Pterostichus (Adelopterus) ambiguus (Fairmaire, 1858)
Female adult
France, 20224, Corse, Calacuccia forêt de Valdu Niellu 12-09-2000
Copyright © Pascal Deschamps

Pterostichus (Adelosia) macer Marsham, 1802
Male adult

E. Ukraine,
Kharkov reg., 1972
16mm.

Copyright © Ruslan Panin

Pterostichus (Adelosia) macer Marsham, 1802
Male adult

Length: 15.4 mm
West Kazakhstan,
Uralskaya area, nearby Volodarskoye village, sands.
15.04.2000
F.G.Bidashko leg. and det.

Copyright © Alexander Anichtchenko

Pterostichus (Anilloferonia) malkini (Hatch, 1953)
Male adult
Image from Labonte, 2013
Copyright © - - -

Pterostichus (Anomostichus) anomostriatus Sciaky, 1995 Paratype
Male adult
China – pr. Sichuan, Emei mt. 2500 4-20.V.1989 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Pterostichus (Aphaonus) compressus Rost, 1892
Male adult
Russia, Krasnodar reg., 6 km NE Soloniki vill.,
Sochi Nat.Park, 28-30.V.2009 Chibilov E. leg.
photo A. Anichtchenko
det.I.A.Solodovnikov
Copyright © Alexander Anichtchenko

Pterostichus (Aphaonus) cylindriformis Reitter, 1887e: 252
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Aphaonus) mariamae Zamotajlov, Solodovnikov & Fominykh, 2015 Holotype
Male adult
Abkhazia, Gulripsh Distr., Tsebelda Plateau near village Ablukhvara, left bank of River Kelasur, slopes of Mt. Bashlug, 650 m, 15.05.-19.07.2012, pitfall traps (D. Fominykh) (ZISP).
Copyright © Kirill Makarov

Pterostichus (Aphaonus) arcanoides Lorenz, 1998: 12
Female adult
Chedym Mt. Range
Copyright © Kirill Makarov

Pterostichus (Aphaonus) svetlanae Zamotajlov, Solodovnikov & Fominykh, 2015 Holotype
Female adult
Abkhazia, Gudauta Distr., vill. Mzara (= Chiryuta) vicinities, valley of River Mzara, Zashirbara Mt. Range, N 43º10'18" / E 040º48'23", 705 m, 15.05.2014 (I. Solodovnikov & S. Solodovnikova) (CSIA).
Copyright © Igor Solodovnikov

Pterostichus (Aphaonus) neilgaimani Chaladze & Kalatozishvili, 2017 Holotype
Male adult
Image from Chaladze & Kalatozishvili, 2017
Copyright © - - -

Pterostichus (Aphaonus) trubilini Zamotajlov, 1997
Male adult
Image courtesy www.zin.ru
Copyright © - - -

Pterostichus (Aphaonus) trubilini Zamotajlov, 1997a: 6
Male adult

Copyright © Vittorino Monzini

Pterostichus (Aphaonus) cylindriformis Reitter, 1887e: 252
Male adult

Copyright © Vittorino Monzini

Pterostichus (Aphaonus) arcanoides Lorenz, 1998: 12
Male+Female adult
Chedym Mt. Range
Copyright © - - -

Pterostichus (Argutor) commutabilis (Motschulsky, 1866)
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada

Copyright © Henri Goulet

Pterostichus (Argutor) praetermissus (Chaudoir, 1868)
Male adult
Canada. Copyright Henri Goulet, Agriculture & Agri-Food Canada det. H. Goulet
Copyright © Henri Goulet

Pterostichus (Argutor) sulcitarsis A. Morawitz, 1862a: 250
Male adult
Collection ground Mino city, Osaka 37179
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Argutor) vernalis Panzer, 1796
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Argutor) sulcitarsis A. Morawitz, 1862a: 250
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Argutor) cursor Dejean, 1828: 243
Female adult
det. K. Makarov
Copyright © Alexander Anichtchenko

Pterostichus (Argutor) leonisi Apfelbeck, 1904: 257
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Argutor) leonisi Apfelbeck, 1904: 257
Male adult

Copyright © Igor Solodovnikov

Pterostichus (Argutor) leonisi Apfelbeck, 1904: 257
Female adult

Copyright © Igor Solodovnikov

Pterostichus (Phonias) stricticollis (Solsky, 1874)
Male adult
Kirghisia, Osh, 18.V.1997, V. Gromenko leg. (cGA). det. I. Kabak, 2021
Copyright © Gianni Allegro

Pterostichus (Asioplatysma) medvedevi Kryzhanovskij & Mikhailov, 1972: 319
Male adult

Copyright © Kirill Makarov

Pterostichus (Badistrinus) laticollis (Motschulsky, 1844)
Male adult

    length: 9,9 mm
    Transbaikalia, Tchita area, Nertchinskyi distr., Nertcha river valley near Pravye Kumaki village
    4.08.1999
    Zinoviev E.leg.

Copyright © Evgeniy Zinoviev

Pterostichus (Eurythoracana) haptoderoides Tschitscherine, 1889
Female adult
length: 8,7 mm; Russia, Far East, the vicinities of Khabarovsk sity., 13.07.1986; Tolmacheva R. leg., Dudko R. det.
Copyright © Evgeniy Zinoviev

Pterostichus (Badistrinus) procephalus Bates, 1873a: 285
Male adult
Collection ground Misaki cho, Chiba Pref. 39611
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Pledarus) neglectus A.Morawitz, 1862
Male adult
Collection ground Ishikari, Hokkaido Pref. 28058
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Eurythoracana) haptoderoides japanensis (Lutshnik, 1922)
Female adult
Collection ground Inami cho, Hyogo Pref. 39361
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Badistrinus) bandotaro Tanaka, 1958c: 78 Holotype
Male adult
Lake Wadanuma in the river-bed of Tonegawa, Chiba Pref., Japan, 9. IV. 1949, K. Tanaka<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-241<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Badistrinus) bandotaro Tanaka, 1958c: 78 Holotype
Male adult
Lake Wadanuma in the river-bed of Tonegawa, Chiba Pref., Japan, 9. IV. 1949, K. Tanaka. Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-241. (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Badistrinus) bandotaro Tanaka, 1958c: 78 Holotype
Male adult
Lake Wadanuma in the river-bed of Tonegawa, Chiba Pref., Japan, 9. IV. 1949, K. Tanaka Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-241 (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Badistrinus) bandotaro Tanaka, 1958c: 78 Holotype
Label
Lake Wadanuma in the river-bed of Tonegawa, Chiba Pref., Japan, 9. IV. 1949, K. Tanaka Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-241 (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Badistrinus) laticollis (Motschulsky, 1844)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Badistrinus) haptoderoides haptoderoides
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Biphonias) neglectoides Budilov, 2022 Holotype
Male adult
Russia: Jewish Autonomous Region, the protected zone of ‘Bastak’ Reserve, 19.X 2016, leg. P. Budilov (FCBV).
Copyright © Pavel Budilov

Pterostichus (Biphonias) neglectus A.Morawitz, 1862
Male adult

Copyright © Pavel Budilov

Pterostichus (Badistrinus) neglectus A.Morawitz, 1862 Lectotype
Male adult
M. Bureja 110-1. L=5.1 mm
Copyright © Kirill Makarov

Pterostichus (Badistrinus) neglectus A.Morawitz, 1862 Lectotype
Label
M. Bureja 110-1.
Copyright © Kirill Makarov

Pterostichus (Badistrinus) neglectus A.Morawitz, 1862
Male adult
Primorye Territory, Khasansky district, 18 km SW Barabash, the floodplain of the Bezymyanny Stream, swamp 43°3’52”N 131°21’49”E, 16.VIII.2020, K. Makarov leg.
Copyright © Kirill Makarov

Pterostichus (Badistrinus) neglectus A.Morawitz, 1862
Male genitalia
Primorye Territory, Khasansky district, 18 km SW Barabash, the floodplain of the Bezymyanny Stream, swamp 43°3’52”N 131°21’49”E, 16.VIII.2020, K. Makarov leg.
Copyright © KirillHat

Pterostichus (Badistrinus) neglectus A.Morawitz, 1862
Male adult
Khabarovsk Territory, Khawar Stream below cordon Bichi, 24.VII.2021, Yu. Sundukov & L. Sundukova leg.
Copyright © Kirill Makarov

Pterostichus (Badistrinus) neglectus A.Morawitz, 1862
Endophallus
Khabarovsk Territory, Khawar Stream below cordon Bichi, 24.VII.2021, Yu. Sundukov & L. Sundukova leg.
Copyright © Kirill Makarov

Pterostichus (Badistrinus) goschi Jedlicka, 1930 Lectotype
Male adult
Nikolsk Ussurijsk Ussurigeb. Mandl
Copyright © Kirill Makarov

Pterostichus (Badistrinus) goschi Jedlicka, 1930
Male adult
Nikolsk Ussurijsk Ussurigeb. Mandl
Copyright © Kirill Makarov

Pterostichus (Badistrinus) goschi Jedlicka, 1930 Lectotype
Male adult
Nikolsk Ussurijsk Ussurigeb. Mandl
Copyright © Kirill Makarov

Pterostichus (Badistrinus) goschi Jedlicka, 1930
Male genitalia
Nikolsk Ussurijsk Ussurigeb. Mandl
Copyright © Kirill Makarov

Pterostichus (Badistrinus) goschi Jedlicka, 1930
Endophallus
Nikolsk Ussurijsk Ussurigeb. Mandl
Copyright © Kirill Makarov

Pterostichus (Badistrinus) goschi Jedlicka, 1930 Lectotype
Label
Nikolsk Ussurijsk Ussurigeb. Mandl
Copyright © Kirill Makarov

Pterostichus (Badistrinus) goschi Jedlicka, 1930 Paralectotypes
Male adult
Nikolsk Ussurijsk Ussurigeb. Mandl
Copyright © Kirill Makarov

Pterostichus (Badistrinus) goschi Jedlicka, 1930 Paralectotypes
Label
Nikolsk Ussurijsk Ussurigeb. Mandl
Copyright © Kirill Makarov

Pterostichus (Badistrinus) goschi Jedlicka, 1930
Male adult
Primorye Territory, Khasansky district, 10.7 km east of Khasan, Cape Ostrovok Falshivy, 42°26’54”N 130°46’47”E 23.V.2022, K. Makarov leg.
Copyright © Kirill Makarov

Pterostichus (Badistrinus) goschi Jedlicka, 1930
Male genitalia
Primorye Territory, Khasansky district, 10.7 km east of Khasan, Cape Ostrovok Falshivy, 42°26’54”N 130°46’47”E 23.V.2022, K. Makarov leg.
Copyright © Kirill Makarov

Pterostichus (Badistrinus) goschi Jedlicka, 1930
Endophallus
Primorye Territory, Khasansky district, 10.7 km east of Khasan, Cape Ostrovok Falshivy, 42°26’54”N 130°46’47”E 23.V.2022, K. Makarov leg.
Copyright © Kirill Makarov

Pterostichus oblongopunctatus (Fabricius, 1787: 202)
Male adult
Pterostichus (Bothriopterus) oblongopunctatus
Chelyabinskaya obl., Katav-Ivanovskiy prov., mountain Maliy Iremal, alt. 1180 m. 19/08/2008.
Leg. E.Chibilov
Copyright © Sunvas

Pterostichus (Bothriopterus) adstrictus Eschscholtz, 1823
Male adult
Canada
Copyright © Henri Goulet

Pterostichus (Bothriopterus) oblongopunctatus (Fabricius, 1787)
Male adult
SE Siberia, Irkutsk Area, near Bratsk city, 30.VII.07, Fominyh D. leg. long=13mm.
Copyright © Ruslan Panin

Pterostichus (Bothriopterus) oblongopunctatus (Fabricius, 1787)
Male adult
<p> long=11.5mm</p>
Copyright © Ruslan Panin

Pterostichus (Bothriopterus) oblongopunctatus (Fabricius, 1787)
Female adult
photo Vlasenko A. S.
Copyright © Andrey Vlasenko

Pterostichus (Bothriopterus) oblongopunctatus (as honshuensis Habu et Baba, 1958) Holotype
Male adult
Mt. Monnai, NIIGATA, 28. VII. 1957, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-038
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Bothriopterus) oblongopunctatus (as honshuensis Habu et Baba, 1958) Holotype
Male adult
Mt. Monnai, NIIGATA, 28. VII. 1957, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-038
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Alexander Anichtchenko

Pterostichus (Bothriopterus) oblongopunctatus (as honshuensis Habu et Baba, 1958) Holotype
Male adult
Mt. Monnai, NIIGATA, 28. VII. 1957, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-038
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Bothriopterus) oblongopunctatus (as honshuensis Habu et Baba, 1958) Holotype
Male adult
Mt. Monnai, NIIGATA, 28. VII. 1957, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-038
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Bothriopterus) subovatus (Motschulsky, 1860)
Male adult
Collection ground Rebun Is. Hokkaido Pref. 27191
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Bothriopterus) subovatus (Motschulsky, 1860)
Endophallus
Collection ground Rebun Is. Hokkaido Pref. 27191
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Bothriopterus) subovatus (Motschulsky, 1860)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Bothriopterus) trinarius (Casey, 1918)
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Pterostichus (Bothriopterus) mariae (Lutshnik, 1921)
Male adult

Copyright © Kirill Makarov

Pterostichus (Bothriopterus) oblongopunctatus (Fabricius, 1787)
Male adult

Copyright © Kirill Makarov

Pterostichus (Bothriopterus) adstrictus Eschscholtz, 1823
Male adult
Russia, Primorskii reg., Lazo env.
Copyright © Kirill Makarov

Pterostichus (Bothriopterus) aeneocupreus (Fairmaire, 1887)
Male adult
Bhutan, near Gangcheyting, [Camp 2] 3689 m, 27° 24 14.0''N, 90° 14' 04.2''E, 9.oct. 2017
Copyright © Joachim Schmidt

Pterostichus (Calopterus) pilosus (Host, 1789)
Male adult

Ukrainian Carpathians,
Transcarpathian Reg., Rakhov Distr., Chornohora range h~1850m, 29.VI-28.VII.2007, Panin R. leg.
male,18 mm.

Copyright © Ruslan Panin

Pterostichus (Calopterus) selmanni prevostii (Dejean, 1828)
Male adult
France, Jura, Lajoux, 1190m, 20/05/2013, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Pterostichus (Calopterus) selmanni roubali Schauberger, 1927
Female adult
CZ; SW Bohemia, Šumava Mts.; 16mm
Copyright © Aleš Sedláček

Pterostichus (Calopterus) pilosus pilosus s.str.
Female adult
CZ; NE Moravia, Štramberk; 15mm
Copyright © Aleš Sedláček

Pterostichus (Calopterus) pilosus pilosus (Host, 1789)
Male adult
W Ukraine, NE Carpathians,, Chornagora Mt Range, h=2000m, alpin zone, 11.VI-1.IX.2015, Panin R. leg 16 mm
Copyright © Ruslan Panin

Pterostichus (Cheporus) burmeisteri Heer, 1838
Male adult

NW Ukrainian Carpathians,
Lviv Prov., Staryi Sambor Distr., near Dobromyl , h=450m 9.VI-25.VII.2007 Panin R. leg.
male,15 mm.

Copyright © Ruslan Panin

Pterostichus (Cheporus) burmeisteri burmeisteri
Female adult
Suisse-Valais-Vissoie, 1700m-17.vi.1980-N. Mal Leg-Coll.RBINS, N. Mal
Copyright © Noël Mal

Pterostichus (Cheporus) burmeisteri burmeisteri
Female adult

Copyright © Noël Mal

Pterostichus (Cheporus) burmeisteri baldensis Schaum, 1862
Female adult
NE Italy, Veneto, Mts Lessini, Vallone del Malera h=1500 m., 28-30.V.2019, Gontarenko A. leg./det.Panin R. L=14 mm
Copyright © Ruslan Panin

Pterostichus (Cheporus) dissimilis (A. & G.B.Villa, 1833)
Male adult
Italy, Valcanale - Bergamo
Copyright © Gianni Allegro

Pterostichus balthasari Jedlička, 1937 Lectotype
Male adult
(NMPC) Image from Dorjderem S, Shi H, Liang H., 2020
Copyright © - - -

Pterostichus (Chinapterus) liupanensis Dorjderem, Shi & Liang, 2020 Holotype Paratype
Male adult

Copyright © Hongliang Shi

Pterostichus (Chinapterus) lianhuaensis Dorjderem, Shi & Liang, 2020 Holotype Paratype
Male adult

Copyright © Hongliang Shi

Pterostichus (Chinapterus) przewalskyi Tschitscherine, 1888 Lectotype
Male adult

Copyright © Hongliang Shi

Pterostichus (Chinapterus) przewalskyi Tschitscherine, 1888
Male adult
Sichuan, Zoigê county, 8km W. of Hongxing, wetland, 3236 m, 34.1236N, 102.6545E, under rock, 2016.VII.25, Shi HL lgt. (IZAS)
Copyright © Hongliang Shi

Pterostichus (Chinapterus) przewalskyi Tschitscherine, 1888
Male genitalia
The genus Pterostichus in China III: a brief review of subgenus Chinapterus Berlov (Coleoptera, Carabidae) with descriptions of two new species
Copyright © Hongliang Shi

Pterostichus (Chinapterus) lianhuaensis Dorjderem, Shi & Liang, 2020
Male genitalia

Copyright © Hongliang Shi

Pterostichus (Chinapterus) liupanensis Dorjderem, Shi & Liang, 2020
Male genitalia

Copyright © Hongliang Shi

Pterostichus (Circinatus) bullatus Allegro & Sciaky, 2010
Male adult

Copyright © Gianni Allegro

Pterostichus (Circinatus) dentifer Allegro & Sciaky, 2010 Holotype
Male adult
China, C Sichuan, Dayi distr., / Chadiping env., 1500 m / 5–7.08.1996 / A. Zamotajlov, A. Miroshnikov & / D. Fedorenko. Image from Hongliang Shi & Hongbin Liang, 2015
Copyright © - - -

Pterostichus (Circinatus) bullatus Allegro & Sciaky, 2010
Female adult
Image from Hongliang Shi & Hongbin Liang, 2015
Copyright © - - -

Pterostichus (Circinatus) yan Shi & Liang, 2015 Holotype
Male adult
Image from Hongliang Shi & Hongbin Liang, 2015
Copyright © - - -

Pterostichus (Circinatus) miao Shi & Liang, 2015 Holotype
Male adult
Image from Hongliang Shi & Hongbin Liang, 2015
Copyright © - - -

Pterostichus (Circinatus) maitreya Shi & Liang, 2015 Holotype
Male adult
Image from Hongliang Shi & Hongbin Liang, 2015
Copyright © - - -

Pterostichus (Circinatus) baenningeri Jedlicka, 1931 Holotype
Male adult
Image from Hongliang Shi & Hongbin Liang, 2015
Copyright © - - -

Pterostichus (Circinatus) ailaoicus Shi & Liang, 2015 Holotype
Male adult
Image from Hongliang Shi & Hongbin Liang, 2015
Copyright © - - -

