Carabus (Acoptolabrus) lopatini Morawitz, 1886
Male+Female adult
Russia,
Sakhalin isl., Tchekhov Mt, VI.1997
female, 40mm
photo & coll. Panin R.
Copyright © Ruslan Panin

Carabus (Acoptolabrus) schrencki schrencki (as hauryi Gehin, 1885 )
Male adult

Copyright © Boleslav Brezina

Carabus (Acoptolabrus) schrencki schrenck (as dongningensis Deuve & Li, 2005)
Male adult
NE China, Heilongjiang prov., Dongning, V.2009<br /> L=25 mm
Copyright © Ruslan Panin

Carabus (Acoptolabrus) constricticollis constricticollis (as grallatorius Roeschke, 1921)
Male adult
NE-Korea:North Hamgyong Prov., Sechon-dong, VII.2004, Li leg.
L=28 mm
Copyright © Ruslan Panin

Carabus (Acoptolabrus) constricticollis pihyonensis Deuve & Li, 1998
Male adult
N-Korea, North Pyongan Prov., Tongnim County, 5-25.V.2010 <br /> 29 mm
Copyright © Ruslan Panin

Carabus (Acoptolabrus) constricticollis Kraatz, 1886
Male adult
China, Liaoning prov., Mt. Guanmen 20.6.2009 (cAA)
Copyright © Alexander Anichtchenko

Carabus (Aulonocarabus) gossarei imanensis Lafer, 1989
Male adult

Copyright © Kirill Makarov

Carabus (Aulonocarabus) gaschkewitschi czarensis Obydov, 1997
Female adult
Russia, CE-Siberia, N Chita reg., Kodar Mt. Range, Big Leprindo Lake env. h~2000m., 2-5.VII.2002 A. Prokhorov leg./det. A. Anichtchenko
L=19 mm
Copyright © Ruslan Panin

Carabus (Aulonocarabus) canaliculatus hailinensis Deuve & Li, 2000
Male adult
NE China, Heilongjiang Prov., Mudanjiang City, Mt.Tai-Gu-Ding-zi h=800m, 15-25.VI.2010. L=29 mm
Copyright © Ruslan Panin

Carabus (Aulonocarabus) gaschkewitschi gaschkewitschi Motschulsky, 1859
Male adult

Copyright © Kirill Makarov

Carabus (Aulonocarabus) kolymensis kodarensis Brinev, 1997
Male adult
Russia, SE Siberia, N Chita reg., Kodar Mt. Rng. h=1800m, Novaya Chara vill., 23.VI.2006. L=17 mm
Copyright © Ruslan Panin

Carabus (Aulonocarabus) truncaticollis fleischeri Reitter, 1898
Female adult
Russia, N Burajatia, Barguzinskiy Mts., the upper reaches of M.Chivyrkuy riv., h=2000 m, 26.VI-12.VII.2007, Vashchenko S leg. L=18.5 mm
Copyright © Ruslan Panin

Carabus (Aulonocarabus) truncaticollis fleischeri Reitter, 1898
Male adult
Russia, N Burajatia, Barguzinskiy Mts., the upper reaches of M.Chivyrkuy riv., h=2000 m, 26.VI-12.VII.2007, Vashchenko S leg. L=17 mm
Copyright © Ruslan Panin

Carabus (Carabus) arvensis arvensis Herbst, 1784
Male+Female adult
W Ukraine, NW Podolian, Lviv reg., Javoriv distr., near Stradch vill., 20-30.IV.2009, Panin R. leg.
Copyright © Ruslan Panin

Carabus (Carabus) arvensis arvensis Herbst, 1784
Male adult
W. Ukraine , NW. Podolian, Lviv reg., Nikolaev distr., near Krasiv vill., 9.II.08, Panin R. leg. L=17mm
Copyright © Ruslan Panin

Carabus (Carabus) granulatus leander Kraatz, 1878
Male+Female adult
Russia, C-Caucasus: Balkaria, Valley of riv. Cherek Balkarskyi, h=1750-1800m 10.VI-20.VII.1992, Zamotajlov A. leg.
female,22mm
Copyright © Ruslan Panin

Carabus (Carabus) granulatus granulatus Linne, 1758
Male adult
W Ukraine,
Lviv Reg., Javoriv distr., near Ivano-Frankovo vill., 30.IV-18.V.2009, Panin R. leg.
male, 21mm
Copyright © Ruslan Panin

Carabus (Carabus) granulatus granulatus Linne, 1758
Female adult
long. 22mm
Copyright © Ruslan Panin

Carabus (Carabus) menetriesi menetriesi Hummel, 1827
Male adult
W-Ukraine, Roztochia, Lviv Reg., Yavorov distr., near Ivano-Frankovo vill., Site Zalyvki, Panin R. leg., 1-20.V.2005, male, 21 mm.
Copyright © Ruslan Panin

Carabus (Carabus) stscheglowi Mannerheim, 1827
Male+Female adult
Central Russia , Voronezh region, near Grafskoe, 12.VIII.1984 long=20mm
Copyright © Ruslan Panin

Carabus (Carabus) arvensis faldermanni Dejean, 1829
Male+Female adult
Russia, Far East, Primorskiy Kray, Ussuriysk Distr., Kaymanovka vill., 11.VII-1.VIII..2011, A. Korshunov leg.
female, 23 mm
Copyright © Ruslan Panin

Carabus (Carabus) stscheglowi Mannerheim, 1827
Male adult

Copyright © Kirill Makarov

Carabus (Carabus) granulatus granulatus Linne, 1758
Male adult
W-Ukraine, Roztochia,
Lviv Reg., Javorov distr., near Ivano-Frankovo vill., Site Zalyvki, 17.IV-3.V.2010, Panin R. leg.
L=20 mm

Copyright © Ruslan Panin

Carabus (Carabus) arvensis arvensis Herbst, 1784 (as noricus Sokolar, 1910)
Female adult
CZ, C Moravia, Dřevohostice - Líšná, 13mm
Copyright © Aleš Sedláček

Carabus (Carabus) granulatus granulatus (as hibernicus Lindroth, 1956)
Female adult
W-Ireland, County Mayo, Letterneevoge (R312),16-30.VI.2013, J.Stypula leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Carabus) arvensis hokkaidoensis Lapouge, 1924
Female adult

Copyright © Kirill Makarov

Carabus (Carabus) granulatus yezoensis Bates, 1883
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Carabus (Carabus) menetriesi menetriesi Hummel, 1827
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Carabus (Carabus) stscheglowi Mannerheim, 1827
Male adult
Russian, Chuvashia, Cheboksary, Zavolzhye distr., , near lake Astrakhanka, L=27 mm
Copyright © Ruslan Panin

Carabus (Carabus) arvensis arvensis Herbst, 1784
Female adult
NW Ukraine, Polesia, Rovno reg., Rokytne distr., near Bil’sk vill., 4-6.VII.2015, Panin R. leg. L=20 mm
Copyright © Ruslan Panin

Carabus (Carabus) arcensis arcensis Herbst, 1784
Female adult
Lower Austria, Baden, Klausen-Leopoldsdorf, 15.XII.2024, M. Egger leg. L=22 mm
Copyright © Ruslan Panin

Carabus (Carabus) menetriesi menetriesi Hummel, 1827
Endophallus
W Ukraine, Lviv city, Landscape Reserve “Peatbog Bilogorshcha”
Copyright © Ruslan Panin

Carabus (Carabus) menetriesi menetriesi Hummel, 1827
Male adult
W Ukraine, Lviv city, Landscape Reserve “Peatbog Bilogorshcha” NW part, 17.IV-14.V.2018, Panin R. leg., L=21.5 mm
Copyright © Ruslan Panin

Carabus (Procechenochilus) kokujewi Semenov, 1898
Female adult
RUSSIA, Karachay-Cherkessia, NW Caucasus Mts., Daut Mt. Range, Iskrovtsev - W.Ullu-Kel Mt. pass circus, h=2800-2600m
Copyright © Alexander Anichtchenko

Carabus (Procechenochilus) ? kokujewi Semenov, 1898
Female adult
Russia, NW Caucasus, Karachay-Cherkessia, Karachaevsk distr., near Uchkula vill., valley Uchkulanichi riv. h=2800-2900m, 25-28.VII.2010, Fominykh D. leg.
female, 23 mm
Copyright © Ruslan Panin

Carabus (Cechenochilus) heydenianus Starck, 1889
Male+Female adult
NW-Caucasus,
Krasnodar reg., Mostovskoy distr., Skirda Mt Range, N slopes Yatyrgvart Mt. h=2300-2500m., 09-10.VII.2008, D. Fominykh leg.
female,19mm/male,18mm
Copyright © Ruslan Panin

Carabus (Cechenochilus) adangensis Gottwald, 1983
Female adult
Russia, NW-Caucasus:
Karacaj-Cherkessija, Abishira–Akhuba Mt. Range h=2000-2650m, Kyafar-Agur Riv. 14–17.VII.2009, Fominykh D. leg.
female,21mm
Copyright © Ruslan Panin

