Brachinus (Neobrachinus) elongatulus Chaudoir, 1876
Male adult
Size variation within the species. Scale bar: 5 mm. Image from Ikagawa & Moore, 2022
Copyright © - - -

Brachinus (Neobrachinus) gebhardis Erwin, 1965
Male adult
Image from Ikagawa & Moore, 2022. Scale bar: 1 cm.
Copyright © - - -

Brachinus (Neobrachinus) elongatulus Chaudoir, 1876
Male adult
Image from Ikagawa & Moore, 2022. Scale bar: 1 cm.
Copyright © - - -

Brachinus (Neobrachinus) mexicanus Dejean, 1831
Male adult
Image from Ikagawa & Moore, 2022. Scale bar: 1 cm.
Copyright © - - -

Brachinus (Neobrachinus) costipennis Motschulsky, 1859
Male adult
Image from Ikagawa & Moore, 2022. Scale bar: 1 cm.
Copyright © - - -

Brachinus (Neobrachinus) lateralis Dejean, 1831
Male adult
Image from Ikagawa & Moore, 2022. Scale bar: 1 cm.
Copyright © - - -

Brachinus (Neobrachinus) favicollis Erwin, 1965
Male adult
Image from Ikagawa & Moore, 2022. Scale bar: 1 cm.
Copyright © - - -

Brachinus (Neobrachinus) pallidus Erwin, 1965
Male adult
U.S.A. Californie Arroyo Macho Igo Alomeda Janvier 1972
Copyright © Pascal Deschamps

Zacotus matthewsii LeConte, 1869
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Zacotus fredericki Nunenmacher, 1944 Holotype
Male adult
Image from FMNH
Copyright © - - -

Zacotus matthewsii LeConte, 1869
Male adult
Washington, Cowitz Co., Ariel forest 14.6.2014
Copyright © Alexander Anichtchenko

Calosoma (Callistenia) moniliatus Le Conte, 1851: 200
Male adult
SE Canada, Nueva Escocia
19.5.1970
male, 17 mm
Copyright © Alexander Anichtchenko

Calosoma (Calosoma) scrutator Fabricius, 1775
Female adult
USA, Illinois, Peoria Co.,<br /> Orange Prairie, 31.5.1990<br /> 32 mm<br /> coll. A.Montes<br /> photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Calosoma) scrutator Fabricius, 1775
Male adult
3 m SE Tishomingo , Tishomingo Co., Mississippi, USA; 24.VI.2005<br /> Eric B. Lago leg.
Copyright © Denis Potanin

Calosoma (Calosoma) wilcoxi Le Conte, 1848
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Calosoma (Calosoma) wilcoxi Le Conte, 1848
Male adult
2 m. W Rawlings, Allegany Co., Maryland, USA; 26 May 1983<br /> J.D. Glaser leg.
Copyright © Denis Potanin

Calosoma (Carabosoma) prominens LeConte, 1853
Male adult
USA Arizona, Cochise Co., Interstate 8 at Texas, Canyon Rest Area, 1500m, 32º00.3N 110º08.4W, 24.7.1996 Kavanaugh D. leg.; 30 mm
Copyright © Alexander Anichtchenko

Calosoma (Carabosoma) peregrinator Guerin-Meneville, 1844
Male adult
Fort Davis, Jeff Davis Co., Texas, USA; July 1998
Copyright © Denis Potanin

Calosoma (Chrysostigma) semilaeve LeConte, 1851: 199
Male adult
Orange Co., California, USA; May 1973
Copyright ©

Calosoma (Callistenia) moniliatus Le Conte, 1851: 200
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Calosoma (Chrysostigma) tepidum LeConte, 1851: 199
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Calosoma (Chrysostigma) simplex LeConte, 1878
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Calosoma (Chrysostigma) semilaeve LeConte, 1851: 199
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Calosoma (Carabosoma) angulatum Chevrolat, 1834
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Calosoma (Carabosoma) eremicola Fall, 1910: 91
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Calosoma (Carabosoma) parvicollis Fall, 1910: 90
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Calosoma (Carabosoma) peregrinator Guerin-Meneville, 1844
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Calosoma (Carabosoma) prominens LeConte, 1853: 400
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Calosoma (Carabosoma) sponsum Casey, 1897: 340
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Scaphinotus (Brennus) interruptus Menetries, 1844
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Scaphinotus (Brennus) crenatus Motschulsky, 1859
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Scaphinotus (Brennus) cristatus Harris, 1839
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Scaphinotus (Brennus) oreophilus Rivers, 1890
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Scaphinotus (Brennus) rugiceps incipiens (Casey, 1897)
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Scaphinotus (Brennus) striatopunctatus (Chaudoir, 1844)
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Scaphinotus (Brennus) punctatus LeConte, 1874
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab (det as subtilis Schaum, 1863), NMNH.; as punctatus det by Anichtchenko A. 2020
Copyright © - - -

