Carabus (Sphodristocarabus) hollbergi Mannerheim, 1827
Male adult
N Azerbaijan,
4km W Ismayilli, h=700m, 9.VII.2008, Pljushch I.G. leg.
male,22mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) adamsi Adams, 1817 (as chiragricus)
Male adult
Russia, Stavropol reg., Kislovodsk vic., 1,5 км ESE Podkumok 905 м, N43*57E42*47, 13-18.05.2006, A.Vlasenko leg.
det. Obydov
Copyright © Dmitri Obydov

Carabus (Sphodristocarabus) adamsi Adams, 1817 (as chiragricus Fischer, 1828)
Male+Female adult
Russia, N. Caucasus, Stavropol reg., Zheleznovodsk, Zheleznaya Mt., h=600 m. 5-11.VI.2011, Е. Shankhiza leg. male - 24 mm, female - 25 mm.
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Carabus (Sphodristocarabus) adamsi Adams, 1817
Endophallus
CN-Georgia, N of Tbilisi, NW of Tianeti, Tvalivi env.,
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) adamsi Adams, 1817
Male adult
CN-Georgia, N of Tbilisi, NW of Tianeti, Tvalivi env., 20.VI.2015, F. Grycz leg. L=24 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) adamsi adamsi (as adamsianus Mandl, 1975)
Male adult
Dagestan. det & Photo I. Belousov
Copyright © Creative Commons

Carabus (Sphodristocarabus) adamsi adamsi (as adamsianus Mandl, 1975)
Female adult
Dagestan. det & Photo I. Belousov
Copyright © Creative Commons

Carabus (Sphodristocarabus) adamsi hollbergi Mannerheim, 1827
Male adult
det & Photo I. Belousov
Copyright © Creative Commons

Carabus (Sphodristocarabus) biroi biroi Menetries, 1832
Male+Female adult
Amasya (Samsun), TURQUIE VI-1987
Copyright © Alain Mollard

Carabus (Sphodristocarabus) blumenthaliensis basilewskianus Breuning & Ruspoli, 1970
Male adult
SW Georgia, Adzharia 10 km Baku 2000m
18.6.98
25 mm
Copyright © Alexander Anichtchenko

Carabus (Sphodristocarabus) blumenthaliensis basilewskianus Breuning & Ruspoli, 1970
Male+Female adult
NE-Turkey,
Artvin Prov., Hopa vers Borcka, Cankutaran Gecidi h=800m, 17.VI.2007 leg. D. Prunier
female, 24mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) blumenthaliensis aequicostalis Staven & Heinz, 1995
Male adult
NE-Turkey:NE of Artvin:Dalis Dağ ,h=1600-1800m, 8-18.VII.2014, C. Reuter leg. L= 25 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) blumenthaliensis aequicostalis Staven & Heinz, 1995
Endophallus

Copyright © Ruslan Panin

Carabus (Sphodristocarabus) bohemani bohemani Menetries, 1832
Female adult
S Azerbaydzan, Lenkoran distr., Vazha riv.,
2 km NW Dashdatuk vill. 38º40*09N 48º42*06E
17.4.2004 Makarov K. leg.
female, 22 mm
Copyright © Alexander Anichtchenko

Carabus (Sphodristocarabus) bohemani bohemani Menetries, 1832
Male adult
Azerbaijan,
Talysh Mts., Polikesh, h~1800m.,2-6.VI.2001 Dolin V.G. leg.
male, 20mm.
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) bohemani esfandiarii Heinz, 1973
Female adult
Hasan-Kif (Mazandaran) IRAN VII-2000
Copyright © Alain Mollard

Carabus (Sphodristocarabus) bohemani esperanzae Heinz, 1970
Male+Female adult
Rasht, (Masuleh) IRAN VI-2000
Copyright © Alain Mollard

Carabus (Sphodristocarabus) elegantulus Motschulsky, 1850 (as astrabadensis Semenov, 1903)
Male+Female adult
Ft.Pashmaky, (Golestan) IRAN VI-2000
Copyright © Alain Mollard

Carabus (Sphodristocarabus) enigmaticus enigmaticus Heinz, 1980
Male+Female adult
Ispir (Erzurum) TURQUIE VI-1991
Copyright © Alain Mollard

Carabus (Sphodristocarabus) enigmaticus citamgineus Lassalle & Charet, 1992
Male+Female adult
Gole (Erzurum) TURQUIE VII-1991
Copyright © Alain Mollard

Carabus (Sphodristocarabus) georgia Cavazzuti, 1984
Male adult

Copyright © Boleslav Brezina

Carabus (Sphodristocarabus) giachinoi Cavazzuti, 1991
Female adult
Ulukoy, (Gumushane) TURQUIE VI-1991
Copyright © Alain Mollard

Carabus (Sphodristocarabus) gilnickii brunneicupreus Deuve, 1994
Male+Female adult
Camlik, (Rize), TURQUIE VI-1997
Copyright © Alain Mollard

Carabus (Sphodristocarabus) gilnickii hasandaghensis Darge & Basquin, 1990
Male+Female adult
Ispir (Erzurum), TURQUIE VI-1997
Copyright © Alain Mollard

Carabus (Sphodristocarabus) heinzi heinzi Breuning, 1964
Male+Female adult
Egribel (Gerisum), TURQUIE VI-1992
Copyright © Alain Mollard

Carabus (Sphodristocarabus) heinzi mateuellus Deuve, 2003
Male adult
Col Harçebeli,(Ordu) TURQUIE V-2001
Copyright © Alain Mollard

Carabus (Sphodristocarabus) kurdicus kurdicus Heinz, 1975
Female adult
Resadiye,(Bitlis) TURQUIE V-1991
Copyright © Alain Mollard

Carabus (Sphodristocarabus) kurdicus kurdicus Heinz, 1975
Female adult

Copyright © Igor Solodovnikov

Carabus (Sphodristocarabus) macrogonus simonianus Breuning & Ruspoli, 1970
Female adult
Turkey, Ordu, Golkoy, 1220 m, 19.7.91
Copyright © Alexander Anichtchenko

Carabus (Sphodristocarabus) macrogonus resplendens Breuning, 1934
Female adult
Turkey; Cayirkent Samsun 12.VII.1996
Copyright © Alexander Anichtchenko

Carabus (Sphodristocarabus) macrogonus macrogonus Chaudoir, 1847
Male adult
NE-Turkey:Trabzon env., Sumela, 17-21.VI.2010, M.Janalik leg.
L=20 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) macrogonus macrogonus Chaudoir, 1847
Endophallus
NE-Turkey:Trabzon env., Sumela,
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) macrogonus pseudokolatensis Heinz & Korge, 1967
Male adult
NE-Turkey::Kackar Dagi Mt, Ayder S of Ardesen h=1600m, 17-21.VI.2010, M.Janalik leg.
L=20 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) macrogonus pseudokolatensis Heinz & Korge, 1967
Endophallus
NE-Turkey::Kackar Dagi Mt, Ayder S of Ardesen h=1600m
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) macrogonus akkusensis Breuning & Ruspoli, 1970 Paratype
Male+Female adult
Akkus,(Ordu) TURQUIE X-1962
Copyright © Alain Mollard

Carabus (Sphodristocarabus) macrogonus diogenei Cavazzuti, 1991
Female adult
Gokçebel,(Sinop) TURQUIE VI-1988
Copyright © Alain Mollard

Carabus (Sphodristocarabus) macrogonus keltepensis Vigna-Taglianti & Bruschi, 1976
Female adult
Mt. Keltepe (Izmit) TURQUIE VI-2005
Copyright © Alain Mollard

Carabus (Sphodristocarabus) macrogonus drannadaghensis Breuning, 1964
Male+Female adult
Drannaz Dag,(Sinop) TURQUIE VII-1993
Copyright © Alain Mollard

