Pterostichus (Adelopterus) ambiguus (Fairmaire, 1858)
Female adult
France, 20224, Corse, Calacuccia forêt de Valdu Niellu 12-09-2000
Copyright © Pascal Deschamps

Pterostichus (Adelosia) macer Marsham, 1802
Male adult

E. Ukraine,
Kharkov reg., 1972
16mm.

Copyright © Ruslan Panin

Pterostichus (Anilloferonia) malkini (Hatch, 1953)
Male adult
Image from Labonte, 2013
Copyright © - - -

Pterostichus (Anomostichus) anomostriatus Sciaky, 1995 Paratype
Male adult
China – pr. Sichuan, Emei mt. 2500 4-20.V.1989 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Pterostichus (Aphaonus) compressus Rost, 1892
Male adult
Russia, Krasnodar reg., 6 km NE Soloniki vill.,
Sochi Nat.Park, 28-30.V.2009 Chibilov E. leg.
photo A. Anichtchenko
det.I.A.Solodovnikov
Copyright © Alexander Anichtchenko

Pterostichus (Aphaonus) cylindriformis Reitter, 1887e: 252
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Aphaonus) mariamae Zamotajlov, Solodovnikov & Fominykh, 2015 Holotype
Male adult
Abkhazia, Gulripsh Distr., Tsebelda Plateau near village Ablukhvara, left bank of River Kelasur, slopes of Mt. Bashlug, 650 m, 15.05.-19.07.2012, pitfall traps (D. Fominykh) (ZISP).
Copyright © Kirill Makarov

Pterostichus (Argutor) commutabilis (Motschulsky, 1866)
Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada

Copyright © Henri Goulet

Pterostichus (Argutor) leonisi Apfelbeck, 1904: 257
Male adult

Copyright © Igor Solodovnikov

Pterostichus (Argutor) leonisi Apfelbeck, 1904: 257
Female adult

Copyright © Igor Solodovnikov

Pterostichus (Asioplatysma) medvedevi Kryzhanovskij & Mikhailov, 1972: 319
Male adult

Copyright © Kirill Makarov

Pterostichus (Badistrinus) laticollis (Motschulsky, 1844)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Badistrinus) haptoderoides haptoderoides
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Bothriopterus) subovatus (Motschulsky, 1860)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Bothriopterus) oblongopunctatus (Fabricius, 1787)
Male adult

Copyright © Kirill Makarov

Pterostichus (Bothriopterus) aeneocupreus (Fairmaire, 1887)
Male adult
Bhutan, near Gangcheyting, [Camp 2] 3689 m, 27° 24 14.0''N, 90° 14' 04.2''E, 9.oct. 2017
Copyright © Joachim Schmidt

Pterostichus (Calopterus) pilosus (Host, 1789)
Male adult

Ukrainian Carpathians,
Transcarpathian Reg., Rakhov Distr., Chornohora range h~1850m, 29.VI-28.VII.2007, Panin R. leg.
male,18 mm.

Copyright © Ruslan Panin

Pterostichus (Calopterus) selmanni prevostii (Dejean, 1828)
Male adult
France, Jura, Lajoux, 1190m, 20/05/2013, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Pterostichus (Calopterus) selmanni roubali Schauberger, 1927
Female adult
CZ; SW Bohemia, Šumava Mts.; 16mm
Copyright © Aleš Sedláček

Pterostichus (Calopterus) pilosus pilosus s.str.
Female adult
CZ; NE Moravia, Štramberk; 15mm
Copyright © Aleš Sedláček

Pterostichus (Calopterus) pilosus pilosus (Host, 1789)
Male adult
W Ukraine, NE Carpathians,, Chornagora Mt Range, h=2000m, alpin zone, 11.VI-1.IX.2015, Panin R. leg 16 mm
Copyright © Ruslan Panin

Pterostichus (Cheporus) burmeisteri Heer, 1838
Male adult

NW Ukrainian Carpathians,
Lviv Prov., Staryi Sambor Distr., near Dobromyl , h=450m 9.VI-25.VII.2007 Panin R. leg.
male,15 mm.

Copyright © Ruslan Panin

Pterostichus (Cheporus) burmeisteri baldensis Schaum, 1862
Female adult
NE Italy, Veneto, Mts Lessini, Vallone del Malera h=1500 m., 28-30.V.2019, Gontarenko A. leg./det.Panin R. L=14 mm
Copyright © Ruslan Panin

Pterostichus balthasari Jedlička, 1937 Lectotype
Male adult
(NMPC) Image from Dorjderem S, Shi H, Liang H., 2020
Copyright © - - -

Pterostichus (Circinatus) bullatus Allegro & Sciaky, 2010
Male adult

Copyright © Gianni Allegro

Pterostichus (Circinatus) dentifer Allegro & Sciaky, 2010 Holotype
Male adult
China, C Sichuan, Dayi distr., / Chadiping env., 1500 m / 5–7.08.1996 / A. Zamotajlov, A. Miroshnikov & / D. Fedorenko. Image from Hongliang Shi & Hongbin Liang, 2015
Copyright © - - -

Pterostichus (Circinatus) bullatus Allegro & Sciaky, 2010
Female adult
Image from Hongliang Shi & Hongbin Liang, 2015
Copyright © - - -

