Carabus (Archicarabus) montivagus montivagus Palliardi, 1825
Endophallus

Copyright © Ruslan Panin

Carabus (Archicarabus) montivagus montivagus Palliardi, 1825
Male+Female adult
SW-Romania: Banat, Caras-Severin Reg., Sasca Montana – Stinapari, 21.V.-5.VI.2010, Krause Rene leg.<br /> female, 26 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) montivagus blandus Frivaldsky, 1865
Female adult
Slovacchia, Plastovce, 16.7.1995 Budka leg. 21 mm
Copyright © Alexander Anichtchenko

Carabus (Archicarabus) montivagus blandus Frivaldsky, 1865
Female adult
female, 23 mm<br /> S-Slovakia, Region de Nitra, Plastovce viil., 1.V.2009, Rudolf Kmeco leg.
Copyright © Ruslan Panin

Carabus (Archicarabus) montivagus vellepiticus Hampe, 1850
Female adult
W Croatia, NP Northern Velebit, Stolak, 44.969492, 14.986169, 23.V-27.VI.2022, L=24 mm
Copyright © Ruslan Panin

Carabus (Archicarabus) montivagus vellepiticus Hampe, 1850
Male adult
W Croatia, NP Northern Velebit, Stolak, 44.969492, 14.986169, 23.V-27.VI.2022, L=23.5 mm
Copyright © Ruslan Panin

Carabus (Carabus) granulatus Linne, 1758
Larva
Carabus (Carabus) granulatus granulatus Linne, 1758
Larva of 2nd instar
L = 16,2 mm
Moskov Reg., Istra Distr., env. Pavlovskaya Sloboda 12.VI.2008 leg. K.Makarov
Copyright © Kirill Makarov

Carabus (Carabus) arvensis arvensis Herbst, 1784
Male+Female adult
W Ukraine, NW Podolian, Lviv reg., Javoriv distr., near Stradch vill., 20-30.IV.2009, Panin R. leg.
Copyright © Ruslan Panin

Carabus (Carabus) arvensis arvensis Herbst, 1784
Male adult
W Ukraine, Male Polissya, Lviv reg., near Kamenka-Bugzkaya, under the bark of oak tree, in the forest, 29.III.08, Panin R. leg. male,17mm
Copyright © Ruslan Panin

Carabus (Carabus) arvensis arvensis Herbst, 1784
Male adult
W. Ukraine , NW. Podolian, Lviv reg., Nikolaev distr., near Krasiv vill., 9.II.08, Panin R. leg. L=17mm
Copyright © Ruslan Panin

Carabus (Carabus) arvensis arvensis Herbst, 1784
Endophallus
W Ukraine, NW Podolian, Lviv reg., Javoriv distr.,near Stradch vill.,
Copyright © Ruslan Panin

Carabus (Carabus) arvensis arvensis Herbst, 1784 (as noricus Sokolar, 1910)
Female adult
CZ, C Moravia, Dřevohostice - Líšná, 13mm
Copyright © Aleš Sedláček

Carabus (Carabus) arvensis arvensis Herbst, 1784
Female adult
NW Ukraine, Polesia, Rovno reg., Rokytne distr., near Bil’sk vill., 4-6.VII.2015, Panin R. leg. L=20 mm
Copyright © Ruslan Panin

Carabus (Carabus) arcensis arcensis Herbst, 1784
Female adult
Lower Austria, Baden, Klausen-Leopoldsdorf, 15.XII.2024, M. Egger leg. L=22 mm
Copyright © Ruslan Panin

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761 (as bohemicus)
Female adult
female, 33mm
Copyright © Boleslav Brezina

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Male adult
male, 26mm
Copyright © Boleslav Brezina

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Male adult
W-Ukraine,
Lviv Reg., Zhydachiv distr., near Vybranivka Vill. Panin R.leg. 12.X.2007
31mm.
Copyright © Ruslan Panin

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Female adult
36 mm
Fundort: Germania, Oberfranken, Kulmbach
leg.det. U.Schmidt, 1977
Foto: U.Schmidt, 2008
Copyright © U.Schmidt

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Male adult
W Ukraine, Lviv city, Sykhiv, Zubra forest, (Osvytskiy Grove), under bark, 49°47'28.10" N, 24° 2'20.09" E, 10.XI.2017, Panin R leg.
Copyright © Ruslan Panin

