Bembidion (Actedium) pallidipenne (Illiger, 1802b)
Male adult
length: 4,1-4,7 mm, Niederlande, Maasvlakte, leg. A.Teunisse, 1989; det. G.Rößler, 1996, Foto: U.Schmidt, 2010
Copyright © U.Schmidt

Bembidion (Andrewesa) patris Schmidt, 2010 Paratype
Male adult
NEPAL Annapurna Mts. / Madi Khola below Sikles / 1400 m, water power st. [station, 28°22N 84°08E] / 10.5.1996 leg. J. Schmidt. Image from Schmidt, 2010
Copyright © - - -

Bembidion (Antiperyphanes) mandibulare (Solier, 1849)
Male adult
Chile: Reg. X, Chiloé: Cucao, DRM voucher V100789. Image from Maddison D.R., 2014. Scale bar 1 mm.
Copyright © - - -

Bembidion (Antiperyphanes) hirtipes (Jeannel, 1962)
Male adult
Argentina, Mendoza, Pampa Palauco. Image from Maddison D.R., 2014.
Copyright © - - -

Bembidion (Antiperyphanes) rufoplagiatum Germain, 1906
Male adult
Argentina, Neuquen, Arroyo Queni at Lago Queni. Image from Maddison D.R., 2014.
Copyright © - - -

Bembidion (Antiperyphanes) zanettii Toledano, 2008
Male adult
Ecuador: Napo: Rio Quijos W of Baeza. Image from Maddison D.R., 2014.
Copyright © - - -

Bembidion (Antiperyphanes) spinolai Solier, 1849
Male adult
Argentina: Chubut: Rio Azul at Lago Puelo. Image from Maddison D.R., 2014.
Copyright © - - -

Bembidion (Antiperyphus) philippii Germain, 1906
Male adult
Argentina: Neuquén: Rio Collón Curá ca 13 km S La Rinconada, DRM voucher V100787. Image from Maddison D.R., 2014. Scale bar 1 mm.
Copyright © - - -

Bembidion (Apteromimus) platyderoides (Wollaston, 1877)
Male adult
St. Helena Island. Image courtesy of David Maddison
Copyright © CC-BY 4.0

Bembidion (Apteromimus) shepherdae Maddison, 2019
Female adult
St Helena
Copyright © David Maddison

Bembidion (Archaeophilochthus) christelae Ortuno & Arillo, 2010 Holotype
Male adult
Baltic Eocene amber
Copyright © Alexander Anichtchenko

Bembidion (Armatocillenus) sauteri (Jedlicka, 1958) Holotype
Female adult
Formosa, Sauter // Anping, 1909.X. // Typus [red] // Armatocillenus sauteri n.sp. [handwritten], det. ING. JEDLIČKA [pink] (HNHM).
Copyright © - - -

Bembidion (INCERTAE) bandotaro Morita, 1991
Female adult
Kuril Islands
photo Makarov K.
Copyright © Kirill Makarov

Bembidion (Asioperyphus) infuscatum Dejean, 1831
Female adult
length: 6,4 mm
West Siberia, KHMAO-Yugra, Khanty-Mansiysk distr., Ob river, Bolshoy Tchukhtinsiy island
4.08.2007.
Zinoviev E.leg.
Copyright © Evgeniy Zinoviev

Bembidion (Asioperyphus) postremum Say, 1830
Female adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Asioperyphus) altestriatum Netolitzky, 1934a: 68
Male adult
Russia, SE Primorie, Lazo Nat. res., 10 km SW Sokol'chi vill.
(Приморский кр., Лазовский зап., 10 км SW пос. Сокольчи, окр. кордона Корпадь 43°15' N, 134°7' E, 23-27.06.10, leg. М.Смирнов)
Image of M. Smirnov (М.Э. Смирнов) from www.zin.ru\Animalia\Coleoptera
Copyright © Maksim Smirnov

Bembidion (Asioperyphus) serorum Netolitzky, 1934
Male adult

Copyright © Kirill Makarov

Bembidion (Asioperyphus) pamiricola pamiricola Lutshnik, 1930: 34
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Asioperyphus) bandotaro Morita, 1991a: 119
Female adult
Kuriles isl. det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Asioperyphus) ovale Motschulsky, 1844: 239
Male adult
Siberia, E Sajan Mts., Sentsa riv, trib. of Oka riv., 1700m, 9-10.7.1998, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Bembidion (Asioperyphus) lunatum Duftschmid, 1812: 211
Male adult
Latvia, Daugavpils distr., Silene, Ilgas, 18.5.1991, A. Barsevskis leg.
Copyright © Alexander Anichtchenko

Bembidion (Asioperyphus) sajanum Shilenkov in Kryzhanovskij & al., 1995
Male adult
Siberia, E Sajan Mts., Sentsa riv, trib. of Oka riv., 1700m, 9-10.7.1998, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Bembidion (Asioperyphus) jorgeberti Neri & Toledano, 2024 Paratype
Male adult
Burjatia, East Sajan Mts, Black Irkut valley, mouth of Susar riv., 1860m, 30.6.2012, leg J. Gebert
Copyright © Jörg Gebert

Peryphus conformis Motschulsky, 1844 Lectotype
Male adult
Mt. Hamar-Dab. (ZMU)
Copyright © Kirill Makarov

Bembidion (Asioperyphus) pseudoinfuscatum Neri, Toledano & Rébl, 2023 Holotype
Male adult
Holotype, ♂, “N 49°38’50.3” E 114°41’27.1 / Mongolei Dornod Aimak / Dashbalbar NE, / Salzsee Steppe / 27.07.2012 670m / leg. Schnitter” [printed]” (CTVR).
Copyright © Qin Yuyao

Bembidion (Australoemphanes) ateradustum Liebherr, 2008
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Bembidion (Bembidion) quadrimaculatum (Linne, 1761)
Male adult
length: 2,5-3,6 mm
Germania, Oberfranken, Kulmbach
leg.det. U.Schmidt, 1972
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Bembidion (Bembidion) quadrimaculatum oppositum Say, 1823
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Bembidion) quadrimaculatum mandli Netolitzky, 1932
Female adult

Copyright © Kirill Makarov

Bembidion (Bembidion) humerale humerale Sturm, 1825: 176
Female adult
W Ukraine, Lviv city, Landscape Reserve “Peatbog Bilogorshcha” NW part, 17.IV-10.VII.2018, Panin R. leg., L =2.95 mm
Copyright © Denis Khrapov

Bembidion (Bembidionetolitzkya) conforme Dejean, 1831
Male adult
length: 4,5-6 mm
Germania, Oberbayern, Oberstdorf
leg.det. U.Schmidt, 2005
Foto: U.Schmidt, 2006
Copyright © U.Schmidt

Bembidion (Bembidionetolitzkya) geniculatum Heer, 1837
Female adult
Ukrainian Carpathians,
Ivano-Frankivsk reg.,Yaremcha distr.,Gorgany Mts. ,h=750 m. 22.VI.08 Panin R. leg.
7mm.
Copyright © Ruslan Panin

Bembidion (Bembidionetolitzkya) cyaneum Chaudoir, 1846a: 203
Male adult
N Кавказ, Карачаево-Черкесия, окр. г.Теберда 1.VII.1994 leg. А. Гусаков
N Caucas, Karachaevo-Cherkessia, Teberda vicinity
Image of M. Smirnov (М.Э. Смирнов) from www.zin.ru\Animalia\Coleoptera
Copyright © Maksim Smirnov

Bembidion (Bembidionetolitzkya) tibiale (Duftschmid, 1812)
Male adult
NW Кавказ, 20 км S г. Горячий Ключ, бер. горной речки 3.04.1996 leg. М. Смирнов
NW Caucas, 20 km S from Goriachii Kluch
Image of M. Smirnov (М.Э. Смирнов) from www.zin.ru\Animalia\Coleoptera
Copyright © Maksim Smirnov

Bembidion (Bembidionetolitzkya) cyaneum Chaudoir, 1846
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Bembidionetolitzkya) varicolor agurense Lutshnik, 1938
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Bembidionetolitzkya) longipes K. Daniel, 1902a: 29
Male adult
France, 05380 Chateauroux les Alpes Le Rabioux, 02-VII-2014
Copyright © Pascal Deschamps

Bembidion (Bembidionetolitzkya) cimmerium Andrewes, 1922c: 238
Male adult
India, Uttarakhand, 5km NW Ghangaria, Valley of Flowers, N30*43'356" E079*85.309 , 3417 m, 06.2011, Anichtchenko A. leg. 5.5 mm. det. J. Schmidt 2015
Copyright © Alexander Anichtchenko

Bembidion (Bembidionetolitzkya) penninum Netolitzky, 1918a: 20
Male adult
Val Cairasca Sempione VI-1915 Mancini
Copyright © Jan Muilwijk

Bembidion (Bembidionetolitzkya) chaklaense Schmidt, 2018 Paratype
Male adult
“TIBET South Centr. 16.VII.07 / 50 km NNE Lhasa, SW of / Chak La pass 4100-4550m / Chakla river valley, lg. Schmidt”, “30°03’36.6N 91°16’51.7E / to / 30°06’10.1N 91°16’38.4E” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) lhai Schmidt, 2018 Paratype
Male adult
“TIBET South Centr. 17-20.VI.07 / Budha Vall. N of Yangpachem / ca. 30°10’56N 90°29’21E / 5000-5200m, leg. J. Schmidt” (CSCHM).
Copyright © Alexander Anichtchenko

Bembidion (Bembidionetolitzkya) pinnigerum Schmidt, 2018 Holotype
Male adult
“TIBET South Centr. 17/18.VII.07 / Reting Tsangpo Vall. E Reting / 4230-4800 m, Kiykiy place / ca. 30°24’15N 91°42’59E lg.J.S.” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) lunepabomaguvi lunepabomaguvi Paratype
Male adult
“INDIA Arunachal Pradesh / 10-15 km W Sela Pass / 34-3500 m, 24.VI.2008 / 27°30N 92°03E lg. Reuter” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) lunepabomaguvi bhutanicum Schmidt, 2018 Paratype
Male adult
“BT Wangdue Phodrang distr. / Phobjikha Vall., 2800 m / 28.IX.2017, leg. J. Schmidt / 27°24’40”N 90°12’12”E” (NBCB).
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) namtso Schmidt, 2018 Paratype
Male adult
“TIBET (South Central) 12.VII.07 / S Namtso, Langma Valley / 5100-5150 m, leg. Schmidt / 30°37’39,1”N90°51’56,5”E” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) retingense Schmidt, 2018 Paratype
Male adult
“CHINA Tibet 1.VII.2015 / side valley S Reting / 4400 m leg. J. Schmidt / 30°16’44”N 91°31’55”E” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) smetanai qingmaiense Schmidt, 2018 Paratype
Male adult
“China, S.W. Sichuan / Qingmai 3000 m / N 28°48’46.7”, E 099°52’09.2” / 3.7.2006, lgt. Janata, M.” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) transhimalayanum Schmidt, 2018 Paratype
Female adult
“S TIBET, ca. 40km SW of / Yangpachem, 13.VII.2010 / Pijin moraine 5000-5500 m / 29°45’36N 90°17’54E” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) wrzecionkoi Toledano & Schmidt, 2008
Male adult
south central Tibet, NE of Gyantse, Yung La pass and above, 4850-5400 m, 28°57’14”N 89°50’04”E, 17.VII.2010
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) yamtso Schmidt, 2018 Holotype
Male adult
“S TIBET 14/15.VI.2011 / E-slope Karo La pass / 4900-5300 m, leg. Schmidt / 28°55’N 90°18’E” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) zugmayeri Schmidt, 2018 Holotype
Male adult
“Sumdschiling / Ebene [plateau] W Tibet”, “Süssw. Tümp. [fresh water pond] / VII/06. Zugm.” (ex coll. W. Marggi, Thun, in CSCHM).
Copyright © Joachim Schmidt

Bembidion (Bembidionetolitzkya) bactrianum K. Daniel, 1902
Male adult
Tadzhikistan, Varzob, Khodja obi Garm, 2000m 21-24.3.2015 (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Bembidionetolitzkya) tibiale (Duftschmid, 1812)
Female adult

Copyright © Kirill Makarov

Bembidion (Bembidionetolitzkya) tibiale (Duftschmid, 1812)
Male adult

Copyright © Kirill Makarov

Bembidion (Bembidionetolitzkya) conforme Dejean, 1831
Male adult
W Ukraine, NE Carpathians, Tiachiv distr., h~400 m, bank of stream Mala Ugolka, 23.VI.2016, Panin R. leg.
Copyright © Ruslan Panin

Bembidion (Bembidionetolitzkya) varicolor varicolor
Male adult
CZECH REPUBLIC - Bystřice nad Olší, 15.III.2020, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar

Bembidion (Bembidionetolitzkya) geniculatum claudejeannei Marggi in Löbl & Smetana, 2003
Male adult
Spain, Sierra Nevada, rio Lanjaron, 1200m, 29.3.2010, Anichtchenko A. leg. (cAA). 5 mm; det. M. Toribio
Copyright © Alexander Anichtchenko

Bembidion (Bembidionetolitzkya) coeruleum Audinet-Serville, 1821
Male adult
Marocco c., Asni, National Park Toubkal
Copyright © Paolo Bonavita

