Carabus (Trachycarabus) besseri Fischer, 1822
Male adult

Copyright © Safronov

Carabus (Trachycarabus) besseri Fischer, 1822
Male+Female adult
W Ukraine.NW Podolian,
Lviv reg., Zolochiv distr., Lysa Gora near Stinka vill., steppe sites, 25.IV-9.V.2009 Panin R. leg
female, 30mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) besseri Fischer, 1822
Endophallus

Copyright © Ruslan Panin

Carabus (Trachycarabus) estreicheri Fischer, 1822
Endophallus
Lviv Region, Brody distr., near Yasenov vill.,
Copyright © Ruslan Panin

Carabus (Trachycarabus) estreicheri Fischer, 1822
Male+Female adult
W UKRAINE, NW Podolian,
Lviv Region, Brody distr., near Yasenov vill., steppe sites, 2-28.V.2010, Panin R. leg.
female, 23mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) estreicheri Fischer, 1822
Male adult

Copyright © Safronov

Carabus (Trachycarabus) estreicheri Fischer, 1822
Female adult
W-UKRAINE, NW Podolian,
Lviv Region, Brody distr., near Yasenov vill., steppe sites, 12.V-2.VI.2007 Panin R. leg.
female, 22mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) estreicheri Fischer, 1822
Female adult
Volgograd Reg., Serafimovka Distr., Ust' -Khoperskaya Station 5.V.2009 leg.O.Sveshnikov
18.5 mm
Copyright © Kirill Makarov

Carabus (Trachycarabus) estreicheri Fischer, 1822 (as jaegeri Ménétriés, 1827)
Female adult
E UKAINE, Lugansk city, Site Luganskoe, 20-30.VII.2010, V.A.Landik leg.
L=21 mm
det.Panin.R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) estreicheri (ab.rufofemoratus Lomnicki, 1892)
Female adult
W Ukraine, NW Podolian, Lviv reg., Brody distr., Makitra Mt, steppe site, 15.VI - 24.IX.2017, Panin R. & Kanarsky Yu. leg.
Copyright © Ruslan Panin

Carabus (Trachycarabus) estreicheri Fischer, 1822 (ab.rufofemoratus Lomnicki, 1892)
Female adult
W Ukraina, Podolian Upland, Lviv reg., Brody distr., Mt. Makitra, steppe site, 19.V-5.VI.2020, Panin R. leg. L=23 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
Male adult
Burjatia, Dzhida distr., near Beloozersk vill. 4-9.VII.1997 Anichtchenko A. leg.
male, 23 mm
Copyright © Alexander Anichtchenko

Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
Male adult
Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
China, Beijing, Mentougou, 2005;
Photo & det. by liuye

Copyright © Ye Liu

Carabus (Trachycarabus) latreillei ?semicoriaceus Kraatz, 1881
Endophallus
NE-China: Liaodong Peninsula, Liaoning prov., Gaizhou distr., Mt. Buyun Shan.
Copyright © Ruslan Panin

Carabus (Trachycarabus) latreillei ? semicoriaceus Kraatz, 1881
Male adult
NE-China: Liaodong Peninsula, Liaoning prov., Gaizhou distr., Mt. Buyun Shan,VI.2007, Hao leg.
L=23 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
Endophallus
N Mongolia, Khövsgöl reg., Lake Khovsgol, h=1640m, 51*:12’N; 100*:45’E, steppificated sites,
Copyright © Ruslan Panin

Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
Male adult
N Mongolia, Khövsgöl reg., Lake Khovsgol, h=1640m, 51*:12’N; 100*:45’E, steppificated sites,
20-25.VII.2010. A.Dergalev leg.
L=22 mm
Copyright © Ruslan Panin

Carabus (Morphocarabus) latreillei mengguicus Rapuzzi, 2023 Holotype
Male adult
China, Inner Mongolia Province, Tongliao, Ka’erqin, Jinbaotun, VIII.20. Image from Rapuzzi, 2023
Copyright © - - -

Carabus (Trachycarabus) latreillei shanxiensis Rapuzzi, 2024 Holotype
Male adult
China, Shanxi, Lu Liang, Kang Cheng Zhen, Jiao Kou Xian; Aug. 2022. Image from Rapuzzi, 2024
Copyright © - - -

Carabus (Trachycarabus) mandibularis mandibularis Fischer, 1827
Male adult
NE Kazakhstan, S Altai Mt., Narymskii mt. rng.
Bolshenarymskoe vill. 26.7.2001, 1800m Klimenko leg.
male, 21 mm
Copyright © Alexander Anichtchenko

Carabus (Trachycarabus) mandibularis buchtarmensis Kryzhanovskij, 1953
Male adult
Kazakhstan, Kalbinsky mt. rng., 600m ,
Ognevka vill., 23.6.98 Klimenko A. leg.
male, 23 mm
Copyright © Alexander Anichtchenko

Carabus (Trachycarabus) mandibularis buchtarmensis Kryzhanovskij, 1953
Male adult
NE-Kazakhstan,
S-Altays, Kalbinky Mt. h=500m., near Ognevka vill., 20.VI.1998.A. Klimenko leg.
male, 23mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) mandibularis buchtarmensis Kryzhanovskij, 1953
Endophallus

Copyright © Ruslan Panin

Carabus (Trachycarabus) mandibularis abakkereiorum Kabak & Hu, 2017 Holotype Paratype
Male+Female adult
(ZIN). China, Xinjiang, Altay Mt. R., Kran Valley, N of Altay Town, 47°55´37''N / 88°08´23´´E, h = 1015 m, 31.VII.2015 I.I. Kabak leg.
Copyright © Ilya Kabak

Carabus (Trachycarabus) mandibularis buchtarmensis Kryzhanovskij, 1953 Paralectotypes
Male genitalia
Aedeagus: from Tshernovaya; Endofallus: from Bolshenarymskoe.
Copyright © Ilya Kabak

Carabus (Trachycarabus) mandibularis abakkereiorum Kabak & Hu, 2017
Male genitalia

Copyright © Ilya Kabak

Carabus (Trachycarabus) planarius Obydov, 1994 Paratype
Female adult
Казахстан, Семипалатинский р-н, окр. пос. Шолпан, 500 м, 20 км. южнее пос. Таскескен, 2.06.1993, М. Данилевский leg.
Copyright © Dmitri Obydov

Carabus (Trachycarabus) planus planus Gehin, 1885
Male+Female adult
S Russia, NW Caucasus,<br /> Karacaj-Cherkessija, near Kart-Djurt vill., upper Kylan riv. h=1300m, 26-27.V.2009 , Fominykh D. leg.<br /> L=24mm<br /> photo/det. Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus planus Gehin, 1885
Male adult
NW Caucasus, Rostov reg.,<br /> Avilo-Uspenskaya vill. 22.5.2000 Arzanov Yu. leg.<br /> male, 25 mm
Copyright © Alexander Anichtchenko

Carabus (Trachycarabus) planus planus Gehin, 1885
Male adult
S Russia,<br /> Krasnodar reg., Anapa distr., B. Utrish vill., env., 11-20.VIII.2008, Khatchikov E. leg.<br /> male, 26mm<br /> photo Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus planus Gehin, 1885
Endophallus
<b>endophalus </b> S Russia,Krasnodar reg., Anapa distr., B. Utrish vill., env.,
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus planus Gehin, 1885
Endophallus
<b>endophalus</b> Karacaj-Cherkessija, near Kart-Djurt vill., upper Kylan riv. h=1300m
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus campicola Reitter, 1896
Male adult
Krym Peninsula, near Sudak. Leg.Vashchenko S. 1-5.V.2004<br /> 27 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus campicola Reitter, 1896
Male+Female adult
S Ukraine, Krym Peninsula,<br /> near Perevalnoe vill., in steppe, 15.IV.2009, Shaporinskiy V. leg.<br /> female, 28 mm<br /> photo/det.Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus campicola Reitter, 1896
Endophallus