Pterostichus (Circinatus) liciniformis Csiki, 1930: 669
Male adult
Lijiang. Image from Hongliang Shi & Hongbin Liang, 2015
Copyright © - - -

Pterostichus (Circinatus) dimorphus Shi & Liang, 2015 Holotype
Male adult
Image from Hongliang Shi & Hongbin Liang, 2015
Copyright © - - -

Pterostichus (Circinatus) dimorphus Shi & Liang, 2015 Paratype
Female adult
Image from Hongliang Shi & Hongbin Liang, 2015
Copyright © - - -

Pterostichus (Circinatus) wangjiani Shi & Liang, 2015 Holotype
Male adult
Image from Hongliang Shi & Hongbin Liang, 2015
Copyright © - - -

Pterostichus (Circinatus) tumulus Shi & Liang, 2015 Holotype
Male adult
Image from Hongliang Shi & Hongbin Liang, 2015
Copyright © - - -

Pterostichus (Circinatus) subtilissimus Sciaky, 1996 Holotype
Male adult
Image from Hongliang Shi & Hongbin Liang, 2015
Copyright © - - -

Pterostichus (Circinatus) xilingensis Allegro & Sciaky, 2010 Holotype
Male adult
Image from Hongliang Shi & Hongbin Liang, 2015
Copyright © - - -

Pterostichus (Circinatus) pohnerti Jedlicka, 1934 Holotype
Male adult
Image from Hongliang Shi & Hongbin Liang, 2015
Copyright © - - -

Pterostichus (Circinatus) zoiai Sciaky, 1996c: 222 Holotype
Male adult
Image from Hongliang Shi & Hongbin Liang, 2015
Copyright © - - -

Pterostichus (Circinatus) beneshi Sciaky, 1996 Holotype
Male adult
Image from Hongliang Shi & Hongbin Liang, 2015
Copyright © - - -

Pterostichus (Circinatus) yuxiaodongi Shi & Liang, 2015 Holotype
Male adult
Image from Hongliang Shi & Hongbin Liang, 2015
Copyright © - - -

Pterostichus (Circinatus) adelphus Shi & Liang, 2015 Holotype
Male adult
(IZAS)11.1 mm, “CHINA, Sichuan Prov., Meigu / county, Dafengding nat. res., / Hongxi station, mixed forest; / N28.65850 E103.06123"; “2541 m, 2012.VI.13 day, under dead. Image from Hongliang Shi & Hongbin Liang, 2015
Copyright © - - -

Pterostichus (Circinatus) agilis Allegro & Sciaky, 2010 Holotype
Male adult
(CRS), teneral, 11.8 mm, “CHINA S Sichuan / Luojishan 2200–2800 / m 16–25.vii.1996 / leg. S.Kasantsev”. Image from Hongliang Shi & Hongbin Liang, 2015
Copyright © - - -

Pterostichus (Circinatus) camelus Shi & Liang, 2015 Holotype
Male adult
Image from Hongliang Shi & Hongbin Liang, 2015
Copyright © - - -

Pterostichus (Circinatus) cavazzutianus cavazzutianus Paratype
Male adult
Image from Hongliang Shi & Hongbin Liang, 2015
Copyright © - - -

Pterostichus (Circinatus) cavazzutianus mianningensis Shi & Liang, 2015 Holotype
Male adult
Image from Hongliang Shi & Hongbin Liang, 2015
Copyright © - - -

Pterostichus (Circinatus) zhygealu Shi & Liang, 2015 Holotype
Male adult
Image from Hongliang Shi & Hongbin Liang, 2015
Copyright © - - -

Pterostichus (Circinatus) beneshi Sciaky, 1996 Holotype
Male adult
China – W Sichuan, Wolong 150 km NW Chengdu, 9.VII.1994 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Pterostichus (Circinatus) phiaoc Fedorenko, 2019 Paratype
Female adult
N-Vietnam, 40 km W of Cao Bang, Phia Oac Mt., 22°36´25´´N/ 105°52´08´E, h~1650–1700 m, deciduous forest, 3– 11.X.2018, leg. D. Fedorenko
Copyright © Dmitri Fedorenko

Pterostichus (Circinatus) caobang Fedorenko, 2019 Holotype
Female adult
(ZMMU) N-Vietnam, 40 km W of Cao Bang, Phia Oac Mt., 22°36´25´N/ 105°52´08´E, h~1650– 1700 m, deciduous forest, 3–11.X.2018, leg. D. Fedorenko
Copyright © Dmitri Fedorenko

Pterostichus (Circinatus) laocaiensis Fedorenko, 2019 Paratype
Male adult
N Vietnam, Lao Cai Prov., Hoang Lien Son Mt. Ridge, env. Fansipan [=Phahg Xi Pang] Mt, Tram Ton, h=1950–2100 [m], 15–30.VII.2007, leg. D. Fedorenko
Copyright © Dmitri Fedorenko

Pterostichus (Circinatus) batxatensis Fedorenko, 2019 Holotype
Male adult
N-Vietnam, 40 km WNW of Lao Cai, env. Y Ty, Bat Xat Natn. Park, h= 1600–1900 m, 22°37´–37.6´N/ 103°37.3´–38.3´E, 16–21.X.2018, leg. D. Fedorenko
Copyright © Dmitri Fedorenko

Pterostichus (Circinatus) hoanglien Fedorenko, 2019 Paratype
Female adult
N[orthern]-Vietnam, 40 km WNW of Lao Cai, env. Y Ty, Bat Xat Natn. Park, h=1600–1900 m, 22°37´–37.6´N/ 103°37.3´–38.3´E, 16–21.X.2018, leg. D. Fedorenko
Copyright © Dmitri Fedorenko

Pterostichus (Circinatus) phangxipang Fedorenko, 2019 Paratype
Female adult
N[orthern] Vietnam, Lao Cai Prov[ince]., Hoang Lien Son Mt. Ridge, env. Fansipan [=Phahg Xi Pang] Mt, Tram Ton, h=1950–2100 [m], 15–30.VII.2007, leg. D. Fedorenko
Copyright © Dmitri Fedorenko

Pterostichus (Circinatus) expolitus Fedorenko, 2019 Holotype
Female adult
N-Vietnam, 40 km W of Cao Bang, Phia Oac Mt., E[astern]-slope, h~1600–1800 m, 22°36´27´N/ 105°52´0´´E, deciduous forest, 22.V–6.VI.2018, leg. A. Abramov
Copyright © Dmitri Fedorenko

Pterostichus (Circinatus) tonkinensis Straneo, 1980
Male adult
Vietnam, 40 km W of Cao Bang, Phia Oac Mt., 22°36´50´´N 105°52´0´´E, h~1800 m, deciduous bamboo forest, 3–11.X.2018 (D. Fedorenko)
Copyright © Dmitri Fedorenko

Pterostichus (Cophosus) cylindricus (Herbst, 1784)
Female adult
CZ; S Moravia, Břeclav; 20mm
Copyright © Aleš Sedláček

Pterostichus (Cryobius) kaninensis Poppius, 1906
Male adult

Copyright © Kirill Makarov

Pterostichus (Cryobius) nigripalpis Poppius, 1906
Male adult

length: 8,8 mm
Russia, North Yamal, Venuyeuo river, tundra.
29.06.1993
A.Ryabitsev leg

Copyright © Evgeniy Zinoviev

Pterostichus (Cryobius) pinguedineus Eschscholtz, 1823
Male adult

Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada

Copyright © Henri Goulet

Pterostichus (Cryobius) unctulatus Duftschmid, 1812
Male adult
Ukrainian Carpathians, Transcarpathian reg., Rakhov Distr., Chornogora range, h~2000m. Panin R.leg 29.VI-28.VII.2007. L=8mm
Copyright © Ruslan Panin

Pterostichus (Cryobius) ventricosus Eschscholtz, 1823
Male adult

length: 7,6 mm
Russia, Polar Urals, Krasnyi Kamen' mountains, tundra.
11.07.1995
A.Ryabitsev leg

Copyright © Evgeniy Zinoviev

Pterostichus (Cryobius) middendorffi J.R. Sahlberg, 1875: 103
Female adult

Copyright © Kirill Makarov

Pterostichus (Cryobius) pinguedineus Eschscholtz, 1823: 106
Female adult

Copyright © Kirill Makarov

Pterostichus (Cryobius) cf. ventricosus Eschscholtz, 1823
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Cryobius) parasimilis G.E. Ball, 1962: 6
Male adult
det. K. Makarov
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) similis Mannerheim, 1852b: 296
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Cryobius) middendorffi J.R. Sahlberg, 1875: 103
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Cryobius) middendorffi J.R. Sahlberg, 1875: 103
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Cryobius) brevicornis (Kirby, 1837)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Cryobius) quadraticollis Chaudoir, 1846a: 140
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Cryobius) bargusinicus Shilenkov, 2000 Paratype
Female adult
Бурятия, 6.08.1997, Курумканский р-н., верх. р. Шаманки, А. Шаврин leg.
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) tunkinensis Shilenkov, 2000 Paratype
Male adult
З. Хамар-Дабан, верх. реки Малый Зангисан, гора Байшинт-Ула, 2600-2800м, 23.5.1999, Анищенко А. leg.
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) abaxoides abaxoides (Dejean, 1828)
Male adult
Spain, C Pirineos, Pleta Llardana, 2600m, 23.8.2005, Anichtchenko A. leg. & det. ; 10mm
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) pumilio (Dejean, 1828)
Male adult
Spain, Navarra, Sierra de Aralar, 9.5.2006, Altxueta, 1000m, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) kavanaughi Shilenkov, 2000 Paratype
Female adult
Восточный Саян, Тункинские гольцы, верх. р. Кынгарги, 2500м, 18-23. VII.1995, Анищенко А. leg. (cAA) [E Sayan Mts., Tunkinskie goltsy, Kyngarga riv.]; 6.5 мм
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) aralarensis Espanol & Mateu, 1945
Female adult
Spain, Mts, Cantabricos, Soto de Sajambre, Canto Cabronero Mt., 1850m, 12.VIII.2008, Anichtchenko A. leg. & det. 7.5mm
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) pusillus sagittalis Jeannel, 1937a: 18
Female adult
Spain, C Pirineos, Pleta Llardana, 2600 m, 23.8.2005, Anichtchenko A. leg; 5.8 mm
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) ehlersi (Heyden, 1881)
Male adult
Spain, Mt. Cantabricos, Picos de Europa, 2200 m, 3 km NW mirador Fuente De, 22.07.2019, A. Anichtchenko & R. Sciaky leg. 9 mm. det. Anichtchenko A.
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) aralarensis asturicus Jeanne, 1969a: 3
Male adult
Spain, Cantabria, 2 km E Puerto de San Glorio, near Vejo riv., 43.072369, -4.733061, 1300 m., 23.07.2019, A. Anichtchenko & R. Sciaky leg. det. Anichtchenko A. 7.5 mm
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) cantabricus cantabricus (Schaufuss, 1862)
Male adult
Spain, Cantabria, Pico Tres Mares, 2000-2100m, 24.07.2019, A. Anichtchenko & R. Sciaky leg.
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) procerulus (Heyden, 1880)
Male adult
Spain, Peña Ubiña, Montes Cantabricos, 2000 m, 28.6.2009, Anichtchenko A. leg. 8 mm.
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) cantabricus vasconicus Aubry, 1963
Male adult
Spain, Sierra de Urbasa, 900-1000 m, 27.5.2006, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) unctulatus Duftschmid, 1812
Male adult

Copyright © Kirill Makarov

Pterostichus (Cryobius) pusillus Dejean, 1828
Female adult
Hautes-Pyrénées Department, Vallon d'Estaragne, 42°48'15.93"N, 0° 8'1.36"E, 2580 m, July 2021, P. Moret leg. & det.
Copyright © Pierre Moret

Pterostichus (Cryobius) pumilio (Dejean, 1828)
Male adult
Hautes-Pyrénées Department, Vallon d'Estaragne, 42°48'49.70"N, 0° 8'45.07"E, 2130 m, July 2021, P. Moret leg. & det.
Copyright © Pierre Moret

Pterostichus (Cryobius) amoenus (Dejean, 1828)
Male adult
Hautes-Pyrénées Department, Cirque de Troumouse, 42° 42' 48.7" N, 0° 06' 43.0" E, 2370 m, July 2017, P. Moret leg. & det.
Copyright © Pierre Moret

Pterostichus (Cryobius) horvatovichi Kirschenhofer, 1991 Holotype
Male adult
Image from: Korean type specimens of insects deposited in Hungarian Natural History Museum
Copyright © - - -

Pterostichus (Cryobius) negligens negligens Sturm, 1824: 60
Male adult

Copyright © Jörg Gebert

Pterostichus (Cryobius) homalonotus sojot Shilenkov, 2000 Paratype
Female adult
Бурятия, Восточный Саян, окр. влк. Кропоткина 2050м, 10-15.7.98, Aнищенко А. leg. (cAA) 7.8mm [Buryatia, W Sayan mts., volcan Kropotkin vic., Anichtchenko A. leg]
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) cf empetricola (Dejean, 1828)
Female adult
Russia, Kamchatka reg., Esso, 500m, 5.7.2002 (cAA) 6 mm
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) lucidus Motschulsky, 1844
Male adult
Hamar-Daban, Malyi Zangisan riv., Baishint-Ula Mt., 2300-2500m, 21.5.1999, Anichtchenko A. leg. (cAA). det. V. Shilenkov, 1999
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) colasi Jeannel, 1937
Male adult
Spain, Lleida, Vielha, Panta de Senet, Barranco de Salenca. Image from Sedlmeier and Faille 2022: First insights into the phylogeny of Cryobius
Copyright © - - -

Pterostichus (Cryobius) kolymensis Erjiomin, 1998 Holotype Paratype
Male adult
A – holotype (Kolyma Lowland, near Chersky settlement, collected by S. Kolesnikov); B – paratype (Kolyma Lowland, near Krestovka River, collected by S. Kiselev); C – paratype (western Chukotka, near Valkumey settlement, collected by S. Kiselev). Originally, this species was designated as longipennis and deposited under this name, but later it was renamed and described as kolymensis (Erjiomin 1998). Image from Zubrii et al, 2023
Copyright © - - -

Pterostichus (Cryobius) fulvescens (Motschulsky, 1844)
Male adult
Burjatia, W Sajan Mts., Tunkinskie Goltsy, Mondy vill. vic., 1500m, 1-7.7.2001, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) kurosawai Tanaka, 1958: 58
Male adult
S Sakhalin isl., Chehov Mt., 8.7.1988, E. Berlov leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) amaroides Dejean, 1828
Female adult
Andorra, 6.1983, Monzini leg. (cAA). 8 mm
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) cf bryanti (Vandyke, 1951)
Male adult
Alaska, Sitka (cAA). 6 mm; det. Anichtchenko A. 2024
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) apenninus (Dejean, 1831)
Female adult
Italy, Emilia, M. Penna vers. N. (PR), 1400m 24.5.1999 (cAA). 7 mm
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) unctulatus (Duftschmid, 1812)
Female adult
Lombardia, Val Trompia, M. Colombine 20.VI. (cAA)
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) shilenkovi Erjiomin & Kabak, 1991 Paratype
Male adult
«В Казахстан, Ю Алтай, гребень хр. Азутау, 30 км СВ Алексеевки, 2100 м, у снега, 26.07.1996, И. Кабак leg.» (~48°37ʹ17ʺN / 85°46ʹ03ʺE, 2165 м), “Paratypus Pterostichus (Cryobius) shilenkovi sp. n., 1991 Erjomin et Kabak”
Copyright © Kirill Makarov

Pterostichus (Cryobius) shilenkovi Erjiomin & Kabak, 1991: 139 Paratype
Male genitalia
Aedeagus and gonocoxites
Copyright © Kirill Makarov

Pterostichus (Cryobius) kuraicus Shilenkov, 2000
Male genitalia
4–5 – endophallus of the specimen from the Keshatau Mt., lateral view, 6 – same, dorsal view; 7–14, 18–19 – P. kuraicus: 7–10, 18 – specimens from the left bank of the Kanas Lake, 11–14, 19 – specimens from the Keshatau Mt.
Copyright © Kirill Makarov

Pterostichus (Cryobius) kuraicus Shilenkov, 2000
Male adult
1 – Xinjiang, Altai Mts, left bank of Kanas Lake, 48°44ʹ13ʺN / 87°04ʹ42ʺE, 2645 m, near snow, 15.07.2019 I.I. Kabak leg.; 2 – Xinjiang, S Altai Mts, Keshtau Mt., ENE Qinghe, 46°46ʹ46ʺN / 90°46ʹ28ʺE, 46°46ʹ16ʺN / 90°46ʹ36ʺE, 3160–3345 m, 12.08.2013, I.I. Kabak leg.
Copyright © Kirill Makarov

Pterostichus (Cylindrocharis) rostratus (Newman, 1838)
Male adult

Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada

Copyright © Henri Goulet

Pterostichus (Eosteropus) aethiops (Panzer, 1796)
Female adult
W-Ukraine, Lviv reg., Zhydachiv distr., near Vybranivka vill., forest, under the bark, Panin R. leg. 12.X.2007 female,14mm
Copyright © Ruslan Panin

Pterostichus (Eosteropus) alacer A. Morawitz, 1862
Male adult

Copyright © Evgeniy Zinoviev

Pterostichus (Eosteropus) mannerheimii Dejean, 1831
Male adult

length: 15,0 mm
Russia, South Yamal, Stchustchya river.
17.07.1995
A.Ryabitsev leg

Copyright © Evgeniy Zinoviev

Pterostichus (Eosteropus) maurusiacus Mannerheim, 1825
Male adult

length: 13,2 mm
Russia, Altai mountains, Chemal distr., Bogatash pass
17.09.1993
Yu.Mikhailov leg., Yu.Dudko det.

Copyright © Evgeniy Zinoviev

Pterostichus (Eosteropus) orientalis Motschulsky, 1844
Male adult

length: 14,2 mm
Russia, Far East, near Ol'ga settl.
1.07.1995
V.Semerikov leg

Copyright © Evgeniy Zinoviev

Pterostichus (Eosteropus) rufitarsis cordatus Letzner, 1842
Female adult
Ukrainian Carpathians, Transcarpathian reg., Rakhov Distr., Chornogora range, h=2000m., Panin R. leg., 29.VI-28.VII.2007 L=12 mm
Copyright © Ruslan Panin

Pterostichus (Eosteropus) tuberculiger Tschitscherine, 1897
Male adult

length: 14,1 mm
Russia, Far East, Khabarovsk area, Komsomol'skyi reservation
9.06.1988
Yu.I.Korobeynikov leg.

Copyright © Evgeniy Zinoviev

Pterostichus (Eosteropus) virescens Gebler, 1833
Male adult

length: 11,5 mm
Russia, Altai mountains, Abakskyi mount. range, 2 km NW from Chelyush cordon. h ~ 1840 m.
3.06.1994
Lomakin D. leg.