Bembidion (Bembidionetolitzkya) toubkalense Bonavita & Vigna Taglianti, 2018 Paratype
Male adult
Morocco, Toubkal
Copyright © Paolo Bonavita

Bembidion (Bembidionetolitzkya) scelio (Antoine, 1945)
Male adult
Maroc, Haut-Atlas, Gorges du Todra 1500 m
Copyright © Paolo Bonavita

Bembidion (Bembidionetolitzkya) cassolai (Bonavita & Vigna Taglianti, 1993)
Male adult
Italy, Sardegna, Medio Campedano, Gonnosfanadiga, 1.5km SW Riu Piras. leg & det. J. Gebert
Copyright © Jörg Gebert

Bembidion (Blepharoplataphus) hastii C.R. Sahlberg, 1827
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Blepharoplataphus) hastii C.R. Sahlberg, 1827
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Chilioperyphus) orregoi Germain, 1906
Male adult
Argentina: Chubut: Rio Azul at Lago Puelo. Scale bar 1 mm. Image from Maddison D.R., 2014.
Copyright © - - -

Bembidion (Chlorodium) splendidum Sturm, 1825: 145
Female adult

Copyright © Kirill Makarov

Bembidion (Chlorodium) pygmaeum ab. bilunulatum Bielz, 1852
Male adult
N Moravia, Týn nad Bečvou; 2,5mm. det. J. Coulon, 2015
Copyright © Aleš Sedláček

Bembidion (Chlorodium) pygmaeum (Fabricius, 1792)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Chlorodium) splendidum luteipes (Motschulsky, 1844)
Male adult

Copyright © Kirill Makarov

Bembidion (Chlorodium) leucolenum Bates, 1873a: 275
Female adult
Kunashir Island, lower reaches of Krivonozhka Stream, 20-22.VI.2013, Yu. & L. Sundukov leg.
Copyright © Kirill Makarov

Bembidion (Chlorodium) leucolenum Bates, 1873a: 275
Male adult
Kunashir Island, watershed of Golovnina and Khlebnikova rivers west of Dubovoe, 28.IX.2015, Yu. Sundukov
Copyright © Kirill Makarov

Bembidion (Chlorodium) difforme (Motschulsky, 1844)
Female adult
Russia, Burjatia, Muhor-Shibirsk distr., 5km N Nikolsk vill., salt marsh, 27.8.1998, Anichtchenko A. leg & det.
Copyright © Alexander Anichtchenko

Cillenus (Cillenus) lateralis Samouelle, 1819
Male adult
Netherlands, Bruinisse, leg. Krift, 1976; det. U.Schmidt, 1982
Photo: U.Schmidt, 2008
Copyright © Udo Schmidt

Bembidion (Cillenus) palosverdes Kavanaugh and Erwin, 1992
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Bembidion (Cillenus) lateralis Samouelle, 1819
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Cillenus (Desarmatocillenus) sumaoi (Morita, 1981)
Male adult
Chiba, Honshu, Japan<br /> (c) Field Museum of Natural History <br /> (Maruyama Munetoshi)
Copyright © Munetoshi Maruyama

Cillenus (Desarmatocillenus) kasaharai Habu, 1978 Holotype
Male adult
Oohara, Iriomote Is., OKINAWA, 11. VI. 1976, S. Kasahara
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-024
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Cillenus (Desarmatocillenus) kasaharai Habu, 1978 Holotype
Male adult
Oohara, Iriomote Is., OKINAWA, 11. VI. 1976, S. Kasahara
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-024
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Cillenus (Desarmatocillenus) kasaharai Habu, 1978 Holotype
Male adult
Oohara, Iriomote Is., OKINAWA, 11. VI. 1976, S. Kasahara
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-024
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Cillenus (Desarmatocillenus) kasaharai Habu, 1978 (Label) Holotype
Male adult
Oohara, Iriomote Is., OKINAWA, 11. VI. 1976, S. Kasahara
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-024
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Cillenus (Desarmatocillenus) yokohamae (Bates, 1883)
Male adult

Copyright © Kirill Makarov

Bembidion (Desarmatocillenus) aestuarii (Habu & Ueno, 1954)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Bembidion (Desarmatocillenus) kasaharai Habu, 1978
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Bembidion (Desarmatocillenus) yakushimanum Sasakawa, 2007
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidion (Desarmatocillenus) foochowense Lindroth, 1980
Male adult
TAIWAN: Lunglungchi, Fulung, Taipei Hsien, 31.vii.2001, Terada lgt.
Copyright © - - -

Bembidion (Diplocampa) transparens (Gebler, 1829)
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Diplocampa) fumigatum (Duftschmid, 1812)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Diplocampa) transparens (Gebler, 1829)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Diplocampa) transparens prostratum (Motschulsky, 1844)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Diplocampa) blandulum Netolitzky, 1910a: 221
Male adult
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin L=3.6 mm
Copyright © Kirill Makarov

Bembidion (Diplocampa) blandulum Netolitzky, 1910a: 221
Female adult
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin L=4.0 mm
Copyright © Kirill Makarov

Bembidion (Diplocampa) osellai Neri & Toledano, 2023 Holotype
Male adult
1 ♂, Holotype, (3.40 mm), Median lobe of the aedeagus, holotype (0.64 mm) “Turchia – vil. Gümüshane / Bayburt / m 1200, 1.VII.75 / Osella leg. [printed]” (PN); the specimen is not in good conditions, lacking the two terminal right antennomeres, fore right leg, median left leg and left hind tarsomeres. We added to the specimen the following label [red, printed]: “Bembidion (Diplocampa) osellai P. Neri & L. Toledano, 2022 – Holotypus”
Copyright © Qin Yuyao

Bembidion (Diplocampa) acutangulum Neri & Toledano, 2023 Holotype Paratype
Female adult
1 ♀, paratype, (3.50 mm); “Armenia: ca. 10 km N Sevan / 40°36’48”N 44°57’29”E / nr. Tsovagyugh village / 22.5.2001, ca. 1940m / Shaverdo & Schillhammer [printed]” (NHMW) Median lobe of the aedeagus, holotype (0.68 mm) (NHMW)
Copyright © Qin Yuyao

Bembidion (Diplocampa) loeffleri Jedlicka, 1963c: 169 Holotype Paratype
Male adult
1 ♂, holotype, (3.80 mm); “Persien / Dr. Löffler // Holotype (pink) // Montage / Morvan / 1974 n. 474 (handwritten) // Mus. Nat. Pragae (red, printed) / inv. 66303 (handwritten) // Diplocampa / löffleri sp. n. (handwritten) / det. Ing. Jedlička (pink, printed)” (NMPC) Median lobe of the aedeagus, ♂ paratype (0.73 mm), “Persien / Dr. Löffler // Paratypus (pink) // Mus. Nat. Pragae (red, printed) / inv. 66304 (handwritten) // löffleri sp. n. (handwritten) / det. Ing. Jedlička (pink, printed)” (NMPC).
Copyright © Qin Yuyao

Bembidion (Diplocampa) blandulum Netolitzky, 1910a: 221 Holotype
Male adult
1 ♂, holotype, (3.80 mm); “Persia sett. / Doria 62 [handwritten] // Mus. Civ. / Genova [printed] // Diplocampa / blandulum Net. [handwritten] / det. Netolitzky [printed] // Holotypus [printed] / Bembidion (Diplo / campa) blandulum / Netolitzky, 1910 [pink, handwritten]” (MCSNG) the specimen is in rather good condition, missing only the left antenna, two right antennomeres and a few tarsomeres of the right median leg.
Copyright © Qin Yuyao

Bembidion (Ecuadion) ricei Maddison & Toledano, 2012 Paratype
Male adult
ECUADOR: Napo: Rio Chalpi Grande, 2800m, 0.3645°S, 78.0852°W, D.R. Maddison voucher V100622
Scale bar is 1 mm
image from (Maddison & Toledano, 2012)
Copyright © ---

Bembidion (Ecuadion) georgeballi Toledano, 2008
Male adult
ECUADOR: Pichincha: Quebrada Lozada, on road to Res. Yanacocha, 3460m, 0.1105°S, 78.5642°W, voucher V100658<br /> Scale bar is 1 mm.<br /> Image from Maddison & Toledano, 2012
Copyright © - - -

Bembidion (Ecuadion) andersoni Toledano, 2008
Male adult
ECUADOR: Pichincha: Reserva Yanacocha, start Andean Snipe Trail, DRM voucher V100655. Image from Maddison D.R., 2014. Scale bar 1 mm.
Copyright © - - -

Bembidion (Ecuadion) chimborazonum (Bates, 1891)
Male adult
ECUADOR: Pichincha: Paso de la Virgen, DRM voucher V100793. Image from Maddison D.R., 2014.
Copyright © - - -

Bembidion (Ecuadion) sanctaemarthae Darlington, 1934
Male adult
ECUADOR: Napo: Rio Chalpi Grande. Image from Maddison D.R., 2014.
Copyright © - - -

Bembidion (Ecuadion) ricei Maddison & Toledano, 2012
Male adult
ECUADOR: Napo: Rio Chalpi Grande. Scale bar 1 mm. Image from Maddison D.R., 2014.
Copyright © - - -

Bembidion (Ecuadion) cotopaxi Moret & Toledano, 2002
Male adult
ECUADOR: Pichincha: km 17 on route 28 W of Papallacta. Scale bar 1 mm. Image from Maddison D.R., 2014.
Copyright © - - -

Bembidion (Ecuadion) walterrossii Toledano, 2008
Male adult
ECUADOR: Napo: Vinillos, 4.1 km S Cosanga. Scale bar 1 mm. Image from Maddison D.R., 2014.
Copyright © - - -

Bembidion (Ecuadion) cotopaxi Moret & Toledano, 2002 Paratype
Female adult
Ecuador, Cotopaxi, Laguna Quilotoa, 3500m, 22.7.2001 P. Moret leg.
Copyright © Alexander Anichtchenko

Bembidion (Ecuadion) agonoides Vigna-Taglianti & Toledano, 2008
Female adult
Ecuador, Provincia Cotopaxi, Otonga Nature Reserve, 0°25’15” S, 79°00’23” W, 1800-2000 m
Copyright © Pierre Moret

Bembidion (Ecuadion) walterrossii Toledano, 2008: 94
Female adult
Ecuador, Provincia Cotopaxi, Otonga Nature Reserve, 0°25’15” S, 79°00’23” W, 1800-2000 m
Copyright © Pierre Moret

Bembidion (Emphanes) tenellum Erichson, 1837
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Emphanes) latiplaga flavibase De Monte, 1956a: 179
Male adult
Morocco, Souss-Massa Draa, Zagora, OuedDraa riv., 1.2.2013. det. M. Toribio, 2017. 2.5 mm
Copyright © Alexander Anichtchenko

Bembidion (Emphanes) bulgani Jedlicka, 1968
Female adult
Russia: Primorskii krai: 20 km NWW Spassk-Dal'ny, Khanka Lake, Khankaysky Nat. Res., kordon Vostochnyi, 3-8.VI 2008, A. Napolov leg.
Copyright © Kirill Makarov

Bembidion (Emphanes) tenellum tenellum Erichson, 1837
Male adult
C Russia, Ivanovo ar., Pestyaki distr., Demidovo vill. env., Lukh riv. 4-5.VII.2014 leg. M. Smirnov ; Фотография с сайта www.zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Bembidion (Emphanes) latiplaga latiplaga Chaudoir, 1850
Female adult
NW-Ukraine, Volhynia, Turiysk env., 17-26.VI.2016, to light, Voytko P. leg./ Panin R. det., L=3.1 mm
Copyright © Denis Khrapov

Bembidion (Emphanes) normannum normannum Dejean, 1831
Male adult
Spain, Guipuzcoa, Playaundi, 20.3.2006, Anichtchenko A. leg. (cAA)
Copyright © Alexander Anichtchenko

Bembidion (Endosomatium) megalops (Wollaston, 1877)
Female adult
St. Helena Island. Image courtesy of David Maddison
Copyright © CC-BY 4.0

Bembidion (Euperyphus) combustum combustum Menetries, 1832: 138
Female adult
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin L=7.0 mm
Copyright © Kirill Makarov

Bembidion (Euperyphus) giganteum giganteum J.R. Sahlberg, 1900
Female adult
S Kazakhstan, N slope Talas Mt.rng., Dzhabagly riv., 1100m 28.7.2004, A.Matalin leg. (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Peryphus) morawitzi Csiki, 1928
Female adult
China, Liaoning, Kuandian co., Banlaing, Ba-che-chuan (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Euperyphus) scapulare lomnickii Netolitzky, 1916: 260
Female adult
W Ukraine, NE Carpathians, Lviv reg., near Rozgirche vill., bank of Stryi river, 11.V.2014, Khrapov D. leg./det.Panin R., L= 5.4 mm
Copyright © Denis Khrapov

Bembidion (Euperyphus) scapulare lomnickii Netolitzky, 1916: 260
Male adult
W-Ukraine, Transcarpathian region, Vynogradiv distr., bank of Tisza river near Mala Kopanya vill., 5-7.VI.2018, Panin.R leg., L=5,8 mm
Copyright © Denis Khrapov

