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New Images added

Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878
Female adult
Russia, Far East, Primorskiy Kray, Chuguevsky distr., 30 km ESE from Chuguevka, 9-14.VIII.2007, S.Shabalin leg.
Copyright © Ruslan Panin

Cicindela (Cicindela) restricta spinigera Eschscholtz, 1829
Unsexed
Kluchevskaya Sopka, near Springs (ZIN)
Copyright © Igor Solodovnikov

Cicindela (Cicindela) restricta spinigera Eschscholtz, 1829
Female adult
Kamchatka (ZIN)
Copyright © Igor Solodovnikov

Cicindela (Cicindela) restricta spinigera Eschscholtz, 1829
Unsexed
Kamchatka, Kluchevskaya Sopka volcan (ZIN)
Copyright © Igor Solodovnikov

Calomera lunulata (Fabricius, 1781)
Female adult
Morocco, Zagora distr., Nesrate vill., Qued Draa, 30.01.01N 005.31.51W, 28.6.2010, Anichtchenko A. leg. (cAA). 14.5 mm
Copyright © Alexander Anichtchenko

Calomera lunulata (Fabricius, 1781)
Male adult
Morocco, Zagora distr., Nesrate vill., Qued Draa, 30.01.01N 005.31.51W, 28.6.2010, Anichtchenko A. leg. (cAA)
Copyright © Alexander Anichtchenko

Calomera lunulata (Fabricius, 1781)
Male adult
Morocco, Zagora distr., Nesrate vill., Qued Draa, 30.01.01N 005.31.51W, 28.6.2010, Anichtchenko A. leg. (cAA). 14 mm
Copyright © Alexander Anichtchenko

Carabus (Morphocarabus) hummeli smaragdulus Kraatz, 1878 (variability)
Male adult
Far East, Promorskij Krai, Ussuriisk reg., Kamenushka, 9-15.6.2008 (cAA)
Copyright © Alexander Anichtchenko

Calochroa (Calostola) bicolor haemorrhoidalis (Wiedemann, 1823)
Male adult
Pakistan, prov. Pendjab, new Fakirabab, 08.07.2025, leg. Ivanov A.S.; det. Anichtchenko A. 2025
Copyright © Igor Solodovnikov

Carabus (Tachypus) cancellatus carinatus (as subcarinatus Kolbe, 1912)
Male adult
E-Netherlands, Gelderland prov., Nijmegen., Berg en Dal vill., 6.XII.2014, H.de Haan leg., L=25 mm
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus durus Reitter, 1896
Female adult
W-Ukraine, Transcarpathian Lowland, Mukachevo distr., near Znayatsevo vill., 12.VI-8.VII.2009, R. Panin leg., L=30 mm
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus cancellatus (as charcoviensis Bernau, 1915)
Female adult
NE-Ukraine, Kharkov reg., Zmiev distr., 1-7.VI.2007, Kovalchuk I. leg., L=27 mm
Copyright © Ruslan Panin

Carabus (Tachypus) cancellatus durus (as carpathicus Kolbe, 1913)
Male adult
W-UKRAINE, Ivano-Frankivsk reg., Gorgany Mts, Mt.Synyak h =1530м., 25.V-17.VII.2014, Panin R. leg., L=28 mm
Copyright © Ruslan Panin

New Taxa added

Species telnovi Wiesner & Anichtchenko, 2025 (Therates, Cicindelini, Cicindelinae) VTN
Species biluoshanensis Deuve, 2025 (Cychrus, Cychrini, Carabinae) CH (Yunnan)
Species sehnalianus Deuve, 2025 (Cychrus, Cychrini, Carabinae) CH (Yunnan)
Species changgui Jiang & Zeng, 2025 (Pterostichus, Pterostichini, Pterostichinae) CH (Chongqing)
Subspecies durus Reitter, 1896 (Carabus, Carabini, Carabinae) PL, RO, SK, UK
Subspecies riparius sensu stricto (Pterostichus, Pterostichini, Pterostichinae) CAN, USA (Alaska; Idaho; Montana; Oregon; Washington)
Species polilloensis Dheurle, 2025 (Neocollyris, Collyridini, Cicindelinae) PLP (Polillo Islands)
Species badzhaliensis Sundukov, 2025 (Pterostichus, Pterostichini, Pterostichinae) RU (Khabarovsky Province)
Species kirilli Sundukov, 2025 (Pterostichus, Pterostichini, Pterostichinae) RU (Khabarovsky Province)
Species storozhenkoi Sundukov, 2025 (Pterostichus, Pterostichini, Pterostichinae) RU (Khabarovsky Province)

