Acinopus (Acinopus) picipes (Olivier, 1795)
Female adult

Copyright © Kirill Makarov

Acinopus (Osimus) ammophilus Dejean, 1829
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Acinopus (Oedematicus) megacephalus (P. Rossi, 1794)
Male adult

Copyright © Kirill Makarov

Acinopus (Haplacinopus) striolatus Zoubkoff, 1833: 317
Male adult
Caspian sea, 22.6.91; 18 mm
Copyright © Alexander Anichtchenko

Acinopus (Oedematicus) gutturosus Buquet, 1840
Male adult
Spain, Algeciras, San Roque,4.2014, Anichtchenko A. leg.; det. Anichtchenko A. 2018
Copyright © Alexander Anichtchenko

Acinopus (Acinopus) giganteus Dejean, 1831
Male adult
Spain, Andalucia, Sierra de las Nieves, Los Quejigales, 6-8.4.2010, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Acinopus (Acmastes) haroldii Schaum, 1863
Male adult
Spain, Cadiz, Tarifa, 10.9.1989, J.L. Torres (cAA) 15 mm
Copyright © Alexander Anichtchenko

Acinopus (Acinopus) sabulosus Fabricius, 1792
Male adult
Marruecos, Oukaimeden distr., Ait-Lekak vill vic., 21.6.2010 1650m, N31.16.58 W007.49.17, Anichtchenko A. leg. & det. (cAA) 12 mm
Copyright © Alexander Anichtchenko

Acinopus (Acinopus) almeriensis Mateu, 1954
Male adult
Spain, Almeria, Cabo de Gata, playa de San Jose 20.4.2004 Anichtchenko A. leg & det. (cAA) 14mm
Copyright © Alexander Anichtchenko

Afromizonus tecospilus Basilewsky, 1947
Female adult
Gambia, Janjangbure env, 19-22.VII.2003 B.Vořišek leg.
Copyright © Rudolf Kmeco

Allosiopelus fulvicollis Ito, 2016 Holotype
Male adult
Coimbatore, alt. 1,400 ft., Madras, Indien, XI.1966, P. S. Natahn leg. (Zoologische Staatssammlung, München.) 8.3 mm
Copyright © - - -

Amblygnathus mexicanus Bates, 1882
Male adult
Louisiana, Webster Parish
Photo I. Sokolov
Copyright © Igor Sokolov

Amblygnathus cephalotes Dejena, 1829
Unsexed
Antilles. Image from Shpeley, Hunting & Ball, 2017
Copyright © - - -

Athrostictus paganus Dejean, 1831
Male adult
Antilles. Image from Shpeley, Hunting & Ball, 2017
Copyright © - - -

Athrostictus sulcatulus Dejean, 1829
Male+Female adult
Iguazu, Misiones Province, I-2010, A. Cicchino Coll. & det.
Copyright © Armando Cicchino

Athrostictus punctatulus (Putzeys, 1878)
Male adult
Image from Messer & Raber, 2021
Copyright © - - -

Axinotoma hulstaerti Basilewsky, 1968
Male adult
Gabon, Makokou 7.7 km SW, Ivindo NP, Ipassa forest research s., 0*30.52N 12*48'21E, 21-22.6.2016, secondary lowland rainforest, MV light, D.Telnov leg.10 mm. det. S. Facchini, 2019
Copyright © Alexander Anichtchenko

Aztecarpalus trochotrichis Ball, 1970 Holotype
Male adult
Mexico, Oaxaca. 88.5 mi s. Valle Nacional. 8000'. V-2-1966; GeorgeE. Ball, D. R. Whitehead collectors.
Copyright © - - -

Aztecarpalus schaefferi Ball, 1970
Female adult
TEXAS: Travis County Camp Mabry Texas ANG N 30.82; W -97.76 16.IV-7.V.2004; J.C. Abbott UTIC Database #205365 (This image was created as part of the Insects Unlocked project at the University of Texas at Austin)
Copyright © - - -

Boeomimetes confusus Basilewsky, 1948
Male adult
10,3 mm
Namibia, Etosha National Park, Okaukuejo, 1400 m, leg. U.Schmidt, 18.-20.III.1994;
det. M.Baehr, 2012 Photo: U.Schmidt, 2013
Copyright © Udo Schmidt

Boeomimetes ephippium (Boheman, 1860)
Female adult
Namibia, Otjozondjupa Reg. 20.5 km NW from Grootfontein 19°26'19"S 17°57'49"E 20.II.2018 leg. A. Rubenyan L = 12.1 mm
Copyright © Kirill Makarov

Bradybaenus scalaris Olivier, 1808
Male adult
Sénégal, Nianing, piège lumineux, 14/09/2009 Pascal Deschamps coll.
Copyright © Pascal Deschamps

