Carabus (Tribax) agnatus agnatus Ganglbauer, 1889
Male adult
NW-Caucasus,
Krasnodar reg., Mostovskoj distr., Skirda Mts. N. slope Yatyrgvart Mt., h=2300-2500m. 09-10.VII.2008, D.Fominyh leg.
male, 23mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Tribax) agnatus pseudagnatus Novotný & Voříšek, 1988
Female adult
NW-Caucasus, Karachay-Cherkessia, Bol'shaya Marka River h=2500-2600m, 12.VIII.2010, I.A. Solodovnikov leg./det.
Copyright © Ruslan Panin

Carabus (Tribax) agnatus pseudagnatus Novotný & Voříšek, 1988
Male adult
W-Caucasus: Abkhasia, Gudauta distr., Mt Anchkho h=2200-2350 m, 14-20.VII.2011, S.Pavluchuk, I. Solodovnikov leg./det. I. A. Solodovnikov.
L=25 mm
Copyright © Ruslan Panin

Carabus (Tribax) agnatus pseudagnatus Novotný & Voříšek, 1988
Endophallus
W-Caucasus: Abkhasia, Gudauta distr., Mt Anchkho h=2200-2350 m,
Copyright © Ruslan Panin

Carabus (Tribax) agnatus agnatus Ganglbauer, 1889
Male adult

Copyright © Kirill Makarov

Carabus (Tribax) agnatus agnatus Ganglbauer, 1889
Female adult

Copyright © Kirill Makarov

Carabus (Tribax) apschuanus apschuanus Rost, 1893
Male+Female adult
Carabus (Tribax) apschuanus apschuanus Rost, 1893    
Russia, Caucasus, Krasnodar reg.,
h=1200 m, 29.VII.2000
A. Zamotajlov leg.
Copyright © Safronov

Carabus (Tribax) apschuanus pseudoplatessa Gottwald, 1982
Male+Female adult
NW-Caucasus, Karachay-Cherkessia, Teberda: Dzhamagat gorge, the left bank of the riv. Dzhamagat, 10.VII-28.VIII.2010 S.Pavluchuk leg.
female, 27 mm
Copyright © Ruslan Panin

Carabus (Tribax) apschuanus pseudoplatessa Gottwald, 1982
Endophallus
endophallus
Copyright © Ruslan Panin

Carabus (Tribax) apschuanus pseudoplatessa Gottwald, 1982
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Carabus (Tribax) apschuanus apschuanus Rost, 1893
Male adult
ABKHASIA, Sukhum distr., N slope of Dou Mt.R., left board of the river Bzyb, (Fagus + ferns + stony taluses (the beginning of a track on Dow's (Dou) pass), H = 740 m, 43°19'3.64" N, 40°52'37.55" E, 10.V-19.VI.2015, Leg. I.A. Solodovnikov, E.V. Tatun
Copyright © Igor Solodovnikov

Carabus (Tribax) biebersteini ulgeni Gottwald, 1982
Male adult
S Russia, Karacaj-Cherkessija,
near Kart-Djurt vill., upper Kylan riv. h=1300m, 26-27.V.2009, Fominykh D. leg.
male, 28mm
Copyright © Ruslan Panin

Carabus (Tribax) biebersteini ingusch Zolotarev, 1913
Male+Female adult
Carabus (Tribax) biebersteini ingusch Zolotarev, 1913
Russia, C.Caucasus, N.Ossetia,
valley of River Khazniaush
10.VI-31.VII.1993
A. Zamotajlov leg.
Copyright © Safronov

Carabus (Tribax) biebersteini constantinowi Starck, 1894
Male adult
Carabus (Tribax) biebersteini constantinowi Starck, 1894
NW-Caucasus, S Russia,
Karacaj-Cherkessija, Nizhniy Arkhyz, 21-30.V.2001, Tikhonov V. leg.
male,30mm
Copyright © Ruslan Panin

