Trichotichnus (Trichotichnus) angustitarsis Kataev, Hongbin & Wrase, 2022 Paratype
Male adult
Baoshan Pref. , mountain range 25 km S Tengchong, 1900 m, 24°48'28"N 98°32'03"E, devasted primary decid. forest, litter, sifted, 2.VI.2007, D.W. Wrase leg.; Image from Kataev et al, 2022
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Trichotichnus (Trichotichnus) anthracinus Landin, 1955
Male adult
Yunnan. Image from Kataev et al, 2022
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Trichotichnus (Trichotichnus) anthracinus Landin, 1955 Holotype
Female adult
N. E. Burma Kambaiti , 7000 ft, 4–8/6 1934, R. Malaise (NRM). Image from Kataev et al, 2022
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Trichotichnus (Trichotichnus) aquilo Andrewes, 1930
Male adult
Left: Ganesh Himal, Nepal. Right: Jinping, Yunnan. Scale bar = 1 mm. Image from Kataev & Schmidt, 2018
Copyright © Joachim Schmidt

Trichotichnus (Trichotichnus) congruus Motschulsky, 1866a: 164
Male adult
Collection ground Ebetsu　city, Hokkaido Pref. 39970
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Trichotichnus (Trichotichnus) congruus Motschulsky, 1866a: 164
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 39970
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Trichotichnus (Trichotichnus) coruscus (as sachalinensis) Holotype
Male adult
Konuma, SAGHALIEN, VII. 1938, T. Takahashi. COL-143
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) coruscus (Tschitscherine, 1895) Holotype
Male adult
Konuma, SAGHALIEN, VII. 1938, T. Takahashi. COL-143
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) coruscus (Tschitscherine, 1895) Holotype
Male adult
Konuma, SAGHALIEN, VII. 1938, T. Takahashi. COL-143
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) coruscus (Tschitscherine, 1895) Holotype
Male adult
Konuma, SAGHALIEN, VII. 1938, T. Takahashi. COL-143
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) coruscus (Tschitscherine, 1895) Holotype
Male adult
Konuma, SAGHALIEN, VII. 1938, T. Takahashi. COL-143
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) cyrtops (Tschitscherine, 1906)
Male adult
Yunnan. Image from Kataev et al, 2022
Copyright © - - -

Trichotichnus (Trichotichnus) dichrous (Dejean, 1829)
Male adult
Great Smoky Mountains National Park
photo I. M. Sokolov
Copyright © Igor Sokolov

Trichotichnus (Trichotichnus) fedorenkoi Kataev, Hongbin & Wrase, 2022 Paratype
Male adult
Tam-Dao. Image from Kataev et al, 2022
Copyright © - - -

Trichotichnus (Trichotichnus) doiinthanonensis N.Ito, 1997
Male adult
YUNNAN, Gongshan, Dulongjiang , 0.5 km WSW Maku, trail, 27.68310°N 98.30038°E, 1845 m, day, 29.VIII.2006, D. Kavanaugh leg. ( IOZ). Image from Kataev et al, 2022
Copyright © - - -

Trichotichnus (Trichotichnus) emarginatus Andrewes, 1930
Male adult
Annapurna
Copyright © Joachim Schmidt

Trichotichnus (Trichotichnus) fukuharai Habu, 1957 Holotype
Male adult
Tokura, Oze, GUNMA, 04. IX. 1952, N. Fukuhara. COL-137
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) fukuharai Habu, 1957 Holotype
Male adult
Tokura, Oze, GUNMA, 04. IX. 1952, N. Fukuhara. COL-137
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) fukuharai Habu, 1957 Holotype
Male adult
Tokura, Oze, GUNMA, 04. IX. 1952, N. Fukuhara. COL-137
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) fukuharai Habu, 1957
Male adult
Yunnan. Image from Kataev et al, 2022
Copyright © - - -

Trichotichnus (Trichotichnus) glabellus Andrewes, 1930
Male adult
Annapurna
Copyright © Joachim Schmidt

Trichotichnus (Trichotichnus) hamulipenis nepalensis Kataev & Hongbin & Wrase, 2022 Holotype
Male adult
NEPAL P: Mahakali / D: Darchula 1 km WSW Batar, Chamliya Khola , 2000 m, 29°52′ 01′′N 80°53′55′′E 11.VI.2005 leg. A. Weigel river-side ( NME)
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Trichotichnus (Trichotichnus) hamulipenis hamulipenis Paratype
Male adult
Gongshan, Dulongjiang, Miliwang , above Bapo , 27.72383°N 98.361°E, 1956 m, 31.X.2004 night, H.B. Liang leg.
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Trichotichnus (Trichotichnus) imafukui Habu, 1961 Holotype
Male adult
Mt. Nishikomaga-dake, NAGANO, 18. VII. 1958, S. Imafuku. COL-142
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) imafukui Habu, 1961 Holotype
Male adult
Mt. Nishikomaga-dake, NAGANO, 18. VII. 1958, S. Imafuku. COL-142
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) imafukui Habu, 1961d: 151 EA: JA Holotype
Male adult
Mt. Nishikomaga-dake, NAGANO, 18. VII. 1958, S. Imafuku. COL-142
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) imafukui Habu, 1961 Holotype
Male adult
Mt. Nishikomaga-dake, NAGANO, 18. VII. 1958, S. Imafuku. COL-142
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) kavanaughi Kataev, Hongbin & Wrase, 2022 Paratype
Male adult
Gongshan, Dulongjiang, Kongdang , light trap, 27.87755°N 98.33612°E, 1511 m, 27.VIII.2006 night, Liu Ye leg.
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Trichotichnus (Trichotichnus) kobayashie Habu, 1957 Holotype
Male adult
Kamikoura, Aogashima Is., TOKYO, 12. XI. 1954, S. Kobayashi. COL-144
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) kobayashie Habu, 1957b: 66 EA: JA Holotype
Male adult
Kamikoura, Aogashima Is., TOKYO, 12. XI. 1954, S. Kobayashi. COL-144
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) kobayashie Habu, 1957b: 66 EA: JA Holotype
Male adult
Kamikoura, Aogashima Is., TOKYO, 12. XI. 1954, S. Kobayashi. COL-144
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) kobayashie Habu, 1957b: 66 EA: JA Holotype
Male adult
Kamikoura, Aogashima Is., TOKYO, 12. XI. 1954, S. Kobayashi. COL-144
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) kurosai Habu, 1973 Holotype
Male adult
Magura, 04. IX. 1941, K. Kosuge. COL-147
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) kurosai Habu, 1973 Holotype
Male adult
Magura, 04. IX. 1941, K. Kosuge. COL-147
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) kurosai Habu, 1973 Holotype
Male adult
Magura, 04. IX. 1941, K. Kosuge. COL-147
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) kurosai Habu, 1973 Holotype
Male adult
Magura, 04. IX. 1941, K. Kosuge. COL-147
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) kurosai Habu, 1973 Holotype
Male adult
Magura, 04. IX. 1941, K. Kosuge. COL-147
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) laevicollis (as carpathicus Schauberger, 1936)
Male adult
W Ukraine, NE Carpathians Mts.,
Transcarpathian reg., Rakhov Distr., Chornogora range, S slope of Petros Mt., h~1650m, 28.VI.2009, Panin R.leg.
L=8 mm
Copyright © Ruslan Panin

Trichotichnus (Trichotichnus) laevicollis (Duftschmid, 1812)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Trichotichnus (Trichotichnus) lewisi Schauberger, 1936: 9
Male adult
Collection ground Kikonai cho, Hokkaido Pref. 34925
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Trichotichnus (Trichotichnus) lewisi Schauberger, 1936: 9
Endophallus
Collection ground Kikonai cho, Hokkaido Pref. 34925
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Trichotichnus (Trichotichnus) liparus Andrewes, 1926
Male adult
Left: Annapurna, below Thulo Bugin; Right: Manaslu, S Bara Pokhari. Scale bar 1 mm. Image from Kataev & Schmidt, 2018
Copyright © Joachim Schmidt

Trichotichnus (Trichotichnus) longitarsis A. Morawitz, 1863: 65
Male adult
Collection ground Ebetsu　city, Hokkaido Pref. 40068
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Trichotichnus (Trichotichnus) longitarsis A. Morawitz, 1863: 65
Endophallus
Collection ground Ebetsu　city, Hokkaido Pref. 40068
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Trichotichnus (Trichotichnus) longitarsis A. Morawitz, 1863: 65
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Trichotichnus (Trichotichnus) miser (Tschitscherine, 1897)
Male adult
Daegu-si, Republic of Korea. Scale bar: 2 mm.
Copyright © Dooyoung Kim

Trichotichnus (Trichotichnus) miser (Tschitscherine, 1897)
Endophallus
Scale bar: 0.5 mm.
Copyright © Dooyoung Kim

Trichotichnus (Trichotichnus) miwai continentalis Kataev & Hongbin & Wrase, 2022 Paratype
Male adult
Wuliangshan Mt. Range , 24°45'02"N 100°30'24"E, 2270 m, 12.VI.2011, I. Belousov, I. Kabak, A. Korolev leg.
Copyright © - - -

Trichotichnus (Trichotichnus) nishioi Habu, 1961
Male adult
Far East, Lazo Distr., midd. course of Socolovka Riv. 42°58'35"N 133°59'29E 28.VII.2005 K.Makarov leg.
L=8.5 mm
Copyright © Kirill Makarov

Trichotichnus (Trichotichnus) nishioi Habu, 1961 Holotype
Male adult
Kotoni, HOKKAIDO, 03. V. 1955, Y. Nishio. COL-138
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) nishioi Habu, 1961 Holotype
Male adult
Kotoni, HOKKAIDO, 03. V. 1955, Y. Nishio. COL-138
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) noctuabundus Habu, 1954b: 56
Male adult
Collection ground Siota cho, Saga pref, 34869
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Trichotichnus (Trichotichnus) noctuabundus Habu, 1954b: 56
Male adult
Collection ground Siota cho, Saga pref, 34869
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Trichotichnus (Trichotichnus) noctuabundus Habu, 1954
Male adult
Vietnam. Image from Kataev et al, 2022
Copyright © - - -