Scaphinotus (Brennus) ventricosus Dejean, 183
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Scaphinotus (Stenocantarus) angusticollis Mannerheim, 1824
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Scaphinotus (Stenocantarus) angusticollis Mannerheim, 1824
Male adult
NW-USA, Washington, Thurston County, Douglas fir forest, 15.X.2005, A. Laforgue leg. L=21mm
Copyright © Ruslan Panin

Calosoma (Calosoma) scrutator Fabricius, 1775
Male adult
USA: Texas, Live Oak County 5 miles South of George West, 24.09.2010, local collector
Copyright © Igor Solodovnikov

Calosoma (Carabosoma) angulatum Chevrolat, 1834
Male adult
Reserva silvestra de Domitila, Pl., Granade prov., Nicaragua, 1-5.6.2005. det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Scaphinotus (Stenocantarus) velutinus (Menetries, 1843)
Male adult
USA, OR, Benton Co., III-V.1996 Mary's Peak, 3600m N slope, Labonte cols. (cAA) 26 mm; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Scaphinotus (Stenocantarus) angusticollis olympiae Van Dyke, 1944 (loc. typ.)
Male adult
USA: WA., Clallam Co, Olympic Nat Park, Sol Duc Hot Springs 25.5.96, M. Kippenhan leg. (cAA) 17 mm; det. A. Anichtchenko, 2020
Copyright © Alexander Anichtchenko

Scaphinotus (Neocychrus) behrensi Roeschke, 1907
Male adult
USA: OR., Benton Co., Mary's Peak 3600 m, N Slope, III-V.1996, J. Labonte cols. (cAA) 18 mm
Copyright © Alexander Anichtchenko

Scaphinotus (Brennus) rugiceps Horn, 1872
Male adult
USA: OR: Josephine Co. / Lone Mountain Rd, 4.3 miles S.W of Hwy 199 / 42° 01.787' N 123° 45.760' W 1719', 18-IV-6-VI-2020, M. Kippenhan (cAA) 15 mm; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Scaphinotus (Brennus) obliquus LeConte, 1868
Male adult
USA, CA., Shasta Co., Castle Crags State Park, 29-29.5.2016, M. Kippenhan leg. (cAA) 15 mm; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Scaphinotus (Stenocantarus) angusticollis Mannerheim, 1824
Female adult
USA, WA., Pierce Co., Penrose S.P. Trails, 7 sept 1996, M. Kippenhan leg (cAA) 19mm; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Scaphinotus (Brennus) cf oreophilus Rivers, 1890
Female adult
USA , CA, Toulumne Co., The Pines CG, Rt 120 West of Buck Meadows, 3267m, 37.49.14N 120.05.7W 19.6.2011 (cAA) 15mm; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Scaphinotus (Brennus) cf oreophilus Rivers, 1890
Male adult
USA , CA, Toulumne Co., The Pines CG, Rt 120 West of Buck Meadows, 3267m, 37.49.14N 120.05.7W 19.6.2011, M. Kippenhan leg. (cAA) 15mm
Copyright © Alexander Anichtchenko

Calosoma (Calosoma) scrutator Fabricius, 1775
Male adult

Copyright © Christensen_Tim

Scaphinotus (Stenocantarus) angusticollis Mannerheim, 1824
Male adult

Copyright © Igor Solodovnikov

Scaphinotus (Brennus) interruptus Menetries, 1844
Male adult
USA California, Sonoma Co., 2.5mi N of Cazadero on King Ridge Rd. Big Austin Creek, 70m, 38.33N 123.06W 21.6.1996 D.H.Kavanaugh leg. (cAA). det. Anichtchenko A. 2021
Copyright © Alexander Anichtchenko

Scaphinotus (Brennus) cristatus Harris, 1839
Female adult
USA California, Monterrey Co., Carmel, 18.8.1974, J.B.Johnson (cAA) 16mm
Copyright © Alexander Anichtchenko

Scaphinotus (Brennus) striatopunctatus (Chaudoir, 1844)
Male adult
USA California, Sonoma Co., 10 mi NW of Santa Rosa, Pepperwood Ranch NR, 22.3.1997 D.H. Kavanaugh leg. (cAA) det. A. Anichtchenko 2021
Copyright © Alexander Anichtchenko

Calosoma (Chrysostigma) cancellatus Eschscholtz, 1833
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Calosoma (Callistenia) dietzi Schaeffer, 1904: 197
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH. Photo processed by Anichtchenko A. 2022
Copyright © - - -