Bembidion (Euperyphus) eques Sturm, 1825
Female adult
Italy, Moggio Udinese, Torrente Aupa, 6.9.2023, 13.2007° E, 46.4084° N, J. Gebert leg.
Copyright © Jörg Gebert

Bembidion (Euperyphus) fluviatile fluviatile Dejean, 1831: 113
Male adult
CZECH REPUBLIC - Polanka nad Odrou, 6.IV.2017, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar

Bembidion (Euperyphus) scapulare tergluense Netolitzky, 1918
Male adult
Italy, Fiume Tagliamento, Cornino, iv.2024, leg. Jörg Gebert
Copyright © Jörg Gebert

Bembidion (Eupetedromus) graciliforme Hayward, 1897
Female adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Eupetedromus) sibiricum Dejean, 1831a: 66
Female adult
S Primorie, Ussurijski Reserve, Mirinov Klyuch 43°38.644' N 132°26.693' E 25.VIII.2007 leg. K.Makarov, A.Zaitzev
L =4.7
Copyright © Kirill Makarov

Bembidion (Eupetedromus) dentellum (Thunberg, 1787)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Eupetedromus) ruthenum Tschitscherine, 1895c: 298
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Eupetedromus) immaturum Lindroth, 1954 Paratype
Male adult
W Newfoundland, Deer Lake, 30.V.51, Lindroth. 4254:2 (MZLU)
Copyright © Lund University

Bembidion (Eurytrachelus) pogonoides Bates, 1883
Female adult
Far East, Lazovsky Nat.Res., shore of Japan Sea, cordon Petrova, Petrovski spring 10.06.1995 leg. Yu.Sundukov
female
L=6,2 mm
Copyright © Maksim Smirnov

Bembidion (Eurytrachelus) nitidum (Kirby, 1837)
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Eurytrachelus) laticolle (Dufischmid, 1812)
Female adult
Orenburg Reg., Ilek Distr., Ilek env., valley of Ural Riv., beach 28.VII-8VIII.2005 S.Vastschenko leg.
L=6.2 mm
Copyright © Kirill Makarov

Bembidion (Eurytrachelus) vitiosum Gemminger & Harold, 1868
Female adult
Russia, Irkutsk reg., Nizhneudinsk distr., Uda riv., Bolshoy Uk waterfall, 22.7.1898, A.Anichtchenko leg. (DUBC).
Copyright © Alexander Anichtchenko

Bembidion (Furcacampa) versicolor (Leconte, 1848)
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Furcacampa) impotens Casey, 1918
Female adult
Great Smoky Mountains National Park
photo I. M. Sokolov
Copyright © Igor Sokolov

Bembidion (Furcacampa) affine Say, 1823
Male adult
Great Smoky Mountains National Park
photo I. M. Sokolov
Copyright © Igor Sokolov

Bembidion (Geocosmius) elescarabajo Maddison, 2023
Male adult

Copyright © David Maddison

Bembidion (Geocosmius) franiae Erwin, 1982
Male adult

Copyright © David Maddison

Bembidion (Gnatholymnaeum) spurcum Blackburn, 1881
Male adult
Hawaii. Image from J.K. Liebherr, 2008
Copyright © - - -

Bembidion (Hirmoplataphus) nigrum Say, 1823
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Hirmoplataphus) hirmocaelum Chaudoir, 1850
Female adult
Siberia, Irkutsk, Irkut riv., 3.6.1997, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Bembidion (Hirmoplataphus) pliculatum Bates, 1883
Male adult
S Sakhalin isl., Susuya riv. leg & det. Sundukov
Copyright © Kirill Makarov

Bembidion (Hirmoplataphus) hirmocaelum Chaudoir, 1850
Male adult
Primorski reg., Kedrovaya riv. det. Sundukov, 2023
Copyright © Kirill Makarov

Bembidion (Hydrium) laevigatum Say, 1830
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Bembidion (Hydrium) laevigatum Say, 1830
Male adult
USA: Utah: Grand Co., Moab, Colorado River, 1190 m, 38.5739°N, 109.5769°W
Copyright © David Maddison

Bembidion (Hydrium) mimbres Maddison, 2020 Paratype
Male adult
USA: New Mexico: Grant Co., Gila River, Billings Vista, 1320 m, 32.8163°N, 108.6032°W, 11.viii.2005. DRM 05.043. D.R. & J.H.A. Maddison
Copyright © David Maddison

Bembidion (Jammuphanes) sinuosum Schmidt & Marggi, 2014 Paratype
Male adult
Pakistan, Kaghan valley, Shogran, 3050m, 19.6.2001. Imagr from Schmidt & Marggi, 2014
Copyright © - - -

Bembidion (Josefia) praeceptor Schmidt, 2018 Paratype
Male adult
“Tibet (South Central) 26.VI.07 / Gyama vall. ca. 60km E Lhasa / 4230-4620 m leg. J. Schmidt / ca. 29°41’48N 91°38’29E”, “south ascent of / Chungenpo Shong side vall” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Leuchydrium) tigrinum Leconte, 1879
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidion (Limnaeoperyphus) quadriimpressum (Motschulsky, 1860)
Male adult
Kunashiri Isl., mouth of Medny Spring 44°16'31"N 145°56'29"E 27.VI.2008 leg. K.Makarov
L=4.3
Copyright © Kirill Makarov

Bembidion (Limnaeoperyphus) quadriimpressum (Motschulsky, 1860)
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidion (Liocosmius) darlingtonielum Cooper & Maddison, 2014
Male adult

Copyright © David Maddison

Bembidion (Liocosmius) cachagua Maddison, 2023 Holotype
Male adult
CAS: “U.S.A.: CALIFORNIA: Monterey Co., Hastings Nat. History Reserve. J.M. Linsdale Colln.”, “C.A.S. Accession 29-V-48 Cachagua Creek”, “David R. Maddison DNA3842 DNA Voucher”
Copyright © David Maddison

Bembidion (Lymnaeum) nigropiceum (Marsham, 1802)
Male adult
Thompson Island, 7.5.2008, Massachusetts
Images J. Rawlins
Copyright © ---

Bembidion (Lymnaeum) laticeps (Leconte, 1858)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Bembidion (Lymnaeum) nigropiceum (Marsham, 1802)
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidion (Plataphus) altaicum Gebler, 1833
Male adult
Primorie reg., Lazo distr., near Lazo vill. Lazovka riv. 20.06.04 leg. M.& L. Smirnov photo from www.zin.ru
Copyright © Maksim Smirnov

Bembidion (Metallina) properans (Stephens, 1828b): 26 (Tachypus)
Female adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Metallina) lampros (Herbst, 1784)
Male adult
length: 2,7-4,1 mm
Germania, Bayern, Oberfranken, Neuenmarkt
leg.det. U.Schmidt, 1972
Photo: U.Schmidt, 2009
Copyright © U.Schmidt

Bembidion (Metallina) lampros (Herbst, 1784)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Metallina) elevatum (Motschulsky, 1844)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Metallina) properans (Stephens, 1828)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Neja) nigricorne Gyllenhal, 1827: 402
Female adult
Czech Republic, Bohemia occ., Kraslice, Tisovec hill, 750-755 m a.s.l., 24.iv.1999, lgt. et det. J. Hejkal
Copyright ©

Bembidion (Neobembidion) constricticolle Hayward, 1897
Male adult

Copyright © David Maddison

Bembidion (Neoemphanes) shimoyamai Habu, 1978
Male adult
Far East, khanka Lake env., Bol.Usachi Riv. near Novokachalinsk 5.VII.2004 R.Filimonov leg.
L=3.5 mm
Copyright © Kirill Makarov

Bembidion (Neoemphanes) shimoyamai Habu, 1978 Holotype
Male adult
Oirase, Nishitsugaru-gun, AOMORI, 10. VIII. 1973, K. Shimoyama
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-036
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Neoemphanes) shimoyamai Habu, 1978 Holotype
Male adult
Oirase, Nishitsugaru-gun, AOMORI, 10. VIII. 1973, K. Shimoyama
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-036
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Neoemphanes) shimoyamai Habu, 1978 Holotype
Male adult
Oirase, Nishitsugaru-gun, AOMORI, 10. VIII. 1973, K. Shimoyama
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-036
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Neoemphanes) shimoyamai Habu, 1978 Holotype
Male adult
Oirase, Nishitsugaru-gun, AOMORI, 10. VIII. 1973, K. Shimoyama
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-036
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Neoemphanes) shimoyamai Habu, 1978 (Label) Holotype
Male adult
Oirase, Nishitsugaru-gun, AOMORI, 10. VIII. 1973, K. Shimoyama
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-036
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Nepha) menetriesii (Kolenati, 1845)
Female adult
Russia, Orenburg reg., Sol'-Iletskii distr., 10 km W Troitsk vill. 6.05.09
leg. M. & L. Smirnov's
Female
photo M. Smirnov from www.zin.ru
Copyright © Maksim Smirnov

Bembidion (Nepha) genei illigeri Netolitzky, 1914
Male adult
length: 4,0-4,9 mm
Germania, Bayern, Oberfranken, Selbitz
leg.det. U.Schmidt, 2001
Foto: U.Schmidtz, 2010
Copyright © U.Schmidt

Bembidion (Nepha) genei illigeri Netolitzky, 1914: 54
Female adult
length: 4,6 mm
South Urals, Chelyabinsk area, Asha distr., the vicinities of Sim town, Sim river valley
14.05.2001.
Rudoiskatel' P.leg.
From the collection of A.Ivanov (Ekaterinburg)
Copyright © Evgeniy Zinoviev

Bembidion (Nepha) tetrasemum Chaudoir, 1846a: 207
Male adult
NW Caucas, Krasnodar reg., Tuapse distr., Podkhrebetovoe vill. vicinity, 19.V.2012 leg. M. Smirnov
Фотография с сайта zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Bembidion (Nepha) caucasicum (Motschulsky, 1844)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Nepha) schmidti subcallosum Wollaston, 1864: 71
Male adult
3,6 mm; Spain, Canary Islands, Tenerife, Teno, Erjos, 1000 m, leg. J.Schoenfeld, 11.VII.2010; det. Lompe, 2012. Photo: U.Schmidt, 2014
Copyright © Udo Schmidt

Bembidion (Nepha) hauserianum Netolitzky, 1918
Female adult
Iran, 20 km SW Yasuj, 5-6.V.2007, Anichtchenko A. leg.
Copyright © Alexander Anichtchenko

Bembidion (Nepha) hauserianum Netolitzky, 1918a: 23
Male adult
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin L=4.1 mm
Copyright © Kirill Makarov

Bembidion (Nepha) hauserianum Netolitzky, 1918a: 23
Female adult
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin L=4.2 mm
Copyright © Kirill Makarov

Bembidion (Nepha) glabrum (Motschulsky, 1850)
Male adult
Новосибирская обл., 9 км З г. Карасук, оз Кротовое. Image from Dudko, Bespalov, Zinovyev & Lyubechanskii, 2018
Copyright © - - -

Bembidion (Nepha) callosum subconnexum De Monte, 1953
Male adult
Spain, Gredos Mts., 2km E from La Mira, 1800m, 4-7.7.2007, A. Anichtchenko leg. & det.
Copyright © Alexander Anichtchenko

Bembidion (Nepha) schmidti alluaudi (Antoine, 1925)
Male adult
Morocco, Grand Atlas, 2200m Oukaimedene env., 16-17..6.2010, Anichtchenko A. leg. (DUBC). det. M. Toribio
Copyright © Alexander Anichtchenko

Bembidion (Nepha) ibericum Piochard de la Brulerie, 1868
Male adult
Spain, Sierra de Guadarrama, Laguna de los Cleveles, 6.VIII.2024, 2000m, Anichtchenko A. leg & det. (cAA).
Copyright © Alexander Anichtchenko

Bembidion (Nepha) genei trinacriae (Bonavita & Vigna-Taglianti, 2010)
Female adult
S Spain, Sierra de las Nieves, Los Quejigales 6-8.4.2010, Anichtchenko A. leg & det. (cAA)
Copyright © Alexander Anichtchenko

Bembidion (Nipponobembidion) ainu Habu & Baba, 1968a: 145 Holotype
Male adult
Mt. Daisetsu, HOKKAIDO, 26. VI. 1958, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-034
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Nipponobembidion) ainu Habu & Baba, 1968a: 145 Holotype
Male adult
Mt. Daisetsu, HOKKAIDO, 26. VI. 1958, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-034
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Nipponobembidion) ainu Habu & Baba, 1968a: 145 Holotype
Male adult
Mt. Daisetsu, HOKKAIDO, 26. VI. 1958, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-034
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Nipponobembidion) ainu Habu & Baba, 1968a: 145 Holotype
Male adult
Mt. Daisetsu, HOKKAIDO, 26. VI. 1958, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-034
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Nipponobembidion) ruruy Makarov & Sundukov, 2014 Holotype
Male adult
Northern Kunashir, 2.5 km NW of Cape Nelyudimyi, N44°29.433', E146°11.783', 4.VIII.2013, leg. K. Makarov & Y. Sundukov” and “HOLOTYPE Bembidion (Nipponobembidion) ruruy Makarov & Sundukov 2014 [printed on red paper]
Copyright © Kirill Makarov