Carabus (Chaetocarabus) intricatus intricatus Linne, 1761
Female adult
W Ukraine, Lviv city, Holosko, Bryukhovytsʹkyi forest 49.886833, 24.012083 under bark, 10.XI.2024, R. Panin leg., L=35 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) catenulatus catenulatus Scopoli, 1763
Male adult
Lombardy - North Italy
Copyright © Monzini

Carabus (Eucarabus) catenulatus catenulatus Scopoli, 1763
Female adult
NW Croatia, Bosanci, 1-12,VI.2004, Baros Slovak leg.<br /> L=33 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) catenulatus catenulatus Scopoli, 1763
Male adult
Italia, Riva San Vitale (Chleti), VI.1996 Rubizzani leg., 29 mm
Copyright © Alexander Anichtchenko

Carabus (Eucarabus) ulrichi rhilensis Kraatz, 1876 (as sokolari Born, 1904)
Male+Female adult
Czech Rep., W Bohemia,
Plzen reg., Tachov Distr., Plana u Marianskych Lazni, 17-29.V.2008, S.I.Budka leg.
female, 26 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) ulrichi rhilensis (as stussineri Géhin, 1885)
Male adult
W Croatia, Ivanščica Mts , Ivanec, 24.VI-7.VII.2006, Krause Rene leg.
L=29 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) catenulatus fontanellae Reitter, 1896
Male adult
W-Croatia, Cres island,VIII.1994, G. Zannna leg. L=33 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) parreyssi parreyssi Palliardi, 1825
Male adult
W-Croatia, NP Northern Velebit, Zavižan,15.V-13.VI.1996, Colombetta leg., L=21 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) parreyssi parreyssi Palliardi, 1825
Female adult
SW-Croatia, Mts Velebit, Mali Halan, h=1390 m, 23.V-20.VI.2022, L=24 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) parreyssi parreyssi Palliardi, 1825
Endophallus
SW-Croatia, Mts Velebit, Mali Halan, h=1390 m,
Copyright © Ruslan Panin

Carabus (Eucarabus) parreyssi parreyssi Palliardi, 1825
Male adult
SW-Croatia, Mts Velebit, Mali Halan, h=1390 m, 23.V-20.VI.2022, L=22.5 mm
Copyright © Ruslan Panin

Carabus (Eucarabus) catenulatus fontanellae Reitter, 1896
Male adult
W-Croatia, Dalmatia, Isl. Rab, 16-30.V.2019, G. Proscia leg.
Copyright © Ruslan Panin

Carabus (Eucarabus) ulrichi rhilensis (as stussineri Géhin, 1885)
Female adult
S Austria, Styria, Leutschach, Schmirnberg, 46.634400, 15.502400, 1-25.VI.2021, M.Egger leg., L=30 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus rilvensis Kolbe, 1887
Female adult
Bulgaria, Stara Zagora distr. Stara Planina Mts., Shipka Mt., 16.08.2005, A. Safronov leg.
Copyright © Safronov

Carabus (Megodontus) croaticus croaticus Dejean, 1826
Male adult
YU- Slovenija, Krim 1000 m, 30.10.1975 Drovenik leg.
Copyright © Alexander Anichtchenko

Carabus (Megodontus) violaceus germarii Sturm, 1815
Female adult
Croatia, Ivanščica Mts , Ivanec, 6-13.VI.2003, Krause Rene leg.
L=36 mm
Copyright © Ruslan Panin

Carabus (Megodontus) croaticus croaticus Dejean, 1826
Male adult
SW Slovenia, Ilirska Bistrica, Sviščaki, Snežnik,17-28.VI.2009, Manfred Egger leg.
L=32 mm
Copyright © Ruslan Panin

Carabus (Megodontus) caelatus schreiberi Kraatz, 1887
Male adult
SW Slovenia, Obrov, Poljane, 14-27.VIII.2010, Manfed Egger leg.
L=33 mm
Copyright © Ruslan Panin

Carabus (Megodontus) croaticus zepcensis (as bosiljevici Drovenik & Pavićević, 1985)
Male adult
CW-Serbia, Zlatar Mts, Nova Varoš, Zlatar Lake env., 1-12.VII.2007, P.Sima leg. L=30 mm
Copyright © Ruslan Panin

Carabus (Megodontus) caelatus dalmatinus (as procerus Reitter, 1885)
Male adult
S Bosnia, Potoci env. h=500 m, VII.2006, Bartoloni leg.
L=38 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus azurescens Dejean, 1826
Endophallus
W-Croatia, Velebit Mts., betw.Karlobag & Brusane vill.,
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus azurescens Dejean, 1826 (loc.typ.)
Male adult
W-Croatia, Velebit Mts., betw.Karlobag & Brusane vill., VI.2002, Manfred Egger leg.
L=30 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus germarii Sturm, 1815
Male adult
N Italy, Trent, Ampola, Mt. Tremalzo h=1700m, 3-13.VI.2010, Manfred Egger leg.
29 mm
Copyright © Ruslan Panin

Carabus (Megodontus) violaceus germarii Sturm, 1815
Endophallus
N Italy, Trent, Ampola, Mt. Tremalzo h=1700m
Copyright © Ruslan Panin

Carabus (Megodontus) caelatus dalmatinus Duftschmid, 1812
Female adult
Bosnia, Mostar; 40 mm
Copyright © Aleš Sedláček

Carabus (Megodontus) croaticus antoniocaldoni Rapuzzi, 2014 Holotype Paratype
Male+Female adult
Serbia, Kopaonik mountain, 1500m, IX.2013, I. Rapuzzi and L. Caldon leg; (Ivan Rapuzzi collection).
Copyright © - - -

Carabus (Megodontus) croaticus croaticus Dejean, 1826
Endophallus
W-Croatia, Velebit Mts., h=950 m, Oltari - Zavizan,
Copyright © Ruslan Panin

Carabus (Megodontus) croaticus croaticus Dejean, 1826
Female adult
W-Croatia, Velebit Mts., h=950 m, Oltari - Zavizan, 30.VI. 2018, L=32 mm
Copyright © Ruslan Panin

Carabus (Megodontus) croaticus croaticus Dejean, 1826
Male adult
W-Croatia, Velebit Mts., h=950 m, Oltari - Zavizan, 30.VI.2018
Copyright © Ruslan Panin

Carabus (Megodontus) caelatus dalmatinus Duftschmid, 1812
Female adult
Croatia, Brač island, 2025, D. Strelnikoff leg. (cAA)
Copyright © Alexander Anichtchenko

Carabus (Megodontus) caelatus dalmatinus Duftschmid, 1812
Male adult
Croatia, Brač island, 2025, D. Strelnikoff leg. (cAA)
Copyright © Alexander Anichtchenko

Carabus (Megodontus) caelatus dalmatinus Duftschmid, 1812
Male adult
Croatia, Brač island, 2025, D. Strelnikoff leg. (cAA)
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) illigeri illigeri (as bjelasnicensis Apfelbeck, 1902)
Male+Female adult
Serbia, Debelo Brdo, 5.VI.2007, S. Sapaly leg. L=32 mm photo Panin R.
Copyright © Ruslan Panin

Carabus (Morphocarabus) illigeri illigeri Dejean, 1826 (as bjelasnicensis Apfelbeck, 1902)
Endophallus
endophallus as bjelasnicensis Apfelbeck, 1902
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens trans illigeri Dejean, 1826
Male+Female adult
W-Croatia, Karlovac, near Bosiljevo vill., h~300m, V.2000, I.Rapuzzi leg. female, 33 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens trans illigeri Dejean, 1826
Endophallus
W-Croatia, Karlovac, near Bosiljevo vill., h~300m,
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens trans scheidleri
Endophallus
NC Croatia, Ivanščica Mts , Ivanec,4-18.VI.2011, Krause Rene leg.
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens trans scheidleri
Male adult
NC Croatia, Ivanščica Mts , Ivanec,4-18.VI.2011, Krause Rene leg.<br /> L=31 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens praecellens Pillardi, 1825
Male adult
Serbia ,
Frushka Gora, Ruma VI.1985
Copyright © Igor Solodovnikov

Carabus (Morphocarabus) praecellens praecellens Pillardi, 1825
Endophallus
Serbia, Fruska Gora Mts, 20-31.V.2006, M.Sima leg.
Copyright © Ruslan Panin

Carabus (Morphocarabus) praecellens praecellens Pillardi, 1825
Male adult
Serbia, Fruska Gora Mts, 20-31.V.2006, M.Sima leg.
L=29 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) illigeri illigeri (as tarensis Toševski, 1990)
Male adult
WC-Serbia: National park Tara Mts.:NE Mitrovac h=1500m, 1-12.VII.2007, Pr.Sima leg. L=30 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) illigeri illigeri (as tarensis Toševski, 1990)
Endophallus
WC-Serbia: National park Tara Mts.:NE Mitrovac h=1500m,
Copyright © Ruslan Panin