Carabus (Cechenochilus) kokujewi Semenov, 1898
Male adult
Russia, Karachai-Cherkessia, vall. Daut Riv., near Glacier Ryndzhi-Murudzhu, h = 3050 m, 22.VIII.2011, leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) kokujewi Semenov, 1898
Female adult
Russia, Karachai-Cherkessia, vall. Daut Riv., near Glacier Ryndzhi-Murudzhu, h = 3100 m, 22.VIII.2011, leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) kokujewi Semenov, 1898
Male adult
Russia, Karachai-Cherkessia, vall. Daut Riv., near Glacier Ryndzhi-Murudzhu, h = 3050 m, 22.VIII.2011, leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) kokujewi Semenov, 1898
Female adult
Russia, Karachai-Cherkessia, vall. Daut Riv., near Glacier Ryndzhi-Murudzhu, h = 2550 m, 23.VIII.2011, leg. I.A. Solodovnikov, S.V. Solodovnikova
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) adangensis Gottwald, 1983
Female adult
Russia, Karachai-Cherkessia, Abischira-Akhuba Mt.R., env. vill. Romantic, alpin zone, h = 2470 m, 10.07-22.08.2019, leg. S.M. Pavlyuchuk, A.P. Tatulyan
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) boeberi boeberi Adams, 1817
Female adult
Грузия, Мцхета-Мтианети, к северу от пос. Гудаури, ЮВ отроги г. Байдарская (нижний пик), по ущелью ручья, альпика, под камнями, точка 2, h = 2700 м, 42.495413° N, 44.487844° E, 10.08.2019, leg. И.А., А.И., С.В. Солодовниковы, О.И. Данилова 26 mm
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) heydenianus Starck, 1889
Male adult
Carabus (Procechenochilus) heydenianus euxinus Semenov, 1896 Abkhasia: Arabika plato (TOPOTYPUS)
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) heydenianus Starck, 1889
Female adult
Carabus (Procechenochilus) heydenianus euxinus Semenov, 1896 Abkhasia: Arabika plato (TOPOTYPUS)
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) boeberi aequaliceps Reitter, 1896
Male adult
Russia, Dagestan, Andiyskiy Mt.R., near Andiyskiye vorota pass, 06.VI–18.VII.2021, leg. D. Fominykh
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) boeberi aequaliceps Reitter, 1896
Female adult
Russia, Dagestan, Andiyskiy Mt.R., near Andiyskiye vorota pass, 06.VI–18.VII.2021, leg. D. Fominykh
Copyright © Igor Solodovnikov

Carabus (Cechenochilus) boeberi felix Semenov & Znojko, 1931
Male adult

Copyright © Kirill Makarov

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Male adult
W Ukraine, Lviv city, Sykhiv, Zubra forest, (Osvytskiy Grove), under bark, 49°47'28.10" N, 24° 2'20.09" E, 10.XI.2017, Panin R leg.
Copyright © Ruslan Panin

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Female adult
W Ukraine, Lviv city, Holosko, Bryukhovytsʹkyi forest 49.886833, 24.012083 under bark, 10.XI.2024, R. Panin leg., L=35 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) smaragdinus mandschuricus Semenov, 1898
Male adult
Russia, Far East, South Primoriye, Kaymanovka village, 10-20.07.2000; P.Gorbunov leg.
Copyright © Anton Menstschikov

Carabus (Coptolabrus) jankowskii jankowskii Oberthur, 1883
Male adult
N.Korea, Chagang Do, Mt. Langlin, 30.07.2005
Copyright © Anton Menstschikov

Carabus (Damaster) blaptoides rugipennis Motschulsky, 1861
Male adult
Kunashir isl.
Copyright © Kirill Makarov

Carabus (Diocarabus) loschnikovi loschnikovi Fischer, 1823
Male adult
Komi Resp., Pecero-Ilych Nat. Res., Zajachij Kamen h~786 62°10'06" N 59°12'53" E 8.VIII.2004 M. Chernjakhovski leg.
L=15.0 mm
Copyright © Evgeniy Zinoviev

Carabus (Diocarabus) loschnikovi tsaganensis Obydov, 2001
Male adult
Russia, W Tuva,
Kyzyl-Tayga Mt Range h=1500 m, Ak-Suk riv vall., 21.VI.2006, A.& N. Menstschikovy leg.
L=16 mm
Copyright © Ruslan Panin

Carabus (Diocarabus) slovtzovi slovtzovi Mannerheim, 1849
Male adult
S Siberia, Burjatia reg., E Sayan Mts., Khugaituma mt., h=1700 m, 1-5. VII.2001
L=14,3 mm

Copyright © Maksim Smirnov

Carabus (Diocarabus) beybienkoi Kryzhanovskij, 1973
Male adult
S Sakhalin, Susunaj Mt.R. 10.08 - 13.09.2003 D. Kochetkov leg.
L=14,5 mm
Copyright © Maksim Smirnov

Carabus (Diocarabus) fraterculus fraterculus Reitter, 1895
Male adult
N-Korea, N Pyongan Prov.,Sakju, Mt Ponsan, 20-29.VI.2009, L. Bieber leg.<br /> L=19 mm
Copyright © Ruslan Panin

Carabus (Diocarabus) aurocinctus Motschulsky, 1846
Female adult
det. K. Makarov
Copyright © Alexander Anichtchenko

Carabus (Diocarabus) opaculus opaculus Putzeys, 1875
Female adult

Copyright © Kirill Makarov

Carabus (Diocarabus) massagetus massagetus Motschulsky, 1846 (alpin form)
Male adult
Russsia, Altay, Ukok upland, Kaldzhin-Kul Lake h=2400-2550 m, 49 *19*N/87*26*E, 16-27.VII.2008, R.Dudko leg. L=17.5 mm
Copyright © Ruslan Panin

Carabus (Diocarabus) fraterculus neochinensis Deuve & Li, 1998
Female adult
NE-China: Liaoning prov., Fengcheng City, Shicheng, 20-28.VI.2011, Li Jingke leg., L=22 mm
Copyright © Ruslan Panin

Carabus (Diocarabus) fraterculus neochinensis Deuve & Li, 1998
Male adult
NE-China: Liaoning prov., Fengcheng City, Shicheng, 20-28.VI.2011, Li Jingke leg., L=20 mm
Copyright © Ruslan Panin

Carabus (Eupachys) glyptopterus Fischer, 1827
Male+Female adult
Russia, SE-Siberia,
Buryat Republic, Ivolginsky Distr., 04.VII.2010, Filippov A.E. leg.
female, 32 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Hemicarabus) tuberculosus Dejean, 1829
Male adult
Far East, Lazovsky Dist., shore of Japan Sea, 3 km NE Preobrazhenie Vill., Ezhovaya (Tasovaya) Bay 13.07.06 M. & L. Smirnov's leg.
male
L=20,0 mm
Copyright © Maksim Smirnov

Carabus (Hemicarabus) nitens Linne, 1758
Male+Female adult
Belarus,
Vitebsk reg., Ushachskiy distr., near Bolshie Dolcy vill. 9.V.1990, I.A.Solodovnikov leg.
female, 18mm.
Copyright © Ruslan Panin

Carabus (Homoeocarabus) maeander maeander s.str.
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Carabus (Homoeocarabus) maeander paludis Gehin, 1885
Female adult
Japan, Hokkaido; Tomakomai city, Benten, 17.V.2015, Takemoto leg., L=21 mm
Copyright © Ruslan Panin

Carabus (Leptocarabus) arboreus pararboreus (Ishikawa, 1992)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Carabus (Limnocarabus) clathratus clathratus Linne, 1761
Female adult
W-Ukraine, Maloe Polesie, Lviv Region, near Kamenka-Bugzkaya, 28.III.2008, Panin R. leg. L=31 mm
Copyright © Ruslan Panin

Carabus (Limnocarabus) clathratus clathratus (as jansoni Kraatz, 1890)
Male adult
W-Ireland, County Mayo, Bellacorick bog, 9.VII.2013, J.Stypula leg. L=27 mm
Copyright © Ruslan Panin

Carabus (Limnocarabus) clathratus clathratus Linne, 1761
Female adult

Copyright © Kirill Makarov

Carabus (Megodontus) violaceus aurolimbatus Dejean, 1829
Male adult
C.UKRAINE,
Poltava reg., near Novi Sanzhary, bank of Vorskla River, 1-2.VII.07, D.Kovalchuk leg.
male 25mm
Copyright © Ruslan Panin

Carabus (Megodontus) schoenherri schoenherri Fischer, 1822 (as sangilenus Obydov, 1997)
Male adult
Tyva, Akademika Obrucheva mt.rng., 15km N Cherbi vill.,
Chailag ur., 7.7.1999 A. & N. Menshikovy leg.
male, 26mm
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Megodontus) exaratus Quensel, 1806
Male adult
Russia, C.Caucasus, N.Ossetia,
valley of River Khazniaush,
h=1400 m, 10.VI-31.VII.1993
A. Zamotajlov leg.
Copyright © Safronov