Pterostichus (Circinatus) beneshi Sciaky, 1996 Holotype
Male adult
China – W Sichuan, Wolong 150 km NW Chengdu, 9.VII.1994 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Pterostichus (Circinatus) tonkinensis Straneo, 1980
Male adult
Vietnam, 40 km W of Cao Bang, Phia Oac Mt., 22°36´50´´N 105°52´0´´E, h~1800 m, deciduous bamboo forest, 3–11.X.2018 (D. Fedorenko)
Copyright © Dmitri Fedorenko

Pterostichus (Cophosus) cylindricus (Herbst, 1784)
Female adult
CZ; S Moravia, Břeclav; 20mm
Copyright © Aleš Sedláček

Pterostichus (Cryobius) kaninensis Poppius, 1906
Male adult

Copyright © Kirill Makarov

Pterostichus (Cryobius) pinguedineus Eschscholtz, 1823
Male adult

Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada

Copyright © Henri Goulet

Pterostichus (Cryobius) middendorffi J.R. Sahlberg, 1875: 103
Female adult

Copyright © Kirill Makarov

Pterostichus (Cryobius) parasimilis G.E. Ball, 1962: 6
Male adult
det. K. Makarov
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) middendorffi J.R. Sahlberg, 1875: 103
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Cryobius) quadraticollis Chaudoir, 1846a: 140
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Cryobius) abaxoides abaxoides (Dejean, 1828)
Male adult
Spain, C Pirineos, Pleta Llardana, 2600m, 23.8.2005, Anichtchenko A. leg. & det. ; 10mm
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) pumilio (Dejean, 1828)
Male adult
Spain, Navarra, Sierra de Aralar, 9.5.2006, Altxueta, 1000m, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) aralarensis Espanol & Mateu, 1945
Female adult
Spain, Mts, Cantabricos, Soto de Sajambre, Canto Cabronero Mt., 1850m, 12.VIII.2008, Anichtchenko A. leg. & det. 7.5mm
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) homalonotus sojot Shilenkov, 2000 Paratype
Female adult
Бурятия, Восточный Саян, окр. влк. Кропоткина 2050м, 10-15.7.98, Aнищенко А. leg. (cAA) 7.8mm [Buryatia, W Sayan mts., volcan Kropotkin vic., Anichtchenko A. leg]
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) lucidus Motschulsky, 1844
Male adult
Hamar-Daban, Malyi Zangisan riv., Baishint-Ula Mt., 2300-2500m, 21.5.1999, Anichtchenko A. leg. (cAA). det. V. Shilenkov, 1999
Copyright © Alexander Anichtchenko

Pterostichus (Cryobius) fulvescens (Motschulsky, 1844)
Male adult
Burjatia, W Sajan Mts., Tunkinskie Goltsy, Mondy vill. vic., 1500m, 1-7.7.2001, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Pterostichus (Cylindrocharis) rostratus (Newman, 1838)
Male adult

Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada

Copyright © Henri Goulet

Pterostichus (Eosteropus) aethiops (Panzer, 1796)
Female adult
W-Ukraine, Lviv reg., Zhydachiv distr., near Vybranivka vill., forest, under the bark, Panin R. leg. 12.X.2007 female,14mm
Copyright © Ruslan Panin

Pterostichus (Eosteropus) rufitarsis cordatus Letzner, 1842
Female adult
Ukrainian Carpathians, Transcarpathian reg., Rakhov Distr., Chornogora range, h=2000m., Panin R. leg., 29.VI-28.VII.2007 L=12 mm
Copyright © Ruslan Panin

Pterostichus (Eosteropus) discrepans A.Morawitz, 1862
Female adult

Copyright © Kirill Makarov

Pterostichus (Eosteropus) orientalis (Motschulsky, 1844)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Eosteropus) tuberculiger tuberculiger
Male adult
S Primorie, Ussuriisk distr., 3km NE Kaimanovka vill. 22-25.8.2007, K. Makarov A.Zaitsev leg. (DUBC) 13mm
Copyright © Alexander Anichtchenko

Pterostichus (Ethira) pilifer (Bates, 1878)
Male adult
Natural History Museum (2014). Dataset: Collection specimens. Resource: Specimens. Natural History Museum Data Portal (data.nhm.ac.uk). © The Trustees of the Natural History Museum, London
Copyright © - - -

Pterostichus (Falsargutor) pseudopedius pseudopedius Reitter, 1887
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Feronidius) ? hungaricus (Dejean, 1828)
Male+Female adult
W. UKRAINE, Transcarpathian region, Mukachevo Distr., riverine forest near Znyatsevo vill., 16-31.VII.2008, Panin R. leg., female, 16 mm
Copyright © Ruslan Panin

Pterostichus (Feronidius) melas (Creutzer, 1799)
Male adult

W-Ukraine,
Lviv Region, Zolochiv distr., near Gologirky, in steppe, 16-29.V.2007 Panin R. leg.
long-17mm

Copyright © Ruslan Panin

Pterostichus (Feronidius) fornicatus (Kolenati, 1845)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Feronidius) hungaricus (Dejean, 1828)
Male adult
Romania, Mts Trascau, Rimetea env., Mt Piatra Secuiului h~1100 m, 27.V-11.VIII.2008, I.Retezár & B.Tallosi leg. L=15 mm
Copyright © Ruslan Panin

Pterostichus (Gutta) kongshuhensis Gueorguiev, 2015 Holotype
Male adult
CHINA, Yunnan Prov., Tengchong County, vil. Kongshuhe near cave Wu Shi Shan 1-2-3 N25.73111 E98.66459, ca. 2500-2600m, 11.11.2013, B. Petrov leg.
Copyright © Borislav Gueorguiev

