Endofallus Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993 Endophallus
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Ruslan Panin
Carabus (Morphocarabus) illigeri illigeri Dejean, 1826 (as bjelasnicensis Apfelbeck, 1902) Endophallus
endophallus as bjelasnicensis Apfelbeck, 1902
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Ruslan Panin
Carabus (Archicarabus) monticola fontanae Born, 1906 Endophallus
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Ruslan Panin
Carabus (Morphocarabus) rothi hampei Kuster, 1846 Endophallus
W Ukraine, Vinogradov distr.
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Ruslan Panin
Carabus (Morphocarabus) rothi hampei Kuster, 1846 Endophallus
endophallus hampei
Vinogradov distr.
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Igor Solodovnikov
Carabus (Morphocarabus) kollari Palliardi, 1825 Endophallus
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Ruslan Panin
Carabus (Trachycarabus) estreicheri Fischer, 1822 Endophallus
Lviv Region, Brody distr., near Yasenov vill.,
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Ruslan Panin
Carabus (Trachycarabus) sibiricus tscherkessicus Korge, 1964 Endophallus
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Ruslan Panin
Carabus (Megodontus) imperialis Fischer, 1823 Endophallus
endophallus
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Safronov
Carabus (Megodontus) schoenherri sajanus Breuning, 1927 Endophallus
endophallus
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Safronov
Carabus (Tribax) titan titan Zolotarev, 1913 Endophallus
endophallus
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Ruslan Panin
Carabus (Tribax) steveni satanas Semenov, 1896 Endophallus
endophallus
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Ruslan Panin
Carabus (Tribax) circassicus kubaniensis Semenov, 1896 Endophallus
endophallus
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Ruslan Panin
Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886 Endophallus
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Ruslan Panin
Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886 Endophallus
aedeagus
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Ruslan Panin
Carabus (Trachycarabus) sibiricus obliteratus Fischer, 1827 Endophallus
endophallus sibiricus obliteratus Fischer, 1827
NE Kazakhstan,
S Altai mts.,S Ulbinsky mt.h=500m.,Turgusun vill., 20.VI.2001
male 25mm.
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Ruslan Panin
Carabus (Trachycarabus) sibiricus haeres (as fossulatus Dejean, 1826) Endophallus
endophallus
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Ruslan Panin
Carabus (Trachycarabus) sibiricus errans Fischer, 1823 Endophallus
endophallus sibiricus errans
SW-Russia,
Rostov reg. env. stanitsa Razdorskaya, A.Ponomarev leg
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Ruslan Panin
Carabus (Trachycarabus) sibiricus errans Fischer, 1823 Endophallus
endophallus sibiricus errans- Lugansk reg., near Slavyanoserbk
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Ruslan Panin
Carabus (Trachycarabus) scabriusculus scabriusculus Olivier, 1795 Endophallus
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Ruslan Panin
Carabus (Trachycarabus) scabriusculus inapertus Motschulsky, 1850 (loc.typ) Endophallus
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Ruslan Panin
Carabus (Trachycarabus) planus planus Gehin, 1885 Endophallus
<b>endophalus </b> S Russia,Krasnodar reg., Anapa distr., B. Utrish vill., env.,
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Ruslan Panin
Carabus (Trachycarabus) planus planus Gehin, 1885 Endophallus
<b>endophalus</b> Karacaj-Cherkessija, near Kart-Djurt vill., upper Kylan riv. h=1300m
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Ruslan Panin
Carabus (Trachycarabus) planus campicola Reitter, 1896 Endophallus
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Ruslan Panin
Carabus (Trachycarabus) mandibularis buchtarmensis Kryzhanovskij, 1953 Endophallus
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Ruslan Panin
Carabus (Trachycarabus) besseri Fischer, 1822 Endophallus
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Ruslan Panin
Carabus (Morphocarabus) wulffiusi wulffiusi Morawitz, 1862 Endophallus
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Ruslan Panin
Carabus (Morphocarabus) illigeri simulator Kraatz, 1876 Endophallus
NE-Serbia,
Krs Mts., Majdanpek env.,h=300m.20-31.V.2006, Pr.Sima leg
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Ruslan Panin
Carabus (Morphocarabus) spasskianus spasskianus Breuning, 1932 Endophallus
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Ruslan Panin
Carabus (Morphocarabus) spasskianus schestopalovi Plutenko, 1995 Endophallus
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Ruslan Panin
Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799 Endophallus
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Ruslan Panin
Carabus (Morphocarabus) excellens excellens Fabricius, 1798 Endophallus
Moldova,<br />
Anenii Noi distr., near Garbovets vill.
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Ruslan Panin
Carabus (Morphocarabus) helleri Ganglbauer, 1893 Endophallus
SE Moravia,
Strani, V.2002, M.Snizek leg.
male bronze colour.
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Ruslan Panin
Carabus (Morphocarabus) helleri Ganglbauer, 1893 Endophallus
SE Moravia,
Strani, V.2002, M.Snizek leg.
male blue colour
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Ruslan Panin
Carabus (Tribax) apschuanus pseudoplatessa Gottwald, 1982 Endophallus
endophallus
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Ruslan Panin
Carabus (Archicarabus) montivagus montivagus Palliardi, 1825 Endophallus
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Ruslan Panin
Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844: 100 Endophallus
endophallus
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Alexander Anichtchenko
Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844 (as melleus Lapouge, 1909) Endophallus
<p>
endophallus odoratus odoratus (as melleus Lapouge, 1909)<br />
<br />
<br />
</p>
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Ruslan Panin
Carabus (Morphocarabus) odoratus bargusinicus Shilenkov, 1996 Endophallus
endophallus
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Ruslan Panin
Carabus (Morphocarabus) monilis Fabricius, 1792 Endophallus
France 39, Foncina le Haut 300m
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Alexander Anichtchenko
Carabus (Morphocarabus) monilis Fabricius, 1792 (as interpositus Géhin, 1880) Endophallus
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Ruslan Panin
Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996 Endophallus
endophallusCopyright ©
Ruslan Panin
Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924 Endophallus
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Alexander Anichtchenko
Carabus (Morphocarabus) aeruginosus zyrjanovskianus Shilenkov & O.Berlov, 1996 Endophallus
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Ruslan Panin
Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822 Endophallus
endophallus
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Ruslan Panin
Carabus (Eucarabus) ulrichi ulrichi Germar, 1824 Endophallus
endophallus ulrichi (as podolicus Semenov, 1898)
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Ruslan Panin
Carabus (Platycarabus) irregularis montandoni Buysson, 1882 Endophallus
endophallus
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Ruslan Panin
Carabus (Microplectes) riedeli Menetries, 1832 Endophallus
endophallusCopyright ©
Ruslan Panin
Carabus (Microplectes) convallium convallium Starck, 1889 Endophallus
Endophallus
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Ruslan Panin
Carabus (Microplectes) convallium argonautarum Semenov, 1898 Endophallus
Endophallus
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Ruslan Panin
Carabus (Pachystus) tamsi tamsi Menetries, 1832 Endophallus
endophallus
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Ruslan Panin
Carabus (Pachystus) glabratus glabratus Paykull, 1790 Endophallus
Endophallus
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Ruslan Panin
Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876 Endophallus
endophalus
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Ruslan Panin
Carabus (Megodontus) violaceus fiorii Born, 1901 Endophallus
endophallus
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Ruslan Panin
Carabus (Megodontus) violaceus baeterrensis Lapouge, 1901 Endophallus
ehdophallus
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Ruslan Panin
Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823 Endophallus
endophallus
Lviv reg., Nikolaev distr., near Krasiv vill.
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Ruslan Panin
Carabus (Megodontus) exaratus Quensel, 1806 Endophallus
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Ruslan Panin
Carabus (Archicarabus) nemoralis nemoralis Muller, 1764 Endophallus
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Ruslan Panin
Carabus (Platycarabus) fabricii malachiticus Thomson, 1875 Endophallus
Endophallus fabricii malachiticus
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Ruslan Panin
Carabus (Neoplectes) daphnis daphnis Kurnakov, 1962 Endophallus
Georgia, Mingrelia, Lecheha vallis, Tehuri superior
Caucasus centr., 1200m, 27.6-7.7.1989 Retezar I. leg.
