Eremosphodrus (Eremosphodrus) rotundicollis Reitter, 1894d: 122
Male adult
Turkmenia, Repetek Reserv.
Copyright © - - -

Eremosphodrus (Rugisphodrus) dvorshaki Casale & Vereschagina, 1986
Female adult
Iran, Semnan, ~24 km NW Imamshehr, Alborz Mts., S slope Shahvar Mt,, 36º32'25" N, 54º44'51" E, ~2839 м, 05.V.2018, Kolesnichenko K. leg.; det. Anichtchenko A. 2019; 14 mm
Copyright © Alexander Anichtchenko

Hystrichosphodrus vailatii Casale & Giachino, 2005 Paratype
Female adult

Copyright © Vittorino Monzini

Laemostenus (Pristonychus) sericeus sericeus Fischer von Waldheim, 1824
Male adult
S Russia, NW-Caucasus, Karachay-Cherkessia,
Skalisty Mnt. Range, Riv.Muzdukh source h=2170 m, in a cave 43:42’08,68 N 42:19’28,10 E, 31.VII.2010, Solodovnikov I.A leg.
L=19mm

Copyright © Ruslan Panin

Laemostenus (Laemostenus) venustus Dejean, 1828: 60 (Pristonychus)
Male adult
S Ukraine,
Crimea Peninsula, Angarskyi Pass, 26.XII.2010, Shaporinskiy V. leg./det. Panin R.
L=15 mm
Copyright © Ruslan Panin

Laemostenus (Antisphodroides) koenigi Reitter, 1887
Male adult
Karachaevo-Cherkessia,
Zelenchukskii distr., near Bukovo vill.,h~2000m., 9-13.VII.2007, S.Dementiev leg.
L=17mm
Copyright © Ruslan Panin

Laemostenus (Antisphodrus) deconincki Casale & Queinnec, 2001
Male adult
Iran, Lorestan, 50 km N Ardimeshk, 2000 m
Sar Takht, 3-4.V.2007 Anichtchenko A. leg.
long 13 mm
Copyright © Alexander Anichtchenko

Laemostenus (Laemostenus) caspius Meneiries, 1832
Male adult
S Azerbajdzhan, Astara Distr., upper course of istisu-Chaj Riv., env. Siov Vill., h~1000 38°26'28" N 18-19.IV.2004 48°39'01" E K.Makarov leg.
L=12.6 mm
Copyright © Kirill Makarov

Laemostenus (Laemostenus) janthinus coeruleus (Dejean, 1828)
Male adult
Italia, Trentino, Mte Pasubio 1600mt
4.IX.2002 A.Degiovanni leg.
male, long 16 mm
Copyright © Alexander Anichtchenko

Laemostenus (Pristonychus) cimmerius cimmerius Fischer von Waldheim, 1823: pl. 36 (Sphodrus)
Male adult
S Ukraine, Krym Peninsula,
Bakhchisaraysk distr.,near Preduschelnoe, h=400m., 22.VII.2009, Shaporinskiy V. leg.
L=19 mm.
Copyright © Ruslan Panin

Laemostenus (Pristonychus) sericeus hepaticus Faldermann, 1836
Male adult
N Azerbaijan,
4km W Ismayilli, h=700m, 9.VII.2008, Pljushch I.G. leg.
L=20mm
det.Gerasimov R.P
Copyright © Ruslan Panin

Laemostenus (Pristonychus) terricola Herbst, 1784
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Laemostenus (Pristonychus) tichyi Kult, 1946
Female adult
W. UKRAINE, NW Podolian Upland, Ternopol Reg., Pidvolochisk distr., near Kamyanki vill., steppe site,, 17.V - 3.VI.2009, Panin R. leg./det.Solodovnikov I. & Panin R. female, 18 mm
Copyright © Ruslan Panin

Laemostenus (Pristonychus) tichyi Kult, 1946
Male adult
W Ukraine, NW Podolian, Ternopol reg., Gusyatyn distr., Monastyrykha vill., 9.VI.1993, Rizun V.B leg./ det. Panin R. & Solodovnikov I.A. 2010 male,16 mm photo Panin R.
Copyright © Ruslan Panin

Laemostenus (Pristonychus) terricola terricola Herbst, 1784: 140 (Carabus)
Female adult
L=16,0 mm
RUSSIA, Tver reg., Likhoslavlskiy distr., Domanikha village, in the basement of the house used as a potato storehouse. VIII.2001
leg. Luts A. N.
det. & photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Laemostenus (Pristonychus) stussineri Ganglbauer, 1896b: 463 (Aechmites)
Female adult
CE-Greece, Mts Kato Olympos, Karia env., 6-17.VII.2011, Krause Rene leg.
L=20 mm


