Carabus (Archicarabus) pseudomonticola Lapouge, 1908 ? nemoralis
Male adult
NE Spain, Catalonia, Sant Jaume de Frontanyà, h=1000 m, V-VI.2010, Pascal Stefani leg.
L=26 мм
Copyright © Ruslan Panin

Carabus (Archicarabus) pseudomonticola Lapouge, 1908
Male adult
Spain, Montseny, Barcelona
19.VI.1986 M. Tomas leg.
male, 21 mm
Copyright © Alexander Anichtchenko

Carabus (Archicarabus) nemoralis prasinotinctus Heyden, 1880
Male+Female adult
AUDE.Pic de Nore, h=1200m.IX.2005, A. Laforgue leg.
female,25mm/male, 24mm
Copyright © Ruslan Panin

Carabus (Archicarabus) nemoralis prasinotinctus Heyden, 1880
Male adult
N Spain, 3 km N Bera de Bidasoa 14.9.2002
Anichtchenko A. leg.
long. 25mm
Copyright © Alexander Anichtchenko

Carabus (Archicarabus) alysidotus Illiger, 1798
Female adult
Italy, Reggio Emilia, Albinea env. h=150 m, VI.2018, A. Cartini leg., L=25 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) monticola monticola (as maritimensis Born, 1923)
Female adult
SE France, Alpes-Maritimes, Col de Braus h=1000 m, 23.V-18.VI.2014, P. Cavazzuti & G. Proscia leg., L= 24 mm
Copyright © Ruslan Panin

Carabus (Carabus) arvensis sylvaticus Dejean, 1826
Female adult
France, Centre reg., Eure-et-Loir, Forкt de Montecot, VI.2009, Pascal Stefani leg.
female, 20 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) arcensis sylvaticus Dejean, 1826 (as sugintensis Brunier, 1922)
Male adult
NW -France, Haut-Rhin Depart., Fulleren, XI.2004, Pascal Stefani leg.
Copyright © Ruslan Panin

Carabus (Carabus) arvensis sylvaticus Dejean, 1826 (as liebmanni Lengerken, 1911 )
Female adult
France 67, Weifersweller 350m, Vosges massif
25.8.91
long. 21mm
Copyright © Alexander Anichtchenko

Carabus (Carabus) arvensis sylvaticus Dejean, 1826
Male adult
France 61, Longny-au-Perche. 30.V.1991
male, 19 mm
Copyright © Alexander Anichtchenko

Carabus (Carabus) arvensis arvensis Herbst, 1784
Male+Female adult
W Ukraine, NW Podolian, Lviv reg., Javoriv distr., near Stradch vill., 20-30.IV.2009, Panin R. leg.
Copyright © Ruslan Panin

Carabus (Carabus) arvensis arvensis Herbst, 1784
Male adult
W Ukraine, Male Polissya, Lviv reg., near Kamenka-Bugzkaya, under the bark of oak tree, in the forest, 29.III.08, Panin R. leg. male,17mm
Copyright © Ruslan Panin

Carabus (Carabus) arvensis arvensis Herbst, 1784
Male adult
W. Ukraine , NW. Podolian, Lviv reg., Nikolaev distr., near Krasiv vill., 9.II.08, Panin R. leg. L=17mm
Copyright © Ruslan Panin

Carabus (Carabus) granulatus granulatus Linne, 1758
Male adult
W Ukraine,
Lviv Reg., Javoriv distr., near Ivano-Frankovo vill., 30.IV-18.V.2009, Panin R. leg.
male, 21mm
Copyright © Ruslan Panin

Carabus (Carabus) granulatus granulatus Linne, 1758
Male adult
23 mm
Germania, Oberfranken, Bad Steben
leg.det. U.Schmidt, 2006
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Carabus (Carabus) granulatus granulatus Linne, 1758
Female adult
Polonia, Verh. Zalesie, V.1994 Strelnikov D. leg.
long. 21mm
Copyright © Alexander Anichtchenko

Carabus (Carabus) granulatus granulatus Linne, 1758
Female adult
long. 22mm
Copyright © Ruslan Panin

Carabus (Carabus) granulatus granulatus Linne, 1758
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Carabus (Carabus) granulatus granulatus Linne, 1758
Male adult
W-Ukraine, Roztochia,
Lviv Reg., Javorov distr., near Ivano-Frankovo vill., Site Zalyvki, 17.IV-3.V.2010, Panin R. leg.
L=20 mm

Copyright © Ruslan Panin

Carabus (Carabus) arvensis arvensis Herbst, 1784 (as noricus Sokolar, 1910)
Female adult
CZ, C Moravia, Dřevohostice - Líšná, 13mm
Copyright © Aleš Sedláček