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Alexander Anichtchenko
Carabus (Neoplectes) daphnis askhicus Belousov, 1992 Endophallus
Georgia, env. Martvilli, Askhi plateau
10-15.7.2001 A.Rubenian leg.
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Alexander Anichtchenko
Carabus (Neoplectes) daphnis askhicus Belousov, 1992 Endophallus
W Georgia, valley of Chkhorodukhu riv., 1000-1200m
14.V-6.VII.1991
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Alexander Anichtchenko
Carabus (Neoplectes) daphnis askhicus Belousov, 1992 Paratype Endophallus
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Ruslan Panin
Carabus (Neoplectes) juenthneri avadcharensis Kurnakov, 1970 Endophallus
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Ruslan Panin
Carabus (Neoplectes) reitteri laevisternis Starck, 1890 Endophallus
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Ruslan Panin
Carabus (Neoplectes) prometheus prometheus Reitter, 1887 (as nubicola Zolotarev, 1913) Endophallus
Endophallus prometheus prometheus (as nubicola Zolotarev, 1913)
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Ruslan Panin
Carabus (Neoplectes) prometheus prometheus Reitter, 1887 Endophallus
Endophallus prometheus prometheus Reitter, 1887
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Ruslan Panin
Carabus (Neoplectes) prometheus prometheus Reitter, 1887 Endophallus
aedeagus
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Ruslan Panin
Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990 Endophallus
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Ruslan Panin
Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990 Endophallus
aedeagus miroshnikovi Psebai env., Nikitino
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Ruslan Panin
Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890 Endophallus
Endophallus
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Ruslan Panin
Carabus (Imaibius) nouristani nouristani (Ledoux, 1977) Endophallus
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Frank Kleinfeld
Carabus (Hygrocarabus) nodulosus Creutzer, 1799 Endophallus
Endophallus
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Ruslan Panin
Carabus (Hygrocarabus) variolosus variolosus Fabricius, 1787 Endophallus
Endophallus
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Ruslan Panin
Carabus (Eucarabus) manifestus pyonganicola Deuve & Li, 1998 Endophallus
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Ruslan Panin
Carabus (Morphocarabus) praecellens trans illigeri Dejean, 1826 Endophallus
W-Croatia, Karlovac, near Bosiljevo vill., h~300m,
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Ruslan Panin
Carabus (Morphocarabus) spasskianus aeruginosiformis Breuning, 1932 Endophallus
Russia, Khakassia,
W-Sayan, Alan Mt range, valley Enisey riv., 5 km S Mayna vill.,
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Ruslan Panin
Carabus (Trachycarabus) sibiricus gvozdevae Obydov, 1997 Endophallus
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Ruslan Panin
Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924 Endophallus
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Ruslan Panin
Carabus (Morphocarabus) spasskianus shoriensis Obydov, 1999 Endophallus
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Ruslan Panin
Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886 Endophallus
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Ruslan Panin
Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988 (as shchessiensis) Paratype Endophallus
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Ruslan Panin
Carabus (Carabus) granulatus granulatus Linne, 1758 Endophallus
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Ruslan Panin
Carabus (Chrysocarabus) auronitens escheri Palliardi, 1825 Endophallus
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Ruslan Panin
Carabus (Chrysocarabus) auronitens festivus Dejean, 1826 Endophallus
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Ruslan Panin
Carabus (Chrysocarabus) auronitens farinesi Dejean, 1826 Endophallus
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Ruslan Panin
Carabus (Chrysocarabus) hispanus Fabricius, 1787 Endophallus
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Ruslan Panin
Carabus (Eucarabus) obsoletus obsoletus Sturm, 1815 Endophallus
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Ruslan Panin
Carabus (Megodontus) violaceus aurolimbatus Dejean, 1829 Endophallus
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Ruslan Panin
Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997 Endophallus
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Ruslan Panin
Carabus (Morphocarabus) rothi hampei Kuster, 1846 (transition to rothi incompsus Kraatz, 1880) Endophallus
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Ruslan Panin
Carabus (Morphocarabus) helleri Ganglbauer, 1893 (as pseudopreyssleri Breuning, 1932) Endophallus
Czech Rep., NE Moravia, Moravian-Silesian Beskids Mt Rng, PR Trojacka, Hodslavice vill.,
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Ruslan Panin
Carabus (Morphocarabus) kollari Palliardi, 1825 (as semetricus Kraatz, 1878) Endophallus
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Ruslan Panin
Carabus (Morphocarabus) spasskianus geblerianus Obydov, 1999 Endophallus
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Ruslan Panin
Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859 Endophallus
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Ruslan Panin
Carabus (Morphocarabus) rothi hampei (as ormayi Reitter, 1896) loc.typ. Endophallus
near Beregovo, 27.IV-22.V..2011,, Panin R. & Chyzhevsky O. leg
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Ruslan Panin
Carabus (Morphocarabus) praecellens trans scheidleri Endophallus
NC Croatia, Ivanščica Mts , Ivanec,4-18.VI.2011, Krause Rene leg.
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Ruslan Panin
Carabus (Morphocarabus) rothi hampei Kuster, 1846 Endophallus
C Romania, Tвrgu Mureş,
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Ruslan Panin
Carabus (Morphocarabus) rothi hampei (as zilahiensis Csiki, 1906) Endophallus
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Ruslan Panin
Carabus (Tomocarabus) bessarabicus ? concretus Fischer, 1823 Endophallus
Steppe Reserve “Provalsky",
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Ruslan Panin
Carabus (Neoplectes) satyrus duripshensis Kurnakov, 1972 Endophallus
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Safronov
Carabus (Morphocarabus) regalis tschusovae Obydov, 2004 Endophallus
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Ruslan Panin
Carabus (Semnocarabus) carbonicolor carbonicolor Morawitz, 1886
Endophallus
Copyright © Alexandr Safronov
Carabus (Sphodristocarabus) varians (as dvorshaki Mandl, 1975)
Endophallus
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Carabus (Neoplectes) apollo tenebricosus Kurnakov, 1962
Endophallus
Copyright © Alexandr Safronov
Carabus (Morphocarabus) rothi rothi Dejean, 1829 Endophallus
C Romania, Sibiu Area, right bank Olt River, Bradu vill., h=380 m,
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Ruslan Panin
Carabus (Morphocarabus) venustus venustus Morawitz, 1862 Endophallus
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Ruslan Panin
Carabus (Pachystus) hungaricus cribellatus Adams, 1812 Endophallus
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Ruslan Panin
Carabus (Pachystus) hungaricus scythus Motschulsky, 1847 Endophallus
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Ruslan Panin
Carabus (Trachycarabus) sibiricus sibiricus Fischer, 1822 Endophallus
near Krasnoyarsk
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Ruslan Panin
Carabus (Mesocarabus) dufouri dufouri Dejean, 1829 Endophallus
Jaen reg., Monte Javalcuz h=1436 m
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Ruslan Panin
Carabus (Mesocarabus) lusitanicus brevis (as sorianus Jeanne, 1972) Endophallus
NC-Spain, Soria, Puerto de Oncala
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Ruslan Panin
Carabus (Carabus) arvensis arvensis Herbst, 1784 Endophallus
W Ukraine, NW Podolian, Lviv reg., Javoriv distr.,near Stradch vill.,
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Ruslan Panin
Carabus (Tribax) puschkini ishikawaianus Breuning & Ruspoli, 1970 Endophallus
NE-Turkey: Artvin:Hopa, Borзka env., h=450 m,
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Ruslan Panin
Carabus (Morphocarabus) praecellens praecellens Pillardi, 1825 Endophallus
Serbia, Fruska Gora Mts, 20-31.V.2006, M.Sima leg.