Copyright © Ruslan Panin

Laemostenus (Microsphodrus) dolini Kabak, 1992
Male adult

Copyright © Kirill Makarov

Laemostenus (Microsphodrus) visai (Dvorak, 1982)
Male adult

Copyright © Kirill Makarov

Laemostenus (Antisphodrus) elongatus elongatus (Dejean, 1828)
Female adult
SW Slovenia, Obrov, Poljane, 5-19.IX.2009, Manfed Egger leg.
L=16 mm
Copyright © Ruslan Panin

Laemostenus (Antisphodrus) schreibersii (Kuster, 1846)
Female adult
NE Italy, Belluno reg., Cansiglio, Val Di Piera, 12.V-22.IX. 2007, Manfed Egger leg.
L=14.5 mm
Copyright © Ruslan Panin

Laemostenus (Pristonychus) lederi arenicus Kalashyan, 1979: 187
Female adult
Armenia, Vayots Dzor Province, 1.8 km E of Areni Vill. h~1170 39°43'43" N 45°12'27"E 20.X.1986 leg. M.Kalashian
L=12.5 mm
Copyright © Kirill Makarov

Laemostenus (Pristonychus) lederi lederi Reitter, 1885f: 358
Female adult
S Azerbaijan, Talysh, Astara Distr., env. Sym Vill. h~950 38°29'15"N 48°36'51"E 13-16.V.2012 leg. D.Kasatkin
L=14.2 mm
Copyright © Kirill Makarov

Laemostenus (Antisphodrus) nusayriyahensis Lohaj & Mlejnek, 2007 Holotype
Female adult
NW Syria, Homs, cave between Mashtal Helu and Uyounl Wadi, 600 m a.s.l., Mts Jebel Ansaria, 31.3.2001, M.Řezáč lgt., deposited in CRL. Image from Roman Lohaj & Roman Mlejnek, 2007
Copyright © - - -

Laemostenus (Antisphodrus) glasunowi Semenov, 1895a: 193
Female adult
Iran, Hasan Keif. Image from Roman Lohaj & Roman Mlejnek, 2007
Copyright © - - -

Laemostenus (Antisphodrus) puchneri Lohaj & Mlejnek, 2007 Holotype
Male adult
Turkei, s.c. Anatolien, Murtici, umg.Gözelsu, Gulendag, 1050 m, A. Puchner, 16. IV.– 7.V.2000, deposited in CWH. Image from Roman Lohaj & Roman Mlejnek, 2007
Copyright © - - -

Laemostenus (Pristonychus) spinifer Schaufuss, 1862
Male adult
N INDIA, Uttarakhand, 13-15.04.2012, Uttarkashi distr. N30°57'41.57", E78°41'54.75" left tributary of Bhagirathi, 2300 m, Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Laemostenus (Pristonychus) piceus Motschulsky, 1850a: 43
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Laemostenus (Actenipus) macropus Chaudoir, 1861a: 10
Male adult
Italy
Copyright © Vittorino Monzini

Laemostenus (Laemostenus) venustus Dejean, 1828: 60 (Pristonychus)
Male adult
W Ukraine, NE Carpathians, Transcarpathian Reg., Tiachiv distr., near Mala Ugolka vill., h=700m, 15.V-27.VI.2015, Panin R. & Kanarsky Yu. leg. L=16mm
Copyright © Ruslan Panin

Laemostenus (Pristonychus) terricola punctatus Dejean, 1828
Male adult
Hungary, Budapest, Télényi, 10-12.VII.2004, I.Retezár leg. L=19 mm
Copyright © Ruslan Panin

Laemostenus (Pristonychus) tichyi Kult, 1946
Male adult
W Ukraine, Ternopol Reg. Zalishchyky distr., Kasperivtsi vill., cellar, 22-23.IV.2016, Bachynsky A. leg./ det. Panin R & Solodovnikov I. L=18 mm
Copyright © Ruslan Panin

Laemostenus (Pristonychus) mannerheimi mannerheimi Kolenati, 1845: 50
Male adult
Armenia, Shirak ar., Amasya env., 40º56' N, 43º47' E, 25.VI.2003, A.Rubenyan. Image from www.zin.ru
Copyright © Maksim Smirnov

Laemostenus (Microsphodrus) rimmae Deuve, 1993: 86
Female adult
Kyrgizstan, 25.6.1998, 2800m, W Tian Shan, N Hang Talaakyi r., 10 km N Tjuzashu pass, Putchkov leg & det. 12 mm
Copyright © Alexander Anichtchenko

Laemostenus (Microsphodrus) loudai (Dvorak, 1982)
Male adult
Kyrgizstan, N Kirgizsky mt. rng., Kara Balta riv., 1600m, 8.4.2007, A. Klimenko leg. 10 mm
Copyright © Alexander Anichtchenko

