Cychrus volkovi O.Berlov, 1997 Holotype Holotype
Male adult

Copyright © Oleg Berlov

Cychrus (Cychrus) aeneus matocqi Deuve, 2016 Holotype
Male adult
Géorgie, Ratcha-Lekhoumi, environs d’Orbéli, Chaîne des Monts de Lekhoumi, 700 mètres (R. Boulben, Th. Deuve, A. Matocq, A. Taghavian leg. (MNHN).
Copyright © Thierry Deuve

Cychrus (Cychrus) aeneus stefanii Deuve, 2019 Holotype
Male adult
Géorgie, Basse-Svanétie, à l’ouest de Lentekhi, versant nord des monts Devashi, env. 900 mètres (R. Boulben et P. Stéfani leg.) (MNHN)
Copyright © Thierry Deuve

Cychrus (Cychrus) aeneus japoshvilii Deuve & Prunier, 2014
Male adult
Georgy: Samtskhe-Dzhavakhetie, near Borjomi: Akhaldaba (TOPOTYPUS)
Copyright © Igor Solodovnikov

Cychrus (Cychrus) aeneus aeneus Fisher-Waldheim, 1824
Male adult
Georgy: env. Bakuriani
Copyright © Igor Solodovnikov

Cychrus (Cychrus) aeneus volkovi O.Berlov, 1997
Male adult

Copyright © Igor Solodovnikov

Cychrus (Cychrus) anatolicus frivaldskyi Roeschke, 1907
Female adult
Turkey, Akkus 900m 3.8.1991 (cAA) det. Anichtchenko A. 2021
Copyright © Alexander Anichtchenko

Cychrus angustatus Hoppe & Hornschuch, 1825
Male adult
Italia, Lombardia (BG), Arera mt., 2100m<br /> Zambla, 19.6.2003 A. Degiovanni leg.<br /> det. V. Monzini, 2014 male, 20 mm
Copyright © Alexander Anichtchenko

Cychrus angustatus Hoppe & Hornschuch, 1825
Male adult
NC-Italy, Bergamo, Mt. Presolana h=1500m, Colere env., 4-15.VI.2010, Manfred Egger leg.
L=23 mm
Copyright © Ruslan Panin

Cychrus attenuatus attenuatus Fabricius, 1792
Male+Female adult
W.UKRAINE, NE Carpathians,
Lviv Reg., Skole distr., Skolivski Beskidy Mts. Zelemyanka Mt.Range h=700m, 26.V -1.VII.2010, Panin R. leg.

Copyright © Ruslan Panin

Cychrus attenuatus attenuatus s.str.
Female adult
CZ, NE Moravia, Moravskoslezské Beskydy Mts.; 13mm
Copyright © Aleš Sedláček

Cychrus (Cychrus) auvrayorum andrei Cavazzuti, 2001
Female adult
China, S Gansu, Min Shan, 5-20 km S Cagan town, 2300-3200m, 33°33.09N 104°20.12E, 7-20.7.2019, leg. V. Patrikeev (cAA). 18 mm; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Cychrus (Cychrus) becvari Deuve, 2018 Holotype
Female adult
Chine, Yunnan, Mont Haba Xueshan, versant sudest, 2500-3800 mètres, 27°20’N-1°11’E, S. Bečvář leg. (MNHN)
Copyright © Thierry Deuve

Cychrus (Cychrus) becvari Deuve, 2018 Paratype
Female adult
Chine, Yunnan, Mont Haba Xueshan, versant sudest, 2500-3800 mètres, 27°20’N-1°11’E (S. Bečvář leg.)
Copyright © Thierry Deuve

Cychrus (Cychrus) angustior Kleinfeld, 2000 Holotype
Female adult
China, W.-Sichuan, Xiaojin: Siguniangshan (31°06'/102°54'), 4000 m; VII. 1999; in coll. Kleinfeld
Copyright © Frank Kleinfeld

Cychrus (Cychrus) cf bruggei taiyangheicus Deuve & Tian, 2007
Male adult
W Sichuan pass E of Jinchuan city 3850 m; leg. V. Patrikeev (cAA). 13.8 mm; det. A. Anichtchenko, 2020
Copyright © Alexander Anichtchenko