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Ruslan Panin
Carabus (Morphocarabus) illigeri versicolor Frivaldsky, 1835 Endophallus
C Bulgaria: Stara planina Mts, Gabrovo
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Ruslan Panin
Carabus (Tribax) puschkini ponticus Deyrolle, 1869 Endophallus
NE-Turkey:Trabzon, Zigana Pass h=1700m,
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Ruslan Panin
Carabus (Morphocarabus) rothi comptus (as szoerenyensis Csiki, 1908) Endophallus
SW Romania, Tarcu Mts, vf. Nedeia h=2000m,
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Ruslan Panin
Carabus (Megodontus) bonvouloiri bonvouloiri (as kaschkarensis Blumenthal & Breuning, 1967) Endophallus
NE-Turke,Rize vill., S Ayder h=1740 m,
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Ruslan Panin
Carabus (Morphocarabus) rothi hampei (as gutiiensis Takács & Lie, 1992) Endophallus
N Romania, E Carpathian, Maramureş Reg., Gutai Mts, cabana Mogosa, 4 km N Baia Sprie,
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Ruslan Panin
Carabus (Sphodristocarabus) macrogonus macrogonus Chaudoir, 1847 Endophallus
NE-Turkey:Trabzon env., Sumela,
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Ruslan Panin
Carabus (Sphodristocarabus) macrogonus pseudokolatensis Heinz & Korge, 1967 Endophallus
NE-Turkey::Kackar Dagi Mt, Ayder S of Ardesen h=1600m
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Ruslan Panin
Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931 Endophallus
NE-China:Liaoning prov.,Fengcheng City, Miaoyang,
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Ruslan Panin
Carabus (Morphocarabus) rothi incompsus Kraatz, 1880 Endophallus
C Romania, Brașov County, Cristian vill., h=380 m
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Ruslan Panin
Carabus (Morphocarabus) rothi alutensis Savulescu, 1972 Endophallus
C Romania, S Carpathians, Rimnicu Vilcea env., Calimaneşti, Cozia Mts.,
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Ruslan Panin
Carabus (Lamprostus) robustus ducalis Schaufuss, 1882 Endophallus
NE-Turkey:Trabzon, Zigana Pass h=2000m
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Ruslan Panin
Carabus (Diocarabus) fraterculus fraterculus Reitter, 1895 Endophallus
N-Korea, N Pyongan Prov.,Sakju, Mt Ponsan,
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Ruslan Panin
Carabus (Sphodristocarabus) varians armeniacus Mannerheim, 1830 Endophallus
NE Turkey, Artvin Prov., Şavşat, Зamlıca.
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Ruslan Panin
Carabus (Pachystus) hungaricus hungaricus Fabricius, 1792 Endophallus
Czech, S Moravia, Pouzdrany
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Ruslan Panin
Carabus (Lamprostus) torosus torosus Frivaldsky, 1835 Endophallus
NE-Bulgaria, Dobrich Region, Kavarna env.,
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Ruslan Panin
Carabus (Semnocarabus) erosus kuldshaensis Ballion, 1878 Endophallus
S-Kazakhstan:Ili River valley,:near Bakanas,
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Ruslan Panin
Carabus (Megodontus) stroganowi transfuga Semenov, 1898 Endophallus
SE-Azerbaijan: Talysh, Kalvaz,
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Ruslan Panin
Carabus (Deroplectes) dokhtouroffi dokhtouroffi Ganglbauer, 1886 Endophallus
NC Tadzhikistan, Pamir, N-slope Peter I.Mt Rng, h=2100m, Ganishou vill.
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Ruslan Panin
Carabus (Morphocarabus) viridilimbatus panini I.Solodovnikov 2017 Holotype Endophallus
Russia, SE Yakutia, Neryungri distr.,
Tokinski Stanovik Mt Rng., Beder riv., (right tributary rivers Algama), 26.VI.1996
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Ruslan Panin
Carabus (Ulocarabus) stschurovskii Solsky, 1874 Endophallus
Tadzhikistan, Pamir-Alay, S slope Gissar Mt.Range, Takob, h-1700m,
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Ruslan Panin
Carabus (Morphocarabus) rothi hampei (as diffinis Csiki, 1906) Endophallus
NW Romania, Apuseni Mts, Trascău Mt Rng, Petresti de Jos, 12 km E Turda,.
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Ruslan Panin
Carabus (Ulocarabus) theanus Reitter, 1895 Endophallus
Tadzhikistan, S slope Gissar Mt.Range, left bank of the river Sorbo, Haidar gorge,
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Ruslan Panin
Carabus (Mesocarabus) (? lusitanicus) gougeleti Reiche, 186 Endophallus
CW Spain, Province Huelva, Arroyomolinos de Leon.
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Ruslan Panin
Carabus (Platycarabus) fabricii ucrainicus Lazorko, 1951 Endophallus
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Ruslan Panin
Carabus (?Oreocarabus) titanus Breuning, 1933 Endophallus
C-China Sichuan prov., Nanping 1500m,
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Ruslan Panin
Carabus (Megodontus) kolbei kolbei Roeschke, 1887 Endophallus
N-Japan, C-Hokkaido, Hidaka distr., Niikappu env.,
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Ruslan Panin
Carabus (Leptocarabus) procerulus Chaudoir, 1862 Endophallus
Japan:S Honshu , Matsubara-ko,
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Ruslan Panin
Carabus (Trachycarabus) latreillei ?semicoriaceus Kraatz, 1881 Endophallus
NE-China: Liaodong Peninsula, Liaoning prov., Gaizhou distr., Mt. Buyun Shan.
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Ruslan Panin
Carabus (Carabus) arvensis hokkaidoensis Lapouge, 1924 Endophallus
N Japan:Hokkaido, Hamamasu env.,
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Ruslan Panin
Carabus (Ohomopterus) japonicus japonicus Motschulsky, 1857 Endophallus
S Japan, Kyushu, Kumamoto Pref., Izumi viil., h=600 m
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Ruslan Panin
Carabus (Eurycarabus) famini fezzanus Lapouge, 1915 Endophallus
Morocco, Middle Atlas Mt Rng, Azrou city h=1500m,
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Ruslan Panin
Carabus (Leptocarabus) procerulus Chaudoir, 1862 Endophallus
Japan, S Honshu, Mie Pref., Hakusan,
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Ruslan Panin
Carabus (Oreocarabus) taedatus agassi Le Conte, 1850 Endophallus
SC-USA, New Mexico, Santa Fe,
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Ruslan Panin
Carabus (Ohomopterus) yamato kitai (Imura, 2004) Endophallus
Japan, S Honshu, Mie Pref., Hakusan,
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Ruslan Panin
Carabus (Trachycarabus) latreillei latreillei Fischer, 1822 Endophallus
N Mongolia, Khövsgöl reg., Lake Khovsgol, h=1640m, 51*:12’N; 100*:45’E, steppificated sites,
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Ruslan Panin
Carabus (Ulocarabus) theanus theanus Reitter, 1895 Endophallus
Tadzhikistan, S slope Gissar Mt.Range, left bank of the river Sorbo, Haidar gorge,
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Ruslan Panin
Carabus (Eucarabus) manfredeggeri Müller 2012 Paratype Endophallus
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Ruslan Panin
Carabus (Imaibius) dardiellus perobscurus Deuve, 1980 Endophallus
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Ruslan Panin
Carabus (Morphocarabus) illigeri illigeri (as tarensis Toševski, 1990) Endophallus
WC-Serbia: National park Tara Mts.:NE Mitrovac h=1500m,
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Ruslan Panin
Carabus (Cupreocarabus) huangi huangi Deuve, 1992 Endophallus
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Ruslan Panin
Carabus (Megodontus) violaceus dryas Gistl, 1857 Endophallus
NE-Greece, Mt. Olympos, Litochoro env.
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Ruslan Panin
Carabus (Lamprostus) saulcyi saulcyi Piochard, 1875 Endophallus
C Lebanon, Jbeil Distr., Ehmej vill., h=1100 m
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Ruslan Panin
Carabus (Pachystus) hortensis hortensis Linne, 1758 Endophallus
W-Ukraine, Male Polissya, near Kamenka-Buzka vill.
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Ruslan Panin
Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847 Endophallus
WC Spain, Cáceres, Hervás, h=1000m
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Ruslan Panin
Carabus (?Oreocarabus) qinlingensis Imura, 1993 Endophallus
China, W-Shaanxi, Taibai Shan Mts, btw.Houzhenzi/Ba Xian Tai, nr.Nan Tian Men h=3200m, 33°55'17"N/107°46'53"E,
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Ruslan Panin
Carabus (Morphocarabus) rothi hampei (as hunyadensis Csiki, 1926) Endophallus
CW Romania, Apuseni Mts, Metalliferous Mt Rng, Hunedoara reg., near Săcărâmb vill.
