Carabidae

Carabidae image gallery

Region

Carabus (Diocarabus) loschnikovi loschnikovi Fischer, 1823
Male adult
Komi Resp., Pecero-Ilych Nat. Res., Zajachij Kamen h~786 62°10'06" N 59°12'53" E 8.VIII.2004 M. Chernjakhovski leg.
L=15.0 mm
Copyright © Evgeniy Zinoviev

Aplothorax burchelli Waterhouse, 1841
Male adult
St. Helena island
Copyright © Boleslav Brezina

Calosoma (Calosoma) inquisitor cupreum Dejean, 1826: 196
Female adult
Armenia, Zangezur range, Meghri suburbs., Vahravar vill., Rivulet vall., h=1400 m Yuriy Shevnin leg.; 23.05.2007; 21 mm.
Copyright © Anton Menstschikov

Calosoma (Calosoma) schayeri schayeri Erichson, 1842
Male adult
Australia N.T. Alice Springs, Airport
19.5.2000 M. Langer leg.
male, 21 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisphaena) reticulatum reticulatum s.str.
Female adult
Germania, or. DDR, Bezirk Frankfurt, 24.V.1989
female, 25 mm
photo & det. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Callistenia) moniliatus Le Conte, 1851: 200
Male adult
SE Canada, Nueva Escocia
19.5.1970
male, 17 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) anthrax Semenov, 1900 Holotype
Male adult
collection of ZIN
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) elegans s.str. (as declivis Dohrn, 1884b)
Male adult
SE Kazakhstan Alma-Ata reg., 30 km N Sary-Ozek 28.4.1998 Klimenko leg. male, 22 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) elegans amethystinus Semenov & Redikorzev, 1928
Male adult
SE Kazakhstan, E Zailiyski mt. rng. Chilik riv.
Malybai vill. 28.4.1999 Klimenko A. leg.
male, 25 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) elegans amethystinus Semenov & Redikorzev, 1928
Male adult
26 mm.
Zailiyski Alatay.
06.1997 Chibilov E. leg.
Copyright © Sunvas

Calosoma (Callisthenes) elegans elegans s.str.
Male adult
SE Kazakhstan, Tchu-Iliyskiy Mts., Kordai pass, h=1500; A. Klimenko leg.; 20.04.1997;
24 mm.
photo A. Menstschicov
Copyright © Anton Menstschikov

Calosoma (Callisthenes) elegans manderstjernae Ballion, 1870
Female adult
Kirgizia, S Kirgizskii mt. rng., 2200m
Kichi-Kimin riv., Ak-Tuz vill. 05.1998 Klimenko leg.
female, 32 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) elegans semenovi Motschulsky, 1859
Female adult
SE Kazakhstan, Zailijsky Mt. rng.,
Talgar env., 1500 m, 21.5.2000
female, 27 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) eversmanni Chaudoir, 1850
Male adult
Syria, Antilibanon
1.3.1991
male, 22 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) fischeri Fischer, 1842
Male adult
Mongolia, Mandalgob, 20 km
N Amardalai vill. 11-12.6.2003 Anichtchenko A. leg
male, 19 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) grumi Semenov, 1900 Holotype
Female adult
collection ZIN
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) breuningi Mandl, 1954
Female adult
E Kazakhstan, S Alakol lake, Koktuma vill., 1-10.5.1999 Nekroshevich S. leg. female, 22 mm; det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) breuningi iliense Kabak, 2022
Male adult
SE Kazakhstan, near Sary Ozek vill., A. Klimenko leg; 22.04.1997; 22 mm.; det. Anichtchenko A. 2022
Copyright © Anton Menstschikov

Calosoma (Callisthenes) kuschakewitschi kuschakewitschi s.str.
Male adult
Karzhantau mt. rng., Susungen riv.,
2300 m, 7.6.1998 Tretjakov leg.
male, 23 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) mirificus Casale, 1979
Female adult
Afganistan, S slope of Paropamiz Mt. rng., near Sabzan pass, 2400m, 22.VI.1967 M.Stoljarov leg.
Collection of ZIN
det. D. Obydov 1999
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) panderi Fischer, 1822
Male adult
NW Kazakhstan, Aktiubinsk reg., Baiganinsk distr.,
near Zharly vill. 10-12.05.1997 leg. M.& L. Smirnov
male, 20 mm
Photo from www.zin.ru
Copyright © Maksim Smirnov

Calosoma (Callisthenes) regelianus A. Morawitz, 1886
Female adult
Tadzikistan, Darvaz Ezgan 2.5.1971
female, 25 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Callitropa) externum Say, 1823: 149
Female adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Calosoma (Calosoma) maximowiczi A. Morawitz, 1863: 20
Female adult
China, Mt. Wangfoshan (1940 m.), Baokang Countri, Hubei; 20-28.06.2008, 30 mm.
Copyright © Anton Menstschikov

Calosoma (Calosoma) oceanicum klynstrai Breuning, 1927
Male adult
INDONESIA, Tanimbar Island, III.2009
photo & coll. N. Belonogov
Copyright © Nil Belonogov

Calosoma (Carabophanus) arrowi Breuning, 1928
Male adult
Ethiopia, Oromia reg., Arsi highlands, Wahoro env., Tiliku Lensho river valley10.-12.II.2010, J.Štastny lgt
photo Jaroslav Stastny
Copyright © Jaroslav Štastny

Calosoma (Castrida) alternans granulatum Perty, 1830
Female adult
Argentina, Entre Rios,
Liebig, nov.1997
long. 32 mm
photo, det. & coll. Anichtchenko A. 2009
Copyright © Alexander Anichtchenko

Calosoma (Castrida) vagans Dejean,
Male adult
Chile, Santiago de Chile, Quilicura
19.9.1991 G. Aceituno leg.
male, 21 mm
Copyright © Alexander Anichtchenko

Calosoma (Ctenosta) bastardi Alluaud, 1925
Female adult

Copyright © Alexander Anichtchenko

Calosoma (Ctenosta) senegalense Dejean, 1831
Female adult
length: 22-27 mm
Kenia Tsavo National Parc
leg. U.Schmidt, 1991; det. W.Lorenz, 2000
Foto: U.Schmidt, 2006
Copyright © U.Schmidt

Calosoma (Carabops) abyssinicum Gestro, 1880
Male adult
Ethiopia, Bale prov., Str. ZW. Sasheme u. Dodola, Mt. Mascall 3100-3300 m
21.VII.2005 Puchner leg.
male, 18 mm
det. Puchner F. 2005
photo & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Ceroglossus buqueti deuvei Jiroux, 1996
Male adult
Chile, VIII Reg., 1000-1250 m, P.N. Nahuelbuta, (Piedra del Aguila) 7-12.XII.1999 Heinz leg.
male, 22 mm
photo, det. & coll. Anichtchenko A. 2009
Copyright © Alexander Anichtchenko

Ceroglossus buqueti andestus Kraatz-Koschlau, 1887
Male adult
CHILE, Liquine; 05/12.02.2005; C.Saavedra leg.; Determination confirmed by Anichtchenko, 2019
Copyright © Anton Menstschikov

Ceroglossus buqueti cherquencoensis Dorsselaer, 1955
Male adult
Chile, IX reg., Cherquenco Vn. Llaima, 800-1000m, 29-30.I.1999 Heinz leg. 21 mm; det. & coll. Anichtchenko A. 2019
Copyright © Alexander Anichtchenko

Ceroglossus buqueti sybarita Gerstaecker, 1858
Female adult
Chile, Chiloe, 160m, 8 km N Butalcura 22-24.XII.1998 det. Heinz
female, 24 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Ceroglossus chilensis cyanicollis Kraatz, 1887
Male adult
Chile (VIII reg), Umg. Lag. de la Laja, 950 m, 19-20.12.1998 Heinz leg. 26 mm; det. Anichtchenko A., 2019
Copyright © Alexander Anichtchenko

Ceroglossus chilensis fallaciosus Kraatz, 1880
Male adult
CHILE, Talca prov., Constitucion tn. env., II.2008. det. Anichtchenko A., 2019
Copyright © Nil Belonogov

Ceroglossus chilensis ficheti Jiroux, 1996
Male adult
Chile (VIII reg), P.N. Nahuelbuta, (Piedra del Aguila) 700 m, 7-12.12.1999, Heinz leg.; 28 mm. Determination confirmed by Anichtchenko, 2019
Copyright © Alexander Anichtchenko

Ceroglossus chilensis gloriosus Gerstaecker, 1858
Male adult
Chile (IX reg.), Malalcahuello, 800 m, 9-10.12.1999 Heinz leg. 25 mm; Determination confirmed by Anichtchenko, 2019
Copyright © Alexander Anichtchenko

Ceroglossus chilensis latemarginatus Kraatz-Koschlau, 1889
Male adult
Chile (VIII. Reg.) Pata de Gallina, 650m, b.Contulmo 7-12.12.1999 Heinz leg. 27 mm; Determination confirmed by Anichtchenko, 2019
Copyright © Alexander Anichtchenko

Ceroglossus chilensis mochae Reed, 1874
Female adult
CHILE, Arauco prov., Isla Mocha. - II.2008. photo Belonogov N.; Determination confirmed by Anichtchenko, 2019
Copyright © Nil Belonogov

Ceroglossus chilensis seladonicus Kraatz-Koschlau, 1887
Female adult
Chile, Malleco prov., Madalcahuello
enero 2003 G. Aceituno leg.
28 mm
Copyright © Alexander Anichtchenko

Ceroglossus chilensis solieri Roeschke, 1900
Female adult
Chile, Chiloe, 160m, 8 km N Butalcura 22-24.XII.1998 det. Heinz
female, 28 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Ceroglossus darwini darwini s.str.
Male adult
Chile, Chiloe, ca. 25 km NW Queilen, 350 m, (n. Castro), 26-27.XII.1999 Leg. Heinz
male, 22 mm
photo, det. & coll. Anichtchenko A. 2009
Copyright © Alexander Anichtchenko

Ceroglossus resplendens Jaffrezic & Rataj, 2006
Female adult
CHILE. X Region, Termas de Carranco; 01/05 February. 2007; C.Saavedra leg.; det. Anichtchenko A. 2019
Copyright © Anton Menstschikov

Ceroglossus speciosus Gerstaecker, 1858
Female adult
Chile, Chiloe, S. Pedro de Piruquiña 350 m, (n. Castro), 24-25.XII.1999 Heinz leg.
female, 23 mm
photo, det. & coll. Anichtchenko A. 2009
Copyright © Alexander Anichtchenko

Calosoma (Teratexis) fabulosum Semenov & Znojko, 1933 Paratype
Male adult
collection of ZIN
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko

Pamborus alternans Latreille, 1817
Female adult
Australia, NSW. Kerewong S.F.
21.12.1978
30 mm
Copyright © Alexander Anichtchenko

Pamborus brisbanensis Castelnau, 1867
Male adult
Museum of Zurich
photo Anichtchenko
Copyright © Alexander Anichtchenko

Pamborus guerini Gory, 1830
Male adult
Museum of Zurich
photo Anichtchenko
Copyright © Alexander Anichtchenko

Pamborus opacus Gehin, 1885
Male adult
Museum of Zurich
photo Anichtchenko
Copyright © Alexander Anichtchenko

Pamborus pradieri darlingtoni Banninger, 1940
Female adult
Australia, N.S.W. New England rng.,
Moonan Flats, 18.12.2000 T.Pollak leg.
21 mm
Copyright © Alexander Anichtchenko

Pamborus pradieri pradieri Chaudoir, 1869
Male adult
Museum of Zurich
photo Anichtchenko
Copyright © Alexander Anichtchenko

Pamborus punctatus Darlington, 1961 Paratype Paratype
Male adult
Museum of Zurich
photo Anichtchenko
Copyright © Alexander Anichtchenko

Pamborus tropicus Darlington, 1961 Paratype Paratype
Male adult
Museum of Zurich
photo Anichtchenko
Copyright © Alexander Anichtchenko

Carabus (Acathaicus) alexandrae alexandrae Semenov, 1887 Holotype Holotype
Male adult
Kwantin and Li-tschara-pu, 22.VI.1885, leg. Potanin Deposited in Zoolog. Museum St. Petersburg. male, long. 22 mm photo B. Brezina
Copyright © Boleslav Brezina

Carabus (Acathaicus) alexandrae cratocephaloides Semenov, 1887
Female adult
female, 28mm
Copyright © Boleslav Brezina

Carabus (Acathaicus) interruptocostatus Semenov, 1887 Holotype Holotype
Female adult

Copyright © Boleslav Brezina

Carabus (Acoptolabrus) lopatini Morawitz, 1886
Male+Female adult
Russia,
Sakhalin isl., Tchekhov Mt, VI.1997
female, 40mm
photo & coll. Panin R.
Copyright © Ruslan Panin

Carabus (Acoptolabrus) schrencki schrencki (as hauryi Gehin, 1885 )
Male adult

Copyright © Boleslav Brezina

Carabus (Acoptolabrus) schrencki kimieae Kezuka & Imura, 1989
Male+Female adult
Mt. Wulongshan, Dandong City,
Liaoning Province, CHINE
25-30.VI.2008
Copyright © Safronov

Carabus (Acrocarabus) callisthenoides Semenov, 1888
Male adult
male, 22mm
Copyright © Boleslav Brezina

Carabus (Acrocarabus) guerini aldarkose Kabak, 1992
Male adult
male, 22mm
Copyright © Boleslav Brezina

Carabus (Acrocarabus) guerini guerini Fischer, 1842
Male adult
SE Kazakhstan, W Dzhungar Mts, Lepsinsk env., 1600 m., 18.06.2000, A.Klimenko
Copyright © Anton Menstschikov

Carabus (Alipaster) barovskii bayanbulak Deuve, 1993 Paratype Paratype
Female adult
female, 17mm
Copyright © Boleslav Brezina

Carabus (Alipaster) infantulus infantulus Morawitz, 1886
Male adult
male, 15mm
Copyright © Boleslav Brezina

Carabus (Apoplesius) deuvei hajdaji Hackel, 2007 Holotype Holotype
Male adult
Turkey- prov. Mus 60 km NW Mus 1630m
3856N 4108E Bulgan Gec. 23-24.06.03 E. & P. Hajdaj leg.
Copyright © Boleslav Brezina

Carabus (Apotomopterus) anxiensis Kleinfeld, 1998 Holotype & Paratype Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) bousquetellus gengmaensis Kleinfeld, 2003 Holotype & Paratype Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) cantonensis pervarius Kleinfeld, 1997 Holotype
Male adult
Frank Kleinfeld
Copyright © - - -

Carabus (Apotomopterus) casaleianus jigongshanicus Kleinfeld, 1998 Holotype & Paratype Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) datianshanicus Kleinfeld, 1997 Holotype & Paratype Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) cantonensis metallaicus Kleinfeld, 1999 Holotype Holotype
Female adult
as pervarius metallaicus
Copyright © Frank Kleinfeld

Carabus (Apotomopterus) davidis neodavidis Kleinfeld & Schutze, 2001 Holotype & Paratype Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) davidis viridithoracis Kleinfeld & Schutze, 2001 Holotype & Paratype Holotype Paratype
Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) davidis yizhangensis Kleinfeld, 2002 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) delavayi s.str. (as medea Kleinfeld, 1999) Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Apotomopterus) delavayi tenuimanus Deuve & Imura, 1990
Female adult
Lijiang, Yunnan, Chine
female
photo Daniel Prunier
Copyright © Daniel Prunier

Carabus (Morphocarabus) illigeri illigeri (as bjelasnicensis Apfelbeck, 1902)
Male+Female adult
Serbia, Debelo Brdo, 5.VI.2007, S. Sapaly leg. L=32 mm photo Panin R.
Copyright © Ruslan Panin

Carabus (Archicarabus) monticola fontanae Born, 1906
Male+Female adult
N-Italy,
Piemonte, Bielmonte, h1350 m, 14-15.VI.2010, Pascal Stefani leg.
female, 23 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) estreicheri Fischer, 1822
Male+Female adult
W UKRAINE, NW Podolian,
Lviv Region, Brody distr., near Yasenov vill., steppe sites, 2-28.V.2010, Panin R. leg.
female, 23mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei Kuster, 1846
Male+Female adult
W.Ukraine.
Transcarpathian region,Vinogradiv distr., near Malaya Kopanya vill., Chorna Gora Mt., h=180m. 7-18.VII.2008. Panin R. leg.
female 33 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei Kuster, 1846
Male adult
W.Ukraine.
Transcarpathian region,Vinogradov distr., near Malaya Kopanya vill., Chorna Gora Mt., h=180m. 7-18.VII.08. Panin R. leg.
male, 28 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860
Male+Female adult
SW -France,
Pyrenees-Atlantiques, Foret de Sare, XII.2009, Pascal Stefani leg.
female, 31 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as delacrei Tarrier, 1975)
Male+Female adult
NE Spain, Catalonia, Sant Jaume de Frontanyа, h=1000 m, V-VI.2010, Pascal Stefani leg.
female, 38 mm
Copyright © Ruslan Panin

Carabus (Carabus) arvensis sylvaticus Dejean, 1826
Female adult
France, Centre reg., Eure-et-Loir, Forкt de Montecot, VI.2009, Pascal Stefani leg.
female, 20 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) arcensis sylvaticus Dejean, 1826 (as sugintensis Brunier, 1922)
Male adult
NW -France, Haut-Rhin Depart., Fulleren, XI.2004, Pascal Stefani leg.
Copyright © Ruslan Panin

Carabus (Chrysocarabus) olympiae Sella, 1855
Male+Female adult
N-Italy,
Piemonte, Bielmonte, 14-15.VI.1981, Pascal Stefani leg.
female, 34 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Coptolabrodes) haeckeli Brezina & Imura, 1997
Male adult
C China, Shaanxi,
Tsinling Mts., Taibaishan Mt., h=3000m, 33:55N/107:44 E, VI.2004, V.Siniaev & his team
female, 31 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus tscherkessicus Korge, 1964
Male adult
S Russia, NW-Caucasus,
Karachay-Cherkessia,Teberda, Mukhinsky (Mukhu) Mt. pass, h=2750-2800 m , 8.VIII.2010, I.A. Solodovnikov leg.
male, 23 mm

Copyright © Ruslan Panin

Carabus (Eupachys) glyptopterus Fischer, 1827
Male+Female adult
Russia, SE-Siberia,
Buryat Republic, Ivolginsky Distr., 04.VII.2010, Filippov A.E. leg.
female, 32 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Coptolabrus) formosus brockmanni (as nili Deuve, 2005 PT) Paratype
Male+Female adult
C China, Shaanxi,
Liupan Mts., Guan Shan Mt. h=800 m, , 34:50N/106:48 E, Team V.Siniaev leg.
female, 38 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792
Female adult
NE-FRANCE,
Lorraine Reg., Moselle, 15.II.2010, Pascal Stefani leg.
female, 26 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792
Male adult
Austria,
Weissenbach stt., III.1979, Kretschmer leg.
male, 24mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Isiocarabus) fiduciarius fiduciarius (as fiduciarius tim) Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Isiocarabus) hunanicola liangshanicus (as dreuxi liangshanicus) HT PT Kleinfeld, 1998 Holotype Paratype
Male+Female adult
Carabus (Isiocarabus) hunanicola liangshanicus (as dreuxi liangshanicus) HT PT
Copyright © Frank Kleinfeld

Carabus (Tribax) titan titan Zolotarev, 1913
Male+Female adult
Russia,NW Caucasus,
Krasnodar reg.,Chernogorie Mnt. Range, Agulova Balka h=950m V-VII.1999, A.S.Zamotajlov leg.
female,32mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Tribax) steveni steveni Menetries, 1832
Male adult
Carabus (Tribax) steveni steveni Menetries, 1832
Karachaevo-Cherkessia,
Hudes riv., near Tsarskie vorota,
V.2009 Anichtchenko A. leg.
male, 25mm
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Tribax) puschkini puschkini Adams, 1817
Male adult
Armenia, Shirak Province, Ashotsk, 18.VI.2001, P.V.Kazaryan leg. male 25mm
Copyright © Ruslan Panin

Carabus (Tribax) circassicus teberdensis Zolotarev, 1913
Male+Female adult
W-Caucasus: Karacaj-Cherkessija,
Abishira–Akhuba Mnt. Range h=2000-2650m, Kyafar-Agur Riv., 14–17.VII.2009, Fominykh D. leg.
female, 25mm
Copyright © Ruslan Panin

Carabus (Tribax) circassicus circassicus Ganglbauer, 1886
Male+Female adult
Abkhasia,
W.slope of Gagrinsky Mt., Krepostnaya River source, h=930-980m., A.S.Zamotajlov leg. 27.VII.2000
female, 29 mm/male, 26mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Tribax) circassicus abagonensis Starck, 1894
Male+Female adult
S Russia, NW Caucasus,
Adygea Rep., Mt.Abago, 26.VI.1999
L=24mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Tribax) biebersteini ulgeni Gottwald, 1982
Male adult
S Russia, Karacaj-Cherkessija,
near Kart-Djurt vill., upper Kylan riv. h=1300m, 26-27.V.2009, Fominykh D. leg.
male, 28mm
Copyright © Ruslan Panin

Carabus (Tribax) apschuanus apschuanus Rost, 1893
Male+Female adult
Carabus (Tribax) apschuanus apschuanus Rost, 1893
Russia, Caucasus, Krasnodar reg.,
h=1200 m, 29.VII.2000
A. Zamotajlov leg.
Copyright © Safronov

Carabus (Tribax) agnatus agnatus Ganglbauer, 1889
Male adult
NW-Caucasus,
Krasnodar reg., Mostovskoj distr., Skirda Mts. N. slope Yatyrgvart Mt., h=2300-2500m. 09-10.VII.2008, D.Fominyh leg.
male, 23mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus sibiricus Fischer, 1822
Male adult

Copyright © Kirill Makarov

Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886
Male+Female adult
W Ukraine, Ternopol reg., Pidvolochisk distr., Supranivka vill. 3.VII.1908. Leg. Stobetskii. S. Photo R. Panin
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886
Male adult
W Ukraine,
Ternopil reg., Pidvolohcisk distr., Supranivka vill.
male, 26mm
The end of XIX the beginning of XX century
photo Panin R.
coll. SMNH Ukraine, Lviv
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus haeres (as fossulatus Dejean, 1826)
Male+Female adult
E.UKRAINE, Kharkov reg., Zmiev distr.,in steppe,10-20.V I.2007, D.Kovalchuk leg. female, 26mm/male, 24mm. photo/det. Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus errans Fischer, 1823
Male+Female adult
female:SE Ukraine,
Donetsk, 25.VII.2009, A.I.Gubin leg. L=30mm.
male:E Ukraine, Lugansk reg., near Slavyanoserbk, 5-15.VIII.2009, S.Konovalov leg. L=26mm.
photo/det.Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus bosphoranus Fischer, 1823
Male adult
Carabus (Trachycarabus) sibiricus bosphoranus Fisch. 1823 S Ukraine, Krym Peninsula, Dolgorukovskaya jaila, 25.VIII.1991 I.A.Solodovnikov leg. male, 23mm 
Copyright © Ruslan Panin

Carabus (Trachycarabus) scabriusculus scabriusculus (as erythropus Fischer, 1822)
Male+Female adult
W.Ukraine, NW Podolian Upland , Lviv Reg., Zolochiv distr., near Luky vill., steppe sites, 7.V-8.VI. 2010 leg. Panin R. female, 22mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus planus Gehin, 1885
Male adult
S Russia, Krasnodar reg., Anapa distr., B. Utrish vill., env., 11-20.VIII.2008, Khatchikov E. leg. male, 26mm photo Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus planus Gehin, 1885
Endophallus
endophalus S Russia,Krasnodar reg., Anapa distr., B. Utrish vill., env.,
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus planus Gehin, 1885
Endophallus
endophalus Karacaj-Cherkessija, near Kart-Djurt vill., upper Kylan riv. h=1300m
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus campicola Reitter, 1896
Male adult
Krym Peninsula, near Sudak. Leg.Vashchenko S. 1-5.V.2004 27 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus campicola Reitter, 1896
Male+Female adult
S Ukraine, Krym Peninsula, near Perevalnoe vill., in steppe, 15.IV.2009, Shaporinskiy V. leg. female, 28 mm photo/det.Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) besseri Fischer, 1822
Male+Female adult
W Ukraine.NW Podolian,
Lviv reg., Zolochiv distr., Lysa Gora near Stinka vill., steppe sites, 25.IV-9.V.2009 Panin R. leg
female, 30mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) scabripennis scabripennis Chaudoir, 1850
Male adult
Carabus (Tomocarabus) scabripennis scabripennis Chaudoir, 1850
C. Armenia,
Kotayk reg., Yeghvard, Arailer Mt., 16.VI.1999, P.V.Kazaryan leg.
male, 20mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Tomocarabus) marginalis Fabricius, 1794
Male+Female adult
E Ukraine,
Kharkov reg.,near Vasishchevo vill.10-20.VI.2006 D.Kovalchuk leg.
female 24mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) decolor Fischer, 1823
Male adult
S Russia, NW-Caucasus,
Karacaj-Cherkessija,Skalisty Mnt. Range, Site Banditskoe h=1400m, 2-3.V.2009,
Fominykh D., Bondarenko A.S. leg.
male,17mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) wulffiusi wulffiusi Morawitz, 1862
Male+Female adult
Russia, Far East,
Primorskiy Kray, near Kamenushka vill., 2-3.VIII.1992, K.E.Dovgaylo leg.
female, 24mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Male+Female adult
NE-Serbia, Krs Mts., Majdanpek env.,h=300m.20-31.V.2006, Pr.Sima leg. female, 32mm/male, 30mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) venustus venustus Morawitz, 1862
Male adult
SE Primorskii reg., Lazovskii Nat.Res., 12 km S Staraya Kamenka vill., near Amerika kor. 43°15N, 134°1E, 19.VII.04 leg. M.& L. Smirnovy photo from www.zin.ru
Copyright © Maksim Smirnov

Carabus (Morphocarabus) spasskianus spasskianus Breuning, 1932
Male+Female adult
SE Siberia, Russia
Irkutsk reg., near Pivovarikha vill., 7.V.07, D. Fominyh leg.
male, 24mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus saragaschensis Obydov, 1999
Male adult
W Sayan Mts., Batenevskii Mt.rng., 12 km E Pervomaiskoe vill.,
Enisey riv., mountain steppe, 400m, 5.VII.2000 A. & N. Menschikovy leg.
female, 26mm

Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
Male adult
Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799
S.Bohemia,
Hluboka nad Vltavou, 24.VI-18.VII.1990, S. Budka leg.
male, 27mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens (as lomnitzkii Reitter, 1896 )
Male adult
W-UKRAINE, NW Podolian, Lviv Reg., Zolochiv Distr., W Ukraine, Lysa Gora steppe massif, 19.V.-7.VI.2007, Panin R. leg. 25 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens Fabricius, 1798 (ab.erythromerus Dejean, 1826)
Male adult
Moldova, Anenii Noi distr., near Garbovets vill. 20.VI.08 Bodnar' I.I.leg. male, 26mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens (as lomnitzkii Reitter, 1896)
Male adult
W Ukraine, NW Podolian, Ternopil reg., Pidvilochisk distr., near Rosokhuvatets & Kamyanki vill., steppe sites, 24.V-15.VI.2008, Panin R. leg.
male, 24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) helleri Ganglbauer, 1893
Male+Female adult
SE Moravia, Strani, V.2002, M.Snizek leg. male bronze colour-26mm male blue colour-28mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi incompsus (as mendax Csiki,1906)
Male+Female adult
W.Ukraine,
Transcarpathian Reg., Rakhov Distr., Chornogora Mt Range, h=1900m. alpine zone, Panin R. leg/det. 28.VI-9.VIII.2008
female, 32 mm
photo endophallus Solodovnikov I.A
Copyright © Ruslan Panin

Carabus (Eucarabus) obsoletus carpathicus Palliardi, 1825
Male+Female adult
female, 25 mm
SW-Romania, Banat, Resita - Valiug, 3-8.VI.2007, Krause R. leg.
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus tibiscinus Csiki, 1905
Male+Female adult
female,28 mm
SE Hungary, Csongrбd, Makу env., 1-8.VIII.2004, Krause R., Slovak, Pittner leg
Copyright © Ruslan Panin

Carabus (Tribax) apschuanus pseudoplatessa Gottwald, 1982
Male+Female adult
NW-Caucasus, Karachay-Cherkessia, Teberda: Dzhamagat gorge, the left bank of the riv. Dzhamagat, 10.VII-28.VIII.2010 S.Pavluchuk leg.
female, 27 mm
Copyright © Ruslan Panin

Carabus (Pachystus) hungaricus mingens Quensel, 1806
Male adult
male, 31 mm
SW-Russia: Stavropol Krai, near Stavropol city , Russkiy Les, 11.VI-28.IX.2010, Pavluchuk S. leg.
Copyright © Ruslan Panin

Carabus (Archicarabus) montivagus montivagus Palliardi, 1825
Male+Female adult
SW-Romania: Banat, Caras-Severin Reg., Sasca Montana – Stinapari, 21.V.-5.VI.2010, Krause Rene leg.
female, 26 mm
Copyright © Ruslan Panin

Carabus (Procechenochilus) kokujewi Semenov, 1898
Female adult
RUSSIA, Karachay-Cherkessia, NW Caucasus Mts., Daut Mt. Range, Iskrovtsev - W.Ullu-Kel Mt. pass circus, h=2800-2600m
Copyright © Alexander Anichtchenko

Carabus (Procechenochilus) ? kokujewi Semenov, 1898
Female adult
Russia, NW Caucasus, Karachay-Cherkessia, Karachaevsk distr., near Uchkula vill., valley Uchkulanichi riv. h=2800-2900m, 25-28.VII.2010, Fominykh D. leg.
female, 23 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) starckianus ganglbaueri Reitter, 1888
Female adult
NW Caucasus, S Russia, Adygea Rep., Confluence of the river Belaya & Kisha h~700m, 24.V-7.VIII.2010, D.Fominyh leg.
female,32 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus andrzejuscii (as wolffii Dejean, 1826)
Male+Female adult
SW-Romania: Banat, Moldova Noua, Ravenska viil., 21.V.-5.VI.2010, Krause Rene leg. female, 32 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus andrzejuscii (as mehelyi Ganglbauer, 1896 loc. typ.)
Male+Female adult
N-Romania, Rodna Mts., Pasul Setref - Sacel 10km E, h=810 m, 8-19.VI.2009, Krause Rene leg. female, 31mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus andrzejuscii Dejean, 1826 (as pygmaeus Petri, 1912)
Male adult
W Ukraine, NE Carpathians, Transcarpathian Reg., Rakhov Distr., Chornogora Mt Range h=2030m, alpine zone, 13-24.VI.2010 Panin R. leg./det. male, 25 mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) convexus s.str. (as pyrenaicus Born, 1907)
Male adult
Pirineos, Canfranc, 1300m, Barranco de Izas
15.6.2005 Anichtchenko A. leg.
male, 16 mm
Copyright © Alexander Anichtchenko

Carabus (Tomocarabus) bessarabicus bessarabicus Fischer, 1823
Male+Female adult
UKRAINE,
Kherson Reg., Golaya Pristan distr. near Rybalche vill. 6.IV.2000
long (female) 20mm
Copyright © Ruslan Panin

Carabus (Teratocarabus) azrael mizunumaianus Imura, 1991
Male adult
China, Liaoning, Benxi, Gongchangung
20.VI.2004 Hong
Copyright © Boleslav Brezina

Carabus (Tanaocarabus) hendrichsi Bolivar y Pieltain, Rotger & Coronado, 1967
Male adult
male, 23mm
Copyright © Boleslav Brezina

Carabus (Morphocarabus) odoratus bargusinicus Shilenkov, 1996
Male+Female adult

 Carabus (Morphocarabus) odoratus bargusinicus Shilenkov, 1996
 Russia, N Burajatia,
 Barguzinskiy Mts., the upper reaches of M.Chiverkuy riv., h=2000m, Vashchenko S. leg.
 female, 21mm/male,19mm
 photo Panin R.

Copyright © Ruslan Panin

Carabus (Morphocarabus) odoratus antropovi Shilenkov 1996
Male+Female adult
Russia, S Tuva, Khorumnug -Tayga mts. Range, h=1600-2300m, Buren riv. valley, 4-26.VI.2000 Vashchenko S & A. leg female, 20mm/male,18mm photo Panin R.
Copyright © Ruslan Panin

Carabus (Tachypus) auratus s.str. (as brunieri Barthe, 1921)
Male adult
SC-FRANCE, Aude, Rennes les Bains, 6.VII.1989, Sunard A. leg.
male, 26mm
Copyright © Ruslan Panin

Carabus (Tachypus) auratus lasserrei Doue, 1855
Male adult
SC-FRANCE, Midi-Pyrуnуes reg. Tarn, Saint Paul cap de Joux, 24.V.2003, A. Laforgue leg.
male, 24mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) monilis Fabricius, 1792 (as interpositus Géhin, 1880)
Male adult
NW France,
Basse-Normandie, Orne, Forкt d'Йcouves, V.2009, Pascal Stefani leg.
male, 29mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) monilis (as interpositus Gehin, 1880)
Male+Female adult
NW France,
Haute-Normandie reg., Bois Les Loges, Etretat, Seine-Maritime,10.V.1988
female, 28mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) monilis (as dupeuxi Deuve, 1974)
Male+Female adult

France,
Charente, Foret de Braconne VII.1998, Laforgue leg.
famale, 27mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) michailovi Kabak, 1992
Male adult
NE-Kazakhstan,
S Altai Mts.,Sarym-Sakty Range, Sarym-Sakty riv., h=2400m. 21.VI.1999., A.Klimenko leg.
male,18mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
Male adult
Russia, S Siberia,
S Khakassia, N Tuva reg., Sailyg-Khem-Taiga mts. range h=2400m, 10.VII.2004
male, 18mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
Male+Female adult
Russia, S Siberia,
S Khakassia, N Tuva reg., Sailyg-Khem-Taiga mts. range h=2000-2600m,10-30.VI.2005, S.Vashchenko & S.Alexeenko leg.
male,17.5mm/female,19mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Morphocarabus) mestscherjakovi pseudoodoratus Brinev, 2002 Paratype Paratype
Female adult
Krasnojarsk reg., Aradan distr., 5 km SW Aradan vill., vall. of Us riv., A.Brinev leg. female, 20 mm photo & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) kozhantschikovi martjanovi Lutshnik, 1924
Female adult
Khakassia, 1800 m, near Cheremushki vill., 12.6.-20.7.2000
female, 21 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
Male adult
W Sayan Mts., Kazhyr-Suk riv., 80 km W Oleniya
Rechka, W part of Oiskii mt. rng., 30.6.-7.7.1999
2100 m, Brinev A. leg.
male, 19 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) hummeli nevelskii Shilenkov, 1996
Male+Female adult
Russia, Sakhalin isl.,
Makarov distr., Lesnaja riv., VII.1992, M. Nesterov leg.
female,23mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) varians varians Fischer, 1827
Male+Female adult
female: Transcaucasia, Armenia, Amasia distr., Karakhachskiy mountain pass h=2270m, 3.VIII.1985 P.V.Kazaryan leg., 22mm male: Armenia, Amasia distr., Arpalach vill. VI.1999, P.V. Kazaryan leg.
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) blumenthaliensis basilewskianus Breuning & Ruspoli, 1970
Male+Female adult
NE-Turkey,
Artvin Prov., Hopa vers Borcka, Cankutaran Gecidi h=800m, 17.VI.2007 leg. D. Prunier
female, 24mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) varians janthinus s.str.
Male+Female adult
NW-Caucasus, Russia, Krasnodar reg.,Tuapse distr., Mt.Semashkho 04.V.2008, D.Fominyh leg. female, 24mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) varians armeniacus Mannerheim, 1830
Female adult
NE Turkey, Artvin Prov., Borçka, Macahel Gecidi h=1450m, 18.VI.2007, D.Prunier leg. female, 26mm
Copyright © Ruslan Panin

Carabus (Semnocarabus) regulus regulus Dohrn, 1882
Male adult
photo M. Smirnov from www.zin.ru
Copyright © Maksim Smirnov

Carabus (Rhigocarabus) xiei sanggarparicus Deuve, 1996 Holotype Paratype
Male adult
Holotype
male, 19mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) sehnali Brezina & Hackel, 2006 Holotype Paratype
Male adult
male, 20mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) sehnali Brezina & Hackel, 2006
Female adult
female, 23mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) gentleman Brezina & Hackel, 2004 Holotype
Male adult
male, 19mm
Copyright © Boleslav Brezina

Carabus (Pseudotribax) ferghanicus Breuning, 1933
Male adult
male, 29mm
Copyright © Boleslav Brezina

Carabus (Pseudocranion) sackeni sackeni Semenov, 1898
Male adult
male, 23mm
Copyright © Boleslav Brezina

Carabus (Pseudocranion) sackeni remondianus Deuve, 1996 Paratype
Female adult
Museum of Institute of Systematic Biology (Daugavpils, Latvia)
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) aeruginosus zyrjanovskianus Shilenkov & O.Berlov, 1996
Male+Female adult
NE KAZAKHSTAN,
S-Altays, Ulbinsky Mt. h=900m., Bobrovka vill., 24.VI.01, A. Klimenko leg.
female,25 mm
Copyright © Ruslan Panin

Carabus (Pseudocoptolabrus) burmanensis Breuning, 1932
Male adult
male, 23mm
Copyright © Boleslav Brezina

Carabus (Morphocarabus) aeruginosus verzhutzkii O.Berlov & Shilenkov, 1996
Male adult
C Altai Mts., Seminskii Mt.rng., Katun riv., 3 km S Ust'-Sema riv.,
4.VII.2002 A. & N. Menschikovy leg.
det. Mentschikov A.
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
Male adult
SE Siberia, E Sayan, Nizhneudinsk, h=500m, 21-26.VI.2006,
male, 26mm
Copyright © Ruslan Panin

Carabus (Mimocarabus) maurus maurus Adams, 1817
Male+Female adult
Armenia,
Ashotsk distr., Tsoghamarg vill., 24-27.V.1999 P.V.Kazaryan leg.
female, 20mm/male,18mm
Copyright © Ruslan Panin

Carabus (Procrustes) coriaceus hopffgarteni Kraatz, 1878
Male adult

Copyright © Safronov

Carabus (Procrustes) chevrolati chevrolati Cristoforis & Jan, 1837
Male+Female adult
female, 31mm: Ashotsk,Vardaghbyur vill., 30.V.1997
male, 30 mm: Amasia, Chaldash vill.,8-11.VIII.1997
P.V. Kazaryan leg.
Copyright © Ruslan Panin

Carabus (Procerus) scabrosus tauricus Bonelli, 1811
Female adult

Copyright © Safronov

Carabus (Cechenochilus) heydenianus Starck, 1889
Male+Female adult
NW-Caucasus,
Krasnodar reg., Mostovskoy distr., Skirda Mt Range, N slopes Yatyrgvart Mt. h=2300-2500m., 09-10.VII.2008, D. Fominykh leg.
female,19mm/male,18mm
Copyright © Ruslan Panin

Carabus (Cechenochilus) adangensis Gottwald, 1983
Female adult
Russia, NW-Caucasus:
Karacaj-Cherkessija, Abishira–Akhuba Mt. Range h=2000-2650m, Kyafar-Agur Riv. 14–17.VII.2009, Fominykh D. leg.
female,21mm
Copyright © Ruslan Panin

Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Holotype
Male adult
Russia, Karachai-Circassian Republic, Abishira-Akhuba Mount Range between passes Agur and Mylgvar, 2785–2830 m.
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Paratype
Male adult
Russia, W Caucasus,
Karacaj-Cherkessija, Abishira-Akhuba Mnt Range, Area of pass Turii h~2650-2850m, 17.VII.2009, Fominykh D. leg.
male,17 mm
photo & coll. Panin R.
Copyright © Ruslan Panin

Carabus (Cechenochilus) boeberi glebi Solodovnikov, Zamotajlov & Fominykh, 2009 Paratype
Male adult
Russia, Karachai-Circassian Republic, Abishira-Akhuba Mount Range between passes Agur and Mylgvar, 2785–2830 m.
Copyright © Igor Solodovnikov

Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Male adult
W.Ukraine, Transcarpathian Reg., Rakhov Distr., Chornogora range., h=2020m. Panin R. leg. 28.VI-9.VIII.2008, male, 19 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Female adult
W Ukraine, NE Carpathians, Transcarpathian Reg., Rakhov Distr., Chornogora Mts Range h=2030m, alpin zone, 13-24.VIII.2010, Panin R. leg., female, 22 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Female adult
Ukrainian Carpathians, Transcarpathian reg., Rakhov Distr., Chornogora range, h=2010m, 29.VI-28.VII.2007, Panin R. leg., female, 23 mm., photo/det.Panin R
Copyright © Ruslan Panin

Carabus (Platycarabus) irregularis montandoni Buysson, 1882
Male adult
W-Ukraine, NW Podolian,
Lviv Reg., Zhydachiv distr., near Vybranivka, under the bark of oak tree, in the forest,12.X.2007 Panin R. leg
L= 22 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) sternbergi Roeschke, 1898
Female adult
Korea, Gyeongi Do,
Bokwang Sa 30.2.1976
female, 28 mm
Copyright © Alexander Anichtchenko

Carabus (Eucarabus) ulrichi s.str. (as volhynicus Lutshnik, 1927)
Male+Female adult
W Ukraine, NW Podolian,
Ternopil reg., Pidvolochisk distr., forest Bogdanivskiy, near Romanove Selo, 7-24.V.2008, Panin R. & Solodovnikov I. leg.
Copyright © Ruslan Panin

Carabus (Pagocarabus) crassesculptus diruptus Morawitz, 1886
Male adult
China, Gansu, Umg. Xiahe 3000-3200m
24-26.6.1993 Heinz leg.
20 mm
Copyright © Alexander Anichtchenko

Carabus (Pagocarabus) crassethoracis Kleinfeld, 1999 Holotype
Male adult

Copyright © Frank Kleinfeld

Carabus (Microplectes) riedeli Menetries, 1832
Female adult
female, 19mm
Copyright © Boleslav Brezina

Carabus (Microplectes) riedeli Menetries, 1832
Male adult
N-Ossetia,
Khazniaush Pass, h=2500-2700m, 9.VI-31.VII.1994, Zamotajlov A.leg.
male, 17mm
Copyright © Ruslan Panin

Carabus (Microplectes) convallium convallium Starck, 1889
Male adult
male, 19mm
Copyright © Boleslav Brezina

Carabus (Microplectes) convallium convallium Starck, 1889
Male+Female adult
female:NW-Caucasus, Russia,
Krasnodar reg., Apsheron Distr., Lagonaki Mt., near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominyh leg.
male:Karachaevo-Cherkessia,the right source of the river Beskess, h=2100m, 10.VIII.1997, I.A. Solodovnikov leg.
female, 21mm/male, 19mm
Copyright © Ruslan Panin

Carabus (Microplectes) convallium argonautarum Semenov, 1898
Male adult
male, 18mm
Copyright © Boleslav Brezina

Carabus (Pachystus) tamsi tamsi Menetries, 1832
Male+Female adult
Transcaucasia, Armenia,
Ashotsk, 18.VI.2001, P.V.Kazaryan leg.
female, 26mm//male, 24mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus lusitanicus Fabricius, 1801(as schaumi Gaubil, 1849)
Male+Female adult
Portugal,
Viseu distr., Gosende, h~1010m, 15.X.2009, D.Prunier leg.
female,26 mm
Copyright © Ruslan Panin

Carabus (Oxycarabus) saphyrinus pseudosaphyrinus Schweiger, 1964 (as rabaronianus)
Male adult
Anatolia, Kastamonu, Yaraligoz verso Catalzeitin,
VI.2001 Sola Claudio leg.
male, 28 mm
Copyright © Alexander Anichtchenko

Carabus (Orinocarabus) sylvestris transsylvanicus Dejean, 1826
Male adult
Ukrainian Carpathians, Transcarpathian reg., Rakhov Distr., Chornogora Mts. Petros Mt. h=1850-2000m. alpine zone, 29.VI-28.VII.2007 Panin R. Leg.
Copyright © Safronov

Carabus (Pachystus) hortensis neumeyeri Schaum, 1856
Female adult
Italy, Basilicata, Serra Calvello (PZ)
1400 m, 7.2003 F. Angelini leg.
female, 25 mm
Copyright © Alexander Anichtchenko

Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
Male adult
SW-France,
Foret d'Iraty Pyrenees-Atlantiques, d'Occabe, 15.VIII.1988
male,30 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Ohomopterus) insulicola insulicola Chaudoir, 1869
Male adult
Japan, Yamanashi pref., Kodachi, Kawaguchiko-cho, Minamitusru distr.
2.IX.2000 T. Kurihara leg.
male, 29 mm
Copyright © Alexander Anichtchenko

Carabus (Ohomopterus) insulicola insulicola Chaudoir, 1869
Female adult
15.VII.1998 JAPAN, Ichihara, env. Chiba
Copyright © Nil Belonogov

Carabus (Ohomopterus) lewisianus lewisianus Breuning, 1932
Male adult
Japan, Yamanashi pref., Koganesawa-rindo, Otsuki city,
13.6.1990 K. Kurihara leg.
male, 20 mm
Copyright © Alexander Anichtchenko

Carabus (Megodontus) violaceus fiorii Born, 1901
Male+Female adult
NW-Italy,
Reggio Emilia, Regnano, 5.IX.1996 leg. Cartini A.
female, 30 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus baeterrensis Lapouge, 1901
Male+Female adult
S France,
Midi-Pyrénées Region, Ariège, Col du Chioula h=1500m, V-VI.2005, Pascal Stefani leg.
photo Panin R.
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus aurolimbatus Dejean, 1829
Male adult
C.UKRAINE,
Poltava reg., near Novi Sanzhary, bank of Vorskla River, 1-2.VII.07, D.Kovalchuk leg.
male 25mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823
Male adult
W-UKRAINE
NW Podolian, Lviv Reg., Zolochiv Distr., near Osovitsa, steppe sites, Panin R. leg. 1.V-6.VI.07
male, 26mm
Copyright © Safronov

Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823
Male+Female adult
W. Ukraine, W. Podolian, Lviv reg., Nikolaev distr., near Krasiv vill., under the bark of oak tree, 8.II.2009, Panin R. leg. female, 29mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823 (as carbonatus Schaufuss, 1882)
Male adult
W.UKRAINE, Ukrainian Carpathians, Lviv Reg., Skolivski Beskidy Mts., near Skole, h=700m, 20.IX.2009, Panin R. leg. male, 33mm
Copyright © Ruslan Panin

Carabus (Megodontus) schoenherri schoenherri Fischer, 1822 (as sangilenus Obydov, 1997)
Male adult
Tyva, Akademika Obrucheva mt.rng., 15km N Cherbi vill.,
Chailag ur., 7.7.1999 A. & N. Menshikovy leg.
male, 26mm
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Megodontus) exaratus Quensel, 1806
Male adult
Russia, C.Caucasus, N.Ossetia,
valley of River Khazniaush,
h=1400 m, 10.VI-31.VII.1993
A. Zamotajlov leg.
Copyright © Safronov

Carabus (Megodontus) exaratus Quensel, 1806
Male+Female adult
NW Caucasus, Russia,
Krasnodar reg., Krasnodar City, 27.IX-07.X.07, D.D.Fominykh leg.
female,29mm/male, 27mm
Copyright © Ruslan Panin

Carabus (Megodontus) danilevskii Obydov, 1993
Male adult
Krasnoyarsk Area, Sayanogorsk Distr, env. Cheremushki Vill., right bank of Enisej Riv., h= 500 - 800 m, 23.07.1999 A. Semin leg.
L=17,0 mm
Copyright © Maksim Smirnov

Carabus (Rhigoidocarabus) zhubajie laojunensis Kleinfeld, 1999 Holotype
Male adult

Copyright © Frank Kleinfeld

Carabus (Archicarabus) pseudomonticola Lapouge, 1908 ? nemoralis
Male adult
NE Spain, Catalonia, Sant Jaume de Frontanyà, h=1000 m, V-VI.2010, Pascal Stefani leg.
L=26 мм
Copyright © Ruslan Panin

Carabus (Pagocarabus) draco Brezina, 1999
Male adult

Copyright © Boleslav Brezina

Carabus (Pagocarabus) caelestinus caelestinus (Imura, 2007) Paratype
Male adult

Copyright © Boleslav Brezina

Carabus (Carabus) gressittianus fuscomarginatus Kleinfeld, 1997 Holotype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Archicarabus) gotschi gotschi Chaudoir, 1846
Male+Female adult
Transcaucasia, Armenia,
Ashotsk distr., Bandivan, 8-21.X.1991 P.V.Kazaryan leg.
female, 20mm
Copyright © Ruslan Panin

Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Male adult
W.Ukraine, Transcarpathian Reg., Rakhov Distr., Chornogora Mt range h=2020m alpine zone, Panin R. leg. 28.VI-9.VIII.2008, male 19 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) daphnis askhicus Belousov, 1992 Paratype
Male adult
SC-Caucasus, W Georgia, valley of riv. Chkhorodukhu, h=1000-1200m, 14.V.-6.VII.1991, Zamotajlov A.S & Miroshnikov leg.
male, 29 mm
photo & collection Panin R.
Copyright © Ruslan Panin

Carabus (Neoplectes) daphnis askhicus Belousov, 1992 Paratype
Male+Female adult
W Georgia, valley of Chkhorodukhu riv., 1000-1200m
14.V-6.VII.1991
male 30mm, female 33mm
photo & collection Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) daphnis daphnis Kurnakov, 1962
Male adult
Georgia, Mingrelia, Lecheha vallis, Tehuri superior
Caucasus centr., 1200m, 27.6-7.7.1989 Retezar I. leg.
male, 28 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) edithae exedithae Gottwald, 1985
Male adult
Karachaevo-Cherkessia, low cource of Daut river,
16-20.V.2009 Anichtchenko A. leg

Copyright © Alexander Anichtchenko

Carabus (Neoplectes) juenthneri atchibachi Zamotajlov, 1991 Paratype
Female adult
Abkhasia,
Valley of river Bzyb near Mt. Achibakhu. h=1500m., A.S. Zamotajlov leg.
female: 33mm, 31mm
coll. R.Panin
Copyright © Ruslan Panin

Carabus (Megodontus) caelatus caelatus Fabricius, 1801
Male adult
Bosnia, Trebevic, Vidikovac-Brus, 1200m
21.11.1985 Mihljevic B. leg.
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) juenthneri avadcharensis Kurnakov, 1970 (loc.typ.)
Male+Female adult
Abkhasia, near Avadhara, Site Pyv h~1850m, 43.490730 40.686433, 1.VI-12.VII.2009, Zamotajlov A.S., Bondarenko A.S. leg.
L=27mm

Copyright © Ruslan Panin

Carabus (Neoplectes) koltzei kobachidzei Kurnakov, 1970
Male+Female adult
Caucasus, E Abkhasia, SW slope of Akiba mt.rng., 1200m, 7.V.-2.VII.1991 Zamotajlov & Miroshnikov leg.
male 37mm, female 38mm
photo & collection Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Cratophyrtus) turcosinensis medvedevi Kryzhanovskij, 1968
Female adult
S Kazakhstan, NW slope of Ugamskii mt.rng., 3 km W Sairamskii peak, 2200m
tributary of Karzhantau riv., 15.6.2004 Matalin A. leg.
20 mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) kratkyi felicitanus Reitter, 1893 (as porcellus)
Male adult
NW Caucasus, Resp. Adygea, Mt.Tybga, 28.VI.1997
male,20mm
Copyright © Ruslan Panin

Carabus (Neoplectes) kratkyi Ganglbauer, 1890
Male+Female adult
W Caucas, 1900m, Khatsavitaya Mt., 4.VI-13.VII.1994
A.Miroshnikov leg.
male 25mm, female 26mm
photo & collection Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) kratkyi Ganglbauer, 1890
Male+Female adult
W Caucas Mts., 2500m, Mt. Yatyrgvarta,
25.VI-12.VIII.1992 Shchurov V. leg.
male 20mm, female 21mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890
Female adult
NW-Caucasus, Russia,
Karacaj-Cherkessija, Abishira-Akhuba range h=2650m, upper Kyafar-Agur Riv., 4-9.VIII.2009, Solodovnikov I.A. leg.
female, 23 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890
Male adult
NW-Caucasus: Karacaj-Cherkessija,
Abishira–Akhuba Mnt. Range h=2000-2650m, Kyafar-Agur Riv., 14–17.VII.2009, Fominykh D. leg.
male, 22mm
Copyright © Ruslan Panin

Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990
Male+Female adult
W.Caucasus,
Psebai env., Nikitino, 2.V-3.VII.1993, leg. A.S.Zamotajlov
female 33mm
Copyright © Ruslan Panin

Carabus (Neoplectes) prometheus prometheus Reitter, 1887
Male adult
NW-Caucasus,Krasnodar reg., Apsheron Distr., Lagonaki Mt., near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominykh leg.
L= 42mm.
Copyright © Ruslan Panin

Carabus (Neoplectes) prometheus prometheus Reitter, 1887
Male+Female adult
NW Caucasus, Russia,
Krasnodar reg.,Krasnoarmeisk distr., Krasny Les, near Podlesnyi vill.,17-22.IV.2008, D.D.Fominykh leg.
female, 40mm
Copyright © Ruslan Panin

Carabus (Neoplectes) protensus olegi Belousov, 1992 Paratype
Male adult
Caucasus, W Georgia, Svanetia, right bank of riv. Magana, 800m, 12.V.-4.VII.1991, Zamotajlov & Miroshnikov leg.
male, 31mm
photo and collection Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) reitteri laevisternis Starck, 1890
Male adult
Abkhasia,W.slope of Gagrinsky Mt., Krepostnaya River source, h=1325m., A.S.Zamotajlov leg. 27.VII.2000
male, 29 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) reitteri laevisternis Starck, 1890
Male adult
W-Caucasus:Sochi env., Aibga Mt. h=2000m., 06.VII.2000
male, 29mm
Copyright © Ruslan Panin

Carabus (Neoplectes) satyrus napraensis Belousov & Zamotajlov, 1993 Paratype
Male adult
Abkhasia, Bzybian Mt. Range near Mt. Chibzharga., h=1900-2000m., A.S. Zamotajlov leg.
male, 28mm
photo & coll.Panin R.
Copyright © Ruslan Panin

Carabus (Neoplectes) starcki Heyden, 1885
Male+Female adult
NW-Caucasus,
Krasnodar reg., Mt. Aishkho, Malaya Laba River, h=2200m., 14.VII.2008, D.Fominyh leg.
female-21mm
Copyright © Ruslan Panin

Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988 (as shchessiensis) Paratype
Male+Female adult
NW Caucasus,
Krasnodar reg., Tuapsinskyi distr. Shessi Mt.
30.V.-25.VI.1993 Mt.Shessi, h=1800m., Imura & A.S.Zamotajlov, 1993 leg.
female, 29mm
Copyright © Ruslan Panin

Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988
Female adult
Russia,W Caucasus,
Krasnodar reg.,Chernogorie mts.,Chernogor mt. h=1700-1800m. 23.V.-30.VII.1999. A.S.Zamotajlov leg.
female 29mm.
Copyright © Ruslan Panin

Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
Male adult
NW-Caucasus, Russia,
Krasnodar reg., Apsheron Distr., Lagonaki Mt. ,near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominykh leg.
male, 28mm
Copyright © Ruslan Panin

Carabus (Neoplectes) starckianus theseus Briansky, 1910
Male+Female adult
NW-Caucasus, Russia,
Krasnodar reg.,Tuapse distr., Mt.Semashkho 04.V.2008, D.Fominyh leg.
female, 32mm
Copyright © Ruslan Panin

Carabus (Aristocarabus) viridifossulatus viridifossulatus Fairmaire, 1887
Female adult
35mm
Copyright © Boleslav Brezina

Carabus (Aristocarabus) viridifossulatus seticollis Deuve & Mourzine, 2000
Male adult
28 mm
Copyright © Boleslav Brezina

Carabus (Aulonocarabus) canaliculatus rufinus Beheim & Breuning, 1943
Female adult

Copyright © Safronov

Carabus (Aulonocarabus) truncaticollis polaris Poppius, 1905
Male adult
Arctic Russia, NW Siberia,
Yamal, 230 N from Labytnangi, Panta-Yakha Riv., 15-29.VI.2002
male,18mm.
Copyright © Ruslan Panin

Carabus (Axinocarabus) fedtschenkoi fedtschenkoi Solsky, 1874
Male adult
27 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) linxiaensis ponggartangensis Deuve, 1995 Holotype
Male adult
17 mm
Copyright © Boleslav Brezina

Carabus (Carabus) arvensis arvensis Herbst, 1784
Male+Female adult
W Ukraine, NW Podolian, Lviv reg., Javoriv distr., near Stradch vill., 20-30.IV.2009, Panin R. leg.
Copyright © Ruslan Panin

Carabus (Carabus) arvensis arvensis Herbst, 1784
Male adult
W. Ukraine , NW. Podolian, Lviv reg., Nikolaev distr., near Krasiv vill., 9.II.08, Panin R. leg. L=17mm
Copyright © Ruslan Panin

Carabus (Carabus) arvensis carpathus Born, 1902
Male+Female adult
Ukrainian Carpathians,
Transcarpathian reg., Rakhov Distr., Chornohora range, h=1850 -2000m., 29.VI-28.VII.2007 Panin R. leg.
18mm.
Copyright © Ruslan Panin

Carabus (Carabus) granulatus leander Kraatz, 1878
Male+Female adult
Russia, C-Caucasus: Balkaria, Valley of riv. Cherek Balkarskyi, h=1750-1800m 10.VI-20.VII.1992, Zamotajlov A. leg.
female,22mm
Copyright © Ruslan Panin

Carabus (Carabus) granulatus granulatus Linne, 1758
Male adult
W Ukraine,
Lviv Reg., Javoriv distr., near Ivano-Frankovo vill., 30.IV-18.V.2009, Panin R. leg.
male, 21mm
Copyright © Ruslan Panin

Carabus (Carabus) granulatus granulatus Linne, 1758
Female adult
long. 22mm
Copyright © Ruslan Panin

Carabus (Carabus) menetriesi menetriesi Hummel, 1827
Male adult
W-Ukraine, Roztochia, Lviv Reg., Yavorov distr., near Ivano-Frankovo vill., Site Zalyvki, Panin R. leg., 1-20.V.2005, male, 21 mm.
Copyright © Ruslan Panin

Carabus (Carabus) stscheglowi Mannerheim, 1827
Male+Female adult
Central Russia , Voronezh region, near Grafskoe, 12.VIII.1984 long=20mm
Copyright © Ruslan Panin

Carabus (Platycarabus) fabricii fabricii Panzer, 1812
Male adult
NE-Italy,
Trentino-Alto Adige Reg,. Bolzano-Bozen Prov.,Val Aurina – Rottal 2050-2150m, 28.VI.1997, I.Rapuzzi leg.
male, 19mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) formosus elytraviridis Kleinfeld, 2007 Holotype Paratype
Male+Female adult
China, Sichuan, Aba pref., Nanping Co., Jiuzhaigou (ca. 33.17N/103.54E), 2000 m, 30.5.-6.6.1990, coll. Kleinfeld.
HT 32 mm, PT 36 mm
Copyright © Frank Kleinfeld

Carabus (Cathaicus) brandti brandti Faldermann, 1835 (as grossegranulatus) Syntypes Lectotype
Male adult
C. (Cathaicus) brandti grossegranulatus Breuning 1943 Lectotypus/SyntypusLectotypus (25 mm) von "South Mandschou". Syntypus (25 mm) "South Mandschou, 30. V. 1937"
Copyright © Frank Kleinfeld

Carabus (Carabus) sculpturatus Menetries, 1832
Male+Female adult
Azerbaijan,
Talysh Mts. Hirkan National Park, Moscow Forest, 8-9.VII.1988, P.V.Kazaryan leg.
female, 22 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) estreicheri Fischer, 1822
Female adult
Volgograd Reg., Serafimovka Distr., Ust' -Khoperskaya Station 5.V.2009 leg.O.Sveshnikov
18.5 mm
Copyright © Kirill Makarov

Carabus (Hygrocarabus) nodulosus Creutzer, 1799
Male+Female adult
Austria, Styria Region (Steiermark), Leibnitz, 31.X.1999, I.Rapuzzi leg. female, 29 mm
Copyright © Ruslan Panin

Carabus (Hygrocarabus) variolosus Fabricius, 1787
Male adult
W-Ukraine, NW Podolian, Lviv reg., Zhydachiv distr.,near Vybranivka vill., 15.V-8.VI.2008, Panin R. leg. male, 27mm
Copyright © Ruslan Panin

Carabus (Hygrocarabus) variolosus Fabricius, 1787
Male+Female adult
W-Ukraine, Lviv Reg., Zhydachiv distr., near Vybranivka vill., Panin R. leg. female, 30mm
Copyright © Ruslan Panin

Carabus (Platycarabus) irregularis irregularis Fabricius, 1792
Male adult
NE-Italy,
Friuli Reg., Udine, Carnia, Pierabech h~1100m, I.Rapuzzi leg.
male, 23 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) manifestus pyonganicola Deuve & Li, 1998
Male+Female adult
N-Korea: Pyonganam-Do, YangTok County,(39;127), 12-19,VI.2006
Copyright © Ruslan Panin

Carabus (Platycarabus) irregularis ramanus Sokolar, 1909 (as hayeki Kobmann, 1925)
Male adult
N Slovenia, Maribor – Dravograd, Pomorje Mts.,13.XI.2005, I.Rapuzzi leg.
Copyright © Ruslan Panin

Carabus (Platycarabus) irregularis ramanus Sokolar, 1909 (as bucephalus Kraatz, 1879)
Male+Female adult
W Slovenia, Goriška region Trnovo, h~1000 m, IX.2002, I.Rapuzzi leg.
female, 29 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens trans illigeri Dejean, 1826
Male+Female adult
W-Croatia, Karlovac, near Bosiljevo vill., h~300m, V.2000, I.Rapuzzi leg. female, 33 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) creutzeri creutzeri Fabricius, 1801 (as trnovensis Bernau, 1914)
Male adult
W Slovenia,
Lokve, Trnovski Gozd h=1100m, 17.IV.2006, leg. I.Rapuzzi
male, 26mm
Copyright © Ruslan Panin

Carabus (Platycarabus) creutzeri creutzeri Fabricius, 1801(as posarinii Rapuzzi, 1999 PT) Paratype
Male+Female adult
Slovenia:Koper:Kras Mt.R.:Brkini Mts.:Suhorije env., VIII.1998, I.Rapuzzi leg.
photo & koll. Panin R.
female, 32 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) creutzeri kircheri Germar, 1824
Male+Female adult
NE-Italy, Veneto Reg., Belluno, Forcella Lavardet h=2000m, VII.2002, I.Rapuzzi leg.
female, 24 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) casaleianus casaleianus Deuve, 1994
Female adult
China, SW Hunan Prov., Chengbu Co, Nan-Shan-Ding (26.34;110,08)1-10.VIII,2006, I.Rapuzzi leg.
L=33 mm
Copyright © Ruslan Panin

Carabus (Diocarabus) lucepunctus ignipunctatus Rapuzzi, 2009 PT Paratype
Male+Female adult
China, SW Hunan Prov., Chengbu Co, Nan-Shan-Ding (26.34;110,08)1-10.VIII,2006, I.Rapuzzi leg.
femele, 18 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) morbillosus alternans Palliardi, 1825
Male adult
S Italy, Sicily, Province Palermo, Piana degli Albanesi h~700m, XI.2010, I.Rapuzzi leg.
L=32 mm
Copyright © Ruslan Panin

Carabus (Aulonocarabus) canaliculatus pseudocareniger Deuve, 1991
Male adult
N Korea, Ryanggang Prov., Mt. Tongjom Ridge h~1861m, (128.60E;41.20N) I. Rapuzzi leg.
L=27 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) manifestus guanmenshanus Imura, 1991
Female adult
NE China, Liaoning, Anshan City, Mt.Qian Shan, 4-16.VII.2005, I.Rapuzzi leg.
L=26 mm
Copyright © Ruslan Panin

Carabus (Ohomopterus) albrechti albrechti Morawitz, 1862
Female adult
Japan, S-Hokkaido:Oshima Peninsula,Pass btw Otoshibe - Assabu, h~250-300 m, 10-20.VII.2010, 42:01’53’Т; 140:22’09’E, I.Rapuzzi & L.Caldon leg.
Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus aeruginosiformis Breuning, 1932
Male adult
Russia, Khakassia,
W-Sayan, Alan Mt range, valley Enisey riv., 5 km S Mayna vill., 15.VII.2000, A.& N. Menstschicovy leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Pagocarabus) erwini erwini s.str.
Male adult

Copyright © Boleslav Brezina

Carabus (Meganebrius) arunensis Heinertz, 1980
Female adult

Copyright © Boleslav Brezina

Carabus (Lipaster) stjernvalli stjernvalli Mannerheim, 1830
Male+Female adult
Armenia,
Ashotsk, 22.VI.1990, P.V.Kazaryan leg.
female 34mm
Copyright © Ruslan Panin

Carabus (Limnocarabus) clathratus clathratus Linne, 1761
Female adult
W-Ukraine, Maloe Polesie, Lviv Region, near Kamenka-Bugzkaya, 28.III.2008, Panin R. leg. L=31 mm
Copyright © Ruslan Panin

Carabus (Leptoplesius) merzbacheri kirgisiensis Mandl, 1967
Female adult
22mm
Copyright © Boleslav Brezina

Carabus (Leptocarabus) yokoae marcilhaci Deuve, 1992
Male adult
CHINA, Shanxi, Tschingling Mts., Taibeichan Nat. reserv.;
Copyright © Anton Menstschikov

Carabus (Leptocarabus) harmandi fujisan Ishikawa, 1972
Female adult
Japan, Yamanashi pref., Fuji-Rinds 1600m, Narusawa vill. 3.V.2005 Kurihara K. leg. 25 mm
Copyright © Alexander Anichtchenko

Carabus (Lasiocoptolabrus) sunwukong sunwukong Imura, 1993 Paratype
Male adult

Copyright © Boleslav Brezina

Carabus (Lasiocoptolabrus) sunwukong baotianmanus (Imura, 2005) Paratype
Male adult

Copyright © Boleslav Brezina

Carabus (Lamprostus) calleyi pseudoprasinus Lapouge, 1912
Male+Female adult
Transcaucasia, Armenia,
Ashotsk distr. Tsoghamarg vill, .21-22.V.1997, P.V.Kazaryan leg
female, 24 mm
Copyright © Ruslan Panin

Carabus (Ischnocarabus) tenuitarsis tenuitarsis Kraatz, 1877
Male adult
male, 24mm
Copyright © Boleslav Brezina

Carabus (Iniopachus) pyrenaeus jeanneli Deuve, 1994
Male adult
1-2: Pirineos, Canfranc, Ibon de Iserias 2200 m 16-17.7.05 3: Pirineos, Monte Perdido, Collata Anisolo 2400m 25.8.05
Copyright © Alexander Anichtchenko

Carabus (Iniopachus) auriculatus ubinensis (Puissegur, 1955)
Female adult
Spain, Montes Cantabricos, Peña Ubiña, 2400m
5-6.VII.2009 Anichtchenko A. leg.
female, 19 mm
Copyright © Alexander Anichtchenko

Carabus (Imaibius) baronii battalensis Heinz & Staven, 1998 Paratype
Male adult

Copyright © Alexander Anichtchenko

Carabus (Pseudocoptolabrus) businskyi Deuve, 1990
Male adult
CH, NW-Yunnan (Lijiang) Yulongxue Shan, E-slope
27:10:50N/100:12:26E/ 4400m, alpine meadows 12.VII.2000
K. & B. Brezina, M.Brychcin, M. Hackel
Copyright © Boleslav Brezina

Carabus (Trachycarabus) sibiricus gvozdevae Obydov, 1997
Male adult
Russia, Khakassia,
W Sayan, Enisey riv., Batenevskiy Mt Range, 12 km E Pervomayskiy vill., 11.VII.2000, A.& N. Menstschicovy leg.
L=21 mm
Copyright © Ruslan Panin

Carabus (Homoeocarabus) maeander maeander s.str.
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Carabus (Cathoplius) aliai Escalera, 1944
Male adult
male, 27mm
Copyright © Boleslav Brezina

Carabus (Cathoplius) asperatus asperatus (Dejean, 1826)
Male adult
male, 24mm
Copyright © Boleslav Brezina

Carabus (Cechenotribax) petri Semenov & Znojko, 1932
Female adult
female, 18mm
Copyright © Boleslav Brezina

Carabus (Cephalornis) potanini (Semenov, 1887)
Male adult
China, S Gansu, 3400m, Minshan mts.,
Wenxian 8.92, J. M. Bousquet
Copyright © Boleslav Brezina

Carabus (Chaetocarabus) intricatus bayardi Solier, 1835
Male+Female adult
S-Italy,
Basilicata Reg., Potenza Prov., Latronico, F. Angelini leg. 14.VI.2002
female, 26mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens costellatus Gehin, 1882 (as subfestivus Oberthur, 1884)
Female adult
W- FRANCE,
Bretagne Reg., Finistrere, Foret de Coatloc'h, XI.2006
female, 26 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens costellatus Gehin, 1882
Male+Female adult
C-FRANCE,
Limousin Reg., Corrиze Dep., Aubazines, X.2005, Pascal Stefani leg.
female,26mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens cupreonitens Chevrolat, 1861
Female adult
NW-FRANCE,
Basse-Normandie Reg., Orne, Forêt de Bellême, 14.XII.2008, Pascal Stefani leg.
female, 26mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens cupreonitens Chevrolat, 1861
Male+Female adult
NW-FRANCE,
Basse-Normandie Reg,, Orne, Foret d' Ecouves, V.2009, Pascal Stefani leg.
male, 25 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens escheri (as laevipennis Seidlitz, 1891)
Male adult
W Ukraine, NE Carpathians Mts.
Transcarpathian, Rakhov Distr., Chornogora range, Petros Mt.h=2010m, 9.VI.2009, R.Panin leg
male, 25 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens escheri Palliardi, 1825 (ab. opacus Haury, 1878)
Male+Female adult
W Ukraine, NE Carpathians Mts., Transcarpathian reg., Rakhov Distr., Chornogora range, h=2020m, alpin zone, 28.VI-22.VIII.2009, Panin R. leg. female, 27mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens farinesi Dejean, 1826 (as barthei Lapouge, 1912)
Male+Female adult
SC -France,
Midi-Pyrйnйes reg., Ariиge, Foret de Belesta, XI.2008, J.Nicolas leg.
L=31 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens montanus Gehin, 1882
Male+Female adult
SC-FRANCE,
Haute-Garonne, Col de Mentй h~1300m., II.1980. Kretschmer leg.
female, 23mm.
photo/det. Panin R.
Copyright © Ruslan Panin

Carabus (Chrysocarabus) solieri liguranus Breuning, 1933
Female adult
NW Italy,
Altare, Savona,15.VI.1995, Cartini A.leg.
female, 33mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) splendens Olivier, 1790
Male+Female adult
SE France,
Midi-Pyrйnйes reg., Ariиge, Vaychis, 22.X.2002, A.Laforgue leg.
female, 30mm.
Copyright © Ruslan Panin

Carabus (Chrysocarabus) hispanus Fabricius, 1787
Male adult
FRANCE,
Tarn, Foret de la Gresigne, h=400m.VII.2005 Laforgue leg.
male-30mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896 (as juberriensis Mollard, 1999)
Male+Female adult
ANDORRA,
E-Pyrenees, Juberri, h=1350-1400m, 22.VI-18.VII.2007
female, 33mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
Male+Female adult
Russia, S Siberia, W Sayan, Ergaki Mt range, Nizhnyaya Buyba river h=1700 m, 19.VII.1999, A.& N. Menstschikovy leg.
Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus shoriensis Obydov, 1999
Male adult
Russia, Khakassia,
Abakanskii Mt Range, Askiz river vall., 7 km E Kazanovka vill., 15.VI.2006, A.& N. Menstschikovy leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Diocarabus) loschnikovi tsaganensis Obydov, 2001
Male adult
Russia, W Tuva,
Kyzyl-Tayga Mt Range h=1500 m, Ak-Suk riv vall., 21.VI.2006, A.& N. Menstschikovy leg.
L=16 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) obsoletus obsoletus Sturm, 1815
Male adult
NW Ukrainian Carpathians, Lviv Reg., Staryi Sambor Distr., near Dobromyl , h=500 m., 15-24.VI.2005, Panin R. leg., male, 28mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997
Male+Female adult
Russia,Tuva,
Akademika Obrucheva Mt range, 15 km N Cherbi vill., Chailang ur. 7.VII.1999, A & N. Menstschikovy leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Hemicarabus) tuberculosus Dejean, 1829
Male adult
Far East, Lazovsky Dist., shore of Japan Sea, 3 km NE Preobrazhenie Vill., Ezhovaya (Tasovaya) Bay 13.07.06 M. & L. Smirnov's leg.
male
L=20,0 mm
Copyright © Maksim Smirnov