Bradybaenus opulentus Boheman, 1848
Female adult
NAMIBIA, Oshana reg., 67,5 km NNW from Outjo, Toshari Lodge territory, H=1205 m S19°33.390' E15°52.679' 2-5.XII.2012 leg. E.Shankhiza
L=12.0 mm
Copyright © Kirill Makarov

Bradybaenus perrieri Jeannel, 1948
Male adult
MADAGASCAR SUD, 27.1.1995 Ambilalialika,P. Schüle det.
Copyright © Peter Schüle

Coleolissus (Coleolissus) biakensis Baehr, 2018 Holotype
Male adult
INDONESIA, Papua Biak, Mniber, 16-22.12.06, 00.43.28S 135.46.01E leg. O. Mehl (CBM). 10.6 mm.
Copyright © Martin Baehr

Coleolissus inessae Kataev, 2021 Holotype
Male adult
INDIA: W Karnataka, W Gats Mts, jog Falls, 530 m, at light, 14°13.240´N 74°48.471´E, 2–5.XI.2013 S. Saluk leg. (ZIN).
Copyright © - - -

Cratacanthus dubius Palisot de Beauvoi, 1811
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Cratognathus grandiceps Boheman, 1860
Male adult
Namibia, Otjiwarongo, Kalkfeld, 30 km east, Mt.Etjo Safari Lodge, 1500 m, leg. U.Schmidt, 15.III.1994
det. M.Baehr, 2012
Photo: U.Schmidt, 2014
7,8 mm
Copyright © Udo Schmidt

Daptus pictus Fischer von Waldheim, 1823
Male adult
Kazakhstan, 20 km NW Akkuduk vill., N 43*06'58 E 054*11'46, h=90m, 24.V.2011, Ivanov A. leg.
det. Anichtchenko A. 2013
Copyright © Alexander Anichtchenko

Dioryche yunnana Kataev, 2002
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Dioryche subrecta Kataev, 2012
Male adult

Copyright © Kirill Makarov

Discoderus parallelus Haldeman, 1843
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Discoderus cyaneopacus (Darlington, 1934)
Unsexed
Haiti. Image from Shpeley, Hunting & Ball, 2017
Copyright © - - -

Discoderus thoracicus Putzeys, 1878
Male adult
Haiti. Image from Shpeley, Hunting & Ball, 2017
Copyright © - - -

Discoderus beauvoisii (Dejean, 1829)
Unsexed
Image from Shpeley, Hunting & Ball, 2017
Copyright © - - -

Euryderus grossus (Say, 1830)
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Harpalinus bekilyanus Jeannel, 1948
Male adult
MADAGASCAR, Tulear Prov., nr. Tsimanampetsotsa NP, P. Schüle det.
Copyright © Peter Schüle

Harpalobrachys leiroides (Motschulsky, 1844)
Female adult
Russia, Burjatia, Tunka valley, Tunka vill. vicinity, 17.VII.1995. Anichtchenko A. leg. 12 mm det. Anichtchenko A.
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) caspius (as roubali Schauberger, 1928)
Male adult
NW Podolian Upland, W UKRAINE,<br /> Lviv Reg., Zolochiv distr., near Gologirki vill., 30.04-19.05.07 Panin R.leg.<br /> L= 15mm
Copyright © Ruslan Panin

Harpalus (Harpalus) laticeps Leconte, 1850
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Harpalus (Harpalus) mitridati Pliginskiy, 1915
Male adult
Rostov Area, Kumyska Vill. 24.III.2006 leg. M.Nabozhenko
L=8.5
Copyright © Kirill Makarov

Harpalus (Plectralidus) eraticus Say, 1823
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Harpalus (Harpalus) affinis (Schrank, 1781)
Female adult
RUSSIA, Tver reg., Likhoslavlskiy distr., 900 m to NW from Domanikha village, grassland. H=161 m. N57*13.964’ E035*45.033’ 7-15.VII.2005 leg., det. & photo - Vlasenko A. S. L=10,4 mm
Copyright © Andrey Vlasenko

Harpalus (Harpalus) egorovi Lafer, 1989
Male adult

Copyright © Kirill Makarov

Harpalus (Harpalus) amplicollis Menetries, 1848
Female adult

Copyright © Kirill Makarov

Harpalus (Harpalus) brevicornis Germar, 1824
Male adult

Copyright © Kirill Makarov

Harpalus (Loboharpalus) rubefactus Bates, 1873
Male adult
det. Makarov K.
Copyright © Kirill Makarov

Harpalus (Zangoharpalus) tinctulus Bates, 1873
Male adult
Collection ground Sapporo city, Hokkaido Pref. pref. 33415
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Loboharpalus) platynotus Bates, 1873
Male adult
Collection ground Suttsu cho, Hokkaido Pref. 33499
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) calathoides Motschulsky, 1844: 200
Male adult