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Ruslan Panin
Carabus (Oreocarabus) latro minshanensis Deuve, 1987 Endophallus
China, N-Sichuan, Jiuzhaigou env.,
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Ruslan Panin
Carabus (Meganebrius) quinlani smetanai Häckel, 2011 Paratype Endophallus
C-Nepal:Dhaulagiri Massif:Baglung Distr.,(1.5km W of Okhale vill., 28.32’152’’N/ 83.48’673’E, h=2600m)
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Ruslan Panin
Carabus (Rhigocarabus) ladygini degensis Deuve, 1989 Endophallus
C-China, NW- Sichuan prov., Garzê Tibetan AP, Pass Garzê – Sêrtar
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Ruslan Panin
Carabus (Megodontus) violaceus azurescens Dejean, 1826 Endophallus
W-Croatia, Velebit Mts., betw.Karlobag & Brusane vill.,
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Ruslan Panin
Carabus (Megodontus) violaceus germarii Sturm, 1815 Endophallus
N Italy, Trent, Ampola, Mt. Tremalzo h=1700m
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Ruslan Panin
Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896 Endophallus
W Ukraine,Transcarpathian Reg., Rakhiv distr., Marmarosh Mts., near Lug vill., h=400 m.
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Ruslan Panin
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906 Endophallus
Transcarpathian region, Uzhgorod distr., 28.IV-23.V.2011
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Ruslan Panin
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906 Endophallus
SE Slovakia, Slanske vrchy Mts, Herlianske sedlo, 8-19.VI.2009, Krause Rene leg.
det.Panin R.
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Ruslan Panin
Carabus (Morphocarabus) zawadzkii zawadzkii, Kraatz 1854 Endophallus
Gorgany Mts. h=700 m, 16.VI.2011,
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Ruslan Panin
Carabus (Morphocarabus) zawadzkii zawadzkii, Kraatz 1854 Endophallus
W.UKRAINE, Ukrainian Carpathians
Lviv Reg., Skole distr., Skolivski Beskidy Mts.
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Igor Solodovnikov
Carabus (Mesocarabus) cantabricus cantabricus Chevrolat, 1840 Endophallus
NW-Spain, Pontevedra prov., A Cañiza viil., Alto de Fontefría h=800m
det.Panin R.
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Ruslan Panin
Carabus (Mesocarabus) dufouri baguenai Breuning, 1926 Endophallus
SE Spain, Almería Province, Carboneras env.
det. R.Panin
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Ruslan Panin
Carabus (Mesocarabus) macrocephalus Dejean, 1826 Endophallus
NW-Spain,Cantabrian Mts, Picos de Europa Mt Rng, Puerto de Panderruedas h=1460 m,
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Ruslan Panin
Carabus (Carabus) arvensis deyrollei Gory, 1839 Endophallus
NW-Spain, Cantabria, Udías env., Monte Corona h=100m
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Ruslan Panin
Carabus (Mesocarabus) latus latus Dejean, 1826 Endophallus
C-Spain, Toledo, Mts de Toledo, Sierra del Castañar Rng, Puerto del Milagro env, h=920m
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Ruslan Panin
Carabus (Semnocarabus) carbonicolor korolkowi Semenov, 1896 Endophallus
NC-Kyrgyzstan, Kyrgyz Mt.Rng , 10 km SW of Bishkek, Chon-Aryk env., h=1100m,
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Ruslan Panin
Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 Endophallus
N Hungary, Pilis Mts., Piliszentlelek,20-27.VII.2011, Krause Rene leg.
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Ruslan Panin
Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as pseudojucundus Retezár, 1974) Endophallus
NW-Hungary::Komarom distr., Tardos, Gerecse Mts.
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Ruslan Panin
Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as vertesensis Retezár, 1974) Endophallus
NW Hungary: Fejer: Vertes Mt Rng, Tatabanya
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Ruslan Panin
Carabus (Mesocarabus) lusitanicus helluo (as albarracinus Ganglbauer, 1886) Endophallus
CE-Spain, Provincia Teruel, Sierra de Javalón Rng, El Cañigral env., h=1460m,
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Ruslan Panin
Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826 Endophallus
CW-Spain:Salamanca, Brincones env., h=750m
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Ruslan Panin
Carabus (Mesocarabus) problematicus harcyniae (as belgicus Lapouge, 1913) Endophallus
Luxembourg, Bollendorf-Pont env., 3.III.1999
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Ruslan Panin
Carabus (Mesocarabus) (? problematicus trans lusitanicus) planiusculus (as xaxarsi Born, 1917) Endophallus
NE Spain, Catalonia,Girona Prov., San Hilario Sacalm env.<br />
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Ruslan Panin
Carabus (Aulonocarabus) kurilensis ? daisetsuzanus Kono, 1936 Endophallus
N-Japan, Hokkaido, Katō District, Kamishihoro, Horoka near Maruyama Bridge,
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Ruslan Panin
Carabus (Tribax) agnatus pseudagnatus Novotný & Voříšek, 1988 Endophallus
W-Caucasus: Abkhasia, Gudauta distr., Mt Anchkho h=2200-2350 m,
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Ruslan Panin
Carabus (Neoplectes) edithae exedithae Gottwald, 1985 Endophallus
NС-Caucasus, Karachaevo-Cherkessia, right bank Pkhiya river, Abishir-Akhuba Mt.Range h=2250-2600 m,
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Ruslan Panin
Carabus (Sphodristocarabus) separatus persa Lapouge, 1924 Endophallus
N-Iran: Mazandaran Province, Elborz Mt.Rng, Vandarbon env.
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Ruslan Panin
Carabus (Morphocarabus) rothi comptus (as borlovensis Lie, 1994) Endophallus
SW Romania,, Banat, Borlova env.,
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Ruslan Panin
Carabus (Limnocarabus) clathratus auraniensis Muller, 1902 Endophallus
W Ukraine, Transcarpathian Reg., 4 km NE Chop, forest Peresh.
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Ruslan Panin
Carabus (Morphocarabus) brinevi Kabak, 2014 Paratype Endophallus
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Ilya Kabak
Carabus (Ophiocarabus) nilkiensis Kabak, 2014 Paratype Endophallus
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Ilya Kabak
Carabus (Mimocarabus) pumilio Kuster, 1846 Endophallus
S-Armenia, Vayots Dzor Region, E Ughedzor vill., h=2630 m
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Ruslan Panin
Carabus (Pachycarabus) staehlini Adams, 1817 Endophallus
S Russia, E Caucasus, Dagestan, Nukatl Mt Rng h=2600m,
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Ruslan Panin
Carabus (Imaibius) boysi boysi Tatum, 1851 Endophallus
NW-India, Himachal Pradesh, Manikaran env., Pulga vill.
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Ruslan Panin
Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014 Endophallus
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Igor Solodovnikov
Carabus (Neoplectes) pseudopshuensis Zamotajlov, 1991 Endophallus
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Igor Solodovnikov
Carabus (Neoplectes) besleticus besleticus Kurnakov, 1972 Endophallus
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Igor Solodovnikov
Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972 Endophallus
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Igor Solodovnikov
Carabus (Neoplectes) besleticus dsychvensis Gottwald, 1985 Endophallus
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Igor Solodovnikov
Carabus (Neoplectes) besleticus duripshensis Kurnakov, 1972 Endophallus
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Igor Solodovnikov
Carabus (Neoplectes) besleticus napraensis Belousov & Zamotajlov, 1993 Paratype Endophallus
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Igor Solodovnikov
Carabus (Neoplectes) besleticus anastasiae Solodovnikov, 2014 Paratype Endophallus
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Igor Solodovnikov
Carabus (Lamprostus) torosus sinopensis Breuning & Ruspoli, 1970 Endophallus
NC-Turkey, Sinop province, Erfelek env., h=200m
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Ruslan Panin
Carabus (Archicarabus) phoenix Lapouge, 1924 Endophallus
C-Lebanon, Byblos /Jbeil Distr., Kfardebian env., h=1200m,
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Ruslan Panin
Carabus (Procrustes) piochardi praestigiator Morawitz, 1886 Endophallus
C-Lebanon, Byblos /Jbeil Distr., Kfardebian env., h=1200m,
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Ruslan Panin
Carabus (Sphodristocarabus) blumenthaliensis aequicostalis Staven & Heinz, 1995 Endophallus
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Ruslan Panin
Carabus (Mesocarabus) riffensis Fairmaire, 1875 Endophallus
Morocco, Rif, Foresta di Ketama, 25.06.1998, 1600m, leg. D. Sechi (immature)
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Alexander Anichtchenko
Carabus (Ophiocarabus) wusunshanicus Kabak, 2015 Endophallus
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Ilya Kabak
Carabus (Ophiocarabus) regeli regeli Morawitz, 1886 Endophallus
Karatau Mt. R.