Carabus (Carabus) granulatus granulatus (as hibernicus Lindroth, 1956)
Female adult
W-Ireland, County Mayo, Letterneevoge (R312),16-30.VI.2013, J.Stypula leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Carabus) arvensis arvensis Herbst, 1784
Female adult
NW Ukraine, Polesia, Rovno reg., Rokytne distr., near Bil’sk vill., 4-6.VII.2015, Panin R. leg. L=20 mm
Copyright © Ruslan Panin

Carabus (Carabus) granulatus granulatus Linne, 1758
Male adult
ugly specimens
Copyright © Igor Solodovnikov

Carabus (Carabus) granulatus granulatus Linne, 1758
Male adult
Carabus (s.str.) granulatus granulatus ab. wimmeli Schulz, 1900 S-Belarus, Brest reg., env. Tomashovka, pine forest, 26.VI-13.VII.2022, Kuznetzov V.A.
Copyright © Igor Solodovnikov

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761 (as bohemicus)
Female adult
female, 33mm
Copyright © Boleslav Brezina

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Male adult
male, 26mm
Copyright © Boleslav Brezina

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Male adult
W-Ukraine,
Lviv Reg., Zhydachiv distr., near Vybranivka Vill. Panin R.leg. 12.X.2007
31mm.
Copyright © Ruslan Panin

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Female adult
36 mm
Fundort: Germania, Oberfranken, Kulmbach
leg.det. U.Schmidt, 1977
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Male adult
W Ukraine, Lviv city, Sykhiv, Zubra forest, (Osvytskiy Grove), under bark, 49°47'28.10" N, 24° 2'20.09" E, 10.XI.2017, Panin R leg.
Copyright © Ruslan Panin

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Female adult
W Ukraine, Lviv city, Holosko, Bryukhovytsʹkyi forest 49.886833, 24.012083 under bark, 10.XI.2024, R. Panin leg., L=35 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860
Male+Female adult
SW -France,
Pyrenees-Atlantiques, Foret de Sare, XII.2009, Pascal Stefani leg.
female, 31 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860
Female adult
Spain,
Navarra, Sierra de Aralar, Altxueta Mt., 5.V.2006
Anichtchenko A. leg.
L= 17mm
Copyright © Alexander Anichtchenko

Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860
Male adult
Spain,
Pais Vasco, Bera de Bidasoa, Larrun 450m,
9.9.2001 Anichtchenko A. leg.
male, 29mm
Copyright © Alexander Anichtchenko

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as delacrei Tarrier, 1975)
Male+Female adult
NE Spain, Catalonia, Sant Jaume de Frontanyа, h=1000 m, V-VI.2010, Pascal Stefani leg.
female, 38 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as delacrei Tarrier, 1975)
Male adult
Spain,
Catalunya, San Jaume de Frontana,
May 2001
Copyright © Ruslan Panin

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826
Female adult
Auriac alt.
500m. (France : Aude) 25.V.200
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792
Female adult
NE-FRANCE,
Lorraine Reg., Moselle, 15.II.2010, Pascal Stefani leg.
female, 26 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792
Male adult
Austria,
Weissenbach stt., III.1979, Kretschmer leg.
male, 24mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792 (as aureus)
Male adult
France 28, Saint-Eliph, V.1991
female, 25mm
Copyright © Alexander Anichtchenko

Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792
Female adult
France 28, Saint-Eliph, V.1991
female, 24 mm

Copyright © Alexander Anichtchenko

Carabus (Chrysocarabus) auronitens costellatus Gehin, 1882 (as subfestivus Oberthur, 1884)
Female adult
W- FRANCE,
Bretagne Reg., Finistrere, Foret de Coatloc'h, XI.2006
female, 26 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens costellatus Gehin, 1882
Male+Female adult
C-FRANCE,
Limousin Reg., Corrиze Dep., Aubazines, X.2005, Pascal Stefani leg.
female,26mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens costellatus Gehin, 1882
Male adult
France 42, Mount de Madeleine, 1300m, les Biefs
18.VIII.1991
male, 19mm
Copyright © Alexander Anichtchenko