Cychrus (Cychrus) angustior Kleinfeld, 2000 Paratype
Female adult

Copyright © - - -

Cychrus (Cychrus) cf busatoi Cavazzuti, 2009
Male adult
China, Yunnan, Dequen, Baima pass. 4500m, 20.6.2016, leg. V. Patrikeev (cAA) det. A. Anichtchenko, 2019
Copyright © Alexander Anichtchenko

Cychrus (Cychrus) busatoi Cavazzuti, 2009 Holotype
Male adult
Colle fra Zhongdian e Wengshui, 3800 m, in picetum, Confine fra Sichuan e Yunnan, Cina sud-occidentale.
Copyright © - - -

Cychrus caraboides Linnaeus, 1758
Male adult
L=17,0 mm
RUSSIA, Moscow city, East distr., National park «Losiniy Ostrov», spruce forest. 9.IX.2000
leg., det. & photo - Vlasenko A. S.
Copyright © Andrey Vlasenko

Cychrus (Cychrus) caraboides (as elongatus Hoppe,)
Male adult
W Ukraine, NE Carpathians, Chornagora Mt Range, Mt Hoverla, h~2030 m, alpin zone, 10.VI.-5,VIII.2017, R. Panin R. L =13 mm
Copyright © Ruslan Panin

Cychrus (Cychrus) chareti Deuve, 1994
Female adult
NW Yunnan, Shangri La pref., Balakow pass, 3900m, 10.7.2016, V. Patrikeev leg. (cAA) 15mm; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Cychrus (Cychrus) cylindricollis Pini, 1871
Male adult

Copyright © Vittorino Monzini

Cychrus (Cychrus) davidiani Imura & Cavazzuti, 2017 Holotype
Female adult
SW. of Guoditang［ 锅底塘］, 27°12’38”N/ 99°21’22”E, 3235 m, 6.4 km ENE. of Weixi［ 维 西, SE. of Weixi-lisuzu Zizhixian［维西傈僳族自治县］, NW. Yunnan, China, 29-V-2016. Image from Imura & Cavazzuti, 2017
Copyright © - - -

Cychrus (Cychrus) dufouri Chaudoir, 1869
Male adult
Spain, Sierra de Aralar, 23.6.2005, A. Anichtchenko leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Cychrus (Cychrus) cf. furumii minshanicus Deuve, 1993 (as simillimus)
Female adult
S Gansu 80 km E Tewo town , 40 km SW of Lazikow pass 2800 m, S slope Qinling shan (cAA) 18 mm; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Cychrus (Cychrus) gorodinskiellus Deuve, 2016 Holotype
Male adult
Chine, Yunnan, Dêqên, monts Hengduan Shan, près du col de Doker La, 3900 mètres. (in private coll. Ingo Brunk)
Copyright © Thierry Deuve

Cychrus (Cychrus) grajus K. Daniel & J. Daniel, 1898d: 61
Female adult
Alpi Graje, Gran Paradiso 2350 m - leg V. Monzini
Copyright © Vittorino Monzini

Cychrus (Cychrus) hampei Gestro, 1875
Male adult
S. Livno, Cvrsnica Planina, Pločno mt., 2100 m, 43°36’N/17°33’E, reg. alp., VI-IX-2018
Copyright © Frank Kleinfeld

Cychrus hemphillii hemphillii s.str.
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Cychrus hemphillii rickseckeri LeConte, 1884
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Cychrus hemphillii rickseckeri LeConte, 1884
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © - - -

Cychrus (Cychrus) horribilis Deuve, 2016 Holotype Paratype
Male+Female adult
Chine, Gansu, 15 km au sud-ouest de Wudu (Longnan), 3500 mètres, 33°19’37,1’’N-104°45’05,9’’E (M. Janata et R.Sehnal), in coll. Ross Sehnal, à Unhošt (République Tchèque).
Copyright © Thierry Deuve