Copyright © Evgeniy Zinoviev

Pterostichus (Eosteropus) discrepans A.Morawitz, 1862
Female adult

Copyright © Kirill Makarov

Pterostichus (Eosteropus) fuligineus A. Morawitz, 1862b: 325
Female adult

Copyright © Kirill Makarov

Pterostichus (Eosteropus) orientalis antiquus (Motschulsky, 1860)
Male adult
Collection ground Fukushima cho, Hokkaido Pref. 31290
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Eosteropus) orientalis antiquus (Motschulsky, 1860)
Endophallus
Collection ground Fukushima cho, Hokkaido Pref. 31290
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Eosteropus) japonicus (as prolongatus Morawitz, 1862)
Male adult
Collection ground Tomakomai, Hokkaido Pref. 30926
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Eosteropus) japonicus (Motschulsky, 1860)
Endophallus
Collection ground Tomakomai, Hokkaido Pref. 30926
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Eosteropus) rengensis Morita, 2010 Holotype
Male adult
Renge Spa~Mt. Shirouma-dake, Itoigawa-shi, Niigata Pref., Japan, 17. VIII. 1967, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-271
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Eosteropus) rengensis Morita, 2010 Holotype
Male adult
Renge Spa~Mt. Shirouma-dake, Itoigawa-shi, Niigata Pref., Japan, 17. VIII. 1967, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-271
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Eosteropus) noborui Morita, 2008 Holotype
Male adult
Mt. Iide-san, Yamagata Pref., Japan, 23. VII. 1959, Y. Asano
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-262
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Eosteropus) noborui Morita, 2008 Holotype
Male adult
Mt. Iide-san, Yamagata Pref., Japan, 23. VII. 1959, Y. Asano
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-262
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Eosteropus) noborui Morita, 2008 Holotype
Male adult
Mt. Iide-san, Yamagata Pref., Japan, 23. VII. 1959, Y. Asano
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-262
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Eosteropus) noborui Morita, 2008 Holotype
Male adult
Mt. Iide-san, Yamagata Pref., Japan, 23. VII. 1959, Y. Asano
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-262
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Eosteropus) noborui Morita, 2008 Holotype
Male adult
Mt. Iide-san, Yamagata Pref., Japan, 23. VII. 1959, Y. Asano
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-262
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Eosteropus) karasawai Tanaka, 1958f: 218 Holotype
Male adult
Shibunoyu, Mt. Yatsugatake, Nagano Pref., Japan, 31. V. 1956, Y. Karasawa
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-244
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Eosteropus) karasawai Tanaka, 1958f: 218 Holotype
Male adult
Shibunoyu, Mt. Yatsugatake, Nagano Pref., Japan, 31. V. 1956, Y. Karasawa
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-244
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Eosteropus) karasawai Tanaka, 1958f: 218 Holotype
Male adult
Shibunoyu, Mt. Yatsugatake, Nagano Pref., Japan, 31. V. 1956, Y. Karasawa
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-244
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Eosteropus) karasawai Tanaka, 1958f: 218 Holotype
Male adult
Shibunoyu, Mt. Yatsugatake, Nagano Pref., Japan, 31. V. 1956, Y. Karasawa
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-244
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Eosteropus) karasawai Tanaka, 1958f: 218 Holotype
Male adult
Shibunoyu, Mt. Yatsugatake, Nagano Pref., Japan, 31. V. 1956, Y. Karasawa
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-244
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Eosteropus) mannerheimii (Dejean, 1831)
Male adult
Chelyabinskaya obl., Katav-Ivanovskiy prov., mountain Maliy Iremal, alt. 1180 m. 19/08/2008.
Leg. E.Chibilov
Copyright © Evgeniy Chibilov

Pterostichus (Eosteropus) alacer A. Morawitz, 1862a: 254
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Eosteropus) orientalis (Motschulsky, 1844)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Eosteropus) japonicus (Motschulsky, 1860)
Male adult
Pterostichus (Eosteropus) prolongatus det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Eosteropus) moestus (Say, 1823)
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Pterostichus (Eosteropus) orientalis nigromontanus Lafer & Budilov, 2015 Holotype
Male adult
Россия: Приморский край, Хасанский р-н, заповедник «Кедровая Падь», 27.VIII 2004, В.Н. Кузнецов (с препаратом вывернутого эндофаллуса) (БПИ).
Copyright © Pavel Budilov

Pterostichus (Eosteropus) orientalis nigromontanus Lafer & Budilov, 2015 Holotype
Endophallus
Россия: Приморский край, Хасанский р-н, заповедник «Кедровая Падь», 27.VIII 2004, В.Н. Кузнецов
Copyright © Pavel Budilov

Pterostichus (Eosteropus) tuberculiger laferi O. Berlov & E. Berlov, 1996
Male adult

Copyright © Kirill Makarov

Pterostichus (Eosteropus) discrepans A.Morawitz, 1862
Male adult
Russia, Amur area, Skovorodinsk distr., Urka river left bank, 8 km N Yerofey Pavlovich vill., 54*02’24.84N 121*58’30.69E, 552m, 28-29.8. 2014 Shavrin A. leg. det. K. Makarov, 2018. 8.5 mm
Copyright © Alexander Anichtchenko

Pterostichus (Eosteropus) tuberculiger tuberculiger
Male adult
S Primorie, Ussuriisk distr., 3km NE Kaimanovka vill. 22-25.8.2007, K. Makarov A.Zaitsev leg. (DUBC) 13mm
Copyright © Alexander Anichtchenko

Pterostichus (Epinialoe) spiculifer ishidai Tanaka, 1961: 44 Holotype
Male adult
Kitazawatôge Pass, Akaishi Mountain-range, Yamanashi Pref., Japan, 15. VIII. 1956, K. Fujii
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-245
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Epinialoe) spiculifer ishidai Tanaka, 1961: 44 Holotype
Male adult
Kitazawatôge Pass, Akaishi Mountain-range, Yamanashi Pref., Japan, 15. VIII. 1956, K. Fujii
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-245
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Epinialoe) spiculifer ishidai Tanaka, 1961: 44 Holotype
Male adult
Kitazawatôge Pass, Akaishi Mountain-range, Yamanashi Pref., Japan, 15. VIII. 1956, K. Fujii
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-245
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Epinialoe) spiculifer ishidai Tanaka, 1961: 44 Holotype
Male adult
Kitazawatôge Pass, Akaishi Mountain-range, Yamanashi Pref., Japan, 15. VIII. 1956, K. Fujii
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-245
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Epinialoe) spiculifer ishidai Tanaka, 1961: 44 Holotype
Male adult
Kitazawatôge Pass, Akaishi Mountain-range, Yamanashi Pref., Japan, 15. VIII. 1956, K. Fujii
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-245
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Epinialoe) spiculifer ikutanii Tanaka, 1961: 45 Holotype
Male adult
Mt. Fuji, Yamanashi Pref., Japan, 9. VI. 1957, Y. Karasawa
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-246
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Epinialoe) spiculifer ikutanii Tanaka, 1961: 45 Holotype
Male adult
Mt. Fuji, Yamanashi Pref., Japan, 9. VI. 1957, Y. Karasawa
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-246
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Epinialoe) spiculifer ikutanii Tanaka, 1961: 45 Holotype
Male adult
Mt. Fuji, Yamanashi Pref., Japan, 9. VI. 1957, Y. Karasawa
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-246
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Epinialoe) spiculifer ikutanii Tanaka, 1961: 45 Holotype
Male adult
Mt. Fuji, Yamanashi Pref., Japan, 9. VI. 1957, Y. Karasawa
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-246
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Epinialoe) spiculifer ikutanii Tanaka, 1961: 45 Holotype
Male adult
Mt. Fuji, Yamanashi Pref., Japan, 9. VI. 1957, Y. Karasawa
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-246
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Ethira) pilifer (Bates, 1878)
Male adult
Natural History Museum (2014). Dataset: Collection specimens. Resource: Specimens. Natural History Museum Data Portal (data.nhm.ac.uk). © The Trustees of the Natural History Museum, London
Copyright © - - -

Pterostichus (Falsargutor) pseudopedius pseudopedius Reitter, 1887
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Falsargutor) pseudopedius pseudopedius Reitter, 1887e: 247
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Feronidius) ? hungaricus (Dejean, 1828)
Male+Female adult
W. UKRAINE, Transcarpathian region, Mukachevo Distr., riverine forest near Znyatsevo vill., 16-31.VII.2008, Panin R. leg., female, 16 mm
Copyright © Ruslan Panin

Pterostichus (Feronidius) melas (Creutzer, 1799)
Male adult

W-Ukraine,
Lviv Region, Zolochiv distr., near Gologirky, in steppe, 16-29.V.2007 Panin R. leg.
long-17mm

Copyright © Ruslan Panin

Pterostichus (Feronidius) melas melas Creutzer, 1799
Male adult

length: 16 mm
South Russia, Stavropol' area, Pyatigorsk town
29.12.2009
Pupysheva N.leg.
From the collection of A.Parkhachev (Ekaterinburg)

Copyright © Evgeniy Zinoviev

Pterostichus (Feronidius) incommodus Schaum, 1858
Female adult
CZ, S Moravia, Bílé Karpaty; 16mm
Copyright © Aleš Sedláček

Pterostichus (Feronidius) fornicatus (Kolenati, 1845)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Feronidius) hungaricus (Dejean, 1828)
Male adult
Romania, Mts Trascau, Rimetea env., Mt Piatra Secuiului h~1100 m, 27.V-11.VIII.2008, I.Retezár & B.Tallosi leg. L=15 mm
Copyright © Ruslan Panin

Pterostichus (Feronidius) melas italicus Dejean, 1828: 274
Male adult

Copyright © Alberto Narro

Pterostichus (Feronidius) melas italicus Dejean, 1828: 274
Male genitalia

Copyright © Alberto Narro

Pterostichus (Gutta) kongshuhensis Gueorguiev, 2015 Holotype
Male adult
CHINA, Yunnan Prov., Tengchong County, vil. Kongshuhe near cave Wu Shi Shan 1-2-3 N25.73111 E98.66459, ca. 2500-2600m, 11.11.2013, B. Petrov leg.
Copyright © Borislav Gueorguiev

Pterostichus (Haplomaseus) caucasicus (Menetries, 1832)
Male adult
Russia, NW-Caucasus, Karacaj-Cherkessija, Skalisty Mnt. Range, Site Banditskoe h=1400m, 2-3.V.2009, Fominykh D., Bondarenko A.S. leg. 13 mm
Copyright © Ruslan Panin

Pterostichus (Haplomaseus) caucasicus (Menetries, 1832)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Huaius) hanwang Tian & He, 2020 Holotype
Male adult
cave Hanwang Dong, Zhongbai, Zengjia, Chaotian, Guangyuan, Sichuan (四川省广元市朝天区曾家镇中柏村汉王洞), 32°34′40.14″N, 106°6′25.32″E, 1210 m, IX-22-2018, leg. Li He (SCAU). Image from Tian & He, 2020
Copyright © Mingyi Tian

Pterostichus (Hypherpes) tristis (Dejean, 1828)
Male adult

Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada

Copyright © Henri Goulet

Pterostichus (Hypherpes) annosus Casey, 1913 Holotype
Male adult
«California» USNM [# 47030]
Copyright © Kipling W. Will

Pterostichus (Hypherpes) arcanus Casey, 1913 Lectotype
Male adult
«S[an]ta Cruz M[oun]t[ain]s, California», in USNM [# 46985].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) baldwini (Casey, 1924) Lectotype
Male adult
«Baldwin Lake (8600 ft.) [San Bernardino County], California». in USNM [# 47035].
Copyright © Kipling W. Will

Pterostichus breviusculus Casey, 1913 Lectotype
Male adult
«Anderson Valley, Mendocino Co[unty], California» , in USNM [# 46992].
Copyright © Kipling W. Will

Pterostichus bucolicus Casey, 1913 Paralectotypes
Male adult
«Humboldt Co[unty], California»
Copyright © Kipling W. Will

Pterostichus (Hypherpes) canallatus Casey, 1913 Lectotype
Male adult
«S[an]ta Barbara [Santa Barbara County], California» , in USNM [# 47026].
Copyright © Kipling W. Will

Pterostichus cuneatulus Casey, 1913 Lectotype
Male adult
«Humboldt Co[unty], California» , in USNM [# 46998].
Copyright © Kipling W. Will

Pterostichus diabolus Casey, 1913 Lectotype
Male adult
«M[oun]t Diablo [Contra Costa County], California» , in USNM [# 47036].
Copyright © Kipling W. Will

Hypherpes alamedae Casey, 1918 Lectotype
Female adult
«Alameda [Alameda County], California» . Lectotype (♀), in USNM [# 47013].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) esuriens Casey, 1913 Lectotype
Male adult
«Hermitage and Guallala, Mendocino Co[unty], California» , in USNM [# 47033].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) gliscans Casey, 1913 Lectotype
Male adult
«San Clemente Island [Los Angeles County], California» , in USNM [# 47025].
Copyright © Kipling W. Will

Pterostichus gregalis Casey, 1913 Lectotype
Male adult
«Humboldt Co[unty], California», in USNM [# 47027].
Copyright © Kipling W. Will

Pterostichus humboldti Casey, 1913 Lectotype
Male adult
«Humboldt Co[unty], California» (original citation), in USNM [# 47012].
Copyright © Kipling W. Will

Hypherpes intectus Casey, 1918 Lectotype
Male adult
«Boulder Co[unty], Colorado» in USNM [# 47006].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) jacobinus Casey, 1913 Lectotype
Female adult
«San Diego [San Diego County], California» , in USNM [# 47024].
Copyright © Kipling W. Will

Hypherpes kansanus Casey, 1918 Lectotype
Male adult
«Kansas (probably western)» , in USNM [# 46988].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) laborans Casey, 1913 Lectotype
Male adult
«Lake Tahoe [Placer County], California» , in USNM [# 47020].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) lacertus Casey, 1913 Lectotype
Female adult
«Hydesville, Valley of Eel River, Humboldt Co[unty], California» , in USNM [# 47038].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) lassulus (Casey, 1920) Lectotype
Female adult
«Olancha, Inyo Co[unty], California», in USNM [# 46999].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) mercedianus (Casey, 1918) Lectotype
Male adult
«Merced Creek [Mariposa County], California» in USNM [# 47031].
Copyright © Kipling W. Will

Pterostichus breviusculus mimus Casey, 1913 Lectotype
Female adult
«Sonoma Co[unty], California» , in USNM [# 46990].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) miscellus Casey, 1913 Lectotype
Male adult
«S[an]ta Barbara [Santa Barbara County], California» , in USNM [# 47019].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) obsidianus Casey, 1913 Lectotype
Male adult
«Monterey [Monterey County], California» , in USNM [# 46982].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) occultus Casey, 1913 Lectotype
Female adult
«Siskiyou Co[unty], California» , in USNM [# 47009].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) ordinarius Casey, 1913 Lectotype
Male adult
«North Fork, Madera Co[unty], California» , in USNM [# 47021].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) ovalipennis Casey, 1913 Lectotype
Female adult
«between Indio and Colton [probably in Riverside County], California» , in USNM [# 47023].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) panticulatus Casey, 1913 Holotype
Female adult
«with scarcely a doubt from California» in USNM [# 47034].
Copyright © Kipling W. Will

Pterostichus parens Casey, 1913 Lectotype
Male adult
«Siskiyou Co[unty], Cal[ifornia]», in USNM [# 47008].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) pergracilis (Casey, 1920) Lectotype
Female adult
«Olancha, Inyo Co[unty], California» , in USNM [# 47032].
Copyright © Alexander Anichtchenko

Hypherpes placerensis Casey, 1918 Lectotype
Male adult
«Placer Co[unty], California» , in USNM [# 47015].
Copyright © Kipling W. Will

Pterostichus plutonicus Casey, 1913 Lectotype
Male adult
«coast regions north of San Francisco, California» , in USNM [# 46991].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) protensiformis (Casey, 1924) Lectotype
Male adult
«Berkeley [Alameda County], California» , in USNM [# 47016].
Copyright © Kipling W. Will

Hypherpes protensipennis Casey, 1918 Lectotype
Male adult
«Berkeley [Alameda County], Cal[ifornia]» , in USNM [# 47029].
Copyright © Kipling W. Will

Pterostichus pugetanus Casey, 1913 Lectotype
Female adult
«Clackamas Co[unty], Oregon», in USNM [# 46995].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) restrictus (Casey, 1918) Lectotype
Male adult
«Boulder Co[unty], Col[orado]», in USNM [# 47001].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) sierranus Casey, 1913 Lectotype
Female adult
«Lake Tahoe and Merced Cr[eek], California» , in USNM [# 47028].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) sponsor Casey, 1913 Lectotype
Female adult
«Monterey [Monterey County], California» , in USNM [# 47037].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) suffusus Casey, 1913 Lectotype
Male adult
«Humboldt Co[unty], California», in USNM [# 46993].
Copyright © Kipling W. Will

Pterostichus tahoensis Casey, 1913 Lectotype
Male adult
«Lake Tahoe [Placer County], California» , in USNM [# 47011].
Copyright © Kipling W. Will

Pterostichus zunianus Casey, 1913 Lectotype
Male adult
«New Mexico» , in USNM [# 47005].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) crenicollis Leconte, 1873
Male adult

Copyright © Kipling W. Will

Pterostichus (Hypherpes) californicus (Dejean, 1828)
Male adult

Copyright © Kipling W. Will

Pterostichus (Hypherpes) lama (Menetries, 1843)
Female adult
USA, WA, Clark Po near Vancouver Lake forest, 2.VI.2012. 25 mm
Copyright © Alexander Anichtchenko

Pterostichus (Hypherpes) lama (Menetries, 1843)
Male adult
USA: Washington, 34 mm
Copyright © Igor Solodovnikov

Pterostichus (Hypherpes) lama (Menetries, 1843)
Female adult
USA: Washington
Copyright © Igor Solodovnikov

Pterostichus (Hypherpes) protensiformis (Casey, 1924)
Male adult

Copyright © Kipling W. Will

Pterostichus (Hypherpes) vicinus Mannerheim, 1843
Male adult
slightly teneral
Copyright © Kipling W. Will

Pterostichus (Hypherpes) serripes (Leconte, 1875)
Male adult

Copyright © Kipling W. Will

Pterostichus (Hypherpes) craterensis (Hatch, 1949)
Male adult

Copyright © Kipling W. Will

Pterostichus (Hypherpes) tarsalis Leconte, 1873
Male adult

Copyright © Kipling W. Will

Pterostichus (Natalianoe) microps Heyden, 1887
Female adult
Russia, Far East, Primorskii Krai, Skotovskii reg., Anisimovka 17-26.VI.2008 A.Napolov (cAA) 19 mm
Copyright © Alexander Anichtchenko

Pterostichus (Koreonialoe) microps Heyden, 1887
Male adult

Copyright © Kirill Makarov

Pterostichus (Koreonialoe) micropoides Yin, Shi & Liang, 2021 Holotype Paratype
Male+Female adult
Paratype from Hunchun city, Jilin
Copyright © - - -

Pterostichus (Koreonialoe) syleus Kirschenhofer, 1997
Male adult
Fengcheng city, Liaoning. Image from Yin, Shi & Liang, 2021
Copyright © - - -

Pterostichus (Koreonialoe) tetralobatus Yin, Shi & Liang, 2021 Holotype
Male adult
Image from Yin, Shi & Liang, 2021
Copyright © - - -

Pterostichus (Koreonialoe) bellatrix (Tschitscherine, 1895)
Male adult
Changbai county, Jilin. Image from Yin, Shi & Liang, 2021
Copyright © - - -

Pterostichus (Koreonialoe) ishikawai Nemoto, 1988
Female adult
Sud Korea, Juwangsan National Park
Copyright © Vittorino Monzini

Pterostichus (Galapterus) galae Farkac & Plutenko, 1996
Male adult

Far East, S Sikhote-Alin, Snezhnaya Mt., source of Ussuri Riv., h = 1400 -1682 m. 1-6.07.2004 Yu.Sundukov leg.
male
L=13,0 mm

Copyright © Maksim Smirnov

Pterostichus (Lenapterus) agonus G.Horn, 1880
Male adult

length: 11,9 mm
Russia, Chukotka, Belyaka sandbar
13.07.1987
P.S.Tomkovich leg., Yu.I.Korobeynikov det.