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Ilya Kabak
Carabus (Lamprostus) hemprichi elonensis Schweiger, 1970 Endophallus
N-Israel, Midlle Galilea, Nazareth env., Hoshaya viil.
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Ruslan Panin
Carabus (Lamprostus) syrus cheikensis Deuve, 1992 Endophallus
N-Israel, Midlle Galilea, Nazareth env., Hoshaya viil.,
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Ruslan Panin
Carabus (Procrustes) paiafa paiafa White, 1845 Endophallus
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Kirill Makarov
Carabus (Carabus) stscheglowi Mannerheim, 1827 Endophallus
Russian, Chuvashia, Cheboksary, Zavolzhye distr., , near lake Astrakhanka,
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Ruslan Panin
Carabus (Morphocarabus) odoratus baeri Menetries, 1851: 45 Endophallus
Russia, SW Taymyr, near Talnakh, 25 km N Norilsk, hilly tundra, hillside overgrown with alder slate, dwarfish birches, blueberry, h = 150-250 м, 24-28.06.2003, leg. S.Saluk
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Igor Solodovnikov
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906 Endophallus
NE-Hungary, 4 km S Sátoraljaújhely, right bank riv. Bodrog (Long-erdő),
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Ruslan Panin
Carabus (Morphocarabus) rothi distinguendus Csiki, 1906 Endophallus
NC-Hungary: E- part of Budapest,Cinkota,
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Ruslan Panin
Carabus (Morphocarabus) rothi distinguendus (as bieneri Mandl, 1965) Endophallus
C-Slovakia, Detva Distr., Stara Huta env. h=700-800m,
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Ruslan Panin
Carabus (Morphocarabus) rothi distinguendus (as subparvulus Mandl, 1965) Endophallus
NC-Hungary:Borzsony Mts, Nograd-Diosjeno,
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Ruslan Panin
Carabus (Lamprostus) erenleriensis erenleriensis Schweiger, 1966 Endophallus
NW-Turkey:Bolu Prov., Abant Lake
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Ruslan Panin
Carabus (Sphodristocarabus) adamsi Adams, 1817 Endophallus
CN-Georgia, N of Tbilisi, NW of Tianeti, Tvalivi env.,
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Ruslan Panin
Carabus (Morphocarabus) odoratus kurmaevi I.Solodovnikov, 2017 Holotype Endophallus
Russia, Siberia, Krasnoyarsk Krai, the North Yenisei district, 55 km of CE of the vill. Bryanka, valley of the river Chirimba, stream Nakhodny, h = 350 m, the mixed forest (birch and pine); on a steep slope along an animal track on a hill brow, 23.VII-20.VIII.2015, leg. A.V. Kurmaev
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Igor Solodovnikov
Carabus (Morphocarabus) viridilimbatus saluki I.Solodovnikov 2017 Paratype Endophallus
Russia, SW Taymyr, near Talnakh, 25 km N Norilsk, hilly tundra, hillside overgrown with alder slate, dwarfish birches, blueberry, h = 150-250 м, 24-28.06.2003, leg. S.Saluk
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Igor Solodovnikov
Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859 Endophallus
Russia, Chita Region, Udokansky district. Kodar Range, 56 °34'42,9 " N, 118 °29'34,95" E, 6/13/2002, leg. A. Pokhorov
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Igor Solodovnikov
Carabus (Orinocarabus) linnaei linnaei Panzer, 1812 Endophallus
W.UKRAINE, Lviv Reg., NE Carpathians, Skolivski Beskidy Mts. Zelemyanka Mt. Rng
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Ruslan Panin
Carabus (Orinocarabus) linnaei hoverlaensis Müller A. & Panin R., 2018 Endophallus
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone,
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Ruslan Panin
Carabus (Neoplectes) ibericus ibericus (as imereticus Retezar, 2011) Paratype Endophallus
NC-Georgia, S-Caucasus: W slope Likhi Mt.Rng., 3 km W Rikoti Pass h~900 m,
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Ruslan Panin
Carabus (Tribax) puschkini platypterus Ganglbauer, 1886 Endophallus
SW Georgia, Lesser Caucasus, Adzhar-Imereti (Meskheti) Mt Range, 10 km S Sairme h~1850 m.
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Ruslan Panin
Carabus (Ophiocarabus) successor successor Reitter, 1896 Endophallus
SE-Kazakhstan, Narynkol distr., 4 km SW Bayankol, h~2400-2500 m
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Ruslan Panin
Carabus (Tribax) puschkini kolenatii (as suramensis Kraatz, 1887) Endophallus
S-Caucasus: NC-Georgia, W slope Likhi Mt.Range, 3 km W Rikoti Pass h~900m
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Ruslan Panin
Carabus (Cryptocarabus) tsharynensis Kabak 1994 Endophallus
SE-Kazakhstan, Almaty Region, W-Zailiyskiy Alatau rng, Belshabdar valley h~1300 m
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Ruslan Panin
Carabus (Sphodristocarabus) separatus separatus Gehin, 1885 Endophallus
N-Iran:Gilan Prov., Asalem - Khalkal road,
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Ruslan Panin
Carabus (Tribax) kasakorum georgiensis Gottwald, 1980 Endophallus
S Russia, C-Caucasus: N Ossetia, Chekhatsirtit Mt range, Mt Tepli h=3000 m
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Ruslan Panin
Carabus (Mesocarabus) cantabricus moroderi (as magdalenai Meyer & Mollard, 1996) Endophallus
NW-Spain, Leon prov., Sierra de Gistredo, Pto. de Magdalena,
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Ruslan Panin
Carabus (Tribax) adelphus rex Gottwald, 1980 Endophallus
Abkhazia, Tkuarchalsky district, Kodorsky Mt.R., it is right board of the Bashishkvara River, beechen forest + rhododendron, h = 1693 m, 15-23.07.2018, leg. I.A. Solodovnikov
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Igor Solodovnikov
Carabus (Tribax) adelphus adelphus Rost, 1892 SW-Caucasus:"Samursakan"(=nr.Gali & Anara); Gali env.:Okumi Endophallus
Georgia, Zemo Svaneti., W slopes of Ingurishi Mt., right bank of Inguri riv., 09.V-20.VI.2018 leg. D. Fominykh
Copyright ©
Igor Solodovnikov
Carabus (Tribax) nacharensis Rost, 1893 Endophallus
Abkhazia, vall. Kluch River, 13.08.1928
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Igor Solodovnikov
Carabus (Tribax) adelphus shtavlerensis Fominykh, Zamotajlov et Titarenko, 2018 Endophallus
Georgia, Zemo Svaneti, left bank of Nenskra River near Tita Vill., Shtavler Mt. Range, NW slopes of Utur Mt., 43°04'N, 42°11'E, 1048 m, 14.IV–13.VI.2016 (D. Fominykh) HOLOTYPE
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Igor Solodovnikov
Carabus (Coptolabrus) elysii xiaoganicus Deuve et Remond, 2016 Endophallus
China, Hubei, Xiaogan, Mt Shuangfeng Shan,
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Ruslan Panin
Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876 Endophallus
NC-Spain, Basque Co, Álava, Sierra de Gorbea.