Laemostenus (Eucryptotrichus) pinicola Graells, 1851
Male adult
Spain, Sierra de Guadarrama, 2000m, pto de Navacerrada, 29.6.2006
Copyright © Alexander Anichtchenko

Laemostenus (Laemostenus) complanatus (Dejean, 1828)
Male adult
Spain, Andalusia, Guadalfeo riv., near Orgiva, 1-4.4.2010, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Laemostenus (Laemostenus) parallelocollis (Reiche & Saulcy, 1855)
Male adult
Libano, Jbail prov., Quartaba, 8.5.2000, I. Rapuzzi leg. det. A. Anichtchenko 2017
Copyright © Alexander Anichtchenko

Laemostenus (Laemostenus) quadricollis L. Redtenbacher, 1843
Female adult
Israel, E Halilaa, Zefat env., Bait Zeitim, 35*28'N 32*99'E, 15-17.3.2006, A. Zamesov leg., det. Anichtchenko 2017
Copyright © Alexander Anichtchenko

Laemostenus (Pristonychus) sericeus Fischer von Waldheim, 1824
Female adult
Russia, N Caucasus, Kabardino-Balkaria rep., Dumala riv., 9.8.2006, Nabozhenko leg. det. Anichtchenko A. 2017
Copyright © Alexander Anichtchenko

Laemostenus (Actenipus) oblongus Dejean, 1828
Female adult
France, Aude, 8.1992, Rennes les Bains
Copyright © Alexander Anichtchenko

Laemostenus (Sphodroides) susicus Antoine, 1941
Female adult
Morocco, Sidi R'Bat, 21.3.2012. det. Anichtchenko A. 2017
Copyright © Alexander Anichtchenko

Laemostenus (Licinopsis) angustula Machado, 1987d: 404
Male adult
Canarias, La Palma, Cueva de los Arreboles, 22.9.1986, J.L. Martin leg.; 14mm
Copyright © Alexander Anichtchenko

Laemostenus (Calathidius) acuminatus (Wollaston, 1862)
Female adult
Canarias, Tenerife, 23.3.1974. det. A. Anichtchenko. 15 mm
Copyright © Alexander Anichtchenko

Laemostenus (Antisphodrus) peleus (Schaufuss, 1861)
Male adult
Spain, Asturias, Fuente del Infierno, 16.9.1988. 14 mm
Copyright © Alexander Anichtchenko

Laemostenus (Laemostenus) luristanus - quadricollis
Male adult
Iran, Lorestan, 50 km N Ardimeshk, 2000m, Sar Takht, 3-4.V.2007, leg. & det. Anichtchenko A.
Copyright © Alexander Anichtchenko

Laemostenus (Sphodroides) cordicollis (Chaudoir, 1854)
Male adult
Iran, Fars, near Sarvestan, 8.5.2007, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Laemostenus (Antisphodrus) malhommei (Antoine, 1954)
Female adult
Morocco, rand Atlas Mts., 2200-2300m, N31*13'09,5" W007*49'29,7", Oukaimedene env., 16-17.6.2010, Anichtchenko A. leg. & det. 13 mm
Copyright © Alexander Anichtchenko

Laemostenus (Laemostenus) magellensis Leoni, 1907b: 53
Male adult
Italy, Abruzzo, Toppe del Tesoro, AQ Roccaraso mt., 2100 m 27.7.2001 Degiovanni leg.
Copyright © Alexander Anichtchenko

Laemostenus (Antisphodrus) esfandiarii Morvan, 1974
Male adult
Iran, Mazandaran, between Marzan Abad & Chalus, 21-22.V.2007, leg. & det. Anichtchenko A. 10 mm
Copyright © Alexander Anichtchenko

Laemostenus (Laemostenus) barbarus (Lucas, 1846)
Female adult
Algeria, Konstantin Mt., grotte, 27.5.1973, V. Orlov leg.
Copyright © Alexander Anichtchenko

Laemostenus (Ceuthosthenes) cf. mauritanicus (Dejean, 1828)
Female adult
2km W Ceuta, Allabacal, 17-18.2.2004, leg. & det. Anichtchenko A.
Copyright © Alexander Anichtchenko

Laemostenus (Pristonychus) ferghanicus Vereschagina & Kabak, 1997: 861
Male adult
Kyrghyzstan, SE part of Ferghana Mt.R., Yassy River, Karateget
Copyright © Ilya Kabak

Laemostenus (Laemostenus) libanensis (Piochard de la Brulerie, 1876)
Male adult
Rayfoun ca. 33°58'N, 35°42'E, 18.XI.2012, mixed decid. forest, 800-900 m pitfall trap, leg. Reuter. Image from Casale & Assmann, 2017
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Laemostenus (Laemostenus) libanensis (Piochard de la Brulerie, 1876) Holotype
Male adult
Image from Casale & Assmann, 2017
Copyright © - - -