Copyright © Evgeniy Zinoviev

Pterostichus (Lenapterus) agonus G.Horn, 1880
Male adult

length: 10,7 mm.
Russia, North part of Gydan peninsula, arctic tundra near Leskino settl. (the beach of Enissey Bay). 07.07.
N.G.Erokhin leg.

Copyright © Evgeniy Zinoviev

Pterostichus (Lenapterus) cancellatus (Motschulsky, 1860)
Male adult

length: 11,9 mm
Russia, Kolyma-Indigirka lowland, low reaches of Kolyma river, Kon'kovaya river.
15.07.1982
Khlebosolov E.I. leg

Copyright © Evgeniy Zinoviev

Pterostichus (Lenapterus) costatus (Menetries, 1851)
Male adult

length: 11,8 mm
Russia, North Yamal, Venuyeuo river, tundra.
25.06.1990
A.Ryabitsev leg

Copyright © Evgeniy Zinoviev

Pterostichus (Lenapterus) punctatissimus (Randall, 1838)
Male adult

Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada

Copyright © Henri Goulet

Pterostichus (Lenapterus) vermiculosus (Menetries, 1851)
Male adult

length: 13,4 mm
South Yamal, middle reaches of Yada-Yakhodyyakha river, riverbank
19.07.1996
Zinoviev E.leg

Copyright © Evgeniy Zinoviev

Pterostichus (Lenapterus) vermiculosus (Menetries, 1851)
Male adult

Copyright © Kirill Makarov

Pterostichus (Lenapterus) agonus G.Horn, 1880
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Lenapterus) subrugosus Straneo, 1955
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Lenapterus) agonus G.Horn, 1880
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Lenapterus) rugosipennis Jedlicka, 1932
Female adult

Copyright © Gianni Allegro

Pterostichus (Leptoferonia) pumilus Casey, 1913
Female adult
Oregon, Portland

Copyright © Igor Sokolov

Pterostichus (Leptoferonia) hackerae Will, 2016 Holotype
Male adult

Copyright © Kipling W. Will

Pterostichus (Leptoferonia) angustus (Dejean, 1828)
Female adult

Copyright © Kipling W. Will

Pterostichus (Leptoferonia) falli Vandyke, 1925
Female adult

Copyright © Kipling W. Will

Pterostichus (Lianoe) dufourii boisgiraudii L. Dufour, 1843
Male adult
France, Pyrenee Ariege, Vall. du Biros, Eylie Cirque de la Plagne, 1100-1500m, 20.6.1993 (DUBC). det. Anichtchenko A. 2021
Copyright © Alexander Anichtchenko

Pterostichus (Lianoe) arribasi Ortuño, 1991
Male adult
Ezcaray, 16.8.97, Valdezcaray, La Rioja, Spain, M. Toribio leg. & det. (cAA). 15 mm
Copyright © Alexander Anichtchenko

Pterostichus (Licentius) cuii Tian & He, 2021 Holotype
Male adult
cave Tangren Dong, Beijiao Cun, Fozizhuang Xiang, Fangshan Qu, Beijing, 39° 47ʹ42ʺN 115°51ʹ55ʺE, 324 m, 20.XII.2020, leg. Qingwu Cui (SCAU).
Copyright © Mingyi Tian

Pterostichus (Licentius) mirabilis Jedlicka, 1939 Holotype
Male adult
Courtesy of Mr. Sunbin Huang
Copyright © - - -

Pterostichus (Lyropedius) lyroderus (Chaudoir, 1846)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Lyrothorax) yoritomus Bates, 1873
Male adult
Collection ground Maizuru city, Kyoto 33744
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Lyrothorax) yoritomus Bates, 1873
Endophallus
Collection ground Maizuru city, Kyoto 33744
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Lyrothorax) fujitai Tanaka & Ishida, 1972 Holotype
Male adult
Mt. Ohdaigahara (Kimmeisui), Nara Pref., Japan, 21. VII. 1953, H. Ishida
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-243
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Lyrothorax) fujitai Tanaka & Ishida, 1972 Holotype
Male adult
Mt. Ohdaigahara (Kimmeisui), Nara Pref., Japan, 21. VII. 1953, H. Ishida
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-243
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Lyrothorax) fujitai Tanaka & Ishida, 1972 Holotype
Male adult
Mt. Ohdaigahara (Kimmeisui), Nara Pref., Japan, 21. VII. 1953, H. Ishida
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-243
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Lyrothorax) fujitai Tanaka & Ishida, 1972 Holotype
Male adult
Mt. Ohdaigahara (Kimmeisui), Nara Pref., Japan, 21. VII. 1953, H. Ishida
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-243
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Lyrothorax) amagisanus Tanaka & Ishida, 1972 Holotype
Male adult
Mt. Amagi-Manjirô, Shizuoka Pref., Japan, 29. VI. 1964, K. Tanaka
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-242
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Lyrothorax) amagisanus Tanaka & Ishida, 1972 Holotype
Male adult
Mt. Amagi-Manjirô, Shizuoka Pref., Japan, 29. VI. 1964, K. Tanaka
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-242
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Lyrothorax) amagisanus Tanaka & Ishida, 1972 Holotype
Male adult
Mt. Amagi-Manjirô, Shizuoka Pref., Japan, 29. VI. 1964, K. Tanaka
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-242
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Lyrothorax) amagisanus Tanaka & Ishida, 1972 Holotype
Male adult
Mt. Amagi-Manjirô, Shizuoka Pref., Japan, 29. VI. 1964, K. Tanaka
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-242
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Lyrothorax) caspius (Menetries, 1832)
Male adult
CZ, SE Moravia, Břeclav; 11mm
(introduced)
Copyright © Aleš Sedláček

Pterostichus (Lyrothorax) fritzei Muilwijk, Kataev & Wrase, 2024 Holotype
Male adult
“Iran Naharkhoran / südl. Gorgan 450–/1000m 14-8-1967 Heinz // HOLOTYPE / Pterostichus (Lyrothorax) / fritzei sp. n. Muilwijk, / Kataev & Wrase des. 2024”
Copyright © Jan Muilwijk

Pterostichus (Lyrothorax) vagus Muilwijk, Kataev & Wrase, 2024 Holotype
Male adult
“IR–Gorgan, Shaskola / Fagus orientalis primeval / 36.77083 N / 5438922 E / 17.7.2017 pitfall / leg. Jahed Razi // HOLOTYPE / Pterostichus (Lyrothorax) vagus sp. n. / Muilwijk, Kataev & Wrase des. 2024”
Copyright © Jan Muilwijk

Pterostichus (Lyrothorax) caspius (Menetries, 1832)
Male adult
Talysh
Copyright © Jan Muilwijk

Pterostichus (Incertae) zhejiangensis Kirschenhofer, 1997 Holotype
Male adult

Copyright © Kipling W. Will

Pterostichus (Melanius) castor Goulet & Bousquet, 1983
Male adult

Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada

Copyright © Henri Goulet

Pterostichus (Melanius) corvinus (Dejean, 1828)
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Pterostichus (Melanius) aterrimus aterrimus
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Melanius) aterrimus aterrimus (Herbst, 1784)
Male adult
W Ukraine, Lviv, Holosko, bank of Natural lake, 24.V-8.VII.2016, Panin R. leg. L=15 mm.
Copyright © Ruslan Panin

Pterostichus (Metallophilus) interruptus (Dejean, 1828)
Male adult
Russia, Primorskii reg, near Vladivostok, Okeanskaya, 29.VII.2000, R. & L. Taliak leg.
Copyright © Denis Potanin

Pterostichus (Metallophilus) interruptus (Dejean, 1828)
Male adult
Russia, Far East, Khabarovsk reg., Bolshoi Khehzir range, Chirki vill., 22.VII.1992, K.E.Dowgailo leg. det.I.A. Solodovnikov. 18mm
Copyright © Ruslan Panin

Pterostichus (Metallophilus) orion (Tschitscherine, 1901)
Male adult

NE Chita reg., Kodar Mt rng., Chara riv.
15 km N Leprindo, 2200m 5-15.7.2001 Brinev leg.
14 mm

Copyright © Alexander Anichtchenko

Pterostichus (Metallophilus) rugosus (Gebler, 1823)
Female adult
Russia, Far East, Khabarovsk reg., Verhnebureinskij distr., Bereinskij Nat. Res., Kordon Strelka, 23-25.VII.2006, leg. A.Barsevskis (DUBC) 15 mm
Copyright © Alexander Anichtchenko

Pterostichus (Metallophilus) mirus (Tschitscherine, 1894)
Female adult
Russia, Siberia, SW Tuva, pass Barlyk, 2100m, 10.VII.2010
14 mm
Copyright © Alexander Anichtchenko

Pterostichus (Metallophilus) orion (Tschitscherine, 1901)
Female adult

Copyright © Kirill Makarov

Pterostichus (Metallophilus) interruptus (Dejean, 1828)
Male adult

Copyright © Kirill Makarov

Pterostichus (Metallophilus) rugosus (Gebler, 1823)
Male adult
Russia, Chita reg., Malye Kovali vill. vic., near Chichatka riv., 22-29.7.1999, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Pterostichus (Metallophilus) rugosus (Gebler, 1823)
Female adult
Russia, Chita reg., Malye Kovali vill. vic., Chichatka riv., 22-29.7.1999, A. Anichtchenko leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Pterostichus (Metallophilus) dandongensis Kirschenhofer, 1997
Male adult
China, Mt Lao-Tu-Ding-Zi 1593m Wuchang, Heilongjiang 1-18. 6.2008 (DUBC). det. Anichtchenko A. 2021
Copyright © Alexander Anichtchenko

Pterostichus (Metallophilus) dandongensis Kirschenhofer, 1997
Female adult
China, Liaoning
Copyright © Alexander Anichtchenko

Pterostichus (Metallophilus) interruptus (Dejean, 1828)
Male adult
Primorskii reg, Shkotovskii distr., Anisimovka vill. (DUBC) det. Anichtchenko A.
Copyright © Alexander Anichtchenko

Pterostichus (Micronialoe) chogyesanus Park; Kwon & Lafer, 1996
Male adult
male
Copyright © Dooyoung Kim

Pterostichus (Micronialoe) bifoveolatus Park; Kwon & Lafer, 1996
Male adult
male
Copyright © Dooyoung Kim

Pterostichus (Micronialoe) spinulosus Kim & Suh, 2024 Holotype
Male adult

Copyright © Dooyoung Kim

Pterostichus (Monoferonia) diligendus (Chaudoir, 1868)
Male adult

Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada

Copyright © Henri Goulet

Pterostichus (Petrophilus) melanarius (Illiger, 1798)
Female adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Pterostichus (Morphnosoma) melanarius (Illiger, 1798)
Female adult
W Ukraine,<br /> Ternopol Reg., Pidvolochisk distr., forest Bogdanivskyi 24.V-15.VI.08 Panin R. leg.<br /> long =19 mm
Copyright © Ruslan Panin

Pterostichus (Feroperis) vladivostokensis Lafer, 1979 : 13
Male adult

Copyright © Kirill Makarov

Pterostichus (Feroperis) procax A. Morawitz, 1862a: 256
Male adult

Copyright © Kirill Makarov

Pterostichus (Morphnosoma) chechcirensis Lafer, 1979: 28 Paratype
Male adult
Habarovsk krai, Hehcir Mt rng., Ryabinin leg.
Copyright © Pavel Budilov

Pterostichus (Morphnosoma) chechcirensis Lafer, 1979: 28 Paratype
Endophallus

Copyright © Pavel Budilov

Pterostichus (Morphnosoma) cf shingarevi shingarevi
Female adult
Russia, Primoskii reg., Spassk-Dalnii distr., 6 km E Evseevka vill. 7-9.8.1999, Anichtchenko A. leg. (cAA). det. O. Berlov; 11 mm
Copyright © Alexander Anichtchenko

Pterostichus (Morphnosoma) petulans Jedlicka, 1938
Male adult
Russia, Primorskii krai, Shkotovskii distr., Anisimovka vill., 17-26.6.2008 (DUBC). 15 mm. det. A. Anichtchenko, 2021
Copyright © Alexander Anichtchenko

Pterostichus (Morphnosoma) jungens (Tschitscherine, 1893)
Male adult
Russia, S Primorie, Chernye Mts., valley of Poima riv., 30.7.1999, A. Anichtchenko leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Pterostichus (Morphnosoma) kurentzovi kurentzovi
Male adult
Russia, S Primorie, Lazo distr., Preobrazhenie vic., 12.8.2007, K. Makarov leg. (DUBC)
Copyright © Alexander Anichtchenko

Pterostichus (Morphnosoma) shingarevi maichensis Lafer, 1979
Male adult
Russia, Primorskii reg, Ussuriisk distr., vic. Kaimanovka vill., 9-15.6.2008 (DUBC) det. A. Anichtchenko, 2021
Copyright © Alexander Anichtchenko

Pterostichus (Morphnosoma) acutidens (Fairmaire, 1889) Syntypes
Male adult
(MNHN). Image from Sun et al, 2018
Copyright © Hongliang Shi

Pterostichus (Morphnosoma) melanodes (Chaudoir, 1878) Syntypes
Female adult
(MNHN). Image from Sun et al, 2018
Copyright © Hongliang Shi

Pterostichus (Morphnosoma) rasilis Park & Kwon, 1996
Male adult
Changbai Mountain, Jilin, China (IZAS). Image from Sun et al, 2018
Copyright © Hongliang Shi

Pterostichus (Morphnosoma) silvestris Sun & Shi, 2018 Holotype
Male adult
Image from Sun et al, 2018
Copyright © Hongliang Shi

Pterostichus (Morphnosoma) silvestris Sun & Shi, 2018 Paratype
Endophallus
Image from Sun et al, 2018
Copyright © Hongliang Shi

Pterostichus (Morphnosoma) silvestris Sun & Shi, 2018
Male genitalia
Image from Sun et al, 2018
Copyright © Hongliang Shi

Pterostichus (Morphnosoma) maryseae Sun & Shi, 2018 Paratype
Endophallus
Image from Sun et al, 2018
Copyright © Hongliang Shi

Pterostichus (Morphnosoma) maryseae Sun & Shi, 2018 Holotype
Male adult
Image from Sun et al, 2018
Copyright © Hongliang Shi

Pterostichus (Morphnosoma) maryseae Sun & Shi, 2018 Holotype
Male adult
Image from Sun et al, 2018
Copyright © Hongliang Shi

Pterostichus (Morphohaptoderus) schuelkei Sciaky & Wrase, 1997: 1102 Holotype
Female adult
body length = 10.8 mm, “CHINA (Shaanxi) / Qin Ling Shan / 108.47 E / 33.51 N / Mt. W pass / autoroute km 70,47km. S Xian 2350–2500 m / 7.–19.VII.1996 / Kleinfeld & Schutze”; “COLL.WRASE / BERLIN”; “HOLOTYPUS / Pterostichus / (Circinatus)/ schuelkei sp. n. / Sciaky & Wrase des. 1997” [red label]
Copyright © Hongliang Shi

Pterostichus (Morphohaptoderus) saueri Sciaky, 1994
Male adult
China, Sichuan, Ganzi pref, Daxue Shan 102.00E 30.03N 5km E Kangding, river valley 3000m 20-23.5.1997 leg A. Putz (cAA) 6.5 mm
Copyright © Alexander Anichtchenko

Pterostichus (Morphohaptoderus) glabellus Fedorenko, 2023 Paratype
Female adult
N[orthern] Vietnam, Lao Cai Prov[ince]./ Hoang Lien Son Mt.ridge,/ env. Fansipan [= Phan Xi Pang] Mt, Tram Ton/ h = 1950-2100 [m],/ 15-30./ leg. VII.2007
Copyright © Dmitri Fedorenko

Pterostichus (Morphohaptoderus) unisetosus Fedorenko, 2023 Holotype
Male adult
(ZMMU): N-Vietnam, 40 km WNW of/ Lao Cai, env. Y Ty, Bat Xat/ Nat[io]n.[al] Park, h=1600-1900 m/ 22°37´-37.6´N/ 103°37.3´-38.3´E, 16-21./ leg. D. Fedorenko X.2018
Copyright © Dmitri Fedorenko

Pterostichus (Morphohaptoderus) primitivus Fedorenko, 2023 Holotype
Female adult
N-Vietnam, 40 km WNW of/ Lao Cai, env. Y Ty, Bat Xat/ Nat[io]n.[al] Park, h=1600-1900 m/ 22°37´-37.6´N/ 103°37.3´-38.3´E, 16-21./ leg. D. Fedorenko X.2018
Copyright © Dmitri Fedorenko

Pterostichus (Morphohaptoderus) huashanus Sciaky, 1994
Female adult
7.5 mm (cAA)
Copyright © Alexander Anichtchenko

Pterostichus (Myosodus) rudestriatus (Reitter, 1883)
Female adult
S Russia, NW-Caucasus, Karachay-Cherkessia, Teberda, Alibek Mountain Pass h=2800 m, 23.VII.2010, Solodovnikov I.A leg. <br L=16 mm</p>
Copyright © Ruslan Panin

Pterostichus (Myosodus) aibgensis Starck, 1890
Male adult

Abkhasia,
near Avadhara, Site Pyv h~1850m, 43.490730 40.686433, 1.VI-12.VII.2009, Zamotajlov A.S. leg.
male,16mm

Copyright © Ruslan Panin

Pterostichus (Myosodus) sodalicius Heyden, 1885
Male adult
Kirgizia, Sary-Chelek 29.6.1993 Klimenko A. leg. (cAA) 18 mm
Copyright © Alexander Anichtchenko

Pterostichus (Myosodus) variabilis (Menetries, 1832)
Female adult

S Russia, NW-Caucasus,
Karacaj-Cherkessija, Abishira-Akhuba Mt. Range, h=2650 m alpine zone, upper Kyafar-Agur Riv., 4-9.VIII.2009, Solodovnikov I.A. leg.
female, 16mm

Copyright © Ruslan Panin

Pterostichus (Myosodus) intricatus roubalianus (Lutshnik, 1928)
Female adult
RUSSIA: Krasnodarsky kray, Tuapse, 500 m, 25.V.1995, N. Yu. Pichugin leg
Copyright © Denis Potanin

Pterostichus (Myosodus) intricatus roubalianus (Lutshnik, 1928)
Female adult
S Russia, W Caucasus,
Tuapse distr., Kashina (=Lysaya) Mt. h=1300m., 10.V.2006, Solodovnikov I.A. leg.
Female, 18 mm
Copyright © Ruslan Panin

Pterostichus (Myosodus) aibgensis Starck, 1890
Male adult

Copyright © Gianni Allegro

Pterostichus (Myosodus) avaricus Kryzhanovskij & Abdurakhmanov, 1983
Male adult

Copyright © Gianni Allegro

Pterostichus (Myosodus) batesi (Tschitscherine, 1894)
Male adult

Copyright © Gianni Allegro

Pterostichus (Myosodus) schoenherri Faldermann, 1836
Male adult
SW Georgia, Lesser Caucasus, Adzhar-Imereti (Meskheti) Rng, Zekari Pass env. h~2100-2200m, 26-30.VI.2006, A.Puchkov leg. L=17 mm
Copyright © Ruslan Panin

Pterostichus (Myosodus) zamotajlovi Belousov, 1991
Male adult
Russia, Karachaevo-Cherkessia, N slopes of Sophia Mt., upper of Sophia riv., h = 2700 m, 07-25.VII.2020 leg. A.P. Tatulyan
Copyright © Igor Solodovnikov

Pterostichus (Myosodus) aibgensis adelaidae Reitter, 1899
Male adult

Copyright © Igor Solodovnikov

Pterostichus (Myosodus) aibgensis adelaidae Reitter, 1899
Female adult

Copyright © Igor Solodovnikov

Pterostichus (Myosodus) cf filipjevi (Lutshnik, 1928)
Male adult

Copyright © Kirill Makarov

Pterostichus (Myosodus) sodalicius Heyden, 1885
Female adult
Uzbekistan, Chatkal Mt. rng., Irtash, 41°10'25.7"N 70°16'20.4"E, 5-7.8.2021, 2400m, A. Anichtchenko leg. & det. (cAA). 15mm
Copyright © Alexander Anichtchenko

Pterostichus (Myosodus) aapsorum Belousov, 1991
Male adult
Abkhazia, N part of Bzybean plateau, SW slopes of Kvansha Mt., alpine zone, h = 2340 m, 11-17.VII-2021 leg. B. Dubinin
Copyright © Igor Solodovnikov

Pterostichus (Myosodus) nivicola (Menetries, 1832)
Male adult
Russia: Dagestan, Akhtynsk distr., N slopes of Shalbuzdag Mt., 6,5 km N of Miskindzha vill., alpin zone, h = 3100 m, 41.352463° N, 47.811141°E, 03-05.08.2022 leg. I.A., A.I., S.V. Solodovnikov’s, O.I. Danilova
Copyright © Igor Solodovnikov

Pterostichus (Myosodus) sodalicius Heyden, 1885
Female adult

Copyright © Igor Solodovnikov

Pterostichus (Myosodus) schoenherri Faldermann, 1836
Female adult
Georgia, Lesser Caucasus, Imereti prov., Meskheti Mt Rng, N slope, Zekari Pass h~2150 2220 m alpine meadows 8-9.VII.2021, E.Rujan leg./det. R. Panin
Copyright © Ruslan Panin

Pterostichus (Neohaptoderus) catei Sciaky & Wrase, 1997 Paratype
Male adult

China,Shaanxi, Qin Ling Shan, 110.06E 34.27N Hua Shan Mt., N Valley, 1200-1400m 118 km E Xian, sifted 18-20.08.1995 A.Pütz leg.
10 mm
photo & coll. Anichtchenko A. 2009

Copyright © Alexander Anichtchenko

Pterostichus (Neohaptoderus) austrinus Fedorenko, 2023 Holotype
Male adult
(ZMMU), labelled: N-Vietnam, 40km W of/ Cao Bang, Phia Oac Mt./ 22°36´25N 105°52´08E/ h~1650-1700 m,/ deciduous forest, 3-11.X./ leg. D.Fedorenko 2018
Copyright © Dmitri Fedorenko

Pterostichus (Neohaptoderus) ignavus (Tschitscherine, 1897)
Male adult
(cAA). 9.5 mm
Copyright © Alexander Anichtchenko

Pterostichus (Daisenialoe) fujimurai fujimurai s.str. Holotype
Male adult
Mt. Daisen, Tottori Pref., Japan, 14. VII. 1956, T. Fujimura
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-041
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Daisenialoe) fujimurai fujimurai s.str. Holotype
Male adult
Mt. Daisen, Tottori Pref., Japan, 14. VII. 1956, T. Fujimura
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-041
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Daisenialoe) fujimurai fujimurai s.str. Holotype
Male adult
Mt. Daisen, Tottori Pref., Japan, 14. VII. 1956, T. Fujimura
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-041
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Daisenialoe) fujimurai fujimurai s.str. Holotype
Male adult
Mt. Daisen, Tottori Pref., Japan, 14. VII. 1956, T. Fujimura
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-041
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Daisenialoe) fujimurai fujimurai s.str. (Label) Holotype
Male adult
Mt. Daisen, Tottori Pref., Japan, 14. VII. 1956, T. Fujimura
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-041
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) taoi Kasahara, 1993
Male adult
Collection ground Seto city, Aichi Pref. 37441
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Nialoe) taoi Kasahara, 1993
Male adult
Collection ground Seto city, Aichi Pref. 37441
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Nialoe) musashiensis Kasahara, 1993
Male adult
Collection ground Hinode cho, Kyoto 37142
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Nialoe) musashiensis Kasahara, 1993
Male adult
Collection ground Hinode cho, Kyoto 37142
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Nialoe) watanabei Tanaka, 1960 Holotype
Male adult
Monomitouge, 630 m, Tanzawa Mts., Kanagawa Pref., Japan, 18. V. 1959, Y. Watanabe
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-250
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) watanabei Tanaka, 1960 Holotype
Male adult
Monomitouge, 630 m, Tanzawa Mts., Kanagawa Pref., Japan, 18. V. 1959, Y. Watanabe
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-250
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) watanabei Tanaka, 1960 Holotype
Male adult
Monomitouge, 630 m, Tanzawa Mts., Kanagawa Pref., Japan, 18. V. 1959, Y. Watanabe
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-250
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) watanabei Tanaka, 1960 Holotype
Male adult
Monomitouge, 630 m, Tanzawa Mts., Kanagawa Pref., Japan, 18. V. 1959, Y. Watanabe
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-250
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) watanabei Tanaka, 1960 Holotype
Male adult
Monomitouge, 630 m, Tanzawa Mts., Kanagawa Pref., Japan, 18. V. 1959, Y. Watanabe
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-250
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) sumondakensis Sasakawa, 2005 Holotype
Male adult
Mt. Sumon, 1100 m, Niigata Pref., Japan, 4. IX. 1969, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-270
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) sumondakensis Sasakawa, 2005 Holotype
Male adult
Mt. Sumon, 1100 m, Niigata Pref., Japan, 4. IX. 1969, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-270
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) sumondakensis Sasakawa, 2005 Holotype
Male adult
Mt. Sumon, 1100 m, Niigata Pref., Japan, 4. IX. 1969, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-270
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) sumondakensis Sasakawa, 2005 Holotype
Endophallus
Mt. Sumon, 1100 m, Niigata Pref., Japan, 4. IX. 1969, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-270
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) sumondakensis Sasakawa, 2005 Holotype
Male adult
Mt. Sumon, 1100 m, Niigata Pref., Japan, 4. IX. 1969, K. Baba
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-270
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) rhanis angustistylis Tanaka, 1958 Holotype
Male adult
Mt. Fuji, Yamanashi Pref., Japan, 9. VI. 1957, Y. Karasawa
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-251
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) rhanis angustistylis Tanaka, 1958 Holotype
Male adult
Mt. Fuji, Yamanashi Pref., Japan, 9. VI. 1957, Y. Karasawa
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-251
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) rhanis angustistylis Tanaka, 1958 Holotype
Male adult
Mt. Fuji, Yamanashi Pref., Japan, 9. VI. 1957, Y. Karasawa
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-251
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) rhanis angustistylis Tanaka, 1958 Holotype
Male adult
Mt. Fuji, Yamanashi Pref., Japan, 9. VI. 1957, Y. Karasawa
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-251
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) rhanis angustistylis Tanaka, 1958 Holotype
Male adult
Mt. Fuji, Yamanashi Pref., Japan, 9. VI. 1957, Y. Karasawa
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-251
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) okutamae Tanaka, 1963 Holotype
Male adult
Mt. Mitô, Okutama, Tokyo, Japan, 21. VIII. 1963, K. Tanaka
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-249
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) okutamae Tanaka, 1963 Holotype
Male adult
Mt. Mitô, Okutama, Tokyo, Japan, 21. VIII. 1963, K. Tanaka
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-249
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) okutamae Tanaka, 1963 Holotype
Male adult
Mt. Mitô, Okutama, Tokyo, Japan, 21. VIII. 1963, K. Tanaka
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-249
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) okutamae Tanaka, 1963 Holotype
Male adult
Mt. Mitô, Okutama, Tokyo, Japan, 21. VIII. 1963, K. Tanaka
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-249
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) okutamae Tanaka, 1963 Holotype
Male adult
Mt. Mitô, Okutama, Tokyo, Japan, 21. VIII. 1963, K. Tanaka
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-249
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Daisenialoe) fujimurai fujimurai s.str. as muranishii Tanaka, 1958 Holotype
Male adult
Mt.Daisen, Tottori Pref., Japan, 22. VII. 1956, T. Muranishi
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-247
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Daisenialoe) fujimurai fujimurai s.str. as muranishii Tanaka, 1958 Holotype
Male adult
Mt.Daisen, Tottori Pref., Japan, 22. VII. 1956, T. Muranishi
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-247
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Daisenialoe) fujimurai fujimurai s.str. as muranishii Tanaka, 1958 Holotype
Male adult
Mt.Daisen, Tottori Pref., Japan, 22. VII. 1956, T. Muranishi
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-247
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Daisenialoe) fujimurai fujimurai s.str. as muranishii Tanaka, 1958 Holotype
Male adult
Mt.Daisen, Tottori Pref., Japan, 22. VII. 1956, T. Muranishi
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-247
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Daisenialoe) fujimurai fujimurai s.str. as muranishii Tanaka, 1958 Holotype
Male adult
Mt.Daisen, Tottori Pref., Japan, 22. VII. 1956, T. Muranishi
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-247
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) mitoyamanus Tanaka, 1971 Holotype
Male adult
Mt. Mitô, Okutama, Tokyo, Japan, 3. VII. 1966, K. Tanaka
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-252
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) mitoyamanus Tanaka, 1971 Holotype
Male adult
Mt. Mitô, Okutama, Tokyo, Japan, 3. VII. 1966, K. Tanaka
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-252
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) mitoyamanus Tanaka, 1971 Holotype
Male adult
Mt. Mitô, Okutama, Tokyo, Japan, 3. VII. 1966, K. Tanaka
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-252
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) mitoyamanus Tanaka, 1971 Holotype
Male adult
Mt. Mitô, Okutama, Tokyo, Japan, 3. VII. 1966, K. Tanaka
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-252
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) mitoyamanus Tanaka, 1971 Holotype
Male adult
Mt. Mitô, Okutama, Tokyo, Japan, 3. VII. 1966, K. Tanaka
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-252
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) mitoyamanus Tanaka, 1971 Holotype
Male adult
Mt. Mitô, Okutama, Tokyo, Japan, 3. VII. 1966, K. Tanaka
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-252
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) fujisanus Tanaka & Suga, 1972 Holotype
Male adult
Mt. Fuji (south side), 1400 m, Shizuoka Pref., Japan, 31. VII. 1971, K. Tanaka & S. Suga
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-253
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) fujisanus Tanaka & Suga, 1972 Holotype
Male adult
Mt. Fuji (south side), 1400 m, Shizuoka Pref., Japan, 31. VII. 1971, K. Tanaka & S. Suga
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-253
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) fujisanus Tanaka & Suga, 1972 Holotype
Male adult
Mt. Fuji (south side), 1400 m, Shizuoka Pref., Japan, 31. VII. 1971, K. Tanaka & S. Suga
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-253
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) fujisanus Tanaka & Suga, 1972 Holotype
Male adult
Mt. Fuji (south side), 1400 m, Shizuoka Pref., Japan, 31. VII. 1971, K. Tanaka & S. Suga
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-253
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) fujisanus Tanaka & Suga, 1972 Holotype
Male adult
Mt. Fuji (south side), 1400 m, Shizuoka Pref., Japan, 31. VII. 1971, K. Tanaka & S. Suga
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-253
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) falcispinus Sasakawa, 2005 Holotype
Male adult
Ishikiri Cave, Nakajo, Niigata Pref., Japan, 12. IX. 1972, K. Baba<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-269<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) falcispinus Sasakawa, 2005 Holotype
Male adult
Ishikiri Cave, Nakajo, Niigata Pref., Japan, 12. IX. 1972, K. Baba<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-269<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) falcispinus Sasakawa, 2005 Holotype
Male adult
Ishikiri Cave, Nakajo, Niigata Pref., Japan, 12. IX. 1972, K. Baba<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-269<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) falcispinus Sasakawa, 2005 Holotype
Male adult
Ishikiri Cave, Nakajo, Niigata Pref., Japan, 12. IX. 1972, K. Baba<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-269<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) falcispinus Sasakawa, 2005 Holotype
Male adult
Ishikiri Cave, Nakajo, Niigata Pref., Japan, 12. IX. 1972, K. Baba<br /> Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-269<br /> (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br /> NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) brunneipennis akaishicus Tanaka, 1958 Holotype
Male adult
Kitazawatôge Pass, Akaishi Mountain-range, Yamanashi Pref., Japan, 15. VIII. 1956, K. Fujii
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-248
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) brunneipennis akaishicus Tanaka, 1958 Holotype
Male adult
Kitazawatôge Pass, Akaishi Mountain-range, Yamanashi Pref., Japan, 15. VIII. 1956, K. Fujii
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-248
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) brunneipennis akaishicus Tanaka, 1958 Holotype
Male adult
Kitazawatôge Pass, Akaishi Mountain-range, Yamanashi Pref., Japan, 15. VIII. 1956, K. Fujii
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-248
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) brunneipennis akaishicus Tanaka, 1958 Holotype
Male adult
Kitazawatôge Pass, Akaishi Mountain-range, Yamanashi Pref., Japan, 15. VIII. 1956, K. Fujii
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-248
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) yahikosanus Sasakawa, 2009
Female adult
Mt. Yahikosan. Image from Sasakawa & Itô, 2022
Copyright © - - -

Pterostichus (Oreophilus) jurinei (Panzer, 1803)
Male adult
Ukrainian Carpathians, Transcarpathian reg., Rakhov Distr., Chornogora range, h~2000m. Panin R. leg. L=12mm
Copyright © Ruslan Panin

Pterostichus (Oreophilus) variolatus variolatus Dejean, 1828
Male adult
S Slovenia, Ilirska Bistrica, Snežnik Mts,15-28.VI.2009, Manfred Egger leg.
L=19 mm

Copyright © Ruslan Panin

Pterostichus (Oreophilus) bicolor Aragona, 1830 [nom.cons.]
Female adult
SE France, Maritime Alps, Saint-Martin-Vésubie, h=1700 m, 28.VI.2000, M. Zelsing leg. L=15 mm
Copyright © Ruslan Panin

Pterostichus (Oreophilus) paulinoi vanvolxemi Putzeys, 1874
Male adult
Portugal, Algarve sierra de Monchique, baragem dos pisoes 20-9-83, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Pterostichus (Oreophilus) impressus (Fairmaire & Laboulbene, 1854)
Male adult
France, Alpes Maritimes, St. Martin Vésubie, Val Madone de Fenestre, 1800 m, B. Thumser leg. 13.07.2004
Copyright © Fabian Bötzl

Pterostichus (Oreophilus) cribratus (Dejean, 1828)
Male adult

Copyright © Gianni Allegro

Pterostichus (Oreophilus) impressus (Fairmaire & Laboulbene, 1854)
Male adult
France : Alpes Maritimes, Le Boreon 26.vii.1978 . (Size : 18 mm)
Copyright © Noël Mal

Pterostichus (Oreophilus) multipunctatus (Dejean, 1828)
Female adult

Copyright © Kirill Makarov

Pterostichus (Oreophilus) variolatus carniolicus Ganglbauer, 1891
Female adult
YU-Nevososo Gasperien, Lirig, 1400m-vi.1968-Coll.RBINS,N. Mal
Copyright © Noël Mal

Pterostichus (Oreophilus) externepunctatus externepunctatus
Male adult

Copyright © Noël Mal

Pterostichus (Oreophilus) impressus (Fairmaire & Laboulbene, 1854)
Male adult

Copyright © Noël Mal

Pterostichus (Oreophilus) xatartii (Dejean, 1828)
Male adult
France : Pyrénées or., Ft. du Carcanet-9.x.1971, P. Picard Leg. - Coll.RBINS, N. Mal
Copyright © Noël Mal

Pterostichus (Oreophilus) externepunctatus Dejean, 1828
Female adult
NW-Italy, Piemonte prov.,Cottian Alps, Mt Albergian, 11.VI.1979, S. Dacatra leg., L=15 mm
Copyright © Ruslan Panin

Pterostichus (Oreophilus) duratii A. & G.B.Villa, 1835
Female adult
La Brigue, Alpes-Maritimes, France
Copyright © Jean Michel Lemaire

Pterostichus (Oreophilus) franzi Negre, 1955
Male adult
SPAIN, Cantabria, 3 km SW Fuente De, Cantijon riv., 1400 m, 21.07.2019, A. Anichtchenko & R. Sciaky leg.
Copyright © Alexander Anichtchenko

Pterostichus (Oreophilus) pinguis (Dejean, 1828)
Male adult
Alagna Valsesia (VC)
Copyright © Gianni Allegro

Pterostichus (Oreophilus) flavofemoratus (Dejean, 1828)
Male adult
Valle di Cogne e Valsavarenche (Aosta)
Copyright © Gianni Allegro

Pterostichus (Oreophilus) duratii A. & G.B.Villa, 1835
Male adult
Italy, Passo della Teglia - Imperia
Copyright © Gianni Allegro

Pterostichus (Oreophilus) parnassius Schaum, 1859
Female adult
Italy, Oropa - Biella
Copyright © Gianni Allegro

Pterostichus (Oreophilus) jurinei (Panzer, 1803)
Female adult

Copyright © Kirill Makarov

Pterostichus (Oreophilus) morio carpathicus Kult, 1944
Female adult
CSFR Hohe Tatra, Solisko 1800-2090m, 23.6.1991 (DUBC)
Copyright © Alexander Anichtchenko

Pterostichus (Oreophilus) bicolor Aragona, 1830
Male adult
H. Penna, Senova, 1400m 18.6.1978 (cAA)
Copyright © Alexander Anichtchenko

Pterostichus (Oreophilus) impressus (Fairmaire & Laboulbene, 1854)
Male adult
France, Orvault, Loire-Atlantique, 16.12.1962 [label error] (DUBC) 17 mm
Copyright © Alexander Anichtchenko

Pterostichus (Oreophilus) cantaber (Chaudoir, 1868)
Male adult
Spain, Santander, Loredo, 8.8.1993 (cAA)
Copyright © Alexander Anichtchenko

Pterostichus (Oreoplatysma) belizini Lutshnik, 1933
Male adult

S Russia, Krasnodar Reg.,
Near Sochi , Solokh-Aul h=500m, 5.VIII.2007
I.A.Solodovnikov leg.
L=9mm
photo Panin R.