Bembidion (Nipponobembidion) ruruy Makarov & Sundukov, 2014 Paratype
Female adult

Copyright © Kirill Makarov

Bembidion (Notaphemphanes) ephippium (Marsham, 1802)
Male adult
S Hungary, Kiskunsági Nemzeti Park; 3,2mm
Copyright © Aleš Sedláček

Bembidion (Notaphemphanes) ephippium (Marsham, 1802)
Female adult

Copyright © Kirill Makarov

Bembidion (Notaphocampa) niloticum Dejean, 1831
Female adult
4,4 mm
Namibia, Etosha National Park, Okaukuejo, 1400 mб leg. U.Schmidt, 18.-20.III.1994;
det. M.Baehr, 2012
Photo: U.Schmidt, 2014
Copyright © Udo Schmidt

Bembidion (Notaphocampa) foveolatum Dejean, 1831
Female adult
Bembidion sobrinum Boheman, 1848 det. M.Baehr, 2014; 4,2 mm. Namibia, Otjiwarongo, Kalkfeld, 30 km East, Mt. Etjo Safari Lodge, 1500 m, leg. U.Schmidt, 15.III.1994; Photo: U.Schmidt, 2016
Copyright © Udo Schmidt

Bembidion (Notaphocampa) niloticum Dejean, 1831
Female adult

Copyright © Kirill Makarov

Bembidion (Notaphus) patruele Dejean, 1831
Female adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Notaphus) semipunctatum (Danovan, 1806)
Female adult

Copyright © Kirill Makarov

Bembidion (Notaphus) obliquum Sturm 1825: 160
Male adult
Russia, Ивановская обл., Пестяковский р-н, окр. дер. Пурежка, бер. р. Лух 24-25.VI.2011 М. Смирнов. Фотография с сайта www.zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Bembidion (Notaphus) contractum Say, 1823
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Bembidion (Notaphus) indistinctum Dejean, 1831
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Bembidion (Notaphus) flohri Bates, 1878
Male adult

Copyright © David Maddison

Bembidion (Notaphus) brownorum Maddison, Sproul & Will, 2023 Holotype
Male adult
EMEC: 39.13841/−122.34621 USA: California: Colusa Co. Antelope Valley, Freshwater Creek uv light pan trap 133 m. 1.vii.2021 K.Will
Copyright © David Maddison

Bembidion (Notaphus) mormon Hayward, 1897
Male adult

Copyright © David Maddison

Bembidion (Notaphus) callens Casey, 1918 Paralectotypes
Female adult
Tuçson [Pima County], Arizona.
Copyright © David Maddison

Bembidion (Notaphus) obtusangulum Leconte, 1863
Male adult

Copyright © David Maddison

Bembidion (Notaphus) varium (Olivier, 1795)
Female adult
CZECH REPUBLIC - Olomouc dist., Skalička, 8.IV.2020, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar

Bembidion (Notholopha) scitulum (Erichson, 1834)
Male adult
CHILE: Reg. VII: Los Niches E of Curicó, DRM voucher V100790; Image from Maddison D.R., 2014. Scale bar 1 mm.
Copyright © - - -

Bembidion (Notholopha) sexfoveatum Germain, 1906
Male adult
CHILE: Reg. IX: 16.3 km E Malalcahuello, Cuesta Las Raices, DRM voucher V100598.. Image from Maddison D.R., 2014. Scale bar 1 mm.
Copyright © - - -

Bembidion (Nothonepha) eburneonigrum Germain, 1906
Male adult
Argentina: Neuquén: Rio Neuquén at Chos Malal, DRM voucher V100785. Image from Maddison D.R., 2014.
Copyright © - - -

Bembidion (Nothonepha) tetrapholeon Maddison, 2014 Paratype
Male adult
Argentina: Neuquén: Arroyo Queñi at Lago Queñi, 830m, 40.1575°S, 71.721°W, / 10–11.ii.2007. DRM 07.035. D.R. Maddison, S.A.Roig. Scale bar 1 mm. Image from Maddison D.R., 2014.
Copyright © - - -

Bembidion (Nothonepha) tetrapholeon Maddison, 2014 Paratype
Female adult
Argentina: Chubut: Rio Azul at Lago Puelo. Scale bar 1 mm. Image from Maddison D.R., 2014.
Copyright © - - -

Bembidion (Nothonepha) lonae Jensen-Haarup, 1910
Male adult
Argentina: Mendoza: Salinas del Diamante. Scale bar 1 mm. Image from Maddison D.R., 2014.
Copyright © - - -

Bembidion (Nothonepha) engelhardti engelhardti
Male adult
Argentina: Neuquén: Rio Salado at route 40. Scale bar 1 mm. Image from Maddison D.R., 2014.
Copyright © - - -

Bembidion (Nothonepha) germainianum (Toledano, 2002)
Male adult
Argentina: Neuquén: Rio Salado at route 40. Scale bar 1 mm. Image from Maddison D.R., 2014.
Copyright © - - -

Bembidion (Nothonepha) tucumanum (Jeannel, 1962)
Male adult
Argentina: Mendoza: Salinas del Diamante. Scale bar 1 mm. Image from Maddison D.R., 2014.
Copyright © - - -

Bembidion (Ocydromus) perspicuum (Leconte, 1848)
Male adult
USA: Colorado: Huerfano Co., Huerfano River at Badito, 37.7285°N 105.0167°W.
Scale bar is 1 mm.
Photo David R. Maddison & Alexander P. Swanson
Copyright © ---

Bembidion (Ocydromus) echigonum Habu & Baba, 1957 Holotype
Male adult
Mt. Amakazari (alt. 800m), NIIGATA, 25. VI. 1955, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-030
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Ocydromus) echigonum Habu & Baba, 1957 Holotype
Male adult
Mt. Amakazari (alt. 800m), NIIGATA, 25. VI. 1955, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-030
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Ocydromus) echigonum Habu & Baba, 1957 Holotype
Male adult
Mt. Amakazari (alt. 800m), NIIGATA, 25. VI. 1955, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-030
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Ocydromus) echigonum Habu & Baba, 1957 (Label) Holotype
Male adult
Mt. Amakazari (alt. 800m), NIIGATA, 25. VI. 1955, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-030
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Ocydromus) negrei Habu, 1958 Holotype
Male adult
Mt. Osore, AOMORI, 01. VIII. 1953, A. Habu Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-033. (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html). Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Ocydromus) negrei Habu, 1958 Holotype
Male adult
Mt. Osore, AOMORI, 01. VIII. 1953, A. Habu<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-033<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Ocydromus) negrei Habu, 1958 Holotype
Male adult
Mt. Osore, AOMORI, 01. VIII. 1953, A. Habu. Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-033 (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html) Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Ocydromus) zolotarewi Reitter, 1910b: 313
Male adult
Russia, NW Caucas, Krasnodar reg., Tuapse distr., Podchrebetovoe vicinity
(NW Кавказ, Краснодарский край, Туапсинский р-н, окр пос. Подхребeтовое 19.V.2012 leg. М. Смирнов)
Image of M. Smirnov (М.Э. Смирнов) from www.zin.ru\Animalia\Coleoptera
Copyright © Maksim Smirnov

Bembidion (Peryphanes) grapii Gyllenhal, 1827
Male adult
NW Caucas, Krasnodar reg., Tuapse distr., Podkhrebetovoe vill. vicinity, 19.V.2012 leg. M. Smirnov <br /> Фотография с сайта zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Bembidion (Ocydromus) scopulinum (Kirby, 1837)
Male adult
Russia, Kunashir isl., Severjanka riv., 27.VIII.2014 (Кунашир, пойма р. Северянка)
Copyright © Kirill Makarov

Bembidion (Ocydromus) scopulinum (Kirby, 1837)
Male adult
Russia, Kunashir isl., Severjanka riv., 27.VIII.2014 (Кунашир, пойма р. Северянка)
Copyright © Kirill Makarov

Bembidion (Peryphanes) yukonum Fall, 1926: 131
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Ocydromus) transsylvanicum Bielz, 1852
Male adult
W Ukraine, NE Carpathians, Transcarpathian Reg., Marmarosh Mts., near Lug vill., bank of Kuziy (Kizi) stream, 400m, 26.IV.2014, Panin R. & Kanarsky Yu. leg. L= 5 mm
Copyright © Ruslan Panin

Bembidion (Ocydromus) decorum caraffai Sainte-Claire Deville, 1906a: 15
Female adult
France Corse
Copyright © Pascal Deschamps

Bembidion (Ocydromus) decorum subconvexum K. Daniel & J. Daniel, 1902: 102
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Ocydromus) saxatile Gyllenhal, 1827: 406
Female adult

Copyright © Kirill Makarov

Bembidion (Ocydromus) hasurada Andrewes, 1924a: 77
Male adult
India, Uttarakhand, 10-12.iV.2012, 14km E Uttarkashi, Anichtchenko A. leg.; 5 mm. det. J. Schmidt, 2015
Copyright © Alexander Anichtchenko

Bembidion (Peryphus) transversale Dejean, 1831
Male adult
Images from David R. Maddison & Alexander P. Swanson, 2010
Copyright © - - -

Bembidion (Ocydromus) saxatile morsum Netolitzky, 1930
Male adult
Tajikistan, Khoja Obi Garm, 15.9.2011, A. Barsevskis leg. (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Ocydromus) corgenoma Maddison, 2020 Paratype
Male adult
USA: Oregon: Benton Co., Corvallis, Willamette River, 44.5491°N, 123.2449°W, 60 m
Copyright © David Maddison

Bembidion (Ocydromus) transversale Dejean, 1831
Male adult
Canada: Alberta: Burbank, junction of Red Deer and Blindman Rivers, 52.3542°N, 113.7556°W (voucher V101454)
Copyright © David Maddison

Bembidion (Ocydromus) erosum (Motschulsky, 1850)
Male adult
USA: California: Del Norte Co., Wilson Creek, 3 m, 41.6051°N, 124.1005°W (V101453)
Copyright © David Maddison

Bembidion (Ocydromus) perspicuum (Leconte, 1848)
Male adult
USA: Arizona: Cochise Co., San Pedro R at Charleston, 31.6239°N, 110.1722°W (voucher V101461)
Copyright © David Maddison

Bembidium haplogonum Chaudoir, 1868 (NEOTYPE)
Male adult
USA: California: Lake Co., North Branch Cache Creek at hwy 20, 305 m 38.9881°N, 122.5400°W
Copyright © David Maddison

Bembidion (Ocydromus) sarpedon Casey, 1918
Male adult
USA: Colorado: Las Animas Co., Cokedale, Reilly Canyon, 37.1346°N, 104.6114°W (V101459)
Copyright © David Maddison

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Sakhalin Isl., valley of the Rogatka River, 46°58'03"N 142°46'24"E, 18.vii.2011, leg. K.Makarov L=5,7
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Sakhalin Isl., valley of the Rogatka River, 46°58'03"N 142°46'24"E, 18.vii.2011, leg. K.Makarov L=5,8
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Kinashir Isl., 1.5 km south-west mouth of the Ozernaya River, 43°52'27"N 145°27'16"E, 26.vii.2011, leg. K.Makarov L=5,2
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., 1.5 km south-west mouth of the Ozernaya River, 43°52'27"N 145°27'16"E, 26.vii.2011, leg. K.Makarov L=5.0
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Kinashir Isl., Zmeinyy (Stolbovskoy) Stream, 44°00'31"N 145°40'49"E, 26.viii.2017, leg. K.Makarov L=5,75 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., Zmeinyy (Stolbovskoy) Stream, 44°00'31"N 145°40'49"E, 26.viii.2017, leg. K.Makarov L=5,6 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Kinashir Isl., mouth of the Medny Stream, 44°16'34"N 145°56'27"E, 16.viii.2017, leg. K.Makarov L=5,85 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., mouth of the Medny Stream, 44°16'34"N 145°56'27"E, 16.viii.2017, leg. K.Makarov L=5,45
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Kinashir Isl., Dokuchaeva Ridge, solfatara field “Bolshoye”, h~400, 44°26'57"N 146°5'28"E, 8.viii.2017, leg. K.Makarov L=5.0 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., Dokuchaeva Ridge, solfatara field “Bolshoye”, h~400, 44°26'57"N 146°5'28"E, 8.viii.2017, leg. K.Makarov L=4,7 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Kinashir Isl., caldera of Golovnin Volcano, shore of Kipyashcheye Lake, 43°51'47"N 145°30'04"E, 24.vi.2008, leg. K.Makarov L=4,55 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., caldera of Golovnin Volcano, shore of Kipyashcheye Lake, 43°51'47"N 145°30'04"E, 24.vi.2008, leg. K.Makarov L=4,75
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Kinashir Isl., caldera of Golovnin Volcano, solfatara field "Cherepakhovoye", 43°52'51"N 145°30'01"E, 28.vii.2011, leg. K.Makarov L=4,8 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., caldera of Golovnin Volcano, solfatara field "Cherepakhovoye", 43°52'51"N 145°30'01"E, 28.vii.2011, leg. K.Makarov L=4,8 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., Mendeleev Volcano, source of Kislyi Stream below the solfatara field, 43°59'46"N 145°45'54"E, 22.ix.2016, leg. Yu.Sundukov L=4,9 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Kinashir Isl., Mendeleev Volcano, source of Kislyi Stream below the solfatara field, 43°59'46"N 145°45'54"E, 22.ix.2016, leg. Yu.Sundukov L=4,8
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Shikotan Isl., the Tserkovnaya Bay, 43°44'59"N 146°41'40"E, 29.vi-10.vii.2012, leg. Yu.Sundukov L=5,3 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Shikotan Isl., the Tserkovnaya Bay, 43°44'59"N 146°41'40"E, 29.vi-10.vii.2012, leg. Yu.Sundukov L=5,6 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Female adult
Yuriy Isl., Shirokaya Bay, 43°25'33"N 146°04'08"E, 2.ix.2016, leg. Yu.Sundukov L=4,8 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) dolorosum (Motschulsky, 1850)
Male adult
Yuriy Isl., Shirokaya Bay, 43°25'33"N 146°04'08"E, 2.ix.2016, leg. Yu.Sundukov L=5 mm
Copyright © Kirill Makarov

Bembidion (Ocydromus) fontinale (Raffray, 1885)
Male adult
Ethiopia Amhara, Mt. Abuna yosef, n-slope, 3800-3950 m, 12°07’52”N 39°11’39”E, 4.iii.2019, leg. D. Hauth, J. Schmidt, Yeshitla M., Yitbarek, W. (CSCHM)
Copyright © Joachim Schmidt

Bembidion (Ocydromus) basistriatum Fairmaire, 1893
Male adult
Ethiopia, Oromia, Mt Enkuolo, NE-slope, 3100-3200m, 07°24’41”N 39°22’26”E, 5-7.Xii.16, leg. J. Schmidt (CSChM)
Copyright © Joachim Schmidt

Bembidion (Ocydromus) melanoceroides Toledano, Bonavita & Schmidt, 2021 Paratype
Male adult
Ethiopia, Oromia, Mt Enkuolo, N-slope, 3350-3600m, 07°22’48”N 39°21’07”E, 6-9.Xii.16, leg. J. Schmidt (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Ocydromus) melanocerum Chaudoir, 1876
Male adult
Ethiopia, Amhara, W-slope Mt. Choke, 2750-2900 m, 10°38’N 37°45’E, 1.iii.2019, leg. D. Hauth, J. Schmidt, Yeshitla M., Yitbarek, W. (CSCHM)
Copyright © Joachim Schmidt

Bembidion (Ocydromus) terryerwini Neri & Toledano, 2021 Paratype
Male adult
N 29°21'14.7" E 57°20'31.7" / Iran, Prov. Kerman, Rāyen, Goroah, / Babtahoune, Bachufer-Schotterbank / 06.06.2014 2700 m / leg. Schnitter
Copyright © Luca Toledano

Bembidion (Ocydromus) hiekei Muller-Motzfeld, 1986b: 171 Paratype
Male adult

Copyright © Luca Toledano

Bembidion (Ocydromus) sokolowskii Fassati, 1957b: 154 Holotype
Male adult
MHNG
Copyright © Luca Toledano

Bembidion (Ocydromus) decorum decorum (Panzer, 1799b)
Male adult
CZECH REPUBLIC - Staré Hamry, 22.IX.2020, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar

Bembidion (Ocydromus) modestum (Fabricius, 1801)
Male adult
CZECH REPUBLIC - Olomouc dist., Skalička, 11.VI.2020, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar

Bembidion (Ocydromus) loebli Schmidt, 2014 Holotype
Male adult
Holotype male, with label data „INDIA Madras / Nilgiri 2100 m / Ootacamund - Pykara / 21.XI.72 / Besuchet Löbl Mussard“ in MHNG
Copyright © Qin Yuyao

Bembidion (Ocydromus) salweenum Schmidt, 2014 Holotype
Male adult
Holotype male, with label data “STIBET 26-29.IX.2010 / Kema SSE Naqu 4450 m / Salween River, banks, alpine / steppe, 31°16’34N 92°06’20E” in ZSM
Copyright © Qin Yuyao

Bembidion (Ocydromus) saxatile himalotibeticum Schmidt, 2014 Holotype
Male adult
Holotype male, with label data “S TIBET 15.VII.2010 / YamTso lake shore near / Nagartse, near Samding / Monastery, 4420 m” in ZSM.
Copyright © Qin Yuyao

Bembidion (Ocydromus) primorjense Neri & Toledano, 2023 Holotype
Male adult
Holotype, ♂, “Russia N Primorskyi reg. / 70 km NW Plastun 18-26 VII / Podnebesnyi pass 600-800 m / A. Plutenko leg. 1996” [printed]” (CTVR)
Copyright © Qin Yuyao

Bembidion (Ocyturanes) parsorum Netolitzky, 1934
Female adult
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin L=4.7 mm
Copyright © Kirill Makarov

Bembidion (Ocyturanes) parsorum Netolitzky, 1934
Female adult

Copyright © Kirill Makarov

Bembidion (Ocyturanes) parsorum Netolitzky, 1934
Male adult
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin L=4.9 mm
Copyright © Kirill Makarov

Bembidion (Ocyturanes) praeustum Dejean, 1831
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidion (Ocyturanes) christophi Schmidt, 2018 Paratype
Male adult
“S TIBET 15.VII.2010 / Tibetan Himalaya / above Kampa La 48-4850 m” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Ocyturanes) gracilipenne Schmidt, 2018 Paratype
Male adult
“S TIBET, N Lhasa Valley / above Chupsang Monast. / 19.IX.2010, 4900m / 29°44’01N 91°07’47E ” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Ocyturanes) lhatseense Schmidt, 2018 Paratype
Male adult
“S TIBET 17.VI.2011 / above pass Tsuo La / 47-4940 m, leg. Schmidt / 29°03’37N 87°58’52E” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Ocyturanes) eucheres Netolitzky, 1943
Male adult
Tajikistan, Khoja Obi, Garm, 15.9.2011, Barsevskis A. leg. (DUBC) det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Ocyturanes) samai Neri & Toledano, 2017 Paratype
Male adult
Afghanistan, Vt. N Salang 1450 m, Khenjan, bord torrent, 11.8.1975, leg. Ledoux (DUBC)
Copyright © Alexander Anichtchenko

Bembidion (Ocyturanes) muilwijki Neri & Toledano, 2017 Paratype
Male adult
S Iran, Sisakht Dena, 2500-3000 m, 13-14.6.1973
Copyright © Alexander Anichtchenko

Bembidion (Ocyturanes) dyscheres Netolitzky, 1943
Female adult
N Iran, C Elburz, Val. Darband Sar, 2500-3000m 18.7.1970 (DUBC). det. L. Toledano
Copyright © Alexander Anichtchenko

Bembidion (Ocyturanes) montislunae Neri & Toledano, 2022 Holotype
Male adult
Oman, Dhofer province / Jabal al Qamar / 20 km N Dhalqut / N 16.71092° E 53.15350° / (1100 m.n.m.) 28.8.2012 / lgt. P. Kučera (CTVR). Image from Paolo Neri & Luca Toledano, 2022
Copyright © - - -

Bembidion (Ocyturanes) ivanloebli Neri & Toledano, 2018 Holotype
Male adult
“Iran, Fars, Sepidan, 21-9-2005, Muilwijk leg.” (CTVR)
Copyright © Luca Toledano

Bembidion (Ocyturanes) waziristanum Andrewes, 1932c: 863
Male adult
Pakistan, W. Himalai mts., 20 km SW Chilez (CTVR)
Copyright © Luca Toledano

Bembidion (Ocyturanes) lobanovi Mikhailov, 1984b: 297
Female adult
N Tajikistan, Fanskie Mts. Khurdak (CTVR). Scale: 1 mm
Copyright © Luca Toledano

Bembidion (Ocyturanes) signatipenne Jacquelin du Val, 1852
Male adult
Imago: NW Turkey, Edremit, Kaz Dağ, Ay i Deresi (PN); Aedeagus specimen from Crete, Rethimno, Plakias, river Megalopotamos (PN).
Copyright © Luca Toledano

Bembidion (Ocyturanes) viduum Netolitzky, 1910a: 215 Holotype
Male adult
Kilikisch. Taurus (NHMW). Scale: 1 mm.
Copyright © Luca Toledano

Bembidion (Ocyturanes) kurdistanicum Netolitzky, 1920
Male adult
Imago: Iran, Mazandaran Prov., Karaj-Chalus road (MNB); Aedeagus: Holotype, Persia Septentr., Kerim (NHMW).
Copyright © Luca Toledano

Bembidion (Ocyturanes) heinzi Korge, 1971: 16 Paratype
Male adult
Anatolia bor., Köse Flußufer (MNB). Aedeagus of Holotype, Anatolia bor., Köse Flußufer (MNB).
Copyright © Luca Toledano

Bembidion (Ocyturanes) antennarium (Morvan, 1972) Holotype
Male adult
Iran, Zagros, Kurang (PM). Scale 1mm
Copyright © Luca Toledano

Bembidion (Ocyturanes) mirzayani (Morvan, 1973) Paratype
Female adult
Iran, Zagros Mts., Sisakht (PM). Scale: 1 mm.
Copyright © Luca Toledano

Bembidion (Ocyturanes) dudichi Csiki, 1928
Male adult
Morocco, Great Atlas mts, 2200-2300m Oukaimeden env., 16.6.2010, Anichtchenko A. leg. (cAA). det. M. Toribio
Copyright © Alexander Anichtchenko

Bembidion (Omoperyphus) kempi Andrewes, 1922b: 178
Male adult
India, Uttarakhand, 13-15.4.2012, Uttarkashi distr., left tributary of Bhagirathi riv., 2150m, N30*57'43" E78*41'54.75", Anichtchenko A. leg. det. J. Schmidt 2015
Copyright © Alexander Anichtchenko

Bembidion (Omoperyphus) hypocrita Dejean, 1831
Male adult
Spain, Jaen prov., Arroyo de la piedra del Aguila, 37.683885, -3.485062, 3-4.5.2014, Anichtchenko A. leg & det. (cAA)
Copyright © Alexander Anichtchenko

Bembidion (Omotaphus) alsium Coquerel, 1866
Male adult
St. Helena Island. Image courtesy of David Maddison
Copyright © CC-BY 4.0

Bembidion (Omotaphus) gorilla Bonavita, Toledano & Vigna Taglianti, 2016
Male adult

Copyright © David Maddison

Bembidion (Omotaphus) scotti Netolitzky, 1931
Male adult

Copyright © David Maddison

Bembidion (Omotaphus) mellissii Wollaston, 1869
Male adult

Copyright © David Maddison

Bembidion (Omotaphus) mixtum Schaum, 1863b: 90 Lectotype
Male adult
43101 [handwritten] // mixtum Sch. [handwritten] / dt. Netolitzky [printed] // Hist. -Coll. (Coleoptera) / Nr. 43101 (=55133) / Bembidion mixtum Klug / Aegypt. Ehrenberg / Zool. Mus. Berlin [blue, printed] // SYNTYPE / Bembidium mixtum / Schaum, 1863 / labelled by MFNB 2015 [red, printed] (MNB).
Copyright © Luca Toledano

Bembidion (Omotaphus) aethiopicum Raffray, 1885
Male adult

Copyright © Luca Toledano

Bembidion (Omotaphus) mixtum Schaum, 1863b: 90
Male adult

Copyright © Paolo Bonavita

Bembidion (Omotaphus) kivuanum (Basilewsky, 1951) Holotype
Male adult
Imago - Holotype: “HOLOTYPUS [red, printed] // édéage en preparation microscop. [hand�written] // I.R.S.A.C. – MUS. Congo, Uvira: Bassin Ulindi / 2850m. 29-IX-1950 / N. Leleup [printed] / (Bord d’un marais) [handwritten] // RMCA ENT / 000019230 “ (MRAC); Aedeagus - paratype.
Copyright © Paolo Bonavita

Bembidion (Omotaphus) kilimanum Alluaud, 1908 Holotype
Male adult
“AFRIQUE ORle ALLEMANDE / KILIMANDJARO / (ZONE DES PRAIRIES) / CH. ALLUAUD I-IV 1904 [white, printed] // TYPE [red, printed] // MUSEUM PARIS / Coll. Ch. ALLUAUD [blue, printed] // Bembid. / (Testediolum / kilimanum / n.sp. Alluaud / [white, handwritten]” (MNHN).
Copyright © Paolo Bonavita

Bembidion (Omotaphus) demeyeri Toledano & Bonavita, 2016 Holotype
Male adult
“Uganda Occidental / Province D’UNYORO / Rég. de l’Albert-Nyanza / CH. ALLUAUD II 1909 [white, printed] // MUSEUM PARIS / Coll. Ch. ALLUAUD [blue, printed] // Bembid. (Notaphus) / Bakeri / type. Alluaud / Alluaud détérm [white, handwritten, with a blue strip on top]” (MNHN)
Copyright © Paolo Bonavita

Bembidion (Omotaphus) gorilla Bonavita, Toledano & Vigna Taglianti, 2016 Holotype
Male adult
? Holotype. “RWERU: 2800/2900M, 20 - VI – 1964, Tamisage, Bambousaise // Coll. Mus. Tervuren, Zaire: P. N. Virunga, Volcan Mikeno, R. P. Celis VI-64” (MRAC)
Copyright © Paolo Bonavita