Laemostenus (Laemostenus) aegyptiacus Schatzmayr, 1936a: 88 Holotype
Female adult
Egypt, North Sinai, Wadi Garraui, South Seluan (MSNM). Image from Casale & Assmann, 2017
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Laemostenus (Laemostenus) sinaiticus Casale & Assmann, 2017 Holotype
Male adult
Egypt, South Sinai Saint Katherina Mountains Wadi Shagg (Tinya); West of Abu Sila Riparian habitats ~ 1600 m asl, 14.12.2009 leg. Th. Assmann” (cAS). Image from Casale & Assmann, 2017
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Laemostenus (Laemostenus) antonrichteri Casale, 1988: 923 Holotype
Male adult
Mount Hermon (=Jebel esh-Sheikh) at 1100-1400 m a.s.l. Image from Casale & Assmann, 2017
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Laemostenus (Laemostenus) antonrichteri Casale, 1988
Female adult
Upper Galilee (Sharakh Cave). Image from Casale & Assmann, 2017
Copyright © - - -

Laemostenus (Pristonychus) terricola terricola (Herbst, 1784)
Female adult
W-Ukraine, Podolian Tovtry, Ternopol reg., Pidvolochysk distr., Shevchenkove vill., steppe sites, 6.V-7.VI.2012, Panin R. leg. L=18 mm
Copyright © Ruslan Panin

Laemostenus (Pristonychus) terricola terricola (Herbst, 1784)
Male adult
W-Ukraine, Podolian Tovtry, Ternopol reg., Pidvolochysk distr., Shevchenkove vill., steppe sites, 6.V-7.VI.2012, Panin R. leg. L=17 mm
Copyright © Ruslan Panin

Laemostenus (Antisphodrus) cavicola Schaum, 1858b: 382
Male adult

Copyright © Vittorino Monzini

Laemostenus (Actenipus) oblongus cebennicus Jeannel, 1937b: 92
Male adult
CS-France, Aveyron (12), mines de Potensac, 26.VI.2007, L=16 mm
Copyright © Ruslan Panin

Laemostenus (Antisphodrus) insubricus Ganglbauer, 1903b: 224
Male adult
N Italy, Lombardia, Opreno, Cave Büs del Boter, 29.X.2011,G.Trezzi leg. L=14 mm
Copyright © Ruslan Panin

Laemostenus (Antisphodrus) iranicus Dvorak, 1982
Male adult
Iran, Esfahan, Dena Mts., 4 km ZO Bizhan pass, 10.vi.2018, 3850m, Muilwijk J. leg. & det.
Copyright © Jan Muilwijk

Laemostenus (Antisphodrus) aequalis Muilwijk & Malek Hosseini, 2016
Female adult
Iran Kohgiluyeh and Boyer-Ahmad Kiler cave Roowdrysh 14.vi.2018 S.Rahideh & J.Muilwijk
Copyright © Jan Muilwijk

Laemostenus (Antisphodrus) hormozi Azadbakhsh & Sciaky, 2018 Holotype
Male adult
Male, Iran, Hormozgan province, Haji-Abad region, 20.III.2017-20.IV.2017, S. Azadbakhsh leg and det.
Copyright © Saeed Azadbakhsh

Laemostenus (Iranosphodrus) rudichae Lohaj & Casale, 2011 Holotype
Male adult
IRAN (Kermān), Highland of Iran, Răyen env., ca 2500 m, 29º35 N/57º26 E, 18.V.2006 P.Rudich & J.Kny (white label, printed) / HOLOTYPUS Laemostenus Iranosphodrus subgen. nov. rudichae sp. nov. R. Lohaj & A. Casale det. 2011 (red label, printed)“, deposited in cDW. Image from Lohaj & Casale, 2011
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Laemostenus (Pristonychus) pretiosus Faldermann, 1837
Male adult
SW Georgia, Lesser Caucasus, Adzhar-Imereti (Meskheti) Rng, 4 km NW of Zekari Pass h~2000 m, 28-30.VI.2013. L= 16.5 mm
Copyright © Ruslan Panin

Laemostenus (Antisphodrus) hormozi Azadbakhsh & Sciaky, 2018 Paratype
Male adult
Iran, Hormozgan prov, Bandar Abbas env., Geno Mts. 1700m, 20.III.-20.IV.2017, S. Azadbakhsh leg.
Copyright © Alexander Anichtchenko

Laemostenus (Antisphodrus) laserae Casale & Wrase, 2018 Holotype
Male adult
IRAN (Hormozgan Prov.) Genu Mts N Bandar Abbas, slope with limestone, 1680 m 27°24ʹ04.5ʺN 56°10ʹ49.6ʺE (under broom bush, under stones/in soil) 26.II.2015 Wrase & Laser [03] (cWR). Image from A. Casale & D.W. Wrase, 2018
Copyright © - - -