Pterostichus (Haplomaseus) caucasicus (Menetries, 1832)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Huaius) hanwang Tian & He, 2020 Holotype
Male adult
cave Hanwang Dong, Zhongbai, Zengjia, Chaotian, Guangyuan, Sichuan (四川省广元市朝天区曾家镇中柏村汉王洞), 32°34′40.14″N, 106°6′25.32″E, 1210 m, IX-22-2018, leg. Li He (SCAU). Image from Tian & He, 2020
Copyright © Mingyi Tian

Pterostichus (Hypherpes) tristis (Dejean, 1828)
Male adult

Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada

Copyright © Henri Goulet

Pterostichus (Hypherpes) lama (Menetries, 1843)
Female adult
USA, WA, Clark Po near Vancouver Lake forest, 2.VI.2012. 25 mm
Copyright © Alexander Anichtchenko

Pterostichus (Hypherpes) lama (Menetries, 1843)
Male adult
USA: Washington, 34 mm
Copyright © Igor Solodovnikov

Pterostichus (Hypherpes) lama (Menetries, 1843)
Female adult
USA: Washington
Copyright © Igor Solodovnikov

Pterostichus (Koreonialoe) microps Heyden, 1887
Male adult

Copyright © Kirill Makarov

Pterostichus (Koreonialoe) micropoides Yin, Shi & Liang, 2021 Holotype Paratype
Male+Female adult
Paratype from Hunchun city, Jilin
Copyright © - - -

Pterostichus (Koreonialoe) ishikawai Nemoto, 1988
Female adult
Sud Korea, Juwangsan National Park
Copyright © Vittorino Monzini

Pterostichus (Galapterus) galae Farkac & Plutenko, 1996
Male adult

Far East, S Sikhote-Alin, Snezhnaya Mt., source of Ussuri Riv., h = 1400 -1682 m. 1-6.07.2004 Yu.Sundukov leg.
male
L=13,0 mm

Copyright © Maksim Smirnov

Pterostichus (Lenapterus) punctatissimus (Randall, 1838)
Male adult

Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada

Copyright © Henri Goulet

Pterostichus (Lenapterus) vermiculosus (Menetries, 1851)
Male adult

Copyright © Kirill Makarov

Pterostichus (Lenapterus) agonus G.Horn, 1880
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Lenapterus) subrugosus Straneo, 1955
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Leptoferonia) pumilus Casey, 1913
Female adult
Oregon, Portland

Copyright © Igor Sokolov

Pterostichus (Leptoferonia) hackerae Will, 2016 Holotype
Male adult

Copyright © Kipling W. Will

Pterostichus (Lianoe) dufourii boisgiraudii L. Dufour, 1843
Male adult
France, Pyrenee Ariege, Vall. du Biros, Eylie Cirque de la Plagne, 1100-1500m, 20.6.1993 (DUBC). det. Anichtchenko A. 2021
Copyright © Alexander Anichtchenko

Pterostichus (Lianoe) arribasi Ortuño, 1991
Male adult
Ezcaray, 16.8.97, Valdezcaray, La Rioja, Spain, M. Toribio leg. & det. (cAA). 15 mm
Copyright © Alexander Anichtchenko

Pterostichus (Licentius) cuii Tian & He, 2021 Holotype
Male adult
cave Tangren Dong, Beijiao Cun, Fozizhuang Xiang, Fangshan Qu, Beijing, 39° 47ʹ42ʺN 115°51ʹ55ʺE, 324 m, 20.XII.2020, leg. Qingwu Cui (SCAU).
Copyright © Mingyi Tian

Pterostichus (Lyropedius) lyroderus (Chaudoir, 1846)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Lyrothorax) caspius (Menetries, 1832)
Male adult
CZ, SE Moravia, Břeclav; 11mm
(introduced)
Copyright © Aleš Sedláček

Pterostichus (Incertae) zhejiangensis Kirschenhofer, 1997 Holotype
Male adult

Copyright © Kipling W. Will

Pterostichus (Melanius) castor Goulet & Bousquet, 1983
Male adult

Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada

Copyright © Henri Goulet

Pterostichus (Melanius) aterrimus aterrimus
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Melanius) aterrimus aterrimus (Herbst, 1784)
Male adult
W Ukraine, Lviv, Holosko, bank of Natural lake, 24.V-8.VII.2016, Panin R. leg. L=15 mm.
Copyright © Ruslan Panin

Pterostichus (Melanius) elongatus (Duftschmid, 1812)
Female adult
Marruecos, Gada de Debdou, 9.6.1993 (cAA)
Copyright © Alexander Anichtchenko

Pterostichus (Metallophilus) orion (Tschitscherine, 1901)
Male adult

NE Chita reg., Kodar Mt rng., Chara riv.
15 km N Leprindo, 2200m 5-15.7.2001 Brinev leg.
14 mm

Copyright © Alexander Anichtchenko

Pterostichus (Metallophilus) mirus (Tschitscherine, 1894)
Female adult
Russia, Siberia, SW Tuva, pass Barlyk, 2100m, 10.VII.2010
14 mm
Copyright © Alexander Anichtchenko

Pterostichus (Metallophilus) interruptus (Dejean, 1828)
Male adult

Copyright © Kirill Makarov

Pterostichus (Metallophilus) rugosus (Gebler, 1823)
Male adult
Russia, Chita reg., Malye Kovali vill. vic., near Chichatka riv., 22-29.7.1999, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Pterostichus (Micronialoe) chogyesanus Park; Kwon & Lafer, 1996
Male adult
male
Copyright © Dooyoung Kim

