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Carabidae of the World
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Carabinae Latreille, 1802
→
Cychrini Laporte, 1834
→
Cychrina sensu stricto
→
Scaphinotus Dejean, 1826
→ Brennus Motschulsky, 1865
Region
Afghanistan (AF)
Albania (AL)
Aleutian isl (ALU)
Algeria (AG)
Andorra (AN)
Angola (ANG)
Antarctica (Antarctica)
Antilles islands (ANT)
Antigua and Barbuda (Antigua and Barbuda)
Barbados (Barbados)
Cayman islands (Cayman islands)
Cuba (Cuba)
Dominica (Lesser Antilles) (Dominica (Lesser Antilles))
Dominican Republic (Hispaniola, Greater Antilles) (Dominican Republic (Greater Antilles))
Grenada (Grenada)
Guadeloupe (Guadeloupe)
Haiti (Haiti)
Jamaica (Jamaica)
Martinique (Martinique)
Puerto Rico (Puerto Rico)
Saint Kitts and Nevis (Saint Kitts and Nevis)
Saint Lucia (Saint Lucia)
St. Vincent (St. Vincent)
Trinidad and Tobago (Trinidad and Tobago)
Arab Emirates (AE)
Argentina (ARG)
Armenia (AR)
Australia (AST)
Christmas Island (Christmas Island)
Lord Howe Island (Lord Howe Island)
Macquarie Island (Macquarie Island)
Tasmania (Tasmania)
Austria (AU)
Azerbaijan (AB)
Azores (AZ)
BALTIC AMBER, Baltic See coast and Amber quarry Jantarny near Kaliningrad, Russia (BAM)
Bahamas (BHM)
Bahrain (BA)
Bangladesh (BGD)
Belgium (BE)
Belize (BLZ)
Bhutan (BT)
Bolivia (BOL)
Bosnia and Herzegovina (BH)
Botswana (BTW)
Brazil (BRZ)
Acre (AC)
Alagoas (AL)
Amazonas (AM)
Arnapá (AP)
Bahia (BA)
Ceará (CE)
District Federal (Brasilia) (DF)
Espirito Santo (ES)
Goiás (GO)
Maranhão (MA)
Mato Grosso (MT)
Mato Grosso do Sul (MS)
Minas Gerais (MG)
Paraná (PR)
Paraîba (PB)
Pará (PA)
Pernambouc (PE)
Piaui (PI)
Rio Grande do Norte (RN)
Rio Grande do Sul (RS)
Rio de Janeiro (RJ)
Rondônia (RO)
Roraima (RR)
Santa Catarina (SC)
Sergipe (SE)
São Paulo (SP)
Tocantins (TO)
Brunei (Brunei)
Bulgaria (BU )
Burundi (BUR)
Byelorussia (BY)
Cambodia (CBD)
Cameroon (CAM)
Canada (CAN)
Canary isls (CI)
Cape Verde (Cape Verde)
Central African Republic (CAR)
Chad (CHD)
Chile (CHL)
Juan Fernández Islands (Juan Fernández Islands)
China (CH)
Anhui (Anhui)
Beijing (Beijing)
Chongqing (Chongqing)
Fujian (Fujian)
Gansu (Gansu)
Guangdong (Guangdong)
Guangxi (Guangxi)
Guizhou (Guizhou)
Hainan (Hainan)
Hebei (Hebei)
Heilongjiang (Heilongjiang)
Henan (Henan)
Honkong (Hongkong)
Hubei (Hubei)
Hunan (Hunan)
Inner Mongolia (Inner Mongolia)
Jiangsu (Jiangsu)
Jiangxi (Jiangxi)
Jilin (Jilin)
Liaoning (Liaoning)
Macau (Macau)
Ningxia (Ningxia)
Qinghai (Qinghai)
Shaanxi (Shaanxi)
Shandong (Shandong)
Shanghai (Shanghai)
Shanxi (Shanxi)
Sichuan (Sichuan)
Tianjin (Tsiensin) (Tianjin)
Xinjiang (Xinjiang)
Xizang (Tibet) (Xizang (Tibet))
Yunnan (Yunnan)
Zhejiang (Zhejiang)
Colombia (COL)
Comoros isl. (COM)
Costa Rica (CRC)
Croatia (CR)
Crozet Islands (CRZ)
Cyprus (CY)
Czech (CZ)
DOMINICAN AMBER, Dominican Republic, Lower Miocene. (DAM)
Democratic Republic of the Congo (CNG)
Denmark (DE)
Diego Garcia isls (DGC)
Djibouti (DJB)
Ecuador (ECD)
Egypt (EG)
Sinai (Sinai)
El Salvador (SVD)
Equatorial Guinea (EQG)
Bioco (Fernando Pó) island (Bioco (Fernando Pó) island)
Eritrea (ERT)
Estonia (EN)
Ethiopia (ETP)
Falkland il. (FLK)
Faroe Islands (FA)
Finland (FI)
France (FR)
Corse (Corse)
French Guiana (GNF)
Gabon (GAB)
Galapagos Islands (GAL)
Georgia (GG)
Abkhazia (Abkhazia)
Ajaria (Ajaria)
South Ossetia (South Ossetia)
Germany (GE)
Great Britain (GB)
Greece (GR)
Aegean Islands (Aegean Islands)
Crete (Crete)
Dodecanese Islands (Dodecanese Islands)
Euboea (Evia) Island (Euboea (Evia) Island)
Ionian Islands (Ionian Islands)
Peloponnese (Peloponnese)
Saronic Islands (Saronic Islands)
Sporades Islands (Sporades Islands)
The Cyclades (The Cyclades)
Guatemala (GTM)
Guyana (GYN)
Honduras (HDR)
Hungary (HU)
Iceland (IC)
India (IND)
Andaman and Nicobar isl. (Andaman and Nicobar isl.)