Bembidion (Omotaphus) mellissii Wollaston, 1869
Male adult
Image from Bonavita et al. 2016, with modifications
Copyright © Paolo Bonavita

Bembidion (Omotaphus) madagascariense Chaudoir, 1876
Male adult

Copyright © Paolo Bonavita

Bembidion (Omotaphus) picturatum Fairmaire, 1898
Male adult

Copyright © Paolo Bonavita

Bembidion (Omotaphus) scotti Netolitzky, 1931
Male adult

Copyright © Paolo Bonavita

Bembidion (Omotaphus) uhligi Toledano & Maddison, 2016 Holotype
Male adult
Holotype? RSA-Western Cape, NE of Botrivier env., 34°13’S, 19°12’E, 24.II.2014 R. Kmeco leg.
Copyright © Paolo Bonavita

Bembidion (Omotaphus) tropicum Chaudoir, 1876
Male adult

Copyright © Paolo Bonavita

Bembidion (Pamirium) zlotini Mikhailov, 1996: 71
Female adult
det. I.Kabak et B.Kataev SE Kazakhstan, Zailiysky Alatau mts. Kok-Zhaylau. N 43°08'15" / E 77°00'08". h=2270 m. 24-26.06.2013. S.Kolov leg. Image of S. Kolov, from www.zin.ru
Copyright © - - -

Bembidion (Pamirium) platypterum Solsky, 1874: 120
Female adult
Kazakhstan, Alma-Ata, 2.06.1990. det. Anichtchenko A. 2016
Copyright © Alexander Anichtchenko

Bembidion (Paraprincidium) karinae Schmidt, 2009 Holotype
Female adult
NEPAL, Prov. Koshi, Tumlingtar, Arun-Ufer [Arun river bank], 300 mNN, 20.X.2002 leg. J. Weipert (Naturkundemuseum Erfurt). Image from Schmidt, 2009
Copyright © - - -

Bembidion (Paraprincidium) ruficolle (Panzer, 1796)
Female adult
Latvia, Jekabpils, Dunava 1.7.2006 (DUBC)
Copyright © Alexander Anichtchenko

Bembidion (Peryphanes) lacunarium (Zimmermann, 1869)
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Peryphanes) geberti Marggi, 2011 Holotype
Male adult
Türkei, Provinz Nevşehir, NE Avanos Kızılırmak, temporärer Flusslauf, 24.5.2001, leg. J. Gebert [38°46.070 N, 34°53.755 E], coll.
Naturhistorisches Museum Bern (NMBE).
Foto: Ortwin Bleich
Copyright © ---

Bembidion (Peryphanes) adygorum Belousov & Sokolov, 1996
Male adult
NW Caucas, Krasnodar reg., Tuapse distr., Podkhrebetovoe vill. vicinity, 19.V.2012 leg. M. Smirnov
Фотография с сайта zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Bembidion (Peryphanes) olegleonidovici Fassati, 1990: 169
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Peryphanes) milleri Jacquelin du Val, 1852
Male adult
4,5 mm; Germany, Rheinland-Pfalz, Eifel, Gerolstein, leg. J.Scheuern, 30.IV.1983; det. F.Koehler, 1994; Photo: U.Schmidt, 2014
Copyright © Udo Schmidt

Bembidion (Peryphanes) dalmatinum dalmatinum Dejean, 1831
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Peryphanes) adygorum Belousov & Sokolov, 1996: 14
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Peryphanes) phryganobium Belousov et Sokolov, 1996:24
Female adult
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin L=6.5 mm
Copyright © Kirill Makarov

Bembidion (Peryphanes) dalmatinum haupti Reitter, 1908a: 110
Male adult
Iran, Kurdistan prov., near Marivan city 18.V.2017 leg. D.Kasatkin L=5.7 mm
Copyright © Kirill Makarov

Bembidion (Peryphanes) dalmatinum haupti Reitter, 1908a: 110
Female adult
Iran, Lorestan prov., Dorud distr., near Cham Chit vill. 13.V.2017 leg. D.Kasatkin L=5.6 mm
Copyright © Kirill Makarov

Bembidion (Peryphanes) lirykense Reitter, 1908
Male adult
Iran, 40km S Ardabil, Goli, 20.5.2005, Anichtchenko A. leg. (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Peryphanes) fraxator Menetries, 1832
Male adult
Russia, Karachaevo-Cherkessia, urochishe Dzhalpak, 17-19.5.2009, A. Anichtchenko leg. (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Peryphanes) sanatum Bates, 1883: 274 Holotype
Male adult
“Type [round, bordered in red] // Japan / G. Lewis / 1910 – 310 [printed] // B. / sanatum / Bates [handwritten]” (NHMUK).
Copyright © Luca Toledano

Bembidion (Peryphanes) sanatum sanatum
Male adult
Japan, Honshu Island, Mount Hikageyama Jyoushu, Gunma Pref., 18.IX.1993 leg. S. Morita
Copyright © Kirill Makarov

Bembidion (Peryphanes) sanatum sanatum
Female adult
Japan, Honshu Island, Mount Hikageyama Jyoushu, Gunma Pref., 18.IX.1993 leg. S. Morita
Copyright © Kirill Makarov

Bembidion (Peryphanes) sanatum iwanai Makarov & Sundukov, 2021 Holotype
Male adult
Kunashir Island, valley of Kislaya River, below hot springs, 18.VI.2011, leg. A. Matalin
Copyright © Kirill Makarov

Bembidion (Peryphanes) sanatum iwanai Makarov & Sundukov, 2021 Paratype
Female adult
Kunashir Island, valley of Kislaya River, below hot springs, 18.VI.2011, leg. A. Matalin
Copyright © Kirill Makarov

Bembidion (Peryphanes) milleri carpathicum J. Muller, 1918b: 88
Male adult
W Ukraine, W Podolian Upland, Rogatyn Opillia, Zaluzhzhya vill, 49°24'33"N 24°35'36"E, clay quarry, 30-31.VIII.2018, Panin R leg., L=4.6mm
Copyright © Denis Khrapov

Bembidion (Peryphanes) cf brunnicorne Dejean, 1831
Male adult
W.Ukraine, Transcarpathian, Polonynian Beskids, Yavirnyk Rng, h=600 m, near Ruskyi Mochar vill.,15.V.2021, V. Khrapova leg./R. Panin det., L=4.7 mm
Copyright © Denis Khrapov

Bembidion (Peryphanes) deletum deletum Audinet-Serville, 1821: 77
Male adult
CZECH REPUBLIC - Beskydy Mts., Kněhyně, 27.VI.2020, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar

Bembidion (Peryphanes) stephensii Crotch, 1866
Female adult
CZECH REPUBLIC - Moravian-Silesian reg., Opava dist., Závada, 25.VI.2020, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar

Bembidion (Testediolum) armeniacum kokandicum Solsky, 1874: 120
Female adult

Copyright © Kirill Makarov

Bembidion (Testediolum) gagates gagates Andrewes, 1924a: 72
Male adult
India, Uttarakhand, 13-15.4.2012, Uttarkashi distr., left tributary of Bhagirathi riv., 2300m, N30*57'41.57" E78*41'54.75", Anichtchenko A. leg. det. J. Schmidt 2015
Copyright © Alexander Anichtchenko

Bembidion (Peryphidium) validum salangense Neri & Toledano, 2021 Paratype
Male adult
“Afghanistan / Lac du Salang 3700 m / 4.8.74 Leg. Ledoux”
Copyright © Luca Toledano

Bembidion (Peryphiolus) caligatum Jeanne & Muller-Motzfeld, 1982
Male adult
Spain, Guadarrama, Cotos, 25.8.2001, Anichtchenko A. leg & det.
Copyright © Alexander Anichtchenko

Bembidion (Peryphiolus) monticola monticola Sturm, 1825: 135
Male adult
CZECH REPUBLIC - Frýdek-Místek dist., Staré Hamry, 22.IX.2020, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar

Bembidion (Peryphodes) ephippigerum (Leconte, 1852)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Bembidion (Peryphophila) problematicum Schmidt, 2018 Holotype
Male adult
“NEPAL, Ghatte / Kirola – Khola. Camp / 3500m. Pasang 12.5.88” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Peryphophila) pseudoclarum Schmidt, 2018 Holotype
Male adult
“NEPAL Manaslu Mts. / Marsyangdi Khola vall. / Bhachok Khola 1000m / 13.4.1999 leg. Schmidt” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Peryphus) tetracolum Say, 1823: 89
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Peryphus) petrosum Gebler, 1833: 275
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Peryphus) koikei Habu & Baba, 1960 Holotype
Male adult
Kurokawa, NIIGATA, 27. VII. 1957, K. Baba<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-029<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Peryphus) koikei Habu & Baba, 1960 Holotype
Male adult
Kurokawa, NIIGATA, 27. VII. 1957, K. Baba<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-029<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Peryphus) koikei Habu & Baba, 1960 Holotype
Male adult
Kurokawa, NIIGATA, 27. VII. 1957, K. Baba<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-029<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (INCERTAE) ehikoense Habu, 1984 Holotype
Male adult
Mt. Hiko, FUKUOKA, 03. VIII. 1948, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-035
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (INCERTAE) ehikoense Habu, 1984 Holotype
Male adult
Mt. Hiko, FUKUOKA, 03. VIII. 1948, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-035
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (INCERTAE) ehikoense Habu, 1984 Holotype
Male adult
Mt. Hiko, FUKUOKA, 03. VIII. 1948, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-035
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (INCERTAE) ehikoense Habu, 1984 (Label) Holotype
Male adult
Mt. Hiko, FUKUOKA, 03. VIII. 1948, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-035
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Peryphus) captivorum Netolitzky, 1943
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Peryphus) subcostatum subcostatum (Motschulsky, 1850)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Peryphus) bruxellense Wesmael, 1835
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Peryphus) obscurellum obscurellum (Motschulsky, 1845)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Peryphus) captivorum Netolitzky, 1943a: 29
Male adult
Россия, SE Приморский кр., Лазовский зап., ~12 км S пос. Старая Каменка, окр. кордона Америка, 43°15'N / 134°01'E, 10.07.04, leg. М. & Л. Смирновы. image from www.zin.ru
Copyright © Maksim Smirnov

Bembidion (Peryphus) tetracolum tetracolum Say, 1823: 89
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Peryphus) bracculatum Bates, 1889b: 212
Female adult
India, Uttarakhand, 5km NW Ghangaria, Valley of Flowers, N30*43'356" E079*85.309 , 3417 m, 06.2011, Anichtchenko A.; 5.5 mm, det. J. Schmidt, 2015
Copyright © Alexander Anichtchenko

Bembidion () eutherum Andrewes, 1923b: 96
Male adult
India, Uttarakhand, 15 km SW New Tehri, 869m, 18-20.4.2012, Anichtchenko A. leg.; det. J. Schmidt 2015
Copyright © Alexander Anichtchenko

Bembidion (Peryphus) jedlickai jedlickai Fassati, 1945: 22
Female adult
Siberia, E Sajan Mts., Sentsa riv, trib. of Oka riv., 1700m, 9-10.7.1998, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Bembidion (Peryphus) bualei bualei Jacquelin du Val, 1852
Male adult
Italy, Lombardia, Brescla, Sabbio Chiese, 20.5.2011, E. Ruzzier leg.
Copyright © Alexander Anichtchenko

Bembidion (Peryphus) hummleri hummleri J. Muller, 1918
Male adult
Morocco, Quarzazate prov., Quwd Drag N 30*30'26" W006*06'01", near Tin-Zoulin vill., 27.6.2010, A. Anichtchenko leg. det. M. Toribio, 2017
Copyright © Alexander Anichtchenko

Bembidion (Peryphus) irregulare Netolitzky, 1934b: 90
Male adult
Shyock valley betw. Karghalik & Saser Brangsa, 4150-4500 m, 24.VI-11.VIII.1930
Copyright © Jan Muilwijk

Bembidion (Peryphus) sejunctum Casey, 1918
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidion (Peryphus) ahrensi Schmidt, 2018 Paratype
Male adult
“Uttar Ganga Ebene / bei Dhorpathan / 3000 m, 6.-8.3.”, “C-NEPAL-Himalaya / Dhaulagiri-Mts. / leg. Ahrens 1994” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Peryphus) aparupa Schmidt, 2018 Paratype
Male adult
“W-NEPAL Baglung Lekh / W Pokhara, 10-15km NW / Baglung, 2350-2550m / leg. Schmidt 10-12.5.04 / 28°18‘36”N 83°31‘56“E” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Peryphus) dhaulagiricum Schmidt, 2018 Holotype
Male adult
“NEPAL W Dhaulagiri / below Dhule 26-2700m / 7.IX.12 leg. J. Schmidt / 28°41‘07N 82°56‘31E” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Peryphus) glabripenne Schmidt, 2018 Paratype
Female adult
“Nepal, District Karnali, / Nähe Pohada 02.06.1997 / 3010-3400 mNN, leg. Grill, / Flußufer / HF, 29°04’N, 82°42’E” (CSCHM)
Copyright © Joachim Schmidt