Pterostichus (Micronialoe) spinulosus Kim & Suh, 2024 Holotype
Male adult

Copyright © Dooyoung Kim

Pterostichus (Monoferonia) diligendus (Chaudoir, 1868)
Male adult

Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada

Copyright © Henri Goulet

Pterostichus (Feroperis) vladivostokensis Lafer, 1979 : 13
Male adult

Copyright © Kirill Makarov

Pterostichus (Morphnosoma) petulans Jedlicka, 1938
Male adult
Russia, Primorskii krai, Shkotovskii distr., Anisimovka vill., 17-26.6.2008 (DUBC). 15 mm. det. A. Anichtchenko, 2021
Copyright © Alexander Anichtchenko

Pterostichus (Morphnosoma) jungens (Tschitscherine, 1893)
Male adult
Russia, S Primorie, Chernye Mts., valley of Poima riv., 30.7.1999, A. Anichtchenko leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Pterostichus (Morphnosoma) melanarius cardioderus (Chaudoir, 1846)
Female adult
Georgia, Racha distr., near Okureshi vill, Khvamli Mt h=1000-1100 m, 19-24,VI.2024, I.Uspenskyi leg./det. R Panin, L=22 mm
Copyright © Ruslan Panin

Pterostichus (Morphohaptoderus) schuelkei Sciaky & Wrase, 1997: 1102 Holotype
Female adult
body length = 10.8 mm, “CHINA (Shaanxi) / Qin Ling Shan / 108.47 E / 33.51 N / Mt. W pass / autoroute km 70,47km. S Xian 2350–2500 m / 7.–19.VII.1996 / Kleinfeld & Schutze”; “COLL.WRASE / BERLIN”; “HOLOTYPUS / Pterostichus / (Circinatus)/ schuelkei sp. n. / Sciaky & Wrase des. 1997” [red label]
Copyright © Hongliang Shi

Pterostichus (Morphohaptoderus) saueri Sciaky, 1994
Male adult
China, Sichuan, Ganzi pref, Daxue Shan 102.00E 30.03N 5km E Kangding, river valley 3000m 20-23.5.1997 leg A. Putz (cAA) 6.5 mm
Copyright © Alexander Anichtchenko

Pterostichus (Morphohaptoderus) glabellus Fedorenko, 2023 Paratype
Female adult
N[orthern] Vietnam, Lao Cai Prov[ince]./ Hoang Lien Son Mt.ridge,/ env. Fansipan [= Phan Xi Pang] Mt, Tram Ton/ h = 1950-2100 [m],/ 15-30./ leg. VII.2007
Copyright © Dmitri Fedorenko

Pterostichus (Morphohaptoderus) unisetosus Fedorenko, 2023 Holotype
Male adult
(ZMMU): N-Vietnam, 40 km WNW of/ Lao Cai, env. Y Ty, Bat Xat/ Nat[io]n.[al] Park, h=1600-1900 m/ 22°37´-37.6´N/ 103°37.3´-38.3´E, 16-21./ leg. D. Fedorenko X.2018
Copyright © Dmitri Fedorenko

Pterostichus (Morphohaptoderus) huashanus Sciaky, 1994
Female adult
7.5 mm (cAA)
Copyright © Alexander Anichtchenko

Pterostichus (Myosodus) rudestriatus (Reitter, 1883)
Female adult
S Russia, NW-Caucasus, Karachay-Cherkessia, Teberda, Alibek Mountain Pass h=2800 m, 23.VII.2010, Solodovnikov I.A leg. <br L=16 mm</p>
Copyright © Ruslan Panin

Pterostichus (Myosodus) lacunosus roubalianus (Lutshnik, 1928)
Female adult
S Russia, W Caucasus, <br /> Tuapse distr., Kashina (=Lysaya) Mt. h=1300m., 10.V.2006, Solodovnikov I.A. leg.<br /> Female, 18 mm
Copyright © Ruslan Panin

Pterostichus (Myosodus) schoenherri Faldermann, 1836
Male adult
SW Georgia, Lesser Caucasus, Adzhar-Imereti (Meskheti) Rng, Zekari Pass env. h~2100-2200m, 26-30.VI.2006, A.Puchkov leg. L=17 mm
Copyright © Ruslan Panin

Pterostichus (Myosodus) zamotajlovi Belousov, 1991
Male adult
Russia, Karachaevo-Cherkessia, N slopes of Sophia Mt., upper of Sophia riv., h = 2700 m, 07-25.VII.2020 leg. A.P. Tatulyan
Copyright © Igor Solodovnikov

Pterostichus (Myosodus) aibgensis adelaidae Reitter, 1899
Male adult

Copyright © Igor Solodovnikov

Pterostichus (Myosodus) aibgensis adelaidae Reitter, 1899
Female adult

Copyright © Igor Solodovnikov

Pterostichus (Myosodus) sodalicius Heyden, 1885
Female adult
Uzbekistan, Chatkal Mt. rng., Irtash, 41°10'25.7"N 70°16'20.4"E, 5-7.8.2021, 2400m, A. Anichtchenko leg. & det. (cAA). 15mm
Copyright © Alexander Anichtchenko

Pterostichus (Myosodus) schoenherri Faldermann, 1836
Female adult
Georgia, Lesser Caucasus, Imereti prov., Meskheti Mt Rng, N slope, Zekari Pass h~2150 2220 m alpine meadows 8-9.VII.2021, E.Rujan leg./det. R. Panin
Copyright © Ruslan Panin

Pterostichus (Myosodus) lacunosus (Chaudoir, 1844)
Male adult
S Georgia, Samtskhe-Javakheti, Akhaltsikhe distr., Atskuri h=1200-1350 m, 1-7.VII.2013, E.Rutjan leg., L=19.5 mm
Copyright © Ruslan Panin

Pterostichus (Neohaptoderus) catei Sciaky & Wrase, 1997 Paratype
Male adult

China,Shaanxi, Qin Ling Shan, 110.06E 34.27N Hua Shan Mt., N Valley, 1200-1400m 118 km E Xian, sifted 18-20.08.1995 A.Pütz leg.
10 mm
photo & coll. Anichtchenko A. 2009

Copyright © Alexander Anichtchenko

Pterostichus (Neohaptoderus) austrinus Fedorenko, 2023 Holotype
Male adult
(ZMMU), labelled: N-Vietnam, 40km W of/ Cao Bang, Phia Oac Mt./ 22°36´25N 105°52´08E/ h~1650-1700 m,/ deciduous forest, 3-11.X./ leg. D.Fedorenko 2018
Copyright © Dmitri Fedorenko

Pterostichus (Neohaptoderus) ignavus (Tschitscherine, 1897)
Male adult
(cAA). 9.5 mm
Copyright © Alexander Anichtchenko

Pterostichus (Daisenialoe) fujimurai fujimurai s.str. Holotype
Male adult
Mt. Daisen, Tottori Pref., Japan, 14. VII. 1956, T. Fujimura
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-041
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Nialoe) fujisanus Tanaka & Suga, 1972 Holotype
Male adult
Mt. Fuji (south side), 1400 m, Shizuoka Pref., Japan, 31. VII. 1971, K. Tanaka & S. Suga
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-253
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Oreophilus) variolatus variolatus Dejean, 1828
Male adult
S Slovenia, Ilirska Bistrica, Snežnik Mts,15-28.VI.2009, Manfred Egger leg.
L=19 mm

Copyright © Ruslan Panin

Pterostichus (Oreophilus) multipunctatus (Dejean, 1828)
Female adult

Copyright © Kirill Makarov

Pterostichus (Oreophilus) externepunctatus Dejean, 1828
Female adult
NW-Italy, Piemonte prov.,Cottian Alps, Mt Albergian, 11.VI.1979, S. Dacatra leg., L=15 mm
Copyright © Ruslan Panin

Pterostichus (Oreophilus) jurinei (Panzer, 1803)
Female adult

Copyright © Kirill Makarov

Pterostichus (Oreophilus) bicolor Aragona, 1830
Male adult
H. Penna, Senova, 1400m 18.6.1978 (cAA)
Copyright © Alexander Anichtchenko

Pterostichus (Oreophilus) impressus (Fairmaire & Laboulbene, 1854)
Male adult
France, Orvault, Loire-Atlantique, 16.12.1962 [label error] (DUBC) 17 mm
Copyright © Alexander Anichtchenko

Pterostichus (Oreoplatysma) cecchiniae Jakobson, 1907
Male adult

Russia, Krasnodar reg., 6 km NE Soloniki vill.,
28-30.V.2009 Anichtchenko A. leg.
male, long. 10 mm

Copyright © Alexander Anichtchenko

Pterostichus (Oreoplatysma) koenigi (Reitter, 1887)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Orientostichus) pulcher Sciaky & Allegro, 2013 Holotype Paratype
Male+Female adult
Label of Holotype male: China, S Sichuan, Ebian, Yizhi Yakou, Xiaoliang Shan-Si He, 2800-3100 m, 19.VII.2007, Cavazzuti P.F. leg. (IZAS). Scale bar 5 mm
Copyright © Gianni Allegro

Pterostichus (Orientostichus) perlutus Jedlicka, 1938a: 5
Male adult
W Sichuan, pass W Kangding, Zheduo mts, 4200m, 12.7.2013, V. Patrikeev leg. (cAA) 16 mm; det. H. Shi, 2020
Copyright © Alexander Anichtchenko

Pterostichus (Orientostichus) jialini Yin, Zhu & Shi, 2023 Holotype
Male adult
Image from Yin, Zhu & Shi, 2023
Copyright © - - -

Pterostichus (Orientostichus) pemphis Yin, Zhu & Shi, 2023 Holotype
Male adult
Image from Yin, Zhu & Shi, 2023
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Pterostichus (Orsonjohnsonus) johnsoni Ulke, 1889
Male adult

USA, Oregon, Clackamas County

Copyright © Igor Sokolov

Pterostichus (Paraferonia) lubricus LeConte, 1852
Male adult

Great Smoky Mountains National Park

Copyright © Igor Sokolov

Pterostichus (Paranialoe) biexcisus subparallelus Straneo, 1955
Male adult
Nishinoshima Is., Mita (Mt. Takuhi-yama). Image from Takashi Shimizu and Ryo Nakamura, 2023
Copyright © - - -

Pterostichus (Parapterostichus) grajus (Dejean, 1828)
Male adult
Italia, Oropa (Bi) Lago Mucrone, 14.6.2003 S. Dacatra leg. (cAA) 12 mm
Copyright © Alexander Anichtchenko