Copyright © Ruslan Panin

Carabus (Trachycarabus) planus retowskianus Mandl, 1955
Male adult
S Ukraine, Zaporizhia reg., Melitopol distr., near Troitskoe vill., 1.VI.2010, I.V. Kovalev leg., L=27 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) planus campicola Dejean, 1831
Female adult
S Ukraine, Crimea, S slope Mt Demerdzhi h=650 m, sibljak, VII.2004, R. Panin leg., L=29 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) scabriusculus scabriusculus (as erythropus Fischer, 1822)
Male+Female adult
W.Ukraine,<br /> NW Podolian Upland , Lviv Reg., Zolochiv distr., near Luky vill., steppe sites, 7.V-8.VI. 2010<br /> leg. Panin R.<br /> female, 22mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) scabriusculus scabriusculus (as erythropus Fischer, 1822)
Male adult
W-UKRAINE<br /> NW Podolian Upland, Lviv Reg., Zolochiv Distr., near Luka vill., in steppe. Panin R. leg. 1.V - 16.VI.07
Copyright © Safronov

Carabus (Trachycarabus) scabriusculus scabriusculus Olivier, 1795
Endophallus

Copyright © Ruslan Panin

Carabus (Trachycarabus) scabriusculus inapertus Motschulsky, 1850 (loc.typ)
Male+Female adult
NE UKAINE,
Kharkov reg., Zmiev distr., in steppe, 15.VI-14.VII.2009, Kovalchuk D. leg.
female, 20mm
photo/det. Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) scabriusculus inapertus Motschulsky, 1850 (loc.typ)
Endophallus

Copyright © Ruslan Panin

Carabus (Trachycarabus) scabriusculus lippii Dejean, 1826
Female adult
SW Ukraine, Odessa reg., Ivanovka distr., steppe ravine near Gudevichevo vill., 22.IV-5.V.2022, A.Gontarenko leg., L=22 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) scabriusculus lippii Dejean, 1826
Male adult
SW Ukraine, Odessa reg., Ivanovka distr., steppe ravine near Gudevichevo vill., 22.IV-5.V.2022, A.Gontarenko leg., L=21.5 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) scabriusculus inapertus Motschulsky, 1850
Male adult
NC Ukraine, Kiev city, Lysa Gora, 1-15.VI.2012, Gerasimov R.leg./det. R.Panin, L=19 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus tscherkessicus Korge, 1964
Male adult
S Russia, NW-Caucasus,
Karachay-Cherkessia,Teberda, Mukhinsky (Mukhu) Mt. pass, h=2750-2800 m , 8.VIII.2010, I.A. Solodovnikov leg.
male, 23 mm

Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus tscherkessicus Korge, 1964
Endophallus

Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus sibiricus Fischer, 1822
Male adult

Copyright © Kirill Makarov

Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886
Male+Female adult
W Ukraine,<br /> Ternopol reg., Pidvolochisk distr., Supranivka vill.<br /> 3.VII.1908. Leg. Stobetskii. S.<br /> Photo R. Panin
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886
Male adult
W Ukraine,
Ternopil reg., Pidvolohcisk distr., Supranivka vill.
male, 26mm
The end of XIX the beginning of XX century
photo Panin R.
coll. SMNH Ukraine, Lviv
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886
Endophallus

Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886
Endophallus
aedeagus
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus obliteratus Fischer, 1827
Male adult
Altai mts., Kalbinskii mt.rng.
Ognevka vill. VII.1994, Klimenko A. leg.
male, 28 mm
Copyright © Alexander Anichtchenko

Carabus (Trachycarabus) sibiricus obliteratus Fischer, 1827
Male adult
NE Kazakhstan,
S Altai Mts., Ulbinsky range h~500m.,Turgusun vill., 20.VI.2001
male 25mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus obliteratus Fischer, 1827
Endophallus
endophallus sibiricus obliteratus Fischer, 1827
NE Kazakhstan,
S Altai mts.,S Ulbinsky mt.h=500m.,Turgusun vill., 20.VI.2001
male 25mm.
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus kolosovi Zinoviev, 1997 loc.typ.
Female adult
S. Ural, Bashkiria, near Mesjagutovo vill., 09.07.2001 A. Menstschicov leg.
photo A. Menstschicov
Copyright © Anton Menstschikov

Carabus (Trachycarabus) sibiricus kolosovi Zinoviev, 1997
Female adult
Russia, North-East Bashkortostan. Ai river, near Sikiyaz-Tamak vill.
12.07.1997, Zinoviev E.leg
length: 21,8 mm
Copyright © Evgeniy Zinoviev