Copyright © Kirill Makarov

Harpalus (Harpalus) cupreus fastuosus Faldermann, 1836
Male adult
NW Caucas, Krasnodar reg., SE Goryachii Kluch, леса между пос. Асфальтовая Гора и Садовый (нежилой) 21.V.2012 leg. М. Смирнов
Image of M. Smirnov (М.Э. Смирнов) from www.zin.ru\Animalia\Coleoptera
Copyright © Maksim Smirnov

Harpalus (Harpalus) crates Bates, 1883
Male adult
SE Maritime Prov., Lazovsky dist., Lazo env., 17-20.VI.2010, leg. М. Smirnov Фотография с сайта zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Harpalus (Harpalus) dispar splendens (Gebler, 1829)
Male adult

Copyright © Kirill Makarov

Harpalus (Cephalophonus) cephalotes Fairmaire & Laboulbene, 1854
Male adult
SE Bulgaria, Primorsko; 16mm
Copyright © Aleš Sedláček

Harpalus (Harpalus) pulvinatus lubricus Reitter, 1900
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) kryzhanovskii Kataev, 1988: 772
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Pseudoophonus) ussuriensis ussuriensis s.str.
Female adult

Copyright © Kirill Makarov

Harpalus (Pseudoophonus) capito A.Morawitz, 1862
Female adult

Copyright © Kirill Makarov

Harpalus (Harpalus) caspius (as roubali Schauberger, 1928)
Male adult
Moravia, Kobylí; 11 mm
Copyright © Aleš Sedláček

Harpalus (Baeticoharpalus) lopezi Serrano & Lencina, 2008
Male adult
Spain, Jaen prov., Sierra Magina, 2000m, 37.728701, -3.460772 3-4.05.2014, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) reflexus anadoluensis Kataev, 1993
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Cryptophonus) tenebrosus Dejean, 1829
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Harpalus (Harpalus) solitaris Dejean, 1829a: 337
Female adult
W-Ukraine, Transcarpathian reg., Chornagora Massif, 8 km NE Bohdan vill., h=800m, 9-29.VI..2014, Panin R. leg. L=10.5 mm
Copyright © Ruslan Panin

Harpalus (Semiophonus) signaticornis (Duftschmid, 1812)
Male adult
France, 77860, Quincy-Voisins, 12-05-2018, leg. & det. P. Deschamps
Copyright © Pascal Deschamps

Harpalus (Harpalus) lederi Tschitscherne, 1899
Female adult
Buryatia, Hamar-Daban Mts., Taltsy riv., 1700m 19-25.5.1997, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Harpalus (Cryptophonus) melancholicus Dejean, 1829
Male adult
Italy, Patro di Moncalvo (Asti).
Copyright © Gianni Allegro

Harpalus (Anophonus) pterostichus (Reitter, 1900)
Male adult
Uzbekistan, Chatkal Mt. rng., Irtash, 41°10'25.7"N 70°16'20.4"E, 5-7.8.2021, 2400m, A. Anichtchenko leg. (cAA). 10mm; det. I. Kabak, 2021
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) nevadensis K. Daniel & J. Daniel, 1898
Male adult
Spain, Sierra Nevada, 5km N Capileira, 1800-2000m, 25.4.2003, Anichtchenko A. leg & det. (cAA) 9 mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) pseudoserripes Reitter, 1900
Female adult
Iran, Khorasan, Torbat-E-Heydariyeh, pass 5 km N Zharf 23.5.2008, Anichtchenko A. leg (cAA) 10 mm; det. Kataev B.
Copyright © Alexander Anichtchenko

Harpalus (Opadius) cordifer Notman, 1919
Female adult
USA, Washington, Thurston Co., 19.8.2005 (cAA) 8.5mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) optabilis Dejean, 1829
Male adult
Burjatia, Selenga riv., Deben vic., 15.6.2003 Anichtchenko A. leg & det. (cAA) 12 mm
Copyright © Alexander Anichtchenko

Harpalus (Harpalus) compressus Motschulsky, 1844: 213
Male adult
SW Ukraine, Odessa reg., Ivanovka distr., Gudevichevo vill., steppe, 13.VI-11.VII.2022, A.Gontarenko leg. L=12 mm
Copyright © Ruslan Panin

Harpalus (Cryptophonus) fulvus Dejean, 1829
Male adult
Italy, Sardinia, Chia (CA), Stagno Campana, 29.VIII.1994
Copyright © Gianni Allegro

Harpalus (Harpalus) asemus Basilewsky, 1946
Male adult
left - Yemen; right - Tanzania. Scale bars = 1.0 mm. Image from Kataev, 2021
Copyright © - - -