Carabus (Pachystus) hortensis hortensis Linne, 1758
Male adult
Germania, Oberfranken, Langenau
leg.det. U.Schmidt, 2002
Foto: U.Schmidt, 2006
Copyright © U.Schmidt

Carabus (Pachystus) hortensis neumeyeri Schaum, 1856
Female adult
Italy, Basilicata, Serra Calvello (PZ)
1400 m, 7.2003 F. Angelini leg.
female, 25 mm
Copyright © Alexander Anichtchenko

Carabus (Pachystus) hortensis chalkidikensis Müller, 2009 Holotype Paratype
Male+Female adult
Griechenland, Makedonien, Chalkidike, Holomondas Berge, zwischen den Orten Amea und Taxiarhis, 932 m, N 40°27'861", E0 23O32'661" leg. et coll. A. Muller (Wien).
Copyright ©

Carabus (Pachystus) hortensis hortensis Linne, 1758
Endophallus
W-Ukraine, Male Polissya, near Kamenka-Buzka vill.
Copyright © Ruslan Panin

Carabus (Pachystus) hortensis hortensis Linne, 1758
Female adult
Natural hybrid Carabus hortensis X Carabus granulatus. Belarus, Minsk reg., 8,7 km of Borisov, near vill. Lyubatovstchina, pine forest (Pinetum pleuroziosum), h = 213 m, 54.31934° N, 28.511486° E, 20-27.09.2018, leg. Klychenok Snezhana
Copyright © Igor Solodovnikov

Carabus (Oreocarabus) hortensis hortensis Linne, 1758
Male adult

Copyright © Kirill Makarov

Carabus (Pachystus) glabratus gibbosus Heyden, 1866
Male adult
N Croatia, Ivanščica Mts, Ivanec, 4-18.VI.2011, Krause Rene leg., L=32 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) irregularis ramanus Sokolar, 1909 (as hayeki Kobmann, 1925)
Male adult
N Slovenia, Maribor – Dravograd, Pomorje Mts.,13.XI.2005, I.Rapuzzi leg.
Copyright © Ruslan Panin

Carabus (Platycarabus) irregularis ramanus Sokolar, 1909 (as bucephalus Kraatz, 1879)
Male+Female adult
W Slovenia, Goriška region Trnovo, h~1000 m, IX.2002, I.Rapuzzi leg.
female, 29 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) creutzeri creutzeri Fabricius, 1801 (as trnovensis Bernau, 1914)
Male adult
W Slovenia,
Lokve, Trnovski Gozd h=1100m, 17.IV.2006, leg. I.Rapuzzi
male, 26mm
Copyright © Ruslan Panin

Carabus (Platycarabus) creutzeri creutzeri Fabricius, 1801
Male adult
Italy, Fruili - UD - Carnia, N. Paulano 29.III.2002 leg. I.Rapuzzi
L=22.0 mm
Copyright © Kirill Makarov

Carabus (Platycarabus) creutzeri creutzeri Fabricius, 1801(as posarinii Rapuzzi, 1999 PT) Paratype
Male+Female adult
Slovenia:Koper:Kras Mt.R.:Brkini Mts.:Suhorije env., VIII.1998, I.Rapuzzi leg.
photo & koll. Panin R.
female, 32 mm
Copyright © Ruslan Panin

Carabus (Platycarabus) irregularis ramanus Sokolar, 1909
Female adult
NW Croatia, Ivanščica Mts , Ivanec, 4-18.VI.2011, Krause Rene leg., L=33 mm
Copyright © Ruslan Panin

Carabus (Procerus) gigas gigas Creutzer, 1799
Male adult
Croatia
Copyright © Stanislav Krejcik

Carabus (Procerus) gigas gigas Creutzer, 1799
Male+Female adult
N Croatia, Ivanščica Mts , Ivanec, 4-18.VI.2011, Krause Rene leg.
female, 54 mm
Copyright © Ruslan Panin

Carabus (Procrustes) coriaceus Linne, 1758
Larva
Latvija photo Andris Bukejs
Copyright © Alexander Anichtchenko

Carabus (Procrustes) coriaceus coriaceus Linne, 1758
Female adult
The neighborhood Nicolo-Berezovskoe republic of
Bashkortostan 5.07
Copyright © Sunvas

