Nesamblyops sp.
Male adult
New Zealand
Scale bars = 0.5mm
image from Sokolov I., 2013. A new genus and eight new species of the subtribe Anillina (Carabidae, Trechinae, Bembidiini) from Mexico, with a cladistic analysis and some notes on the evolution of the genus.
Copyright © Igor Sokolov

Anillinus affabilis (Brues, 1902) Lectotype
Male adult
Image from Sokolov I.M., Reddell J.R. & Kavanaugh D.H., 2014 Scale = 0.5 mm.
Copyright © Igor Sokolov

Illaphanus pteridophyticus Giachino, 2005 Holotype
Male adult

Copyright © Alexander Anichtchenko

Scotodipnus alpinus Baudi di Selve, 1871
Female adult
Piemonte, Valle Po, Crissolo (CN), 27.V.2000, 1250m, fagus, leg. F. Angelini
Copyright © Alexander Anichtchenko

Nesamblyops sp.
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Winklerites stevanovici Hlavac & Margini, 2017 Paratype
Male adult
Srbija [p], Dimitrovgrad, Petrlaš, Džemanska prop[ast], 02.05.2002 [h], M. Stevanović leg. Image from HLAVÁČ & MAGRINI, 2017
Copyright © - - -

Rhegmatobius quadricollis (Ehlers, 1883)
Unsexed
Majella, Abruzzo, Italy
Copyright © Vittorino Monzini

Microtyphlus saxatilis Magrini, Leo & Fancello, 2004
Male adult
Italia, Sardegna, Perda Longa (NU) Baunei.
Copyright © Augusto Degiovanni

Anillaspis explanata (Horn, 1888)
Female adult
USA: California: El Dorado Co., Alabaster Cave (CMNH). Scale bar 1 mm; Image from LaBonte & Maddison, 2023
Copyright © - - -

Megastylulus pivai Giachino & Sciaky, 2003 Holotype
Male adult
leg. E. Piva - Isl. St.Lucia Caraibi
Copyright © Vittorino Monzini

Tasmanitachoides major Baehr, 1990 Holotype
Male adult

Copyright © Museums Victoria

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Image from Sallenave, Erwin, and Roig-Juñent, 2008
Copyright © - - -

Andinodontis muellermotzfeldi Toledano & Erwin, 2010
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidarenas setiventris Negre, 1973
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides sp.
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides erwini Maddison & Porch, 2021 Holotype
Male adult
Australia: Tasmania: River Forth at C136, 41.4712°S 146.1366°E, 126 m, 14.i.2019. DRM 19.012. D.R. Maddison & N.A. Porch. Scale bar 1mm
Copyright © David Maddison

Andinodontis maveetyae Erwin & Maddison, 2010 Holotype
Male adult
Perú: Cusco, Rocotal, 3.2 km E Tres Cruces, 13.1123° S, 71.5834° W, 2082 m, 27 June 2010 (S. MAVEETY) (MHN-UNSAAC: ADP124815)
Copyright © Terry Erwin

Bembidion (Limnaeoperyphus) quadriimpressum (Motschulsky, 1860)
Male adult
Kunashiri Isl., mouth of Medny Spring 44°16'31"N 145°56'29"E 27.VI.2008 leg. K.Makarov
L=4.3
Copyright © Kirill Makarov

Bembidion (Hirmoplataphus) nigrum Say, 1823
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Trepanedoris) atripes (Motschulsky, 1844)
Male adult

Copyright © Kirill Makarov

Bembidion (Diplocampa) fumigatum (Duftschmid, 1812)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Ocys harpaloides (Audinet-Serville, 1821)
Male adult
Spain
Copyright © Kirill Makarov

Bembidion (Terminophanes) kaschmirense Netolitzky, 1920
Female adult
India, Uttarakhand, 5 km NE Govind Ghat, road to Gangaria, 9-10.6.2011, Anichtchenko A. leg. det. J. Schmidt 2015 8 mm
Copyright © Alexander Anichtchenko

Bembidion (Terminophanes) mckinleyi scandicum Lindroth, 1943
Female adult
E Siberia, Lake Baikal, Malyi Kochergat vill., Goloustnaya riv., 16.VII.1991, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Bembidion (Bembidionetolitzkya) namtso Schmidt, 2018 Paratype
Male adult
“TIBET (South Central) 12.VII.07 / S Namtso, Langma Valley / 5100-5150 m, leg. Schmidt / 30°37’39,1”N90°51’56,5”E” (CSCHM).
Copyright © Joachim Schmidt

Sinechostictus (Pseudolimnaeum) lederi (Reitter, 1888)
Female adult

Copyright © Kirill Makarov

Bembidion (Nesocidium) corticarium Sharp, 1903
Female adult
Halalea For. Res., Namolokama Mtn., 1295-1340 m el.. Image from Liebherr, 2008
Copyright © - - -

Horologion speokoites Valentine, 1932 Holotype
Male adult
"Arbuckle's Cave Maxwelton, W.Va. July 12 1931 J.M. Valentine / On bit of wet decaying wood, muddy floor of lower level / Type gen. Type sp. / U.S.N.M. Type 44255"
Copyright © Curt Harden

Lissopogonus tonkinensis Zamotajlov & Sciaky, 1996: 40
Male adult
N Vietnam, Lao Cai Province 250 km from Hanoi bearing 310°, SaPa vill env., h~1250 Hoang Lien Son Nat. Park 27.VI-2.VII.1998 leg.A.Napolov
L= 4,8
Copyright © Kirill Makarov

Lissopogonus cf. borneensis Baehr, 2001
Male adult
Malaysia, Borneo, Sarawak, Miri div., Bario 10 km NE, 3 km N Pa Lungan, 7-8.III.2014, R. Cibulskis leg. det. A. Anichtchenko 2015
Copyright © Alexander Anichtchenko

Lissopogonus morvani Deuve, 2009
Female adult
Nepal, Kosi Mure, 2000-2100m, 27.30N 87.16E, 6-12.6.2001. 7 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus Andrewes, 1923
Male adult
Nepal, 12km NE Chainpur, 1400m, 29.36.17N, 81.17.54E, 28.6.2009. 5.8 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus laosensis Deuve, 2009
Male adult
Laos, Nong Khiaw vic, 20.553945, 102.622681, 1.V.2026, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Patrobus assimilis Chaudoir, 1844
Female adult
CZ, N Bohemia; Krušné Hory Mts., Výsluní; 8mm
Copyright © Aleš Sedláček

Penetretus temporalis Bedel, 1909
Female adult
Spain, Cadiz prov., Algeciras vic., h=50m, 36.080429, -5.428613, 11-12.05.2014, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Dimorphopatrobus ludmilae Casale et Sciaky, 1994: 47
Female adult
Tibet, Motuo, Hanmi, 8.2013. det. A. Anichtchenko, 2015 17 mm
Copyright © Alexander Anichtchenko

Patrobus atrorufus (Stroem, 1768)
Male adult

Copyright © Kirill Makarov

Deltomerus (Deltomerus) jelineki Zamotajlov, 2001
Female adult
Iran Alburz prov Shemshak 2750m 12.06.17
Copyright © Jan Muilwijk

Platidiolus cf. borealis Zamotajlov, 2001
Female adult
Russia, Khabarovsk reg., Verhnebureinski distr. Bureya river valley, Ussomah river, 22.7.2006 (anal sternite with 3 pairs of setae). det. Anichtchenko A. 2018
Copyright © Alexander Anichtchenko

Penetretus nebrioides (Vuillefroy, 1866)
Male adult
SPAIN, Cantabria, 2 km E Puerto de San Glorio, near Vejo riv., 43.072369, -4.733061, 1300 m., 23.07.2019, A. Anichtchenko & R. Sciaky leg.; det. Anichtchenko A. 2019
Copyright © Alexander Anichtchenko

Prodiplous businskyi (Casale & Sciaky, 2003) Paratype
Male adult
China, SE Tibet, Zayu co., Irawaddy-Salween divide East of Tsema-la pass, 4300 m, 18.VI.1999 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Deltomerus (Deltomerus) corax Peyerimhoff, 1922
Male adult
Morocco, Grand Atlas, 2200-2300m, N31.13.09.5 W07.49.29.7 16-17.6.2010 Anichtchenko A. leg. & det. (cAA) 12 mm
Copyright © Alexander Anichtchenko