Cychrus (Cychrus) horribilis Deuve, 2016
Female adult
China, S Gansu, Min Shan, 5-20 km S Cagan town, 2300-3200m, 33°33.09N 104°20.12E, 7-20.7.2019, leg. V. Patrikeev (cAA). 15.5 mm; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Cychrus (Cychrus) cf inexpectatior Deuve, 1991 or davidi
Female adult
China, Yunnan, Dequen, near Haba Snow mt. 27.26'57N 99.59'00E, leg. V. Patrikeev. det. Anichtchenko A., 2020
Copyright © Alexander Anichtchenko

Cychrus (Cychrus) italicus Bonelli, 1810
Female adult
Basilicata, Pignola (PZ), 15.9.2002, 900-1200m, F. Angelini leg.
Copyright © Alexander Anichtchenko

Cychrus (Cychrus) italicus Bonelli, 1810
Male adult
N Italy, Piemonte, Alessandria, Forotondo h=1300 m, Fabbrica Curone, 6.V-16.VII.2016, G.Proscia leg., L=25 mm
Copyright © Ruslan Panin

Cychrus (Cychrus) kabaki Imura & Cavazzuti, 2017 Holotype Paratype
Male+Female adult
a: holotype, ♂; b: paratype, ♀; ca. 3 km NE. of Mt. E’erze’e［俄尔则俄］, 28°32’58”N/102°21’22”E, ca. 3900 m, on the Xiaoxiangling［小相岭］Mts., western end of Yuexi Xian［越西县］, S. Sichuan, China, 18-VII-2009. Image from Imura & Cavazzuti, 2017
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Cychrus (Cychrus) kalabianus Deuve, 2019 Holotype
Male adult
Chine, Sichuan occidental, montagnes à 17 km au nord-nord-ouest de Ganzi (= Garzê), 31º46’N/99º56’E, 4700-4800 mètres (pelouses alpines et éboulis), in coll. J. Kaláb, à Jinačovice (République Tchèque).
Copyright © Thierry Deuve

Cychrus (Cychrus) kaznakovi kaznakovi s.str.
Female adult
N Sichuan, Langmusi vill. 3700 5.8.2015 (cAA) 16.5 mm; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Cychrus (Cychrus) kaznakovi goergneri Korell & Kleinfeld, 1988
Male adult
S Gansu, 30 km E of Labrang monastery, 3700m 15.7.2012 (cAA) 14 mm; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Cychrus (Cychrus) koiwayai yunnanicola Deuve, 2018 Holotype
Male adult
Chine, Nord-Ouest du Yunnan, Monts Hengduan Shan, au nord du Mont Meili Xueshan (Kawagarbo), à l’ouest du village de Shuaqu, 3348-4494 mètres, 28°35’37’’N-98°41’25’’E, in coll. M. Janata, à Prague (République Tchèque).
Copyright © Thierry Deuve

Cychrus koltzei Roeschke, 1907
Male adult

Copyright © Kirill Makarov

Cychrus (Cychrus) koltzei Roeschke, 1907
Male adult
Russia, S Primorie, Ussuriisk distr., 3 km NE Kaimanovka vill., 22-26.8.2007, Makarov K. leg (DUBC)
Copyright © Alexander Anichtchenko

Cychrus (Cychrus) leoninus Imura & Cavazzuti, 2017 Holotype Paratype
Male+Female adult
a: holotype, ♂; b: paratype, ♀; south side of Mt. Shizi Shan［狮子山］, 27°53’29~39”N/103°14’13~15”E, 3980~4077 m, northern end of Jinyang Xian［金阳县］, S. Sichuan, China, 17 VI-2010. Image from Imura & Cavazzuti, 2017
Copyright © - - -

Cychrus (Cychrus) maoxianicus Deuve & Mourzine, 2000
Male adult
C Sichuan, Jiyuding shan, 30 km N of Wenchuan town, 3250 m 2.7.17, leg. V. Patrikeev. 11 mm; det. A. Anichtchenko, 2020
Copyright © Alexander Anichtchenko