Andhra Pradesh (Andhra Pradesh)
Arunachal Pradesh (Arunachal Pradesh)
Assam (Assam)
Bihar (Bihar)
Chhattisgarh (Chhattisgarh)
Delhi (Delhi)
Goa (Goa)
Gujarat (Gujarat)
Haryana (Haryana)
Himachal Pradesh (Himachal Pradesh)
Jammu and Kashmir (Jammu and Kashmir)
Jharkhand (Jharkhand)
Karnataka (Karnataka)
Kerala (Kerala)
Ladakh (Ladakh)
Madhya Pradesh (Madhya Pradesh)
Maharashtra (Maharashtra)
Manipur (Manipur)
Meghalaya (Meghalaya)
Mizoram (Mizoram)
Nagaland (Nagaland)
Odisha (Odisha)
Puducherry (Puducherry)
Punjab (Punjab)
Rajasthan (Rajasthan)
Sikkim (Sikkim)
Tamil Nadu (Tamil Nadu)
Telangana (Telangana)
Tripura (Tripura)
Uttar Pradesh (Uttar Pradesh)
Uttarakhand (Uttarakhand)
West Bengal (West Bengal)
Indonesia (IDS)
Borneo (Kalimantan) (Borneo)
Java (Java)
Lesser Sunda Islands (Lesser Sunda Ils.)
Maluku Islands (Maluku Isl.)
Papua (Papua)
Riau Islands (Riau Islands)
Sulawesi (Celebes) (Sulawesi)
Sumatra (Sumatra)
Timor (Timor)
Iran (IN)
Iraq (IQ)
Ireland (IR)
Israel (IS)
Italy (IT)
Sardegna (Sardegna)
Sicily (Sicily)
Japan (JA)
Hokkaido (Hokkaido)
Honshu (Honshu)
Kyushu (Kyushu)
Ryukyu Islands (Ryukyu Islands)
Shikoku (Shikoku)
Jordan (JO)
Kashmir (KA)
Kazakhstan (KZ)
Kenya (KEN)
Kerguelen Islands (Kerguelen Islands)
Kuwait (KU)
Kyrgyzstan (KI)
Laos (LAO)
Latvia (LA)
Lebanon (LE)
Lesotho (LSO)
Libya (LB)
Liechtenstein (LS)
Lithuania (LT)
Luxenbourg (LU)
Macedonia (MC)
Madagascar (MAD)
Madeira Arch. (MR)
Malawi (MLW)
Malaysia (MLS)
Sabah (Borneo) (Sabah (Borneo))
Sarawak (Borneo) (Sarawak (Borneo))
Maldives (MLD)
Malta (MA)
Mauritania (MAU)
Mauritius (MRC)
Melanesia (Melanesia)
Bismarck Archipelago (Bismarck Archipelago)
Fiji (Fiji)
New Caledonia (New Caledonia)
Papua New Guinea (Papua New Guinea)
Solomon Islands (Solomon Islands)
Vanuatu ils (Vanuatu ils)
Mexico (MEX)
Aguascalientes (Aguascalientes)
Baja California (Baja California)
Baja California Sur (Baja California Sur)
Campeche (Campeche)
Chiapas (Chiapas)
Chihuahua (Chihuahua)
Coahuila (Coahuila)
Colima (Colima)
Durango (Durango)
Federal District (Federal District)
Guanajuato (Guanajuato)
Guerrero (Guerrero)
Hidalgo (Hidalgo)
Jalisco (Jalisco)
Mexico State (Mexico State)
Michoacan (Michoacan)
Morelos (Morelos)
Nayarit (Nayarit)
Nuevo Leon (Nuevo Leon)
Oaxaca (Oaxaca)
Puebla (Puebla)
Queretaro (Queretaro)
Quintana Roo (Quintana Roo)
San Luis Potosi (San Luis Potosi)
Sinaloa (Sinaloa)
Sonora (Sonora)
Tabasco (Tabasco)
Tamaulipas (Tamaulipas)
Tlaxcala (Tlaxcala)
Veracruz (Veracruz)
Yucatan (Yucatan)
Zacatecas (Zacatecas)
Micronesia (MIC)
Caroline islands (Caroline islands)
Guam (Guam)
Kiribati (Kiribati)
Marshall ils (Marshall ils)
North Marians isl. (North Marians isl.)