Calosoma (Callistenia) discors Le Conte, 1857
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH. Photo processed by Anichtchenko A. 2022
Copyright © - - -

Calosoma (Callistenia) subaeneus Chaudoir, 1869
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH. Photo processed by Anichtchenko A. 2022
Copyright © - - -

Calosoma (Callistenia) latipennis Horn, 1870
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH. Photo processed by Anichtchenko A. 2022
Copyright © - - -

Calosoma (Callistenia) luxatus Say, 1823
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH. Photo processed by Anichtchenko A. 2022
Copyright © - - -

Calosoma (Callistenia) wilkesi Le Conte, 1851
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH. Photo processed by Anichtchenko A. 2022
Copyright © - - -

Calosoma (Callistenia) monticola Casey, 1913
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH. Photo processed by Anichtchenko A. 2022
Copyright © - - -

Calosoma (Callistenia) schaefferi Breuning, 1928
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH. Photo processed by Anichtchenko A. 2022
Copyright © - - -

Calosoma (Callistenia) placerus Gidaspow, 1959
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH. Photo processed by Anichtchenko A. 2022
Copyright © - - -

Calosoma (Callistenia) subasperatus Schaeffer, 1915
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH. Photo processed by Anichtchenko A. 2022
Copyright © - - -

Calosoma (Carabosoma) peregrinator Guerin-Meneville, 1844
Male adult
Texas Canyon, Cochise County, Arizona 3.7.2013 (DUBC) 23 mm
Copyright © Alexander Anichtchenko

Calosoma (Callistenia) luxatus Say, 1823 (zimmermanni LeConte, 1848)
Female adult
USA OR, Jackson Co., FR300, 0.18mi W of rd. 20, 42.02.342N 122.54.937, 20.6.2021 M. Kippenhan leg. (cAA) 15.5 mm; det. A. Anichtchenko 2023
Copyright © Alexander Anichtchenko

Calosoma (Callistenia) latipennis Horn, 1870: 70 Holotype
Female adult
MCZ
Copyright © Harvard College

Carabus zimmermani LeConte, 1847 Holotype
Female adult
Museum of Comparative Zoology, Harvard University, Cambridge, Ma. (https://mczbase.mcz.harvard.edu/)
Copyright © Harvard College

Calosoma striatulum LeConte, 1859 Holotype
Male adult
Museum of Comparative Zoology, Harvard University, Cambridge, Ma. (https://mczbase.mcz.harvard.edu/)
Copyright © Harvard College

Calosoma (Callistenia) luxatus Say, 1823
Female adult
USA Utah, Iron Co., 18 mi East Cedar City, Jet Hwy 14-143, 9.6.2001 M. Kippenhan leg (cAA) 15 mm
Copyright © Alexander Anichtchenko

Calosoma (Callistenia) luxatus Say, 1823
Female adult
USA OR, Jackson Co., Tubs Springs State Wayside Hwy 66, 42.06.93N 122.26.5W, 9.6.2013 M. Kippenhan leg. (cAA) 18.5 mm
Copyright © Alexander Anichtchenko

Calosoma (Chrysostigma) morrisoni Horn, 1885
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Omus ambiguus Schaupp, 1884
Male adult
USA California, Shasta co., Shasta-Trinity NF, 7 ml W Castella 9-10.VII.1991 1100m D.A. Pollock leg. (cAA). 18 mm; det. Mike Kippenhan 2022
Copyright © Alexander Anichtchenko

Megacephala (Tetracha) carolina carolina s.str.
Male adult
USA Texas, Falls Co., Falls on the
Brazos River 8.10.1985
Copyright © Alexander Anichtchenko

Cicindela (Cicindela) longilabris perviridis Schaupp, 1884
Female adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Amblycheila schwarzi W.Horn, 1903
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Megacephala (Tetracha) carolina carolina s.str.
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Cylindera (Cylindera) lunalonga (Schaupp, 1884)
Male adult
USA: CA: Lassen Co, near Westwood, VI-2004
Copyright © Michael Kippenhan

Cylindera (Cylindera) terricola continua (Knaus, 1922)
Male adult
USA: NV: Nye Co., Beatty, VI-2003
Copyright © Michael Kippenhan

Cylindera (Cylindera) lemniscata lemniscata s.str.
Female adult
8.5 mm
Copyright © Kirill Makarov

Cicindela (Cicindelidia) psilogramma Bates, 1890 Paratype
Female adult
Copyright: Royal Belgian Institute of natural Sciences (RBINS) / DIGIT-3 Belspo Licence: CC
Copyright © - - -

Cicindela (Cicindelidia) psilogramma Bates, 1890 Paratype
Female adult
Copyright: Royal Belgian Institute of natural Sciences (RBINS) / DIGIT-3 Belspo Licence: CC
Copyright © - - -

Omus angustocylindricus W.Horn, 1913 Paratype
Male adult
Copyright: Royal Belgian Institute of natural Sciences (RBINS) / DIGIT-3 Belspo Licence: CC
Copyright © - - -

Omus angustocylindricus W.Horn, 1913 Paratype
Male adult
Copyright: Royal Belgian Institute of natural Sciences (RBINS) / DIGIT-3 Belspo Licence: CC
Copyright © - - -

Omus californicus lecontei G.H. Horn, 1872 Lectotype
Male adult
MCZ
Copyright © - - -

Cicindela (Cicindela) bellissima Leng, 1902
Female adult
USA, Oregon, Lincoln co., Newton (on ocean beach)
Copyright © Riccardo Sciaky

Cicindela (Cicindela) depressula Casey, 1897
Female adult
USA, Washington, Mt. Saint Helens. det. R. Sciaky
Copyright © Riccardo Sciaky

Cicindela (Cicindela) nebraskana Casey, 1909
Male adult
USA, Grasslands NP East Block (cRS). det. R. Sciaky
Copyright © Riccardo Sciaky

Cicindela (Cicindela) willistoni pseudosenilis Horn, 1900
Male adult
USA, California, Inyo co., Owens lake
Copyright © Riccardo Sciaky

Cicindela (Cicindela) tenuicincta Schaupp, 1884
Male adult
USA, California, Modoc co.
Copyright © Riccardo Sciaky

Eunota gabbi (G. Horn, 1866)
Male adult
Mexico, Sonora, Bahia de Cholla, Golfo de Califor., 14.6.1996, J. Shetterly (cAA) 10 mm
Copyright © Alexander Anichtchenko

Eunota californica mojavi (Cazier, 1937)
Female adult
Mexico, Sonora, Bahia de Cholla, Golfo de California, 14.6.1996, at lights, Shetterly et al (cAA) 12.5 mm
Copyright © Alexander Anichtchenko

Omus audouini Reiche, 1838
Male adult
USA: OR: Multnomah Co. / Portland, Mount Tabor Park / 45° 30.946' N 122° 35.842' W 526' / 22-IV-2020, M. Kippenhan (cAA)
Copyright © Alexander Anichtchenko

Megacephala (Tetracha) carolina carolina
Male adult

Copyright © Christensen_Tim

Cicindela (Cicindela) bellissima Leng, 1902
Male adult
USA, Oregon, Tillamook, 45.456216 Lon: -123.844014; Mar 23, 2013
Copyright © Christensen_Tim

Cicindela (Cicindelidia) tenuisignata (LeConte, 1851)
Female adult
Mexico, Durango, 78km E Hwy 45 XV Hidalgo, Preson de Doña Licha, 1259m, 26*20.374N 104*23.377W, 5.8.2014, J.A. Shetterly leg & det. (cAA). 11 mm
Copyright © Alexander Anichtchenko

Cicindela (Cicindela) hirticollis corpuscula Rumpp, 1962
Male adult
Image from Knisley, Kippenhan & Brzoska, 2014
Copyright © - - -

Cicindela (Cicindela) hirticollis siuslawensis Graves, 1988
Female adult
Image from Knisley, Kippenhan & Brzoska, 2014
Copyright © - - -

Cicindela (Cicindela) tranquebarica viridissima Fall, 1910
Male adult
Image from Knisley, Kippenhan & Brzoska, 2014
Copyright © - - -

Omus submetallicus G.Horn, 1868
Male adult
Image from Knisley, Kippenhan & Brzoska, 2014
Copyright © - - -

Eunota gabbi (G. Horn, 1866)
Male adult
Image from Knisley, Kippenhan & Brzoska, 2014
Copyright © - - -

Cicindela (Cicindela) willistoni (Leconte, 1879)
Male adult
Image from Knisley, Kippenhan & Brzoska, 2014 (as latesignata). det. A. Harman, 2026
Copyright © - - -

Cicindela (Cicindela) hirticollis gravida LeConte, 1851
Male adult
Image from Knisley, Kippenhan & Brzoska, 2014
Copyright © - - -

Parvindela terricola continua (Knaus, 1922)
Male adult
Image from Knisley, Kippenhan & Brzoska, 2014
Copyright © - - -

Cicindela (Cicindela) tranquebarica joaquinensis Knisley & Haines, 2007
Female adult
Image from Knisley, Kippenhan & Brzoska, 2014
Copyright © - - -

Cicindela (Cicindela) depressula depressula
Male adult
USA, OR Clackamas Co., Mt. Hood, Timberline L., 45.19.98N 121.42.57W 12.VII.2004 M. Kippenhan leg. (cAA) 12 mm
Copyright © Alexander Anichtchenko

