Carabus (Archicarabus) nemoralis lamadridae Born, 1895 (as layrei Tarrier, 1975)
Male adult
NW-Spain, León Province, Acebo, V.2007, Pascal Stefani leg.
Copyright © Ruslan Panin

Carabus (Archicarabus) pseudomonticola Lapouge, 1908 ? nemoralis
Male adult
NE Spain, Catalonia, Sant Jaume de Frontanyà, h=1000 m, V-VI.2010, Pascal Stefani leg.
L=26 мм
Copyright © Ruslan Panin

Carabus (Archicarabus) pseudomonticola Lapouge, 1908
Male adult
Spain, Montseny, Barcelona
19.VI.1986 M. Tomas leg.
male, 21 mm
Copyright © Alexander Anichtchenko

Carabus (Archicarabus) nemoralis prasinotinctus Heyden, 1880
Male+Female adult
AUDE.Pic de Nore, h=1200m.IX.2005, A. Laforgue leg.
female,25mm/male, 24mm
Copyright © Ruslan Panin

Carabus (Archicarabus) nemoralis prasinotinctus Heyden, 1880
Male adult
N Spain, 3 km N Bera de Bidasoa 14.9.2002
Anichtchenko A. leg.
long. 25mm
Copyright © Alexander Anichtchenko

Carabus (Archicarabus) steuartii Deyrolle, 1852
Male adult
Portugal, S. Martinho, 27.5.1987
male, 18 mm
Copyright © Alexander Anichtchenko

Carabus (Archicarabus) steuartii Deyrolle, 1852
Female adult
NW-Spain, Pontevedra prov., Serra do Candán Mt range, 1.III.1992, S. Devesa leg./det. S. Sobrino
Copyright © Ruslan Panin

Carabus (Carabus) granulatus Linne, 1758
Larva
Carabus (Carabus) granulatus granulatus Linne, 1758
Larva of 2nd instar
L = 16,2 mm
Moskov Reg., Istra Distr., env. Pavlovskaya Sloboda 12.VI.2008 leg. K.Makarov
Copyright © Kirill Makarov

Carabus (Carabus) arvensis deyrollei Gory, 1839
Male adult
Spain, Asturias, 1-8.7.1999
long. 17mm
Copyright © Alexander Anichtchenko

Carabus (Carabus) arvensis deyrollei Gory, 1839
Female adult
NW Spain, Galicia Reg., Pontevedra Prov., A Lama, Xesta, h~650 m, 1.X.1994
L=17 mm
Copyright © Ruslan Panin

Carabus (Carabus) arvensis deyrollei Gory, 1839
Female adult
NW Spain, Lugo env./Sarria/; 15mm
Copyright © Aleš Sedláček

Carabus (Carabus) arvensis deyrollei Gory, 1839
Endophallus
NW-Spain, Cantabria, Udías env., Monte Corona h=100m
Copyright © Ruslan Panin

Carabus (Carabus) arvensis deyrollei Gory, 1839
Male adult
NW-Spain,
Cantabria, Udías env., Monte Corona h=100m,14.VI.2002, Javier de Diego leg.
L=16 mm
Copyright © Ruslan Panin

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761 (as bohemicus)
Female adult
female, 33mm
Copyright © Boleslav Brezina

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Male adult
male, 26mm
Copyright © Boleslav Brezina

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Male adult
W-Ukraine,
Lviv Reg., Zhydachiv distr., near Vybranivka Vill. Panin R.leg. 12.X.2007
31mm.
Copyright © Ruslan Panin

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Female adult
36 mm
Fundort: Germania, Oberfranken, Kulmbach
leg.det. U.Schmidt, 1977
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Male adult
W Ukraine, Lviv city, Sykhiv, Zubra forest, (Osvytskiy Grove), under bark, 49°47'28.10" N, 24° 2'20.09" E, 10.XI.2017, Panin R leg.
Copyright © Ruslan Panin

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Female adult
W Ukraine, Lviv city, Holosko, Bryukhovytsʹkyi forest 49.886833, 24.012083 under bark, 10.XI.2024, R. Panin leg., L=35 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860
Male+Female adult
SW -France,
Pyrenees-Atlantiques, Foret de Sare, XII.2009, Pascal Stefani leg.
female, 31 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860
Female adult
Spain,
Navarra, Sierra de Aralar, Altxueta Mt., 5.V.2006
Anichtchenko A. leg.
L= 17mm
Copyright © Alexander Anichtchenko

Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860
Male adult
Spain,
Pais Vasco, Bera de Bidasoa, Larrun 450m,
9.9.2001 Anichtchenko A. leg.
male, 29mm
Copyright © Alexander Anichtchenko

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as delacrei Tarrier, 1975)
Male+Female adult
NE Spain, Catalonia, Sant Jaume de Frontanyа, h=1000 m, V-VI.2010, Pascal Stefani leg.
female, 38 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as delacrei Tarrier, 1975)
Male adult
Spain,
Catalunya, San Jaume de Frontana,
May 2001
Copyright © Ruslan Panin

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826
Female adult
Auriac alt.
500m. (France : Aude) 25.V.200
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens montanus Gehin, 1882
Male+Female adult
SC-FRANCE,
Haute-Garonne, Col de Mentй h~1300m., II.1980. Kretschmer leg.
female, 23mm.
photo/det. Panin R.
Copyright © Ruslan Panin

Carabus (Chrysocarabus) lineatus lateralis Chevrolat, 1840
Male adult
Spain, Villoslada de Cameros, Logroño, 19.6.87
F. Rodriguez leg.
male, 24mm
Copyright © Alexander Anichtchenko