Carabus (Trachycarabus) sibiricus haeres (as fossulatus Dejean, 1826)
Female adult
<b>Carabus (Trachycarabus) sibiricus fossulatus Dejean, 1826</b><br /> NE Ukraine,<br /> Kharkov reg.,1974<br /> female, 27mm.
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus haeres (as fossulatus Dejean, 1826)
Male+Female adult
E.UKRAINE,<br /> Kharkov reg., Zmiev distr.,in steppe,10-20.V I.2007, D.Kovalchuk leg.<br /> female, 26mm/male, 24mm.<br /> photo/det. Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus haeres (as fossulatus Dejean, 1826)
Endophallus
endophallus
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus errans Fischer, 1823
Female adult
Carabus (Trachycarabus) sibiricus errans Fischer, 1823
S.Russia, Rostov-Don reg., Avilo-Uspenskaya env.
V.2000
Copyright © Safronov

Carabus (Trachycarabus) sibiricus errans Fischer, 1823
Male adult
SW-Russia,
Rostov reg. env. stanitsa Razdorskaya, A. Ponomarev leg. 9-15.V.2001
male 25mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus errans Fischer, 1823
Male+Female adult
female:SE Ukraine,
Donetsk, 25.VII.2009, A.I.Gubin leg. L=30mm.
male:E Ukraine, Lugansk reg., near Slavyanoserbk, 5-15.VIII.2009, S.Konovalov leg. L=26mm.
photo/det.Panin R.
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus errans Fischer, 1823
Endophallus
endophallus sibiricus errans- Lugansk reg., near Slavyanoserbk
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus bosphoranus Fischer, 1823
Female adult
Carabus (Trachycarabus) sibiricus bosphoranus Fischer, 1823
S Ukraine,
Krym Peninsula, Nikitskaya Jaila, S.Mosyanin leg. 10.VIII.2000
female 23mm.
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus bosphoranus Fischer, 1823
Male adult
Carabus (Trachycarabus) sibiricus bosphoranus Fisch. 1823<br /> S Ukraine, Krym Peninsula,<br /> Dolgorukovskaya jaila, 25.VIII.1991 I.A.Solodovnikov leg.<br /> male, 23mm<br />
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus gvozdevae Obydov, 1997
Male adult
Russia, Khakassia,
W Sayan, Enisey riv., Batenevskiy Mt Range, 12 km E Pervomayskiy vill., 11.VII.2000, A.& N. Menstschicovy leg.
L=21 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus tscherkessicus Korge, 1964
Male adult

Copyright © Kirill Makarov

Carabus (Trachycarabus) sibiricus sibiricus Fischer, 1822
Male adult
E Siberia, Russia, Krasnoyarsk Reg., near Krasnoyarsk, on dry steppe slopes, 7.VIII,2011, M.Ivanov leg.
L=21 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus sibiricus Fischer, 1822
Endophallus
near Krasnoyarsk
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus haeres Fischer, 1823
Female adult

Copyright © Kirill Makarov

Carabus (Trachycarabus) sibiricus haeres Fischer, 1823
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Carabus (Trachycarabus) sibiricus obliteratus Fischer, 1827 Paratype
Female adult
China, Xinjiang, “Prov. Xinjiang: Ake Tei A Zhu Pai 1750 m, 5.VIII.1960, leg. Wan Su Yung, China Naturwiss. Institut”; “Paratypus Carabus (Trachycarabus) ♀ sibiricus pseudobliteratus nov. ssp. det. Korell et Kleinfeld, 1982”. (MFNB)
Copyright © Ilya Kabak

Carabus (Trachycarabus) sibiricus haeres (trans s. errans)
Female adult
C Ukraine, S of Poltava city, edge forest-lane, 2-26.IX.2010, O. Pozharov leg. /det. R.Panin. L=30 mm
Copyright © Ruslan Panin

Carabus (Trachycarabus) sibiricus seriatoporus Fischer von Waldheim, 1828
Male adult
окр. Якутска, пос. Бестях на р. Лена, 30.VI.1962, Сладков leg.
Copyright © Dmitri Obydov