Carabus (Sphodristocarabus) macrogonus schweigerinae Schweiger, 1962
Male+Female adult
Lac d’Abant, (Bolu) TURQUIE VIII-1993
Copyright © Alain Mollard

Carabus (Sphodristocarabus) scovitzi czipkai Heinz, 1980
Male adult
Turkey, Artvin, Kafkasor, 1600 m
4.7.1996 Staven and Skoupy leg.
male, 22 mm
Copyright © Alexander Anichtchenko

Carabus (Sphodristocarabus) scovitzi hedini Heinz, 1973
Male+Female adult
Borçka (Artvin) TURQUIE V-2005
Copyright © Alain Mollard

Carabus (Sphodristocarabus) scovitzi sarigoelensis Deuve & Charet, 1992
Male+Female adult
Kaçkar (Artvin) TURQUIE VII-1992
Copyright © Alain Mollard

Carabus (Sphodristocarabus) scovitzi scovitzi Faldermann, 1835
Endophallus
Georgia, Adzharia, Adzharo-Imeretinskiy Mt.R., near Tchidila pass, alpin zone, h=2350 m, 05.VI.2016 leg. D. Fominykh
Copyright © Igor Solodovnikov

Carabus (Sphodristocarabus) scovitzi scovitzi Faldermann, 1835
Male adult
Georgia, Adzharia, Adzharo-Imeretinskiy Mt.R., near Tchidila pass, alpin zone, h=2350 m, 05.VI.2016 leg. D. Fominykh
Copyright © Igor Solodovnikov

Carabus (Sphodristocarabus) separatus persa Lapouge, 1924
Endophallus
N-Iran: Mazandaran Province, Elborz Mt.Rng, Vandarbon env.
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) separatus persa Lapouge, 1924
Male adult
N-Iran: Mazandaran Province, Elborz Mt.Rng, Vandarbon env., 10.VII.2002 L=23 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) separatus bararii Deuve, 2000
Male adult
N-Iran: Mazendaran: Ramsar env., deciduous forest, h~305 m, 16-24.VI.2017, Václav Čutka leg. L=22 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) separatus bararii Deuve, 2000
Female adult
N-Iran: Mazendaran: Ramsar env., deciduous forest, h~305 m, 16-24.VI.2017, Václav Čutka leg. L=23,5 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) separatus separatus Gehin, 1885
Male adult
N-Iran:Gilan Prov., Asalem - Khalkal road, 25-30.V.2018, Václav Čutka leg. L= 21 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) separatus separatus Gehin, 1885
Female adult
N-Iran: Gilan Prov., Asalem - Khalkal road, 25-30.V.2018, Václav Čutka leg. L= 23 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) separatus separatus Gehin, 1885
Male adult
N-Iran:Gilan Prov., Asalem - Khalkal road, 25-30.V.2018, Václav Čutka leg. L= 22 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) separatus separatus Gehin, 1885
Endophallus
N-Iran:Gilan Prov., Asalem - Khalkal road,
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) separatus haerii Heinz, 1973
Female adult
N-Iran:Gilan Prov., Rudbar, Harzevil env.,23-24.V.2018, Václav Čutka leg. L=24 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) separatus haerii Heinz, 1973
Male adult
N-Iran: Gilan Prov., E of Rudbar, Barehsar env., 19-23.VI.2017, Václav Čutka leg. L=22 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) tokatensis almushensis Mandl, 1975
Male adult
Turkey, Sivas, 20 km N Zara, N Arapca, 1750m; 11.VII.1993
Copyright © Alexander Anichtchenko

Carabus (Sphodristocarabus) tokatensis tokatensis Roeschke, 1898
Male+Female adult
Yilidiz Dag, (Tokat) TURQUIE VI-1985
Copyright © Alain Mollard

Carabus (Sphodristocarabus) tokatensis simonae Cavazzuti, 1984
Male+Female adult
Pulumur, (Tunceli) TURQUIE VI-1988
Copyright © Alain Mollard

Carabus (Sphodristocarabus) tokatensis kosensis Cavazzuti, 1984
Female adult
Kosedagi-Geç (Gumushane) TURQUIE VII-2000
Copyright © Alain Mollard

Carabus (Sphodristocarabus) tokatensis kosensis Cavazzuti, 1984
Endophallus
CE-Turkey: Gümüshane: Köse Dag(pass N of Köse), h = 1700 m, VII.2000, leg. St. Il. Remond (TOPOTYPUS)
Copyright © Igor Solodovnikov

Carabus (Sphodristocarabus) tokatensis kosensis Cavazzuti, 1984
Male adult
CE-Turkey: Gümüshane: Köse Dag(pass N of Köse), h = 1700 m, VII.2000, leg. St. Il. Remond (TOPOTYPUS)
Copyright © Igor Solodovnikov

Carabus (Sphodristocarabus) varians varians Fischer, 1827
Male+Female adult
female: Transcaucasia, Armenia, Amasia distr., Karakhachskiy mountain pass h=2270m, 3.VIII.1985 P.V.Kazaryan leg., 22mm<br /> male: Armenia, Amasia distr., Arpalach vill. VI.1999, P.V. Kazaryan leg.
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) varians rugatus Breuning, 1934
Female adult

Copyright © Ruslan Panin

Carabus (Sphodristocarabus) varians janthinus s.str.
Male+Female adult
NW-Caucasus, Russia, Krasnodar reg.,Tuapse distr., Mt.Semashkho 04.V.2008, D.Fominyh leg.<br /> female, 24mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) varians armeniacus Mannerheim, 1830
Female adult
NE Turkey,<br /> Artvin Prov., Borçka, Macahel Gecidi h=1450m, 18.VI.2007, D.Prunier leg.<br /> female, 26mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) varians janthinus (as dvorshaki)
Male adult
Abkhasia, Gagra, 12.VI.1986<br /> photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Sphodristocarabus) varians varians Fischer, 1827
Male+Female adult
N. Armenia, Idzhevan distr., Acharkut vill. env., h=800 m.<br /> 26-28.VII.2008 S. Andreev leg.<br /> male - 22 mm, female - 23 mm.
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Carabus (Sphodristocarabus) varians janthinus s.str. (as dvorshaki Mandl, 1975)
Male+Female adult
W. Caucasus, Abkhazia, Gudauta distr..,<br /> Duripsh vill. env., Khipsta riv., h=400-500 м.<br /> 03-13.VI.2009 A. Safronov leg.<br /> body lenght 20-25 мм.
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Carabus (Sphodristocarabus) varians (as dvorshaki Mandl, 1975)
Endophallus

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Carabus (Sphodristocarabus) armeniacus (as laevilineatus)
Female adult

Copyright © Kirill Makarov

Carabus (Sphodristocarabus) varians armeniacus Mannerheim, 1830
Male adult
NE Turkey, Artvin Prov., Şavşat, Çamlıca,18.VI.1993, M. Janata leg.<br /> L=22 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) varians armeniacus Mannerheim, 1830
Endophallus
NE Turkey, Artvin Prov., Şavşat, Зamlıca.
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) varians janthinus Ganglbauer, 1887
Female adult
Carabus (Sphrodistocarabus) janthinus dvorschaki Mandl, 1975 Abkhazia, Gudautsky district, S- Bzybsky Ridge, a watershed hill at the head of the river Khipsta, 43°17'56.63" N, 40°40'16.74" E, h= 1490 m, mature beech-maple forest with butterbur, moist soil, 05.07-17.08.2017, leg. I.A. Solodovnikov, S.V. Solodovnikova, E.V. Tatun
Copyright © Igor Solodovnikov

Carabus (Sphodristocarabus) varians armeniacus Mannerheim, 1830
Male adult
C Georgia, Imereti, Caucasus, Likhi Mt Range, Rikoti Pass h=900-1000 m, 14-16.V.2017, E. Rutjan E. leg., L=22 mm
Copyright © Ruslan Panin