Penetretus andalusicus (Reitter, 1897)
Male adult
Spain, Sierra Nevada, Lagunillos de la Virgen, 2850m, 18.5.2005, A. Anichtchenko leg. & det (cAA)
Copyright © Alexander Anichtchenko

Pogonus (Pogonoidius) punctulatus Dejean, 1828
Male adult
Volgograd Reg., Pallas Distr., Elton Vill., on UV-light 16-18.VII.2006 K.Makarov leg.
L=6.4 mm
Copyright © Kirill Makarov

Pogonus (Pogonus) iridipennis Nicolai, 1822
Male adult
Volgograd Reg., Pallas Distr., N shore of Elton Lake, right bank of Khara Riv., mouth 13.V.2006 A.Matalin leg.
L=6.7 mm
Copyright © Kirill Makarov

Diodercarus arrowi Lutshnik, 1931: 528
Male adult
Iran, Hormozgan province, Bandar Abbas env., 12.III.2015, S. Azadbakhsh leg.
Copyright © Alexander Anichtchenko

Pogonus (Pogonus) minutus Dejean, 1828
Male adult
Pogonus senegalensis Dejean, 1831 det. M.Baehr, 2015. Size: 5,3 mm. Namibia, Etosha National Park, Okaukuejo, 1400 m leg. U.Schmidt, 18.-20.III.1994; Photo: U.Schmidt, 2016
Copyright © Udo Schmidt

Diplochaetus planatus (G.Horn, 1876)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Thalassotrechus barbarae (G.Horn, 1892)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Pogonus (Pogonus) fennelli Hudson, 2000
Female adult
Australia 7.5 mm; det. K. Will, 2020
Copyright © Alexander Anichtchenko

Iranopogonus freyellus Jedlicka, 1959 Holotype
Male adult
Iran, Sistan va Baluchestan Province, Chabahar env., IX.1937, leg. Band (NHMB). Image of Matthias Borer (NHMB) from Azadbakhsh, 2019
Copyright © - - -

Sirdenus (Syrdenopsis) grayii (Wollaston, 1862)
Female adult
Uzbekistan, Kyzyl Kumy, 25km SW Bukhara, 9.6.1994 Klimenko A. leg. (cAA)
Copyright © Alexander Anichtchenko

Sirdenus (Sirdenus) filiformis (Dejean, 1828)
Male adult
Italie, Sardaigne, Sestu 11-05-1982
Copyright © Pascal Deschamps

Phrypeus rickseckeri (Hayward, 1897)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Sinozolus cf. yuae Deuve, 1997
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Chaltenia patagonica Roig-Junent et Cicchino, 2001
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachyta (Tachyta) nana (Gyllenhal, 1810)
Male adult
Far East, Lazovskij Nat. Res., Proselochnaja Riv. near nouth, N slope of Tumannaja Mt., h~350-500 43°00'08"N 134°06'56"E 9.VII.2005 K.Makarov leg.
L=2.7 mm
Copyright © Kirill Makarov

Elaphropus (Tachylopha) spenceri (Sloane, 1896) Paratype
Male adult

Copyright © Museums Victoria

Micratopus sp.
Male adult
Panama, Darien Nat. Park, Rancho Frio, 10.I.2017, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Elaphropus (Nototachys) occidentalis Boyd & Erwin, 2016 Paratype
Male adult
Perú, Madre de Dios, Pakitza
Copyright © Terry Erwin

Elaphropus (Idiotachys) acutifrons Boyd & Erwin, 2016 Holotype
Female adult
Brazil, Santarem
Copyright © Terry Erwin

Meotachys (Scolistichus) riparius Boyd & Erwin, 2016 Paratype
Female adult
Brazil, Amazonas, Rio Solimões, Ilha de Marchantaria
Copyright © Terry Erwin