Palau archipelago (Palau archipelago)
Moldavia (MD)
Mongolia (MG)
Montenegro (ME)
Mordor (RU)
Altai Republic and Altaisky Province (Altai Republic and Altaisky Province)
Amurskaya Area (Amurskaya Area)
Bashkiria, E of Chelyabinsk regs. (S Ural mts. Bashkiria, E of Chelyabinsk regs.)
Buryat Republic (Buryat Republic)
Chukot autonomous region (Chukotka)
East Sayan Mountains (East Sayan Mountains)
Ingushetia, Chechnya, Daghestan (E Caucas) (Ingushetia, Chechnya, Daghestan (E Caucas))
Irkutsk Area (Irkutsk Area)
Kabardino-Balkaria, Ossetia (C Caucas) (Kabardino-Balkaria, Ossetia (C Caucas))
Kaliningrad, Briansk, Smolensk, Tula, Orlov, Kaluga, Moskow, Riaz (C of Russian plains)
Kamchatskaya Area (Kamchatka)
Karachaevo-Cherkessia (W Caucas) (Karachaevo-Cherkessia)
Karelia, Leningrad reg., Pskov reg., Kostroma reg., Perm reg., Komi, Pechora plains. (N of Russian plains)
Khabarovsky Province (Khabarovsky Province)
Khamar-Daban Mountains (Khamar-Daban Mountains (E Siberia))
Komi, Yamalo-Nenetsk, Hanty-Mansi (N Ural mts.)
Krasnodar reg. (W Caucas)
Krasnoyarsky Province (Krasnoyarsk)
Kuriles Islands (Kuriles Islands)
Magadanskaya Area (Magadan)
Omsk, S of Tyumen, W Novosibirsk regs. (S of W Siberia)
Perm, Sverdlovsk regs. (C Ural mts)
Primorsky District (Primorie) (Maritime Prov)
Sakhalin Island (Sakhalin Island)
Tuva Republic (Tuva)
Voronezh, Rostov, Saratov, Kalmykia, Volgograd, Astrahan regs. (S of Russian plains)
West Sayan Mountains (West Sayan)
Yakut-Saha Republic (Yakut-Saha Republic)
Zabaykalsky Krai (ex Chita reg.) (Zabaykalsky Krai)
plains of Krasnodar reg., S Rostov reg., Stavropol reg. (Ciscaucas reg. (plains of Krasnodar reg., S Rostov)
taiga from Ural mts to Yenisei riv. (Middle of W Siberia)
tundra of Tyumen, Yamalo-Nenetsk regs. (N of W Siberia)
Morocco (MO)
Mozambique (MZB)
Myanmar (Burma) (BRM)
Namibia (NAM)
Nepal (NP)
Netherlands (NL)
New Zealand (NZL)
Antipodes Islands (Antipodes Islands)
Auckland Islands (Auckland Islands)
Campbell Island (Campbell Island)
Chatham Islands (Chatham Islands)
Nicaragua (NCG)
North Korea (NC)
Norway (NR)
Svalbard (Spitzbergen) (SB)
Oman (OM)
Pakistan (PA)
Panama (PNM)
Paraguay (PRG)
Peru (PER)
Philippines (PLP)
Basilan (Basilan)
Cebu (Cebu)
Guimaras (Guimaras)
Leyte (Leyte)
Luzon (Luzon)
Marinduque (Marinduque)
Masbate (Masbate)
Mindanao (Mindanao)
Mindoro (Mindoro)
Negros (Negros)
Palawan (Palawan)
Panay (Panay)
Polillo Islands (Polillo Islands)
Romblon (Romblon)
Samar (Samar)
Sulu Archipelago (Sulu Archipelago)
Poland (PL)
Polynesia (POL)
Cook Islands (Cook Islands)
Hawaiian Islands (HAW)
Marquesas Islands (Marquesas Islands)
Niue (Niue)
Norfolk ils (Norfolk ils)
Pitcairn ils (Pitcairn ils)
Rapa Nui (Easter Island) (Rapa Nui)
Samoa (Samoa)
Society Islands (Society Islands)
Tahiti (Tahiti)
Tokelau (TKL)
Tonga (Tonga)
Tuvalu ils (Tuvalu ils)
Portugal (PT)
Qatar (QA)
Republic of South Africa (RSA)
Republic of the Congo (CG)
Reunion isl. (Reunion isl.)