Bembidion (Peryphus) cordicolle Jacquelin da Val, 1852
Male adult
Turkey, Kemer distr., Mts.Taurus, Ulupinar riv.,36*27'01N 30*26'14E 21.9.2014, I. Enuschenko leg. (DUBC) det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Peryphus) persicum Menetries, 1832
Male adult
Tajikistan, Khoja Obi, Garm, 15.9.2011, Barsevskis A. leg. (DUBC) det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Peryphus) abbreviatum pulpani Fassati, 1955
Male adult
Tadzhikistan, Ayni distr., Sughd prov., Iskanderkul lake, 2300m 5.7.2012, A. Barsevskis leg. (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Asioperyphus) notatum Andrewes, 1922b: 175
Female adult
India, Uttarakhand, 8 km NW Jaspur, lake shore, 22-23.4.2012, Anichtchenko A. leg. (DUBC). det. L. Toledano
Copyright © Alexander Anichtchenko

Bembidion (Peryphus) baleense Toledano, Bonavita & Schmidt, 2021 Holotype
Male adult
Ethiopia, Oromia, Bale Mts., Sebsebe Washia Forest, Salgen Vall. 3130 m, 07°02’08”n 39°36’06”e, 4.ii.2019, leg. R.Emmerich, J. Schmidt, Yeshitla M. (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Peryphus) damota Toledano, Bonavita & Schmidt, 2021 Holotype
Male adult
Ethiopia, Sodo, Mt. Damot, 2380 m, 06°53’26”N 37°46’51”E, 3.Xii.2017, leg. D. Hauth, J. Schmidt, Yeshitla M.” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Peryphus) scottustulatum Netolitzky, 1937
Male adult
Ethiopia, Amhara, n-slope Mt. Choke, 3800-3950 m, 10°43’16”N 37°51’15”E, 26.ii.2019, leg. D. Hauth, J. Schmidt, Yeshitla M., Yitbarek, W. (CSChM)
Copyright © Joachim Schmidt

Bembidion (Peryphus) psuchrum Andrewes, 1922c: 241
Male adult
Inde, Himachal Pradesh, Mahari, 3500 m (CTVR)
Copyright © Luca Toledano

Bembidion (Peryphus) tricuspis Neri & Toledano, 2018 Paratype
Male adult
“SW Iran, Hoseiniyeh / 28 km NNW Andimeshk / 12-13. 4. 1977, 360 m // Loc. no. 286 / Exped Nat Mus / Praha” (NMPC).
Copyright © Luca Toledano

Bembidion (Peryphus) subcostatum vau Netolitzky, 1913: 213
Male adult
CZECH REPUBLIC - Olomouc reg., Přerov dist., Skalička, 8.IV.2020, V. Rýznar lgt., det. et coll.
Copyright © Vojtěch Rýznar

Bembidion (Peryphus) andreae (Fabricius, 1787)
Female adult
Italy, Tagliamento, Ronchis, iv.2024, leg. Jörg Gebert
Copyright © Jörg Gebert

Bembidion (Peryphus) bualei polonicum J. Muller, 1930
Male adult
Ahrenshoop, Steilufer s. Ort 12.4047E, 54.3726N, 5.2025, leg. & det. J. Gebert
Copyright © Jörg Gebert

Bembidion (Peryphus) subflavescens Antoine, 1944
Male adult
Morocco, Souss-Massa, Draa, Zagora, Oued Draa riv., 1.2.2013 (cAA). det. M. Toribio 2017
Copyright © Alexander Anichtchenko

Bembidion (Peryphus) maritimum (Stephens, 1839)
Female adult
Spain, Guipuzcoa, Playaundi, 20.3.2006, Anichtchenko A. leg & det. (cAA)
Copyright © Alexander Anichtchenko

Bembidion (Peryphus) femoratum femoratum Sturm, 1825
Female adult
Latvia, Jekabpils distr, Dunava, 23.7.2008 (DUBC)
Copyright © Alexander Anichtchenko

Bembidion (Peryphus) petrosum petrosum
Male adult
E Siberia, Lake Baikal, Baikalo-Lenskii NP., Bereg Buryh Medvedei, Pokoiniki, 27.6.1995, Anichtchenko A. leg & det. (cAA)
Copyright © Alexander Anichtchenko

Bembidion (Philochthus) pallidiveste Carret, 1905
Male adult
Volgogrda Area, Pallasovka Distr., near Elton Vill. 4.V.2009 leg. K.Makarov
L = 3.8
Copyright © Kirill Makarov

Bembidion (Philochthus) guttula (Fabricius, 1792)
Male adult
length: 2,8-3,8 mm
Germania, Bayern, Oberfranken, Rudolfstein
leg.det. U.Schmidt, 2001
Foto: U.Schmidt, 2010
Copyright © U.Schmidt

Bembidion (Philochthus) biguttatum (Fabricius, 1779)
Male adult

Copyright © Kirill Makarov

Bembidion (Philochthus) guttula (Fabricius, 1792)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Philochthus) inoptatum Schaum, 1857a: 149
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Philochthus) mannerheimii C.R. Sahlberg, 1827
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Philochthus) cf. neresheimeri J. Muller, 1929
Female adult
Ивановская обл., г. Иваново, парк, 1905 г., 29.05.2014, М. Смирнов leg.; det. K.Makarov. Image from www.zin.ru\Animalia\Coleoptera
Copyright © Maksim Smirnov

Bembidion (Philochthus) lunulatum (Geoffroy, 1785)
Male adult
Turkey, Marmaris, Hisarönü, leg. U.Schmidt, 18.V.1997; det. Marggi, 2014; Photo: U.Schmidt, 2014. 3,8 mm
Copyright © Udo Schmidt

Bembidion (Philochthus) biguttatum (Fabricius, 1779)
Male adult
Russia, Ivanovo, "Kharinka" city park, 30.V.2015, M. Smirnov leg.
Copyright © Maksim Smirnov

Bembidion (Philochthus) ellipticocurtum Netolitzky, 1935c: 167
Female adult
Lebanon, Jbail prov., Laqlouq, 8.5.2000. det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Philochthus) neresheimeri J. Muller, 1929: 101
Male adult
W Ukraine, Lviv city, Sykhiv, waste-ground, 49°47'29"N 24°3'18"E, 7.IV.2018, Panin R. leg., L=3.1mm
Copyright © Denis Khrapov

Bembidion (Philochthus) vicinum Lucas, 1846
Male adult
Morocco 65 km N Essaouria, 5 km N Jamaa Shoim 31°57'02"N 009°25'25"E 22-24.III.2011 leg. A.Gusakov. L= 3.7 mm
Copyright © Kirill Makarov

Bembidion (Philochthus) vicinum Lucas, 1846: 86
Female adult
Morocco 65 km N Essaouria, 5 km N Jamaa Shoim 31°57'02"N 009°25'25"E 22-24.III.2011 leg. A.Gusakov. L= 4.0 mm
Copyright © Kirill Makarov

Bembidion (Phyla) obtusum Audinet-Serville, 1821: 83
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Phyla) phoeniceum Huber & Marggi, 1997: 778
Female adult
Turkey, Kirish distr., Camuyuva riv. 28.9.2014, I. Enischenko leg. (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Phyla) incommodum Netolitzky, 1926
Male adult
Iran, 40 km S Ardabil, Goli, 20.5.2005, Anichtchenko A. leg. (DUBC). det. L. Toledano
Copyright © Alexander Anichtchenko

Bembidion (Plataphus) simplex Hayward, 1897
Male adult
Great Smoky Mountains National Park. photo I. M. Sokolov
Copyright © Igor Sokolov

Bembidion (Plataphus) gebleri (Gebler, 1833)
Male adult
Far East, Lazovskij Nat. Res., Formozova spring 43°00'06"N 134°05'13"E 10.VII.2005 K.Makarov leg.
L=4.5 mm
Copyright © Kirill Makarov

Bembidion (Plataphus) prasinum (Duftschmid, 1812)
Female adult

Copyright © Kirill Makarov

Bembidion (Plataphodes) tetraporum kurokawense Habu & Baba, 1958 Holotype
Male adult
Kurokawa, NIIGATA, 27. VII. 1957, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-025
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Plataphodes) tetraporum kurokawense Habu & Baba, 1958 Holotype
Male adult
Kurokawa, NIIGATA, 27. VII. 1957, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-025
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Plataphodes) tetraporum kurokawense Habu & Baba, 1958 Holotype
Male adult
Kurokawa, NIIGATA, 27. VII. 1957, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-025
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Plataphodes) tetraporum kurokawense Habu & Baba, 1958 (Label) Holotype
Male adult
Kurokawa, NIIGATA, 27. VII. 1957, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-025
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Plataphodes) fujiyamai Habu, 1958 Holotype
Male adult
Mt. Shozu-dake, Hakuba, NAGANO, 31. VIII. 1949, I. Fujiyama
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-026
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Plataphodes) fujiyamai Habu, 1958 Holotype
Male adult
Mt. Shozu-dake, Hakuba, NAGANO, 31. VIII. 1949, I. Fujiyama
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-026
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Plataphodes) fujiyamai Habu, 1958 Holotype
Male adult
Mt. Shozu-dake, Hakuba, NAGANO, 31. VIII. 1949, I. Fujiyama
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-026
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Plataphodes) fujiyamai Habu, 1958 (Label) Holotype
Male adult
Mt. Shozu-dake, Hakuba, NAGANO, 31. VIII. 1949, I. Fujiyama
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-026
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Alexander Anichtchenko

Bembidion (Plataphodes) daisetsuzanum Habu, 1958 Holotype
Male adult
Mt. Daisetsu, HOKKAIIDO, 15. VII. 1952, T. Shirozu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-027
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Plataphodes) daisetsuzanum Habu, 1958 Holotype
Male adult
Mt. Daisetsu, HOKKAIIDO, 15. VII. 1952, T. Shirozu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-027
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Plataphodes) daisetsuzanum Habu, 1958 Holotype
Male adult
Mt. Daisetsu, HOKKAIIDO, 15. VII. 1952, T. Shirozu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-027
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Plataphodes) daisetsuzanum Habu, 1958 Holotype
Male adult
Mt. Daisetsu, HOKKAIIDO, 15. VII. 1952, T. Shirozu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-027
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Plataphodes) tetraporum Bates, 1883: 270
Female adult
Kuriles isl., Kunashir
Copyright © Kirill Makarov

Bembidion (Plataphus) gebleri persuasum Netolitzky, 1938a: 38
Male adult
SE Приморский кр., Лазовский зап., побережье Японского моря, бухта Тачингоуза, 27.06.04 leg. М.& Л.Смирновы
Russia, SE Primorie, Lazo Nat. res., seashore, Tachingouz bay
Image of M. Smirnov (М.Э. Смирнов) from www.zin.ru\Animalia\Coleoptera
Copyright © Maksim Smirnov

Bembidion (Plataphus) infuscatipenne Netolitzky, 1938
Male adult
Russia, SE Primorie, Lazo Nat. res., ~10 км SW Sokolchi vill., Korpad' vicinity 43°15'N / 134°7'E, 23-27.06.10, leg. M. Smirnov
Фотография с сайта zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Bembidion (Plataphus) lucillum lucillun Bates, 1883
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion () pseudolucillum Netolitzky, 1938a: 37 EA: JA
Male adult
Kunashir Island, creek mouth near Moristyi Island, 17.VIII. 2013, K. Makarov
Copyright © Kirill Makarov

Bembidion () pseudolucillum Netolitzky, 1938a: 37 EA: JA
Female adult
Kunashir Island, creek mouth near Moristyi Island, 17.VIII. 2013, K. Makarov
Copyright © Kirill Makarov

Bembidion () pseudolucillum Netolitzky, 1938a: 37 EA: JA
Female adult
Kunashir Island, creek mouth near Moristyi Island, 17.VIII. 2013, K. Makarov
Copyright © Kirill Makarov

Bembidion () pseudolucillum Netolitzky, 1938a: 37 EA: JA
Female adult
Kunashir Island, creek mouth near Moristyi Island, 17.VIII. 2013, K. Makarov
Copyright © Kirill Makarov

Bembidion (Plataphus) coelestinum (Motschulsky, 1844)
Female adult
N Mongolia, Tuul-Gol R., Engin Aimak Tev, 14.8.2013, I. Enuschenko leg. (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Plataphus) difficile (Motschulsky, 1844)
Male adult
Russia, Tuva rep.,Todzhynsk distr., E Sayan mts., Arzhkhan-Khem, 2200 m, 21.6.2017, I. Enuschenko leg. (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Plataphus) pseudolucillum Netolitzky, 1938a: 37
Male adult
Kibune, Kyoto (NHMW)
Copyright © Luca Toledano

Bembidion (INCERTAE) amurense Motschulsky, 1859
Male adult
photo Makarov K.
Copyright © Kirill Makarov

Bembidion (INCERTAE) umeyai Habu, 1959 Holotype
Male adult
Jozankei, HOKKAIDO, 18. IX. 1954, K. Umeya
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-031
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (INCERTAE) umeyai Habu, 1959 Holotype
Male adult
Jozankei, HOKKAIDO, 18. IX. 1954, K. Umeya
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-031
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (INCERTAE) umeyai Habu, 1959 Holotype
Male adult
Jozankei, HOKKAIDO, 18. IX. 1954, K. Umeya
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-031
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (INCERTAE) umeyai Habu, 1959 (Label) Holotype
Male adult
Jozankei, HOKKAIDO, 18. IX. 1954, K. Umeya
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-031
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Asioperyphus) gotoense Habu, 1973 Holotype
Male adult
Goto, Tomie, NAGASAKI, 02. V. 1970, M. Ezima
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-037
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Asioperyphus) gotoense Habu, 1973 Holotype
Male adult
Goto, Tomie, NAGASAKI, 02. V. 1970, M. Ezima
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-037
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Asioperyphus) gotoense Habu, 1973 Holotype
Male adult
Goto, Tomie, NAGASAKI, 02. V. 1970, M. Ezima
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-037
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Asioperyphus) gotoense Habu, 1973 (Label) Holotype
Male adult
Goto, Tomie, NAGASAKI, 02. V. 1970, M. Ezima
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-037
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Asioperyphus) xanthochiton Andrewes, 1922b: 176
Female adult
India, Uttarakhand, 14 km NW New Tehri, 950m, N30*28'25.3" E 078*20'56.2", 10.4.2012, A. Anichtchenko leg. det. J. Schmidt 2015
Copyright © Alexander Anichtchenko

Bembidion (Ocydromus) phaedrum Andrewes, 1923b: 97
Female adult
India, Uttarakhand, 15 km SW New Tehri, 869 m, N30*15'874" E 078*21'587", 18-20.4.2012, A. Anichtchenko leg.; 5 mm; det. J. Schmidt 2015
Copyright © Alexander Anichtchenko

Bembidion (Politophanes) inventor Schmidt, 2018 Holotype
Male adult
“E. NEPAL: KOSI / Val. Induwa Kola / 2000 m, 18.IV.84 / Löbl - Smetana” (MHNG).
Copyright © Joachim Schmidt

Bembidion (Princidium) compactum Andrewes, 1922c: 240
Female adult
India, Uttarakhand, 10 km NNE of Ramnagar, gorge of left trib. of Kosi River, 400-450 m,
29°29' N, 79°09,5' E, R. Dudko, I. Lyubechanskii, 19-20.04.2012
det. R. Dudko 2012
Copyright © Roman Dudko

Bembidion (Princidium) punctulatum punctulatum s.str.
Male adult
Germany, Bavaria, Ascholding, Isar, leg. det. U.Schmidt, 3.V.1975 Photo: U.Schmidt, 2012
Copyright © Udo Schmidt

Bembidion (Princidium) punctulatum Drapiez, 1820
Male adult
Pechoro-Ilychskii N.P. (Печеро-Илычский зап)
Copyright © Kirill Makarov

Bembidion (Pseudophilochthus) nubigena (Wollaston, 1877)
Male adult
St. Helena Island. Image courtesy of David Maddison
Copyright © CC-BY 4.0

Bembidion (Pseudophilochthus) evanescens (Wollaston, 1877)
Female adult

Copyright © David Maddison

Bembidion (Pseudophilochthus) gemmulipenne (Wollaston, 1877)
Male adult

Copyright © David Maddison

Bembidion (Pseudophilochthus) fossor (Wollaston, 1877)
Male adult

Copyright © David Maddison

Bembidion (Pseudophilochthus) trechoides (Wollaston, 1877)
Male adult

Copyright © David Maddison

Bembidion (Pseudophilochthus) sublimbatum (Wollaston, 1877)
Male adult

Copyright © David Maddison

Bembidion (Pseudophilochthus) dicksoniae (Wollaston, 1877)
Male adult

Copyright © David Maddison

Bembidion (Pseudophilochthus) grayanus (Wollaston, 1877)
Male adult

Copyright © David Maddison

Sakagutia umi Sasakawa, 2007
Male adult
Far East, Lazo Distr., 4 km VE Preobrazhenie Vill., Ezhovaja Bay 42°55'59N 26.VII.2005 133°59'29E K.Makarov leg.<br /> L=5.5 mm
Copyright © Kirill Makarov

Sakagutia umi Sasakawa, 2007
Male adult
SE Приморье, Лазовский р-н., побережье Японского моря, 3 км NE пос. Преображение бухта Ежовая (Тасовая), литораль, 12.07.2010, leg. М. Смирнов. Image from www.zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Bembidion (Sakagutia) umi Sasakawa, 2007
Male adult
S. Korea, Gyeongsanbuk-do, Pohang-si, collected by digging the pebbles near the estuary
Copyright © Dooyoung Kim

Bembidion (Semicampa) schueppelii Dejean, 1831
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Semicampa) chaudoirii Chaudoir, 1850b: 179
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Semicampa) gilvipes Sturm, 1825
Female adult
Latvia, Daugavpils loki, Daugava 29.4.2008 (DUBC)
Copyright © Alexander Anichtchenko

Bembidion (Sloanephila) tahitiense Liebherr & Maddison, 2013 Holotype
Male adult
(MNHN): French Polynesia: Tahiti Nui / Mt. Mauru lava tube / el. 705 m 18-IX-2006 lot 01 / 17˚38.017´S 149˚21.284´W / pyr. fog moss/liverworts / along stream J.K. Liebherr // HOLOTYPE / Bembidion / tahitiense / Liebherr & / Maddison 2012 (black-bordered red label).
Copyright © David Maddison

Bembidion (Sloanephila) jacksoniense Guerin Meneville, 1830
Female adult
Wiluna W.A. / ix 25 1931 //Australia / Harvard Exp. / Darlington (MCZ). Scale bars 0.5 mm.
Copyright © David Maddison

Bembidion (Taiwanobembidion) aliense Habu, 1973
Male adult

Copyright © David Maddison

Bembidion (Talanes) subfasciatum Chaudoir, 1850b: 187
Female adult
Krasnodar Reg., Taman Peninsula, Tuzlo Lake 6.V.2009
2.7 mm
Copyright © Kirill Makarov

Bembidion (Talanes) aspericolle (Germar, 1829)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Terminophanes) pulcherrimum (Motschulsky, 1850)
Male adult
Caucasus, Elbrus, Itkol (CTVR). Image from Toledano L. & Gueorguiev B., 2015
Copyright © - - -

Bembidion (Terminophanes) kaschmirense Netolitzky, 1920
Female adult
India, Uttarakhand, 5 km NE Govind Ghat, road to Gangaria, 9-10.6.2011, Anichtchenko A. leg. det. J. Schmidt 2015 8 mm
Copyright © Alexander Anichtchenko

Bembidion (Terminophanes) mckinleyi scandicum Lindroth, 1943
Female adult
E Siberia, Lake Baikal, Malyi Kochergat vill., Goloustnaya riv., 16.VII.1991, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Bembidion (Terminophanes) reibnitzi Schmidt, 2018 Paratype
Male adult
“S TIBET Kyi Chu river / betw. Lhasa and airport / ca. 3650 m, 22.IX.2010 / leg. J. Schmidt” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Terminophanes) terminale kirgisorum Netolitzky, 1934
Male adult
Tajikistan, Ayni distr., Sughd prov., Sarytag, 9km S lake Iskanderkul, 2380m, 6.7.2012, A. Barsevskis leg. (DUBC) det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Terminophanes) consummatum Bates, 1873
Male adult
Russia, E Siberia, Irkutsk reg., Bolshaya Rechka vill, 3.8.1990, Anichtchenko A. leg. & det
Copyright © Alexander Anichtchenko

Bembidion (Testediolum) julianum De Monte, 1943
Male adult
Italy
Copyright © Augusto Degiovanni

Bembidion (Testedium) bipunctatum (Linne, 1761)
Male adult
length: 3,6-4,8 mm
Austria, Schladminger Tauern, Mariapfarr
leg.det. U.Schmidt, 1984
Foto: U.Schmidt, 2010
Copyright © U.Schmidt

Bembidion (Testedium) laetum Brulle, 1836: pl.2
Female adult
Morocco, 65 km N Essaouria, 5 km N Jamaa Shoim, 23-25.III.2011, A. Gusakov. Фотография с сайта zin.ru\Animalia\Coleoptera.
Copyright © Maksim Smirnov

Bembidion (Testedium) laetum Brulle, 1836
Female adult
Morocco 65 km N Essaouria, 5 km N Jamaa Shoim 31°57'02"N 009°25'25"E 22-24.III.2011 leg. A.Gusakov. L= 4.0mm
Copyright © Kirill Makarov

Bembidion (Testedium) laetum Brulle, 1836
Male adult
Morocco 65 km N Essaouria, 5 km N Jamaa Shoim 31°57'02"N 009°25'25"E 22-24.III.2011 leg. A.Gusakov. L= 3.9 mm
Copyright © Kirill Makarov

Bembidion (Trepanedoris) concretum Casey, 1918
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Trepanedoris) atripes (Motschulsky, 1844)
Male adult

Copyright © Kirill Makarov

Bembidion (Trepanedoris) faraticolle (Jeanne, 1995) Holotype
Female adult
Image courtesy Biological Museum, Lund University
Copyright © - - -

Bembidion (Trepanedoris) doris (Panzer, 1797)
Male adult

Copyright © Kirill Makarov

Bembidion (Trepanes) articulatum (Panzer, 1796)
Female adult
length: 3,0-3,9 mm
Germania, Bayern, Oberfranken, Kulmbach
leg.det. U.Schmidt, 1992
Foto: U.Schmidt, 2010
Copyright © U.Schmidt

Bembidion (Trepanes) octomaculatum (Goeze, 1777)
Male adult

Copyright © Kirill Makarov

Bembidion (Trepanes) articulatum (Panzer, 1796)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Trepanes) octomaculatum (Goeze, 1777)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Trichoplataphus) inaense Habu, 1956 Holotype
Male adult
Tenryu River-side, Iijima, Kamiina, NAGANO, 06. X. 1945, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-028
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Trichoplataphus) inaense Habu, 1956 Holotype
Male adult
Tenryu River-side, Iijima, Kamiina, NAGANO, 06. X. 1945, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-028
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Trichoplataphus) inaense Habu, 1956 Holotype
Male adult
Tenryu River-side, Iijima, Kamiina, NAGANO, 06. X. 1945, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-028
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Trichoplataphus) inaense Habu, 1956 (Label) Holotype
Male adult
Tenryu River-side, Iijima, Kamiina, NAGANO, 06. X. 1945, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-028
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Bembidion (Trichoplataphus) ozarkense Maddison & Hildebrandt, 2011 Paratype
Male adult
USA: Missouri: Carter Co., Current River at Van Buren, voucher DRMV100311
Copyright © - - -

Bembidion (Trichoplataphus) rolandi Fall, 1922
Male adult
USA: Virginia: Rockbridge Co., Maury River at Glasgow, voucher DRMV100193
Copyright © ---

Bembidion (Trichoplataphus) parconaturaviva Toledano, 2010: 394
Male adult
SE Maritime Prov., Lazovsky dist., ~10 кm SW Sokoltchi vill., 23-27.VI.2010 leg.М. Smirnov. Фотография с сайта www.zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов. det. K. Makarov 2016
Copyright © Maksim Smirnov

Bembidion (Trichoplataphus) kara Andrewes, 1921d: 250
Female adult
India, Uttarakhand, 15 km SW New Tehri, 869 m, N30*15'874" E 078*21'587", 18-20.4.2012, A. Anichtchenko leg.; 6 mm; det. J. Schmidt, 2015
Copyright © Alexander Anichtchenko

Bembidion (Trichoplataphus) deplanatum A. Morawitz, 1862a: 262
Male adult
E Siberia, Irkutsk, Irkut riv., 3.VI.1997, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Bembidion (Trichoplataphus) oxyglymma Bates, 1883
Male adult
Japan, Honshu Pref., Gifu, Tohuyama Dam, Ibigawa river, 16.VII.1992, leg. B. Kuznetsov. det. L. Toledano
Copyright © Alexander Anichtchenko

Bembidion (Zeactedium) giachinoi Toledano, 2005
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
Copyright © - - -

Zecillenus karikari Larochelle & Larivière, 2015
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
Copyright © - - -

Bembidion (Zecillenus) alacre (Broun, 1921)
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
Copyright © - - -

Bembidion (Zecillenus) albescens (Bates 1878)
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
Copyright © - - -

Bembidion (Zecillenus) tepaki Larochelle & Larivière, 2015
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
Copyright © - - -

Bembidion (Zecillenus) tillyardi (Brookes, 1927)
Female adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
Copyright © - - -

Bembidion (Zecillenus) waimarama Larochelle & Larivière, 2015
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
Copyright © - - -

Bembidion (Zecillenus) puponga Larochelle & Larivière, 2015
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
Copyright © - - -

Bembidion (Zecillenus) chalmeri (Broun, 1886)
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
Copyright © - - -

Bembidion (Zemetallina) wanakense Lindroth, 1976
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78.
Copyright © - - -

Bembidion (Zemetallina) chalceipes Bates, 1878
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78.
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Bembidion (Zemetallina) bullerense Larochelle & Larivière, 2015
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78.
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Bembidion (Zemetallina) hokitikense Bates, 1878
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78.
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Bembidion (Zeperyphodes) nesophilum Broun, 1886
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
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Bembidion (Zeperyphodes) callipeplum Bates, 1878
Female adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
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Bembidion (Zeplataphus) tairuense Bates, 1878
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
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