Carabus (Hemicarabus) nitens Linne, 1758
Male+Female adult
Belarus,
Vitebsk reg., Ushachskiy distr., near Bolshie Dolcy vill. 9.V.1990, I.A.Solodovnikov leg.
female, 18mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei Kuster, 1846 (as dacicus Csiki, 1906)
Female adult
NW Romania , Bistrita Nasaud, Beclean env., 17.VI.2009, S.Budka leg.
female, 31 mm
Copyright © Ruslan Panin

Carabus (Heterocarabus) marietti marietti (as ativsengueni Schweiger, 1962)
Male adult
male, 17mm
Copyright © Boleslav Brezina

Carabus (Goniocarabus) sogdianus Semenov, 1898
Male adult
male, 21mm
Copyright © Boleslav Brezina

Carabus (Gnathocarabus) kusnetzovi Semenov, 1903
Female adult
female, 21mm
Copyright © Boleslav Brezina

Carabus (Fulgenticarabus) flutschi koreus Deuve, 2006
Male adult
N Korea, Samjiyon, Ryang Gang Province
6-12.6.2006 leg. Wong
Copyright © Boleslav Brezina

Carabus (Megodontus) kolbei aino Rost, 1908
Male adult

Copyright © Kirill Makarov

Carabus (Pachycarabus) koenigi Ganglbauer, 1886
Male adult
NW Caucasus, S Russia, Adygea Rep., Lagonaki plateau, Site Murzikal h=1800-2000m, 24.V-18.IX.2010, D.Fominykh leg.
L= 22 mm
Copyright © Ruslan Panin

Carabus (Carabus) granulatus interstitialis Duftschmid, 1812
Male adult
Italy, Emilia reg., Modena, Castelnuovo Rangone, 31.XII.1991, Sola C. leg.
L=19 mm
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
Male adult
Italy-Switzerland, Riviera – Preonzo, 18.II.1998, C.Sola leg.
L=26 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) castanopterus A. & J.B. Villa, 1833
Female adult
NC-Italy - S-Switzerland, Val Colla – Gazzirola, 22.VI.1999, C. Sola leg.
L=21mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) helleri Ganglbauer, 1893 (as pseudopreyssleri Breuning, 1932)
Male adult
Czech Rep., NE Moravia, Moravian-Silesian Beskids Mt Rng, PR Trojacka, Hodslavice vill., 29.V.12.VI.2010, Krause Rene leg./det.Panin R. L=30 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) kollari Palliardi, 1825 (as semetricus Kraatz, 1878)
Male adult
SW-Romania:Banat, Bocsa/Coltan, 21.V-5.VI.2010, Krause Rene leg.
L=29 mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) convexus gracilior Gehin, 1885
Male adult
SE-Bulgaria, Burgas Prov., Primorsko env., 18-30.VI.2000, Krause Rene leg.
L=18 mm
Copyright © Ruslan Panin

Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
Female adult
SW Slovakia, Bratislava env., IV.2004, Krause Rene leg.
L=32 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) sylvestris sylvestris Panzer, 1793 (as slabai Niedl, 1968)
Female adult
Czech, WS Bohemia,
Plzen Reg., Domazlice Distr., Babylon, h=500m, 30.VIII.2008, S.I. Budka leg.
L=24mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus violaceus Linne, 1758
Male adult
S.Bohemia, Hluboka nad Vltavou, h=380 m, 6.VIII.2004, S. Budka leg.
L=28 mm
Copyright © Ruslan Panin

Carabus (Carabus) arvensis deyrollei Gory, 1839
Female adult
NW Spain, Galicia Reg., Pontevedra Prov., A Lama, Xesta, h~650 m, 1.X.1994
L=17 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) wiedemanni wiedemanni Menetries, 1836
Male adult
SE-Bulgaria:Burgas env. Tsarevo (Michurin), 10-15.VII.1981, J.Slaba leg. L=22 mm
Copyright © Ruslan Panin

Calosoma (Charmosta) lugens Chaudoir, 1869
Male adult

Copyright © Kirill Makarov

Calosoma (Campalita) auropunctatum auropunctatum Herbst, 1784: 131
Female adult

Copyright © Kirill Makarov

Calosoma (Campalita) auropunctatum dzungaricum Gebler, 1833: 274
Male adult
W Kazakhstan, Emba riv., near Mijaly vill., 02.07.2009, P.Gorbunov leg.; 27 mm.
Copyright © Anton Menstschikov

Carabus (Aulonocarabus) gossarei gossarei Haury, 1879
Male adult

Copyright © Kirill Makarov

Carabus (Apoplesius) omphreodes Reitter, 1898
Male adult

Copyright © Kirill Makarov

Carabus (Mesocarabus) problematicus harcyniae Sturm, 1815 (as obenbergerianus Breuning, 1927)
Male adult
Czech, Bohemia, Plzeň Reg., Domažlice Distr., Babylon, h=500 m, 30.VIII.2008, S.I. Budka leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus germarii Sturm, 1815
Female adult
Croatia, Ivanščica Mts , Ivanec, 6-13.VI.2003, Krause Rene leg.
L=36 mm
Copyright © Ruslan Panin

Carabus (Chaetocarabus) intricatus krueperi Reitter, 1896
Male adult
CE-Greece: Pieria, Stomio (Larissa) Ossa Mt. 7-14.VII.2009, Krause Rene leg.
L=35 mm
Copyright © Ruslan Panin

Carabus (Procrustes) coriaceus banaticus Redtenbacher, 1849
Male adult
SW Romania, Banat, Băile Herculane env., Domogled valley, 24-30.V.2008, Krause Rene leg.
L=35 mm
Copyright © Ruslan Panin

Carabus (Procrustes) chevrolati thirki Chaudoir, 1857
Male adult
NE Turkey, Trabzon, Zigana Pass, h~2000m, V. Slovak leg.
L=27 mm
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus graniger Palliardi, 1825
Male adult
SW-Romania, Banatului Montan, Resita - Valiug, 29.V-9.VI.2005. Krause R. leg.
L=25 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) catenulatus catenulatus Scopoli, 1763
Female adult
NW Croatia, Bosanci, 1-12,VI.2004, Baros Slovak leg.
L=33 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825 (as superbus Kraatz, 1878)
Male+Female adult
SW-Romania,
Banat, Moldova Noua, Ravenska viil., 29.V-12.VI.2006, Krause Rene leg.
female,31mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei macairei Dejean, 1826
Female adult
SW-Romania, Banat, Resita - Valiug, 29.V- 9.VI.2005, Krause R. , Baros, Pittner leg.
L=19 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) lineatus lateralis Chevrolat, 1840 (as quasilineatus Tarrier, 1975)
Male+Female adult
NW Spain,
E Asturias, Monte de Moal h~1000m., 5.VI.2008, Pascal Stefani leg.
female, 27mm.
Copyright © Ruslan Panin

Carabus (Trachycarabus) estreicheri Fischer, 1822 (as jaegeri Ménétriés, 1827)
Female adult
E UKAINE, Lugansk city, Site Luganskoe, 20-30.VII.2010, V.A.Landik leg.
L=21 mm
det.Panin.R.
Copyright © Ruslan Panin

Carabus (Goniocarabus) sogdianus Semenov, 1898
Male adult
N Tajikistan, SW Pamir-Alay, Zarafshan Range, Fann Mts, Marguzor Lake circ., Padrut vill., 16.V.1997
L=21 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus geblerianus Obydov, 1999
Male adult
Russia, NW Altay, Barnaul env.,23.07.1998, D.Lomakin leg.

Copyright © Ruslan Panin

Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859
Male adult
Russia, SE Siberia, Transbaikalia, N Chita reg., Kodar Mt. Range, Big Leprindo Lake env.h~1800m., 1.VII.2006, S. Rybalkin leg. L=20 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus tscherkessicus Korge, 1964
Male adult

Copyright © Kirill Makarov

Carabus (Morphocarabus) rothi hampei (as ormayi Reitter, 1896) loc.typ.
Male adult
W. UKRAINE,
Transcarpathian reg., near Beregovo, 27.IV-22.V..2011,, Panin R. & Chyzhevsky O. leg.
L=28 mm
Copyright © Ruslan Panin

Carabus (Deroplectes) staudingeri Ganglbauer, 1886
Female adult
N Tajikistan, SW Pamir-Alay, Fann Mts h=1800m, Padrut vill., 2-3.V.1997
L=28 mm
Copyright © Ruslan Panin

Carabus (Euleptocarabus) porrecticollis porrecticollis Bates, 1883
Male adult
Japan: Niigata pref., Chayagahara Jyouetsu City,
18.XI.2006 A.Toba leg.
Copyright © Alexander Anichtchenko

Carabus (Eotribax) eous eous Morawitz, 1889
Male adult
Kirghyzstan, N Terskei Ala Too, Borskaun riv., h=2500 m., 7-14.07.2000
male
L=17 mm
Copyright © Maksim Smirnov

Carabus (Eocechenus) kaznakovi nangqenicus Deuve, 1995 Paratype
Male adult

Copyright © Boleslav Brezina

Carabus (Eccoptolabrus) exiguus nivium Breuning, 1933
Female adult
female, 20mm
Copyright © Boleslav Brezina

Carabus (Eccoptolabrus) exiguus exiguus Semenov, 1898 Holotype
Male adult
male, 18 mm
Copyright © Frank Kleinfeld

Carabus (Megodontus) vietinghoffi bowringi Chaudoir, 1863
Male adult
Russia, Far East, Primorskiy Kray, Ussuriysk Distr., Kaymanovka vill , 26.VII-3.VIII.2011, A. Korshunov leg.
L=29
Copyright © Ruslan Panin

Carabus (Megodontus) obovatus Fischer, 1827
Male adult
SW Siberia, Russia, Kemerevo reg., Krapivino distr., Azhendarovo, forest, 26-31.VII.2011, D. Efimov leg.
L=18 mm
Copyright © Ruslan Panin

Carabus (Carabus) arvensis faldermanni Dejean, 1829
Male+Female adult
Russia, Far East, Primorskiy Kray, Ussuriysk Distr., Kaymanovka vill., 11.VII-1.VIII..2011, A. Korshunov leg.
female, 23 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens (f.i. frivaldszkyi Kraatz, 1887)
Female adult
W-UKRAINE, NW Podolian Upland, Lviv Reg., Zolochiv Distr., Lysa Gora steppe massif, 14-30.V.2008, Panin. R. leg./det. L=26 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens Fabricius, 1798
Male adult
C- UKRAINE, NW Odessa reg., Kodyma distr., near Aleksandrivka vill.,18.V.2003, Demidov G.N. leg. L=26 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) fabricii fabricii (as fassatii Smetana, 1955)
Male adult
C-Slovakia:Nízke Tatry Mts., 4.VI.2011, Krause Rene leg.
L=19 mm
Copyright © Ruslan Panin

Carabus (Diocarabus) slovtzovi slovtzovi Mannerheim, 1849
Male adult
S Siberia, Burjatia reg., E Sayan Mts., Khugaituma mt., h=1700 m, 1-5. VII.2001
L=14,3 mm

Copyright © Maksim Smirnov

Carabus (Diocarabus) beybienkoi Kryzhanovskij, 1973
Male adult
S Sakhalin, Susunaj Mt.R. 10.08 - 13.09.2003 D. Kochetkov leg.
L=14,5 mm
Copyright © Maksim Smirnov

Carabus (Deroplectes) arcanus arcanus Semenov, 1898
Male adult

Copyright © Boleslav Brezina

Carabus (Morphocarabus) praecellens trans scheidleri
Male adult
NC Croatia, Ivanščica Mts , Ivanec,4-18.VI.2011, Krause Rene leg. L=31 mm
Copyright © Ruslan Panin

Carabus (Procrustes) coriaceus hopffgarteni (as emgei Ganglbauer, 1888)
Male adult
CE-Greece: Larissa reg. Stomio, Ossa Mt. 6-17.VII.2011, Krause Rene leg.
L=32 mm
Copyright © Ruslan Panin

Carabus (Chaetocarabus) arcadicus arcadicus Gistl, 1850
Female adult
CE-Greece, Kato Olympos, Karia env., 6-17.VII.2011, Krause Rene leg. L=37 mm
Copyright © Ruslan Panin

Carabus (Procerus) gigas gigas Creutzer, 1799
Male+Female adult
N Croatia, Ivanščica Mts , Ivanec, 4-18.VI.2011, Krause Rene leg.
female, 54 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus inflatus Kraatz, 1878
Male adult
SE–France, Var departament, Foret royale -Saint-Paul-en-Foret vill.,18.VI.2002, Thelot J.P. leg.
L=26 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) solieri solieri Dejean, 1826
Female adult
SE France, Alpes-Maritimes, Ascros, VI.2011
L=30 mm
Copyright © Ruslan Panin

Carabus (Cychrostomus) pseudoprosodes Semenov & Znojko, 1932
Male adult
male, 19mm
Copyright © Boleslav Brezina

Carabus (Cychrostomus) anchocephalus lanzhouensis Deuve, 1991 Paratype
Male adult
male, 21mm
Copyright © Boleslav Brezina

Carabus (Cychrostomus) anchocephalus anchocephalus Reitter, 1896
Male adult
male, 22mm
Copyright © Boleslav Brezina

Carabus (Platycarabus) depressus lucens Schaum, 1856
Female adult
SE-France: Alpi Marittime, Monte Viso, 14.VII.2010
L=23 mm
Copyright © Ruslan Panin

Carabus (Oreocarabus) luetgensi getschmanni Lapouge, 1924
Male adult
NW-Spain, León Province, Acebo, h=1000m, V.2007, Pascal Stefani leg.
L=22 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus planiusculus Haury in Gehin, 1885
Male adult
NE Spain, Catalonia, Sant Jaume de Frontanyà, h=1000 m, V-VI.2010, Pascal Stefani leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Tachypus) auratus lotharingus Dejean, 1826
Male adult
SC France: Herault departament, Foret de Saint Gervais, V. 2009 leg. J. Nicolas
L=27mm
Copyright © Ruslan Panin

Carabus (Eucarabus) obsoletus nagyagensis Seidlitz, 1888
Female adult
NW-Romania, Mts Ses (Plopis),Cluj Reg., Bucea env, h=580 m, 8.VI.2011, Sl.Budka leg.
L=26 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei Kuster, 1846
Male adult
C Romania, Târgu Mureş, Ernei, 46°35′471″N 24°37’351″E, h=300 m,20.VIII.2004, Sl.Budka leg.
L=30 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei (as zilahiensis Csiki, 1906)
Male+Female adult
NW Romania, Zalău env., 31.VII.2003, Sl.Budka leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) fabricii fabricii (as fassatii Smetana, 1955)
Male adult
C-Slovakia:Nizke Tatry Mts., h=1750 m,.9.VI.2007, V.Slovak leg.
L=20 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) aumonti maroccanus Bedel, 1895
Male adult
N Morocco (Spain:Melilla), hipodromo, X.1969, leg. Pardo Alcaide
L=38 mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) bessarabicus ? concretus Fischer, 1823
Male adult
E Ukraine, Lugannsk Reg., Sverdlovsk distr., Steppe Reserve “Provalsky", 3.V-3.VI.2011, leg.Glotov S./det.Panin R. female, 22 mm
Copyright © Ruslan Panin

Carabus (Pachystus) hungaricus hungaricus Fabricius, 1792
Female adult
C Hungary, Pest reg., Örkény, 9-10.V.1997, S.Budka leg.
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei (as hunyadensis Csiki, 1926)
Male adult
CW Romania, Apuseni Mts, Muntii Metaliferi, Hunedoara reg., near Sacaramb vill., 6.VI.2011, S.Budka leg.
L=37 mm
Copyright © Ruslan Panin

Carabus (Megodontus) vietinghoffi bowringi (as lazoensis Obydov, 1999)
Male+Female adult
Russia, Far East, Primorsky reg., S. Sichote-Alin Mts.R., Kavalerovo vill. env.,
5-25.VII.1992, S. Kunitzyn leg.
male - 28 mm, female - 30 mm.
Copyright © Alexandr Safronov

Carabus (Megodontus) vietinghoffi bowringi Chaudoir, 1863
Male+Female adult
Russia, Far East, Primorsky reg., Khasansky distr.,
Zanadvorovka vill., Gusevsky pit.
M - 19-21.VI.2007, body lenght 29 mm.
F - 15.VII.2008, body lenght 30 mm.
S. Veriga leg.
Copyright © Alexandr Safronov

Carabus (Neoplectes) satyrus duripshensis Kurnakov, 1972
Male+Female adult
W.Caucasus, Abkhasia, Gudauta distr.,
Duripsh village env., Khipsta valley, h=500-700 m.
07-12.VI.2009 A.Safronov leg.
Body lenght 30-33 mm.
Copyright © Alexandr Safronov

Carabus (Cyclocarabus) aulacocnemus galinae Obydov, 2003 Paratype
Male+Female adult
W-Tianshan:Karatau Mts.,N-part:Boialdyr River valley,
N of Ken-su, 600-750m, 22-V-2000 М. Danilevsky leg.
male - 20 мм, female - 19 мм.
Copyright © Alexandr Safronov

Carabus (Cyclocarabus) namanganensis namanganensis Heyden, 1886
Male+Female adult
Kazakhstan, S. Karatau Mts. R., Mashat vill. env.
23-IV-1993 М. Danilevsky leg.
male - 21 мм, female - 22 мм.
Copyright © Alexandr Safronov

Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878 (as viridipennis Kraatz, 1886, nec Lewis,18
Male+Female adult
Russia, Far East,Primorsky reg., S. Sikhote-Alin Mts. R., Kavalerovo vill. env.,
5-25.VII.1992, С. Kunitzyn leg.
male - 25 mm, female - 25 mm.
Copyright © Alexandr Safronov

Carabus (Morphocarabus) hummeli pusongensis Imura, 1993
Male+Female adult
Russia, Far East, Primorsky ter., Khasansky distr., Zanadvorovka vill.,
Gusevsky pit, S.Veriga leg.
male - 13.VII.2008, female 19-21.VI.2007
male - 28 мм, female - 28 мм.
Copyright © Alexandr Safronov

Carabus (Morphocarabus) hummeli hummeli Fischer, 1823
Male+Female adult
Russia, Far East, Khabarovsk reg., Obluchye env.
M. Khingan Mts. R.
10-20.VII.1992 S. Andreev leg.
male - 26 mm, female - 27 mm.
Copyright © Alexandr Safronov

Carabus (Pantophyrtus) turcomanorum turcomanorum Thieme, 1881
Male+Female adult
Kyrgyzstan, Kyrgyzskyi Mts. R., Uzun Bulak Gorge, h=1800 ms.
01-09.VI.2002 loc. coll.
male - 28 mm, female - 33 mm.
Copyright © Alexandr Safronov

Carabus (Cupreocarabus) kangdingi kangdingi Korell, Kleinfeld & Gorgner, 1992 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Cupreocarabus) brezinai Deuve, 1994
Male adult

Copyright © Boleslav Brezina

Carabus (Ctenocarabus) galicianus Gory, 1839
Male adult
Spain,
Sierra de la Cabrera Baja, 8 km SW Truchillas, 9.VII.2008, Anichtchenko A. leg.
L= 20 mm
Copyright © Alexander Anichtchenko

Carabus (Cryptocechenus) sifanicus Semenov, 1898 (as oxydocupreus)
Male adult
male, 17mm
Copyright © Boleslav Brezina

Carabus (Cryptocarabus) subparallelus Ballion, 1878
Female adult
Kazakhstan,
near Alma-Ata, V.2009, Pak O.leg.
female, 17mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) adamsi Adams, 1817 (as chiragricus Fischer, 1828)
Male+Female adult
Russia, N. Caucasus, Stavropol reg., Zheleznovodsk, Zheleznaya Mt., h=600 m. 5-11.VI.2011, Е. Shankhiza leg. male - 24 mm, female - 25 mm.
Copyright ©

Carabus (Sphodristocarabus) varians varians Fischer, 1827
Male+Female adult
N. Armenia, Idzhevan distr., Acharkut vill. env., h=800 m. 26-28.VII.2008 S. Andreev leg. male - 22 mm, female - 23 mm.
Copyright ©

Carabus (Cryptocarabus) lindemanni Ballion, 1878
Female adult
SE Kazakhstan, Alma-Ata env., 16.04. 2000
L=16,0 mm
Copyright © Maksim Smirnov

Carabus (Cechenotribax) petri Semenov & Znojko, 1932
Male adult
SE Kazakhstan, N-Tianshan:N-slopes Ketmen Mt.Rng. h=2250-2500m., near Ketmen vill.,1-5.VI.1990, Dolin V.G. leg.
L=13.5 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) regalis tschusovae Obydov, 2004
Male+Female adult
NE-Kazakhstan, W Altay, Ivanovsky Mt.Rng, near Ridder (Leninogorsk), h-1300-1500m, 4-6.VII.2003
female, 25 mm
det.Panin R.
Copyright © Ruslan Panin

Carabus (Neoplectes) satyrus duripshensis Kurnakov, 1972
Male adult
W.Caucasus, Abkhasia, Gudauta distr.,
Duripsh village env., Khipsta valley, h=500-700 m.
07-12.VI.2009 A.Safronov leg.
male - 30 mm.
Copyright © Alexandr Safronov

Carabus (Morphocarabus) regalis jenissoni Dejean, 1831 (as cyanescens Kraatz, 1882, nec Sturm, 1815)
Male+Female adult
Kazakhstan, E. Kazakhstan area,
Zyryanovsk env., near Putintzevo vill.
10-VI-2005 M. Danilevsky leg.
male - 24 mm, female - 25 mm.
Copyright © Alexandr Safronov

Carabus (Ophiocarabus) regeli obscurior Semenov, 1888
Male+Female adult
Kyrgyzstan, NE Tian-Shan, E. Terskey Alatau Mts. R.,
Kok-kia River, h=2800 м.
7-8.VI.1997 М. Danilevsky leg.
Body lenght 15-18 mm.
Copyright © Alexandr Safronov

Carabus (Semnocarabus) carbonicolor carbonicolor Morawitz, 1886
Male+Female adult
Kyrgyzstan, NC Tianshan, Kyrgyz Mt. R., Tuyuk riv., h=2000 м.
25-VI-1997 М. Danilevsky leg.
male - 20 mm, female - 21 mm.
Copyright © Alexandr Safronov

Carabus (Pachycarabus) roseri soganliensis Heinz & Korge, 1964
Male+Female adult
Turkey, Rize prov., Ikizdere distr., Rize Daglari,
47 km to SE from lyidere, Sivrikaya Koyu, h=2025 m.
30.VI-5.VII.2011, A. Vlasenko leg.
male - 23 mm, female - 24 mm.
Copyright © Alexandr Safronov

Carabus (Neoplectes) juenthneri avadcharensis Kurnakov, 1970
Male+Female adult
W. Caucasus, Abkhasia, 8 km NE of Ritza Lake, Avadkhara valley, h=1700 m 10-VI-2009 A.Safronov leg.
Copyright ©

Carabus (Procrustes) chevrolati thirki Chaudoir, 1857
Male+Female adult
Turkey, Rize prov., Ikizdere distr., Rize Daglari,
47 km to SE from lyidere, Sivrikaya Koyu, h=2025 m.
30.VI-5.VII.2011, A. Vlasenko leg.
male - 28 mm, female - 31 mm.
Copyright © Alexandr Safronov

Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
Male+Female adult
SW Siberia, Kemerovo reg., Krapivino distr., 8 km SSW Saltymakovo vill., h=150 m. 1-20.VII.2010 D. Efimov leg. male - 25 mm, female - 26 mm.
Copyright ©

Carabus (Megodontus) danilevskii Obydov, 1993
Male+Female adult
Russia, SW Siberia, Krasnoyarsk reg., W. Sayan, near Cheryomushki.
26-29.VII.1996 loc. coll.
male - 18mm, female - 20 mm.
Copyright © Alexandr Safronov

Carabus (Megodontus) kantaikensis Gehin, 1885
Female adult
Russia, SW Siberia, W. Sayan, Krasnoyarsk reg.,
S of Abakan, near Cheryomushki, Dzhoysky Pass.
07-VII-1994 M. Danilevsky leg.
body lenght 18 mm.
Copyright © Alexandr Safronov

Carabus (Megodontus) obovatus Fischer, 1827
Male adult
SW Siberia, Kemerovo reg., Krapivino distr.,
8 km SSW Saltymakovo vill., h=150 m.
20-30.VI.2010 D. Efimov, A. Korshunov leg.
body lenght 18 mm.
Copyright © Alexandr Safronov

Carabus (Megodontus) imperialis Fischer, 1823
Male adult
NE Kazakhstan, Zyrjanovsk distr., Putincevo vill. env.
25-VI-2005 M. Danilevsky leg.
male 24 mm, female 26 mm.
Copyright © Safronov

Carabus (Morphocarabus) helleri Ganglbauer, 1893
Male+Female adult
CZECH, SE Moravia, Velka nad Velickou. 20-V-2005, V. Slovak leg. male - 26 mm, female - 30 mm.
Copyright © Alexandr Safronov

Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825
Male+Female adult
Male - SW Romania, Banat - Bocsa/Coltan,
21.V-05.VI.2010, R. Krause leg., L=29 mm.
Female - SW Romania, Banat - Saska Montana, Stinapari,
25-VI-2010, R. Krause leg., L=31 mm.
Copyright © Alexandr Safronov

Carabus (Megodontus) violaceus violaceus Linne, 1758 (as intermarinus Blumenthal & Staven, 1985)
Male+Female adult
Poland, Gdansk distr., Gdynya env., Reservat «Cisowa»,
5-9.08.1998 I. Solodovnikov leg.
female - 26 mm.
Copyright © Alexandr Safronov

Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860 (as ursuius Lapouge, 1911)
Male+Female adult
FRANCE (64), Ursuya Mt.
XII-2009, P. Stefani leg.
male - 30 mm, female - 32 mm.
Copyright © Alexandr Safronov

Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896
Male adult
Anyos (ANDORRE), h=1450ms,
5-VI-2010, P. Stefani leg.
body lenght - 32 mm.
Copyright © Alexandr Safronov

Carabus (Tribax) biebersteini otcharensis Kurnakov, 1970
Male adult
W. Caucasus, Abkhazia, Gudauta distr.,
Duripsh vill. env., Khipsta River, h=700 m,
11-06-2009 A. Safronov leg.
male - 40 mm
Copyright © Alexandr Safronov

Carabus (Neoplectes) polychrous polychrous Rost, 1892
Female adult
W.Caucasus, Abkhasia, Gudauta distr., Duripsh village env., Khipsta valley, h=500-700 m. 8-11.06.2009 A. Safronov leg. lenght 40-43 mm.
Copyright ©

Carabus (Cratophyrtus) kaufmanni segregatus Morawitz, 1886
Female adult
female, 19mm
Copyright © Boleslav Brezina

Calosoma (Carabosoma) eremicola Fall, 1910: 91
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Carabus (Apotomopterus) dechambreianus dechambreianus Deuve & Li, 2000
Male+Female adult
Gailin, Guanxi prov., CHINA,
17-V-2011 loc. coll.
male - 36 mm, female - 38 mm.
Copyright © Alexandr Safronov

Carabus (Apotomopterus) shun floridus Cavazzuti & Ratti, 1998
Male+Female adult
CHINA, Guangdong prov., Mts. Nanling Reserve, 1-7.VII.2011 loc. coll. male - 38 mm, female - 40 mm.
Copyright © Alexandr Safronov

Carabus (Neoplectes) reitteri gagrinus Starck, 1894 (as gegicus Gottwald, 1985)
Male+Female adult
W.Caucasus, Abkhasia, Gagra distr., Gega riv. valley,
13-15.07.2005 D. Kasatkin. leg.
male - 30 mm, female - 34 mm.
Copyright © Alexandr Safronov

Carabus (Morphocarabus) rothi hampei as telekii Csiki, 1937 (? hampei )
Female adult
CW Romania, Arad distr., Zarandulu Mt Range, near Ilteu vill., h=276 m, 46°66′487″N 22°20’455″E, 25.VI.2005, Sl.Budka leg.
L=30 mm

Copyright © Ruslan Panin

Carabus (Cratocephalus) solskyi Ballion, 1878
Male adult
SE Kazakhstan, Ketmen Mt. rng., Tuyuk vill., 16.06.97 leg. S.Toropov
Copyright © Maksim Smirnov

Carabus (Cratocechenus) jacobsoni Semenov, 1898
Male adult
S Kazakhstan, Almaty reg., Big Lake, h=2800 m, 25.VI.96
male
L=14 mm
Copyright © Maksim Smirnov

Carabus (Cratocechenus) akinini akinini Morawitz, 1886 (as oitalensis ) Paratype
Male adult
male, 20mm
Copyright © Boleslav Brezina

Carabus (Cratocechenus) akinini akinini Morawitz, 1886
Female adult
female, 21mm
Copyright © Boleslav Brezina

Carabus (Cratocarabus) puer Morawitz, 1886
Female adult
female, 16mm
Copyright © Boleslav Brezina

Carabus (Coptolabrus) smaragdinus mandschuricus Semenov, 1898
Male adult
Russia, Far East, South Primoriye, Kaymanovka village, 10-20.07.2000; P.Gorbunov leg.
Copyright © Anton Menstschikov

Carabus (Coptolabrus) smaragdinus longipennis Chaudoir, 1863
Male+Female adult

Copyright © Alexandr Safronov

Carabus (Coptolabrus) smaragdinus longipennis (as mucdenensis Hauser, 1921)
Male adult
Mt. Wulongshan, Dandong City,
Liaoning Province, CHINE
25-30.VI.2008
Copyright © Alexandr Safronov

Carabus (Coptolabrus) pustulifer pustulifer Lucas, 1869
Female adult
Chine, C.Sichuan, Ya'an, 07.2005 Local people
Copyright © Igor Solodovnikov

Carabus (Coptolabrus) pustulifer guerryi Born, 1903
Female adult
N Yunnan: Gaoligongshan, h =1300 m, 06.2003 Local people
Copyright © Igor Solodovnikov

Carabus (Coptolabrus) pustulifer (as cyanelongatus Deuve, 2006)
Female adult
Zhagyab, Quamdu City, Tibet 8-15.VI.2008
Copyright ©

Carabus (Chrysocarabus) lineatus salmantinus Bolivar, 1922
Male adult
CW Spain, Cáceres, San Martín de Trevejo,24.VI.2011, A. Diez leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) rugosus boeticus Deyrolle, 1852
Male adult
SC Spain, Jaen reg., Puente de la Sierra, 10.VII.2011, M.Lopez leg.
L=28 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi rothi Dejean, 1829
Male+Female adult
C Romania, Sibiu Area, right bank Olt River, Bradu vill., h=380 m, 19.VIII.2001, S.Budka leg.
L=28 mm
Copyright © Ruslan Panin

Ceroglossus suturalis suturalis Fabricius, 1775
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Carabus (Morphocarabus) venustus venustus Morawitz, 1862
Male adult
Russia, Far East, Primorskiy reg., Ussuriysk Distr., Kaymanovka vill., 6-15.VIII.2011,
A. Korshunov leg.
L= 21.5 mm
Copyright © Ruslan Panin

Carabus (Pachystus) hungaricus cribellatus Adams, 1812
Male adult
E Siberia, Russia, Krasnoyarsk Reg., near Krasnoyarsk, forest-steppe,1.VII-20.VIII,2011,M.Ivanov leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Pachycarabus) roseri soganliensis Heinz & Korge, 1964
Female adult
NE-Turkey: Rize Prov.Ovit Pass (40.37;40.46) h-2700m,15.VII.2008, I.Rapuzzi & Caldon leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus sibiricus Fischer, 1822
Male adult
E Siberia, Russia, Krasnoyarsk Reg., near Krasnoyarsk, on dry steppe slopes, 7.VIII,2011, M.Ivanov leg.
L=21 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) latus latus Dejean, 1826
Female adult
CW Spain, Cáceres, San Martín de Trevejo, 24.VI.2011, A. Diez leg./det.Panin R.
L=27 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) dufouri dufouri Dejean, 1829
Male adult
SC Spain, Jaen reg., Monte Javalcuz h=1436 m, 6.X.2010, M.A. Lopez leg.
L=27 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus harcyniae (as procedens Csiki, 1927)
Male adult
England, North Yorkshire, Fylingdales Moor, Grouse Hill, VII.2006 L=24 mm
Copyright © Ruslan Panin

Carabus (Pachystus) hungaricus scythus Motschulsky, 1847
Female adult
S Ukraine, Zaporizhzhya reg., Khortytsa island, 16-20.VII.2011, M. Mulenko leg., det. Panin R.
L=28 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus brevis (as sorianus Jeanne, 1972)
Male adult
NC-Spain, Soria, Puerto de Oncala, 2.VI.2006
L=24 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) principalis Bates, 1889
Male+Female adult
Zi-Ding-Shan, h=1406 m, Guiyang County, Binzhou City, Hunan, CHINE
19-29.IV.2008
Copyright © Alexandr Safronov

Carabus (Carabus) stscheglowi Mannerheim, 1827
Male adult

Copyright © Kirill Makarov

Carabus (Coptolabrus) lafossei tiantai Kleinfeld, 1997 Holotype
Female adult

Copyright © Frank Kleinfeld

Carabus (Coptolabrus) lafossei pseudocoelestis Kleinfeld, 1999 Holotype
Male adult

Copyright © Frank Kleinfeld

Carabus (Coptolabrus) jankowskii jankowskii Oberthur, 1883
Male adult
N.Korea, Chagang Do, Mt. Langlin, 30.07.2005
Copyright © Anton Menstschikov

Carabus (Coptolabrus) jankowskii benxiensis Imura, 1991
Female adult
CHINA, Jilin prov., Mt. Longgang., 15.07.2005
Copyright © Anton Menstschikov

Carabus (Coptolabrus) jankowskii benxiensis Imura, 1991
Male adult
NE-China,
Liaoning Prov., Donggou, Mt. Guanmen, 15.VI.2009
male, 36mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) formosus grumorum Semenov, 1898
Female adult
female, 30mm
Copyright © Boleslav Brezina

Carabus (Coptolabrus) formosus brockmanni Kleinfeld & Schutze, 1997 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Coptolabrus) formosus sunpanensis Semenov, 1898 (as rostandi Deuve, 1991)
Male adult
China, N Sichuan province, Venchuan, h=1400 m., 30.VI.2007, A. Gorodinski leg.;
L=33 mm
Copyright © Anton Menstschikov

Carabus (Coptolabrus) elysii pulcher Kleinfeld, 1997 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Coptolabrus) elysii magnificens Kleinfeld, 1999 Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Coptolabrus) elysii elysii Thomson, 1856 (as idai Kleinfeld, 2000) Holotype Paratype
Male+Female adult

Copyright © Frank Kleinfeld

Carabus (Coptolabrus) elysii elysii Thomson, 1856
Female adult
China, Sichuan province, Wanyuan; 15.06.2009; Local people leg.;
L=32 mm.
Copyright © Anton Menstschikov

Carabus (Coptolabrus) augustus pustuliferus Kleinfeld, 1998 Holotype
Male adult

Copyright © Frank Kleinfeld

Carabus (Lipaster) stjernvalli osellai Breuning & Ruspoli, 1970
Male+Female adult
NE-Turkey: Artvin:Hopa, Borçka env., h=750 m, 9.VI-7.VII.2011, M.Janalik leg.
female, 42 mm
Copyright © Ruslan Panin

Carabus (Tribax) puschkini ishikawaianus Breuning & Ruspoli, 1970
Male adult
NE-Turkey: Artvin:Hopa, Borçka env., h=450 m, 29.VI-7.VII.2011, M.Janalik leg.
L=33 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens praecellens Pillardi, 1825
Male adult
Serbia, Fruska Gora Mts, 20-31.V.2006, M.Sima leg.
L=29 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) illigeri versicolor Frivaldsky, 1835
Male+Female adult
C Bulgaria:Stara planina Mts, Gabrovo,12-15.VI.2007, M. Janalik leg. female, 31 mm
Copyright © Ruslan Panin

Carabus (Tribax) puschkini ponticus Deyrolle, 1869
Male adult
NE-Turkey:Trabzon, Zigana Pass h=1700m,14-24.VI.2010, M.Janalik leg.
L=28 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi comptus (as szoerenyensis Csiki, 1908)
Male adult
SW Romania, Tarcu Mts, vf. Nedeia h=2000m, 12.VI.2008, J.Kynkor leg.
L=22 mm
Copyright © Ruslan Panin

Carabus (Carabus) arvensis carpathus (as pseudonoricus Kenyery, 1983)
Male adult
Slovenia: Julske Alpy, Podvelka, h=450m, 24.X.2010, M.Janalik leg.
L=19 mm
Copyright © Ruslan Panin

Carabus (Megodontus) bonvouloiri bonvouloiri (as kaschkarensis Blumenthal & Breuning, 1967)
Male adult
NE-Turke,Rize vill., S Ayder h=1740 m,28.VI-6.VII.2011, M.Janalik leg.
L=32 mm
Copyright © Ruslan Panin

Carabus (Acoptolabrus) schrencki schrenck (as dongningensis Deuve & Li, 2005)
Male adult
NE China, Heilongjiang prov., Dongning, V.2009 L=25 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei (as gutiiensis Takács & Lie, 1992)
Male adult
N Romania, E Carpathian, Maramureş Reg., Gutâi Mts, cabana Mogosa,4 km N Baia Sprie, 9-17.VI.2010, J.Kynkor leg.
L=33 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) obsoletus nagyagensis (as paranagyagensis Lie, 1990)
Male adult
W Romania, Arad reg., Zarand Mts (Muntii Zarandului), Nadas-Barzavah h=142 m, J. Kynkor leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) macrogonus macrogonus Chaudoir, 1847
Male adult
NE-Turkey:Trabzon env., Sumela, 17-21.VI.2010, M.Janalik leg.
L=20 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) macrogonus pseudokolatensis Heinz & Korge, 1967
Male adult
NE-Turkey::Kackar Dagi Mt, Ayder S of Ardesen h=1600m, 17-21.VI.2010, M.Janalik leg.
L=20 mm
Copyright © Ruslan Panin

Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931
Male adult
NE-China:Liaoning prov., Fengcheng City, Miaoyang, 12-17.VI.2011
L=26 mm
Copyright © Ruslan Panin

Carabus (Scambocarabus) kruberi laobeiensis Deuve & Li, 2000
Male adult
NE-China: Liaoning prov., Fengcheng City, Miaoyang, 12-17.VI.2011
L=27 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) fabricii fabricii (as feichtauensis Müller & Sach, 1999)
Male adult
NC Austria, E-Alps, Windischgarsten, Mt Hoher Nock, VII.2004, Manfred Egger leg.
L=18 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) fabricii fabricii (as koralpicus Sokolar, 1910)
Female adult
S Austria, Carinthia reg.(Kärnten), Wolfsberg, Koralpe Mt Range,18.VI.2005, Manfred Egger leg.
L=22 mm
Copyright © Ruslan Panin

Carabus (Procrustes) anatolicus anatolicus Chaudoir, 1857
Male adult
Cyprus, Troodos, 17.2.1V.2010,. Manfred Egger leg.
L=27 mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) convexus moreanus Reitter, 1896
Female adult
Greece:, N Peloponnes, Xylokastro, Zemeno,h=1000m, 6.VI.1997, Manfred Egger leg.
L=21 mm
Copyright © Ruslan Panin

Carabus (Procrustes) banoni Dejean, 1829
Male adult
Greece,W Crete Island, Mili gorge ,12.V.2009, Manfred Egger leg.
L=26 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) rugosus rugosus Fabricius, 1792
Male adult
N Morocco, El Biutz –Taghramt,20.I.2005, Manfred Egger leg.
L=29 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) alpestris hoppei Germar, 1824
Male adult
W Austria, Tirol, Innsbruck, Hafelekar Mts h=2200m, Manfred Egger leg.
L=19 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) alpestris alpestris Sturm, 1815
Female adult
C Austria, E-Alps, Windischgarsten, Mt Hoher Nock, VII.2004, Manfred Egger leg.
L=20 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei linnaei Panzer, 1812
Male adult
W-Ukraine, NW podolian,
Lviv Reg., Zhydachiv distr., near Vybranivka, 17.II.2007,Panin R. leg.
female, 21 mm
Copyright © Ruslan Panin

Carabus (Megodontus) croaticus croaticus Dejean, 1826
Male adult
SW Slovenia, Ilirska Bistrica, Sviščaki, Snežnik,17-28.VI.2009, Manfred Egger leg.
L=32 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) morbillosus constantinus Kraatz, 1899
Male adult
NW Tunisia; Tebarka, E Sejenane,26.I.2004, Manfred Egger leg.
L=33 mm
Copyright © Ruslan Panin

Carabus (Megodontus) caelatus schreiberi Kraatz, 1887
Male adult
SW Slovenia, Obrov, Poljane, 14-27.VIII.2010, Manfed Egger leg.
L=33 mm
Copyright © Ruslan Panin

Carabus (Oreocarabus) carinthiacus Sturm, 1815
Male adult
NE-Italy, Bolzano, Brenner, Zirog h=2000 m, 26.VII.1992, Manfred Egger leg.
L=20 mm
Copyright © Ruslan Panin

Carabus (Damaster) blaptoides oxuroides Schaum, 1862
Female adult
Japan, C-Honshu, Ofuse-cho, Kamitakai, Nagano Pref., 12.VI.2006, Ohshima leg.
L=34 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) fabricii fabricii (as poloniensis Géhin, 1876)
Female adult
N-Slovakia, Orava Beskids Mts, Mt Babia Góra h=1720 m, 5.VII.20.VIII.2010, Rudolf Kmeco leg.
L=22 mm
Copyright © Ruslan Panin

Carabus (Chaetocarabus) arcadicus merlini Schaum, 1861
Male adult
S-Greece:Peloponnesos: Taygetos Mt.Rng., 17-23.V.2005, V. Sima leg.
L=30 mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) convexus dilatatus (as perplexus Schaum, 1857)
Male adult
CE-Greece: Larissa reg., Stomio, Ossa Mt., 6-17.VII.2011, Krause Rene leg.
L=19 mm
Copyright © Ruslan Panin

Carabus (Procrustes) coriaceus mopsucrenae Peyron, 1858
Male adult
SC-Turkey, Adana Prov., Doganbeyli, 8.V.2001, Snizhek M. leg.
L=28 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) ulrichi rhilensis (as stussineri Géhin, 1885)
Male adult
W Croatia, Ivanščica Mts , Ivanec, 24.VI-7.VII.2006, Krause Rene leg.
L=29 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) ulrichi ulrichi (as intercessor Sokolar, 1912)
Male adult
N Hungary, Pilis Mts., Piliszentlelek, 20-27.VII.2011, Krause Rene leg.
L=28 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi incompsus Kraatz, 1880
Male+Female adult
C Romania, Brașov County, Cristian vill., h=380 m,V-VI.2008, M.Pasek leg.
female, 27 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi (? incompsus as spectabilis Csiki, 1906)
Female adult
NC Romania, Bistrita Nasaud reg., Călimani Mts, Colibita vill., mixed forest, meadows h=800m, 15-21.V.2011, M.Pasek leg. L=34 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi incompsus (as mendax Csiki, 1906)
Female adult
W Ukraine, NE Carpathians,
Transcarpathian reg., Chornogora Mt Range, h=1100m,13.VI.2011, Panin R. leg./det.
L=33 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi alutensis Savulescu, 1972
Male+Female adult
C Romania, S Carpathians, Rimnicu Vilcea env., Calimaneşti, Cozia Mts., VII.2007, I. Marvan leg.
L=29 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi comptus (as borlovensis Lie, 1994)
Male adult
SW Romania, S Carpathians,
Banat , Tarcu Mts, Borlova env., 19-31.V.2005, P. Sima leg.
L=24 мм
Copyright © Ruslan Panin

Carabus (Lamprostus) robustus ducalis Schaufuss, 1882
Male adult
NE-Turkey:Trabzon, Zigana Pass h=2000m,14-24.VI.2010, V. Slovák leg.
L=32 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) ulrichi arrogans Schaum, 1858
Male adult
NE-Serbia:Majdanpek, 9-11,V.2010, L.Brhel leg.
L=28 mm
Copyright © Ruslan Panin

Carabus (Diocarabus) fraterculus fraterculus Reitter, 1895
Male adult
N-Korea, N Pyongan Prov.,Sakju, Mt Ponsan, 20-29.VI.2009, L. Bieber leg.
L=19 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) varians armeniacus Mannerheim, 1830
Male adult
NE Turkey, Artvin Prov., Şavşat, Çamlıca,18.VI.1993, M. Janata leg. L=22 mm
Copyright © Ruslan Panin

Carabus (Pachystus) hungaricus hungaricus Fabricius, 1792
Male adult
Czech, S Moravia, Pouzdrany, 8,VIII.2005, V. Slovak lgt.
L=27 mm
Copyright © Ruslan Panin

Carabus (Megodontus) planicollis verae Csiki, 1905
Male+Female adult
♂ C-Romania:S Carpathians, Fagaraş Mts, near Avrig Lake, h=2100 m, 5.VI.2005
♀ C-Romania:S Carpathians, Fagaraş Mts, Saratii-Valley h=1700 m, 6.VI.2005
female, 28 мм
Copyright © Ruslan Panin

Carabus (Lamprostus) torosus torosus Frivaldsky, 1835
Male adult
NE-Bulgaria, Dobrich Region, Kavarna env.
L=31 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) obsoletus obsoletus Sturm, 1815 (as prunneri Mallasz, 1901)
Male adult
NW Romania, Apuseni Mts, Trascău Mt Rng, Lungesti vill., 25 km W Turda,.VI.2012
L=26 mm
Copyright © Ruslan Panin

Carabus (Deroplectes) dokhtouroffi dokhtouroffi Ganglbauer, 1886
Male adult
NC Tadzhikistan, Pamir, N-slope Peter I.Mt Rng, h=2100m,Ganishou vill., 10-12.VI.2006, Ivanova E. leg.
L=28 mm
Copyright © Ruslan Panin

Carabus (Megodontus) stroganowi transfuga Semenov, 1898
Male adult
SE-Azerbaijan: Talysh, Kalvaz, 21-30.VI., Dolin V.G. leg.
L=32 mm
Copyright © Ruslan Panin

Carabus (Semnocarabus) erosus kuldshaensis Ballion, 1878
Male adult
S-Kazakhstan:Ili River valley,:near Bakanas,3-5.V.2002
L=21 mm
Copyright © Ruslan Panin

Carabus (Procrustes) talyschensis talyschensis Menetries, 1832
Female adult
Azerbaijan, Talysh Mts., Zuvand, Kalvaz vill., h=1650 m, 15.VII.2003, Dolin V.G. leg.
L=32 mm
Copyright © Ruslan Panin

Carabus (Goniocarabus) grombczewskii grombczewskii Semenov, 1891
Male adult
NC Tadzhikistan, Pamir, N-slope Peter I.Mt Rng, h=2100m, Ganishou vill.,12.VII.2006, Pak O. leg.
L=19 mm
Copyright © Ruslan Panin

Carabus (Ulocarabus) stschurovskii Solsky, 1874
Male adult
Tadzhikistan, Pamir-Alay, S slope Gissar Mt.Range, Takob, h-1700m, 10.V.1996, Dolin V.G. leg.
det. Panin R.Yu.
L=22 mm
Copyright © Ruslan Panin

Calosoma (Charmosta) investigator Illiger, 1798
Female adult

Copyright © Kirill Makarov

Calosoma (Calosoma) inquisitor inquisitor Linne, 1758: 414
Male adult

Copyright © Kirill Makarov

Calosoma (Callisthenes) panderi Fischer, 1822
Male adult
NW Kazakhstan, Aktiubinsk reg., Baiganinsk distr., near Zharly vill. 10-12.05.1997 leg. M.& L. Smirnov (MPU)
Copyright © Kirill Makarov

Carabus (Morphocarabus) rothi hampei (as diffinis Csiki, 1906)
Male adult
NW Romania, Apuseni Mts, Trascău Mt Rng, Petresti de Jos, 12 km E Turda,. 6.VIII.2008
L=27 mm
Copyright © Ruslan Panin

Carabus (Ulocarabus) stschurovskii Solsky, 1874
Female adult
Tadzhikistan, Pamir, N-slope Peter I.Mt Rng,h=2100m,Ganishou vill., Dolin V.G. leg.
det.Panin R.Yu.
L=24 mm
Copyright © Ruslan Panin

Carabus (Procerus) scabrosus caucasicus Adams, 1817
Male adult
N Caucasus,
Stavropol Krai, Pyatigorsk, Mt.Mashuk, VI.2007 Tikhonov Yu. leg.
44mm
Copyright © Ruslan Panin

Carabus (Procerus) gigas duponcheli Dejean, 1831 (as kusdasi Mandl, 1966)
Male adult
S-Greece: Peloponnes, Mt.Chelmos, 20.V.2007.
L=46 mm
Copyright © Ruslan Panin

Carabus (Aulonocarabus) gaschkewitschi czarensis Obydov, 1997
Female adult
Russia, CE-Siberia, N Chita reg., Kodar Mt. Range, Big Leprindo Lake env. h~2000m., 2-5.VII.2002 A. Prokhorov leg./det. A. Anichtchenko
L=19 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus gougeleti Reiche, 186
Male adult
CW Spain, Province Huelva, Arroyomolinos de León,30.X.2011, T. Alcantara leg.; L=23 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) fabricii ucrainicus Lazorko, 1951
Male adult
Ukrainian Carpathians, Ivano-Frankivsk reg., Yaremcha distr., Gorgany Mts., h=1590 m, 22.VII-3.VIII.08, Panin R. leg., male, 23mm
Copyright © Ruslan Panin

Carabus (Platycarabus) fabricii ucrainicus Lazorko, 1951
Male adult
Ukrainian Carpathians, Ivano-Frankivsk reg., Gorgany Mts., h=1590 m, 17.VI-26.VII.2011, Panin R. leg. male, 23mm
Copyright © Ruslan Panin

Carabus (Megodontus) kolbei kolbei Roeschke, 1887
Male adult
N-Japan, C-Hokkaido, Hidaka distr., Niikappu env., 27.VIII.1989, Sasaki leg,
L=27 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) latreillei ? semicoriaceus Kraatz, 1881
Male adult
NE-China: Liaodong Peninsula, Liaoning prov., Gaizhou distr., Mt. Buyun Shan,VI.2007, Hao leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Megodontus) kolbei hidakamontanus Ishikawa, 1966
Male adult
N-Japan, C-Hokkaido, Hidaka Mt.Rng, Mt.Apoidak near Samani town, 20.VIII.1982,T. Akaishi leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Acoptolabrus) constricticollis constricticollis (as grallatorius Roeschke, 1921)
Male adult
NE-Korea:North Hamgyong Prov., Sechon-dong, VII.2004, Li leg.
L=28 mm
Copyright © Ruslan Panin

Carabus (Eurycarabus) famini lucasi Gaubil, 1849
Male adult
N-Tunisia, Bizerte reg., Menzel Bourguiba city,h=5 m, XII.1981, S. Remond leg.
L=25 mm
Copyright © Ruslan Panin

Carabus (Leptocarabus) procerulus Chaudoir, 1862
Male adult
Japan:S Honshu, Matsubara-ko, VIII.1997, S. Remond leg.
L=30 mm
Copyright © Ruslan Panin

Carabus (Ohomopterus) japonicus japonicus Motschulsky, 1857
Male adult
S Japan, Kyushu, Kumamoto Pref., Izumi viil., h=600 m,VII.2001, S. Remond leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Piocarabus) titanus Breuning, 1933
Male adult
C-China Sichuan prov., Nanping, h=1500m, VI.2000, S. Remond leg. L=34 mm
Copyright © Ruslan Panin

Carabus (Eurycarabus) famini fezzanus Lapouge, 1915
Male adult
Morocco, Middle Atlas Mt Rng, Azrou city h=1500m, IV.1999, S. Remond leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Leptocarabus) procerulus Chaudoir, 1862
Male adult
Japan, S Honshu, Mie Pref., Hakusan, VII.2001, S. Remond leg. L=33 mm
Copyright © Ruslan Panin

Carabus (Oreocarabus) taedatus agassi Le Conte, 1850
Male adult
SC-USA, New Mexico, Santa Fe,VIII.1999, Sehnal R. leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Ohomopterus) yamato kitai (Imura, 2004)
Male adult
Japan, S Honshu, Mie Pref., Hakusan, VII.2001, S. Remond leg.
L=21 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
Male adult
N Mongolia, Khövsgöl reg., Lake Khovsgol, h=1640m, 51*:12’N; 100*:45’E, steppificated sites,
20-25.VII.2010. A.Dergalev leg.
L=22 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) fabricii ucrainicus Lazorko, 1951
Female adult
W-UKRAINE, Ivano-Frankivsk reg., Gorgany Mts, Syvula Mt., h =1820 м., 25.VII.2011,Panin R. leg., L=27 mm 
Copyright © Ruslan Panin

Carabus (Carabus) granulatus granulatus Linne, 1758
Male adult
W-Ukraine, Roztochia,
Lviv Reg., Javorov distr., near Ivano-Frankovo vill., Site Zalyvki, 17.IV-3.V.2010, Panin R. leg.
L=20 mm

Copyright © Ruslan Panin

Carabus (Procrustes) piochardi piochardi Gehin, 1883
Male adult
C-Lebanon, Faraya vill., Mt. Djebel Sannin h=1800m, X.2012, C. Reuter leg. L=31 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus islandicus Lindroth, 1968
Female adult
SE-Iceland:Höfn env.:Hornafjördur, 24-28.VIII.2008, C. Reuter leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) pustulifer shangzhouicus Deuve & Li, 2003
Male adult
C-Chine, SE-Shaanxi Province, Shahezi, Shangzhou City, 6-18.VII.2010 L=43 mm
Copyright © Ruslan Panin

Carabus (Pseudocranion) lazikouensis lazikouensis (as kundaensis Deuve, 2003) Paratype
Female adult
C-China, S-Gansu: SSW of Minxian: Qilian Shan: W Kunda, 34:10N/103:47E, h=2800-3000m, forest, 7.VII.2002, Belousov & Kabak leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) buddaicus gansuicus Deuve, 1991
Male adult
China, NC-Sichuan (ZOIGE), LANGMUSI (=DOGCANGLHAMO), 34:03:48N/102:38:08E, h=4100m, alpine meadows, screes, 16-18.VII.1999, B.Brezina leg.
L=20 mm
Copyright © Ruslan Panin

Carabus (Ulocarabus) theanus delerei Mandl, 1955
Male adult
N-Afghanistan, Hindukush Mt.Rng, NE Salang Pass h=2300 m, 35°27'494"N, 68°58'399"E, IV.2011, C. Reuter leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Ulocarabus) theanus theanus Reitter, 1895
Male adult
Tadzhikistan, S slope Gissar Mt.Rng, left bank of the river Sorbo, Haidar gorge,1.VI.1984, Dolin V.G. leg./det.Panin R.Yu.
L=21 mm
Copyright © Ruslan Panin

Carabus (Ulocarabus)? theanus theanus Reitter, 1895
Male adult
NC Tadzhikistan, Pamir, N-slope Peter I.Mt Rng, h=2100m, Ganishou vill., 10-12.VI.2006, Ivanova E. leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Aulonocarabus) canaliculatus hailinensis Deuve & Li, 2000
Male adult
NE China, Heilongjiang Prov., Mudanjiang City, Mt.Tai-Gu-Ding-zi h=800m, 15-25.VI.2010
L=29 mm
Copyright © Ruslan Panin

Carabus (Megodontus) croaticus zepcensis (as bosiljevici Drovenik & Pavićević, 1985)
Male adult
CW-Serbia, Zlatar Mts, Nova Varoš, Zlatar Lake env., 1-12.VII.2007, P.Sima leg. L=30 mm
Copyright © Ruslan Panin

Carabus (Lamprostus) saulcyi saulcyi Piochard, 1875 (as chambouqensis)
Female adult
N-Lebanon, Fnaпdek, Djebel Qammoua, h=1100-1200m, 29.X.2012, C. Reuter leg.
L=27 mm
Copyright © Ruslan Panin

Carabus (Meganebrius) kadoudali (Morvan, 1981)
Male+Female adult
C-Nepal, SW Dhaulagiri, W Jaljala h=3150 m, 28°30'05"N, 83°11'48"E, 23.IX.2012, J. Schmidt leg.
female, 20 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) illigeri versicolor Frivaldsky, 1835 (ab.)
Female adult
C Bulgaria: Stara Planina Mts, Schipka pass h=1450m, 12-15.VI.2007, M.Janalik leg. L=31 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Female adult
N- Romania, E-Carpathians, Rodna Mts, h=2000m, 7-8.VI.2012, Cartuz Z.leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) manfredeggeri Müller 2012 Paratype
Male+Female adult
N-Korea, Chagang Prov., Manpo City, Songhak-ri, 2-16.VII.2011
female, 21 mm
male, 19 mm
Copyright © Ruslan Panin

Carabus (Megodontus) caelatus dalmatinus (as procerus Reitter, 1885)
Male adult
S Bosnia, Potoci env. h=500 m, VII.2006, Bartoloni leg.
L=38 mm
Copyright © Ruslan Panin

Carabus (Imaibius) dardiellus perobscurus Deuve, 1980
Male adult
N India.Kashmir, Loran, W part Pir Panchal range,h=2400-2500m, 33°49'"N, 74°20'E
L=25 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) smaragdinus dachangensis Lu & Müller, 2009
Female adult
NE-China, Liaoning Prov., Changhai County, Dachangshan Dao Isl.,12.VI.2011
L=40 mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) rumelicus syriensis Breuning, 1943
Male adult
N Lebanon, Fnaпdek, Djebel Qammoua, h=1100-1200m, C. Reuter leg.
L=17.5 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) illigeri illigeri (as tarensis Toševski, 1990)
Male adult
WC-Serbia: National park Tara Mts.:NE Mitrovac h=1500m, 1-12.VII.2007, Pr.Sima leg. L=30 mm
Copyright © Ruslan Panin

Carabus (Cupreocarabus) huangi huangi Deuve, 1992
Male adult
C-China, E Tibet, N-Sichuan, Hongyuan,valley 10 km SE Sanggarpar, h=4200m, 32.300053, 102.584031, alpine meadow / scree, 19.VII.1995, B.Brezina leg. L=21 mm
Copyright © Ruslan Panin

Carabus (Acoptolabrus) mirabilissimus rigoutianus Deuve & Li, 1998
Male adult
N-Korea, South Pyongan prov., Mt. Maeng-san, 6.VI.2010
L=25 mm
Copyright © Ruslan Panin

Carabus (Pseudocranion) taibaishanicus taibaishanicus Deuve, 1989
Female adult
China, W-Shaanxi, Taibai Shan Mts, btw.Houzhenzi/Ba Xian Tai, nr. Nan Tian Men h=3200m,
33°55'17"N/ 107°46'53"E, 25-30.VII.2011, Li Wuyang leg.
L=29 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus dryas Gistl, 1857
Male adult
NE-Greece, Mt. Olympus, Litochoro env., 20-30.V.2005, Sima leg.
L=27 mm
Copyright © Ruslan Panin

Carabus (Lamprostus) saulcyi saulcyi Piochard, 1875
Male adult
C Lebanon, Jbeil Distr., Ehmej vill., h=1100 m.V.2012, C. Reuter leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Imaibius) dardiellus ? subpunctulus Roeschke, 1907
Male adult
N India, Kashmir, Pahalgam, Aru Valley h=2400m, VII.2007
L=25 mm
Copyright © Ruslan Panin

Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847
Male adult
WC Spain, Cáceres, Hervás, h=1000m, V.2007, Pascal Stefani leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (?Oreocarabus) qinlingensis Imura, 1993
Male adult
China, W-Shaanxi, Taibai Shan Mts, btw. Houzhenzi/Ba Xian Tai, nr.Nan Tian Men h=3200m, 33°55'17"N/107°46'53"E, 25-30.VII.2011, Li Wuyang leg.
L=19 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi hampei (as hunyadensis Csiki, 1926)
Male adult
CW Romania, Apuseni Mts, Metalliferous Mt Rng, Hunedoara reg., near Săcărâmb vill., 45°97'472"N/23°02'630"E, h=680 m, mixed forest, meadows, 15-20.V.2011, B.Brezina leg.
L=36 mm
Copyright © Ruslan Panin

Carabus (Meganebrius) pseudoharmandi Mandl, 1965
Female adult
E-Nepal, Kosi zone, Khandbari,(1.5km SW of Chichila vill., 27*45’843’’N/ 87*22’473’E, h=1940m), damaged primary forest, 13 .V.2010, B.Brezina & M.Häckel leg.
L=23 mm

Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens ( as jasilkowskii Born,1905)
Male adult
NE Romania, 5 km W Suceava, 47°63′940″N 26°18′795″E,h=340m, agricultural field, 15-20.V.2011, B. Brezina leg. L=24 mm
Copyright © Ruslan Panin

Carabus (Meganebrius) arunensis Heinertz, 1980
Male adult
E-Nepal, Kosi zone, Khandbari, 1.5km SW of Chichila vill., 27*47’175’’N/ 87*23’622’E, h=1880m, damaged primary forest, 11.V.2010, B.Brezina & M.Häckel leg.
L=27 mm

Copyright © Ruslan Panin

Carabus (Ophiocarabus) ballionis Kraatz, 1879
Male adult
NW-China, Xinjiang prov., 40 km N Narat, h=1500, 12-12.VI.2004, Benes & Kaiser leg.
L=22 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) smaragdinus liaodongensis (Li, 1992)
Male adult
NE-China, E-Liaoning: Anshan: Mt.Qian Shan, 9-14.VI.2011. L=39 mm
Copyright © Ruslan Panin

Carabus (Oreocarabus) latro minshanensis Deuve, 1987
Male adult
China, N-Sichuan, Jiuzhaigou env., 13.VI.2001,E. Kučera leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Lamprostus) syrus pseudopinguis Heinz & Staven, 2000
Male adult
NW-Jordan, Ajlun env., oak forests on dry limestone, h=897m, 32°21′33.1″N/ 035°43’47.4″E, 4.V.2010, Schnitter/Schellhorn leg.
L=26 mm
Copyright © Ruslan Panin

Carabus (Meganebrius) quinlani smetanai Häckel, 2011 Paratype
Male+Female adult
C-Nepal:Dhaulagiri Massif:Baglung Distr., (1.5km W of Okhale vill., 28.32’152’’N/ 83.48’673’E, h=2600m) damaged primary forest, 19.V.2010, B.Brezina & M.Häckel leg.
male, 21mm
female, 23 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus vlasuljensis Apfelbeck, 1894
Male adult
WC-Serbia, Borak - Gornje Babine, 1-10.VII.2007, M. Sima leg.
L=28 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) indigestus confucius Breuning, 1933
Female adult
C-China, E Tibet,W-Sichuan prov., Kangding, Zheduo Pass h=4300m 30.VI.2005 , V.Benes leg,
L=17 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) ladygini degensis Deuve, 1989
Male adult
C-China, NW- Sichuan prov., Garzê Tibetan AP, Pass Garzê – Sêrtar, 19.VII.2007, V.Benes leg.
L=17 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus azurescens Dejean, 1826 (loc.typ.)
Male adult
W-Croatia, Velebit Mts., betw.Karlobag & Brusane vill., VI.2002, Manfred Egger leg.
L=30 mm
Copyright © Ruslan Panin

Carabus (Apoplesius) pseudodepressus Machard, 1988
Female adult
Turkey, Prov. Bitlis, S near Tatvan, 23.IV.2009 leg. M. & S. Nabozhenko
L=19.0 mm
Copyright © Kirill Makarov

Carabus (Megodontus) caelatus caelatus (as malissorum Apfelbeck, 1919)
Male adult
Montenegro, National park Durmitor; near Zabljak vill., Mt. Sedlo h=2200 m, 6-22.VIII.2010, Franko Sandel leg.
29 мм
Copyright © Ruslan Panin

Carabus (Rhigocarabus) buddaicus linxiaicus Deuve, 1995
Male adult
China, S-Gansu (LABRANG):, valley.E of Ponggartang, 35:12-13N/102:54-55E, h=3000-3700m, alpine meadows, coniferous forests,10.-14.VII.1999, B.Brezina leg.
L=20 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus germarii Sturm, 1815
Male adult
N Italy, Trent, Ampola, Mt. Tremalzo h=1700m, 3-13.VI.2010, Manfred Egger leg.
29 mm
Copyright © Ruslan Panin

Calosoma (Carabomorphus) brachycerum (Gersteacker, 1884)
Male adult
N Tanzania, 105 km W of Arusha and 5 km S of Kwa Kuchinga vill. 3°41'S 25°59' E 31.VIII-2.IX.2006 leg. A.Sokolov
L=23 mm
Copyright © Kirill Makarov

Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
Male adult
W-UKRAINE, Transcarpathian region, Rakhiv distr., Chornogora Mt Range, h=800 m, near Bogdan vill., 10.VI.2012, Panin R. leg.
L= 33 мм
Copyright © Ruslan Panin

Carabus (Cupreocarabus) lixianensis lixianensis Deuve, 1990
Male adult
China, E Tibet, N-Sichuan, (Barkam),valley SW of Barkam, h=3200m, 31*53’ N; 102*12’E, mixed/coniferous forest, 22.VII.1995, K.& B.Brezina
L=23 mm
Copyright © Ruslan Panin

Carabus (Tachypus) cristoforii Spence, 1823
Female adult
S France, Pyrénés Mts., Tarascon; 15mm
Copyright © Aleš Sedláček

Calosoma (Callisthenes) breviusculus Mannerheim, 1830
Male adult
Turkey, Buglan; 21mm
Copyright © Aleš Sedláček

Calosoma (Microcallisthenes) relictum Apfelbeck, 1918
Male adult
Macedonia, Bistra planina Mts.; 13mm
Copyright © Aleš Sedláček

Calosoma (Microcallisthenes) relictum Apfelbeck, 1918
Female adult
Macedonia, Bistra planina Mts.; 13mm
Copyright © Aleš Sedláček

Calosoma (Microcallisthenes) relictum Apfelbeck, 1918
Male adult
Macedonia, Bistra planina Mts.; 14mm
Copyright © Aleš Sedláček

Calosoma (Calosoma) sycophanta Linne, 1758
Male adult
CZ; S Moravia, Břeclav; 26mm
Copyright © Aleš Sedláček

Carabus (Megodontus) bonvouloiri graciliformis Breuning, 1964
Female adult
NC Turkey, Gölköy env., Harcbeli pass, 37mm
Copyright © Aleš Sedláček

Carabus (Pachystus) cribratus staneki Sterba, 1931
Male adult
NC Turkey, Küre Mts., 60km N Kastamonu; 25mm
Copyright © Aleš Sedláček

Carabus (Pachystus) cribratus staneki Sterba, 1931
Female adult
NC Turkey, Küre Mts., 60km N Kastamonu; 27mm
Copyright © Aleš Sedláček

Carabus (Tachypus) auratus honnoratii Dejean, 1826
Male adult
SE France, Carpentras; 23mm
Copyright © Aleš Sedláček

Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906 loc.typ.
Male+Female adult
W-UKRAINE, Transcarpathian region, Uzhgorod distr., h=250 m, 8.VII-25.VIII.2011 Panin R. leg./det. female, 33 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906 loc.typ.
Male+Female adult
W-UKRAINE, Transcarpathian region, Uzhgorod distr., 28.IV-23.V.2011 Panin R. leg./det. female, 33 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Male adult
SE Slovakia, Slanske vrchy Mts, Herlianske sedlo, 8-19.VI.2009, Krause Rene leg.
L=30 mm
det.Panin R.
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Female adult
NW Ukrainian Carpathians,
Lviv Region, Staryi Sambor Distr., near Dobromyl , h=450m, 9.VI-25.VII.07, Panin R. leg./det.Solodovnikov I. & Panin R.
female, 35mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Female adult
NW Ukrainian Carpathians
Lviv Region, Staryi Sambor Distr., near Dobromyl , h=500m 9.VI-25.VII.2007, Panin R. leg./det.Solodovnikov I. & Panin R.
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii zawadzkii, Kraatz 1854
Male adult
W.Ukraine, Ivano-Frankivsk reg.,
Yaremche distr., near Tatariv vill., Gorgany Mts. h=700 m, 16.VI.2011, Panin R. leg.
L=31 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii zawadzkii, Kraatz 1854
Male+Female adult
W.UKRAINE, Ukrainian Carpathians
Lviv Reg., Skole distr., Skolivski Beskidy Mts. Zelemyanka Mt. h=1000m. 9-20.VII.07 Panin R. leg.
female, 35mm.
Copyright © Ruslan Panin

Carabus (Ophiocarabus) successor koeksuensis Kabak, 2013 Paratype
Male adult
China, Xinjiang, Koeksu Bas., Tshon-Aidak, SSE Tekes, 42* 32' 47'' N / 82* 05' 56'' E -- 42* 32' 51'' N / 82* 05' 36'' E, 3330-3550 m, 11.07.2012, Kabak I.I. leg.
Copyright © Ilya Kabak

Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Male adult
NE Serbia, Majdanpek; male, 33mm
Copyright © Aleš Sedláček

Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Female adult
NE Serbia, Kučevo; female, 33mm
Copyright © Aleš Sedláček

Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Female adult
NE Serbia, Majdanpek; 35mm
Copyright © Aleš Sedláček

Carabus (Morphocarabus) helleri Ganglbauer, 1893
Male adult
CZ; CE Moravia, Bílé Karpaty Mts.; 24mm
Copyright © Aleš Sedláček

Carabus (Morphocarabus) helleri Ganglbauer, 1893
Male adult
CZ; CE Moravia, Bílé Karpaty Mts.; 25mm
Copyright © Aleš Sedláček

Carabus (Heterocarabus) marietti marietti Cristoforis & Jan, 1837
Female adult
as ativsengueni Schweiger, 1962
NW Turkey, Bolu, Abant Mts.; 16mm

Copyright © Aleš Sedláček

Carabus (Nesaeocarabus) abbreviatus Brulle, 1835
Male adult
Canarias Isle, NC Tenerife; 22mm
Copyright © Aleš Sedláček

Carabus (Carabus) arvensis venetianus Bernau, 1914
Male adult
N Italy, Veneto, Dolomiten Mts.; 13mm
Copyright © Aleš Sedláček

Carabus (Carabus) arvensis venetianus Bernau, 1914
Female adult
N Italy, Veneto, Dolomiten Mts.; 15mm
Copyright © Aleš Sedláček

Carabus (Carabus) arvensis arvensis Herbst, 1784 (as noricus Sokolar, 1910)
Female adult
CZ, C Moravia, Dřevohostice - Líšná, 13mm
Copyright © Aleš Sedláček

Carabus (Procrustes) banoni Dejean, 1829
Male adult
Crete, Agia Irini (Sélinos); 31mm
Copyright © Aleš Sedláček

Carabus (Macrothorax) morbillosus macilentus Lapouge, 1899
Female adult
SE Spain, Murcia reg., P.N. de Calblanque, 1.V.2003, T.Gazurek leg.
L=34 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) ? lafossei coelestis Stewart, 1845
Female adult
CE-China, Shangai, 3.V.2007 L=34 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) cantabricus cantabricus Chevrolat, 1840 (loc. typ.)
Male adult
NW-Spain, Pontevedra prov., A Cañiza viil., Alto de Fontefría h=800m, 31.XII.2007, T.Gazurek leg./det.Panin R. L=27 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) baguenai Breuning, 1926
Male adult
SE Spain, Almería Province, Carboneras env., 25.XI.2006, T.Gazurek leg./det. R.Panin L=26 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) rugosus celtibericus Germar, 1824
Female adult
SW-Portugal, Setúbal Distr., Cercal do Alentejo/ Tanganheira vill. 8.XII.2006, T.Gazurek leg.
L=31 m
Copyright © Ruslan Panin

Carabus (Archicarabus) steuartii Deyrolle, 1852
Female adult
NW-Spain, Pontevedra prov., Serra do Candán Mt range, 1.III.1992, S. Devesa leg./det. S. Sobrino
Copyright © Ruslan Panin

Carabus (Ctenocarabus) galicianus Gory, 1839
Female adult
NW-Spain, Pontevedra prov., Campo Lameiro env., 15.VIII.1994, S.Sobrino leg.
L=28 mm
Copyright © Ruslan Panin

Carabus (Ctenocarabus) melancholicus costatus Germar, 1824
Female adult
C-Spain, Madrid region, Rozas de Puerto Real env.,21.XII.2008, T.Gazurek leg.
L=27 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) creutzeri kircheri Germar, 1824
Male adult
S Austria, Karnische Alpen; male, 24mm
Copyright © Aleš Sedláček

Carabus (Platycarabus) creutzeri kircheri Germar, 1824
Female adult
S Austria, Karnische Alpen; 25mm
Copyright © Aleš Sedláček