Rodriguez isls. (ROD)
Romania (RO)
Rwanda (RWA)
Santa Helena isl (SH)
Sao Tome e Principe (STM)
Saudi Arabia (SA)
Serbia (SRB)
Seychelles (SEY)
Singapore (SGP)
Slovakia (SK)
Slovenia (SL)
Somalia (SOM)
South Georgia and the South Sandwich Islands (SGSSI)
South Korea (SC)
Spain (SP)
Balearic Islands (Balearic Islands)
Sri Lanka (SRL)
St. Paul isls (STP)
Sudan (SUD)
Surinam (SUR)
Swaziland (SWZ)
Sweden (SV)
Switzerland (SZ)
Syria (SY)
Tadzhikistan (TD)
Taiwan (Formosa) (Taiwan)
Tanzania (TAN)
Zanzibar isl. (Zanzibar isl.)
Thailand (TAI)
Tunisia (TU)
Turkey (TR)
Turkmenistan (TM)
Uganda (UGD)
Ukraine (UK)
Crimea (Crimea)
United States (USA)
Alabama (Alabama)
Alaska (Alaska)
Arizona (Arizona)
Arkansas (Arkansas)
California (California)
Colorado (Colorado)
Connecticut (Connecticut)
Delaware (Delaware)
Florida (Florida)
Georgia (Georgia)
Idaho (Idaho)
Illinois (Illinois)
Indiana (Indiana)
Iowa (Iowa)
Kansas (Kansas)
Kentucky (Kentucky)
Louisiana (Louisiana)
Maine (Maine)
Maryland (Maryland)
Massachusetts (Massachusetts)
Michigan (Michigan)
Minnesota (Minnesota)
Mississippi (Mississippi)
Missouri (Missouri)
Montana (Montana)
Nebraska (Nebraska)
Nevada (Nevada)
New Hampshire (New Hampshire)
New Jersey (New Jersey)
New Mexico (New Mexico)
New York (New York)
North Carolina (North Carolina)
North Dakota (North Dakota)
Ohio (Ohio)
Oklahoma (Oklahoma)
Oregon (Oregon)
Pennsylvania (Pennsylvania)
Rhode Island (Rhode Island)
South Carolina (South Carolina)
South Dakota (South Dakota )
Tennessee (Tennessee)
Texas (Texas)
Utah (Utah)
Vermont (Vermont)
Virginia (Virginia)
Washington (Washington)
West Virginia (West Virginia)
Wisconsin (Wisconsin)
Wyoming (Wyoming)
Uruguay (URG)
Uzbekistan (UZ)
Venezuela (VNZ)
Vietnam (VTN)
West Africa (WAF)
Benin (Benin)
Burkina Faso (Burkina Faso)
Côte d'Ivoire (Côte d'Ivoire)
Gambia (Gambia)
Ghana (Ghana)
Guinea (Guinea)
Guinea Bissau (Guinea Bissau)
Liberia (Liberia)
Mali (Mali)
Niger (Niger)
Nigeria (Nigeria)
Senegal (Senegal)
Sierra Leone (Sierra Leone)
Togo (Togo)
Yemen (YE)
Socotra (Socotra)
Zambia (ZMB)
Zimbabwe (ZIM)
Scaphinotus (Brennus) crenatus Motschulsky, 1859
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright ©
- - -
crenatus
taxon page
(1 photos)
Scaphinotus (Brennus) cristatus Harris, 1839
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright ©
- - -
Scaphinotus (Brennus) cristatus Harris, 1839
Female adult
USA California, Monterrey Co., Carmel, 18.8.1974, J.B.Johnson (cAA) 16mm
Copyright ©
Alexander Anichtchenko
cristatus
taxon page
(2 photos)
Scaphinotus (Brennus) interruptus Menetries, 1844
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright ©
- - -
Scaphinotus (Brennus) interruptus Menetries, 1844
Male adult
USA California, Sonoma Co., 2.5mi N of Cazadero on King Ridge Rd. Big Austin Creek, 70m, 38.33N 123.06W 21.6.1996 D.H.Kavanaugh leg. (cAA). det. Anichtchenko A. 2021
Copyright ©
Alexander Anichtchenko
interruptus
taxon page
(2 photos)
Scaphinotus (Brennus) marginatus var. fallax Roeschke, 1907
Male adult
