Pheropsophus (Stenaptinus) javanus (Dejean, 1825)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Brachinus (Brachynolomus) scotomedes L. Redtenbacher, 1867
Female adult
Japan, Higashihagio Town, Omuta City, Fukuoka Pref., 19.6.2011, S. Nishimoto leg.
Copyright © Alexander Anichtchenko

Eobroscus (Eobroscus) lutshniki Roubal, 1928d: 90
Male adult
Far East of Russia, S Sakhalin isl., Aniva dist., Taranai vill., 30.07.2006, O. Legesin leg.
Anichtchenko A. det.
Copyright © Alexander Anichtchenko

Craspedonotus tibialis Schaum, 1863
Male adult

Copyright © Kirill Makarov

Carabus (Eucarabus) manifestus pyonganicola Deuve & Li, 1998
Male+Female adult
N-Korea: Pyonganam-Do, YangTok County,(39;127), 12-19,VI.2006
Copyright © Ruslan Panin

Carabus (Eucarabus) manifestus guanmenshanus Imura, 1991
Female adult
NE China, Liaoning, Anshan City, Mt.Qian Shan, 4-16.VII.2005, I.Rapuzzi leg.
L=26 mm
Copyright © Ruslan Panin

Carabus (Hemicarabus) tuberculosus Dejean, 1829
Male adult
Far East, Lazovsky Dist., shore of Japan Sea, 3 km NE Preobrazhenie Vill., Ezhovaya (Tasovaya) Bay 13.07.06 M. & L. Smirnov's leg.
male
L=20,0 mm
Copyright © Maksim Smirnov

Calosoma (Charmosta) lugens Chaudoir, 1869
Male adult

Copyright © Kirill Makarov

Carabus (Diocarabus) fraterculus fraterculus Reitter, 1895
Male adult
N-Korea, N Pyongan Prov.,Sakju, Mt Ponsan, 20-29.VI.2009, L. Bieber leg.<br /> L=19 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) cartereti omisanensis Lassalle, 1999
Male adult
South Korea, Gangwon Province, Kyebang-san, h=1100m, VI.1998 L=24 mm
Copyright © Ruslan Panin

Carabus (Coptolabrus) jankowskii fusanus Born, 1907
Female adult
South Korea, S Gyeongsang Prov., Busan, Mt Geumwonsan, 17.VI.2017, L=37 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) manifestus guanmenshanus Imura, 1991
Male adult
China, Liaoning, Tieling County, Sandagou env., h=850 m, 13-18.VI.2011
Copyright © Ruslan Panin

Carabus (Diocarabus) fraterculus neochinensis Deuve & Li, 1998
Female adult
NE-China: Liaoning prov., Fengcheng City, Shicheng, 20-28.VI.2011, Li Jingke leg., L=22 mm
Copyright © Ruslan Panin

Carabus (Diocarabus) fraterculus neochinensis Deuve & Li, 1998
Male adult
NE-China: Liaoning prov., Fengcheng City, Shicheng, 20-28.VI.2011, Li Jingke leg., L=20 mm
Copyright © Ruslan Panin

Cylindera (Eugrapha) elisae s.str. (as amurensis)
Female adult
Far East, Lazovskij Nat. Res., Proselochnaja Riv. near nouth, beach 43°00'34"N 134°07'43"E 17-21.VIII.2007 K.Makarov & A.Zaitsev leg.<br /> L=10.0 mm
Copyright © Kirill Makarov

Cicindela (Sophiodela) chinensis flammifera W.Horn, 1921
Female adult
S Korea, Aebonsan (=Yebongsan, Suwon-si, Gyeonggi-do) Mt., 22.V.2014, L=26 mm
Copyright © Ruslan Panin

Abroscelis anchoralis punctatissima (Schaum, 1863)
Female adult
Japan, Ishikawa (JWGC). det. J. Wiesner
Copyright © Peter Schüle

Cylindera (Cylindera) gracilis (Pallas, 1773)
Male adult
Russia, Tuva, Kaa-Khemsky Dist., near Boyarovka vill., Koptu river valley, h=1000 m, 8-9.VII.2003, S. Vashchenko leg.
Copyright © Ruslan Panin

Planetes (Planetes) puncticeps Andrewes, 1919d: 480
Male adult
Collection ground Tsushima, Nagasaki pref. 35695
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Drypta (Drypta) ussuriensis Jedlicka, 1963
Male adult
SE Приморский кр., Лазовский зап., ~10 км SW пос. Сокольчи, окр. кордона Корпадь 43°15 N,134°7 E 23-27.06.10 leg. М.Смирнов
Image of M. Smirnov (М.Э. Смирнов) from www.zin.ru\Animalia\Coleoptera
Copyright © Maksim Smirnov

Dendrocellus geniculata (Klug, 1834)
Male adult
9,2 mm Thailand, Phang Nga Prov., Khao Lak, Khao Lak Beach, at light leg. U.Schmidt, 29.IV.2015; det. M.Baehr, 2015; Photo: U.Schmidt, 2015
Copyright © Udo Schmidt

Dendrocellus geniculatus (Klug, 1834)
Male adult
Philippines, Mindanao, Bukidnon, Kalatungan, IX.2014. det. A, Anichtchenko, 2017
Copyright © Alexander Anichtchenko

Galerita (Galerita) orientalis Schmidt-Goebel, 1846
Male adult
Higashihagio, Omuta City, Fukuoka pref., Japan, 21 may 2016, S. Nishimoto leg.
Copyright © Alexander Anichtchenko

Elaphrus (Elaphroterus) punctatus Motschulsky, 1844
Female adult
S Siberia, Tuva rep., Such vill., Mezel riv., 10.6.2002
Copyright © Alexander Anichtchenko

Elaphrus (Elaphrus) comatus Goulet, 1983
Male genitalia
China, Liaoning, Kuandian Co., Banlaing, Ba-che-chuan
Copyright © Alexander Anichtchenko

Elaphrus (Neoelaphrus) sibiricus Motschulsky, 1844
Male adult
SE Siberia, Burijatia, E Sayan Mts., Mondy vill. 800m, 8.6.2002 (cAA). det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Loxoncus (Loxoncus) circumcinctus (Motschulsky, 1858)
Female adult
SE Primorskii reg., Lazo Natural res., seashore
Nachingouz bay, 6.07.06 leg. Yu. Sundukov
female, 9 mm
Photo from www.zin.ru
Copyright © Maksim Smirnov

Harpalomimetes orbicollis N. Ito, 1995b: 278 Holotype
Male adult
Watarase, Fujioka, Tochigi Pref., Japan, 21. V. 1968, A. Habu. COL-267
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Platymetopus flavilabris (Fabricius, 1798)
Male adult
Collection ground Higashisonogi cho, NagasakiPref. 35330
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Zangoharpalus) tinctulus Bates, 1873
Male adult
Collection ground Sapporo city, Hokkaido Pref. pref. 33415
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Harpalus (Loboharpalus) platynotus Bates, 1873
Male adult
Collection ground Suttsu cho, Hokkaido Pref. 33499
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Acupalpus (Stenolophidius) inornatus Bates, 1873
Male adult
Acupalpus (Palcuapus) inornatus Bates, 1873 Collection ground Kamogawa city, Chiba Pref. 34922 (C) Working Group for Biological Indicator Ground Beetles Database, Japan Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Stenolophus (Stenolophus) propinquus A.Morawitz, 1862
Male adult
Russia, SE Primorie, Lazo Nat. res., ~10 км SW Sokolchi vill., Korpad' vicinity 43°15'N / 134°7'E, 23-27.06.10, leg. M. Smirnov
Фотография с сайта zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Harpalus (Harpalus) crates Bates, 1883
Male adult
SE Maritime Prov., Lazovsky dist., Lazo env., 17-20.VI.2010, leg. М. Smirnov Фотография с сайта zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Harpalus (Pseudoophonus) capito A.Morawitz, 1862
Female adult

Copyright © Kirill Makarov

Harpalus (Nephoharpalus) pallidipennis A. Morawitz, 1862
Male adult
Image from Kataev, 2023
Copyright © - - -

Calleida (Callidiola) lepida L.Redtenbacher, 1867
Male adult
Japan, Ehime pref., Komeno-machi, Matsuyama-shi
8.VII.2003 Kurihara T. leg.
male, long. 11 mm
Copyright © Alexander Anichtchenko

Odacantha (Odacantha) puziloi Solsky, 1875
Male adult
Russia, Primorskii reg.,
Khanka lake. 3.VI.2008
long. 7 mm
Copyright © Alexander Anichtchenko

Paradromius (Manodromius) ruficollis (Motschulsky, 1844)
Male adult
Kunashiri Isl., Calder of Golovnina Volcano, env. Ozernyi post 43°52'26"N 145°28'56"E 12.VIII.2009 leg.K.Makarov, A.Zaitzev
5.5 mm
Copyright © Kirill Makarov

Lachnolebia cribricollis (A.Morawitz, 1862)
Male adult

Copyright © Kirill Makarov

Orionella lewisii (Bates, 1873) (obenbergeri Jedlicka, Type) Holotype
Female adult
Mt. Minoo, Osaka, 26.X.1944, leg. Ueno (NMPC)
Copyright © Alexander Anichtchenko

Mochtherus tetraspilotus (W.S.Macleay, 1825)
Female adult
India, Uttarakhand, left side of Kosi riv., 5 km N Ramnagar, N29*432' E79*140', 19-23.06.2011 Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Lachnoderma asperum Bates, 1883
Male adult
CHINA, Taiwan, Nantou, Puli, Shizaitou;
1995.VII.25; Chen C.C. coll.
scale bars = 2.0 mm
Image from Shi H., Zhou H. & Liang H., 2013
Copyright © - - -

Dromius (Dromius) quadraticollis A.Morawitz, 1862
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Parena (Crossoglossa) laesipennis (Bates, 1873)
Male adult
Taiwan
Copyright © Hongliang Shi

Anaulacus (Aephnidius) adelioides W.S.MacLeay, 1825
Female adult
Philippines, Mindanao, S Cotobato, Lake Holon, 5-10.10.2019, Anichtchenko A. leg & det. (cAA)
Copyright © Alexander Anichtchenko

Diplocheila (Isorembus) zeelandica L. Redtenbacher, 1867: 5
Male adult
Collection ground Shimamaki mura, Hokkaido Pref. 33049<br /> (C) Working Group for Biological Indicator Ground Beetles Database, Japan<br /> Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Diplocheila (Neorembus) latifrons latifrons
Female adult
THAILAND, Province Nakhon Ratchasima (Korat), near Nong Bun Nak, h = 200 m, 14°41'25"N 102°27'45.7"E, 08.02-08.03.2008 , leg. A.V. Korshunov
Copyright © Igor Solodovnikov

Nebria (Orientonebria) coreica Solsky, 1875
Male adult

Copyright © Kirill Makarov

Nebria (Sadonebria) chinensis Bates, 1872
Male adult
China, Liaoning prov., Kuandian co., Banlaing, Ba-che-chuan 6.2006 13 mm
Copyright © Alexander Anichtchenko

Nebria (Reductonebria) ochotica R.F. Sahlberg, 1844
Female adult
Russia, Khabarovsk reg., Verhnebureinskii diistr., Bureya riv vall., Ussomah riv. 22.7.2006
Copyright © Alexander Anichtchenko

Chlaenius (Achlaenius) sericimicans Chaudoir, 1876
Male adult
China, Hunan pro.
Photo & det. by liuye
Copyright © Ye Liu