Harpalus (Opadius) cordatus (Leconte, 1853)
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Glanodes) obliquus G.Horn, 1880
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Smirnovia) kandaharensis Jedlicka, 1956
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Idioharpalus) numidicus Bedel, 1893
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Hemipangus) badakschanus Jedlicka, 1956
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Loxophonus) agakhaniantzi (Mikhailov, 1972)
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Brachyharpalus) kazanensis Jedlicka, 1958
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Brachypangus) antonowi (Tschitscherine, 1898)
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Hypsinephus) salinus Dejean, 1829
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Pharalus) indianus Csiki, 1932
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Aristoharpalus) ingenuus Tschitscherine, 1898
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Anophonus) cyanopterus (Tschitscherine, 1897)
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Megaharpalus) stoetznerianus Schauberger, 1932
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Oreoharpalus) famelicus Tschitscherine, 1898
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Hypsoharpalus) arnoldii Kataev, 1988
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Mesoharpalus) kiritshenkoi Kataev, 1990
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Eremoharpalus) medvedevi Kataev, 2006
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Nephoharpalus) pallidipennis A. Morawitz, 1862
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Diaharpalus) vittatus Gebler, 1833
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Harpalobius) viridanus Motschulsky, 1844
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Bactroharpalus) obnixus Casey, 1924
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Homaloharpalus) tangutorum Kataev, 1993
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Asioharpalus) nigrans A. Morawitz, 1862
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Anamblystus) torridoides Reitter, 1900
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Platyharpalus) ventralis Leconte, 1848
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Psammoharpalus) kozlovi Kataev, 1993
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Acardystus) flavescens (Piller & Mitterpacher, 1783)
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Euharpalops) fraternus Leconte, 1852
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Calathoderus) potanini Tschitscherine, 1906
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Epiharpalus) punctipennis Mulsant, 1852
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Caucasoharpalus) aeneipennis (Faldermann, 1836)
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Drymoharpalus) atratus Latreille, 1804
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Meroharpalus) fulvilabris Mannerheim, 1853
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Sinoharpalus) hiekei Kataev & Wrase, 2010
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Macroharpalus) animosus Casey, 1924
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Afroharpalus) natalensis Boheman, 1848
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Hyloharpalus) tibeticus Andrewes, 1930
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Zangoharpalus) praticola Bates, 1891
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Plectralidus) retractus Leconte, 1863
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Megapangus) katiae F.Battoni, 1985
Male adult
Image from Kataev, 2023
Copyright © - - -

Harpalus (Acardystus) flavescens (Piller & Mitterpacher, 1783)
Female adult
W Ukraine, Lviv city, Holosko, Bryukhovytsʹkyi forest, on the sands, 17.V-25.VI.2024, R.Panin leg., L=13 mm
Copyright © Ruslan Panin

Hartonymus hoodi Casey, 1914
Male adult
United States, Illinois, Mason, Sand Ridge State Forest Campground, 148m, 40.3901, -89.8858, Davidson, R. 26 Oct 2010
Copyright © Smithsonian NMNH

Heteracantha depressa Brulle, 1834
Male adult
Algeria, Ghardala
19.5.1960 G. Ledoux leg.
20 mm
Copyright © Alexander Anichtchenko

Hyphaereon (Hyphaereon) lautulus Andrewes, 1929 Holotype
Male adult
7.5mm. Natural History Museum (2014). Dataset: Collection specimens. Resource: Specimens. Natural History Museum Data Portal (data.nhm.ac.uk). © The Trustees of the Natural History Museum, London
Copyright © - - -

Kareya tibiale (Laferte-Senectere, 1853)
Female adult
Cameroon. Scale 1 mm. Image from Kataev, 2015
Copyright © - - -

Kareya laosensis (Jedlicka, 1966)
Male+Female adult
Laos. Scale 1 mm. Image from Kataev, 2015
Copyright © - - -

Xenodochus exaratus (Dejean, 1829)
Female adult
Sénégal, Nianing piège lumineux 21/09/09, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Meroctenus (Meroctenus) crenulatus orientalis Kataev, 2017
Male adult
Pakistan, Kabul river. Scale bar 5mm
Copyright © - - -

Meroctenus (Meroctenus) mediocris (Andrewes, 1936) Lectotype
Male adult
Scale bar 5 mm
Copyright © - - -

Microderes (Neopangus) breviformis (Tschitscherine, 1898)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Microderes (Microderes) brachypus (Steven, 1809)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Neoaulacoryssus cupripennis (Gory, 1833)
Female adult
Antilles. Image from Shpeley, Hunting & Ball, 2017
Copyright © - - -

Neodiachipteryx davidsoni Shpeley, Hunting & Ball, 2017 Holotype
Male adult
DOMINICAN/ REPUBLIC/ Sabaneta Prov./ Santiago Rodrigues/ Zamba/ August 2, 1980 (CMNH). Image from Shpeley, Hunting & Ball, 2017
Copyright © - - -

Neodiachipteryx carinigera (Putzeys, 1878)
Male adult
Antilles, Hispaniola. Image from Shpeley, Hunting & Ball, 2017
Copyright © - - -

Neophygas microcephalus (Faldermann, 1835)
Male adult
Mongolia, Ovorhangai aimak, near Hayrhandulaan, 18.7.2006, Anichtchenko A. leg.; 9 mm
Copyright © Alexander Anichtchenko

Nesarpalus (Nesacinopus) solitarius (Wollaston, 1863)
Male adult
Spain, Canaries, Lanzarote, Teguise 18-03-2014, Pascal Deschamps coll. & det.
Copyright © Pascal Deschamps

Nesarpalus (Nesacinopus) fortunatus (Wollaston, 1863)
Male adult
12,5 mm; Spain, Canary Islands, Gran Canaria, Ayacata, 5 km NE leg. J.Schoenfeld, 18.IV.2008; det. M.Baehr, 2008; Photo: U.Schmidt, 2014
Copyright © Udo Schmidt

Nipponoharpalus discrepans (A.Morawitz, 1862)
Male adult
Collection ground Chitose city, Hokkaido Pref. pref. 34168
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Omostropus mandibularis (Roth, 1851)
Female adult
RSA, Nothern prov., Louis Trichardt Ben Lavin Nat. Res., 29.I.2000. det. S. Facchini 2003. 8 mm
Copyright © Alexander Anichtchenko

Ooidius madecassus (Jeannel, 1948)
Female adult
MADAGASCAR, Tulear Prov., nr. Tsimanampetsotsa NP, P. Schüle det.
Copyright © Peter Schüle

Ooidius dorsiger (Klug, 1853)
Female adult
11,3 mm. Namibia, Etosha National Park, Okaukuejo, 1400 m, leg. U.Schmidt, 19.III.1994; det. M.Baehr, 2014. Photo: U.Schmidt, 2016
Copyright © Udo Schmidt

Ophoniscus nepalensis Kataev & Wrase, 2012 Holotype
Male adult

Copyright © Kirill Makarov

Ophonus (Metophonus) gammeli (Schauberger, 1932)
Male adult
Stavropol Reg., env. Zheleznovodsk 1.06.2007 S.Svetlov leg.
male
L=11,0 mm
Copyright © Maksim Smirnov

Ophonus (Ophonus) stictus Stephens, 1828
Male adult

Copyright © Kirill Makarov

Ophonus (Hesperophonus) cribricollis (Dejean, 1829)
Male adult
SK, S Slovakia, Kamenica nad Hronom; 8mm
Copyright © Aleš Sedláček

Ophonus (Hesperophonus) azureus (Fabricius, 1775)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Ophonus (Hesperophonus) convexicollis Menetries, 1832
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Ophonus (Metophonus) rufibarbis (Fabricius, 1792)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Ophonus (Metophonus) cordatus (Duftschmid, 1812)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Ophonus (Metophonus) puncticeps Stephens, 1828
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Ophonus (Ophonus) stictus Stephens, 1828
Female adult
W Ukraine, NW Podolian Upland, Lviv reg., Brody distr., Makitra Mt, steppe site, 15.V-15.VI.2017, Panin R. & Kanarsky Yu. leg. L=15 mm
Copyright © Ruslan Panin

Ophonus (Brachyophonus) vignai Sciaky, 1987 Paratype
Female adult
Turchia, vil. Antalya, dint. Catallar 28.IV.1973 Leg. Vigna Taglianti (coll. R. Sciaky)
Copyright © Riccardo Sciaky

Ophonus (Incisophonus) incisus (Dejean, 1829)
Female adult
France Gard dep., Nimes 3.11.2013 (cAA) 14 mm
Copyright © Alexander Anichtchenko

Ophonus (Hesperophonus) chlorizans Solsky, 1874
Female adult
Kirgizstan, Chandalash mt. rng, Chakmaksuu riv. 2500m, 5.6.2000 (DUBC) 7. 5mm
Copyright © Alexander Anichtchenko

Ophonus (Metophonus) cunii Fairmaire, 1880
Male adult
Spain, Andalicia, Sierra de Grazalema, Puerto de las Palomas, 10.4.2010, Anichtchenko A. leg. & det. (cAA) 9.5 mm
Copyright © Alexander Anichtchenko

Ophonus (Metophonus) rupicola (Sturm, 1818)
Male adult
Russia, NW Caucasus, Krasnodar reg., Taman env., 23.4.2000 (DUBC)
Copyright © Alexander Anichtchenko

Ophonus (Metophonus) berberus Antoine, 1925
Male adult
Morocco, Atlas mts., pass near Oukaimeden, N31.10.33 W 007.50.38, 2800-2900m, 18-20.6.2010, Anichtchenko A. leg. & det. (cAA). 7 mm
Copyright © Alexander Anichtchenko