Cychrus (Cychrus) maoxianicus Deuve & Mourzine, 2000
Male adult
C Sichuan, Jiyuding shan, 30 km N of Wenchuan town, 3250 m 2.7.17, leg. V. Patrikeev. 11 mm; det. A. Anichtchenko, 2020 (hind angles of pronotum)
Copyright © Alexander Anichtchenko

Cychrus (Cychrus) marcilhaci marcilhaci
Male adult
China, S Gansu, 7 km S Tewo town, 18.07.2012, 3000m 34.01N 103.12E, V. Patrikeev leg. (cAA). det. M. Häckel, 2019
Copyright © Alexander Anichtchenko

Cychrus (Cychrus) cf. minshanicola Deuve, 1987
Male adult
S Gansu 7 km S Tewo town 18.07.12 3000 m, 34.01N 103.12E (cAA) 15 mm; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Cychrus (Cychrus) morawitzi morawitzi
Male adult

Copyright © Kirill Makarov

Cychrus (Cychrus) naviauxi Deuve & Mourzine, 1998 (as fedotovi)
Female adult
S Gansu Min Shan 5-20km S Cagan, 2300-3200m, 7-20.7.19 (cAA) 15.5 mm; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Cychrus (Cychrus) naviauxi Deuve & Mourzine, 1998 (as fedotovi)
Male adult
S Gansu Min Shan 5-20km S Cagan, 2300-3200m, 7-20.7. (cAA) det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Cychrus (Cychrus) pangi Deuve & Tian, 2004
Male adult
C Sichuan, Jiyuding Shan, 30km N of Wenchuan, 3250m, 2.7.17 , leg. V. Patrikeev. det. A. Anichtchenko, 2020
Copyright © Alexander Anichtchenko

Cychrus (Cychrus) pangi Deuve & Tian, 2004
Male adult
C Sichuan, Jiyuding Shan, 30km N of Wenchuan, 3250m, 2.7.17 (hind angles of pronotum)
Copyright © Alexander Anichtchenko

Cychrus (Cychrus) paraxiei Cavazzuti, 2009 Holotype
Male adult
Zhongdian, Yunnan nord-occidentale, Cina sud-occidentale 27-30.06.2006
Copyright © - - -

Cychrus (Cychrus) puetzi Kleinfeld, Korell & Wrase, 1996 Holotype
Female adult
Shaanxi, Qin-Ling-Shan 108.47E/33.51N, 2500 - 2600 m
Copyright © Frank Kleinfeld

Cychrus (Cychrus) puetzi Kleinfeld, Korell & Wrase, 1996 Paratype
Male adult
Shaanxi, Qin-Ling-Shan 108.47E/33.51N, 2500 - 2600 m
Copyright © Frank Kleinfeld

Cychrus (Cychrus) reuteri Kleinfeld, 2018 Holotype Paratype
Male+Female adult
China, W.-Sichuan, Gongga Shan, 29°35’N/101°60’E, 3000-3100 m, 10/-12-VI-2018, leg.
Copyright © Frank Kleinfeld

Cychrus inessae O.Berlov, 1997 Holotype
Male adult

Copyright © Oleg Berlov

Cychrus (Cychrus) rosti belousovi Berlov, 1997
Male adult

Copyright © Igor Solodovnikov

Cychrus (Cychrus) rosti rosti
Male adult
Abkhasia, env. Dzhampal
Copyright © Igor Solodovnikov

Cychrus (Cychrus) rosti rosti
Female adult
Abkhasia: Sukhum distr., Birtzkha Mt.
Copyright © Igor Solodovnikov

Cychrus (Cychrus) rosti retezari Deuve & Prunier, 2014
Male adult
Abkhasia: Ochamchyra distr., near vill. Nardzhkheu, h = 520 m
Copyright © Igor Solodovnikov

Cychrus (Cychrus) rosti tillyi Berlov, 1997
Male adult
Abkhasia: vall. riv. Yupshara
Copyright © Igor Solodovnikov

Cychrus (Cychrus) rosti tillyi Berlov, 1997
Female adult
Abkhasia: vall. riv. Yupshara
Copyright © Igor Solodovnikov

Cychrus (Cychrus) rosti milyaevi Berlov, 1997
Male adult
Abkhasia: vall. riv. Mzara
Copyright © Igor Solodovnikov

Cychrus (Cychrus) rosti dhordaini Prunier, 2012 Holotype
Male adult
Muhazstate 70m, NE Kobuleti, Batumi, Adjarie, Georgie 13.IX.2007
Copyright © - - -

Cychrus (Cychrus) rosti retezari Deuve & Prunier, 2014
Male adult
Svanétie, environs d’Idliani (R. Boulben et P. Stéfani), rive droite de l’Enguri
Copyright © Thierry Deuve

Cychrus (Cychrus) sabdensis tatsienlu Deuve, 2013
Male adult
W Sichuan, Zeduo pass, 4000m, 20-30.6.2013, leg. V. Patrikeev (cAA). 15.5 mm; det. A. Anichtchenko, 2020
Copyright © Alexander Anichtchenko

Cychrus schmidti Chaudoir, 1837
Male adult
NW-Slovenia, Julian Alps,Mt. Mangart h=2670m, VIII.1995, Manfred Egger leg. L=15 mm
Copyright © Ruslan Panin

Cychrus (Cychrus) schmidti Chaudoir, 1837
Male adult
Slovenia Mt Canino, 08.05 (cAA) 14mm
Copyright © Alexander Anichtchenko

Cychrus (Cychrus) cf semelai Deuve, 1997
Male adult
N-W Yunnan, Dali, Cang shan 4000 m. det. M. Häckel, 2019
Copyright © Alexander Anichtchenko

Cychrus semigranosus pindicus Maran, 1940
Male adult
CE-Greece: Larissa reg., Stomio, Ossa Mt. 6-17.VII.2011, Krause Rene leg. L=21 mm;
Copyright © Ruslan Panin

Cychrus (Cychrus) semigranosus (as montanus Winkler, 1939)
Female adult
W Ukraina, Chernivtsi Reg., Hertsa Distr., 23-25.VI.2017, Kanarsky Yu. leg. L=19 mm
Copyright © Ruslan Panin

Cychrus (Cychrus) semigranosus pindicus Maran, 1940
Male adult
Greece, Thessalie, Ossa Mt., 15.7.2002 (DUBC)
Copyright © Alexander Anichtchenko

Cychrus (Cychrus) semigranosus semigranosus Palliardi, 1825
Male adult
N Ukraine, Kiev city, NP Holosiivsky, 12.VIII - 9.IX.2023, R. Nerush leg./det. R Panin, L=17 mm
Copyright © Ruslan Panin

Cychrus (Cychrus) shangrilanus Imura & Cavazzuti, 2017 Holotype Paratype
Male+Female adult
Mt. Tianbao Shan [天宝山], 27°37’05”N/99°56’50”E, 4025 m, NE. of Xiaozhongdian [小中甸], SC. Xianggelila Xian [香格里拉县], NW. Yunnan, 23-V-2015. Image from Imura & Cavazzuti, 2017
Copyright © - - -

Cychrus (Cychrus) shangrilanus Imura & Cavazzuti, 2017
Endophallus
Image from Imura & Cavazzuti, 2017
Copyright © - - -

Cychrus (Cychrus) signatus signatus
Male adult
Georgia, Adjara, env. Batumi, Makkhinjauri, abandoned house above railway tunnel, h = 107 m
Copyright © Igor Solodovnikov

Cychrus (Cychrus) sinicus Deuve, 1989
Male adult
Shaanxi, Tsinling Mts, Taibaishan, 2200m, 33.55N 107.44E, 6.2004 (cAA).
Copyright © Alexander Anichtchenko

Cychrus (Cychrus) songpanensis subtatzaopin Deuve, 2017 Holotype
Male adult
Chine, Sichuan, col entre Dawujiao et Nanping, 3600 m., 32°52’N-104°40’E, in coll. M. Janata, Prague (République Tchèque).
Copyright © Thierry Deuve