Parvindela terricola susanagreae (Kippenhan, 2007)
Male adult
USA, CA, Inyo County, Klondike Lake, SE shore 37.12.2N 118.18.06W, 19.VI.2011 M. Kippenhan (cAA) 11mm
Copyright © Alexander Anichtchenko

Cicindela (Cicindela) purpurea hatchi Leffler, 1980
Female adult
OR, Benton Co., 20 sept. 1997, 29th Street and Walnut, Corvalis, M. Kippenhan leg & det. (cAA)
Copyright © Alexander Anichtchenko

Eunota californica pseudoerronea x mojavi
Female adult
USA, CA, San Bernardino Co., Baker, Soda Dry Lake, 285m, 35.09.79N 116.06.42W 21.6.2003 Kippenhan leg. & det. (cAA) 12mm
Copyright © Alexander Anichtchenko

Eunota californica pseudoerronea x mojavi
Male adult
USA, CA, San Bernardino Co., Baker, Soda Dry Lake, 285m, 35.09.79N 116.06.42W 21.6.2003 Kippenhan leg. & det. (cAA) 12mm
Copyright © Alexander Anichtchenko

Cicindela (Cicindela) senilis Horn, 1867 Lectotype
Male adult
© President and Fellows of Harvard College
Copyright © Harvard College

Cicindela (Cicindela) senilis and timbisha syn. nov.
Male+Female adult
C. timbisha, syn. nov. (top row) males (A–B), females (C–D) (USA: California: Inyo Co.); C. senilis (bottom row), males (E–F), females (G–H) (USA: California: Solano Co.). Image from Duran, Chambers, Nelson & Roman, 2023
Copyright © - - -

Cicindela (Cicindela) arida A.C.Davis, 1928 Holotype
Male adult

Copyright © Smithsonian NMNH

Elaphrus (Elaphrus) viridis Horn, 1878
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Elaphrus (Elaphroterus) purpurans Hausen, 1891
Male adult
Photographer: Beatriz Gimenez de Cordoba
SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © ---

Stenomorphus californicus rufipes Leconte, 1858
Male adult
Arkansas, Logan County. Photo I. Sokolov
Copyright © Igor Sokolov

Selenophorus (Selenophorus) schaefferi (Csiki, 1932)
Female adult
Image from Messer & Raber, 2021
Copyright © - - -

Selenophorus (Selenophorus) aequinoctialis Dejean, 1829
Male adult
Image from Messer & Raber, 2021
Copyright © - - -

Discoderus (Discoderus) amoenus LeConte, 1863
Male adult
Image from Messer & Raber, 2025
Copyright © - - -

Discoderus (Discoderus) impotens (LeConte, 1858)
Male adult
Image from Messer & Raber, 2025
Copyright © - - -

Discoderus (Selenalius) cordicollis G.Horn, 1891
Male adult
Image from Messer & Raber, 2025
Copyright © - - -

Discoderus (Paraselenalius) robustus G.Horn, 1883
Male adult
Image from Messer & Raber, 2025.
Copyright © - - -

Discoderus (Paraselenalius) obsidianus Casey, 1914
Male adult
Image from Messer & Raber, 2025.
Copyright © - - -

Tetragonoderus (Crossonychus) pallidus G.Horn, 1868 Lectotype
Male adult
"Cal.[ifornia]"
Copyright © - - -

Infernophilus castaneus (G.Horn, 1882) Lectotype
Male adult
MCZ
Copyright © - - -

Calybe (Ega) laetula (LeConte, 1851)
Male adult
det. A. Anichtchenko, 2018
Copyright © Alexander Anichtchenko

Lebia (Lebia) tuckeri (Casey, 1920)
Male adult
United States, New Mexico, Eddy, Sitting Bull Falls, 32.2433, -104.697, Wappes, J. E. 30 Jun 2000
Copyright © Smithsonian NMNH

Chlaenius (Agostenus) interruptus G.Horn, 1876
Male adult

Copyright © Erich Kirschenhofer

Chlaenius (Callistometus) ruficauda Chaudoir, 1856
Male adult
Mexico, Sonora, 26*54.897N 99*09.487W 1563m road from Alamos to Guiracoba 4.8.2016. J.A. Shetterly leg. (cAA) 12 mm; det. Anichtchenko A. 2021
Copyright © Alexander Anichtchenko

Chlaenius sericinitens Chaudoir, 1856 Holotype
Female adult
California (MNHN)
Copyright © Erich Kirschenhofer

Chlaenius (Chlaeniellus) variabilipes Eschscholtz, 1833
Male adult
BMNH
Copyright © Erich Kirschenhofer