Trichotichnus (Trichotichnus) schmidti Kataev, Hongbin & Wrase, 2022 Paratype
Male adult
LEFT & middle: Baoshan Co., Gaoligong Mt. R., Nankang Pass , 24.82647°N 98.76787°E, 2100–2200 m, along road, 4.X.2012, B. Kataev leg. (ZIN); RIGHT: Baoshan Pref. , mountain range 25 km S Tengchong, 1900 m, 24°48'21"N 98°32'05"E, cleft with devasted primary forest, litter, mushrooms sifted, 30.VIII.2009, M. Schülke leg. Image from Kataev et al, 2022
Copyright © - - -

Trichotichnus (Trichotichnus) shibatai Habu, 1973: 403 Holotype
Male adult
Iwawaki, OSAKA, 18. VI. 1966, H. Nomura. COL-145
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) shibatai Habu, 1973: 403 Holotype
Male adult
Iwawaki, OSAKA, 18. VI. 1966, H. Nomura. COL-145
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) shibatai Habu, 1973: 403 Holotype
Male adult
Iwawaki, OSAKA, 18. VI. 1966, H. Nomura. COL-145
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) shibatai Habu, 1973: 403 Holotype
Male adult
Iwawaki, OSAKA, 18. VI. 1966, H. Nomura. COL-145
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) shibatai Habu, 1973: 403 Holotype
Male adult
Iwawaki, OSAKA, 18. VI. 1966, H. Nomura. COL-145
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) subangulatus Ito, 2021
Male adult
Baoshan Pref., Gaoligong Shan, 65 km NNE Tengchong, 1750 m, 25°35'20"N 98°40'21"E, sec. mixed forest, overgrown stone debris, litter and moss sifted, 27.VIII.2009, D.W. Wrase leg.
Copyright © - - -

Trichotichnus (Trichotichnus) subrobustus Kataev, Hongbin & Wrase, 2022 Holotype
Male adult
CHINA, Yunnan Provin., Gongshan, Dulongjiang Bapo, Mabiluo, riverside 27.76208°N, 98.34567°E ”, “ 1503 m, 2004.10.27 day Liang Hongbin collector California Academy & IOZ., Chinese. Acad. Sci. ( IOZ). Image from Kataev et al, 2022
Copyright © - - -

Trichotichnus (Trichotichnus) teradai Habu, 1980 Holotype
Male adult
Tzuchung, TAIWAN, 25. V. 1977, K. Terada. COL-141
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) teradai Habu, 1980 Holotype
Male adult
Tzuchung, TAIWAN, 25. V. 1977, K. Terada. COL-141
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) teradai Habu, 1980 Holotype
Male adult
Tzuchung, TAIWAN, 25. V. 1977, K. Terada. COL-141
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) teradai Habu, 1980 Holotype
Male adult
Tzuchung, TAIWAN, 25. V. 1977, K. Terada. COL-141
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) teradai Habu, 1980 Holotype
Male adult
Tzuchung, TAIWAN, 25. V. 1977, K. Terada. COL-141
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) vespertinus Habu, 1954b: 57
Male adult
Collection ground Ashikaga city,Tochigi Pref.Ashikaga～shi 36079
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Trichotichnus (Trichotichnus) vespertinus Habu, 1954b: 57
Endophallus
Collection ground Ashikaga city,Tochigi Pref.Ashikaga～shi 36079
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Trichotichnus (Trichotichnus) vespertinus Habu, 1954b: 57
Male adult
Daegu-si, Republic of Korea. Scale bar: 3 mm.
Copyright © Dooyoung Kim

Trichotichnus (Trichotichnus) vespertinus Habu, 1954b: 57
Endophallus
Scale bar: 0.6 mm.
Copyright © Dooyoung Kim

Trichotichnus (Trichotichnus) vulpeculus Say, 1823
Male adult
Great Smoky Mountains National Park
photo I. M. Sokolov
Copyright © Igor Sokolov

Trichotichnus (Trichotichnus) yukihikoi Habu, 1961 Holotype
Male adult
Hakone, Kojiri, KANAGAWA, 03. X. 1958, Y. Hirano. COL-146
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) yukihikoi Habu, 1961 Holotype
Male adult
Hakone, Kojiri, KANAGAWA, 03. X. 1958, Y. Hirano. COL-146
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) yukihikoi Habu, 1961 Holotype
Male adult
Hakone, Kojiri, KANAGAWA, 03. X. 1958, Y. Hirano. COL-146
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) yukihikoi Habu, 1961 Holotype
Male adult
Hakone, Kojiri, KANAGAWA, 03. X. 1958, Y. Hirano. COL-146
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Trichotichnus (Trichotichnus) yukihikoi Habu, 1961 Holotype
Male adult
Hakone, Kojiri, KANAGAWA, 03. X. 1958, Y. Hirano. COL-146
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake