Carabus (Carabus) donjuan Imura, 2011 Holotype
Endophallus
Image from Imura, 2011
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Carabus (Carabus) koidei yanyuan Cavazzuti & Rapuzzi, 2010
Endophallus
Image from Imura, 2011
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Carabus (Acoptopterus, ?Carabus) yunnanus yanyuanicus Cavazzuti, 1996
Endophallus
China, S-Sichuan, S Yanyuan, Chuandongzi Mt h=3300-3600 m
Copyright © Ruslan Panin

Carabus (Apotomopterus) protenes baoxingensis Deuve & Mourzine, 2003
Endophallus
China: С-Sichuan:Qionglai Shan Mts., Baoxing env. (TOPOTYPUS)
Copyright © Igor Solodovnikov

Carabus (Apotomopterus) delavayi parvipes Cavazzuti, 1996
Endophallus
S-Sichuan: S of Yanyuan, Chuandongzi Massif h=3200 m, 27.351206, 101.507036
Copyright © Ruslan Panin

Carabus (Apotomopterus) ludivinae wuliangensis Cavazzuti, 1997
Endophallus

Copyright © Ruslan Panin

Carabus (Apotomopterus) protenes ascendens Semenov, 1898
Endophallus
China, Chongqing, N Jinfo Shan, Shanwangping h=1400m, 29.101944, 107.166389
Copyright © Ruslan Panin

Carabus (Apotomopterus) korellianus Kleinfeld, 2002
Endophallus
China, SW-Yunnan, Dehong Distr., Ruili, Pijiazhai vill., h=1800 m, 24.048111, 97.738222
Copyright © Ruslan Panin

Carabus (Apotomopterus) madefactus Cavazzuti, 1997
Endophallus
China, CW-Yunnan, Nánjiàn Yízú Zìzhìxiàn, Mt Lingbao h=2400 m, 24.768611, 100.500000
Copyright © Ruslan Panin

Carabus (Apotomopterus) mecynodes pinchuanensis Imura, 1999
Endophallus
China, S-Sichuan, 40-50 km E Xichang (to Zhaojue) 27°51'N 102°30'E, h=2700-3000m, Pine forest
Copyright © Ruslan Panin

Carabus (Apotomopterus) patroclus apricalis Cavazzuti, 1997
Endophallus
China, S Sichuan, W of Pingchuanzhen, near Galuo vill., h=2400 m
Copyright © Ruslan Panin

Carabus (Apotomopterus) microtatos lingbaonis Imura, 2019
Endophallus
China, CW-Yunnan, Nánjiàn Yízú Zìzhìxiàn, Mt Lingbao h=2400 m, 24.768611, 100.500000
Copyright © Ruslan Panin

Carabus (Apotomopterus) bousquetellus bousquetellus Deuve, 1998
Endophallus
S-China, CW-Yunnan, W of Lincang, Daxue Shan h=2600m, 23.935556, 100.257778
Copyright © Ruslan Panin

Carabus (Apotomopterus) patroclus patroclus Breuning, 1933
Endophallus
China, S Sichuan, Pass 56 km SW Miannin, near Wawo h=2440 m
Copyright © Ruslan Panin

Carabus (Coptolabrus) elysii xiaoganicus Deuve et Remond, 2016
Endophallus
China, Hubei, Xiaogan, Mt Shuangfeng Shan,
Copyright © Ruslan Panin

Carabus (Coptolabrus) lafossei tungchengensis Li, 1993
Endophallus
China, Hubei, Shangzhi co., Shimenyan 8-11.6.2009 (cAA)
Copyright © Alexander Anichtchenko

Carabus (Coptolabrus) augustus ignigena Hauser, 1914
Endophallus

Copyright © Alexander Anichtchenko

Carabus (Cupreocarabus) huangi huangi Deuve, 1992
Endophallus

Copyright © Ruslan Panin

Carabus (Cupreocarabus) balangicus balangicus Cavazzuti, 2002
Endophallus
Chine, C-Sichuan, Xiaojin, Wolong valley: Balang Shan Pass h=3448 m, 30.884617, 102.973117
Copyright © Ruslan Panin

Carabus (Diocarabus) fraterculus fraterculus Reitter, 1895
Endophallus
N-Korea, N Pyongan Prov.,Sakju, Mt Ponsan,
Copyright © Ruslan Panin

Carabus (Eucarabus) sternbergi sternbergi Roeschke, 1898
Endophallus
Scale bar = 5mm
Copyright © Jong-Bong Choi