Carabus (Tribax) biebersteini vlasenkoi Obydov, 2011 Holotype Paratype
Male+Female adult
Russia, North Caucasus,Stavropol Region, 33 km from Kislovodsk City to the north-west, Predgornyi Dist.,8 km from st. Bekeshevskaya to the south-west, Forestry “Ugolnaya Dacha”,Pastbischnyi Mt. Ridge, left coast of Kuma River, 800 m, N44º02’ E42°21’, 11-17.05.2006, A. Vlasenko leg.
Copyright © - - -

Carabus (Tribax) circassicus teberdensis Zolotarev, 1913
Male+Female adult
W-Caucasus: Karacaj-Cherkessija,
Abishira–Akhuba Mnt. Range h=2000-2650m, Kyafar-Agur Riv., 14–17.VII.2009, Fominykh D. leg.
female, 25mm
Copyright © Ruslan Panin

Carabus (Tribax) circassicus kubaniensis Semenov, 1896
Male adult
N Caucasus, Krasnodar reg., 30 km S Maykop
august 1993
male, 26 mm
Copyright © Alexander Anichtchenko

Carabus (Tribax) circassicus kubaniensis Semenov, 1896
Male+Female adult
Russia, NW Caucasus,
Krasnodar reg., Chernogorie mts., 24.V-29.VII.1999. A.S. Zamotajlov leg.
female 29mm/male 25mm
Copyright © Ruslan Panin

Carabus (Tribax) circassicus kubaniensis Semenov, 1896
Endophallus
endophallus
Copyright © Ruslan Panin

Carabus (Tribax) circassicus circassicus Ganglbauer, 1886
Male+Female adult
Abkhasia,
W.slope of Gagrinsky Mt., Krepostnaya River source, h=930-980m., A.S.Zamotajlov leg. 27.VII.2000
female, 29 mm/male, 26mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Tribax) circassicus abagonensis Starck, 1894
Male+Female adult
S Russia, NW Caucasus,
Adygea Rep., Mt.Abago, 26.VI.1999
L=24mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Tribax) circassicus circassicus (as tshchaltensis Novotný & Voříšek
Male adult
W-Caucasus: Abkhasia, Gudauta distr., E slope Mt Anchkho h=2200-2350 m, 14-20.VII.2011, S.Pavluchuk, I. Solodovnikov leg./det. I. A. Solodovnikov.
25 mm

Copyright © Ruslan Panin

Carabus (Tribax) kasakorum kasakorum Semenov, 1896
Male adult
male, 26mm
Copyright © Boleslav Brezina

Carabus (Tribax) kasakorum kasakorum Semenov, 1896
Male adult
male, 27mm
Copyright © Boleslav Brezina

Carabus (Tribax) kasakorum kasakorum Semenov, 1896
Female adult
female, 29mm
Copyright © Boleslav Brezina

Carabus (Tribax) kasakorum kasakorum Semenov, 1896
Female adult
female, 29mm
Copyright © Boleslav Brezina

Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
Male adult
male, 23mm
Copyright © Boleslav Brezina

Carabus (Tribax) kasakorum kasakorum Semenov, 1896
Male adult
Pr. Wladikavkas, montes, coll. A. Zolotarew
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Tribax) kasakorum kasakorum Semenov, 1896
Male adult
Montes prope Wladikavkas, 6000 f., coll. A. Zolotarew
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Female adult
Caucasus, N. Osetia, 60 km S Alagir, 1850 m, 12.IX.1990, A. Klimenko leg.<br /> photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Female adult
Cuaucas, N Osetia, Ardon riv., Alagir, Duravtuan, 28.IV.1982, A.Sokolov leg.<br /> photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
Endophallus
S Russia, C-Caucasus: N Ossetia, Chekhatsirtit Mt range, Mt Tepli h=3000 m
Copyright © Ruslan Panin

Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
Male adult
Russia, C-Caucasus: N Ossetia, Chekhatsirtit Mt range, Mt Tepli h=3000 m, 21.VII.2009. /det. Solodovnikov I.A, L=20.5 mm
Copyright © Ruslan Panin