Ophonus (Metophonus) laticollis Mannerheim, 1825
Male adult
Russia, NW Caucasus, Novorossiisk distr., Markoth Mts., 23.5.2005 (DUBC) det. Anichtchenko A. 2021; 10.5mm
Copyright © Alexander Anichtchenko

Ophonus (Metophonus) subsinuatus Rey, 1886
Male adult
WС Ukraine, Vinnytsa reg., Lityn distr., Petryk vill., to light, 5.VI.2020, S. Kokhan leg.,/det.R.Panin, L=7,6mm
Copyright © Denis Khrapov

Oxycentrus (Oxycentrus) argutoroides (Bates, 1873)
Male adult
Collection ground Tsushima, Nagasaki pref. 35695<br /> (C) Working Group for Biological Indicator Ground Beetles Database, Japan<br /> Prohibited secondary use of all data
Copyright ©

Oxycentrus (Oxycentrus) minutopunctatus N.Ito, 1998
Female adult
Philippines, Mindanao, Aliwagwag, 7.740420, 126.301397, light trap, 14-18.vi.2024, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Oxycentrus (Oxycentrus) minutopunctatus N.Ito, 1998
Male adult
Philippines, Mindanao, Aliwagwag falls, Cateel riv., Davao Oriental, 7.740266, 126.301608, 200m, 4-7.v.2023, Anichtchenko A. leg. & det. (cAA).
Copyright © Alexander Anichtchenko

Oxycentrus (Oxycentrus) planibasis N.Ito, 1998
Female adult
Philippines, Mindanao, Ugwad Falls (Kapuka), Caraga, Davao Or., 260m, 7.440359, 126.434027, 29-30.IV.2023, Anichtchenko A. leg. & det. (cAA).
Copyright © Alexander Anichtchenko

Pangus scaritides (Sturm, 1818)
Male adult
SK, S Slovakia, Velké Kosihy; 8mm
Copyright © Aleš Sedláček

Parasiopelus ornatus (Peringuey, 1892)
Male adult
Zimbabwe, Masvingo province, 95km NE Beitbridge, Bubi river, 21*42S 30*29E, 12-27.XII.2005. det. P. Schule
Copyright © Alexander Anichtchenko

Paraulacoryssus puertoricensis Mutchler, 1934
Unsexed
Puerto Rico. Image from Shpeley, Hunting & Ball, 2017
Copyright © - - -

Parophonus (Hyparpalus) holosericeus (Dejean, 1829)
Female adult
Sénégal, Nianing, piège lumineux, 18/09/2009, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Parophonus (Parophonus) planicollis (Dejean, 1829)
Female adult

Copyright © Alexander Anichtchenko

Parophonus (Parophonus) mendax (P.Rossi, 1790)
Male adult
Bulgaria. Chernomerets, 24.XI.2006, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Parophonus (Parophonus) hispanus (Rambur, 1838)
Male adult
Spain, Andalicia, San Pedro de Alcantara, 7.4.2012, Anichtchenko A. leg. & det. (cAA) 7.5 mm
Copyright © Alexander Anichtchenko

Parophonus (Ophonomimus) interstitialis (Reitter, 1889)
Male adult
S Tadzhikistan, Kurgan-Tube reg., 20km S Dzhilikul, 11.5.1999 (DUBC) 8 mm
Copyright © Alexander Anichtchenko

Parophonus (Ophonomimus) hirsutulus (Dejean, 1829)
Male adult
S Tadzhikistan, Kurgan-Tube reg., 20km S Dzhilikul, 11.5.1999 (DUBC) 8.5mm
Copyright © Alexander Anichtchenko

Parophonus (Orphanixus) pierroni Jeannel, 1948
Male adult
Zimbabwe, Matabele South, Bubi riv., 95km NE Beitbridge 507m, 25.11.2006 (cAA) det Facchini ; 8mm
Copyright © Alexander Anichtchenko

Penthophonus glasunovi (Tschitscherine, 1898)
Female adult
Iran, Khamadan prov., 1 km E Ylfan vill. 34.740476°N 48.621117°E 10-11.V.2017 leg. D.Kasatkin L=11.5 mm
Copyright © Kirill Makarov

Piosoma setosum LeConte, 1848
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Platymetopus flavilabris (Fabricius, 1798)
Male adult
Collection ground Higashisonogi cho, NagasakiPref. 35330
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Platymetopus tessellatus Dejean, 1829
Female adult
Sénégal, M’Bour, piège lumineux, 5/10/2010, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Platymetopus quadrimaculatus Dejean, 1829
Male adult
Vietnam. Image from Kataev, 2023
Copyright © - - -