Loricera (Loricera) foveata LeConte, 1851
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Nebria (Catonebria) gebleri Dejean, 1831
Female adult
USA Montana, Mineral Co. Trovd Creec
13.5 mil NE of Hoodoo pass 5.8.1996
11 mm
Copyright © Alexander Anichtchenko

Nebria (Reductonebria) acuta Lindroth, 1961 Paratype
Male adult
Museum of Zurich
photo Anichtchenko A.
Copyright © Alexander Anichtchenko

Nebria (Vancouveria) virescens Horn, 1870
Male adult
Canada, Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Notiophilus semiopacus Eschscholtz, 1833
Male adult

Copyright © Arvids Barsevskis

Notiophilus sierranus Casey, 1920
Male adult

Copyright © Arvids Barsevskis

Notiophilus sylvaticus Eschscholtz, 1833
Male adult

Copyright © Arvids Barsevskis

Leistus (Neoleistus) longipennis Casey, 1920
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Nebria (Reductonebria) sahlbergi sahlbergi s.str.
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Nebria (Reductonebria) sahlbergi triad Kavanaugh, 1979
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Nebria (Catonebria) praedicta Kavanaugh & Schoville, 2009
Male adult
USA, CA: Trinity Co, Thompson Peak, 2150-2500 m, 41°01' N, 123°03' W, 16.05.2013, R. Dudko, S. Schoville leg.
Copyright © Roman Dudko

Omophron (Omophron) dentatum LeConte, 1852
Male adult
USA, California, Groveland, Co. Tuolumne 17-19.IV.1978 C. v. Nidek. 6,6 mm Photo U. Valainis
Copyright © - - -

Omophron (Omophron) solidum Casey, 1897
Male adult
USA, California, Humboldt Co., Van Duzen park, 8 mi E of Carolla 17-21.VI.1982 C. v. Nidek. Photo U.Valainis
Copyright © Alexander Anichtchenko

Omophron (Omophron) obliteratum G. Horn, 1870
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Omophron (Omophron) gilae LeConte, 1852
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Omophron (Omophron) dentatum LeConte, 1852
Female adult
S Francisco California, H.G.C. (BMNH)
Copyright © Alexander Anichtchenko

Promecognathus crassus LeConte, 1868
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Promecognathus laevissimus Dejean, 1829
Male adult
Photographer: Beatriz Gimenez de Cordoba
SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © ---

Promecognathus crassus LeConte, 1868
Male adult
USA, Washington, Clarck Co., Vancuver lake, 12.vi.2012
Copyright © Alexander Anichtchenko

Pterostichus (Hypherpes) annosus Casey, 1913 Holotype
Male adult
«California» USNM [# 47030]
Copyright © Kipling W. Will

Pterostichus (Hypherpes) arcanus Casey, 1913 Lectotype
Male adult
«S[an]ta Cruz M[oun]t[ain]s, California», in USNM [# 46985].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) baldwini (Casey, 1924) Lectotype
Male adult
«Baldwin Lake (8600 ft.) [San Bernardino County], California». in USNM [# 47035].
Copyright © Kipling W. Will

Pterostichus breviusculus Casey, 1913 Lectotype
Male adult
«Anderson Valley, Mendocino Co[unty], California» , in USNM [# 46992].
Copyright © Kipling W. Will

Pterostichus bucolicus Casey, 1913 Paralectotypes
Male adult
«Humboldt Co[unty], California»
Copyright © Kipling W. Will

Pterostichus (Hypherpes) canallatus Casey, 1913 Lectotype
Male adult
«S[an]ta Barbara [Santa Barbara County], California» , in USNM [# 47026].
Copyright © Kipling W. Will

Pterostichus cuneatulus Casey, 1913 Lectotype
Male adult
«Humboldt Co[unty], California» , in USNM [# 46998].
Copyright © Kipling W. Will

Pterostichus diabolus Casey, 1913 Lectotype
Male adult
«M[oun]t Diablo [Contra Costa County], California» , in USNM [# 47036].
Copyright © Kipling W. Will

Hypherpes alamedae Casey, 1918 Lectotype
Female adult
«Alameda [Alameda County], California» . Lectotype (♀), in USNM [# 47013].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) esuriens Casey, 1913 Lectotype
Male adult
«Hermitage and Guallala, Mendocino Co[unty], California» , in USNM [# 47033].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) gliscans Casey, 1913 Lectotype
Male adult
«San Clemente Island [Los Angeles County], California» , in USNM [# 47025].
Copyright © Kipling W. Will

Pterostichus gregalis Casey, 1913 Lectotype
Male adult
«Humboldt Co[unty], California», in USNM [# 47027].
Copyright © Kipling W. Will

Pterostichus humboldti Casey, 1913 Lectotype
Male adult
«Humboldt Co[unty], California» (original citation), in USNM [# 47012].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) jacobinus Casey, 1913 Lectotype
Female adult
«San Diego [San Diego County], California» , in USNM [# 47024].
Copyright © Kipling W. Will

Hypherpes kansanus Casey, 1918 Lectotype
Male adult
«Kansas (probably western)» , in USNM [# 46988].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) laborans Casey, 1913 Lectotype
Male adult
«Lake Tahoe [Placer County], California» , in USNM [# 47020].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) lacertus Casey, 1913 Lectotype
Female adult
«Hydesville, Valley of Eel River, Humboldt Co[unty], California» , in USNM [# 47038].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) lassulus (Casey, 1920) Lectotype
Female adult
«Olancha, Inyo Co[unty], California», in USNM [# 46999].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) mercedianus (Casey, 1918) Lectotype
Male adult
«Merced Creek [Mariposa County], California» in USNM [# 47031].
Copyright © Kipling W. Will

Pterostichus breviusculus mimus Casey, 1913 Lectotype
Female adult
«Sonoma Co[unty], California» , in USNM [# 46990].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) miscellus Casey, 1913 Lectotype
Male adult
«S[an]ta Barbara [Santa Barbara County], California» , in USNM [# 47019].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) obsidianus Casey, 1913 Lectotype
Male adult
«Monterey [Monterey County], California» , in USNM [# 46982].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) occultus Casey, 1913 Lectotype
Female adult
«Siskiyou Co[unty], California» , in USNM [# 47009].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) ordinarius Casey, 1913 Lectotype
Male adult
«North Fork, Madera Co[unty], California» , in USNM [# 47021].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) ovalipennis Casey, 1913 Lectotype
Female adult
«between Indio and Colton [probably in Riverside County], California» , in USNM [# 47023].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) panticulatus Casey, 1913 Holotype
Female adult
«with scarcely a doubt from California» in USNM [# 47034].
Copyright © Kipling W. Will

Pterostichus parens Casey, 1913 Lectotype
Male adult
«Siskiyou Co[unty], Cal[ifornia]», in USNM [# 47008].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) pergracilis (Casey, 1920) Lectotype
Female adult
«Olancha, Inyo Co[unty], California» , in USNM [# 47032].
Copyright © Alexander Anichtchenko

Hypherpes placerensis Casey, 1918 Lectotype
Male adult
«Placer Co[unty], California» , in USNM [# 47015].
Copyright © Kipling W. Will

Pterostichus plutonicus Casey, 1913 Lectotype
Male adult
«coast regions north of San Francisco, California» , in USNM [# 46991].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) protensiformis (Casey, 1924) Lectotype
Male adult
«Berkeley [Alameda County], California» , in USNM [# 47016].
Copyright © Kipling W. Will

Hypherpes protensipennis Casey, 1918 Lectotype
Male adult
«Berkeley [Alameda County], Cal[ifornia]» , in USNM [# 47029].
Copyright © Kipling W. Will

Pterostichus pugetanus Casey, 1913 Lectotype
Female adult
«Clackamas Co[unty], Oregon», in USNM [# 46995].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) sierranus Casey, 1913 Lectotype
Female adult
«Lake Tahoe and Merced Cr[eek], California» , in USNM [# 47028].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) sponsor Casey, 1913 Lectotype
Female adult
«Monterey [Monterey County], California» , in USNM [# 47037].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) suffusus Casey, 1913 Lectotype
Male adult
«Humboldt Co[unty], California», in USNM [# 46993].
Copyright © Kipling W. Will

Pterostichus tahoensis Casey, 1913 Lectotype
Male adult
«Lake Tahoe [Placer County], California» , in USNM [# 47011].
Copyright © Kipling W. Will

Pterostichus (Hypherpes) crenicollis Leconte, 1873
Male adult

Copyright © Kipling W. Will

Pterostichus (Hypherpes) californicus (Dejean, 1828)
Male adult

Copyright © Kipling W. Will

Pterostichus (Leptoferonia) angustus (Dejean, 1828)
Female adult

Copyright © Kipling W. Will

Pterostichus (Hypherpes) protensiformis (Casey, 1924)
Male adult

Copyright © Kipling W. Will

Pterostichus (Hypherpes) vicinus Mannerheim, 1843
Male adult
slightly teneral
Copyright © Kipling W. Will

Pterostichus (Hypherpes) serripes (Leconte, 1875)
Male adult

Copyright © Kipling W. Will

Pterostichus (Leptoferonia) falli Vandyke, 1925
Female adult

Copyright © Kipling W. Will

Pterostichus (Hypherpes) tarsalis Leconte, 1873
Male adult

Copyright © Kipling W. Will

Dyschirius (Dyschiriodes) alticola Lindroth, 1961
Male adult
Photographer: Beatriz Gimenez de Cordoba
SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © ---

Dyschirius (Dyschiriodes) gibbipennis Leconte, 1857
Male adult
Photographer: Beatriz Gimenez de Cordoba
SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © ---

Phrypeus rickseckeri (Hayward, 1897)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Ocydromus) perspicuum (Leconte, 1848)
Male adult
USA: Colorado: Huerfano Co., Huerfano River at Badito, 37.7285°N 105.0167°W.
Scale bar is 1 mm.
Photo David R. Maddison & Alexander P. Swanson
Copyright © ---

Thalassotrechus barbarae (G.Horn, 1892)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Bembidion (Cillenus) palosverdes Kavanaugh and Erwin, 1992
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Phrypeus rickseckeri (Hayward, 1897)
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Trechus (Trechus) humboldti Vandyke, 1945
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidion (Lionepha) australerasa Maddison, 2020 Holotype
Male adult
USA: California: Amador Co., Oyster Lake, Silver Lake Campground
Copyright © David Maddison

Bembidion (Lionepha) lindrothi Maddison & Sproul, 2020 Holotype
Male adult
USA: California: Tulare Co., outlet of Emerald Lake
Copyright © David Maddison

Bembidion (Lionepha) pseudoerasum Lindroth, 1963
Male adult
USA: California: El Dorado Co., Strawberry Creek at Sciots Camp
Copyright © David Maddison

Bembidion (Ocydromus) erosum (Motschulsky, 1850)
Male adult
USA: California: Del Norte Co., Wilson Creek, 3 m, 41.6051°N, 124.1005°W (V101453)
Copyright © David Maddison

Bembidion (Ocydromus) perspicuum (Leconte, 1848)
Male adult
USA: Arizona: Cochise Co., San Pedro R at Charleston, 31.6239°N, 110.1722°W (voucher V101461)
Copyright © David Maddison

Bembidium haplogonum Chaudoir, 1868 (NEOTYPE)
Male adult
USA: California: Lake Co., North Branch Cache Creek at hwy 20, 305 m 38.9881°N, 122.5400°W
Copyright © David Maddison

Bembidion (Notaphus) brownorum Maddison, Sproul & Will, 2023 Holotype
Male adult
EMEC: 39.13841/−122.34621 USA: California: Colusa Co. Antelope Valley, Freshwater Creek uv light pan trap 133 m. 1.vii.2021 K.Will
Copyright © David Maddison

Bembidion (Liocosmius) cachagua Maddison, 2023 Holotype
Male adult
CAS: “U.S.A.: CALIFORNIA: Monterey Co., Hastings Nat. History Reserve. J.M. Linsdale Colln.”, “C.A.S. Accession 29-V-48 Cachagua Creek”, “David R. Maddison DNA3842 DNA Voucher”
Copyright © David Maddison

Anillaspis explanata (Horn, 1888)
Female adult
USA: California: El Dorado Co., Alabaster Cave (CMNH). Scale bar 1 mm; Image from LaBonte & Maddison, 2023
Copyright © - - -

Trechus (Trechus) santaluciaensis Donabauer, 2014 Paratype
Male adult
Monterey County, Big Sur, 28.IV.2013, leg. M. Donabauer; Image from Donabauer, 2014
Copyright © - - -

Trechus (Trechus) conformis Jeannel, 1927
Male adult
Gualale, near campground at Gualale river, 2.V.2013, leg. M. Donabauer. Image from Donabauer, 2014
Copyright © - - -

Trechus (Trechus) humboldti Vandyke, 1945
Male adult
Image from Donabauer, 2014
Copyright © - - -

Bembidion (Trepanedoris) clemens clemens Casey, 1918
Male adult
USA: Arizona: Yavapai Co., Prescott)
Copyright © David Maddison

Bembidion (Trepanedoris) endeca Maddison & Sproul, 2026 Paratype
Male adult
Dairy Creek, Harney Co., Oregon
Copyright © David Maddison

Bembidion (Trepanedoris) connivens (Leconte, 1852)
Male adult
USA: California: Tehama Co., Red Bluff
Copyright © David Maddison

Bembidion (Trepanedoris) altipaludis Maddison, 2026 Paratype
Female adult
USA: Oregon: Deschutes Co., Three Creek Meadow 1925m 44.1153°N 121.6263°W
Copyright © David Maddison

Bembidion (Trepanedoris) ampliceps Casey, 1918
Male adult
USA: California: Kern Co., Stable Creek
Copyright © David Maddison