Geocharidius phineus Erwin, 1982
Male adult
Guatemala
image from Sokolov I., 2013. A new genus and eight new species of the subtribe Anillina (Carabidae, Trechinae, Bembidiini) from Mexico, with a cladistic analysis and some notes on the evolution of the genus.
Copyright © Igor Sokolov

Anillinus langdoni Sokolov & Carlton, 2004
Male adult
USA-TN, Sevier Co, GSMNP, secondary site, 490 m, ex leaf et log litter, 35°43.157' N, 83°23.309' W, 19.04.2011, J.-S. Park det. I. Sokolov, 2013
Copyright © Roman Dudko

Scotodipnus (Binaghites) armellinii (Ganglbauer, 1900)
Male adult
Veneto Italia
Copyright © Erminio Piva

Rhegmatobius quadricollis (Ehlers, 1883)
Unsexed
Majella, Abruzzo, Italy
Copyright © Vittorino Monzini

Anillinus albrittonorum Sokolov & Schnepp, 2021 Paratype
Female adult
6 mi NW High Springs, Columbia County, Florida. Image from Sokolov & Schnepp, 2021
Copyright © Igor Sokolov

Scotodipnus (Scotodipnus) glaber (Baudi di Selve, 1869)
Male adult
Monte Chiappo, PV, Italia
Copyright © Vittorino Monzini

Microtyphlus saxatilis Magrini, Leo & Fancello, 2004
Male adult
Italia, Sardegna, Perda Longa (NU) Baunei.
Copyright © Augusto Degiovanni

Rhegmatobius bastianinii Magrini & Casale, 2015 Paratype
Male adult
Italia, Lazio (LT) M.te S. Angelo, Formia, 1200m.
Copyright © Augusto Degiovanni

Nesamblyops tararua Sokolov, 2023
Male adult
Judd Ridge, Tararua Range, Wellington, NO
Copyright © Igor Sokolov

Rhegmatobius strictus (Baudi di Selva, 1891)
Male adult
Sardegna, Gennargentu Mte., Spada Mt., 1000, 27.4.2003 A. Degiovanni leg. (cAA).
Copyright © Alexander Anichtchenko

Tasmanitachoides major Baehr, 1990 Holotype
Male adult

Copyright © Museums Victoria

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Image from Sallenave, Erwin, and Roig-Juñent, 2008
Copyright © - - -

Andinodontis muellermotzfeldi Toledano & Erwin, 2010
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidarenas setiventris Negre, 1973
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides sp.
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides erwini Maddison & Porch, 2021 Holotype
Male adult
Australia: Tasmania: River Forth at C136, 41.4712°S 146.1366°E, 126 m, 14.i.2019. DRM 19.012. D.R. Maddison & N.A. Porch. Scale bar 1mm
Copyright © David Maddison

Andinodontis maveetyae Erwin & Maddison, 2010 Holotype
Male adult
Perú: Cusco, Rocotal, 3.2 km E Tres Cruces, 13.1123° S, 71.5834° W, 2082 m, 27 June 2010 (S. MAVEETY) (MHN-UNSAAC: ADP124815)
Copyright © Terry Erwin

Bembidion (Furcacampa) versicolor (Leconte, 1848)
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Amerizus (Amerizus) teles Belousov & Dudko, 2010 Paratype
Male adult
NE Altay Mts., 35-40 km SSE Artybash, Evrechala Mt., 1850-2100 м, tundra, 51°28' N, 87°26' E, 20-22.05.2003, R.Yu.Dudko & I.I.Lubechanskii leg.
Copyright © Igor Belousov

Bembidion (Peryphanes) dalmatinum dalmatinum Dejean, 1831
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Notaphus) contractum Say, 1823
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Bembidion (Leuchydrium) tigrinum Leconte, 1879
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidion (Terminophanes) reibnitzi Schmidt, 2018 Paratype
Male adult
“S TIBET Kyi Chu river / betw. Lhasa and airport / ca. 3650 m, 22.IX.2010 / leg. J. Schmidt” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Peryphanes) milleri carpathicum J. Muller, 1918b: 88
Male adult
W Ukraine, W Podolian Upland, Rogatyn Opillia, Zaluzhzhya vill, 49°24'33"N 24°35'36"E, clay quarry, 30-31.VIII.2018, Panin R leg., L=4.6mm
Copyright © Denis Khrapov

Bembidion (Peryphanes) cf brunnicorne Dejean, 1831
Male adult
W.Ukraine, Transcarpathian, Polonynian Beskids, Yavirnyk Rng, h=600 m, near Ruskyi Mochar vill.,15.V.2021, V. Khrapova leg./R. Panin det., L=4.7 mm
Copyright © Denis Khrapov

Odontium (Bracteon) conicolle Motschulsky, 1844: 273
Female adult
Burjatia, Barguzin depr., Alla riv., 8.8.1997, Anichtchenko A. leg & det.
Copyright © Alexander Anichtchenko

Bembidion (Emphanes) gobiense Jedlicka, 1964
Male adult
Mongolia, salt lake between Ulaanbaatar and Harhorin, 10.66.2006, Anichtchenko A. leg. (cAA). det. L. Toledano, 2026
Copyright © Alexander Anichtchenko

Horologion speokoites Valentine, 1932 Holotype
Male adult
"Arbuckle's Cave Maxwelton, W.Va. July 12 1931 J.M. Valentine / On bit of wet decaying wood, muddy floor of lower level / Type gen. Type sp. / U.S.N.M. Type 44255"
Copyright © Curt Harden

Lissopogonus tonkinensis Zamotajlov & Sciaky, 1996: 40
Male adult
N Vietnam, Lao Cai Province 250 km from Hanoi bearing 310°, SaPa vill env., h~1250 Hoang Lien Son Nat. Park 27.VI-2.VII.1998 leg.A.Napolov
L= 4,8
Copyright © Kirill Makarov

Lissopogonus cf. borneensis Baehr, 2001
Male adult
Malaysia, Borneo, Sarawak, Miri div., Bario 10 km NE, 3 km N Pa Lungan, 7-8.III.2014, R. Cibulskis leg. det. A. Anichtchenko 2015
Copyright © Alexander Anichtchenko

Lissopogonus morvani Deuve, 2009
Female adult
Nepal, Kosi Mure, 2000-2100m, 27.30N 87.16E, 6-12.6.2001. 7 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus Andrewes, 1923
Male adult
Nepal, 12km NE Chainpur, 1400m, 29.36.17N, 81.17.54E, 28.6.2009. 5.8 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus laosensis Deuve, 2009
Male adult
Laos, Nong Khiaw vic, 20.553945, 102.622681, 1.V.2026, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Diplous (Platidius) dolini Zamotajlov, 2005
Female adult
S Primorie, Lazovsky Reserve, Formosova Spring 43°00'06"N 134°05'13"E 10.VII.2005 leg. K.Makarov
L=9.3
Copyright © Kirill Makarov

Deltomerus (Deltomerus) pseudoplatynus pseudoplatynus s.str.
Female adult
NW-Caucasus,
Krasnodar reg., Apsheron Distr., Lagonaki Mt., near Guamka vill., h=1100m. 06-08.VI.2008 D.Fominykh leg.
long - 11mm.
Copyright © Ruslan Panin

Patrobus atrorufus (Stroem, 1768)
Male adult

Copyright © Kirill Makarov

Platidiolus rufus Chaudoir, 1878
Male adult
Russia, Hamar-Daban Mt. rng., Komarinskiy Mt., upper flow of Slyudyanka riv. 15.7.2013, A. Shavrin leg. det. Anichtchenko A. 2018
Copyright © Alexander Anichtchenko

Penetretus nebrioides (Vuillefroy, 1866)
Male adult
SPAIN, Cantabria, 2 km E Puerto de San Glorio, near Vejo riv., 43.072369, -4.733061, 1300 m., 23.07.2019, A. Anichtchenko & R. Sciaky leg.; det. Anichtchenko A. 2019
Copyright © Alexander Anichtchenko

Himalopenetretus franzi (Zamotajlov & Sciaky, 1998) Holotype
Male adult
India, Uttar Pradesh; 17 km SE of Gangotri, Tapovan, 4500 m, 26.VI.1996 (coll. R. Sciaky)
Copyright © Riccardo Sciaky

Grandipenetretus major (Zamotajlov et Wrase, 1997) Paratype
Male adult
China, Shaanxi, Qin Ling Shan, 47 km S Xian, 26-29.VIII.1995 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Parapenetretus (Robustopenetretus) saueri Zamotajlov & Sciaky, 1996 Holotype
Male adult
China – W Sichuan, Emei Shan. 2800 25-31.VI.1992 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Patrobus septentrionis volgensis Zamotajlov & Isaev, 2006
Male adult

Copyright © Igor Solodovnikov

Diplous (Diplous) sibiricus atratus Habu, 1951
Male adult

Copyright © Kirill Makarov

Pogonistes convexicollis Chaudoir, 1871a: 33
Male adult
Volgogrda Area, Pallasovka Distr., N coast of Elton Lake, Lakeside near the mouth of Khara River, Salt-marsh 20.IX.2006 leg. K.Makarov
L=4.0
Copyright © Kirill Makarov

Pogonistes rufoaeneus Dejean, 1828: 19 (Pogonus)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pogonus (Pogonus) orientalis virens Menetries, 1849
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pogonus (Pogonus) minutus Dejean, 1828
Male adult
Pogonus senegalensis Dejean, 1831 det. M.Baehr, 2015. Size: 5,3 mm. Namibia, Etosha National Park, Okaukuejo, 1400 m leg. U.Schmidt, 18.-20.III.1994; Photo: U.Schmidt, 2016
Copyright © Udo Schmidt

Diplochaetus planatus (G.Horn, 1876)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Thalassotrechus barbarae (G.Horn, 1892)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Pogonus (Pogonus) fennelli Hudson, 2000
Female adult
Australia 7.5 mm; det. K. Will, 2020
Copyright © Alexander Anichtchenko

Sirdenus (Sirdenus) filiformis (Dejean, 1828)
Male adult
Italie, Sardaigne, Sestu 11-05-1982
Copyright © Pascal Deschamps

Pogonus (Pogonus) chalceus viridanus Dejean, 1828
Male adult
Spain, Alicante, Urbanova, Antiguas salinas, 12.1.2024 (DUBC)
Copyright © Alexander Anichtchenko

Pogonus (Pogonus) syriacus Chaudoir, 1871
Female adult
Cyprus, Limassol distr., Akrotiri env, 7.4.2012 R. Ambrus leg. (cAA). 7 mm; det. Anichtchenko A. 2026
Copyright © Alexander Anichtchenko

Phrypeus rickseckeri (Hayward, 1897)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Sinozolus cf. yuae Deuve, 1997
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Chaltenia patagonica Roig-Junent et Cicchino, 2001
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Elaphropus (Tachyura) incurvus (Say, 1830)
Female adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Elaphropus (Tachyuropsis) exaratus (Bates, 1873)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Elaphropus (Idiotachys) acutifrons Boyd & Erwin, 2016 Holotype
Female adult
Brazil, Santarem
Copyright © Terry Erwin

Elaphropus (Ammotachys) marchantarius Boyd & Erwin, 2016 Paratype
Male adult
Venezuela, San Carlos de Rio Negro
Copyright © Terry Erwin

Stigmatachys uvea Boyd & Erwin, 2016 Holotype
Female adult
Perú, Loreto, Campamento San Jacinto
Copyright © Terry Erwin

Eotachys (Eotachys) enormis Dostal, 2015 Holotype
Male adult
Madagascar, Andasibe National Park, Lokato, near Andasibe, 10.X.2007, leg. Zd. Mracek (in coll. Dostal, Vienna). Scale bar 1 mm
Copyright © Alexander Dostal

Pericompsus (Pericompsus) laetulus LeConte, 1851
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachyura (Tachylopha) leleupi (Basilewsky In Basilewsky & Straneo, 1950)
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachyura (Tachyura) stevensi (Andrewes, 1925)
Female adult
India, Uttarakhand, 5 km SW Rishikesh, left bank of Ganga riv., 9.4.2012, Anichtchenko A. leg. (cAA). det. M. Baehr
Copyright © Alexander Anichtchenko

Tachyura (Tachylopha) ovatus (Motschulsky, 1851)
Male adult
Laos central, Ban Gnang, Nam Nimboun riv., 17.958915, 104.746499, 20-24.XI.2025, Anichtchenko A. leg & det. (cAA). 2.2 mm
Copyright © Alexander Anichtchenko

Queinnectrechus micrangulus Belousov & Kabak, 2003 Paratype Paratype
Male adult
China, Sichuan, SSW of Shimian, E slope of Mt "4977" W of Lijipin 3000-3900m
5.07.2000 Belousov & Kabak leg.
long.
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko

Trechus (Trechus) championi Jeannel, 1920d: 109
Male adult
India, Uttarakhand, 13-15.04.2012, Uttarkashi distr., N30°57'41.57", E78°41'54.75" left tributary of Bhagirathi riv., 2300 m, Anichtchenko A. leg. det. Anichtchenko A.
Copyright © Alexander Anichtchenko

Perileptus (Perileptus) areolatus areolatus Creutzer, 1799
Male adult

Copyright © Kirill Makarov

Anophthalmus schmidtii insignis J. Muller, 1912: 299
Male adult
W-Slovenia, Dinaric Alps, Trnovo Forest Plateau, Lokve env., 2.V-16.X.2010, M. Egger leg. L=8 mm
Copyright © Ruslan Panin

Puertrechus mengsaensis Belousov & Kabak, 2014 Paratype
Female adult
CH, Yunnan Province, Mt. E Mengsa Town, 23º42´26´´ N / 99º47´26´´ E, H=3045 m, 30.05.2010, Belousov & Kabak leg. Image from IGOR A. BELOUSOV & ILYA I. KABAK, 2014
Copyright © - - -

Pseudanophthalmus ciliaris orlindae Barr, 1959 Paratype
Female adult
Kent. VI.25.1957, Cook Cave, nr. Adairville, Logan Co., TC Barr. Type N 4260: 2 (MZLU)
Copyright © Lund University

Guiaphaenops deuvei Tian, Feng & Wei, 2017 Holotype
Male adult
cave near Yangli Cun, Jiayou Zhen, Lingyun Xian, Baise Shi, Guangxi, 24°28'39"N, 106°37'52"E, 643 m, VII-25-2012, Mingyi Tian, Weixin Liu, Feifei Sun & Haomin Yin leg., in South China Agricultural University, Guangzhou, China (SCAU). Image from Tian, Feng & Wei, 2017
Copyright © - - -

Queinnectrechus (Gaoligongtrechus) balli Deuve and Kavanaugh, 2016 Holotype
Male adult
CHINA, Yunnan Province, Gaoligong Shan, Nujiang Prefecture, Nujiang State Nature Reserve, Dulong/Gongshan Yakou [= Qiqi/Dulong divide] area, 21 airkm w of Gongshan,”/ “N27.69655°/ E98.45389°, 3300-3680m, 16-17 July 2000, Stop#00-24C, D.H. Kavanaugh, C.E. Griswold, Liang H.-B., D. Ubick, & Dong D.-Z. collectors (IOZ). Image from Deuve et al, 2016
Copyright © - - -

Junnanotrechus wrasei Belousov & Kabak, 2014 Holotype Paratype
Male+Female adult
China (N Yunnan), Dali Bai Nat. Aut. Pref., Diancang Shan, pass 43 km NW Dali, 3104 m, 25 ° 59 ´ 33.5 ´´ N / 99 ° 52 ´ 12.5 ´´ E, (pasture & shrubs, litter, moss and mushrooms sifted), 23.08. 2009, (M. Schülke) ( ZISP)
Copyright © - - -

Youtrechus zijunae Tian, Huang & Jia, 2023 Holotype
Male adult
29.73 °N / 114.24 °E, 160 m, a limestone cave called Feixian Dong, Xian’an District, Xianning, Hubei, 2019. VII. 13, Mingyi Tian, Mengzhen Chen, Zijun Ma & Jingli Cheng leg., in SCAU.
Copyright © Mingyi Tian

Sloaneana tasmaniae (Sloane, 1915)
Male adult
Grove, S.J. (2010). Tasmanian Forest Insect Collection web-site
Copyright © Simon Grove

Idacarabus troglodytes Lea, 1910 Syntypes
Male adult

Copyright © Museums Victoria

Brachydema victoriae Sloane, 1915 Syntypes
Unsexed

Copyright © Museums Victoria

Oopterus patulus (Broun, 1881)
Male adult
scale bar 1mm
Copyright © - - -

Oopterus discoideus Larochelle & Larivière, 2017 Paratype
Female adult
“[NEW ZEALAND FD] Wolfe Flat 600m-680m (typed) / Turret Ra [= Range] (typed) / Manapouri Exp. Jan. 70 J. McBurney (typed) / moss (typed) 70/69. Scale bar 1mm
Copyright © - - -

Pseudoopterus otiraensis Larochelle & Larivière, 2017 Paratype
Male adult
NEW ZEALAND WD Arthur’s Pass NP, Kellys Creek x Otira River [= Kelly Creek near Otira River Junction] 4248S 17134E 350m 5.III. 2009 Larochelle and Larivière (typed) / Floodplain Podoc.-brleaf for. [= Podocarp-broadleaf forest] (Fuchsia-Horopito dom. [= dominant]): wet clayish soil. Under large, deeply embedded stones. Scale bar 1mm
Copyright © - - -

Maungazolus parvulus (Broun, 1903)
Male adult

Copyright © - - -

Maungazolus acutus Larochelle & Larivière, 2017 Paratype
Male adult
NEW ZEALAND FD Takahe Valley Murchison Mts [= Mountains] 11 Dec 1972 A.C. Eyles (typed) / Litter (typed) 72/271
Copyright © - - -

Pseudoopterus plicaticollis (Blanchard, 1843)
Male adult
Scale bar 1mm
Copyright © - - -

Merizodus soledadinus (Guerin-Meneville, 1830)
Male adult

Copyright © - - -