Carabus (Megodontus) exaratus Quensel, 1806
Male+Female adult
NW Caucasus, Russia,
Krasnodar reg., Krasnodar City, 27.IX-07.X.07, D.D.Fominykh leg.
female,29mm/male, 27mm
Copyright © Ruslan Panin

Carabus (Megodontus) danilevskii Obydov, 1993
Male adult
Krasnoyarsk Area, Sayanogorsk Distr, env. Cheremushki Vill., right bank of Enisej Riv., h= 500 - 800 m, 23.07.1999 A. Semin leg.
L=17,0 mm
Copyright © Maksim Smirnov

Carabus (Megodontus) kolbei aino Rost, 1908
Male adult

Copyright © Kirill Makarov

Carabus (Megodontus) vietinghoffi bowringi Chaudoir, 1863
Male adult
Russia, Far East, Primorskiy Kray, Ussuriysk Distr., Kaymanovka vill , 26.VII-3.VIII.2011, A. Korshunov leg.
L=29
Copyright © Ruslan Panin

Carabus (Megodontus) obovatus Fischer, 1827
Male adult
SW Siberia, Russia, Kemerevo reg., Krapivino distr., Azhendarovo, forest, 26-31.VII.2011, D. Efimov leg.
L=18 mm
Copyright © Ruslan Panin

Carabus (Megodontus) vietinghoffi bowringi (as lazoensis Obydov, 1999)
Male+Female adult
Russia, Far East, Primorsky reg., S. Sichote-Alin Mts.R., Kavalerovo vill. env.,
5-25.VII.1992, S. Kunitzyn leg.
male - 28 mm, female - 30 mm.
Copyright © Alexandr Safronov

Carabus (Megodontus) vietinghoffi bowringi Chaudoir, 1863
Male+Female adult
Russia, Far East, Primorsky reg., Khasansky distr.,
Zanadvorovka vill., Gusevsky pit.
M - 19-21.VI.2007, body lenght 29 mm.
F - 15.VII.2008, body lenght 30 mm.
S. Veriga leg.
Copyright © Alexandr Safronov

Carabus (Megodontus) danilevskii Obydov, 1993
Male+Female adult
Russia, SW Siberia, Krasnoyarsk reg., W. Sayan, near Cheryomushki.
26-29.VII.1996 loc. coll.
male - 18mm, female - 20 mm.
Copyright © Alexandr Safronov

Carabus (Megodontus) kantaikensis Gehin, 1885
Female adult
Russia, SW Siberia, W. Sayan, Krasnoyarsk reg.,
S of Abakan, near Cheryomushki, Dzhoysky Pass.
07-VII-1994 M. Danilevsky leg.
body lenght 18 mm.
Copyright © Alexandr Safronov

Carabus (Megodontus) obovatus Fischer, 1827
Male adult
SW Siberia, Kemerovo reg., Krapivino distr.,
8 km SSW Saltymakovo vill., h=150 m.
20-30.VI.2010 D. Efimov, A. Korshunov leg.
body lenght 18 mm.
Copyright © Alexandr Safronov

Carabus (Megodontus) imperialis Fischer, 1823
Male adult
NE Kazakhstan, Zyrjanovsk distr., Putincevo vill. env.
25-VI-2005 M. Danilevsky leg.
male 24 mm, female 26 mm.
Copyright © Safronov

Carabus (Megodontus) violaceus aurolimbatus Dejean, 1829
Male adult

Copyright © Kirill Makarov

Carabus (Megodontus) kolbei taikinus Ishikawa, 1993
Male adult
N-Japan, SC-Hokkaido:Tokachi, Hiroo Distr., Taiki-cho env., 28.VIII-2.IX.2017, Teruo Hirabayashi leg. L=24 mm
Copyright © Ruslan Panin

Carabus (Megodontus) obovatus Fischer, 1827
Male adult
C.obovatus taizhensis Obydov, 2006 Khakassia: Kuznetskyi Alatau, env. vill. Priiskovyi, Bobrovaya Mt., h = 1200-1300 m
Copyright © Igor Solodovnikov

Carabus (Megodontus) obovatus Fischer, 1827
Endophallus
C.obovatus taizhensis Obydov, 2006 Khakassia: Kuznetskyi Alatau, env. vill. Priiskovyi, Bobrovaya Mt., h = 1200-1300 m
Copyright © Igor Solodovnikov

Carabus (Megodontus) kolbei taikinus Ishikawa, 1993
Female adult
Japan, SC-Hokkaido:Tokachi, Hiroo Distr., Taiki-cho env., 3-7.IX.2019, S.Shinichi leg.
Copyright © Ruslan Panin

Carabus (Megodontus) leachi leachi Fischer, 1823
Female adult
NE Kazakhstan, W Altay, near Shemonaikha city., flood plain, 18-28.VII.2018, S.Vashchenko leg., L=22.5 mm
Copyright © Ruslan Panin

Carabus (Megodontus) leachi leachi Fischer, 1823
Male adult
NE Kazakhstan, W Altay, near Shemonaikha city., flood plain, 18-28.VII.2018, S.Vashchenko leg., L=21.5 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus harcyniae Sturm, 1815 (as obenbergerianus Breuning, 1927)
Male adult
Czech, Bohemia, Plzeň Reg., Domažlice Distr., Babylon, h=500 m, 30.VIII.2008, S.I. Budka leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus harcyniae (as procedens Csiki, 1927)
Male adult
England, North Yorkshire, Fylingdales Moor, Grouse Hill, VII.2006 L=24 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus harcyniae (as procedens Csiki, 1927)
Male adult
W-Ireland, County Mayo, Clare Isl., 17-28.VI.2002, S. McCormack leg.<br /> L=25 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus harcyniae (as procedens Csiki, 1927)
Female adult
N-Ireland, Fermanagh Distr., Lough Navar Forest, 17.VII.2013, D.Novak & J.Stypula leg.<br /> L=27mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus harcyniae (as belgicus Lapouge, 1913)
Male adult
Luxembourg, Bollendorf-Pont env., 3.III.1999 /det.R Panin
L=24 mm
Copyright © Ruslan Panin

Carabus (Microplectes) riedeli Menetries, 1832
Female adult
female, 19mm
Copyright © Boleslav Brezina

Carabus (Microplectes) riedeli Menetries, 1832
Male adult
N-Ossetia,
Khazniaush Pass, h=2500-2700m, 9.VI-31.VII.1994, Zamotajlov A.leg.
male, 17mm
Copyright © Ruslan Panin

Carabus (Microplectes) convallium convallium Starck, 1889
Male adult
male, 19mm
Copyright © Boleslav Brezina

Carabus (Microplectes) convallium convallium Starck, 1889
Male+Female adult
female:NW-Caucasus, Russia,
Krasnodar reg., Apsheron Distr., Lagonaki Mt., near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominyh leg.
male:Karachaevo-Cherkessia,the right source of the river Beskess, h=2100m, 10.VIII.1997, I.A. Solodovnikov leg.
female, 21mm/male, 19mm
Copyright © Ruslan Panin

Carabus (Microplectes) convallium argonautarum Semenov, 1898
Male adult
male, 18mm
Copyright © Boleslav Brezina

Carabus (Morphocarabus) wulffiusi wulffiusi Morawitz, 1862
Male+Female adult
Russia, Far East,
Primorskiy Kray, near Kamenushka vill., 2-3.VIII.1992, K.E.Dovgaylo leg.
female, 24mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) venustus venustus Morawitz, 1862
Male adult
SE Primorskii reg., Lazovskii Nat.Res., 12 km S Staraya Kamenka vill., near Amerika kor. 43°15N, 134°1E, 19.VII.04 leg. M.& L. Smirnovy photo from www.zin.ru
Copyright © Maksim Smirnov

Carabus (Morphocarabus) spasskianus spasskianus Breuning, 1932
Male+Female adult
SE Siberia, Russia
Irkutsk reg., near Pivovarikha vill., 7.V.07, D. Fominyh leg.
male, 24mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens (as lomnitzkii Reitter, 1896 )
Male adult
W-UKRAINE, NW Podolian, Lviv Reg., Zolochiv Distr., W Ukraine, Lysa Gora steppe massif, 19.V.-7.VI.2007, Panin R. leg. 25 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens Fabricius, 1798 (ab.erythromerus Dejean, 1826)
Male adult
Moldova, Anenii Noi distr., near Garbovets vill. 20.VI.08 Bodnar' I.I.leg. male, 26mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens (as lomnitzkii Reitter, 1896)
Male adult
W Ukraine, NW Podolian, Ternopil reg., Pidvilochisk distr., near Rosokhuvatets & Kamyanki vill., steppe sites, 24.V-15.VI.2008, Panin R. leg.
male, 24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) odoratus bargusinicus Shilenkov, 1996
Male+Female adult

    Carabus (Morphocarabus) odoratus bargusinicus Shilenkov, 1996
    Russia, N Burajatia,
    Barguzinskiy Mts., the upper reaches of M.Chiverkuy riv., h=2000m, Vashchenko S. leg.
    female, 21mm/male,19mm
    photo Panin R.

