Geocharidius phineus Erwin, 1982
Male adult
Guatemala
image from Sokolov I., 2013. A new genus and eight new species of the subtribe Anillina (Carabidae, Trechinae, Bembidiini) from Mexico, with a cladistic analysis and some notes on the evolution of the genus.
Copyright © Igor Sokolov

Nesamblyops sp.
Male adult
New Zealand
Scale bars = 0.5mm
image from Sokolov I., 2013. A new genus and eight new species of the subtribe Anillina (Carabidae, Trechinae, Bembidiini) from Mexico, with a cladistic analysis and some notes on the evolution of the genus.
Copyright © Igor Sokolov

Zapotecanillus montanus Sokolov, 2013 Paratype
Male adult
MEXICO, Oaxaca, 52miles N of Oaxaca, Ber.202, sink litter, 17 May 1971 S.B.Peck collector (CMNH)
Scale bar = 0.5mm
Copyright © Igor Sokolov

Hypotyphlus degiovannii Magrini, 2013 Holotype
Male adult
Italia, Lazio, Montagna di Scauri (Latina), 50 m s.l.m., 1.V.2008, P. Magrini leg. (CM).
Copyright © Paolo Magrini

Pelodiaetodes (Pelodiaetodes) constricticollis Sokolov, 2015 Paratype
Male adult
NZ, Northland, Radar Bush. Image from Sokolov, 2015
Copyright © - - -

Anillus angelae Magrini & Vanni, 1989
Male adult
Arnale cave, Coreno Ausonio, Italia
Copyright © Vittorino Monzini

Rhegmatobius bastianinii Magrini & Casale, 2015 Paratype
Male adult
Italia, Lazio (LT) M.te S. Angelo, Formia, 1200m.
Copyright © Augusto Degiovanni

Nesamblyops townsendi Sokolov, 2023
Male adult
Tennyson Inlet, Marlborough Sounds, SO
Copyright © Igor Sokolov

Anillaspis explanata (Horn, 1888)
Female adult
USA: California: El Dorado Co., Alabaster Cave (CMNH). Scale bar 1 mm; Image from LaBonte & Maddison, 2023
Copyright © - - -

Stylulus (Stylulus) isabelae Giachino & Sciaky, 2003 Holotype
Male adult

Copyright © Vittorino Monzini

Tasmanitachoides major Baehr, 1990 Holotype
Male adult

Copyright © Museums Victoria

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Image from Sallenave, Erwin, and Roig-Juñent, 2008
Copyright © - - -

Andinodontis muellermotzfeldi Toledano & Erwin, 2010
Female adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Argentinatachoides balli Sallanave, Erwin & Roig-Junient, 2008
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidarenas setiventris Negre, 1973
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides sp.
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tasmanitachoides erwini Maddison & Porch, 2021 Holotype
Male adult
Australia: Tasmania: River Forth at C136, 41.4712°S 146.1366°E, 126 m, 14.i.2019. DRM 19.012. D.R. Maddison & N.A. Porch. Scale bar 1mm
Copyright © David Maddison

Andinodontis maveetyae Erwin & Maddison, 2010 Holotype
Male adult
Perú: Cusco, Rocotal, 3.2 km E Tres Cruces, 13.1123° S, 71.5834° W, 2082 m, 27 June 2010 (S. MAVEETY) (MHN-UNSAAC: ADP124815)
Copyright © Terry Erwin

Bembidion (Testediolum) armeniacum kokandicum Solsky, 1874: 120
Female adult

Copyright © Kirill Makarov

Bembidion (Semicampa) schueppelii Dejean, 1831
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Peryphus) obscurellum obscurellum (Motschulsky, 1845)
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Bembidion (Zecillenus) albescens (Bates 1878)
Male adult
Image from Larochelle A. & Larivière M.-C., 2015. Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini). Insecta Mundi 0415: 1–78
Copyright © - - -

Bembidion (Testedium) laetum Brulle, 1836: pl.2
Female adult
Morocco, 65 km N Essaouria, 5 km N Jamaa Shoim, 23-25.III.2011, A. Gusakov. Фотография с сайта zin.ru\Animalia\Coleoptera.
Copyright © Maksim Smirnov

Bembidion (Pamirium) platypterum Solsky, 1874: 120
Female adult
Kazakhstan, Alma-Ata, 2.06.1990. det. Anichtchenko A. 2016
Copyright © Alexander Anichtchenko

Bembidion (Nepha) hauserianum Netolitzky, 1918
Female adult
Iran, 20 km SW Yasuj, 5-6.V.2007, Anichtchenko A. leg.
Copyright © Alexander Anichtchenko

Bembidion (Leuchydrium) tigrinum Leconte, 1879
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Bembidion (Taiwanobembidion) aliense Habu, 1973
Male adult

Copyright © David Maddison

Bembidion (Omotaphus) mellissii Wollaston, 1869
Female adult
RSA, 3km NW Nieuwoudtville, Willems riv. 27.11.2023 (cAA). det. Anichtchenko A. 2026
Copyright © Alexander Anichtchenko

Horologion speokoites Valentine, 1932 Holotype
Male adult
"Arbuckle's Cave Maxwelton, W.Va. July 12 1931 J.M. Valentine / On bit of wet decaying wood, muddy floor of lower level / Type gen. Type sp. / U.S.N.M. Type 44255"
Copyright © Curt Harden

Lissopogonus tonkinensis Zamotajlov & Sciaky, 1996: 40
Male adult
N Vietnam, Lao Cai Province 250 km from Hanoi bearing 310°, SaPa vill env., h~1250 Hoang Lien Son Nat. Park 27.VI-2.VII.1998 leg.A.Napolov
L= 4,8
Copyright © Kirill Makarov

Lissopogonus cf. borneensis Baehr, 2001
Male adult
Malaysia, Borneo, Sarawak, Miri div., Bario 10 km NE, 3 km N Pa Lungan, 7-8.III.2014, R. Cibulskis leg. det. A. Anichtchenko 2015
Copyright © Alexander Anichtchenko

Lissopogonus morvani Deuve, 2009
Female adult
Nepal, Kosi Mure, 2000-2100m, 27.30N 87.16E, 6-12.6.2001. 7 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus Andrewes, 1923
Male adult
Nepal, 12km NE Chainpur, 1400m, 29.36.17N, 81.17.54E, 28.6.2009. 5.8 mm; det. Anichtchenko A. 2025
Copyright © Alexander Anichtchenko

Lissopogonus glabellus laosensis Deuve, 2009
Male adult
Laos, Nong Khiaw vic, 20.553945, 102.622681, 1.V.2026, Anichtchenko A. leg. & det. (cAA)
Copyright © Alexander Anichtchenko

Diplous (Platidius) dolini Zamotajlov, 2005
Female adult
S Primorie, Lazovsky Reserve, Formosova Spring 43°00'06"N 134°05'13"E 10.VII.2005 leg. K.Makarov
L=9.3
Copyright © Kirill Makarov

Archipatrobus flavipes (Motschulsky, 1864)
Male adult
Collection ground Kobe city, Hyogo Pref. 33645<br /> (C) Working Group for Biological Indicator Ground Beetles Database, Japan<br /> Prohibited secondary use of all data
Copyright © Katsuro Yahiro

Dimorphopatrobus ludmilae Casale et Sciaky, 1994: 47
Female adult
Tibet, Motuo, Hanmi, 8.2013. det. A. Anichtchenko, 2015 17 mm
Copyright © Alexander Anichtchenko

Himalopenetretus franzi (Zamotajlov & Sciaky, 1998) Holotype
Male adult
India, Uttar Pradesh; 17 km SE of Gangotri, Tapovan, 4500 m, 26.VI.1996 (coll. R. Sciaky)
Copyright © Riccardo Sciaky

Grandipenetretus major (Zamotajlov et Wrase, 1997) Paratype
Male adult
China, Shaanxi, Qin Ling Shan, 47 km S Xian, 26-29.VIII.1995 (Coll. R. Sciaky)
Copyright © Riccardo Sciaky

Himalopenetretus burangensis Yan, Shi & Liang, 2020 Holotype
Male adult
“Xizang, Ngari Pref., Burang County, Burang Town. N of Tinkar pass, alpine meadow. 30.2085N, 81.0661E, 4800m.”; “under stone, 2019. VII. 15, Shi HL lgt., Exp. BJFU 2019”; “DNA sample series number SHL2019--Xiz006”; “HOLOTYPE♂ Himalopenetretus burangensis sp. nov., des. YAN & SHI, 2020” [red label]. (IZAS). Image from Yan, Shi & Liang, 2020
Copyright © - - -

Troglopatrobus zhouchaoi Deuve, He & Tian, 2020 Holotype
Male adult
cave Lianhua Dong, Tangba Cun, Bailu Zhen, Pengzhou Shi, Chengdu, Sichuan, 31°10ʹ5852ʺN 103°56ʹ09.13ʺE, 1064 m in altitude, 20.VI.2019, leg. Li He, Yuan Li & Chao Zhou (SCAU). Image from Deuve, He & Tian, 2020
Copyright © - - -

Patrobus septentrionis volgensis Zamotajlov & Isaev, 2006
Male adult

Copyright © Igor Solodovnikov

Deltomerodes conaensis Yan, H. Shi, Liang & J. Shi, 2021 Holotype
Male adult
Image from Yan, H. Shi, Liang & J. Shi, 2021
Copyright © - - -

