Calosoma (Callisthenes) anthrax Semenov, 1900 Holotype
Male adult
collection of ZIN
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) grumi Semenov, 1900 Holotype
Female adult
collection ZIN
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) anthrax Semenov, 1900
Female adult
Inner Mongolia (China), Dalai-Nor Lake, VI.1957, Svetlovidova leg.
Copyright © Dmitri Obydov

Calosoma (Callisthenes) anthrax Semenov, 1900
Female adult
E-Mongolia: Bujur-Noor Lake, 29.VII.1976, Pechocki lg. (TOPOTYPUS)
Copyright © Igor Solodovnikov

Calosoma (Callisthenes) breuningi Mandl, 1954
Female adult
E Kazakhstan, S Alakol lake, Koktuma vill., 1-10.5.1999 Nekroshevich S. leg. female, 22 mm; det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) breuningi iliense Kabak, 2022
Male adult
SE Kazakhstan, near Sary Ozek vill., A. Klimenko leg; 22.04.1997; 22 mm.; det. Anichtchenko A. 2022
Copyright © Anton Menstschikov

Calosoma (Callisthenes) karelini vladimiri Obydov, 1998
Male adult
Kazakhstan, Taldy-Kurgan env., Kyzyl-Zhar Mts., 400 m, 15.IV.2000, M. Danilevsky leg.
photo from zin.ru
Copyright © Dmitri Obydov

Calosoma (Callisthenes) breuningi iliense Kabak, 2022 Paratype
Male adult
W of Kerbulak, 44°03ʹ03ʹʹ N, 77°01ʹ12ʹʹ E, 640 m, 25.IV.2017, I. I. Kabak leg.
Copyright © Ilya Kabak

Calosoma (Callisthenes) breuningi iliense Kabak, 2022 Paratype
Female adult
«SE Kazakhstan, right bank of Ili River, near Malaysary, 30.IV–1.V.1985, G. V. Nikolaev, A. S. Badenko leg.» (ca. 44°17ʹ30ʹʹ N, 76°57ʹ16ʹʹ E, 510 m),
Copyright © Ilya Kabak

Calosoma (Callisthenes) breuningi iliense Kabak, 2022 Paratype
Male adult
«SE Kazakhstan, right bank of Ili River, near Malaysary, 30.IV–1.V.1985, G. V. Nikolaev, A. S. Badenko leg.» (ca. 44°17ʹ30ʹʹ N, 76°57ʹ16ʹʹ E, 510 m),
Copyright © Ilya Kabak

Calosoma (Callisthenes) breuningi breuningi
Female adult
SE Kazakhstan, Almaty region, Ak-Eshki lake, 42 km NNE Taldy-Kurgan, H = 456 m, sands, 45°21ʹ44ʺ N, 78°37ʹ04ʺ E, 21.IV.2017, S. V. Kolov leg.
Copyright © Ilya Kabak

Calosoma (Callisthenes) breuningi breuningi
Female adult
SE Kazakhstan, Alakol lake, Koktuma env.
Copyright © Ilya Kabak

Calosoma (Callisthenes) breuningi breuningi
Female adult
«Sergiopol» (patria falsa!), (ZIN)
Copyright © Ilya Kabak

Calosoma (Callisthenes) breuningi iliense Kabak, 2022
Endophallus
W of Kerbulak, 44°03ʹ03ʹʹ N, 77°01ʹ12ʹʹ E, 640 m, 25.IV.2017, I. I. Kabak leg.
Copyright © Ilya Kabak

Calosoma (Callisthenes) breviusculus Mannerheim, 1830
Male adult
Armenia, near Ashtarik<br /> 15.6.1997<br /> male, 22 mm<br /> photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) substriatus Motschulsky, 1859
Female adult
Azerbaijan, Talysh, Gasmalian, 20.V.1992, V. Siniaev leg.
photo from zin.ru
Copyright © Dmitri Obydov

Calosoma (Callisthenes) substriatus Motschulsky, 1859 Lectotype
Male adult
"Callisthenes substriatus Motsch. Mt. Saganlug Persia"
Copyright © Dmitri Obydov

Calosoma (Callisthenes) substriatus Motschulsky, 1859
Male adult
Azerbaijan, Talysh, Gasmalian, 20.V.1992, V. Siniaev leg.
photo from zin.ru
Copyright © Dmitri Obydov

Calosoma (Callisthenes) pumicatus pumicatus s.str. Holotype
Female adult
"Plateau Persan Occid. de Zendjan a Ardebil, alt. moy 300 m (J. de Morgan)".
Copyright © Dmitri Obydov

