Carabidae

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Afranillus taxon page
Afrodipnus taxon page
Angustanillus taxon page
Anillaspis explanata (Horn, 1888)
Female adult
USA: California: El Dorado Co., Alabaster Cave (CMNH). Scale bar 1 mm; Image from LaBonte & Maddison, 2023
Copyright © - - -
Anillaspis taxon page (1 photos)
Anillinus langdoni Sokolov & Carlton, 2004
Male adult
USA-TN, Sevier Co, GSMNP, secondary site, 490 m, ex leaf et log litter, 35°43.157' N, 83°23.309' W, 19.04.2011, J.-S. Park det. I. Sokolov, 2013
Copyright © Roman Dudko

Anillinus affabilis (Brues, 1902) Lectotype
Male adult
Image from Sokolov I.M., Reddell J.R. & Kavanaugh D.H., 2014 Scale = 0.5 mm.
Copyright © Igor Sokolov

Anillinus sinuatus (Jeannel, 1963) Holotype
Female adult
Image from Sokolov I.M., Reddell J.R. & Kavanaugh D.H., 2014 Scale = 0.5 mm. (Deposited in NMNH)
Copyright © Igor Sokolov

Anillinus depressus (Jeannel, 1963) Holotype
Female adult
Image from Sokolov I.M., Reddell J.R. & Kavanaugh D.H., 2014 Scale = 0.5 mm. (NMNH)
Copyright © Igor Sokolov

Anillinus forthoodensis Sokolov & Reddell, 2014 Holotype
Male adult
TX: Bell Co., Talking Crows Cave, Fort Hood, 4.V.2006, J. Fant, M. Reyes \ Texas Memorial Museum Invertebrate Zool Coll #45.781 \ Holotype Anillinus forthoodensis Sokolov & Reddell 2014 [red label] \ CAS Type No. 18872
Copyright © Igor Sokolov

Anillinus affabilis (Brues, 1902)
Male adult
TEXAS, Travis County, Tooth Cave
Copyright © Igor Sokolov

Anillinus acutipennis Sokolov & Reddell, 2014 Paratype
Female adult
TX: Bell Co., Hidden Pit Cave, Fort Hood, 27.X.2007, J. Reddell, M. Reyes \ Texas Memorial Museum Invertebrate Zool Coll #60.107
Copyright © Igor Sokolov

Anillinus comalensis Sokolov & Kavanaugh, 2014 Paratype
Female adult
TX: Comal Co., 7mi W New Braunfels, 27.I.1995, A. G. Grubbs
Copyright © Igor Sokolov

Anillinus wisemanensis Sokolov & Kavanaugh, 2014 Holotype
Male adult
TX: Hays Co., Wiseman Sink, 28.IV.1995, A. G. Grubbs \ Texas Memorial Museum Invertebrate Zool Coll #27.149 \ Holotype Anillinus wisemanensis Sokolov & Kavanaugh 2014 [red label] \ CAS Type No. 18873
Copyright © Igor Sokolov

Anillinus sinuatus (Jeannel, 1963)
Male adult
TEXAS, Bexar County
Copyright © Igor Sokolov

Anillinus felicianus Sokolov, 2021 Holotype
Male adult
(NMNH): \ USA-LA: West Feliciana Par., ~4mi SW Jackson at 30.794°N, 91.254°W, mixed pine-hardwood forest, soil washing/berlese Sokolov I.M. 19–25 Apr 2018
Copyright © Igor Sokolov

Anillinus relictus Sokolov, 2021 Holotype
Male adult
(CMNH): ALABAMA: Blount Co., Tidwell Hollow Nature Trail east of Oneonta. T. N. King April 1 1972 \ 4/1/72 o [handwritten] \ THOMAS C. BARR COLLECTION 2011 Acc. No. 38,014
Copyright © Igor Sokolov

Anillinus albrittonorum Sokolov & Schnepp, 2021 Paratype
Female adult
6 mi NW High Springs, Columbia County, Florida. Image from Sokolov & Schnepp, 2021
Copyright © Igor Sokolov

Anillinus bexarensis Sokolov, 2022 Paratype
Female adult
Up the Creek Cave, Bexar County, Texas ; 1.7 mm
Copyright © Igor Sokolov

Anillinus reddelli Sokolov, 2022 Paratype
Female adult
Beck Horse Cave, Williamson County, Texas
Copyright © Igor Sokolov

