Zapotecanillus montanus Sokolov, 2013 Paratype
Male adult
MEXICO, Oaxaca, 52miles N of Oaxaca, Ber.202, sink litter, 17 May 1971 S.B.Peck collector (CMNH)
Scale bar = 0.5mm
Copyright © Igor Sokolov

Nesamblyops sp.
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Pelodiaetus sulcatipennis Jeannel, 1937
Male adult
New Zealand, Otago, Outram
Copyright © Igor Sokolov

Pelodiaetus nunni Sokolov, 2019 Paratype
Male adult
NZ, Canterbury, Ahuriri Scenic Reserve. 1.27 mm
Copyright © Igor Sokolov

Anillus angelae Magrini & Vanni, 1989
Male adult
Arnale cave, Coreno Ausonio, Italia
Copyright © Vittorino Monzini

Anillinus albrittonorum Sokolov & Schnepp, 2021 Paratype
Female adult
6 mi NW High Springs, Columbia County, Florida. Image from Sokolov & Schnepp, 2021
Copyright © Igor Sokolov

Nesamblyops townsendi Sokolov, 2023
Male adult
Tennyson Inlet, Marlborough Sounds, SO
Copyright © Igor Sokolov

Nesamblyops tararua Sokolov, 2023
Male adult
Judd Ridge, Tararua Range, Wellington, NO
Copyright © Igor Sokolov

Medusapyga alsea LaBonte, 2023 Paratype
Female adult
“OREGON, Benton Co., Prairie Peak, 11 km S Alsea, J.R. LaBonte, 9 V 1993, elev. 810 m, 44.2844°N 123.5858°W, under stones". Image from LaBonte & Maddison, 2023
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Rhegmatobius strictus (Baudi di Selva, 1891)
Male adult
Sardegna, Gennargentu Mte., Spada Mt., 1000, 27.4.2003 A. Degiovanni leg. (cAA).
Copyright © Alexander Anichtchenko

Tasmanitachoides major Baehr, 1990 Holotype
Male adult

Copyright © Museums Victoria

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Image from Sallenave, Erwin, and Roig-Juñent, 2008
Copyright © - - -

Andinodontis muellermotzfeldi Toledano & Erwin, 2010
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidarenas setiventris Negre, 1973
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides sp.
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides erwini Maddison & Porch, 2021 Holotype
Male adult
Australia: Tasmania: River Forth at C136, 41.4712°S 146.1366°E, 126 m, 14.i.2019. DRM 19.012. D.R. Maddison & N.A. Porch. Scale bar 1mm
Copyright © David Maddison

Andinodontis maveetyae Erwin & Maddison, 2010 Holotype
Male adult
Perú: Cusco, Rocotal, 3.2 km E Tres Cruces, 13.1123° S, 71.5834° W, 2082 m, 27 June 2010 (S. MAVEETY) (MHN-UNSAAC: ADP124815)
Copyright © Terry Erwin

Bembidion (Phyla) obtusum Audinet-Serville, 1821: 83
Male adult
Henri Goulet Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Bembidion (Philochthus) pallidiveste Carret, 1905
Male adult
Volgogrda Area, Pallasovka Distr., near Elton Vill. 4.V.2009 leg. K.Makarov
L = 3.8
Copyright © Kirill Makarov

Sinechostictus (Sinechostictus) ruficornis Sturm, 1825
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Zeperyphodes) nesophilum Broun, 1886
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
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Bembidion (Trichoplataphus) parconaturaviva Toledano, 2010: 394
Male adult
SE Maritime Prov., Lazovsky dist., ~10 кm SW Sokoltchi vill., 23-27.VI.2010 leg.М. Smirnov. Фотография с сайта www.zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов. det. K. Makarov 2016
Copyright © Maksim Smirnov

Bembidion (Peryphus) bracculatum Bates, 1889b: 212
Female adult
India, Uttarakhand, 5km NW Ghangaria, Valley of Flowers, N30*43'356" E079*85.309 , 3417 m, 06.2011, Anichtchenko A.; 5.5 mm, det. J. Schmidt, 2015
Copyright © Alexander Anichtchenko

Bembidion (Peryphophila) pseudoclarum Schmidt, 2018 Holotype
Male adult
“NEPAL Manaslu Mts. / Marsyangdi Khola vall. / Bhachok Khola 1000m / 13.4.1999 leg. Schmidt” (CSCHM).
Copyright © Joachim Schmidt

Bembidion (Ocydromus) saxatile morsum Netolitzky, 1930
Male adult
Tajikistan, Khoja Obi Garm, 15.9.2011, A. Barsevskis leg. (DUBC). det. L. Toledano, 2019
Copyright © Alexander Anichtchenko

Bembidion (Liocosmius) cachagua Maddison, 2023 Holotype
Male adult
CAS: “U.S.A.: CALIFORNIA: Monterey Co., Hastings Nat. History Reserve. J.M. Linsdale Colln.”, “C.A.S. Accession 29-V-48 Cachagua Creek”, “David R. Maddison DNA3842 DNA Voucher”
Copyright © David Maddison

Bembidion (Armatocillenus) sauteri (Jedlicka, 1958) Holotype
Female adult
Formosa, Sauter // Anping, 1909.X. // Typus [red] // Armatocillenus sauteri n.sp. [handwritten], det. ING. JEDLIČKA [pink] (HNHM).
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Horologion speokoites Valentine, 1932 Holotype
Male adult
"Arbuckle's Cave Maxwelton, W.Va. July 12 1931 J.M. Valentine / On bit of wet decaying wood, muddy floor of lower level / Type gen. Type sp. / U.S.N.M. Type 44255"
Copyright © Curt Harden

Lissopogonus tonkinensis Zamotajlov & Sciaky, 1996: 40
Male adult
N Vietnam, Lao Cai Province 250 km from Hanoi bearing 310°, SaPa vill env., h~1250 Hoang Lien Son Nat. Park 27.VI-2.VII.1998 leg.A.Napolov
L= 4,8
Copyright © Kirill Makarov

Lissopogonus cf. borneensis Baehr, 2001
Male adult
Malaysia, Borneo, Sarawak, Miri div., Bario 10 km NE, 3 km N Pa Lungan, 7-8.III.2014, R. Cibulskis leg. det. A. Anichtchenko 2015
Copyright © Alexander Anichtchenko

Lissopogonus morvani Deuve, 2009
Female adult
Nepal, Kosi Mure, 2000-2100m, 27.30N 87.16E, 6-12.6.2001. 7 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus Andrewes, 1923
Male adult
Nepal, 12km NE Chainpur, 1400m, 29.36.17N, 81.17.54E, 28.6.2009. 5.8 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus laosensis Deuve, 2009
Male adult
Laos, Nong Khiaw vic, 20.553945, 102.622681, 1.V.2026, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Patrobus cinctus Motschulsky, 1860
Female adult
Jewish Aut. Reg., Obluchie Distr., 3-5 km upper Radde Vill. on Lagar Riv., forest 24.07.2004 I.Melnik leg.
L=9.5 mm
Copyright © Kirill Makarov

Diplous (Platidius) dolini Zamotajlov, 2005
Female adult
S Primorie, Lazovsky Reserve, Formosova Spring 43°00'06"N 134°05'13"E 10.VII.2005 leg. K.Makarov
L=9.3
Copyright © Kirill Makarov

Archipatrobus flavipes (Motschulsky, 1864)
Male adult
Collection ground Kobe city, Hyogo Pref. 33645<br /> (C) Working Group for Biological Indicator Ground Beetles Database, Japan<br /> Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Patrobus atrorufus (Stroem, 1768)
Male adult

Copyright © Kirill Makarov

Deltomerus (Deltomerus) jelineki Zamotajlov, 2001
Female adult
Iran Alburz prov Shemshak 2750m 12.06.17
Copyright © Jan Muilwijk

Platidiolus cf. borealis Zamotajlov, 2001
Female adult
Russia, Khabarovsk reg., Verhnebureinski distr. Bureya river valley, Ussomah river, 22.7.2006 (anal sternite with 3 pairs of setae). det. Anichtchenko A. 2018
Copyright © Alexander Anichtchenko

Grandipenetretus major (Zamotajlov et Wrase, 1997) Paratype
Male adult
China, Shaanxi, Qin Ling Shan, 47 km S Xian, 26-29.VIII.1995 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Parapenetretus (Robustopenetretus) saueri Zamotajlov & Sciaky, 1996 Holotype
Male adult
China – W Sichuan, Emei Shan. 2800 25-31.VI.1992 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Himalopenetretus burangensis Yan, Shi & Liang, 2020 Holotype
Male adult
“Xizang, Ngari Pref., Burang County, Burang Town. N of Tinkar pass, alpine meadow. 30.2085N, 81.0661E, 4800m.”; “under stone, 2019. VII. 15, Shi HL lgt., Exp. BJFU 2019”; “DNA sample series number SHL2019--Xiz006”; “HOLOTYPE♂ Himalopenetretus burangensis sp. nov., des. YAN & SHI, 2020” [red label]. (IZAS). Image from Yan, Shi & Liang, 2020
Copyright © - - -

Penetretus andalusicus (Reitter, 1897)
Male adult
Spain, Sierra Nevada, Lagunillos de la Virgen, 2850m, 18.5.2005, A. Anichtchenko leg. & det (cAA)
Copyright © Alexander Anichtchenko

Cardiaderus chloroticus (Fischer von Waldheim, 1823)
Male adult
Volgogrda Area, Pallasovka Distr., N coast of Elton Lake, right bank of Khara River, 3 km upstream of the mouth, Salina on fl oodplain terrace 20.IX.2006 leg. K.Makarov
L=7.0
Copyright © Kirill Makarov

Pogonus (Pogonoidius) punctulatus Dejean, 1828
Male adult
Volgograd Reg., Pallas Distr., Elton Vill., on UV-light 16-18.VII.2006 K.Makarov leg.
L=6.4 mm
Copyright © Kirill Makarov

Diplochaetus planatus (G.Horn, 1876)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Thalassotrechus barbarae (G.Horn, 1892)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Iranopogonus freyellus Jedlicka, 1959 Holotype
Male adult
Iran, Sistan va Baluchestan Province, Chabahar env., IX.1937, leg. Band (NHMB). Image of Matthias Borer (NHMB) from Azadbakhsh, 2019
Copyright © - - -

Pogonopsis pallida Bedel, 1898 Holotype
Male adult
Tunisia (MNHN) Image from Azadbakhsh, 2019
Copyright © - - -

Sirdenus (Syrdenopsis) grayii (Wollaston, 1862)
Female adult
Uzbekistan, Kyzyl Kumy, 25km SW Bukhara, 9.6.1994 Klimenko A. leg. (cAA)
Copyright © Alexander Anichtchenko

Pogonus (Pogonus) chalceus viridanus Dejean, 1828
Male adult
Spain, Alicante, Urbanova, Antiguas salinas, 12.1.2024 (DUBC)
Copyright © Alexander Anichtchenko

Pogonus (Pogonus) biroi Csiki, 1907
Male adult
India, Goa, Palm Beach, 20-23.4.2013 (cAA). 5.5 mm; det. Anichtchenko A. 2026
Copyright © Alexander Anichtchenko

Pogonus (Pogonus) syriacus Chaudoir, 1871
Female adult
Cyprus, Limassol distr., Akrotiri env, 7.4.2012 R. Ambrus leg. (cAA). 7 mm; det. Anichtchenko A. 2026
Copyright © Alexander Anichtchenko

Phrypeus rickseckeri (Hayward, 1897)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Sinozolus cf. yuae Deuve, 1997
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Chaltenia patagonica Roig-Junent et Cicchino, 2001
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Elaphropus (Elaphropus) latissimus (Motschulsky, 1851)
Female adult
Kunashiri Isl., Calder of Golovnina Volcano, S coast od Goryachee Lake 43°52'19"N 145°29'43"E 12.VIII.2008 leg.K.Makarov, A.Zaitzev
2.0 mm
Copyright © Kirill Makarov

Lymnastis galilaeus Piochard de la Brullerie, 1876
Male adult

Copyright © Gianni Allegro

Elaphropus (Tachylopha) spenceri (Sloane, 1896) Paratype
Male adult

Copyright © Museums Victoria

Setitachys macrops Baehr, 2016 Paratype
Female adult
Qld. Bentinck Is. Rokoti, 16 June 1963. P. Aitken, N. B. Tindale
Copyright © Martin Baehr

Tachyxysta howdenorum Boyd & Erwin, 2016 Holotype
Male adult
México, Chiapas, El Aguacero.
Copyright © Terry Erwin

Meotachys (Scolistichus) riparius Boyd & Erwin, 2016 Paratype
Female adult
Brazil, Amazonas, Rio Solimões, Ilha de Marchantaria
Copyright © Terry Erwin

Meotachys (Hylotachys) ballorum Boyd & Erwin, 2016 Paratype
Female adult
Brazil, Amazonas, Tapurucuara
Copyright © Terry Erwin

Tachys (incertae) mulwalensis Sloane, 1900
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Eotachys (Jeannelys) euryodes (Bates, 1892)
Unsexed
Thailand, Prov. Phang Nga, Khao Lak, Similana Resort leg. U.Schmidt, 14.XI.2007; det. M.Baehr, 2008; Photo: U.Schmidt, 2021
Copyright © Udo Schmidt

Elaphropus (Elaphropus) globulus (Dejean, 1831)
Male adult
Spain, Granada, rio Darro, barranco de Teatino, 20.V.2005, Anichtchenko A. leg & det. (cAA). 1.8 mm
Copyright © Alexander Anichtchenko

Trechus (Epaphius) nigricornis Motschulsky, 1844
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Duvalius (Duvalius) gestroi gestroi Dodero, 1900
Male adult
photo by Mauro Agosti
Copyright © - - -

Shenaphaenops humeralis Ueno, 1999j: 630 Holotype
Male adult
Cave Shendong Migong (Shendong in the original description), Muqiao, Laoyingshan, Shuicheng County, northwestern Guizhou, 26°35'15"N/ 104°59'47"E, 1910 m in altitude, VIII-22-2014, Mingyi Tian leg. Scale bar: 2.0 mm. Image from Mingyi Tian et al., 2016
Copyright © - - -

Kenodactylus audouini Guerin-Meneville, 1830
Male adult
Argentina, PN Tierra del Fuego, Bahia Lapataia, 12-15.I.2003, T. Ueno; det. T. Ueno, 2007
Copyright © Alexander Anichtchenko

Doderotrechus ghilianii ghilianii Fairmaire, 1859a: 26
Male adult
Piemonte, Valle Po, Crissolo (TO), Gr. Buco di Valenza, 5.V.1995 (Collection R. Sciaky)
Copyright © Riccardo Sciaky

Trechobembix baldiensis baldiensis
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Wanoblemus huangshanicus Tian & Li, 2020 Holotype
Male adult
cave Shenxian Dong, Qiaoshan, Xinming, Huangshan, Anhui, 30°23'9.55"N, 118°14'7.66"E, 366 m in altitude, 2016-IV-22, leg. Wenbo Li (SCAU). Image from Fang et al, 2020
Copyright © - - -

Guizhaphaenopsodes solidior Tian & X. Huang, 2021 Holotype
Male adult
cave Tangle Dong, suburb of Jishou, Xiangxi Tujia & Miao Autonomous Prefecture, west Hunan Province, 28°18’21”N /109°39’15”E, 270 m in altitude, 2018-I-15, leg. Xinglong Huang (SCAU).
Copyright © Mingyi Tian

Uenotrechus gejianbangi Tian et Wei, 2017 Holotype
Male adult
( SCAU), Cave Ji Dong (also called Shuiku Dong ), Neiwen Tun , Sancai Cun , Shuiyuan Zhen , Huanjiang Maonan Autonomous County, Guangxi Zhuang Autonomous Region, 24°49.430'N / 108°05.504'E, 290 m, XII-02-2016, leg. Arnaud Faille, Jujian Chen and Sunbin Huang.
Copyright © Mingyi Tian

Canarobius chusyae Machado, 1987c: 324
Male adult

Copyright © Creative Commons

Sloaneana tasmaniae (Sloane, 1915)
Male adult
Grove, S.J. (2010). Tasmanian Forest Insect Collection web-site
Copyright © Simon Grove

Idacarabus troglodytes Lea, 1910 Syntypes
Male adult

Copyright © Museums Victoria

Zolus helmsi (Sharp, 1886)
Female adult

Copyright © - - -

Zolus subopacus Broun, 1915
Male adult

Copyright © - - -

Zolus wongi Larochelle & Larivière, 2017 Paratype
Female adult
[NEW ZEALAND BR] Mt Dewar 1000m J.C. Watt (typed) / under bark [of] logs (typed) / Paparoa Range Dec. 1969 (NZAC). Scale bar 1mm
Copyright © - - -

Oopterus discoideus Larochelle & Larivière, 2017 Paratype
Female adult
“[NEW ZEALAND FD] Wolfe Flat 600m-680m (typed) / Turret Ra [= Range] (typed) / Manapouri Exp. Jan. 70 J. McBurney (typed) / moss (typed) 70/69. Scale bar 1mm
Copyright © - - -

Pseudoopterus otiraensis Larochelle & Larivière, 2017 Paratype
Male adult
NEW ZEALAND WD Arthur’s Pass NP, Kellys Creek x Otira River [= Kelly Creek near Otira River Junction] 4248S 17134E 350m 5.III. 2009 Larochelle and Larivière (typed) / Floodplain Podoc.-brleaf for. [= Podocarp-broadleaf forest] (Fuchsia-Horopito dom. [= dominant]): wet clayish soil. Under large, deeply embedded stones. Scale bar 1mm
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Maungazolus acutus Larochelle & Larivière, 2017 Paratype
Male adult
NEW ZEALAND FD Takahe Valley Murchison Mts [= Mountains] 11 Dec 1972 A.C. Eyles (typed) / Litter (typed) 72/271
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Pseudoopterus latipennis (Broun, 1903)
Male adult
Scale bar 1mm
Copyright © - - -

Percodermus niger Sloane, 1920
Female adult

Copyright © Creative Commons