Nothoderis rufotestacea Hayward, 1900
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachyura (Tachyphanes) patruelis Baehr, 2016 Holotype
Male adult
Turkey, Pr. Siirt Botan Cayi-Tal 500 m, 23.4.1983 Leg. W. Schacht (Zoologische Staatssammlung München).
Copyright © Martin Baehr

Liotachys antennatus Bates, 1871
Female adult
Bolivia: Santa Cruz Prov. Andrez Ibanez Potrerillo de Guenda, Terebinto. / 3-4 October 1994 / leg. P. Bettella / det. R. Davidson. Carnegie Museum of Natural History, Pittsburg, PA, USA
Copyright © Curt Harden

Tachyura (Tachylopha) ovatus (Motschulsky, 1851)
Male adult
Laos central, Ban Gnang, Nam Nimboun riv., 17.958915, 104.746499, 20-24.XI.2025, Anichtchenko A. leg & det. (cAA). 2.2 mm
Copyright © Alexander Anichtchenko

Trechus (Epaphius) nigricornis Motschulsky, 1844
Male adult
det. Makarov K.
Copyright © Kirill Makarov

Shiqianaphaenops majusculus Ueno, 1999c: 47 Holotype
Male adult
Cave Feng Dong, Tangshan, Shiqian County, 27°29'10"N/108°15'23"E, 700 m in altitude, Mingyi Tian & Jingli Cheng leg.Scale bar: 2.0 mm. Image from Mingyi Tian et al., 2016
Copyright © - - -

Omalodera discoidalis discoidalis
Unsexed
Image from Chilean Coleoptera Biodiversity
Copyright © - - -

Eocnides fragilis Ueno, 1989a: 14
Male adult
Bingzhongluo Township (Niwaluo He, just below Nu Jiang Road, 1630 m, N28.05140°/ E98.59319°, 8 October 2002, D.H. Kavanaugh, P.E. Marek and D.Z. Dong collectors. Image from Deuve et al, 2016
Copyright © - - -

Doderotrechus ghilianii ghilianii Fairmaire, 1859a: 26
Male adult
Piemonte, Valle Po, Crissolo (TO), Gr. Buco di Valenza, 5.V.1995 (Collection R. Sciaky)
Copyright © Riccardo Sciaky

Oxytrechus cf. lallemandi Jeannel, 1927
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Typhlotrechus velebiticus (Ganglbauer, 1904c)
Male adult

Copyright © Vittorino Monzini

Canarobius chusyae Machado, 1987c: 324
Male adult

Copyright © Creative Commons

Trechus (Abunetrechus) bipartitus Raffray, 1880
Male adult
Ethiopia, Amhara, Mt. Abuna Yosef, N-slope, 3800–3950 m, 12°07'52"N, 39°11'39"E, 4.III.2019, leg. D. Hauth, J. Schmidt, Yeshitla M., Yitbarek W
Copyright © Joachim Schmidt

Pheggomisetes globiceps breiti Mandl, 1942
Male adult
Bulgaria, Gintsi, cave Dineva, 24.3.1995 B. Gueorguiev leg. (cAA)
Copyright © Alexander Anichtchenko

Thayerella newtoni Baehr, 2016 Holotype
Female adult

Copyright © Martin Baehr

Brachydema victoriae Sloane, 1915 Syntypes
Unsexed

Copyright © Museums Victoria

Merizodus sp.
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Zolus helmsi (Sharp, 1886)
Female adult

Copyright © - - -

Oopterus patulus (Broun, 1881)
Male adult
scale bar 1mm
Copyright © - - -

Oopterus discoideus Larochelle & Larivière, 2017 Paratype
Female adult
“[NEW ZEALAND FD] Wolfe Flat 600m-680m (typed) / Turret Ra [= Range] (typed) / Manapouri Exp. Jan. 70 J. McBurney (typed) / moss (typed) 70/69. Scale bar 1mm
Copyright © - - -

Synteratus ovalis Broun, 1909
Unsexed
Scale bar 1mm
Copyright © - - -

Maungazolus parvulus (Broun, 1903)
Male adult

Copyright © - - -

Percodermus niger Sloane, 1920
Female adult

Copyright © Creative Commons

Merizodus soledadinus (Guerin-Meneville, 1830)
Male adult

Copyright © - - -