Calosoma (Callisthenes) pumicatus pumicatus (as zarudnianum) Holotype
Male adult
Holotypus of Calosoma zarudnianum:
"Calosoma zarudnianum m, sp. n., A. Semenov-Tian-Shansky det. 21", "W. Pers., 13.V.04, Zarudnyi leg.".
Copyright © Dmitri Obydov

Calosoma (Callisthenes) pumicatus schelkovnikovi Zaitzev, 1918 Lectotype
Male adult
"ms. Sehend 11. 600' 18.V.16", "Mus Caucas. 100-16. Exp. Urmiana", "Calosoma schelkovnikovi Zaitz. (type)", "Received personally from F.A. Zaitzev in 1920".
Copyright © Dmitri Obydov

Calosoma (Callisthenes) breviusculus Mannerheim, 1830
Male adult
Turkey, Buglan; 21mm
Copyright © Aleš Sedláček

Calosoma (Callisthenes) breviusculus Mannerheim, 1830
Female adult
ARMENIE, Aragatsotn route du lac Kari, 28-06-2014, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Calosoma (Callisthenes) breviusculus pumicatus Lapouge, 1907
Female adult
Iran: Ardabil prov. Noor Lake (TOPOTYPUS)
Copyright © Igor Solodovnikov

Calosoma (Callisthenes) elegans semenovi Motschulsky, 1859
Female adult
SE Kazakhstan, Zailijsky Mt. rng.,
Talgar env., 1500 m, 21.5.2000
female, 27 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) elegans semenovi Motschulsky, 1859 Lectotype
Male adult
"elegans Kirsch. Stett. Zeit., 1859, 107. Semenof", "Neotypus, C. semenovi Motsch., Kryzhanovskij, 1966".
Copyright © Dmitri Obydov

Calosoma (Callisthenes) elegans keminensis Obydov, 2001 Holotype
Male adult
Kirghizia, Kemin (Bystrovka) env., 1300 m, Kyzyl-Oktyabr, 16.V.2000, M. Danilevsky leg.
Copyright © Dmitri Obydov

Calosoma (Callisthenes) elegans danilevskii Obydov, 1997d
Female adult
SE-Kazakhstan, Almaty reg., C Trans-Ili Alatau N-slopes, Kaskalen river Valley, Izvestkovy vill., h=1700 m, 6.V.2015, D.Kovalchuk leg./det R.Panin, L=32 mm
Copyright © Ruslan Panin

Calosoma (Callisthenes) elegans kashense Kabak, 2022 Paratype
Male adult
China, Xinjiang, left bank of Kash Riv., E of Nilki Town, foothills, 43°50ʹ17ʺ N, 82°48ʹ07ʺ E, H = 1270 m, 09.V.2017, I. I. Kabak leg. (ZIN)
Copyright © Ilya Kabak

Calosoma (Callisthenes) subtilestriatum Mandl, 1954 Holotype
Male adult
«Kuldsha», «marginatum det. Ganglb.», «Typus», «Calosoma subtilestriatum, Typus, det. Ing. K. Mandl», (NHMW).
Copyright © Ilya Kabak

Calosoma (Callisthenes) elegans gongliuense Kabak & Liang, 2023 Holotype Paratype
Male+Female adult
China, Xinjiang, Yili, Gongliu County, Wutu Vill., Institute of Zoology, Chinese Academy of Sciences” (in Chinese); “1982.IV.5;” “0262;” “102(E) 770020” (ca. 43°23ʹ52ʺ N / 82°14ʹ12ʺ E, H = 860 m) (IOZ).
Copyright © Ilya Kabak

Callisthenes declivis Dohrn, 1884 Lectotype
Male adult
Kazakhstan: “Turkestan,” “Type,” “elegans var. declivisDohrn,” “Mus. Zool. Polonicus Warszawa 12/45,” “Inst. Zool. P.A.N. Warszawa Cotypus Nr. 1253,” “Lectotypus Callisthenes declivis Dohrn, 1884 design. I. Kabak, 2004” (= Callisthenes elegans elegans Kirsch, 1859). (MZPW)
Copyright © Ilya Kabak

Calosoma (Callisthenes) elegans semenovi Motschulsky, 1859
Male adult
SE Kazakhstan, near Almaty, Komsomol vill., h=650 m, 43.458889, 76.748333, 3. V.2015
Copyright © Ruslan Panin

Calosoma (Callisthenes) elegans amethystinus Semenov & Redikorzev, 1928
Male adult
SE-Kazakhstan, Almaty reg., near Kazakhstan vill., 43.469167, 77.787222, 19.IV.2019, R. Rakhimov leg., L=29 mm
Copyright © Ruslan Panin