Copyright © Ruslan Panin

Pterostichus (Oreoplatysma) cecchiniae Jakobson, 1907
Male adult

Russia, Krasnodar reg., 6 km NE Soloniki vill.,
28-30.V.2009 Anichtchenko A. leg.
male, long. 10 mm

Copyright © Alexander Anichtchenko

Pterostichus (Oreoplatysma) koenigi (Reitter, 1887)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Orientostichus) pulcher Sciaky & Allegro, 2013 Holotype Paratype
Male+Female adult
Label of Holotype male: China, S Sichuan, Ebian, Yizhi Yakou, Xiaoliang Shan-Si He, 2800-3100 m, 19.VII.2007, Cavazzuti P.F. leg. (IZAS). Scale bar 5 mm
Copyright © Gianni Allegro

Pterostichus (Orientostichus) distinctissimus Jedlicka, 1940: 13
Male adult
Taiwan: Hualien, Guanyuan, 2300 m, 27.VI.2006, leg. Y.-F. Hsu. (17 mm). det. J. Marrero. Taiwan Agricultural Research Institute (TARI) collection
Copyright © Jose Marrero

Pterostichus (Orientostichus) perlutus Jedlicka, 1938a: 5
Male adult
W Sichuan, pass W Kangding, Zheduo mts, 4200m, 12.7.2013, V. Patrikeev leg. (cAA) 16 mm; det. H. Shi, 2020
Copyright © Alexander Anichtchenko

Pterostichus (Orientostichus) jialini Yin, Zhu & Shi, 2023 Holotype
Male adult
Image from Yin, Zhu & Shi, 2023
Copyright © - - -

Pterostichus (Orientostichus) pemphis Yin, Zhu & Shi, 2023 Holotype
Male adult
Image from Yin, Zhu & Shi, 2023
Copyright © - - -

Pterostichus (Orientostichus) condylus Yin, Zhu & Shi, 2023 Holotype
Male adult
Image from Yin, Zhu & Shi, 2023 Scale bar: 5 mm.
Copyright © - - -

Pterostichus (Orientostichus) liyuani Yin, Zhu & Shi, 2023 Holotype
Male adult
Image from Yin, Zhu & Shi, 2023. Scale bar: 5 mm.
Copyright © - - -

Pterostichus (Orientostichus) leo Yin, Zhu & Shi, 2023 Holotype
Male adult
Image from Yin, Zhu & Shi, 2023
Copyright © - - -

Pterostichus (Orientostichus) orbicollis Yin, Zhu & Shi, 2023 Holotype
Male adult
Image from Yin, Zhu & Shi, 2023
Copyright © - - -

Pterostichus (Orsonjohnsonus) johnsoni Ulke, 1889
Male adult

USA, Oregon, Clackamas County

Copyright © Igor Sokolov

Pterostichus (Paraferonia) lubricus LeConte, 1852
Male adult

Great Smoky Mountains National Park

Copyright © Igor Sokolov

Pterostichus (Paranialoe) biexcisus subparallelus Straneo, 1955
Male adult
Nishinoshima Is., Mita (Mt. Takuhi-yama). Image from Takashi Shimizu and Ryo Nakamura, 2023
Copyright © - - -

Pterostichus (Parapterostichus) grajus (Dejean, 1828)
Male adult
Italia, Oropa (Bi) Lago Mucrone, 14.6.2003 S. Dacatra leg. (cAA) 12 mm
Copyright © Alexander Anichtchenko

Pterostichus urengaicus Jurecek, 1924: 144
Male adult
Russia, S.Ural, Chelyab. reg, Katav-Ivanovsk. prov., Serpievskiy wildlife preserve, Lavki mount. 28.08.2007. Chibilov leg.
Copyright © Sunvas

Pterostichus (Petrophilus) eximius A. Morawitz, 1862
Male adult

length: 12,3 mm
Russia, South Sakhalin, Aniva distr., 6 Km East from Novoaleksandrovka settl.
14.07.1990
Basarukin A. leg.

Copyright © Evgeniy Zinoviev

Pterostichus (Petrophilus) dauricus Gebler, 1832
Male adult

length: 14,2 mm
Russia, Chita area, Nerchinskyi distr., Nercha river valley near Pravye Kumaki village
4.08.1999
Zinoviev E.leg.

Copyright © Evgeniy Zinoviev

Pterostichus (Petrophilus) dilutipes Motschulsky, 1844
Male adult

length: 12,3 mm
West Siberia, KHMAO-Yugra, Nizhnevartovsk distr., Glubokiy Sabun river valley,
riparian coniferous forest
12.07.2004
Belskaya E.A. le

Copyright © Evgeniy Zinoviev

Pterostichus (Petrophilus) elmbergi Poppius, 1908
Male adult

length: 11,7 mm
Russia, Altai mountains, northern part of Tchulyshman plateau; upstreams of Bayaz river, coniferous (siberian pine and fir) forest.
28.06.1993
Lomakin D. leg.

Copyright © Evgeniy Zinoviev

Pterostichus (Petrophilus) eximius A. Morawitz, 1862
Male adult

Length: 13,2 mm
Transbaikalia, Tchita area, Krasnotchikoyskiy distr., Tchkoy river near Nizhniy Naryn village,
Coniferous forest,
21.07.1999
Zinoviev E. leg and det.

Copyright © Evgeniy Zinoviev

Pterostichus (Petrophilus) foveolatus Duftschmid, 1812
Male+Female adult

    Ukrainian Carpathians,
    Transcarpathian reg., Rakhov Distr., Chornogora range, h=1850-2000m. Panin R.leg 29.VI-28.VII.07
    L=15mm

Copyright © Ruslan Panin

Pterostichus (Petrophilus) kokeilii archangelicus Poppius, 1907
Male adult

Length 13,3 mm
Russia, Polar Urals, Krasnyi Kamen' mountain,
11.07.1995
A.Ryabitsev leg

Copyright © Evgeniy Zinoviev

Pterostichus (Petrophilus) magus magus Mannerheim, 1825
Male adult

length: 12,7 mm
Sverdlovsk (Ekaterinburg), coniferous (pine) forest in the sity.
20.05.1981
Zinoviev E. leg

Copyright © Evgeniy Zinoviev

Pterostichus (Petrophilus) magus mongolicus Motschulsky, 1844
Female adult

SE Siberia,
Irkutsk Area, Khamar-Daban Mountains, 6-8.VI.2007, D.Fominyh leg.
female,15mm

Copyright © Evgeniy Zinoviev

Pterostichus (Petrophilus) montanus Motschulsky, 1844
Male adult

KMAO-Yugra, Nizhnevartovsk distr., Glubokiy Sabun River valley
12.07.2004
Belskaya E.A. leg

Copyright © Evgeniy Zinoviev

Pterostichus (Petrophilus) septentrionis Chaudoir, 1868
Male adult
SE Siberia,
Irkutsk Area, Khamar-Daban Mountains, 6-8.VI.2007 D.Fominyh leg.
long=13mm
Copyright © Ruslan Panin

Pterostichus (Petrophilus) seriatus Chaudoir, 1850
Male adult

Length 12,6 mm.
Russia, South-East Altai, Tchikhatchev mount. range, 3 km SW from Thernaya mountain, h=2800 m., mount. tundra. 11.07.1996.
R.Dudko leg. and det.

Copyright © Evgeniy Zinoviev

Pterostichus (Petrophilus) subaeneus Chaudoir, 1850
Male adult
length: 11,1 mm Russia, Altai mountains, northern part of Tchulyshman plateau; Tongzyak mountain subalpine tundra near 28.06.1993, Lomakin D. leg.
Copyright © Evgeniy Zinoviev

Pterostichus (Petrophilus) tomensis (Gebler, 1847)
Male adult

Kemerovo Reg., env. Osinniki, source of Krasnaya Kaltank Riv., Populus tremulae-fores, W slope , h = 450 м 28.V-6.VI.03 Yu.Sundukov leg.
L=18,0 mm

Copyright © Maksim Smirnov

Pterostichus (Petrophilus) triseriatus Gebler, 1847
Male adult
length: 11,8 mm Kemerovskaya distr., Kuznetskiy Alatay Mt. range, Dvuglavaya Mt. h=1600 m 17.07.1995. Yu. Mikhailov leg., R.Dudko det.
Copyright © Evgeniy Zinoviev

Pterostichus (Petrophilus) uralensis Motschulsky, 1850
Male adult

Russia, Middle Urals, Sverdlovskaya area, Kamensk distr, near Maminskoye settl. Coniferous forest.
3.07.2003.
Markov N.I.leg

Copyright © Evgeniy Zinoviev

Pterostichus (Petrophilus) urengaicus Jurecek, 1924
Male adult

length: 13 mm
South Urals, Malyi Iremel' mountain
15.07.1984
Kashevarov B.N. leg

Copyright © Evgeniy Zinoviev

Pterostichus (Petrophilus) coracinus (Newman, 1838)
Male adult
Great Smoky Mountains National Park
photo I. M. Sokolov
Copyright © Igor Sokolov

Pterostichus (Petrophilus) lachrymosus (Newman, 1838)
Female adult
Great Smoky Mountains National Park
photo I. M. Sokolov
Copyright © Igor Sokolov

Pterostichus (Petrophilus) novus Straneo, 1944
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Pterostichus (Petrophilus) stygicus (Say, 1823)
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Pterostichus (Petrophilus) thunbergi A. Morawitz, 1862b: 325
Male adult
Collection ground Makkari, Hokkaido Pref. 29357
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Petrophilus) turanensis Jedlicka, 1959
Male adult
Russia, E Sayan Mts., Tunkinskie Goltsy, Kyngarga riv., 2200 m, 18-23.VII.1995, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Pterostichus (Petrophilus) montanus (Motschulsky, 1844)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Petrophilus) montanus (Motschulsky, 1844)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Petrophilus) kokeilii archangelicus Poppius, 1907
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Petrophilus) seriatus (Chaudoir, 1850)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Petrophilus) tundrae (Tschitscherine, 1894)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Petrophilus) elmbergi Poppius, 1908d: 22
Male adult

Copyright © Kirill Makarov

Pterostichus (Petrophilus) tundrae (Tschitscherine, 1894)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Petrophilus) dilutipes (Motschulsky, 1844)
Male adult

Copyright © Kirill Makarov

Pterostichus (Petrophilus) triseriatus (Gebler, 1847)
Male adult
Tyva, Kyzyl-Taiga mt. rng., 1500m, Ak-Sug riv. 21.6.2006, A. Menshikov leg. (cAA) det. A. Anichtchenko, 2021; 10.5 mm
Copyright © Alexander Anichtchenko

Pterostichus (Petrophilus) subaeneus (Chaudoir, 1850)
Male adult
Russia, E Siberia, Irkutsk reg., Nizhneudinsk distr., 9km SW Telezhnyi vill., 1040m, 28.7.1998, A. Anichtchenko leg & det. (cAA)
Copyright © Alexander Anichtchenko

Pterostichus (Petrophilus) septentrionis (Chaudoir, 1868)
Male adult
E Siberia, Hamar-Daban mts., Osinovka riv., 2000m, 28-31.7.1995, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Pterostichus (Petrophilus) eximius A. Morawitz, 1862
Male adult

Copyright © Kirill Makarov

Pterostichus (Petrophilus) kokeilii archangelicus Poppius, 1907
Male adult

Copyright © Kirill Makarov

Pterostichus (Petrophilus) urengaicus Jurecek, 1924
Male adult

Copyright © Kirill Makarov

Pterostichus (Petrophilus) dauricus (Gebler, 1832)
Female adult
Russia, Chita area, Kyrenskiy distr., valley of Kyra riv., Kyra env., 16.7.2009, Shavrin A. leg (cAA). 14 mm. det. A. Anichtchenko, 2021
Copyright © Alexander Anichtchenko

Pterostichus (Petrophilus) sutschanensis Jedlicka, 1962
Male adult
Russia, S Primorie, Ussuriisk distr., 3 km NE Kaimanovka vill., 22-26.8.2007, K. Makarov leg & det. (DUBC) 10 mm
Copyright © Alexander Anichtchenko

Pterostichus (Petrophilus) kapanovi Kabak, 2021 Holotype
Male adult
China, Xinjiang, Altai Mt. R., N of Kom (= Hemu) village, 48°37ʹ36ʺN / 87°25ʹ40ʺE, H = 3015 m, alpine meadows, 2.08.2015 (I.I. Kabak).
Copyright © Ilya Kabak

Pterostichus (Petrophilus) huguai Kabak, 2021 Holotype
Male adult
China, Xinjiang, Altai Mts, left bank of Kanas Lake, 48°44ʹ35ʺN / 87°06ʹ03ʺE, H = 2710 m, alpine meadows, 15.07.2019 (I.I. Kabak).
Copyright © Ilya Kabak

Pterostichus (Petrophilus) mongoloaltaicus Kabak, 2021 Holotype Paratype
Male+Female adult
China, Xinjiang, S Altai Mts, Keshtau Mt., ENE Qinghe, 46°48ʹ30ʺN / 90°44ʹ05ʺE – 46°46ʹ46ʺN / 90°46ʹ28ʺE, H = 2920–3160 m, 12.08.2013 (I.I. Kabak).
Copyright © Ilya Kabak

Pterostichus (Petrophilus) magoides (Straneo, 1937) Holotype Paratype
Female adult
«Altai Sibir.», «N. 15», «Museo Storia Naturale Milano», «magoides Stran.» (MSNM)
Copyright © Ilya Kabak

Pterostichus (Petrophilus) magoides (Straneo, 1937)
Male adult
Left: S Altai, Markakol Lake vic. Right: E Kazakhstan, SW Altai Mts., Rakhmanovskoe Forestry, Arasan River, W of Rakhmanovskie Klyuchi, 1736 m, 49°31ʹ55ʺN / 86°27ʹ40ʺE, 1.05.2017
Copyright © Ilya Kabak

Pterostichus (Petrophilus) magoides (Straneo, 1937)
Male genitalia
Kazakhstan, Markakol lake
Copyright © Ilya Kabak

Pterostichus (Petrophilus) findelii (Dejean, 1828)
Male adult
Roumanie, Faragasului 02-07-2012
Copyright © Pascal Deschamps

Pterostichus (Phonias) corrusculus Leconte, 1873
Male adult

Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada

Copyright © Henri Goulet

Pterostichus (Phonias) eobius (Tschitscherine, 1899)
Male adult

Copyright © Kirill Makarov

Pterostichus (Phonias) diligens (Sturm, 1824)
Male adult

Copyright © Kirill Makarov

Pterostichus (Phonias) sasajii Morita, 2007
Male adult
Collection ground Chitose city, Hokkaido Pref. 27527
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Phonias) longinquus Bates, 1873
Male adult
Collection ground Inami cho, Hyogo Pref. 39361
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Phonias) longinquus Bates, 1873
Male adult
Collection ground Inami cho, Hyogo Pref. 39361
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Phonias) shavrini O.Berlov & Anistschenko, 1999 Paratype
Male adult

Copyright © Alexander Anichtchenko

Pterostichus (Phonias) kutensis Poppius, 1905
Male adult
Хабаровский край, Верхнебуреинский район, окр. пос. Софийск. Лиственничная тайга, h~1000м, 4.09.2013 (Russia, Habarovsk prov., Verhnebureinsk distr., Sofiisk vill.)
Copyright © Pavel Budilov

Pterostichus (Phonias) kutensis Poppius, 1905
Endophallus
Хабаровский край, Верхнебуреинский район, окр. пос. Софийск. Лиственничная тайга, h~1000м, 4.09.2013 (Habarovsk prov., Verhnebureinsk distr., Sofiisk vill.)
Copyright © Pavel Budilov

Pterostichus (Phonias) longinquus Bates, 1873
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Phonias) strenuus (Panzer, 1796)
Female adult
Russia, Moscow det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Phonias) diligens (Sturm, 1824)
Male adult
Russia, Moskow det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Phonias) diligens (Sturm, 1824)
Female adult
Россия, Песчанное. det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Phonias) kutensis Poppius, 1905
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Phonias) jankowskyi (Tschitscherine, 1897)
Female adult
Russian Far East, Primorskii krai, Khasanskii district, 9 km SSE Barabash, middle course of Kedrovaya River N43°06'24" E131°32'17" h~120m asl 22-30.V.2015 leg. K.Makarov
Copyright © Kirill Makarov

Pterostichus (Phonias) ovoideus (Sturm, 1824)
Female adult
France, 77860, Quincy-Voisins, 17-05-2008
Copyright © Pascal Deschamps

Pterostichus (Phonias) burkhan O.Berlov & Anistschenko, 2005 Paratype
Male adult
Russia, Chita reg., 23km SW Borzia vill. Borzinskoe lake, 1.6.2002, Anichtchenko A. leg. (cAA)
Copyright © Alexander Anichtchenko

Pterostichus (Biphonias) ripensis (Motschulsky, 1866) Lectotype
Male adult
"Kisi // fl. Amur" (ZMUM). Image from Sundukov, 2013
Copyright © - - -

Pterostichus (Biphonias) ripensis (Motschulsky, 1866) Lectotype
Male genitalia
Image from Sundukov, 2013
Copyright © - - -

Pterostichus (Phonias) batdaiensis Fedorenko, 2023 Paratype
Male adult
N-Vietnam, Lao Cai Prov[ince]./ env. Sa Pa/ near Cat Cat river/ 22.33399°N 103.82281°E/ h = 1320 m 20–23.IV./ A. Prosvirov leg. 2013
Copyright © Dmitri Fedorenko

Pterostichus (Phonias) lutschniki Jedlicka, 1962
Male adult
South Primorye, Kedrovaya Pad nature reserve, mouth of Kedrovaya River, swamp 43°04'47"N 131°36'42"E 1.VI.2019 leg. K. Makarov L= 7.1
Copyright © Kirill Makarov

Pterostichus (Phonias) lutschniki Jedlicka, 1962
Male genitalia
South Primorye, Kedrovaya Pad nature reserve, mouth of Kedrovaya River, swamp 43°04'47"N 131°36'42"E 1.VI.2019 leg. K. Makarov L= 7.1
Copyright © Kirill Makarov

Pterostichus (Phonias) lutschniki Jedlicka, 1962
Endophallus
South Primorye, Kedrovaya Pad nature reserve, mouth of Kedrovaya River, swamp 43°04'47"N 131°36'42"E 1.VI.2019 leg. K. Makarov L= 7.1
Copyright © Kirill Makarov

Pterostichus (Phonias) lutschniki Jedlicka, 1962
Male adult
South Primorye, Kedrovaya Pad nature reserve, mouth of Kedrovaya River, swamp 43°04'47"N 131°36'42"E 1.VI.2019 leg. K. Makarov L= 7.1
Copyright © Kirill Makarov

Pterostichus (Phonias) lutschniki Jedlicka, 1962
Female adult
Primorsky Kray, Khasan City, on light 25.V.70 Nazarov Yu.N.
Copyright © Kirill Makarov

Pterostichus (Phonias) lutschniki Jedlicka, 1962
Female adult
Primorsky Kray, Khasan City, on light 25.V.70 Nazarov Yu.N.
Copyright © Kirill Makarov

Pterostichus (Phonias) lutschniki Jedlicka, 1962
Male genitalia
Hirotuka Kanagawa-Ken 23.jul.1958 S.Nomura L= 7.4 mm
Copyright © Kirill Makarov

Pterostichus (Phonias) lutschniki Jedlicka, 1962
Endophallus
Hirotuka Kanagawa-Ken 23.jul.1958 S.Nomura L= 7.4 mm
Copyright © Kirill Makarov

Pterostichus (Phonias) lutschniki Jedlicka, 1962
Female adult
South Primorye, Khasansky district, 7.5 km SE of Khasan 42°24'47"N 130°44'12"E 24-25.V.2022 leg.K.Makarov, A.Matalin L= 7.8 mm
Copyright © Kirill Makarov

Pterostichus (Phonias) lutschniki Jedlicka, 1962
Male adult
South Primorye, Khasansky district, 7.5 km SE of Khasan 42°24'47"N 130°44'12"E 24-25.V.2022 leg.K.Makarov, A.Matalin L= 7.2 mm
Copyright © Kirill Makarov

Pterostichus (Phonias) lutschniki Jedlicka, 1962
Endophallus
South Primorye, Khasansky district, 7.5 km SE of Khasan 42°24'47"N 130°44'12"E 24-25.V.2022 leg.K.Makarov, A.Matalin L= 7.2 mm
Copyright © Kirill Makarov

Pterostichus (Phonias) lutschniki Jedlicka, 1962
Male adult
South Primorye, Khasansky district, 7.5 km SE of Khasan 42°24'47"N 130°44'12"E 24-25.V.2022 leg.K.Makarov, A.Matalin L= 7.6 mm
Copyright © Kirill Makarov

Pterostichus (Phonias) lutschniki Jedlicka, 1962
Male genitalia
South Primorye, Khasansky district, 7.5 km SE of Khasan 42°24'47"N 130°44'12"E 24-25.V.2022 leg.K.Makarov, A.Matalin L= 7.6 mm
Copyright © Kirill Makarov

Pterostichus (Phonias) lutschniki Jedlicka, 1962
Male adult
Primorsky Kray, Khasan City, on light 25.V.70 Nazarov Yu.N.
Copyright © Kirill Makarov

Pterostichus (Phonias) lutschniki Jedlicka, 1962
Male adult
Primorsky Kray, Khasan City, on light 25.V.70 Nazarov Yu.N.
Copyright © Kirill Makarov

Pterostichus (Phonias) lutschniki Jedlicka, 1962
Male genitalia
Primorsky Kray, Khasan City, on light 25.V.70 Nazarov Yu.N.
Copyright © Kirill Makarov

Pterostichus (Phonias) lutschniki Jedlicka, 1962
Endophallus
Primorsky Kray, Khasan City, on light 25.V.70 Nazarov Yu.N.
Copyright © Kirill Makarov

Pterostichus (Phonias) subitus Csiki, 1930 Lectotype
Male+Female adult
Lectotype, left, male: «Samarkand, Reitter»; «velox m. Typ. Tschitscherin det.», «Lectotypus Pterostichus velox Tschitr., design. I. Kabak, 2023». Female, right: Uzbekistan, Lower Amu-Darya Valley,SE of Urgench, Kasilat, SW of Xonqa, 41°25ʹ59.2ʺN / 60°43ʹ05.2ʺE, 102 m, 20.05.2023.
Copyright © Kirill Makarov

Pterostichus (Phonias) subitus Csiki, 1930
Male genitalia

Copyright © Kirill Makarov

Pterostichus (Platypterus) lineatopunctatus L.Miller, 1850
Male adult
Upper Austria, NE-Alps,Windischgarsten, Rettenbach, Mt. Hoher Nock, h=1960 m,15.VI.2012, Manfred Egger leg.

Copyright © Ruslan Panin

Pterostichus (Platypterus) panzeri (Panzer, 1803)
Female adult
Austria, 47.32N 14.30E, path to Sparafeld Mt, 18.6.2017, L. Minkina leg (cAA) 14 mm
Copyright © Alexander Anichtchenko

Pterostichus (Platysma) niger niger s.str.
Male adult

NW Ukrainian Carpathians,
Lviv Prov., Staryi Sambor Distr., near Dobromyl , h=450m, 9.VI-25.VII.2007 Panin R. leg.

Copyright © Ruslan Panin

Pterostichus (Platysma) niger niger s.str.
Male adult

length: 22,8 mm
West Siberia, KHMAO-Yugra, the vicinities of Khanty-Mansiysk town
1.07.2004.
Zinoviev E.leg.

Copyright © Evgeniy Zinoviev

Pterostichus (Platysma) niger niger s.str.
Male adult

Chelyabinsk, 4.03.2011

Chibilov leg.

Copyright © Sunvas

Pterostichus (Platysma) eschscholtzii (Germar, 1824)
Male adult

Copyright © Kirill Makarov

Pterostichus (Platysma) leptis Bates, 1873
Female adult
Collection ground Erimo cho, Hokkaido Pref. 30912
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Platysma) leptis Bates, 1873
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Platysma) planicola (Tschitscherine, 1899)
Female adult
Russia, Dagestan, Makhachkala, Karaman‐2, H = ‐25 m, 43°02'35''N / 47°26'15''E, 20.06.2015, Iljina E.V. leg. Image from Kabak et al, 2020
Copyright © Ilya Kabak

Pterostichus (Plectes) drescheri (Fischer, 1817)
Male adult
length: 18,2 mm Russia, West Siberia, low reaches of Ob river, near Khashgort settl., 14.07.1981, Urazova A.leg.
Copyright © Evgeniy Zinoviev

Pterostichus (Plectes) consors (Tschitscherine, 1893)
Female adult
NE Kazakhstan, S Alai Mts., Kalbinsky mt. rng., Ognevka vill., 16.6.2000 (DUBC). det. K. Makarov, 2021
Copyright © Alexander Anichtchenko

Pterostichus (Plectes) drescheri (Fischer, 1817)
Male adult
Russia, W Sayan Mts., Cheremushki vill. 3.8.1998 (DUBC) 18 mm; det. Anichtchenko A.
Copyright © Alexander Anichtchenko

Pterostichus (Pledarus) gibbicollis (Motschulsky, 1844)
Male adult
length: 8,6 mm; Russia, Chita area, Nerchinskyi distr., Nercha river valley near Pravye Kumaki village, 4.08.1999, Zinoviev E.leg.
Copyright © Evgeniy Zinoviev

Pterostichus (Pledarus) gibbicollis (Motschulsky, 1844)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Pledarus) larisae Sundukov, 2013
Female adult
Russian Far East, Primorskii krai, Lazovskii district, 2 km NW Preobrazhenie, mouth of Sokolovka River 28.07.2005 leg. K.Makarov
Copyright © Kirill Makarov

Pterostichus (Pledarus) gibbicollis (Motschulsky, 1844)
Female adult

Copyright © Kirill Makarov

Pterostichus (Pseudethira) gerdi Schmidt, 2009 Holotype
Male adult
„NEPAL P: Seti/D: Bajhang / 30 km NE Chainpur, Ghat- / ganga Khola NE Shima“, „N29°45.08, E81°23.30 / 2400 m, 25.VI.2009 / leg. A. Weigel forest #30a“ (NME).<br /> Photo of Johannes Reibnitz (SMNS).
Copyright © - - -

Pterostichus (Pseudethira) ganesh J.Schmidt, 2005 Paratype
Male adult
Nepal, Ganesh Himal, W of Pansang pass, 3200-3500m, 18.6.2000 (cAA); 15mm
Copyright © Alexander Anichtchenko

Pterostichus (Pseudoferonina) lolo Bergdahl, 2011 Holotype
Male adult
deposited in CAS, labeled: “USA: IDAHO,<br /> Clearwater Co.; Cottonwood Creek @ USFS Rd 250; Orogrande Ck. Rd.; NF Clearwater<br /> R. watershed; elev. ~870 m (2860 ft); 12.5 MI. ESE Headquarters, 3.5 MI. SW<br /> of Bungalow; 46°33'10"N, 115°33'08"W; T38N, R7E, S34; 17 July 2009; #73-2009;<br /> J. C. Bergdahl, coll.”/ “HOLOTYPE Pterostichus lolo Bergdahl designated 2011” [red<br /> label].<br /> Photo of James C. Bergdahl & David H. Kavanaugh / ZooKeys 104: 77–96 (2011)
Copyright © - - -

Pterostichus (Pseudoferonina) bousqueti Bergdahl, 2011 Holotype
Male adult
deposited in CAS, labeled: “USA: Boise Co., Pine Flat Creek at South Fork Payette River Road, 6 km W of Lowman, 44°04'03"N at<br /> 115°40'56"W, T9N, R7E, S31, elev. 1160 m (3805 f ), South Fork Payette River Basin, Boise National Forest, 05 May 1999, #26-1999, J. C. Bergdahl, coll.”/ “HOLOTYPE Pterostichus bousqueti Bergdahl designated 2011” [red label].<br /> Photo of James C. Bergdahl & David H. Kavanaugh / ZooKeys 104: 77–96 (2011)
Copyright © - - -

Pterostichus (Pseudomaseus) anthracinus (Illiger, 1798)
Male adult
W UKRAINE, Lviv Reg., near Kamenka-Bugskaya, in forest under the bark oak 29.III.08 Panin R. leg. L=12mm
Copyright © Ruslan Panin

Pterostichus (Pseudomaseus) gracilis (Dejean, 1828)
Male adult

length: 9,3 mm
Russia, West Siberia, KHMAO-Yugra, Khanty-Mansiysk distr., Ob river, Bolshoy Tchukhtinsiy island
4.08.2007.
Zinoviev E.leg

Copyright © Evgeniy Zinoviev

Pterostichus (Pseudomaseus) luctuosus (Dejean, 1828)
Male adult

Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada

Copyright © Henri Goulet

Pterostichus (Pseudomaseus) minor (Gyllenhal, 1827)
Male adult

W UKRAINE,
Lviv Reg., near Kamenka-Bugskaya, 29.III.2008 Panin R. leg.
L=8mm

Copyright © Ruslan Panin

Pterostichus (Pseudomaseus) rotundangulus A.Morawitz, 1862
Female adult
Collection ground Inami cho, Hyogo Pref. 39361
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Pseudomaseus) anthracinus (Illiger, 1798)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Pseudomaseus) nigrita (Paykull, 1790)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Pseudomaseus) rotundangulus A.Morawitz, 1862
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Pseudomaseus) latoricaensis Pulpan, 1965
Female adult

Copyright © Boleslav Brezina

Pterostichus (Pseudomaseus) fuscicornis (Reiche & Saulcy, 1855)
Male adult
Iran, East Azerbaijan prov., near Kandovan vill. 37.748092°N 46.286895°E 21-22.V.2017 leg.D.Kasatkin L=10.9 mm
Copyright © Kirill Makarov

Pterostichus (Pseudomaseus) fuscicornis (Reiche & Saulcy, 1855)
Endophallus
Iran, East Azerbaijan prov., near Kandovan vill. 37.748092°N 46.286895°E 21-22.V.2017 leg.D.Kasatkin
Copyright © Kirill Makarov

Pterostichus (Pseudomaseus) gracilis (Dejean, 1828)
Female adult

Copyright © Kirill Makarov

Pterostichus (Pseudomaseus) minor (Gyllenhal, 1827)
Male adult

Copyright © Kirill Makarov

Pterostichus (Pseudorites) nicaeensis (A. Villa & G.B. Villa, 1835)
Male adult
La Brigue, Alpes-Maritimes, France
Copyright © Jean Michel Lemaire

Pterostichus (Pseudorites) nicaeensis (A. Villa & G.B. Villa, 1835)
Female adult
Italy, Passo della Guardia - Imperia
Copyright © Gianni Allegro

Pterostichus (Pseudosteropus) illigeri (Panzer, 1803)
Female adult
Austria, UTM VN96, 47.30N 14.53E, path to Eisenerzer Reichenstein 21.6.2019, L. Minkina leg. (cAA) 10 mm
Copyright © Alexander Anichtchenko

Pterostichus (Pterostichus) hagenbachii Sturm, 1824
Male adult
France, Jura, Lajoux, 1190m, 20/05/2013, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Pterostichus (Aphaon) rhilensis kourili Maran, 1933c: 74
Female adult
Bulgaria, Pirin mountains, Vihren, 08.07.2010, E. Svec lgt.; 12 mm, V. Zieris det. 2012, collection Boetzl
Copyright © Fabian Bötzl

Pterostichus (Pterostichus) fasciatopunctatus (Creutzer, 1799)
Female adult
Germany, Bavaria, Passau, Laufenbach dell, F. Bötzl leg. & det., 08.09.2014
Copyright © Fabian Bötzl

Pterostichus (Pterostichus) rutilans (Dejean, 1828)
Male adult
NW Italy, Piemonte,(VC), Oropa, Lago del Mucrone (Lake), h~1900m, 22.VII.1984, L. Diotti leg. L=13 mm
Copyright © Ruslan Panin

Pterostichus (Pterostichus) rufipes (Dejean, 1828)
Male adult
France, 30450, Concoules, 27-05-2017
Copyright © Pascal Deschamps

Pterostichus (Pterostichus) fasciatopunctatus (Creutzer, 1799)
Female adult
Italia : Friuli, Cosizza-viii.1976 - Coll.RBINS, N. Mal
Copyright © Noël Mal

Pterostichus (Pterostichus) fasciatopunctatus (Creutzer, 1799)
Female adult

Copyright © Noël Mal

Pterostichus (Pterostichus) micans Heer, 1838
Male adult
Italy, Lombardia, BG, Castione della Presolana, 8-19.7.2010, 900m, R. Poloni leg. 15 mm
Copyright © Alexander Anichtchenko

Pterostichus (Pterostichus) cristatus cristatus
Male adult
France, 77860, Quincy-Voisins, 20-06-2009
Copyright © Pascal Deschamps

Pterostichus (Pterostichus) pedemontanus Ganglbauer, 1891
Female adult
Italy, Val Sessera - Biella
Copyright © Gianni Allegro

Pterostichus (Pterostichus) focarilei Casale & Giachino, 1985 Paratype
Male adult
Italy, Massiccio del Sirino-Monte del Papa
Copyright © Gianni Allegro

Pterostichus (Pterostichus) andreinii Dodero, 1922
Male adult
Italy, Monte Cucco - Perugia
Copyright © Gianni Allegro

Pterostichus (Pterostichus) dux L.Schaufuss, 1862
Female adult
Spain, Mts Cantabricos, Macizo Central, Picu Urriellu, 2000m, 13.8.2008 (cAA) leg & det. Anichtchenko A.; 18 mm;
Copyright © Alexander Anichtchenko

Pterostichus (Pterostichus) walteri Reitter, 1883f: 224
Female adult

Copyright © Kipling W. Will

Pterostichus (Pterostichus) auratus Heer, 1837
Male adult

Copyright © Vittorino Monzini

Pterostichus (Pterostichus) impressicollis (Fairmaire & Laboulbene, 1854)
Female adult
Italia, Toscana Size: 15mm Leg e Coll. A. Degiovanni
Copyright © Augusto Degiovanni

Pterostichus (Pterostichus) rutilans (Dejean, 1828)
Female adult
14mm; Italia, Piemonte, Leg., e coll., A. Degiovanni
Copyright © Augusto Degiovanni

Pterostichus (Pterostichus) auratus Heer, 1837
Male adult
16mm; Italia, Piemonte Leg., e coll., A. Degiovanni
Copyright © Augusto Degiovanni

Pterostichus (Pterostichus) honnoratii (Dejean, 1828)
Male adult
France, Vanduse, Mt. Serein, 13.5.1983 (cAA). 14 mm
Copyright © Alexander Anichtchenko

Pterostichus (Pterostichus) rhilensis rhilensis Rottenberg, 1874
Male adult
Image from Donabauer, 2020
Copyright © - - -

Pterostichus (Qianius) lii Jia & Tian, 2021 Holotype
Male adult
cave Hei Dong, Longjiang Cun, Shenxi Zhen (Town), Honghuagang Qu (District), Zunyi Shi (City), Guizhou Province, 27°36ʹ46ʺN 106°58ʹ08ʺE, 822 m, 22.VIII.2020, leg. Yunchun Li and Yucheng Lin (SCAU).
Copyright © Mingyi Tian

Pterostichus (Rhagadus) microcephalus Motschulsky, 1860
Male adult
length: 10,0 mm. Russia, Far East, Primorskiy area, Partizanskiy distr., 35 km NE from Sergeevka settl. 01.09.1986 B.Kataev leg. and det.
Copyright © Evgeniy Zinoviev

Pterostichus (Rhagadus) takaosanus Habu, 1958 Holotype
Male adult
near Mt. Takao, TOKYO, 02. V. 1943, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-039
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Rhagadus) takaosanus Habu, 1958 Holotype
Male adult
near Mt. Takao, TOKYO, 02. V. 1943, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-039
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Rhagadus) takaosanus Habu, 1958 Holotype
Male adult
near Mt. Takao, TOKYO, 02. V. 1943, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-039
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Rhagadus) takaosanus Habu, 1958 (Label) Holotype
Male adult
near Mt. Takao, TOKYO, 02. V. 1943, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-039
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Rhagadus) brittoni Habu, 1958 Holotype
Male adult
Katashina, near Oze, GUNMA, 25. VII. 1954, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-040
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Rhagadus) brittoni Habu, 1958 Holotype
Male adult
Katashina, near Oze, GUNMA, 25. VII. 1954, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-040
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Rhagadus) brittoni Habu, 1958 Holotype
Male adult
Katashina, near Oze, GUNMA, 25. VII. 1954, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-040
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Rhagadus) brittoni Habu, 1958 (Label) Holotype
Male adult
Katashina, near Oze, GUNMA, 25. VII. 1954, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-040
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Rhagadus) thorectes Bates, 1873a: 287
Male adult
Collection ground Taka cho, Hyogo Pref. 39536
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Rhagadus) thorectes Bates, 1873a: 287
Endophallus
Collection ground Taka cho, Hyogo Pref. 39536
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Rhagadus) quelpartensis Habu, 1985: 75 Holotype
Male adult
Reihokigan, Quelpart Is., KOREA, 19. V. 1968 Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-052 (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html) NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Rhagadus) quelpartensis Habu, 1985: 75 Holotype
Male adult
Reihokigan, Quelpart Is., KOREA, 19. V. 1968 Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-05 (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html) NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Rhagadus) quelpartensis Habu, 1985: 75 Holotype
Male adult
Reihokigan, Quelpart Is., KOREA, 19. V. 1968 Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-052 (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html) NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Rhagadus) quelpartensis Habu, 1985: 75 Holotype
Male adult
Reihokigan, Quelpart Is., KOREA, 19. V. 1968 Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-052 (http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html) NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Rhagadus) latemarginatus (Straneo, 1936) as straneoi Habu, 1958 Holotype
Male adult
Mt. Hiko, FUKUOKA, 29. X. 1951, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-050
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Rhagadus) latemarginatus (Straneo, 1936) as straneoi Habu, 1958 Holotype
Male adult
Mt. Hiko, FUKUOKA, 29. X. 1951, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-050
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Rhagadus) latemarginatus (Straneo, 1936) as straneoi Habu, 1958 Holotype
Male adult
Mt. Hiko, FUKUOKA, 29. X. 1951, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-050
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Rhagadus) solskyi (Chaudoir, 1878)
Female adult

Copyright © Kirill Makarov

Pterostichus (Rhagadus) microcephalus (Motschulsky, 1860)
Male adult
Russia, Far East, Khosan (DUBC) 9 mm; det. K. Makarov, 2022
Copyright © Alexander Anichtchenko

Pterostichus (Sinoreophilus) scalptus Sciaky & Wrase, 1997
Male adult

Copyright © Gianni Allegro

Pterostichus (Sinoreophilus) lanista (Tschitscherine, 1898)
Male adult
C Sichuan, Jiyuding Shan, 30km N Wenchuan, 3550m, 2.7.2017, leg. V. Patrikeev (cAA). det. Hongliang Shi, 2020
Copyright © Alexander Anichtchenko

Pterostichus (Sinoreophilus) potanini Tschitscherine, 1889
Female adult
China, N Sichuan, Langmusi vill. 3700 5.8.2015, leg. V. Patrikeev (cAA) det. Hongliang Shi, 2020
Copyright © Alexander Anichtchenko

Pterostichus (Sinoreophilus) potanini Tschitscherine, 1889
Male adult
China, S Gansu, Langmusi vill., 3750m, 5.08.2015 (cAA) 14 mm
Copyright © Alexander Anichtchenko

Pterostichus (Sinosteropus) horstweiperti Schmidt & Gueorguiev, 2013 Paratype
Male adult
NEPAL Arun Vall. to Makalu / Kalo Pokhari 4090m / 29.VI.2013 leg. S. Tamang / 27°41’35N 87°12’18E
Copyright © Borislav Gueorguiev

Pterostichus (Lianoe) pseudopachinus Nakane, 1963
Male adult
Collection ground Odaigahara, Nara Pref. 36755
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Lianoe) masumotoi Tanaka; Morita & Suga, 1987 Holotype
Male adult
Mt. Fuji (south side), 1600 m, Shizuoka Pref., Japan, 31. VII-7. VIII. 1971, K. Tanaka & K. Suga
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-253
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Lianoe) masumotoi Tanaka; Morita & Suga, 1987 Holotype
Male adult
Mt. Fuji (south side), 1600 m, Shizuoka Pref., Japan, 31. VII-7. VIII. 1971, K. Tanaka & K. Suga
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-253
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Lianoe) masumotoi Tanaka; Morita & Suga, 1987 Holotype
Male adult
Mt. Fuji (south side), 1600 m, Shizuoka Pref., Japan, 31. VII-7. VIII. 1971, K. Tanaka & K. Suga
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-253
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Lianoe) masumotoi Tanaka; Morita & Suga, 1987 Holotype
Male adult
Mt. Fuji (south side), 1600 m, Shizuoka Pref., Japan, 31. VII-7. VIII. 1971, K. Tanaka & K. Suga
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-253
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Lianoe) masumotoi Tanaka; Morita & Suga, 1987 Holotype
Male adult
Mt. Fuji (south side), 1600 m, Shizuoka Pref., Japan, 31. VII-7. VIII. 1971, K. Tanaka & K. Suga
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-253
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Pterostichus) shiibanus Habu, 1958 Holotype
Male adult
Shiiba, MIYAZAKI, 23. IX. 1950, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-049
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Pterostichus) shiibanus Habu, 1958 Holotype
Male adult
Shiiba, MIYAZAKI, 23. IX. 1950, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-049
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Pterostichus) shiibanus Habu, 1958 Holotype
Male adult
Shiiba, MIYAZAKI, 23. IX. 1950, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-049
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Pterostichus) shiibanus Habu, 1958 Holotype
Male adult
Shiiba, MIYAZAKI, 23. IX. 1950, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-049
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Steropanus) cavifrons Fedorenko, 2018 Holotype
Male adult
Vietnam, Gia Lai Province, ~50 km N of An Khe, Kon Chu Rang Nat[ure]. Reserv[e]., 14°30–31'N/108°32'E, h=1000– 1040 m, pitfall traps, 24.V–2.VI.2016, leg. D. Fedorenko
Copyright © Dmitri Fedorenko

Pterostichus (Steropanus) asulcatus Fedorenko, 2018 Holotype
Male adult
(ZMMU): ‘N-Vietnam, Dien Bien Prov[ince]., Muong Nhe Nat[ure] Reserve, 22°21'54''N/ 102°14'35''E, h=800 m, Exped[ition]. Russ[ia].-V[iet]n[am]. Tropic[al]. C[ente]r, 11–20.VI.2014, A.Abramov, A.Shchinov leg.’.
Copyright © Dmitri Fedorenko

Pterostichus (Steropanus) glymmiger (Andrewes, 1937)
Male adult
Vietnam, Lao Cai Prov., Sa Pa Distr., Fan Si Pan [Phang Xi Pang] Mt., 1400–1500 m, 26.V.–6.VI.1999, N.L.Orlov [leg.], 22°18'56''N 103°49'35''E;
Copyright © Dmitri Fedorenko

Pterostichus (Steropanus) kataevi Fedorenko, 2018 Holotype
Female adult
(ZISP) labelled: ‘CH[ina], Yunnan, Jinping, Shilicun Forest Station, 1960 m, 22°46'28.4''N 103°17'49.3''E, 16.XII.2003, B. Kataev leg. (5)’
Copyright © Dmitri Fedorenko

Pterostichus (Steropanus) violaceus (Straneo, 1949) Holotype
Male adult
Vietnam, Kon Tum Prov[ince], 2–3 km W of Ngoc Linh Mt., 15°05'N, 107°57'E, h=1600–2150 m, pitfall traps, 19.III–IV.2006, leg. A. Anichkin
Copyright © Dmitri Fedorenko

Pterostichus (Steropanus) sulcatipennis Fedorenko, 2018 Holotype
Male adult
Vietnam, Kon Tum Prov[ince], 2–3 km W of Ngoc Linh Mt., 15°05'N, 107°57'E, h=1600–2150 m, pitfall traps, 19.III–IV.2006, leg. A. Anichkin
Copyright © Dmitri Fedorenko

Pterostichus (Steropanus) obliteratus Fedorenko, 2018 Holotype
Male adult
N Vietnam, Lao Cai Prov[ince]., Hoang Lien Son Mt. ridge, env. Fansipan [Phang Xi Pang] Mt, Tram Ton, h=1950–2100 [m], 15–30.VII.2007, leg. D.Fedorenko
Copyright © Dmitri Fedorenko

Pterostichus (Steropanus) pseudoviolaceus Fedorenko, 2018 Holotype
Male adult
Vietnam, Kon Tum Prov[ince], 2–3 km W of Ngoc Linh Mt., 15°05'N, 107°57'E, h=1600–2150 m, pitfall traps, 19.III– IV.2006, leg. A. Anichkin
Copyright © Dmitri Fedorenko

Pterostichus (Steropanus) alveolatus Fedorenko, 2018 Holotype
Unsexed
Vietnam, Kon Tum Prov[ince]., Kon Plong Distr[ict]., 14°45'13''N/ 108°17'40''E, env. ngok Boc I Mt., h = 1300–1500 m, 8–23.IV.2015, D.Fedorenko leg. (Elytra only)
Copyright © Dmitri Fedorenko

Pterostichus (Steropanus) securipenis Fedorenko, 2020 Holotype
Male adult
(ZMMU): Vietnam, Quang Nam Prov[ince]., Nam Gian Distr[ict]., Song Thanh Nat[io]n[al]. Park, 15°33´48´´N/ 107°23´22´´E, h=1050 m, 23.IV–11.V.2019, leg. D. Fedorenko.
Copyright © Dmitri Fedorenko

Pterostichus (Steropus) ebenus (Quensel, 1806)
Male adult
SC Spain, Jaen provincia, Linares, 30.IX.2001, M.A. Lopez Vergara leg.
L=21 mm
Copyright © Ruslan Panin

Pterostichus (Steropus) gallega (Fairmaire, 1859)
Female adult
France, Sainte Engrace, (64) 27-05-2007, Pascal Deschamps coll. & det.
Copyright © Pascal Deschamps

Pterostichus (Steropus) insidiatrix (Brulerie, 1872)
Male adult
Spain, Teruel, Montes Universales, Pto del Portillo 1780m 31.5.2006, Anichtchenko A. leg & det. (cAA) 19mm
Copyright © Alexander Anichtchenko

Pterostichus (Steropus) ghilianii Putzeys, 1846
Male adult
Spain, Sierra de Guadarrama, pto Somosierra, 1500m 3.6.2006, Anichtchenko A. leg & det. (DUBC) 15 mm
Copyright © Alexander Anichtchenko

Pterostichus (Steropus) ghilianii Putzeys, 1846
Female adult
Spain, Sierra de Guadarrama, pto Somosierra, 1500m 3.6.2006, A. Anichtchenko leg & det. (DUBC). 16 mm
Copyright © Alexander Anichtchenko

Pterostichus (Steropus) gallega (Fairmaire, 1859)
Female adult
Spain, Navarra, Bera de Bidasoa, Larrun Mt. 850m 9-15.9.2001, A. Anichtchenko leg. & det. (cAA) 17 mm
Copyright © Alexander Anichtchenko

Pterostichus (Steropus) madidus (Fabricius, 1775)
Male adult
Belgium, Limburg, Forest reserve of Kolmont, 24.4.2016 (DUBC). det. Anichtchenko A., 2021. 14 mm
Copyright © Alexander Anichtchenko

Pterostichus (Steropus) gallega (Fairmaire, 1859)
Male adult
Spain, Navarra, Bera de Bidasoa, Larrun Mt. 850m 9-15.9.2001, A. Anichtchenko leg. & det. (cAA) 19 mm
Copyright © Alexander Anichtchenko

Pterostichus (Steropus) gallega (Fairmaire, 1859)
Male adult
Spain, Asturias, Rioseco, 11.8.87 (DUBC) 16 mm
Copyright © Alexander Anichtchenko

Pterostichus (Tausternus) hoii Fedorenko, 2020 Paratype
Male adult

Copyright © Dmitri Fedorenko

Pterostichus (Troglorites) breuili mendizabali Jeannel, 1921
Male adult
Spain, Guipuzcoa, Albistur, (Tolosa) cueva Mendicute, M.Toribio leg.

Copyright © Alexander Anichtchenko

Pterostichus (Troglorites) breuili mendizabali Jeannel, 1921
Male adult
Spain, Guipuzcoa, Albistur, (Tolosa) cueva Mendicute, M.Toribio leg.
10 mm
Copyright © Alexander Anichtchenko

Pterostichus (Troglorites) ochsi (Fagniez, 1921)
Female adult
Courmes, Alpes-Maritimes, France
Copyright © Jean Michel Lemaire

Pterostichus (Troglorites) breuili breuili Jeannel, 1919
Male adult
Alli, Navarra, Spain, Marc Curti leg.
Copyright © Jean Michel Lemaire

Pterostichus (Troglorites) breuili mendizabali Jeannel, 1921
Female adult

Copyright © Vittorino Monzini

Pterostichus (Troglorites) breuili breuili Jeannel, 1919
Male adult

Copyright © Vittorino Monzini

Pterostichus (Jedlickaia) sterbai Jedlicka, 1934 Paratype
Male adult
Pe Yen Tsin, Yunnan (coll. R. Sciaky)
Copyright © Riccardo Sciaky

Pterostichus (Tschitscherinea) krali Sciaky, 1997 Holotype
Male adult
China, Yunnan, Habashan mts., SE slopes, 4-5.VI.1995 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Pterostichus (Tubuliphallus) megacephalus Sciaky & Allegro, 2013 Holotype
Male adult
China, C Sichuan, Xiling Snow Mts., 2100-3100 m, 1-3.VIII.1996 (CSc)
Copyright © Gianni Allegro

Pterostichus (Vietoderus) laevibasis Fedorenko, 2023 Paratype
Male adult
N-Vietnam, 40km W of / CaoBang, Phia Oac Mt./ E-slope, h=1600-1800 m/ 22°36´27´´N105°52´0´´E/ 22.V-6.VI.2018/ leg. A.Abramov.
Copyright © Dmitri Fedorenko

Pterostichus (Vietosteropus) abramovi Fedorenko, 2017 Holotype
Male adult
(ZMMU): ‘Vietnam, Dak Lak Prov [ince]., Chu Yang Sin Nat[io]n[al]. Park, 12º222403N 108º212113E, 1.5 km W [of ] Chu Pan Phan Mt, h=1650 m, 30.III–11.IV.2012.
Copyright © Dmitri Fedorenko

Pterostichus (Vietosteropus) annamita chuyangsin Fedorenko, 2017 Holotype
Male adult
(ZMMU): ‘Vietnam, Dak Lak Prov.[ince], Chu Yang Sin Natn. Park, 12º232483N/ 108º202593E, Krong Kmar riv[er]., upper flow, h=1200–1650 m, 30.III–14.IV.2012, D. Fedorenko leg.
Copyright © Dmitri Fedorenko

Pterostichus (Vietosteropus) semiopacus Fedorenko, 2017 Holotype
Male adult
(ZMMU): ‘S[outh] Vietnam, Lam Dong Prov[ince]., Bi Doup — Nui Ba [Nature] Res[erve], 12º072N 108º392203E, Bi Doup Mt., N[orthern] slope, h=1700–1900 m, 15–16.IV.2008, leg. D. Fedorenko.
Copyright © Dmitri Fedorenko

Pterostichus (Vietosteropus) bidoupensis Fedorenko, 2017 Holotype
Male adult
(ZMMU) : ‘S[outh] Vietnam, Lam Dong Prov[ince]., Bi Doup — Nui Ba [Nature] Res[erve], 12º102443N 108º402443E, env. Long Lanh, h=1400–1600 m, 1–2.IV.2008, leg. D. Fedorenko
Copyright © Dmitri Fedorenko

Pterostichus (Vietosteropus) ngokboci Fedorenko, 2017 Holotype
Male adult
(ZMMU): Vietnam, Kon Tum Prov[ince]., Kon Plong Distr[ict]., 14º452N 108º172513E, env. ngok Boc I Mt., h =1300–1400 m, 8–10.VI.2016, D. Fedorenko leg.
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Pterostichus (Vietosteropus) konchurang Fedorenko, 2017 Holotype
Male adult
(ZMMU): Vietnam, Gia Lai Province, ~50 km N of An Khe, Kon Chu Rang Nat[ure]. Reserv[e]., 14º312N 108º323E, h=1000–1040 m, 24.V–2.VI.2016, D. Fedorenko leg.
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Pterostichus (Vietosteropus) dalatensis Fedorenko, 2017
Endophallus

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Pterostichus (Vietosteropus) kontumensis Fedorenko, 2017
Endophallus

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Pterostichus (Vietosteropus) abramovi Fedorenko, 2017
Male adult

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Pterostichus (Vietosteropus) sulcicollis Fedorenko, 2017 Paratype
Female adult
Vietnam, Kon Tum Prov[ince], 2–3 km W of Ngoc Linh Mt., 15º052N, 107º572E, h=1600–2150 m, pitfall traps, 19.III–9.IV.2006, leg. A. Anichkin
Copyright © Dmitri Fedorenko

Pterostichus (Vietosteropus) anichkini Fedorenko, 2017 Holotype
Male adult
Vietnam, Kon Tum Prov[ince], 2-3 km W of Ngoc Linh Mt., 15º052N, 107º572E, h=1600–2150 m, pitfall traps, 19.III–9.IV.2006, leg. A. Anichkin
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Pterostichus (Vietosteropus) gialaiensis Fedorenko, 2017 Holotype
Male adult
(ZISP): Vietnam, Gialai-Kontum [= Gia Lai — Kon Tum province], Buonloi [=Buon Loi], 15 XI 1993, Gorokhov
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Pterostichus (Vietosteropus) kontumensis Fedorenko, 2017 Holotype
Male adult
Vietnam, Kon Tum Prov[ince], 2-3 km W of Ngoc Linh Mt., 15º052N, 107º572E, h=1600–2150 m, pitfall traps, 19.III–9.IV.2006, leg. A. Anichkin
Copyright © Dmitri Fedorenko

Pterostichus (Vietosteropus) dalatensis Fedorenko, 2017 Holotype
Male adult
S[outh] Vietnam, Lam Dong Prov[ince]., Bi Doup — Nui Ba [Nature] Res[erve], 12º102443N 108º402443E, env. Long Lanh, h=1400–1600 m, 20– 21.IV.2008, leg. D. Fedorenko
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Pterostichus (Vietosteropus) honbaensis Fedorenko, 2017 Holotype
Male adult
Vietnam, Khanh Hoa Province, Hon Ba Mt, ~1400 m, IV.2003, leg. A.V.Borisenko
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Pterostichus (Vietosteropus) cavicollis Straneo, 1984 Paratype
Female adult
(MSNM): ‘Tonkin, Montes Mauson, April, May, 2-30002, H.Fruhstorfer’, ‘Allotypus, Pterostich., cavicollis [hw]’ [red], ‘Pterostichus cavicollis n.sp. [hw], det. S.L. Straneo 1983, Allotypus. Photo of Fabrizio Rigato.
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Pterostichus (Vietosteropus) ngoclinhensis Fedorenko, 2017 Holotype
Male adult
Vietnam, Kon Tum Prov[ince], 2–3 km W of Ngoc Linh Mt., 15º052N, 107º572E, h=1600–2150 m, pitfall traps, 19.III– 9.IV.2006, leg. A. Anichkin
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Pterostichus (Vietosteropus) chupanphan Fedorenko, 2017 Holotype
Male adult
Vietnam, Dak Lak Prov.[ince], Chu Yang Sin Natn. Park, 12º222403N/ 108º212113E, 1.5 km W [of ] Chu Pan Phan Mt, h=1650 m, 30.III– 11.IV.2012, D. Fedorenko leg.
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Pterostichus (Vietosteropus) konplongensis Fedorenko, 2017 Holotype
Male adult
Vietnam, Kon Tum Prov[ince]., Kon Plong Distr[ict]., 14º442203N 108º192E, env. ngok Boc I Mt., h=1100–1200 m, 8–23.IV.2015, D. Fedorenko leg.
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Pterostichus (Vietosteropus) annamita s.str. (Straneo, 1939)
Endophallus

Copyright © Dmitri Fedorenko