Carabus (Chrysocarabus) auronitens cupreonitens Chevrolat, 1861
Female adult
NW-FRANCE,
Basse-Normandie Reg., Orne, Forêt de Bellême, 14.XII.2008, Pascal Stefani leg.
female, 26mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens cupreonitens Chevrolat, 1861
Male+Female adult
NW-FRANCE,
Basse-Normandie Reg,, Orne, Foret d' Ecouves, V.2009, Pascal Stefani leg.
male, 25 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens farinesi Dejean, 1826 (as barthei Lapouge, 1912)
Male+Female adult
SC -France,
Midi-Pyrйnйes reg., Ariиge, Foret de Belesta, XI.2008, J.Nicolas leg.
L=31 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens festivus Dejean, 1826
Male adult
male, 22mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) auronitens montanus Gehin, 1882
Male+Female adult
SC-FRANCE,
Haute-Garonne, Col de Mentй h~1300m., II.1980. Kretschmer leg.
female, 23mm.
photo/det. Panin R.
Copyright © Ruslan Panin

Carabus (Chrysocarabus) solieri clairi Gehin, 1885
Male adult
France, Caussols, VII.1984
male, 26 mm
Copyright © Alexander Anichtchenko

Carabus (Chrysocarabus) solieri liguranus Breuning, 1933
Female adult
NW Italy,
Altare, Savona,15.VI.1995, Cartini A.leg.
female, 33mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) solieri solieri Dejean, 1826
Male adult
SE-France: Alpes-Maritimes
Copyright © Safronov

Carabus (Chrysocarabus) splendens Olivier, 1790
Male+Female adult
SE France,
Midi-Pyrйnйes reg., Ariиge, Vaychis, 22.X.2002, A.Laforgue leg.
female, 30mm.
Copyright © Ruslan Panin

Carabus (Chrysocarabus) splendens Olivier, 1790
Male adult
Spain, Pais Vasco, Bera de Bidasoa, 15.9.2001
Anichtchenko A. leg.
male, 30mm
Copyright © Alexander Anichtchenko

Carabus (Chrysocarabus) hispanus Fabricius, 1787
Male adult
FRANCE,
Tarn, Foret de la Gresigne, h=400m.VII.2005 Laforgue leg.
male-30mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) hispanus Fabricius, 1787
Male adult
Auriac alt. 500m.
(France : Aude) 25.V.2000
Copyright © Pascal Stefani

Carabus (Chrysocarabus) solieri solieri Dejean, 1826
Female adult
SE France, Alpes-Maritimes, Ascros, VI.2011
L=30 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) lineatus troberti Kraatz, 1860 (as ursuius Lapouge, 1911)
Male+Female adult
FRANCE (64), Ursuya Mt.
XII-2009, P. Stefani leg.
male - 30 mm, female - 32 mm.
Copyright © Alexandr Safronov

Carabus (Chrysocarabus) auronitens festivus Dejean, 1826 (ab. holochrysum)
Female adult
SC-FRANCE, Tarn, Le Linas h=550 m, XI.2008, Pascal Stefani leg. L=24 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as aragonensis Lapouge,1913)
Male adult
Male adult ,Sierra de Montseny (Barcelone) Spain.
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as frontanyaensis Mollard,1999)
Female adult
N of Boreda, (Lerida) Spain.
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as brevicollis Lapouge,1910)
Female adult
Corbières (Aude) France.
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as brevicollis var. viridis)
Female adult
Rennes les bains (Aude) France.
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as delacrei Tarrier,1975)
Female adult
Nostra Signora de Queralt, Berga (Barcelona) Spain.
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as occideneissimus Meyer,1995)
Male adult
Collado de Boixols, (Lérida) Spain.
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as urgelensis Mollard,1999)
Female adult
Sierra del Cadi,Toloriu, (Lérida) Spain.
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as jeannei)
Female adult
Sierra de la Mussara (Tarragona) Spain.
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as merangensis Mollard,1999)
Male adult
Méranges, near Puigcerda, (Catalunya) Spain
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as monserraticus Csiki,1917)
Female adult
Sierra de Montserrat, (Barcelone) Spain
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as palensis Mollard,1999)
Male adult
Col de N’hug, (N-Catalunya) Spain.
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as quillansis Mollard,1999)
Male adult
Granès (Aude) France.
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (f. typ.)
Female adult
Prats de Mollo (Pyr.Or) France
Copyright © Alain Mollard

Carabus (Chrysocarabus) rutilans rutilans Dejean, 1826 (as soredensis Mollard,1999)
Male adult
Ft.de Sorède, Mts. Albères (Pyr.Or.) France
Copyright © Alain Mollard

