Carabus (Calocarabus) gologensis Deuve e Tian 2004
Male adult
18 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) gratus gratus Semenov, 1887
Female adult
17 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) gratus nishidai Deuve & Imura, 1993 Paratype
Male adult

Copyright © Boleslav Brezina

Carabus (Calocarabus) gratus nishidai Deuve & Imura, 1993 Paratype
Female adult
19 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) guinanensis guinanensis Deuve, 1991
Male adult
16 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) guinanensis guinanensis Deuve, 1991
Female adult
17 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) guinanensis maqenicola Kalab, 2005 Paratype
Male adult
China, SE Qinghai mts. 15 km E Maqen, 34.27N/100.24E 4300m
J.Kalab leg., 22-24.7.2004 alpine meadows, screes
Copyright © Pierfranco Cavazzuti

Carabus (Calocarabus) linxiaensis linxiaensis Deuve, 1992
Male adult
17 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) linxiaensis linxiaensis Deuve, 1992
Female adult
18 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) linxiaensis linxiaensis Deuve, 1992
Male adult

Copyright © Boleslav Brezina

Carabus (Calocarabus) przewalskii somoli Deuve, 2007 Holotype
Male adult
16 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) przewalskii somoli Deuve, 2007
Female adult
19 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) przewalskii przewalskii Morawitz, 1886 (as elnarae)
Male adult
CINA, Qinghai Nord-Or. Passo fre Daliang e Obo
3600 m, 11.VII.2005 Cavazzuti leg.
Copyright © Pierfranco Cavazzuti

Carabus (Calocarabus) przewalskii przewalskii Morawitz, 1886
Male adult
17 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) przewalskii przewalskii Morawitz, 1886
Female adult

Copyright © Boleslav Brezina

Carabus (Calocarabus) przewalskii przewalskii Morawitz, 1886
Male adult

Copyright © Boleslav Brezina

Carabus (Calocarabus) przewalskii przewalskii Morawitz, 1886
Female adult
19 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) przewalskii sewertzowi Semenov, 1898 Holotype
Male adult
17 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) przewalskii sewertzowi Semenov, 1898
Male adult
19 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) przewalskii sewertzowi Semenov, 1898
Female adult
20 mm
Copyright © Boleslav Brezina

Carabus (Calocarabus) przewalskii paulusianus Deuve, 2007
Male adult
19 mm
Copyright © Boleslav Brezina

Carabus (Cryptocechenus) kalabi Deuve, 1990
Male adult
male, 18mm
Copyright © Boleslav Brezina

Carabus (Cryptocechenus) kalabi Deuve, 1990 (as cuprifuscatus) Paratype
Female adult

Copyright © Boleslav Brezina

Carabus (Calocarabus) przewalskii menyuanensis Deuve, Liu & Liang, 2024 Holotype
Male adult
Chine, province du Qinghai, canton de Menyuan, ville de Zhugu, Sigou, 2602 m, 37,1353°N 102,3887°E (Yizhou Liu & Wenqi Yin leg., 26.VII.2020, dans des pièges), in IOZ. Image from Deuve 2024
Copyright © - - -

Carabus (Coptolabrus) formosus grumorum Semenov, 1898
Female adult
female, 30mm
Copyright © Boleslav Brezina

Carabus (Coptolabrus) formosus grumorum Semenov, 1898
Male adult
male, 24mm
Copyright © Boleslav Brezina

Carabus (Cychrostomus) anchocephalus anchocephalus (as facetus Semenov, 1903)
Female adult
L=24mm
Copyright © Boleslav Brezina

Carabus (Cychrostomus) anchocephalus anchocephalus Reitter, 1896
Male adult
male, 22mm
Copyright © Boleslav Brezina

Carabus (Cychrostomus) anchocephalus anchocephalus Reitter, 1896
Male adult
China, Shanxi pro., Jiaochen
Copyright © Ye Liu

Carabus (Cychrostomus) anchocephalus farinosus Rapuzzi, 2023 Holotype
Male adult
China, Inner Mongolia Province, 30 Km NW Hohhot, 3/7.VI.2010, M. Janata legit. Image from Rapuzzi, 2023
Copyright © - - -

Carabus (Eccoptolabrus) exiguus ochotonarum Brezina, 1996 Holotype Paratype
Female adult

Copyright © Frank Kleinfeld

Carabus (Eccoptolabrus) exiguus ochotonarum Brezina, 1996
Female adult
CH. S-Gansu (Labrang) valley E of Ponggartang
35:14N/102:55E. ca. 2700m conoferous forest
9.VII.1995 K. & B. Brezina
Copyright © Frank Kleinfeld

Carabus (Eccoptolabrus) exiguus ochotonarum Brezina, 1996 Paratype
Male adult

Copyright © Frank Kleinfeld

Carabus (Eccoptolabrus) exiguus evae Brezina et Hackel, 2007 Holotype
Male adult
China, S-Qinghai, 5km S Baima, 30.VI.-1.VII.2006 V. Banes
long of imago 19 mm
Copyright © Boleslav Brezina

Carabus (Eccoptolabrus) exiguus alucinatus Cavazzuti, 2006 Holotype
Female adult
female, 21mm
Copyright © Boleslav Brezina

Carabus (Pagocarabus) wagae folwarcznyi Deuve, 1997 Paratype
Male adult
male, 21 mm
Copyright © Boleslav Brezina

Carabus (Pagocarabus) ? wagae Fairmaire, 1882
Male adult
China, E Tibet, pas Dongdola 5000 m, 4.7.1995
Copyright © Alexander Anichtchenko

Carabus (Eocechenus) mouthiezianoides Deuve & Kalab, 1992 Paratype
Male adult
male, 18mm
Copyright © Boleslav Brezina

Carabus (Eocechenus) kaznakovi nangqenicus Deuve, 1995 Paratype
Male adult

Copyright © Boleslav Brezina

Carabus (Eocechenus) ditomoides Deuve, 1991
Male adult
male, 19mm
Copyright © Boleslav Brezina

Carabus (Pagocarabus) wagae wagae Fairmaire, 1882 Holotype
Female adult
Thibet MNHN) http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec146
Copyright © - - -

Carabus (Pagocarabus) crassesculptus diruptus Morawitz, 1886
Male adult
China, Gansu, Umg. Xiahe 3000-3200m
24-26.6.1993 Heinz leg.
20 mm
Copyright © Alexander Anichtchenko

Carabus (Pagocarabus) crassesculptus tonghua Deuve & Mourzine, 2001
Male adult
China, NW Sishuan, 3500m, Hongyuan, Somang Qu riv. grassland, 9-29.6.2004 (cAA). 20 mm
Copyright © Alexander Anichtchenko

Carabus (Piocarabus) reitterianus Breuning, 1934
Male adult
China, Gansu, ca 25 km E Xiahe 2700-2900m, Wagatan Timber<br /> 30.7.-2.8.1993 Heinz leg.<br /> male, 23 mm
Copyright © Alexander Anichtchenko

Carabus (Piocarabus) vladimirskyi Dejean & Boisduval, 1830
Male adult
China, Beijing, Jiufeng mountain, 2007, Liu ye leg.;<br /> Photo & det. by liuye
Copyright © Ye Liu

Carabus (Piocarabus) vladimirskyi Dejean & Boisduval, 1830
Male adult
Buryatia, Kyahta distr. [Khyagh], 5 km SE Chikoi vill., 16.8.1998, Anichtchenko A. leg.
Copyright © Alexander Anichtchenko

Carabus (Rhigocarabus) thais thais Heinz, 1997
Male adult
male, 16mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) thais thais Heinz, 1997
Male adult
male, 19mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) thais thais Heinz, 1997
Male adult
male, 20mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) roborowskii roborowskii Semenov, 1887
Male adult
China, E Tibet, Chegarla 4800m, 20 km E
of Rioche, 26.6.97 A. Wrzecionko leg.
Copyright © Alexander Anichtchenko

Carabus (Rhigocarabus) morawitzianus breviculus Cavazzuti & Ratti, 1998 Paratype
Male adult
male, 15 mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) dacatraianus Deuve, 1995
Male adult
China, E Tibet, Angenla 5000-5100m
45 km W of Bachen 29.6-2.7.1997 Wrzecionko leg.
16 mm
Copyright © Alexander Anichtchenko

Carabus (Rhigocarabus) cateniger cateniger s.str. (as sommitalis)
Male adult
male, 16 mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) cateniger cateniger Morawitz, 1886
Female adult
female, 19mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) cateniger cateniger Morawitz, 1886
Male adult
male, 18mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) cateniger dabanicola Deuve, 1998
Male adult
male, 16mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) cateniger dabanicola Deuve, 1998
Female adult
female, 18mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) buddaicus bocsiki Deuve, 2007 Holotype
Male adult
male, 20 mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) buddaicus angenlaicus Deuve, 2007 Paratype
Male adult
male, 18 mm
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) morawitzianus breviculus Cavazzuti & Ratti, 1998 Paratype
Male adult
China, N Tibet, Tangula Shan Mts., Tangula, pass 5100-5300m, 8-11.7.1997 A. Wrzecionko leg.
Copyright © Alexander Anichtchenko

Carabus (Rhigocarabus) buddaicus gansuicus Deuve, 1991
Male adult
China, NC-Sichuan (ZOIGE), LANGMUSI (=DOGCANGLHAMO), 34:03:48N/102:38:08E, h=4100m, alpine meadows, screes, 16-18.VII.1999, B.Brezina leg.
L=20 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) buddaicus linxiaicus Deuve, 1995
Male adult
China, S-Gansu (LABRANG):, valley.E of Ponggartang, 35:12-13N/102:54-55E, h=3000-3700m, alpine meadows, coniferous forests,10.-14.VII.1999, B.Brezina leg. L=20 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) morawitzianus morawitzianus Semenov, 1887
Male adult
C China, Qinghai, Zadoi County, Toramarkog h~4600 m, VI.2001, P. Cavazzuti leg. L=17 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) morawitzianus morawitzianus Semenov, 1887
Male adult
Qinghai, Anemaqen mts pass, 20 km N of Maduo town, 4500 m, 6.7.2012, V. Patrikeev leg.; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Carabus (Rhigocarabus) buddaicus gansuicus Deuve, 1991
Male adult
N Sichuan, Langmusi vill. 3700 m, 1-25.7.2018, Leg. V. Patrikeev. det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Carabus (Rhigocarabus) thais thais Heinz, 1997
Male adult
E Qinghai-Gansu border, Dalia pass 50km W Lixia town 3800m 20.7.2019 (cAA) det. Anichtchenko 2020
Copyright © Alexander Anichtchenko