Carabus (Eucarabus) xiuyanensis Deuve & Li, 1998
Endophallus
NE-China: SE-Liaoning, Xiuyan County, Dayingzi, Mt. Laopingding,
Copyright © Ruslan Panin

Carabus (Eucarabus) billbergi kuandianicus Rapuzzi, 2007
Endophallus
NE China, Liaoning, Kuandian County, Dachuantou env.,
Copyright © Ruslan Panin

Carabus (Hemicarabus) tuberculosus Dejean, 1829
Endophallus
Carabus (Hemicarabus) tuberculosus n. strophium F.-W., 1842 SE Kazakhstan: NW Tarbogatai Mt.R.
Copyright © Igor Solodovnikov

Carabus (Isiocarabus) miaorum Lassalle & Prunier, 1993
Endophallus
China, NW-Guangxi, Mao'er Shan, h~1600 m 25°52'N 110°29'E
Copyright © Ruslan Panin

Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931
Endophallus
NE-China:Liaoning prov.,Fengcheng City, Miaoyang,
Copyright © Ruslan Panin

Carabus (Meganebrius) everesti everesti Andrewes, 1929
Endophallus
Kemba Kharka
Copyright © Joachim Schmidt

Carabus (Morphocarabus) wulffiusi wulffiusi Morawitz, 1862
Endophallus

Copyright © Ruslan Panin

Carabus (Morphocarabus) brinevi Kabak, 2014 Paratype
Endophallus

Copyright © Ilya Kabak

Carabus (Ophiocarabus) nilkiensis Kabak, 2014 Paratype
Endophallus

Copyright © Ilya Kabak

Carabus (Ophiocarabus) wusunshanicus Kabak, 2015
Endophallus

Copyright © Ilya Kabak

Carabus (Ophiocarabus) successor successor Reitter, 1896
Endophallus
SE-Kazakhstan, Narynkol distr., 4 km SW Bayankol, h~2400-2500 m
Copyright © Ruslan Panin

Carabus (Ophiocarabus) ernsti ulastaiensis Kabak, 2019 Paratype
Endophallus
China, Xinjiang, S slopes of Boro-Horo Mt. R., ridge between Kaptshik and Zekku rivers, NE of Ulastai (= Wulasitai) Vill., alpine meadows, 43°56'07"N / 83°18'33"E – 43°56'22"N / 83°18'30"E, 3135–3280 m, 17.07.2017 leg I.I. Kabak
Copyright © Kirill Makarov

Carabus (Ophiocarabus) successor eoketmenicus Kabak, 2016 Holotype Paratype
Endophallus
endophallus - paratypes; aedeagus - holotype
Copyright © Ilya Kabak

Carabus (Ophiocarabus) kurdaiensis Kabak, 2016 Paratype
Endophallus

Copyright © Ilya Kabak

Carabus (Ophiocarabus) ernsti Kabak, 2001
Endophallus
7–8, 12–13 – C. ernsti ulastaiensis, paratypes; 9–10, 14–15 – C. ernsti ernsti Kabak, 2002: 9, 14 – from the locality in mountains to the West of the Saryk River, 10, 15 – from the locality in right tributary of the Saryk River; 11,16 – C. ernsti nilkiensis Kabak, 2014: 11 – holotype, 12 – paratype. ml – median lobe; pp – praeputial pad; rbll – right basolateral lobe.
Copyright © Ilya Kabak

Carabus (?Oreocarabus) qinlingensis Imura, 1993
Endophallus
China, W-Shaanxi, Taibai Shan Mts, btw.Houzhenzi/Ba Xian Tai, nr.Nan Tian Men h=3200m, 33°55'17"N/107°46'53"E,
Copyright © Ruslan Panin

Carabus (Oreocarabus) latro minshanensis Deuve, 1987
Endophallus
China, N-Sichuan, Jiuzhaigou env.,
Copyright © Ruslan Panin

Carabus (?Oreocarabus) titanus Breuning, 1933
Endophallus
C-China Sichuan prov., Nanping 1500m,
Copyright © Ruslan Panin

Carabus (Pseudocoptolabrus) watanabei valdemar Deuve, 2016 Paratype
Endophallus
Chine, Yunnan, Dêqên, monts Hengduan Shan, près du col de Doker La, 3600 m
Copyright © Thierry Deuve

Carabus (Pseudocoptolabrus) belousovianus Imura, 2011
Endophallus
Image from Imura, 2011
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Carabus (Rhigocarabus) ladygini degensis Deuve, 1989
Endophallus
C-China, NW- Sichuan prov., Garzê Tibetan AP, Pass Garzê – Sêrtar
Copyright © Ruslan Panin