Copyright © Ruslan Panin

Carabus (Morphocarabus) odoratus antropovi Shilenkov 1996
Male+Female adult
Russia, S Tuva, Khorumnug -Tayga mts. Range, h=1600-2300m, Buren riv. valley, 4-26.VI.2000 Vashchenko S & A. leg<br /> female, 20mm/male,18mm<br /> photo Panin R.
Copyright © Ruslan Panin

Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
Male adult
Russia, S Siberia,
S Khakassia, N Tuva reg., Sailyg-Khem-Taiga mts. range h=2400m, 10.VII.2004
male, 18mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996
Male+Female adult
Russia, S Siberia,
S Khakassia, N Tuva reg., Sailyg-Khem-Taiga mts. range h=2000-2600m,10-30.VI.2005, S.Vashchenko & S.Alexeenko leg.
male,17.5mm/female,19mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Morphocarabus) mestscherjakovi pseudoodoratus Brinev, 2002 Paratype Paratype
Female adult
Krasnojarsk reg., Aradan distr., 5 km SW Aradan vill., vall. of Us riv., A.Brinev leg.<br /> female, 20 mm<br /> photo & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) kozhantschikovi martjanovi Lutshnik, 1924
Female adult
Khakassia, 1800 m, near Cheremushki vill., 12.6.-20.7.2000<br /> female, 21 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
Male adult
W Sayan Mts., Kazhyr-Suk riv., 80 km W Oleniya<br /> Rechka, W part of Oiskii mt. rng., 30.6.-7.7.1999<br /> 2100 m, Brinev A. leg.<br /> male, 19 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) hummeli nevelskii Shilenkov, 1996
Male+Female adult
Russia, Sakhalin isl.,
Makarov distr., Lesnaja riv., VII.1992, M. Nesterov leg.
female,23mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) aeruginosus verzhutzkii O.Berlov & Shilenkov, 1996
Male adult
C Altai Mts., Seminskii Mt.rng., Katun riv., 3 km S Ust'-Sema riv.,
4.VII.2002 A. & N. Menschikovy leg.
det. Mentschikov A.
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
Male adult
SE Siberia, E Sayan, Nizhneudinsk, h=500m, 21-26.VI.2006,
male, 26mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus aeruginosiformis Breuning, 1932
Male adult
Russia, Khakassia,
W-Sayan, Alan Mt range, valley Enisey riv., 5 km S Mayna vill., 15.VII.2000, A.& N. Menstschicovy leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924
Male+Female adult
Russia, S Siberia, W Sayan, Ergaki Mt range, Nizhnyaya Buyba river h=1700 m, 19.VII.1999, A.& N. Menstschikovy leg.
Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus shoriensis Obydov, 1999
Male adult
Russia, Khakassia, Abakanskii Mt Range, Askiz river vall., 7 km E Kazanovka vill., 15.VI.2006, A.& N. Menstschikovy leg. L=24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997
Male+Female adult
Russia,Tuva, Akademika Obrucheva Mt range, 15 km N Cherbi vill., Chailang ur. 7.VII.1999, A & N. Menstschikovy leg. L=23 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus geblerianus Obydov, 1999
Male adult
Russia, NW Altay, Barnaul env.,23.07.1998, D.Lomakin leg.

Copyright © Ruslan Panin

Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859
Male adult
Russia, SE Siberia, Transbaikalia, N Chita reg., Kodar Mt. Range, Big Leprindo Lake env.h~1800m., 1.VII.2006, S. Rybalkin leg. L=20 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens (f.i. frivaldszkyi Kraatz, 1887)
Female adult
W-UKRAINE, NW Podolian Upland,<br /> Lviv Reg., Zolochiv Distr., Lysa Gora steppe massif, 14-30.V.2008, Panin. R. leg./det.<br /> L=26 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens Fabricius, 1798
Male adult
C- UKRAINE, NW Odessa reg., Kodyma distr., near Aleksandrivka vill.,18.V.2003, Demidov G.N. leg.<br /> L=26 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878 (as viridipennis Kraatz, 1886, nec Lewis,18
Male+Female adult
Russia, Far East,Primorsky reg., S. Sikhote-Alin Mts. R., Kavalerovo vill. env.,
5-25.VII.1992, С. Kunitzyn leg.
male - 25 mm, female - 25 mm.
Copyright © Alexandr Safronov

Carabus (Morphocarabus) hummeli pusongensis Imura, 1993
Male+Female adult
Russia, Far East, Primorsky ter., Khasansky distr., Zanadvorovka vill.,
Gusevsky pit, S.Veriga leg.
male - 13.VII.2008, female 19-21.VI.2007
male - 28 мм, female - 28 мм.
Copyright © Alexandr Safronov

Carabus (Morphocarabus) hummeli hummeli Fischer, 1823
Male+Female adult
Russia, Far East, Khabarovsk reg., Obluchye env.
M. Khingan Mts. R.
10-20.VII.1992 S. Andreev leg.
male - 26 mm, female - 27 mm.
Copyright © Alexandr Safronov

Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
Male+Female adult
SW Siberia, Kemerovo reg., Krapivino distr.,<br /> 8 km SSW Saltymakovo vill., h=150 m.<br /> 1-20.VII.2010 D. Efimov leg.<br /> male - 25 mm, female - 26 mm.
Copyright ©

Carabus (Morphocarabus) venustus venustus Morawitz, 1862
Male adult
Russia, Far East, Primorskiy reg., Ussuriysk Distr., Kaymanovka vill., 6-15.VIII.2011, <br /> A. Korshunov leg.<br /> L= 21.5 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) wulffiusi dekraatzi Kraatz, 1881
Male adult
NE-China, Heilongjiang, Wuchang city, Mt Zheng-nan-gou, 1-11.VI.2011<br /> L=22 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens ( as jasilkowskii Born,1905)
Male adult
NE Romania, 5 km W Suceava, 47°63′940″N 26°18′795″E,h=340m, agricultural field, 15-20.V.2011, B. Brezina leg. L=24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens (as moldaviensis Born, 1903)
Female adult
E-Romania, Vaslui reg.,15 km SE Vaslui, Crasna env., V-VI,2013, M,Pasek leg. L=26 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) henningi henningi Fischer, 1817
Male adult

Copyright © Kirill Makarov

Carabus (Morphocarabus) venustus venustus Morawitz, 1862
Male adult

Copyright © Kirill Makarov

Carabus (Morphocarabus) odoratus baeri Menetries, 1851: 45
Male adult
Russia, SW Taymyr, near Talnakh, 25 km N Norilsk, hilly tundra, hillside overgrown with alder slate, dwarfish birches, blueberry, h = 150-250 м, 24-28.06.2003, leg. S.Saluk
Copyright © Igor Solodovnikov

Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859
Female adult
Russia, SE Siberia, N Chita reg., Kodar Mt. Range, Big Leprindo Lake env., h~1800 m., L=21 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) viridilimbatus panini I.Solodovnikov 2017 Holotype
Male adult
Russia, SE Yakutia, Neryungri distr., Tokinski Stanovik Mt Rng., Beder riv., (right tributary rivers Algama), 26.VI.1996, Krasnov M. leg., 17 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens (ab.goldeggi Duftschmid, 1812)
Male adult
W-Ukraine, Podolian Tovtry, Ternopol reg., Pidvolochysk distr., Shevchenkove vill., steppe sites, 6.V-7.VI.2012, Panin R. leg. L=24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
Male adult
Russia, S Tuva, Khorumnug-Taiga Mts, Buren riv. Valley, Buren-Bay-Khaak vill. h=900 m, 17-24.VI.2002, Vashchenko A.& B.leg.,/ det. Panin R. L=25 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) henningi cf. saldaitisi Obydov 1997
Female adult
Russia, S Tuva, E Tanu-Ola Mts., Cha-Ova-Art Pass, h=1700m, 28.VI-3.VII.2000, Vashchenko S. leg. L=20 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) henningi cf saldaitisi Obydov 1997
Male adult
Russia, S Tuva, E Tanu-Ola Mts., Cha-Ova-Art Pass, h=1700m, 28.VI-3.VII.2000, Vashchenko S. leg. L=19.5 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) henningi pelevini Obydov 2006
Female adult
E Kazakhstan, S Altay, Sarym-Sakty Mt Range, 12 km SE Ornek vill. h=1200m, 10.VII.1998, L=21 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) henningi pelevini Obydov 2006a: 99
Male adult
E Kazakhstan, S Altay, Sarym-Sakty Mt Range, 12 km SE Ornek vill. h=1200m, 10.VII.1998, L=20 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) henningi henningi Fischer, 1817
Male adult
Russia, S Siberia, NW Buryatia, E Sayans Mts., Tunka Range, near Mondy vill., h=2400 m, alpine tundra, 1-15.VI.2001, Mishustin R. leg./det. R.Panin
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens (as lomnitzkii Reitter, 1896)
Male adult
W Ukraine, NW Podolian Upland, Lviv reg., Zolochiv distr., Lysa Gora steppe massif, 11.VII-14.VIII. 2021, Panin R. & D. Khrapov leg., L=25 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens (as lomnitzkii Reitter, 1896)
Male adult
W Ukraine, NW Podolian Upland, Lviv reg., Zolochiv distr., Lysa Gora steppe massif, 11.VII-14.VIII. 2021, Panin R. & D. Khrapov leg., L=25 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens (as lomnitzkii Reitter, 1896)
Female adult
W Ukraina, Podolian Upland, steppe reserve Kasova Gora near Burshtyn vill., 15.VI -28.VII.2020, R. Panin & D. Khrapov leg., L=27 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) excellens excellens (as lomnitzkii Reitter, 1896)
Male adult
W Ukraina, Podolian Upland, steppe reserve Kasova Gora near Burshtyn vill., 15.VI -28.VII.2020, R. Panin & D. Khrapov leg., L=24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus cf putus Motschulsky, 1846
Female adult
Russia, SW Siberia, C-Altays Mts, Kuray Mt Rng NE Aktash vill., h=2100-3000 m, 8-13.VII.2022, E.Filippov leg./det.R Panin, L= 22 mm (alpine petrophilic population close to the form schestopalovi Plutenko, 1995)
Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus cf putus Motschulsky, 1846
Male adult
Russia, SW Siberia, C-Altays Mts, Kuray Mt Rng NE Aktash vill., h=2100-3000 m, 8-13.VII.2022, E.Filippov leg./det.R Panin, L= 23 mm (alpine petrophilic population close to the form schestopalovi Plutenko, 1995)
Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus putus Motschulsky, 1846 (loc.typ.)
Female adult
Russia, W Altay, 18 km N of Zmeinogorsk, 23.VI.1998, A.& N. Menstschikovy leg. L=24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) spasskianus putus Motschulsky, 1846 (loc.typ.)
Male adult
Russia, W Altay, 18 km N of Zmeinogorsk, 23.VI.1998, A.& N. Menstschikovy leg. L=23 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) odoratus antropovi Shilenkov 1996
Female adult
Russia, S Siberia, Tuva, W Tannu-Ola Mts. 35 km NE Khandagayty vill., Teeli Pass 2200-2800 m, 23-25.VI.2021, A.& E.Filippovs leg. L=19 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) odoratus antropovi Shilenkov 1996
Male adult
Russia, S Siberia, Tuva, W Tannu-Ola Mts. 35 km NE Khandagayty vill., Teeli Pass 2200-2800 m, 23-25.VI.2021, A.& E.Filippovs leg. L=19 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822
Male adult
Russia, NE Altay, Turochak Distr., Lake Teletskoe N part, near Yaylyu vill. 23-25.VII.2019, Ivliev P. leg./ det. R. Panin, L=24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) aeruginosus eschscholtzi Mannerheim, 1827
Female adult
NE Kazakhstan, W Altay Mts, near Shemonaikha city., flood plain, 25-28.VII.2018, S.Vashchenko leg. L=25 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) aeruginosus eschscholtzi Mannerheim, 1827
Male adult
NE Kazakhstan, W Altay Mts, near Shemonaikha city., flood plain, 25-28.VII.2018, S.Vashchenko leg., L=22.5 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924
Male adult
Russia, W Sayan Mts., Kazyr-Suk river, W slope of Oyskiy Mt. rng., ?h=2200 m 7-15.VII.1999, Brinev A. leg., L=19 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) aeruginosus aeruginosus (as salechardensis Obydov, 1999)
Female adult
N Russia, Polar Ural, Yamalo-Nenets reg., Harp station envir., 15.VII.1995, G. Chekura leg.
Copyright © Ruslan Panin

Carabus (Morphocarabus) henningi oviformis Beheim & Breuning, 1943
Male adult
Russia, Polar Ural, Yamalo-Nenets reg.,Harp station envir., 15.VII.1995, G. Chekura leg. L=19 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878 (variability)
Male adult
Far East, Promorskij Krai, Ussuriisk reg., Kamenushka, 9-15.6.2008 (cAA)
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
Female adult
Russia, Far East, Primorskiy Kray, Chuguevsky distr., 30 km ESE from Chuguevka, 9-14.VIII.2007, S.Shabalin leg.
Copyright © Ruslan Panin

Carabus (Neoplectes) starckianus ganglbaueri Reitter, 1888
Female adult
NW Caucasus, S Russia, Adygea Rep., Confluence of the river Belaya & Kisha h~700m, 24.V-7.VIII.2010, D.Fominyh leg.
female,32 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) edithae exedithae Gottwald, 1985
Male adult
Karachaevo-Cherkessia, low cource of Daut river, 16-20.V.2009 Anichtchenko A. leg
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) juenthneri atchibachi Zamotajlov, 1991 Paratype
Female adult
Abkhasia,
Valley of river Bzyb near Mt. Achibakhu. h=1500m., A.S. Zamotajlov leg.
female: 33mm, 31mm
coll. R.Panin
Copyright © Ruslan Panin

Carabus (Neoplectes) juenthneri avadcharensis Kurnakov, 1970 (loc.typ.)
Male+Female adult
Abkhasia, near Avadhara, Site Pyv h~1850m, 43.490730 40.686433, 1.VI-12.VII.2009, Zamotajlov A.S., Bondarenko A.S. leg.
L=27mm

Copyright © Ruslan Panin

Carabus (Neoplectes) kratkyi felicitanus Reitter, 1893 (as porcellus)
Male adult
NW Caucasus, Resp. Adygea, Mt.Tybga, 28.VI.1997
male,20mm
Copyright © Ruslan Panin

Carabus (Neoplectes) kratkyi Ganglbauer, 1890
Male+Female adult
W Caucas, 1900m, Khatsavitaya Mt., 4.VI-13.VII.1994
A.Miroshnikov leg.
male 25mm, female 26mm
photo & collection Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) kratkyi Ganglbauer, 1890
Male+Female adult
W Caucas Mts., 2500m, Mt. Yatyrgvarta,
25.VI-12.VIII.1992 Shchurov V. leg.
male 20mm, female 21mm
Copyright © Alexander Anichtchenko

Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890
Female adult
NW-Caucasus, Russia,
Karacaj-Cherkessija, Abishira-Akhuba range h=2650m, upper Kyafar-Agur Riv., 4-9.VIII.2009, Solodovnikov I.A. leg.
female, 23 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890
Male adult
NW-Caucasus: Karacaj-Cherkessija,
Abishira–Akhuba Mnt. Range h=2000-2650m, Kyafar-Agur Riv., 14–17.VII.2009, Fominykh D. leg.
male, 22mm
Copyright © Ruslan Panin

Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990
Male+Female adult
W.Caucasus,
Psebai env., Nikitino, 2.V-3.VII.1993, leg. A.S.Zamotajlov
female 33mm
Copyright © Ruslan Panin

Carabus (Neoplectes) prometheus prometheus Reitter, 1887
Male adult
NW-Caucasus,Krasnodar reg., Apsheron Distr., Lagonaki Mt., near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominykh leg.
L= 42mm.
Copyright © Ruslan Panin

Carabus (Neoplectes) prometheus prometheus Reitter, 1887
Male+Female adult
NW Caucasus, Russia,
Krasnodar reg.,Krasnoarmeisk distr., Krasny Les, near Podlesnyi vill.,17-22.IV.2008, D.D.Fominykh leg.
female, 40mm
Copyright © Ruslan Panin

Carabus (Neoplectes) reitteri laevisternis Starck, 1890
Male adult
Abkhasia,W.slope of Gagrinsky Mt., Krepostnaya River source, h=1325m., A.S.Zamotajlov leg. 27.VII.2000
male, 29 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) reitteri laevisternis Starck, 1890
Male adult
W-Caucasus:Sochi env., Aibga Mt. h=2000m., 06.VII.2000
male, 29mm
Copyright © Ruslan Panin

Carabus (Neoplectes) starcki Heyden, 1885
Male+Female adult
NW-Caucasus,
Krasnodar reg., Mt. Aishkho, Malaya Laba River, h=2200m., 14.VII.2008, D.Fominyh leg.
female-21mm
Copyright © Ruslan Panin

Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988 (as shchessiensis) Paratype
Male+Female adult
NW Caucasus,
Krasnodar reg., Tuapsinskyi distr. Shessi Mt.
30.V.-25.VI.1993 Mt.Shessi, h=1800m., Imura & A.S.Zamotajlov, 1993 leg.
female, 29mm
Copyright © Ruslan Panin

Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988
Female adult
Russia,W Caucasus,
Krasnodar reg.,Chernogorie mts.,Chernogor mt. h=1700-1800m. 23.V.-30.VII.1999. A.S.Zamotajlov leg.
female 29mm.
Copyright © Ruslan Panin

Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
Male adult
NW-Caucasus, Russia,
Krasnodar reg., Apsheron Distr., Lagonaki Mt. ,near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominykh leg.
male, 28mm
Copyright © Ruslan Panin

