Calosoma (Callisthenes) anthrax Semenov, 1900 Holotype
Male adult
collection of ZIN
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) grumi Semenov, 1900 Holotype
Female adult
collection ZIN
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) anthrax Semenov, 1900
Female adult
Inner Mongolia (China), Dalai-Nor Lake, VI.1957, Svetlovidova leg.
Copyright © Dmitri Obydov

Calosoma (Callisthenes) anthrax Semenov, 1900
Female adult
E-Mongolia: Bujur-Noor Lake, 29.VII.1976, Pechocki lg. (TOPOTYPUS)
Copyright © Igor Solodovnikov

Calosoma (Callisthenes) breuningi Mandl, 1954
Female adult
E Kazakhstan, S Alakol lake, Koktuma vill., 1-10.5.1999 Nekroshevich S. leg. female, 22 mm; det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) breuningi Mandl, 1954
Male adult
E.Kazakhstan, Alakol lake, Koktuma vill., 07.V.1999
Copyright © Anton Menstschikov

Calosoma (Callisthenes) karelini breuningi Mandl, 1954 Holotype
Male adult
"Sergiopol", "Cal. Breuningi, Typus, m., det. Ing. K. Mandl"
Copyright © Dmitri Obydov

Calosoma (Callisthenes) karelini vladimiri Obydov, 1998
Male adult
Kazakhstan, Taldy-Kurgan env., Kyzyl-Zhar Mts., 400 m, 15.IV.2000, M. Danilevsky leg.
photo from zin.ru
Copyright © Dmitri Obydov

Calosoma (Callisthenes) breuningi breuningi Holotype
Male adult
«Sergiopol», «Panderi v. n. aenea (St. blau)», «Karel. subsp. det. Ganglb.», «Callisthenes Karelini, Fisch.», «Typus», «Cal. Breuningi m. Typus, det. Ing. K. Mandl», «Breuningi m.» (NHMW).
Copyright © Ilya Kabak

Calosoma (Callisthenes) breuningi iliense Kabak, 2022 Paratype
Male adult
W of Kerbulak, 44°03ʹ03ʹʹ N, 77°01ʹ12ʹʹ E, 640 m, 25.IV.2017, I. I. Kabak leg.
Copyright © Ilya Kabak

Calosoma (Callisthenes) breuningi iliense Kabak, 2022 Paratype
Female adult
«SE Kazakhstan, right bank of Ili River, near Malaysary, 30.IV–1.V.1985, G. V. Nikolaev, A. S. Badenko leg.» (ca. 44°17ʹ30ʹʹ N, 76°57ʹ16ʹʹ E, 510 m),
Copyright © Ilya Kabak

Calosoma (Callisthenes) breuningi iliense Kabak, 2022 Paratype
Male adult
«SE Kazakhstan, right bank of Ili River, near Malaysary, 30.IV–1.V.1985, G. V. Nikolaev, A. S. Badenko leg.» (ca. 44°17ʹ30ʹʹ N, 76°57ʹ16ʹʹ E, 510 m),
Copyright © Ilya Kabak

Calosoma (Callisthenes) breuningi breuningi
Female adult
SE Kazakhstan, Almaty region, Ak-Eshki lake, 42 km NNE Taldy-Kurgan, H = 456 m, sands, 45°21ʹ44ʺ N, 78°37ʹ04ʺ E, 21.IV.2017, S. V. Kolov leg.
Copyright © Ilya Kabak

Calosoma (Callisthenes) breuningi breuningi
Male adult
Сев. предг. Джунгарского Алатау у с. Уч-Арал Ала-Кульск р-на, 7.V.1952, Проценко» (ca. 46°09ʹ25ʺ N, 80°58ʹ55ʺ E, 410 m), (ZIN)
Copyright © Ilya Kabak

Calosoma (Callisthenes) breviusculus Mannerheim, 1830
Male adult
Armenia, near Ashtarik<br /> 15.6.1997<br /> male, 22 mm<br /> photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) substriatus Motschulsky, 1859
Female adult
Azerbaijan, Talysh, Gasmalian, 20.V.1992, V. Siniaev leg.
photo from zin.ru
Copyright © Dmitri Obydov

Calosoma (Callisthenes) substriatus Motschulsky, 1859 Lectotype
Male adult
"Callisthenes substriatus Motsch. Mt. Saganlug Persia"
Copyright © Dmitri Obydov

Calosoma (Callisthenes) substriatus Motschulsky, 1859
Male adult
Azerbaijan, Talysh, Gasmalian, 20.V.1992, V. Siniaev leg.
photo from zin.ru
Copyright © Dmitri Obydov

Calosoma (Callisthenes) pumicatus pumicatus s.str. Holotype
Female adult
"Plateau Persan Occid. de Zendjan a Ardebil, alt. moy 300 m (J. de Morgan)".
Copyright © Dmitri Obydov

Calosoma (Callisthenes) pumicatus pumicatus (as zarudnianum) Holotype
Male adult
Holotypus of Calosoma zarudnianum:
"Calosoma zarudnianum m, sp. n., A. Semenov-Tian-Shansky det. 21", "W. Pers., 13.V.04, Zarudnyi leg.".
Copyright © Dmitri Obydov

Calosoma (Callisthenes) pumicatus schelkovnikovi Zaitzev, 1918 Lectotype
Male adult
"ms. Sehend 11. 600' 18.V.16", "Mus Caucas. 100-16. Exp. Urmiana", "Calosoma schelkovnikovi Zaitz. (type)", "Received personally from F.A. Zaitzev in 1920".
Copyright © Dmitri Obydov

Calosoma (Callisthenes) breviusculus Mannerheim, 1830
Male adult
Turkey, Buglan; 21mm
Copyright © Aleš Sedláček

Calosoma (Callisthenes) breviusculus Mannerheim, 1830
Female adult
ARMENIE, Aragatsotn route du lac Kari, 28-06-2014, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Calosoma (Callisthenes) breviusculus pumicatus Lapouge, 1907
Female adult
Iran: Ardabil prov. Noor Lake (TOPOTYPUS)
Copyright © Igor Solodovnikov

Calosoma (Callisthenes) elegans karagaicus Lapouge, 1924
Male adult
KIRGHIZSTAN, Inner Tian-Shan Mts., near Naryn tn., Naryn riv. Valley, 18-20.V.1999, Leg. Irtlatch A.
Copyright © Anton Menstschikov

Calosoma (Callisthenes) elegans danilevskii Obydov, 1997d Paratype
Male adult
Kazakhstan, Kaskelen Riv. Val., 1500 m, 9.VI.1989, M. Danilevsky leg.<br /> from zin.ru
Copyright © Dmitri Obydov

Calosoma (Callisthenes) elegans danilevskii Obydov, 1997d Paratype
Female adult
Kazakhstan, Kaskelen Riv. Val., 1500 m, 9.VI.1989, M. Danilevsky leg.
Copyright © Dmitri Obydov

Calosoma (Callisthenes) elegans ballioni Solsky, 1874
Male adult
Kazakhstan, Karaoi, 550 m, 7.5.1991, 24.IV.1994, M. Danilevsky leg.
Copyright © Dmitri Obydov

Calosoma (Callisthenes) elegans valentinae Obydov, 1997 Paratype
Female adult
Kazakhstan, Kapchagai, left bank of Ili Riv., 25. IV.1994, M. Danilevsky leg.
Copyright © Dmitri Obydov

Calosoma (Callisthenes) elegans saryakensis Kabak, 1992 Holotype
Male adult
Koksengir mt. environs, Karaganda dist., 21.V.1958, Tobias leg.
Copyright © Dmitri Obydov

Calosoma (Callisthenes) elegans danilevskii Obydov, 1997d
Female adult
SE-Kazakhstan, Almaty reg., C Trans-Ili Alatau N-slopes, Kaskalen river Valley, Izvestkovy vill., h=1700 m, 6.V.2015, D.Kovalchuk leg./det R.Panin, L=32 mm
Copyright © Ruslan Panin

Calosoma (Callisthenes) elegans kashense Kabak, 2022 Paratype
Male adult
China, Xinjiang, left bank of Kash Riv., E of Nilki Town, foothills, 43°50ʹ17ʺ N, 82°48ʹ07ʺ E, H = 1270 m, 09.V.2017, I. I. Kabak leg. (ZIN)
Copyright © Ilya Kabak

Calosoma (Callisthenes) elegans rostislavi Semenov, 1906
Male adult
CH, Xinjiang, S foothills of Boro-Horo Mt. R., left bank of Piliktshi Riv., E of Karagatsh Vill., placor, 44°04ʹ38ʹʹ N, 81°30ʹ49ʹʹ E – 44°04ʹ48ʹʹ N, 81°30ʹ55ʹʹ E, 990–1040 m, 07.V.2017, I. I. Kabak leg.
Copyright © Ilya Kabak