Carabus (Tribax) kasakorum scintillans Rapuzzi, 2018 Holotype
Male adult
Russia, Caucasus, Ingushetia, South slopes of Skalistyi Mountain, Range, Beyni village environs, 2,800 m, 21.V/19.VII.2018, Ivan Rapuzzi collection. Image from Rapuzzi, 2018
Copyright © - - -

Carabus (Tribax) kasakorum kasakorum Semenov, 1896
Male adult
Russia: N.Osetia: Pastbyshchniy Mt.R., S slopes of Khoshkharanrag Mt.R., right bank of Amusadon riv., h=1500 m
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum kasakorum Semenov, 1896
Female adult
Russia: N.Osetia: Pastbyshchniy Mt.R., S slopes of Khoshkharanrag Mt.R., right bank of Amusadon riv., h=1500 m
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Male adult
Russia, Caucasus, Ingushetia, South slopes of Skalistyi Mt Range, Stolovaya Mt., near Beyni, alpin zone, h = 2800 m, N 42°51’35.30",E 44°42’13.70", h = 2800m, 21.V–19.VII.2018, D. Fominykh leg.
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Female adult
Russia, Caucasus, Ingushetia, South slopes of Skalistyi Mt Range, Stolovaya Mt., near Beyni, alpin zone, h = 2800 m, N 42°51’35.30",E 44°42’13.70", h = 2800m, 21.V–19.VII.2018, D. Fominykh leg.
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Endophallus
Russia, Caucasus, Ingushetia, South slopes of Skalistyi Mt Range, Stolovaya Mt., near Beyni, alpin zone, h = 2800 m, N 42°51’35.30",E 44°42’13.70", h = 2800m, 21.V–19.VII.2018, D. Fominykh leg. (TOPOTYPUS)
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Male adult
Russia, Caucasus, Ingushetia, Skalistyi Mt. Range, S slopes of Tsoreylam Mt. Range, 5 km W of Mt. Khakhalgi, N 42°50’07.60", E 45°05’02.20", h=2500 m, 25.V–21.VII.2017, D. Fominykh leg.
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Endophallus
Russia, Caucasus, Ingushetia, Skalistyi Mt. Range, S slopes of Tsoreylam Mt. Range, 5 km W of Mt. Khakhalgi, N 42°50’07.60", E 45°05’02.20", h=2500 m, 25.V–21.VII.2017, D. Fominykh leg.
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Female adult
Russia, Chechnya, Caucasus, Skalystyi Mt. Range upper river Gekhi, Mt. Ekkyrkort, N 42°52', E 45°26', h=2230 m, 26.V–21.VII.2018, D.Fominykh leg.
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
Endophallus
Russia: N.Osetia, Bokovoy Mt.R., near Arkhonskiy pass., alpin zone, h = 2700 m, 23.V-21VII.2018, D. Fominykh leg.
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
Female adult
Russia: N.Osetia, Bokovoy Mt.R., near Arkhonskiy pass., alpin zone, h = 2700 m, 23.V-21VII.2018, D. Fominykh leg.
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
Male adult
Russia: N.Osetia, Bokovoy Mt.R., near Arkhonskiy pass., alpin zone, h = 2700 m, 23.V-21VII.2018, D. Fominykh leg.
Copyright © Igor Solodovnikov

Carabus (Tribax) nothus planipennis Chaudoir, 1846
Female adult
Dagestan, Huizah., near Motchoh, h=1800, 27.07.1993, Ilina leg.
photo A. Menstschicov
Copyright © Anton Menstschikov

Carabus (Tribax) nothus planipennis Chaudoir, 1846
Male adult
male, 22mm
Copyright © Boleslav Brezina

Carabus (Tribax) nothus planipennis Chaudoir, 1846
Male adult
male, 21mm
Copyright © Boleslav Brezina

Carabus (Tribax) nothus planipennis Chaudoir, 1846
Male adult
Dagestan, Bogosskii mt. rng, Inhokvari-Honoh
3500 m, 15-25.6.85 Abdurakhmanov leg.
Copyright © Alexander Anichtchenko

Carabus (Tribax) nothus nothus Adams, 1817
Male adult
male, 24mm
Copyright © Boleslav Brezina

Carabus (Tribax) nothus nothus Adams, 1817
Female adult
female, 24mm
Copyright © Boleslav Brezina

Carabus (Tribax) nothus nothus Adams, 1817
Female adult
female, 25mm
Copyright © Boleslav Brezina

Carabus (Tribax) nothus fausti Dohrn, 1873
Male adult
male, 24mm
Copyright © Boleslav Brezina

Carabus (Tribax) nothus (as abdurachmanovi)
Male adult
Carabus (Tribax) nothus (as abdurachmanovi)
male, 22mm
Copyright © Boleslav Brezina

Carabus (Tribax) nothus (as abdurachmanovi)
Female adult
Carabus (Tribax) nothus (as abdurachmanovi)
female, 23mm
Copyright © Boleslav Brezina

Carabus (Tribax) nothus planipennis Chaudoir, 1846
Male adult
Georgia, Lagodehi, Ninigora, 2750 м, near snow 24.V.1979, I.Belousov leg.
photo from www.zin.ru
Copyright © Dmitri Obydov

Carabus (Tribax) kasakorum daghestanicola Rapuzzi, 2018 Holotype
Female adult
Russia, East Caucasus, Daghestan, Andiysky Mountain Range, Kharami pass, 2,500 m, 5.VII.2010, S. Rybalkin legit, Ivan Rapuzzi collection. Image from Rapuzzi, 2018
Copyright © - - -

Carabus (Tribax) nothus sharoiensis Fominykh, Zamotajlov, Avtaeva et Titarenko, 2019 Paratype
Male adult

Copyright © Igor Solodovnikov

Carabus (Tribax) nothus fausti Dohrn, 1873
Male adult
Carabus (Microtribax) fausti fausti Dohrn, 1873 Russia: Daghestan, Kyabyak Mt.R.
Copyright © Igor Solodovnikov

Carabus (Tribax) nothus fausti Dohrn, 1873
Female adult
Carabus (Microtribax) fausti fausti Dohrn, 1873 Russia: Daghestan, Kyabyak Mt.R.
Copyright © Igor Solodovnikov

Carabus (Tribax) steveni steveni Menetries, 1832
Male adult
Carabus (Tribax) steveni steveni Menetries, 1832
Karachaevo-Cherkessia,
Hudes riv., near Tsarskie vorota,
V.2009 Anichtchenko A. leg.
male, 25mm
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Tribax) steveni steveni Menetries, 1832
Male adult
Carabus (Tribax) steveni steveni Menetries, 1832
C Caucasus, Kabardino-Balkaria,
Elbrusskyi distr., riv. Malka, upper reaches riv. Sultan-Bashi, h=2400-2600 m, 26, VII, 2008. Leg. I.A.Solodovnikov.
male, 20,5 mm
Copyright © Igor Solodovnikov

Carabus (Tribax) steveni steveni Menetries, 1832
Female adult
Carabus (Tribax) steveni steveni Menetries, 1832
C Caucasus, Kabardino-Balkaria,
Elbrusskyi distr., riv. Malka, upper reaches riv. Sultan-Bashi, h=2400-2600 m, 26, VII, 2008. Leg. I.A.Solodovnikov.
female 24,5 mm
Copyright © Igor Solodovnikov

Carabus (Tribax) steveni schamyli Hampe, 1852
Female adult
S Russia, N Ossetia, Alagir distr., near Nizhniy Tsey vill.,VII.1981
female, 25mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Tribax) steveni schamyli Hampe, 1852
Male adult
C Caucasus, N Ossetia,
N slope of Pastbistshnyi Mt.R., Tagardon env., h=1100m, 7-9.08.1997, leg. A.G.Koval
male 25,5 mm
Copyright © Igor Solodovnikov