Pterostichus (Petrophilus) tomensis (Gebler, 1847)
Male adult

Kemerovo Reg., env. Osinniki, source of Krasnaya Kaltank Riv., Populus tremulae-fores, W slope , h = 450 м 28.V-6.VI.03 Yu.Sundukov leg.
L=18,0 mm

Copyright © Maksim Smirnov

Pterostichus (Petrophilus) turanensis Jedlicka, 1959
Male adult
Russia, E Sayan Mts., Tunkinskie Goltsy, Kyngarga riv., 2200 m, 18-23.VII.1995, Anichtchenko A. leg. & det.
Copyright © Alexander Anichtchenko

Pterostichus (Petrophilus) montanus (Motschulsky, 1844)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Petrophilus) tundrae (Tschitscherine, 1894)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Petrophilus) triseriatus (Gebler, 1847)
Male adult
Tyva, Kyzyl-Taiga mt. rng., 1500m, Ak-Sug riv. 21.6.2006, A. Menshikov leg. (cAA) det. A. Anichtchenko, 2021; 10.5 mm
Copyright © Alexander Anichtchenko

Pterostichus (Petrophilus) subaeneus (Chaudoir, 1850)
Male adult
Russia, E Siberia, Irkutsk reg., Nizhneudinsk distr., 9km SW Telezhnyi vill., 1040m, 28.7.1998, A. Anichtchenko leg & det. (cAA)
Copyright © Alexander Anichtchenko

Pterostichus (Petrophilus) septentrionis (Chaudoir, 1868)
Male adult
E Siberia, Hamar-Daban mts., Osinovka riv., 2000m, 28-31.7.1995, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Pterostichus (Phonias) corrusculus Leconte, 1873
Male adult

Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada

Copyright © Henri Goulet

Pterostichus (Phonias) eobius (Tschitscherine, 1899)
Male adult

Copyright © Kirill Makarov

Pterostichus (Phonias) longinquus Bates, 1873
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Phonias) strenuus (Panzer, 1796)
Female adult
Russia, Moscow det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Phonias) diligens (Sturm, 1824)
Male adult
Russia, Moskow det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Phonias) jankowskyi (Tschitscherine, 1897)
Female adult
Russian Far East, Primorskii krai, Khasanskii district, 9 km SSE Barabash, middle course of Kedrovaya River N43°06'24" E131°32'17" h~120m asl 22-30.V.2015 leg. K.Makarov
Copyright © Kirill Makarov

Pterostichus (Phonias) burkhan O.Berlov & Anistschenko, 2005 Paratype
Male adult
Russia, Chita reg., 23km SW Borzia vill. Borzinskoe lake, 1.6.2002, Anichtchenko A. leg. (cAA)
Copyright © Alexander Anichtchenko

Pterostichus (Platypterus) lineatopunctatus L.Miller, 1850
Male adult
Upper Austria, NE-Alps,Windischgarsten, Rettenbach, Mt. Hoher Nock, h=1960 m,15.VI.2012, Manfred Egger leg.

Copyright © Ruslan Panin

Pterostichus (Platypterus) panzeri (Panzer, 1803)
Female adult
Austria, 47.32N 14.30E, path to Sparafeld Mt, 18.6.2017, L. Minkina leg (cAA) 14 mm
Copyright © Alexander Anichtchenko

Pterostichus (Platysma) niger niger s.str.
Male adult

NW Ukrainian Carpathians,
Lviv Prov., Staryi Sambor Distr., near Dobromyl , h=450m, 9.VI-25.VII.2007 Panin R. leg.

Copyright © Ruslan Panin

Pterostichus (Platysma) eschscholtzii (Germar, 1824)
Male adult

Copyright © Kirill Makarov

Pterostichus (Plectes) consors (Tschitscherine, 1893)
Female adult
NE Kazakhstan, S Alai Mts., Kalbinsky mt. rng., Ognevka vill., 16.6.2000 (DUBC). det. K. Makarov, 2021
Copyright © Alexander Anichtchenko

Pterostichus (Plectes) drescheri (Fischer, 1817)
Male adult
Russia, W Sayan Mts., Cheremushki vill. 3.8.1998 (DUBC) 18 mm; det. Anichtchenko A.
Copyright © Alexander Anichtchenko

Pterostichus (Pledarus) gibbicollis (Motschulsky, 1844)
Female adult

Copyright © Kirill Makarov

Pterostichus (Pseudethira) ganesh J.Schmidt, 2005 Paratype
Male adult
Nepal, Ganesh Himal, W of Pansang pass, 3200-3500m, 18.6.2000 (cAA); 15mm
Copyright © Alexander Anichtchenko

Pterostichus (Pseudoferonina) lolo Bergdahl, 2011 Holotype
Male adult
deposited in CAS, labeled: “USA: IDAHO,<br /> Clearwater Co.; Cottonwood Creek @ USFS Rd 250; Orogrande Ck. Rd.; NF Clearwater<br /> R. watershed; elev. ~870 m (2860 ft); 12.5 MI. ESE Headquarters, 3.5 MI. SW<br /> of Bungalow; 46°33'10"N, 115°33'08"W; T38N, R7E, S34; 17 July 2009; #73-2009;<br /> J. C. Bergdahl, coll.”/ “HOLOTYPE Pterostichus lolo Bergdahl designated 2011” [red<br /> label].<br /> Photo of James C. Bergdahl & David H. Kavanaugh / ZooKeys 104: 77–96 (2011)
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Pterostichus (Pseudomaseus) luctuosus (Dejean, 1828)
Male adult

Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada

Copyright © Henri Goulet

Pterostichus (Pseudomaseus) anthracinus (Illiger, 1798)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Pterostichus (Pseudorites) nicaeensis (A. Villa & G.B. Villa, 1835)
Female adult
Italy, Passo della Guardia - Imperia
Copyright © Gianni Allegro

Pterostichus (Pseudosteropus) illigeri (Panzer, 1803)
Female adult
Austria, UTM VN96, 47.30N 14.53E, path to Eisenerzer Reichenstein 21.6.2019, L. Minkina leg. (cAA) 10 mm
Copyright © Alexander Anichtchenko

Pterostichus (Pterostichus) hagenbachii Sturm, 1824
Male adult
France, Jura, Lajoux, 1190m, 20/05/2013, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Pterostichus (Aphaon) rhilensis kourili Maran, 1933c: 74
Female adult
Bulgaria, Pirin mountains, Vihren, 08.07.2010, E. Svec lgt.; 12 mm, V. Zieris det. 2012, collection Boetzl
Copyright © Fabian Bötzl

Pterostichus (Pterostichus) fasciatopunctatus (Creutzer, 1799)
Female adult
Germany, Bavaria, Passau, Laufenbach dell, F. Bötzl leg. & det., 08.09.2014
Copyright © Fabian Bötzl

Pterostichus (Pterostichus) rutilans (Dejean, 1828)
Male adult
NW Italy, Piemonte,(VC), Oropa, Lago del Mucrone (Lake), h~1900m, 22.VII.1984, L. Diotti leg. L=13 mm
Copyright © Ruslan Panin

Pterostichus (Pterostichus) rufipes (Dejean, 1828)
Male adult
France, 30450, Concoules, 27-05-2017
Copyright © Pascal Deschamps

Pterostichus (Pterostichus) dux L.Schaufuss, 1862
Female adult
Spain, Mts Cantabricos, Macizo Central, Picu Urriellu, 2000m, 13.8.2008 (cAA) leg & det. Anichtchenko A.; 18 mm;
Copyright © Alexander Anichtchenko

Pterostichus (Pterostichus) honnoratii (Dejean, 1828)
Male adult
France, Vanduse, Mt. Serein, 13.5.1983 (cAA). 14 mm
Copyright © Alexander Anichtchenko

Pterostichus (Qianius) lii Jia & Tian, 2021 Holotype
Male adult
cave Hei Dong, Longjiang Cun, Shenxi Zhen (Town), Honghuagang Qu (District), Zunyi Shi (City), Guizhou Province, 27°36ʹ46ʺN 106°58ʹ08ʺE, 822 m, 22.VIII.2020, leg. Yunchun Li and Yucheng Lin (SCAU).
Copyright © Mingyi Tian

Pterostichus (Rhagadus) solskyi (Chaudoir, 1878)
Female adult

Copyright © Kirill Makarov

Pterostichus (Sinoreophilus) scalptus Sciaky & Wrase, 1997
Male adult

Copyright © Gianni Allegro

Pterostichus (Sinoreophilus) lanista (Tschitscherine, 1898)
Male adult
C Sichuan, Jiyuding Shan, 30km N Wenchuan, 3550m, 2.7.2017, leg. V. Patrikeev (cAA). det. Hongliang Shi, 2020
Copyright © Alexander Anichtchenko

Pterostichus (Sinoreophilus) potanini Tschitscherine, 1889
Female adult
China, N Sichuan, Langmusi vill. 3700 5.8.2015, leg. V. Patrikeev (cAA) det. Hongliang Shi, 2020
Copyright © Alexander Anichtchenko

Pterostichus (Sinosteropus) horstweiperti Schmidt & Gueorguiev, 2013 Paratype
Male adult
NEPAL Arun Vall. to Makalu / Kalo Pokhari 4090m / 29.VI.2013 leg. S. Tamang / 27°41’35N 87°12’18E
Copyright © Borislav Gueorguiev

Pterostichus (Lianoe) masumotoi Tanaka; Morita & Suga, 1987 Holotype
Male adult
Mt. Fuji (south side), 1600 m, Shizuoka Pref., Japan, 31. VII-7. VIII. 1971, K. Tanaka & K. Suga
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba. COL-253
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
NIAES © Prohibited secondary use of all data
Copyright © Hiraku Yoshitake

Pterostichus (Steropanus) cavifrons Fedorenko, 2018 Holotype
Male adult
Vietnam, Gia Lai Province, ~50 km N of An Khe, Kon Chu Rang Nat[ure]. Reserv[e]., 14°30–31'N/108°32'E, h=1000– 1040 m, pitfall traps, 24.V–2.VI.2016, leg. D. Fedorenko
Copyright © Dmitri Fedorenko

Pterostichus (Steropanus) glymmiger (Andrewes, 1937)
Male adult
Vietnam, Lao Cai Prov., Sa Pa Distr., Fan Si Pan [Phang Xi Pang] Mt., 1400–1500 m, 26.V.–6.VI.1999, N.L.Orlov [leg.], 22°18'56''N 103°49'35''E;
Copyright © Dmitri Fedorenko

Pterostichus (Steropanus) pseudoviolaceus Fedorenko, 2018 Holotype
Male adult
Vietnam, Kon Tum Prov[ince], 2–3 km W of Ngoc Linh Mt., 15°05'N, 107°57'E, h=1600–2150 m, pitfall traps, 19.III– IV.2006, leg. A. Anichkin
Copyright © Dmitri Fedorenko

Pterostichus (Steropanus) securipenis Fedorenko, 2020 Holotype
Male adult
(ZMMU): Vietnam, Quang Nam Prov[ince]., Nam Gian Distr[ict]., Song Thanh Nat[io]n[al]. Park, 15°33´48´´N/ 107°23´22´´E, h=1050 m, 23.IV–11.V.2019, leg. D. Fedorenko.
Copyright © Dmitri Fedorenko

Pterostichus (Steropus) ebenus (Quensel, 1806)
Male adult
SC Spain, Jaen provincia, Linares, 30.IX.2001, M.A. Lopez Vergara leg.
L=21 mm
Copyright © Ruslan Panin

Pterostichus (Steropus) insidiatrix (Brulerie, 1872)
Male adult
Spain, Teruel, Montes Universales, Pto del Portillo 1780m 31.5.2006, Anichtchenko A. leg & det. (cAA) 19mm
Copyright © Alexander Anichtchenko

Pterostichus (Steropus) ghilianii Putzeys, 1846
Male adult
Spain, Sierra de Guadarrama, pto Somosierra, 1500m 3.6.2006, Anichtchenko A. leg & det. (DUBC) 15 mm
Copyright © Alexander Anichtchenko

Pterostichus (Steropus) madidus (Fabricius, 1775)
Male adult
Belgium, Limburg, Forest reserve of Kolmont, 24.4.2016 (DUBC). det. Anichtchenko A., 2021. 14 mm
Copyright © Alexander Anichtchenko

Pterostichus (Steropus) gallega (Fairmaire, 1859)
Male adult
Spain, Navarra, Bera de Bidasoa, Larrun Mt. 850m 9-15.9.2001, A. Anichtchenko leg. & det. (cAA) 19 mm
Copyright © Alexander Anichtchenko

Pterostichus (Tausternus) hoii Fedorenko, 2020 Paratype
Male adult

Copyright © Dmitri Fedorenko

Pterostichus (Troglorites) breuili mendizabali Jeannel, 1921
Male adult
Spain, Guipuzcoa, Albistur, (Tolosa) cueva Mendicute, M.Toribio leg.
10 mm
Copyright © Alexander Anichtchenko

Pterostichus (Troglorites) ochsi (Fagniez, 1921)
Female adult
Courmes, Alpes-Maritimes, France
Copyright © Jean Michel Lemaire

Pterostichus (Jedlickaia) sterbai Jedlicka, 1934 Paratype
Male adult
Pe Yen Tsin, Yunnan (coll. R. Sciaky)
Copyright © Riccardo Sciaky

Pterostichus (Tschitscherinea) krali Sciaky, 1997 Holotype
Male adult
China, Yunnan, Habashan mts., SE slopes, 4-5.VI.1995 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Pterostichus (Tubuliphallus) megacephalus Sciaky & Allegro, 2013 Holotype
Male adult
China, C Sichuan, Xiling Snow Mts., 2100-3100 m, 1-3.VIII.1996 (CSc)
Copyright © Gianni Allegro

Pterostichus (Vietoderus) laevibasis Fedorenko, 2023 Paratype
Male adult
N-Vietnam, 40km W of / CaoBang, Phia Oac Mt./ E-slope, h=1600-1800 m/ 22°36´27´´N105°52´0´´E/ 22.V-6.VI.2018/ leg. A.Abramov.
Copyright © Dmitri Fedorenko

Pterostichus (Vietosteropus) abramovi Fedorenko, 2017 Holotype
Male adult
(ZMMU): ‘Vietnam, Dak Lak Prov [ince]., Chu Yang Sin Nat[io]n[al]. Park, 12º222403N 108º212113E, 1.5 km W [of ] Chu Pan Phan Mt, h=1650 m, 30.III–11.IV.2012.
Copyright © Dmitri Fedorenko

Pterostichus (Vietosteropus) annamita chuyangsin Fedorenko, 2017 Holotype
Male adult
(ZMMU): ‘Vietnam, Dak Lak Prov.[ince], Chu Yang Sin Natn. Park, 12º232483N/ 108º202593E, Krong Kmar riv[er]., upper flow, h=1200–1650 m, 30.III–14.IV.2012, D. Fedorenko leg.
Copyright © Dmitri Fedorenko

Pterostichus (Vietosteropus) kontumensis Fedorenko, 2017 Holotype
Male adult
Vietnam, Kon Tum Prov[ince], 2-3 km W of Ngoc Linh Mt., 15º052N, 107º572E, h=1600–2150 m, pitfall traps, 19.III–9.IV.2006, leg. A. Anichkin
Copyright © Dmitri Fedorenko

Pterostichus (Vietosteropus) subirideus Fedorenko, 2023 Paratype
Male adult
Vietnam, Ha Tinh Prov[ince]./ Vu Quang nat[io]n[al]. Park,/ Kim Quang env., h~200 m/ 18°17´38´´N 105°22´06´´E/ 24.V-1.VI.2022/ leg. D. Fedorenko
Copyright © Dmitri Fedorenko

Pterostichus (Wraseiellus) crassiapex Shi & Sciaky, 2013 Paratype
Male adult
China – W Hubei, road Badong-Yesanguan, Tiechanghuang, 1300 m, 27.VI-11.VII.2003 (coll. R. Sciaky)
Copyright © Riccardo Sciaky