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Ruslan Panin
Carabus (Oreocarabus) luetgensi pseudosteuarti Lapouge, 1924 Endophallus
Portugal, Guarda, Sierra de Estrela h=800 m,
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Ruslan Panin
Carabus (Neoplectes) compressus rebellis Reitter, 1895 Endophallus
Georgia, Kvemo-Svaneti, S slopes of Svanetskiy Mt.R., S slope of Latpari pass, h=2225m, 17.VI -13.VII.2017 leg. D.Fominykh
Copyright ©
Kirill Makarov
Carabus (Neoplectes) polychrous svetlanae Solodovnikov, 2015 Endophallus
Copyright ©
Igor Solodovnikov
Carabus (Neoplectes) polychrous rousianus Gottwald, 1985 Endophallus
Copyright ©
Igor Solodovnikov
Carabus (Neoplectes) polychrous khipstaensis Solodovnikov, 2015 Endophallus
Copyright ©
Igor Solodovnikov
Carabus (Pseudocoptolabrus) watanabei valdemar Deuve, 2016 Paratype Endophallus
Chine, Yunnan, Dêqên, monts Hengduan Shan, près du col de Doker La, 3600 m
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Thierry Deuve
Carabus (Morphocarabus) excellens ciscarpathicus Müller A., Panin R. & Kanarskiy Yu., 2019 Holotype Endophallus
W-Ukraine, Ciscarpathian, Lviv Region, Stryi distr., Biletsky flood-plain forest, near Pyatnychany vill.
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Ruslan Panin
Carabus (Neoplectes) martviliensis Retezar & Djavelidze, 1992 Endophallus
Zeda Gordi env., 980 m and Doberazeni env., 600 m.
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Kirill Makarov
Carabus (Neoplectes) C. martviliensis mzekalae Endophallus
Melouri env., 475 m and Dzedzileti env. , 770 m
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Kirill Makarov
Carabus (Orinocarabus) sylvestris transsylvanicus Dejean, 1826 Endophallus
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030-2040 m alpin zone,
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Ruslan Panin
Carabus (Rhigocarabus) gigolo Heinz & Brezina, 1996 Endophallus
China, Tewo. det. Anichtchenko, A. 2020
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Alexander Anichtchenko
Carabus (Deroplectes) klapperichianus coiffaitianus Deuve, 1990 Endophallus
Tadzhikistan: Badakhshan: Ishkashim Mt.R.: Kuh-i-Lal, VI.1979 leg. I.K. Lopatin
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Igor Solodovnikov
Carabus (Rhigocarabus) buddaicus gansuicus Deuve, 1991 Endophallus
N Sichuan, Langmusi vill 3700 m, 1-25.7.2018
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Alexander Anichtchenko
Carabus (Rhigocarabus) roborowskii langmusiensis Deuve, 1996 Endophallus
N Sichuan, near Langmusi, 3700m 1-25.7.2018, V. Patrikeev leg. det. Anichtchenko A. 2020
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Alexander Anichtchenko
Carabus (Ohomopterus) uenoi Ishikawa, 1960 Endophallus
Copyright ©
Jong-Bong Choi
Carabus (Ophiocarabus) ernsti ulastaiensis Kabak, 2019 Paratype Endophallus
China, Xinjiang, S slopes of Boro-Horo Mt. R., ridge between Kaptshik and Zekku rivers, NE of Ulastai (= Wulasitai) Vill., alpine meadows, 43°56'07"N / 83°18'33"E – 43°56'22"N / 83°18'30"E, 3135–3280 m, 17.07.2017 leg I.I. Kabak
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Kirill Makarov
Carabus (Cryptocarabus) sacarum Kabak, 1998 Endophallus
SE-Kazakhstan, Almaty reg., C-Zailiyskiy Alatau N-slopes, Kaskalen river Valley, Izvestkovy vill., h=1700 m,
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Ruslan Panin
Carabus (Eucarabus) angustus dopyeong Rapuzzi, 2015 Endophallus
scale bar = 5mm
Copyright ©
Jong-Bong Choi
Carabus (Eucarabus) sternbergi sternbergi Roeschke, 1898 Endophallus
Scale bar = 5mm
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Jong-Bong Choi
Carabus (Eucarabus) sternbergi gureopensis J.K. Park & J. Park, 2013 Paratype Endophallus
Scale bar = 5mm
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Jong-Bong Choi
Carabus (Eucarabus) cartereti cartereti Deuve, 1982 Endophallus
Scale bar = 5mm
Copyright ©
Jong-Bong Choi
Carabus (Eucarabus) nitididorsus nitididorsus Ishikawa & Kim, 1983 Endophallus
Scale bar = 5mm
Copyright ©
Jong-Bong Choi
Carabus (Eucarabus) nitididorsus paegunsanus Imura, 1997 Endophallus
Scale bar = 5mm
Copyright ©
Jong-Bong Choi
Carabus (Eucarabus) nitididorsus baegyangsanus (Choi & Park, 2019) Holotype Endophallus
Scale bar = 5mm
Copyright ©
Jong-Bong Choi
Carabus (Eucarabus) namhaedoensis Kwon & Lee, 1984 Endophallus
Scale bar = 5mm
Copyright ©
Jong-Bong Choi
Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822 Endophallus
Russia, S Tuva, Khorumnug-Taiga Mts, Buren riv. Valley, Buren-Bay-Khaak vill. h=900 m
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Ruslan Panin
Carabus (Aulonocarabus) truncaticollis fleischeri Reitter, 1898 Endophallus
Russia, N Burajatia, Barguzinskiy Mts., the upper reaches of M.Chivyrkuy riv., h=2000 m
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Ruslan Panin
Carabus (Mesocarabus) lusitanicus helluo (as alicantinus Ganglbauer, 1886) Endophallus
SE-Spain, Murcia reg., Jumilla, Sierra del Carche, h=1300 m,
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Ruslan Panin
Carabus (Orinocarabus) bertolinii Kraatz, 1878 Endophallus
NE-Italy, Trentino prov. Dolomites Alps, Plateau Pale di San Martino h=2500 m,
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Ruslan Panin
Carabus (Eucarabus) xiuyanensis Deuve & Li, 1998 Endophallus
NE-China: SE-Liaoning, Xiuyan County, Dayingzi, Mt. Laopingding,
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Ruslan Panin
Carabus (Euleptocarabus) porrecticollis porrecticollis Bates, 1883 Endophallus
Japan, C Honshu, Nagano Pre., Toyooka town,
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Ruslan Panin
Carabus (Eucarabus) billbergi kuandianicus Rapuzzi, 2007 Endophallus
NE China, Liaoning, Kuandian County, Dachuantou env.,
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Ruslan Panin
Carabus (Tribax) macropus Chaudoir, 1877 Endophallus
Russia, Ingushetia, S slopes of Skalistyi Mt.R., Khahalgi Mt., alpin zone, h = 2500 m
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Igor Solodovnikov
Carabus (Carabus) vinctus Weber, 1801 Endophallus
LOUISIANA, East Baton Rouge Parish, 1.5mi W Denham Springs, 30.482 ̊ N 90.981 ̊ W, bottomland hardwood forest, 4 Sep 2017
Copyright ©
Igor Sokolov
Carabus (Cechenochilus) heydenianus Starck, 1889 Endophallus
Carabus (Procechenochilus) heydenianus euxinus Semenov, 1896 Abkhasia: Arabika plato (TOPOTYPUS)
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Igor Solodovnikov
Carabus (Megodontus) obovatus Fischer, 1827 Endophallus
C.obovatus taizhensis Obydov, 2006 Khakassia: Kuznetskyi Alatau, env. vill. Priiskovyi, Bobrovaya Mt., h = 1200-1300 m
Copyright ©
Igor Solodovnikov
Carabus (Megodontus) kolbei hidakamontanus Ishikawa, 1966 Endophallus
E-Hokkaido: Samani distr., Horomankyo env.