Carabus (Tribax) steveni schamyli Hampe, 1852
Male adult
C Caucasus, N Ossetia,
N slope of Pastbistshnyi Mt.R., Tagardon env., h=1100m, 7-9.08.1997, leg. A.G.Koval
female 27 mm
Copyright © Igor Solodovnikov

Carabus (Tribax) steveni schamyli Hampe, 1852
Male adult
N Osetia, B. Balkary, 1700 m, 2.9.85
Abdurahmanov leg.
male, 24 mm
Copyright © Alexander Anichtchenko

Carabus (Tribax) steveni satanas Semenov, 1896
Male adult
Carabus (Tribax) steveni satanas Semenov, 1896
S Russia, C-Caucasus:
Balkaria, Valley of riv. Cherek Balkarskyi, h=1750-1800m 10.VI-20.VII.1992 Leg. Zamotajlov A.
male, 23 mm
photo Panin R., det. Solodovnikov I.A
Copyright © Ruslan Panin

Carabus (Tribax) steveni satanas Semenov, 1896
Male+Female adult
Carabus (Tribax) steveni satanas Semenov, 1896
Russia, C.Caucasus, Kabardino-Balkaria,
valley of River Cherek Balkarsky, h=1800 m
10.VI-20.VII.1992
A. Zamotajlov leg.
Copyright © Safronov

Carabus (Tribax) steveni satanas Semenov, 1896
Endophallus
endophallus
Copyright © Ruslan Panin

Carabus (Tribax) steveni schamyli Hampe, 1852
Female adult
C Caucasus, North Ossetia, the road on the village of Coban (Koban), the left river bank Gizeldon, the deciduous wood, h = 895-905 m, N 42°55'32.91", E 44°31'53.53", 20.05-03.07.2017, leg. A.P. Tatulyan
Copyright © Igor Solodovnikov

Carabus (Tribax) steveni schamyli Hampe, 1852
Female adult
C Caucasus, North Ossetia, the road on the village of Coban (Koban), the left river bank Gizeldon, the deciduous wood, h = 895-905 m, N 42°55'32.91", E 44°31'53.53", 20.05-03.07.2017, leg. A.P. Tatulyan
Copyright © Igor Solodovnikov

Carabus (Tribax) steveni schamyli Hampe, 1852
Male adult
C Caucasus, North Ossetia, the road on the village of Coban (Koban), the left river bank Gizeldon, the deciduous wood, h = 895-905 m, N 42°55'32.91", E 44°31'53.53", 20.05-03.07.2017, leg. A.P. Tatulyan
Copyright © Igor Solodovnikov

Carabus (Tribax) steveni schamyli Hampe, 1852
Male adult
C Caucasus, North Ossetia, the road on the village of Coban (Koban), the left river bank Gizeldon, the deciduous wood, h = 895-905 m, N 42°55'32.91", E 44°31'53.53", 20.05-03.07.2017, leg. A.P. Tatulyan
Copyright © Igor Solodovnikov

Carabus (Tribax) titan titan Zolotarev, 1913
Male+Female adult
Russia,NW Caucasus,
Krasnodar reg.,Chernogorie Mnt. Range, Agulova Balka h=950m V-VII.1999, A.S.Zamotajlov leg.
female,32mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Tribax) titan titan Zolotarev, 1913
Endophallus
endophallus
Copyright © Ruslan Panin

Carabus (Tribax) titan titan Zolotarev, 1913
Male adult
Carabus (Tribax) titan titan Zolotarev, 1913
Tuapse distr., Anastasievka vill., Semiglavaya mt.,
Pshekaho riv., 20.7.1995
male, 30 mm
Copyright © Alexander Anichtchenko

Carabus (Tribax) titan abadzechus Zamotajlov, 1988
Male adult
Carabus (Tribax) titan abadzechus Zamotajlov, 1988
W Caucasus, Adygea, Asish-Tau Mt.R., vall riv. Bzykha, Fagus forest, h=1450m, 14-22.08.2008, I.A.Solodovnikov, O.Beljavskyi
male, 25 mm

Copyright © Igor Solodovnikov