Subspecies roxinai Deuve, 2009 (Carabus, Carabini, Carabinae) BRM
Subspecies punctifera sensu stricto (Cymindis, Lebiini, Lebiinae) USA (Arizona; New Mexico; Texas), MEX
Subspecies toltec Hilchie & Ball, 2024 (Cymindis, Lebiini, Lebiinae) MEX
Species crenatoverpa Hilchie & Ball, 2024 (Cymindis, Lebiini, Lebiinae) MEX (Nayarit)
Species cuyuteca Hilchie & Ball, 2024 (Cymindis, Lebiini, Lebiinae) MEX (Jalisco)
Species apache Hilchie & Ball, 2024 (Cymindis, Lebiini, Lebiinae) USA (Arizona)
Species geminata Hilchie & Ball, 2024 (Cymindis, Lebiini, Lebiinae) MEX (Morelos)
Species huichilobos Hilchie & Ball, 2024 (Cymindis, Lebiini, Lebiinae) MEX (Oaxaca)
Species rugofrons Hilchie & Ball, 2024 (Cymindis, Lebiini, Lebiinae) MEX (Jalisco)
Species tonatiuh Hilchie & Ball, 2024 (Cymindis, Lebiini, Lebiinae) MEX (Oaxaca)
Species yaqui Hilchie & Ball, 2024 (Cymindis, Lebiini, Lebiinae) USA (Arizona)
Species zacapa Hilchie & Ball, 2024 (Cymindis, Lebiini, Lebiinae) GTM
Species zapotec Hilchie & Ball, 2024 (Cymindis, Lebiini, Lebiinae) MEX (Oaxaca)
Species mcclevei Shpeley & Devender 2022 (Calathus, Sphodrini, Platyninae) MEX (Sonora)
Species tigrinus Shpeley & Devender 2022 (Calathus, Sphodrini, Platyninae) MEX (Sonora)
Species balli Shpeley & Van Devender, 2022 (Amblygnathus, Harpalini, Harpalinae) MEX (Sonora)
Subspecies boulardi sensu stricto (Tachypeles, Ozaenini, Paussinae) GYN
Subspecies nigrostriatus Deuve, 2025 (Tachypeles, Ozaenini, Paussinae) GNF
Species hudsonioides Deuve, 2025 (Tachypeles, Ozaenini, Paussinae) PER
Species obandoi Deuve, 2025 (Tachypeles, Ozaenini, Paussinae) CRC
Species zonensis Deuve, 2025 (Tachypeles, Ozaenini, Paussinae) PNM
Species quesadai Deuve, 2025 (Tachypeles, Ozaenini, Paussinae) CRC
Species reductops Deuve, 2025 (Tachypeles, Ozaenini, Paussinae) PNM
Species minibrunneus Deuve, 2025 (Tachypeles, Ozaenini, Paussinae) GYN
Species constantiniana Deuve, 2025 (Filicerozaena, Ozaenini, Paussinae) PER
Species gaujoni Deuve, 2025 (Filicerozaena, Ozaenini, Paussinae) ECD
Species woytkowskii Deuve, 2025 (Filicerozaena, Ozaenini, Paussinae) PER
Subspecies filiformis sensu stricto (Proozaena, Ozaenini, Paussinae)
Subspecies nigrescens Deuve, 2025 (Proozaena, Ozaenini, Paussinae) PER
Species belizensis Deuve, 2025 (Proozaena, Ozaenini, Paussinae) BLZ
Species matogrossensis Deuve, 2025 (Goniotropis, Ozaenini, Paussinae) BRZ
Species sinuatus Roux, 2025 (Archastes, Nebriini, Nebriinae) CH (Sichuan)
Species relegatus Roux, 2025 (Archastes, Nebriini, Nebriinae) CH (Sichuan)
Subspecies grandis sensu stricto (Cicindela, Cicindelini, Cicindelinae) ANG
Subspecies pseudocinctus (W. Horn, 1913) (Cicindela, Cicindelini, Cicindelinae) TAN
Subspecies quadrisignata sensu stricto (Megacephala, Megacephalini, Cicindelinae) CAM, WAF (Nigeria), CAR, CHD
Subspecies rivalieri Basilewsky, 1966 (Megacephala, Megacephalini, Cicindelinae) WAF (Benin; Burkina Faso; Togo)
Subspecies bergeri Peeters & I.Rapuzzi, 2025 (Carabus, Carabini, Carabinae) AU
Species uludaghensis Machard, 2025 (Platyderus, Sphodrini, Platyninae) TR
Species tauricus Machard, 2025 (Platyderus, Sphodrini, Platyninae) TR
Species gracilis Machard, 2025 (Platyderus, Sphodrini, Platyninae) TR
Species drouxi Machard, 2025 (Platyderus, Sphodrini, Platyninae) TR
Species montisabanti Machard, 2025 (Platyderus, Sphodrini, Platyninae) TR
Species eregliensis Machard, 2025 (Platyderus, Sphodrini, Platyninae) TR
Species narlicaensis Machard, 2025 (Platyderus, Sphodrini, Platyninae) TR
Subspecies durmitorensis sensu stricto (Duvalius, Trechini, Trechinae) BH, ME
