikov
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014
Endophallus

Copyright © Igor Solodovnikov

Carabus (Neoplectes) satyrus s.str.
Endophallus

Copyright © Igor Solodovnikov

Carabus (Neoplectes) pseudopshuensis Zamotajlov, 1991
Male+Female adult
Abkhazia, Sukhum Distr., SE of Village Pskhu, hamlet Bitaga, Mt. Bzybskaya, 550-650 m, beech forest site, 12.V-30.VI.2010, leg. I. Solodovnikov, D. Fominykh
Copyright © Igor Solodovnikov

Carabus (Neoplectes) pseudopshuensis Zamotajlov, 1991
Male+Female adult
Abkhazia, Sukhum Distr., SE of Village Pskhu, hamlet Bitaga, Mt. Bzybskaya, 550-650 m, beech forest site, 12.V-30.VI.2010, leg. I. Solodovnikov, D. Fominykh
Copyright © Igor Solodovnikov

Carabus (Neoplectes) pseudopshuensis Zamotajlov, 1991
Male adult
1 male - Abkhazia, Sukhum Distr., vicinities of Village Sanchara, River Baul, beech forest site, 20.VII.1991, leg. M.N. Maksimenkov 2 male - Abkhazia, Sukhum Distr., SE of Village Pskhu, hamlet Bitaga, Mt. Bzybskaya, 550-650 m, beech forest site, 12.V-30.VI.2010, leg. I. Solodovnikov, D. Fominykh
Copyright © Igor Solodovnikov

Carabus (Neoplectes) pseudopshuensis Zamotajlov, 1991
Male+Female adult
Abkhazia, Sukhum Distr., N Slopes of Bzybian Mt. Range near Village Serebryanyi, 500-600 m, 25.IV-05.VII.2013, leg. D. Fominykh
Copyright © Igor Solodovnikov

Carabus (Neoplectes) pseudopshuensis Zamotajlov, 1991
Male+Female adult
Abkhazia, Sukhum Distr., N Slopes of Bzybian Mt. Range near Village Serebryanyi, 500-600 m, 25.IV-05.VII.2013, leg. D. Fominykh
Copyright © Igor Solodovnikov

Carabus (Neoplectes) pseudopshuensis Zamotajlov, 1991
Male+Female adult
Abkhazia, Sukhum Distr., N Slopes of Bzybian Mt. Range near Village Serebryanyi, 500-600 m, 25.IV-05.VII.2013, leg. D. Fominykh
Copyright © Igor Solodovnikov

Carabus (Neoplectes) pseudopshuensis Zamotajlov, 1991
Endophallus

Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus besleticus Kurnakov, 1972
Male+Female adult
Abkhazia, Sukhum Distr., vicinities of Village Kaman, N slope of Mt. Birtskha, 160 m, beech, linden, box, elder forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus besleticus Kurnakov, 1972
Male+Female adult
Abkhazia, Sukhum Distr., vicinities of Village Kaman, N slope of Mt. Birtskha, 160 m, beech, linden, box, elder forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus besleticus Kurnakov, 1972
Male+Female adult
Abkhazia, Sukhum Distr., vicinities of Village Kaman, N slope of Mt. Birtskha, 160 m, beech, linden, box, elder forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus besleticus Kurnakov, 1972
Male+Female adult
Abkhazia, Sukhum Distr., vicinities of Village Kaman, N slope of Mt. Birtskha, 160 m, beech, linden, box, elder forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus besleticus Kurnakov, 1972
Endophallus

Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972
Male+Female adult
Abkhazia, Gudauta Distr, vicinities of Mtsara (Chiryuta), Zashirbara Mt. Range, source of River Mtsara, beech, blackberry, fern site, 575-690 m, 11.V-05.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972
Male+Female adult
Abkhazia, Gudauta Distr, vicinities of Mtsara (Chiryuta), Zashirbara Mt. Range, source of River Mtsara, beech, blackberry, fern site, 575-690 m, 11.V-05.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972
Male+Female adult
Abkhazia, Gudauta Distr, vicinities of Mtsara (Chiryuta), Zashirbara Mt. Range, source of River Mtsara, beech, blackberry, fern site, 575-690 m, 11.V-05.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972
Male+Female adult
Abkhazia, Gudauta Distr, vicinities of Mtsara (Chiryuta), Zashirbara Mt. Range, source of River Mtsara, beech, blackberry, fern site, 575-690 m, 11.V-05.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972
Endophallus

Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus duripshensis Kurnakov, 1972
Male+Female adult
Abkhazia, Gudauta Distr., valley of River Khipsta, vicinities of Village Tvanaarkhu, karst crater with cave, beech, blackberry site, 425 m, N 43º14’ E 040º39’9”, 08.V-05.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus duripshensis Kurnakov, 1972
Endophallus

Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus napraensis Belousov & Zamotajlov, 1993 Paratype
Male+Female adult
Abkhazia, Gudauta Distr., Bzybian Mt. Range, vicinities of Mt. Chibzharga, 1900-2000 m, 20.VI-26.VII.1992, leg. I. Belousov, A. Zamotajlov, A. Miroshnikov
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus napraensis Belousov & Zamotajlov, 1993 Paratype
Female adult
Abkhazia, Gudauta Distr., Bzybian Mt. Range, vicinities of Mt. Chibzharga, 1900-2000 m, 20.VI-26.VII.1992, leg. I. Belousov, A. Zamotajlov, A. Miroshnikov
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus napraensis Belousov & Zamotajlov, 1993 Paratype
Endophallus

Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus anastasiae Solodovnikov, 2014 Holotype Paratype
Female adult
Abkhasia, Gudauta distr., S slope Bzybian Mt.R., upper reach. Riv. Khipsta, 43°18' N / 40°41' E, h = 1907-1915 m, subalpine meadows with karst blocks, 07-12.08.2014, leg. I.A. Solodovnikov, E.V. Tatun
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus anastasiae Solodovnikov, 2014 Paratype
Female adult
Abkhasia, Gudaura distr., S slope Bzybian Mt.R., upper reach. Riv. Khipsta, 43°18' N / 40°41' E, h = 1907-1915 m, subalpine meadows with karst blocks, 07-12.08.2014, leg. I.A. Solodovnikov, E.V. Tatun
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus anastasiae Solodovnikov, 2014 Paratype
Male adult
Abkhasia, Gudauta distr., S slope Bzybian Mt.R., upper reach. Riv. Khipsta, 43°18' N / 40°41' E, h = 1984-1990 m, subalpine meadows with big karst blocks, 07-12.08.2014, leg. I.A. Solodovnikov, E.V. Tatun
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus anastasiae Solodovnikov, 2014 Paratype
Female adult
Abkhasia, Gudauta distr., S slope Bzybian Mt.R., upper reach. Riv. Khipsta, 43°18' N / 40°41' E, h = 1984-1990 m, subalpine meadows with big karst blocks, 07-12.08.2014, leg. I.A. Solodovnikov, E.V. Tatun
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus anastasiae Solodovnikov, 2014 Paratype
Endophallus

Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014 Paratype
Female adult
Abkhazia, Bzybsky Mt.Rng, right bank of river Reshevie (left tributary of river Bzyb) h~670-720 м, 13-14.V.2010, D.Fominykh & I.Solodovnikov leg. L=41 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) starcki Heyden, 1885
Male+Female adult
NW-Caucasus,
Krasnodar reg., Mt. Aishkho, Malaya Laba River, h=2200m., 14.VII.2008, D.Fominyh leg.
female-21mm
Copyright © Ruslan Panin

Carabus (Neoplectes) starcki Heyden, 1885
Male adult
Caucasian Nat. Park, S slope
Pseashkho mt. 2500m 21.7.1975, Ovtsharenko leg.
20 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) starcki Heyden, 1885
Male adult
NW Caucasus, Adygeya, Pshekho-Su Mt. env., h=1500 m., 21.07.2006, D.Safronov leg.
Copyright © Safronov

Carabus (Neoplectes) starcki Heyden, 1885 (as parvulus)
Male adult
Prov. Kuban, mt. Aischa, VI.1906, Shaposhnikov leg.
det. D.V. Obydov

Copyright © Dmitri Obydov

Carabus (Neoplectes) starckianus ganglbaueri Reitter, 1888
Endophallus

Copyright © Alexander Anichtchenko

Carabus (Neoplectes) starckianus ganglbaueri Reitter, 1888
Male adult
Russia, NW Caucasus, Krasnodar reg., Adygeia distr., Guzeripl env. 20.5.2003
male, 26 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) starckianus ganglbaueri Reitter, 1888
Female adult
NW Caucasus, S Russia, Adygea Rep., Confluence of the river Belaya & Kisha h~700m, 24.V-7.VIII.2010, D.Fominyh leg.
female,32 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) starckianus adelaidae Starck, 1889
Male adult
Russia, Caucasus, Chvizhepse River Val., 27.V.1994, P. Gorbachev leg.
Photo from www.zin.ru
det. D.V. Obydov
Copyright © Dmitri Obydov

Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988 (as shchessiensis) Paratype
Male+Female adult
NW Caucasus,
Krasnodar reg., Tuapsinskyi distr. Shessi Mt.
30.V.-25.VI.1993 Mt.Shessi, h=1800m., Imura & A.S.Zamotajlov, 1993 leg.
female, 29mm
Copyright © Ruslan Panin

Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988
Female adult
Russia,W Caucasus,
Krasnodar reg.,Chernogorie mts.,Chernogor mt. h=1700-1800m. 23.V.-30.VII.1999. A.S.Zamotajlov leg.
female 29mm.
Copyright © Ruslan Panin

Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988
Male adult
West. Caucasus, Shakhe Valley, 1.VI.1989, A. Shamaev leg.
det. D.V. Obydov

Copyright © Dmitri Obydov

Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
Male adult
NW-Caucasus, Russia,
Krasnodar reg., Apsheron Distr., Lagonaki Mt. ,near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominykh leg.
male, 28mm
Copyright © Ruslan Panin

Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
Male+Female adult
NW-Caucasus, Russia,
Krasnodar reg.,the right bank of Mzymta river, Engelmanovs glades, h=1200m., A.S.Zamotajlov leg.
female-30mm\male-28mm
Copyright © Ruslan Panin

Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
Female adult

Copyright © Safronov

Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
Male adult
Rusia, W Caucasus, Sochi distr., Aishkha mts.
900m, 15.7.2003 Gorbachev leg.
30 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) starckianus starckianus Ganglbauer, 1886
Male adult
Russia, Krasnodar reg., 6 km NE Soloniki vill.,
Sochi Nat.Park, 28-30.V.2009 Anichtchenko A. leg.

Copyright © Alexander Anichtchenko

Carabus (Neoplectes) starckianus starckianus Ganglbauer, 1886
Female adult
W Caucas, Sochi distr., near Lazorevskoe vill., 9.06.1993, P.Gorbachev leg.
det. D.V. Obydov


Copyright © Dmitri Obydov

Carabus (Neoplectes) starckianus starckianus Ganglbauer, 1886 (as phlagochensis)
Female adult
Caucasus, Sotschi, Flagoch.
det. D.V. Obydov
from www.zin.ru

Copyright © Dmitri Obydov

Carabus (Neoplectes) starckianus theseus Briansky, 1910
Male+Female adult
NW-Caucasus, Russia,
Krasnodar reg.,Tuapse distr., Mt.Semashkho 04.V.2008, D.Fominyh leg.
female, 32mm
Copyright © Ruslan Panin

Carabus (Neoplectes) starckianus theseus Briansky, 1910
Male adult
Tuapse distr., Anastasievka vill., Semiglavaya mt.
riv. Pshekaho 20.7.1995
30 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) starckianus theseus Briansky, 1910
Female adult
Tuapse distr., Anastasievka vill., Semiglavaya mt.
bass. Pshekaho riv. 20.7.1995
female, 30 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
Endophallus

Copyright © Ruslan Panin

Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988 (as shchessiensis) Paratype
Endophallus

Copyright © Ruslan Panin

Carabus (Neoplectes) starckianus starckianus (as phlagochensis)
Female adult
Russia, Кrasnodar Reg., Dagomys, Flagokh, N43-44' / E39-40', 11.VII.2010, N. Pichuginleg.
Specimen from the collection N. U. Pichugin (Vladimir).
Copyright © Dmitri Obydov

Carabus (Neoplectes) starckianus olgae Semenov, 1898 Lectotype
Male adult
[Circassia. Rost. Car. imperatot St.] [44] [Plectes Olgae m. A. Semenov det., ♂ Typ. un. X.96] red label этикетка [LECTOTYPUS Gottwald, 1983 design], yellow label [Zoological Institute Russian Academy of Sciences St Peterburg]
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus olgae Semenov, 1898
Label

Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus olgae Semenov, 1898 Paralectotypes
Male adult
• [Circassien. Rost. [18]98 Pl. obtusus var. 5.] [5] [Plectes Olgae m. A. Semenov det., ♂ I.98] red label [PARALECTOTYPUS Gottwald, 1983 design], yellow label [Zoological Institute Russian Academy of Sciences St Peterburg]
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus olgae Semenov, 1898
Label

Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus starckianus Ganglbauer, 1886
Male adult
Russia, Krasnodar area, Sochi, Sochi distr., right bank of Riv. Persiyanka, NE slope Phlagokh Mt., 03.05-02.07.2013, Leg. I.A. Solodovnikov, S.V. Solodovnikova (TOPOTYPUS as phlagochensis Imura & Zamotajlov, 1993)
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus theseus Briansky, 1910
Male adult
Russia, Krasnodar Territory, Tuapse district, env. vill. Psebe, left bank of the Psebe River, hornbeam + oak + walnut, h = 146 m, 03.05-01.07.2014, leg. I.A. Solodovnikov, S.V. Solodovnikova, S.M. Pavlyuchuk
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus theseus Briansky, 1910
Female adult
Russia, Krasnodar Territory, Tuapse district, env. vill. Psebe, left bank of the Psebe River, hornbeam + oak + walnut, h = 146 m, 03.05-01.07.2014, leg. I.A. Solodovnikov, S.V. Solodovnikova, S.M. Pavlyuchuk
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
Female adult
Russia, Krasnodar Territory, West-Caucasus, Apsheronsky district, Guam ridge, env. Mezmay village, by the road, beech, oak, blackberry, h = 1200-1250 m, 03.IV-14.VI.2020, leg. S.M. Pavlyuchuk, A.P. Tatulyan
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
Female adult
Russia, Krasnodar Territory, West-Caucasus, Apsheronsky district, Guam ridge, env. Mezmay village, by the road, beech, oak, blackberry, h = 1200-1250 m, 03.IV-14.VI.2020, leg. S.M. Pavlyuchuk, A.P. Tatulyan
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
Male adult
Russia, Krasnodar Territory, West-Caucasus, Apsheronsky district, Guam ridge, env. Mezmay village, by the road, beech, oak, blackberry, h = 1200-1250 m, 03.IV-14.VI.2020, leg. S.M. Pavlyuchuk, A.P. Tatulyan
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
Male adult
Russia, Krasnodar Territory, West-Caucasus, Apsheronsky district, Guam ridge, env. Mezmay village, by the road, beech, oak, blackberry, h = 1200-1250 m, 03.IV-14.VI.2020, leg. S.M. Pavlyuchuk, A.P. Tatulyan
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988 Paratype
Male adult
Krasnodar area, Babuk-Aul, 19.06.1983 , leg. A. Miroshnikov
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
Female adult
W Caucasus, NW slope Aibga Mt.R. vall. riv. Galion, h = 650 m, Fagus, Alnus viridis forest, 18-24.VI.1999, I.A. Solodovnikov leg.
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
Male adult
W Caucasus, Aibga Mt.R., 02.VIII.1994, A.S. Zamotailov leg.
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
Male adult
Russia, W Caucasus, Krasnaya Polyana, Atchischkho Mt.R., h = 1200 m, 10.06.1995, N.Yu. Pichugin leg.
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus adelaidae Starck, 1889
Female adult
Russia, Krasnodar area, Adler, Ader distr., right bank vall. Riv. Msymta, near vill. Chvesipse, h = 800 m, 06.VII.1999, B. Saratovskikh leg.
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus adelaidae Starck, 1889
Female adult
Russia, Krasnodar area, Adler, Ader distr., right bank vall. Riv. Msymta, near vill. Chvesipse (Fagus+Quercus+Castanea forest) [Sochi national park], h = 720 m, 03.V-08.VII.2016, I.A. Solodovnikov, S.V.Solodovnikova, V.M.Kotsur leg.
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus adelaidae Starck, 1889
Female adult
Russia, Krasnodar area, Adler, Ader distr., right bank vall. Riv. Msymta, near vill. Chvesipse (Fagus+Quercus+Castanea forest) [Sochi national park], h = 720 m, 03.V-08.VII.2016, I.A. Solodovnikov, S.V.Solodovnikova, V.M.Kotsur leg.
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus adelaidae Starck, 1889
Male adult
Russia, Krasnodar area, Adler, Ader distr., right bank vall. Riv. Msymta, near vill. Chvesipse (Fagus+Quercus+Castanea forest) [Sochi national park], h = 720 m, 03.V-08.VII.2016, I.A. Solodovnikov, S.V.Solodovnikova, V.M.Kotsur leg.
Copyright © Igor Solodovnikov

Carabus (Archiplectes) prometheus zolotarevi Zamotajlov, 1988
Male adult
Paratypus, male Prov. Kuban distr., Maikop, Laba Min., Ch. Shaposhnikov leg.
from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Archiplectes) prometheus zolotarevi Zamotajlov, 1988 Paratype
Male adult
Paratypus, male - Fl. Min. Laba, VII.1918, Shaposhnikov leg.
from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Neoplectes) zolotarevi Zamotajlov, 1988
Female adult
Carabus (Archiplectes) zolotarevi zolotarevi Zamotajlov, 1988 Russia: Krasnodar reg., SW slopes of Malye Balkany Mt.R., env. Allous pass, h = 1960 m, 08-18.VII.2023 leg. D. Fominykh
Copyright © Igor Solodovnikov

Carabus (Neoplectes) zolotarevi Zamotajlov, 1988
Endophallus
Carabus (Archiplectes) zolotarevi zolotarevi Zamotajlov, 1988 Russia: Krasnodar reg., SW slopes of Malye Balkany Mt.R., env. Allous pass, h = 1960 m, 08-18.VII.2023 leg. D. Fominykh
Copyright © Igor Solodovnikov

Carabus (Neoplectes) zolotarevi Zamotajlov, 1988
Male adult
Carabus (Archiplectes) zolotarevi zolotarevi Zamotajlov, 1988 Russia: Krasnodar reg., SW slopes of Malye Balkany Mt.R., env. Allous pass, h = 1960 m, 08-18.VII.2023 leg. D. Fominykh
Copyright © Igor Solodovnikov