Carabus (Neoplectes) starckianus theseus Briansky, 1910
Male+Female adult
NW-Caucasus, Russia,
Krasnodar reg.,Tuapse distr., Mt.Semashkho 04.V.2008, D.Fominyh leg.
female, 32mm
Copyright © Ruslan Panin

Carabus (Neoplectes) juenthneri avadcharensis Kurnakov, 1970
Male+Female adult
W. Caucasus, Abkhasia, 8 km NE of Ritza Lake,<br /> Avadkhara valley, h=1700 m<br /> 10-VI-2009 A.Safronov leg.
Copyright ©

Carabus (Neoplectes) edithae exedithae Gottwald, 1985
Male adult
NС-Caucasus, Karachaevo-Cherkessia, Abishir-Akhuba Mt.Range h=2250-2600 m, right bank Pkhiya river, 10-13.VI.2013, S. Pavluchuk leg.<br /> L=25 mm
Copyright © Ruslan Panin

Carabus (Neoplectes) reitteri reitteri Retowski, 1885
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Carabus (Neoplectes) edithae Reitter, 1893
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Carabus (Neoplectes) starckianus theseus Briansky, 1910
Male adult
Russia, Krasnodar Territory, Tuapse district, env. vill. Psebe, left bank of the Psebe River, hornbeam + oak + walnut, h = 146 m, 03.05-01.07.2014, leg. I.A. Solodovnikov, S.V. Solodovnikova, S.M. Pavlyuchuk
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus theseus Briansky, 1910
Female adult
Russia, Krasnodar Territory, Tuapse district, env. vill. Psebe, left bank of the Psebe River, hornbeam + oak + walnut, h = 146 m, 03.05-01.07.2014, leg. I.A. Solodovnikov, S.V. Solodovnikova, S.M. Pavlyuchuk
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
Female adult
Russia, Krasnodar Territory, West-Caucasus, Apsheronsky district, Guam ridge, env. Mezmay village, by the road, beech, oak, blackberry, h = 1200-1250 m, 03.IV-14.VI.2020, leg. S.M. Pavlyuchuk, A.P. Tatulyan
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
Female adult
Russia, Krasnodar Territory, West-Caucasus, Apsheronsky district, Guam ridge, env. Mezmay village, by the road, beech, oak, blackberry, h = 1200-1250 m, 03.IV-14.VI.2020, leg. S.M. Pavlyuchuk, A.P. Tatulyan
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
Male adult
Russia, Krasnodar Territory, West-Caucasus, Apsheronsky district, Guam ridge, env. Mezmay village, by the road, beech, oak, blackberry, h = 1200-1250 m, 03.IV-14.VI.2020, leg. S.M. Pavlyuchuk, A.P. Tatulyan
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus kaljuzhnyji Zamotajlov, 1988
Male adult
Russia, Krasnodar Territory, West-Caucasus, Apsheronsky district, Guam ridge, env. Mezmay village, by the road, beech, oak, blackberry, h = 1200-1250 m, 03.IV-14.VI.2020, leg. S.M. Pavlyuchuk, A.P. Tatulyan
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
Female adult
W Caucasus, NW slope Aibga Mt.R. vall. riv. Galion, h = 650 m, Fagus, Alnus viridis forest, 18-24.VI.1999, I.A. Solodovnikov leg.
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
Male adult
W Caucasus, Aibga Mt.R., 02.VIII.1994, A.S. Zamotailov leg.
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886
Male adult
Russia, W Caucasus, Krasnaya Polyana, Atchischkho Mt.R., h = 1200 m, 10.06.1995, N.Yu. Pichugin leg.
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus adelaidae Starck, 1889
Female adult
Russia, Krasnodar area, Adler, Ader distr., right bank vall. Riv. Msymta, near vill. Chvesipse, h = 800 m, 06.VII.1999, B. Saratovskikh leg.
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus adelaidae Starck, 1889
Female adult
Russia, Krasnodar area, Adler, Ader distr., right bank vall. Riv. Msymta, near vill. Chvesipse (Fagus+Quercus+Castanea forest) [Sochi national park], h = 720 m, 03.V-08.VII.2016, I.A. Solodovnikov, S.V.Solodovnikova, V.M.Kotsur leg.
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus adelaidae Starck, 1889
Female adult
Russia, Krasnodar area, Adler, Ader distr., right bank vall. Riv. Msymta, near vill. Chvesipse (Fagus+Quercus+Castanea forest) [Sochi national park], h = 720 m, 03.V-08.VII.2016, I.A. Solodovnikov, S.V.Solodovnikova, V.M.Kotsur leg.
Copyright © Igor Solodovnikov

Carabus (Neoplectes) starckianus adelaidae Starck, 1889
Male adult
Russia, Krasnodar area, Adler, Ader distr., right bank vall. Riv. Msymta, near vill. Chvesipse (Fagus+Quercus+Castanea forest) [Sochi national park], h = 720 m, 03.V-08.VII.2016, I.A. Solodovnikov, S.V.Solodovnikova, V.M.Kotsur leg.
Copyright © Igor Solodovnikov

Carabus (Neoplectes) reitteri reitteri Retowski, 1885
Male adult

Copyright © Kirill Makarov

Carabus (Pachystus) hortensis hortensis Linne, 1758
Female adult
Natural hybrid Carabus hortensis X Carabus granulatus. Belarus, Minsk reg., 8,7 km of Borisov, near vill. Lyubatovstchina, pine forest (Pinetum pleuroziosum), h = 213 m, 54.31934° N, 28.511486° E, 20-27.09.2018, leg. Klychenok Snezhana
Copyright © Igor Solodovnikov

Carabus (Pachycarabus) koenigi Ganglbauer, 1886
Male adult
NW Caucasus, S Russia, Adygea Rep., Lagonaki plateau, Site Murzikal h=1800-2000m, 24.V-18.IX.2010, D.Fominykh leg.
L= 22 mm
Copyright © Ruslan Panin

Carabus (Pachycarabus) staehlini Adams, 1817
Male adult
S Russia, E-Caucasus, Dagestan, Nukatl Mt Rng h=2600m, 25.VII.1997, V. Savizky leg. L=21 mm
Copyright © Ruslan Panin

Carabus (Pachycarabus) koenigi Ganglbauer, 1886
Male adult

Copyright © Kirill Makarov

Carabus (Pachycarabus) staehlini staehlini Adams, 1817
Male adult

Copyright © Kirill Makarov

Carabus (Pachycarabus) koenigi Ganglbauer, 1886
Endophallus
NW Caucasus, Republic Adygea, Mt.Tybga h=2100 m,
Copyright © Ruslan Panin

Carabus (Pachycarabus) koenigi Ganglbauer, 1886
Male adult
NW Caucasus, Republic Adygea, Mt.Tybga h=2100 m, 28.VI.2006, Tret’yakov Yu. leg., L=21 mm
Copyright © Ruslan Panin

Carabus (Pachystus) hungaricus mingens Quensel, 1806
Male adult
male, 31 mm
SW-Russia: Stavropol Krai, near Stavropol city , Russkiy Les, 11.VI-28.IX.2010, Pavluchuk S. leg.
Copyright © Ruslan Panin

Carabus (Pachystus) hungaricus cribellatus Adams, 1812
Male adult
E Siberia, Russia, Krasnoyarsk Reg., near Krasnoyarsk, forest-steppe,1.VII-20.VIII,2011,M.Ivanov leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Pachystus) hungaricus scythus Motschulsky, 1847
Female adult
S Ukraine, Zaporizhzhya reg., Khortytsa island, 16-20.VII.2011, M. Mulenko leg., det. Panin R.
L=28 mm
Copyright © Ruslan Panin

Carabus (Pachystus) hungaricus scythus Motschulsky, 1847 (loc.typ.)
Male adult
S-Ukraine, near Dnipropetrovsk, 6-15.IX.2013, A. Sukhenko leg.<br /> L=28 mm
Copyright © Ruslan Panin

Carabus (Pachystus) glabratus glabratus Paykull, 1790
Male adult

Copyright © Kirill Makarov

Carabus (Pachystus) cribratus cribratus Quensel, 1806
Male adult
NW Armenia, Amasia, (Shirak prov.), 20-24.VIII.1998, Kazaryan P.V. leg.
Copyright © Ruslan Panin

Carabus (Piocarabus) vladimirskyi Dejean & Boisduval, 1830
Male adult
Buryatia, Kyahta distr. [Khyagh], 5 km SE Chikoi vill., 16.8.1998, Anichtchenko A. leg.
Copyright © Alexander Anichtchenko

Carabus (Procerus) scabrosus caucasicus Adams, 1817
Male adult
N Caucasus,
Stavropol Krai, Pyatigorsk, Mt.Mashuk, VI.2007 Tikhonov Yu. leg.
44mm
Copyright © Ruslan Panin