Carabus (Mesocarabus) macrocephalus Dejean, 1826
Male adult
NW-Spain,Cantabrian Mts, Picos de Europa Mt Rng, Puerto de Panderruedas h=1460 m, 25.IX.2004,T.Gazurek leg.
L=28 mm
Copyright © Ruslan Panin

Carabus (Carabus) arvensis deyrollei Gory, 1839
Male adult
NW-Spain,
Cantabria, Udías env., Monte Corona h=100m,14.VI.2002, Javier de Diego leg.
L=16 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) alpestris adamellicola Ganglbauer, 1904
Male adult
N-Italy, Brescia prov., Bagolino, Adamello Mts, Passo del Maniva env., h=2000m, VIII.2011, M.Monzini leg.
L=19 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) latus cf. molossoides Lassalle, 1984
Male adult
C-Spain, Toledo, Mts de Toledo, Sierra del Castañar Rng, Puerto del Milagro env, h=920m, 7.II.2009, T.Gazurek leg./det.R. Panin.
L=24 mm
Copyright © Ruslan Panin

Carabus (Pachystus) cavernosus cavernosus Frivaldsky, 1837
Female adult
W Bulgaria, Cerovo, 31 mm
Copyright © Aleš Sedláček

Carabus (Oreocarabus) carinthiacus Sturm, 1815
Female adult
NW Slovenia, Trnovski Gozd; 24 mm
Copyright © Aleš Sedláček

Carabus (Orinocarabus) fairmairei fairmairei Thomson, 1875
Male adult
N Italy, Alpi Graie; 20 mm
Copyright © Aleš Sedláček

Carabus (Carabus) granulatus granulatus (as hibernicus Lindroth, 1956)
Female adult
W-Ireland, County Mayo, Letterneevoge (R312),16-30.VI.2013, J.Stypula leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) fairmairei heteromorphus Daniel, 1896
Female adult
NW-Italy, Aosta region, Champorcher env., Mt.Dondenaz, VIII.2008, V.Monzini leg.
L=19 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) morbillosus constantinus (as galloprovincialis Lapouge, 1910)
Female adult
SE-France, Var, near Le Muy, VIII.1980, Michon leg.
L=31 mm
Copyright © Ruslan Panin

Carabus (Pachystus) hungaricus scythus Motschulsky, 1847 (loc.typ.)
Male adult
S-Ukraine, near Dnipropetrovsk, 6-15.IX.2013, A. Sukhenko leg.
L=28 mm
Copyright © Ruslan Panin

Carabus (Acoptolabrus) gehinii gehinii Fairmaire, 1886
Female adult
N-Japan, SW-Hokkaido, Chitose env.,VI.2011.
L=34 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) planatus Chaudoir, 1843
Male adult
S Italy, N Sicily, Nebrodi; 33 mm
Copyright © Aleš Sedláček

Carabus (Orinocarabus) fairmairei fenestrellanus Beuthin, 1899
Male adult
NW-Italy, Piemonte prov., Cottian Alps, Mt Albergian h=2450 m.
L=20 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) morbillosus morbillosus Fabricius, 1792 (as cheminorum Deuve, 1988)
Female adult
SE-France, Var, Fréjus env., 26.II.1997,
L=31 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus harcyniae (as procedens Csiki, 1927)
Male adult
W-Ireland, County Mayo, Clare Isl., 17-28.VI.2002, S. McCormack leg. L=25 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus harcyniae (as procedens Csiki, 1927)
Female adult
N-Ireland, Fermanagh Distr., Lough Navar Forest, 17.VII.2013, D.Novak & J.Stypula leg. L=27mm
Copyright © Ruslan Panin

Carabus (Limnocarabus) clathratus clathratus (as jansoni Kraatz, 1890)
Male adult
W-Ireland, County Mayo, Bellacorick bog, 9.VII.2013, J.Stypula leg. L=27 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus purpurascens Fabricius, 1787 (as osismius Lapouge, 1922)
Male adult
W-FRANCE, Brittany , Finistère, Botvarec Forest, 19.XI.2006, Pascal Stefani leg.
L=29 mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) convexus moreanus Reitter, 1896
Male adult
S Greece, Taygetos Mts.; 13mm
Copyright © Aleš Sedláček

Carabus (Tomocarabus) convexus dilatatus Dejean, 1826
Male adult
NE Serbia, Majdanpek; 13mm
Copyright © Aleš Sedláček

Carabus (Tomocarabus) convexus gracilior Gehin, 1885
Female adult
SW Romania, Banat; 15mm
Copyright © Aleš Sedláček

Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825
Male adult
SW Romania, Banat - Eibenthal; 29 mm
Copyright © Aleš Sedláček

Carabus (Eucarabus) ulrichi fastuosus Palliardi, 1825
Female adult
SW Romania, Banat - Eibenthal; 33 mm
Copyright © Aleš Sedláček

Carabus (Eucarabus) ulrichi arrogans Schaum, 1858
Male adult
NE Serbia, Majdanpek; 23 mm
Copyright © Aleš Sedláček

Carabus (Eucarabus) ulrichi arrogans Schaum, 1858
Female adult
NE Serbia, Majdanpek; 29 mm
Copyright © Aleš Sedláček

Carabus (Semnocarabus) carbonicolor callosus Semenov, 1896
Male+Female adult
KAZAKHSTAN,
Bolshoe Almaatinskoe Lake,30.VI.1992/ det.Panin R.
female 19mm
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus durus Reitter, 1896 (as carpathicus Kolbe, 1913)
Male+Female adult
Ukrainian Carpathians,
Ivano-Frankivsk reg., Gorgany Mts., 20km SW Nadvorna, 22.VI-3.VIII.20Panin R. leg.
female, 26mm
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus durus Reitter, 1896 (as ungensis Csiki, 1905)
Male+Female adult
W Ukraine,
Transcarpathian reg., Vinogradov distr., near Malaya Kopanya vill.,7-18.VII.2008, Panin R. leg.
female, 32mm.
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus durus Reitter, 1896 (? hybrid)
Female adult
W. UKRAINE, NE Carpathians,
Marmarosh Mts h=450m, Rakhiv distr., near Dilove vill.,19-23.VII.2010, Panin R. leg.
L=29 mm
Copyright © Ruslan Panin

Carabus (Aulonocarabus) truncaticollis kodaricola Deuve, 1998
Female adult
SE-Siberia:N-Chita reg., Kodar Mt.Rng.,h=1800m, Leprindo vill., 1.VII.2006,
L=17 mm
Copyright © Ruslan Panin

Carabus (Cratocephalus) solskyi Ballion, 1878
Female adult
SE-KAZAKHSTAN, Ketmen Mt Rng, h=1800m, Tujuk vill.,15.V.2004
L=36 mm
Copyright © Ruslan Panin

Carabus (Cratocechenus) ovtchinnikovi ovtchinnikovi Gottwald, 1987
Male adult
N-Kyrgyzstan, Kyungey Ala-Too Mt Rng, Cholpon-Ata env., h=2500m, 19.VI.2000,
L=19 mm
Copyright © Ruslan Panin

Carabus (Semnocarabus) carbonicolor korolkowi Semenov, 1896
Male adult
NC-Kyrgyzstan, Kyrgyz Mt.Rng , 10 km SW of Bishkek, Chon-Aryk env, h=1100m, 24.V.1998, /det. Panin R.
22 mm

Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens jucundus Csiki, 1906
Male adult
N Hungary, Pilis Mts., Piliszentlelek, 20-27.VII.2011, Krause Rene leg./det. R.Panin
L=30 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as pseudojucundus Retezár, 1974)
Male adult
NW-Hungary::Komarom distr., Tardos, Gerecse Mts., 20.-27.V.2011, Krause Rene leg.
L=26 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as vertesensis Retezár, 1974)
Male adult
NW Hungary: Fejer: Vértes Mt Rng, Tatabanya, 28.VII-2.VIII.1996
L=29 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) augustus tenuiorithorax Deuve & Li, 2006
Male+Female adult
SE-China, SW-Jiangxi, Mt Guan, V.2009, local coll. female, 47 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) formosus bousqueti Deuve, 1993
Male adult
C-China, S-Gansu prov., Wenxian County, Mt.Guanzigou,1-14.V.2010, local coll. leg.
L=32 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens (as moldaviensis Born, 1903)
Female adult
E-Romania, Vaslui reg.,15 km SE Vaslui, Crasna env., V-VI,2013, M,Pasek leg. L=26 mm
Copyright © Ruslan Panin

Carabus (Acoptolabrus) constricticollis pihyonensis Deuve & Li, 1998
Male adult
N-Korea, North Pyongan Prov., Tongnim County, 5-25.V.2010 29 mm
Copyright © Ruslan Panin

Carabus (Acoptolabrus) ? leechi huisaensis Deuve & Li, 1999
Male adult
NC-Korea, Ryanggang Prov., Puksubaek-san, 4-10.V.2010
28 mm

Copyright © Ruslan Panin

Carabus (Carabus) arvensis hokkaidoensis Lapouge, 1924
Female adult

Copyright © Kirill Makarov

Carabus (Tachypus) vagans Olivier, 1795
Female adult
SE-France, Var: Le Muy env., 30.VII.2008., A. Paulian leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Eurycarabus) famini piraticus (as mamorensis (Kocher, 1964)
Female adult
NW-Morocco, betw.Meknes/Kenitra, Foret de Mamora, II.2008
L= 27 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) lafossei saturatus Hauser, 1913
Male adult
SE-China, Jiangxi prov.,Yushan County, Mt Wangjing, VI.2012
40 мм
Copyright © Ruslan Panin

Carabus (Chrysocarabus) lineatus lineatus (as pseudonumancianus Mollard, 2010)
Male adult
NC-Spain, Sierra de Cantabria , Sierra de Toloño Rng, Puerto de Herrera , h=1100m, 19.I,2007 25 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) pustulifer (as cyanelongatus Deuve, 2006)
Male adult
C-Chine, Tibet, Qamdo, Gonjo County, Ke Ri Xiang env., VI.2012, Alain Mollard leg. L=43 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus helluo (as albarracinus Ganglbauer, 1886)
Male adult
CE-Spain, Provincia Teruel, Sierra de Javalón Rng, El Cañigral env., h=1460m, X.2010, A. Mollard leg. L=23 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826
Male adult
CW-Spain: Salamanca, Brincones env., h=750m, X.2003 L=25 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) davidis davidis Deyrolle & Fairmaire, 1878
Male adult
SE-China, Jiangxi prov., Yushan County, Mt Wangjing, VI,2012
L=38 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus harcyniae (as belgicus Lapouge, 1913)
Male adult
Luxembourg, Bollendorf-Pont env., 3.III.1999 /det.R Panin
L=24 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) (? problematicus trans lusitanicus) planiusculus (as xaxarsi Born, 1917)
Male adult
NE Spain, Catalonia, Girona Prov., San Hilario Sacalm env., 20.II.1988, A. Mollard leg. L=25 m
Copyright © Ruslan Panin

Carabus (Orinocarabus) bertolinii Kraatz, 1878
Male adult
NE-Italy, Dolomites-Alps, Trentino prov., Rolle Pass env., Malga Fosse h=2000m, 8-28.VI.2013, Manfred Egger leg.
L=19 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) formosus brockmanni Kleinfeld & Schutze, 1997
Female adult
C-China, S-Shaanxi: Shangzhou City, Shahezi,1-14.VI.,2013, local coll. leg.
L=39 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) smaragdinus dolichognathus Deuve, 1995
Male adult
NW-Korea, North Pyongan Province, Mt. Donggorim, 1-20.VI.2013, local coll. leg. L=37 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) augustus augustus Bates, 1888
Male adult
SE-China, Jiangxi prov., Yudu County, Ganzhou city, Yun Fengzang h=1000m, 5-20.V.2011
L=43 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) alpestris dolomitanus Mandl, 1956
Male adult
NE-Italy, South Tyrol, Dolomites-Alps, 500 m left of Würzjoch, Gömaalm - at the foot Mt.Peitlerkofel h=2100m, 28.VII-17.VIII.2013, Manfred Egger leg.
l=18 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) monticola roccae Born, 1923
Male adult
NW-Italy, Liguria:Savona env., Monte San Giorgio, 30.X.1977, Monauzzi leg.
L=23 mm

Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei linnaei (as scopolii Dejean, 1826)
Female adult
W Ukraine, NW-Podolian, near Lviv city, Vynnykivskyi forest, 20.IV.-13.V.2011, R. Panin leg.
L=22 mm
Copyright © Ruslan Panin

Carabus (Tribax) circassicus circassicus (as tshchaltensis Novotný & Voříšek
Male adult
W-Caucasus: Abkhasia, Gudauta distr., E slope Mt Anchkho h=2200-2350 m, 14-20.VII.2011, S.Pavluchuk, I. Solodovnikov leg./det. I. A. Solodovnikov.
25 mm

Copyright © Ruslan Panin

Carabus (Nesaeocarabus) faustus cabrerai Enderlein, 1929
Male adult
Spain, Canarias Isl.,Teneriffe: NW-part:Teno Massif, Erjos env., h=1000m, 18-21.III.2013, V.Slovak leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi rothi (as equistriatus Kraatz, 1878)
Female adult
C Romania, Sibiu reg., left bank Olt River, Avrig env., 12.VI.2013, V. Slovak leg.
L=30 mm
Copyright © Ruslan Panin

Carabus (Aulonocarabus) kurilensis ? daisetsuzanus Kono, 1936
Male adult
N-Japan, Hokkaido, Katō District,Kamishihoro, Horoka near Maruyama Bridge,19.VI.2013, T. Hirabayashi leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Tribax) agnatus pseudagnatus Novotný & Voříšek, 1988
Female adult
NW-Caucasus, Karachay-Cherkessia, Bol'shaya Marka River h=2500-2600m, 12.VIII.2010, I.A. Solodovnikov leg./det.
Copyright © Ruslan Panin

Carabus (Tribax) agnatus pseudagnatus Novotný & Voříšek, 1988
Male adult
W-Caucasus: Abkhasia, Gudauta distr., Mt Anchkho h=2200-2350 m, 14-20.VII.2011, S.Pavluchuk, I. Solodovnikov leg./det. I. A. Solodovnikov.
L=25 mm
Copyright © Ruslan Panin

Carabus (Tribax) agnatus agnatus Ganglbauer, 1889
Male adult

Copyright © Kirill Makarov

Carabus (Neoplectes) edithae exedithae Gottwald, 1985
Male adult
NС-Caucasus, Karachaevo-Cherkessia, Abishir-Akhuba Mt.Range h=2250-2600 m, right bank Pkhiya river, 10-13.VI.2013, S. Pavluchuk leg.
L=25 mm
Copyright © Ruslan Panin

Carabus (Oreocarabus) taedatus bicanaliceps Casey, 1920
Male adult
NW-USA, Washington, Thurston County, Mima Mound Prairie, 15.IX.2005, A. Laforgue leg. L=23 мм
Copyright © Ruslan Panin

Carabus (Eucarabus) catenulatus fontanellae Reitter, 1896
Male adult
W-Croatia, Cres island,VIII.1994, G. Zannna leg. L=33 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) cartereti omisanensis Lassalle, 1999
Male adult
South Korea, Gangwon Province, Kyebang-san, h=1100m, VI.1998 L=24 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) separatus persa Lapouge, 1924
Male adult
N-Iran: Mazandaran Province, Elborz Mt.Rng, Vandarbon env., 10.VII.2002 L=23 mm
Copyright © Ruslan Panin

Carabus (Megodontus) caelatus dalmatinus Duftschmid, 1812
Female adult
Bosnia, Mostar; 40 mm
Copyright © Aleš Sedláček

Calosoma (Ctenosta) guineense Imhoff, 1843
Male adult
Rep. Centre Africaine: Bangui env. IV.1974 28 mm
Copyright © Ruslan Panin

Calosoma (Calosoma) frigidum Kirby, 1837: 19
Female adult
SC-Canada, Manitoba prov., Winnipeg, King’s Park, 23.VI.2008, Todd Lawton leg. L=24 mm
Copyright © Ruslan Panin

Calosoma (Campalita) chinense Kirby, 1819: 379
Female adult
North Korea, Ryanggang Province, Mt P'urun-bong, 24.VIII.1994 L=33 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823
Male adult
W-Ukraine, Podolian Tovtry, Ternopol reg., Pidvolochysk distr., between Chagari-Zbarazhki & Shevchenkove vill., steppe sites, 6.V-7.VI.2012, Panin R. leg. L=25 mm
Copyright © Ruslan Panin

Carabus (Aulonocarabus) gaschkewitschi gaschkewitschi Motschulsky, 1859
Male adult

Copyright © Kirill Makarov

Carabus (Carabus) granulatus yezoensis Bates, 1883
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Carabus (Carabus) menetriesi menetriesi Hummel, 1827
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Carabus (Diocarabus) aurocinctus Motschulsky, 1846
Female adult
det. K. Makarov
Copyright © Alexander Anichtchenko

Carabus (Diocarabus) opaculus opaculus Putzeys, 1875
Female adult

Copyright © Kirill Makarov

Carabus (Leptocarabus) arboreus pararboreus (Ishikawa, 1992)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Carabus (Morphocarabus) henningi henningi Fischer, 1817
Male adult

Copyright © Kirill Makarov

Carabus (Morphocarabus) venustus venustus Morawitz, 1862
Male adult

Copyright © Kirill Makarov

Carabus (Chrysocarabus) auronitens escheri Palliardi, 1825 (as laevipennis Seidlitz, 1891)
Male adult
W Ukraine, NE Carpathians,Transcarpathian Region, Rakhiv Distr., Chornogora Mts Range, h=2000 m, alpin zone, V-28.VI..2014, Panin R. leg.
Copyright © Ruslan Panin

Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Female adult
W Ukraine, NE Carpathians, Transcarpathian Region, Chornogora Mt Range, h=2000 m, alpin zone, 28.VII-11.IX.2014, Panin R. leg. L=22 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) reitteri reitteri Retowski, 1885
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Carabus (Procrustes) paiafa acuticollis Motschulsky, 1858
Male adult
SC-Turkey: Alania (Irmasan), rte 695 au km 25 Akseki, h= 1225 m, 02.X.2003
Copyright © Igor Solodovnikov

Carabus (Imaibius) wittmerorum Heinertz, 1978
Male adult
N. Pakistan: Frontier prov., Miandam, Swat vall., h=1800-2600 m, leg.S.Remond (TOPOTYPUS)
Copyright © Igor Solodovnikov

Carabus (Imaibius) barysomus hazarensis Deuve, 1982 (TOPOTYP)
Male adult
N-Pakistan: Mansehra prov., Hazara, Kaghan valley: Shocgran env., h = 2750 m L= 42 mm
Copyright © Igor Solodovnikov

Carabus (Damaster) blaptoides capito Lewis, 1881
Male adult
Japan: Niigato pref., Sado Isl., Sado, Yamato
Copyright © Igor Solodovnikov

Carabus (Cryptocarabus) ? turkestanus turkestanus Breuning, 1929
Male adult
Kyrgyzstan, W Kungei Ala-Too Mt Rng, Chon-Kemin riv., Kyzyl-Bairak vill., 25.IV.2010. L= 18 mm
Copyright © Ruslan Panin

Carabus (Aulonocarabus) kolymensis kodarensis Brinev, 1997
Male adult
Russia, SE Siberia, N Chita reg., Kodar Mt. Rng. h=1800m, Novaya Chara vill., 23.VI.2006. L=17 mm
Copyright © Ruslan Panin

Carabus (Mimocarabus) pumilio Kuster, 1846
Male adult
S-Armenia, Vayots Dzor Region, E Ughedzor vill., h=2630 m, 9.VI.2014, L=16 mm
Copyright © Ruslan Panin

Carabus (Pachycarabus) imitator imitator Reitter, 1883
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Carabus (Neoplectes) satyrus satyrus Kurnakov, 1962
Male+Female adult
Abkhazia, Gulripsh Distr., NW slopes of Gurzul Mt. Range near Merkheul Village, 170-230 m, left tributary of Machara River, hornbeam, alder, rhododendron (Rhododendron ponticum) forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Neoplectes) satyrus satyrus Kurnakov, 1962
Male adult
Abkhazia, Gulripsh Distr., NW slopes of Gurzul Mt. Range near Merkheul Village, 170-230 m, left tributary of Machara River, hornbeam, alder, rhododendron (Rhododendron ponticum) forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Neoplectes) edithae Reitter, 1893
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Carabus (Pachycarabus) staehlini Adams, 1817
Male adult
S Russia, E-Caucasus, Dagestan, Nukatl Mt Rng h=2600m, 25.VII.1997, V. Savizky leg. L=21 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus helluo Dejean, 1826
Female adult
SE-Spain, Prov.Alicante, Port de la Carrasqueta, 26.XII.1987, M.Toribio leg. L=25 mm
Copyright © Ruslan Panin

Carabus (Imaibius) boysi boysi Tatum, 1851
Male adult
NW-India, Himachal Pradesh, Manikaran env., Pulga vill., 4.VII.2008. L=26 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014 Holotype Paratype
Male adult
Abkhazia, N Slopes of Bzybian Mt. Range, right bank of River Reshevie (= Reshava) (left tributary of River Bzyb), 700-1000 m, 13.V-02.VII.2010, leg. D. Fominykh, I. Solodovnikov
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014 Paratype
Male adult
Abkhazia, N Slopes of Bzybian Mt. Range, right bank of River Reshevie (= Reshava) (left tributary of River Bzyb), 700-1000 m, 13.V-02.VII.2010, leg. D. Fominykh, I. Solodovnikov
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014 Paratype
Male adult
Abkhazia, N Slopes of Bzybian Mt. Range, right bank of River Reshevie (= Reshava) (left tributary of River Bzyb), 700-1000 m, 13.V-02.VII.2010, leg. D. Fominykh, I. Solodovnikov
Copyright © Igor Solodovnikov

Carabus (Neoplectes) besleticus besleticus Kurnakov, 1972
Male+Female adult
Abkhazia, Sukhum Distr., vicinities of Village Kaman, N slope of Mt. Birtskha, 160 m, beech, linden, box, elder forest site, 05.V-04.VII.2013, leg. I. Solodovnikov, S. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Eucarabus) obsoletus nagyagensis (as fossulifer Fleischer, 1893)
Male adult
NW-Romania, Bihor, Mts Seș (Plopis), 20 km N Aleșd, Pădurea Neagră env., VI.2014, J.Kynkor leg. L=24 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) obsoletus obsoletus (as csikii Mallasz, 1901)
Female adult
NE Carpathians, Marmarosh Mts., Transcarphatian Reg., near Lug vill., h=350 m, 26.IV-8.VI.2014, R.Panin & Yu. Kanarsky leg. L=30 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) obsoletus obsoletus (as csikii Mallasz, 1901)
Female adult
NE Carpathians, Marmarosh Mts., Transcarphatian Reg., near Lug vill., h=350 m, 26.IV-8.VI.2014, R.Panin & Yu. Kanarsky leg. L=32 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) gemmifer pseudoformosus Deuve, 2006
Female adult
China, Tibet AR, Qamdo, Gonjo County, Kè Rì Xiāng vill.(Kerri Township), h=3100m, VI.2012, A. Mollard leg. L=38 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) pedemontanus raynaudianus Deuve & Simard, 1977
Male adult
SE France, Alpes-Maritimes, Isola env., Colle Santa-Anna h=2309 m, 44*13',127" N, 07*06',242"E, 31-7.VIII..2012. L=21 mm
Copyright © Ruslan Panin

Carabus (Iniopachus) pyrenaeus pyrenaeus (as cephalicus Csiki, 1927)
Male adult
SW France, Pyrenees-Atlantiques, Mt.Cirque d'Esturou h=1800m, 29.VIII.2014, Pascal Stefani leg. L=20 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus aurichalceus Kraatz, 1879
Male adult
NC-Spain, Cantabria, Santander, Cueto env.XII.2013 L=28 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens festivus Dejean, 1826 (ab. holochrysum)
Female adult
SC-FRANCE, Tarn, Le Linas h=550 m, XI.2008, Pascal Stefani leg. L=24 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus mixtus (as neomixtus Tarrier, 1975)
Male adult
L=29mm SE-France, Drôme, Foret de Saou, VII.2014, Pascal Stefani leg.
Copyright © Ruslan Panin

Ceroglossus chilensis villaricensis Kraatz-Koschlau, 1885
Male adult
CHILE, Villarica, local collector leg. 26 мм.; Determination confirmed by Anichtchenko, 2019
Copyright © Sergey Kakunin

Carabus (Tomocarabus) convexus erzeliki Schweiger, 1962
Female adult
NC-Turkey, Sinop province, Erfelek env., h=200m, 10-27.VII.2014, C. Reuter leg. L=20 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) phoenix Lapouge, 1924
Male+Female adult
C-Lebanon, Byblos /Jbeil Distr., Kfardebian env., h=1200m, XI.2014, C. Reuter leg. female,23mm
Copyright © Ruslan Panin

Carabus (Lamprostus) torosus sinopensis Breuning & Ruspoli, 1970
Male adult
NC-Turkey, Sinop province, Erfelek env., h=200m, 10-27.VII.2014, C. Reuter leg. L=31 mm
Copyright © Ruslan Panin

Carabus (Diocarabus) massagetus massagetus Motschulsky, 1846 (alpin form)
Male adult
Russsia, Altay, Ukok upland, Kaldzhin-Kul Lake h=2400-2550 m, 49 *19*N/87*26*E, 16-27.VII.2008, R.Dudko leg. L=17.5 mm
Copyright © Ruslan Panin

Carabus (Procrustes) piochardi praestigiator Morawitz, 1886
Male adult
C-Lebanon, Byblos /Jbeil Distr., Kfardebian env., h=1200m, XI.2014, C. Reuter leg. L=31 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) blumenthaliensis aequicostalis Staven & Heinz, 1995
Male adult
NE-Turkey:NE of Artvin:Dalis Dağ ,h=1600-1800m, 8-18.VII.2014, C. Reuter leg. L= 25 mm
Copyright © Ruslan Panin

Carabus (Procrustes) coriaceus rugifer Kraatz, 1877
Female adult
W.UKRAINE, NE Carpathians, Lviv Reg., Skole distr., Zelemyanka Mt.Range h=700m, 26.V-1.VII.2010, Panin R. leg. L=43 mm
Copyright © Ruslan Panin

Calosoma (Orinodromus) deckeni (Gersteacker, 1867)
Male adult
04/1912, Tanzania, Mount Kilimanjaro, Bismarck Hügel, 2700m. Image from: Mal, N., Drumont, A., Kerkhof, S. & P. Grootaert, 2012. Pictorial beetle collection of the Royal Belgian Institute of Natural Sciences.
Copyright © - - -

Carabus (Mesocarabus) lusitanicus gougeleti (as valderepisensis Branes, 1988)
Male adult
Spain, Jaen prov., Despeñaperros, Arroyo de los Arcos, 38.381847, -3.515651, 1-2.05.2014, Anichtchenko A. leg. & det.; 18 mm
Copyright © Alexander Anichtchenko

Carabus (Limnocarabus) clathratus clathratus Linne, 1761
Female adult

Copyright © Kirill Makarov

Carabus (Megodontus) violaceus aurolimbatus Dejean, 1829
Male adult

Copyright © Kirill Makarov

Carabus (Euleptocarabus) porrecticollis Bates, 1883
Female adult
Atsushiokanomachikano, Kitakata City, Fukushima Pref., Japan, 29.III.2014, T. Nakamura leg.
Copyright © Alexander Anichtchenko

Carabus (Lamprostus) hemprichi elonensis Schweiger, 1970
Male adult
N-Israel, Midlle Galilea, Nazareth env., Hoshaya viil., 3-12..III.2015, B.Vas’ko leg./det. Panin R.
Copyright © Ruslan Panin

Carabus (Lamprostus) syrus cheikensis Deuve, 1992
Male adult
N-Israel, Midlle Galilea, Nazareth env., Hoshaya viil., 13.XII.2012, Vas’ko, leg./det. Panin R. L=30мм
Copyright © Ruslan Panin

Carabus (Neoplectes) reitteri gagrinus Starck, 1894
Male adult
ABKHASIA, Gagra distr., Mamdsyshkha Mt., near Gagra, Fagus forest + karst, h = 1300-1330 m, biotope N 23, 43º18’186" N / 040º18’44" E, 12.05-05.07.2014, Leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Neoplectes) reitteri gagrinus Starck, 1894
Female adult
ABKHASIA, Gagra distr., Mamdsyshkha Mt., near Gagra, Fagus forest + karst, h = 1300-1330 m, biotope N 23, 43º18’186" N / 040º18’44" E, 12.05-05.07.2014, Leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Neoplectes) reitteri gagrinus Starck, 1894
Female adult
ABKHASIA, Gagra distr., Mamdsyshkha Mt., near Gagra, Fagus forest + karst, h = 1300-1330 m, biotope N 23, 43º18’186" N / 040º18’44" E, 12.05-05.07.2014, Leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Apotomopterus) breuningianus breuningianus Le Moult, 1930
Female adult
SC-China, Guangxi Reg., Pingle County, Mt. Shizishan h=1290 m,5-20,VII.2011, local coll. leg. L=43 mm
Copyright © Ruslan Panin

Carabus (Leptocarabus) arboreus fujisanus Bates, 1883
Male adult
Japan, CE-Honshu, Yamanashi pref., Mt. Fujisan h~1600m, 23.IX.1982, Y. Imura leg. L=23 mm
Copyright © Ruslan Panin

Carabus (Pachystus) cavernosus variolatus Costa, 1839
Male adult
C-Italy:Abruzzo reg., Gran Sasso d'Italia Massif, h~1600 m, VIII.1997, Seichert leg. L=26 mm
Copyright © Ruslan Panin

Carabus (Carabus) stscheglowi Mannerheim, 1827
Male adult
Russian, Chuvashia, Cheboksary, Zavolzhye distr., , near lake Astrakhanka, L=27 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus violaceus trans violaceus purpurascens Fabricius,1787
Male+Female adult
E-Netherlands, Gelderland, Winterswijk Woold env.,15.II.2015, H.de Haan leg.
Copyright © Ruslan Panin

Carabus (Procrustes) coriaceus coriaceus Linne, 1758
Female adult
Belgique : Prov. Luxembourg, Harnoncourt 31.vii.1975 - M. Grotz Leg. (Size : 36 mm)
Copyright © Noël Mal

Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896
Habitat
Andorra, Anyos, ii.2012, J. Nicolas Leg. & Photos
Copyright © Noël Mal

Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896
Male adult
Andorra, Leg. Gerts, vi.1966, 32 mm
Copyright © Noël Mal

Carabus (Apotomopterus) penelope penelope Kleinfeld, 1997 Paratype
Male adult
Chine, C-Fujian: SE of Yong’an, Mt.Tianbaoyan, 25°58’ N / 117°22’ E, h = 1100 m, VII.1997, leg. Wong
Copyright © Igor Solodovnikov

Carabus (Chaetocarabus) arcadicus arcadicus Gistl, 1850
Male adult
Graecia, Pindus, Mt. Velouchi (Timfristos), 1200m, vi.1970-A. Sermet Leg., 30 mm
Copyright © Noël Mal

Calosoma (Calosoma) sycophanta Linne, 1758
Male adult
Turkey : Antalya, Yukarikaraman 3.iv.2000 - M. Grotz Leg. (Size : 33 mm) Coll. N. Mal
Copyright © Noël Mal

Calosoma (Calosoma) sycophanta Linne, 1758
Female adult
Turkey : Antalya, Yukarikaraman 3.iv.2000 - M. Grotz Leg. (Size : 33 mm) Coll. N. Mal
Copyright © Noël Mal

Carabus (Tachypus) cancellatus intermedius Dejean, 1826
Female adult
NW -Serbia,Fruška Gora,Vrdnik env. h=380m, 9-10.VII.2013, M. Janalik leg. L=29 mm
Copyright © Ruslan Panin

Carabus (Carabus) arvensis carpathus Born, 1902
Male adult
W Ukraine, NE Carpathians,Transcarpathian reg., Vihorlat, near Strychava vill., Mt Knyaginytsya h= 500 m, 18.V-22.VI. 2014, Yu.Kanarsky leg. L=18 mm
Copyright © Ruslan Panin

Carabus (Carabus) arvensis arvensis Herbst, 1784
Female adult
NW Ukraine, Polesia, Rovno reg., Rokytne distr., near Bil’sk vill., 4-6.VII.2015, Panin R. leg. L=20 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) augustus ignigena Hauser, 1914
Female adult
China : Guizhoh, Sinan vi.2006 (Size : 35 mm) Coll. N. Mal
Copyright © Noël Mal

Carabus (Megodontus) violaceus exasperatus Duftschmid, 1812
Male adult
Hungary, Budapest, Mt Buda, Harsbokor-hill,16-19.V.1992, I.Retezár leg., L=27 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) odoratus baeri Menetries, 1851: 45
Male adult
Russia, SW Taymyr, near Talnakh, 25 km N Norilsk, hilly tundra, hillside overgrown with alder slate, dwarfish birches, blueberry, h = 150-250 м, 24-28.06.2003, leg. S.Saluk
Copyright © Igor Solodovnikov

Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Male adult
NE-Hungary, 4 km S Sátoraljaújhely, right bank riv. Bodrog (Long-erdő), 8.VI.1992, leg./det. I.Retezár leg. L=28 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi distinguendus Csiki, 1906
Male adult
NC-Hungary: E- part of Budapest, Cinkota,28.VIII-17.IX.1980, I.Retezár leg. L=27 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus problematicus Herbst, 1786
Female adult
W-Hungary, NE-Alps, Mts. Kőszeg, Velem env., Borha-forrás, 26.VII.1981, Gy. Somorijai leg. L=26 mm
Copyright © Ruslan Panin

Carabus (Pachycarabus) swaneticus kvirensis Belousov, 1992
Male adult
SC-Caucasus: NW-Georgia, Mingrelia, Egrissky Mt.Rng., h~1800-2000 m, 23-26.VII.1990, I.Retezár leg. L=20 mm
Copyright © Ruslan Panin

Carabus (Pachystus) cavernosus rozneri Retezár & Gy. Szél, 2013 Paratype
Male adult
Romania, Apuseni : Trascău Mts. L=27 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi distinguendus (as bieneri Mandl, 1965)
Male adult
C-Slovakia, Detva Distr., Stara Huta env. h=700-800m, 3.VI.-5.VII.2012, V.Slovák lgt. L=29 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) rothi distinguendus (as subparvulus Mandl, 1965)
Male+Female adult
NC-Hungary:Borzsony Mts, Nograd-Diosjeno,1-14.V.2004, Krause Rene leg. female, 32 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859
Female adult
Russia, SE Siberia, N Chita reg., Kodar Mt. Range, Big Leprindo Lake env., h~1800 m., L=21 mm
Copyright © Ruslan Panin

Calosoma (Ctenosta) grandidieri Maindron, 1900
Female adult
MADAGASCAR, Tulear Prov., nr. Tsimanampetsotsa NP, P. Schüle det.
Copyright © Peter Schüle

Carabus (Lamprostus) erenleriensis erenleriensis Schweiger, 1966, loc.typ.
Male adult
NW-Turkey:Bolu Prov., Abant Lake, 25.VIII.1990 L=35 mm
Copyright © Ruslan Panin

Carabus (Imaibius) stoliczkanus brauni Heinz, 1993
Female adult
NE Pakistan, Azad Kashmir, Bagh Distr., Sudhan-Gali h~2300 m,15.VIII.1994, S Dacatra leg. L=37 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) arrowi yubeicus Deuve & Tian, 1999
Female adult
S-China, Guangxi, Jinxiu Co., Mt. Dayao,28.VII.2007, L=40 mm
Copyright © Ruslan Panin

Carabus (Ctenocarabus) melancholicus melancholicus Fabricius, 1798
Male adult
NW Morocco, Kénitra, Souk El Arbaa env., II.1985, V.Olivella leg. L=25 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) concolor lepontinus Born, 1908
Male adult
NW-Italy, Piemonte (VB), Lepontine Alps, Mt. Zeda , 6.VII.1998, L Diotti leg., L=20 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) concolor concolor Fabricius, 1792
Female adult
NW-Italy, Breuil-Cervinia h~2500 m, 2.IX.1971, Georgeon leg. L=21 mm
Copyright © Ruslan Panin

Carabus (Oxycarabus) saphyrinus pirithous Reitter, 1896
Male adult
NW-Turkey,Istanbul (Asian part),Polonezköy env., 2.VI.2009. S. Dacatra leg. L=27 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) morawitzianus morawitzianus Semenov, 1887
Male adult
C China, Qinghai, Zadoi County, Toramarkog h~4600 m, VI.2001, P. Cavazzuti leg. L=17 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) pedemontanus germanae Casale & Cavazzuti, 1977
Male adult
NW-Italy, Piemonte, Imperia prov., Mt.Saccarello h~2000m, 30.V.1986. L=21 mm
Copyright © Ruslan Panin

Calosoma (Castrida) leleuporum (Basilewsky, 1968) Holotype
Male adult
Picture: Camille Locatelli, Copyright: RBINS / DIGIT-3 Belspo Licence: CC BY NC ND http://virtualcollections.naturalsciences.be
Copyright © - - -

Carabus (Orinocarabus) fairmairei pelvicus Cavazzuti, 1973
Male adult
NW-Italy, Cuneo, Maira Vall., Acceglio env., La Colletta h~2800 m, 12.VII.2007, R. Monguzzi leg. L=19 mm
Copyright © Ruslan Panin

Carabus (Isiocarabus) miaorum Lassalle & Prunier, 1993
Male adult
S-China, N-Guangxi, Mao'er Shan, h~1600 m, V.2009, L= 26 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) italicus rostagnoi Luigioni, 1904
Female adult
C-Italy, Lazio, Fondi env.,10.II.1979, L = 25 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) victor victor Fischer, 1836
Female adult
NE-Turkey, Giresun, Sehitler Geçidi (Pass) h~2000 m, 23.VII.2001, S. Dacatra leg., L=19 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) parreyssi parreyssi Palliardi, 1825
Male adult
W-Croatia, NP Northern Velebit, Zavižan,15.V-13.VI.1996, Colombetta leg., L=21 mm
Copyright © Ruslan Panin

Carabus (Lamprostus) torosus giresuni Breuning, 1964 (as paululus Heinz & Korge, 1964)
Male adult
NE-Turkey, Giresun, Sehitler Geçidi (Pass) h~2000 m,24.VII.2001, S. Dacatra leg., L=26 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) creutzeri baldensis Schaum, 1857
Male adult
NC-Italy: Trento, Sega di Ala, IX.1999, A. Vicini leg. , L=24 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) rossii Dejean, 1826
Female adult
NC Italy, Emilia reg., Bologna, San Martino, 29.V.2016, G..L.Agnoli leg. L=30 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei folgariacus Born, 1913
Male adult
NE-Italy, Trentino-Alto Adige (Südtirol), N of vill. San Michele vill., 13-17.VI.2016, R. Janak leg. L=20 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens kraussi Lapouge, 1898
Female adult
C-Austria, Styria, Ennstal Alps, Totes Gebirge Mt Rng, Mt Loser h~1100 m, 16-20.VII.2014, R.Janak leg. L=27 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) adamsi Adams, 1817
Male adult
CN-Georgia, N of Tbilisi, NW of Tianeti, Tvalivi env., 20.VI.2015, F. Grycz leg. L=24 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) ignimitella guangxicola Deuve, 1998
Male adult
S-China, Guangxi prov., Jingxiu, Mt. Dayao Shan, 1-17.V.2006, L=47 mm
Copyright © Ruslan Panin

Carabus (Lamprostus) robustus triumphans Cavazzuti, 1987
Male adult
NE-Turkey:Trabzon, Tonya env., VIII.2004. L=41 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) latreilleanus Csiki, 1927
Female adult
NW-Italy, Piemonte, Biella, Biellese Alps Rng, Mt.Camino, 25.VI.1997, V.Monzini leg. L=18 mm
Copyright © Ruslan Panin

Carabus (Lamprostus) torosus nordmanni Chaudoir, 1848 (as enricoi )
Male adult
NE-Turkey:Ordu prov.,3 km N Gölköy h~1100 m, VII.2006, L=37 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) morbillosus lampedusae Born, 1925
Male adult
S Italy, Isl. Lampedusa, Baia dei Conigli, VI.2003, L=29 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014 Paratype
Female adult
Abkhazia, Bzybsky Mt.Rng, right bank of river Reshevie (left tributary of river Bzyb) h~670-720 м, 13-14.V.2010, D.Fominykh & I.Solodovnikov leg. L=41 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) breuningi Csiki, 1927
Female adult
NW-Spain, Prov. Zamora, San Martín de Castañeda, 5.VII.1984, M.Toribio leg. L=28 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) viridilimbatus panini I.Solodovnikov 2017 Holotype
Male adult
Russia, SE Yakutia, Neryungri distr., Tokinski Stanovik Mt Rng., Beder riv., (right tributary rivers Algama), 26.VI.1996, Krasnov M. leg., 17 mm
Copyright © Ruslan Panin

Calosoma (Caminara) denticolle Gebler, 1833: 274
Female adult

Copyright © Kirill Makarov

Carabus (Tachypus) auratus honnoratii Dejean, 1826
Male+Female adult
France : Vaucluse, Mt. Ventoux-Mt. Serein, 1400-1800m, vii.1969-N. Mal Leg.-Coll.RBINS, N. Mal - including f.i. ventouxensis and nicolasi
Copyright © Noël Mal

Carabus (Megodontus) kolbei taikinus Ishikawa, 1993
Male adult
N-Japan, SC-Hokkaido:Tokachi, Hiroo Distr., Taiki-cho env., 28.VIII-2.IX.2017, Teruo Hirabayashi leg. L=24 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) estreicheri (ab.rufofemoratus Lomnicki, 1892)
Female adult
W Ukraine, NW Podolian, Lviv reg., Brody distr., Makitra Mt, steppe site, 15.VI - 24.IX.2017, Panin R. & Kanarsky Yu. leg.
Copyright © Ruslan Panin

Calosoma (Elgonorites) elgonense Burgeon, 1928
Male adult
Kenya, M. Elgon, vers. S, 3600 m., 6.IX.1986. (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Carabus (Coptolabrus) ignimitella antaeus Hauser, 1914 (as guidongicus)
Female adult
China, E-Hunan, Zhuzhou, Yanling County, Wanyang Shan Mts, VI.2003 , L= 47 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) wulffiusi liaoningensis Deuve & Imura, 1990
Female adult
NE-China:E-Liaoning: NP Kuandian Tianqiaogou , VI.2010, L=24
Copyright © Ruslan Panin

Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931
Male adult
NE-China:E-Liaoning: NP Kuandian Tianqiaogou , VI.2010, L=25 mm, det. Panin R.
Copyright © Ruslan Panin

Carabus (Coptolabrus) jankowskii fusanus Born, 1907
Female adult
South Korea, S Gyeongsang Prov., Busan, Mt Geumwonsan, 17.VI.2017, L=37 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) formosus wanxianicus Imura, 1995
Male adult
China, NE-Sichuan, Dazhou, Wanyuan env., 6.VI.2005. L = 33mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
Female adult
NE-Hungary. (photo by Imre Retezár)
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
Male adult
NE-Hungary. (photo by Imre Retezár)
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Male adult
W Ukraine, Transcarpathian, Mts E of Uzgorod, Veliki Lazy h~270-300 m, 8.IV-23.V.2011, Panin R. leg. L=28 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) nankotaizanus Kano, 1932
Male adult
NE-Taiwan, Yilan County, V.2017, L=39 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) jankowskii fusanus (as obtusipennis)
Female adult
SC-Korea: Chiri-san Massif, Mt. Jirisan, 28.VI.2011, L=35 mm
Copyright © Ruslan Panin

Carabus (Homoeocarabus) maeander paludis Gehin, 1885
Female adult
Japan, Hokkaido; Tomakomai city, Benten, 17.V.2015, Takemoto leg., L=21 mm
Copyright © Ruslan Panin

Carabus (Axinocarabus) fedtschenkoi kondarensis Kryzhanovskij, 1953
Female adult
NW-Tadzhikistan: Gissar Mt.Rng, Varzob river ravine, Gushari env., h~1400-1500m, 24.III.2010, L=29 mm
Copyright © Ruslan Panin

Carabus (Leptocarabus) arboreus fujisanus Bates, 1883
Female adult
Japan, Honshu, Mt. Fuji h~1900m, Shizuoka pref., Fujinomiya city, 23.VI.2007, M. Matsunaga leg., L=25 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) irregularis irregularis (as jurassicus Born, 1911)
Male+Female adult
France, Ain (01), Jura Mts, Grand Colombier, Forêt d' Arvières h~1250 m, 25.X.2014, Ya. Vasseur leg., L=23 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) komarowi komarowi Reitter, 1882
Female adult
NW Georgia, Upper Svaneti, Mestia distr., Lakhamula env. h~1250 m, 19.VI.2014, V.Sima leg. L=33 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) mellyi martviliensis Retezar & Djavelidze, 1992
Female adult
W Georgia, Martvili Distr., 5 km N T'aleri, 11-18.VI.2014, V. Sima leg., L=50 mm
Copyright © Ruslan Panin

Carabus (Lamprostus) calleyi pseudoprasinus Lapouge, 1912
Male adult
S Georgia, Tsalka (Paravani) 41°29′18.7″N, 43°51′12.3″E, 8-9.V.2016, Pavel Turek leg. / det. I. Solodovnikov, L=23 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) gemmifer mesites Fairmaire, 1897
Male adult
China, Shimian 1200m Shimian Yaan Sichuan VII.2017 (DUBC). det. Anichtchenko A. 2018
Copyright © Alexander Anichtchenko

Carabus (Orinocarabus) linnaei linnaei Panzer, 1812
Male adult
W.UKRAINE, Lviv Reg., NE Carpathians, Skolivski Beskidy Mts. Zelemyanka Mt. Rng h=600m, 15.VII-12VIII.2014, Panin R. leg. L=19 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei linnaei Panzer, 1812
Female adult
W Ukraine, Transcarpathian Reg., Rakhiv distr., Marmarosh Mts., h~400 m, 19-23.VII.2010, Reserve "Kuziy", Panin R. leg. L=23 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei linnaei Panzer, 1812
Male adult
W-UKRAINE, Ivano-Frankivsk region, S Gorgany Mts, Synyk Mt. h =1630 м., 15.V-18.VI.2010, Panin R. leg. L=19 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei linnaei (as scopolii Dejean, 1826)
Male adult
W Ukraine, Lviv reg., NW Podolian, forest near Vybranivka vill., 12.X.2007, Panin R. leg. L=19 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI.-5,VIII.2017, R. Panin R. & Yu.Kanarsky leg.
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI.-5,VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=17 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Female adult
Male adult W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI.-5,VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=18 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI.-5,VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=17.5 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Holotype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=17.5 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L =17 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=16.5 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=17 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Female adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=17 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Female adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=18 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Female adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=19 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=19 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Female adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=20 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=16.5 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=19 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Gutyn-Tomnatyk h~2015 m alpin zone, 8.VI-28.VII.2011, R. Panin R. leg. L=14 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Female adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Gutyn-Tomnatyk h~2015 m, alpin zone, 8.VI-28.VII.2011, R. Panin R. leg. L=18 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) ibericus ibericus (as imereticus Retezar, 2011) Paratype
Male adult
S-Caucasus: NC-Georgia, W slope Likhi Mt. Rng., 3 km W of Rikoti Pass h~900 m, 15.VI - 8.VII.2006, A.Puchkov leg., L=29.5 mm
Copyright © Ruslan Panin

Carabus (Megodontus) persianus espeyliensis Deuve, 2001
Male adult
N-Iran, Gilan Prov., Deylaman Distr., Balarud Mts, Espili env., 18-24.VI.2017, Václav Čutka leg., L= 32 mm
Copyright © Ruslan Panin

Carabus (Carabus) sculpturatus (as gilanicus Mandl, 1967)
Male adult
N-Iran:Gilan Prov., Asalem Distr., Siahbil, deciduous forest (very wet) h= -10 m, 20-24.VI.2017, V. Čutka leg., L=19 mm
Copyright © Ruslan Panin

Carabus (Carabus) sculpturatus (as gilanicus Mandl, 1967)
Female adult
N-Iran:Gilan Prov., Asalem Distr., Siahbil, deciduous forest (very wet) h= -10 m, 20-24.VI.2017, V. Čutka leg., L=22 mm
Copyright © Ruslan Panin

Carabus (Tribax) puschkini platypterus Ganglbauer, 1886
Male adult
SW Georgia, Lesser Caucasus, Adzhar-Imereti (Meskheti) Mt Range, 10 km S Sairme h~1850 m, 27.VI-2.VII.2013, A.Puchkov & Martynov leg. L=27 mm
Copyright © Ruslan Panin

Carabus (Ophiocarabus) successor successor Reitter, 1896
Male adult
SE-Kazakhstan, Narynkol distr., 4 km SW Bayankol, h~2400-2500 m, 2-14.VI.2007, Shumov leg. L=17.5 mm
Copyright © Ruslan Panin

Carabus (Ophiocarabus) successor successor Reitter, 1896
Female adult
SE-Kazakhstan, Narynkol distr., 4 km SW Bayankol, h~2400-2500 m, 2-14.VI.2007, Shumov leg. L=17 mm
Copyright © Ruslan Panin

Carabus (Meganebrius) quinlani quinlani Mandl, 1965
Male adult
C-Nepal, Gandaki, Manang Distr., Timang env., h~2500-2600m, old open forest, grassland, 3-5.V.2007, Fouqué R. leg. L=19 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) separatus bararii Deuve, 2000
Male adult
N-Iran: Mazendaran: Ramsar env., deciduous forest, h~305 m, 16-24.VI.2017, Václav Čutka leg. L=22 mm
Copyright © Ruslan Panin

Carabus (Pagocarabus) sichuanicola sichuanicola (as drolmae Lassalle, 2003)
Female adult
China, Sichuan, NW Chengdu, Mt. Jiajin Shan h~4000 m,VI. 2016, local collector, L= 24 mm
Copyright © Ruslan Panin

Carabus (Cupreocarabus) alpherakii alpherakii Semenov, 1898
Female adult
China, W Sichuan, Litang env., h~5000 m,29.VII -3.VIIII.1995, J. Schneider leg. L=23 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) separatus bararii Deuve, 2000
Female adult
N-Iran: Mazendaran: Ramsar env., deciduous forest, h~305 m, 16-24.VI.2017, Václav Čutka leg. L=23,5 mm
Copyright © Ruslan Panin

Carabus (Tribax) puschkini kolenatii (as suramensis Kraatz, 1887)
Male adult
S-Caucasus: NC-Georgia, W slope Likhi Mt.Range, 3 km W Rikoti Pass h~900m 11-14.VI -2006, A.Puchkov leg. L=27 mm
Copyright © Ruslan Panin

Carabus (Cryptocarabus) tsharynensis Kabak 1994
Male adult
SE-Kazakhstan, Almaty Region, W-Zailiyskiy Alatau rng, Belshabdar valley h~1300 m, 25-27.VI.2002, A.Puchkov leg. L=17 mm
Copyright © Ruslan Panin

Carabus (Cryptocarabus) tsharynensis Kabak 1994
Female adult
E-Kazakhstan, Almaty Region, W-Zailiyskiy Alatau, Turgen distr., Tauturgen vill., h~1200 m, 8-13.VI.2002, A.Puchkov leg. L=19 mm
Copyright © Ruslan Panin

Carabus (Megodontus) persianus javahardehensis Deuve, 2000
Male adult
N-Iran, Mazandaran Prov., Ramsar Co., Javahar Deh env., 16-25.VI.2017, Václav Čutka leg. L=28 mm
Copyright © Ruslan Panin

Carabus (Megodontus) persianus javahardehensis Deuve, 2000
Female adult
N-Iran, Mazandaran Prov., Ramsar Co., Javahar Deh env., 16-25.VI.2017, Václav Čutka leg. L=30 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) separatus separatus Gehin, 1885
Male adult
N-Iran:Gilan Prov., Asalem - Khalkal road, 25-30.V.2018, Václav Čutka leg. L= 21 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) separatus separatus Gehin, 1885
Female adult
N-Iran: Gilan Prov., Asalem - Khalkal road, 25-30.V.2018, Václav Čutka leg. L= 23 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) separatus separatus Gehin, 1885
Male adult
N-Iran:Gilan Prov., Asalem - Khalkal road, 25-30.V.2018, Václav Čutka leg. L= 22 mm
Copyright © Ruslan Panin

Carabus (Hemicarabus) serratus Say, 1825
Male adult
E-USA,Virginia, Leesburg env., Ball's Bluff Battlefield Regional Park, 27.VIII-4.IX.2015,V. Čutka leg. L=17 mm
Copyright © Ruslan Panin

Carabus (Hemicarabus) serratus Say, 1825
Female adult
E-USA, Virginia, Leesburg env., Ball's Bluff Battlefield Regional Park, 27.VIII-4.IX.2015,V. Čutka leg. L=18 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) separatus haerii Heinz, 1973
Female adult
N-Iran:Gilan Prov., Rudbar, Harzevil env.,23-24.V.2018, Václav Čutka leg. L=24 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) separatus haerii Heinz, 1973
Male adult
N-Iran: Gilan Prov., E of Rudbar, Barehsar env., 19-23.VI.2017, Václav Čutka leg. L=22 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) dufouri Dejean, 1829
Female adult
S Spain, Granada, Sierra Arana, Píñar env., 14.X.2018, F. Garcia leg. L=27 mm
Copyright © Ruslan Panin

Carabus (Piocarabus) sui Imura & Zhou, 1998
Female adult
CE-China, Henan, Mt Wangwu, V.2007, local collector. L= 33 mm
Copyright © Ruslan Panin

Carabus (Oreocarabus) ghiliani La Ferte-Senectere, 1847
Female adult
C-Spain: Sierra de Guadarrama, Puerto de Cotos (Madrid), 20.IX.2018, F.Rodriguez leg., L=22 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) breuningi Csiki, 1927 (loc.typ.)
Male adult
NW-Spain, Orense, Orensano Central Massif, Monte de Cabeza de Manzaneda, 25.VII.1988, F. Rodrigues leg. L=25 mm
Copyright © Ruslan Panin

Carabus (Alipaster) pupulus pupulus Morawitz, 1889
Female adult
S Kyrgyzstan, Inner Tian Shan, Dzhany-Dzher Mt range, upper Mudyuryum riv. Ak-Baytal gorge h~3300 m, 15.VII.2011, S.Toporov leg. L=16 mm
Copyright © Ruslan Panin

Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
Male adult
Russia, C-Caucasus: N Ossetia, Chekhatsirtit Mt range, Mt Tepli h=3000 m, 21.VII.2009. /det. Solodovnikov I.A, L=20.5 mm
Copyright © Ruslan Panin

Carabus (Goniocarabus) perelloi Casale, 1979
Male adult
N-Afghanistan, Hindukush, Provinz Baghlan, N of Kabul, N Salang Pass, subalp., h = 2200 m, 35°27'.494N / 68°58.399'E, Mai 2011, leg. Christoph Reuter (TOPOTYPUS)
Copyright © Igor Solodovnikov

Carabus (Rhigocarabus) turnai turnai Deuve, 1994
Male adult
China, Tibet, W-Sichuan, pass btw Sêrtar / Luhuo h=4200 m, 28.VI.2011, V.Patrikeev leg.L=19 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) turnai turnai Deuve, 1994
Female adult
China, Tibet, W-Sichuan, pass btw Sêrtar / Luhuo h=4200 m, 28.VI.2011, V.Patrikeev leg. L=21 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) elysii xiaoganicus Deuve et Remond, 2016
Male adult
China, Hubei, Xiaogan, Mt Shuangfeng Shan, VII.2016 L=37 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Paratype
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg.
Copyright © Ruslan Panin

Carabus (Coptolabrus) gemmifer gemmifer (as loczyi Csiki, 1927)
Female adult
S-China, W-Yunnan: Dali city, Mt Jizushan, VI.2018, L=34 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) davidis helmutschuetzei Kleinfeld, 1995
Male adult
SE-China, SW-Jiangxi:S of Yongxin: Mt.Jinggangshan,V.2012, L=35 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
Male adult
NC-Spain, Basque Co, Álava, Sierra de Gorbea, VI.2014, L=29 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) smaragdinus furumiellus Deuve, 1994
Female adult
NE-China, E-Liaoning, Benxi, Mt Liuershan, VIII.1998. L=39 mm
Copyright © Ruslan Panin

Carabus (Oreocarabus) luetgensi pseudosteuarti Lapouge, 1924
Female adult
Portugal, Guarda, Sierra de Estrela h=800 m, 26.VI-4.VIII.2015, L=26 mm
Copyright © Ruslan Panin

Carabus (Oreocarabus) luetgensi pseudosteuarti Lapouge, 1924
Male adult
Portugal, Guarda, Sierra de Estrela h=800 m, 26.VI-4.VIII.2015, L=24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
Female adult
W Ukraine, Transcarpathian lowland, near Chop, 11.VI-19.VII.2019, Panin R leg. L=29 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
Male adult
W Ukraine, Transcarpathian lowland, near Chop, 11.VI-19.VII.2019, Panin R leg. L=30 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
Female adult
W Ukraine, Transcarpathian lowland, near Chop, 11.VI-19.VII.2019, Panin R leg. L=31 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
Female adult
W Ukraine, Transcarpathian lowland, near Chop, 11.VI-19.VII.2019, Panin R leg. L=29 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
Male adult
W Ukraine, Transcarpathian lowland, near Chop, 11.VI-19.VII.2019, Panin R leg. L=28 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii ronayi Csiki, 1906
Female adult
W Ukraine, Transcarpathian lowland, near Chop, 11.VI-19.VII.2019, Panin R leg. L=30 mm
Copyright © Ruslan Panin

Ceroglossus chiloensis sybarita Gerstaecker, 1858
Female adult
Chile, Isla Chiloe, Puntra, 1.1988. det. Anichtchenko A. 2019
Copyright © Alexander Anichtchenko

Ceroglossus chiloensis chiloensis
Male adult
Chile, Ahoni, Isla Chiloe, IV.1988. 20 mm; det. Anichtchenko A. 2019
Copyright © Alexander Anichtchenko

Ceroglossus chilensis mochae Reed, 1874
Female adult
Chile, Isla Mocha, 3.1988. det. Anichtchenko A., 2019
Copyright © Alexander Anichtchenko

Ceroglossus buqueti deuvei Jiroux, 1996
Male adult
Chile, W.Temuco, feb.1986. 22 mm; det. Anichtchenko A. 2019
Copyright © Alexander Anichtchenko

Ceroglossus chilensis fallaciosus Kraatz, 1880
Female adult
Chile, Concepcion, 12.1986. 26 mm; det. Anichtchenko A. 2019
Copyright © Alexander Anichtchenko

Ceroglossus suturalis puxel Jiroux, 1996
Male adult
Llanquihue, Chile, 2.1986. 20 mm. det. Anichtchenko A. 2019
Copyright © Alexander Anichtchenko

Ceroglossus buqueti subnitens Kraatz-Koschlau, 1885
Female adult
Chile. Tolhuaca 4.2.1988. det. Anichtchenko A. 2019
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Male adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 4-24.V.2010, Panin R. leg. L= 29 mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicusn (as rareulensis Born, 1907)
Male adult
NE-Romania, Rarau Mts. h-1300m, VI.2014, J. Kynkor leg. L=23 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus (as polonicus Lomnicki, 1886)
Female adult
W Ukraine, Lviv Region, Zhydahciv distr., Borodchytsi vill.,the right bank of Dniester river, 11-16.VII.2012, Panin. R leg. L=27 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Holotype
Male adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 28.V-9.VI.2013, Panin R. leg. L=29 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Female adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 28.V-9.VI.2013, Panin R. leg. L= 28 mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Male adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 18.V-12.VI.2014, L=28 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Male adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 27.IV-13.VI.2018, Panin R. leg. L= 28 mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Male adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 27.IV-13.VI.2018, Panin R. leg. L= 29 mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Female adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 20.V-25.VI.2008, Panin R. leg. L= 31 mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Female adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 20.V-25.VI.2008, Panin R. leg. L= 30 mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Male adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 12.V-2.VI.2009, Panin R. leg. L= 29 mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Female adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., 27.IV-13.VI.2018, Panin R. leg. L= 31 mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Male adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill., Panin R. leg. L= 28 mm.
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens (ab.goldeggi Duftschmid, 1812)
Male adult
W-Ukraine, Podolian Tovtry, Ternopol reg., Pidvolochysk distr., Shevchenkove vill., steppe sites, 6.V-7.VI.2012, Panin R. leg. L=24 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) sylvestris transsylvanicus Dejean, 1826
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg. L=21 mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) sylvestris transsylvanicus Dejean, 1826 (aberrant)
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone, 10.VI-5.VIII.2017, R. Panin R. & Yu.Kanarsky leg.
Copyright © Ruslan Panin

Calosoma (Castrida) granatense granatense Gehin, 1885
Male adult
Ecuador, 28.9.85, Galapagos, Isabela Sud, Sierra Negra Sud, 490m Pierre Moret leg. & det.
Copyright © Alexander Anichtchenko

Calosoma (Blaptosoma) atrovirens Chaudoir, 1869
Male adult
Mexico, SLP, Hwy 80, 3k. E Ciudad del Maiz, 20.VII.1991, leg. Shetterly et al; 22 mm; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Calosoma (Blaptosoma) porosifrons Bates, 1891
Female adult
Mexico, Durango, Otinapa road 23*59.2N 104*58.5W, 6.7.2010, leg. J.A. Shetterly et al; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Calosoma (Blaptosoma) viridisulcatum Chaudoir, 1863
Male adult
Mexico, Jal., N Guadalajara, Hwy 80, 20.18.744N 103.54.015W, 1664m, km 49, 30.VI.2010, leg. J.A. Shetterly et al; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Calosoma (Blaptosoma) laeve Dejean, 1826
Male adult
Mexico, Mich, 11 k E Zacapu, Hwy 15, 19*45.840 N 101*43.733 W, 2021 m, 28.6.2010, leg. J.A. Shetterly et al; 26 mm; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Carabus (Deroplectes) klapperichianus coiffaitianus Deuve, 1990
Male adult
Tadzhikistan: Badakhshan: Ishkashim Mt.R.: Kuh-i-Lal, VI.1979 leg. I.K. Lopatin (TOPOTYPUS)
Copyright © Igor Solodovnikov

Carabus (Deroplectes) klapperichianus coiffaitianus Deuve, 1990
Female adult
Tadzhikistan: Badakhshan: Ishkashim Mt.R.: Kuh-i-Lal, VI.1979 leg. I.K. Lopatin (TOPOTYPUS)
Copyright © Igor Solodovnikov

Carabus (Deroplectes) klapperichianus salangicus Kleinfeld & Reuter, 2010 Paratype
Female adult
N-Afghanistan, Hindukush Mt.Rng, Salang Pass, Nordseite, 25°27'.494N, 68°58.399'E, ca. reg. subalp., h = 2200-2400 m, V.2010, leg. Christoph Reuter
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) kokujewi Semenov, 1898
Male adult
Russia, Karachai-Cherkessia, vall. Daut Riv., near Glacier Ryndzhi-Murudzhu, h = 3050 m, 22.VIII.2011, leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) kokujewi Semenov, 1898
Female adult
Russia, Karachai-Cherkessia, vall. Daut Riv., near Glacier Ryndzhi-Murudzhu, h = 3100 m, 22.VIII.2011, leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov

Calosoma (Calosoma) sycophanta Linne, 1758
Male adult
Italy: Abruzzo, Pallano, h = 1000m, 18.06.1993, leg. Facovone
Copyright © Igor Solodovnikov

Calosoma (Calosoma) sycophanta Linne, 1758
Female adult
Calosoma (s.str.) sycophanta severum f.i. cupreum Letzner, 1850 Turkey, Pozanti, 30.IV.2009 , leg. O.Boilly
Copyright © Igor Solodovnikov

Calosoma (Calosoma) scrutator Fabricius, 1775
Male adult
USA: Texas, Live Oak County 5 miles South of George West, 24.09.2010, local collector
Copyright © Igor Solodovnikov

Calosoma (Calosoma) sycophanta Linne, 1758
Male adult
Italy: Abruzzo, Pallano, h = 1000m, 18.06.1993, leg. Facovone
Copyright © Igor Solodovnikov

Calosoma (Calosoma) aurocinctum Chaudoir, 1850
Female adult
MEXICO, Sian Ka’an, Quintana Roo, 19.VI.2015, leg. Julio Medin
Copyright © Igor Solodovnikov

Calosoma (Carabosoma) angulatum Chevrolat, 1834
Male adult
Reserva silvestra de Domitila, Pl., Granade prov., Nicaragua, 1-5.6.2005. det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Calosoma (Ctenosta) planicolle Chaudoir, 1869
Male adult
C Tanzania, Chikuyu, Dodoma, 1.2015 (DUBC)
Copyright © Alexander Anichtchenko

Carabus (Cechenochilus) kokujewi Semenov, 1898
Male adult
Russia, Karachai-Cherkessia, vall. Daut Riv., near Glacier Ryndzhi-Murudzhu, h = 3050 m, 22.VIII.2011, leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) kokujewi Semenov, 1898
Female adult
Russia, Karachai-Cherkessia, vall. Daut Riv., near Glacier Ryndzhi-Murudzhu, h = 2550 m, 23.VIII.2011, leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) adangensis Gottwald, 1983
Female adult
Russia, Karachai-Cherkessia, Abischira-Akhuba Mt.R., env. vill. Romantic, alpin zone, h = 2470 m, 10.07-22.08.2019, leg. S.M. Pavlyuchuk, A.P. Tatulyan
Copyright © Igor Solodovnikov

Carabus (Rhigocarabus) morawitzianus morawitzianus Semenov, 1887
Male adult
Qinghai, Anemaqen mts pass, 20 km N of Maduo town, 4500 m, 6.7.2012, V. Patrikeev leg.; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
Female adult
W Ukraine, Transcarpathian, Marmarosh Mts., near Lug vill., h=400 m, 28.IV -26.VII.2017, Panin R. leg., L= 38 mm
Copyright © Ruslan Panin

Calosoma (Carabomimus) morelianum Bates, 1891
Female adult
Mexico, Santa Martha de Ocuilan, 2760m, 19*04'09"N 99*23'17"W. 19.6.1995, J. Shetterly et al leg. ; det. Anichtchenko A., 2020; 14mm
Copyright © Alexander Anichtchenko

Calosoma (Carabomimus) politum diminutum Bates, 1891
Male adult
Mexico, MX, Salazar, Alt. 3000 m, 19*18.194N, 99*23.566W, 24.6.2000, leg. J.A. Shetterly et al.; 15 mm; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Calosoma (Carabomimus) striatulum s.str.
Male adult
Mexico, VC, Hwy 140, NE Perote, km 140, 19*37'50"N 97*10'17"W, 2450 m, 21.6.2000, J. Shetterly et al leg.; 15 mm; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Calosoma (Blaptosoma) haydeni punctulicolle Bates, 1891
Male adult
Mexico, Chihuahua, S Hy 24, San Francisco Mesa, nr. mine 2300 m, 24*49.671N 105*50.956W, 25.7.2005, leg. J.A. Shetterly (cAA). det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Carabus (Cryptocarabus) sacarum Kabak, 1998
Male adult
SE-Kazakhstan, Almaty reg., C-Zailiyskiy Alatau N-slopes, Kaskalen river Valley, Izvestkovy vill., h=1700 m, 6.V.2015, D.Kovalchuk leg./det. R.Panin L=18 mm
Copyright © Ruslan Panin

Carabus (Cryptocarabus) sacarum Kabak, 1998
Female adult
SE-Kazakhstan, Almaty reg., C-Zailiyskiy Alatau N-slopes, Kaskalen river Valley, Izvestkovy vill., h=1700 m, 6.V.2015, D.Kovalchuk leg./det. R.Panin L=20 mm
Copyright © Ruslan Panin

Calosoma (Callisthenes) elegans danilevskii Obydov, 1997d
Female adult
SE-Kazakhstan, Almaty reg., C Trans-Ili Alatau N-slopes, Kaskalen river Valley, Izvestkovy vill., h=1700 m, 6.V.2015, D.Kovalchuk leg./det R.Panin, L=32 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
Male adult
Russia, S Tuva, Khorumnug-Taiga Mts, Buren riv. Valley, Buren-Bay-Khaak vill. h=900 m, 17-24.VI.2002, Vashchenko A.& B.leg.,/ det. Panin R. L=25 mm
Copyright © Ruslan Panin

Carabus (Aulonocarabus) truncaticollis fleischeri Reitter, 1898
Female adult
Russia, N Burajatia, Barguzinskiy Mts., the upper reaches of M.Chivyrkuy riv., h=2000 m, 26.VI-12.VII.2007, Vashchenko S leg. L=18.5 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) henningi cf. saldaitisi Obydov 1997
Female adult
Russia, S Tuva, E Tanu-Ola Mts., Cha-Ova-Art Pass, h=1700m, 28.VI-3.VII.2000, Vashchenko S. leg. L=20 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) henningi cf saldaitisi Obydov 1997
Male adult
Russia, S Tuva, E Tanu-Ola Mts., Cha-Ova-Art Pass, h=1700m, 28.VI-3.VII.2000, Vashchenko S. leg. L=19.5 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus haeres (trans s. errans)
Female adult
C Ukraine, S of Poltava city, edge forest-lane, 2-26.IX.2010, O. Pozharov leg. /det. R.Panin. L=30 mm
Copyright © Ruslan Panin

Carabus (Aulonocarabus) truncaticollis fleischeri Reitter, 1898
Male adult
Russia, N Burajatia, Barguzinskiy Mts., the upper reaches of M.Chivyrkuy riv., h=2000 m, 26.VI-12.VII.2007, Vashchenko S leg. L=17 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) henningi pelevini Obydov 2006
Female adult
E Kazakhstan, S Altay, Sarym-Sakty Mt Range, 12 km SE Ornek vill. h=1200m, 10.VII.1998, L=21 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) henningi pelevini Obydov 2006a: 99
Male adult
E Kazakhstan, S Altay, Sarym-Sakty Mt Range, 12 km SE Ornek vill. h=1200m, 10.VII.1998, L=20 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) henningi henningi Fischer, 1817
Male adult
Russia, S Siberia, NW Buryatia, E Sayans Mts., Tunka Range, near Mondy vill., h=2400 m, alpine tundra, 1-15.VI.2001, Mishustin R. leg./det. R.Panin
Copyright © Ruslan Panin

Carabus (Eucarabus) ulrichii (as podolicus Semenov, 1898 loc.typ. )
Male adult
CW-UKRAINE, Vinnytsia Distr., Zarvantsi vill., 1-25.V.2020, Kokhan S. leg./det. R.Panin,
Copyright © Ruslan Panin

Carabus (Apotomopterus) patroclus vicarius Cavazzuti, 1996
Male adult
China, S Sichuan, / Yunnan border, W of Lugu lake, S of Yongning vill., h=2700m, 11-18.VI.2017, L=31 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) arrowi ghirettii Cavazzuti, 1995
Female adult
S-China, NE-Guizhou prov., Fanjing Shan, 1800-2000 m, 5-11.VI.14, 27.54°N 108.42°E , L=40 mm
Copyright © Ruslan Panin

Carabus (Isiocarabus) fiduciarius fiduciarius Thomson, 1856
Female adult
CE-China, Hubei, Xiaogan City, Shuangfeng Mountain, 26.VI.2016, L=30 mm
Copyright © Ruslan Panin

Carabus (Carabus) andreiianus Deuve, 2007
Female adult
C-China, N-Sichuan, Pingwu Co. h=1500 m, 27.VI.2011, L=24 mm
Copyright © Ruslan Panin

Carabus (Carabus) koidei koidei Imura, 1996
Male adult
C-China, S-Sichuan, Pass btw. Yanyaun / Muli, h=3244 m, 27.68638N; 101.22335E,,11-18.VI.2017, Ch. Reuter leg. L=22 mm
Copyright © Ruslan Panin

Carabus (Cupreocarabus) mollardianus Deuve, 2016
Female adult
Chine, Sichuan, Lushan, Mt. Jiajin Shan, 10.VII.2016 local collector leg./ det. R.Panin, L=22 mm
Copyright © Ruslan Panin

Carabus (Cupreocarabus) ? siguniangensis jiachingensis Lassalle, 2003
Male adult
Chine, Sichuan, Lushan, Mt. Jiajin Shan, 10.VII.2016 local collector leg., L=21 mm
Copyright © Ruslan Panin

Carabus (Sigenacarabus) trachynodes moxiensis Deuve, 1992
Male adult
China, W-Sichuan, W of Moxi, Daxue Range, Mt.Gongga Shan h=3000-3100m, 29°34′29.6″N /101°59′44.0″E, 10-12.VI.2018, Ch. Reuter leg. L=21 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) ludivinae ludivinae Deuve, 1996
Male adult
S-China, SW-Yunnan: Lincang city, Mt. Daxue, 18.IV.2017, L=29 mm
Copyright © Ruslan Panin

Carabus (Meganebrius) tuberculipennis ganjalaensis Deuve, 2000
Male adult
Nepal, Helambu, GanjaLa Pass S-slope, Dhukpu Kharka h=4100m, 28°03'50"N, 85°34'21"E, 12.VIII.2010, S.Tamang leg., L=21 mm
Copyright © Ruslan Panin

Carabus (s.str.) morphocaraboides zhaojueanus Imura, 1999
Female adult
China, S-Sichuan, SE Mianning, Pass btw Mianshanzhen & Yuexi 28°30’N/102°26’E, 22-23.VI.2017, Ch.Reuter leg., L=22 mm; det. B. Brezina, 2020
Copyright © Ruslan Panin

Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
Male adult
W Ukraine, Transcarpathian Reg., 4 km NE Chop, forest Peresh, Panin R. leg. L=30 mm
Copyright © Ruslan Panin

Carabus (Pagocarabus) crassesculptus eleganticolor Deuve, 1997
Female adult
S Gansu, Min Shan, 5-20 km S Cagan, 2300-3200m 7-20.7.2019 V. Patrikeev leg. (cAA). det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Carabus (Macrothorax) rugosus laufferi Breuning, 1927
Female adult
C-Spain: Avila, El Hornillo env., 27.V.2017, J. Ferrero leg. 30 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) rugosus levantinus Lauffer, 1905
Male adult
SE-Spain, Murcia, Jumillla, Sierra del Carche, 6.VII.2019, F. Garcia leg., L=29 mm
Copyright © Ruslan Panin

Carabus (s.str.) morphocaraboides zhaojueanus Imura, 1999
Male adult
China, S-Sichuan, road 56 km E of Xichang to Zhaojue, Mt.WenLin, Pass h=3200 m, 27°52’N/102°30E, 5-20.VI.2017, Ch.Reuter leg.; det B. Brezina, 2020
Copyright © Ruslan Panin

Carabus (Ctenocarabus) melancholicus submeridionalis Breuning, 1975
Male adult
S- Spain, Cádiz, Tarifa, El Cabrito, 28.XII.2019, F. Garcia leg. L=26 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus helluo (as alicantinus Ganglbauer, 1886)
Male adult
SE-Spain, Murcia reg., Jumilla, Sierra del Carche, h =1300 m, 30.XI.2019, F. Garcia leg.
Copyright © Ruslan Panin

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Male adult
W Ukraine, Lviv city, Sykhiv, Zubra forest, (Osvytskiy Grove), under bark, 49°47'28.10" N, 24° 2'20.09" E, 10.XI.2017, Panin R leg.
Copyright © Ruslan Panin

Carabus (Macrothorax) planatus Chaudoir, 1843
Female adult
S Italy, N Sicily, Nebrodi Mts, Portella di Fémmina Morta h=1500 m, 2.IX.1989, V.Aliquo leg. L=33 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) smaragdinus branickii Taczanowski, 1887
Male adult
N Korea, S. Hamgyong Prov., Mt.Chail Peak, 1-16.VII.2010, l=38 mm
Copyright © Ruslan Panin

Carabus (Pachystus) hortensis preslii Dejean, 1830
Male adult
S-Greece:, N Peloponnes, Erymanthos Reg., Kalentzi, 26.VI.2007, Manfred Egger leg. L=26 mm
Copyright © Ruslan Panin

Carabus (Acoptolabrus) schrencki minpongsanensis Deuve & Li, 2003
Male adult
North Korea, Chagang Prov., Manpo, Songhak-ri (villl.,) 1-16.VII.2011, L=26 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) manifestus guanmenshanus Imura, 1991
Male adult
China, Liaoning, Tieling County, Sandagou env., h=850 m, 13-18.VI.2011
Copyright © Ruslan Panin

Carabus (Orinocarabus) bertolinii Kraatz, 1878
Male adult
NE-Italy, Trentino prov. Dolomites Alps, Plateau Pale di San Martino h=2500 m, 17-19.VIII.2019, M. Egger leg.
Copyright © Ruslan Panin

Carabus (Ohomopterus) dehaanii dehaanii Chaudoir, 1848
Female adult
Japan, S Honshu, Hyogo Pref., Nishiwaki city, 20.VI.2010, Shinichi S. leg., L=37 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) alysidotus Illiger, 1798
Female adult
Italy, Reggio Emilia, Albinea env. h=150 m, VI.2018, A. Cartini leg., L=25 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) billbergi kuandianicus Rapuzzi, 2007
Female adult
NE China, Liaoning, Kuandian County, Dachuantou env., 20-29.VI.2015,
Copyright © Ruslan Panin

Carabus (Eucarabus) xiuyanensis Deuve & Li, 1998
Male adult
NE-China: SE-Liaoning, Xiuyan County, Dayingzi, Mt. Laopingding, 16.VI.2015, L=24.5 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) fruhstorferi Roeschke, 1900
Male adult
Japan, Tsushima Isl., 20.IX.2018, S. Shinichi leg., L=35 mm
Copyright © Ruslan Panin

Carabus (Damaster) blaptoides oxuroides Schaum, 1862
Female adult
Japan, C Honshu, Nagano Prefecture, Nagano city, 3.IV.2012, S. Shinichi leg. L=42 mm. det. Anichtchenko A. 2021
Copyright © Ruslan Panin

Carabus (Eucarabus) xiuyanensis Deuve & Li, 1998
Female adult
NE-China: SE-Liaoning, Xiuyan County, Dayingzi, Mt. Laopingding, 16.VI.2015,
Copyright © Ruslan Panin

Carabus (Euleptocarabus) porrecticollis porrecticollis Bates, 1883
Male adult
Japan, C Honshu, Nagano Pre., Toyooka town, 28.XII.2019, S. Shinichi leg. L=30 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) billbergi kuandianicus Rapuzzi, 2007
Male adult
NE China, Liaoning, Kuandian County, Dachuantou env., 20-29.VI.2015, L=23 mm
Copyright © Ruslan Panin

Carabus (Damaster) blaptoides rugipennis Motschulsky, 1861
Male adult
Kunashir isl.
Copyright © Kirill Makarov

Calosoma (Ctenosta) bastardi Alluaud, 1925
Female adult
S-Madagaskar, Tulear prov., near Betioky town, h = 150 m, 23°38'11" S, 44°20'9" E, 27.XII.2019, leg. A. Kozlov & Yu. Kovaleva
Copyright © Igor Solodovnikov

Calosoma (Ctenosta) grandidieri Maindron, 1900
Male adult
S-Madagaskar, Tulear prov., near Betioky town, h = 150 m, 23°38'11" S, 44°20'9" E, 27.XII.2019, leg. A. Kozlov & Yu. Kovaleva
Copyright © Igor Solodovnikov

Pamborus alternans Latreille, 1817
Female adult
Australia, Queenland state, O’reily, Lamington Nat. Park, XII.2017
Copyright © Igor Solodovnikov

Carabus (Tribax) macropus Chaudoir, 1877
Male adult
Russia, Ingushetia, S slopes of Skalistyi Mt.R., Khahalgi Mt., alpin zone, h = 2500 m, 25.V-19.VII.2018 leg. D.Fominykh
Copyright © Igor Solodovnikov

Carabus (Tribax) macropus Chaudoir, 1877
Female adult
Russia, Ingushetia, S slopes of Skalistyi Mt.R., Khahalgi Mt., alpin zone, h = 2500 m, 25.V-19.VII.2018 leg. D.Fominykh
Copyright © Igor Solodovnikov

Carabus (Tribax) macropus Chaudoir, 1877
Endophallus
Russia, Ingushetia, S slopes of Skalistyi Mt.R., Khahalgi Mt., alpin zone, h = 2500 m
Copyright © Igor Solodovnikov

Carabus (Tribax) steveni schamyli Hampe, 1852
Female adult
C Caucasus, North Ossetia, the road on the village of Coban (Koban), the left river bank Gizeldon, the deciduous wood, h = 895-905 m, N 42°55'32.91", E 44°31'53.53", 20.05-03.07.2017, leg. A.P. Tatulyan
Copyright © Igor Solodovnikov

Carabus (Tribax) steveni schamyli Hampe, 1852
Female adult
C Caucasus, North Ossetia, the road on the village of Coban (Koban), the left river bank Gizeldon, the deciduous wood, h = 895-905 m, N 42°55'32.91", E 44°31'53.53", 20.05-03.07.2017, leg. A.P. Tatulyan
Copyright © Igor Solodovnikov

Carabus (Tribax) steveni schamyli Hampe, 1852
Male adult
C Caucasus, North Ossetia, the road on the village of Coban (Koban), the left river bank Gizeldon, the deciduous wood, h = 895-905 m, N 42°55'32.91", E 44°31'53.53", 20.05-03.07.2017, leg. A.P. Tatulyan
Copyright © Igor Solodovnikov

Carabus (Tribax) steveni schamyli Hampe, 1852
Male adult
C Caucasus, North Ossetia, the road on the village of Coban (Koban), the left river bank Gizeldon, the deciduous wood, h = 895-905 m, N 42°55'32.91", E 44°31'53.53", 20.05-03.07.2017, leg. A.P. Tatulyan
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus theseus Briansky, 1910
Male adult
Russia, Krasnodar Territory, Tuapse district, env. vill. Psebe, left bank of the Psebe River, hornbeam + oak + walnut, h = 146 m, 03.05-01.07.2014, leg. I.A. Solodovnikov, S.V. Solodovnikova, S.M. Pavlyuchuk
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus theseus Briansky, 1910
Female adult
Russia, Krasnodar Territory, Tuapse district, env. vill. Psebe, left bank of the Psebe River, hornbeam + oak + walnut, h = 146 m, 03.05-01.07.2014, leg. I.A. Solodovnikov, S.V. Solodovnikova, S.M. Pavlyuchuk
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
Female adult
Russia, Krasnodar Territory, West-Caucasus, Apsheronsky district, Guam ridge, env. Mezmay village, by the road, beech, oak, blackberry, h = 1200-1250 m, 03.IV-14.VI.2020, leg. S.M. Pavlyuchuk, A.P. Tatulyan
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
Female adult
Russia, Krasnodar Territory, West-Caucasus, Apsheronsky district, Guam ridge, env. Mezmay village, by the road, beech, oak, blackberry, h = 1200-1250 m, 03.IV-14.VI.2020, leg. S.M. Pavlyuchuk, A.P. Tatulyan
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
Male adult
Russia, Krasnodar Territory, West-Caucasus, Apsheronsky district, Guam ridge, env. Mezmay village, by the road, beech, oak, blackberry, h = 1200-1250 m, 03.IV-14.VI.2020, leg. S.M. Pavlyuchuk, A.P. Tatulyan
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
Male adult
Russia, Krasnodar Territory, West-Caucasus, Apsheronsky district, Guam ridge, env. Mezmay village, by the road, beech, oak, blackberry, h = 1200-1250 m, 03.IV-14.VI.2020, leg. S.M. Pavlyuchuk, A.P. Tatulyan
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
Female adult
W Caucasus, NW slope Aibga Mt.R. vall. riv. Galion, h = 650 m, Fagus, Alnus viridis forest, 18-24.VI.1999, I.A. Solodovnikov leg.
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
Male adult
W Caucasus, Aibga Mt.R., 02.VIII.1994, A.S. Zamotailov leg.
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
Male adult
Russia, W Caucasus, Krasnaya Polyana, Atchischkho Mt.R., h = 1200 m, 10.06.1995, N.Yu. Pichugin leg.
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus adelaidae Starck, 1889
Female adult
Russia, Krasnodar area, Adler, Ader distr., right bank vall. Riv. Msymta, near vill. Chvesipse, h = 800 m, 06.VII.1999, B. Saratovskikh leg.
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus adelaidae Starck, 1889
Female adult
Russia, Krasnodar area, Adler, Ader distr., right bank vall. Riv. Msymta, near vill. Chvesipse (Fagus+Quercus+Castanea forest) [Sochi national park], h = 720 m, 03.V-08.VII.2016, I.A. Solodovnikov, S.V.Solodovnikova, V.M.Kotsur leg.
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus adelaidae Starck, 1889
Female adult
Russia, Krasnodar area, Adler, Ader distr., right bank vall. Riv. Msymta, near vill. Chvesipse (Fagus+Quercus+Castanea forest) [Sochi national park], h = 720 m, 03.V-08.VII.2016, I.A. Solodovnikov, S.V.Solodovnikova, V.M.Kotsur leg.
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus adelaidae Starck, 1889
Male adult
Russia, Krasnodar area, Adler, Ader distr., right bank vall. Riv. Msymta, near vill. Chvesipse (Fagus+Quercus+Castanea forest) [Sochi national park], h = 720 m, 03.V-08.VII.2016, I.A. Solodovnikov, S.V.Solodovnikova, V.M.Kotsur leg.
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) boeberi boeberi Adams, 1817
Female adult
Грузия, Мцхета-Мтианети, к северу от пос. Гудаури, ЮВ отроги г. Байдарская (нижний пик), по ущелью ручья, альпика, под камнями, точка 2, h = 2700 м, 42.495413° N, 44.487844° E, 10.08.2019, leg. И.А., А.И., С.В. Солодовниковы, О.И. Данилова 26 mm
Copyright © Igor Solodovnikov

Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993
Male adult
Uzbekistan, Chatkal Mt. rng., Irtash, 41°10'25.7"N 70°16'20.4"E, 5-7.8.2021, 2400m, A. Anichtchenko leg. (cAA). 27mm
Copyright © Alexander Anichtchenko

Carabus (Pantophyrtus) longipedatus Belousov & Kabak, 1993
Female adult
Uzbekistan, Chatkal Mt. rng., Irtash, 41°10'25.7"N 70°16'20.4"E, 5-7.8.2021, 2400m, A. Anichtchenko leg. (cAA). 30 mm
Copyright © Alexander Anichtchenko

Carabus (Cechenochilus) boeberi tschitscherini Semenov, 1896
Male adult
Russia, C Caucasus, Ingushetia, S slopes of Skalistyi Mt.R.
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) boeberi tschitscherini Semenov, 1896
Female adult
Russia, C Caucasus, Ingushetia, S slopes of Skalistyi Mt.R.
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) heydenianus Starck, 1889
Male adult
Carabus (Procechenochilus) heydenianus euxinus Semenov, 1896 Abkhasia: Arabika plato (TOPOTYPUS)
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) heydenianus Starck, 1889
Female adult
Carabus (Procechenochilus) heydenianus euxinus Semenov, 1896 Abkhasia: Arabika plato (TOPOTYPUS)
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) boeberi aequaliceps Reitter, 1896
Male adult
Russia, Dagestan, Andiyskiy Mt.R., near Andiyskiye vorota pass, 06.VI–18.VII.2021, leg. D. Fominykh
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) boeberi aequaliceps Reitter, 1896
Female adult
Russia, Dagestan, Andiyskiy Mt.R., near Andiyskiye vorota pass, 06.VI–18.VII.2021, leg. D. Fominykh
Copyright © Igor Solodovnikov

Carabus (Pachystus) hortensis hortensis Linne, 1758
Female adult
Natural hybrid Carabus hortensis X Carabus granulatus. Belarus, Minsk reg., 8,7 km of Borisov, near vill. Lyubatovstchina, pine forest (Pinetum pleuroziosum), h = 213 m, 54.31934° N, 28.511486° E, 20-27.09.2018, leg. Klychenok Snezhana
Copyright © Igor Solodovnikov

Carabus (Megodontus) obovatus Fischer, 1827
Male adult
C.obovatus taizhensis Obydov, 2006 Khakassia: Kuznetskyi Alatau, env. vill. Priiskovyi, Bobrovaya Mt., h = 1200-1300 m
Copyright © Igor Solodovnikov

Carabus (Megodontus) obovatus Fischer, 1827
Endophallus
C.obovatus taizhensis Obydov, 2006 Khakassia: Kuznetskyi Alatau, env. vill. Priiskovyi, Bobrovaya Mt., h = 1200-1300 m
Copyright © Igor Solodovnikov

Carabus (Acoptolabrus) gehinii munakatai (Ishikawa, 1968)
Male adult
Japan, SW-Hokkaido, Yakumo town, 4.VI.2017, S.Shinichi leg., L=30 mm
Copyright © Ruslan Panin

Carabus (Damaster) blaptoides blaptoides (Kollar, 1836)
Female adult
Japan, SW Honshu, Shimane Pref., Unnan city, 16.I2016, S.Shinichi leg., L=48 mm
Copyright © Ruslan Panin

Carabus (Megodontus) kolbei taikinus Ishikawa, 1993
Female adult
Japan, SC-Hokkaido:Tokachi, Hiroo Distr., Taiki-cho env., 3-7.IX.2019, S.Shinichi leg.
Copyright © Ruslan Panin

Carabus (Megodontus) kolbei kosugei Nakane, 1955
Male adult
Japan, SW-Hokkaido, Otaru city, 2-16.VIII.2020, S.Shinichi leg.
Copyright © Ruslan Panin

Carabus (Megodontus) kolbei hidakamontanus Ishikawa, 1966
Male adult
SE-Hokkaido: Samani distr.
Copyright © Igor Solodovnikov

Carabus (Megodontus) kolbei hidakamontanus Ishikawa, 1966
Male adult
E-Hokkaido: Samani distr., Horomankyo env.
Copyright © Igor Solodovnikov

Carabus (Megodontus) kolbei hidakamontanus Ishikawa, 1966
Female adult
E-Hokkaido: Samani distr., Horomankyo env.
Copyright © Igor Solodovnikov

Carabus (Megodontus) kolbei hidakamontanus Ishikawa, 1966
Endophallus
E-Hokkaido: Samani distr., Horomankyo env.
Copyright © Igor Solodovnikov

Carabus (Ohomopterus) dehaanii dehaanii Chaudoir, 1848
Male adult
Japan, S Honshu, Mie Pref., Inabe city, 21.II.2021, Shinichi S. leg., L=33 mm
Copyright © Ruslan Panin

Carabus (Tribax) nothus sharoiensis Fominykh, Zamotajlov, Avtaeva et Titarenko, 2019 Paratype
Male adult

Copyright © Igor Solodovnikov

Carabus (Neoplectes) ibericus lafertei Chaudoir, 1846
Female adult
SC-Georgia, Samtskhe–Javakheti, Meskheti Mt Range, Borjomi, Likani env., h=897 m, 41°50.125'N 43°19.905'E, VI.2018, D. Prunier leg., L=37 mm
Copyright © Ruslan Panin

Carabus (Procrustes) coriaceus ressli Mandl, 1961
Male adult
SC-Turkey: Hatay Prov, Dörtyol, Nur Mts, Topaktaş h=900-1200 m, 12.V.2005, D. Prunier leg.
Copyright © Ruslan Panin

Carabus (Procrustes) coriaceus ressli Mandl, 1961
Female adult
SC-Turkey: Hatay Prov, Dörtyol, Nur Mts, Topaktaş h=900-1200 m, 12.V.2005, D. Prunier leg. L=35 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus antiquus Dejean, 1826
Female adult
SW Portugal, Beja Distr., 3,7 km NW Brunheiras, road CM1072, 37°46'31.9"N 8°47'02.2"W, 15.III.2011, D. Prunier leg., L=26 mm
Copyright © Ruslan Panin

Carabus (Tribax) nothus fausti Dohrn, 1873
Male adult
Carabus (Microtribax) fausti fausti Dohrn, 1873 Russia: Daghestan, Kyabyak Mt.R.
Copyright © Igor Solodovnikov

Carabus (Tribax) nothus fausti Dohrn, 1873
Female adult
Carabus (Microtribax) fausti fausti Dohrn, 1873 Russia: Daghestan, Kyabyak Mt.R.
Copyright © Igor Solodovnikov

Carabus (Diocarabus) fraterculus neochinensis Deuve & Li, 1998
Female adult
NE-China: Liaoning prov., Fengcheng City, Shicheng, 20-28.VI.2011, Li Jingke leg., L=22 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) protenes baoxingensis Deuve & Mourzine, 2003
Male adult

Copyright © Igor Solodovnikov

Carabus (Apotomopterus) protenes baoxingensis Deuve & Mourzine, 2003
Female adult

Copyright © Igor Solodovnikov

Carabus (Apotomopterus) protenes baoxingensis Deuve & Mourzine, 2003
Endophallus
China: С-Sichuan:Qionglai Shan Mts., Baoxing env. (TOPOTYPUS)
Copyright © Igor Solodovnikov

Carabus (Tribax) lederi lederi Reitter, 1882
Male adult
Georgia, Kvemo-Svaneti, Svanetian Mt.R., near Latpari pass., alpin zone, 2700-3000m (TOPOTYPUS)
Copyright © Igor Solodovnikov

Carabus (Tribax) lederi lederi Reitter, 1882
Female adult
Georgia, Kvemo-Svaneti, Svanetian Mt.R., near Latpari pass., alpin zone, 2700-3000m (TOPOTYPUS)
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum kasakorum Semenov, 1896
Male adult
Russia: N.Osetia: Pastbyshchniy Mt.R., S slopes of Khoshkharanrag Mt.R., right bank of Amusadon riv., h=1500 m
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum kasakorum Semenov, 1896
Female adult
Russia: N.Osetia: Pastbyshchniy Mt.R., S slopes of Khoshkharanrag Mt.R., right bank of Amusadon riv., h=1500 m
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Male adult
Russia, Caucasus, Ingushetia, South slopes of Skalistyi Mt Range, Stolovaya Mt., near Beyni, alpin zone, h = 2800 m, N 42°51’35.30",E 44°42’13.70", h = 2800m, 21.V–19.VII.2018, D. Fominykh leg.
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Female adult
Russia, Caucasus, Ingushetia, South slopes of Skalistyi Mt Range, Stolovaya Mt., near Beyni, alpin zone, h = 2800 m, N 42°51’35.30",E 44°42’13.70", h = 2800m, 21.V–19.VII.2018, D. Fominykh leg.
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Endophallus
Russia, Caucasus, Ingushetia, South slopes of Skalistyi Mt Range, Stolovaya Mt., near Beyni, alpin zone, h = 2800 m, N 42°51’35.30",E 44°42’13.70", h = 2800m, 21.V–19.VII.2018, D. Fominykh leg. (TOPOTYPUS)
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Male adult
Russia, Caucasus, Ingushetia, Skalistyi Mt. Range, S slopes of Tsoreylam Mt. Range, 5 km W of Mt. Khakhalgi, N 42°50’07.60", E 45°05’02.20", h=2500 m, 25.V–21.VII.2017, D. Fominykh leg.
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Endophallus
Russia, Caucasus, Ingushetia, Skalistyi Mt. Range, S slopes of Tsoreylam Mt. Range, 5 km W of Mt. Khakhalgi, N 42°50’07.60", E 45°05’02.20", h=2500 m, 25.V–21.VII.2017, D. Fominykh leg.
Copyright © Igor Solodovnikov

Carabus (Macrothorax) rugosus celtibericus Germar, 1824
Male adult
SW-Portugal, Serra do Cercal h=217 m, 37°47'15.6"N 8°42'02.0"W, road N390, 19.III.2011, D. Prunier leg., L=27 mm
Copyright © Ruslan Panin

Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Female adult
Russia, Chechnya, Caucasus, Skalystyi Mt. Range upper river Gekhi, Mt. Ekkyrkort, N 42°52', E 45°26', h=2230 m, 26.V–21.VII.2018, D.Fominykh leg.
Copyright © Igor Solodovnikov

Carabus (Diocarabus) fraterculus neochinensis Deuve & Li, 1998
Male adult
NE-China: Liaoning prov., Fengcheng City, Shicheng, 20-28.VI.2011, Li Jingke leg., L=20 mm
Copyright © Ruslan Panin

Carabus (Ctenocarabus) melancholicus costatus Germar, 1824
Male adult
NW Portugal, Viana do Castelo distr., Sierra de Agra, 41°47'52.3"N 8°41'27.6"W, h=786 m, 3.VII.2016, D. Prunier leg.
Copyright © Ruslan Panin

Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
Female adult
Russia: N.Osetia, Bokovoy Mt.R., near Arkhonskiy pass., alpin zone, h = 2700 m, 23.V-21VII.2018, D. Fominykh leg.
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
Male adult
Russia: N.Osetia, Bokovoy Mt.R., near Arkhonskiy pass., alpin zone, h = 2700 m, 23.V-21VII.2018, D. Fominykh leg.
Copyright © Igor Solodovnikov

Carabus (Tribax) puschkini kolenatii Chaudoir, 1846
Female adult
C-Georgia, Trialeti Mt Range, 2 km SE of Borjomi h=916 m, 41°48'35.3"N 43°26'11.8"E, 17.VI.2013, D. Prunier leg./det. Panin R., L=29 mm
Copyright © Ruslan Panin

Carabus (Tribax) puschkini kolenatii Chaudoir, 1846
Male adult
C-Georgia, Trialeti Mt Range, 2 km SE of Borjomi h=916 m, 41°48'35.3"N 43°26'11.8"E, 17.VI.2013, D. Prunier leg./det. Panin R., L=28 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus trabuccarius Fairmaire, 1857
Male adult
NE Spain, Catalonia, Tarragona, Sierra de Montsant h=800 m, Albarca env.,7.XI.2018, T. Rafales leg., L=25 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus trabuccarius Fairmaire, 1857
Female adult
NE Spain, Catalonia, Tarragona, Sierra de Montsant h=800 m, Albarca env.,7.XI.2018, T. Rafales leg., L=27 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus lusitanicus (as vieirae Oliveira, 1875)
Female adult
CW-Portugal: Leiria distr., near Marinha Grande, 39°45'57.8"N 8°58'10.3"W, h-79 m, 19.IX.2016, D. Prunir leg., L=26 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens (as lomnitzkii Reitter, 1896)
Male adult
W Ukraine, NW Podolian Upland, Lviv reg., Zolochiv distr., Lysa Gora steppe massif, 11.VII-14.VIII. 2021, Panin R. & D. Khrapov leg., L=25 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus lusitanicus (as vieirae Oliveira, 1875)
Male adult
CW-Portugal: Leiria distr., near Marinha Grande, 39°45'57.8"N 8°58'10.3"W, h-79 m, 19.IX.2016, D. Prunir leg., L=24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens (as lomnitzkii Reitter, 1896)
Male adult
W Ukraine, NW Podolian Upland, Lviv reg., Zolochiv distr., Lysa Gora steppe massif, 11.VII-14.VIII. 2021, Panin R. & D. Khrapov leg., L=25 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens (as lomnitzkii Reitter, 1896)
Female adult
W Ukraina, Podolian Upland, steppe reserve Kasova Gora near Burshtyn vill., 15.VI -28.VII.2020, R. Panin & D. Khrapov leg., L=27 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens (as lomnitzkii Reitter, 1896)
Male adult
W Ukraina, Podolian Upland, steppe reserve Kasova Gora near Burshtyn vill., 15.VI -28.VII.2020, R. Panin & D. Khrapov leg., L=24 mm
Copyright © Ruslan Panin

Carabus (Relictocarabus) meurguesianus Ledoux, 1990
Male adult
Image from molbiol.ru
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Carabus (Shunichiocarabus) uenoianus uenoianus Imura, 1995
Male adult

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Calosoma (Callistenia) monticola Casey, 1913
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH. Photo processed by Anichtchenko A. 2022
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Calosoma (Calopachys) omiltemium Bates, 1891
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH. Photo processed by Anichtchenko A. 2022
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Carabus (Trachycarabus) scabriusculus lippii Dejean, 1826
Female adult
SW Ukraine, Odessa reg., Ivanovka distr., steppe ravine near Gudevichevo vill., 22.IV-5.V.2022, A.Gontarenko leg., L=22 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) scabriusculus lippii Dejean, 1826
Male adult
SW Ukraine, Odessa reg., Ivanovka distr., steppe ravine near Gudevichevo vill., 22.IV-5.V.2022, A.Gontarenko leg., L=21.5 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii zawadzkii, Kraatz 1854
Male adult
W Ukraine, NE Carpathians, Lviv region, Skole Beskids Mts, Zelemyanka Mt. Range h=915 m, 22.VI-17.VII.2022, Panin R. & Kanarsky Yu. leg., L= 32 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) wiedemanni congruus Motschulsky, 1865
Female adult
Turkey, Cankiri, Cerkes - Kursunlu, Atkaracalar 1250m 10.7.1996 Leg Staven and Skoupy (DUBC)
Copyright © Alexander Anichtchenko

Carabus (Procrustes) chevrolati internatus Heinz & Korge, 1964
Female adult
Turkey, Bolu, Gerede, Esentepe 1350m, 1.7.1994, leg. K. Staven
Copyright © Alexander Anichtchenko

Carabus (Archicarabus) wiedemanni wiedemanni (as mussardianus Breuning, 1965)
Female adult
NC-Turkey, Sinop province, Erfelek env., h=200m, 10-27.VII.2014, C. Reuter leg. L=24 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) wiedemanni (as mussardianus Breuning, 1965)
Male adult
NC-Turkey, Sinop province, Erfelek env., h=200m, 10-27.VII.2014, C. Reuter leg. L=23 mm
Copyright © Ruslan Panin

Carabus (Cechenochilus) boeberi felix Semenov & Znojko, 1931
Male adult

Copyright © Kirill Makarov

Calosoma (Charmosta) investigator Illiger, 1798
Male adult

Copyright © Kirill Makarov

Carabus (Neoplectes) reitteri reitteri Retowski, 1885
Male adult

Copyright © Kirill Makarov

Carabus (Pachystus) glabratus glabratus Paykull, 1790
Male adult

Copyright © Kirill Makarov

Carabus (Pachycarabus) koenigi Ganglbauer, 1886
Male adult

Copyright © Kirill Makarov

Carabus (Pachycarabus) staehlini staehlini Adams, 1817
Male adult

Copyright © Kirill Makarov

Carabus (Platycarabus) irregularis ramanus Sokolar, 1909
Female adult
NW Croatia, Ivanščica Mts , Ivanec, 4-18.VI.2011, Krause Rene leg., L=33 mm
Copyright © Ruslan Panin

Calosoma (Callitropa) macrum LeConte, 1853
Female adult
Texas Live Oak Co. 20.9.2016 (cAA)
Copyright © Alexander Anichtchenko

Calosoma (Carabosoma) marginalis Casey, 1897
Female adult
Texas Live Oak Co. 20.9.2016 (cAA)
Copyright © Alexander Anichtchenko

Carabus (Sphodristocarabus) varians janthinus Ganglbauer, 1887
Female adult
Carabus (Sphrodistocarabus) janthinus dvorschaki Mandl, 1975 Abkhazia, Gudautsky district, S- Bzybsky Ridge, a watershed hill at the head of the river Khipsta, 43°17'56.63" N, 40°40'16.74" E, h= 1490 m, mature beech-maple forest with butterbur, moist soil, 05.07-17.08.2017, leg. I.A. Solodovnikov, S.V. Solodovnikova, E.V. Tatun
Copyright © Igor Solodovnikov

Carabus (Trachycarabus) planus retowskianus Mandl, 1955
Male adult
S Ukraine, Zaporizhia reg., Melitopol distr., near Troitskoe vill., 1.VI.2010, I.V. Kovalev leg., L=27 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) estreicheri Fischer, 1822 (ab.rufofemoratus Lomnicki, 1892)
Female adult
W Ukraina, Podolian Upland, Lviv reg., Brody distr., Mt. Makitra, steppe site, 19.V-5.VI.2020, Panin R. leg. L=23 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus campicola Dejean, 1831
Female adult
S Ukraine, Crimea, S slope Mt Demerdzhi h=650 m, sibljak, VII.2004, R. Panin leg., L=29 mm
Copyright © Ruslan Panin

Carabus (Pachystus) glabratus gibbosus Heyden, 1866
Male adult
N Croatia, Ivanščica Mts, Ivanec, 4-18.VI.2011, Krause Rene leg., L=32 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus cf putus Motschulsky, 1846
Female adult
Russia, SW Siberia, C-Altays Mts, Kuray Mt Rng NE Aktash vill., h=2100-3000 m, 8-13.VII.2022, E.Filippov leg./det.R Panin, L= 22 mm (alpine petrophilic population close to the form schestopalovi Plutenko, 1995)
Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus cf putus Motschulsky, 1846
Male adult
Russia, SW Siberia, C-Altays Mts, Kuray Mt Rng NE Aktash vill., h=2100-3000 m, 8-13.VII.2022, E.Filippov leg./det.R Panin, L= 23 mm (alpine petrophilic population close to the form schestopalovi Plutenko, 1995)
Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus putus Motschulsky, 1846 (loc.typ.)
Female adult
Russia, W Altay, 18 km N of Zmeinogorsk, 23.VI.1998, A.& N. Menstschikovy leg. L=24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus putus Motschulsky, 1846 (loc.typ.)
Male adult
Russia, W Altay, 18 km N of Zmeinogorsk, 23.VI.1998, A.& N. Menstschikovy leg. L=23 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) odoratus antropovi Shilenkov 1996
Female adult
Russia, S Siberia, Tuva, W Tannu-Ola Mts. 35 km NE Khandagayty vill., Teeli Pass 2200-2800 m, 23-25.VI.2021, A.& E.Filippovs leg. L=19 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) odoratus antropovi Shilenkov 1996
Male adult
Russia, S Siberia, Tuva, W Tannu-Ola Mts. 35 km NE Khandagayty vill., Teeli Pass 2200-2800 m, 23-25.VI.2021, A.& E.Filippovs leg. L=19 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Male adult
SE Poland, Subcarpathia Prov., Nadleśnictwo Kołaczyce N of Jasło vill. M. Kosibowicz leg./det. R Panin, VII.2016, L=29 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Male adult
SE Poland, N Carpathians, Subcarpathia Prov., Nadleśnictwo Baligród, (209a) 22.VII.2021, M.Kosibowicz leg./det. R.Panin, L=31 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Male adult
SE Poland, N Carpathians, Subcarpathia Prov., National Park Magura, VII.2016, M.Ranocha leg./ det. R. Panin, L=30 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Male adult
SE Poland, N Carpathians, Subcarpathia Prov., Nadleśnictwo Lesko, VII.2017 M.Ranocha leg./ det. R. Panin, L=31 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
Male adult
Russia, NE Altay, Turochak Distr., Lake Teletskoe N part, near Yaylyu vill. 23-25.VII.2019, Ivliev P. leg./ det. R. Panin, L=24 mm
Copyright © Ruslan Panin

Calosoma (Callistenia) luxatus Say, 1823 (zimmermanni LeConte, 1848)
Female adult
USA OR, Jackson Co., FR300, 0.18mi W of rd. 20, 42.02.342N 122.54.937, 20.6.2021 M. Kippenhan leg. (cAA) 15.5 mm; det. A. Anichtchenko 2023
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) aeruginosus eschscholtzi Mannerheim, 1827
Female adult
NE Kazakhstan, W Altay Mts, near Shemonaikha city., flood plain, 25-28.VII.2018, S.Vashchenko leg. L=25 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) aeruginosus eschscholtzi Mannerheim, 1827
Male adult
NE Kazakhstan, W Altay Mts, near Shemonaikha city., flood plain, 25-28.VII.2018, S.Vashchenko leg., L=22.5 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792
Male adult
C-Poland, Łódź sity, Łagiewniki Forest, in dead logs of Birch, 18-30.XII.2013, D. Nawak leg., L=25 mm
Copyright © Ruslan Panin

Carabus (Megodontus) leachi leachi Fischer, 1823
Female adult
NE Kazakhstan, W Altay, near Shemonaikha city., flood plain, 18-28.VII.2018, S.Vashchenko leg., L=22.5 mm
Copyright © Ruslan Panin

Carabus (Megodontus) leachi leachi Fischer, 1823
Male adult
NE Kazakhstan, W Altay, near Shemonaikha city., flood plain, 18-28.VII.2018, S.Vashchenko leg., L=21.5 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
Male adult
Russia, W Sayan Mts., Kazyr-Suk river, W slope of Oyskiy Mt. rng., ?h=2200 m 7-15.VII.1999, Brinev A. leg., L=19 mm
Copyright © Ruslan Panin

Calosoma (Chrysostigma) calidus Fabricius, 1775
Female adult
Canada, Manitoba, 3km S Aweme, 12.6.2010 (DUBC)
Copyright © Alexander Anichtchenko

Calosoma (Castrida) argentinensis Csiki, 1927
Female adult
(DUBC)
Copyright © Alexander Anichtchenko

Carabus (Eucarabus) parreyssi parreyssi Palliardi, 1825
Female adult
SW-Croatia, Mts Velebit, Mali Halan, h=1390 m, 23.V-20.VI.2022, L=24 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) parreyssi parreyssi Palliardi, 1825
Endophallus
SW-Croatia, Mts Velebit, Mali Halan, h=1390 m,
Copyright © Ruslan Panin

Carabus (Eucarabus) parreyssi parreyssi Palliardi, 1825
Male adult
SW-Croatia, Mts Velebit, Mali Halan, h=1390 m, 23.V-20.VI.2022, L=22.5 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) monticola monticola (as maritimensis Born, 1923)
Female adult
SE France, Alpes-Maritimes, Col de Braus h=1000 m, 23.V-18.VI.2014, P. Cavazzuti & G. Proscia leg., L= 24 mm
Copyright © Ruslan Panin

Carabus (Pachystus) cavernosus variolatus Costa, 1839
Female adult
C-Italy: Abruzzo reg., Gran Sasso massif, Campo Imperatore, h=1900 m, VIII.2021, G.Proscia leg., L=27 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) montivagus vellepiticus Hampe, 1850
Female adult
W Croatia, NP Northern Velebit, Stolak, 44.969492, 14.986169, 23.V-27.VI.2022, L=24 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) montivagus vellepiticus Hampe, 1850
Male adult
W Croatia, NP Northern Velebit, Stolak, 44.969492, 14.986169, 23.V-27.VI.2022, L=23.5 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) solieri bonnetianus Colas, 1936 (loc.typ.)
Female adult
SE-France, Var, Tanneron, Saint-Cassien des Bois, 18.IV.2019, L=32 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) solieri bonnetianus Colas, 1936 (loc.typ.)
Male adult
SE-France, Var, Tanneron, Saint-Cassien des Bois, 18.IV.2019, L= 31 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) solieri clairi Gehin, 1885 (as devillei Tarrier, 1972)
Male adult
SE-France, Alpes-Maritimes, Beuil env., h=1550m, 14.VII.1986
Copyright © Ruslan Panin

Carabus (Procrustes) coriaceus excavatus Charpentier, 1825
Male adult
W Croatia, Velebit Mts, Jablanac Stirovaca h=998 m, 44.71124, 14.95673,3.VI-7.VII.2017, G. Proscia leg., L= 37 mm
Copyright © Ruslan Panin

Carabus (Procrustes) coriaceus excavatus Charpentier, 1825
Female adult
W Croatia, Velebit Mts, Jablanac Stirovaca h=998 m, 44.71124, 14.95673,3.VI-7.VII.2017, G. Proscia leg., L= 38 mm
Copyright © Ruslan Panin

Carabus (Megodontus) croaticus croaticus Dejean, 1826
Female adult
W-Croatia, Velebit Mts., h=950 m, Oltari - Zavizan, 30.VI. 2018, L=32 mm
Copyright © Ruslan Panin

Carabus (Megodontus) croaticus croaticus Dejean, 1826
Male adult
W-Croatia, Velebit Mts., h=950 m, Oltari - Zavizan, 30.VI.2018
Copyright © Ruslan Panin

Carabus (Chaetocarabus) intricatus bayardi (as silaensis Haury, 1881)
Endophallus
S-Italy, Calabria, Sila Grande Massif, Camigliatello h=1040 m, 19.XI.2009, G. Proscia leg. L=31 mm
Copyright © Ruslan Panin

Carabus (Chaetocarabus) intricatus bayardi (as silaensis Haury, 1881)
Male adult
S-Italy, Calabria, Sila Grande Massif, Camigliatello h=1040 m, 19.XI.2009, G. Proscia leg. L=28 mm
Copyright © Ruslan Panin

Carabus (Cupreocarabus) balangicus balangicus Cavazzuti, 2002
Male adult
Chine, C-Sichuan, Xiaojin Co, Wolong valley, Balang Shan Pass h=3448 m, 30.884617, 102.973117, 9.VI.-1.VII.2011, G. Proscia leg. L=19 mm
Copyright © Ruslan Panin

Carabus (Cupreocarabus) balangicus balangicus Cavazzuti, 2002
Male adult
Chine, C-Sichuan, Xiaojin Co, Wolong valley, Balang Shan Pass h=3448 m, 30.884617, 102.973117, 9.VI.-1.VII.2011, G. Proscia leg. L=20 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) cateniger cateniger (as sommitalis Cavazzuti, 2006)
Female adult
China, C-Gansu, Mts Qilian h=3378 m, road G227, Yangxiongzi env. 38.050917, 100.887267, 18-23.VI.2011, G.Proscia leg., L=19 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) cateniger cateniger (as sommitalis Cavazzuti, 2006)
Male adult
China, C-Gansu, Mts Qilian h=3378 m, road G227, Yangxiongzi env. 38.050917, 100.887267, 18-23.VI.2011, G.Proscia leg., L=17 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) cateniger dabanicola Deuve, 1998
Male adult
China, Qinghai, Xining city, 55km NE of Datong, Daban Shan h=3936 m, 37.348450, 101.397600, 17-23.VI.2011, G.Proscia leg., L= 16 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) cateniger dabanicola Deuve, 1998
Female adult
China, Qinghai, Xining city, 55km NE of Datong, Daban Shan h=3936 m, 37.348450, 101.397600, 17-23.VI.2011, G.Proscia leg., L= 18.5 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) roborowskii roborowskii Semenov, 1887
Male adult
China, E-Qinghai, Mts NW of Tongren city, 35.562778, 101.843667, h=3870 m, 15-26.VI.2011, G. Proscia leg./det.R.Panin, L=20 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) roborowskii roborowskii Semenov, 1887
Female adult
China, E-Qinghai, Mts NW of Tongren city, 35.562778, 101.843667, h=3870 m, 15-26.VI.2011, G. Pros cia leg./det.R.Panin, L=20 mm
Copyright © Ruslan Panin

Carabus (Cupreocarabus) huangi huangi Deuve, 1992
Male adult
China, E Tibet, N-Sichuan, 53 km NE of Barkam, Pass Sandaoban env., h=4293 m, 32.232950, 102.598533, 11-29.VI.2011, G. Proscia leg., L=21 mm
Copyright © Ruslan Panin

Carabus (Cupreocarabus) huangi huangi Deuve, 1992
Female adult
China, E Tibet, N-Sichuan, 53 km NE of Barkam, Pass Sandaoban env., h=4293 m, 32.232950, 102.598533, 11-29.VI.2011, G. Proscia leg., L=21.5 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) catenulatus fontanellae Reitter, 1896
Male adult
W-Croatia, Dalmatia, Isl. Rab, 16-30.V.2019, G. Proscia leg.
Copyright © Ruslan Panin

Carabus (Trachycarabus) scabriusculus inapertus Motschulsky, 1850
Male adult
NC Ukraine, Kiev city, Lysa Gora, 1-15.VI.2012, Gerasimov R.leg./det. R.Panin, L=19 mm
Copyright © Ruslan Panin

Carabus (Morvanicarabus) scheibei scheibei Eidam, 1937
Female adult
NE-Afghanistan, Hindu Kush, Nuristan Prov., near Parun, 35°19'N 70°52'E h=2900 m, Pine forest, VI.2022, local collector/det. Kleinfeld, L= 24 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) riffensis Fairmaire, 1875
Female adult
N-Morocco, Riff Mts h=1650 m, 26 km E of Bab Berred, 34.961333, -4.669417, Cedar forest, C. Reuter leg. L=33 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) riffensis Fairmaire, 1875
Endophallus
N-Morocco, Riff Mts h=1650 m, 26 km E of Bab Berred
Copyright © Ruslan Panin

Carabus (Mesocarabus) riffensis Fairmaire, 1875
Male adult
N-Morocco, Riff Mts h=1650 m, 26 km E of Bab Berred, 34.961333, -4.669417, Cedar forest, C. Reuter leg. L=27 mm
Copyright © Ruslan Panin

Carabus (Isiocarabus) miaorum Lassalle & Prunier, 1993
Male adult
China, NW-Guangxi, Mao'er Shan, h~1600 m 25°52'N 110°29'E, 28.V-9.VI.2009, C.Reuter leg.
Copyright © Ruslan Panin

Carabus (Apotomopterus) delavayi parvipes Cavazzuti, 1996
Endophallus
S-Sichuan: S of Yanyuan, Chuandongzi Massif h=3200 m, 27.351206, 101.507036
Copyright © Ruslan Panin

Carabus (Apotomopterus) delavayi parvipes Cavazzuti, 1996
Male adult
S-Sichuan: S of Yanyuan, Chuandongzi Massif h=3200 m, 27.351206, 101.507036, 8-19.VI.2017, C.Reuter leg., L=32 mm
Copyright © Ruslan Panin

Carabus (Imaibius) caschmirensis fortetuberculatus Heinz, 1983 Paratype
Male adult

Copyright © Alexander Anichtchenko

Carabus (Lamprostus) syrus moukarzelae Deuve, 2002
Female adult
S-Lebanon: Marjaayoun, 18.XI.-11.XII.2017, C. Reuter leg., L=35 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) protenes ascendens Semenov, 1898
Male adult
China, Chongqing, N Jinfo Shan, Shanwangping h=1400m, 29.101944, 107.166389, 2-8.VI.2018, C.Reuter leg./det. F.Kleinfeld, L=26 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) protenes ascendens Semenov, 1898
Endophallus
China, Chongqing, N Jinfo Shan, Shanwangping h=1400m, 29.101944, 107.166389
Copyright © Ruslan Panin

Carabus (Apotomopterus) laoshanicus Imura, 1995
Male adult
China, E Guangxi, Jinxiu Yao County, Tongmu vill., local collector, L=36 mm
Copyright © Ruslan Panin

Carabus (Imaibius) nouristani nouristani (Ledoux, 1977)
Female adult
NE-Afghanistan: Nuristan: Darah-I-Nur valley, 2900m 34°45'N 70°40'E, VIII.2012, C.Reuter leg., L=26 mm
Copyright © Ruslan Panin

Carabus (Pachystus) cribratus cribratus Quensel, 1806
Male adult
NW Armenia, Amasia, (Shirak prov.), 20-24.VIII.1998, Kazaryan P.V. leg.
Copyright © Ruslan Panin

Carabus (Apotomopterus) korellianus Kleinfeld, 2002
Male adult
China, SW-Yunnan, Dehong Distr., Ruili, Pijiazhai vill., h=1800 m, 24.048111, 97.738222, 9-12.VI.2023, C.Reuter leg., L=31 mm
Copyright © Ruslan Panin

Carabus (Imaibius) nouristani nouristani (Ledoux, 1977)
Male adult
NE-Afghanistan: Nuristan: Darah-I-Nur valley, 2900m 34°45'N 70°40'E, VIII.2012, C.Reuter leg., L=26 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) madefactus Cavazzuti, 1997
Male adult
China, CW-Yunnan, Nánjiàn Yízú Zìzhìxiàn, Mt Lingbao h=2400 m, 24.768611, 100.500000, 6-15.VI.2023, C.Reuter leg./det. F.Kleinfeld, L=27 mm
Copyright © Ruslan Panin

Carabus (Sigenacarabus) trachynodes moxiensis Deuve, 1992
Male adult
China, W-Sichuan, Daxue Range, Mt.Gongga Shan h=3000-3100m, Hailuogou Glacier Forest Park, 29.574889, 101.995556, 10-12.VI.2018, Ch. Reuter leg., L=21 mm
Copyright © Ruslan Panin

Carabus (Acoptopterus, ?Carabus) yunnanus yanyuanicus Cavazzuti, 1996
Endophallus
China, S-Sichuan, S Yanyuan, Chuandongzi Mt h=3300-3600 m
Copyright © Ruslan Panin

Carabus (Apotomopterus) mecynodes pinchuanensis Imura, 1999
Endophallus
China, S-Sichuan, 40-50 km E Xichang (to Zhaojue) 27°51'N 102°30'E, h=2700-3000m, Pine forest
Copyright © Ruslan Panin

Carabus (Acoptopterus) yunnanus yanyuanicus Cavazzuti, 1996 (? or oblongior ssp.)
Male adult
China, S-Sichuan, S Yanyuan, Chuandongzi Mt h=3300-3600 m, 16-20.VI.2023, C.Reuter leg./det.Klienfeld, 2023, L=21.5 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) rhododendron Deuve & Imura, 1991
Male adult
China, N Yunnan: Baimaxue Shan, S of Deqen, h=4300m, 28°20'N 99°03'E, 16-20.VI.2016, C.Reuter leg., L=18 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) mecynodes pinchuanensis Imura, 1999
Female adult
China, S-Sichuan, 40-50 km E Xichang (to Zhaojue) 27°51'N 102°30'E, h=2700-3000m, Pine forest, 5-20.VI.2017, C.Reuter leg., L=33 mm
Copyright © Ruslan Panin

Carabus (Piocarabus) vladimirskyi Dejean & Boisduval, 1830
Male adult
Buryatia, Kyahta distr. [Khyagh], 5 km SE Chikoi vill., 16.8.1998, Anichtchenko A. leg.
Copyright © Alexander Anichtchenko

Carabus (Apotomopterus) patroclus apricalis Cavazzuti, 1997
Endophallus
China, S Sichuan, W of Pingchuanzhen, near Galuo vill., h=2400 m
Copyright © Ruslan Panin

Carabus (Apotomopterus) patroclus apricalis Cavazzuti, 1997
Male adult
China, S Sichuan, W of Pingchuanzhen, near Galuo vill., h=2400 m, 7-18.VI.2017, C.Reuter leg./det. F.Kleinfeld, L=30 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) microtatos lingbaonis Imura, 2019
Endophallus
China, CW-Yunnan, Nánjiàn Yízú Zìzhìxiàn, Mt Lingbao h=2400 m, 24.768611, 100.500000
Copyright © Ruslan Panin

Carabus (Apotomopterus) microtatos lingbaonis Imura, 2019
Male adult
China, CW-Yunnan, Nánjiàn Yízú Zìzhìxiàn, Mt Lingbao h=2400 m, 24.768611, 100.500000, 6-15.VI.2023, C.Reuter leg., L=24 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) bousquetellus bousquetellus Deuve, 1998
Endophallus
S-China, CW-Yunnan, W of Lincang, Daxue Shan h=2600m, 23.935556, 100.257778
Copyright © Ruslan Panin

Carabus (Eurycarabus) famini atlantis Alluaud, 1923
Female adult
C Morocco, Haut Atlas, SW of Oukaïmeden h=2900m, 31.173644, -7.844442, 29.IV-11.V.2024, C.Reuter leg., L=21 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) mecynodes pinchuanensis Imura, 1999
Male adult
China, S-Sichuan, 40-50 km E Xichang (to Zhaojue) 27°51'N 102°30'E, h=2700-3000m, Pine forest, 5-20.VI.2017, C.Reuter leg., L= 29 mm
Copyright © Ruslan Panin

Carabus (Lamprostus) syrus cheikensis Deuve, 1992
Female adult
SE Lebanon, Rashaya Distr., Tannoura, oak forest, h=700 m, 11-19.IV.2016, C. Reuter leg., L= 32 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) microtatos lingbaonis Imura, 2019
Female adult
China, CW-Yunnan, Nánjiàn Yízú Zìzhìxiàn, Mt Lingbao h=2400 m, 24.768611, 100.500000, 6-15.VI.2023, C.Reuter leg., L=25 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) patroclus patroclus Breuning, 1933
Male adult
China, S Sichuan, Pass 56 km SW Miannin, near Wawo h=2440 m, 21-24. VI.2017, C.Reuter leg./det. F.Kleinfeld, L=29 mm
Copyright © Ruslan Panin

Carabus (Apotomopterus) bousquetellus bousquetellus Deuve, 1998
Male adult
S-China, CW-Yunnan, W of Lincang, Daxue Shan h=2600m, 23.935556, 100.257778, 7-13.VI.2023, C.Reuter leg./det. F.Kleinfeld, L=24 mm
Copyright © Ruslan Panin

Carabus (Acoptolabrus) constricticollis Kraatz, 1886
Male adult
China, Liaoning prov., Mt. Guanmen 20.6.2009 (cAA)
Copyright © Alexander Anichtchenko

Carabus (Coptolabrus) gemmifer mesites Fairmaire, 1897
Female adult
China, Sichuan, Xichang City, Muli Co., Ke-Er-xiang 2-12.7.2009 (cAA) 32 mm
Copyright © Alexander Anichtchenko

Carabus (Coptolabrus) ignimitella angulicollis Hauser, 1911
Male adult
China Hunan, Guidong pr., 20.5.2009 (cAA) 44 mm
Copyright © Alexander Anichtchenko

Carabus (Sphodristocarabus) varians armeniacus Mannerheim, 1830
Male adult
C Georgia, Imereti, Caucasus, Likhi Mt Range, Rikoti Pass h=900-1000 m, 14-16.V.2017, E. Rutjan E. leg., L=22 mm
Copyright © Ruslan Panin

Calosoma (Callisthenes) elegans elegans (as valentinae Obydov, 1997)
Male adult
SE-Kazakhstan, Almaty reg., W of Zarechnoe vill., h=600 m, 43.715278, 76.972778, 5.IV.2018, L=23 mm
Copyright © Ruslan Panin

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Female adult
W Ukraine, Lviv city, Holosko, Bryukhovytsʹkyi forest 49.886833, 24.012083 under bark, 10.XI.2024, R. Panin leg., L=35 mm
Copyright © Ruslan Panin

Calosoma (Callisthenes) elegans semenovi Motschulsky, 1859
Male adult
SE Kazakhstan, near Almaty, Komsomol vill., h=650 m, 43.458889, 76.748333, 3. V.2015, L=26 mm
Copyright © Ruslan Panin

Carabus (Piocarabus) titanus Breuning, 1933
Female adult
China Sichuan prov., Nanping, h=1500m, VI.2000, S. Remond leg.
L=38 mm
Copyright © Ruslan Panin

Calosoma (Callisthenes) elegans semenovi Motschulsky, 1859
Male adult
SE Kazakhstan, near Almaty, Komsomol vill., h=650 m, 43.458889, 76.748333, 3. V.2015
Copyright © Ruslan Panin

Carabus (Cyclocarabus) zarudnyi Semenov & Znojko, 1932
Female adult
NE-Uzbekistan, W-Tian-Shan, NW slopes Pskem Mt. Rng, 8-14 km E Pskem, 2100-2500 m, 16.VI.1998, A. Puchkov leg., L=21 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) ibericus lafertei Chaudoir, 1846
Endophallus
Georgia, Lesser Caucasus, Imereti prov., Meskheti Mt Rng, N slope, Zekari Pass h~2150 -2250 m alpine meadows
Copyright © Ruslan Panin

Carabus (Neoplectes) ibericus lafertei Chaudoir, 1846
Male adult
Georgia, Lesser Caucasus, Imereti prov., Meskheti Mt Rng, N slope, Zekari Pass h~2150 -2250 m alpine meadows 25.VII.2021, E.Rujan leg./det. R. Panin, L=33 mm
Copyright © Ruslan Panin

Carabus (Cyclocarabus) karaterekensis hemicallisthenes Kabak & Kalab, 1997
Male adult
NW- Kyrgyzstan, W-Tian-Shan, Chandalash Mt.Rng, Chakmak-Suu vill., 3.VI.2000, L=17 mm
Copyright © Ruslan Panin

Carabus (Cyclocarabus) brosciformis Semenov, 1896
Male adult
NE-Uzbekistan, W-Tian-Shan, Chatkal Mt Range, Tashkent reg., Yangiabad env., h=2100m, 30.IV.2002, L=23.5 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Paratype
Female adult
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky forest, near Pyatnychany vill., 28.V-9.VI.2013, Panin R. leg. L=30 mm
Copyright © Ruslan Panin
Carabini taxon page (4242 photos)
Cychrus sehnali Haeckel, 2007 Holotype
Male adult

Copyright © Boleslav Brezina

Cychropsis (Sinocychropsis) tryznai Haeckel & Sehnal, 2007 Holotype
Male adult
China - W Sichuan, 5.6.2006 West of Zhier (Zier) 4118 M
alpine forest N 28.20.507. E 101.28.014 leg. R Sehnal & M. Tryzna
Holotype
Copyright © Boleslav Brezina

Cychrus angustatus Hoppe & Hornschuch, 1825
Male adult
Italia, Lombardia (BG), Arera mt., 2100m Zambla, 19.6.2003 A. Degiovanni leg. det. V. Monzini, 2014 male, 20 mm
Copyright © Alexander Anichtchenko

Cychrus belousovi O.Berlov, 1997 Holotype
Male adult

Copyright © Oleg Berlov

Cychrus tuberculatus Harris, 1839
Female adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Cychropsis (Shuocychropsis) paramontana Sehnal & Hackel, 2006 Holotype Holotype
Male adult

Copyright © Boleslav Brezina

Scaphinotus (Steniridia) guyoti LeConte, 1866
Male adult
USA Georgia, Brasstown, Towns Co.,
Bald Mt., 16.7.2000 Sehnal leg.
male, 23 mm
Copyright © Alexander Anichtchenko

Scaphinotus (Nomaretus) bilobus Say, 1825
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Sphaeroderus canadensis Chaudoir, 1861
Male adult
Canada Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Sphaeroderus stenostomus lecontei Dejean, 1826
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Sphaeroderus nitidicollis (as brevoorti LeConte, 1848)
Female adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Cychrus caraboides Linnaeus, 1758
Male adult
L=17,0 mm
RUSSIA, Moscow city, East distr., National park «Losiniy Ostrov», spruce forest. 9.IX.2000
leg., det. & photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Scaphinotus (Maronetus) hubbardi Schwarz, 1895
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Scaphinotus (Neocychrus) angulatus Harris, 1839
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Scaphinotus (Scaphinotus) viduus Dejean, 1826
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Cychrus attenuatus attenuatus Fabricius, 1792
Male+Female adult
W.UKRAINE, NE Carpathians,
Lviv Reg., Skole distr., Skolivski Beskidy Mts. Zelemyanka Mt.Range h=700m, 26.V -1.VII.2010, Panin R. leg.

Copyright © Ruslan Panin

Cychrus semigranosus pindicus Maran, 1940
Male adult
CE-Greece: Larissa reg., Stomio, Ossa Mt. 6-17.VII.2011, Krause Rene leg. L=21 mm;
Copyright © Ruslan Panin

Cychrus angustatus Hoppe & Hornschuch, 1825
Male adult
NC-Italy, Bergamo, Mt. Presolana h=1500m, Colere env., 4-15.VI.2010, Manfred Egger leg.
L=23 mm
Copyright © Ruslan Panin

Cychrus attenuatus attenuatus s.str.
Female adult
CZ, NE Moravia, Moravskoslezské Beskydy Mts.; 13mm
Copyright © Aleš Sedláček

Scaphinotus (Stenocantarus) angusticollis Mannerheim, 1824
Male adult
NW-USA, Washington, Thurston County, Douglas fir forest, 15.X.2005, A. Laforgue leg. L=21mm
Copyright © Ruslan Panin

Scaphinotus (Scaphinotus) viduus Dejean, 1826
Male adult
E-USA, West Virginia, Pendleton Co., Allegheny Mts. Mt. Spruce Knob, 20-27.VI.2012, T. Lawton leg. L=27 mm
Copyright © Ruslan Panin

Cychrus (Cychrus) morawitzi morawitzi
Male adult

Copyright © Kirill Makarov

Cychrus (Cychrus) semigranosus (as montanus Winkler, 1939)
Female adult
W Ukraina, Chernivtsi Reg., Hertsa Distr., 23-25.VI.2017, Kanarsky Yu. leg. L=19 mm
Copyright © Ruslan Panin

Cychrus (Cychrus) caraboides (as elongatus Hoppe,)
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030 m, alpin zone, 10.VI.-5,VIII.2017, R. Panin R. L =13 mm
Copyright © Ruslan Panin

Scaphinotus (Steniridia) aeneicollis Beutenmuller, 1903
Male adult
CE USA, N Carolina, Yancey Co., S of Eskota, 17.V-30.VI.2013, T. Lawton leg., L=21 mm
Copyright © Ruslan Panin

Scaphinotus (Scaphinotus) elevatus coloradensis Vandyke, 1907
Male adult
CS-Canada, Manitoba, 10 km E of Libau, 21.VIII.2016, T. Lawton leg., L=18 mm
Copyright © Ruslan Panin

Scaphinotus (Nomaretus) fissicollis LeConte, 1853
Male adult
USA, Arizona, Logan Co., 13 miles SW of Paris, 13.X-23.XI.2014, T. Lawton leg., L=14 mm.
Copyright © Ruslan Panin

Scaphinotus (Steniridia) andrewsii mutabilis (Casey, 1920)
Male adult
CE USA, W Virginia, Pendleton Co., Mts Allegheny, Mt Spruce Knob, 2.VI-1.VII.2014, T. Lawton leg., L=22 mm
Copyright © Ruslan Panin

Cychrus (Cychrus) spinicollis spinicollis
Female adult
SPAIN, Cantabria, 2 km E Puerto de San Glorio, near Vejo riv., 43.072369, -4.733061, 1300 m., 23.07.2019, A. Anichtchenko & R. Sciaky leg.
Copyright © Alexander Anichtchenko

Cychrus (Cychropsis) draconis jiudingensis Hackel & Sehnal, 2014
Female adult
C Sichuan, Jiyuding Shan, 30km N Wenchuan, 3250m, 2.7.2017, leg. V. Patrikeev. det. A. Anichtchenko, 2020
Copyright © Alexander Anichtchenko

Scaphinotus (Scaphinotus) petersi catalinae Vandyke, 1924 (Topotype)
Male adult
USA, Arizona, Pima County, Mt. Lemmon, N 32°26.448', W 110°47.032', ele: 9013', 07.VIII, 2017, leg. Todd Lawton
Copyright © Igor Solodovnikov

Scaphinotus (Stenocantarus) velutinus (Menetries, 1843)
Male adult
USA, OR, Benton Co., III-V.1996 Mary's Peak, 3600m N slope, Labonte cols. (cAA) 26 mm; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Scaphinotus (Stenocantarus) angusticollis olympiae Van Dyke, 1944 (loc. typ.)
Male adult
USA: WA., Clallam Co, Olympic Nat Park, Sol Duc Hot Springs 25.5.96, M. Kippenhan leg. (cAA) 17 mm; det. A. Anichtchenko, 2020
Copyright © Alexander Anichtchenko

Scaphinotus (Neocychrus) behrensi Roeschke, 1907
Male adult
USA: OR., Benton Co., Mary's Peak 3600 m, N Slope, III-V.1996, J. Labonte cols. (cAA) 18 mm
Copyright © Alexander Anichtchenko

Scaphinotus (Brennus) marginatus var. fallax Roeschke, 1907
Male adult
USA, OR., Yamhill Co., Crawford Beck Vineyard, Rd 153 E of Amity, 45.06.805N 123.08.539W 30.3.-22.5.2019, M. Kippenhan leg. (cAA) 14mm; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Scaphinotus (Brennus) marginatus Fischer, 1822
Female adult
USA, OR., Multnomah Co., Portland, Oaks Bottom WA, 7.5.2004 M. Kippenhan (cAA) 14 mm; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Scaphinotus (Brennus) rugiceps Horn, 1872
Male adult
USA: OR: Josephine Co. / Lone Mountain Rd, 4.3 miles S.W of Hwy 199 / 42° 01.787' N 123° 45.760' W 1719', 18-IV-6-VI-2020, M. Kippenhan (cAA) 15 mm; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Scaphinotus (Brennus) obliquus LeConte, 1868
Male adult
USA, CA., Shasta Co., Castle Crags State Park, 29-29.5.2016, M. Kippenhan leg. (cAA) 15 mm; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Scaphinotus (Stenocantarus) angusticollis Mannerheim, 1824
Female adult
USA, WA., Pierce Co., Penrose S.P. Trails, 7 sept 1996, M. Kippenhan leg (cAA) 19mm; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Scaphinotus (Stenocantarus) hatchi Beer, 1971
Male adult
USA OR Lane Co., N Shore Waldo lake, 27.6.97 M Kippenhan leg. (cAA) 17mm
Copyright © Alexander Anichtchenko

Scaphinotus (Brennus) cf oreophilus Rivers, 1890
Female adult
USA , CA, Toulumne Co., The Pines CG, Rt 120 West of Buck Meadows, 3267m, 37.49.14N 120.05.7W 19.6.2011 (cAA) 15mm; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Scaphinotus (Pseudonomaretus) relictus Horn, 1881
Male adult
Canada, BRITISH COLUMBIA (BC), W of Creston, N 49°08.494', W 116°46.381', ele: 5822', 24.VII.2015, leg. Todd Lawton
Copyright © Igor Solodovnikov

Scaphinotus (Stenocantarus) hatchi Beer, 1971
Male adult
USA: Oregon, Lane County, env. Waldo Lake, N 43°44.835', W 122°00.566', ele: 4272', 19.VII.2015, leg. Todd Lawton (TOPOTYPUS)
Copyright © Igor Solodovnikov

Scaphinotus (Brennus) marginatus Fischer, 1822
Male adult
Canada, BRITISH COLUMBIA (BC), Wynndel, Duck Creek Road, N 49°11.852', W 116°32.294', ele: 1917', 24.VII-04.IX.2015, leg. Todd Lawton
Copyright © Igor Solodovnikov

Scaphinotus (Stenocantarus) angusticollis Mannerheim, 1824
Male adult

Copyright © Igor Solodovnikov

Scaphinotus (Scaphinotus) viduus Dejean, 1826
Male adult

Copyright © Igor Solodovnikov

Scaphinotus (Scaphinotus) viduus Dejean, 1826
Female adult

Copyright © Igor Solodovnikov

Scaphinotus (Scaphinotus) irregularis (Beutenmüller, 1903)
Female adult

Copyright © Igor Solodovnikov

Scaphinotus (Scaphinotus) elevatus coloradensis Vandyke, 1907
Male adult

Copyright © Igor Solodovnikov

Scaphinotus (Scaphinotus) elevatus coloradensis Vandyke, 1907
Female adult

Copyright © Igor Solodovnikov

Scaphinotus (Steniridia) guyoti LeConte, 1866
Male adult

Copyright © Igor Solodovnikov

Scaphinotus (Steniridia) guyoti LeConte, 1866
Female adult

Copyright © Igor Solodovnikov

Cychrus (Cychrus) starcki zamotailovi Berlov, 1997
Male adult
Abkhasia, Gudauta distr., env. vill. Ludzava, vall. Ryaphi Riv. (Pitzunda), 10-17.VII.2011, Soodovnikov I.A.
Copyright © Igor Solodovnikov

Cychrus (Cychrus) signatus signatus
Male adult
Georgia, Adjara, env. Batumi, Makkhinjauri, abandoned house above railway tunnel, h = 107 m
Copyright © Igor Solodovnikov

Cychrus (Cychrus) rosti belousovi Berlov, 1997
Male adult

Copyright © Igor Solodovnikov

Cychrus (Cychrus) rosti rosti
Male adult
Abkhasia, env. Dzhampal
Copyright © Igor Solodovnikov

Cychrus (Cychrus) rosti rosti
Female adult
Abkhasia: Sukhum distr., Birtzkha Mt.
Copyright © Igor Solodovnikov

Cychrus (Cychrus) aeneus japoshvilii Deuve & Prunier, 2014
Male adult
Georgy: Samtskhe-Dzhavakhetie, near Borjomi: Akhaldaba (TOPOTYPUS)
Copyright © Igor Solodovnikov

Cychrus (Cychrus) aeneus aeneus Fisher-Waldheim, 1824
Male adult
Georgy: env. Bakuriani
Copyright © Igor Solodovnikov

Cychrus (Cychrus) rosti retezari Deuve & Prunier, 2014
Male adult
Abkhasia: Ochamchyra distr., near vill. Nardzhkheu, h = 520 m
Copyright © Igor Solodovnikov

Cychrus (Cychrus) aeneus volkovi O.Berlov, 1997
Male adult

Copyright © Igor Solodovnikov

Cychrus (Cychrus) rosti tillyi Berlov, 1997
Male adult
Abkhasia: vall. riv. Yupshara
Copyright © Igor Solodovnikov

Cychrus (Cychrus) rosti tillyi Berlov, 1997
Female adult
Abkhasia: vall. riv. Yupshara
Copyright © Igor Solodovnikov

Cychrus (Cychrus) rosti tillyi Berlov, 1997
Male adult
Abkhasia: vall. riv. Yupshara
Copyright © Igor Solodovnikov

Cychrus (Cychrus) rosti milyaevi Berlov, 1997
Male adult
Abkhasia: vall. riv. Mzara
Copyright © Igor Solodovnikov

Cychrus (Cychrus) starcki starki
Male adult
Russia: W-Caucasus, Lasorevskoe distr., vall. Psezuapse Riv.
Copyright © Igor Solodovnikov

Scaphinotus (Brennus) interruptus Menetries, 1844
Male adult
USA California, Sonoma Co., 2.5mi N of Cazadero on King Ridge Rd. Big Austin Creek, 70m, 38.33N 123.06W 21.6.1996 D.H.Kavanaugh leg. (cAA). det. Anichtchenko A. 2021
Copyright © Alexander Anichtchenko

Scaphinotus (Steniridia) ridingsii monongahelae Leng, 1917
Male adult
USA W Virginia , Spruce Knob Mt, 4.7.1996 (cAA)
Copyright © Alexander Anichtchenko

Cychrus (Cychropsis) nepalensis Mandl, 1965
Male adult
Nepal, Lamtang, Dunche, near Songta vill. 3500m 15.5.2000 (cAA). det. Anichtchenko A. 2021
Copyright © Alexander Anichtchenko

Scaphinotus (Steniridia) andrewsii germari (Chaudoir, 1861)
Male adult
USA Virginia, Smoth co., Whitetop Mt. 10.7.1995 (cAA) 17 mm; det. Anichtchenko A. 2021
Copyright © Alexander Anichtchenko

Scaphinotus (Steniridia) andrewsii mutabilis (Casey, 1920) (as valentinei Barr)
Male adult
USA W Virginia, Spruce Knob Mt., 4.7.96 Sehnal leg. (cAA) 20mm
Copyright © Alexander Anichtchenko

Sphaeroderus canadensis canadensis
Male adult
USA Virginia, Rockimgham Co., Rawley Springs 4.9.2003 (cAA)
Copyright © Alexander Anichtchenko

Scaphinotus (Steniridia) tricarinatus (Casey, 1914) Topotype
Male adult
USA, N.C. Blue Ridge Parkway 430mi, 8.7.1995, Sehnal leg. (cAA) 18mm; det. A.Anichtchenko, 2021
Copyright © Alexander Anichtchenko

Scaphinotus (Scaphinotus) kelloggi Dury, 1912 Topotype
Male adult
USA, New Mexico, Grant Co., Mimbres Mts., 2 km W Emory Pass, Iron Creek Camp 2.8.1993, 2310 m, R.L. Davidson & R.E. Acciavatti leg. (cAA). R. Davidson det. 17 mm
Copyright © Alexander Anichtchenko

Cychrus (Cychrus) schmidti Chaudoir, 1837
Male adult
Slovenia Mt Canino, 08.05 (cAA) 14mm
Copyright © Alexander Anichtchenko

Cychrus (Shuocychropsis) huangcao Hackel & Sehnal, 2014 Holotype
Female adult
China, Sichuan, VIII – 2012, North of Shimian, Huangcao Shan, 3500 m; in collection of R. Sehnal
Copyright © Martin Häckel

Cychrus (Shuocychropsis) discoidicollis Deuve, 2017 Holotype
Male adult
Chine, S Sichuan, Mt 12 km SE Yanyuan, 4000-4200 m, 27°21’N-101°34’E, pelouses alpines et pierriers, in coll. J. Kaláb, à Jinačovice (République Tchèque)
Copyright © Thierry Deuve

Scaphinotus (Nomaretus) cavicollis LeConte, 1859
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH. Photo processed by Anichtchenko A. 2022
Copyright © - - -

Scaphinotus (Scaphinotus) petersi petersi
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH. Photo processed by Anichtchenko A. 2022
Copyright © - - -

Cychrus (Cychrus) semigranosus semigranosus Palliardi, 1825
Male adult
N Ukraine, Kiev city, NP Holosiivsky, 12.VIII - 9.IX.2023, R. Nerush leg./det. R Panin, L=17 mm
Copyright © Ruslan Panin

Scaphinotus (Scaphinotus) mexicanus Bates, 1882
Male adult
Mexico, Durango, Las Adjuntas W El Salto, Sierra Madre occ., 2600m, 25.vii.1990, Heinz leg. (cAA)
Copyright © Alexander Anichtchenko
Cychrini taxon page (246 photos)