Cychrus (Cychrus) songpanensis tatzaopin Deuve, 1996
Male adult
N Sichuan, pass between Nanping and Pingwu, 3400m 20-25.6.18 (wide apice of aedeagus) (cAA) 14.5 mm; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Cychrus (Cychrus) spinicollis spinicollis
Female adult
SPAIN, Cantabria, 2 km E Puerto de San Glorio, near Vejo riv., 43.072369, -4.733061, 1300 m., 23.07.2019, A. Anichtchenko & R. Sciaky leg.
Copyright © Alexander Anichtchenko

Cychrus (Cychrus) spinicollis spinicollis
Male adult
SPAIN, Cantabria, 2 km E Puerto de San Glorio, near Vejo riv., 43.072369, -4.733061, 1300 m., 23.07.2019, A. Anichtchenko & R. Sciaky leg.
Copyright © Alexander Anichtchenko

Cychrus (Cychrus) starcki starki
Male adult
Russia, Krasnodar reg., 6 km NE Soloniki, 28-30.5.2009, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Cychrus (Cychrus) starcki zamotailovi Berlov, 1997
Male adult
Abkhasia, Gudauta distr., env. vill. Ludzava, vall. Ryaphi Riv. (Pitzunda), 10-17.VII.2011, Soodovnikov I.A.
Copyright © Igor Solodovnikov

Cychrus (Cychrus) starcki starki
Male adult
Russia: W-Caucasus, Lasorevskoe distr., vall. Psezuapse Riv.
Copyright © Igor Solodovnikov

Cychrus (Cychrus) thibetanus thibetanus
Male adult
Kangding, Garze, Sichuan, China, Xinduqiao, 30.01'53N 101.29'54E, V. Patrikeev leg. det. M. Häckel, 2019
Copyright © Alexander Anichtchenko

Cychrus (Cychrus) tubaemons Imura & Cavazzuti, 2017 Holotype Paratype
Male+Female adult
SE. of Mt. Laba Shan [喇叭山], ca. 1.7 km W. of Shangliju [上利苴], 26°45’20”N/99°37’16”E, 2755 m, WSW. of Shitoubaizu Xiang [石头白族乡] in SW. Lijinag-shi [丽江市], 25-VI.2014. Image from Imura & Cavazzuti, 2017
Copyright © - - -

Cychrus (Cychrus) tubaemons Imura & Cavazzuti, 2017
Endophallus
Image from Imura & Cavazzuti, 2017
Copyright © - - -

Cychrus tuberculatus Harris, 1839
Female adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Cychrus (Cychrus) cf. yulongxuicus Deuve, 1990
Male adult
NW Yunnan Lidjiang pref. S Yulong shan 4000 m (cAA) 14 mm; det. Anichtchenko A. 2020
Copyright © Alexander Anichtchenko

Cychrus (Cychrus) yunnanus yunnanus (= gorodinskii)
Male adult
China, 20 km SE Dequen, Yunnan, leg. V. Patrikeev (cAA). det. Anichtchenko A. 2019
Copyright © Alexander Anichtchenko

Cychrus (Cychrus) yunnanus Fairmaire, 1887 Holotype Paratype
Female adult
1: ssp. yunnanus (holotype from “Yunnan” and its labels; 2: ssp. senis Imura & Cavazzuti, 2017 (Mt. Laojun Shan, paratype); 3-5: ssp. gorodinskii (3: Jizong; 4: Xugongqingshangcun; 5: NE. of Dêqên). Image from Imura & Cavazzuti, 2017
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Cychrus (Cychrus) yunnanus Fairmaire, 1887 Holotype Paratype
Male adult
6-8: ssp. senis Imura & Cavazzuti, 2017 (6: Mt. Xuebang Shan, paratype; 7: Mt. Laojun Shan, holotype; 8: Mt. Liming-laojun Shan, paratype); 9-13: ssp. gorodinskii (9: Shanghuoshan; 10: Jizong; 11: Xugongqingshangcun; 12: Pass Baimaxue Shan Yakou; 13: NW. of Dêqên) Image from Imura & Cavazzuti, 2017
Copyright © - - -