Anillinus hirsutus Sokolov, 2012 Paratype
Male adult
Paratype male, CMNH
Copyright © Curt Harden
Anillinus taxon page (16 photos)
Anillodes taxon page
Anillopsidius taxon page
Anillopsis taxon page
Anillotarsus taxon page
Anillus angelae Magrini & Vanni, 1989
Male adult
Arnale cave, Coreno Ausonio, Italia
Copyright © Vittorino Monzini
Anillus taxon page (3 photos)
Argiloborus taxon page
Bafutyphlus taxon page
Bhutanillus taxon page
Bulirschia taxon page
Bylibaraphanus taxon page
Caeconannus taxon page
Caecoparvus taxon page
Carayonites taxon page
Cryptocharidius mandibularis M.Etonti & Mateu, 1992
Male adult
Peru, Dept. Ancash, Cordillera Blanca, Punta Olimpica Pass, m 4400 a.s.l., G. Allegro legit
Copyright © Gianni Allegro
Cryptocharidius taxon page (1 photos)
Cryptorites taxon page
Dicropterus taxon page
Elgonotyphlus taxon page
Erwinanillus taxon page
Externalillus taxon page
Geocharidius phineus Erwin, 1982
Male adult
Guatemala
image from Sokolov I., 2013. A new genus and eight new species of the subtribe Anillina (Carabidae, Trechinae, Bembidiini) from Mexico, with a cladistic analysis and some notes on the evolution of the genus.
Copyright © Igor Sokolov
Geocharidius taxon page (1 photos)
Geocharis taxon page
Gilesdytes taxon page
Gracilanillus taxon page
Gregorydytes taxon page
Hesperanillus taxon page
Honduranillus taxon page
Hygranillus taxon page
Hypotyphlus degiovannii Magrini, 2013 Holotype
Male adult
Italia, Lazio, Montagna di Scauri (Latina), 50 m s.l.m., 1.V.2008, P. Magrini leg. (CM).
Copyright © Paolo Magrini
Hypotyphlus taxon page (1 photos)
Iason taxon page
Iberanillus taxon page
Illaphanus pteridophyticus Giachino, 2005 Holotype
Male adult

Copyright © Alexander Anichtchenko
Illaphanus taxon page (1 photos)
Kimberleytyphlus taxon page
Leleupanillus taxon page
Lusotyphlus taxon page
Magnanillus taxon page
Malagasydipnus taxon page
Malagasytyphlus taxon page
Medusapyga hehalis LaBonte, 2023 Paratype
Female adult
WASHINGTON, Thurston Co., 11.3 km NW Littlerock, elev. ~400 m, D.J. Cox and J.R. LaBonte, 6 May 2007, 46.9600°N 123.13450°W. Image from LaBonte & Maddison, 2023
Copyright © - - -

Medusapyga alsea LaBonte, 2023 Paratype
Female adult
“OREGON, Benton Co., Prairie Peak, 11 km S Alsea, J.R. LaBonte, 9 V 1993, elev. 810 m, 44.2844°N 123.5858°W, under stones". Image from LaBonte & Maddison, 2023
Copyright © - - -
Medusapyga taxon page (2 photos)
Megastylulus pivai Giachino & Sciaky, 2003 Holotype
Male adult
leg. E. Piva - Isl. St.Lucia Caraibi
Copyright © Vittorino Monzini
Megastylulus taxon page (1 photos)
Mexanillus taxon page
Typhlocharis carmenae Zaballos & Ruiz-Tapiador, 1995
Male adult
Spain, Toledo, Los Navalucillos 23.I.1997
Zaballos & Ruiz-Tapiador leg.
long 1.3mm
Copyright © Alexander Anichtchenko
Microcharidius taxon page
Microdipnites taxon page
Microdipnodes taxon page
Microdipnus taxon page
Microtyphlus saxatilis Magrini, Leo & Fancello, 2004
Male adult
Italia, Sardegna, Perda Longa (NU) Baunei.
Copyright © Augusto Degiovanni
Microtyphlus taxon page (1 photos)
Mystroceridius taxon page
Neodipnus taxon page
Neoillaphanus taxon page
Nesamblyops sp.
Male adult
New Zealand
Scale bars = 0.5mm
image from Sokolov I., 2013. A new genus and eight new species of the subtribe Anillina (Carabidae, Trechinae, Bembidiini) from Mexico, with a cladistic analysis and some notes on the evolution of the genus.
Copyright © Igor Sokolov

Nesamblyops sp.
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Nesamblyops townsendi Sokolov, 2023
Male adult
Tennyson Inlet, Marlborough Sounds, SO
Copyright © Igor Sokolov

Nesamblyops tararua Sokolov, 2023
Male adult
Judd Ridge, Tararua Range, Wellington, NO
Copyright © Igor Sokolov

Nesamblyops parvulus Sokolov, 2023 Paratype
Female adult
Mount Stokes, Marlborough Sounds, SO
Copyright © Igor Sokolov
Nesamblyops taxon page (5 photos)
Nothanillus taxon page
Orthotyphlus taxon page
Paranillopsis taxon page
Paranillus taxon page
Parillaphanus taxon page
Parvocaecus taxon page
Pelocharis taxon page
Pelodiaetodes (Pelodiaetodes) constricticollis Sokolov, 2015 Paratype
Male adult
NZ, Northland, Radar Bush. Image from Sokolov, 2015
Copyright © - - -
Pelodiaetodes taxon page (1 photos)
Pelodiaetus sulcatipennis Jeannel, 1937
Male adult
New Zealand, Otago, Outram
Copyright © Igor Sokolov

Pelodiaetus nunni Sokolov, 2019 Paratype
Male adult
NZ, Canterbury, Ahuriri Scenic Reserve. 1.27 mm
Copyright © Igor Sokolov
Pelodiaetus taxon page (2 photos)
Pelonomites taxon page
Perucharidius taxon page
Pilbaradytes taxon page
Pilbaranillus taxon page
Pilbaraphanus taxon page
Prioniomus taxon page
Pseudanillus taxon page
Pseudillaphanus taxon page
Rhegmatobius bastianinii Magrini & Casale, 2015 Paratype
Male adult
Italia, Lazio (LT) M.te S. Angelo, Formia, 1200m.
Copyright © Augusto Degiovanni
Rhegmatobius taxon page (2 photos)
Scotodipnus alpinus Baudi di Selve, 1871
Female adult
Piemonte, Valle Po, Crissolo (CN), 27.V.2000, 1250m, fagus, leg. F. Angelini
Copyright © Alexander Anichtchenko

Scotodipnus (Binaghites) affinis affinis (Baudi di Selva, 1871)
Male adult
Italy, Emilia, M. Penna (PR), 1400m, 24.5.1999, leg. F. Angelini
Copyright © Alexander Anichtchenko

Scotodipnus (Binaghites) armellinii (Ganglbauer, 1900)
Male adult
Veneto Italia
Copyright © Erminio Piva

Scotodipnus (Scotodipnus) glaber (Baudi di Selve, 1869)
Male adult
Monte Chiappo, PV, Italia
Copyright © Vittorino Monzini
Scotodipnus taxon page (3 photos)
Selenodipnus taxon page
Serranillus jeanneli Barr, 1996
Male adult
USA, Georgia, White County
Copyright © Igor Sokolov
Serranillus taxon page (1 photos)
Serratotyphlus taxon page
Stylulus (Stylulus) isabelae Giachino & Sciaky, 2003 Holotype
Male adult

Copyright © Vittorino Monzini
Stylulus taxon page (2 photos)
Tasmanillus taxon page
Typhlocharis taxon page (2 photos)
Typhlonesiotes taxon page
Winklerites stevanovici Hlavac & Margini, 2017 Paratype
Male adult
Srbija [p], Dimitrovgrad, Petrlaš, Džemanska prop[ast], 02.05.2002 [h], M. Stevanović leg. Image from HLAVÁČ & MAGRINI, 2017
Copyright © - - -
Winklerites taxon page (1 photos)
Zapotecanillus montanus Sokolov, 2013 Paratype
Male adult
MEXICO, Oaxaca, 52miles N of Oaxaca, Ber.202, sink litter, 17 May 1971 S.B.Peck collector (CMNH)
Scale bar = 0.5mm
Copyright © Igor Sokolov

Zapotecanillus oaxacanus Sokolov, 2013 Paratype
Male adult
MEXICO. Oaxaca 18.7mi S Valle Nacional 5200’ 17 Aug.1973\ A.Newton Collector
Scale bar = 0.5mm
Copyright © Igor Sokolov

Zapotecanillus nanus Sokolov, 2013 Paratype
Female adult
MEXICO, Oaxaca, 18.7mi N Valle National, 5200’ 17 Aug.1973\ A.Newton Collector
Copyright © Igor Sokolov

Zapotecanillus ixtlanus Sokolov, 2013 Paratype
Male adult
MEXICO, Oaxaca, 37mi S Valle National
Copyright © Igor Sokolov

Zapotecanillus iviei Sokolov, 2013 Paratype
Male adult
OAXACA, 2 mi S. Cerro Pelon, 8–9000 ft. 03 JUL 1982 M.A. Ivie colr.
Copyright © Igor Sokolov

Zapotecanillus kavanaughi Sokolov, 2013 Paratype
Male adult
MEXICO, Oaxaca, 14km N San Juan
Copyright © Igor Sokolov

Zapotecanillus pecki Sokolov, 2013 Paratype
Male adult
MEXICO, Oaxaca, 3.5mi S Suchixtepec
Copyright © Igor Sokolov

Zapotecanillus longinoi Sokolov, 2013 Paratype
Male adult
MEXICO, Chiapas, Sierra Morena
Copyright © Igor Sokolov
Zapotecanillus taxon page (8 photos)
Zeanillus taxon page
Zoianillus taxon page