Subspecies bulbosus Janák, 2025 (Duvalius, Trechini, Trechinae) ME
Species ignotus N.Ito, 2017 (Oxycentrus, Harpalini, Harpalinae) IDS (Borneo), MLS
Species angustipennis N.Ito, 2013 (Oxycentrus, Harpalini, Harpalinae) CBD
Species laetus N.Ito, 2013 (Oxycentrus, Harpalini, Harpalinae) LAO
Species shpeleyi Messer & Raber, 2025 (Discoderus, Harpalini, Harpalinae) MEX (Oaxaca)
Species nonarcuatus Messer & Raber, 2025 (Discoderus, Harpalini, Harpalinae) MEX (San Luis Potosi)
Species pueblaensis Messer & Raber, 2025 (Discoderus, Harpalini, Harpalinae) MEX (Guerrero; Oaxaca; Puebla)
Species circulus Messer & Raber, 2025 (Discoderus, Harpalini, Harpalinae) USA (New Mexico; Texas), MEX (Chihuahua; Coahuila; Durango)
Species amblyderus (Bates, 1878) (Discoderus, Harpalini, Harpalinae) MEX
Subgenus Selenalius Casey, 1914 (Discoderus, Harpalini, Harpalinae)
Subgenus Paraselenalius Messer & Raber, 2025 (Discoderus, Harpalini, Harpalinae)
Genus Discoderus LeConte, 1853 (Harpalini, Harpalinae)
Species elongatulus (Dejean, 1831) (Tachys, Bembidiini, Trechinae) FR, IT (Sardegna), PT, SP, AG, MO, TU
Subspecies montreuilianus Deuve, 2014 (Carabus, Carabini, Carabinae) GG
Subgenus Trachycarabus Géhin, 1885 (Carabus, Carabini, Carabinae)
Species bartolozzii Allegro, 2025 (Pseudaptinus, Zuphiini, Dryptinae) ECD
Species moreti Allegro, 2025 (Pseudaptinus, Zuphiini, Dryptinae) ECD
Species gracilis Allegro, 2025 (Pseudaptinus, Zuphiini, Dryptinae) ECD
Species mateui Ryznar & Anichtchenko, 2025 (Syntomus, Lebiini, Lebiinae) IND (Sikkim; Uttarakhand; West Bengal), NP
Species kratkyi Machado, 2025 (Orzolina, Bembidiini, Trechinae) CI
Subspecies pseudomanifestus Rapuzzi, 2009 (Carabus, Carabini, Carabinae) NC
Subspecies bedelianus sensu stricto (Orthomus, Pterostichini, Pterostichinae) MR
Subspecies isambertoi Serrano & Boieiro, 2025 (Orthomus, Pterostichini, Pterostichinae) MR
Subspecies bostockii sensu stricto (Megacephala, Megacephalini, Cicindelinae) AST
Subspecies viridisericea (Sumlin, 1992) (Megacephala, Megacephalini, Cicindelinae) AST
Subspecies basalis sensu stricto (Megacephala, Megacephalini, Cicindelinae) AST
Subspecies phaeoxantha (W. Horn, 1894) (Megacephala, Megacephalini, Cicindelinae) AST
Subspecies australasiae sensu stricto (Megacephala, Megacephalini, Cicindelinae) AST
Subspecies nickerli (Srnka, 1895) (Megacephala, Megacephalini, Cicindelinae) AST
Subspecies horii sensu stricto (Thopeutica, Cicindelini, Cicindelinae) IDS (Sulawesi)
Subspecies paucirufescens Cassola & Brzoska, 2008 (Thopeutica, Cicindelini, Cicindelinae) IDS (Sulawesi)
Subspecies eximia sensu stricto (Thopeutica, Cicindelini, Cicindelinae) IDS (Sulawesi)
Subspecies sangihensis Cassola, 1991 (Thopeutica, Cicindelini, Cicindelinae) IDS (Sulawesi)
Species khasanensis Sundukov & Makarov, 2025 (Amara, Zabrini, Pterostichinae) RU (Maritime Prov)
Subspecies cordimeatus Cavazzuti, 1997 (Carabus, Carabini, Carabinae) CH (Sichuan)
Subspecies emituo Imura, 2017 (Carabus, Carabini, Carabinae) CH (Yunnan)
Subspecies sue Imura, 2012 (Carabus, Carabini, Carabinae) JA (Shikoku)
Species obliquealba (Motschulsky, 1864) (Cylindera, Cicindelini, Cicindelinae) BRZ
Species acompsa (Chaudoir, 1852) (Cylindera, Cicindelini, Cicindelinae) BRZ
Species ocskayi (Gistl, 1837) (Cylindera, Cicindelini, Cicindelinae) BRZ
Species amayai Moravec & Šafránek, 2025 (Cylindera, Cicindelini, Cicindelinae) BOL
Subspecies dalong Deuve & Kaláb, 2009 (Carabus, Carabini, Carabinae) CH (Sichuan)
Genus Gadorites Ortuño & Muñoz-Santiago, 2025 (Pterostichini, Pterostichinae)
Species heterocostatus Deuve & Schmidt, 2023 (Carabus, Carabini, Carabinae) NP

More taxa

stop z-russism! Stop war!

Published: 14 September 2025 11:100 commentsPublished by: Alexander Anichtchenko

stop z-russism! Stop war!

Mess in genus Carabus

Published: 09 August 2025 21:290 commentsPublished by: Alexander Anichtchenko

Over the last 40 years, a huge number of taxa have been described in the genus Carabus.
Unfortunately, most of them were described by amateurs, often for self-love or commercial purposes. Most types deposited in private collections, which makes a classical revision of the genus with synonymization of fictitious taxa in accordance with the Code of Zoological Nomenclature almost impossible.
In our database, all weak species, subspecies and obvious synonyms in Carabus will be listed as synonyms.

The survival

Published: 24 July 2025 18:230 commentsPublished by: Alexander Anichtchenko

The survival and development of the site directly depends on your subscriptions and donations. Please, click subscribe

Maximum likelihood phylogenetic hypothesis for the placement of tiger beetles within Carabidae

Published: 20 July 2023 23:462 commentsPublished by: Alexander Anichtchenko

Circular cladogram representing the maximum likelihood phylogeny inferred in IQ-TREE using the Comp_DT dataset comprising nine gene fragments and about 11kb of molecular data. The branch lengths have been modified to ease the visualization of relationships
between subfamilies within Carabidae (image from H. Gough, 2020) Maximum likelihood phylogenetic hypothesis for the placement of tiger beetles within Carabidae

Maximum likelihood phylogenetic hypothesis for the placement of tiger beetles within Carabidae

Good Buy year of Tiger and happy new year of Rabbit!

Published: 23 December 2022 20:440 commentsPublished by: Alexander Anichtchenko

Dear Friends, Colleagues, Contributors and Visitors! I wish you Merry X-mas and happy new year of the Rabbit 2023!
Sincerely, Alexander
Good Buy year of Tiger and happy new year of Rabbit!

Good Buy year of Tiger and happy new year of Rabbit!

Happy New Year of the Tiger!

Published: 23 December 2021 22:210 commentsPublished by: Alexander Anichtchenko

Dear Friends, Colleagues, Contributors and Visitors! I wish you Merry X-mas and happy new year of the Tiger 2022!
Sincerely, Alexander
Happy New Year of the Tiger!

Coléoptères as example of profanation of science

Published: 23 May 2021 20:47Category: Our news1 commentPublished by: Alexander Anichtchenko

Two things are infinite: the universe and human stupidity; and I'm not sure about the universe. Albert Einstein
TARRIER (M.), 2021. – Nouvelles sous-espèces de France et d’Espagne du genre Carabus L., 1758 (Coleoptera, Carabidae)- Paru le 29 juin 2021 - Issued on June 29, 2021
Summary
Descriptions of nineteen new subspecies in the genus Carabus L., 1758, from
France and Spain: Carabus (Ctenocarabus) melancholicus tietarensis n. subsp., C.
(Oreocarabus) guadarramus hirschfelderi n. subsp., C. (Oreocarabus) guadarramus
sagranus n. subsp., C. (Mesocarabus) lusitanicus problematicoides n. subsp., C. (s. str.)
deyrollei neoreductus n. subsp., C. (Archicarabus) pseudomonticola hispanicola n. subsp.,
C. (Archicarabus) nemoralis litigiosus n. subsp., C. (Archicarabus) nemoralis
montisdiniensis n. subsp., C. (Chrysocarabus) solieri devillei n. subsp., C.
(Chrysocarabus) solieri strobinoi n. subsp., C. (Chrysocarabus) solieri teillonensis n.
subsp., C. (Chrysocarabus) solieri axiomorphus n. subsp., C. (Chrysocarabus) solieri piezus n. subsp., C. (Chrysocarabus) solieri dujardini n. subsp., C. (Chrysocarabus)
solieri rouyricus n. subsp., C. (Chrysocarabus) lateralis subsalmantinus n. subsp., C.
(Chrysocarabus) lateralis sanabriensis n. subsp., C. (Chrysocarabus) lateralis opulentior
n. subsp., C. (Macrothorax) rugosus baeticoides n. subsp.

Stop describing fake species!

Published: 16 May 2021 10:310 commentsPublished by: Alexander Anichtchenko

Stop describing fake species!
Under each stone does not live another species new to science.
If you follow the logic of some "specialists", then Homo sapiens can be divided into several species, and dogs into several genera.

Happy New Year of the Bull!

Published: 23 December 2020 17:43Category: Our news1 commentPublished by: Alexander Anichtchenko

Dear Friends, Colleagues, Contributors and Visitors! I wish you Merry X-mas and happy new year 2021!
Sincerely, Alexander
Happy New Year of the Bull!

Lost in Determination

Published: 13 November 2020 03:29Category: Site News0 commentsPublished by: Alexander Anichtchenko

Taxonomy in 21st Century: Hobby or Business?

Published: 12 February 2020 16:41Category: Site News0 commentsPublished by: Alexander Anichtchenko

Recently was published excellent paper of Fominykh, Zamotajlov, Khomitskiy & Titarenko, 2020 "Contribution to the knowledge of Carabus (Lipaster) stjernvalli Mannerheim, 1830 in the Caucasus, with description of two new subspecies (Coleoptera: Carabidae: Carabini)", "dedicated to study of variability of species Carabus (Lipaster) stjernvalli", although, in fact, the sole purpose of the work was to describe new taxa in order to sell their Paratypes.
But, this work perfectly proves that there are no differences between the previously described subspecies, as well as the newly described ones. New synonymy in the species Carabus (Lipaster) stjernvalli proposed here 12.2.2020 by A. Anichtchenko