Carabus (Procrustes) coriaceus coriaceus Linne, 1758
Male adult
length: 32.42 mm
Germania, Oberfranken, Weismain
leg.det. U.Schmidt, 1983
Foto: U.Schmidt, 2005
Copyright © U.Schmidt

Carabus (Procrustes) coriaceus coriaceus Linne, 1758
Female adult
Belgique : Prov. Luxembourg, Harnoncourt 31.vii.1975 - M. Grotz Leg. (Size : 36 mm)
Copyright © Noël Mal

Carabus (Procrustes) coriaceus excavatus Charpentier, 1825
Male adult
W Croatia, Velebit Mts, Jablanac Stirovaca h=998 m, 44.71124, 14.95673,3.VI-7.VII.2017, G. Proscia leg., L= 37 mm
Copyright © Ruslan Panin

Carabus (Procrustes) coriaceus excavatus Charpentier, 1825
Female adult
W Croatia, Velebit Mts, Jablanac Stirovaca h=998 m, 44.71124, 14.95673,3.VI-7.VII.2017, G. Proscia leg., L= 38 mm
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
Male adult
Italy-Switzerland, Riviera – Preonzo, 18.II.1998, C.Sola leg.
L=26 mm
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
Male adult
relict lowland populations of blue-purple color, of the the river Ticino park - Italy - Lombardy
Copyright © Monzini

Carabus (Tachypus) cancellatus intermedius Dejean, 1826
Female adult
NW -Serbia,Fruška Gora,Vrdnik env. h=380m, 9-10.VII.2013, M. Janalik leg. L=29 mm
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
Male adult
Carabus (s.str.) cancellatus emarginatus n.monzinii Monzini, 1983 25,5 mm
Copyright © Igor Solodovnikov

Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
Endophallus
Carabus (Tachypus) cancellatus emarginatus n. dahli Heer, 1841
Copyright © Igor Solodovnikov

Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
Male adult
Carabus (Tachypus) cancellatus emarginatus n. dahli Heer, 1841
Copyright © Igor Solodovnikov

Carabus (Tachypus) cancellatus emarginatus Duftschmid, 1812
Female adult
Carabus (Tachypus) cancellatus emarginatus n. dahli Heer, 1841
Copyright © Igor Solodovnikov

Carabus (Tachypus) cancellatus intermedius (as maximus Haury, 1880)
Male adult
SW Hungary, Bátaapáti, Com. Baranya, 6-9.VI.2003, Gy. Szel leg/det. I.Retezar 2010, L=25 mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) convexus s.str. (as pyrenaicus Born, 1907)
Male adult
Pirineos, Canfranc, 1300m, Barranco de Izas
15.6.2005 Anichtchenko A. leg.
male, 16 mm
Copyright © Alexander Anichtchenko

Carabus (Tomocarabus) convexus s.str. (as paganetti Born, 1905)
Male adult
Lazio, Roma, Bosco di Nettuno
22.5.1986 C. Iacovone leg.
male, 20 mm
Copyright © Alexander Anichtchenko

Carabus (Tomocarabus) convexus convexus Fabricius, 1775
Male adult
W.Ukraine,
Transcarpathian region,Vinogradov distr., near Malaya Kopanya vill., Chorna Gora Mt., h=180m. 7-18.VII.08. Panin R. leg.
male 18 mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) convexus s.str. (ab. niger)
Male adult
W Ukraine, NW Podolia, Lviv Region, Zolochiv distr., near Gologirky vill., in steppe 1-19.V.2007 Panin R. leg.
16mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) convexus moreanus Reitter, 1896
Female adult
Greece:, N Peloponnes, Xylokastro, Zemeno,h=1000m, 6.VI.1997, Manfred Egger leg. L=21 mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) convexus dilatatus (as perplexus Schaum, 1857)
Male adult
CE-Greece: Larissa reg., Stomio, Ossa Mt., 6-17.VII.2011, Krause Rene leg.
L=19 mm
Copyright © Ruslan Panin

Carabus (Tomocarabus) convexus moreanus Reitter, 1896
Male adult
S Greece, Taygetos Mts.; 13mm
Copyright © Aleš Sedláček

Carabus (Tomocarabus) convexus dilatatus Dejean, 1826
Male adult
NE Serbia, Majdanpek; 13mm
Copyright © Aleš Sedláček