Laemostenus (Arabosphodrus) balkenohli Sciaky, 1996b: 207
Female adult
Oman. Image from A. Casale & D.W. Wrase, 2018
Copyright © - - -

Laemostenus (Laemostenus) luristanus Casale, 1988
Female adult
Iran, Fars, 7 km W Dašt-e Aržan. Image from A. Casale & D.W. Wrase, 2018
Copyright © - - -

Laemostenus (Antisphodrus) schreibersii (Kuster, 1846)
Male adult
Colli Berici, Vicenza, Italia. Photo of Erminio Piva
Copyright © Vittorino Monzini

Laemostenus (Pristonychus) turkestanicus Semenov, 1891
Male adult
Tadjikistan, Peter 1 Mt.R., Ganishou env.
Copyright © Ilya Kabak

Laemostenus (Actenipus) caussolensis J. Ochs, 1937: 63
Female adult
grotte de la Statue, Halpes maritimes, France
Copyright © Vittorino Monzini

Laemostenus (Antisphodrus) mirzayani (Morvan, 1973)
Male adult
Iran, Zagros Mountains, Esfahan Fereydunshahr, Mei-danak Kamaran road, Kamaran pass Ferdum Mountains 3150–3400 m, 2–3.VII.2017, leg. Rutjan E.
Copyright © Jan Muilwijk

Laemostenus (Antisphodrus) murastyii Muilwijk & Lohaj, 2019 Holotype
Female adult
W. Iran, Isfahan prov., Fereydunshahr, Kamaran pass, 3650 m, 2–3.VII.2017, leg. Murastyi D. / Laemostenus murastyii sp. nov. Muilwijk & Lohaj / Holotype [red label].
Copyright © Jan Muilwijk

Laemostenus (Antisphodrus) navaricus Vuillefroy, 1893
Male adult
F, Arbailles, gr. Istaerdy
Copyright © Vittorino Monzini

Laemostenus (Pristonychus) zhentangensis Zhu, Shi & Liang, 2021 Holotype
Male adult
Image from Zhu, Shi & Liang, 2021
Copyright © - - -

Laemostenus (Pristonychus) zhamensis Zhu, Shi & Liang, 2021 Holotype
Male adult
Image from Zhu, Shi & Liang, 2021
Copyright © - - -

Laemostenus (Antisphodroides) levushkini Vareschagina, 1986: 560
Female adult

Copyright © Igor Solodovnikov

Laemostenus (Pristonychus) terricola terricola (Herbst, 1784)
Female adult
Belarus, Vitebsk reg., Vitebsk distr., vill. Novoselki
Copyright © Igor Solodovnikov

Laemostenus (Actenipus) pippiai C. Fiori, 1961
Male adult
Sardegna
Copyright © Vittorino Monzini

Laemostenus (Microsphodrus) tacitus (Dvorak, 1982)
Male adult
Uzbekistan, Chatkal Mt. rng., Irtash, 41°10'25.7"N 70°16'20.4"E, 5-7.8.2021, 2400m, A. Anichtchenko leg. (cAA). det. I. Kabak, 2021
Copyright © Alexander Anichtchenko

Laemostenus (Stenosphodrus) subtilissimus Casale & Sciaky, 1999
Female adult
Tadzhikistan, Zeravshan Mt. Rng., Marguzor lake, 5 km Hurdak, 1950/2400 m - S. Bajdek leg.
Copyright © Vittorino Monzini

Laemostenus (Antisphodrus) levantinus Bolivar y Pieltain, 1919
Male adult

Copyright © Vittorino Monzini

Laemostenus (Antisphodrus) bozdagensis Lohaj & Anlas, 2021 Holotype
Male adult
TURKEY, Manisa, Bozdağlar, Alaşehir, environs Dağarlar village, 570 m, MSS-1 trap, 38°11‘49“N, 28°28‘51“E, 04.-28.X.2017, S. Anlaş lgt.
Copyright © - - -

Laemostenus (Antisphodrus) binboga Lohaj & Anlas, 2021 Holotype
Male adult
TURKEY, Kayseri, Sarız, Binboğa Dağlari, 2465 m, 20 MSS, 38°20‘03“N, 36°31‘28“E, 08.06.2016 - 02.11.2017, S. Anlaş lgt.
Copyright © - - -

Laemostenus (Pristonychus) bellicosus Casale & Wrase, 2012 Holotype
Female adult
SYRIA occ., Crac des Chevaliers 34° 45’ N 36° 17’ E, 27.VI.1998 leg. P. Kabátek, 610 m (cWR). Image from Casale & Wrase, 2012
Copyright © - - -

Laemostenus (Pristonychus) eggeri Casale & Wrase, 2012 Paratype
Male adult
SYRIEN, nord-west Aleppo, ca. 500-550 m, umg. HARIM u. BASHMISHLI, I. Puchner 1.V.-20.V.99 (cWR). Image from Casale & Wrase, 2012
Copyright © - - -

Laemostenus (Laemostenus) phoenicius Casale & Wrase, 2012 Paratype
Female adult
Libanon: Maїfouq 800 m 4/13/IV/1997 (ne. Jbaїl) Heinz leg. Image from Casale & Wrase, 2012
Copyright © - - -

Laemostenus (Laemostenus) croyi Casale & Wrase, 2012 Paratype
Female adult
Turkey (Antalya) E Taurus Mts. Kizildag, env. 1800 m 37.03N/32.21E (subalpine) 6.-15.VI.2004 P. Croy (cDW). Image from Casale & Wrase, 2012
Copyright © - - -

Laemostenus (Antisphodrus) monguzzii Casale, 1988
Male adult
Val Trompia, BS, Lombardia, Italia
Copyright © Vittorino Monzini

Laemostenus (Actenipus) vignai Casale, 1988: 537
Female adult

Copyright © Vittorino Monzini

Laemostenus (Actenipus) angustatus Dejean, 1828
Male adult

Copyright © Vittorino Monzini

Laemostenus (Pristonychus) cf capitatus Chaudoir, 1854: 144
Male adult
NW Turkey, Bursa prov., Unçukuru vill., h=500 m, 1-14.XI.2018, S.Vashchenko leg., L=18.5 mm
Copyright © Ruslan Panin

Laemostenus (Antisphodrus) cavicola cavicola Schaum, 1858b: 382 (Sphodrus)
Female adult
Slovenia, Poljane, Obrov, Mts Ćićarija, Cave Polina Peč (Cad. No. 938), II.2016, L=17.5 mm
Copyright © Ruslan Panin

Laemostenus (Antisphodrus) cavicola cavicola Schaum, 1858b: 382 (Sphodrus)
Male adult
Slovenia, Poljane, Obrov, Mts Ćićarija, Cave Polina Peč (Cad. No. 938), II.2016, L=18 mm
Copyright © Ruslan Panin

Laemostenus (Pristonychus) cadilhaci Antoine, 1951
Male adult
Maroc, Ifni El Caid, 20-07-1984
Copyright © Pascal Deschamps

Laemostenus (Actenipus) acutangulus (Schaufuss, 1862)
Male adult
Accettura bosco Gallipoli, Basilicata, leg F. Angelini
Copyright © Vittorino Monzini

Laemostenus (Stenosphodrus) subtilissimus Casale & Sciaky, 1999a: 198
Male adult
NW Tajikistan, Gissar Mts, Fanny Mt Rng, like Khurdak, h=1800-2000 m, 12-24.V.1997
Copyright © Ruslan Panin

Laemostenus (Antisphodrus) mirzayani (Morvan, 1973)
Female adult
CW Iran, Ferdum Mts, Esfahan prov., Fereydunshahr - Meidanak road, Kamaran pass, h= 3150–3400m, 2–3.VII.2017, leg. Rutjan E./det. J. Muilwijk, L=15 mm
Copyright © Ruslan Panin

Pseudotaphoxenus aralensis Kabak, 2009 Holotype
Male adult
Kazakhstan, Aralsk, Nikolaev, 28.6.1980; ca. 46°49’N / 61°40’E
Copyright © Ilya Kabak

Pseudotaphoxenus rugupennis Faldermann, 1836
Male adult
Mongolia, 50 km N Mandalgob, Baga Gazryn Chuluu mts.,
1500m, 7-9.6.2003 Anichtchenko A. leg.
long. 26 mm
Copyright © Alexander Anichtchenko

Pseudotaphoxenus dauricus Fischer von Waldheim, 1823
Male adult
length: 20,6 mm
Transbaikalia, Tchita area, Krasnotchikoyskyi distr., near Albituy settl.
20.07.1999
Zinoviev E.leg.
Copyright © Evgeniy Zinoviev

Pseudotaphoxenus dauricus Fischer von Waldheim, 1823
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pseudotaphoxenus rufitarsis Fischer von Waldheim, 1823 : pl. 36
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pseudotaphoxenus planicollis Gebler, 1833
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pseudotaphoxenus taschkensis Jedlicka, 1952
Female adult
E-Uzbekistan, Aktash, Karzhantau Mt Rng, h~1400-1600m, 30.VI-4.VII.1991, I.Retezár leg. L=19 mm
Copyright © Ruslan Panin

Pseudotaphoxenus juvencus juvencus Ballion, 1871
Male adult
E Tajikistan, Dushanbe, Yavroz env., 6-9.V.1985, J. Kalab leg./det.Casale, L=16 mm
Copyright © Ruslan Panin