Carabus (Chrysocarabus) solieri liguranus Breuning, 1933
Male adult

Copyright © Igor Solodovnikov

Carabus (Chrysocarabus) auronitens auronitens Fabricius, 1792
Male adult
C-Poland, Łódź sity, Łagiewniki Forest, in dead logs of Birch, 18-30.XII.2013, D. Nawak leg., L=25 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) solieri bonnetianus Colas, 1936 (loc.typ.)
Female adult
SE-France, Var, Tanneron, Saint-Cassien des Bois, 18.IV.2019, L=32 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) solieri bonnetianus Colas, 1936 (loc.typ.)
Male adult
SE-France, Var, Tanneron, Saint-Cassien des Bois, 18.IV.2019, L= 31 mm
Copyright © Ruslan Panin

Carabus (Chrysocarabus) solieri clairi Gehin, 1885 (as devillei Tarrier, 1972)
Male adult
SE-France, Alpes-Maritimes, Beuil env., h=1550m, 14.VII.1986
Copyright © Ruslan Panin

Carabus (Eurycarabus) genei Gene, 1839
Male adult
Corse de Sud, env. de
Rondinara, VI.1998 Tingaud M. leg.
male, 23 mm

Copyright © Alexander Anichtchenko

Carabus (Hemicarabus) nitens Linne, 1758
Male adult
male, 15mm
Copyright © Boleslav Brezina

Carabus (Hemicarabus) nitens Linne, 1758
Male+Female adult
Belarus,
Vitebsk reg., Ushachskiy distr., near Bolshie Dolcy vill. 9.V.1990, I.A.Solodovnikov leg.
female, 18mm.
Copyright © Ruslan Panin

Carabus (Hemicarabus) nitens Linne, 1758
Male adult
Russia, Ural, riv. Rezh, vill. Kamenka; 10.VII.2009, N. Menstschicova leg.
male, 14 mm.,
Copyright © Anton Menstschikov

Carabus (Hemicarabus) nitens Linne, 1758
Male adult
16,6 mm, England, Dorset, Studland
leg.det. J.W.Yerbury, 18.-29.VIII.2006
Coll. Oxford University Museum of Natural History
Photo: U.Schmidt, 2011
Copyright © Udo Schmidt

Carabus (Hemicarabus) nitens Linne, 1758
Female adult

Copyright © Kirill Makarov

Carabus (Hemicarabus) nitens Linne, 1758
Male adult
Belarus, Vitebsk reg., Miory distr. Swamp «Mokh», 09.05.2011, leg. G.G. Sushko
Copyright © Igor Solodovnikov

Carabus (Hemicarabus) nitens Linne, 1758
Male adult

Copyright © Igor Solodovnikov

Carabus (Iniopachus) pyrenaeus jeanneli Deuve, 1994
Male adult
1-2: Pirineos, Canfranc, Ibon de Iserias 2200 m 16-17.7.05<br /> 3: Pirineos, Monte Perdido, Collata Anisolo 2400m 25.8.05
Copyright © Alexander Anichtchenko

Carabus (Iniopachus) pyrenaeus pyrenaeus (as cephalicus Csiki, 1927)
Male adult
SW France, Pyrenees-Atlantiques, Mt.Cirque d'Esturou h=1800m, 29.VIII.2014, Pascal Stefani leg. L=20 mm
Copyright © Ruslan Panin

Carabus (Iniopachus) pyrenaeus costulus Gehin, 1885
Male+Female adult
Mt. Canigou, 2400m (Pyrénées Orientales, Pyrénées) FRANCE, VI-1985
Copyright © Alain Mollard

Carabus (Iniopachus) pyrenaeus vallierensis Colas, 1965
Male+Female adult
Mt.Vallier (Ariege,Pyrénées) FRANCE, VI-1998
Copyright © Alain Mollard

Carabus (Limnocarabus) clathratus clathratus Linne, 1761
Female adult
W-Ukraine, Maloe Polesie, Lviv Region, near Kamenka-Bugzkaya, 28.III.2008, Panin R. leg. L=31 mm
Copyright © Ruslan Panin

Carabus (Limnocarabus) clathratus clathratus Linne, 1761
Male adult
length: 20-36 mm, Germania, Mecklenburg, Wismar, leg. F.Wolf, 1998; det. A.Skale, Photo: U.Schmidt, 2009
Copyright © U.Schmidt

Carabus (Limnocarabus) clathratus clathratus (as clathratus foveolatoseriatus)
Female adult
The Chelyabinsk area, Kizilskyi township, Cheka mountain (steppe). 14-15/07/2008
Copyright © Sunvas

Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
Male adult
Transcaucasia, Armenia,
Amasia distr., Arpalach vill., VI-VII.1999, P.V.Kazaryan leg.
male, 25mm
Copyright © Ruslan Panin

Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
Female adult
SW Slovakia, Bratislava env., IV.2004, Krause Rene leg.
L=32 mm
Copyright © Ruslan Panin

Carabus (Limnocarabus) clathratus clathratus (as jansoni Kraatz, 1890)
Male adult
W-Ireland, County Mayo, Bellacorick bog, 9.VII.2013, J.Stypula leg. L=27 mm
Copyright © Ruslan Panin

Carabus (Limnocarabus) clathratus auraniensis Muller, 1902 (as antonellii Luigioni, 1921)
Male adult
Gaggiano Milan, Italy
Copyright © Vittorino Monzini

Carabus (Limnocarabus) clathratus clathratus Linne, 1761
Female adult

Copyright © Kirill Makarov

Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
Male adult
W Ukraine, Transcarpathian Reg., 4 km NE Chop, forest Peresh, Panin R. leg. L=30 mm
Copyright © Ruslan Panin

Carabus (Limnocarabus) clathratus arelatensis Lapouge, 1903
Female adult
Aigues-Tortes, Camargue, France VI-1984
Copyright © Alain Mollard

Carabus (Macrothorax) morbillosus constantinus Kraatz, 1899
Female adult

Copyright © Safronov

Carabus (Macrothorax) morbillosus constantinus Kraatz, 1899
Male adult
NW Tunisia; Tebarka, E Sejenane,26.I.2004, Manfred Egger leg.
L=33 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) morbillosus constantinus (as galloprovincialis Lapouge, 1910)
Female adult
SE-France, Var, near Le Muy, VIII.1980, Michon leg.
L=31 mm
Copyright © Ruslan Panin

Carabus (Macrothorax) morbillosus morbillosus Fabricius, 1792 (as cheminorum Deuve, 1988)
Female adult
SE-France, Var, Fréjus env., 26.II.1997,
L=31 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus fulgens Charpentier, 1825
Male adult
NC-Spain,
Navarra Reg., Sierra de Andнa, IX.2004, Pascal Stefani leg.
male, 30 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus fulgens Charpentier, 1825 (as gerundensis Born, 1916)
Male adult
Spain, Gerona, Garriguella, XI.1990 J. Soler leg.
male, 31mm
Copyright © Alexander Anichtchenko

Carabus (Megodontus) violaceus mixtus Gehin, 1876
Female adult
France, Eloise 74, pont de Gresin 14.07.1991
female, 29mm
Copyright © Alexander Anichtchenko

Carabus (Megodontus) violaceus fiorii Born, 1901
Male+Female adult
NW-Italy,
Reggio Emilia, Regnano, 5.IX.1996 leg. Cartini A.
female, 30 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus baeterrensis Lapouge, 1901
Male+Female adult
S France,
Midi-Pyrénées Region, Ariège, Col du Chioula h=1500m, V-VI.2005, Pascal Stefani leg.
photo Panin R.
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus purpurascens Fabricius, 1787 (as osismius Lapouge, 1922)
Male adult
W-FRANCE, Brittany , Finistère, Botvarec Forest, 19.XI.2006, Pascal Stefani leg.
L=29 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus mixtus (as neomixtus Tarrier, 1975)
Male adult
L=29mm SE-France, Drôme, Foret de Saou, VII.2014, Pascal Stefani leg.
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus violaceus trans violaceus purpurascens Fabricius,1787
Male+Female adult
E-Netherlands, Gelderland, Winterswijk Woold env.,15.II.2015, H.de Haan leg.
Copyright © Ruslan Panin

Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
Male adult
SW-France,
Foret d'Iraty Pyrenees-Atlantiques, d'Occabe, 15.VIII.1988
male,30 mm
photo Panin R.
Copyright © Ruslan Panin

Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876 Loc.typ.
Female adult
Spain, Navarra, Sierra de Aralar,
Altxueta 1200-1300m 6.1.2001
Copyright © Alexander Anichtchenko

Carabus (Mesocarabus) problematicus harcyniae Sturm, 1815 (as obenbergerianus Breuning, 1927)
Male adult
Czech, Bohemia, Plzeň Reg., Domažlice Distr., Babylon, h=500 m, 30.VIII.2008, S.I. Budka leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus inflatus Kraatz, 1878
Male adult
SE–France, Var departament, Foret royale -Saint-Paul-en-Foret vill.,18.VI.2002, Thelot J.P. leg.
L=26 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus planiusculus Haury in Gehin, 1885
Male adult
NE Spain, Catalonia, Sant Jaume de Frontanyà, h=1000 m, V-VI.2010, Pascal Stefani leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus harcyniae (as procedens Csiki, 1927)
Male adult
England, North Yorkshire, Fylingdales Moor, Grouse Hill, VII.2006 L=24 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus harcyniae (as procedens Csiki, 1927)
Male adult
W-Ireland, County Mayo, Clare Isl., 17-28.VI.2002, S. McCormack leg.<br /> L=25 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus harcyniae (as procedens Csiki, 1927)
Female adult
N-Ireland, Fermanagh Distr., Lough Navar Forest, 17.VII.2013, D.Novak & J.Stypula leg.<br /> L=27mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus harcyniae (as belgicus Lapouge, 1913)
Male adult
Luxembourg, Bollendorf-Pont env., 3.III.1999 /det.R Panin
L=24 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) (? problematicus trans lusitanicus) planiusculus (as xaxarsi Born, 1917)
Male adult
NE Spain, Catalonia, Girona Prov., San Hilario Sacalm env., 20.II.1988, A. Mollard leg.<br /> L=25 m
Copyright © Ruslan Panin

Carabus (Mesocarabus) problematicus inflatus (as solidus)
Female adult
Ripoll, Barcelone, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) problematicus planiusculus (as xaxarsi)
Male adult
Sant Sadurni, Gerone, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) problematicus planiusculus Haury in Gehin, 1885
Male adult
La Seo de Urgel, Gerone, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) problematicus planiusculus (as bepmalei Lapouge, 1904)
Female adult
Pic de Bassibes, Huesca, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) problematicus planiusculus (as navarrensis Breuning, 1933)
Male adult
Mt.Urculo, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) problematicus planiusculus (as trapeti Bleuse, 1885)
Female adult
Panticosa, Huesca, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) problematicus planiusculus (as lagardei Mollard, 1996) Paratype
Female adult
Sierra de Andia, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) problematicus planiusculus (as andorranus Lapouge, 1902)
Female adult
Col d'envalira, Pas de la Case, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
Female adult
Sierra de Aralar, Navarra, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) cantabricus brabeus (as barcelecoanus)
Male adult
Col de Velate, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) cantabricus brabeus (as barcelecoanus)
Female adult
Col de Velate, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
Male adult
Col de Velate, Navarre, Spain. det. A. Mollard; Copyright © Magellanes
Copyright © - - -

Carabus (Mesocarabus) problematicus problematicus Herbst, 1786
Female adult
W-Hungary, NE-Alps, Mts. Kőszeg, Velem env., Borha-forrás, 26.VII.1981, Gy. Somorijai leg. L=26 mm
Copyright © Ruslan Panin

Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876
Male adult
NC-Spain, Basque Co, Álava, Sierra de Gorbea, VI.2014, L=29 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) monilis Fabricius, 1792
Male adult
France 39, Foncina le Haut 300m, 23.6.1979; 29 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) monilis Fabricius, 1792 (as taunicus)
Male adult
W Germany, Vogelsberg Mt., 700m, 13.5.1985
male, 24 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) monilis Fabricius, 1792 (as interpositus Géhin, 1880)
Male adult
NW France,
Basse-Normandie, Orne, Forкt d'Йcouves, V.2009, Pascal Stefani leg.
male, 29mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) monilis (as interpositus Gehin, 1880)
Male+Female adult
NW France,
Haute-Normandie reg., Bois Les Loges, Etretat, Seine-Maritime,10.V.1988
female, 28mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) monilis (as amoenus Baudet-Lafarge, 1836)
Female adult
C France,
Puy-de-Dome, Col de la Croix Saint Robert h=1300m, VII.2008, Pascal Stefani leg.
female, 24 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) monilis (as helveticus Heer, 1837 Holotype) Holotype
Male adult
Carabus (Morphocarabus) monilis (as helveticus Heer, 1837 Holotype)
?hybrid of monilis/cancellatus
Museum of Zurich
photo Anichtchenko
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) monilis (as dupeuxi Deuve, 1974)
Male+Female adult

France,
Charente, Foret de Braconne VII.1998, Laforgue leg.
famale, 27mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) monilis Fabricius, 1792
Male adult
24 mm, England, Warwickshire
leg.det. D.J.Mann, 15.IV.1993
Photo: U.Schmidt, 2011
Copyright © Udo Schmidt

Carabus (Morphocarabus) monilis Fabricius, 1792
Female adult
France, Dröme, Ft. de Saou, 17.vii.1975-M. Grotz Leg.-Coll.RBINS, N. Mal as monilis saouensis nicodi
Copyright © Noël Mal