Calosoma (Callisthenes) elegans elegans (as valentinae Obydov, 1997)
Male adult
SE-Kazakhstan, Almaty reg., W of Zarechnoe vill., h=600 m, 43.715278, 76.972778, 5.IV.2018, L=23 mm
Copyright © Ruslan Panin

Calosoma (Callisthenes) eversmanni Chaudoir, 1850
Male adult
Syria, Antilibanon
1.3.1991
male, 22 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) peksi Heinz & Pavesi, 1995
Female adult
Turquie (Izmir), Boz-dag( ob. Bozdag(ko'y 1500-1700 m, 2-3.V.1996, Heinz leg.
Copyright © Dmitri Obydov

Calosoma (Callisthenes) peksi Heinz & Pavesi, 1995
Male adult
Turquie (Izmir), Boz-dag( ob. Bozdag(ko'y 1500-1700 m, 2-3.V.1996, Heinz leg.
Copyright © Dmitri Obydov

Calosoma (Callisthenes) fischeri Fischer, 1842
Male adult
Mongolia, Mandalgob, 20 km
N Amardalai vill. 11-12.6.2003 Anichtchenko A. leg
male, 19 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) fischeri Fischer, 1842
Female adult
Mongolia, 10 km N Mandal-Gobi
Copyright © Igor Solodovnikov

Calosoma (Callisthenes) kuschakewitschi kuschakewitschi s.str.
Male adult
Karzhantau mt. rng., Susungen riv.,
2300 m, 7.6.1998 Tretjakov leg.
male, 23 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) kuschakewitschi plasoni Born, 1917
Female adult
Kirgizia, Chon-Aryk. 23.V.1998<br /> V. Dolin leg.
Copyright © Denis Potanin

Calosoma (Callisthenes) kuschakewitschi kuschakewitschi s.str.
Male adult

Copyright © Kirill Makarov

Calosoma (Callisthenes) glasunovi Semenov, 1900 Lectotype
Male adult
"Trkst. Mnt. Nurata, Sintab, Glasunov 1892", "Calosoma glasunovi m. Typ., XII.99, A. Semenov det."
Copyright © Dmitri Obydov

Calosoma (Callisthenes) glasunovi Semenov, 1900
Female adult
Uzbekistan, Nuratau Mt. Ridge, 1300 m, 6 km from Farish Village to the south, 6-7.V.2002, male, D. Obydov leg.
Copyright © Dmitri Obydov

Calosoma (Callisthenes) glasunovi Semenov, 1900
Female adult
Uzbekistan, Nuratau Mt. Ridge, 1300 m, 6 km from Farish Village to the south, 6-7.V.2002, female, D. Obydov leg.
Copyright © Dmitri Obydov

Calosoma (Callisthenes) kuschakewitschi batesoni Semenov & Redikorzev, 1928 Paratype
Male adult
"Bateson 1887 Inter Kazalinsk et Karkaralinsk (Desert. Kirgis.)"
Copyright © Dmitri Obydov

Calosoma (Callisthenes) kuschakewitschi solskyanus Obydov, 1999 Holotype
Male adult
South Kazakhstan, Ulanbel env., 19.4.1977, A. Shatalkin leg.
Copyright © Dmitri Obydov

Calosoma (Callisthenes) kuschakewitschi solskyanus Obydov, 1999
Female adult
Kazakhstan, 80 km from Merke to the north, 6.V.1994, J. Kadlec leg.
Copyright © Dmitri Obydov

Calosoma (Callisthenes) kuschakewitschi sidzhakensis Obydov, 2008 Holotype
Male adult
Uzbekistan-Kazakhstan border, Ugamsky Mt. Ridge., 2100 m, 15 km N-W Sidzhak Village, 1.VI.1978.
Copyright © Dmitri Obydov

Calosoma (Callisthenes) marginatus Gebler, 1829
Female adult
NE Kazakhstan, N Zaisan lake, 20.6.2000
Narymskii mt.rng. Klimenko A. leg.
female, 18 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) pseudocarabus Semenov et Redikorzev, 1928
Male adult
E Kazakstan, NE Saur, 2 km. W m. Tas, h=2500; 10.06.2001, Ivanov A.V. leg., 14 mm., male
photo, det. & coll. Menstschikov
Copyright © Anton Menstschikov

Calosoma (Callisthenes) mirificus Casale, 1979
Female adult
Afganistan, S slope of Paropamiz Mt. rng., near Sabzan pass, 2400m, 22.VI.1967 M.Stoljarov leg.
Collection of ZIN
det. D. Obydov 1999
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) panderi Fischer, 1822
Male adult
NW Kazakhstan, Aktiubinsk reg., Baiganinsk distr.,
near Zharly vill. 10-12.05.1997 leg. M.& L. Smirnov
male, 20 mm
Photo from www.zin.ru
Copyright © Maksim Smirnov

Calosoma (Callisthenes) pavlovskii Kryzhanovskij, 1955
Male adult
Kirgizstan, Talas Mt. rng., Besh-Tash
riv., 1000m, 15.5.2003
male, 20 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) pavlovskii Kryzhanovskij, 1955 Holotype
Male adult
"Talas, IV.08, E. Fischer", "Callisthenes pavlovskii sp. nov., Holotype, O. Kryzhanovskij det.".
Copyright © Dmitri Obydov

Calosoma (Callisthenes) akkolicus Obydov et Gottwald, 2002 Holotype
Male adult
Kazakhstan, Karatau, Akkol env., 9-10.V.1978, male, J. Gottwald leg.
Copyright © Dmitri Obydov

Calosoma (Callisthenes) akkolicus Obydov et Gottwald, 2002 Paratype
Female adult
Kazakhstan, Karatau, Akkol env., 9-10.V.1978, female, J. Gottwald leg.
Copyright © Dmitri Obydov

Calosoma (Callisthenes) akkolicus korelli Obydov, 2004 Paratype
Female adult
Kirghizia, Kyzyl-Sai Village env., V.1992, M. Pejcha leg.
Copyright © Dmitri Obydov

Calosoma (Callisthenes) panderi Fischer, 1822
Male adult
NW Kazakhstan, Aktiubinsk reg., Baiganinsk distr., near Zharly vill. 10-12.05.1997 leg. M.& L. Smirnov (MPU)
Copyright © Kirill Makarov

Calosoma (Callisthenes) cyaneosterum Mandl, 1954 Holotype
Male adult
«Songarei», «Panderi det. Ganglb.», «Collectio Kaufmann», «Typus», «Calosoma cyaneosterum m. Typus, det. Ing. K. Mandl» (NHMW)
Copyright © Ilya Kabak

Calosoma (Callisthenes) panderi Fischer, 1822
Male adult
«Окр. Уральска, IV.1907, Б. Уваров» (ca. 51°14ʹ N, 51°23ʹ E, H = 30 m)
Copyright © Ilya Kabak

Calosoma (Microcallisthenes) relictum Apfelbeck, 1918c
Male adult
Macedonia, Shar mt., Peak Tserapashina
1850-2530m 9.7.1995
male, 13 mm
Copyright © Alexander Anichtchenko

Calosoma (Microcallisthenes) relictum Apfelbeck, 1918c
Male adult
Macedonia, Shar mt., Peak Titov Vrah
2747 m, 14.7.1995 V.Sakalian leg.
male, 13 mm
Copyright © Alexander Anichtchenko

Calosoma (Microcallisthenes) relictum Apfelbeck, 1918
Male adult
Macedonia, Bistra planina Mts.; 13mm
Copyright © Aleš Sedláček

Calosoma (Microcallisthenes) relictum Apfelbeck, 1918
Female adult
Macedonia, Bistra planina Mts.; 13mm
Copyright © Aleš Sedláček

Calosoma (Microcallisthenes) relictum Apfelbeck, 1918
Male adult
Macedonia, Bistra planina Mts.; 14mm
Copyright © Aleš Sedláček

Calosoma (Callisthenes) regelianus A. Morawitz, 1886
Female adult
Tadzikistan, Darvaz Ezgan 2.5.1971
female, 25 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) klapperichi Mandl, 1955
Male adult
Afganistan, Bad. Shiva, 14.VII.1954. Topotypus
Copyright © Dmitri Obydov

Calosoma (Callisphaena) reticulatum reticulatum s.str.
Female adult
Germania, or. DDR, Bezirk Frankfurt, 24.V.1989
female, 25 mm
photo & det. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Callisphaena) reticulatum reticulatum
Male adult
Russia, Dagestan, 22 mm
Copyright © Igor Solodovnikov

Calosoma (Callisphaena) reticulatum reticulatum
Female adult
Russia, Dagestan, 26 mm
Copyright © Igor Solodovnikov

Calosoma (Callisthenes) usgentensis usgentensis Solskyi, 1874
Female adult
S Kirgizia, Dzhalal-Abad reg.,
Suzak vill., 1000m, 25.5.1999
male, 27 mm
photo, det. & coll. A. Anichtchenko
Copyright © Alexander Anichtchenko