Carabus (Leptocarabus) arboreus exarboreus (Ishikawa, 1992)
Male adult
E.Russia, Far East, Sakhalin Isl.,<br /> forest of M.Tym river<br /> 11-IX-2000 . Egorov leg.<br /> male 27mm, female 30mm.
Copyright © Safronov

Carabus (Leptocarabus) arboreus arboreus Lewis, 1882
Male adult
N-Japan, SW-Hokkaido, Chitose sity, 25.V-8.VI.2013, T. Hirabayashi leg.<br /> 32 mm
Copyright © Ruslan Panin

Carabus (Leptocarabus) arboreus pararboreus (Ishikawa, 1992)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Carabus (Leptocarabus) arboreus pararboreus (Ishikawa, 1992)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Carabus (Leptocarabus) arboreus fujisanus Bates, 1883
Male adult
Japan, CE-Honshu, Yamanashi pref., Mt. Fujisan h~1600m, 23.IX.1982, Y. Imura leg. L=23 mm
Copyright © Ruslan Panin

Carabus (Leptocarabus) arboreus fujisanus Bates, 1883
Female adult
Japan, Honshu, Mt. Fuji h~1900m, Shizuoka pref., Fujinomiya city, 23.VI.2007, M. Matsunaga leg., L=25 mm
Copyright © Ruslan Panin

Carabus (Leptocarabus) arboreus akitanus (Ishikawa, 1992)
Male adult
Muriyoshi (Akita), JAPON, VIII-2001; 26 mm
Copyright © Alain Mollard

Carabus (Leptocarabus) arboreus babai Ishikawa, 1968
Male adult
Hiychiyama (Niigata, Honshu), JAPON, XI-1984; 20 mm
Copyright © Alain Mollard

Carabus (Leptocarabus) arboreus exilis Bates, 1883
Male adult
Sado Isle ( Honshu), JAPON, X-1994; 27 mm
Copyright © Alain Mollard

Carabus (Leptocarabus) arboreus gracillimus Bates, 1883
Female adult
Mt.Tate-yama, 2000m (Toyama, Honshu), JAPON, X-1994; 20 mm
Copyright © Alain Mollard

Carabus (Leptocarabus) arboreus karibanus (Ishikawa, 1992)
Female adult
Kuromatsunai (Hokkaido), JAPON, X-1994; 30 mm
Copyright © Alain Mollard

Carabus (Leptocarabus) arboreus kitakamisanus (Ishikawa, 1992)
Male adult
Niisato (Iwate, Honshu), JAPON, VIII-2001; 30 mm
Copyright © Alain Mollard

Carabus (Leptocarabus) arboreus nepta (Ishikawa, 1984)
Male adult
North, Hakkoda-san(Aomori, Honshu), JAPON, VI-1986 ; 25 mm
Copyright © Alain Mollard

Carabus (Leptocarabus) arboreus parexilis Nakane, 1961
Male adult
Towada-Cho (Honshu), JAPON, X-1990; 30 mm
Copyright © Alain Mollard

Carabus (Leptocarabus) arboreus ogurai Ishikawa, 1969
Female adult
Tabayama (Yamanashi, Honshu), JAPON, X-1990; 24 mm
Copyright © Alain Mollard

Carabus (Leptocarabus) arboreus pronepta (Ishikawa, 1992)
Male adult
Hakkode-san (Aomori, Honshu), JAPON, XI-1990; 24 mm
Copyright © Alain Mollard

Carabus (Leptocarabus) arboreus shimoheiensis (Ishikawa, 1992)
Female adult
Kitakami (Iwate, Honshu), JAPON, XI-1986; 23 mm
Copyright © Alain Mollard

Carabus (Leptocarabus) arboreus shinanensis Born, 1922
Male adult
Mt. Yatsugatake (Nagano, Honshu), JAPON, VII-1985; 23 mm
Copyright © Alain Mollard

Carabus (Leptocarabus) arboreus tenuiformis Bates, 1883
Female adult
Mt. Osaki-Yama,1900m (Tochigi, Honshu), JAPON, VI-1988; 26 mm
Copyright © Alain Mollard

Carabus (Leptocarabus) harmandi fujisan Ishikawa, 1972
Female adult
Japan, Yamanashi pref., Fuji-Rinds 1600m, Narusawa vill. 3.V.2005 Kurihara K. leg. 25 mm
Copyright © Alexander Anichtchenko

Carabus (Leptocarabus) harmandi harmandi Lapouge, 1909 Lectotype
Male adult
Honshu, Tochigi, Mts near Nikko. Muséum national d’Histoire naturelle, Paris (France) Collection: Insects - Coleoptera (EC) Specimen MNHN-EC-EC549
Copyright © MNHN

Carabus (Leptocarabus) hiurai Kamiyoshi & Mizoguchi, 1960
Male adult
Collection ground Hojyo city, Ehime Pref. 34358
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Carabus (Leptocarabus) koreanus Reitter, 1895
Male adult
Habitus of A. koreanus ssp. and male aedeagus tips: A. koreanus koreanus; B: A. koreanus minor; C: A. koreanus mineti; D: A. koreanus rectokoreanus (Scale bars: 1 cm). Image from T. Kwon et al 2023
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Carabus (Leptocarabus) kumagai kumagai Kimura & Komiya, 1974
Male adult
Collection ground Otsu city, Shiga Pref. 35330
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Carabus (Leptocarabus) kyushuensis cerberus Ishikawa, 1962
Female adult
Fuchu (Mitsugi,Hiroshima) JAPON, VII-2001. 34 mm
Copyright © Alain Mollard

Carabus (Leptocarabus) kyushuensis nakatomii Ishikawa, 1966
Male adult
Tombara (Honshu,Shimane) JAPON, VII-2001. 36 mm
Copyright © Alain Mollard

Carabus (Leptocarabus) kyushuensis kyushuensis Nakane, 1960
Female adult
Izumi, (Honshu, Kumamoto) JAPON, VII-2001. 31 mm
Copyright © Alain Mollard

Carabus (Leptocarabus) procerulus Chaudoir, 1862
Male adult
Japan, Yamanashi pref., Otawa-rindo,
Narusawa vill., 9.IX.2002 T. Kurihara leg
male, 31 mm
Copyright © Alexander Anichtchenko

Carabus (Leptocarabus) procerulus Chaudoir, 1862
Endophallus
Japan:S Honshu , Matsubara-ko,
Copyright © Ruslan Panin

Carabus (Leptocarabus) procerulus Chaudoir, 1862
Male adult
Japan:S Honshu, Matsubara-ko, VIII.1997, S. Remond leg.
L=30 mm
Copyright © Ruslan Panin

Carabus (Leptocarabus) procerulus Chaudoir, 1862
Endophallus
Japan, S Honshu, Mie Pref., Hakusan,
Copyright © Ruslan Panin

Carabus (Leptocarabus) procerulus Chaudoir, 1862
Male adult
Japan, S Honshu, Mie Pref., Hakusan, VII.2001, S. Remond leg.<br /> L=33 mm
Copyright © Ruslan Panin

Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931
Male adult
NE-China:Liaoning prov., Fengcheng City, Miaoyang, 12-17.VI.2011
L=26 mm
Copyright © Ruslan Panin

Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931
Endophallus
NE-China:Liaoning prov.,Fengcheng City, Miaoyang,
Copyright © Ruslan Panin

Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931
Male adult
NE-China:E-Liaoning: NP Kuandian Tianqiaogou , VI.2010, L=25 mm, det. Panin R.
Copyright © Ruslan Panin

Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931 Neotype
Male adult
Chongjin Muséum national d’Histoire naturelle, Paris (France). Collection: Insects - Coleoptera (EC). Specimen MNHN-EC-EC518
Copyright © MNHN

Carabus (Leptocarabus) seishinensis seunglaki (Kwon & Lee, 1984)
Male adult
Taebaeck-san (Kangwon-sud) KOREE, VII-1998; 21 mm
Copyright © Alain Mollard

Carabus (Leptocarabus) semiopacus Reitter, 1895
Female adult
N Korea, Nangnim, 14.VII.1981
female, long. 24 mm
Copyright © Alexander Anichtchenko

Carabus (Leptocarabus) semiopacus sstr (left); semiopacus gwangyangensis (right)
Male adult
Image from T. Kwon et al (2023)
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Carabus (Leptocarabus) semiopacus Reitter, 1895
Male adult
1A and 3B: A. (A.) semiopacus semiopacus; 2A and 4B: A. (A.) semiopacus gwangyangensis Kwon & Park, 2023 (A: right lateral view; B: left and right basal lobes), (Scale bars: 1 mm). Image from T. Kwon et al, 2023
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Carabus (Leptocarabus) vogtianus cf horvatovichi Deuve, 1992
Male adult
R. of Korea, Geonggido/Gangwondo provincial border, Pass North from Dopyeong, 600 m, 3/13.VII.2012, I. Rapuzzi and L. Caldon leg. Image from Rapuzzi, 2015
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Carabus (Leptocarabus) vogtianus horvatovichi Deuve, 1992
Male adult

Copyright © Jong-Bong Choi

Carabus (Leptocarabus) vogtianus horvatovichi Deuve, 1992 Holotype
Male adult
Image from: Korean type specimens of insects deposited in Hungarian Natural History Museum.
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Carabus (Leptocarabus) yokoae marcilhaci Deuve, 1992
Male adult
CHINA, Shanxi, Tschingling Mts., Taibeichan Nat. reserv.;
Copyright © Anton Menstschikov

Carabus (Leptocarabus) yokoae marcilhaci Deuve, 1992
Male adult
China, Shanxi, Taschingling Mts., Taibeishan Nat. res.
2000 m., 5-15.08.2005
Copyright © Alexander Anichtchenko

Carabus (Leptocarabus) yokoae xunyangbanus Imura, 1998
Male adult
Qinling Shan, (Xi’an, Shanxi) CHINE VI-1992; 27 mm
Copyright © Alain Mollard

Carabus (Leptocarabus) yokoae nanjiangensis Imura & Su, 2000
Female adult
Gangwu Shan (Taoyuan, Sichuan) CHINE VI-1992; 27 mm
Copyright © Alain Mollard