Carabus (Morphocarabus) monilis tanarguensis Maguerre & Stefani, 2021 (syn.) Holotype Paratype
Male+Female adult
Holotype, m (21 mm), environs du Col de la Croix de Bauzon, 1300 m, communes de La Souche, Borne et Mayres, Ardèche, France (07/2003), in coll. D. Maguerre, à Mons-en-Barœul, France. Paratype f (23 mm), au dessus du Col de Meyrand, 1400 mètres, communes de Loubaresse et de Borne, Ardèche, France, (07/2004), in coll. P. Stéfani.
Copyright © - - -

Carabus (Pachystus) hortensis hortensis Linne, 1758
Male adult
Germania, Oberfranken, Langenau
leg.det. U.Schmidt, 2002
Foto: U.Schmidt, 2006
Copyright © U.Schmidt

Carabus (Pachystus) hortensis hortensis Linne, 1758
Female adult
Natural hybrid Carabus hortensis X Carabus granulatus. Belarus, Minsk reg., 8,7 km of Borisov, near vill. Lyubatovstchina, pine forest (Pinetum pleuroziosum), h = 213 m, 54.31934° N, 28.511486° E, 20-27.09.2018, leg. Klychenok Snezhana
Copyright © Igor Solodovnikov

Carabus (Oreocarabus) hortensis hortensis Linne, 1758
Male adult

Copyright © Kirill Makarov

Carabus (Orinocarabus) fairmairei Thomson, 1875
Female adult
SE France,
Grand Belvédère du refuge du Mont Viso, 5-20.VII.1991
female, 20mm.
photo Panin R.
Copyright © Ruslan Panin

Carabus (Orinocarabus) pedemontanus maurinensis Deuve, 2002 Holotype
Male adult

Copyright © Frank Kleinfeld

Carabus (Orinocarabus) sylvestris sylvestris Panzer, 1793 (as slabai Niedl, 1968)
Female adult
Czech, WS Bohemia,
Plzen Reg., Domazlice Distr., Babylon, h=500m, 30.VIII.2008, S.I. Budka leg.
L=24mm
Copyright © Ruslan Panin

Carabus (Orinocarabus) fairmairei stecki Born, 1902
Female adult
France, Hautes-Alpes, env. Lake Foreant, near Col de D'Agnel, ca 2700-2800 m 17.VIII.2013 leg. B.Kataev L=18.0 mm
Copyright © Kirill Makarov

Carabus (Orinocarabus) fairmairei stecki Born, 1902
Male adult
France, Hautes-Alpes, env. Lake Foreant, near Col de D'Agnel, ca 2700-2800 m 17.VIII.2013 leg. B.Kataev L=16.5 mm
Copyright © Kirill Makarov

Carabus (Orinocarabus) pedemontanus raynaudianus Deuve & Simard, 1977
Male adult
SE France, Alpes-Maritimes, Isola env., Colle Santa-Anna h=2309 m, 44*13',127" N, 07*06',242"E, 31-7.VIII..2012. L=21 mm
Copyright © Ruslan Panin

Carabus (Pachystus) glabratus glabratus Paykull, 1790
Female adult
W UKRAINE<br /> Lviv region, Stryi distr., near Bilche vill., 9-20.VI.2005, Panin R. leg.<br /> female 30 mm
Copyright © Ruslan Panin

Carabus (Pachystus) glabratus glabratus Paykull, 1790
Male adult
W Ukraine, NW Podolian, Lviv reg., Zolochiv distr., near Stinka vill., Lysa Gora, 26.IV.2009 R.Panin leg., 29mm
Copyright © Ruslan Panin

Carabus (Pachystus) glabratus glabratus Paykull, 1790
Female adult
Germania, Oberfranken, Kulmbach leg.det. U.Schmidt, 1974 Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Carabus (Pachystus) glabratus glabratus Paykull, 1790
Male adult

Copyright © Kirill Makarov

Carabus (Platycarabus) depressus lucens Schaum, 1856
Female adult
Italia, Alpi Cozie, Col. delle Finestre,
2150m, VII.1986 Ratti leg.
female, 24 mm
Copyright © Alexander Anichtchenko

Carabus (Platycarabus) depressus depressus Bonelli, 1811
Female adult
Italia, Bellamonte, 2200m
VI.1992 Rubizzani leg.
female, 20 mm
Copyright © Alexander Anichtchenko

Carabus (Platycarabus) irregularis irregularis Fabricius, 1792
Male adult
NE-Italy,
Friuli Reg., Udine, Carnia, Pierabech h~1100m, I.Rapuzzi leg.
male, 23 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) depressus lucens Schaum, 1856
Female adult
SE-France: Alpi Marittime, Monte Viso, 14.VII.2010
L=23 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) irregularis irregularis (as jurassicus Born, 1911)
Male+Female adult
France, Ain (01), Jura Mts, Grand Colombier, Forêt d' Arvières h~1250 m, 25.X.2014, Ya. Vasseur leg., L=23 mm
Copyright © Ruslan Panin

Carabus (Procrustes) coriaceus coriaceus Linne, 1758
Female adult
The neighborhood Nicolo-Berezovskoe republic of
Bashkortostan 5.07
Copyright © Sunvas

Carabus (Procrustes) coriaceus coriaceus Linne, 1758
Male adult
length: 32.42 mm
Germania, Oberfranken, Weismain
leg.det. U.Schmidt, 1983
Foto: U.Schmidt, 2005
Copyright © U.Schmidt

Carabus (Procrustes) coriaceus coriaceus Linne, 1758
Female adult
Belgique : Prov. Luxembourg, Harnoncourt 31.vii.1975 - M. Grotz Leg. (Size : 36 mm)
Copyright © Noël Mal

Carabus (Tachypus) cristoforii Spence, 1823 (as nicolasi Reitter, 1888)
Male adult
S-France, Hautes-Pyrenees, Pic de Nere h=2200m,15.VII.1999,
male, 14mm
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus fusus Palliardi, 1825
Male+Female adult
France, Riveres Charente, leg. Laforgue VI.1996
female, 24mm
Copyright © Ruslan Panin

Carabus (Tachypus) auratus auratus Linne, 1761
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Carabus (Tachypus) auratus s.str. (as brunieri Barthe, 1921)
Male adult
SC-FRANCE, Aude, Rennes les Bains, 6.VII.1989, Sunard A. leg.
male, 26mm
Copyright © Ruslan Panin

Carabus (Tachypus) auratus auratus Linne, 1761
Female adult
30 mm
Germania, Oberfranken, Pegnitz
leg.det. U.Schmidt, 1972
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Carabus (Tachypus) auratus lasserrei Doue, 1855
Male adult
SC-FRANCE, Midi-Pyrуnуes reg. Tarn, Saint Paul cap de Joux, 24.V.2003, A. Laforgue leg.
male, 24mm
Copyright © Ruslan Panin

Carabus (Tachypus) auratus lotharingus Dejean, 1826
Male adult
SC France: Herault departament, Foret de Saint Gervais, V. 2009 leg. J. Nicolas
L=27mm
Copyright © Ruslan Panin

Carabus (Tachypus) cristoforii Spence, 1823
Female adult
S France, Pyrénés Mts., Tarascon; 15mm
Copyright © Aleš Sedláček

Carabus (Tachypus) auratus honnoratii Dejean, 1826
Male adult
SE France, Carpentras; 23mm
Copyright © Aleš Sedláček

Carabus (Tachypus) vagans Olivier, 1795
Female adult
SE-France, Var: Le Muy env., 30.VII.2008., A. Paulian leg.
L=24 mm
Copyright © Ruslan Panin

Carabus (Tachypus) auratus honnoratii Dejean, 1826
Male+Female adult
France : Vaucluse, Mt.Ventoux-Mt. Serein, vii.1969, N. Mal Leg., Coll.RBINS-N. Mal incl. ventouxensis and Nicolasi
Copyright © Noël Mal

Carabus (Tachypus) auratus honnoratii Dejean, 1826
Male+Female adult
France : Vaucluse, Mt. Ventoux-Mt. Serein, 1400-1800m, vii.1969-N. Mal Leg.-Coll.RBINS, N. Mal - including f.i. ventouxensis and nicolasi
Copyright © Noël Mal

Carabus (Tomocarabus) convexus s.str. (as pyrenaicus Born, 1907)
Male adult
Pirineos, Canfranc, 1300m, Barranco de Izas
15.6.2005 Anichtchenko A. leg.
male, 16 mm
Copyright © Alexander Anichtchenko

Carabus (Tomocarabus) convexus s.str. (as paganetti Born, 1905)
Male adult
Lazio, Roma, Bosco di Nettuno
22.5.1986 C. Iacovone leg.
male, 20 mm
Copyright © Alexander Anichtchenko

Carabus (Tomocarabus) convexus convexus Fabricius, 1775
Male adult
W.Ukraine,
Transcarpathian region,Vinogradov distr., near Malaya Kopanya vill., Chorna Gora Mt., h=180m. 7-18.VII.08. Panin R. leg.
male 18 mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) convexus s.str. (ab. niger)
Male adult
W Ukraine, NW Podolia, Lviv Region, Zolochiv distr., near Gologirky vill., in steppe 1-19.V.2007 Panin R. leg.
16mm
Copyright © Ruslan Panin