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Igor Solodovnikov
Carabus (Ohomopterus) dehaanii dehaanii Chaudoir, 1848 Endophallus
Japan, S Honshu, Mie Pref., Inabe city
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Ruslan Panin
Carabus (Procrustes) coriaceus ressli Mandl, 1961 Endophallus
SC-Turkey:Hatay Prov, Dörtyol, Nur Mts, Topaktaş h=900-1200 m,
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Ruslan Panin
Carabus (Ctenocarabus) melancholicus costatus Germar, 1824 Endophallus
NW Portugal, Viana do Castelo distr., Sierra de Agra
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Ruslan Panin
Carabus (Apotomopterus) protenes baoxingensis Deuve & Mourzine, 2003 Endophallus
China: С-Sichuan:Qionglai Shan Mts., Baoxing env. (TOPOTYPUS)
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Igor Solodovnikov
Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896 Endophallus
Russia, Caucasus, Ingushetia, South slopes of Skalistyi Mt Range, Stolovaya Mt., near Beyni, alpin zone, h = 2800 m, N 42°51’35.30",E 44°42’13.70", h = 2800m, 21.V–19.VII.2018, D. Fominykh leg. (TOPOTYPUS)
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Igor Solodovnikov
Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896 Endophallus
Russia, Caucasus, Ingushetia, Skalistyi Mt. Range, S slopes of Tsoreylam Mt. Range, 5 km W of Mt. Khakhalgi, N 42°50’07.60", E 45°05’02.20", h=2500 m, 25.V–21.VII.2017, D. Fominykh leg.
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Igor Solodovnikov
Carabus (Macrothorax) rugosus celtibericus Germar, 1824 Endophallus
SW-Portugal, Serra do Cercal h=217 m, 37°47'15.6"N 8°42'02.0"W, road N390,
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Ruslan Panin
Carabus (Tribax) kasakorum georgiensis Gottwald, 1980 Endophallus
Russia: N.Osetia, Bokovoy Mt.R., near Arkhonskiy pass., alpin zone, h = 2700 m, 23.V-21VII.2018, D. Fominykh leg.
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Igor Solodovnikov
Carabus (Mesocarabus) lusitanicus trabuccarius Fairmaire, 1857 Endophallus
NE Spain, Catalonia, Tarragona, Sierra de Montsant h=800 m, Albarca env.,
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Ruslan Panin
Carabus (Mesocarabus) lusitanicus lusitanicus (as vieirae Oliveira, 1875) Endophallus
CW-Portugal: Leiria distr., near Marinha Grande, 39°45'57.8"N 8°58'10.3"W, h-79 m
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Ruslan Panin
Calosoma (Callisthenes) breuningi iliense Kabak, 2022 Endophallus
W of Kerbulak, 44°03ʹ03ʹʹ N, 77°01ʹ12ʹʹ E, 640 m, 25.IV.2017, I. I. Kabak leg.
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Ilya Kabak
Carabus (Tribax) apschuanus galianus Belousov & Zamotajlov, 1993 Endophallus
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Igor Solodovnikov
Carabus (Tribax) edmundi Semenov, 1896 Endophallus
Russia, S-Dagestan
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Igor Solodovnikov
Carabus (Tribax) osseticus abanosicus Deuve, 2013 Endophallus
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Igor Solodovnikov
Carabus (Tribax) osseticus osseticus Adams, 1817 Endophallus
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Igor Solodovnikov
Carabus (Tachypus) cancellatus cancellatus Illiger, 1798 Endophallus
Carabus (Tachypus) cancellatus tuberculatus f.conspersus Lapouge, 1902 (Blaс form)
N-Belarus, Mogilev reg., Bykhov distr., 5 km N Voronino Vill., mixed-pine forest, 20.V-03.VI.2007, Komyagunova N.P.
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Igor Solodovnikov
Carabus (Sphodristocarabus) scovitzi scovitzi Faldermann, 1835 Endophallus
Georgia, Adzharia, Adzharo-Imeretinskiy Mt.R., near Tchidila pass, alpin zone, h=2350 m, 05.VI.2016 leg. D. Fominykh
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Igor Solodovnikov
Carabus (Sphodristocarabus) tokatensis kosensis Cavazzuti, 1984 Endophallus
CE-Turkey: Gümüshane: Köse Dag(pass N of Köse), h = 1700 m, VII.2000, leg. St. Il. Remond (TOPOTYPUS)
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Igor Solodovnikov
Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812 Endophallus
Carabus (Tachypus) cancellatus emarginatus n. dahli Heer, 1841
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Igor Solodovnikov
Carabus (Morphocarabus) odoratus antropovi Shilenkov 1996 Endophallus
Russia, S Siberia, Tuva, W Tannu-Ola Mts. 35 km NE Khandagayty vill., Teeli Pass 2200-2800 m
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Ruslan Panin
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906 Endophallus
SE Poland, Subcarpathia Prov., Nadleśnictwo Kołaczyce N of Jasło vill., M. Kosibowicz leg./det. R Panin
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Ruslan Panin
Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822 Endophallus
Russia, NE Altay, Turochak Distr., Lake Teletskoe N part, near Yaylyu vill.
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Ruslan Panin
Carabus (Morphocarabus) aeruginosus eschscholtzi Mannerheim, 1827 Endophallus
NE Kazakhstan, W Altay Mts, near Shemonaikha city., flood plain,
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Ruslan Panin
Carabus (Meganebrius) santostamangi santostamangi Endophallus
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Joachim Schmidt
Carabus (Meganebrius) thudamensis Deuve & Schmidt, 2017 Holotype Endophallus
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Joachim Schmidt
Carabus (Meganebrius) everesti everesti Andrewes, 1929 Endophallus
Kemba Kharka
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Joachim Schmidt
Carabus (Meganebrius) salpansis Deuve, 1984 Endophallus
Kemba Kharka
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Joachim Schmidt
Carabus (Meganebrius) indicus indicus Fairmaire, 1889 Endophallus
Tonglu
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Joachim Schmidt
Carabus (Rhigocarabus) casanova (Imura & Brezina, 2008) Endophallus
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Boleslav Brezina
Carabus (Rhigocarabus) gigolo Heinz & Brezina, 1996 Endophallus
Langmusi
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Boleslav Brezina
Carabus (Rhigocarabus) gentleman Brezina & Hackel, 2004 Endophallus
Jiangjing
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Boleslav Brezina
Carabus (Rhigocarabus) gigoloides priapeius Cavazzuti, 2006 Endophallus
Yeliguan
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Boleslav Brezina
Carabus (Rhigocarabus) nianjuaensis Imura, 2016 Endophallus
Image from Imura, 2016
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Carabus (Rhigocarabus) kozaburoi Imura, 1999 Paratype Endophallus
Gaoligong Shan, N. Myanmar. Image from Imura, 2016
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Carabus (Rhigocarabus) itzingeri rugulosior Deuve, 1996 Holotype Endophallus
NW-Yunnan: Hengduan Shan: Yanmen env.: 28:06N/98:52E. Image from Imura, 2016
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Carabus (Rhigocarabus) itzingeri tanakai Imura, 1997 Endophallus
Image from Imura, 2016
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Carabus (Rhigocarabus) itzingeri choguy Imura, 2001 Holotype Endophallus
Image from Imura, 2016
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Carabus (Rhigocarabus) rhododendron xuebangshanensis Imura, 2016 Holotype Endophallus
Image from Imura 2016
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Carabus (Rhigocarabus) rhododendron xinshanchangensis Imura, 2016 Holotype Endophallus
Image from Imura, 2016
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Carabus (Rhigocarabus) rhododendron lidimaensis Imura, 2016 Holotype Endophallus
Image from Imura, 2016
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Carabus (Rhigocarabus) rhododendron s.str. Holotype Endophallus
Image from Imura, 2016
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Carabus (Carabus) donjuan Imura, 2011 Holotype Endophallus
Image from Imura, 2011
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Carabus (Pseudocoptolabrus) belousovianus Imura, 2011 Endophallus
Image from Imura, 2011
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Carabus (Carabus) koidei yanyuan Cavazzuti & Rapuzzi, 2010 Endophallus
Image from Imura, 2011
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Carabus (Hemicarabus) nitens Linne, 1758 Endophallus
Belarus, Vitebsk reg., Miory distr. Swamp «Mokh», 09.05.2011, leg. G.G. Sushko
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Igor Solodovnikov
Carabus (Hemicarabus) tuberculosus Dejean, 1829 Endophallus
Carabus (Hemicarabus) tuberculosus n. strophium F.-W., 1842 SE Kazakhstan: NW Tarbogatai Mt.R.