Carabus (Procrustes) coriaceus coriaceus Linne, 1758
Female adult
Belgique : Prov. Luxembourg, Harnoncourt 31.vii.1975 - M. Grotz Leg. (Size : 36 mm)
Copyright © Noël Mal

Carabus (Sphodristocarabus) varians janthinus s.str.
Male+Female adult
NW-Caucasus, Russia, Krasnodar reg.,Tuapse distr., Mt.Semashkho 04.V.2008, D.Fominyh leg.<br /> female, 24mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) adamsi Adams, 1817 (as chiragricus Fischer, 1828)
Male+Female adult
Russia, N. Caucasus, Stavropol reg., Zheleznovodsk, Zheleznaya Mt., h=600 m. 5-11.VI.2011, Е. Shankhiza leg. male - 24 mm, female - 25 mm.
Copyright ©

Carabus (Sphodristocarabus) adamsi Adams, 1817
Male adult
CN-Georgia, N of Tbilisi, NW of Tianeti, Tvalivi env., 20.VI.2015, F. Grycz leg. L=24 mm
Copyright © Ruslan Panin

Carabus (Sphodristocarabus) varians janthinus Ganglbauer, 1887
Female adult
Carabus (Sphrodistocarabus) janthinus dvorschaki Mandl, 1975 Abkhazia, Gudautsky district, S- Bzybsky Ridge, a watershed hill at the head of the river Khipsta, 43°17'56.63" N, 40°40'16.74" E, h= 1490 m, mature beech-maple forest with butterbur, moist soil, 05.07-17.08.2017, leg. I.A. Solodovnikov, S.V. Solodovnikova, E.V. Tatun
Copyright © Igor Solodovnikov

Carabus (Tachypus) cancellatus tibiscinus Csiki, 1905
Male+Female adult
female,28 mm<br /> SE Hungary, Csongrбd, Makу env., 1-8.VIII.2004, Krause R., Slovak, Pittner leg
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus s.str. (as sulinensis Born, 1902)
Male+Female adult
S UKRAINE, <br /> Krym Peninsula, Staryi Krym, 17.V.2002<br /> L= 26 mm.
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus cancellatus Illiger, 1798
Male adult
W-Ukraine, Podolian Upland (Lviv Opillia) W of Vybranivka vill., under the bark, 8.II.2008, R.Panin leg., L=22 mm
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus cancellatus (as charcoviensis Bernau, 1915)
Female adult
NE-Ukraine, Kharkov reg., Zmiev distr., 1-7.VI.2007, Kovalchuk I. leg., L=27 mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) marginalis Fabricius, 1794
Male+Female adult
E Ukraine,
Kharkov reg.,near Vasishchevo vill.10-20.VI.2006 D.Kovalchuk leg.
female 24mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) decolor Fischer, 1823
Male adult
S Russia, NW-Caucasus,
Karacaj-Cherkessija,Skalisty Mnt. Range, Site Banditskoe h=1400m, 2-3.V.2009,
Fominykh D., Bondarenko A.S. leg.
male,17mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) convexus s.str. (as pyrenaicus Born, 1907)
Male adult
Pirineos, Canfranc, 1300m, Barranco de Izas
15.6.2005 Anichtchenko A. leg.
male, 16 mm
Copyright © Alexander Anichtchenko

Carabus (Tomocarabus) bessarabicus ? concretus Fischer, 1823
Male adult
E Ukraine, Lugannsk Reg., Sverdlovsk distr., Steppe Reserve “Provalsky", 3.V-3.VI.2011, leg.Glotov S./det.Panin R.<br /> female, 22 mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) marginalis Fabricius, 1794
Endophallus
N Ukraine, Kiev reg., Vyshgorod disrt., near Khotyanivka vill.,
Copyright © Ruslan Panin

Carabus (Tomocarabus) marginalis Fabricius, 1794
Male adult
N Ukraine, Kiev reg., Vyshgorod disrt., near Khotyanivka vill., 1.VII-7.IX..2025, E. Rutjan leg.l=22 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) estreicheri Fischer, 1822
Male+Female adult
W UKRAINE, NW Podolian,
Lviv Region, Brody distr., near Yasenov vill., steppe sites, 2-28.V.2010, Panin R. leg.
female, 23mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus tscherkessicus Korge, 1964
Male adult
S Russia, NW-Caucasus,<br /> Karachay-Cherkessia,Teberda, Mukhinsky (Mukhu) Mt. pass, h=2750-2800 m , 8.VIII.2010, I.A. Solodovnikov leg.<br /> male, 23 mm<br />
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus sibiricus Fischer, 1822
Male adult

Copyright © Kirill Makarov

Carabus (Trachycarabus) sibiricus haeres (as fossulatus Dejean, 1826)
Male+Female adult
E.UKRAINE,<br /> Kharkov reg., Zmiev distr.,in steppe,10-20.V I.2007, D.Kovalchuk leg.<br /> female, 26mm/male, 24mm.<br /> photo/det. Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus errans Fischer, 1823
Male+Female adult
female:SE Ukraine,
Donetsk, 25.VII.2009, A.I.Gubin leg. L=30mm.
male:E Ukraine, Lugansk reg., near Slavyanoserbk, 5-15.VIII.2009, S.Konovalov leg. L=26mm.
photo/det.Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus bosphoranus Fischer, 1823
Male adult
Carabus (Trachycarabus) sibiricus bosphoranus Fisch. 1823<br /> S Ukraine, Krym Peninsula,<br /> Dolgorukovskaya jaila, 25.VIII.1991 I.A.Solodovnikov leg.<br /> male, 23mm<br />
Copyright © Ruslan Panin

Carabus (Trachycarabus) scabriusculus scabriusculus (as erythropus Fischer, 1822)
Male+Female adult
W.Ukraine,<br /> NW Podolian Upland , Lviv Reg., Zolochiv distr., near Luky vill., steppe sites, 7.V-8.VI. 2010<br /> leg. Panin R.<br /> female, 22mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus planus Gehin, 1885
Male adult
S Russia,<br /> Krasnodar reg., Anapa distr., B. Utrish vill., env., 11-20.VIII.2008, Khatchikov E. leg.<br /> male, 26mm<br /> photo Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus planus Gehin, 1885
Endophallus
<b>endophalus </b> S Russia,Krasnodar reg., Anapa distr., B. Utrish vill., env.,
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus planus Gehin, 1885
Endophallus
<b>endophalus</b> Karacaj-Cherkessija, near Kart-Djurt vill., upper Kylan riv. h=1300m
Copyright © Ruslan Panin

Carabus (Trachycarabus) estreicheri Fischer, 1822
Female adult
Volgograd Reg., Serafimovka Distr., Ust' -Khoperskaya Station 5.V.2009 leg.O.Sveshnikov
18.5 mm
Copyright © Kirill Makarov

Carabus (Trachycarabus) sibiricus (as gvozdevae Obydov, 1997)
Male adult
Russia, Khakassia, <br /> W Sayan, Enisey riv., Batenevskiy Mt Range, 12 km E Pervomayskiy vill., 11.VII.2000, A.& N. Menstschicovy leg.<br /> L=21 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) estreicheri Fischer, 1822 (as jaegeri Ménétriés, 1827)
Female adult
E UKAINE, Lugansk city, Site Luganskoe, 20-30.VII.2010, V.A.Landik leg.
L=21 mm
det.Panin.R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus tscherkessicus Korge, 1964
Male adult

Copyright © Kirill Makarov

Carabus (Trachycarabus) sibiricus sibiricus Fischer, 1822
Male adult
E Siberia, Russia, Krasnoyarsk Reg., near Krasnoyarsk, on dry steppe slopes, 7.VIII,2011, M.Ivanov leg.
L=21 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
Male adult
N Mongolia, Khövsgöl reg., Lake Khovsgol, h=1640m, 51*:12’N; 100*:45’E, steppificated sites,
20-25.VII.2010. A.Dergalev leg.
L=22 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) estreicheri (ab.rufofemoratus Lomnicki, 1892)
Female adult
W Ukraine, NW Podolian, Lviv reg., Brody distr., Makitra Mt, steppe site, 15.VI - 24.IX.2017, Panin R. & Kanarsky Yu. leg.
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus haeres (trans s. errans)
Female adult
C Ukraine, S of Poltava city, edge forest-lane, 2-26.IX.2010, O. Pozharov leg. /det. R.Panin. L=30 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) estreicheri Fischer, 1822 (ab.rufofemoratus Lomnicki, 1892)
Female adult
W Ukraina, Podolian Upland, Lviv reg., Brody distr., Mt. Makitra, steppe site, 19.V-5.VI.2020, Panin R. leg. L=23 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) scabriusculus inapertus Motschulsky, 1850
Male adult
NC Ukraine, Kiev city, Lysa Gora, 1-15.VI.2012, Gerasimov R.leg./det. R.Panin, L=19 mm
Copyright © Ruslan Panin

Carabus (Tribax) titan titan Zolotarev, 1913
Male+Female adult
Russia,NW Caucasus,
Krasnodar reg.,Chernogorie Mnt. Range, Agulova Balka h=950m V-VII.1999, A.S.Zamotajlov leg.
female,32mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Tribax) steveni steveni Menetries, 1832
Male adult
Carabus (Tribax) steveni steveni Menetries, 1832
Karachaevo-Cherkessia,
Hudes riv., near Tsarskie vorota,
V.2009 Anichtchenko A. leg.
male, 25mm
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Tribax) circassicus teberdensis Zolotarev, 1913
Male+Female adult
W-Caucasus: Karacaj-Cherkessija,
Abishira–Akhuba Mnt. Range h=2000-2650m, Kyafar-Agur Riv., 14–17.VII.2009, Fominykh D. leg.
female, 25mm
Copyright © Ruslan Panin

Carabus (Tribax) circassicus circassicus Ganglbauer, 1886
Male+Female adult
Abkhasia,
W.slope of Gagrinsky Mt., Krepostnaya River source, h=930-980m., A.S.Zamotajlov leg. 27.VII.2000
female, 29 mm/male, 26mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Tribax) circassicus abagonensis Starck, 1894
Male+Female adult
S Russia, NW Caucasus,
Adygea Rep., Mt.Abago, 26.VI.1999
L=24mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Tribax) biebersteini ulgeni Gottwald, 1982
Male adult
S Russia, Karacaj-Cherkessija,
near Kart-Djurt vill., upper Kylan riv. h=1300m, 26-27.V.2009, Fominykh D. leg.
male, 28mm
Copyright © Ruslan Panin

Carabus (Tribax) apschuanus apschuanus Rost, 1893
Male+Female adult
Carabus (Tribax) apschuanus apschuanus Rost, 1893    
Russia, Caucasus, Krasnodar reg.,
h=1200 m, 29.VII.2000
A. Zamotajlov leg.
Copyright © Safronov

Carabus (Tribax) agnatus agnatus Ganglbauer, 1889
Male adult
NW-Caucasus,
Krasnodar reg., Mostovskoj distr., Skirda Mts. N. slope Yatyrgvart Mt., h=2300-2500m. 09-10.VII.2008, D.Fominyh leg.
male, 23mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Tribax) apschuanus pseudoplatessa Gottwald, 1982
Male+Female adult
NW-Caucasus, Karachay-Cherkessia, Teberda: Dzhamagat gorge, the left bank of the riv. Dzhamagat, 10.VII-28.VIII.2010 S.Pavluchuk leg.
female, 27 mm
Copyright © Ruslan Panin

Carabus (Tribax) circassicus circassicus (as tshchaltensis Novotný & Voříšek
Male adult
W-Caucasus: Abkhasia, Gudauta distr., E slope Mt Anchkho h=2200-2350 m, 14-20.VII.2011, S.Pavluchuk, I. Solodovnikov leg./det. I. A. Solodovnikov.
25 mm

Copyright © Ruslan Panin

Carabus (Tribax) agnatus pseudagnatus Novotný & Voříšek, 1988
Female adult
NW-Caucasus, Karachay-Cherkessia, Bol'shaya Marka River h=2500-2600m, 12.VIII.2010, I.A. Solodovnikov leg./det.
Copyright © Ruslan Panin

Carabus (Tribax) agnatus pseudagnatus Novotný & Voříšek, 1988
Male adult
W-Caucasus: Abkhasia, Gudauta distr., Mt Anchkho h=2200-2350 m, 14-20.VII.2011, S.Pavluchuk, I. Solodovnikov leg./det. I. A. Solodovnikov.
L=25 mm
Copyright © Ruslan Panin

Carabus (Tribax) agnatus agnatus Ganglbauer, 1889
Male adult

Copyright © Kirill Makarov

Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
Male adult
Russia, C-Caucasus: N Ossetia, Chekhatsirtit Mt range, Mt Tepli h=3000 m, 21.VII.2009. /det. Solodovnikov I.A, L=20.5 mm
Copyright © Ruslan Panin

Carabus (Tribax) steveni schamyli Hampe, 1852
Female adult
C Caucasus, North Ossetia, the road on the village of Coban (Koban), the left river bank Gizeldon, the deciduous wood, h = 895-905 m, N 42°55'32.91", E 44°31'53.53", 20.05-03.07.2017, leg. A.P. Tatulyan
Copyright © Igor Solodovnikov

Carabus (Tribax) steveni schamyli Hampe, 1852
Female adult
C Caucasus, North Ossetia, the road on the village of Coban (Koban), the left river bank Gizeldon, the deciduous wood, h = 895-905 m, N 42°55'32.91", E 44°31'53.53", 20.05-03.07.2017, leg. A.P. Tatulyan
Copyright © Igor Solodovnikov

Carabus (Tribax) steveni schamyli Hampe, 1852
Male adult
C Caucasus, North Ossetia, the road on the village of Coban (Koban), the left river bank Gizeldon, the deciduous wood, h = 895-905 m, N 42°55'32.91", E 44°31'53.53", 20.05-03.07.2017, leg. A.P. Tatulyan
Copyright © Igor Solodovnikov

Carabus (Tribax) steveni schamyli Hampe, 1852
Male adult
C Caucasus, North Ossetia, the road on the village of Coban (Koban), the left river bank Gizeldon, the deciduous wood, h = 895-905 m, N 42°55'32.91", E 44°31'53.53", 20.05-03.07.2017, leg. A.P. Tatulyan
Copyright © Igor Solodovnikov

Carabus (Tribax) nothus sharoiensis Fominykh, Zamotajlov, Avtaeva et Titarenko, 2019 Paratype
Male adult

Copyright © Igor Solodovnikov

Carabus (Tribax) nothus fausti Dohrn, 1873
Male adult
Carabus (Microtribax) fausti fausti Dohrn, 1873 Russia: Daghestan, Kyabyak Mt.R.
Copyright © Igor Solodovnikov

Carabus (Tribax) nothus fausti Dohrn, 1873
Female adult
Carabus (Microtribax) fausti fausti Dohrn, 1873 Russia: Daghestan, Kyabyak Mt.R.
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum kasakorum Semenov, 1896
Male adult
Russia: N.Osetia: Pastbyshchniy Mt.R., S slopes of Khoshkharanrag Mt.R., right bank of Amusadon riv., h=1500 m
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum kasakorum Semenov, 1896
Female adult
Russia: N.Osetia: Pastbyshchniy Mt.R., S slopes of Khoshkharanrag Mt.R., right bank of Amusadon riv., h=1500 m
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Male adult
Russia, Caucasus, Ingushetia, South slopes of Skalistyi Mt Range, Stolovaya Mt., near Beyni, alpin zone, h = 2800 m, N 42°51’35.30",E 44°42’13.70", h = 2800m, 21.V–19.VII.2018, D. Fominykh leg.
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Female adult
Russia, Caucasus, Ingushetia, South slopes of Skalistyi Mt Range, Stolovaya Mt., near Beyni, alpin zone, h = 2800 m, N 42°51’35.30",E 44°42’13.70", h = 2800m, 21.V–19.VII.2018, D. Fominykh leg.
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Endophallus
Russia, Caucasus, Ingushetia, South slopes of Skalistyi Mt Range, Stolovaya Mt., near Beyni, alpin zone, h = 2800 m, N 42°51’35.30",E 44°42’13.70", h = 2800m, 21.V–19.VII.2018, D. Fominykh leg. (TOPOTYPUS)
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Male adult
Russia, Caucasus, Ingushetia, Skalistyi Mt. Range, S slopes of Tsoreylam Mt. Range, 5 km W of Mt. Khakhalgi, N 42°50’07.60", E 45°05’02.20", h=2500 m, 25.V–21.VII.2017, D. Fominykh leg.
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Endophallus
Russia, Caucasus, Ingushetia, Skalistyi Mt. Range, S slopes of Tsoreylam Mt. Range, 5 km W of Mt. Khakhalgi, N 42°50’07.60", E 45°05’02.20", h=2500 m, 25.V–21.VII.2017, D. Fominykh leg.
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum tschetschenicus Semenov, 1896
Female adult
Russia, Chechnya, Caucasus, Skalystyi Mt. Range upper river Gekhi, Mt. Ekkyrkort, N 42°52', E 45°26', h=2230 m, 26.V–21.VII.2018, D.Fominykh leg.
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
Female adult
Russia: N.Osetia, Bokovoy Mt.R., near Arkhonskiy pass., alpin zone, h = 2700 m, 23.V-21VII.2018, D. Fominykh leg.
Copyright © Igor Solodovnikov

Carabus (Tribax) kasakorum georgiensis Gottwald, 1980
Male adult
Russia: N.Osetia, Bokovoy Mt.R., near Arkhonskiy pass., alpin zone, h = 2700 m, 23.V-21VII.2018, D. Fominykh leg.
Copyright © Igor Solodovnikov

Carabus (Tribax) osseticus abanosicus Deuve, 2013
Male adult

Copyright © Igor Solodovnikov

Carabus (Tribax) osseticus osseticus Adams, 1817
Male adult

Copyright © Igor Solodovnikov