USA, OR., Yamhill Co., Crawford Beck Vineyard, Rd 153 E of Amity, 45.06.805N 123.08.539W 30.3.-22.5.2019, M. Kippenhan leg. (cAA) 14mm; det. Anichtchenko A. 2020
Copyright ©
Alexander Anichtchenko
Scaphinotus (Brennus) marginatus Fischer, 1822
Female adult
USA, OR., Multnomah Co., Portland, Oaks Bottom WA, 7.5.2004 M. Kippenhan (cAA) 14 mm; det. Anichtchenko A. 2020
Copyright ©
Alexander Anichtchenko
Scaphinotus (Brennus) marginatus Fischer, 1822
Male adult
Canada, BRITISH COLUMBIA (BC), Wynndel, Duck Creek Road, N 49°11.852', W 116°32.294', ele: 1917', 24.VII-04.IX.2015, leg. Todd Lawton
Copyright ©
Igor Solodovnikov
marginatus
taxon page
(3 photos)
Scaphinotus (Brennus) obliquus LeConte, 1868
Male adult
USA, CA., Shasta Co., Castle Crags State Park, 29-29.5.2016, M. Kippenhan leg. (cAA) 15 mm; det. Anichtchenko A. 2020
Copyright ©
Alexander Anichtchenko
obliquus
taxon page
(1 photos)
Scaphinotus (Brennus) oreophilus Rivers, 1890
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright ©
- - -
Scaphinotus (Brennus) cf oreophilus Rivers, 1890
Female adult
USA , CA, Toulumne Co., The Pines CG, Rt 120 West of Buck Meadows, 3267m, 37.49.14N 120.05.7W 19.6.2011 (cAA) 15mm; det. Anichtchenko A. 2020
Copyright ©
Alexander Anichtchenko
Scaphinotus (Brennus) cf oreophilus Rivers, 1890
Male adult
USA , CA, Toulumne Co., The Pines CG, Rt 120 West of Buck Meadows, 3267m, 37.49.14N 120.05.7W 19.6.2011, M. Kippenhan leg. (cAA) 15mm
Copyright ©
Alexander Anichtchenko
oreophilus
taxon page
(3 photos)
Scaphinotus (Brennus) punctatus LeConte, 1874
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab (det as subtilis Schaum, 1863), NMNH.; as punctatus det by Anichtchenko A. 2020
Copyright ©
- - -
punctatus
taxon page
(1 photos)
Scaphinotus (Brennus) rugiceps incipiens (Casey, 1897)
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright ©
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Scaphinotus (Brennus) rugiceps rugiceps s.str.
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright ©
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Scaphinotus (Brennus) rugiceps Horn, 1872
Male adult
USA: OR: Josephine Co. / Lone Mountain Rd, 4.3 miles S.W of Hwy 199 / 42° 01.787' N 123° 45.760' W 1719', 18-IV-6-VI-2020, M. Kippenhan (cAA) 15 mm; det. Anichtchenko A. 2020
Copyright ©
Alexander Anichtchenko
rugiceps
taxon page
(3 photos)
Scaphinotus (Brennus) striatopunctatus (Chaudoir, 1844)
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright ©
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Scaphinotus (Brennus) striatopunctatus (Chaudoir, 1844)
Male adult
USA California, Sonoma Co., 10 mi NW of Santa Rosa, Pepperwood Ranch NR, 22.3.1997 D.H. Kavanaugh leg. (cAA) det. A. Anichtchenko 2021
Copyright ©
Alexander Anichtchenko
striatopunctatus
taxon page
(2 photos)
Scaphinotus (Brennus) ventricosus Dejean, 183
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright ©
- - -
ventricosus
taxon page
(1 photos)
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