Platymetopus figuratus pictus Andrewes, 1923
Unsexed
India, Rajastan. Image from Kataev, 2023
Copyright © - - -

Platymetopus figuratus figuratus
Female adult
RSA. (cAA). det. S. Facchini 2024
Copyright © Alexander Anichtchenko

Pseudodiachipteryx expansipennis Burgeon, 1936
Female adult
Mozambique, 18-19.XII.2005, Sofala Province, Gorongosa Res.; det. P. Schule
Copyright © Alexander Anichtchenko

Pseudoselenophorus imitator Peringuey, 1896
Female adult
Mozambique, Maputo province, Ponta do Ouro to Salamanga, 23.XI.2004. det. P. Schule
Copyright © Alexander Anichtchenko

Selenophorus (Selenophorus) gagatinus Dejean, 1829
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Selenophorus (Selenophorus) hylacis (Say, 1823)
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Selenophorus (Selenophorus) palliatus (Fabricius, 1798)
Male adult
Florida, Walton County
Photo I. Sokolov
Copyright © Igor Sokolov

Selenophorus (Celiamorphus) spinosus Shpeley, Hunting & Ball, 2017 Holotype
Male adult
BRAZIL: Amazonas/ Benjamin Constant/ Rio Javary/ II-15-III-15-1942; August Robaus/ Collector (AMNH). Image from Shpeley, Hunting & Ball, 2017
Copyright © - - -

Selenophorus (Celiamorphus) yucatanus Putzeys, 1878
Male adult
Antilles. Image from Shpeley, Hunting & Ball, 2017
Copyright © - - -

Selenophorus (Selenophorus) galapagoensis G.R.Waterhouse, 1845
Male adult
Ecuador, Galapagos, San Cristobal Nord, Las Goteras, 490m P. Moret leg. & det. (DUBC)
Copyright © Alexander Anichtchenko

Siopelophonus congestus Kataev, 2015 Paratype
Male adult
Nepal. Scale 1 mm.
Copyright © - - -

Siopelus (Pseudosiopelus) pulchellus (Dejean, 1829)
Female adult
Sénégal, M’Bour, piège lumineux, 24/09/2009, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Siopelus (Aulacoryssus) aciculatus (Dejean, 1829)
Female adult
Sénégal, M’Bour, piège lumineux, 18/10/2010, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Siopelus (Neosiopelus) punctatellus (Reiche, 1847)
Male adult
7,2 mm. Namibia, Outjo, 25 km south Okaukuejo, Ongave Lodge, 1000 m, leg. U.Schmidt, 21.III.1994; det. M.Baehr, 2012 Photo: U.Schmidt, 2014
Copyright © Udo Schmidt

Siopelus (Neosiopelus) melancholicus (Boheman, 1848)
Male adult
6,8 mm; Kenya, Tsavo National Park, Voi Lodge, 500-600 m, leg. U.Schmidt, 25.XI.1991; det. M.Baehr, 2014; Photo: U.Schmidt, 2016
Copyright © Udo Schmidt

Siopelus (Africobatus) harpaloides (Guerin-Meneville, 1847)
Male adult
RSA, Transvaal, Louis Trichardt, 17-19.I.1999. S. Facchini det. 2003 ; 7mm
Copyright © Alexander Anichtchenko

Siopelus (Laparhetes) gracilis (Harold, 1879)
Female adult
ZAMBIA, 19./30.XI.14 NW Province\ Chimfunsi Wildlifereserve\ P. Schüle leg.
Copyright © Peter Schüle

Siopelus (Siopelus) usambarensis Facchini, 2021 Holotype
Male adult
Coll. I.R.Sc.N.B., Tanzania, Tanga Region, Amani Nature Reserve, 06.X.2013, Agroforestry area, 05°06.239'S, 38°37.509'E, 994 m, Leg. L. Geeraert (RBINS).
Copyright © Sergio Facchini

Siopelus (Siopelus) microcephalus Facchini, 2021 Holotype
Male adult
Guinea, Seredou, lux, 4.5.1975, leg. Zott (MFNB).
Copyright © Sergio Facchini

Siopelus (Orinophonus) malawicus Facchini, 2021 Holotype
Male adult
Coll. Mus. Tervuren, Malawi South. Reg., Mount Mulanje, Lichenya Plateau, 2000 m, XI.1981, R. Jocqué (MRAC).
Copyright © Sergio Facchini

Siopelus (Pseudosiopelus) piceus Facchini, 2017
Female adult
RSA, 28°56'48S, 29°12'23E, Maloti-Drakensberg NP, path to Cathedral peak, 1450-2140m, 1.12.2023, L. Minkina leg. (cAA). 6 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Stenomorphus californicus rufipes Leconte, 1858
Male adult
Arkansas, Logan County. Photo I. Sokolov
Copyright © Igor Sokolov

Trichopselaphus subiridescens Chaudoir, 1843
Female adult
Paraguay, Dep. Alto Parana, Limoy, 1-10.XI.2006, leg. U. Drechsel. det. A. Anichtchenko, 2017. 12 mm
Copyright © Alexander Anichtchenko

Trichopselaphus calima Arenas-Clavijo, 2024 Paratype
Male adult
Colombia, Valle del Cauca, Trujillo, Andin´apolis, finca Gratinianos, 2388 msnm 4°10’18,01” N, 76°25’25,57”O, 04-17 diciembre 2021, en bosque, trampa de caıda, Arnulfo Gomez. Image from Arenas-Clavijo, 2024
Copyright © - - -

Trichotichnus (Trichotichnus) nishioi Habu, 1961
Male adult
Far East, Lazo Distr., midd. course of Socolovka Riv. 42°58'35"N 133°59'29E 28.VII.2005 K.Makarov leg.
L=8.5 mm
Copyright © Kirill Makarov

Trichotichnus (Trichotichnus) longitarsis A. Morawitz, 1863: 65
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Trichotichnus (Trichotichnus) laevicollis (Duftschmid, 1812)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Trichotichnus (Bottchrus) newtoni Kataev, 2016 Paratype
Male adult
Mindanao. Scale 1mm. Image Aleksey V. Kovalev.
Copyright © - - -

Trichotichnus (Amaroschesis) smetanai Kataev & N. Ito, 1999 Paratype
Male adult
China (W. Sichuan), Daxue Shan, Hailuogou Glacier park (Gongga Shan), Camp 1, 2100 m, 29.36N / 102.04E, 27. /28./31.V.97, Wrase (MNHN, http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec7443)
Copyright © - - -

Trichotichnus (Bottchrus) ivani Kataev & Schmidt, 2018 Holotype
Male adult
E-NEPAL Taplejung Distr. / W above Gurja 21-2500 m / 23-24.V.2016 leg. Schmidt / 27°19’N 87°33’E (cJS).
Copyright © Joachim Schmidt

Trichotichnus (Iridessus) tonklii Kirschenhofer, 1992b: 39
Male adult
Image from Kataev, 2020
Copyright © - - -

Trichotichnus (Parairidessus) perforatus Kataev, 2020 Holotype
Male adult
INDIA occ., 7–11.x.2005 / Maharashtra state / MULSHI env. F. Kantner leg / 40 km W of Pune” (SMNS). Image from Kataev, 2020
Copyright © - - -

Trichotichnus (Parairidessus) saluki Kataev, 2020 Holotype
Male adult
INDIA: W Karnataka, / W Ghats Mts, Jog Falls, / 530 m, at light, / 14°13.240ʹN 74°48.471ʹE, / 2–5.XI.2013 S. Saluk leg. (ZIN). Image from Kataev, 2020
Copyright © - - -

Trichotichnus (Bottchrus) cyanescens N. Ito, 1998
Male adult
Nepal, Arun Valley to Makalu , Dhabeli , 27°37'22"N 87°18'06"E ", 2340 m, 16.VI.2013, S. Tamang leg.
Copyright © Joachim Schmidt

Trichotichnus (Bottchrus) gupchiensis Kataev & Schmidt, 2017 Paratype
Male adult
NEPAL Manaslu Mts. SE slope, W Gupchi Danda 2200–2300 m, 28°08'37"N 84°44'42"E, 18.V.2006 leg. J. Schmidt
Copyright © Joachim Schmidt

Trichotichnus (Neotichnus) riedeli Kataev, 2022 Holotype
Female adult
Papua N. G. / Morobe Prov. / leg. A. Riedel, ca. 10 km S Garaina, / Saureri, 1800–2150 m, / 26.III.1998 (ZSM). Image from Kataev, 2022
Copyright © - - -

Trichotichnus (Allotichnus) beroni Kataev, 2022 Holotype
Male adult
“Indonesia, Sumatra, Mt. Kerinci / Nat. Park Kerinci — Seblat / Rhododendron forest, 3000–3100 m, / 25.V.1994, P. Beron et V. Beshkov leg.” (NMNHS). Image from Kataev, 2022
Copyright © - - -

Trichotichnus (Trichotichnus) lewisi Schauberger, 1936: 9
Male adult
Japan, Aichi-ken, Toyota-shi 2.8.2015 (cAA). 13.5 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Trichotichnus (Lampetes) semirugosus Darlington, 1968 Holotype
Female adult

Copyright © Harvard College

Typsiharpalus bonvouloiri (Vuillefroy, 1866)
Female adult
Spain, Sierra de Guadarrama, Cotos, 15-7-2009, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Typsiharpalus azruensis Antoine, 1925
Male adult
Maroc, Moyen Atlas, Ifrane, Juin 2005
Copyright © Pascal Deschamps