Carabus (Rhigocarabus) gigolo Heinz & Brezina, 1996
Endophallus
China, Tewo. det. Anichtchenko, A. 2020
Copyright © Alexander Anichtchenko

Carabus (Rhigocarabus) buddaicus gansuicus Deuve, 1991
Endophallus
N Sichuan, Langmusi vill 3700 m, 1-25.7.2018
Copyright © Alexander Anichtchenko

Carabus (Rhigocarabus) roborowskii langmusiensis Deuve, 1996
Endophallus
N Sichuan, near Langmusi, 3700m 1-25.7.2018, V. Patrikeev leg. det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Carabus (Rhigocarabus) casanova (Imura & Brezina, 2008)
Endophallus

Copyright © Boleslav Brezina

Carabus (Rhigocarabus) gigolo Heinz & Brezina, 1996
Endophallus
Langmusi
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) gentleman Brezina & Hackel, 2004
Endophallus
Jiangjing
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) gigoloides priapeius Cavazzuti, 2006
Endophallus
Yeliguan
Copyright © Boleslav Brezina

Carabus (Rhigocarabus) nianjuaensis Imura, 2016
Endophallus
Image from Imura, 2016
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Carabus (Rhigocarabus) itzingeri rugulosior Deuve, 1996 Holotype
Endophallus
NW-Yunnan: Hengduan Shan: Yanmen env.: 28:06N/98:52E. Image from Imura, 2016
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Carabus (Rhigocarabus) itzingeri tanakai Imura, 1997
Endophallus
Image from Imura, 2016
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Carabus (Rhigocarabus) itzingeri choguy Imura, 2001 Holotype
Endophallus
Image from Imura, 2016
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Carabus (Rhigocarabus) rhododendron xuebangshanensis Imura, 2016 Holotype
Endophallus
Image from Imura 2016
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Carabus (Rhigocarabus) rhododendron xinshanchangensis Imura, 2016 Holotype
Endophallus
Image from Imura, 2016
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Carabus (Rhigocarabus) rhododendron lidimaensis Imura, 2016 Holotype
Endophallus
Image from Imura, 2016
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Carabus (Rhigocarabus) rhododendron s.str. Holotype
Endophallus
Image from Imura, 2016
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Carabus (Rhigocarabus) roborowskii roborowskii Semenov, 1887
Endophallus
China, E-Qinghai, Mts NW of Tongren city, 35.562778, 101.843667, h=3870 m
Copyright © Ruslan Panin

Carabus (Semnocarabus) carbonicolor carbonicolor Morawitz, 1886
Endophallus

Copyright © Alexandr Safronov

Carabus (Semnocarabus) erosus karascharensis Eidam Lectotype Paralectotypes
Endophallus
aedeagus - lectotype. Endophallus - paralectotype.
Copyright © Kirill Makarov

Carabus (Semnocarabus) cicatricosulus pseudoerosus Mandl, 1955
Endophallus
left tributary of Kuqa River, E of Lake Kyrghyz-Artukul (= Dalongchi), 42°27'15''N, 83°22'00''E, H ~ 2550–2600 m, 7–8.VI.2002, I. Kabak leg
Copyright © Kirill Makarov

Carabus (Semnocarabus) erosus erosus Motschulsky, 1865
Endophallus
SE-Kazakhstan, 70 km E of Almaty, E Trans-Ili Alatau N-slopes, valley Assy (Asy) riv. Assy plateau h=1700 m, 12.V.2015
Copyright © Ruslan Panin

Carabus (Sigenacarabus) trachynodes moxiensis Deuve, 1992
Endophallus
China, W-Sichuan, Daxue Range, Mt.Gongga Shan h=3000-3100m, Hailuogou Glacier Forest Park, 29.574889, 101.995556
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus obliteratus Fischer, 1827
Endophallus
endophallus sibiricus obliteratus Fischer, 1827
NE Kazakhstan,
S Altai mts.,S Ulbinsky mt.h=500m.,Turgusun vill., 20.VI.2001
male 25mm.
Copyright © Ruslan Panin

Carabus (Trachycarabus) latreillei ?semicoriaceus Kraatz, 1881
Endophallus
NE-China: Liaodong Peninsula, Liaoning prov., Gaizhou distr., Mt. Buyun Shan.
Copyright © Ruslan Panin

Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
Endophallus
N Mongolia, Khövsgöl reg., Lake Khovsgol, h=1640m, 51*:12’N; 100*:45’E, steppificated sites,
Copyright © Ruslan Panin