Pseudotaphoxenus michajlovi (Vereschagina, 1988)
Male adult

Copyright © Kirill Makarov

Pseudotaphoxenus onerosus Vereschagina & Kabak, 2000
Male adult

Copyright © Kirill Makarov

Pseudotaphoxenus tianshanicus Semenov, 1908: 120
Male adult
Kyrghyzstan, S slpoe of Kunghei Alatoo Mt.R., Ananievo env.
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Pseudotaphoxenus dostali Casale, 1988: 282
Male adult
SE Kazakhstan, Zhetyzhol Mt.R., 4 km SE of Sunkar (=Pokrovka) Vill.
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Pseudotaphoxenus igori Vereschagina & Kabak, 1996: 131
Male adult
SE Kazakhstan, Ketmegne Mt.R., Tuyuk env.
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Pseudotaphoxenus pongraczi Jedlicka, 1952b : 87
Male adult
SE Kazakhstan, Zailijskij Alatau Mt.R., Dolan River
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Pseudotaphoxenus pseudocollaris Casale, 1988: 293
Male adult
SE Kazakhstan, Almaty City, Kok-Tiube Mt.
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Pseudotaphoxenus strigitarsis (Jedlicka, 1958)
Male adult
Tajikistan, Ramitskoe l-vo, 25.5.1954
Copyright © Alexander Anichtchenko

Pseudotaphoxenus achillei Kabak, 2018 Holotype
Male adult
China, Xinjiang, right bank of Tekes River, S of Karatogai Vill., 43°11'46''N / 82°15'38''E, h = 1290 m, 6.V.2017 I.I. Kabak leg. (ZIN)
Copyright © Ilya Kabak

Pseudotaphoxenus tarantsha Kabak, 2018 Holotype Paratype
Male+Female adult
(ZIN). China, Xinjiang, S foothills of Boro-Horo Range, left bank of Piliktshi River, E of Karagatsh Vill., placor, 44°04'38''N / 81°30'49''E – 44°04'48''N / 81°30'55''E, 990–1040 m, 7.V.2017 (I.I. Kabak leg.).
Copyright © Ilya Kabak

Pseudotaphoxenus kazakhstanicus Kabak, I.Solodovnikov, 2016 Holotype
Male adult
Kazakhstan, “W Kazakhstan, Aktobe Prov., Aktolagai Range, h = 170–220 m, Mt. 238 m, 8–11.05.2004, I.A. Solodovnikov, K.E. Dovgailo” [ca. 47°27’N / 55°07’E], «Pitfall traps along gorge with Spiraea and Cytisus” [both labels are in Russian]
Copyright © Igor Solodovnikov

Pseudotaphoxenus kazakhstanicus Kabak, I.Solodovnikov, 2016 Paratype
Female adult
Kazakhstan, Karaganda Prov., “Koksengir Mts., 40 km S of Zhana-Arka, Karag[anda Prov.], G. Medvedev, 29.V.961” [ca. 540 m, 48°22’N / 71°31’E] [in Russian]
Copyright © Igor Solodovnikov

Pseudotaphoxenus taschkensis Jedlicka, 1952b: 87
Male adult
Uzbekistan, Chatkal Mt. rng., Aksay river, snow spot, 41°30'46.0"N 70°02'32.9"E, 2200m, 9-10.8.2021, Anichtchenko A. leg. & det. (cAA). 15 mm
Copyright © Alexander Anichtchenko

Pseudotaphoxenus dignus (Vereschagina, 1988)
Female adult
Uzbekistan, Chatkal Mt. rng., Irtash, 41°10'25.7"N 70°16'20.4"E, 5-7.8.2021, 2400m, A. Anichtchenko leg. (cAA). 17mm. det. I. Kabak, 2021
Copyright © Alexander Anichtchenko

Pseudotaphoxenus subcylindricus Semenov, 1891
Male adult

Copyright © Vittorino Monzini

Pseudotaphoxenus giorgiofiorii Casale & Vereschagina, 1986: 163
Male adult
Turkmenistan, Kushka Badenis
Copyright © Vittorino Monzini

Pseudotaphoxenus originalis Schaufuss, 1865: 99
Male adult
Mongolia, Umnegovi Aimak, 32 km west of Dalanzadgad 43°32'57"N 104°01'26"E 3.VIII.2019 leg. A. Rubenyan L=20.1 mm
Copyright © Kirill Makarov

Pseudotaphoxenus pseudocollaris Casale, 1988: 293
Male adult
SE Kazakhstan, Almaty reg, Kaskelen gorge h=1600 m, 43.100000, 76.589167, 19.IV.2019
Copyright © Ruslan Panin

Reflexisphodrus refleximargo Reitter, 1894
Male adult
det. Makarov K.
Copyright © Kirill Makarov

Reflexisphodrus formosus (Semenov, 1895)
Male adult
Russia, Buryatia, Selenginskii distr., 10 km N Deben vill., 14-15.8.1998, A. Shavrin leg. det. Anichtchenko A.
Copyright © Alexander Anichtchenko

Sphodropsis ghilianii caprai Binaghi, 1939
Male adult

Copyright © Vittorino Monzini

Sphodrus leucophthalmus Linne, 1758
Female adult
W Austria, Hall in Tirol, 11.IV.1952, Richler leg.
L=26 mm

Copyright © Ruslan Panin

Sphodrus leucophthalmus Linne, 1758
Male adult
Spain, Almeria, Punto de Sabinar, 30SWF 2962
Dalias, 24.1.1990
L= 26 mm
Copyright © Alexander Anichtchenko

Sphodrus leucophthalmus Linne, 1758: 413 (Carabus)
Female adult
22 mm, England, Reading
leg.det.W.Holland, 1901
Photo: U.Schmidt, 2010
Copyright © Udo Schmidt

Sphodrus trochanteribus Mateu, 1990a: 31
Male adult
C.Pakistan :Mussafargarkh
Copyright © Igor Solodovnikov

Sphodrus trochanteribus Mateu, 1990a: 31
Male adult
Pakistan, Punjab Prov., Muzaffargarh 1.XI-25.XII.1994 leg. V.Semin L = 25.5 mm
Copyright © Kirill Makarov

Stenolepta cylindrica Semenov, 1889e: 688
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Taphoxenus (Lychnifugus) cellarum cellarum M.F. Adams, 1817: 296 (Carabus)
Male adult
Azerbaijan,
Gobustan Karadagh, 05.IV.1986, Kazaryan P.V. leg.
L=24mm
photo Panin R.\det.Solodovnikov I.A.
Copyright © Ruslan Panin

Taphoxenus (Taphoxenus) gigas Fischer von Waldheim, 1823: pl. 36 (Sphodrus)
Male adult
N-Kazakhstan, Kostanay Reg., Zhetikara env., virgin land steppe, 3-25.VII.2013, Kozionov leg.
L=29 mm

Copyright © Ruslan Panin

Taphoxenus (Taphoxenus) trochanteratus Emden, 1954
Male adult
Turkmenia, Kopet-Dag Mt.R., Dushak
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Taphoxenus (Lychnifugus) ziegleri Casale & Assmann, 2017 Holotype
Male adult
Jordanien 762m sü Madaba Bodenfalle W. Ziegler 13.3.2015 ; Olivenhain bei AUM N31°39'30 E35°47'47 (cAS). 25 mm; Image from Casale & Assmann, 2017
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Taphoxenus (Lychnifugus) meridionalis Casale, 1982a: 146 Paratype
Male adult
Syria, Homs. Image from Casale & Assmann, 2017
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Taphoxenus (Lychnifugus) cellarum (Adams, 1817)
Male adult
Azerbaijan, Baku. Image from Casale & Assmann, 2017
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Taphoxenus (Lychnifugus) cerberus cerberus (Ganglbauer, 1905)
Female adult
Amasya, Anatolia, Turkey. Image from Casale & Assmann, 2017
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Taphoxenus (Lychnifugus) elburzensis Casale & Wrase, 2018 Holotype
Male adult
Iran, Prov. Tehran, Elburz Mts, Kuh-e Demavend, 10 km W of Reine 52°02ʹ24ʺE, 35°53ʹ22ʺN 2200–2650 m, 14.V.1998 leg. Gy. Fábián et K. Székety”/“Holotypus ♂ T. (L.) elburzensis sp. n. A. Casale & D. Wrase 2018” (cCA). Image from A. Casale & D.W. Wrase, 2018
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Taphoxenus (Lychnifugus) murzini Casale & Wrase, 2018 Holotype
Male adult
Iran, W. Fars Prov. NW Sepidan, Poolad Kaf 30.366888N, 51.899993E, 3100 m, 9.VI.2014 S. Murzin leg.”/“Holotypus ♂ T. (L.) murzini sp. n. A. Casale & D. Wrase 2018” (cWR). Image from A. Casale & D.W. Wrase, 2018
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Taphoxenus (Lychnifugus) sahendensis (Morvan, 1981)
Male adult
NW Iran: Azerbaijan: Sahand Mts. Image from A. Casale & D.W. Wrase, 2018
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Taphoxenus (Taphoxenus) trochanteratus Emden, 1954
Male adult
C-Afghanistan, Bamyan prov., Bandi-Amir env., 3000 m, 14-19.07.2013, O.Pak lg.
Copyright © Igor Solodovnikov