Chlaenius (Achlaenius) variicornis A. Morawitz, 1863
Male adult
Photo from www.zin.ru
Copyright © Kirill Makarov

Chlaenius (Chlaeniellus) circumductus A. Morawitz, 1862
Male adult
China, Heilongjiang Pro., Ussuri river, 2009.5.20, liuye coll.<br /> photo & det. by liuye
Copyright © Ye Liu

Chlaenius (Chlaeniostenus) circumdatus Brulle, 1835
Male adult
China, Shichuan pro., 2007, Bi W.X. coll.;
photo by Bi Wenxuan
det. by liuye
Copyright © Bi W.X.

Chlaenius (Chlaenites) spoliatus motschulskyi Andrewes, 1928
Male adult
China, Shandong pro., Binzhou, Bank of Huanghe river, 2009.10, Liu ye coll.
photo & det. by liuye
Copyright © Ye Liu

Chlaenius (Epomis) nigricans Wiedemann, 1821: 110
Male adult
China, Jiangxi pro., 1975.5.16, Zhang Y.W. coll.;
photo & det. by liuye
Copyright © Ye Liu

Lachnocrepis japonica Bates, 1873
Male adult

Copyright © Kirill Makarov

Peronomerus auripilis Bates, 1883: 235
Male adult
Collection ground Taiki cho, Hokkaido Pref. 34157
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Panagaeus (Panagaeus) japonicus Chaudoir, 1861c: 356
Male adult
SE Maritime Prov., Lazovsky dist., Japan Sea coast, Tachingousa bay (Prosyolotchnaya) env. 2-5.VII.10 leg.М. Smirnov
Фотография с сайта zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Panagaeus (Panagaeus) robustus A. Morawitz, 1862
Male adult

Copyright © Kirill Makarov

Agonum (Glaucagonum) sylphis (Bates, 1873)
Male adult
Kunashir isl., Ozernaya riv., 43 53.07N 145 27.44E 19.VII.2008 leg. K.Makarov
Copyright © Kirill Makarov

Euplynes batesi Harold, 1877
Male adult
S Primorie,, 20 km NNW Spassk-Dalny, Khanka Lake Reserve, kordon “Vostochny” 3-8.VI.2008, leg. A.Napolov
L=6.8
Copyright © Kirill Makarov

Agonum (Lissagonum) lampros (Bates, 1873)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Metacolpodes buchanani Hope, 1831: 21
Male adult

Copyright © Kirill Makarov

Xestagonum xestum (Bates, 1883)
Male adult
JAPAN, Nagano-ken,Sakae-mura,Sakai, 36.8319°N, 138.6145°E, alt. 854 m, 6. V. 2018, S. Uchida leg.
Copyright © Shûta Uchida

Pristosia proxima A. Morawitz, 1862
Male adult
S Primorie, Lazovsky Reserve, kordon “Korpad” 43°15'17"N 134°07'59"E Barber traps 7-8.VIII.2007 leg. K.Makarov, A.Zaitzev
L=12.5
Copyright © Kirill Makarov

Synuchus (Synuchus) nitidus Motschulsky, 1861
Male adult
Kunashiri Isl., Stolbovskie Springs south of Stolbchaty Cape 44°00'26"N 145°40'59"E h~42 23.VIII.2008 leg.I.Melnik
L=14.3
Copyright © Kirill Makarov

Caelostomus (Caelostomus) picipes (W.S.Macleay, 1825)
Male adult
India, Uttarakhand, 10 km NNE of Ramnagar, gorge of left trib. of Kosi River, 400-450 m,
29°29' N, 79°09,5' E, 19-20.04.2012, R. Dudko, I. Lyubechanskii
det. R. Dudko 2012
Copyright © Roman Dudko

Amara (Curtonotus) gigantea (Motschulsky, 1844)
Male adult

Copyright © Kirill Makarov

Trigonotoma lewisii Bates, 1873
Male adult
Japan, Chiba penins., Takeoka, Boso, 3.XII.2009. det. A. Anichtchenko
Copyright © Alexander Anichtchenko

Cosmodiscus platynotus Bates, 1873
Female adult
Vietnam, Lam Dong Province, Bi Doup-Nui Ba Nature Reserve, env. Long Lanh, 12°10´44´´N/108°40´44´´E, h=1400–1600 m, at light, 7–20.IV.2008 or 28.IV–10.V.2009
Copyright © Dmitri Fedorenko

Pterostichus (Koreonialoe) ishikawai Nemoto, 1988
Female adult
Sud Korea, Juwangsan National Park
Copyright © Vittorino Monzini

Pterostichus (Micronialoe) chogyesanus Park; Kwon & Lafer, 1996
Male adult
male
Copyright © Dooyoung Kim

Pterostichus (Micronialoe) spinulosus Kim & Suh, 2024 Holotype
Male adult

Copyright © Dooyoung Kim

Pterostichus (Phaenoraphis) sagittus Park, 2013 Holotype
Male adult
5-6,VI,2012, Mt. Jeombongsan, Jindong-ri,Girin-myeon, Inje-gun, Gawangwondo Prov. Coll.D.P.Lyu. Image from Park et al, 2013
Copyright © - - -

Rhysodes comes (Lewis, 1888)
Unsexed
Primorje reg., Ussuriyskiy distr., near Kamenushka vill. 10.8.1993
Copyright © Alexander Anichtchenko

Dyschiriodes (Eudyschirius) globosus Herbst, 1784: 142
Male adult

Copyright © Kirill Makarov

Pogonus (Pogonoidius) punctulatus Dejean, 1828
Male adult
Volgograd Reg., Pallas Distr., Elton Vill., on UV-light 16-18.VII.2006 K.Makarov leg.
L=6.4 mm
Copyright © Kirill Makarov

Diplous (Platidius) dolini Zamotajlov, 2005
Female adult
S Primorie, Lazovsky Reserve, Formosova Spring 43°00'06"N 134°05'13"E 10.VII.2005 leg. K.Makarov
L=9.3
Copyright © Kirill Makarov

Bembidion (Odontium) chloropus Bates, 1883: 277
Male adult
Far East, Lazovskij Nat. Res., Proselochnaja Riv. near nouth, beach 43°00'34"N 134°07'43"E 17-21.VIII.2007 K.Makarov & A.Zaitsev leg. L=5.6 mm
Copyright © Kirill Makarov

Archipatrobus flavipes (Motschulsky, 1864)
Male adult
Collection ground Kobe city, Hyogo Pref. 33645<br /> (C) Working Group for Biological Indicator Ground Beetles Database, Japan<br /> Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Cillenus (Desarmatocillenus) yokohamae (Bates, 1883)
Male adult

Copyright © Kirill Makarov

Bembidion (Notaphocampa) niloticum Dejean, 1831
Female adult
4,4 mm
Namibia, Etosha National Park, Okaukuejo, 1400 mб leg. U.Schmidt, 18.-20.III.1994;
det. M.Baehr, 2012
Photo: U.Schmidt, 2014
Copyright © Udo Schmidt

Sakagutia umi Sasakawa, 2007
Male adult
SE Приморье, Лазовский р-н., побережье Японского моря, 3 км NE пос. Преображение бухта Ежовая (Тасовая), литораль, 12.07.2010, leg. М. Смирнов. Image from www.zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright © Maksim Smirnov

Bembidion (Odontium) chloropus Bates, 1883: 277
Male adult

Copyright © Kirill Makarov

Bembidion (Notaphocampa) niloticum Dejean, 1831
Female adult

Copyright © Kirill Makarov

Bembidion (Desarmatocillenus) aestuarii (Habu & Ueno, 1954)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison