Brachinus (Brachynolomus) stenoderus Bates, 1873
Endophallus
Collection ground Hobetsu cho, Hokkaido Pref. 33485
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Brachinus (Brachynolomus) scotomedes L. Redtenbacher, 1867
Endophallus
Collection ground Tochigi Pref.Fujioka～machi 34784
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pheropsophus (Stenaptinus) javanus (Dejean, 1825)
Endophallus

Copyright © Dmitri Fedorenko

Carabus (Eucarabus) manifestus pyonganicola Deuve & Li, 1998
Endophallus

Copyright © Ruslan Panin

Carabus (Diocarabus) fraterculus fraterculus Reitter, 1895
Endophallus
N-Korea, N Pyongan Prov.,Sakju, Mt Ponsan,
Copyright © Ruslan Panin

Carabus (Eucarabus) angustus dopyeong Rapuzzi, 2015
Endophallus
scale bar = 5mm
Copyright © Jong-Bong Choi

Carabus (Eucarabus) sternbergi sternbergi Roeschke, 1898
Endophallus
Scale bar = 5mm
Copyright © Jong-Bong Choi

Carabus (Eucarabus) sternbergi gureopensis J.K. Park & J. Park, 2013 Paratype
Endophallus
Scale bar = 5mm
Copyright © Jong-Bong Choi

Carabus (Eucarabus) cartereti cartereti Deuve, 1982
Endophallus
Scale bar = 5mm
Copyright © Jong-Bong Choi

Carabus (Eucarabus) nitididorsus nitididorsus Ishikawa & Kim, 1983
Endophallus
Scale bar = 5mm
Copyright © Jong-Bong Choi

Carabus (Eucarabus) nitididorsus paegunsanus Imura, 1997
Endophallus
Scale bar = 5mm
Copyright © Jong-Bong Choi

Carabus (Eucarabus) namhaedoensis Kwon & Lee, 1984
Endophallus
Scale bar = 5mm
Copyright © Jong-Bong Choi

Carabus (Hemicarabus) tuberculosus Dejean, 1829
Endophallus
Carabus (Hemicarabus) tuberculosus n. strophium F.-W., 1842 SE Kazakhstan: NW Tarbogatai Mt.R.
Copyright © Igor Solodovnikov

Galerita (Galerita) orientalis Schmidt-Goebel, 1846
Endophallus
Higashihagio, Omuta City, Fukuoka pref., Japan, 21 may 2016, S. Nishimoto leg.
Copyright © Alexander Anichtchenko

Trichotichnus (Trichotichnus) vespertinus Habu, 1954b: 57
Endophallus
Collection ground Ashikaga city,Tochigi Pref.Ashikaga～shi 36079
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Trichotichnus (Trichotichnus) lucidus (A. Morawitz, 1863)
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 39970
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Trichotichnus (Trichotichnus) congruus Motschulsky, 1866a: 164
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 39970
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Stenolophus (Stenolophus) agonoides Bates, 1883
Endophallus
Collection ground Kobe city, HyogoPref. 38129
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Stenolophus (Stenolophus) propinquus A.Morawitz, 1862
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 39970
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Stenolophus (Egadroma) difficilis (Hope, 1845)
Endophallus
Collection ground Kamogawa city, ChibaPref. 34922
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) simplicidens Schauberger, 1929
Endophallus
Collection ground Toride city, Ibaraki pref. 28346
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) eous Tschitscherine, 1901
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 38963
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) roninus Bates, 1873
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 39324
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) tridens A.Morawitz, 1862
Endophallus
Collection ground Hokkaido Pref. Fukushima～cho 34916
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) griseus (Panzer, 1796)
Endophallus
Collection ground Hokkaido Pref. Matsumae～cho 32398
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Zangoharpalus) tinctulus Bates, 1873
Endophallus
Collection ground Sapporo city, Hokkaido Pref. pref. 33415
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) pallidipennis A. Morawitz, 1862a: 260
Endophallus
Collection ground Higashisonogi cho, Nagasaki Pref. 35357
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) crates Bates, 1883: 239
Endophallus
Collection ground Chinzei cho, Saga pref. 35168
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) corporosus (Motschulsky, 1861)
Endophallus
Collection ground Horonobe cho, Hokkaido Pref. 33807
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) chalcentus Bates, 1873a: 263
Endophallus
Collection ground Minobu cho, Yamanashi Pref. 35617
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Harpalus) bungii Chaudoir, 1844: 451
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 2000/pit
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Loboharpalus) platynotus Bates, 1873
Endophallus
Collection ground Suttsu cho, Hokkaido Pref. 33499
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Pseudoophonus) jureceki (Jedlicka, 1928)
Endophallus
Collection ground Sapporo city, Hokkaido Pref. 34554
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Bradycellus (Tachycellus) subditus (Lewis, 1879)
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 39249
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Anisodactylus (Anisodactylus) tricuspidatus A.Morawitz, 1863
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 39970
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Anisodactylus (Pseudanisodactylus) punctatipennis A.Morawitz, 1862
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 39970
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Anisodactylus (Pseudanisodactylus) signatus (Panzer, 1796)
Endophallus
Collection ground Mukawa cho, Hokkaido Pref. 37843
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Acupalpus (Palcuapus) inornatus Bates, 1873
Endophallus
Collection ground Kamogawa city, ChibaPref. 34922
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Acupalpus (Setacupalpus) hilaris Tschitscherine, 1899
Endophallus
Russia, Evreiskaya AO, Birobidzhan vic., 3.11.2013. leg. Budilov. P.
Copyright © Pavel Budilov

Trichotichnus (Trichotichnus) vespertinus Habu, 1954b: 57
Endophallus
Scale bar: 0.6 mm.
Copyright © Dooyoung Kim

Trichotichnus (Trichotichnus) miser (Tschitscherine, 1897)
Endophallus
Scale bar: 0.5 mm.
Copyright © Dooyoung Kim

Trichotichnus (Bottchrus) nanus Habu, 1954c: 233
Endophallus
Scale bar: 0.5 mm.
Copyright © Dooyoung Kim

Anisodactylus (Anisodactylus) tricuspidatus A.Morawitz, 1863
Endophallus
Primorye Territory, Khasan Distr., vicinity of Slavyanka Vill., Cape Busse, 42°52'42"N 131°27'58"E, 13.VI.2023, I. Melnik leg. L=11.0mm
Copyright © Kirill Makarov

Anisodactylus (Pseudanisodactylus) punctatipennis A.Morawitz, 1862
Endophallus
Primorye Territory, Khasan Distr., 12.5 km WNW of Barabash Vill., Ovchinnikovo, 43°13'56"N 131°21'23"E, h~110 m, 10–13.VIII.2020, K. Makarov, A. Matalin leg. L=11.1 mm
Copyright © Kirill Makarov

Odacantha (Heliocasnonia) aegrota (Bates, 1883)
Endophallus
Collection ground Kujuu machi, Ooita pref. 35966
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Anaulacus (Aephnidius) adelioides W.S.MacLeay, 1825
Endophallus
Collection ground Hokuto city, YamanashiPref. 37119
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Coptodera (Coptoderina) subapicalis Putzeys, 1877a: 84
Endophallus
Collection ground Shonai machi, OoitaPref. 34095<br /> (C) Working Group for Biological Indicator Ground Beetles Database, Japan<br /> Prohibited secondary use of all data
Copyright ©

Lebia (Poecilothais) bifenestrata A. Morawitz, 1862a: 245
Endophallus
Collection ground Misugi machi, Mie pref. 34126
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Lebia (Poecilothais) retrofasciata Motschulsky, 1864: 227
Endophallus
Collection ground Toyoura cho, Hokkaido Pref. 34482
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Coptodera (Coptoderina) japonica Bates, 1883
Endophallus
Taiwan. Image from Hunting & Yang, 2019
Copyright © - - -

Diplocheila (Isorembus) zeelandica L. Redtenbacher, 1867: 5
Endophallus
Collection ground Shimamaki mura, Hokkaido Pref. 33049
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Chlaenius (Chlaeniostenus) circumdatus xonthopleurus Chaudoir, 1856
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 38879
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Chlaenius (Achlaenius) kurosawai Kasahara, 1986d: 147
Endophallus
Collection ground Shimamoto cho, OosakaPref. 35471
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Badister (Badister) pictus Bates, 1873
Endophallus
Collection ground Noda city, iChiba pref. 34517
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Dischissus mirandus Bates, 1873: 244
Endophallus
Collection ground Tsushima, Nagasaki pref. 34430
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Dischissus japonicus Andrewes, 1933: 5
Endophallus
Collection ground Tsushima, Nagasaki pref. 35695
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Panagaeus (Panagaeus) japonicus Chaudoir, 1861c: 356
Endophallus
Collection ground Kaminokuni cho, Hokkaido Pref. 35988
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Panagaeus (Panagaeus) robustus A. Morawitz, 1862b: 323
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 36670
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Tinoderus singularis Bates, 1873a: 245
Endophallus
Collection ground Fujioka machi, TochigiPref. 34784
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Peronomerus auripilis Bates, 1883: 235
Endophallus
Collection ground Taiki cho, Hokkaido Pref. 34157
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Metacolpodes buchanani Hope, 1831: 21
Endophallus
Russia, Evreiskaya AO., Birobidjan 31.07.2013
Copyright © Pavel Budilov

Amara (Bradytus) sinuaticollis A. Morawitz, 1862
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 38963
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Curtonotus) macronota Solsky, 1875
Endophallus
Collection ground Tomakomai city, Hokkaido Pref. 37172
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Curtonotus) hiogoensis (Bates, 1873)
Endophallus
Collection ground Tsushima, NagasakiPref. 35695
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Curtonotus) gigantea (Motschulsky, 1844)
Endophallus
Collection ground Kagoshima city, Kagoshima Pref. 38979
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Amara) obscuripes Bates, 1873a: 294
Endophallus
Collection ground Shibecha cho, Hokkaido Pref. 38493
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Amara) congrua A. Morawitz, 1862b: 326
Endophallus
Collection ground Ise city, Mie Pref. 37125
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Pterostichus (Bothriopterus) subovatus (Motschulsky, 1860)
Endophallus
Collection ground Rebun Is. Hokkaido Pref. 27191
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Amara (Amara) silvestrii Baliani, 1937: 179
Endophallus
China, Gaoligong Shan. Image from Kavanaugh, Hieke, Liang & Dong, 2014
Copyright © - - -

Amara (Curtonotus) gigantea (Motschulsky, 1844)
Endophallus
Russia, Primorskii reg.
Copyright © Alexander Anichtchenko