Carabus (Rhigocarabus) cateniger copulans Semenov, 1898
Male adult
E Qinghai, Laji Shan, near Garang, 3900m, 19.6.2011, Patrikeev V. leg (cAA) det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Carabus (Rhigocarabus) cateniger papaoensis Cavazzuti, 2006
Male adult
E Qinghai, near Babao town, 3800m 5.8.12 Patrikeev V. leg. (cAA) det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Carabus (Rhigocarabus) cateniger cateniger Morawitz, 1886
Male adult
NW Gansu, 80 km S Jayuguan, Tulai shan, Qionglai mts, 10.7.2014, 4200 m, Patrikeev leg (cAA)
Copyright © Alexander Anichtchenko

Carabus (Rhigocarabus) cateniger cateniger (as sommitalis Cavazzuti, 2006)
Female adult
China, C-Gansu, Mts Qilian h=3378 m, road G227, Yangxiongzi env. 38.050917, 100.887267, 18-23.VI.2011, G.Proscia leg., L=19 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) cateniger cateniger (as sommitalis Cavazzuti, 2006)
Male adult
China, C-Gansu, Mts Qilian h=3378 m, road G227, Yangxiongzi env. 38.050917, 100.887267, 18-23.VI.2011, G.Proscia leg., L=17 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) cateniger dabanicola Deuve, 1998
Male adult
China, Qinghai, Xining city, 55km NE of Datong, Daban Shan h=3936 m, 37.348450, 101.397600, 17-23.VI.2011, G.Proscia leg., L= 16 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) cateniger dabanicola Deuve, 1998
Female adult
China, Qinghai, Xining city, 55km NE of Datong, Daban Shan h=3936 m, 37.348450, 101.397600, 17-23.VI.2011, G.Proscia leg., L= 18.5 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) roborowskii roborowskii Semenov, 1887
Male adult
China, E-Qinghai, Mts NW of Tongren city, 35.562778, 101.843667, h=3870 m, 15-26.VI.2011, G. Proscia leg./det.R.Panin, L=20 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) roborowskii roborowskii Semenov, 1887
Female adult
China, E-Qinghai, Mts NW of Tongren city, 35.562778, 101.843667, h=3870 m, 15-26.VI.2011, G. Pros cia leg./det.R.Panin, L=20 mm
Copyright © Ruslan Panin

Carabus (Rhigocarabus) degensis ladyginioides Deuve, Liu & Liang, 2024 Holotype
Male adult
Chine, province de Qinghai, canton de Madoi, col de Bayan Har, 4827 m, 34,1267°N 97,6575°E, pelouses alpines et éboulis (Hongliang Shi, Zhongbao Lu, Weifeng Yan & Pingzhou Zhu leg., 30.VII.2017, dans des pièges), in IOZ. Image from Deuve et al, 2024
Copyright © - - -

Carabus (Rhigocarabus) roborowskii sininensis Semenov, 1898 (as lajiensis Deuve & Imura, 1991)
Male adult
E Qinghai, E Laji Shan, N of Hualun, 3900m, 28.7.2019 Patrikeev leg (cAA). det. Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Rhigocarabus) ladygini jigzhianus Cavazzuti, 2006
Female adult
S Quinghai, Guoluo mts., NW part, 4200-4500m, 11.7.17 Patrikeev V. leg (cAA). det. Anichtchenko A.
Copyright © Alexander Anichtchenko

Carabus (Rhigocarabus) buddaicus cholaensis Deuve, 2008 Holotype
Male adult
W-Sichuan: Chola Shan Pass, 4500m (in coll Benes, Prague)
Copyright © Thierry Deuve

Carabus (Rhigocarabus) buddaicus angenlaicus Deuve, 2008 Holotype
Male adult
E-Tibet: 30km E of Yanga, Angen La Pass 4800m (in coll Wrzecionko)
Copyright © Thierry Deuve

Carabus (Rhigocarabus) buddaicus obenbergeri Semenov & Znojko, 1932
Male adult
Qinghai, S of Kukunor lake, pass near Heka, 25.7.2010 (cAA)
Copyright © Alexander Anichtchenko

Carabus (Scambocarabus) sculptipennis Chaudoir, 1887
Male adult
China, Beijing, Xiaolongmeng, 2005, Liu ye coll.;
22mm; photo by Liuye
Copyright © Ye Liu