Deltomerus (Deltomerus) corax Peyerimhoff, 1922
Male adult
Morocco, Grand Atlas, 2200-2300m, N31.13.09.5 W07.49.29.7 16-17.6.2010 Anichtchenko A. leg. & det. (cAA) 12 mm
Copyright © Alexander Anichtchenko

Pogonus (Pogonus) iridipennis Nicolai, 1822
Male adult
Volgograd Reg., Pallas Distr., N shore of Elton Lake, right bank of Khara Riv., mouth 13.V.2006 A.Matalin leg.
L=6.7 mm
Copyright © Kirill Makarov

Pogonistes rufoaeneus Dejean, 1828: 19 (Pogonus)
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pogonus (Pogonoidius) cumanus Lutshnik, 1916c: 76
Male adult
det. K. Makarov
Copyright © Kirill Makarov

Pogonus (Pogonus) orientalis virens Menetries, 1849
Female adult
det. K. Makarov
Copyright © Kirill Makarov

Diplochaetus planatus (G.Horn, 1876)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Thalassotrechus barbarae (G.Horn, 1892)
Male adult
Creative Commons CC-BY 3.0 license
Copyright © David Maddison

Iranopogonus freyellus Jedlicka, 1959 Holotype
Male adult
Iran, Sistan va Baluchestan Province, Chabahar env., IX.1937, leg. Band (NHMB). Image of Matthias Borer (NHMB) from Azadbakhsh, 2019
Copyright © - - -

Pogonopsis pallida Bedel, 1898 Holotype
Male adult
Tunisia (MNHN) Image from Azadbakhsh, 2019
Copyright © - - -

Sirdenus (Sirdenus) filiformis (Dejean, 1828)
Male adult
Italie, Sardaigne, Sestu 11-05-1982
Copyright © Pascal Deschamps

Pogonus (Pogonus) biroi Csiki, 1907
Male adult
India, Goa, Palm Beach, 20-23.4.2013 (cAA). 5.5 mm; det. Anichtchenko A. 2026
Copyright © Alexander Anichtchenko

Phrypeus rickseckeri (Hayward, 1897)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Sinozolus cf. yuae Deuve, 1997
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Chaltenia patagonica Roig-Junent et Cicchino, 2001
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachyura (Amaurotachys) notaphoida (Bates, 1886)
Male adult
India, Uttarakhand, 5 km NNE of Ramnagar, valley of Kosi River, 400-420 m, 29°26' N, 79°08,5' E, 21.04.2012, R. Dudko, I. Lyubechanskii
Copyright © Roman Dudko

Tachyura ascendens (Alluaud, 1917)
Female adult
TANZANIA, Kilimanjaro, Nkuansa, grassland,1660m, 3.7.2011, R. Felix det. as Elaphropus (Tachyura) exiguus (BOHEMAN 1848)
Copyright © Peter Schüle

Elaphropus (Idiotachys) acutifrons Boyd & Erwin, 2016 Holotype
Female adult
Brazil, Santarem
Copyright © Terry Erwin

Elaphropus (Ammotachys) marchantarius Boyd & Erwin, 2016 Paratype
Male adult
Venezuela, San Carlos de Rio Negro
Copyright © Terry Erwin

Meotachys (Hylotachys) ballorum Boyd & Erwin, 2016 Paratype
Female adult
Brazil, Amazonas, Tapurucuara
Copyright © Terry Erwin

Stigmatachys uvea Boyd & Erwin, 2016 Holotype
Female adult
Perú, Loreto, Campamento San Jacinto
Copyright © Terry Erwin

Pericompsus (Pericompsus) laetulus LeConte, 1851
Male adult
Creative Commons CC-BY 3.0
Copyright © David Maddison

Tachyta (Paratachyta) quadrinotata Baehr, 2016 Holotype
Female adult
320 Ilam Distr., betw. Hodia Khola Valley to Sktia, 250–500 m, Shorea forest, dry, 7 Apr 1988, MARTENS & SCHAWALLER leg. / Nepal-Expeditionen JOCHEN MARTENS (SMNS).
Copyright © Martin Baehr

Tachys (Paratachys) frischi Coulon & Felix, 2009
Female adult
Iran, Khuzestan, 30 km E Shushtar, 5.5.2007, A. Anichtchenko leg. (cAA). det. J. Muilwijk, 2021
Copyright © Alexander Anichtchenko

Tachyura (Tachyura) stevensi (Andrewes, 1925)
Female adult
India, Uttarakhand, 5 km SW Rishikesh, left bank of Ganga riv., 9.4.2012, Anichtchenko A. leg. (cAA). det. M. Baehr
Copyright © Alexander Anichtchenko

Ushijimaella lucida Belousov et Kabak, 2002
Male adult
China, Sichuan, WSW of Lixian, Qunlaishan Mt.R., W of Mt. "5892", h~3000 11.VII.2000 leg. I.Belousov, I.Kabak <br /> L= 3,6 mm
Copyright © Kirill Makarov

Duvalius (Hungarotrechus) bokori gelidus Csiki, 19 12c: 510
Male adult
SK, SE Slovakia, Slovenský kras; 3mm
Copyright © Aleš Sedláček

Geotrechus orcinus Linder, 1859: 72
Male adult
France, gr.te de Gargas, H.P., 15-4-1962, leg. M. Cabidoche
Copyright © Vittorino Monzini

Duvalius (Duvalius) doderoi Gestro, 1885
Male adult
Tann-a de Raixe cave, Carsi GE 12/4/2014, leg. V.Monzini. Photo Riccardo Sciaky
Copyright © Vittorino Monzini

Guizhaphaenopsodes solidior Tian & X. Huang, 2021 Holotype
Male adult
cave Tangle Dong, suburb of Jishou, Xiangxi Tujia & Miao Autonomous Prefecture, west Hunan Province, 28°18’21”N /109°39’15”E, 270 m in altitude, 2018-I-15, leg. Xinglong Huang (SCAU).
Copyright © Mingyi Tian

Orotrechus jamae C. Etonti et M. Etonti, 1979: 27
Male adult
NE Italy, Belluno, Prealps, Cansiglio plateau, Bus dell'Ors, 19.V.2013, L=6.2mm
Copyright © Denis Khrapov

Zhijinaphaenops jingliae Deuve & Tian, 2015
Male adult
Guizhou: Guiyang, Xifeng, Xishan, Shenli, Mafen Dong cave (息烽县西山镇马粪洞), 27°04'N, 106°37'E, 1157 m, 2020–II–29, leg. Guangyuan Cheng
Copyright © Mingyi Tian

Trechus (Trechus) minitrechus Schmidt & Faille, 2018 Paratype
Male adult
ETHIOPIA, Oromia, Mt Enkuolo NE slope, alt. 3200–3300 m, 07°23′45″ N, 39°22′27″ E, 8 Dec. 2016
Copyright © Joachim Schmidt

Velebitaphaenops giganteus Casale & Jalzic, 2012
Female adult
Image from cavebiology.net
Copyright © Creative Commons

Yinaphaenops guilinensis Li, Liu & Huang, 2024 Holotype
Male adult
cave Guizai Dong, Lingchuan, Guilin, Guangxi, China, 26-IV-2024, leg. Sun-Bin Huang, Ming-Zhi Zhao, Ran Li & Rong Chen (SCAU). Image from Li, Liu & Huang, 2024
Copyright © - - -

Thayerella newtoni Baehr, 2016 Holotype
Female adult

Copyright © Martin Baehr

Idacarabus troglodytes Lea, 1910 Syntypes
Male adult

Copyright © Museums Victoria

Brachydema victoriae Sloane, 1915 Syntypes
Unsexed

Copyright © Museums Victoria

Zolus wongi Larochelle & Larivière, 2017 Paratype
Female adult
[NEW ZEALAND BR] Mt Dewar 1000m J.C. Watt (typed) / under bark [of] logs (typed) / Paparoa Range Dec. 1969 (NZAC). Scale bar 1mm
Copyright © - - -

Oopterus patulus (Broun, 1881)
Male adult
scale bar 1mm
Copyright © - - -

Oopterus discoideus Larochelle & Larivière, 2017 Paratype
Female adult
“[NEW ZEALAND FD] Wolfe Flat 600m-680m (typed) / Turret Ra [= Range] (typed) / Manapouri Exp. Jan. 70 J. McBurney (typed) / moss (typed) 70/69. Scale bar 1mm
Copyright © - - -

Synteratus ovalis Broun, 1909
Unsexed
Scale bar 1mm
Copyright © - - -

Pseudoopterus otiraensis Larochelle & Larivière, 2017 Paratype
Male adult
NEW ZEALAND WD Arthur’s Pass NP, Kellys Creek x Otira River [= Kelly Creek near Otira River Junction] 4248S 17134E 350m 5.III. 2009 Larochelle and Larivière (typed) / Floodplain Podoc.-brleaf for. [= Podocarp-broadleaf forest] (Fuchsia-Horopito dom. [= dominant]): wet clayish soil. Under large, deeply embedded stones. Scale bar 1mm
Copyright © - - -

Maungazolus acutus Larochelle & Larivière, 2017 Paratype
Male adult
NEW ZEALAND FD Takahe Valley Murchison Mts [= Mountains] 11 Dec 1972 A.C. Eyles (typed) / Litter (typed) 72/271
Copyright © - - -

Merizodus soledadinus (Guerin-Meneville, 1830)
Male adult

Copyright © - - -