Calosoma (Callisthenes) eversmanni Chaudoir, 1850
Male adult
Syria, Antilibanon
1.3.1991
male, 22 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) peksi Heinz & Pavesi, 1995
Female adult
Turquie (Izmir), Boz-dag( ob. Bozdag(ko'y 1500-1700 m, 2-3.V.1996, Heinz leg.
Copyright © Dmitri Obydov

Calosoma (Callisthenes) peksi Heinz & Pavesi, 1995
Male adult
Turquie (Izmir), Boz-dag( ob. Bozdag(ko'y 1500-1700 m, 2-3.V.1996, Heinz leg.
Copyright © Dmitri Obydov

Calosoma (Callisthenes) fischeri Fischer, 1842
Male adult
Mongolia, Mandalgob, 20 km
N Amardalai vill. 11-12.6.2003 Anichtchenko A. leg
male, 19 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) fischeri Fischer, 1842
Female adult
Mongolia, 10 km N Mandal-Gobi
Copyright © Igor Solodovnikov

Calosoma (Callisthenes) kuschakewitschi kuschakewitschi s.str.
Male adult
Karzhantau mt. rng., Susungen riv.,
2300 m, 7.6.1998 Tretjakov leg.
male, 23 mm
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) kuschakewitschi plasoni Born, 1917
Female adult
Kirgizia, Chon-Aryk. 23.V.1998<br /> V. Dolin leg.
Copyright © Denis Potanin

Calosoma (Callisthenes) kuschakewitschi kuschakewitschi s.str.
Male adult

Copyright © Kirill Makarov

Calosoma (Callisthenes) glasunovi Semenov, 1900 Lectotype
Male adult
"Trkst. Mnt. Nurata, Sintab, Glasunov 1892", "Calosoma glasunovi m. Typ., XII.99, A. Semenov det."
Copyright © Dmitri Obydov

Calosoma (Callisthenes) glasunovi Semenov, 1900
Female adult
Uzbekistan, Nuratau Mt. Ridge, 1300 m, 6 km from Farish Village to the south, 6-7.V.2002, male, D. Obydov leg.
Copyright © Dmitri Obydov

Calosoma (Callisthenes) glasunovi Semenov, 1900
Female adult
Uzbekistan, Nuratau Mt. Ridge, 1300 m, 6 km from Farish Village to the south, 6-7.V.2002, female, D. Obydov leg.
Copyright © Dmitri Obydov

Calosoma (Callisthenes) kuschakewitschi batesoni Semenov & Redikorzev, 1928 Paratype
Male adult
"Bateson 1887 Inter Kazalinsk et Karkaralinsk (Desert. Kirgis.)"
Copyright © Dmitri Obydov

Calosoma (Callisthenes) kuschakewitschi solskyanus Obydov, 1999 Holotype
Male adult
South Kazakhstan, Ulanbel env., 19.4.1977, A. Shatalkin leg.
Copyright © Dmitri Obydov

Calosoma (Callisthenes) kuschakewitschi solskyanus Obydov, 1999
Female adult
Kazakhstan, 80 km from Merke to the north, 6.V.1994, J. Kadlec leg.
Copyright © Dmitri Obydov

Calosoma (Callisthenes) kuschakewitschi sidzhakensis Obydov, 2008 Holotype
Male adult
Uzbekistan-Kazakhstan border, Ugamsky Mt. Ridge., 2100 m, 15 km N-W Sidzhak Village, 1.VI.1978.
Copyright © Dmitri Obydov

Calosoma (Callisthenes) marginatus Gebler, 1829
Female adult
NE Kazakhstan, N Zaisan lake, 20.6.2000
Narymskii mt.rng. Klimenko A. leg.
female, 18 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) pseudocarabus Semenov et Redikorzev, 1928
Male adult
E Kazakstan, NE Saur, 2 km. W m. Tas, h=2500; 10.06.2001, Ivanov A.V. leg., 14 mm., male
photo, det. & coll. Menstschikov
Copyright © Anton Menstschikov

Calosoma (Callisthenes) mirificus Casale, 1979
Female adult
Afganistan, S slope of Paropamiz Mt. rng., near Sabzan pass, 2400m, 22.VI.1967 M.Stoljarov leg.
Collection of ZIN
det. D. Obydov 1999
photo Anichtchenko A. 2010
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) panderi Fischer, 1822
Male adult
NW Kazakhstan, Aktiubinsk reg., Baiganinsk distr.,
near Zharly vill. 10-12.05.1997 leg. M.& L. Smirnov
male, 20 mm
Photo from www.zin.ru
Copyright © Maksim Smirnov

Calosoma (Callisthenes) pavlovskii Kryzhanovskij, 1955
Male adult
Kirgizstan, Talas Mt. rng., Besh-Tash
riv., 1000m, 15.5.2003
male, 20 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) pavlovskii Kryzhanovskij, 1955 Holotype
Male adult
"Talas, IV.08, E. Fischer", "Callisthenes pavlovskii sp. nov., Holotype, O. Kryzhanovskij det.".
Copyright © Dmitri Obydov

Calosoma (Callisthenes) akkolicus Obydov et Gottwald, 2002 Holotype
Male adult
Kazakhstan, Karatau, Akkol env., 9-10.V.1978, male, J. Gottwald leg.
Copyright © Dmitri Obydov

Calosoma (Callisthenes) akkolicus Obydov et Gottwald, 2002 Paratype
Female adult
Kazakhstan, Karatau, Akkol env., 9-10.V.1978, female, J. Gottwald leg.
Copyright © Dmitri Obydov

Calosoma (Callisthenes) akkolicus korelli Obydov, 2004 Paratype
Female adult
Kirghizia, Kyzyl-Sai Village env., V.1992, M. Pejcha leg.
Copyright © Dmitri Obydov

Calosoma (Callisthenes) panderi Fischer, 1822
Male adult
NW Kazakhstan, Aktiubinsk reg., Baiganinsk distr., near Zharly vill. 10-12.05.1997 leg. M.& L. Smirnov (MPU)
Copyright © Kirill Makarov

Calosoma (Callisthenes) cyaneosterum Mandl, 1954 Holotype
Male adult
«Songarei», «Panderi det. Ganglb.», «Collectio Kaufmann», «Typus», «Calosoma cyaneosterum m. Typus, det. Ing. K. Mandl» (NHMW)
Copyright © Ilya Kabak

Calosoma (Callisthenes) panderi Fischer, 1822
Male adult
«Окр. Уральска, IV.1907, Б. Уваров» (ca. 51°14ʹ N, 51°23ʹ E, H = 30 m)
Copyright © Ilya Kabak

Calosoma (Microcallisthenes) relictum Apfelbeck, 1918c
Male adult
Macedonia, Shar mt., Peak Tserapashina
1850-2530m 9.7.1995
male, 13 mm
Copyright © Alexander Anichtchenko

Calosoma (Microcallisthenes) relictum Apfelbeck, 1918c
Male adult
Macedonia, Shar mt., Peak Titov Vrah
2747 m, 14.7.1995 V.Sakalian leg.
male, 13 mm
Copyright © Alexander Anichtchenko

Calosoma (Microcallisthenes) relictum Apfelbeck, 1918
Male adult
Macedonia, Bistra planina Mts.; 13mm
Copyright © Aleš Sedláček

Calosoma (Microcallisthenes) relictum Apfelbeck, 1918
Female adult
Macedonia, Bistra planina Mts.; 13mm
Copyright © Aleš Sedláček

Calosoma (Microcallisthenes) relictum Apfelbeck, 1918
Male adult
Macedonia, Bistra planina Mts.; 14mm
Copyright © Aleš Sedláček

Calosoma (Callisthenes) regelianus A. Morawitz, 1886
Female adult
Tadzikistan, Darvaz Ezgan 2.5.1971
female, 25 mm
photo, det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Callisthenes) klapperichi Mandl, 1955
Male adult
Afganistan, Bad. Shiva, 14.VII.1954. Topotypus
Copyright © Dmitri Obydov

Calosoma (Callisphaena) reticulatum reticulatum s.str.
Female adult
Germania, or. DDR, Bezirk Frankfurt, 24.V.1989
female, 25 mm
photo & det. Anichtchenko A.
Copyright © Alexander Anichtchenko

Calosoma (Callisphaena) reticulatum reticulatum
Male adult
Russia, Dagestan, 22 mm
Copyright © Igor Solodovnikov

Calosoma (Callisphaena) reticulatum reticulatum
Female adult
Russia, Dagestan, 26 mm
Copyright © Igor Solodovnikov

Calosoma (Callisthenes) usgentensis usgentensis Solskyi, 1874
Female adult
S Kirgizia, Dzhalal-Abad reg.,
Suzak vill., 1000m, 25.5.1999
male, 27 mm
photo, det. & coll. A. Anichtchenko
Copyright © Alexander Anichtchenko