Carabus (Chrysocarabus) lineatus lineatus Dejean, 1826
Female adult
N-Spain,
Navarra, Sierra de Urbasa, Puerto de Opacua, III.2003
female, 27mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) lineatus salmantinus Bolivar, 1922
Male adult
Spain, Candelario, Salamanca, 19.XI.1988
leg. E. Carabajal
long. 24mm
Copyright © Alexander Anichtchenko

Carabus (Chrysocarabus) splendens Olivier, 1790
Male+Female adult
SE France,
Midi-Pyrйnйes reg., Ariиge, Vaychis, 22.X.2002, A.Laforgue leg.
female, 30mm.
Copyright © Ruslan Panin

Carabus (Chrysocarabus) splendens Olivier, 1790
Male adult
Spain, Pais Vasco, Bera de Bidasoa, 15.9.2001
Anichtchenko A. leg.
male, 30mm
Copyright © Alexander Anichtchenko

Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896 (as juberriensis Mollard, 1999)
Male+Female adult
ANDORRA,
E-Pyrenees, Juberri, h=1350-1400m, 22.VI-18.VII.2007
female, 33mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) lineatus lateralis Chevrolat, 1840 (as quasilineatus Tarrier, 1975)
Male+Female adult
NW Spain,
E Asturias, Monte de Moal h~1000m., 5.VI.2008, Pascal Stefani leg.
female, 27mm.
Copyright © Ruslan Panin

Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860 (as ursuius Lapouge, 1911)
Male+Female adult
FRANCE (64), Ursuya Mt.
XII-2009, P. Stefani leg.
male - 30 mm, female - 32 mm.
Copyright © Alexandr Safronov

Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896
Male adult
Anyos (ANDORRE), h=1450ms,
5-VI-2010, P. Stefani leg.
body lenght - 32 mm.
Copyright © Alexandr Safronov

Carabus (Chrysocarabus) lineatus salmantinus Bolivar, 1922
Male adult
CW Spain, Cáceres, San Martín de Trevejo,24.VI.2011, A. Diez leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) lineatus lineatus (as pseudonumancianus Mollard, 2010)
Male adult
NC-Spain, Sierra de Cantabria , Sierra de Toloño Rng, Puerto de Herrera , h=1100m, 19.I,2007 25 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896
Habitat
Andorra, Anyos, ii.2012, J. Nicolas Leg. & Photos
Copyright © Noël Mal

Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896
Male adult
Andorra, Leg. Gerts, vi.1966, 32 mm
Copyright © Noël Mal

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as aragonensis Lapouge,1913)
Male adult
Male adult ,Sierra de Montseny (Barcelone) Spain.
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as frontanyaensis Mollard,1999)
Female adult
N of Boreda, (Lerida) Spain.
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as brevicollis Lapouge,1910)
Female adult
Corbières (Aude) France.
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as brevicollis var. viridis)
Female adult
Rennes les bains (Aude) France.
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as delacrei Tarrier,1975)
Female adult
Nostra Signora de Queralt, Berga (Barcelona) Spain.
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as occideneissimus Meyer,1995)
Male adult
Collado de Boixols, (Lérida) Spain.
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as urgelensis Mollard,1999)
Female adult
Sierra del Cadi,Toloriu, (Lérida) Spain.
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as jeannei)
Female adult
Sierra de la Mussara (Tarragona) Spain.
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as merangensis Mollard,1999)
Male adult
Méranges, near Puigcerda, (Catalunya) Spain
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as monserraticus Csiki,1917)
Female adult
Sierra de Montserrat, (Barcelone) Spain
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as palensis Mollard,1999)
Male adult
Col de N’hug, (N-Catalunya) Spain.
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as quillansis Mollard,1999)
Male adult
Granès (Aude) France.
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (f. typ.)
Female adult
Prats de Mollo (Pyr.Or) France
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as soredensis Mollard,1999)
Male adult
Ft.de Sorède, Mts. Albères (Pyr.Or.) France
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans opulentus Oberthür, 1898
Female adult
East-Sierra de Aigues-Tortes, Tahull,Bohi (Lerida) Spain
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans opulentus Oberthür, 1898 (as alinsis, Mollard 1999)
Male adult
Alins
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans opulentus Oberthür, 1898 (as turbonensis, Mollard 1999)
Male adult
Massif du Turbon, (Huesca) Spain
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans opulentus Oberthür, 1898 (as cotiellensis, Mollard 1999)
Female adult
East-Mt. Cotiella (Huesca) Spain
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896 (as casteillensis, Mollard 1999)
Female adult
Cap del Prat d’Orient,Castells (Lerida) Spain
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896 (as benesensis, Mollard 1999)
Female adult
S-Sierra de Aigues-Tortes, Malpas-Benes
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896 (as pervesensis, Mollard 1999)
Female adult
Col de Pervers, Sierra de St. Gervas
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896 (as basetaensis, Mollard 1999)
Female adult
Guils, Sierra de la Baseta (Lerida) Spain
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896 (as capdellensis, Mollard 1999)
Female adult
La Torre de Capdella (Lerida) Spain
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans perignitus Reitter, 1896 (as juberriensis, Mollard 1999)
Male adult
Cesters, 1600m, Andorra
Copyright © Alain Mollard

Carabus (Ctenocarabus) galicianus Gory, 1839
Male adult
Spain,
Sierra de la Cabrera Baja, 8 km SW Truchillas, 9.VII.2008, Anichtchenko A. leg.
L= 20 mm
Copyright © Alexander Anichtchenko

Carabus (Ctenocarabus) galicianus Gory, 1839
Female adult
NW-Spain, Pontevedra prov., Campo Lameiro env., 15.VIII.1994, S.Sobrino leg.
L=28 mm
Copyright © Ruslan Panin

Carabus (Ctenocarabus) melancholicus costatus Germar, 1824
Female adult
C-Spain, Madrid region, Rozas de Puerto Real env.,21.XII.2008, T.Gazurek leg.
L=27 mm
Copyright © Ruslan Panin

Carabus (Ctenocarabus) melancholicus melancholicus Fabricius, 1798
Male adult
NW Morocco, Kénitra, Souk El Arbaa env., II.1985, V.Olivella leg. L=25 mm
Copyright © Ruslan Panin

Carabus (Ctenocarabus) melancholicus costatus Germar, 1824
Male adult

Copyright © Noël Mal

Carabus (Ctenocarabus) melancholicus submeridionalis Breuning, 1975
Male adult
S- Spain, Cádiz, Tarifa, El Cabrito, 28.XII.2019, F. Garcia leg. L=26 mm
Copyright © Ruslan Panin

Carabus (Ctenocarabus) melancholicus costatus Germar, 1824
Endophallus
NW Portugal, Viana do Castelo distr., Sierra de Agra
Copyright © Ruslan Panin

Carabus (Ctenocarabus) melancholicus costatus Germar, 1824
Male adult
NW Portugal, Viana do Castelo distr., Sierra de Agra, 41°47'52.3"N 8°41'27.6"W, h=786 m, 3.VII.2016, D. Prunier leg.
Copyright © Ruslan Panin

Carabus (Hemicarabus) nitens Linne, 1758
Male adult
male, 15mm
Copyright © Boleslav Brezina

Carabus (Hemicarabus) nitens Linne, 1758
Male+Female adult
Belarus,
Vitebsk reg., Ushachskiy distr., near Bolshie Dolcy vill. 9.V.1990, I.A.Solodovnikov leg.
female, 18mm.
Copyright © Ruslan Panin

Carabus (Hemicarabus) nitens Linne, 1758
Male adult
Russia, Ural, riv. Rezh, vill. Kamenka; 10.VII.2009, N. Menstschicova leg.
male, 14 mm.,
Copyright © Anton Menstschikov

Carabus (Hemicarabus) nitens Linne, 1758
Male adult
16,6 mm, England, Dorset, Studland
leg.det. J.W.Yerbury, 18.-29.VIII.2006
Coll. Oxford University Museum of Natural History
Photo: U.Schmidt, 2011
Copyright © Udo Schmidt

Carabus (Hemicarabus) nitens Linne, 1758
Female adult

Copyright © Kirill Makarov

Carabus (Hemicarabus) nitens Linne, 1758
Male adult
Belarus, Vitebsk reg., Miory distr. Swamp «Mokh», 09.05.2011, leg. G.G. Sushko
Copyright © Igor Solodovnikov

Carabus (Hemicarabus) nitens Linne, 1758
Endophallus
Belarus, Vitebsk reg., Miory distr. Swamp «Mokh», 09.05.2011, leg. G.G. Sushko
Copyright © Igor Solodovnikov

Carabus (Hemicarabus) nitens Linne, 1758
Male adult

Copyright © Igor Solodovnikov

Carabus (Iniopachus) pyrenaeus jeanneli Deuve, 1994
Male adult
1-2: Pirineos, Canfranc, Ibon de Iserias 2200 m 16-17.7.05<br /> 3: Pirineos, Monte Perdido, Collata Anisolo 2400m 25.8.05
Copyright © Alexander Anichtchenko

Carabus (Iniopachus) auriculatus ubinensis (Puissegur, 1955)
Female adult
Spain, Montes Cantabricos, Peña Ubiña, 2400m
5-6.VII.2009 Anichtchenko A. leg.
female, 19 mm
Copyright © Alexander Anichtchenko

Carabus (Iniopachus) auriculatus auriculatus Putzeys, 1872
Female adult
Spain, Picos de Europa, Macizo Central, Las Colladinas,
ref. Hermoso 2100m, 8.8.2003
Copyright © Alexander Anichtchenko

Carabus (Iniopachus) pyrenaeus pyrenaeus (as cephalicus Csiki, 1927)
Male adult
SW France, Pyrenees-Atlantiques, Mt.Cirque d'Esturou h=1800m, 29.VIII.2014, Pascal Stefani leg. L=20 mm
Copyright © Ruslan Panin

Carabus (Iniopachus) pyrenaeus cerdanus Lapouge, 1924
Male+Female adult
Cirque des Peyssons, ANDORRE, VI-1985
Copyright © Alain Mollard

Carabus (Macrothorax) morbillosus macilentus Lapouge, 1899
Male adult
Spain, Alicante, Santa Pola, 20.2.1988 M. Godoy leg.
male, 29 mm
Copyright © Alexander Anichtchenko

Carabus (Macrothorax) rugosus boeticus Deyrolle, 1852
Female adult
Spain, Cadiz, Novo Sancti Petri, 10.6.2003
female, 32 mm
Copyright © Alexander Anichtchenko

Carabus (Macrothorax) rugosus laufferi Breuning, 1927
Female adult
Spain, Gredos Mts., 2 km E from La Mira Mt.
4-7.7.2007 Anichtchenko A. leg.
female, 29 mm
Copyright © Alexander Anichtchenko

Carabus (Macrothorax) rugosus celtibericus Germar, 1824
Male adult
Portugal, Vila Fresca de Azeitas,
Santana, 13.12.2001
male, 30 mm
Copyright © Alexander Anichtchenko

Carabus (Macrothorax) rugosus boeticus Deyrolle, 1852
Male adult
SC Spain, Jaen reg., Puente de la Sierra, 10.VII.2011, M.Lopez leg.
L=28 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) morbillosus macilentus Lapouge, 1899
Female adult
SE Spain, Murcia reg., P.N. de Calblanque, 1.V.2003, T.Gazurek leg.
L=34 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) rugosus celtibericus Germar, 1824
Female adult
SW-Portugal, Setúbal Distr., Cercal do Alentejo/ Tanganheira vill. 8.XII.2006, T.Gazurek leg.
L=31 m
Copyright © Ruslan Panin

Carabus (Macrothorax) rugosus laufferi Breuning, 1927
Female adult
C-Spain: Avila, El Hornillo env., 27.V.2017, J. Ferrero leg. 30 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) rugosus levantinus Lauffer, 1905
Male adult
SE-Spain, Murcia, Jumillla, Sierra del Carche, 6.VII.2019, F. Garcia leg., L=29 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) rugosus celtibericus Germar, 1824
Male adult
SW-Portugal, Serra do Cercal h=217 m, 37°47'15.6"N 8°42'02.0"W, road N390, 19.III.2011, D. Prunier leg., L=27 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) rugosus celtibericus Germar, 1824
Endophallus
SW-Portugal, Serra do Cercal h=217 m, 37°47'15.6"N 8°42'02.0"W, road N390,
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus fulgens Charpentier, 1825
Male adult
NC-Spain,
Navarra Reg., Sierra de Andнa, IX.2004, Pascal Stefani leg.
male, 30 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus fulgens Charpentier, 1825 (as gerundensis Born, 1916)
Male adult
Spain, Gerona, Garriguella, XI.1990 J. Soler leg.
male, 31mm
Copyright © Alexander Anichtchenko

Carabus (Megodontus) violaceus aurichalceus Kraatz, 1879
Male adult
Spain, Pais Vasco, Aranzazu, 6.I.2005
Anichtchenko A. leg.
male, 27 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Megodontus) violaceus aurichalceus Kraatz, 1879
Male adult
NC-Spain, Cantabria, Santander, Cueto env.XII.2013 L=28 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) macrocephalus Dejean, 1826
Male adult
Spain, Picos de Europa, Macizo Central,
Llagu Bajero 1890m 6.8.2003
Copyright © Alexander Anichtchenko

Carabus (Mesocarabus) lusitanicus lusitanicus Fabricius, 1801
Male adult
Portugal, Vale de Estrela, Guarda, VI.1976
Copyright © Alexander Anichtchenko

Carabus (Mesocarabus) lusitanicus lusitanicus Fabricius, 1801(as schaumi Gaubil, 1849)
Male+Female adult
Portugal,
Viseu distr., Gosende, h~1010m, 15.X.2009, D.Prunier leg.
female,26 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus brevis Dejean, 1826
Female adult
Spain, Sierra Guadarrama, Montes Carpetinos,
Pto. de Somosierra 1600-1700m, 3.VI.2006
Copyright © Alexander Anichtchenko

Carabus (Mesocarabus) breuningi Csiki, 1927
Male adult
Spain, San Martin de Castaneda, Zamora<br /> 11 oct 1984,h~1500 m
Copyright © Alexander Anichtchenko

Carabus (Mesocarabus) lusitanicus gougeleti Reiche, 1863 (as alcaracinus Kraatz, 1886)
Male adult
Spain, Riopa (Albacete)
Copyright © Alexander Anichtchenko

Carabus (Mesocarabus) dufouri dufouri Dejean, 1829 (as papillatus Lapouge, 1902)
Male adult
Spain, Algeciras, Molino de las Cuevas
5-6.4.2003 Anichtchenko A. leg.
Copyright © Alexander Anichtchenko

Carabus (Mesocarabus) baguenai Breuning, 1926
Male adult
Spain, Almeria, Sierra de Baza 10.III.1995<br /> male<br /> photo Anichtchenko A.<br />
Copyright © Alexander Anichtchenko

Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
Male adult
SW-France,
Foret d'Iraty Pyrenees-Atlantiques, d'Occabe, 15.VIII.1988
male,30 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876 Loc.typ.
Female adult
Spain, Navarra, Sierra de Aralar,
Altxueta 1200-1300m 6.1.2001
Copyright © Alexander Anichtchenko

Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
Endophallus
endophalus
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus planiusculus Haury in Gehin, 1885
Male adult
NE Spain, Catalonia, Sant Jaume de Frontanyà, h=1000 m, V-VI.2010, Pascal Stefani leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) latus latus Dejean, 1826
Female adult
CW Spain, Cáceres, San Martín de Trevejo, 24.VI.2011, A. Diez leg./det.Panin R.
L=27 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) dufouri dufouri Dejean, 1829
Endophallus
Jaen reg., Monte Javalcuz h=1436 m
Copyright © Ruslan Panin

Carabus (Mesocarabus) dufouri dufouri Dejean, 1829
Male adult
SC Spain, Jaen reg., Monte Javalcuz h=1436 m, 6.X.2010, M.A. Lopez leg.
L=27 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus brevis (as sorianus Jeanne, 1972)
Endophallus
NC-Spain, Soria, Puerto de Oncala
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus brevis (as sorianus Jeanne, 1972)
Male adult
NC-Spain, Soria, Puerto de Oncala, 2.VI.2006
L=24 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus gougeleti Reiche, 186
Male adult
CW Spain, Province Huelva, Arroyomolinos de León,30.X.2011, T. Alcantara leg.; L=23 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) (? lusitanicus) gougeleti Reiche, 186
Endophallus
CW Spain, Province Huelva, Arroyomolinos de Leon.
Copyright © Ruslan Panin

Carabus (Mesocarabus) cantabricus cantabricus Chevrolat, 1840
Endophallus
NW-Spain, Pontevedra prov., A Cañiza viil., Alto de Fontefría h=800m
det.Panin R.
Copyright © Ruslan Panin

Carabus (Mesocarabus) cantabricus cantabricus Chevrolat, 1840 (loc. typ.)
Male adult
NW-Spain, Pontevedra prov., A Cañiza viil., Alto de Fontefría h=800m, 31.XII.2007, T.Gazurek leg./det.Panin R.<br /> L=27 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) dufouri baguenai Breuning, 1926
Endophallus
SE Spain, Almería Province, Carboneras env.
det. R.Panin
Copyright © Ruslan Panin

Carabus (Mesocarabus) baguenai Breuning, 1926
Male adult
SE Spain, Almería Province, Carboneras env., 25.XI.2006, T.Gazurek leg./det. R.Panin <br /> L=26 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) macrocephalus Dejean, 1826
Male adult
NW-Spain,Cantabrian Mts, Picos de Europa Mt Rng, Puerto de Panderruedas h=1460 m, 25.IX.2004,T.Gazurek leg.
L=28 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) macrocephalus Dejean, 1826
Endophallus
NW-Spain,Cantabrian Mts, Picos de Europa Mt Rng, Puerto de Panderruedas h=1460 m,
Copyright © Ruslan Panin

Carabus (Mesocarabus) latus cf. molossoides Lassalle, 1984
Male adult
C-Spain, Toledo, Mts de Toledo, Sierra del Castañar Rng, Puerto del Milagro env, h=920m, 7.II.2009, T.Gazurek leg./det.R. Panin.
L=24 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) latus latus Dejean, 1826
Endophallus
C-Spain, Toledo, Mts de Toledo, Sierra del Castañar Rng, Puerto del Milagro env, h=920m
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus helluo (as albarracinus Ganglbauer, 1886)
Male adult
CE-Spain, Provincia Teruel, Sierra de Javalón Rng, El Cañigral env., h=1460m, X.2010, A. Mollard leg.<br /> L=23 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus helluo (as albarracinus Ganglbauer, 1886)
Endophallus
CE-Spain, Provincia Teruel, Sierra de Javalón Rng, El Cañigral env., h=1460m,
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826
Male adult
CW-Spain: Salamanca, Brincones env., h=750m, X.2003<br /> L=25 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826
Endophallus
CW-Spain:Salamanca, Brincones env., h=750m
Copyright © Ruslan Panin

Carabus (Mesocarabus) (? problematicus trans lusitanicus) planiusculus (as xaxarsi Born, 1917)
Male adult
NE Spain, Catalonia, Girona Prov., San Hilario Sacalm env., 20.II.1988, A. Mollard leg.<br /> L=25 m
Copyright © Ruslan Panin

Carabus (Mesocarabus) (? problematicus trans lusitanicus) planiusculus (as xaxarsi Born, 1917)
Endophallus
NE Spain, Catalonia,Girona Prov., San Hilario Sacalm env.<br />
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus helluo Dejean, 1826
Female adult
SE-Spain, Prov.Alicante, Port de la Carrasqueta, 26.XII.1987, M.Toribio leg. L=25 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) latus antiquus Dejean, 1826 Holotype
Female adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC36 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec36
Copyright © - - -

Carabus (Mesocarabus) latus antiquus Dejean, 1826
Label
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC36 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec36
Copyright © - - -

Carabus (Mesocarabus) lusitanicus brevis Dejean, 1826 Lectotype
Female adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC32 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec32
Copyright © - - -

Carabus (Mesocarabus) lusitanicus brevis Dejean, 1826
Label
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC32 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec32
Copyright © - - -

Carabus (Mesocarabus) lusitanicus egesippei La Ferté-Sénectere, 1847 Holotype
Female adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC154 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec154
Copyright © - - -

Carabus (Mesocarabus) lusitanicus egesippei La Ferté-Sénectere, 1847
Label
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC154 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec154
Copyright © - - -

Carabus (Mesocarabus) lusitanicus mertolaensis Breuning, 1975 (= brevis) Holotype
Female adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC130 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec130
Copyright ©

Carabus (Mesocarabus) lusitanicus mertolaensis (error of label?)
Label
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC130 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec130
Copyright ©

Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826 Holotype
Male adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC34 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec34
Copyright © - - -

Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826
Label
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC34 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec34
Copyright © - - -

Carabus (Mesocarabus) lusitanicus latus Dejean, 1826 Lectotype
Male adult
"Navalmoral" [Navalmoral de la Mata]. Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC35 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec35
Copyright © - - -

Carabus (Mesocarabus) latus latus Dejean, 1826
Label
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC35 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec35
Copyright © - - -

Carabus (Mesocarabus) lusitanicus trypalsi Breuning, 1975 Paratype
Female adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC132 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec132
Copyright © - - -

Carabus (Mesocarabus) lusitanicus trypalsi Breuning, 1975
Label
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC132 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec132
Copyright © - - -

Carabus (Mesocarabus) lusitanicus trypalsi Breuning, 1975 Holotype
Male adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC133 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec133
Copyright ©

Carabus (Mesocarabus) lusitanicus trypalsi Breuning, 1975
Label
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC133 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec133
Copyright © - - -

Carabus (Mesocarabus) lusitanicus helluo Dejean, 1826 Holotype
Female adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC31 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec31
Copyright © - - -

Carabus (Mesocarabus) lusitanicus helluo Dejean, 1826
Label
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC31 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec31
Copyright © - - -

Carabus (Mesocarabus) dufouri papillatus Lapouge, 1902 Lectotype
Male adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC576 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec576
Copyright © - - -

Carabus (Mesocarabus) dufouri papillatus Lapouge, 1902
Label
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC576 http://coldb.mnhn.fr/catalognumber/mnhn
Copyright © - - -

Carabus (Mesocarabus) lusitanicus gougeleti (as valderepisensis Branes, 1988)
Male adult
Spain, Jaen prov., Despeñaperros, Arroyo de los Arcos, 38.381847, -3.515651, 1-2.05.2014, Anichtchenko A. leg. & det.; 18 mm
Copyright © Alexander Anichtchenko

Carabus (Mesocarabus) cantabricus moroderi (as rodriguezianus Meyer & Mollard in Deuve, 2003)
Female adult
Poza de la Sal, Burgos, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) problematicus planiusculus (as xaxarsi)
Male adult
Sant Sadurni, Gerone, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) problematicus planiusculus Haury in Gehin, 1885
Male adult
La Seo de Urgel, Gerone, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) problematicus planiusculus (as bepmalei Lapouge, 1904)
Female adult
Pic de Bassibes, Huesca, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) problematicus planiusculus (as navarrensis Breuning, 1933)
Male adult
Mt.Urculo, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) problematicus planiusculus (as trapeti Bleuse, 1885)
Female adult
Panticosa, Huesca, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) problematicus planiusculus (as lagardei Mollard, 1996) Paratype
Female adult
Sierra de Andia, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) problematicus planiusculus (as andorranus Lapouge, 1902)
Female adult
Col d'envalira, Pas de la Case, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) lusitanicus lusitanicus (as vieirae Oliveira, 1875)
Male adult
Guia, Leiria, Portugal. det. A. Anichtchenko; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) lusitanicus lusitanicus (as schaumi Gaubil, 1849)
Male adult
S.de Gata, Extramadure, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) lusitanicus castilianus (as fuentei Breuning, 1926)
Male adult
Retortillo, Salamanque, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) lusitanicus castilianus (as bolivari Breuning, 1926)
Male adult
Ledrada, Salamanque, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826
Female adult
Villasdardo, Salamanque, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) lusitanicus castilianus (as sayagoensis Mollard 2006) Paratype
Male adult
Moral de Sayago, Zamora, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) lusitanicus brevis (as olmedoensis Mollard, 2007) Paratype
Female adult
Olmedo, Valladolid, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826
Male adult
S.de Candelario, Salamanca ("Avila" error). Spain. det. A. Anichtchenko, 2016; (in Mollard, 2013 as lusitanicus brevis) Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) lusitanicus antiquus Dejean, 1826
Female adult
Portugal, Ribatejo, Coruche, 13.x.1974, A. Zuzarte leg., 25 mm, Coll. N. Mal
Copyright © Noël Mal

Carabus (Mesocarabus) lusitanicus latus (as molossoides Lassalle, 1984)
Male adult
Cabezabellosa, Extramadure, det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) lusitanicus gougeleti (as nevadensis Brañes, 1989) Paratype
Male adult
Guadix, Granada, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) lusitanicus gougeleti Reiche, 1863
Female adult
S.Norte de Sevilla, Huelva, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) lusitanicus antiquus Dejean, 1826
Male adult
Aliseda, Extramadure, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) lusitanicus helluo Dejean, 1826
Female adult
Puerto de Tortosa, Tarragone, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) lusitanicus trabuccarius Fairmaire, 1857
Female adult
Gaia, Barcelona, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) lusitanicus trabuccarius (as galardonensis Meyer & Mollard, 1996) Paratype
Female adult
Puerto de Monrepos, Huesca, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) cantabricus moroderi (as magdalenai Meyer & Mollard, 1996) Paratype
Female adult
Pto. de Magdalena, Leon, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) cantabricus moroderi Breuning, 1926
Male adult
Foncebadon, Leon, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) cantabricus moroderi (as almanzaensis Meyer & Mollard in Deuve, 2003) Paratype
Female adult
Almaza, Leon, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) breuningi Csiki, 1927
Male adult
Villanueva, Orense, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) cantabricus cantabricus (as asturicus Born, 1925)
Female adult
Pravia, Asturies, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) cantabricus cantabricus (as liebei Eidam, 1925)
Male adult
Fonsagrada, Galice, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) cantabricus cantabricus Chevrolat, 1840 (loc. typ.)
Female adult
La Caniza, Galice, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) cantabricus moroderi (as joaquini Breuning & Toulgoet, 1974)
Male adult
Pena Ubina, Leon, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) cantabricus amayensis Lassalle, 1984
Male adult
Pto.de Amaya, Burgos, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) macrocephaloides (Jeanne, 1972)
Female adult
Sierra de Andia, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) macrocephaloides (Jeanne, 1972)
Female adult
Sierra de Andia, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) macrocephaloides (Jeanne, 1972)
Female adult
Sierra de Andia, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) macrocephaloides (Jeanne, 1972)
Female adult
Sierra de Andia, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
Female adult
Sierra de Aralar, Navarra, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) cantabricus brabeus (as barcelecoanus)
Male adult
Col de Velate, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) cantabricus brabeus (as barcelecoanus)
Female adult
Col de Velate, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
Male adult
Col de Velate, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) lusitanicus brevis (as segovianus Mollard, 2011) Paratype
Male adult
S.de Gorda, Soria, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) lusitanicus gougeleti (as alcaracinus)
Female adult
Alcaraz, Albacete, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) lusitanicus helluo (as alicantinus)
Female adult
S.de Bernia, Alicante, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) lusitanicus trabuccarius (as leyrensis)
Female adult
Sierra de Leyre, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) lusitanicus trabuccarius (as trypalsi)
Male adult
Plan du Trypals, Huesca, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) lusitanicus trabuccarius (as solanensis)
Male adult
S.de Bolave, Huesca, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) lusitanicus trabuccarius (as troncedoensis) Paratype
Female adult
Troncedo, Huesca, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) lusitanicus trabuccarius (as benabarrensis) Paratype
Male adult
Benabarre, Huesca, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) lusitanicus brevis (as pironensis) Paratype
Female adult
Pennasrubias, Segovia, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) lusitanicus brevis (as lermaensis) Paratype
Female adult
Espinosa de Cervera, Burgos, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) lusitanicus brevis (as logronicus)
Male adult
Pto.de Piqueras, La Rioja, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) lusitanicus trabuccarius (as catalonicus)
Male adult
Ft.de Roquerolles, Prades, Tarragone, SPAIN
Copyright © Alain Mollard

Carabus (Mesocarabus) lusitanicus brevis (as segovianus)
Male adult
Sierra de Pardos, South Calatayud, Saragosse, SPAIN
Copyright © Alain Mollard

Carabus (Mesocarabus) lusitanicus helluo (as albarracinus)
Female adult
Altos de Cabrera, Pineda de Ciguela, Cuenca, SPAIN
Copyright © Alain Mollard

Carabus (Mesocarabus) lusitanicus cf. antiquus-gougeleti
Male adult
Bicada, South Beja, PORTUGAL
Copyright © Alain Mollard

Carabus (Mesocarabus) leptopus Thomson, 1875 Holotype
Female adult
3957:2 (MZLU)
Copyright © Lund University

Carabus (Mesocarabus) leptopus
Label

Copyright © Lund University

Carabus (Mesocarabus) breuningi Csiki, 1927
Female adult
NW-Spain, Prov. Zamora, San Martín de Castañeda, 5.VII.1984, M.Toribio leg. L=28 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) dufouri Dejean, 1829
Female adult
S Spain, Granada, Sierra Arana, Píñar env., 14.X.2018, F. Garcia leg. L=27 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) breuningi Csiki, 1927 (loc.typ.)
Male adult
NW-Spain, Orense, Orensano Central Massif, Monte de Cabeza de Manzaneda, 25.VII.1988, F. Rodrigues leg. L=25 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) cantabricus moroderi (as magdalenai Meyer & Mollard, 1996)
Endophallus
NW-Spain, Leon prov., Sierra de Gistredo, Pto. de Magdalena,
Copyright © Ruslan Panin

Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
Endophallus
NC-Spain, Basque Co, Álava, Sierra de Gorbea.
Copyright © Ruslan Panin

Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
Male adult
NC-Spain, Basque Co, Álava, Sierra de Gorbea, VI.2014, L=29 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus helluo (as alicantinus Ganglbauer, 1886)
Endophallus
SE-Spain, Murcia reg., Jumilla, Sierra del Carche, h=1300 m,
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus helluo (as alicantinus Ganglbauer, 1886)
Male adult
SE-Spain, Murcia reg., Jumilla, Sierra del Carche, h =1300 m, 30.XI.2019, F. Garcia leg.
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus antiquus Dejean, 1826
Female adult
SW Portugal, Beja Distr., 3,7 km NW Brunheiras, road CM1072, 37°46'31.9"N 8°47'02.2"W, 15.III.2011, D. Prunier leg., L=26 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus trabuccarius Fairmaire, 1857
Endophallus
NE Spain, Catalonia, Tarragona, Sierra de Montsant h=800 m, Albarca env.,
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus trabuccarius Fairmaire, 1857
Male adult
NE Spain, Catalonia, Tarragona, Sierra de Montsant h=800 m, Albarca env.,7.XI.2018, T. Rafales leg., L=25 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus trabuccarius Fairmaire, 1857
Female adult
NE Spain, Catalonia, Tarragona, Sierra de Montsant h=800 m, Albarca env.,7.XI.2018, T. Rafales leg., L=27 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus lusitanicus (as vieirae Oliveira, 1875)
Female adult
CW-Portugal: Leiria distr., near Marinha Grande, 39°45'57.8"N 8°58'10.3"W, h-79 m, 19.IX.2016, D. Prunir leg., L=26 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus lusitanicus (as vieirae Oliveira, 1875)
Endophallus
CW-Portugal: Leiria distr., near Marinha Grande, 39°45'57.8"N 8°58'10.3"W, h-79 m
Copyright © Ruslan Panin

Carabus (Mesocarabus) lusitanicus lusitanicus (as vieirae Oliveira, 1875)
Male adult
CW-Portugal: Leiria distr., near Marinha Grande, 39°45'57.8"N 8°58'10.3"W, h-79 m, 19.IX.2016, D. Prunir leg., L=24 mm
Copyright © Ruslan Panin

Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847 (as guadalupensis) Paratype
Male adult
ESP - Caceres, S. Guadalupe,
800 m XI.2002
Copyright © Alexander Anichtchenko

Carabus (Oreocarabus) ghiliani La Ferte-Senectere, 1847
Male adult
Spain, Segovia, Sierra de Somosierra,
La Pinilla, 1800m 4.4.2005
male, 19 mm
Copyright © Alexander Anichtchenko

Carabus (Oreocarabus) luetgensi luetgensi s.str.
Male adult
Spain, Zamora, San Martin de Castaneda
10.5.1997
male, 20 mm
Copyright © Alexander Anichtchenko

Carabus (Oreocarabus) luetgensi getschmanni Lapouge, 1924
Male adult
NW-Spain, León Province, Acebo, h=1000m, V.2007, Pascal Stefani leg.
L=22 mm
Copyright © Ruslan Panin

Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847
Male adult
WC Spain, Cáceres, Hervás, h=1000m, V.2007, Pascal Stefani leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847
Endophallus
WC Spain, Cáceres, Hervás, h=1000m
Copyright © Ruslan Panin

Carabus (Oreocarabus) luetgensi luetgensi
Female adult
Maetzu, Alava, Spain. det. A. Mollard. Copyright © Magellanes
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Carabus (Oreocarabus) luetgensi getschmanni Lapouge, 1924
Male adult
Pto.de Opakua, Alava, Spain. det. A. Mollard. Copyright © Magellanes
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Carabus (Oreocarabus) ghiliani La Ferte-Senectere, 1847
Male adult
Pto.de Navafria, Madrid, Spain. det.A.Mollard; Copyright © Magellanes
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Carabus (Oreocarabus) ghiliani (as negrei Breuning, 1966)
Male adult
Sierra de Gredos, Avila, Spain. det.A.Mollard; Copyright © Magellanes
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Carabus (Oreocarabus) luetgensi luetgensi (f. typ.)
Female adult
Sierra de Marao, Vila Real, Portugal; det. A. Mollard. Copyright © Magellanes
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Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847
Female adult
los Santos, Salamanque, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847
Male adult
Villamadana, Palencia, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Oreocarabus) guadarramus (as duerensis Raynaud, 1974)
Female adult
Burgo de Osma, Soria, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Oreocarabus) guadarramus (as duerensis Raynaud, 1974)
Male adult
Sierra de Cucalon, Teruel, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Oreocarabus) guadarramus (as andiaensis Forel & Leplat, 1998)
Male adult
Sierra de Andia, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Oreocarabus) guadarramus (as guadalupensis Mollard, 2003) Paratype
Male adult
S.de Guadalupe, Extramadure, Spain. det.A.Mollard; Copyright © Magellanes
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Carabus (Oreocarabus) guadarramus (as sagranus)
Female adult
Sierra de la Sagra, Granada, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Oreocarabus) guadarramus (as hirschfelderi Branes, 1987)
Male adult
Puerto Viejo, Salamanque, Spain. det. A. Mollard; Copyright © Magellanes
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Carabus (Oreocarabus) ghiliani La Ferte-Senectere, 1847
Female adult
C-Spain: Sierra de Guadarrama, Puerto de Cotos (Madrid), 20.IX.2018, F.Rodriguez leg., L=22 mm
Copyright © Ruslan Panin

Carabus (Oreocarabus) luetgensi pseudoguadarramus Mollard, 2006 Paratype
Female adult
Montes de Herrera (Valladolid) Espagne, XI-2005
Copyright © Alain Mollard

Carabus (Tachypus) cristoforii Spence, 1823 (as nicolasi Reitter, 1888)
Male adult
S-France, Hautes-Pyrenees, Pic de Nere h=2200m,15.VII.1999,
male, 14mm
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus fusus Palliardi, 1825
Male+Female adult
France, Riveres Charente, leg. Laforgue VI.1996
female, 24mm
Copyright © Ruslan Panin

Carabus (Tachypus) auratus auratus Linne, 1761
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Carabus (Tachypus) auratus s.str. (as brunieri Barthe, 1921)
Male adult
SC-FRANCE, Aude, Rennes les Bains, 6.VII.1989, Sunard A. leg.
male, 26mm
Copyright © Ruslan Panin

Carabus (Tachypus) auratus auratus Linne, 1761
Female adult
30 mm
Germania, Oberfranken, Pegnitz
leg.det. U.Schmidt, 1972
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Carabus (Tachypus) cristoforii Spence, 1823
Female adult
S France, Pyrénés Mts., Tarascon; 15mm
Copyright © Aleš Sedláček

Carabus (Tomocarabus) convexus s.str. (as pyrenaicus Born, 1907)
Male adult
Pirineos, Canfranc, 1300m, Barranco de Izas
15.6.2005 Anichtchenko A. leg.
male, 16 mm
Copyright © Alexander Anichtchenko

Carabus (Tomocarabus) convexus s.str. (as paganetti Born, 1905)
Male adult
Lazio, Roma, Bosco di Nettuno
22.5.1986 C. Iacovone leg.
male, 20 mm
Copyright © Alexander Anichtchenko

Carabus (Tomocarabus) convexus convexus Fabricius, 1775
Male adult
W.Ukraine,
Transcarpathian region,Vinogradov distr., near Malaya Kopanya vill., Chorna Gora Mt., h=180m. 7-18.VII.08. Panin R. leg.
male 18 mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) convexus s.str. (ab. niger)
Male adult
W Ukraine, NW Podolia, Lviv Region, Zolochiv distr., near Gologirky vill., in steppe 1-19.V.2007 Panin R. leg.
16mm
Copyright © Ruslan Panin