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Igor Solodovnikov
Carabus (Hemicarabus) serratus Say, 1825 Endophallus
E-USA,Virginia, Leesburg env., Ball's Bluff Battlefield Regional Park
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Igor Solodovnikov
Carabus (Eucarabus) parreyssi parreyssi Palliardi, 1825 Endophallus
SW-Croatia, Mts Velebit, Mali Halan, h=1390 m,
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Ruslan Panin
Carabus (Megodontus) croaticus croaticus Dejean, 1826 Endophallus
W-Croatia, Velebit Mts., h=950 m, Oltari - Zavizan,
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Ruslan Panin
Carabus (Chaetocarabus) intricatus bayardi (as silaensis Haury, 1881) Endophallus
S-Italy, Calabria, Sila Grande Massif, Camigliatello h=1040 m
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Ruslan Panin
Carabus (Chaetocarabus) intricatus bayardi (as silaensis Haury, 1881) Endophallus
S-Italy, Calabria, Sila Grande Massif, Camigliatello h=1040 m, 19.XI.2009, G. Proscia leg. L=31 mm
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Ruslan Panin
Carabus (Cupreocarabus) balangicus balangicus Cavazzuti, 2002 Endophallus
Chine, C-Sichuan, Xiaojin, Wolong valley: Balang Shan Pass h=3448 m, 30.884617, 102.973117
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Ruslan Panin
Carabus (Rhigocarabus) roborowskii roborowskii Semenov, 1887 Endophallus
China, E-Qinghai, Mts NW of Tongren city, 35.562778, 101.843667, h=3870 m
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Ruslan Panin
Carabus (Neoplectes) zolotarevi Zamotajlov, 1988 Endophallus
Carabus (Archiplectes) zolotarevi zolotarevi Zamotajlov, 1988
Russia: Krasnodar reg., SW slopes of Malye Balkany Mt.R., env. Allous pass, h = 1960 m, 08-18.VII.2023 leg. D. Fominykh
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Igor Solodovnikov
Carabus (Deroplectes) arcanus arcanus Semenov, 1898 Endophallus
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Igor Solodovnikov
Carabus (Deroplectes) arcanus hissarianus Semenov, 1900 Endophallus
Tajikistan, Gissar ridge, Kondara, Kwak, h = 1900 m, 7,VI.1970, I.K. Lopatin leg.
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Igor Solodovnikov
Carabus (Deroplectes) sphinx dardazicus Deuve, 1990 Endophallus
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Igor Solodovnikov
Carabus (Cechenochilus) boeberi buschi Semenov & Znojko, 1931 Endophallus
Russia: Kabardino-Balkaria: u.r. Adylsu Riv.
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Igor Solodovnikov
Carabus (Megodontus) tuvensis Shilenkov, 1996 Endophallus
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Igor Solodovnikov
Carabus (Mesocarabus) riffensis Fairmaire, 1875 Endophallus
N-Morocco, Riff Mts h=1650 m, 26 km E of Bab Berred
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Ruslan Panin
Carabus (Isiocarabus) miaorum Lassalle & Prunier, 1993 Endophallus
China, NW-Guangxi, Mao'er Shan, h~1600 m 25°52'N 110°29'E
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Ruslan Panin
Carabus (Apotomopterus) delavayi parvipes Cavazzuti, 1996 Endophallus
S-Sichuan: S of Yanyuan, Chuandongzi Massif h=3200 m, 27.351206, 101.507036
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Ruslan Panin
Carabus (Apotomopterus) ludivinae wuliangensis Cavazzuti, 1997 Endophallus
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Ruslan Panin
Carabus (Apotomopterus) protenes ascendens Semenov, 1898 Endophallus
China, Chongqing, N Jinfo Shan, Shanwangping h=1400m, 29.101944, 107.166389
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Ruslan Panin
Carabus (Pachystus) cribratus cribratus Quensel, 1806 Endophallus
NW Armenia, Amasia, (Shirak prov.)
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Ruslan Panin
Carabus (Apotomopterus) korellianus Kleinfeld, 2002 Endophallus
China, SW-Yunnan, Dehong Distr., Ruili, Pijiazhai vill., h=1800 m, 24.048111, 97.738222
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Ruslan Panin
Carabus (Apotomopterus) madefactus Cavazzuti, 1997 Endophallus
China, CW-Yunnan, Nánjiàn Yízú Zìzhìxiàn, Mt Lingbao h=2400 m, 24.768611, 100.500000
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Ruslan Panin
Carabus (Sigenacarabus) trachynodes moxiensis Deuve, 1992 Endophallus
China, W-Sichuan, Daxue Range, Mt.Gongga Shan h=3000-3100m, Hailuogou Glacier Forest Park, 29.574889, 101.995556
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Ruslan Panin
Carabus (Acoptopterus, ?Carabus) yunnanus yanyuanicus Cavazzuti, 1996 Endophallus
China, S-Sichuan, S Yanyuan, Chuandongzi Mt h=3300-3600 m
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Ruslan Panin
Carabus (Apotomopterus) mecynodes pinchuanensis Imura, 1999 Endophallus
China, S-Sichuan, 40-50 km E Xichang (to Zhaojue) 27°51'N 102°30'E, h=2700-3000m, Pine forest
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Ruslan Panin
Carabus (Apotomopterus) patroclus apricalis Cavazzuti, 1997 Endophallus
China, S Sichuan, W of Pingchuanzhen, near Galuo vill., h=2400 m
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Ruslan Panin
Carabus (Apotomopterus) microtatos lingbaonis Imura, 2019 Endophallus
China, CW-Yunnan, Nánjiàn Yízú Zìzhìxiàn, Mt Lingbao h=2400 m, 24.768611, 100.500000
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Ruslan Panin
Carabus (Apotomopterus) bousquetellus bousquetellus Deuve, 1998 Endophallus
S-China, CW-Yunnan, W of Lincang, Daxue Shan h=2600m, 23.935556, 100.257778
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Ruslan Panin
Carabus (Apotomopterus) patroclus patroclus Breuning, 1933 Endophallus
China, S Sichuan, Pass 56 km SW Miannin, near Wawo h=2440 m
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Ruslan Panin
Carabus (Semnocarabus) erosus karascharensis Eidam Lectotype Paralectotypes Endophallus
aedeagus - lectotype. Endophallus - paralectotype.
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Kirill Makarov
Carabus (Semnocarabus) cicatricosulus pseudoerosus Mandl, 1955 Endophallus
left tributary of Kuqa River, E of Lake Kyrghyz-Artukul (= Dalongchi), 42°27'15''N, 83°22'00''E, H ~ 2550–2600 m, 7–8.VI.2002, I. Kabak leg
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Kirill Makarov
Carabus (Ophiocarabus) successor eoketmenicus Kabak, 2016 Holotype Paratype Endophallus
endophallus - paratypes; aedeagus - holotype
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Ilya Kabak
Carabus (Ophiocarabus) kurdaiensis Kabak, 2016 Paratype Endophallus
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Ilya Kabak
Carabus (Ophiocarabus) ernsti Kabak, 2001 Endophallus
7–8, 12–13 – C. ernsti ulastaiensis, paratypes; 9–10, 14–15 – C. ernsti ernsti Kabak, 2002: 9, 14 – from the locality in mountains to the West of the Saryk River, 10, 15 – from the locality in right tributary of the Saryk River; 11,16 – C. ernsti nilkiensis Kabak, 2014: 11 – holotype, 12 – paratype. ml – median lobe; pp – praeputial pad; rbll – right basolateral lobe.
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Ilya Kabak
Carabus (Coptolabrus) lafossei tungchengensis Li, 1993 Endophallus
China, Hubei, Shangzhi co., Shimenyan 8-11.6.2009 (cAA)
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Alexander Anichtchenko
Carabus (Tribax) puschkini kraatzi Chaudoir, 1877 Endophallus
Georgia, Samtskhe-Javakheti, env. Uraveli vill., 14.VI.2024 leg. D. Fominykh
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Igor Solodovnikov
